Zootaxa 4521 (1): 001–051 ISSN 1175-5326 (print edition) http://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2018 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4521.1.1 http://zoobank.org/urn:lsid:zoobank.org:pub:D2096E74-49D8-4235-B26C-2C97170DBDC7 Taxonomy of coccids (Hemiptera: Coccidae: Coccus L.) associated with Crematogaster ants (Hymenoptera: Formicidae) in the stems of Macaranga plants (Euphorbiaceae) in Southeast Asia PENNY J. GULLAN1, TAKUMASA KONDO2, BRIGITTE FIALA3 & SWEE-PECK QUEK4 1Division of Ecology & Evolution, Research School of Biology, College of Science, The Australian National University, Acton, Can- berra, A.C.T. 2601, Australia. E-mail:
[email protected] 2Corporación Colombiana de Investigación Agropecuaria (Agrosavia), Centro de Investigación Palmira, Calle 23, Carrera 37 Con- tinuo al Penal, Palmira, Valle, Colombia. E-mail:
[email protected] 3Department of Animal Ecology and Tropical Biology, Biocenter, University of Würzburg, Am Hubland, 97074 Würzburg, Germany. E- mail:
[email protected] 4Department of Integrative Biology, University of California, Berkeley 94720, U.S.A. E-mail:
[email protected] Table of content Abstract . 1 Introduction . 2 Materials and methods . 4 Taxonomy . 8 Coccus Linnaeus . 8 Diagnosis for Coccus species associated with Macaranga. 9 Key to adult females of Coccus species associated with Macaranga . 12 Coccus caviramicolus Morrison . 13 Coccus circularis Morrison. 14 Coccus heckrothi Gullan & Kondo sp. n. 18 Coccus lambirensis Gullan & Kondo sp. n. 20 Coccus macarangae Morrison . 22 Coccus macarangicolus Takahashi . 25 Coccus penangensis Morrison . 30 Coccus pseudotumuliferus Gullan & Kondo sp. n. 34 Coccus secretus Morrison . 39 Coccus tumuliferus Morrison .