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Working List of Invasive Vascular of Wyoming ─ III (Vernacular names from selected major works) Dec 2017

Compiled by R.L. Hartman and B.E. Nelson With assistance from R.D. Dorn, W. Fertig, B. Heidel

The following list contains 372 taxa introduced to Wyoming from outside ; included are invasives recognized by the Wyoming state government as noxious (; 28; although Ambrosa tomentosa is native). A number of these repesent escapes from cultivation and are limited to one or a few collections. Nomenclature is based on R.D. Dorn, 2001, Vascular Plants of Wyoming; where updated, Dorn’s synonyms are in square brackets [ ]. Other synonyms found in the list of sources below are not included. Likewise, family names and their circumscriptions follow Dorn; where defined differently by the Angiosperm Working Group IV (APG IV), clarification follows in parenthese. Dorn does not indicate the typical variety or subspecies unless a second infraspecific taxon is recognized. We have included the typical infraspecies where appropriate. The lack of hyphenation, word separation, or capitalization may not reflect the appearance of the vernacular names in the works cited. Sources for vernacular names: 1 Weed Science Society of America. 2010. Composite List of Weeds. 2 Kartesz, J.T., The Biota of North America Program (BONAP). 2015. Taxonomic Data Center. 3 P. Rice. 2000. Invaders Database System. Univ. of Montana. Release 14 Feb 2000 (not available for update). 4 Flora of the Great Plains Association. 1986. Flora of the Great Plains. Univ. Oklahoma Press. 5 C.L. Hitchcock & A. Cronquist. 2018. Flora of the Pacific Northwest. Univ. Washington Press. 6 H.A. Gleason & A. Cronquist. 1991. Manual of Vascular Plants of Northeastern United States and Adjacent Canada. Ed. 2. New York Botanical Garden. 7 Plants Database, Natural Resources Conservation Service, USDA.

Amaranthaceae Amaranthus albus L. tumble pigweed1, tumbleweed2,4,5,6, white pigweed6, prostrate pigweed7 Apiaceae (Umbelliferae) Anthriscus caucalis M. Bieb. bur chervil1,6,7, burr chervil2,3 Carum carvi L. common caraway1,3, caraway2,4,5,6,7 Conium maculatum L. poison hemlock1,2,3,4,5,6,7 Daucus carota L. wild carrot1,3,4,5,6, Queen Anne’s lace2,4,5,6,7 Falcaria vulgaris Bernh. sickleweed1,2,7 Pastinaca sativa L. wild parsnip1,2,3,4,5,7, common parsnip5, parsnip6 Torilis japonica DC. Japanese hedge parsley1,6, erect hedge parsley2,7, hedge parsley4 Asparagaceae Asparagus officinalis L. asparagus2,3,4,5,6, garden asparagas7 Muscari botryoides (L.) P. Miller Not in Dorn grape hyacinth1,4,6, common grape hyacinth2,3,7 (Compositae) Achillea ptarmica L. Not in Dorn sneezewort yarrow1, pearl yarrow2, sneezeweed6,7 Acroptilon repens (L.) DC. Russian knapweed1,3,4,5,6, hardheads2,7, Turkestan thistle5, Russian tarthistle5 [ repens L.] Ambrosia tomentosa Nutt. Native skeletonleaf bursage1,3,6, skeletonleaf burr ragweed2,7, perennial bursage4 Anthemis arvensis L. corn chamomile1,2,3,4,5,6,7, field chamomile5, field dogfennel5 Anthemis cotula L. mayweed chamomile1,3,5, stinking chamomile2,7, dog fennel4,6, stinking mayweed5 Arctium lappa L. great burdock1,3,4,5,6, greater burrdock2, greater burdock7 Arctium minus (Hill) Bernh. common burdock1,3,4,5,6, lesser burrdock2, lesser burdock7 Artemisia abrotanum L. southern wormwood1,2,3, southernwood4,6,7 Artemisia absinthium L. absinth wormwood1,3, oldman2, wormwood4,5, common wormwood6, absinthium5,6,7 Artemisia annua L. annual wormwood1,2,3,5,6, sweet sagewort4,7 Carduus acanthoides L. plumeless thistle1,3,4,6, spiny plumeless thistle2,7, acanthus thistle5 Carduus nutans L. musk thistle1,3,4,5,6, nodding plumeless thistle2,7, nodding thistle4 Carthamus tinctorius L. safflower2,3,4,5,6,7 L. cornflower1,3,4,5,6, garden cornflower2,7, bachelor’s button4,5,6 Centaurea diffusa Lam. diffuse knapweed1,3,4,7, white knapweed2, tumble knapweed5,6, bushy knapweed5 Centaurea iberica Trev. ex Spreng. Iberian starthistle1,3,4, Iberian knapweed2,7 Centaurea macrocephala Puschk. ex Willd. bighead knapweed1,7 globe knapweed2 Not in Dorn Centaurea ×moncktonii C.E. Britton Not in Dorn meadow knapweed7 Centaurea montana L. Not in Dorn perennial coneflower1,7, mountain starthistle2,5 Centaurea nigrescens Willd. Not in Dorn Vochin knapweed1, Tyrol knapweed2,7 short fringed knapweed5,6

1 Centaurea scabiosa L. greater knapweed2,7, scabious starthistle3, greater centaurea4, hardheads6 Centaurea solstitialis L. yellow starthistle1,2,3,4,5,6,7, St. Barnaby’s thistle5 Centaurea stoebe L. spotted knapweed1,2,3,4,5,6,7 ssp. micranthos (S.G. Gmelin ex Gugler) Hayek [Centaurea maculosa Lam.] Centaurea virgata Lam. Not in Dorn squarrose knapweed1,2,3,7 ssp. squarrosa (Boiss.) Gugler Chondrilla juncea L. Not in Dorn rush skeletonweed1,2,3,7, gumsuccory5, skeletonweed6 Cichorium intybus L. chicory1,2,3,4,5,6,7, wild succory5, bluesailors5 Cirsium arvense (L.) Scop. Canada thistle1,3,4,5,6,7, Canadian thistle2, field thistle4, creeping thistle5 Cirsium vulgare (Savi) Ten. bull thistle1,2,3,4,5,6,7, common thistle5, spear thistle5 Cosmos bipinnatus Cav. garden cosmos2,7, common cosmos6 Cota tinctoria (L.) J. Guy yellow chamomile1,3,4,5,6, golden chamomile2,7 [Anthemis tinctoria L.] Crepis tectorum L. narrowleaf hawksbeard1,2,7, rooftop hawksbeard3,5, annual hawksbeard5 Echinops sphaerocephalus L. great globe thistle1,2,3,7, globe thistle6 Galinsoga quadriradiata Ruiz & Pav. hairy galinsoga1, shaggy soldier2,7, fringed quickweed3,4, quickweed5, garden pest5, common quickweed6 Gnaphalum uliginosum L. low cudweed1,4,6, marsh cudweed2,5,7 Guizotia abyssinica (L. f.) Cass. Not in Dorn Nigerseed2,6, ramtilla7 Hieracium aurantiacum L. orange hawkweed1,3,5,7, devil’s paintbrush2, kingdevil5, golden mouseear hawkweed5, orangered kingdevil6 Hieracium caespitosum Dumort. yellow hawkweed1, meadow hawkweed2,3,5,7, yellow kingdevil6 Hieracium ×flagellare Willd. whiplash hawkweed2,6, hawkweed7 Hieracium praealtum Vill. ex Gochnat tall hawkweed1, kingdevil2,7, glaucous kingdevil6 Jacobaea vulgaris Gaertn. tansy ragwort1,3,5,6, stinking willie2,7 [Senecio jacobaea L.] Lactuca serriola L. prickly lettuce1,2,3,4,5,6,7 Leucanthemum maximum (Ramond) DC. Portuguese daisy6, max chrysanthemum7, Shasta dasy2 [Chrysanthemum lacustre Brot., misapplied] Leucanthemum vulgare Lam. oxeye daisy1,2,3,4,5,6,7, marguerite4,5, moondaisy5 [Chrysanthemum leucanthemum L.] Logfia arvensis (L.) Holub field cottonrose7, field filago3,5, field false cottonrose2 Matricaria chamomilla L. wild chamomile1,2,5, chamomile6, German chamomile7 [Matricaria recutita L.] Onopordum acanthium L. Scotch thistle1,2,3,4,5,6,7 cotton thistle5 Scorzonera laciniata L. cutleaf vipergrass2,7, self salsify3, false salsify4 Senecio vulgaris L. common groundsel1,3,5,6, old-man-in-the-spring2,5,7, groundsel4 Sonchus arvensis L. ssp. a. field sowthistle2,4, perennial sowthistle1,3,5,6, field milkthistle5, spiny sowthistle7 Sonchus arvensis L. marsh sowthistle1,5, field sowthistle2,4, perennial sowthistle3,6, moist sowthistle7 ssp. uliginosus (M. Bieb.) Nyman [Sonchus uliginosus M. Bieb.] Sonchus asper (L.) Hill spiny sowthistle1,3,7, spinyleaf sowthistle2, prickly sowthistle4,5,6 Sonchus oleraceus L. annual sowthistle1,3, common sowthistle2,4,5,6,7 Tagetes erecta L. Aztec marigold2, French marigold6, Aztec sowthistle7 [Tagetes patula L.] Tanacetum balsamita L. costmary chrysanthemum1,3, costmary2,4,5,6,7, mint geranium4,5 Tanacetum parthenium (L.) Sch. Bip. feverfew1,2,3,4,5,6,7, featherfew5 Tanacetum vulgare L. common tansy1,2,3,4,5,6,7 Taraxacum erythrospermum Andrz. ex Besser redseed dandelion2,3,4,5,6, rock dandelion7 [Taraxacum laevigatum (Willd.) DC.] Taraxacum officinale Weber ex F.H. Wigg. dandelion1,3, common dandelion2,4,5,6,7 Tragopogon dubius Scop. western salsify1,3,4, meadow goat’s beard2, goat’s beard4, yellow salsify5,7, fistulous goat’s beard6 Tragopogon porrifolius L. common salsify1,3, salsify2,4,5,6,7, vegetable oyster4,5,6 Tragopogon pratensis L. meadow salsify1,3,4,5, Jack-go-to-bed-at-noon2,5,7, showy goat’s beard6 [Tragopogon lamottei Rouy] Tripleurospermum inodoratum (L.) Sch. Bip. false chamomile1, false mayweed2,7, scentless chamomile3,6, wild chamomile4, [Matricaria maritima L., misapplied] scentless mayweed5 Berberidaceae Berberis thunbergii DC. Japanese barberry1,2,3,6,7

2 Boraginaceae Anchusa arvensis (L.) M. Bieb. small bugloss1,2,3,6,7 Asperugo procumbens L. catchweed1,3,5,6, German madwort2,7, madwort4,5,6 Buglossoides arvensis (L.) I.M. Johnst. corn gromwell1,2,3,5,6,7 [Lithospermum arvense L.] Cynoglossum officinale L. houndstongue1,3,4,6, gypsy flower2,7, common houndstongue5 Echium vulgare L. blueweed1,3,4,5,6, common viper’s bugloss2,5,7, viper’s bugloss4, snakeflower5, bluethistle5 Lappula squarrosa (Retz.) Dumort. European sticktight1, bristly sheepburr2, blue stickseed 4,5, tworow stickseed6, European stickseed7 Myosotis arvensis (L.) Hill field forget-me-not1,3,5,7, rough forget-me-not2, field scorpion grass5,6 Myosotis micrantha Pall. ex Lehm. blue scorpion grass2,3,6, strict forget-me-not7 Myosotis scorpioides L. true forget-me-not1,2,3, common forget-me-not5, water scorpion grass6 Myosotis sylvatica Hoffm. woodland forget-me-not2,7, wood-forget-me-not3,5, garden forget-me-not4,6 Brassicaceae (Cruciferae) Alliaria petiolata (M. Bieb.) Cavara & Grande garlic mustard1,2,4,5,7, hedge garlic5 Not in Dorn Alyssum alyssoides (L.) L. yellow alyssum1,3, pale madwort2,7, pale alyssum4,5 Alyssum desertorum Stapf var. d. dwarf alyssum1,3, desert madwort2,7, alyssum4, desert alyssum5 Alyssum simplex Rudolphi field alyssum1, European madwort2, smallflowered alyssum3, alyssum7 [Alyssum parviflorum Fisch. ex M. Bieb. var. micranthum (C.A. Mey.) Dorn] Armoracia rusticana P. Gaertn., B. Mey., & horseradish1,2,3,4,5,6,7 Scherb. Barbarea vulgaris R. Br. yellow rocket1,3,5,6, garden yellowrocket2,7, wintercress4, bitter wintercress5 Berteroa incana (L.) DC. hoary alyssum1,3,6,7, hoary false madwort2, hoary false alyssum4, berteroa5 Brassica juncea (L.) Czern. Indian mustard1,3,4,5,6, Chinese mustard2,5,6, brown mustard6 Brassica nigra (L.) Koch Not in Dorn black mustard1,2,4,5,6,7 Brassica rapa L. birdsrape mustard1,3, rape2, wild turnip4, field mustard5,7, common mustard5, bird rape5 Camelina microcarpa Andrez. ex DC. smallseed falseflax1,3,4,6, littlepod falseflax2,5,7, hairy falseflax5 Camelina sativa (L.) Crantz largeseed falseflax1,3,6, gold-of-pleasure2,4,5, falseflax5,7 Capsella bursa-pastoris (L.) Medik. shepherd’s purse1,2,3,4,5,6,7 Chorispora tenella (Pall.) DC. blue mustard1,3,4,5,6, crossflower2,7, chorispora5 Conringia orientalis (L.) Dumort. haresear mustard1,2,3,4,5,6,7, haresear cabbage5, treacle mustard5 Descurainia sophia (L.) Webb ex Prantl flixweed1,3,4,5, Sophia2,6,7, tansy mustard6 Draba nemorosa L. wood whitlograss1, woodland whitlowgrass2, yellow whitlowort4, wood draba 5,7 Draba verna L. spring whitlowgrass1,2,3,5, whitlowgrass6, spring draba7 Erucastrum gallicum (Willd.) O.E. Schulz dog mustard1,3,4,5,6, common dog mustard2,7 Erysimum cheiranthoides L. wallflower mustard1,3, wormseed wallflower2,4,7, treacle mustard5, wormseed mustard5,6 Erysimum repandum L. bushy wallflower1,3,4,6, spreading wallflower2,5,7, treacle mustard6 Euclidium syriacum (L.) R. Br. Syrian mustard1,2,3,7, euclidium5 Hesperis matronalis L. damesrocket1,3,4,6,7, mother-of-the-evening2, Damask violet5, dame’s violet5, sweet rocket5 Isatis tinctoria L. dyer’s woad1,2,3,5,7, woad6 Lepidium appelianum Al-Shehbaz hairy whitetop1,3,7, globepod pepperwort2, globepod hoarycress5,6, whitetop4 [Cardaria pubescens (C.A. Mey.) Jarm.] Lepidium campestre (L.) R. Br. field pepperweed1,3,7, creamanther field pepperwort2, field peppergrass4,5, field cress5,6, pepperwort5 Lepidium chalapensis L. lenspodded whitetop1,2,7, lenspod pepperwort2, Chalapa hoarycress3,5, lenspodded [Cardaria chalapensis (L.) Hand.-Mazz.] hoary cress4,6 Lepidium draba L. hoary cress1,3,4, heartpod pepperwort2, heartpod hoarycress2,5,6, hoary pepperwort5, [Cardaria draba (L.) Desv.] whitetop7 Lepidium latifolium L. perennial pepperweed1,3,6, broadleaf pepperwort2,5,7, pepperwort5 Lepidium perfoliatum L. clasping pepperweed1,3,6,7, clasping pepperwort2, clasping peppergrass4,5 Lepidium sativum L. garden cress1,3,4,5,6, garden pepperwort2, garden pepperweed7 Lobularia maritima (L.) Desv. sweet alyssum1,3,5,6,7 sweet alison2,5 Nasturtium microphyllum Boenn. ex Reichenb. onerow yellowcress2,7 Not in Dorn Nasturtium officinale R. Br. watercress1,2,3,4,5,6,7

3 Raphanus sativus L. radish1,2,4,6, wild radish3,5, cultivated radish7 Rorippa austriaca (Crantz) Bess. Not in Dorn Austrian fieldcress1,3,4, Austrian yellowcress2,7, field yellowcress6 Sinapsis arvensis L. wild mustard1,3,5,7, corn mustard2, charlock4,5,6 Sisymbrium altissimum L. tumblemustard1, tall hedgemustard2, tall tumblemustard3,7, tumbling mustard4,6, Jim hill mustard5,6 Sisymbrium irio L. Not in Dorn London rocket1,2,4,6,7 Sisymbrium loeselii L. tall hedge mustard1,3,4,6, false London rocket2, Loesel tumblemustard5, small tumbleweed mustard7 Strigosella africana (L.) Botsch. Malcolm stock1,3, African addersmouth2, malcolmia5, African mustard7 [Malcolmia africana (L.) R. Br.] Thlaspi arvense L. field pennycress1,2,3,4,5,6,7, fanweed5, stinkweed5 Campanulaceae Campanula rapunculoides L. creeping bellflower1,2,3,4,5, rover bellflower4,5,6, rampion bellflower7 Cannabaceae Cannabis sativa L. marijuana1,3,4,7, hemp2,4,5,6, Indian hemp6 Caprifoliaceae (inc. in Adoxaceae, APG IV; Viburnum remains) Lonicera ×bella Zabel Not in Dorn showy fly honeysuckle7 Lonicera morrowii A. Gray Morrow’s honeysuckle2,7 Lonicera tatarica L. Tatarian honeysuckle1,3,4,6,7, twinsisters2 Viburnum lantana L. wayfaring tree2,6,7 Caryophyllaceae Agrostemma githago L. corn cockle1,3,4,6, common corn cockle2,5,7, corn campion5 Arenaria serpyllifolia L. thymeleaf sandwort1,2,3,4,5,6,7 Cerastium fontanum Baumg. common mouseear chickweed2,4, mouseear chickweed1,3,5,6, common ssp. vulgare (Hartm.) Greuter & Burdet chickweed5, big chickweed5,7 Cerastium glomeratum Thuill. Sticky chickweed1,3,5, sticky mouseear chickweed2, clammy chickweed6 [Cerastium viscosum L., misapplied] Cerastium tomentosum L. snow-in-summer2,3,7 Dianthus armeria L. ssp. a. Deptford pink1,2,3,4,5,6,7, grass pink5 Dianthus barbatus L. ssp. b. sweet William2,3,5,6,7 Dianthus deltoides L. ssp. d. maiden pink2,3,5,6,7 Gypsophila paniculata L. babysbreath1,3,4,5,6,7, tall babysbreath2 Gypsophila scorzonerifolia Ser. garden babysbreath2,7 Holosteum umbellatum L. ssp. u. spurry1,3, jagged chickweed2,4,5,6,7 Saponaria officinalis L. bouncingbet1,3,4,6,7, bouncingbett2,5, bouncing bettsoapwort5,6 Silene cserei Baumg. biennial campion1, Balkan catchfly2,7, smooth catchfly4 Silene dichotoma Ehrh. ssp. d. hairy catchfly1,3, forked catchfly2,4,5,7 Silene latifolia Poir. white campion1,3,4,5,6, bladder campion2,7, white cockle4,6, evening campion5 Silene noctiflora L. nightflowering catchfly1,2,3,4, sticky cockle4,5,6, nightflowering silene5,7, catchfly6 Silene pendula L. Not in Dorn nodding catchfly1,7, nodding campion2 Silene vulgaris (Moench) Garcke bladder campion1,3,4,5,6, maiden’s tears2,7 Spergula arvensis L. corn spurry1,2,3,4,5,7, stickwort5, starwort5 Spergularia echinosperma (Čelak.) Asch.&Graebn. bristleseed sandspurry2,7 Spergularia media (L.) C. Presl var. m. stout sandspurry3,5, satin flower2, media sandspurry7 Spergularia rubra (L.) J. Presl & C. Presl red sandspurry1,3,5,7, ruby sandspurry2, roadside sandspurry6 Stellaria media (L.) Vill. common chickweed1,2,4,6, chickweed3,5,7 Vaccaria hispanica (Mill.) Rauschert cowcockle1,2,3,4,5, cowherb4,5,6, cow soapwort7 Chenopodiaceae (inc. in Amaranthaceae, APG IV) Atriplex heterosperma Bunge twoscale saltbush2,7, weedy orache3 Atriplex hortensis L. garden orach1,3,7, garden orache2,5, orache4,5,6, sea purslane5 Atriplex patula L. spreading orach1,3, halberdleaf orache2, common orache5, spear orache5, spearscale6, spear saltbush7 Atriplex rosea L. red orach1,3, tumbling orache2, red orache4,5,6, red scale4, tumbling saltweed7 Bassia hyssopifolia (Pall.) Kuntze fivehook bassia1,3,4,6, fivehorn smotherweed2,7 Kochia scoparia (L.) Scrad. kochia1,3,4, Mexican fireweed2, fireweed4, summer cypress4,5,6, mock cypress4,5, [Bassia sieversiana (Pall.) W.A. Weber] burning bush5,7 red belvedere5 Chenopodium album L. common lambsquarter1,3, lamb’s quarters2,4,5,6,7, white goosefoot5, pigweed5,6 Chenopodium glaucum L. var. g. oakleaf goosefoot1,2,3,4,5,6,7, glaucous goosefoot5 Dysphania botrys (L.) Mosyakin & Clements Jerusalem oak goosefoot1,3,5,7, Jerusalem oak2,4,5,6, feather geranium5,6 [Chenopodium botrys L.] Halogeton glomeratus (M. Bieb.) C.A. Mey. halogeton1,3,5, saltlover2,7

4 Salsola collina Pall. slender Russian thistle2,3,7, tumbleweed4, katune6 Salsola tragus L. Russian thistle1,3,4,5,6, prickly Russian thistle2,7, tumbleweed4,5, wind witch5, saltwort6 Suaeda linifolia Pall. pinleaf seepweed2,7 Convolvulaceae Convolvulus arvensis L. field bindweed1,2,3,4,6,7, field morningflory5, small bindweed5 Cuscuta approximata Bab. smallseed alfalfa dodder1,5, alfalfa dodder2,7, clustered dodder3,5, legume dodder6 var. urceolata (Kuntze) Yunck.] Cuscuta epithymum L. clover dodder1,2,3,7, common dodder5, thyme dodder5, legume dodder6 Crassulaceae Sedum acre L. mossy stonecrop1,2,3, wall pepper5, golden carpet6, goldmoss stonecrop7 Cucurbitaceae Cucurbita pepo L. var. p. Not in Dorn field pumpkin2,7 Dipsacaceae (inc. in Caprifoliaceae, APG IV) Dipsacus fullonum L. common teasel1,3,4, Fuller’s teasel2,6,7, gypsy combs6 Knautia arvensis (L.) J.M. Coult. bluebuttons1,2,3,4,6, bluecaps4, field scabious4,5,7 Elaeagnaceae Elaeagnus angustifolia L. Russian olive1,2,3,4,5,6,7, oleaster6 agraria M. Bieb. urban spurge2,3,7 Euphorbia cyparissias L. cypress spurge1,2,3,4,5,6,7 Euphorbia helioscopia L. sun spurge1,3, mad woman’s milk2,7, summer spurge5, wartweed6 Euphorbia myrsinites Not in Dorn myrtle spurge1,2,7 Euphorbia virgata Waldst. & Kit.] leafy spurge1,3,4,6,7, wolf’s milk2, Esula spurge5, Russian leafy spurge7 [Euphorbia esula L. var. e.] [Euphorbia esula L. var. uralensis (Fisch. ex Link) Dorn] Fabaceae (Leguminosae) Astragalus cicer L. chickpea milkvetch2,3,5,6,7 Astragalus tibetanus Benth. ex Bunge Tibetan milkvetch2, Tibet milkvetch7 Caragana arborescens Lam. Siberian peashrub2,4,7, caragana3 Coronilla varia L. trailing crownvetch1,2, purple crownvetch2, crown vetch4,5,6,7 Lathyrus latifolius L. everlasting peavine1,3,5, everlasting pea2,4,6, perennial sweetpea4, perennial peavine5, perennial pea7 Lathyrus sylvestris L. flat peavine1,3,5, narrowleaf vetchling2, narrowleaved everlasting peavine5, flat pea 7 Lotus corniculatus L. birdsfoot trefoil1,3,4,5,6, garden birdsfoot trefoil2 Medicago falcata L. alfalfa2,4, sickle medic3, yellow lucerne5, sickle medic5, yellow alfalfa6,7 [Medicago sativa L. var. falcata (L.) Doell] Medicago lupulina L. black medic1,3,5, black medick2,4,6,7, hop clover5 Medicago polymorpha L. burclover1,3, toothed medick2, burclover5, hairy medic5,7, smooth burclover6 Medicago sativa L. ssp. glomerata (Balb.) Rouy alfalfa1,7 Not in Dorn Medicago sativa L. var. s. alfalfa2,3,4,5,6,7, lucerne5, blue alfalfa6 Medicago sativa L. ssp. ×varia (E.B. Martyn) alfalfa1,4,5,6,7 Arcang. Not in Dorn Melilotus albus Medik. white sweetclover1,3,4,5,6, yellow sweetclover2, sweetclover7 Melilotus officinalis (L.) Pall. yellow sweetclover1,2,3,4,5,6, sweetclover7 Onobrychis viciifolia Scop. sainfoin2,3,4,6,7, saintfoin5, sandfain5, holy clover5 Oxytropis riparia Litv. Ruby Valley locoweed2, riparian crazyweed3, Oxus locoweed7 Phaseolus vulgaris L. . kidney bean2,7, green bean3, common bean6 Sphaerophysa salsula (Pall.) DC. swainsonpea1,3, red bladdervetch2, swainsona5, alkali swainsonpea7 Trifolium aureum Pollich hop clover1,3, greater hop clover2, palmate hop clover6, golden clover7 Trifolium campestre Schreb. large hop clover1,3, lesser hop clover2, low hop clover4, pinnate hop clover6, field clover7 Trifolium dubium Sibth. Not in Dorn small hop clover1,3,4, sucking clover2,7, least hop clover5, little hop clover6 Trifolium fragiferum L. strawberry clover1,3,4,5,6,7, strawberryhead clover2 Trifolium hybridum L. Alsike clover2,3,4,5,6,7 Trifolium pratense L. red clover1,2,3,4,5,6,7 Trifolium repens L. white clover1,2,3,4,5,6,7, ladino clover4, Dutch clover5 Vicia cracca L. bird vetch1,3,4,5,6,7, purplewhite tufted vetch2, tufted vetch5, cat peas5, tinegrass5

5 Vicia villosa var. glabrescens Not in Dorn hairy vetch1,4,5, winter vetch2,5,6,7, woollypod vetch4, woolly vetch5, Vicia villosa Roth var. v. hairy vetch1,3,4,5,6, winter vetch2,5,7, woollypod vetch4, woolly vetch5,6 Fumariaceae (inc. in Papaveraceae, APG IV) Fumaria officinalis L. fumitory1,4,6, common fumitory3,5, drug fumitory2,7 Gentianaceae Centaurium pulchellum (Sw.) Druce ex B.L. Rob. branched centaury2,7, centaury4, branching centaury6 & Fernald Geraniaceae Erodium cicutarium (L.) L’Her. ex Aiton redstem filaree1,3,6, redstem storksbill2,7, filaria4 Geranium pusillum L. smallflower geranium1,3, smallflower cranesbill2,5,6, small cranesbill4, small geranium7 Grossulariaceae Ribes rubrum L. northern red currant2, cultivated red currant3,5, garden red currant6, cultivated current7 Hemerocallidaceae Hemerocallis fulva (L.) L. tawny daylily1,3, orange daylily2,7, daylily4,6 Hyperiaceae Hypericum perforatum L. ssp. p. common St. Johnswort1,2,4,5,6,7, St. Johnswort3, Klamath weed5 Juncaceae Juncus compressus Jacq. roundfruit rush2,7, compressed rush3 Liliaceae (see Alliaceae, Amaryllidaceae, Asparagaceae, Hemerocallidaceae) (Labiatae) Acinos arvensis (Lam.) Dandy Not in Dorn mother-of-thyme1,6, basil thyme2,3,5,7 [Satureja acinos (L.) Scheele] Dracocephalum thymiflorum L. thymeleaf dragonhead2,3,5,6,7 Galeopsis ladanum L. red hempnettle1,2,7, hempnettle6 Glechoma hederacea L. ground ivy1,2,3,4,5,7, Gill-over-the-ground5,6, field balm5, creeping Charlie5 Lallemantia peltata (L.) Fisch. & C.A. Mey. lion’s heart2,7 Lamium amplexicaule L. var. a. henbit1,3,4,6, giraffehead2, common henbit5, common deadnettle5, henbit deadnettle7 Leonurus cardiaca L. motherwort1,2,3,4,5,6, lion’s tail6, common motherwort7 vulgare L. white horehound1,2,3, common horehound4, horehound5,6,7 Mentha spicata L. spearmint1,2,3,4,5,6,7, mackerelmint5 Nepeta cataria L. catnip1,2,3,4,5,6,7, catmint5 Nepeta grandiflora M. Bieb. Caucasus catnip2,7 Nepeta racemosa Lam. racemed catnip2,7 Salvia nemorosa L. woodland sage2,7, violet sage3, sage4 Linaceae Linum usitatissimum L. cultivated flax2,3,5, common flax4,5,6,7, linseed5 Lythraceae Lythrum salicaria L. purple loosestrife1,2,3,4,5,6,7, longpurples5, spiked willowherb5 Abutilon theophrasti Medik. velvetleaf1,2,3,4,6,7, pie marker6 Alcea rosea L. hollyhock1,2,3,4,6,7 Hibiscus trionum L. Venice mallow1,3,4, -of-an-hour2,4,5,6,7, modesty5 Lavatera thuringiaca L. gay mallow2, tree lavatera7 moschata L. musk mallow1,2,3,5,6,7 Malva neglecta Wallr. common mallow1,4,6,7, dwarf mallow2,3,5, cheeses6 Malva parviflora L. little mallow1,3,6, smallwhorl mallow2, smallfruited mallow4, alkali mallow5, cheeseweed5, cheeseweed mallow7 Malva pusilla Smith low mallow2, common mallow3,4,7, dwarf mallow6 [Malva rotundifolia L.] Malva sylvestris L. high mallow1,2,3,4,6,7, common mallow5 Malva verticillata L. curled mallow1, largewhorl mallow2, clustered mallow3,4,7, whorled mallow6 Moraceae Morus alba L. white mulberry1,2,3,4,5,6,7 Oleaceae Syringa vulgaris L. Not in Dorn common lilac1,2,4,6,7, lilac3 Papaveraceae Papaver somniferum L. opium poppy2,3,4,5,6,7 lanceolata L. buckhorn plantain1,3,5, English plantain2,4,5,6, buckhorn4, ribwort5, ribgrass6, narrow

6 plantain7 Plantago major L. broadleaf plantain1, greater plantain2, common plantain4,5,6,7, nippleseed plantain5 (Gramineae) Aegilops cylindrica Host jointed goatgrass1,2,3,4,6,7, goatgrass5 Agropyron cristatum (L.) Gaertn. var. c. crested wheatgrass2,3,4,5,6,7 Agropyron cristatum (L.) Gaertn. clustered wheatgrass2, crested wheatgrass2,3, desert wheatgrass7 var. desertorum (Fisch. ex Link) Dorn Agropyron cristatum (L.) Gaertn. Siberian wheatgrass2,7 var. fragile (Roth) Dorn Agropyron triticeum Gaertn. annual false wheatgrass2, annual wheatgrass3,7 Agrostis capillaris L. colonial bentgrass1,3,5,7, colonial bent2, Rhode Island bentgrass6 Agrostis gigantea Roth Not in Dorn redtop1,4,6,7. black bentgrass2 Agrostis stolonifera L. creeping bentgrass1,3,5,6,7, spreading bent2,6 redtop4,5, fiorin5 Aira caryophyllea L. silver hairgrass1,3,5,6,7, common silver hairgrass2 Alopecurus arundinaceus Poir. creeping meadow foxtail2,7, creeping foxtail3,4 Alopecurus geniculatus L. water foxtail1,3,5,6,7, marsh meadow foxtail2, marsh foxtail4,6 Alopecurus pratensis L. meadow foxtail1,3,4,5,6,7, field meadow foxtail2 Anthoxanthum odoratum L. Not in Dorn sweet vernalgrass1,5,6,7, large sweet vernalgrass2 Apera interrupta (L.) P. Beauv. dense silkybent2,7, interrupted apera3,5 Arrhenatherum elatius (L.) J. Presl & C. Presl tall oatgrass1,2,3,4,5,6,7 var. e. Not in Dorn fatua L. wild oat1,2,3,5,4,6,7 Avena sativa L. oats1,6, oat2, common oats3,5,7, cultivated oats4 Bromus briziformis Fisch. & C.A. Mey. rattlesnake brome1,2,3,7, rattlegrass5, rattle chess5, rattlesnake grass5, rattlesnake chess6 Bromus commutatus Schrad. hairy chess1,2,3,4,5,6, meadow brome5, hairy brome5, bald brome7 Bromus inermis Leyss. smooth brome1,2,3,4,5,6,7, Hungarian brome5 Bromus hordeaceus L. soft brome1,2,7, soft chess6 Bromus japonicus Thunb. ex Murray Japanese brome1,3,4,5, field brome2, Japanese chess5,6 Bromus secalinus L. cheat1,4,5,6, rye brome2,3,5, chess5,6, field brome7 Bromus squarrosus L. corn brome2,7, largeflowered brome3 Bromus tectorum L. downy brome1,3,4, cheat grass2,5,7, downy chess5,6, June grass6 Crypsis alopecuroides (Piller & Mitterp.) Schrad. foxtail pricklegrass2,7, fox crypsis3 Dactylis glomerata L. orchardgrass1,2,3,4,5,6,7, cock’s foot grass5 Digitaria ischaemum (Schreb.) Schreb. ex Muhl. smooth crabgrass1,2,3,4,5,6,7 Digitaria sanguinalis (L.) Scop. large crabgrass1,3, hairy crabgrass2,4,5,7, northern crabgrass6 Echinochloa crus-galli (L.) P. Beauv. barnyardgrass1,4,6,7, large barnyardgrass2,3,5, watergrass5 Echinochloa frumentacea Link Not in Dorn Japanese millet1,2, billion-dollar grass6,7 Elymus elongatus (Host) Runem. tall wheatgrass1,3,4,6,7, Eurasian quackgrass2 var. ponticus (Podp.) Dorn Elymus hispidus (Opiz) Melderis var. h. intermediate quackgrass2,3,4,5, intermediate wheatgrass7 Elymus hispidus (Opiz) Melderis intermediate quackgrass2,3,4,5, intermediate wheatgrass7 var. ruthenicus (Griseb.) Dorn Elymus junceus Fisch. Russian wildrye2,3,4,7 Elymus multicaulis Kar. & Kir. manystem lymegrass2, creeping wildrye3,5, beardless wildrye5, manystem wildrye7 Elymus racemosus Lam. mammoth lymegrass2, Volga wildrye3, Sibarian wildrye5, mammoth wildrye7 Elymus repens (L.) Gould quackgrass1,3,4,5,6, creeping wildrye2, quitchgrass5,7, couchgrass5 Eragrostis cilianensis (All.) Vignolo ex Janch. stinkgrass1,2,3,4,5,6,7, candygrass5 Eragrostis minor Host little lovegrass2,3,4,7 Eragrostis pilosa (L.) P. Beauv. India lovegrass1,3,4,5,6,7, Indian lovegrass2, tufted eragrostis5 var. perplexa (L.H. Harv.) S.D. Koch Hordeum distichon L. barley1,7, common barley2, tworow barley5,6 Hordeum murinum L. hare barley1,7, wall barley2, charming barley3,5 ssp. leporinum (Link) Arcang. [Hordeum leporinum Link] Hordeum vulgare L. ssp. v. barley1,6, common barley2,3,7, cultivated barley4,5 Lolium multiflorum Lam. Italian ryegrass1,3,4,5,6,7, perennial ryegrass2, Australian ryegrass5 Lolium perenne L. perennial ryegrass1,2,3,4,5,7, English ryegrass5,6 Lolium persicum Boiss. & Hohen. ex Boiss. Persian darnel1,3,6, Persian ryegrass2 Panicum miliaceum L. ssp. m. wild proso millet1,3, hog millet2, broomcorn millet4,5,6,7, proso millet4,6 Phalaris canariensis L. canarygrass1,3,4,5,6, common canarygrass2, annual canarygrass7

7 Phleum pratense L. var. p. timothy1,3,4,6,7, common timothy2,5 Poa annua L. annual bluegrass1,2,3,4,5,7, speargrass6 Poa bulbosa L. ssp. vivipara (Koeler) Arcang. bulbous bluegrass1,2,3,5,6,7 Poa compressa L. Canada bluegrass1,3,4,5,6,7, flatstem bluegrass2 Poa pratensis L. Kentucky bluegrass1,2,3,4,5,6,7 Poa trivialis L. roughstalk bluegrass1,2,3,5, rough bluegrass6,7 Polypogon monspeliensis (L.) Desf. rabbitfoot polypogon1,3,5, annual rabbitsfoot grass2,7, rabbitfoot grass4,6, annual beardgrass5 Puccinellia distans (L.) Parl. spreading alkaligrass2, weeping alkaligrass3,5,7, European alkaligrass5,6 Schedonorus arundinaceus (Schreb.) Dumort. tall fescue1,3,4,5,6,7, tall false ryegrass2, alta fescue5,6, reed fescue5 [Festuca arundinacea Schreb.] Schedonorus pratensis (Huds.) P. Beauv. meadow fescue1,3,4,5,6,7, meadow false ryegrass2, English fescue5 [Festuca pratensis Huds.] Secale cereale L. cultivated rye2,3,5, rye4,6, cereal rye7 Setaria italica (L.) P. Beauv. foxtail millet1,3,4,5,6,7, Italian bristlegrass2, Hungarian grass5 Setaria pumila (Poir.) Roem. & Schult. ssp p. yellow foxtail1,3,4,7, yellow bristlegrass2, yellow foxtailgrass6 [Setaria glauca (L.) P. Beauv.] Setaria verticillata (L.) P. Beauv. bristly foxtail1,3,4,5, rough bristlegrass2, bur bristlegrass5, bur foxtailgrass6, hooked bristlegrass7 Setaria viridis (L.) P. Beauv. var. v. green foxtail1,3,4, green bristlegrass2,5,7, green foxtailgrass6 Sorghum halepense (L.) Pers. Johnsongrass1,2,3,4,5,6,7 Taeniatherum caputa-medusae (L.) Nevski medusahead1,2,7 Not in Dorn Triticum aestivum L. bread wheat2, cultivated wheat3,5, wheat4, common wheat6,7 (Leers) Coss. & Durieu ventenata1,2,3,5, North African grass2,7 Polygonaceae Fallopia convolvulus (L.) Á. Löve wild buckwheat1, black bindweed2,3,5,6,7, climbing buckwheat4, wild buckwheat4, [Polygonum convolvulus L.] climbing bindweed5, knot bindweed5, ivy bindweed5, dullseed5, cornbind5 Persicaria hydropiper (L.) Spach Not in Dorn marshpepper smartweed1,3,5, mild waterpepper2, water pepper4,6, smartweed5, marshpepper knotweed7 Persicaria maculosa Gray ladysthumb1,2,3,4,6, spotted ladysthumb2,5,7, heartweed5 [Polygonum persicaria L.] Polygonum aviculare L. prostrate knotweed1,3,5,7 yard knotweed2, knotweed4, doorweed5 Rheum rhabarbarum L. Not in Dorn garden rhubarb2,7, rhubarb3,4 Rumex acetosella L. red sorrel1,3,5,6, common sheep sorrel2,7 sheep sorrel4,5, cow sorrel5, mountain sorrel5, field sorrel5, horse sorrel5, sour weed5 Rumex crispus L. curly dock1,2,3,4,5,6,7, sour dock5, yellow dock5 Rumex patientia L. spinach dock1,3, patience dock2,4,5,6,7 Rumex stenophyllus Ledeb. narrowleaf dock1,2,3,7 Portulacaceae Portulaca oleracea L. common purslane1,4,5,6, little hogweed2,7 Potamogetonaceae Potamogeton crispus L. curlyleaf pondweed1,3, curly pondweed2,6,7, curly muckweed4, curled pondweed5 Ranunculaceae Ranunculus acris L. tall buttercup1,2,3,4,5,7, meadow buttercup5,6, common buttercup6, blisterplant6 Ranunculus repens L. creeping buttercup1,2,3,5,6,7 Ranunculus testiculatus Crantz bur buttercup1,3,6, curveseed butterwort2,7, hornseed buttercup5 Resedaceae Reseda lutea L. yellow mignonette1,3,5,7, yellow upright mignonette2, wild mignonette4, reseda4 Rhamnaceae Frangula alnus Mill. glossy false buckthorn2, tall buckhthorn3, European alder buckthorn6, glossy [Rhamnus frangula L.] buckthorn7 Rhamnus cathartica L. European buckthorn1,2,3, common buckthorn4,6,7 Alchemilla filicaulis var. f. Not in Dorn thinstem lady's mantle2,7 lucidus Schltdl. hairystem cotoneaster2, shiny cotoneaster7 [Cotoneaster aculifolia Turcz.] Malus bacata (L.) Borkh. Not in Dorn Siberian crabapple2, Manchurian crab apple7 Malus pumila Mill. cultvated apple2,5, common apple3, apple4,6, paradise apple7 Potentilla argentea L. var. a. silvery cinquefoil1,3,4,5,7, silverleaf cinquefoil2, silvery fivefingers6 Potentilla inclinata Vill. Not in Dorn ashy cinquefoil2,7

8 Potentilla recta L. sulfur cinquefoil1,2,3,4,7, erect cinquefoil5, sulpher fivefingers6 Poterium sanguisorba L. salad burnet1,2,3,6, garden burnet5, small burnet5,7 var. polygamum (Waldst. & Kit.) Vis. [Sanguisorba minor Scop.] Prunus avium L. sweet cherry1,2,3,4,5,6,7 Rosa rubignosa var. r. sweetbriar rose1,3,7, sweetbriar,2,4,5,6, eglantine5,6 [Rosa eglanteria L.] Rubus laciniatus Willd. cutleaf blackberry1,2,3,7, evergreen blackberry5,6 Rubus idaeus L. var. i. Not in Dorn common red raspberry2, American red raspberry7 Sorbus aucuparia L. European mountain ash1,2,3,5,6,7, Rowan tree5,6, Rubiaceae Galium mollugo L. smooth bedstraw1,3, false babysbreath7, wild madder5, white bedstraw2,5, great hedge bedstraw5 Galium verum L. yellow bedstraw1,3,4,5,6, yellow spring bedstraw2,7, lady’s bedstraw5 Salicaceae Populus alba L. white poplar1,2,3,6,7, silver poplar4,6 Salix fragilis L. crack willow1,2,3,4,6,7 Salix fragilis L. × S. fragilis L. Sapindaceae Acer ginnala Maxim. Amur maple2,7 Scrophulariaceae (inc. in Plantaginaceae, APG IV; Verbascum remains) Digitalis purpurea L. foxglove1,3,5, purple foxglove2,7, common foxglove6 Linaria dalmatica (L.) Mill. var. d. Dalmatian toadflax1,2,3,5,6,7 Linaria vulgaris Mill. yellow toadflax1,3, greater-butter-and-eggs2, butter-and-eggs4,5,6,7 Verbascum thapsus L. common mullein1,3,4,5,6,7, great mullein2, flannel mullein5 anagallis-aquatica L. water speedwell1,3,4,5,6,7, blue water speedwell2, water pimpernel5 Veronica arvensis L. corn speedwell1,2,3,4,6,7, wall speedwell5, common speedwell5 Veronica biloba L. twolobe speedwell2,7, bilobed speedwell3,5 Veronica officinalis L. common speedwell1,4,5, common gypsyweed2,7, Paul’s betony 5 Veronica persica Poir. Persian speedwell1,3,5, birdseye speedwell2,4,6,7 Veronica verna L. spring speedwell2,3,7 Solanaceae Hyoscyamus niger L. black henbane1,2,3,5,7, henbane4,6, hog’s bean5 Lycium barbarum L. matrimonyvine1,2,3,4,5,6,7, boxthorn5 Solanum dulcamara L. bittersweet nightshade1,3, climbing nightshade2,4,5,7, bittersweet4,5,6, felonwort5, blue bindweed5 Solanum physalifolium Rusby hairy nightshade1,3,5,6, groundcherry nightshade2, viscid nightshade4, hoe var. nitidibaccatum (Bitter) Edmonds nightshade7 Tamaricaceae Tamarix chinensis Lour. Chinese tamarisk1, fivestamen tamarisk2,7, salt cedar4, tamarisk5,6 Tamarix ramosissima Ledeb. Not in Dorn salt ceder1,4,7, tamarisk6 Ulmaceae Ulmus pumila L. Sibarian elm1,2,3,4,6,7, Chinese elm6 Valerianaceae (inc. in Adoxaceae, APG IV) Valeriana officinalis L. common valerian1,3, allheal2,5, garden heliotrope5,6, garden valerian5,7 Violaceae Viola tricolor L. var. t. pansy1, johnny jumpup2,6,7 Zygophyllaceae Tribulus terrestris L. puncturevine1,2,3,4,5,6,7, goat head4, land caltrop5, ground burnut5

Invasive taxa attributed to Wyoming but without documentation or in error: Eruca vesicaria (L.) Cav. Invaders garden rocket1,3,5,6, rocket salad2,4,5,7 ssp. sativa (Mill.) Thellung Fagopyrum esculentum Moench. Under cultivation buckwheat1,4,6,7, garden buckwheat2, fagopyrum3 Hippophae rhamnoides L. Under cultivation sea buckthorn2,3, seaberry7 Melilotus indicus (L.) All. Invaders Indian sweetclover1,2,3, smallflowered yellow sweetclover5, annual yellow sweetclover7 Spergularia diandra (Guss.) Held. & Sart. Invaders alkali sandspurry,2,3,5,7 Tragopogon miscellus Ownbey Multiple origin, U.S.A. Moscow salsify2,7, hybrid salsify3,5

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