DOCSLIB.ORG

Fossil Records of Rhinoceroses (Rhinocerotoidea Gray, 1821

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Fossil Records of Rhinoceroses (Rhinocerotoidea Gray, 1821 Bulletin of the Natural History Museum - Plovdiv Bull. Nat. Hist. Mus. Plovdiv, 2017, vol. 2: 1-7 Fossil records of Rhinoceroses (Rhinocerotoidea Gray, 1821), Chalicotheres (Chalicotherioidea Gill, 1872) and Brontotheres (Brontotherioidea (Marsh, 1873) (Peryssodactyla Owen, 1848 - Mammalia Linnaeus, 1758) in Bulgaria Zlatozar N. Boev* National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, BULGARIA * Corresponding author: [email protected]; [email protected] Abstract. The fossil record of the Rhinoceroses, Chalicotheres and Brontotheres in Bulgaria covers 34 taxa from a total of 53 localities dated from the Oligocene to the Late Pleistocene: Rhinocerotoidea (27 taxa), Chalicotherioidea (5), and Brontotherioidea (2). Most widely spread were Aceratherium incisivum (established in 15 localitiees), Dihoplus schleiermacheri (13), Coelodonta antiquitatis (6), and Ancylotherium pentelicum (5). The majority of the localities (27) are concentrated along the largest Bulgarian rivers - Struma (14 localities), Iskar (6). Maritsa (5), and Mesta (2), as well as the Sofia Valley (9) and the Burgas lowland (3). Key words: Fossil, perissodactyls, Neogene, Miocene. Introduction and Neogene (Miocene and Pliocene), data from Chalicotheres (Chalicotherioidea Gill, 1872) Bulgaria and Balkans complete the information and Brontotheres (Brontotherioidea (Marsh, on their final former presence in Europe before 1873) are fossil group of peryssodactyls, while the drastic range restriction to Southern regions Rhinocerotoidea (Gray, 1821) survived through of Aftica and Asia today (Rhinoceratidae), or rhinoceroses (Rhinocerotidae Gray, 1820) with 4 total extinction (Chalicotheriidae and genera and 5 species (DINERSTEIN, 2011): white Brontotheriidae). Present study aims to gather all rhinoceros Ceratotherium simum Burchell, 1817), scattered information on the fossil history of black rhinoceros Diceros bicornis (Linnaeus, 1758), these three superfamilies of Peryssodactyla in Sumatran rhinoceros Dicerorhinus sumatrensis Bulgaria. (Fischer, 1814)[3], Javan rhinoceros Rhinoceros sondaicus Desmarest, 1822[3], and Indian Material and Methods rhinoceros Rhinoceros unicornis Linnaeus, 1758. All For the goal of the study, i. e. gathering the they have Old World subtropical and tropical numerous scattered published (and distribution in Africa and Asia. unpublished) data for the fossil remains of these Territory of Bulgaria, Balkans and all Europe three superfamilies of Peryssodactyla lies on out of the recent range of Rhinocerotidae. throughout all the country, we have checked Although many records of fossil rhinoceroses many sources as scientific publications, popular- from Europe documented very well the former science publications, and museum collections. distribution of these perissodactyls in the Sometimes a collected specimen was published Western Palearctic in the Paleogene (Oligocene) in several publications under different names, © Bull. Nat. Hist. Mus. Plovdiv Regional Natural History Museum – Plovdiv http://rnhm.org/en/ University of Plovdiv Publishing House Fossil records of Rhinoceroses, Chalicotheres and Brontotheres (Peryssodactyla, Mammalia) in Bulgaria but we accepted the last identification or the Lower Pliocene (BAKALOV & NIKOLOV, 1962); newest taxonomical referring. Gaber – 2. (Sofia Region). Coal mine Garmen. Oligocene – Lower Pliocene (BAKALOV & Results and Discussion NIKOLOV, 1962); Gaber – 3. (Sofia Region). Coal mine Oreshets Oligocene – Lower Pliocene Taxa account (BAKALOV & NIKOLOV, 1962); Stanyantsi (Sofia Region): Maeotian (NIKOLOV, 1985) – Late Superfamily RHINOCEROTOIDEA Gray, Miocene. Late Miocene (Turolian, 5.80-5.35 Mya 1821 /Miocene-Pliocene boundary/) (BOHME et al., 2013). Udescribed and unpublished finds Family AMYNODONTIDAE Scott & (GERAADS et al., 2011); Oreshets (Vidin Region): Osborn, 1883 Oligocene-Miocene (NIKOLOV, 1985); Dolni Izvor (Haskovo Region): Oligocene (NIKOLOV, 1. Cadurcodon ardynensis Osborn, 1924: 1963). Kamеno (Burgas region). Oligocene. Sand 4. Aceratherium sp.: Noevtsi (Pernik quarry (NIKOLOV, 1963, 1977a). Region): Maeotian (NIKOLOV, 1985) – Late Miocene; Kromidovo (Blagoevgrad Region): Family HYRACODONTIDAE Cope, 1879 Late Miocene (NIKOLOV, 1985); Slatino – 1. 2. Paraceratherium bugtiense Pilgrim, 1908: (Blagoevgrad Region). Quarry (SPASSOV et al., Dobrinishte (Blagoevgrad Region): Oligocene 2006). Lower part of the Upper Miocene (NIKOLOV, 1985). Listed as Indricotherium (GERAADS & SPASSOV, 2009). transouralicum Pavlova, 1922 5. Chilotherium zernowi (Borissiak, 1915): Kalimantsi (Blagoevgrad Region) NH 59: Family RHINOCEROTIDAE Gray, 1821 Pontian (BAKALOV & NIKOLOV, 1962), Middle Turolian – Late Miocene (GERAADS & SPASSOV, Subfamily Aceratheriinae Dollo, 1885 2002, 2009). Listed as Aceratherium zernowi Borissiak, 1911 (BAKALOV & NIKOLOV, 1962, 3. Aceratherium incisivum (Cuvier, 1822): SPASSOV, 2002). Merichleri (Haskovo Region): Lover Pliocene 6. Chilotherium cf. sarmaticum Korotkevich, (BAKALOV & NIKOLOV, 1962); Oligocene 1970: Oranovo - pr. quartrer Oranovo of town (NIKOLOV, 1985); Ezerovo (Plovdiv Region): of Simitli (Blagoevgrad Region). Quarry Maeotian - Pontian (NIKOLOV, 1985) – Late (SPASSOV et al., 2006; GERAADS & SPASSOV, Miocene; Аhmatovo (Plovdiv Region): Pontian 2009). (NIKOLOV, 1985); Dimitrovgrad (Haskovo 7. Chilotherium (Eochilotherium) cf. kiliasi Region) – Durhan quarry near town. Oligocene Geraads, Koufos, 1990: Kromidovo (BAKALOV & NIKOLOV, 1962); Oligocene- (Blagoevgrad Region). Late Miocene (GERAADS Miocene (NIKOLOV, 1985); Troyanovo – 1 & SPASSOV, 2009). (Haskovo Region). Coal mine. Oligocene- 8. Chilotherium cf. kowalevskii Pavlow, 1913: Miocene (NIKOLOV, 1985); Dobrich (Haskovo Yambol (Yambol Region). Maeotian to Dacian- Region): Oligocene-Miocene (NIKOLOV, 1985); Romanian, Late Miocene (GERAADS & Burgas (Burgas Region): Maeotian - Pontian SPASSOV, 2009). (NIKOLOV, 1985). Precise locality unknown. 9. Chilotherium sp.: Ahmatovo (Plovdiv Maeotian and Pontian, probably mixed fauna Region). Second half/the end of the middle from different beds (Miocene; NIKOLOV, Turolian (MN12) – Late Miocene (GERAADS & 1977a): (BAKALOV & NIKOLOV, 1962) SPASSOV, 2009); Stanyantsi (Sofia Region). Late (BAKALOV, 1927); Hrabarsko (Sofia Region). Miocene (Turolian, 5.80-5.35 Mya /Miocene- Late Miocene (Pontian; BAKALOV & NIKOLOV, Pliocene boundary/) (BOHME et al., 2013); 1962). Abandoned coal mines; Gnilyanska Sarafovo (Burgas region). Miocene formation (NIKOLOV, 1985). Cranial fragment (JAKUBOWSKI & KARASZEWSKI, 1972). with right maxilla and teeth; Aldomirovtsi (Sofia 10. Acerorhinus sp.: Kalimantsi (Blagoevgrad Region): Pontian (NIKOLOV, 1985) – Late Region). Middle Turolian – Late Miocene Miocene; Gaber – 1. (Sofia Region). Coal mine (GERAADS & SPASSOV, 2009). Beli Breg (former Bolshevik) near the Gaber 11. Brachypotherium sp.: Ahmatovo village. (Pontian; NIKOLOV, 1985). Oligocene – (GERAADS & SPASSOV, 2009); Kalimantsi 2 Zlatozar N. Boev Middle Turolian – Late Miocene (GERAADS & 20. Dihoplus schleiermacheri (Kaup 1832): SPASSOV, 2009). Kalimantsi (Blagoevgrad Region) NH 59: Lower Miocene – Pontian (BAKALOV & NIKOLOV, Subfamily Rhinocerotinae Owen, 1845 1962); Middle Turolian – Late Miocene (GERAADS & SPASSOV, 2009). Listed as 12. Dicerorhinus kirchbergensis (Jäger, 1839): Dicerorhinus schleiermacheri Kaup, 1834; Hrabarsko Dobroslavtsi (Sofia Region): Pleistocene. Listed (Sofia Region). Upper Pliocene – Lavant as Rhinoceros mercki Jäger, 1839 (BAKALOV & (BAKALOV & NIKOLOV, 1962); Late Miocene NIKOLOV, 1962); Mladenova cave near Chiren (Pontian; BAKALOV & NIKOLOV, 1962). village (Vratsa Region). Listed as Rhinoceros mercki Abandoned coal mines; Gnilyanska formation Jäger, 1839 (NIKOLOV, 1977b; 1983). (NIKOLOV, 1985). Cranial fragment with right 13. Rhinoceros sp.: Nova cave (near Lyuti maxilla and teeth. Listed as Dicerorhinus Brod village; Pleven Region): Pleistocene schleiermacheri Kaup, 1834; Katina (Sofia Region): (NIKOLOV, 1983). Pontian (NIKOLOV, 1985) – Late Miocene. 14. Coelodonta antiquitatis (Blumenbach, Listed as Dicerorhinus schleiermacheri Kaup, 1834; 1807): Temnata Dupka cave (Lovech Region): Dragovishtitsa (Sofia Region): Upper Pliocene – Late Pleistocene 1,6 Mya (VASIL POPOV – Lavant (BAKALOV & NIKOLOV, 1962); unpubl. data). Listed as Rhinoceros tichorinus Oligocene. Exact locality unknown. Listed as (POPOV, 1936); Bacho Kiro cave (Gabrovo Dicerorhinus schleiermacheri Kaup, 1834. Region): final of Late Pleistocene (50 000-10 000 (NIKOLOV, 1985; Prolesha (Sofia Region): ВР; BOCHEŃSKI, 1982). Listed as Rhinoceros Upper Pliocene – Lavant (BAKALOV & tichorhinus Cuvier, 1812 from Malkata cave NIKOLOV, 1962); Dacian (NIKOLOV, 1985) – (NIKOLOV, 1977b; 1983); Unknown site – 1. Late Miocene; Byala Slatina (Vratsa Region): (Lovech Region). Listed as “one of the Upper Pliocene – Lavant (BAKALOV & Karlukovo caves”. Listed as Rhinoceros tichorhinus NIKOLOV, 1962); Pliocene (NIKOLOV, 1985). Cuvier, 1812 (NIKOLOV, 1983); Unknown site – Listed as Dicerorhinus schleiermacheri Kaup, 1834; 2. Pirgovo (Ruse Region). Caves near Pirgovo Kula (Vidin Regon): Upper Pliocene (BAKALOV village. Pleistocene (KOVACHEV, 1906; & NIKOLOV, 1962); Maeotian (NIKOLOV, 1985) NIKOLOV, 1977). Listed as Rhinoceros tichorhinus – Late Miocene. Listed as Dicerorhinus Cuvier, 1812 (KOVACHEV, 1906); Mirizlivka schleiermacheri Kaup, 1834; Altimir (Vratsa cave (Vidin Region) FP 43. Wurmian – Late Region): Upper Pliocene (BAKALOV & Pleistocene (POPOV, 1933). Listed as Rhinoceros NIKOLOV,
Load more

Recommended publications
  • New Materials of Chalicotherium Brevirostris (Perissodactyla, Chalicotheriidae)
    Geobios 45 (2012) 369–376 Available online at www.sciencedirect.com Original article New materials of Chalicotherium brevirostris (Perissodactyla, Chalicotheriidae) § from the Tunggur Formation, Inner Mongolia Yan Liu *, Zhaoqun Zhang Laboratory of Evolutionary Systematics of Vertebrate, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 142, Xizhimenwai Street, PO Box 643, Beijing 100044, PR China A R T I C L E I N F O A B S T R A C T Article history: Chalicotherium brevirostris was named by Colbert based on a skull lacking mandibles from the late Middle Received 23 March 2011 Miocene Tunggur Formation, Tunggur, Inner Mongolia, China. Here we describe new mandibular Accepted 6 October 2011 materials collected from the same area. In contrast to previous expectations, the new mandibular Available online 14 July 2012 materials show a long snout, long diastema, a three lower incisors and a canine. C. brevirostris shows some sexual dimorphism and intraspecific variation in morphologic characters. The new materials differ Keywords: from previously described C. cf. brevirostris from Cixian County (Hebei Province) and the Tsaidam Basin, Tunggur which may represent a different, new species close to C. brevirostris. The diagnosis of C. brevirostris is Middle Miocene revised. Chalicotheriinae Chalicotherium ß 2012 Elsevier Masson SAS. All rights reserved. 1. Abbreviations progress in understanding the Tunggur geology and paleontology (Qiu et al., 1988). The most important result of this expedition so far is
    [Show full text]
  • Navigation Map of Bulgaria Including Offroadmap by Offroad-Bulgaria.Com Version 2021 Q1
    Navigation Map of Bulgaria Including OFFRoadMap by OFFRoad-Bulgaria.com Version 2021 Q2 The purpose of this map is to provide accessible, accurate and up-to-date information for your GPS devices. Despite all efforts made by the creators to achieve this goal, the roads and the data included in this digital map are intended to be used as guidance only and should not be used solely for navigation. The creators of this map make no warranty as to the accuracy or completeness of the map data. In no event will the creators of this map be liable for any damages whatsoever, including but not limited to loss of revenue or profit, lost or damaged data, and expenses, arising in any way from or consequential upon the use of, or the inability to use this digital map. Contents: - Registering your map - Usage details - OFRM Geotrade 2021 Q2 variants - Coverage >>>>> REGISTRATION <<<<< To register your OFRM Geotrade map, please visit out website www.karta.bg. Click on “Create profile” in the top right corner of the screen and create your personal account. When done, the Support page will load automatically. Click on the button “Register OFRM Geotrade” and enter the 25-symbol map serial number and GPS model to activate your map’s update subscription (if your map includes one). To obtain the 25-symbol serial number, connect your GPS device to your computer via USB cable. If you have a GPS device with preloaded OFRM map, you will find the serial number in file “serial.txt” in the root folder of your device’s base memory or in the file “gmapsupp.unl” in folder “Garmin” (or folder “Map” on the newer models of the nüvi series and the new Drive series) of your device’s base memory.
    [Show full text]
  • Espaço E Economia, 10 | 2017 Alternative Tourism in Bulgaria – General Characteristics 2
    Espaço e Economia Revista brasileira de geografia econômica 10 | 2017 Ano V, número 10 Alternative tourism in Bulgaria – general characteristics Turismo alternativo na Bulgária – características gerais Le tourisme alternatif en Bulgarie : traits générales Turismo alternativo en Bulgaria: características generales. Milen Penerliev Electronic version URL: http://journals.openedition.org/espacoeconomia/2921 DOI: 10.4000/espacoeconomia.2921 ISSN: 2317-7837 Publisher Núcleo de Pesquisa Espaço & Economia Electronic reference Milen Penerliev, « Alternative tourism in Bulgaria – general characteristics », Espaço e Economia [Online], 10 | 2017, Online since 17 July 2017, connection on 19 April 2019. URL : http:// journals.openedition.org/espacoeconomia/2921 ; DOI : 10.4000/espacoeconomia.2921 This text was automatically generated on 19 April 2019. © NuPEE Alternative tourism in Bulgaria – general characteristics 1 Alternative tourism in Bulgaria – general characteristics Turismo alternativo na Bulgária – características gerais Le tourisme alternatif en Bulgarie : traits générales Turismo alternativo en Bulgaria: características generales. Milen Penerliev 1 Alternative tourism is a form of tourism which represents an alternative to the conventional mass tourism. Its forms are small-scale, low-density, divided into forms practiced in urban (religious, cultural, business) and rural areas. Alternative tourism is an attempt to preserve, protect and improve the quality of the existing resource base, which is fundamental for tourism itself. Alternative tourism is featured by active encouragement and care for the development of additional andspecific attractions as well the infrastructure, which are based on the local resources, while at the same time aiding the latter. It has an impact on the quality of life in the region, improves the infrastructure and increases the educational and cultural level of the local community.
    [Show full text]
  • 1 I. ANNEXES 1 Annex 6. Map and List of Rural Municipalities in Bulgaria
    I. ANNEXES 1 Annex 6. Map and list of rural municipalities in Bulgaria (according to statistical definition). 1 List of rural municipalities in Bulgaria District District District District District District /Municipality /Municipality /Municipality /Municipality /Municipality /Municipality Blagoevgrad Vidin Lovech Plovdiv Smolyan Targovishte Bansko Belogradchik Apriltsi Brezovo Banite Antonovo Belitsa Boynitsa Letnitsa Kaloyanovo Borino Omurtag Gotse Delchev Bregovo Lukovit Karlovo Devin Opaka Garmen Gramada Teteven Krichim Dospat Popovo Kresna Dimovo Troyan Kuklen Zlatograd Haskovo Petrich Kula Ugarchin Laki Madan Ivaylovgrad Razlog Makresh Yablanitsa Maritsa Nedelino Lyubimets Sandanski Novo Selo Montana Perushtitsa Rudozem Madzharovo Satovcha Ruzhintsi Berkovitsa Parvomay Chepelare Mineralni bani Simitli Chuprene Boychinovtsi Rakovski Sofia - district Svilengrad Strumyani Vratsa Brusartsi Rodopi Anton Simeonovgrad Hadzhidimovo Borovan Varshets Sadovo Bozhurishte Stambolovo Yakoruda Byala Slatina Valchedram Sopot Botevgrad Topolovgrad Burgas Knezha Georgi Damyanovo Stamboliyski Godech Harmanli Aitos Kozloduy Lom Saedinenie Gorna Malina Shumen Kameno Krivodol Medkovets Hisarya Dolna banya Veliki Preslav Karnobat Mezdra Chiprovtsi Razgrad Dragoman Venets Malko Tarnovo Mizia Yakimovo Zavet Elin Pelin Varbitsa Nesebar Oryahovo Pazardzhik Isperih Etropole Kaolinovo Pomorie Roman Batak Kubrat Zlatitsa Kaspichan Primorsko Hayredin Belovo Loznitsa Ihtiman Nikola Kozlevo Ruen Gabrovo Bratsigovo Samuil Koprivshtitsa Novi Pazar Sozopol Dryanovo
    [Show full text]
  • Bulgaria 2013 International Religious Freedom Report
    BULGARIA 2013 INTERNATIONAL RELIGIOUS FREEDOM REPORT Executive Summary The constitution and other laws and policies generally protect religious freedom and, in practice, the government mostly respected religious freedom. The constitution designates Eastern Orthodox Christianity as the “traditional” religion, exempting its branch in the country, the Bulgarian Orthodox Church (BOC), from the court registration required for all other religious groups. Members of some minority religious groups expressed concern that the government did not proactively intervene to prevent societal abuses. Reports of intolerance by security services and local authorities continued. There were reports of societal abuses and discrimination based on religious affiliation, belief, or practice. Discrimination, harassment, and general public intolerance of some religious groups remained a persistent problem. Anti-Semitic incidents and vandalism against mosques and places of worship persisted. The U.S. embassy regularly communicated with government officials and religious leaders to discuss and promote religious freedom, and facilitated dialogue among religious groups. Section I. Religious Demography The U.S. government estimates the population at 7 million (July 2013 estimate). According to the 2011 census, 76 percent of the population identifies itself as Orthodox Christian. Orthodox Christianity, Hanafi Sunni Islam, Judaism, and Roman Catholicism all hold a historic place in the country’s culture. Muslims are the second-largest religious group, estimated at 10 percent of the population. Groups that together constitute about 2 percent of the population include Catholics, Armenian Christians, Jews, evangelical Protestants, Jehovah’s Witnesses, the Church of Jesus Christ of Latter-day Saints (Mormons), and others. Nearly 12 percent of respondents did not identify with any of the above faiths.
    [Show full text]
  • 71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
    ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas.
    [Show full text]
  • Distribution of Entomopathogenic Nematodes of the Genus Heterorhabditis (Rhabditida: Heterorhabditidae) in Bulgaria
    13 Gradinarov_173 7-01-2013 9:34 Pagina 173 Nematol. medit. (2012), 40: 173-180 173 DISTRIBUTION OF ENTOMOPATHOGENIC NEMATODES OF THE GENUS HETERORHABDITIS (RHABDITIDA: HETERORHABDITIDAE) IN BULGARIA D. Gradinarov1*, E. Petrova**, Y. Mutafchiev***, O. Karadjova** * Department of Zoology and Anthropology, Faculty of Biology, Sofia University “St. Kliment Ohridski”, 8 Dragan Tzankov Blvd., 1164 Sofia, Bulgaria ** Institute of Soil Science, Agrotechnology and Plant Protection “N. Pushkarov”, Division of Plant Protection, Kostinbrod, Bulgaria *** Institute of Biodiversity and Ecosystem Research, 2 Gagarin Str., 1113 Sofia, Bulgaria Received: 21 September 2012; Accepted: 21 November 2012. Summary. The results from studies on entomopathogenic nematodes of the genus Heterorhabditis Poinar, 1976 (Rhabditida: Het- erorhabditidae) in Bulgaria, conducted during 1994-2010 are summarized. Of the 1,227 soil samples collected, 3.5% were positive for the presence of Heterorhabditis spp. Specimens belonging to the genus were obtained from 43 soil samples collected at 27 lo- calities in different regions of the country. Heterorhabditids were established at altitudes from 0 to 1175 m, in habitats both along the Black Sea coast and inland. The prevalent species was H. bacteriophora Poinar, 1976. Its identity was confirmed by detailed morphometric studies and molecular analyses of four recently obtained isolates. Inland, H. bacteriophora prefers alluvial soils in river valleys under herbaceous and woody vegetation. It was also found in calcareous soils with pronounced fluctuations in the temperature and water conditions. The presence of the species H. megidis Poinar, Jackson et Klein, 1987 in Bulgaria needs further confirmation. Key words: Heterorhabditis bacteriophora, morphology, molecular identification, habitat preferences. Entomopathogenic nematodes (EPNs) (Rhabditida: processing of 1,227 soil samples collected during the pe- Steinernematidae, Heterorhabditidae) are obligate para- riod November, 1994 to October, 2010 from different sites of a wide range of soil insects.
    [Show full text]
  • І Railway Line Kalotina Zapad
    TABLE ON CONSTANT SPEED REDUCTIONS IN THE TRAIN TIMETABLE 2020/2021 POSITION SPEED Railway TRAC FRONT N FROM STATION TO STATION reductio REASONS FOR REDUCTION section K No 1 from km to km (m) KR n І railway line Kalotina Zapad - Svilengrad 1 Sf Sofia Voluyak 1 0+000 0+800 800 60 25 Switches at Sofia station 2 Sf Sofia Voluyak 2 0+000 0+800 800 60 25 Switches at Sofia station 3 Sf Voluyak station 8+348 8+448 100 40 5 Worsen technical condition of the rail-sleeper grid and worsen technical condition by axle and level 4 Sf Dragoman station 42+050 42+540 490 70 25 Worsen technical condition of the railway of tracks NN 1, 2, 3 (main), 4, 5 and switches 5 Sf Dragoman station 42+100 42+620 520 40 15 Worsen technical condition of the permanent way of tracks NN 1, 2, 3 (main), 4, 6 and switches 6 Sf Sofia Poduyane 1 0+000 1+100 1 100 60 25 Switches at Sofia station 7 Sf Sofia Poduyane 2 0+000 1+100 1 100 60 25 Switches at Sofia station 8 Sf Iskar station 8+275 9+560 1 285 40 25 Worsen technical condition of the railway of tracks NN 5, 7, 9, 11 and 13 9 Sf Verinsko Ihtiman 1 55+623 56+514 891 60 50 Worsen technical condition of the railway at Ihtiman station 10 Sf Verinsko Ihtiman 2 55+740 56+514 774 60 50 Worsen technical condition of the railway at Ihtiman station 11 Sf Ihtiman stop Nemirovo 1 56+514 57+845 1 331 60 50 Worsen technical condition of the railway at Ihtiman station 12 Sf Ihtiman stop Nemirovo 2 56+514 57+845 1 331 60 50 Worsen technical condition of the railway at Ihtiman station 13 Sf stop Nemirovo Kostenets 1 74+000 74+349 349
    [Show full text]
  • An Early Miocene Dome-Skulled Chalicothere
    PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, NY 10024 Number 3486, 45 pp., 26 ®gures, 8 tables October 27, 2005 An Early Miocene Dome-Skulled Chalicothere from the ``Arikaree'' Conglomerates of Darton: Calibrating the Ages of High Plains Paleovalleys Against Rocky Mountain Tectonism ROBERT M. HUNT, JR.1 CONTENTS Abstract ....................................................................... 2 Introduction .................................................................... 2 Geologic Setting ................................................................ 3 Systematic Paleontology ........................................................ 12 Age of the Dome-Skulled Chalicothere ........................................... 28 The ``Arikaree'' Conglomerates of N.H. Darton ................................... 32 Post-Laramide Evolution of the Rocky Mountains ................................. 39 Acknowledgments ............................................................. 42 References .................................................................... 42 1 Department of Geosciences, W436 Nebraska Hall, University of Nebraska, Lincoln, NE 68588-0549 ([email protected]). Copyright q American Museum of Natural History 2005 ISSN 0003-0082 2 AMERICAN MUSEUM NOVITATES NO. 3486 ABSTRACT Fragmentary skeletal remains discovered in 1979 in southeastern Wyoming, associated with a mammalian fauna of early Hemingfordian age (;18.2 to 18.8 Ma), represent the oldest known occurrence of dome-skulled chalicotheres
    [Show full text]
  • A Checklist of the Herpetofauna in the Bulgarian Part of Hadzhidimovo Gorge (South-Western Bulgaria)
    ECOLOGIA BALKANICA 2019, Vol. 11, Issue 1 June 2019 pp. 17-26 A Checklist of the Herpetofauna in the Bulgarian Part of Hadzhidimovo Gorge (South-Western Bulgaria) George N. Manolev, Lilia V. Philipova, Alexander N. Pulev*, Lidia G. Sakelarieva South-West University “Neofit Rilski”, Faculty of Mathematics and Natural Sciences, Department of Geography, Ecology and Environmental Protection, 66 Ivan Mihailov Str., 2700 Blagoevgrad, BULGARIA *Corresponding author: [email protected] Abstract. The Bulgarian part of Hadzhidimovo Gorge (the Mesta River basin, south-western Bulgaria) was studied in March-October 2018 in order to collect data about the local herpetofauna. A preliminary checklist of the amphibians and reptiles is presented, based on bibliographic records and the field research. The list includes 25 species. The representatives of the class Amphibia are 8 (the half of them are new for the area), and the reptiles are 17 (5 of them are reported for the first time). The field records for all new species (except Pelophylax ridibundus complex) as well as for the most of the known species interesting from zoogeographic point of view are specified. It is of particular interest the discovery of Triturus ivanbureschi, Podarcis muralis, Ablepharus kitaibelii, and Platyceps najadum in the gorge. Key words: Amphibia, Reptilia, checklist, Hadzhidimovo Gorge, Bulgaria. Introduction observations. All the amphibians reported till The territory of Bulgaria has been studied to now belong to the order Anura (BURESCH & a very different extent as regards the ZONKOV, 1942; PETROV et al., 2006; NATURA 2000, herpetofauna. In one of his publications BESHKOV 2013). The reptilians are representatives of the (1993) points to the less studied areas in the order Testudines – 2 species (DOMOZETSKI, 2013; country.
    [Show full text]
  • Identity, Nationalism, and Cultural Heritage Under Siege Balkan Studies Library
    Identity, Nationalism, and Cultural Heritage under Siege Balkan Studies Library Editor-in-Chief Zoran Milutinović (University College London) Editorial Board Gordon N. Bardos (Columbia University) Alex Drace-Francis (University of Amsterdam) Jasna Dragović-Soso (Goldsmiths, University of London) Christian Voss, (Humboldt University, Berlin) Advisory Board Marie-Janine Calic (University of Munich) Lenard J. Cohen (Simon Fraser University Radmila Gorup (Columbia University) Robert M. Hayden (University of Pittsburgh) Robert Hodel (Hamburg University) Anna Krasteva (New Bulgarian University) Galin Tihanov (Queen Mary, University of London) Maria Todorova (University of Illinois) Andrew Wachtel (Northwestern University) VOLUME 14 The titles published in this series are listed at brill.com/bsl Identity, Nationalism, and Cultural Heritage under Siege Five Narratives of Pomak Heritage—From Forced Renaming to Weddings By Fatme Myuhtar-May LEIDEN | BOSTON Cover illustration: Pomak bride in traditional attire. Ribnovo, Rhodope Mountains, Bulgaria. Photo courtesy Kimile Ulanova of Ribnovo. Library of Congress Cataloging-in-Publication Data Myuhtar-May, Fatme. Cultural heritage under siege : five narratives of Pomak heritage : from forced renaming to weddings / by Fatme Myuhtar-May. pages cm. — (Balkan studies library, ISSN 1877-6272 ; volume 14) Includes bibliographical references and index. ISBN 978-90-04-27207-1 (hardback : acid-free paper) — ISBN 978-90-04-27208-8 (e-book) 1. Pomaks—Bulgaria—Social conditions. 2. Pomaks—Bulgaria—Social life and customs. 3. Pomaks— Bulgaria—Case studies. 4. Pomaks—Bulgaria—Biography. 5. Culture conflict—Bulgaria. 6. Culture conflict—Rhodope Mountains Region. 7. Bulgaria—Ethnic relations. 8. Rhodope Mountains Region— Ethnic relations. I. Title. DR64.2.P66M98 2014 305.6’970499—dc23 2014006975 This publication has been typeset in the multilingual ‘Brill’ typeface.
    [Show full text]
  • Middle Miocene Paleoenvironmental Reconstruction of the Central Great Plains from Stable Carbon Isotopes in Large Mammals Willow H
    University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Dissertations & Theses in Earth and Atmospheric Earth and Atmospheric Sciences, Department of Sciences 7-2017 Middle Miocene Paleoenvironmental Reconstruction of the Central Great Plains from Stable Carbon Isotopes in Large Mammals Willow H. Nguy University of Nebraska-Lincoln, [email protected] Follow this and additional works at: http://digitalcommons.unl.edu/geoscidiss Part of the Geology Commons, Paleobiology Commons, and the Paleontology Commons Nguy, Willow H., "Middle Miocene Paleoenvironmental Reconstruction of the Central Great Plains from Stable Carbon Isotopes in Large Mammals" (2017). Dissertations & Theses in Earth and Atmospheric Sciences. 91. http://digitalcommons.unl.edu/geoscidiss/91 This Article is brought to you for free and open access by the Earth and Atmospheric Sciences, Department of at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Dissertations & Theses in Earth and Atmospheric Sciences by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. MIDDLE MIOCENE PALEOENVIRONMENTAL RECONSTRUCTION OF THE CENTRAL GREAT PLAINS FROM STABLE CARBON ISOTOPES IN LARGE MAMMALS by Willow H. Nguy A THESIS Presented to the Faculty of The Graduate College at the University of Nebraska In Partial Fulfillment of Requirements For the Degree of Master of Science Major: Earth and Atmospheric Sciences Under the Supervision of Professor Ross Secord Lincoln, Nebraska July, 2017 MIDDLE MIOCENE PALEOENVIRONMENTAL RECONSTRUCTION OF THE CENTRAL GREAT PLAINS FROM STABLE CARBON ISOTOPES IN LARGE MAMMALS Willow H. Nguy, M.S. University of Nebraska, 2017 Advisor: Ross Secord Middle Miocene (18-12 Mya) mammalian faunas of the North American Great Plains contained a much higher diversity of apparent browsers than any modern biome.
    [Show full text]