476. Naticid Gastropods from the Miyazaki Group

Home , Euspira pallida, Glossaulax, Lunatia, Mammilla (gastropod), Mammilla melanostoma, Natica

Trans. Proc. Palacont. Soc. Japan, N.S., No. 55. pp. 281-293, pls. 42, .13, Sept. 20, 1964

476. NATICID GASTROPODS FROM THE MIYAZAKI GROUP

(PALAEONTOLOGICAL STUDY OF TIIE MIYAZAKI GROUP-X)

TSUGIO SHUTO

Department of Geology, Faculty of Science, Kyushu University

宮崎層群座タマガイ類:宮崎層群産化石軟体動物の研究の一部として腹足綱 Naticidae 科の4属9種.うち2踵は新種,の記載をする。また従来の慣用に従う一,二の既知種の分類について,問題があるのでそれについて私見を述べる。首藤次男

Here are the descriptions of nine spe- Superfamily Naticacea cies of Naticidae from several horizons of the Miyazaki group. Of nine species Family Naticidae six are known in our country. two are Subfamily Polinicinae new to science, and one is possibly new species. Genus Polinices MONTFORT, 1810 I am greatly indebted to Professor Tatsuro MATSUMOTO of Kyushu Uni- (type-species : Polinices albus MONTFORT versity for his kind suggestions and =Nerita mantilla LINNE by advices during the field survey and original designation) laboratory work. I wish to express my cordial thanks to Professor Ryuzo To- Subgenus Neverita Risso. 1926 RIYAMA of the same university for his (type-species : Neverita josephina valuable advices and thorough criticisms on the subject at the type script. Risso by monotypy) I also appreciate Drs. Tadashige HABE Polinices (Neverita) sagamiensis PILSBRY of National Science Museum of Tokyo and Katura OYAMA of the Geological Pl. 42, Figs. 2, 8 and 14. Survey of Japan for their valuable ad- 1904. Polinices sagamiensis PILSBRY, Proc. vices and discussions at the field of the Acad. Nat. Sci. Phitad.. Vol. 56. p. 23, molluscan taxonomy. This study was pl. 4, fs. 37, 37a. partly financed by the aid of the science 1922. Polinices pouisianus, YOKOYAMA.Jour. research expenditure fund by the Minis- Coll. Sci., Imp. Univ. Tokyo, Vol. .11, try of Education. Art. 1, p. 83, pl. 14, f. 12 (not of 12i:c t. u z). 1935. Polinices sagamiensis. OTUKA, Butt. Systematic Description Earthq. Res. Inst., Vol. 13, p. 866, pl. 53. I. 26. Received June 20, 1961; read Jan. 19, 1961 1951. Polinices sagamiensis, (ill RA SE .) TAM, at Fukuoka. Handb. Illust. Shells, pl. 91, 1. 8.

281 282 Tsugio SHUTO

1954. Polinices (Mammillaria) sagamiensis. 1961. Polinices sagamiensis. HAYASAKA,Sci. TANI and OVAMA, Palaeont. Soc. Rep., Tohoku Univ.. 2nd Ser., Vol. 33, Japan, Spec. Pap., No. 2. p. 17, pl. 2.1. no. 1. p. 75, p1. 9, fs. 12a, 12b. f. 12. 1954. Polinices sagamiensis, KIRA. Colored Material: -GK-L 7987 (loc. Nihonma- Illust. Shells Japan, I. p. 35, pl. 17. f. tsu). 7988-7990 (loc. Toriyama). All the 15. specimens were collected by myself.

Measurements:—-

Remarks_: -The specimens from Tori- Neverita josephina Risso, the type-spe- yama are quite identical to the living cies of Neverita, has depressed shell species, but that from Nihonmatsu with crescent umbilical furrow. The slightly differs from others in being difference in the outline of the shells provided with an obsoletely and shal- between these two type-species is very lowly grooved band close to the upper distinct in reality. but they are reason- suture. This band starts at the one- ably congeneric on the basis of the third of the third whorl. It is minutely several intermediate species. and spirally striated and bordered below Horizon : -The fifth and sixth fossil by rather a distinct step-like ridge. The horizon of the Miyazaki group (Lower early two and one-third whorls are Pliocene). separated from the succeeding whorl by Localities :-1) Road side small cut- oblique and weak ridge which is more ting at Nihonmatsu, Takanabe machi. distinct than the normal growth-lines. Koyu gun ; 2) Railroad cutting at the This character is not observed on ordi- entrance of a gorge near Toriyama, nary specimens of P. sagamiensis, both Kawaminami machi. Koyu gun. Miya- living and fossil. and suggestsa possible zaki Prefecture. specific discrepancy between this and others. I am, however. inclined to treat Subgenus Glossaulax PILSBRI, 1929 this specimen as an abnormal variety of P. sagamiensis, because subsutural (type-species : Natica reclusiana grooved band is examined only at a DESHAYES by original single specimen. designation) The present species, although it has been described by many authors as Polinices (Glossaulax) hyugensis n. sp. Polinices MONTFORT, 1810. is quite Pt. 42, Figs. 3, 5, 13 and 15; similar to the type species of Neverita PI. 43, Figs. 9. 10 and 12, Risso, 1826. in general characters. espe- Text-figs. 1 and 2. cially in the outline and umbilical feature. Polinices mantilla (LINNE), the Material:—Holotype : GK-L 8009 (loc. type species of Polinices, has tall shell Hagenoshita. coll. T. SHUTO) ; paratypes : with almost filled umbilicus. while GK-L 8001. 8002. 8008 (loc. Hagenoshita, 476. Naticid Gastropods from the Miyazaki Group 283 coll. T. MATSUMOTO et al.). 8010 (loc . Nakabyu. coll. T. SHUTO), and 8035 to Hagenoshita. coll. T. SHUTO), 8005. 8006 8040 (loc . Yamaji. coll. T. SHUTO). Many (loc. Nihonmatsu, coll. T. SIIUTO). 8011 other specimens from Toriyama are im- (loc. Kugino. coll. T. SHUTO), 8012 (loc. perfectly preserved and not registered.

Measurements:—

* Number of the protoconch whorls

is oblique, roundly convexed. and deeply umbilicate. The umbilicus is narrowed by a distinct umbilical cord and partly covered by the strong parietal callus. The parietal callus extends backward at its lower part along the edge of the umbilical furrow and is fused with the Fig. 1. Umbilical region of (1) Poli- umbilical cord but bordered by a shal- nices (Glossanlax) hagenoshitensis n. sp. low groove between them. The umbili- (GK-L 8004) and (2) P. (G.) hyugensis cal furrow is separated from the basal n. sp. surface by rather a sharp ridge. which The umbilicus is bordered by rather terminates at the anterior extremity of the sharp edge and devoid of the inner the aperture. There is no spiral inner spiral furrow in both species. While they furrow on the wall of the umbilical are quite distinguishable each other in furrow, although many spiral striae are the feature of the callus. observed. The aperture is semicircular with narrowly rounded anterior end. Diagnosis: -The shell is moderate in The inner lip is almost straight and size. obliquely hemispherical. smooth, slightly oblique. The outer lip is sim- and rather solid. The spire is very ple, sharp. slightly sinuate. and ante- low. small. and depressed-conical. The currently oblique. The whorls are about body whorl is large. hemispherical. very five of which early two and a half volu- gently sloped below the suture showing tions are perfectly smooth and shining an extraconical profile with the pleural protoconch (text-fig. 2). The protoconch angle larger than the apical one. The consists of more convexed volutions suture is slightly depressed. The surface separated by deeper suture than in the of the shell is ornamented by the close conch whorls and bordered by more or lines of growth which are distinct below less distinct groove between the first the suture and at the base. The base conch-whorl. 284 Tsugio SHUTO

the former shows some critical differ- ences from the latter. That is to say, the former is (1) much smaller than the latter ; (2) the present species has not inner spiral furrow inside the umbilical furrow. which is evidently observed on the adolescent and younger specimens of the latter. Horizon : -The first. second. (third). fourth. and fifth fossil horizons of the Fig, 2. Protoconch and carly tcleo- Miyazaki group (Middle Miocene to conch of Polinices (Glossaulax) hyugensis Lowest Pliocene). n. sp. Localities :-1) Small cutting along the national highway No. 10 at Hagenoshita. Comparison : -The present species is Uwaye mura ; 2) small cutting along the closely allied to P. (G.) vesicalis (PHILIPPI) national highway No. 10 at Nihonmatsu. in general characters. but is distinguish- Takanabe machi ; 3) road side small ed from the latter in having the distinct cliff north of Yamaji, Mino mura. Koyu ridge separating the basal surface and gun ; 4) river shore at Nakabyu, Yatsu- the umbilical furrow. white umbilical shiro mura ; 5) 300 m south of Kugino, callus. and more triangularly pointed Aya machi ; Higashi-morogata gun, Mi- lowest part of the parietal callus along yazaki Prefecture. the ridge of the furrow. Furthermore the anteriorly elongated columella of Polinices (Glossaulax) the present species gives a wider ap- hagenoshitensis n. sp. pearance to the aperture than in P. vesicalis. P1. 42, Fig. 10. Text-fig. 1. The present species also closely re- sembles P. (C.) didyma (R(DING). flouri- Material:—Holotype : GK-L 8003, para- shing in Japanese waters. in general, but types : GK-L 7993. 8004. and 8007.

Measurements:—

Diagnosis: -The shell is moderate in tinct. The surface of the shell is orna- size. oblique, hemispherical. round, mented with dense growth-lines. which smooth, and solid. The spire is small. are distinct below the suture and at the very low, and depressed-conical. The base and subobsolete at the periphery. body whorl is large. occupying more The protoconch is much smoother and than 90 percent of the total height of more roundly convex between sutures the shell, and very gently sloped just than the conch and consists of more below the suture with slightly concave than two and a half volutions. of which part. The suture is impressed and dis- the first is a minute dome and the 47 6. Naticid Gastropods from the Miyazaki Group 285 others are minute and round tube. The Locality: -Cutting along the national end of the protoconch is marked by a highway No. 10 at Hagenoshita. Uwaye recurrently oblique line. The umbilicus mura. Koyu gun. Miyazaki Prefecture. is separated by moderately sharp edge from the basal surface of the body whorl and narrowed by a large umbili- Polinices (Glossaulax) aff. cal cord. which is provided with the reiniana (DUNKER) larger callus at the outer end than cord P1. 42, Fig. 1. Text-fig. 3. itself. The parietal callus is thick and moderately wide and somewhat extends 1877. Neverita reiniana, DUNKER, Malak. backward at its lower part along the Blatt., Vol., 24, p. 71. edge of the umbilical furrow. It is. 1882. Nererila reiniana. DUNKER, Index however. not pointed as is seen in the Moll. Mar. Japan, p. 62. p1. 4, fs. 15, 16. preceeding species beyond the limit of 1886. Natica reiniana, TRYON, Man. Conch., the umbilical callus, but slightly nar- Vol. 8. p. 35, pl. 13, f. 12. rower than the umbilical callus. There 1920. Polinices (Neverita) ample, YOKOYA- is a shallow groove on the surface be- MA, Jour. Coll. Sci., Imp. Unit. Tokyo, tween the parietal and the umbilical Vol. .39, Art. 6. p. 77, pl. 5, f. 6. callus almost perpendicular to the colu- 1954. Polinices (Glossaulax) reiniana var., mellar lip. TAKI and OYAMA, Palaeont. Soc. Comparison : -The present new species Japan Spec. Pap., ,No. 2. p. 17, pl. 6, is quite similar in size and outline of f. 6. the shell to the preceeding species. but 1958. Neverita reiniana. OZAKI. Bull. Nat. differs in the features of the protoconch Sci. Mus. (Tokyo). Vol. 4, No. 1, p. and the umbilicus. I examined the 143. 2 text-fs. specimens in hand to check whether 1961. Neverita (Glossaulax) reiniana, HABE, Colored Illust. Shells Japan, II, p. 38. the characteristics in question are mere pl. 17, f. 11. fluctuating variation or distinct features. 1961. Nererita (Glossaulax) reiniana, HAYA- The result shows that these two species SAKA, Sci. Rep. Tohoku Unir., 2nd Ser. are sharply separated and there is no Vol. 33, No. 1, p. 76, pl. 9, fs. 18a, intermediate form concerning with the 18b. mentioned characteristics. Horizon: -The fourth fossil horizon of Material: -CK-L 7991 and 7992 (coll. the Miyazaki group (Lowest Pliocene). T. SHUTO)•

Measurements:—

Remarks : -The present specimens are the species of Glossaulax it closely„rre- provided with a groove separating the sembles P. (G.) reiniana (DUNKER)with parietal and the umbilical callus and rather tall spire, roundly convexed readily identified to Glossaulax. Among horls. and solid test. It. W however, 286 Tsugio SHUTO slightly differs from the living speci- Uwaye mura, Koyu gun, Miyazaki Pre- mens of the species in having larger fecture. umbilical cord and callus and in being devoid of the spiral groove inside the umbilical furrow (text-fig. 3). Genus Mantmilla SCHUMACHER. 1817 (type-species : Mammilla fasciata SCHUMACHER=Narita melanostoma GMELIN by monotypy)

ammilla melanostomaM (GMELIN)

Pl. 42, Figs. 4 and 6.

1791. Nerita melanostoma, GMELIN,Systemu Fig. 3. Umbilical region of (1) Poli- naturae, ed. 13, p. 3671. nices (Glossaulax) reiniana (DUNKER) 1852. Nalica metanostoma, PHILIPPI (in (recent) and (2) P. (C.) aff. reiniana MARTINl-CHEMNITZ)Conch. Cab., Vol. (DUNKER) (GK-L 8992). 2, Notice, p. 30, pl. 4. fs. 5, 6, 15. 16. Note the similar callus and different 1956. Natica metanostoma, REEVE, Conch. umbilicus. Recent specimens has the dis- Icon.. Vol. 9, Natica, pl. 18. sp. 78. tinct inner furrow within the umbilicus. 1886. Nalica melanostoma, TRYON, Man. Conch.. Vol. 8. p. 50, pl. 21, fs. 13, 14; The characteristics of the umbilical p1. 22, f. 21. region of these specimens closely re- 1925. Nalica (Mammilla) melanostoma, COSSMANN, Essais de Paleoconch. semble those of P. hyugensis n. sp.. Comp., Vol. 13, p. 129, pl. 1, fs. 36, although their spires are generally more 37: pl. 2, fs. 3, 4. elevated and the tests are much heavier 1934. Polinices (Polinices) melanostoma, than the latter. There are three possi- LADD. Bernice P. Bishop Mus., Bull. bilities concerning with the taxonomic No. 119, p. 211. position of these specimens. They may 1935. Polinices (Polinices) melanostoma, represent. a varietal group in P. reiniana; NOMURA,Sci. Rep., Tohoku Imp. Univ.. they may be a stout form in P. hyu- 2nd Ser., Vol. 18, No. 2, p. 203. p1. 9, gensis ; or they may be a off-shoot of P. f. 32. hyugensis and the ancestor of P. reiniana. 1941. Polinices (Mammilia) melanostoma, Considering that the elevation of the ALTENA, Leidse Geol. Meded., Vol. 12, spire is rather variable and the speci- p. 67. 1951. Polynices opacus, (HIRASE) TAKI, men GK-L 7992 is hardly distinguished Handb. Illust. Shells, pl. 91, f. 1. from P. hyugensis in its outline though 1954. Polinices opacus, KIRA, Colored Illust. the test is much more solid. I perfer to Shells Japan, I. p. 35, pl. 17, f. 13. the second possibility. 1960. Mammilla melanostoma, MACNEH, Horizon :-The fourth fossil horizon U.S. Geol. Surr., Prof. Pap., No. 339. of the Miyazaki group (Lowest Pliocene). p. 54, p1. 8, f. 1: p1. 12, f. 19. Locality: -Small cutting along the national highway No. 10 at Hagenoshita. Material:-GK-L 8033 (coll. T. SHUTOI. 476. Naticid Gostropods from the .Miyazaki Group 287

Measurements:—

Remarks: - The present Mammilla zaki Prefecture. specimen is featured by depressedly pear-shaped tall outline, slit-like very Marnmila maura (BRUGUIERE) narrow umbilicus, and almost smooth surface except for minute growth lines P1. 42, Figs. 7 and 9. and safely identified to M. melanostoma 1792. Natica maura, BRUGCIERE,Encyclo- (GMELIN). Its shell is slightly taller pedie methodique pl. 453. fs. 4a, b. and its umbilicus is much narrower 1852. Natica maura. PHILIPPI (in MARTINI- than those of M. maura BRUGUIERE, CIIErINITZ). Conch. Cab., Vol. 2, Natica, which is a close ally to it. M. kurodai p. 58, pl. 9, f. 6. TAKI (=Natica Macrosloma PHILPPI in 1856. Natica maura, REEVE, Conch. Icon., MARTINI and CHEMNITZ, 1852. p. 55, pl. Vol. 9, Natica, pl. 7, f 25. 9. f. 1. TAKI. 1943. p. 111. text-fs. 1. 2) 1886. Natica maura, TRYON, Man. Conch., is closely allied to the present species. Vol. 8, p. 51, pl. 22, f. 24. 1951. Polinices maurus, (HIRASE) TAKI, but the former has wider umbilicus. Handb. Illust. Shells. pl. 91, f. 3. stronger growth-lines, and somewhat 1954. Polinices (Mammilla) maurus, KIRA, wider shell than the latter. Colored Must. Shells Japan, I, p. 35, Comparing with M. simiae (DESHAYES) pl. 17, f. 12. {1842, p. 652), living in south Japan. the present specimen has much heavier Material: -GK-L 8034 (loc. Yamaji, callus. MI-5070, coll. T. SHUTo) and 8045 (loc. Horizon : -The secondfossil horizon of Hagenoshita, coil. T. SIIUTO). The the Miyazaki group (Middle Miocene). specimen GK-L 8034 is slightly broken Locality: -Road side cutting north of at the labrum and 8015 lost the apical Yamaji, Mino mura. Koyu gun. Miya- part of the protoconch.

Measurements:—

Remarks: -The present specimens are Horizon: -The second and fourth fossil featured by the broad shell. deep and horizon of the Miyazaki group (Middle distinct umbilicus. moderately distinct Miocene to Lowest Pliocene). growth-lines, and dark brown uniform col- Localities : -Small cutting along the oration. On the basis of these charac- national highway No. 10 at Hageno- teristics they are identified to M. maura shita. Uwave mura ; road side 600 m (BRUGUIERE), although the former is north of Yamaji, Mino mura, Koyu-gun, much smaller than the living specimens. Miyazaki Prefecture. 288 Tsugio SHUTO

1921. ? Euspira pallida, DALL, U.S. Nat. Mus., Bull., Vol. 112, p. 164, pl. 14. Genus Lunatia GRAY, 1847 f. 5. 1945. ? Lunalia pallida, ABBOTT, American (type-species : Natica ampularia Sea shells., p. 190. f. 43c. SOWERBY by original designation) 1960. Euspira cf. pallida, MACNEIL. U.S. Geol. Sum, Prof. Pap., No. 339, p. 57. Lunatic pallida BRODERIP and SOWERBY p1. 2. fs. 20, 26. Pl. 43, Figs. 4, 6. 7, 8. 11 and 13. Material: -GK-L 8018 to 8023, 8025, 1829. ? Natica pallida. BRODERIP and Sow- 8026, 8029 (loc. Nihonmatsu, coll. T. ERBY, Zool. Jour., Vol. 4. p. 372 (in SHUTO). 8013 to 8017, 8027. 8028 (loc. DALL, 1921). agenoshita, coll. T. MATSUMOTO et H al.). 1886. ? Notice pallida, TRYON, Man. Conch., and 8032 (loc. Hagenoshita, coll. T. Vol. 8, p. 37. pl. 14, fs. 26 to 28. SHUTO).

Measurements:—

Remarks: -The present thick-shelled Euspira AGASSIZ, 1838, and Lunatia specimens are featured at first by mod- GRAY. 1847. I, however. consider they erately shouldered outline with the are reasonably included in Lunatic, be- maximum convexity at higher part of cause they have solid shell, being devoid the whorls. In other words the whorls of the shallow groove between the are narrowly rounded just below the parietal and umbilical callus which are horizontal or gently sloped subsutural the important characteristics of Euspira. part (shoulder). Secondly the umbilicus New Zealand genus. Uberella FINLAY, is moderately narrow but very deep. 1928, with the type-species, Natica and not separated from the basal sur- rib-ea HUTTON, is also closely allied to face by a sharp angulation. The parietal these genera, especially to Lunatia and callus is thick, solid, wide. and partly it seems there is no critical distinction truncated at the margin of the umbili- between them as far as the descriptions cus, where it merges partly to the nar- and figures are concerned. rower columellar callus. The eversion " Uberella yokoyamai (KURODA and is also thickly deposited with the callous ABE) " (TAKI and OYAMA, 1954. p. H 17, material. There is no funicle in the pl. 5. f. 1), which is originally reported umbilicus. Thirdly the suture is im- as Gennaeosinum (?) (KURODA and HABE. pressed. On the basis of these charac- 1952, p. 59), is distinguished from the teristics they are identified to " ?Natica present specimens in having the maxi- pallida BRODERIP and SOWERBY ". mum convexity at lower position than They are very closely similar to the latter. Although this difference 476. Naticid Gastropods from. the Miyazaki Group 289 looks very slight, it is markedly per- sistent to serve as the criterion for PI. 42. Fig. 11; PI. 43. Figs. 1. 2. 3 and 5. interspecific separation. 1920. Natica janthostoma, YOKOYANIA,Jour. The present specimens are very Coll. Sci., Imp. Univ., Tokyo. Vol. 39, closely allied to " Natica gendinganensis Art. 6, p. 76, pl. 5, fs. 3 and 4 (not MARTIN (1899, p. 262, p1. 29, fs. 627, of DESHAYES). 627a) in general feature. The only dif- 1934. Natica (Tectonatica) janthostoma. KI- ference is that the latter has more pro- NoSHITA and ISAnHAYA,Rep. Fish. duced anterior end of the aperture and Surv., Hokkaido Fish. Experim. Stat., eversion and somewhat smaller ratio of No. 33, p. 7. pl. 4, f. 27. the diameter of the whorl to the dia- 1935. Natica (Tectonatica) janthostoma, No- meter of the whorl-tube of the same RA, Saito Ho-on Kai Mus. Res. MT Bull. No. 5, p. 122, pl. 7, fs. 4 to 6. whorl. I prefer to deal with it as a 1935. Natica (Tectonatica) janthostoma, No- subspecies of L. pallida. MURAand HATAL. ibid., No. 6. p. 128, Horizon : -The fourth and fifth fossil p1. 9, f. 4. horizon of the Miyazaki group (Lower 1949. Tectonatica janthostomoides, KURODA Pliocene). and IIAHE, Venus. Vol. 15, Nos. 5-8, Localities:-1) Small cutting along the p. 7, text-fs. la to d. national highway No. 10 at Hageno- 1954. Natica (Tectonatica) serera. KIRA, shita, Uwaye mura ; 2) cutting along Colored Must. Shells Japan, I, p. 35, the national highway No. 10 at Nihon- pl. 17. f. 18. 1958. Natica janthostomoides, OZAKI. Bull. matsu. Takanabe machi, Koyu gun, Nat. Sci. Mus., (Tokyo). Vol. 4. No. 1.. Miyazaki Prefecture. p. 144, p1. 19. fs. 1 and 2. 1961. Tectonatica janthostomoides, HAYA- Genus Tectonatica SACCO. 1890 AkA, Sci. Rep. Tohoku Univ., 2nd Ser.,S Vol. 33. No. 1, p. 77, pl. 9, Is. 20a and b. (type-species: Natica tectula BONELLI by monotypy) Material: -GK-L 7994 to 7996. coll. T. MATSUMOTO et al.. 7997 to 8000 and Tectonatica janthostomoides 8031, coll. T. SHUTTO. All the specimens KURODA and IIABE came from a single locality.

Measurements:—

The largest specimen. GK-L 7994, more closed umbilicus than the living attains about 20 mm in maximum dia- ones which are provided with narrow meter. but apparently open umbilical furrow Remarks: -The present specimens around the umbilical callus and cord. generally have much smaller shell with Of course the degree of closure at the 290 Tsugio SHUTO

umbilicus is not invariably fixed but the whorls=5.0. somewhat variable and some specimens. Description: -The shell is moderately as GK-L 7995, show typical umbilicus small, moderately thick, and globose. of T. janthostomoides. The present The spire is moderately elevated. The specimens may represent a group which whorls are about five in number, and is biased but not so different from the round. The body whorl is large. round- typical one as to be separated specifi- ly convex at the side slightly above the cally. middle, round at the base, and not um- Horizon: -The fourth fossil horizon of bilicate. The aperture is semicircular. the Miyazaki group (Lowest Pliocene). The outer lip is not observed because Locality: -Small cutting along the of fracturing. The labrum profile is national highway No. 10 at Hageno moderately oblique judging from the shita, Uwaye mura, Koyu gun, Miya- growth lines. The inner lip is moder- zaki Prefecture. ately curved and covered by the distinct callus. The parietal callus is wide, thick, and perfectly fused with the um- Tectonatica sp. bilical one which entirely closes the Text-fig. 4. umbilicus. The callus of the lower part of the columellar lip is also thick and Material: -GK-L 8030. coll. T. SHUTO. continued to the produced eversion. Measurement: -H=13.6 mm. number of Remarks and comparison : -The glo-

Explanation of Plate 42

Fig. 1. Polinices (Glossaulax) aff. reiniana (DUNKER)

GK-L 7991 (•~1.6), loc. Hagenoshita. Figs. 2. 8. 14. P. (Neeerite) sagamiensis PILSBRY 2...GK-L 7987 (•~2.2), loc. Nihonmatsu,

8...GK-L 7988 (•~1.3), loc. Toriyama,

14...GK-L 7989 (•~0.9), loc. Toriyama. Figs. 3, 5, 13 and 15. P. (Glossaulax) hyugensis n. sp.

3...paratype: GK-L 8008 (•~2.3), Inc. Hagenoshita,

5...holotype: GK-L 8009 (•~2), loc. Hagenoshita, 13...paratype: GK-L 8012 (•~2.3), loc. Nakabyu,

15...(•~2.3), same specimen as fig. 13. Figs. 4 and 6. Mannnilla melanostoma (GMELIN)

4...GK-L 8003 (•~3), loc. Yamaji, 6...(•~3), same specimen as fig. 4.

Figs. 7 and 9. M. maura (BRUGUIERE)

7...GK-L 8045 (•~3). loc. Hagenoshita, 9...(•~3), same specimen as fig. 7.

Fig. 10. Polinices (Glossaulax) hagenoshitensis n. sp.

paratype GK-L 8004 (•~2.5), loc. Hagenoshita. Fig. 11. Tectonatica janihostomoides KURODA and HARE

GK-L 7998 (•~3). loc. Hagenoshita.

Fig. 12. Lunatia pallida (BROUERIP and SoWERBY)

GK-L 8018 (•~3). loc. Nihonmatsu. SHUTO : Naticids from the Miyazaki Group Plate 4

1 2 3

5 6 4

8 9 7

12 10 11

13 15 476. Naticid Gastropods from. the Miyazaki Group 291

bose shell with moderately elevated feature of the former. Glossaulax spire and utterly closed umbilicus indi- should be separated from Neverita. cates that the specimen belongs to atica pallida BRODERIP andN SOWERBY Tectonatica. Although whole species of is considered to be included in Lunatia Tectonatica known in our country have GRAY, 1847. on the basis of the shell the umbilical callus fused with the character. The species seems to be a parietal callus. their umbilical callus is characteristic to the Lower Pliocene of apparently separated from the parietal Indo-southwest Pacific region. That is one by the remarkably contracted part to say, besides in Japan it also occurs between them. While in the present in the Lower Pliocene of Okinawa Island specimen, they are perfectly fused to- and its close ally, L. gendinganensis gether and not separated each other. (MARTIN) (1905, p. 262) in the Lower This peculiarity together with the oc- Pliocene Bantam bed of Java. currence of the specimen from the

higher horizon than the preceeding Works Cited species supports the idea that the specimen may not be included in the ABBOTT, R. T. (195-1) : American Seashells. range of variation of T. janthostomoides pp. 1-541, pls. 1-40, Nostrand Co., New and may be a new species. The esta- York. ALTENA, C. O. van Regteren (1941): The blishment of the new species is, however, marine mollusca of the Kendeng beds suspended until) plenty specimens are (East Java) : Gastropoda, pt. 2. Leidse examined. Geol. Meded., Vol. 12, pp. 1-86. Horizon: -The fourth fossil horizon of BRUGUIERE,J. G. (1792) : Encyclopedie me- the Miyazaki group (Lowest Pliocene). thodique, Vol. 1, pt. 2, pp. 345-758. Locality: -Cutting along the national COSSMANN,Maurice (1925): Essais de paleo- highway No. 10 at Nihonmatsu. Taka- conchologie comparee. Vol. 13, pp. 1-345, nabe machi, Koyu gun. Miyazaki Pre- pls. 1-11. fecture. DAII., Wilhelm II. (1921) : Summary of the marine shell-bearing mollusks of the northwest coast of America, from San Concluding Remarks Diego. California, to the Polar Sea most- ly contained in the collection of the Some revisions of the systematic posi- United States National Museum, with tion of certain common Japanese naticid illustrations of hitherto unfigured spe- gastropods are given in this report. cies. U.S. Nat. Mils. Bull., No. 122. pp. Polinices sagamiensis PILSBRY was 1-217, pls. 1-22. usually classified into Polinices s. s. by DESHAYES,G. P. (1842) : Histoire naturelle Japanese authors, but the former is des animaux sans vertebres. Vol. 8. readily distinguished from the latter in DUNKER. Wilhelm (1877) : Mollusca non- nulla nova maris japonici. Malak. Blatt., depressed shell and is better to be Bd. 24. pp. 67-75. included in Neverita Risso, 1926. (1882): Index- molluscorum maris japo- Glossaulax is usually included in nici. pp. 1-301, pls. 1-16, Kassel. .Neverita also by Japanese authors, but GMELIN,J. F. (1788-1793) : Systema naturae, the latter is devoid of the shallow ed. 13. groove separating the parietal and the GRAY, J. F. (1847) : List of the genera of umbilical callus which is the typical recent mollusca, their synonyma and 292 Tsugio SNUTO

types. Proc. Zool. Soc. London, pp. 129- MACNEIL, F. Stearns (1960) : Tertiary and 206. Quaternary gastropoda of Okinawa. U. S. IBABE, Tadashige (1961) : Colored illustra- Geol. Surv., Prof. Pap., No. 339, pp. 1-148, tions of shells of Japan. II, pp. 1-183, pls. 1-19. pls. 1-66 (in Japanese), Hoikusha, Osaka. MARTIN. K. (1899) : Die Fossilien von Java: AYASANA,Shozo (1961) : The geologyH and Samml. Geol. Reichs Mus. Leiden, neue paleontology of the Atsumi peninsula. Folge, Bd. 1, Mt. 1, Heft 5-10, pp. 133- Aichi Prefecture, Japan. Sci. Rep., To- 332, pls. 21-45. hoku Univ., 2nd Ser., Vol. 33, No. 1, pp. NOMURA,Sitihei (1935a) : On some Tertiary 1-103, pls. 1-12. mollusca from northeast Honshu. Japan. KINOSHIITA,Toraichiro and ISAUAYA,Takao pt. 2, Fossil mollusca from the vicinity (1934) : Catalogue of the mollusks of of Ogino, Yama-gun, Hukusima Prefec- okkaido. Japan. Rep. Fish. Surv. Hok-H ture. Saito Ho-on Kai Mus. Res. Bull., kaido Fish. Experim. Stat., No. 33, pp. No. 5, pp. 101-130, pls. 5-7. 1-19, pls. 1-15 (in Japanese). (1935b) : Catalogue - of the Tertiary RA, Tetsuaki (1954) :KI Colored illustrations and Quaternary mollusca from the island of shells of Japan. I. pp. 1-172. pls. 1-67, of Taiwan (Formosa) in the Institute (in Japanese). Hoikusha, Osaka. of Geology and Paleontology, Tohoku KURODDA.Tokubei and HABE, Tadashige Imperial University, Sendai, Japan, pt. (1949): On Natica janthostoma and an 2, Scaphopoda and Gastropoda. Sci. Rep. allied species which was confused with Tohoku Imp. Univ.. 2nd Ser., Vol. 18, No. it. Venus (Japan. Jour. Matac.), Vol. 15. 2. pp. 53-228, pls. 7-10. No. 58, pp. 67-72 (in Japanese). and HATAI, Kotora- (1935) : Pliocene and - (1952) : Check- list and biblio- mollusca from the Daisyaka shell-beds graphy of the recent marine mollusca in the vicinity of Daisyaka, Aomori-Ken of Japan, pp. 1-210, Tokyo. Northeast Honsyu, Japan. Saito Ho-on LADD, H. S. (1934) : Geology of Vitilevu, Kai Allis. Res. Bull., No. 6. pp. 8-142, Fuiji. Bernice P. Bishop illus. Bull.. No. pls. 9-13. 119. OLDROYD, 1. S. (1927) : The marine shells

Explanation of Plate 43

Figs. 1, 2, 3 and 5. Tectonatica janthostomoides KURODA and HABE 1...GK-L 8031 (•~3); loc. Hagenoshita,

2...GK-L 7997 (•~2.5). loc. Hagenoshita,

3...(•~3), same specimen as fig. 1, 5...GK-L 7998 (•~3). loc. Hagenoshita.

Figs. 4, 6, 7, 8, 11 and 13. Lunatia pallida (BROBERIP and SoWERBY) 4...GK-L 8027 (•~3), loc. Hagenoshita,

6...GK-L 8026 (•~3). loc. Nihonmatsu,

7...(•~3). same specimen as fig. 6, 8...GK-K 8012 (•~3), loc. Nihonmatsu, 11...GK-L 8028 (•~3), loc. Hagenoshita,

13...(•~3), same specimen as Fig. 11.

Figs. 9, 10 and 12. Polinices (Glossaulax) hyugensis n. sp. 9...paratype: GK-L 8005 (•~2.5), loc. Nihonmatsu,

10...(•~2.5), same specimen as Fig. 9. 12...paratype: GK-L 8002 (•~2). loc: Hagenoshita. SHUTO: Naticids from the Miyazaki Group Plate 43

1 2 3

4 6

9 7

11 13 476. Naticid Gastropods from the Miyazaki Group 29

of the west coast of north America ; 9, Natica, pls. 1-30. Scaphopoda and Gastropoda. Stanford ROI ING, P. F. (1798) : Museum Boltenianum Univ. Press., Vol. 2, pt. 1. pp. 1-297, pls. sive catalogus cimelionum e tribus 1-29. pt. 2, pp. 1-304, pls. 30-72, pt. 3, regnis naturae. Hamburg. pp. 1-339, pls. 73-108. SACCO, F. (1890) : Molluschi dei terreni OTUKA, Yanosuke (1935) : The Oti-graben terziari del Piedmonte e della Liguria. in southern Noto peninsula, Japan, pt. 3, ScHUMACHER,C. J. (1817) : Essai d'un nou- Bull. Earthq. Res. Inst.. Vol. 13, No. 1, veau systeme de habitations des vers pp. 846-909, pls. 53-57. testaces. pp. 1.287, pls. 1-22. OZAKI, Hiroshi (1958) : Stratigraphical and TAM, Isao (HIRASE, Shintaro) (1951) : Hand- paleontological studies on the Neogene book of Illustrations of Shells. pp. 1-46, and Pleistocene formations of the Tyosi pls. 1-134. Bunkyokaku, Tokyo. district. Bull. Nat. Sri. .Mus. (Tokyo), and OYAMA,Katura (1954)- : The Plio- No. 42, pp. 1-182, pls. 1-24. cene and later faunas from the Kwanto PHILIPPI, R. A. (1849) : Centuria tertia test- region in Japan. Paleont. Soc. Japan, aceorum novorum. Zeitschr. f. Malak., Spec. Pap.. No. 2, pp. 1-68, pls. 1-49. f. 1848, Bd. 5, pp. 151-176, pp. 186-192 TAM, Iwao (1943): The mollusca of the Bd. 6. pp. 17-26, pp. 33-35. inland sea of Japan. pt. 2. Japan. Jour. (1852): Die Gattungen- Natica and alac., Vol. 13, Nos. 1-1, pp. 111-118.M Amaura in MARTINI and CHEMNITZ, text-figs. 1-14. Systematische Conchylien-Cabinet, Bd. 2. TYRON, G. W. (1886) : Manual of concho- PILSBRY,H. A. (1904): New Japanese marine logy, Vol. 8. mollusca ; Gastropoda. Proc. Acad. Nat. WENZ, W. (1943) : in O.H. SCHINDEWOLF, Sci. Philad., Vol. 56, pp. 3-37. pls. 1-6. Handbuch der Palaozoologie, Bd. 6, (1929) : ;Veverita reclusiana (DESHAYES)- Gastropoda (Prosobranchia), pp. 1201- and its allis. Nautilus. Vol. 15. pp. 109- 1596. 133, pl. 6. YOKOYAMA,Matajiro (1920) : Fossils from -and VANATTA,-and E.G. (1909) : Notes on the Miura peninsula and its immediate Polinices didyma . with description of a north. Jour. Coll. Sci., Imp. Univ. Tokyo. new Australian species. Proc. Acad. Vol. 39. Art. 6, pp. 1-193, pls. 1-20. Nat. Sci. Naiad., Vol. 60, pp. 555-559, (1922) : Fossils from the upper- Musa- pl. 29. shino of Kazusa and Shimosa. ibid., REEVE, L. A. (1856) : Conchologia Iconica, Vol. 4.1, Art. 1, pp. 1-200. pls. 1-17.

Ava machi 綾町 Nakabyu 中別府 Hagenoshita 兀の下 Nihonmatsu 二本松 Higashi-morogato gun 東諸県郡 Takanabe machi 商鍋町 Kawaminami machi 川南町 Toriyama 通山 Koyu gun 児湯郡 Uwavc mura 上江村 Kugino 久木野 Yamaji 山路 Mino mura 三納村 Yatsushiro mura 八代村