Hominid/Human Evolution
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Defining the Genus Homo
Defining the Genus Homo Mark Collard and Bernard Wood Contents Introduction ..................................................................................... 2108 Changing Interpretations of Genus Homo ..................................................... 2109 Is Genus Homo a “Good” Genus? ............................................................. 2114 Updating Wood and Collard’s (1999) Review of Genus Homo .............................. 2126 Conclusion ...................................................................................... 2137 Cross-References ............................................................................... 2138 References ...................................................................................... 2138 Abstract The definition of the genus Homo is an important but under-researched topic. In this chapter we show that interpretations of Homo have changed greatly over the last 150 years as a result of the incorporation of new fossil species, the discovery of fossil evidence that changed our perceptions of its component species, and reassessments of the functional capabilities of species previously allocated to Homo. We also show that these changes have been made in an ad hoc fashion. Criteria for recognizing fossil specimens of Homo have been outlined on a M. Collard (*) Human Evolutionary Studies Program and Department of Archaeology, Simon Fraser University, Burnaby, BC, Canada Department of Archaeology, University of Aberdeen, Aberdeen, UK e-mail: [email protected] B. Wood Center for the Advanced -
COVID-19 and Human Rights: We Are All in This Together
COVID-19 and Human Rights We are all in this together APRIL 2020 Human rights are critical – for the response and the recovery They put people at the centre and produce better outcomes Human rights are key in shaping the pandemic response, both for the public health emergency and the broader impact on people’s lives and livelihoods. Human rights put people centre-stage. Responses that are shaped by and respect human rights result in better outcomes in beating the pandemic, ensuring healthcare for everyone and preserving human dignity. But they also focus our attention on who is suffering most, why, and what can be done about it. They prepare the ground now for emerging from this crisis with more equitable and sustainable societies, development and peace. Why are human rights equip States and whole societies to respond to so important to the threats and crises in a way that puts people at the centre. Observing the crisis and its impact COVID-19 response? through a human rights lens puts a focus on how it is affecting people on the ground, partic- The world is facing an unprecedented crisis. ularly the most vulnerable among us, and what At its core is a global public health emer- can be done about it now, and in the long term. gency on a scale not seen for a century, Although this paper presents recommenda- requiring a global response with far-reaching tions, it is worth underlining that human rights consequences for our economic, social and are obligations which States must abide by. political lives. -
Homo Habilis
COMMENT SUSTAINABILITY Citizens and POLICY End the bureaucracy THEATRE Shakespeare’s ENVIRONMENT James Lovelock businesses must track that is holding back science world was steeped in on surprisingly optimistic governments’ progress p.33 in India p.36 practical discovery p.39 form p.41 The foot of the apeman that palaeo ‘handy man’, anthropologists had been Homo habilis. recovering in southern Africa since the 1920s. This, the thinking went, was replaced by the taller, larger-brained Homo erectus from Asia, which spread to Europe and evolved into Nean derthals, which evolved into Homo sapiens. But what lay between the australopiths and H. erectus, the first known human? BETTING ON AFRICA Until the 1960s, H. erectus had been found only in Asia. But when primitive stone-chop LIBRARY PICTURE EVANS MUSEUM/MARY HISTORY NATURAL ping tools were uncovered at Olduvai Gorge in Tanzania, Leakey became convinced that this is where he would find the earliest stone- tool makers, who he assumed would belong to our genus. Maybe, like the australopiths, our human ancestors also originated in Africa. In 1931, Leakey began intensive prospect ing and excavation at Olduvai Gorge, 33 years before he announced the new human species. Now tourists travel to Olduvai on paved roads in air-conditioned buses; in the 1930s in the rainy season, the journey from Nairobi could take weeks. The ravines at Olduvai offered unparalleled access to ancient strata, but field work was no picnic in the park. Water was often scarce. Leakey and his team had to learn to share Olduvai with all of the wild animals that lived there, lions included. -
Hands-On Human Evolution: a Laboratory Based Approach
Hands-on Human Evolution: A Laboratory Based Approach Developed by Margarita Hernandez Center for Precollegiate Education and Training Author: Margarita Hernandez Curriculum Team: Julie Bokor, Sven Engling A huge thank you to….. Contents: 4. Author’s note 5. Introduction 6. Tips about the curriculum 8. Lesson Summaries 9. Lesson Sequencing Guide 10. Vocabulary 11. Next Generation Sunshine State Standards- Science 12. Background information 13. Lessons 122. Resources 123. Content Assessment 129. Content Area Expert Evaluation 131. Teacher Feedback Form 134. Student Feedback Form Lesson 1: Hominid Evolution Lab 19. Lesson 1 . Student Lab Pages . Student Lab Key . Human Evolution Phylogeny . Lab Station Numbers . Skeletal Pictures Lesson 2: Chromosomal Comparison Lab 48. Lesson 2 . Student Activity Pages . Student Lab Key Lesson 3: Naledi Jigsaw 77. Lesson 3 Author’s note Introduction Page The validity and importance of the theory of biological evolution runs strong throughout the topic of biology. Evolution serves as a foundation to many biological concepts by tying together the different tenants of biology, like ecology, anatomy, genetics, zoology, and taxonomy. It is for this reason that evolution plays a prominent role in the state and national standards and deserves thorough coverage in a classroom. A prime example of evolution can be seen in our own ancestral history, and this unit provides students with an excellent opportunity to consider the multiple lines of evidence that support hominid evolution. By allowing students the chance to uncover the supporting evidence for evolution themselves, they discover the ways the theory of evolution is supported by multiple sources. It is our hope that the opportunity to handle our ancestors’ bone casts and examine real molecular data, in an inquiry based environment, will pique the interest of students, ultimately leading them to conclude that the evidence they have gathered thoroughly supports the theory of evolution. -
Microremains from El Miron Cave Human Dental Calculus Suggest A
Journal of Archaeological Science 60 (2015) 39e46 Contents lists available at ScienceDirect Journal of Archaeological Science journal homepage: http://www.elsevier.com/locate/jas Microremains from El Miron Cave human dental calculus suggest a mixed planteanimal subsistence economy during the Magdalenian in Northern Iberia * Robert C. Power a, , Domingo C. Salazar-García b, c, d, e, Lawrence G. Straus f, g, Manuel R. Gonzalez Morales g, Amanda G. Henry a a Research Group on Plant Foods in Hominin Dietary Ecology, Max Planck Institute for Evolutionary Anthropology, Leipzig, Germany b Department of Archaeology, University of Cape Town, Cape Town, South Africa c Departament de Prehistoria i Arqueologia, Universitat de Valencia, Valencia, Spain d Aix Marseille Universite, CNRS, Ministere de la culture et de la communication, LAMPEA UMR 7269, 13090 Aix-en-Provence, France e Department of Human Evolution, Max-Planck Institute for Evolutionary Anthropology, Leipzig, Germany f Department of Anthropology, University of New Mexico, Albuquerque, NM, USA g Instituto Internacional de Investigaciones Prehistoricas de Cantabria, Universidad de Cantabria, Santander, Spain article info abstract Article history: Despite more than a century of detailed investigation of the Magdalenian period in Northern Iberia, our Available online 13 April 2015 understanding of the diets during this period is limited. Methodologies for the reconstruction of Late Glacial subsistence strategies have overwhelmingly targeted animal exploitation, thus revealing only a Keywords: portion of the dietary spectrum. Retrieving food debris from calculus offers a means to provide missing Upper Palaeolithic information on other components of diet. We undertook analysis of human dental calculus samples from Archaeobotany Magdalenian individuals (including the “Red Lady”) at El Miron Cave (Cantabria, Spain), as well as several Palaeolithic diet control samples, to better understand the less visible dietary components. -
Human Evolution Timeline 1
Name: Human Origins Web Inquiry 1. Got to the website: http://humanorigins.si.edu/evidence 2. Read the paragraph under “Evidence of Evolution.” 3. In the maroon box on the left, click on “Timeline Interactive” 4. Start by clicking on the red bands at the bottom of the timeline, beginning with Homo sapiens and the rest of the Hominids. Read the information boxes that pop up when you click. 5. When you come across a species you find particularly interesting, record that information (including dates) in the box below. Record facts of at least 5 interesting hominid species, including Homo sapiens. 6. Then begin to explore the rest of the timeline. As you go, record at least 5 other interesting species. 7. Use the magnifier tool at the bottom of the timeline. Click on “color key” to identify the different color dots. Each dot represents a specific piece of evidence scientists have discovered in their study of evolution. 8. Click on various colored dots to learn about tools, events, geology, climate information, behavior, and skeletal adaptations discovered by scientists that add to the evidence for human evolution. Record 7 pieces of evidence you find interesting or particularly important in the box on the back. Make sure to put down different types of evidence (different colors). 9. Take 15-20 minutes to explore the whole timeline and record interesting facts. 10. Build your own timeline on the next page, filling in the interesting species and evidence you recorded in their proper place in the timeline. Hominids Species Dates Interesting fact(s) Homo sapiens Name: Evidence Type of Date Interesting fact(s) Evidence Name: Human Evolution Timeline 1. -
The Adaptive Significance of Human Language
University of Tennessee, Knoxville TRACE: Tennessee Research and Creative Exchange Senior Thesis Projects, 1993-2002 College Scholars 2000 The Adaptive Significance of Human Language Nathan Oesch Follow this and additional works at: https://trace.tennessee.edu/utk_interstp2 Recommended Citation Oesch, Nathan, "The Adaptive Significance of Human Language" (2000). Senior Thesis Projects, 1993-2002. https://trace.tennessee.edu/utk_interstp2/52 This Project is brought to you for free and open access by the College Scholars at TRACE: Tennessee Research and Creative Exchange. It has been accepted for inclusion in Senior Thesis Projects, 1993-2002 by an authorized administrator of TRACE: Tennessee Research and Creative Exchange. For more information, please contact [email protected]. The Adaptive Significance of Human Language Nathan Oesch Department of Psychology University of Tennessee, Knoxville noesch @ utk. edu Abstract Many experts have argued that human language is fundamentally incompatible with the principles of traditional Darwinian evolutionary theory. According to conventional Darwinian explanations, specific traits evolved among species according to gradual and incremental genetic changes, each of which that were in some way so favorable to the survival and reproduction of ancestral generations that they were ultimately preserved within successive generations of those species. Human language, it has been said, is simply to complex to be explained as a result of Darwinian explanations, since each successive step in the evolution of language would confer no obvious survival benefits to its recipients. According to this idea, language is such an "all-or none system," that it could not possibly have existed in any immediately beneficial intermediate forms and thus could not have evolved according to conventional Darwinian modes of explanation. -
Neither Chimpanzee Nor Human, Ardipithecus Reveals the Surprising Ancestry of Both Tim D
SPECIAL FEATURE: PERSPECTIVE PERSPECTIVE SPECIAL FEATURE: Neither chimpanzee nor human, Ardipithecus reveals the surprising ancestry of both Tim D. Whitea,1, C. Owen Lovejoyb, Berhane Asfawc, Joshua P. Carlsona, and Gen Suwad,1 aDepartment of Integrative Biology, Human Evolution Research Center, University of California, Berkeley, CA 94720; bDepartment of Anthropology, School of Biomedical Sciences, Kent State University, Kent, OH 44242–0001; cRift Valley Research Service, Addis Ababa, Ethiopia; and dThe University Museum, The University of Tokyo, Hongo, Bunkyo-ku Tokyo 113-0033, Japan Edited by Neil H. Shubin, University of Chicago, Chicago, IL, and approved September 10, 2014 (received for review April 25, 2014) Australopithecus fossils were regularly interpreted during the late 20th century in a framework that used living African apes, especially chimpanzees, as proxies for the immediate ancestors of the human clade. Such projection is now largely nullified by the discovery of Ardipithecus. In the context of accumulating evidence from genetics, developmental biology, anatomy, ecology, biogeography, and geology, Ardipithecus alters perspectives on how our earliest hominid ancestors—and our closest living relatives—evolved. human evolution | Australopithecus | hominid | Ethiopia “...the stock whence two or more species have chimpanzees, can serve as adequate repre- (5). Indeed, a widely used textbook still pro- sprung, need in no respect be intermediate sentations of the ancestral past. claims that, “Overall, Au. afarensis seems very between those species.” much like a missing link between the living Background T. H. Huxley, 1860 (1) Africanapesandlaterhomininsinitsdental, ’ Darwin s human evolution scenario attemp- cranial, and skeletal morphology” (6). Charles Darwin famously suggested that ted to explain hominid tool use, bipedality, Australopithecus can no longer be legiti- Africa was humanity’s most probable birth enlarged brains, and reduced canine teeth (2). -
Human Evolution: a Paleoanthropological Perspective - F.H
PHYSICAL (BIOLOGICAL) ANTHROPOLOGY - Human Evolution: A Paleoanthropological Perspective - F.H. Smith HUMAN EVOLUTION: A PALEOANTHROPOLOGICAL PERSPECTIVE F.H. Smith Department of Anthropology, Loyola University Chicago, USA Keywords: Human evolution, Miocene apes, Sahelanthropus, australopithecines, Australopithecus afarensis, cladogenesis, robust australopithecines, early Homo, Homo erectus, Homo heidelbergensis, Australopithecus africanus/Australopithecus garhi, mitochondrial DNA, homology, Neandertals, modern human origins, African Transitional Group. Contents 1. Introduction 2. Reconstructing Biological History: The Relationship of Humans and Apes 3. The Human Fossil Record: Basal Hominins 4. The Earliest Definite Hominins: The Australopithecines 5. Early Australopithecines as Primitive Humans 6. The Australopithecine Radiation 7. Origin and Evolution of the Genus Homo 8. Explaining Early Hominin Evolution: Controversy and the Documentation- Explanation Controversy 9. Early Homo erectus in East Africa and the Initial Radiation of Homo 10. After Homo erectus: The Middle Range of the Evolution of the Genus Homo 11. Neandertals and Late Archaics from Africa and Asia: The Hominin World before Modernity 12. The Origin of Modern Humans 13. Closing Perspective Glossary Bibliography Biographical Sketch Summary UNESCO – EOLSS The basic course of human biological history is well represented by the existing fossil record, although there is considerable debate on the details of that history. This review details both what is firmly understood (first echelon issues) and what is contentious concerning humanSAMPLE evolution. Most of the coCHAPTERSntention actually concerns the details (second echelon issues) of human evolution rather than the fundamental issues. For example, both anatomical and molecular evidence on living (extant) hominoids (apes and humans) suggests the close relationship of African great apes and humans (hominins). That relationship is demonstrated by the existing hominoid fossil record, including that of early hominins. -
Downloaded from Brill.Com09/27/2021 09:14:05PM Via Free Access 218 Rode-Margono & Nekaris – Impact of Climate and Moonlight on Javan Slow Lorises
Contributions to Zoology, 83 (4) 217-225 (2014) Impact of climate and moonlight on a venomous mammal, the Javan slow loris (Nycticebus javanicus Geoffroy, 1812) Eva Johanna Rode-Margono1, K. Anne-Isola Nekaris1, 2 1 Oxford Brookes University, Gipsy Lane, Headington, Oxford OX3 0BP, UK 2 E-mail: [email protected] Keywords: activity, environmental factors, humidity, lunarphobia, moon, predation, temperature Abstract Introduction Predation pressure, food availability, and activity may be af- To secure maintenance, survival and reproduction, fected by level of moonlight and climatic conditions. While many animals adapt their behaviour to various factors, such nocturnal mammals reduce activity at high lunar illumination to avoid predators (lunarphobia), most visually-oriented nocturnal as climate, availability of resources, competition, preda- primates and birds increase activity in bright nights (lunarphilia) tion, luminosity, habitat fragmentation, and anthropo- to improve foraging efficiency. Similarly, weather conditions may genic disturbance (Kappeler and Erkert, 2003; Beier influence activity level and foraging ability. We examined the 2006; Donati and Borgognini-Tarli, 2006). According response of Javan slow lorises (Nycticebus javanicus Geoffroy, to optimal foraging theory, animal behaviour can be seen 1812) to moonlight and temperature. We radio-tracked 12 animals as a trade-off between the risk of being preyed upon in West Java, Indonesia, over 1.5 years, resulting in over 600 hours direct observations. We collected behavioural and environmen- and the fitness gained from foraging (Charnov, 1976). tal data including lunar illumination, number of human observ- Perceived predation risk assessed through indirect cues ers, and climatic factors, and 185 camera trap nights on potential that correlate with the probability of encountering a predators. -
Using Sub-Species Level Phylogenies
1 Reconstructing the ancestral phenotypes of great apes and 2 humans (Homininae) using sub-species level phylogenies 3 Keaghan Yaxley1 and Robert Foley2 4 1Leverhulme Centre for Human Evolutionary Studies, University of Cambridge, UK 5 2Leverhulme Centre for Human Evolutionary Studies, University of Cambridge, UK 6 7 8 1 9 Abstract 10 By their close affinity, the African great apes are of interest to the study of human evolution. 11 While numerous researchers have described the ancestors we share with these species with 12 reference to extant great apes, few have done so with phylogenetic comparative methods. 13 One obstacle to the application of these techniques is the within-species phenotypic variation 14 found in this group. Here we leverage this variation, modelling common ancestors using 15 Ancestral State Reconstructions (ASRs) with reference to subspecies level trait data. A 16 subspecies level phylogeny of the African great apes and humans was estimated from full- 17 genome mtDNA sequences and used to implement ASRs for fifteen continuous traits known 18 to vary between great ape subspecies. While including within-species phenotypic variation 19 increased phylogenetic signal for our traits and improved the performance of our ASRs, 20 whether this was done through the inclusion of subspecies phylogeny or through the use of 21 existing methods made little difference. Our ASRs corroborate previous findings that the Last 22 Common Ancestor (LCA) of humans, chimpanzees and bonobos was a chimp-like animal, 23 but also suggest that the LCA of humans, chimpanzees, bonobos and gorillas was an animal 24 unlike any extant African great ape. -
Photopigments and Color Vision in the Nocturnal Monkey, Aotus GERALD H
Vision Res. Vol. 33, No. 13, pp. 1773-1783, 1993 0042-6989/93 $6.00 + 0.00 Printed in Great Britain. All rights reserved Copyright 0 1993 Pergamon Press Ltd Photopigments and Color Vision in the Nocturnal Monkey, Aotus GERALD H. JACOBS,*? JESS F. DEEGAN II,* JAY NEITZ,$ MICHAEL A. CROGNALE,§ MAUREEN NEITZT Received 6 November 1992; in revised form 3 February 1993 The owl monkey (Aotus tridrgutus) is the only nocturnal monkey. The photopigments of Aotus and the relationship between these photopigments and visual discrimination were examined through (1) an analysis of the tlicker photometric electroretinogram (ERG), (2) psychophysical tests of visual sensitivity and color vision, and (3) a search for the presence of the photopigment gene necessary for the production of a short-wavelength sensitive (SWS) photopigment. Roth electrophysiological and behavioral measurements indicate that in addition to a rod photopigment the retina of this primate contains only one other photopigment type-a cone pigment having a spectral peak cu 543 nm. Earlier results that suggested these monkeys can make crude color discriminations are interpreted as probably resulting from the joint exploitation of signals from rods and cones. Although Aotus has no functional SWS photopigment, hybridization analysis shows that A&us has a pigment gene that is highly homologous to the human SWS photopigment gene. Aotus trivirgatus Cone photopigments Monkey color vision Monochromacy Photopigment genes Evolution of color vision INTRODUCTION interest to anyone interested in visual adaptations for two somewhat contradictory reasons. On the one hand, The owl monkey (A&us) is unique among present study of A&us might provide the possibility of docu- day monkeys in several regards.