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INTERTIDAL ZONATION Introduction to Oceanography Spring 2017 The
INTERTIDAL ZONATION Introduction to Oceanography Spring 2017 The Intertidal Zone is the narrow belt along the shoreline lying between the lowest and highest tide marks. The intertidal or littoral zone is subdivided broadly into four vertical zones based on the amount of time the zone is submerged. From highest to lowest, they are Supratidal or Spray Zone Upper Intertidal submergence time Middle Intertidal Littoral Zone influenced by tides Lower Intertidal Subtidal Sublittoral Zone permanently submerged The intertidal zone may also be subdivided on the basis of the vertical distribution of the species that dominate a particular zone. However, zone divisions should, in most cases, be regarded as approximate! No single system of subdivision gives perfectly consistent results everywhere. Please refer to the intertidal zonation scheme given in the attached table (last page). Zonal Distribution of organisms is controlled by PHYSICAL factors (which set the UPPER limit of each zone): 1) tidal range 2) wave exposure or the degree of sheltering from surf 3) type of substrate, e.g., sand, cobble, rock 4) relative time exposed to air (controls overheating, desiccation, and salinity changes). BIOLOGICAL factors (which set the LOWER limit of each zone): 1) predation 2) competition for space 3) adaptation to biological or physical factors of the environment Species dominance patterns change abruptly in response to physical and/or biological factors. For example, tide pools provide permanently submerged areas in higher tidal zones; overhangs provide shaded areas of lower temperature; protected crevices provide permanently moist areas. Such subhabitats within a zone can contain quite different organisms from those typical for the zone. -
A 16-Year Record of Green and Hawksbill Turtle Nesting Activity at Chagar Hutang Turtle Sanctuary, Redang Island, Malaysia
Indian Ocean Turtle Newsletter No. 12 A 16-year record of green and hawksbill turtle nesting activity at Chagar Hutang Turtle Sanctuary, Redang Island, Malaysia Eng-Heng Chan1 Turtle Conservation Centre Terengganu, Malaysia Email: [email protected] Introduction South China Sea Redang Island Kuala Redang Island (5º 44’ – 5º 50’ N and 102º 59’ – 103º 5’ E) Terengganu is located in the South China Sea, off the east coast of N State of Terengganu Peninsular Malaysia (Fig. 1). It has a land area of about Chagar Hutang Turtle Sanctuary 25 square km and is 45 km north northeast of Kuala Terengganu. It is a popular tourist destination and also Mak Kepit T. Mat Delah Teluk home to the largest aggregation of nesting green turtles Pasir Bujang Dalam Mak Simpan in Peninsular Malaysia. However, overall nesting Che Keling Berjaya Redang Beach Resort Paku density in Peninsular Malaysia is low, compared to Besar Island REDANG Pasir ISLAND Panjang Paku Kecil the Sabah Turtle Islands Park (STIP) in East Malaysia. Island Lima Island T. Betigi Average annual nesting density over the last five years T. Kalong Pasir Kecil Kerengga Kecil Island for green turtles in the STIP was 6,500 and 2,300 for Kg. Lama Kerengga Besar Terengganu (Chan, 2009). Total number of egg clutches Island Turtle MARINE PARK HQ Sanctuaries deposited on Redang Island account for 50-60% of the Pinang Island Ekor Tebu total recorded for the whole of Terengganu State. Island Ling Island 00 11 2 2 33 km km Turtle nesting beaches in Redang Island (Fig. 1) were declared sanctuaries only as recently as 2005. -
Fishes of Terengganu East Coast of Malay Peninsula, Malaysia Ii Iii
i Fishes of Terengganu East coast of Malay Peninsula, Malaysia ii iii Edited by Mizuki Matsunuma, Hiroyuki Motomura, Keiichi Matsuura, Noor Azhar M. Shazili and Mohd Azmi Ambak Photographed by Masatoshi Meguro and Mizuki Matsunuma iv Copy Right © 2011 by the National Museum of Nature and Science, Universiti Malaysia Terengganu and Kagoshima University Museum All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means without prior written permission from the publisher. Copyrights of the specimen photographs are held by the Kagoshima Uni- versity Museum. For bibliographic purposes this book should be cited as follows: Matsunuma, M., H. Motomura, K. Matsuura, N. A. M. Shazili and M. A. Ambak (eds.). 2011 (Nov.). Fishes of Terengganu – east coast of Malay Peninsula, Malaysia. National Museum of Nature and Science, Universiti Malaysia Terengganu and Kagoshima University Museum, ix + 251 pages. ISBN 978-4-87803-036-9 Corresponding editor: Hiroyuki Motomura (e-mail: [email protected]) v Preface Tropical seas in Southeast Asian countries are well known for their rich fish diversity found in various environments such as beautiful coral reefs, mud flats, sandy beaches, mangroves, and estuaries around river mouths. The South China Sea is a major water body containing a large and diverse fish fauna. However, many areas of the South China Sea, particularly in Malaysia and Vietnam, have been poorly studied in terms of fish taxonomy and diversity. Local fish scientists and students have frequently faced difficulty when try- ing to identify fishes in their home countries. During the International Training Program of the Japan Society for Promotion of Science (ITP of JSPS), two graduate students of Kagoshima University, Mr. -
SEATRU Newsletter 2010
Vol. 2, No.1 December 2010 SEATRU Newsletter SEATRU Updates on Conservation, Research and Outreach Programs 2010 Message from the Team Leader This year seems to have flown by in a blink. Without realizing it, we are almost at the end of 2010. This year was the second year for my team with SEATRU. With hard work, dedication and commitments from my team members, I can conclude that this year went off better compared to last year. Off course we have encountered few obstacles along the way, but when we work as a team definitely we could overcome each hurdle. The best thing was, as of the previous year we had fun in performing our work, and we have ended this season with joy and laughter! This year the nesting at Chagar Hutang had increased compared to last year. Monitoring was conducted for the whole year from January to December 2010. We have recorded a total of 676 nesting, of which 661 were green turtle nests and 15 were hawksbill turtle nests. From these a total of 40,000 healthy hatchlings were returned back to the sea. Our sincere thanks to all SEATRU volunteers (Slot A to Slot Z) for helping us in monitoring and protecting turtle nests at Chagar Hutang. It has been 18 years since SEATRU first started the sea turtle conservation program at Chagar Hutang. The unit is still very much active and dedicated in saving the sea turtles even though project leaders have changed hands. It is because of your support and action that motivate us to keep on going. -
Life in the Intertidal Zone
Life in the Rocky Intertidal Zone Tidepools provide a home for many animals. Tidepools are created by the changing water level, or tides. The high energy waves make this a harsh habitat, but the animals living here have adapted over time. When the earth, sun and moon align during the full and new moon we have extreme high and low tides. Generally, there are two high tides and two low tides a day. An example of low and high tide is seen on the right. There are three zones within the tidepools: the high zone, the middle zone, and the low zone. Animals are distributed based on their adaptations to different living (competition and predation) and non living (wave action and water loss) factors. The tidepools at Cabrillo are protected and have been monitored by the National Park Service since 1990. You may notice bolts in Low Tide the rocky intertidal, these are used to assist scientists in gathering data to monitor changes. Tidepool Etiquette: Human impact can hurt the animals. As you explore the tidepools, you may touch the animals living here, but only as gently as you would touch your own eyeball. Some animals may die if moved even a few inches from where they are found. Federal law prohibits collection and removal of any shells, rocks and marine specimens. Also, be aware of the changing tides, slippery rocks and unstable cliffs. Have fun exploring! High Tide High Zone The high zone is covered by the highest tides. Often this area is only sprayed by the (Supralittoral or crashing waves. -
Spatial Patterns of Macrozoobenthos Assemblages in a Sentinel Coastal Lagoon: Biodiversity and Environmental Drivers
Journal of Marine Science and Engineering Article Spatial Patterns of Macrozoobenthos Assemblages in a Sentinel Coastal Lagoon: Biodiversity and Environmental Drivers Soilam Boutoumit 1,2,* , Oussama Bououarour 1 , Reda El Kamcha 1 , Pierre Pouzet 2 , Bendahhou Zourarah 3, Abdelaziz Benhoussa 1, Mohamed Maanan 2 and Hocein Bazairi 1,4 1 BioBio Research Center, BioEcoGen Laboratory, Faculty of Sciences, Mohammed V University in Rabat, 4 Avenue Ibn Battouta, Rabat 10106, Morocco; [email protected] (O.B.); [email protected] (R.E.K.); [email protected] (A.B.); [email protected] (H.B.) 2 LETG UMR CNRS 6554, University of Nantes, CEDEX 3, 44312 Nantes, France; [email protected] (P.P.); [email protected] (M.M.) 3 Marine Geosciences and Soil Sciences Laboratory, Associated Unit URAC 45, Faculty of Sciences, Chouaib Doukkali University, El Jadida 24000, Morocco; [email protected] 4 Institute of Life and Earth Sciences, Europa Point Campus, University of Gibraltar, Gibraltar GX11 1AA, Gibraltar * Correspondence: [email protected] Abstract: This study presents an assessment of the diversity and spatial distribution of benthic macrofauna communities along the Moulay Bousselham lagoon and discusses the environmental factors contributing to observed patterns. In the autumn of 2018, 68 stations were sampled with three replicates per station in subtidal and intertidal areas. Environmental conditions showed that the range of water temperature was from 25.0 ◦C to 12.3 ◦C, the salinity varied between 38.7 and 3.7, Citation: Boutoumit, S.; Bououarour, while the average of pH values fluctuated between 7.3 and 8.0. -
Intertidal Sand and Mudflats & Subtidal Mobile
INTERTIDAL SAND AND MUDFLATS & SUBTIDAL MOBILE SANDBANKS An overview of dynamic and sensitivity characteristics for conservation management of marine SACs M. Elliott. S.Nedwell, N.V.Jones, S.J.Read, N.D.Cutts & K.L.Hemingway Institute of Estuarine and Coastal Studies University of Hull August 1998 Prepared by Scottish Association for Marine Science (SAMS) for the UK Marine SACs Project, Task Manager, A.M.W. Wilson, SAMS Vol II Intertidal sand and mudflats & subtidal mobile sandbanks 1 Citation. M.Elliott, S.Nedwell, N.V.Jones, S.J.Read, N.D.Cutts, K.L.Hemingway. 1998. Intertidal Sand and Mudflats & Subtidal Mobile Sandbanks (volume II). An overview of dynamic and sensitivity characteristics for conservation management of marine SACs. Scottish Association for Marine Science (UK Marine SACs Project). 151 Pages. Vol II Intertidal sand and mudflats & subtidal mobile sandbanks 2 CONTENTS PREFACE 7 EXECUTIVE SUMMARY 9 I. INTRODUCTION 17 A. STUDY AIMS 17 B. NATURE AND IMPORTANCE OF THE BIOTOPE COMPLEXES 17 C. STATUS WITHIN OTHER BIOTOPE CLASSIFICATIONS 25 D. KEY POINTS FROM CHAPTER I. 27 II. ENVIRONMENTAL REQUIREMENTS AND PHYSICAL ATTRIBUTES 29 A. SPATIAL EXTENT 29 B. HYDROPHYSICAL REGIME 29 C. VERTICAL ELEVATION 33 D. SUBSTRATUM 36 E. KEY POINTS FROM CHAPTER II 43 III. BIOLOGY AND ECOLOGICAL FUNCTIONING 45 A. CHARACTERISTIC AND ASSOCIATED SPECIES 45 B. ECOLOGICAL FUNCTIONING AND PREDATOR-PREY RELATIONSHIPS 53 C. BIOLOGICAL AND ENVIRONMENTAL INTERACTIONS 58 D. KEY POINTS FROM CHAPTER III 65 IV. SENSITIVITY TO NATURAL EVENTS 67 A. POTENTIAL AGENTS OF CHANGE 67 B. KEY POINTS FROM CHAPTER IV 74 V. SENSITIVITY TO ANTHROPOGENIC ACTIVITIES 75 A. -
Wave Energy and Intertidal Productivity (Leaf Area Index/Myfilus Californianus/Postelsia/Rocky Shore/Zonation) EGBERT G
Proc. Natl. Acad. Sci. USA Vol. 84, pp. 1314-1318, March 1987 Ecology Wave energy and intertidal productivity (leaf area index/Myfilus californianus/Postelsia/rocky shore/zonation) EGBERT G. LEIGH, JR.*, ROBERT T. PAINEt, JAMES F. QUINNt, AND THOMAS H. SUCHANEKt§ *Smithsonian Tropical Research Institute, Apartado 2072, Balboa, Panama; tDepartment of Zoology NJ-15, University of Washington, Seattle, WA 98195; tDivision of Environmental Studies, University of California at Davis, Davis, CA 95616; and §Bodega Marine Laboratory, P. 0. Box 247, Bodega Bay, CA 94923 Contributed by Robert T. Paine, October 20, 1986 ABSTRACT In the northeastern Pacific, intertidal zones we estimate standing crop and productivity in different zones of the most wave-beaten shores receive more energy from of exposed and sheltered rocky shores of Tatoosh Island (480 breaking waves than from the sun. Despite severe mortality 19' N, 1240 40' W), showing that intertidal productivity is from winter storms, communities at some wave-beaten sites much higher in wave-beaten settings. Finally, we consider produce an extraordinary quantity of dry matter per unit area the various roles waves may play in enhancing the produc- of shore per year. At wave-beaten sites of Tatoosh Island, WA, tivity of intertidal organisms. sea palms, Postelsia palmaeformis, can produce >10 kg of dry matter, or 1.5 x 108 J. per m2 in a good year. Extraordinarily METHODS productive organisms such as Postelsia are restricted to wave- The Power Supplied by Breaking Waves. Calculating the beaten sites. Intertidal organisms cannot transform wave power carried by waves in the open ocean. Waves generated energy into chemical energy, as photosynthetic plants trans- anywhere in the ocean-dissipate most of their energy against form solar energy, nor can intertidal organisms "harness" some shore as surf (13, 14). -
Intertidal Zonation of Two Gastropods, Nerita Plicata and Morula Granulata, in Moorea, French Polynesia
INTERTIDAL ZONATION OF TWO GASTROPODS, NERITA PLICATA AND MORULA GRANULATA, IN MOOREA, FRENCH POLYNESIA VANESSA R. WORMSER Integrative Biology, University of California, Berkeley, California 94720 USA Abstract. Intertidal zonation of organisms is a key factor in ecological community structure and the existence of fundamental and realized niches. The zonation of two species of gastropods, Nerita plicata and Morula granulata were investigated using field observations and lab experimentation. The Nerita plicata were found on the upper limits of the intertidal zone while the Morula granulata were found on the lower limits. The distribution of each species was observed and the possible causes of this zonation were examined. Three main factors, desiccation, flow resistance and shell size were tested for their zonation. In the field, shell measurements of each species were made to see if a vertical shell size gradient existed; the results showed an upshore shell size gradient for each species. In the lab, experiments were run to see if the zonation preference found in the field existed in the lab as well. This experiment confirmed that a zonation between these species does in fact exist. Additional experiments were run to test desiccation and flow resistance between each species. A difference in desiccation rates and flow resistance were found with the Nerita plicata being more resistant to both flow and desiccation. The findings of this study provide an understanding on why zonation between these two species could exist as well as why zonation is important within an intertidal community and ecosystems as a whole. Key words: community structure; gastropod; zonation; intertidal; morphometrics; Morula granulata; Nerita plicata; Mo’orea, French Polynesia; INTRODUCTION The main goal of an ecological survey is to because of the high species diversity, the explore and understand the key dynamic convenience of the habitat as well as the easy relationships among organisms living in a collection of the sessile organisms that inhabit community (Elton 1966). -
Kampus UMT Bukit Kor UMT Terima Dua TNC Baharu Meneroka Lautan Menjana Khazanah Ilmuwan UMT Wakili Malaysia Ke ASEAN Food Conference Dari Minda Ketua Pengarang
UNIVERSITI MALAYSIA TERENGGANU | www.umt.edu.my INFOKUSBil. 45 JUN 2013 Lawatan NC ke Amerika Syarikat Kampus UMT Bukit Kor UMT terima dua TNC baharu Meneroka lautan Menjana Khazanah Ilmuwan UMT wakili Malaysia ke ASEAN Food Conference Dari Minda Ketua Pengarang Assalamualaikum, Salam Sejahtera dan Salam 1Malaysia. Syukur Alhamdulillah kerana kita masih diizinkan-Nya untuk bertemu dalam INFOKUS edisi JUN 2013. Tahun 2013 dilakari dengan detik paling bersejarah bagi negara kecintaan kita. Teraju negara dimeterai dengan kejayaan kepimpinan yang ikhlas bekerja untuk kesejahteraan rakyat dan kejayaan ini atas kehendak-Nya jua. Syukur Alhamdulillah. Syabas kita ucapkan kepada YAB Dato’ Seri Najib Tun Abdul Razak, Perdana Menteri Malaysia dan barisan kepimpinan beliau yang bekerja keras untuk memastikan kejayaan digapai. Penerokaan beliau hingga ke akar umbi kehidupan rakyat jelata “Pepatah Melayu sering melagukan… menghasilkan bibit cinta terhadap negara yang Orang berbudi kita berbahasa, orang mendambakan keamanan dan kedamaian demi keutuhan sebuah kehidupan. memberi kita merasa… Namun fenomena Syabas dan tahniah juga kita hulurkan kini amat berbeza…orang berbudi sering untuk YAB Tan Sri Muhyiddin Yassin, Timbalan Perdana Menteri Malaysia yang diberi mandat dilupa walaupun dekat di mata…apa yang untuk menerajui Kementerian Pelajaran dan nyata hanyalah dosa-dosa yang kadangkala Pengajian Tinggi dan YB Dato Seri Idris Jusoh yang telah dilantik sebagai Menteri Pendidikan II ditimbang tara tanpa pembela…Namun yang baharu. Kami warga pendidik mengalu-alukan taburlah jasa walaupun tidak dikenang”. kehadiran kedua-dua pemimpin berkaliber ini ke dunia pendidikan yang amat mencabar demi untuk mendidik anak bangsa meniti kejayaan hidup dalam (Noraien Mansor, Jun 2013) pelbagai bidang yang mereka terokai. -
Rocky Intertidal, Mudflats and Beaches, and Eelgrass Beds
Appendix 5.4, Page 1 Appendix 5.4 Marine and Coastline Habitats Featured Species-associated Intertidal Habitats: Rocky Intertidal, Mudflats and Beaches, and Eelgrass Beds A swath of intertidal habitat occurs wherever the ocean meets the shore. At 44,000 miles, Alaska’s shoreline is more than double the shoreline for the entire Lower 48 states (ACMP 2005). This extensive shoreline creates an impressive abundance and diversity of habitats. Five physical factors predominantly control the distribution and abundance of biota in the intertidal zone: wave energy, bottom type (substrate), tidal exposure, temperature, and most important, salinity (Dethier and Schoch 2000; Ricketts and Calvin 1968). The distribution of many commercially important fishes and crustaceans with particular salinity regimes has led to the description of “salinity zones,” which can be used as a basis for mapping these resources (Bulger et al. 1993; Christensen et al. 1997). A new methodology called SCALE (Shoreline Classification and Landscape Extrapolation) has the ability to separate the roles of sediment type, salinity, wave action, and other factors controlling estuarine community distribution and abundance. This section of Alaska’s CWCS focuses on 3 main types of intertidal habitat: rocky intertidal, mudflats and beaches, and eelgrass beds. Tidal marshes, which are also intertidal habitats, are discussed in the Wetlands section, Appendix 5.3, of the CWCS. Rocky intertidal habitats can be categorized into 3 main types: (1) exposed, rocky shores composed of steeply dipping, vertical bedrock that experience high-to- moderate wave energy; (2) exposed, wave-cut platforms consisting of wave-cut or low-lying bedrock that experience high-to-moderate wave energy; and (3) sheltered, rocky shores composed of vertical rock walls, bedrock outcrops, wide rock platforms, and boulder-strewn ledges and usually found along sheltered bays or along the inside of bays and coves. -
Intertidal Zone
INTERTIDAL ZONE The Intertidal Zone The Beach Starts in the Mountains • Glaciation, terranes, uplift, volcanism and erosion have brought rocks, sediment and sand down from the mountains through 10,000 streams to Puget Sound and created the beaches we see there. What Lives Where and Why? • Rock, sand, wave action, tidepools, exposure, temperature, light, dessication, food supply, and predation drive what can live in a particular part of the intertidal zone. • Example: Purple sea stars live under overhanging rocks and and climb up at high tide to eat barnacles and mussels on the rock. During summer storms or winter, they stay in low intertidal areas and do not feed. The level of barnacles and mussels on the rock reflects the predation of sea stars. Reproduction of sea stars is determined by the amount of light and food supply. Purple Sea Star Echinoderm Barnacles Arthropod Intertidal Zone above water at low tide & under water at high tide • May include many different types of habitats: rocks, cliffs, sand, wetlands, clay, shale, tidepools, stream outlets • Organisms in the intertidal zones are adapted to an environment of harsh extremes; sometimes submerged, exposed to sun, dessication, wave action, variable salinity, temperature extremes INTERTIDAL ZONE CARKEEK BEACH 11.0 ft -0.4 ft Intertidal Zonation on rocks at low tide Species ranges are compressed into very narrow bands Glacial erratic at Carkeek Park at low tide - encrusted with mussels, barnacles, sea stars, chitons, seaweed Intertidal Zone Organisms Life Zones of the Beach • Splash