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Catalog of the Family Diploglossidae

Harold De Lisle

2017

Cover: fasciatus Photo by Daniel R. Lordelo

Back cover: warreni Photo by Milo Ko řínek

Email: [email protected]

Family Diploglossidae

The family Diploglossidae is distributed in the West Indies, Central America, and South America. The family contains three genera Celestus has 33 and Diploglossus with 14 species. Both share a similar snout, elongated cylindrical body with short well developed limbs. The Ophiodes has 5 species. As the Latin name suggests, they have a snake-like body, with not front limbs and only small rudimentary hind limbs.

The family species have a body covered by cycloid scales that are underlain by osteoderms having a distinctive, peaked, gliding surface.

The family contains both egglayers and viviparous species. The larger species can produce more than 20 offspring.

Savage et al (2008) list the members of the Diploglossidae with the claw as the definitive character: exposed claws in Celestus and claws sheated in the other genera.

Pyron & Burbrink (2013) found Diploglossus and Celestus are strongly supported as paraphyletic with respect to each other and to Ophiodes , but they had data on only six taxa.

Abbreviations of museums holding the types are as follows

AMNH - American Museum of Natural History, Division of Vertebrate Zoology (Herpetology), 79th Street and Central Park West, New York, New York 10024, USA. BMNH – The Natural History Museum, Department of Zoology, Cromwell Road, London SW75BD, United Kingdom. CM - Carnegie Museum of Natural History, Pittsburgh, Pennsylvania, USA. ENCB - Escuela Nacional de Ciencias Biológicas Campus Santo Tomás, 11350 Ciudad de México, CDMX, Mexico FMNH - Field Museum, Division of Amphibians and , Roosevelt Road at Lake ShoreDrive, Chicago, Illinois 60605, USA HMZ - Hamburg University Zoology Museum, Hamburg, Germany KU - University of Kansas, Museum of Natural History, Division of Herpetology, Lawrence, Kansas 66045, USA . LACM - Natural History Museum of Los Angeles County, Los Angeles, California 900 Exposition Blvd, Los Angeles, CA 90007 MACN - Museo Argentino de Ciencias Naturales "Bernardino Rivadavia", Av. Angel Gallardo 470, C1405DJR, Buenos Aires, Argentina MCZ - Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts 02138, USA. MNHNP - Museum National dHistoire Naturelle, Laboratoire des Amphibiens et Reptiles, 25 Rue Cuvier, 75005 Paris, France MRSN - Museo Regionale di Scienze Naturali di Torino, Via Giovanni Giolitti 36, Torino, Italy MVUP - Museo de Vertebrados de la Universidad de Panamá, Panama City, Panama CHFURG – Colecáõ Herpetológica da Fundacáõ Universidade do Rio Grande, Av. Itália km 8 Bairro , Carreiros, Rio Grande, Rio Grande do Sul, Brasil RCSOM – Royal College of Surgeons of England Museum, London RMNH - Naturalis (Nationaal Natuurhistorisch Museum; formerly Rijksmuseum van Natuurlijke Historie), Postbus 9517, 2300RA Leiden, The Netherlands. USNM - National Museum of Natural History, Division of Amphibians and Reptiles, Washington, D.C. 20560, USA . UTA- University of Texas at Arlington, Department of Biological Sciences, UTA Box 19489, Arlington, Texas 76019, USA. ZMB - Universität Humboldt, Zoologisches Museum, Invalidenstrasse 43, 10115 Berlin, Germany. ZSM - Zoologische Staatssammlung München, Pasing-Obermenzing, Münchhausenstraße 21, 81247 München, Germany

Diploglossidae Bocourt, 1873 Bocourt, M.F. 1873 : Recherches Zoologiques pour servir à l'Histoire de la Faune de l'Amérique Centrale et du Mexique. Mission Scientifique au Mexique et dans l'Amérique Centrale , Recherches zoologiques. Part 2, sect. 1 . In: Duméril, Bocourt & Mocquard, 1870-1909, Études sur les reptiles , vol. 2-15, p. 33-860. Type genus: Diploglossus Wiegmann, 1834

Celestus Gray. 1839 Gray, J.E. 1839. A catalogue of the slender-tongued saurians, with descriptions of many new genera and species. Annals and Magazine of Natural History ser 2, 1: 288-293. Type species: Celestus striatus Gray, 1838: 288

Celestus adercus Savage, Lips & Ibáñez, 2008 Celestus adercus Savage, Lips & Ibáñez, 2008: 851 Type locality: abandoned sawmill site on the continental divide, 9.7 km NNW of El Copé (8°40’04” N, 80°35’6” W), Parque Nacional General de División Omar Torrijos Herrera, La Pintada District, Coclé Province, Panama. Holotype: MVUP 1894 Distribution: Serranía de Tabasará, Panama

Celestus adercus Barry Zimmer

Celestus agasepsoides (Thomas, 1971)

Diploglossus agasepsoides Thomas, 1971:2. Type locality: Barreras, , República Dominicana Holotype: USNM 166964 Distribution: Sierra Martin Garcia and Peninsula de Barahona, República Dominicana

Celestus agasepsoides Miguel Landestoy

Celestus anelpistus (Schwartz, Graham & Duval, 1979) Diploglossus anelpistus Schwartz, Graham & Duval, 1979:3 Type locality: Ingenio Catarey, "Come Hombre," , San Cristóbal Province, República Dominicana Holotype: USNM 197336 Distribution: Villa Altagracia, San Cristóbal Province, República Dominicana Last seen 1977

Celestus atitlanensis Smith, 1950 Celestus atitlanensis Smith, 1950: 195 Type locality: Atitlán, Guatemala Holotype: MNHNP 0.5206 Distribution: Mexico: Chiapas State; Pacific slope of Guatemala; El Salvador

Celestus badius Cope, 1868

Celestus badius Cope, 1868: 126 Type locality: Navassa Island Syntypes: USNM 25817-18 Distribution: USA: Navassa Island

Celestus badius Robert Powell

Celestus barbouri Grant, 1940

Celestus barbouri Grant, 1940: 101 Type locality: Mandeville, Manchester Parish, Jamaica Holotype: MCZ 45169 Distribution: central Jamaica: St. James, Trelawny, Manchester, St. Ann parishes

Celestus barbouri Alejandro Sanchez

Celestus carraui (Inchaústegui, Schwartz, and Henderson, 1985) Diploglossus carraui Inchaustegui, Schwartz, and Henderson, 1985:

196 Type locality: Comedero, La Isabela, , República Dominicana Holotype: USNM 197369. Distribution: northern República Dominicana: to Salcedo Province

Celestus carraui R Sadjak

Celestus costatus (Cope, 1862) Celestus costatus costatus (Cope, 1862) Panolopus costatus Cope, 1862: 494 Type locality: Near Jérémie, Département de la Grand'Anse, Haiti Holotype: MCZ 3606 Synonyms : Celestus phoxinus Cope, 1868 Type locality: Near Jérémie, Département de la Grand'Anse, Haiti Holotype: MCZ 12457 Diploglossus ohlendorffi Fischer, 1886 Type locality: Haiti Holotype: HZM, destroyed in WWII Diploglossus nuchalis Boulenger, 1898 Type locality: unknown Holotype: BMNH 1897.3.16.1 Distribution: Haiti: Tiburon Peninsula

Celestus costatus aenetergum Schwartz & Jacobs, 1989 Celestus costatus aenetergum Schwartz & Jacobs, 1989: 193 Type locality: Isla Catalinita, , República Dominicana Holotype: USNM 197323 Distribution: Isla Catalinita, La Altagracia Province, República Dominicana

Celestus costatus chalcorhabdus (Schwartz, 1964) Diploglossus costatus chalcorhabdus Schwartz, 1964: 37 Type locality: 0.9 mi. SE El Macao, La Altagracia Province, República Dominicana Holotype: MCZ 77158 Distribution: extreme eastern República Dominicana: La Altagracia Province to La Romana Province

Celestus costatus emys (Schwartz, 1971) Diploglossus costatus emys Schwartz, 1971: 163 Type locality: Île de la Tortue, Haiti Holotype: USNM 167300 Distribution: Haiti: Île de la Tortue

Celestus costatus leionotus (Schwartz, 1964) Diploglossus costatus leionotus Schwartz, 1964: 28 Type locality: 15 km SE San Juan, San Juan Province, República Dominicana Holotype: MCZ 77154 Distribution: República Dominicana: Valle de San Juan and intermontane valleys in Sierra de , east to Llanos de Azua and southern slopes of Cordillera Central.

Celestus costatus melanchrous (Schwartz, 1964) Diploglossus costatus melanchrous Schwartz, 1964: 34 Type locality: 8 km E Gasper Hernández. , República Dominicana Holotype: MCZ 77157 Distribution: north-central and eastern Haiti, east to Peninsula de Samaná, República Dominicana; inland to Hato Mayor, , San Cristóbal, Peravia, Monseñor Nouel, and La Vega provinces.

Celestus costatus neiba (Schwartz, 1964)

Diploglossus costatus neiba Schwartz, 1964: 30 Type locality: 19 km SW , Elías Piña Province, República Dominicana Holotype: MCZ 77155 Distribution: Sierra de Neiba, Elías Piña & Independencia Provinces, República Dominicana Celestus costatus neiba Mel Rivera

Celestus costatus nesobous (Schwartz, 1964) Diploglossus costatus nesobous Schwartz, 1964: 23 Type locality: western end, Ile-á-Vache, Haiti Holotype: MCZ 77153 Distribution: Haiti: Ile-á-Vache

Celestus costatus oreistes (Schwartz, 1964) Diploglossus costatus oreistes Schwartz, 1964: 25 Type locality: Oriani, Department du Sud-Est, Haiti Holotype: MCZ 74940 Distribution: southeastern Haiti:Morne de Cayette, Montagne Noire, Massif de la Selle (and associated lowlands); east into República Dominicana: Sierra de Baoruco to east coast of Peninsula de Barahona

Celestus costatus psychonothes (Schwartz, 1964) Diploglossus costatus psychonothes Schwartz, 1964: 32 Type locality: 1 mi S Constanza, , República Dominicana Holotype: MCZ 77156 Distribution: República Dominicana: interior Cordillera Central

Celestus costatus saonae (Schwartz, 1971) Diploglossus costatus saonae Schwartz, 1971: 161 Type locality: 0.5 mi W Mano Juan, Isla Saona, República Dominicana Holotype: CM 52285 Distribution: República Dominicana: Isla Saona

Celestus crusculus (Garman, 1887) Celestus crusculus crusculus (Garman, 1887) Diploglossus crusculus Garman, 1887:22 Type locality: Kingston, Kingston Parish, Jamaica Holotype: MCZ 6051 Distribution: coastal areas of Jamaica

Celestus crusculus cundalli Grant, 1940 Celestus crusculus cundalli Grant, 1940: 157 Type locality: Mandeville, Manchester Parish, Jamaica Holotype: MCZ 45163 Distribution: interior of Jamaica

Celestus crusculus cundalli J P Burgess

Celestus crusculus maculatus (Garman, 1888) Diploglossus maculatus Garman, 1888: 5 Type locality: Cayman Brac, Cayman Islands Holotype: MCZ 6231 Distribution: Cayman Islands: Little Cayman I., Cayman Brac

Celestus crusculus molesworthi Grant, 1940 Celestus crusculus molesworthi Grant, 1940: 157 Type locality: Near Buff Bay, Portland Parish, Jamaica Holotype: MCZ 45184 Distribution: coastal northeastern Jamaica between Buff Bay and Boston Bay

Celestus curtissi Grant, 1951 Celestus curtissi curtissi Grant, 1951 Celestus curtissi Grant, 1951: 68 Type locality: Trou Forban, Département de l'Ouest, Haiti Holotype: USNM 11733 Distribution: Haiti: Plaine de Cul de Sac, north to St.Marc, Île de la Gonâve; República Dominicana: east into Valle de Neiba

Celestus curtissi aporus (Schwartz, 1964) Diploglossus curtissi aporus Schwartz, 1964: 45 Type locality: 13.1 mi SW Enriquillo, , República Dominicana Holotype: MCZ 77159 Distribution: República Dominicana: Peninsula de Barahona, west to Dominico-Haitian border, north to Barahona

Celestus curtissi diastatus (Schwartz, 1964) Diploglossus curtissi diastatus Schwartz, 1964: 42 Type locality: Bombardopolis, Departement du Nord-Ouest, Haiti Holotype: MCZ 63402 Distribution: Haiti: Presqu'île du Nord-Ouest, south to Ça Soleil and Terre Neuve; Île de la Tortue.

Celestus curtissi hylonomus (Schwartz, 1964) Diploglossus curtissi hylonomus Schwartz, 1964: 49 Type locality: 0.5 mi NW Boa de Yuma, La Altagracia Province, República Dominicana Holotype: MCZ 77160 Distribution: República Dominicana: southeastern coast, from Rio Ozama to Cabo Engaño Celestus curtissi hylonomus Carlos De Soto M

Celestus cyanochloris Cope, 1894

Celestus cyanochloris Cope, 1894: 200 Type locality: Volcán Irazú, Costa Rica Holotype: AMNH R16290 Distribution: Costa Rica: Cordilleras de Tilarán

Celestus cyanochloris J Savage

Celestus darlingtoni Cochran, 1939

Celestus darlingtoni Cochran, 1939: 2 Type locality: Valle Nuevo, in the Cordillera Central southeast of Constanza, La Vega Province, República Dominicana. Holotype: MCZ 44374. Distribution: República Dominicana: Cordillera Central, La Vega and Santiago Provinces.

Celestus darlingtoni Miguel Landestoy

Celestus duquesneyi Grant, 1940 Celestus duquesneyi Grant, 1940: 157 Type locality: Portland Point, Clarendon Parish, Jamaica Holotype: MCZ 45194 Distribution: Jamaica: Portland Peninsula

Celestus enneagrammus (Cope, 1860)

Siderolamprus enneagrammus Cope, 1860: 368 Type locality: Jalapa, Veracruz, Mexico Holotype: MCZ 2848 Synonyms: Diploglossus steindachneri Cope, 1864 Type locality: Orizaba, Veracruz, Mexico Holotype: USNM 6842 Diploglossus chalybaeus Cope, 1867 Type locality: Orizaba, Veracruz, Mexico Holotype: USNM 6603 Celestus enneagrammus Eric Centenero A Distribution: Mexico: eastern foothills of Veracruz, Oaxaca, Puebla, Chiapas States

Celestus fowleri (Schwartz, 1971) Diploglossus fowleri Schwartz, 1971:3 Type locality: Windsor, Trelawny Parish, Jamaica Holotype: MCZ 125601 Distribution: Jamaica: Trelawny Parish Last see 1970

Celestus haetianus (Cochran, 1927) Celestus haetianus haetianus (Cochran, 1927) Wetmorena haetiana Cochran, 1927: 91 Type locality: Mont Cabalo, Massif de la Selle, Département du Sud-Est, Haiti. Holotype: USNM 72600 Distribution: Haiti: Morne la Viste, Pic la Selle, Montagne Noire

Celestus haetianus mylicus (Schwartz, 1965) Wetmorena haetiana mylica Schwartz, 1965: 45 Type locality: 24 km SW Barahona, , República Dominicana. Holotype: MCZ 77049 Distribution: República Dominicana : eastern Sierra de Baoruco

Celestus haetianus surdus (Schwartz, 1965) Wetmorena haetiana surda Schwartz, 1965: 41 Type locality: For ȇt des Pins, Department du Sud-Est, Haiti Holotype: MCZ 77040 Distribution: Haiti and República Dominicana: eastern Massif de la Selle Celestus haetianus surdus ?

Celestus hewardi Gray, 1845 Celestus hewardi Gray, 1845: 118 Type locality: Jamaica Syntypes: BMNH 1946.12.3.88-90 Synonyms: Celestus impressus Cope, 1868 Type locality: Jamaica Lecotype: ANSP 9225, designated by Schwartz, 1964 Diploglossus variegatus Peters, 1874 Type locality: Jamaica Holotype: ZMB 8029 Distribution: interior Jamaica: Westmoreland, St. James, Trelawy, Manchester, St. Ann, St. Thomas parishes

Celestus hylaius Savage & Lips, 1993

Celestus hylaius Savage & Lips, 1993: 827 Type locality: La Selva Biological Station, Canton de Sarapiquí, Heredia, Costa Rica Holotype: LACM 138541 Distribution: northeastern Costa Rica

Celestus hylaius Timothy D. Paine

Celestus ingridae (Werler & Campbell, 2004) Diploglossus ingridae Werler & Campbell, 2004: 328 Type locality: near village of Santa Martha (18°20’14’’ N, 94°53’25’’ W) on the western slope of Volcán Santa Martha, Sierra de Los Tuxtlas, Veracruz, Mexico Holotype: UTA R-52521 Distribution: Mexico: Veracruz State: Sierra de Los Tuxtlas Last seen 1979

Celestus laf Lotzkat, Hertz and Köhler, 2016

Celestus laf Lotzkat, Hertz and Köhler, 2016: 965 Type locality: grounds of the Lost and Found ecohostel (8.67462°N, 82.21958°W), Reserva Forestal La Fortuna, Chiriquí, Panama Holotype: SMF 90177 Distribution: Chiriquí, Panama

Celestus laf Sebastian Lotzkat

Celestus legnotus (Campbell & Camarillo, 1994) Diploglossus legnotus Campbell & Camarillo, 1994: 195 Type locality: near Tepango de Rodríguez (ca. 20° 00' N, 97° 49' W), Sierra Madre Oriental, Puebla, México. Holotype: ENCB 16096 Distribution: Mexico: Sierra Madre Oriental: northern Puebla, west-central Veracruz, Mexico

Celestus macrotus Thomas & Hedges, 1989

Celestus macrotus Thomas & Hedges, 1989: 886 Type locality: ca. 15 km W (by road) Gros Cheval, northeastern slopes of Pic la Selle, Département de l'Ouest, Haiti. Holotype: USNM 286917 Distribution: Haiti: Massif de la Selle; República Dominicana: Sierra de Baoruco Celestus macrotus Carlos De Soto M

Celestus marcanoi (Schwartz & Inchaústegui, 1976) Diploglossus marcanoi Schwartz & Inchaústegui, 1976: 242 Type locality: Valle de Bao, Cordillera Central (road to Pico Duarte), Santiago Province, República Dominicana. Holotype: USNM 197299 Distribution: República Dominicana: Cordillera Central, Santiago Province Last seen 1984

Celestus microblepharis (Underwood, 1959) Diploglossus microblepharis Underwood, 1959: 2 Type locality: Boscobel, St. Mary Parish, Jamaica Holotype: MCZ 55764 Distribution: Jamaica: St. Mary Parish Last seen 1952

Celestus occiduus (Shaw, 1802) Lacerta occidua Shaw, 1802: 288 Type locality: Jamaica Holotype: BMNH XV.1 18a Synonyms: Scincus gallivasp Daudin, 1802 Type locality: Jamaica Holotype: MNHNP 0.1227 Diploglossus shawii Duméril & Bibron, 1839 Type locality: Jamaica Holotype: MNHNP 0.1227 Diploglossus cliftii Duméril & Bibron,1839 Type locality: Jamaica Holotype: RCSOM 1180 Macrogongylus brauni Werner, 1901 Type locality: Jamaica Holotype: not located Distribution: Jamaica, now probably extinct. Last seen 1840

Celestus orobius Savage & Lips, 1993

Celestus orobius Savage & Lips, 1993: 823 Type locality: near Hortensia, Palma and Fortuna on the Carretera Interamericana, Canton Pérez-Zeledón, Cordillera de Talamanca, San José, Costa Rica Holotype: LACM 138540 Distribution: Costa Rica: Cordillera de Talamanca Celestus orobius T Castoe

Celestus rozellae Smith, 1942

Celestus rozellae Smith, 1942: 372 Type locality: vicinity of Palenque, Chiapas, Mexico Holotype: USNM 113526 Distribution: Mexico: eastern Chiapas, southern Veracruz, eastern Oaxaca, eastern Tabacso States; northern Guatemala; central Belize Celestus rozellae Humberto Bahena B Celestus sepsoides (Gray, 1852) Sauresia sepsoides Gray, 1852: 282 Type locality: Santo Domingo, República Dominicana Holotype: BMNH 1946.8.29.29 Synonym: Embryopus habichii Weinland,1863 Type locality: Jérémie, Département de la Grand'Anse, Haiti Holotype: ZMB 1310 Distribution: Haiti: Tibubon Peninsula, Massif de la Hotte, Massif de la Selle, Ile de la Gon ȃve, Ile Grande Cayemite; República Dominicana: Sierra de Baoruco, Cordillera Central, Cordillera Celestus sepsoides Carlos De Soto M Septentrional

Celestus stenurus (Cope, 1862) Celestus stenurus stenurus (Cope, 1862) Diploglossus stenurus Cope, 1862: 188 Type locality: near Jérémie, Département de la Grand'Anse, Haiti Holotype: MCZ 3612 Distribution: Haiti: Tiburon Peninsula; Île-à-Vache

Celestus stenurus alloeides (Schwartz, 1964) Diploglossus stenurus alloeides Schwartz, 1964 18 Type locality: 6 km E Sánchez, Samaná Province, República Dominicana Holotype: MCZ 77152 Distribution: República Dominicana: Peninsula de Samaná

Celestus stenurus rugosus Cope, 1879

Celestus rugosus Cope, 1879: 272 Type locality: Puerto Plata, Puerto Plata Province, República Dominicana Holotype: USNM 10260 Distribution: northern & central República Dominicana: Valle de , Cordillera Central, western Celestus stenurus rugosus Alejandro Sanchez Sierra de Neiba, Cayos Siete Hermanos; Haiti: northern coast

Celestus stenurus weinlandi Cope, 1868 Celestus weinlandi Cope, 1868: 125 Type locality: Ile Gon ȃve, Haiti Holotype: USNM 12145 Distribution: Haiti: from Gonaïves and Ennery southeast along Golfe de la Gonâve to Plaine de Cul de Sac, Montagnes du Trou-d’Eau, northern slope of Massif de la Selle, Île à Cabrit; República Dominicana:Valle de Neiba, Llanos de Azua

Celestus warreni (Schwartz, 1970)

Diploglossus warreni Schwartz, 1970:780 Type locality: Palmiste, Île de la Tortue, Département du Nord-Ouest, Haiti Holotype: AMNH 103214 Distribution: northwestern Haiti, Départements du Nord and Dept. de l’Artibonite, and Île de la Tortue

Celestus warreni R Sajdak

Diploglossus Wiegmann, 1834 Wiegmann AFA. 1834. Herpetologica Mexicana , seu descriptio amphibiorum Novae Hispaniae, quae itineribus comitis de Sack, Ferdinandi Deppe et Chr. Guil. Schiede in Museum Zoologicum Berolinense pervenerunt. Pars prima, saurorum species amplectens. Adiecto systematis saurorum prodromo, additsque multis in hunc amphibiorum ordinem observationibus. Berlin: C.G. Lüderitz. vi + 54 pp. + Plates I-X. Type species: Tiliqua fasciatus Gray, 1831

Diploglossus bilobatus (O'Shaughnessy, 1874)

Celestus bilobatus O'Shaughnessy, 1874: 257 Type locality: Costa Rica Holotype: BMNH 1946.8.7.91 Distribution: lowland Costa Rica; northeastern Panama; southeastern Nicaragua

Diploglossus bilobatus Donald Filipiak

Diploglossus bivittatus Boulenger, 1895

Diploglossus bivittatus Boulenger, 1895: 732 Type locality: Hacienda Rosa de Jericho [= Santa Rosa de Jericó], Matagalpa, Nicaragua Holotype: BMNH 1946.8.29.37 Distribution: southeastern Guatemala, western El Salvador (Cordillera de Apaneca), southwestern Honduras (Sierra Madre), northwestern Nicaragua Celestus bivittatus LD Wilson

Diploglossus delasagra (Cocteau, 1838)

Scincus delasagra Cocteau, 1838: 110 Type locality: Cuba Syntypes: MNHNP 0.2856, 0.2858-59, RMNH 3626 Distribution: Cuba: Pinar del Río, La Habana, Artemisa, Mayabeque, Matanzas, Villa Clara, Diploglossus delasagra ? Cienfuegos, Sancti Spíritus, Ciego de Ávila, Camagüey, Granma, Santiago de Cuba, Holguín, Guantánamo provinces, Isla de la Juventud

Diploglossus fasciatus (Gray, 1831)

Tiliqua fasciatus Gray, 1831: 71 Type locality: unknown Syntypes: BMNH XV.126a- XV.126b Synonyms: Diploglossus houttuynii Cocteau, 1839 Type locality: Brazil Syntypes: MNHNP 0.5208, 0. 0.261 Diploglossus resplendens Barbour, 1909 Type locality: juntion of the Rio Kaka and Rio Beni, eastern Bolivia Holotype: MCZ 7286 Distribution: Brazil: Acre, Minas Gerais, Diploglossus fasciatus ? Rio de Janeiro States; Boliva: Cochabamba, Beni, La Paz Depts.; southeastern Peru

Diploglossus garridoi Thomas & Hedges, 1998 Diploglossus garridoi Thomas & Hedges, 1998: 98 Type locality: at El Manguito, Granma Province, Cuba Holotype: MNHNCu 4420 Distribution: Cuba: Granma Province: Sierra Maestra

Diploglossus lessonae Peracca, 1890

Diploglossus lessonae Peracca, 1890: 1 Type locality: Brazil Holotype: MRSN R2122 Synonym: Diploglossus tenuifasciatus Parker, 1924 Type locality: Natal, Rio Grande do Norte, Brazil Holotype: BMNH 1946.8.29.39 Distribution: Rio Grande do Norte, Paraiba, Pernambuco, Ceará States

Diploglossus lessonae Bryan Suson

Diploglossus microcephalus (Hallowell, 1856) Eurepis microcephalus Hallowell, 1856: 155 Type locality: Mexico Holotype: ANSP 9531 Distribution: Mexico: unknown locale

Diploglossus microlepis (Gray, 1831) Tiliqua microlepis Gray, 1831: 71 Type locality: unknown Holotype: BMNH XV.117.a Distribution: unknown Last seen 1831

Diploglossus millepunctatus O'Shaughnessy , 1874

Diploglossus millepunctatus O'Shaughnessy , 1874: 301 Type locality: Northwest coast of South America Holotype: CAS 62582 Synonym: Celestus hancocki Slevin, 1928 Type locality: Malpelo Island, Colombia Holotype: CAS 62582 Distribution: Colombia: Valle del Cauca Prov., Malpelo Island Diploglossus millepunctatus Oliver Kuepper

Diploglossus monotropis (Kuhl, 1820)

Scincus monotropis Kuhl, 1820: 128 Type locality: unknown Holotype: unknown Synonym: Camelia jamaicensis Gray, 1839 Type locality: unknown Holotype: BMNH 1946.8.7.90 Distribution: southeastern Nicaragua; eastern Costa Rica; Panama; western Colombia; western Ecuador

Diploglossus monotropis T Hibbitts

Diploglossus montanus Schmidt, 1933

Diploglossus montanus Schmidt, 1933 Type locality: Sierra de Merendon, west of San Pedro, Honduras Holotype: FMNH 5066 Distribution: Sierra de Merendón: northwestern Honduras, eastern Guatemala

Celestus montanus Silviu Petrovan

Diploglossus montisilvestris Myers, 1973 Diploglossus montisilvestris Myers, 1973: 3 Type locality: southeastern slope of Cerro Pirre, Darien, Panama Holotype: KU 1113660 Distribution: eastern Panama: Serranía de Pirre

Diploglossus montisserrati Underwood, 1964

Diploglossus montisserrati Underwood, 1964: 3 Type locality: Woodlands Spring, Montserrat Holotype: MCZ 76924 Distribution: Lesser Antilles: Montserrat

Diploglossus montisserrati A Ogrodwczyk

Diploglossus nigropunctatus Barbour & Shreve, 1937 Diploglossus delasagra nigropunctatus Barbour & Shreve, 1937: 378 ` Type locality: mountains north of Imias, Guantánamo Prov., Cuba Holotype: MCZ 42504 Distribution: Cuba: Holguín, Guantánamo Provinces

Diploglossus owenii Duméril & Bibron, 1839 Diploglossus owenii Duméril & Bibron, 1839: 594 Type locality: Guatemala Holotype: RCSOM, not located Distribution: Guatemala

Diploglossus pleii Duméril & Bibron, 1839

Diploglossus pleii Duméril & Bibron, 1839: 605 Type locality: Puerto Rico (Martinique, in error). Holotype: MNHNP 0.2860 Distribution: interior and northern Puerto Rico: Cordillera Jaicoa, Sierra de Panduras

Diploglossus pleii Alfredo D.Colón A.

Diploglossus scansorius (McCranie & Wilson, 1996) Celestus scansorius McCranie & Wilson, 1996: 260 Type locality: 2.5 airline km NNE La Fortuna (15° 25’N, 87°19’W), Cordillera Nombre de Dios, Yoro, Honduras Holotype: USNM 335049 Distribution: Honduras: Cordillera Nombre de Dios

Ophiodes Wagler, 1828 Wagler, J. 1828. Auszüge aus seinem Systema Amphibiorum. Isis von Oken 21: 740-744 Type species: Pygopus striatus Spix, 1824

Ophiodes enso Entiasupe-Neto, Marquez-Quintela, Regnet, Teixeira, Silveira & Loebmann, 2017 Ophiodes enso Entiasupe-Neto, Marquez-Quintela, Regnet, Teixeira, Silveira & Loebmann, 2017: 516 Type locality: Laranjal Beach (31.76668°S 52.21668°W), Patos Lagoon estuary, of Pelotas, state of Rio Grande do Sul, Brazil. Holotype: CHFURG 3589 Distribution: Brazil: Rio Grande do Sul State: Lagoa dos Patos estuary, and Ilha dos Marinheiros (an islet of Rio Grande)

Ophiodes fragilis (Raddi, 1826)

Seps fragilis Raddi, 1826: 39 Type locality: Río de Janeiro, Brasil Holotype: not located Synonym: Ophiodes yacupoi Gallardo, 1966 Type locality: Yerbal San Martín, Puerto Libertad, Misiones, Argentina Holotype: MACN 12721 Distribution: Brazil: coastal southern Bahia , Ophiodes striatus Pedro H Bernado Minas Gerais, Mato Grosso do Sul States; Argentina: northeastern Misiones Prov.; Paraguay

Ophiodes intermedius Boulenger, 1894

Ophiodes intermedius Boulenger, 1894: 343 Type locality: near Asunción, Paraguay Syntypes: BMNH 1946.8.31.1-4 Distribution: Bolivia: Beni, Cochabamba, Santa Cruz Depts.; Paraguay; Argentina: Chubut, Entre Ríos, Formosa Provinces; Uruguay

Ophiodes intermedius Konrad Mebert

Ophiodes luciae Cacciali & Scott, 2015 Ophiodes luciae Cacciali & Scott, 2015: 43 Type locality: Departamento Presidente Hayes, 35 km NE and across the Paraguay River from Antequera (23°53’18”S, 57°19’23”W) , (Departamento San Pedro), Paraguay Holotype: MNHNP 0.9685 Distribution: Paraguay, Dept. Presidente Hayes Last see 1985

Ophiodes striatus (Spix, 1824)

Pygopus striatus Spix, 1824: 25 Type locality: Rio de Janeiro, Brazil. Lectotype: ZSM 593/0 Synonyms: Pygopus cariococca Spix, 1824 Type locality: Cariococca, Corcovado Mountain, Rio de Janeiro, Brazil Holotype: ZSM 590/0, Pygopus gronovii Wagler, 1828 Type locality: Brazil Holotype: not located Ophiodes grilli Boulenger, 1913 Type locality: Curityba, Parana, Brazil Syntypes: BMNH 1913.9.10.10-11 Distribution: Brazil: (Rio Grande do Sul, Minas Gerais, Espírito Santo, Ceará, Mato Grosso, Goiás, Bahia States; southern Paraguay; Uruguay Ophiodes striatus Paula H Valdujo Ophiodes vertebralis Bocourt, 1881

Ophiodes vertebralis Bocourt, 1881: 459 Type localities: southern Brazil and Uruguay Syntypes: MNHNP 0.5558-9 (Uruguay), 0.6159, 1999.8065 (Brazil) Distribution: Brazil: Rio Grande do Sul State; Argentina: Santiago del Estero Prov.; Uruguay Ophiodes vertebralis Doumecq Milieu

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TAXON INDEX

A adercus (see Celestus ) aenetergum (see Celestus costatus ) agasepsoides (see Celestus ) alloeides (see Celestus stenurus ) anelpistus (see Celestus ) aporus (see Celestus curtissi ) atitlanensis (see Celestus )

B badius (see Celestus ) barbouri (see Celestus ) bilobatus (see Diploglossus ) bivittatus (see Celestus ) brauni (see Celestus occiduus )

C cariococca (see Ophiodes striatus ) carraui (see Celestus ) chalcorhabdus (see Celestus costatus ) chalybaeus (see Celestus enneagrammus ) cliftii (see Celestus occiduus ) costatus (see Celestus ) crusculus (see Celestus ) cundalli (see Celestus crusculus ) curtissi (see Celestus ) cyanochloris (see Celestus )

D darlingtoni (see Celestus ) delasagra (see Diploglossus ) diastatus (see Celestus curtissi ) duquesneyi (see Celestus ) E emys (see Celestus costatus ) enneagrammus (see Celestus ) enso (see Ophiodes )

F fasciatus (see Diploglossus ) fowleri (see Celestus ) fragilis (see Ophiodes )

G gallivasp (see Celestus occiduus ) garridoi (see Diploglossus ) grilli (see Ophiodes striatus ) gronovii (see Ophiodes striatus ) H habichii (see Celestus sepsoides ) haetianus (see Celestus ) hancocki (see Diploglossus millepunctatus ) hewardi (see Celestus ) houttuynii (see Diploglossus fasciatus ) hylaius (see Celestus ) hylonomus (see Celestus curtissi )

I impressus (see Celestus hewardi ) ingridae (see Celestus ) intermedius (see Ophiodes )

J jamaicensis (see Diploglossus monotropis )

L laf (see Celestus ) legnotus (see Celestus ) leionotus (see Celestus costatus ) lessonae (see Diploglossus ) luciae (see Ophiodes )

M macrotus (see Celestus ) maculatus (see Celestus crusculus ) marcanoi (see Celestus ) melanchrous (see Celestus costatus ) microblepharis (see Celestus ) microcephalus (see Diploglossus ) microlepis (see Diploglossus ) millepunctatus (see Diploglossus ) molesworthi (see Celestus crusculus ) monotropis (see Diploglossus ) montanus (see Celestus ) montisilvestris (see Diploglossus ) montisserrati (see Diploglossus ) mylicus (see Celestus haetianus )

N neiba (see Celestus costatus ) nesobous (see Celestus costatus ) nigropunctatus (see Diploglossus ) nuchalis (see Celestus costatus )

O occiduus (see Celestus ) ohlendorffi (see Celestus costatus ) oreistes (see Celestus costatus ) orobius (see Celestus ) owenii (see Diploglossus )

P phoxinus (see Celestus costatus ) pleii (see Diploglossus ) psychonothes (see Celestus costatus )

R resplendens (see Diploglossus fasciatus ) rozellae (see Celestus ) rugosus (see Celestus stenurus ) S saonae (see Celestus costatus ) scansorius (see Celestus ) sepsoides (see Celestus ) shawii (see Celestus occiduus ) steindachneri (see Celestus enneagrammus ) stenurus (see Celestus ) striatus (see Ophiodes ) surdus (see Celestus haetianus )

T tenuifasciatus (see Diploglossus lessonae )

V variegatus (see Celestus haetianus ) vertebralis (see Ophiodes )

W warreni (see Celestus ) weinlandi (see Celestus stenurus )

Y yacupoi (see Ophiodes fragilis )