Debris the Science and Changing Perceptions of Dead Trees by Alexander Evans and Robert Perschel

Total Page:16

File Type:pdf, Size:1020Kb

Debris the Science and Changing Perceptions of Dead Trees by Alexander Evans and Robert Perschel An Appreciation of Debris The Science and Changing Perceptions of Dead Trees By Alexander Evans and Robert Perschel oarse woody debris. To most people, two Coarse woody debris (CWD) is wood in the forest that’s no of the three words in that phrase have longer part of a living tree. Rather than being dead, this wood negative connotations. Debris is the worst is an important source of life in the forest. Foresters who value offender, suggesting random junk strewn it for its ecological functions have been searching for new terms about. When the debris is coarse, it’s even for CWD, and you’ll hear it referred to as biomass, dead wood, worse – it’s unrefined, it’s big. coarse woody material, and even retained organic matter. VIRGINIA BARLOW 44 Northern Woodlands / Winter 2008 Harry Dwyer, a forester in Fayette, Maine, likens the lan- as a seed bed for trees and plants, a mulch layer, and a slow- guage predicament to that of another formerly maligned mate- release fertilizer. Decomposing fungi depend on dead wood for rial: garden waste. He said, “You could look at a pile of rotting nutrients and moisture, and in turn many trees rely on mutu- vegetables as garbage, or you could note its value and call it alistic relationships with mycorrhizal fungi. Nitrogen fixation compost.” Digging into a rotten log in his woodlot, Dwyer in CWM by microbes (both anaerobic and microaerophyllic displays some of the dark, wet material in his hand. “It doesn’t bacteria) provides this important element in both terrestrial matter what you call this – coarse woody debris, coarse woody and aquatic ecosystems. material, or wildlife habitat – it still functions as a critical com- In general, northern forests have less dead wood in them ponent of forest health. But the words we use do make a differ- now than they did before Europeans arrived. Old growth for- ence, particularly now that everyone is talking about removing ests tend to have more logs, snags, and other dead wood than biomass from the forest for fuel.” managed or younger forests. While dead wood accumulates in Let’s call it coarse woody material or CWM. forests over time, it doesn’t last forever. On average, snags typi- When trees die on their own, they remain part of the forest cally stand about 10 years, while fallen logs may last twice that community – as standing dead trees (snags), stumps, down logs, long. Timber harvests tend to temporarily increase the amount or fallen branches. This CWM is gradually broken down into of dead wood because branches and the unmerchantable parts smaller and smaller parts by fungi, insects, and microorgan- of felled trees are left on the ground, and just the more valuable isms, until it’s fully incorporated into the forest soil. Scientific parts of a tree are removed. research is gradually revealing the important role dead wood While harvests may produce a large amount of dead wood plays in forest ecosystems, from wildlife habitat to carbon stor- in the short term, they can also result in fewer trees left to age, though anyone who has ever disturbed an old wood pile die and become dead wood in the future. Bill Keeton, a forest knows dead wood is alive with animals and plants. Moving an ecologist at the University of Vermont, advocates for a balance old log might reveal a black rat snake using it for a nest site, between removing the maximum volume of wood products and or searching the dead wood for dinner. Above the forest floor, retaining some biomass to maintain the structural complexity snags are high-demand housing for wildlife. When a pile- of forest. To increase dead wood and other old-growth charac- ated woodpecker leaves a cavity in a dead tree, a whole host of animals – from other birds to squirrels to wasps to northern Inset: While tops left in the woods after a logging job might look long-eared bats – might move in. Logs that fall in streams and messy, they serve an important ecological role. These tops will pro- rivers form critical aquatic habitat by ponding water and storing tect new tree seedlings from browsing deer before eventually rot- sediments. When water cascades over a fallen log, it can dig out ting and becoming fertilizer. Bottom: On the other hand, tree tops a small pool that will shelter young trout. can be chipped and converted into heat and electricity, and it’s That’s not all. Coarse woody material serves as a carbon advantageous environmentally to burn local renewable resources JIM BLOCK sink (as much as six percent of the carbon stored in rather than fossil fuels from afar. The key, as always, is finding a Northeastern forests is in dead wood); it also serves balance between human needs and the overall health of a forest ecosystem. Northern Woodlands / Winter 2008 45 teristics in younger forests, Keeton favors harvest prescriptions crates for shipment over here but one was left behind on the that include retaining all trees greater than 24 inches in diam- dock. While Cotta’s perspective did not survive forestry’s initial eter, releasing crowns to accelerate growth of larger trees, and translation to American soil, it has gradually returned to the girdling (or felling) diseased, dying, or poorly formed trees to profession in this country, aided by luminaries such as Aldo create snags or downed logs. This ecological forestry approach Leopold, Rachel Carson, and E. O. Wilson. What was once con- values dead wood left in the forest along with the timber taken sidered debris is now valued for the carbon it stores, the wildlife out in a harvest. habitat it provides, and the nutrients it holds. At the same time, though, society increasingly has its eyes on European roots of forestry wood that has little economic value but has great value as a fuel Forestry in the United States was originally built on ideas source. Low-value trees, slash, and tops are being heralded as imported from Europe in the early 1900s. When early foresters “woody biomass,” a potentially important part of the solution to like Gifford Pinchot brought the foundation of scientific forestry our dependence on fossil fuels. over from France and Germany, they also brought an emphasis Woody biomass has long been a useful, if under-utilized, on the sustained yield of timber. Timber was the most important byproduct of forest management activities. Foresters and log- forest product in their minds, so it drove most decisions about gers have prided themselves on their ability to put much of a forest management. Consequently, dead wood in the forest came felled tree to use. But interest in removing woody biomass has to be considered “debris.” A dead tree on the ground was consid- recently accelerated because of the rising cost of other fuels, ered a waste of resources or a potential fire hazard. concerns about carbon emissions from fossil fuels, and risks in Another theme of early European forestry, however, included western states of catastrophic wildfires. For example, the U.S. a more holistic perspective. Forestry textbooks like those written Department of Energy has set a goal to increase biofuel use in by Heinrich Cotta in the first part of the nineteenth century, are the U.S. 25 times by 2030. While biomass can be produced agri- distinctly different from the kind of forestry brought to America culturally, much of the biomass to fuel this increase in renew- by Pinchot. Early pioneers like Cotta talked about using nature able energy would come from forests. In fact, a U.S. Department as the guide and practicing forestry that preserves the soil’s fertility. WATTS PAT It’s as if European forestry was packed up in Snags and downed trees create high-demand housing for wildlife, including this northern flicker (right) and this chipmunk (below). DRAKE FLEEGE 46 Northern Woodlands / Winter 2008 of Energy report estimates that U.S. forests could yield 368 mil- lished guidelines that suggest how and how much woody biomass lion dry tons of useable biomass per year. can be responsibly removed from forests. A recent experimental So the forest-policy discussion of woody debris has become harvest on the Superior National Forest, in Minnesota, showed increasingly complex. On the one hand, optimizing the utiliza- that when harvesters followed state biomass guidelines, there tion of wood – a naturally renewable resource – can help replace were adequate snags and coarse woody material left behind. A our reliance on foreign oil and can be a substitute for oil and report on impacts of biomass harvesting from Massachusetts has other fossil fuels, thus helping to combat global climate change. good news about calcium, a particularly important soil nutrient. On the other hand, as the allure of using woody biomass for When only 50 percent of the biomass is removed, the current energy increases, the removal of additional biomass raises ques- level of calcium would be replenished in 71 years. tions about how much wood can be taken from a forest before A good general guideline for northeastern forests comes from the forest suffers negative consequences. the Society for the Protection of New Hampshire Forests’ manual When timber harvests remove whole trees for energy pro- Good Forestry in the Granite State: “Manage for coarse woody duction, the risk of nutrient loss increases because more leaves material by retaining material that currently exists and allow- and needles are removed from the forests. Foliage and small, ing its accumulation where it is currently missing.” The Forest fine branches contain much higher concentrations of nutrients Stewardship Council’s standard for green-certified forests in the than stem wood.
Recommended publications
  • AN ABSTRACT of the THESIS of Douglas William Havlina for The
    AN ABSTRACT OF THE THESIS OF Douglas William Havlina for the degree of Master of Science in Rangeland Resources presented on January 27, 1995 Title: Fire Effects on Vegetation Diversity, Structure, and Successional Dynamics in Shrub-Steppe and Mixed Conifer Environments of the Hells Canyon, Idaho. Abstract approved: The effects of fire severity on post-fire succession and shrub demography were quantified in shrub-steppe grasslands and subalpine forests in the Hells Canyon of the Payette National Forest, Idaho. Following the 1988 Eagle Bar Fire, species frequency, community diversity, fire adaptations, and stand ages were quantified at 12 plots in burned and unburned forests. Pre-fire composition was dominated by Pseudotsuga menziesii, Pinus contorta, Abies grandis, and Abies lasiocarpa in the overstory. Pre- fire basal area ranged from 41 to 80 m 2 ha-1 . Pre-fire stand ages ranged from 71 years in Abies lasiocarpa forests to > 200 years in Pseudotsuga menziesii stands. Fire scenarios which were sampled consisted of: (1) complete stand-replacement fires; (2) partial stand- replacement fires; and (3) unburned forests (control). Partial stand-replacement forests were characterized by significantly higher mean species diversity and richness (H'=3.16, S=36) than complete stand-replacement (H'=2.78, S=27) or unburned forests (H=2.26, S=15). Vegetation preferentially established in burned areas along a gradient of fire severity according to adapted fire guilds. Single-storied forests dominated by Pinus contorta and Abies lasiocarpa typified stand-replacement fire scenarios, with mean ages ranging from 99 to 159 years corresponding to fire return intervals common in this type.
    [Show full text]
  • Influence of Coarse Woody Debris on Seedlings and Saplings in A
    INFLUENCE OF COARSE WOODY DEBRIS ON SEEDLINGS AND SAPLINGS IN A PINUS PALUSTRIS WOODLAND by ALEXANDRA LOGAN JUSTIN L. HART, COMMITTEE CHAIR MATTHEW C. LAFEVOR ARVIND A.R. BHUTA A THESIS Submitted in partial fulfillment of the requirements for the degree of Master of Science in the Department of Geography in the Graduate School of The University of Alabama TUSCALOOSA, ALABAMA 2020 Copyright Alexandra Logan 2020 ALL RIGHTS RESERVED 2 ABSTRACT Coarse woody debris (CWD) has beneficial effects on plant growth and establishment. Longleaf pine (Pinus palustris Mill.) stands support relatively low amounts of CWD — 2 to 30 m3 ha-1. In April 2011, an EF3 tornado passed through the Oakmulgee Ranger District of the Talladega National Forest in the Fall Line Hills of Alabama. This disturbance resulted in the large addition of CWD to a longleaf pine woodland, and a rare opportunity to analyze how CWD can influence a managed, pine woodland. The goal of this study was to examine the effect of CWD on woody plant richness, density, and growth rate (quantified by height) in a longleaf pine woodland that experienced a catastrophic wind disturbance. A total of three 1 m2 quadrats were established against either side of a piece of CWD (> 3 m in length and ≥ 10 cm in diameter). Another quadrat was established at least 3 m away from the focal CWD piece. For each plot, the presence and height of every woody plant (< 5 cm dbh) were recorded. Sapling density, oak and hickory density, and organic matter were all found to be significantly higher in quadrats adjacent to CWD than away (all p < 0.05).
    [Show full text]
  • Environmental Limits Page 1
    POST Report 370 January 2011 Living with Environmental Limits Page 1 Summary Human well-being is dependent upon assessment approaches. However, where renewable natural resources. Agricultural there is a risk of thresholds being systems, for example, depend upon plant breached and potentially irreversible productivity, soil, the water cycle, the impacts occurring, additional policy nitrogen, sulphur and phosphorus nutrient safeguards to maintain natural resource cycles and a stable climate. Renewable systems within environmental limits are natural resources can be subject to required. biological and physical thresholds beyond which irreversible changes in benefit Managing ecosystems to maximise one provision may occur. These are difficult to particular benefit, such as food provision, define and many are likely to be identified can result in declines in other benefits. only once crossed. An environmental limit The evidence base is not yet sufficient to is usually interpreted as the point or range determine the most effective ways to of conditions beyond which there is a maintain benefit provision within significant risk of thresholds being environmental limits, but a range of policy exceeded and unacceptable changes responses are seeking to optimise multiple occurring.1 benefit provision, including: Biodiversity loss, climate change and a agri-environment schemes range of other pressures are affecting generic measures to enhance renewable natural resources. If biodiversity, which may increase governments do not effectively monitor the the capacity of natural resource use and degradation of natural resource systems to adapt to environmental systems in national account frameworks, change the probability of costs arising from the use of ecological processes to exploiting natural resources beyond increase overall natural system environmental limits is not taken into resilience to address problems account.
    [Show full text]
  • Human Involvement in Food Webs*
    EG35CH01-Strong ARI 13 September 2010 10:20 Human Involvement in Food Webs∗ Donald R. Strong1 and Kenneth T. Frank2 1Department of Evolution and Ecology, University of California, Davis, California 95616; email: [email protected] 2Bedford Institute of Oceanography, Ocean Sciences Division, Dartmouth, Nova Scotia B2Y 4A2, Canada; email: [email protected] Annu. Rev. Environ. Resour. 2010. 35:1–23 Key Words First published online as a Review in Advance on bottom-up, fisheries, intraguild predation, mesopredator, top-down, July 1, 2010 trophic cascade The Annual Review of Environment and Resources is online at environ.annualreviews.org Abstract This article’s doi: Human involvement in food webs has been profound, bringing about 10.1146/annurev-environ-031809-133103 enormous and disproportionate losses of large apex predators on land Copyright c 2010 by Annual Reviews. and in water. The losses have modified or even eliminated concatena- All rights reserved tions of indirect interactions propagating from predators to herbivores 1543-5938/10/1121-0001$20.00 to plants, inter alia. Food webs are a synthesis of bottom-up energy and ∗This article was co-authored by an employee of nutrient flow from plant producers to consumers and top-down regula- a British Commonwealth government as part of tion of producers by consumers. The trophic cascade is the simplest top- his official duties and is therefore subject to down interaction and accounts for a great deal of what is known about Crown Copyright. food webs. In three-link cascades, predators suppress herbivores, re- leasing plants. In longer cascades, predators can suppress smaller meso- by UNIVERSITY OF IDAHO LIBRARY on 03/21/11.
    [Show full text]
  • Bibliography
    Bibliography Abella, S. R. 2010. Disturbance and plant succession in the Mojave and Sonoran Deserts of the American Southwest. International Journal of Environmental Research and Public Health 7:1248—1284. Abella, S. R., D. J. Craig, L. P. Chiquoine, K. A. Prengaman, S. M. Schmid, and T. M. Embrey. 2011. Relationships of native desert plants with red brome (Bromus rubens): Toward identifying invasion-reducing species. Invasive Plant Science and Management 4:115—124. Abella, S. R., N. A. Fisichelli, S. M. Schmid, T. M. Embrey, D. L. Hughson, and J. Cipra. 2015. Status and management of non-native plant invasion in three of the largest national parks in the United States. Nature Conservation 10:71—94. Available: https://doi.org/10.3897/natureconservation.10.4407 Abella, S. R., A. A. Suazo, C. M. Norman, and A. C. Newton. 2013. Treatment alternatives and timing affect seeds of African mustard (Brassica tournefortii), an invasive forb in American Southwest arid lands. Invasive Plant Science and Management 6:559—567. Available: https://doi.org/10.1614/IPSM-D-13-00022.1 Abrahamson, I. 2014. Arctostaphylos manzanita. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory, Fire Effects Information System (Online). plants/shrub/arcman/all.html Ackerman, T. L. 1979. Germination and survival of perennial plant species in the Mojave Desert. The Southwestern Naturalist 24:399—408. Adams, A. W. 1975. A brief history of juniper and shrub populations in southern Oregon. Report No. 6. Oregon State Wildlife Commission, Corvallis, OR. Adams, L. 1962. Planting depths for seeds of three species of Ceanothus.
    [Show full text]
  • The Ecological Role of Coarse Woody Debris an Overview of The
    WORKING PAPER 30 The Ecological Role of Coarse Woody Debris An Overview of the Ecological Importance of CWD in BC Forests 1997 Ministry of Forests Research Program The Ecological Role of Coarse Woody Debris An Overview of the Ecological Importance of CWD in BC Forests Victoria Stevens Ministry of Forests Research Program The use of trade, firm, or corporation names in this publication is for the information and convenience of the reader. Such use does not constitute an official endorsement or approval by the Government of British Columbia of any product or service to the exlusion of any others that may also be suitable. Contents of this report are presented for discussion purposes only. Citation Stevens, Victoria. 1997. The ecological role of coarse woody debris: an overview of the ecological importance of CWD in B.C. forests. Res. Br., B.C. Min. For., Victoria, B.C. Work. Pap. 30/1997. Compiled by Victoria Stevens for B.C. Ministry of Forests Research Branch 31 Bastion Square Victoria, B.C. V8W 3E7 Copies of this report may be obtained, depending upon supply, from: B.C. Ministry of Forests Forestry Division Services Branch Production Resources 595 Pandora Avenue, 1st Floor Victoria, B.C. V8W 3E7 © 1997 Province of British Columbia The contents of this report may not be cited in whole or in part without the approval of the Director of Research, B.C. Ministry of Forests, Victoria, B.C. ACKNOWLEDGEMENTS Plunging into a subject as complex as ecological roles of coarse woody debris is not an activity lending itself to solitude. This paper has been at least partially digested by a large number of readers and greatly improved by the suggestions of many.
    [Show full text]
  • Contrasting Distribution and Foraging Patterns of Herbivorous and Detritivorous Fshes Across Multiple Habitats in a Tropical Seascape
    Marine Biology (2019) 166:51 https://doi.org/10.1007/s00227-019-3498-0 ORIGINAL PAPER Contrasting distribution and foraging patterns of herbivorous and detritivorous fshes across multiple habitats in a tropical seascape M. Eggertsen1 · D. H. Chacin2 · C. Åkerlund1 · C. Halling1 · C. Berkström1,3 Received: 21 September 2018 / Accepted: 8 March 2019 / Published online: 16 March 2019 © The Author(s) 2019 Abstract Understanding drivers behind patterns of functionally important groups of fshes is crucial for successful management and conservation of tropical seascapes. Herbivorous fshes are the most prominent consumers of marine primary production which can have profound efects on reef resilience. We explored environmental variables afecting distribution and foraging patterns of herbivorous and detritivorous fsh assemblages (siganids, acanthurids and parrotfsh) across distinct shallow-water habitats (coral reefs, macroalgae beds and seagrass meadows) during September–November 2016 at Mafa Island, Tanzania (8°00′S, 39°41′E). We performed underwater visual census to quantify fsh assemblages, measured habitat features, deployed mac- roalgal assays and conducted inventories of grazing scars. Multi-dimensional scaling and mixed-efects linear models were used to evaluate diferences in fsh assemblages and environmental variables infuencing abundance and foraging patterns of fshes. Fish communities of focal functional groups difered among habitats. Abundance of herbivores and detritivores as well as relative browsing and scraping was highest on coral reefs compared to macroalgae and seagrass meadows. Adult fsh were more abundant on coral reefs while juveniles were abundant in macroalgal beds. Coral cover and crustose coralline algal cover had a positive efect on the abundance of fsh in coral reef areas, while macroalgal cover had a negative efect.
    [Show full text]
  • Chronic Over Browsing and Biodiversity Collapse
    Journal of the Torrey Botanical Society 138(2), 2011, pp. 220–224 Chronic over browsing and biodiversity collapse in a forest understory in Pennsylvania: Results from a 60 year-old deer exclusion plot Chandra Goetsch and Jennifer Wigg Department of Biological Sciences, University of Pittsburgh, A234 Langley Hall, Pittsburgh, PA 15260 Alejandro A. Royo and Todd Ristau USDA Forest Service Northern Research Station, Forestry Sciences Lab, P.O. Box 267, Irvine, PA 16329-0267 Walter P. Carson1 Department of Biological Sciences, University of Pittsburgh, A234 Langley Hall, Pittsburgh, PA 15260 GOETSCH, C., J. WIGG (Department of Biological Sciences, University of Pittsburgh, A234 Langley Hall, Pittsburgh, PA 15260), A. A. ROYO,T.RISTAU (USDA Forest Service Northern Research Station, Forestry Sciences Lab, P.O. Box 267, Irvine, PA 16329-0267), AND W. P. CARSON (Department of Biological Sciences, University of Pittsburgh, Pittsburgh, PA 15260). Chronic over browsing and biodiversity collapse in a forest understory in Pennsylvania: results from a 60-year-old deer exclusion plot. J Torrey Bot. Soc. 138: 220–224. 2011.—We evaluated the impact of chronic deer over browsing on the diversity and abundance of understory forbs and shrubs within a forest stand in the Allegheny High Plateau Region of Pennsylvania by comparing vegetation inside a 60-year-old exclosure to vegetation within an adjacent reference site. This is the oldest known exclosure in the Eastern Deciduous Forest. Browsing caused the formation of an extremely low diversity herbaceous understory dominated by a single fern species, caused the local extirpation of shrubs, and drove forbs to extremely low abundance (, 0.2% cover m22 vs.
    [Show full text]
  • Student Worksheet Provide an Example of an Herbivore, Carnivore
    Vocabulary and Concepts Review: Student Worksheet Provide an example of an herbivore, carnivore and omnivore that lives in the Great Plains of North America (can be living, or extinct): ____________________________________________________________________________________ What is hypsodonty? List 3 mammals that have hypsodont dentition and aspects of their diet that correspond with needing such specialized teeth. ____________________________________________________________________________________ What is the difference between a savanna and a prairie? Which is characteristic of the North American Great Plains? ____________________________________________________________________________________ Are shrubs and plants short in height more common in open grasslands or wooded forests? How might a diet of these short plants affect tooth wear in herbivores? _______________________________________________________________________________ Why might silica phytoliths be better preserved in the fossil record than blades of grass or leaves? _____________________________________________________________________________________ Provide an example (excluding what was learned about mammalian teeth) of how an organisms’ morphology reflects evolutionary pressures as a result of changes in its environment. _____________________________________________________________________________________ What is the difference between browsing and grazing (food type, eating habit)? _____________________________________________________________________________________ When
    [Show full text]
  • Restore Large Carnivores to Save Struggling Ecosystems
    Restore large carnivores to save struggling ecosystems What? It’s just a flesh wound. Steve Jurvetson We are losing our large carnivores. In ecosystems around the world, the decline of large predators such as lions, bears, dingoes, wolves, and otters is changing landscapes, from the tropics to the Arctic. Habitat loss, persecution by humans and loss of prey have combined to inflict great losses on these populations. In fact more than 75% of the 31 largest carnivore species are declining, and 17 species now occupy less than half their former ranges. Southeast Asia, southern and East Africa, and the Amazon are among areas in which multiple large carnivore species are declining. And with only a few exceptions, large carnivores have already been exterminated from much of the developed world, including areas of Western Europe, and the eastern United States. Top dogs keep ecosystems in order Many of these large carnivore species are endangered and some are at risk of extinction, either in specific regions or entirely. Ironically, they are vanishing just as we are learning about their important ecological effects, which is what led us to write a new paper in the journal Science to document their role. From a review of published reports, we singled out seven species that have been studied for their important ecological role and widespread effects, known as trophic cascades. These are the African lion, leopard, Eurasian lynx, cougar, gray wolf, sea otter and dingo. Based on field research, my Oregon State University co-author Robert Beschta and I documented the impact of cougars and wolves on the regeneration of forest tree stands and riverside vegetation in Yellowstone and other national parks in western North America.
    [Show full text]
  • Does Wolf Presence Reduce Moose Browsing Intensity in Young Forest Plantations?
    Utah State University DigitalCommons@USU Aspen Bibliography Aspen Research 1-30-2018 Does Wolf Presence Reduce Moose Browsing Intensity in Young Forest Plantations? Suzanne T. S. van Beeck Calkoen Swedish University of Agricultural Sciences Dries P. J. Kuijper Polish Academy of Sciences Håkan Sand Swedish University of Agricultural Sciences Navinder J. Singh Swedish University of Agricultural Sciences Sip E. van Wieren Wageningen University Joris P. G. M. Cromsigt Swedish University of Agricultural Sciences Follow this and additional works at: https://digitalcommons.usu.edu/aspen_bib Part of the Agriculture Commons, Ecology and Evolutionary Biology Commons, Forest Sciences Commons, Genetics and Genomics Commons, and the Plant Sciences Commons Recommended Citation van Beeck Calkoen, S. T., Kuijper, D. P., Sand, H. , Singh, N. J., van Wieren, S. E. and Cromsigt, J. P. (2018), Does wolf presence reduce moose browsing intensity in young forest plantations?. Ecography, 41: 1776-1787. doi:10.1111/ecog.03329 This Article is brought to you for free and open access by the Aspen Research at DigitalCommons@USU. It has been accepted for inclusion in Aspen Bibliography by an authorized administrator of DigitalCommons@USU. For more information, please contact [email protected]. doi: 10.1111/ecog.03329 41 1776– 1787 ECOGRAPHY Research Does wolf presence reduce moose browsing intensity in young forest plantations? Suzanne T. S. van Beeck Calkoen, Dries P. J. Kuijper, Håkan Sand, Navinder J. Singh, Sip E. van Wieren and Joris P. G. M. Cromsigt S. T. S. van Beeck Calkoen (http://orcid.org/0000-0002-2574-9056) ([email protected]), N. J. Singh and J.
    [Show full text]
  • Drivers and Impacts of Large Herbivore Foraging Behaviour in a Rewilded Landscape
    Drivers and impacts of large herbivore foraging behaviour in a rewilded landscape Thomas Dando 177355 Conservation Biology MRes 2017/2018 Supervised by Dr Christopher Sandom Abstract Rewilding is increasingly being proposed in conservation discourse, as a means to reverse present ecological declines. Despite its increasing prominence however, there is little known of its ecological outcomes. The historic role of large herbivores in shaping a wild landscape is a particularly important area of research in the context of modern rewilding in temperate Europe, but opportunities to deduce the extent of herbivore influence has been rare. The Knepp Wildland Project offers an opportunity to address this gap. The deliberate abandonment of agricultural land, has allowed self-willed habitat to develop, this has been complemented by the reintroduction of large herbivore species, present in near-natural grazing conditions. This project aimed to ascertain the drivers and impact of large herbivores forage selection and intensity within this rewilding project. A network of thirty-eight camera traps were active for twenty-five trap days in summer and winter to record herbivore movements, as well as seven 2km walked line transects, active over the same study periods. Five 1m2 vegetation quadrats were sampled within each camera trapped field to ascertain floral diversity, while LiDAR data was collected within 30m2 patches to record structural heterogeneity and vegetation height. Herbivore foraging was found to be non-uniform, with overlaps in forage selection between study species creating clear pressure points, conversely other areas are developing under minimal pressure. No sequential patterns occurred when considering when the fields were taken out of agriculture, indicating this is not a driver in forage selection.
    [Show full text]