The Singing Athlete Chapter 2

Total Page:16

File Type:pdf, Size:1020Kb

The Singing Athlete Chapter 2 2. BRAIN BASICS Think about the best singing experience of your life: it may have taken place in a performance, a voice lesson, or just driving around town. Can you remember the euphoria you felt when, suddenly, everything worked the way it should? This state of effortless flow is what you are seeking every time you open your mouth to sing. And yet, no matter how vocally skilled you get, there are inevitably days when singing feels challenging. The issues you experience on a “bad voice day” may not be noticeable to those who hear you, but you’ll be aware that you’re not at the top of your game. If you know how well you can sing, why isn’t it perfect every time? The reason is simple: humans are survival-based organisms. Staying alive is more important than performing well. The majority of your brain doesn’t care if you can float a high C; it cares that you survive the attempt. On a subconscious level, your brain is constantly assessing your environment for threatening signals. Is that person walking toward you friendly or crazy? Do you hear the sound of fireworks or gunshots? Are you smelling a cozy bonfire or a forest ablaze? These assessments occur in an ancient, reflexive part of your nervous system that we will call the first brain (Fig. 2.1). This first brain, which includes the brainstem and the cerebellum, is always asking the question, “Is it safe?” Let’s say you’ve been struggling to belt a high note without strain. One day, all the stars align, and it happens. When it finally comes out of your mouth, you’re not even sure how you got there, but you think, “Yes! I’ve figured this out!” The next day, you get up and vocalize, excited to go for the same phrase again. It cracks. You sing it again. It cracks again. You’re crestfallen; you can’t understand how your voice can be so inconsistent. From a neural perspective, this inconsistency makes sense. The lion’s share of your brain isn’t interested in high-level performance. It’s invested in keeping you alive right this second, not tomorrow or twenty years from now. Depending on the health of your nervous system, your first brain may have even interpreted the novelty of your big, beautiful note as a threat. The next day, when you approached the same phrase, it may have impaired the coordination of your vocal folds to dissuade you from ever trying that freaky thing again. So, what can you do about this? How do elite performers get around the brain’s survival mechanism to consistently sing their best? The first step is to understand threat and how the nervous system responds to it. THE THREAT BUCKET Have you ever felt like you were going to die when performing a song? On the one hand, it’s ridiculous; clearly, no mortal danger is going to come from going flat on the last note of “Bring Him Home.” On the other hand, the feelings of dread that can creep up as you approach a problematic vocal passage can feel real and terrifying. From your nervous system’s perspective, there may not be much difference between falling down a flight of stairs and nearing the climax of “She Used to Be Mine.” Both situations can produce a threat response. Threat is anything your brain thinks might be dangerous. As you go through your day, your brain is looking for patterns and making predictions based on what it perceives. When new information enters your field of awareness, the brain compares this novel stimulus to previous experiences and to what is expected in this environment. If you see a deer in the woods, you may marvel at the beauty of nature. If you see one in your kitchen, some different thoughts will probably arise. Based on the clarity of its predictions, the brain can react to potential threats as vehemently as real ones (e.g., performance anxiety). One of the goals of brain-based training is to improve your nervous system's predictive capacity. To form accurate plans, your brain needs trustworthy information and proper fuel, and all of your body systems need to participate. Erroneous data from any area can increase threat and degrade performance. Some things that can make prediction difficult are: • Lack of sleep • Injuries that haven’t been rehabbed completely • Dehydration • Underdeveloped motor control/weak muscles • Poor dietary choices • Issues with vision/hearing/smell/taste • Stress • Lack of sensation/numbness in areas of the body You probably read at least a couple of those items and thought, “Yeah, that’s me.” Maybe you’ve worn glasses since junior high, or you’ve sprained your ankle three times, and it still bothers you. When these issues begin to compound with other stressors, some nasty consequences can show up. Let’s imagine that your brain has a container in it called the threat bucket (Fig. 2.2). The level of water in this metaphorical receptacle reflects the total amount of threat coming your way. Didn’t sleep well last night? Threat increases, and the bucket starts to fill. Got up too late to eat breakfast and grabbed a donut as you ran out the door? More water in the bucket. Your train stalled underground for a half-hour and made you late to work? Now the level is rising fast. As threats start to accumulate in the bucket, your first brain will begin to get very scared; it believes that if the water reaches the top, you could die. So, it will create a spigot, and liquid will drain out in the form of some kind of output. Some of the possible threat-bucket outputs are: • Pain • Sickness • Breathing problems (asthma/ allergies/reflux) • Fatigue • Anxiety/depression • Loss of flexibility (physical or vocal) • Poor balance/dizziness/motion sickness • Migraines • Hormonal problems Nothing on this list is very much fun. But from a neural perspective, it makes sense why your first brain might give you these outputs. If you’re in mortal danger, the best strategy to keep you alive may be to stop moving. Think of an antelope and a lion on the African plains; if the predator gives chase, the prey will attempt to flee. If the lion succeeds in capturing the antelope, the animal will go into a state of shutdown, basically playing dead. This ancient reflex is based on the hope that a sudden change in vital energy will be enough to create a brief distraction, allowing a final possibility of escape. If you have ever had migraines, chronic fatigue, or frequent illnesses, you can understand how hard it is to move in the throes of these maladies. Even though there is no lion with its jaws around your neck, your threatened first brain reacts like that of the captured antelope. It thinks that the pain, illness, or fatigue it’s giving you is necessary for your survival. Performers are especially prone to threat-bucket outputs that are outside the realm of body pain. Almost all professionals have had to go onstage injured at some point, ignoring the brain’s advice to stop moving. If the signals continue to be disregarded, the brain may think, “Wow, I gave you a ton of ankle pain, but you still tapped in 42nd Street every night for weeks. Since pain didn’t work, it’s time to try something else to keep you still.” It may hinder your immune system, give you depression, or mess with your hormones. Unresolved high threat eventually leads to a shutdown of some sort. To avoid these unpleasant outputs and sing your best, you have to lower the level in your threat bucket. This is why many other body systems beyond your two little vocal folds can have a substantial impact on your voice. Threat levels can be reduced through improvements in the following systems: • Movement maps (moving your joints through a full and controlled range of motion) • Sensory capability (e.g., vision, hearing, taste, smell, touch) • Balance • Respiration habits One of the goals of this book is to help you discover drills that lower threat in your singing and your life. NEUROPLASTICITY The term neuroplasticity refers to changes in brain structure or function that are either directly measured in individual neurons or inferred from measures taken across neuronal populations (Fig. 2.3). In essence, it means that the brain can change. You wouldn’t be reading this book if you didn’t believe in this concept: education is neuroplasticity in action. Until fairly recently, it was thought that neuroplasticity was exclusively the claim of youth. We now know that to be untrue, and that your brain can and will change until the day you die.1 Neuroplasticity isn’t good or bad; it just is. You can make positive plastic changes that improve performance and reduce pain, but you can just as easily have negative plasticity. Here’s an example: Maybe you found a song that you learned and fell in love with; everything felt smooth, and the words seemed like they were written for you. When you took this winning new tune to an audition, you forgot to give a proper tempo, and the accompanist played too slowly. The dragging pace led to a lack of breath coordination and a cracked high note, which made the director involuntarily wince. Ugh. The next thing you know, your new favorite song goes in the garbage; the negative plasticity of the performance experience would make singing it again too painful.
Recommended publications
  • Validation of the Cortical Homunculus Using Functional Mri
    VALIDATION OF THE CORTICAL HOMUNCULUS USING FUNCTIONAL MRI Rushil Anirudh Arizona State University School of Electrical, Computer & Energy Engineering ABSTRACT within the motor strip that correspond to the motor function that is being studied. The motor strip in the frontal lobe of the Human Brain is re- sponsible for all bodily movements or “motor functions”. In this paper we attempt to map the motor cortex of the Human Brain using fMRI data. The advantages of mapping it are sev- eral fold ranging from therapy to epilepsy and stroke patients, to understanding the development of these motor functions from birth. We try to recreate the homunculus, a concept which showed how the real estate of the motor cortex was distributed across these motor functions. Index Terms— fMRI, Primary Motor Cortex, Homuncu- lus. 1. INTRODUCTION The Human brain is the most important organ of the body, however we are still not entirely clear about how or why it works the way it does. While understanding the semantics behind our actions will require us to understand the entire human brain, we attempt to solve a much more low level problem of what happens when humans engage in day to day actions. To be more precise, this involves studying the Pri- mary Motor Cortex or the Motor Strip which is responsible Fig. 1. The idea of the Cortical homunculus as created by for all our motor functions, from our tongue to walking, run- Wilder Penfield. ning etc. Penfield and Boldrey [1] formulated the concept of the Cortical Homunculus which is a pictorial representation of the anatomical divisions of the motor strip.
    [Show full text]
  • Somatotopic Mapping of the DALL
    King’s Research Portal DOI: 10.1093/cercor/bhy050 Document Version Publisher's PDF, also known as Version of record Link to publication record in King's Research Portal Citation for published version (APA): Dall'Orso, S., Steinweg, J., Allievi, A. G., Edwards, A. D., Burdet, E., & Arichi, T. (2018). Somatotopic Mapping of the Developing Sensorimotor Cortex in the Preterm Human Brain. Cerebral Cortex, 28(7), 2507-2515. https://doi.org/10.1093/cercor/bhy050 Citing this paper Please note that where the full-text provided on King's Research Portal is the Author Accepted Manuscript or Post-Print version this may differ from the final Published version. If citing, it is advised that you check and use the publisher's definitive version for pagination, volume/issue, and date of publication details. And where the final published version is provided on the Research Portal, if citing you are again advised to check the publisher's website for any subsequent corrections. General rights Copyright and moral rights for the publications made accessible in the Research Portal are retained by the authors and/or other copyright owners and it is a condition of accessing publications that users recognize and abide by the legal requirements associated with these rights. •Users may download and print one copy of any publication from the Research Portal for the purpose of private study or research. •You may not further distribute the material or use it for any profit-making activity or commercial gain •You may freely distribute the URL identifying the publication in the Research Portal Take down policy If you believe that this document breaches copyright please contact [email protected] providing details, and we will remove access to the work immediately and investigate your claim.
    [Show full text]
  • Krubitzer * and Tony J
    Special Issue: Time in the Brain Review The Combinatorial Creature: Cortical Phenotypes within and across Lifetimes 1, 2 Leah A. Krubitzer * and Tony J. Prescott The neocortex is one of the most distinctive structures of the mammalian brain, Highlights yet also one of the most varied in terms of both size and organization. Multiple There are multiple time-scales that are relevant for understanding how a given processes have contributed to this variability, including evolutionary mecha- phenotype emerges. Brains change nisms (i.e., alterations in gene sequence) that alter the size, organization, and across large, evolutionary time-scales, shorter time-scales such as genera- connections of neocortex, and activity dependent mechanisms that can also tions, and within the life of an individual. modify these same features. Thus, changes to the neocortex can occur over different time-scales, including within a single generation. This combination of Any given phenotype is a combination of genes involved in brain and body devel- genetic and activity dependent mechanisms that create a given cortical opment, behavior,and the environmentin phenotype allows the mammalian neocortex to rapidly and flexibly adjust to which an individual develops. A similar different body and environmental contexts, and in humans permits culture to phenotype in different species may be due to homology, but can also be the impact brain construction. result of a different combination of factors. Brain, Body, and Environment Interactions There are several constraints that restrict In this review we discuss the evolution and development of mammalian neocortex. We examine the avenues along which evolution of the brain and body can proceed.
    [Show full text]
  • Of the Human Cortical Somatosensory System
    bioRxiv preprint doi: https://doi.org/10.1101/675611; this version posted June 21, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY-NC-ND 4.0 International license. The "creatures" of the human cortical somatosensory system N. Saadon-Grosman,1,2* Y. Loewenstein,3,4 † S. Arzy 1,2† 1Neuropsychiatry Lab, Department of Neurology, Hadassah Hebrew University Medical Center, Jerusalem, 9112001, Israel. 2 Department of Medical Neurobiology, Faculty of Medicine, Hadassah Hebrew University Medical School, 9112001, Jerusalem, Israel. 3 The Edmond and Lily Safra Center for Brain Sciences, The Alexander Silberman Institute of Life Sciences, 4 Dept. of Cognitive Sciences and The Federmann Center for the Study of Rationality, The Hebrew University, 9190401 , Jerusalem, Israel. † These authors contributed equally to this work. * Correspondence: [email protected] bioRxiv preprint doi: https://doi.org/10.1101/675611; this version posted June 21, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY-NC-ND 4.0 International license. Abstract Penfield’s description of the “homunculus”, a “grotesque creature” with large lips and hands and small trunk and legs depicting the representation of body-parts within the primary somatosensory cortex (S1), is one of the most prominent contributions to the neurosciences. Since then, numerous studies have identified additional body- parts representations outside of S1.
    [Show full text]
  • How Does the Brain Produce Movement?
    p CHAPTER 10 How Does the Brain Produce Movement? The Hierarchical Control The Basal Ganglia and of Movement the Cerebellum The Forebrain and Movement Initiation The Basal Ganglia and Movement Force The Brainstem and Species-Typical Movement Focus on Disorders: Tourette’s Syndrome Focus on Disorders: Autism The Cerebellum and Movement Skill The Spinal Cord and Movement Execution Focus on Disorders: Paraplegia The Organization of the Somatosensory System The Organization of the Somatosensory Receptors and Sensory Perception Motor System Dorsal-Root Ganglion Neurons The Motor Cortex The Somatosensory Pathways to the Brain The Corticospinal Tracts Spinal-Cord Responses to Somatosensory Input The Motor Neurons The Vestibular System and Balance The Control of Muscles Exploring the Somatosensory The Motor Cortex and Skilled System Movements The Somatosensory Homunculus Investigating Neural Control of Skilled Movements The Effects of Damage to the Somatosensory Cortex The Control of Skilled Movements in Other Species The Somatosensory Cortex and Complex How Motor Cortex Damage Affects Skilled Movement Movements Kevork Djansezian/AP Photo Micrograph: Dr. David Scott/Phototake 354 I p amala is a female Indian elephant that lives at the In one way, however, Kamala uses this versatile trunk zoo in Calgary, Canada. Her trunk, which is really very unusually for an elephant (Onodera & Hicks, 1999). K just a greatly extended upper lip and nose, con- She is one of only a few elephants in the world that paints sists of about 2000 fused muscles. A pair of nostrils runs its with its trunk (Figure 10-1). Like many artists, she paints length and fingerlike projections are located at its tip.
    [Show full text]
  • 1592813808Somatosensory Pat
    Name: Ezenwobi Chiamaka Anne Matric number: 18/mhs07/020 Department: pharmacology Course code: PHS 212 Question: discuss the somatosensory pathway The somatosensory system is a part of the sensory nervous system. The somatosensory system is a complex system of sensory neurons and neural pathways that responds to changes at the surface or inside the body. The axons (as afferent nerve fibers) of sensory neurons connect with, or respond to, various receptor cells. These sensory receptor cells are activated by different stimuli such as heat and nociception, giving a functional name to the responding sensory neuron, such as a chemoreceptor which carries information about temperature changes. Other types include mechanoreceptors, thermoreceptors, and nociceptors which send signals along a sensory nerve to the spinal cord where they may be processed by other sensory neurons and then relayed to the brain for further processing. Sensory receptors are found all over the body including the skin, epithelial tissues, muscles, bones and joints, internal organs, and the cardiovascular system. A somatosensory pathway will typically consist of three neurons: primary, secondary, and tertiary. 1. In the periphery, the primary neuron is the sensory receptor that detects sensory stimuli like touch or temperature. The cell body of the primary neuron is housed in the dorsal root ganglion of a spinal nerve or, if sensation is in the head or neck, the ganglia of the trigeminal or cranial nerves. 2. The secondary neuron acts as a relay and is located in either the spinal cord or the brainstem. This neuron’s ascending axons will cross, or decussate, to the opposite side of the spinal cord or brainstem and travel up the spinal cord to the brain, where most will terminate in either the thalamus or the cerebellum.
    [Show full text]
  • Parietal Lobes: Evolution, Neuroanatomy, and Function
    Parietal Lobes: Evolution, Neuroanatomy, and Function Charles J. Vella, PhD May 3, 2017 Intent of this lecture Parietal lobe syndromes are mostly domain of neurology, not NP But these syndromes will affect NP performance on multiple tests There are lots of anatomical references: you can always use the pdf online as reference text There are lots of unusual, sometimes rare, syndromes My objective is to have you be aware of when parietal lobe functioning is involved in something you are seeing clinically in a patient The discovery of the Parietal Lobe Franciscus Sylvius (1614-1672) Physician, physiologist, anatomist, and chemist Professor at the University of Leidon (1641) Was in the chemistry lab at the University when he discovered the deep cleft (Sylvian Fissure). History of Parietal Lobe Discovery In 1874 Bartholow recorded odd sensation from legs on stimulating post central gyrus through skull wounds Djerine – alexia , agraphia -- angular gyrus lesion Hugo Liepmann--- ideomotor & ideational apraxia in (L) sided lesion Cushing in 1909 --- Electrical stimulation in conscious human beings –– mainly tactile hallucinations; first homunculus Critchley (1953) – 1st monograph on “ The Parietal Lobes” Siegel et al. (2003) – “The Parietal Lobes” Macdonald Critchley: Parietal Lobes Parietal Lobes No independent existence as anatomical / physiological unit Operates in conjunction with brain as a whole Strategically situated between other lobes Greater variety of clinical manifestations than rest of the hemisphere Ignored in
    [Show full text]
  • Structure Processing
    Name: Opoola Temitope Susan Matric No: 18/MHS02/164 Department: Nursing Course code: PHS212 Date: Tuesday 19th, June, 2020 Discuss The somatosensory pathways Somatosensory function is the ability to interpret bodily sensation. Sensation takes a number of forms, including touch, pressure, vibration, temperature, itch, tickle, and pain. The somatosensory system allows us to interpret sensory messages received from our body and consists of sensory receptors located in the skin, tissues, and joints; the nerve cell tracts in the body and spinal cord; and brain centers that process and modulate incoming sensory information The somatosensory system is distributed throughout all major parts of our body. It is responsible for sensing touch, temperature, posture, limb position, and more. It includes both sensory receptor neurons in the periphery (eg., skin, muscle, and organs) and deeper neurons within the central nervous system. Structure A somatosensory pathway consist of three neurons: primary, secondary, and tertiary. 1. In the periphery, the primary neuron is the sensory receptor that detects sensory stimuli like touch or temperature. The cell body of the primary neuron is housed in the dorsal root ganglion of a spinal nerve or, if sensation is in the head or neck, the ganglia of the trigeminal or cranial nerves. 2. The secondary neuron acts as a relay and is located in either the spinal cord or the brainstem. This neuron’s ascending axons will cross, or decussate, to the opposite side of the spinal cord or brainstem and travel up the spinal cord to the brain, where most will terminate in either the thalamus or the cerebellum.
    [Show full text]
  • Name: Ikemefuna Dumebi Matric Number: 18/Mhs02/092 Level: 200Lvl Department: Nursing Science Course: Phs 212 (Physiology) Questi
    NAME: IKEMEFUNA DUMEBI MATRIC NUMBER: 18/MHS02/092 LEVEL: 200LVL DEPARTMENT: NURSING SCIENCE COURSE: PHS 212 (PHYSIOLOGY) QUESTION Discuss the somatosensory pathways ANSWERS The somatosensory system is composed of the neurons that make sensing touch, temperature, and position in space possible. The somatosensory system is distributed throughout all major parts of our body. It is responsible for sensing touch, temperature, posture, limb position, and more. It includes both sensory receptor neurons in the periphery (eg., skin, muscle, and organs) and deeper neurons within the central nervous system. Structure A somatosensory pathway will typically consist of three neurons: primary, secondary, and tertiary. 1. In the periphery, the primary neuron is the sensory receptor that detects sensory stimuli like touch or temperature. The cell body of the primary neuron is housed in the dorsal root ganglion of a spinal nerve or, if sensation is in the head or neck, the ganglia of the trigeminal or cranial nerves. 2. The secondary neuron acts as a relay and is located in either the spinal cord or the brainstem. This neuron’s ascending axons will cross, or decussate, to the opposite side of the spinal cord or brainstem and travel up the spinal cord to the brain, where most will terminate in either the thalamus or the cerebellum. 3. Tertiary neurons have cell bodies in the thalamus and project to the postcentral gyrus of the parietal lobe, forming a sensory homunculus in the case of touch. Regarding posture, the tertiary neuron is located in the cerebellum. Processing The primary somatosensory area of the human cortex is located in the postcentral gyrus of the parietal lobe.
    [Show full text]
  • The Use of High-Density EEG to Map out Cortical Motor Activity and Reorganization Following Lower
    The use of high-density EEG to map out cortical motor activity and reorganization following lower- limb amputation Valur Guðnason1 Ásgeir Alexandersson2, Kyle Edmunds3, Kristleifur Kristjánsson2, Þorvaldur Ingvarsson1,2, Paolo Gargiulo3,4 1University of Iceland, 2Össur, 3Reykjavik University, 4The National University Hospital of Iceland Thesis for a degree of Bachelor of Science University of Iceland Faculty of Medicine School of Health Sciences 1 Thesis for the degree of Bachelor of Science. All rights reserved. No part of this publication may be reproduced or transmitted, in any form or by any means, without written permission. © Valur Guðnason 2016 Printed at: Háskólaprent, Reykjavík, Iceland 2016 2 3 Abstract The use of high-density EEG to map out cortical motor activity and reorganization following lower-limb amputation Author: Valur Guðnason Supervisors: Ásgeir Alexandersson, Kyle Edmunds, Kristleifur Kristjansson, Þorvaldur Ingvarsson, Paolo Gargiulo Introduction Studies have shown that after amputation, changes occur in the sensory and motor cortex. These changes are called cortical reorganization, where adjacent cortical areas occupy the cortical area of the amputated limb. High-density electroencephalography (EEG) has been used to observe cortical reorganization in the motor cortex following upper limb amputation. The aim of this study was to use high-density EEG to map out motor cortical activity and cortical reorganization following lower limb amputation. Materials and methods One healthy and one left transfemoral amputee participated in the study. Using a 256-electrode, high-density EEG system, EEG signals were acquired while participants performed sets of motor tasks. The amputated participant was asked to try to voluntarily execute each movement for the missing limb, not simply imagine the movement.
    [Show full text]
  • General Organization of Somatosensory System the Somatosensory System Is Composed of the Neurons That Make Sensing Touch, Temperature, and Position in Space Possible
    Somatosensory System Boundless General Organization of Somatosensory System The somatosensory system is composed of the neurons that make sensing touch, temperature, and position in space possible. 1. fig. 1 shows a dorsal root ganglion Sensory nerves of a dorsal root ganglion are depicted entering the spinal cord. Our somatosensory system consists of primary, secondary, and tertiary neurons. Sensory receptors housed in dorsal root ganglia project to secondary neurons of the spinal cord that decussate and project to the thalamus or cerebellum. Tertiary neurons project to the postcentral gyrus of the parietal lobe, forming a sensory humunculus. A sensory homunculus maps sub-regions of the cortical poscentral gyrus to certain parts of the body. Decussate: Where nerve fibers obliquely cross from one lateral part of the body to the other. Postcentral Gyrus: A prominent structure in the parietal lobe of the human brain and an important landmark that is the location of the primary somatosensory cortex, the main sensory receptive area for the sense of touch. Thalamus: Either of two large, ovoid structures of gray matter within the forebrain that relay sensory impulses to the cerebral cortex. Source URL: https://www.boundless.com/physiology/peripheral-nervous-system-pns/somatosensory-system/ Saylor URL: http://www.saylor.org/courses/psych402/ Attributed to: [Boundless] www.saylor.org Page 1 of 16 2. fig. 2 shows the sensory homunculus of the human brain 3. fig. 3 shows the sagittal MRI of the human brain The thalamus is marked by a red arrow in this MRI cross-section. Our somatosensory system is distributed throughout all major parts of our body.
    [Show full text]
  • ENCODING of SALTATORY TACTILE VELOCITY in the ADULT OROFACIAL SOMATOSENSORY SYSTEM Rebecca Custead University of Nebraska-Lincoln, [email protected]
    University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Public Access Theses and Dissertations from the Education and Human Sciences, College of (CEHS) College of Education and Human Sciences 7-2016 ENCODING OF SALTATORY TACTILE VELOCITY IN THE ADULT OROFACIAL SOMATOSENSORY SYSTEM Rebecca Custead University of Nebraska-Lincoln, [email protected] Follow this and additional works at: http://digitalcommons.unl.edu/cehsdiss Part of the Behavioral Neurobiology Commons, Biomechanics Commons, Digestive, Oral, and Skin Physiology Commons, Medical Neurobiology Commons, Molecular and Cellular Neuroscience Commons, Neurology Commons, Other Kinesiology Commons, Other Medicine and Health Sciences Commons, Other Neuroscience and Neurobiology Commons, and the Rehabilitation and Therapy Commons Custead, Rebecca, "ENCODING OF SALTATORY TACTILE VELOCITY IN THE ADULT OROFACIAL SOMATOSENSORY SYSTEM" (2016). Public Access Theses and Dissertations from the College of Education and Human Sciences. 273. http://digitalcommons.unl.edu/cehsdiss/273 This Article is brought to you for free and open access by the Education and Human Sciences, College of (CEHS) at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Public Access Theses and Dissertations from the College of Education and Human Sciences by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. ENCODING OF SALTATORY TACTILE VELOCITY IN THE ADULT OROFACIAL SOMATOSENSORY SYSTEM by Rebecca Custead A DISSERTATION Presented to the Faculty of The Graduate College at the University of Nebraska In Partial Fulfillment of Requirements For the degree of Doctor of Philosophy Major: Human Sciences Under the Supervision of Professor Steven M. Barlow PhD Lincoln, Nebraska July, 2016 ENCODING OF SALTATORY TACTILE VELOCITY IN THE ADULT OROFACIAL SOMATOSENSORY SYSTEM Rebecca Custead, Ph.D.
    [Show full text]