371 Genus Neurellipes Bethune-Baker

AFROTROPICAL BUTTERFLIES 17th edition (2018). MARK C. WILLIAMS. http://www.lepsocafrica.org/?p=publications&s=atb

Genus Neurellipes Bethune-Baker, 1910

Transactions of the Entomological Society of London 43: 66 (1-84). Type-species: Lycaenesthes lusones Hewitson, by original designation.

= Neurypexina Bethune-Baker, 1910. Transactions of the Entomological Society of London 43: 64 (1-84). Type-species: Lycaenesthes lyzanius Hewitson, by original designation.

= Pseudoliptena Stempffer, 1946. Revue Francaise d’Entomologie 13: 8 (8-19). Type- species: Pseudoliptena bitje Stempffer, [1866], by monotypy.

The genus Neurellipes belongs to the Family Lycaenidae Leach, 1815; Subfamily Polyommatinae Swainson, 1827; Tribe Lycaenesthini Toxopeus, 1929. The other genera in the Tribe Lycaenesthini in the Afrotropical Region are Anthene, Cupidesthes, Triclema, and Monile.

Neurellipes (Hairtails) is a purely Afrotropical genus containing 49 species. The genus was revised by Libert in 2010.

Species groups in the genus Neurellipes (Libert, 2010: 236)

N. lusones group: N. lusones complex: N. lusones, N. onias, N. sonia, N. alixae N. fulvimacula complex: N. fulvimacula, N. kampala, N. tanzaniensis Species group: N. katera, N. lychnaptes

N. lychnides group: N. lychnides, N. larseni, N. cuypersi

N. leptines group: N. leptines subgroup: N. leptines complex: N. leptines, N. makala, N. rufomarginata, N. ngoko, N. radiata, N. scintillula, N. helpsi, N. bakeri, N. tessmanni, N. discimacula N. mahota complex: N. mahota, N. gola, N. ducarmei, N. georgiadisi N. juba complex: N. juba, N. pyroptera, N. erythropoecilus, N. zenkeri N. lachares subgroup: N. lachares complex: N. lachares, N. emkopoti, N. ramnika, N. atewa N. xanthopoecilus complex: N. xanthopoecilus, N. versatilis, N. lycotas N. lysicles subgroup: N. lysicles Unplaced: N. buchholzi

N. lyzanius group: N. lyzanius complex: N. lyzanius, N. ferenczi, N. flavomaculatus, N. boormani, N. lamprocles, N. mirellae, N. inferna, N. darwini, N. dubia

*Neurellipes lusones (Hewitson, 1874) Large Red-spot Hairtail

Large Red-spot Hairtail (Neurellipes lusones). Left – male upperside. Right – male underside.

Lycaenesthes lusones Hewitson, 1874. Transactions of the Entomological Society of London 1874: 347 (343-355). Neurellipes lusones Hewitson, 1874. d’Abrera, 2009: 800. Neurellipes lusones (Hewitson, 1874). Libert, 2010: 240.

Alternative common name: Large Red-spot Ciliate Blue. Type locality: Gabon: “Gaboon”. Holotype (male), Gabon (Rogers); N.H.M., London. Neallotype (female). Gabon?; N.H.M., London (Libert, 2010). Diagnosis: Close to Neurellipes chryseostictus but is larger and on the underside the white lines are straighter. Male forewing upperside orange patch smaller than in Anthene katera and larger than in

2 Anthene kampala; on underside the white lines narrower than in both these species (Congdon & Collins, 1998). Distribution: Gambia, Guinea, Sierra Leone, Ivory Coast (Collins et al., 2003), Ghana (Collins et al., 2003), Togo, Nigeria (south and Cross River loop) (Larsen, 2005a), Cameroon (Collins et al., 2003), Equatorial Guinea, Gabon, Congo, Central African Republic, Democratic Republic of Congo). Recorded, in error, from Angola, Tanzania, Sudan and Uganda (Libert, 2010). Specific localities: Guinea – Macenta (Libert, 2010); Nzerekore (Libert, 2010); Ziela in the Nimba Mountains (Libert, 2010). Sierra Leone – Baranga (Libert, 2010); Gola (Libert, 2010); Kholifa (Libert, 2010); Kondembaia in the Loma Mountains (Libert, 2010). Ivory Coast – Assinie (Libert, 2010); Bassam (Libert, 2010); Nimba Mountains (Libert, 2010); Yapo (Libert, 2010). Ghana – Ankasa (Libert, 2010); Bia (Libert, 2010); Cape Three Points (Libert, 2010); Tano Ofin (Libert, 2010); Ho (Libert, 2010); Kpandu (Libert, 2010); Lipke (Libert, 2010); Loloby (Libert, 2010). Togo – Misahoe (Libert, 2010). Nigeria – Bassa Pr. (Libert, 2010); Calabar (Libert, 2010); Oban (Libert, 2010); Owenna (Libert, 2010); Umuhaia (Libert, 2010); Uwet (Libert, 2010). Cameroon – Eningo (Plötz, 1880); Agoncho (Plötz, 1880); Victoria (Plötz, 1880); Rumpi Hills (Helps, vide Larsen, 2005a); Dja (Libert, 2010); Dek (Libert, 2010); Yaounde (Libert, 2010); Moloundou (Libert, 2010); Dalugene (Libert, 2010); Batouri (Libert, 2010). Equatorial Guinea – Benito (Libert, 2010). Gabon – Bitam (Libert, 2010); Ogooue (Libert, 2010); Fernan Vaz (Libert, 2010); Lastourville (Libert, 2010); Mimongo (Libert, 2010); Bateke Plateau (Libert, 2010); Langoue (Vande weghe, 2010); camp Nouna (Vande weghe, 2010); camp PPG (Vande weghe, 2010); Ekouyi (Vande weghe, 2010). Congo – Souanke (Libert, 2010); Brazzaville (Libert, 2010); Ngo (Libert, 2010). Central African Republic – Dzanga (Noss, 1998). Democratic Republic of Congo – Mbata Siala (Libert, 2010). Habitat: Forest. Habits: An uncommon forest butterfly (Larsen, 2005a). Larsen collected numbers at damp patches in Ghana (Collins et al., 2003). Early stages: Nothing published. Larval food: Nothing published.

maeander Plötz, 1880 (as sp. of Lycaenesthes). Stettiner Entomologische Zeitung 41: 202 (189-206). Cameroon: “Eningo; Agoncho; Victoria”. Considered a synonym of Anthene lusones lusones (Hewitson, 1874) by Larsen (2005a), syn. nov. Female co-type selected as lectotype. Considered to be a nomen dubium by Libert (2010: 255).

*Neurellipes onias (Hulstaert, 1924)

Lycaenesthes onias Hulstaert, 1924. Revue de Zoologie et de Botanique Africaine 12: 183 (112-122, 173-194). Anthene onias (Hulstaert, 1924). Ackery et al., 1995: 626. Anthene onias Hulstaert, 1924. d’Abrera, 2009: 796. Neurellipes onias (Hulstaert, 1924). Libert, 2010: 243 n. comb.

Type locality: [Democratic Republic of Congo]: “Région de Sasa”. Holotype (male), (Colmant); M.R.A.C., Tervuren. Distribution: Cameroon, Angola, Central African Republic, Democratic Republic of Congo, Uganda, Sudan (south), Tanzania (north-west). Specific localities: Cameroon – Mount Febe near Yaounde (Libert, 2010); Ebogo (Libert, 2010); Njore (Libert, 2010); Goyoum (Libert, 2010). Central African Republic – Bangui (Libert, 2010); Moloukou (Libert, 2010). Democratic Republic of Congo – Sasa Region (TL); Maluku near Kinshasa (Libert, 2010); Bombo

3 Lumene near Kinshasa (Libert, 2010); Bolongo near Mbandaka (Libert, 2010); Kapanga (Libert, 2010); Lusambo (Libert, 2010); Kindu (Libert, 2010); Kafakumba (Libert, 2010); Lupweshi (Libert, 2010); Sandoa (Libert, 2010). Uganda – Budongo (Libert, 2010); Bwamba (Libert, 2010); Kalinzu (Libert, 2010); Kayonza (Libert, 2010); Bugalla (Libert, 2010); Buvuma, Sesse Isles (Libert, 2010). Sudan – 4 05 N, 32 05 E (Libert, 2010); Congo-Nile divide (Libert, 2010); Didinga Mountains (Libert, 2010). Tanzania – Bulembe (Libert, 2010); Kikuru (Libert, 2010); Minziro (Libert, 2010). Early stages: Nothing published. Larval food: Nothing published.

kalinzu Stempffer, 1950 (as sp. of Neurypexima [sic]). Revue Française d’Entomologie 17: 143 (135- 149). Type locality: Uganda: “Kalinzu Forest, W. Ankole (Ouganda)”. Diagnosis: Similar to Anthene chryseostictus but underside paler and with distinct submarginal line on both wings, angled on hindwing (Congdon & Collins, 1998). Distribution: Uganda (west), Tanzania (north-west). Specific localities: Uganda – Kalinzu Forest (TL). Tanzania – Minziro Forest (rare) (Congdon & Collins, 1998). Habitat: Forest. Early stages: Nothing published. Larval food: Nothing published. Given as Neurypexina kalinzu Stempffer, 1950 by d’Abrera, 2009: 801. Formally synonymized with Neurellipes onias (Hulstaert, 1924) by Libert, 2010: 243.

*Neurellipes sonia Libert, 2010

Neurellipes sonia Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 244.

Type locality: Democratic Republic of Congo: Paulis, Uele. Holotype (male) 8 February 1960 (M. Fontaine); M.R.A.C., Tervuren. Distribution: Democratic Republic of Congo. Specific localities: Democratic Republic of Congo – Paulis, Uele (TL); Yopole (Libert, 2010); Lume (Libert, 2010); Avakubi (Libert, 2010); Beni (Libert, 2010); Mapimbi (Libert, 2010); Mangina (Libert, 2010); Mazumbi (Libert, 2010); Biakatu (Libert, 2010); Okapi Reserve near Epulu (Libert, 2010). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

*Neurellipes alixae Libert, 2010

Neurellipes alixae Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 245.

Type locality: Cameroon: N’kongsamba. Holotype (male) (H. Stempffer); M.N.H.N., Paris. Distribution: Cameroon, Gabon, Congo, Democratic Republic of Congo. Specific localities: Cameroon – N’kongsamba (TL); near Mount Manengouba (Libert, 2010); Bitje (Libert, 2010); Dja (Libert, 2010). Gabon – Ogooue (Libert, 2010); Fernan Vaz (Libert, 2010). Congo – Ketta Forest near Ouesso (Libert, 2010). Democratic Republic of Congo – N’gombe near Kinshasa (Libert, 2010); Lukolela (Libert, 2010). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published.

4 Associated ants: Nothing published.

*Neurellipes fulvimacula (Mabille, 1890) Medium Red-spot Hairtail

Lycaena fulvimacula Mabille, 1890. Annales de la Société Entomologique de France (6) 10: 24 (17-51). Synonym of Anthene lusones (Hewitson, 1874). Ackery et al., 1995. Anthene lusones fulvimacula Mabille, 1890. Collins et al., 2003: 94. Anthene lusones fulvimacula Mabille, 1890. Larsen, 2005: 242. Neurellipes fulvimacula (Mabille, 1890). Libert, 2010: 247 stat. rev.

Alternative common name: Medium Red-spot Ciliate Blue. Type locality: Ivory Coast: “Assinnie”. Neotype (male), Yapo Forest, Ivory Coast, 5 July 1998 (H. Warren-Gash); genitalia Libert 109-338; M.N.H.N., Paris. Neallotype (female), same data but genitalia Libert 109-338 (Libert, 2010). Distribution: Senegal, Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana (Collins et al., 2003), Nigeria, Togo, Cameroon, Gabon, Central African Republic, Democratic Republic of Congo, Uganda (west), Tanzania (north-west). Specific localities: Senegal – Sengalkam near Dakar (Libert, 2010). Guinea – Macenta (Libert, 2010); Nzerekore (Libert, 2010); Seredou (Libert, 2010). Sierra Leone – Baranga (Libert, 2010); Gola (Libert, 2010); Panguma (Libert, 2010); Bumbuna (Libert, 2010). Liberia – Yamein (Libert, 2010); Zorzor (Libert, 2010); Mission Franco Liberienne (Libert, 2010). Ivory Coast – Assinie (TL); Yapo Forest (Libert, 2010); Nimba Mountains (Libert, 2010). Ghana – Ankasa National Park (Collins et al., 2003); Bia (Libert, 2010); Bibiani (Libert, 2010); Kumasi (Libert, 2010); Tano Ofin (Libert, 2010); Sagamase (Libert, 2010); Boukro (Libert, 2010); Ho (Libert, 2010); Lipke (Libert, 2010); Kyabobo (Libert, 2010). Nigeria – Bassa Province (TL of chryseostictus); Umuhaia (Libert, 2010). Togo – Cameroon – Korup (Larsen, 2005a – as chryseostictus); Nyassoso (Libert, 2010); Nkongsamba (Libert, 2010); Yaounde (Libert, 2010); Nguila (Libert, 2010); Ebolowa (Libert, 2010); Lolodorf (Libert, 2010); Nazare near Sangmelima (Libert, 2010); Efulen (Libert, 2010); Kribi (Libert, 2010); Maan (Libert, 2010). Gabon – Fernan Vaz (Libert, 2010); Ivindo (Libert, 2010); Ogooue (Libert, 2010); Oyem (Libert, 2010); Ipassa (Vande weghe, 2010); Langoue (Vande weghe, 2010); Nouna (Vande weghe, 2010). Central African Republic – Bangui (Libert, 2010); Moloukou (Libert, 2010). Democratic Republic of Congo – Mbata Siala (Libert, 2010); Bambesa (Libert, 2010); Beni (Libert, 2010); Biakatu (Libert, 2010); Kasai district (Libert, 2010); Kapanga (Libert, 2010); Sandoa (Libert, 2010). Uganda – Semuliki N.P. (Davenport & Howard, 1996); Budongo (Libert, 2010); Bwamba (Libert, 2010); Kagera (Libert, 2010); Buddu (Libert, 2010); Katera (Libert, 2010); Lake Nagogabu (Libert, 2010). Tanzania – Bulembe, in Minziro Forest (Congdon & Collins, 1998 – as chryseostictus); Kikuru Forest (Congdon & Collins, 1998 – as chryseostictus); Bukoba (Libert, 2010). Habitat: Forest. Habits: A fairly common butterfly (Larsen, 2005a). Usually found on vegetation along the edges of forest or near water (Congdon & Collins, 1998). Both sexes are fond of settling on large green leaves in dense forest and appear to be somewhat inactive (Larsen, 2005a) [notes for the synonym chryseostictus]. Early stages: Nothing published. Larval food: Nothing published.

Note: D’Abrera (2009: 800) considers fulvimacula to be a synonym of lusones but does not formally emend its taxonomy.

5 chryseostictus Bethune-Baker, 1910 (as sp. of Neurellipes). Transactions of the Entomological Society of London 43: 67 (1-84). Type locality: Sierra Leone; Nigeria: “Bassa Province (N. Nigeria)”. Distribution: Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, Nigeria (south and Cross River loop), Cameroon, Congo, Democratic Republic of Congo (Mongala, Uele, Lualaba), Uganda, Tanzania (north-west). Specific localities: Nigeria – Bassa Province (TL). Cameroon – Korup (Larsen, 2005a). Tanzania – Bulembe, in Minziro Forest (Congdon & Collins, 1998); Kikuru Forest (Congdon & Collins, 1998). Common name: Medium Red-spot Ciliate Blue. Habitat: Forest. Habits: A fairly common butterfly (Larsen, 2005a). Usually found on vegetation along the edges of forest or near water (Congdon & Collins, 1998). Both sexes are fond of settling on large green leaves in dense forest and appear to be somewhat inactive (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published. Given as Neurellipes chryseostictus Bethune-Baker, 1910 by d’Abrera, 2009: 800. Formally synonymized with Neurellipes fulvimacula (Mabille, 1890) by Libert, 2010: 247.

*Neurellipes kampala (Bethune-Baker, 1910) Kampala Hairtail

Lycaenesthes kampala Bethune-Baker, 1910. Transactions of the Entomological Society of London 43: 52 (1-84). Anthene kampala (Bethune-Baker, 1910). Ackery et al., 2009: 796. Anthene kampala Bethune-Baker, 1910. d’Abrera, 2009: 796. Neurellipes kampala (Bethune-Baker, 1910). Libert, 2010: 248 n. comb.

Alternative common name: Kampala Ciliate Blue. Type locality: [Uganda]: “Kampala”. Holotype (male), April 1900 (F.J. Jackson); N.H.M., London. Neallotype (female), Entebbe, Uganda, 12-20 January 1912 (S.A. Neave); N.H.M., London (Libert, 2010). Distribution: Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria, Cameroon; Equatorial Guinea, Gabon, Congo, Central African Republic, Democratic Republic of Congo, Uganda. Habitat: Forest (Larsen, 2005a). Habits: Early stages: Nothing published. Larval food: Nothing published.

Neurellipes kampala kampala (Bethune-Baker, 1910)

Type locality: [Uganda]: “Kampala”. Holotype (male), April 1900 (F.J. Jackson); N.H.M., London. Neallotype (female), Entebbe, Uganda, 12-20 January 1912 (S.A. Neave); N.H.M., London (Libert, 2010). Distribution: Uganda. Specific localities: Uganda – Kampala (TL); Kamengo (Libert, 2010); Mabira (Libert, 2010); Msarosaro (Libert, 2010); Semuliki N.P. (H. Selb, unpublished, 2016).

Neurellipes kampala incerta Libert, 2010

Neurellipes kampala incerta Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 251.

Type locality: Ivory Coast: Tai Forest. Holotype (male) 2 January 2000 (A.E. Warren-Gash); genitalia Libert 109-336; M.N.H.N., Paris. Distribution: Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria, Cameroon; Equatorial Guinea,

6 Gabon, Congo, Central African Republic. Specific localities: Sierra Leone – Moyamba (Libert, 2010); Panguma (Libert, 2010). Liberia – Kpaine (Libert, 2010); Vaa (Libert, 2010). Ivory Coast – Tai Forest (TL); Anguededou (Libert, 2010); Douekoue (Libert, 2010); Gagnoa (Libert, 2010); Man (Libert, 2010); Mount Peko (Libert, 2010); Mount Tonkoui (Libert, 2010); Tai (Libert, 2010). Ghana – Accra (Libert, 2010); Akyem (Libert, 2010); Amedzofe (Libert, 2010); Bia (Libert, 2010); Hahoe (Libert, 2010); Mpasaso (Libert, 2010); Sagamase (Libert, 2010); Tano Ofin (Libert, 2010). Nigeria – Mkpot 1 in the Oban Hills (Larsen, 2005a); Lagos (Libert, 2010); Ibadan (Libert, 2010); Okomu (Libert, 2010); Ikom (Libert, 2010); Calabar (Libert, 2010). Cameroon – Mamfe (Libert, 2010); Maan (Libert, 2010); Bertoua (Libert, 2010); Moloundou (Libert, 2010). Equatorial Guinea – Alen (Libert, 2010); Bioko (Libert, 2010). Gabon – Fernan Vaz (Libert, 2010); Lambarene (Libert, 2010). Congo – Ouesso (Libert, 2010); Kelle (Libert, 2010). Central African Republic – Bangui (Libert, 2010); Moloukou (Libert, 2010).

Neurellipes kampala triangularis Libert, 2010

Neurellipes kampala triangularis Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 251.

Type locality: Uganda: Budongo. Holotype (male) July 1939 (T.H.E. Jackson); N.H.M., London. Allotype (female) same data but July 1939. Distribution: Central African Republic, Democratic Republic of Congo, Uganda. Specific localities: Central African Republic – Boukoko near Mbaiki (Libert, 2010); Bangui (Libert, 2010). Democratic Republic of Congo – Mbata Siala (Libert, 2010); Kinshasa (Libert, 2010); Eala (Libert, 2010); Kuma (Libert, 2010); Yakoma (Libert, 2010); Kisangani (Libert, 2010); Paulis (Libert, 2010); Beni (Libert, 2010); Lusambo (Libert, 2010); Tshiole (Libert, 2010); Kapanga (Libert, 2010). Uganda – Budongo (TL); Bugoma (Libert, 2010); Bwamba (Libert, 2010); Itwara (Libert, 2010); Toro (Libert, 2010).

*Neurellipes tanzaniensis Libert, 2010

Neurellipes tanzaniensis Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 252.

Type locality: Tanzania: Kele Hill, Minziro Forest, Budongo River. Holotype (male) 1250 m, 17 December 1994 (T.C.E. Congdon, M. Hussein); genitalia Libert 109-204; A.B.R.I., Nairobi. Distribution: Tanzania. Specific localities: Tanzania – Kele Hill, Minziro Forest (TL). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

*Neurellipes katera (Talbot, 1937) 7

Anthene katera Talbot, 1937. Transactions of the Royal Entomological Society of London 86: 68 (59-72). Anthene katera Talbot, 1937. Ackery et al., 1995: 623. Anthene katera Talbot, 1937. d’Abrera, 2009: 796. Neurellipes katera (Talbot, 1937). Libert, 2010: 252 n. comb.

Type locality: [Uganda]: “Katera”. Holotype (male), Katera, Uganda, November 1935 (T.H.E. Jackson); N.H.M., London. Neallotype (female), same data but August 1935 (Libert, 2010). Distribution: Democratic Republic of Congo (north-east), Uganda, Tanzania (north-west). Specific localities: Democratic Republic of Congo – Beni (Libert, 2010); Epulu (Libert, 2010); Mambasa (Libert, 2010); Virunga National Park (Libert, 2010); Kapanga (Libert, 2010); Katako-Kombe (Libert, 2010); Lodja (Libert, 2010); Kuma (Libert, 2010). Uganda – Katera (TL); Buddu (Libert, 2010). Tanzania – Kere Hill, and on the path between Minziro Village and Bulembe, in Minziro Forest (uncommon) (Congdon & Collins, 1998). Habitat: Forest. Habits: Males have been found perched on low vegetation near water as well as along forest paths (Congdon & Collins, 1998). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes lychnaptes (Holland, 1891) Red Forewing Hairtail

Lycaenesthes lychnaptes Holland, 1891. Psyche, a Journal of Entomology. Cambridge, Mass. 6: 51 (50-53). Anthene lychnaptes (Holland, 1891). Ackery et al., 1995: 625. Anthene lychnaptes Holland, 1891. d’Abrera, 2009: 796. Neurellipes lychnaptes (Holland, 1891). Libert, 2010: 253 n. comb.

Alternative common name: Red Forewing. Type locality: Gabon: “upper waters of the Ogové River”. Holotype (male), (A.C. Good); C.M.N.H., Pittsburgh (Libert, 2010). Distribution: Ivory Coast, Ghana, Nigeria, Cameroon, Gabon. Habitat: Wet forests (Larsen, 2005a – for fulvus). Habits: A very rare butterfly, usually found perched on a large green leaf at the side of a forest path (Larsen, 2005a – for fulvus). Early stages: Nothing published. Larval food: Nothing published.

Neurellipes lychnaptes lychnaptes (Holland, 1891)

Type locality: Gabon: “upper waters of the Ogové River”. Holotype (male), (A.C. Good); C.M.N.H., Pittsburgh (Libert, 2010). Distribution: Nigeria (east), Cameroon, Gabon. Specific localities: Nigeria – Oban Hills (Larsen, 2005a – for fulvus). Cameroon – Mamfe (Larsen, 2005a – TL for fulvus); Ebogo (Libert, 2010); Edea (Libert, 2010); Ekomtolo (Libert, 2010); Lolodorf (Libert, 2010). Gabon – Upper Ogove River (TL); Ndjole (Vande weghe, 2010).

lychnoptera Grose-Smith & Kirby, 1893 (as sp. of Lycaenesthes). In: Grose-Smith & Kirby,1892-7. Rhopalocera exotica, being illustrations of new, rare and unfigured species of butterflies 2: 97 (261 pp.). London. Gabon. An unjustified emendation of Lycaenesthes lychnaptes Holland.

fulvus Stempffer, 1962 (as sp. of Anthene (Neurellipes)). Bulletin de l’Institut Français d’Afrique Noire (A) 24: 1180 (1135-1181). Type locality: Cameroon: “Cameroun anglais, Mamfé”. The female appears to be unknown (Larsen, 2005a). Distribution: Ivory Coast, Ghana, Nigeria (south and Cross River loop), Cameroon (west). Specific localities: Ivory Coast – Banco (Larsen, 2005a); Tiassale (Larsen, 2005a); Yapo (Larsen, 2005a). Ghana – Kumasi (Larsen, 2005a); Atewa Range (Larsen, 2005a); Kakum (Larsen, 2005a); Ankasa (Larsen, 2005a). Nigeria – Oban Hills (Larsen, 2005a). Cameroon – Mamfe (TL). Common name: Red Forewing. Habitat: Wet forests (Larsen, 2005a). Habits: A very rare butterfly, usually found perched on a large green leaf at the side of a forest path (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published. Given as Neurellipes fulvus Stempffer, 1962 by d’Abrera, 2009: 800. Treated as a synonym of Neurellipes lychnaptes by Libert, 2010: 254.

Neurellipes lychnaptes anochreus Libert, 2010

Neurellipes lychnaptes anochreus Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 255.

Type locality: Ghana: Bia, western Ghana. Holotype (male) February-March, 2006 (ABRI coll.); A.B.R.I., Nairobi. Allotype (female) Tano Ofin, western Ghana, January, 2004 (ABRI coll.); A.B.R.I., Nairobi. Distribution: Ivory Coast, Ghana. Specific localities: Ivory Coast – Banco (Larsen, 2005a – for fulvus); Tiassale (Larsen, 2005a – for fulvus); Yapo (Larsen, 2005a – for fulvus); Abidjan (Larsen, 2010). Ghana – Bia (TL); Kumasi (Larsen, 2005a – for fulvus); Atewa Range (Larsen, 2005a – for fulvus); Kakum (Larsen, 2005a – for fulvus); Ankasa (Larsen, 2005a – for fulvus); Tano Ofin (Libert, 2010); Kibi (Larsen, 2010).

*Neurellipes lychnides (Hewitson, [1878]) Brown Hairtail

Lycaenesthes lychnides Hewitson, [1878]. In: Hewitson, [1862-78]. Illustrations of diurnal lepidoptera. Lycaenidae: 224 (1: 1-228; 2: 95 pp.; Supplement: 1-48). London. Anthene lychnides (Hewitson, 1878). Ackery et al., 1995: 625. Anthene lychnides Hewitson, 1878. d’Abrera, 2009: 796. Neurellipes lychnides (Hewitson, 1878). Libert, 2010: 257 n. comb.

Alternative common name: Brown Ciliate Blue. Type locality: Nigeria: “Old Calabar”. Holotype (male), genitalia Libert 106-819; N.H.M., London. Neallotype (female), ?Nigeria; N.H.M., London (Libert, 2010). Distribution: Nigeria (south), Cameroon (west). Specific localities: Nigeria – Calabar (TL); Oni (Libert, 2010). Cameroon – Itoki, Korup (Aurivillius, 1895); Funge (Libert, 2010); Mokundange (Libert, 2010); Mumyenge (Libert, 2010); Victoria (Libert, 2010); Bipindi (Libert, 2010). Habitat: Forest. Habits: This appears to be an exceedingly rare species, which has not been found in more than a century (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

9 *Neurellipes larseni Libert, 2010

Neurellipes larseni Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 258.

Type locality: Cameroon: “Ebogo, April 1984 (M. Libert)”. Holotype (male) M.N.H.N., Paris. Allotype female, Mount Kala, Cameroon, December 1986 (M. Libert); genitalia Libert 106-034; M.N.H.N., Paris. Distribution: Nigeria (east), Cameroon, Gabon. Specific localities: Nigeria – Mkpot (Libert, 2010). Cameroon – Ebogo (TL); Mount Kala (Libert, 2010); Mount Ngoekele near Yaounde (Libert, 2010); Dja (Libert, 2010); Sangmelima (Libert, 2010); Manyemen (Libert, 2010); Kienke River (Libert, 2010); Mbebe (Libert, 2010); Kribi (Libert, 2010); Bongola near Campo (Libert, 2010). Gabon – Lake Azingo (Libert, 2010); Ogooue (Libert, 2010). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

*Neurellipes cuypersi Libert, 2010

Neurellipes cuypersi Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 258.

Type locality: Democratic Republic of Congo: Lukolela, 7 December 2001 (F. Cuypers); genitalia Libert, 106-769. Holotype (male) in the M.R.A.C., Tervuren. Distribution: Congo, Democratic Republic of Congo (Equateur). Specific localities: Congo – Etoumbi (Libert, 2010). Democratic Republic of Congo – Lukolela (TL); Kalamba (Libert, 2010). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

*Neurellipes leptines (Hewitson, 1874) Leptines Hairtail

Lycaenesthes leptines Hewitson, 1874. Transactions of the Entomological Society of London 1874: 348 (343-355). Anthene leptines (Hewitson, 1874). Ackery et al., 1995: 624. Anthene leptines Hewitson, 1874. d’Abrera, 2009: 795. Neurellipes leptines (Hewitson, 1874). Libert, 2010: 261 n. comb.

Alternative common name: Leptines Ciliate Blue. Type locality: “Congo”. Holotype (male); N.H.M., London. Neallotype (female), Ebogo, Cameroon, February 1986 (M. Libert); genitalia Libert 107-168; M.N.H.N., Paris (Libert, 2010). Distribution: Cameroon, Equatorial Guinea, Gabon, Congo, Central African Republic, Democratic Republic of Congo (Equateur, Uele, Sankuru). Recorded, in error, from Nigeria (Libert, 2010). Habitat: Forest. Habits: A scarce butterfly (Larsen, 2005a).

10 Early stages: Nothing published. Larval food: Nothing published.

Neurellipes leptines leptines (Hewitson, 1874)

Type locality: “Congo”. Holotype (male); N.H.M., London. Neallotype (female), Ebogo, Cameroon, February 1986 (M. Libert); genitalia Libert 107-168; M.N.H.N., Paris (Libert, 2010). Distribution: Cameroon, Equatorial Guinea, Gabon, Congo. Specific localities: Cameroon – Owong (Aurivillius, 1923 – TL of arnoldi); Ebogo (Libert, 2010); Metet (Libert, 2010); Bitje (Libert, 2010); Dja (Libert, 2010); Lolodorf (Libert, 2010); Maan (Libert, 2010); Mintom II (Libert, 2010). Equatorial Guinea – Makomo, Campo district (Libert, 2010). Gabon – Ivindo (Libert, 2010); Oyem (Libert, 2010); Ogowe (Libert, 2010); Kongou (Vande weghe, 2010).

arnoldi Aurivillius, 1923. In: Schultze & Aurivillius, 1923 (as var. of Lycaenesthes leptines). Ergebnisse der Zweiten Deutschen Zentral-Afrika Expedition 1(17): 1236 (1113-1242). Cameroon: “Süd-Kamerun: Owöng”. Invalid; junior primary homonym of Lycaenesthes arnoldi Jones, 1918 [Lycaenidae].

Neurellipes leptines extensa Libert, 2010

Neurellipes leptines extensa Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 262.

Type locality: Democratic Republic of Congo: Middle Lowa Valley, near Walikali, 3500 feet, March 1924 (T.A. Barns). Holotype (male) in N.H.M., London. Allotype (female) same data but February. Distribution: Central African Republic, Democratic Republic of Congo. Specific localities: Central African Republic – Dzanga (Noss, 1998); Bangui (Libert, 2010). Democratic Republic of Congo – Middle Lowa Valley, near Walikali (TL); Embangobango (Libert, 2010); Itoko (Libert, 2010); Katako-Kombe (Libert, 2010); Kuma (Libert, 2010); Pania (Libert, 2010); Simba-Lokolenge (Libert, 2010); Yahuma (Libert, 2010); Epulu (Libert, 2010); Beni (Libert, 2010).

*Neurellipes makala (Bethune-Baker, 1910)

Lycaenesthes makala Bethune-Baker, 1910. Transactions of the Entomological Society of London 43: 58 (1-84). Anthene makala (Bethune-Baker, 1910). Ackery et al., 1995: 625. Anthene makala Bethune-Baker, 1910. d’Abrera, 2009: 795. Neurellipes makala (Bethune-Baker, 1910). Libert, 2010: 263 n. comb.

Type locality: [Democratic Republic of Congo]: “Makala”. Holotype (male), May 1906 (Powell- Cotton); N.H.M., London. Neallotype (female), Paulis, Uele, Democratic Republic of Congo, 16 January 1960; M.R.A.C., Tervuren (Libert, 2010). Distribution: Nigeria (east), Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo (Uele), Uganda. Specific localities:

11 Nigeria – Eko Esai (Libert, 2010); ?Oban Hills (Libert, 2010). Cameroon – Bekoko (Libert, 2010); Maan (Libert, 2010); Moloundou (Libert, 2010); Ebogo (Libert, 2010); Yaounde (Libert, 2010); Bitje (Libert, 2010). Gabon – Nouna (Vande weghe, 2010). Congo – Kelle (Libert, 2010); Sembe (Libert, 2010); Ketta (Libert, 2010). Central African Republic – Dzanga (Noss, 1998); Bangui (Libert, 2010); Boukoko - Mbaiki (Libert, 2010); Moloukou (Libert, 2010). Democratic Republic of Congo – Makala (TL); Kimuenza near Kinshasa (Libert, 2010); Bokote (Libert, 2010); Simba-Lokolenge (Libert, 2010); Lusambo (Libert, 2010); Sankuru (Libert, 2010); Paulis (Libert, 2010); Kamituga (Libert, 2010). Uganda – Bwamba (Libert, 2010). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes rufomarginata (Bethune-Baker, 1910)

Lycaenesthes rufomarginata Bethune-Baker, 1910. Transactions of the Entomological Society of London 43: 54 (1-84). Anthene rufomarginata (Bethune-Baker, 1910). Ackery et al., 1995: 627. Anthene rufomarginata Bethune-Baker, 1910. d’Abrera, 2009: 796. Neurellipes rufomarginata (Bethune-Baker, 1910). Libert, 2010: 264 n. comb.

Type locality: [Democratic Republic of Congo]: “Makala (Congo Free State); Lake Albert Edward”. Lectotype (male), Makala, Democratic Republic of Congo, June 1906 (Powell-Cotton); N.H.M., London. Paralectotype (female), ?Lake Albert Edward, Democratic Republic of Congo, March, N.H.M., London (Libert, 2010). Distribution: Democratic Republic of Congo (Ituri, north-eastern Tshopo). Specific localities: Democratic Republic of Congo – Makala (TL); Lake Albert Edward (Bethune-Baker, 1910); Kisangani (Libert, 2010); Banalia (Libert, 2010); Medje (Libert, 2010); Beni (Libert, 2010); Mapimbi (Libert, 2010). Habitat: Forest. Habits: Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes ngoko (Stempffer, 1962)

Anthene ngoko Stempffer, 1962. Bulletin de l’Institut Français d’Afrique Noire (A) 24: 1174 (1135-1181). Anthene ngoko Stempffer, 1962. Ackery et al., 1995: 626. Anthene ngoko Stempffer, 1962. d’Abrera, 2009: 796. Neurellipes ngoko (Stempffer, 1962). Libert, 2010: 265 n. comb.

Neurellipes ngoko ngoko (Stempffer, 1962)

12 Anthene ngoko Stempffer, 1962. Bulletin de l’Institut Français d’Afrique Noire (A) 24: 1174 (1135-1181). Anthene ngoko Stempffer, 1962. Ackery et al., 1995: 626. Anthene ngoko Stempffer, 1962. d’Abrera, 2009: 796. Neurellipes ngoko ngoko (Stempffer, 1962). Libert, 2010: 265 n. comb.

Type locality: Congo: “Ouesso, forêt de Ketta”. Holotype (male), Ketta Forest, Ouesso, Congo, October 1959 (T.H.E. Jackson); genitalia Stempffer 5389; N.H.M., London. Neallotype (female), Ebogo, Cameroon, February 1989 (M. Libert); genitalia Libert 107-172; M.N.H.N., Paris (Libert, 2010). Distribution: Cameroon, Congo (north), Central African Republic. Specific localities: Cameroon – Mount Kala (Libert, 2010); Ebogo (Libert, 2010); Meyila (Libert, 2010); Nkoloveng (Libert, 2010); Yaounde (Libert, 2010); Metet (Libert, 2010); Bitje (Libert, 2010); Mintom II (Libert, 2010). Congo – Ketta Forest, Ouesso (TL); Sembe (Libert, 2010). Central African Republic – Bangui (Libert, 2010); Moloukou (Libert, 2010); Yakoli (Libert, 2010).

Neurellipes ngoko fontainei Libert, 2010

Neurellipes ngoko fontainei Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 266.

Type locality: Democratic Republic of Congo: Embangobango, Sankuru, 5 August 1950 (Dr Fontaine); genitalia Libert 106-399. Holotype (male) in the M.R.A.C., Tervuren. Distribution: Democratic Republic of Congo. Specific localities: Democratic Republic of Congo – Embangobango, Sankuru (TL).

*Neurellipes radiata (Bethune-Baker, 1910) Red-edged Hairtail

Lycaenesthes radiata Bethune-Baker, 1910. Transactions of the Entomological Society of London 43: 55 (1-84). Anthene radiata (Bethune-Baker, 1910). Ackery et al., 1995: 626. Anthene radiata Bethune-Baker, 1910. d’Abrera, 2009: 796. Neurellipes radiata (Bethune-Baker, 1910). Libert, 2010: 267 n. comb.

Neurellipes radiata. Male, Ghana. Left – upperside; right – underside. Images courtesy Torben Larsen.

Neurellipes radiata. Female, Ghana, upperside. Images courtesy Torben Larsen.

Alternative common name: Red-edged Ciliate Blue. Type locality: Sierra Leone. Holotype (male), Tani, Sierra Leone, 2 July 1903; N.H.M., London. Neallotype (female), St Paul River, Zorzor Road, Liberia, 18 May 1965 (R.M. Fox); C.M.N.H., Pittsburgh (Libert, 2010). Distribution: Sierra Leone, Liberia (north-west), Ivory Coast (Warren-Gash, pers. comm., 2002), Ghana. Specific localities: Sierra Leone – Tani (TL); Gola (Libert, 2010); near Freetown (Libert, 2010). Liberia – Zorzor (Larsen, 2005a). Ivory Coast – Alepe in eastern Ivory Coast, on the Comoe River (Collins et al., 2003); Tai forest (Larsen, 2005a); Blolequin (Libert, 2010); Douekoue (Libert, 2010). Ghana – Takoradi (Maessen, vide Larsen, 2005a). Habitat: Moist primary forest (Larsen, 2005a). Habits: A rare, local, species (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes scintillula (Holland, 1891) Golden Hairtail

Lycaenesthes scintillula Holland, 1891. Psyche, a Journal of Entomology. Cambridge, Mass.6: 50 (50-53). Anthene scintillula (Holland, 1891). Ackery et al., 1995: 627. Anthene scintillula Holland, 1891. d’Abrera, 2009: 794. Neurellipes scintillula (Holland, 1891). Libert, 2010: 268 n. comb.

Neurellipes scintillula scintillula. Male. Left – upperside; right – underside. Ipassa, Ivindo N.P., Gabon. 18 June 2016. J. Dobson. Images M.C. Williams ex Dobson collection.

14 Alternative common name: Golden Ciliate Blue. Type locality: Gabon: “upper waters of the Ogové River” [false locality]. Holotype (male), Bule Country, Cameroon; C.M.N.H., Pittsburgh. Neallotype (female), Sembe, Souanke district, Congo, March 1960 (T.H.E. Jackson); N.H.M., London (Libert, 2010). Distribution: Sierra Leone, Ivory Coast, Ghana, Nigeria, Cameroon, Equatorial Guinea, Gabon, Congo, Democratic Republic of Congo, Uganda. Habitat: Forest. Habits: A generally scarce species, especially in West Africa (Larsen, 2005a). Males are known to defend territories from perches on large green leaves. The flight is moderately fast (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

Neurellipes scintillula scintillula (Holland, 1891)

Lycaenesthes scintillula Holland, 1891. Psyche, a Journal of Entomology. Cambridge, Mass.6: 50 (50-53). Anthene scintillula scintillula (Holland, 1891). Ackery et al., 1995: 627. Anthene scintillula scintillula Holland, 1891. d’Abrera, 2009: 794. Neurellipes scintillula scintillula (Holland, 1891). Libert, 2010: 268.

Neurellipes scintillula scintillula. Male. Left – upperside; right – underside. Ipassa, Ivindo N.P., Gabon. 18 June 2016. J. Dobson. Images M.C. Williams ex Dobson collection.

Type locality: Gabon: “upper waters of the Ogové River” [false locality]. Holotype (male), Bule Country, Cameroon; C.M.N.H., Pittsburgh. Neallotype (female), Sembe, Souanke district, Congo, March 1960 (T.H.E. Jackson); N.H.M., London (Libert, 2010). Distribution: Nigeria (east), Cameroon, Equatorial Guinea, Gabon, Congo, Democratic Republic of Congo (Mongala, Uele, Tshopo, Equateur, Sankuru, Maniema), Uganda (west). Specific localities: Nigeria – Oban Hills (Larsen, 2005a). Cameroon – Ntale (Libert, 2010); Barombi (Libert, 2010); Maan (Libert, 2010); between Yokadouma and Moloundou (Libert, 2010); Yaounde (Libert, 2010); Ebogo (Libert, 2010). Equatorial Guinea – Alen (Libert, 2010); Benito (Libert, 2010). Gabon – Upper Ogove River (TL); Ivindo National Park (Libert, 2010); Oyem (Libert, 2010); Akaka (Vande weghe, 2010); Langoue (Vande weghe, 2010); Kongou (Vande weghe, 2010); camp Nouna (Vande weghe, 2010). Congo – Sembe (Libert, 2010); Kelle (Libert, 2010); Goumina (Libert, 2010). Democratic Republic of Congo – Lukolela (Libert, 2010); Abumombazi (Libert, 2010); Luebo (Libert, 2010); Lusambo - Pania (Libert, 2010); Kivu (Libert, 2010); Mawambi (Libert, 2010); Kansenye (Libert, 2010); Irangi (Libert, 2010). Uganda – Semuliki N.P. (Davenport & Howard, 1996); Bwamba (Libert, 2010); Bugoma (Libert, 2010).

15 Neurellipes scintillula aurea (Bethune-Baker, 1910)

Lycaenesthes aurea Bethune-Baker, 1910. Transactions of the Entomological Society of London 43: 59 (1-84). Anthene aurea (Bethune-Baker, 1910). Ackery et al., 1995: 621. Anthene scintillula aurea Bethune-Baker, 1901 [sic]. Larsen & Collins, 1998: 374. Anthene scintillula aurea Bethune-Baker, 1910. d’Abrera, 2009: 794. Neurellipes scintillula aurea (Bethune-B., 1910). Libert, 2010: 271 n. comb.

Type locality: Ghana: “Begoro”. Holotype (male), Begoro, 1500 ft, Ghana, March 1906 (G.C. Dudgeon); N.H.M., London. Neallotype (female), Bia National Park, western Ghana, March 2003 (ABRI collection); A.B.R.I., Nairobi (Libert, 2010). Distribution: Sierra Leone, Ivory Coast, Ghana, Nigeria (west). Specific localities: Sierra Leone – Mabang (Libert, 2010); Moyamba (Libert, 2010); Gola (Libert, 2010). Ivory Coast – Yapo (Plowes, vide Larsen, 2005a); Tai Forest (Warren-Gash, vide Larsen, 2005a). Ghana – Begoro (TL); Atewa Range (Maessen, vide Larsen, 2005a); Bia National Park (ABRI, vide Larsen, 2005a); Ho (Libert, 2010); Kibi (Libert, 2010). Nigeria – Sapoba in the Niger Delta (Larsen, 2005a).

*Neurellipes helpsi (Larsen, 1994) Creamy Hairtail

Anthene helpsi Larsen, 1994. Lambillionea 94 (4) (Tome II): 550 (550-552). Anthene helpsi Larsen, 1994. d’Abrera, 2009: 795. Neurellipes helpsi (Larsen, 1994). Libert, 2010: 272 n. comb.

Neurellipes helpsi. Male (holotype), Ghana. Left – upperside; right – underside. Images courtesy Torben Larsen.

Alternative common name: Creamy Ciliate Blue. Type locality: Ghana: “Atewa Range, Kibi, Ghana, 9. I. 1993 (T.G.P. Helps leg.) (In the Natural History Museum, London).” Distribution: Guinea, ?Ivory Coast, Ghana. Specific localities: Guinea – Ziama Forest (Libert, 2010); Seredou (Libert, 2010). Ivory Coast – Banco Forest near Abidjan (Belcastro, vide Larsen, 2005a; record requires confirmation). The single male from Côte d’Ivoire came to Belcastro by a complicated process and given the amount of collecting since undertaken there and in Ghana, the species is probably endemic to the Atewa Range in Ghana (Larsen, pers. comm. 2007). Ghana – Kibi, Atewa Range (TL). Habitat: Dense upland evergreen forest (Larsen, 2005a). Habits: This appears to be a rare and localized species (Larsen, 2005a).

16 Early stages: Nothing published. Larval food: Nothing published.

Note: Libert (2010) states that the population in Guinea probably represents an undescribed subspecies of helpsi.

*Neurellipes bakeri (Druce, 1910)

Lycaenesthes bakeri Druce, 1910. Proceedings of the Zoological Society of London 1910: 374 (356-378). Anthene bakeri (Druce, 1910). Ackery et al., 1995: 621. Anthene bakeri Druce, 1910. d’Abrera, 2009: 796. Neurellipes bakeri (Druce, 1910). Libert, 2010: 272 n. comb.

Type locality: [Democratic Republic of Congo]: “Upper Kasai district”. Holotype (female), (Landbeck); N.H.M., London (Libert, 2010). Known, with certainty, only from the female holotype (Libert, 2010). Distribution: Democratic Republic of Congo. Specific localities: Democratic Republic of Congo – Upper Kasai district (TL). Early stages: Nothing published. Larval food: Nothing published.

ja Bethune-Baker, 1910 (as sp. of Lycaenesthes). Transactions of the Entomological Society of London 43: 58 (1-84). Cameroon: “Bitje; Ja River”. [False locality].

*Neurellipes tessmanni Libert, 2010

Neurellipes tessmanni Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 273.

Type locality: Equatorial Guinea: Benito, 16-31 August 1906 (G. Tessmann). Holotype (male) in the Z.M.H.U., Berlin. Distribution: Cameroon, Equatorial Guinea, Democratic Republic of Congo. Specific localities: Cameroon – Ebogo (Libert, 2010). Equatorial Guinea – Benito (TL). Democratic Republic of Congo – Simba-Lokolenge (Libert, 2010). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

*Neurellipes discimacula (Joicey & Talbot, 1921)

Lycaenesthes discimacula Joicey & Talbot, 1921. Bulletin of the Hill Museum, Witley 1: 96 (40-166). Anthene discimacula (Joicey & Talbot, 1921). Ackery et al., 1995: 622. Anthene discimacula Joicey & Talbot, 1921. d’Abrera, 2009: 794. Neurellipes discimacula (Joicey & Talbot, 1921). Libert, 2010: 274 n. comb.

Type locality: [Democratic Republic of Congo]: “between Epule and Duye Rivers, N. Ituri Valley”. Holotype (male), March 1920 (T.A. Barns); N.H.M., London. Neallotype (female), Beni, Kivu, Democratic Republic of Congo, 8 February 1991 (R. Ducarme); collection Ducarme (Libert, 2010). Distribution: Gabon, Democratic Republic of Congo (Ituri, Tshopo, Sankuru). Specific localities:

17 Gabon – Mpassa, Bateke Plateau (Libert, 2010); camp PPG (Vande weghe, 2010). Democratic Republic of Congo – Between Epule and Duye Rivers (TL); Beni (Libert, 2010); Epulu (Libert, 2010); Kisangani (Libert, 2010); Lusambo (Libert, 2010); Pania (Libert, 2010); Kasai River (Libert, 2010). Habitat: Habits: Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes mahota (Grose-Smith, 1887) Mahota Hairtail

Lycaenesthes mahota Grose-Smith, 1887. Annals and Magazine of Nateural History (5) 19: 65 (62-66). Anthene mahota (Grose-Smith, 1887). Ackery et al., 1995: 625. Anthene mahota Grose-Smith, 1887. d’Abrera, 2009: 794. Neurellipes mahota (Grose-Smith, 1887). Libert, 2010: 275 n. comb.

Alternative common name: Mahota Ciliate Blue. Type locality: [West Africa]: “Delagoa Bay”. [False locality]. Holotype (male), ?Cameroon; N.H.M., London (Libert, 2010). Distribution: Nigeria (Cross River loop), Cameroon, Gabon, Congo. Erroneously recorded from South Africa by Trimen & Bowker, 1889 (MCW). Recorded, in error, from Liberia, Ivory Coast, Equatorial Guinea and Democratic Republic of Congo (Libert, 2010). Specific localities: Nigeria – Oban Hills (Larsen, 2005a); Aningege (Libert, 2010); Calabar (Libert, 2010); Ikom (Libert, 2010); Uwet (Libert, 2010). Cameroon – Korup (Larsen, 2005a); Ebogo (Libert, 2010); Yaounde (Libert, 2010); Dja (Libert, 2010); Mintom II (Libert, 2010); Moloundou (Libert, 2010). Gabon – Andok Foula (Libert, 2010); Ogooue (Libert, 2010); Fougamou (Libert, 2010); Kinguele (Vande weghe, 2010); Langoue (Vande weghe, 2010); Kongou (Vande weghe, 2010); camp Nouna (Vande weghe, 2010). Congo – Kelle (Libert, 2010); Sembe (Libert, 2010); Ketta (Libert, 2010). Habitat: Dense forest (Larsen, 2005a). Habits: Specimens are found in ones and twos, fluttering weakly along forest paths, and frequently

18 settling (Larsen, 2005a). The slow flight of the ‘red’ anthenes is probably an indication that they are part of a mimicry complex (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes gola Libert, 2010

Neurellipes gola Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 277.

Type locality: Sierra Leone: Lalehun, Gola North, 1-5 December 2008 (ABRI coll.); genitalia Libert 109-197. Holotype (male) in A.B.R.I., Nairobi. Allotype (female), same data as holotype. Distribution: Sierra Leone, Liberia, Ivory Coast. Specific localities: Sierra Leone – Lalehun, Gola North (TL). Liberia – Yamein (Libert, 2010). Ivory Coast – Koadguezon (Libert, 2010). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

*Neurellipes ducarmei Libert, 2010

Neurellipes ducarmei Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 279.

Type locality: Democratic Republic of Congo: Mangina, April 1994 (R. Ducarme); genitalia Libert 107-674. Holotype (male) in M.R.A.C., Tervuren. Distribution: Cameroon, Gabon, Congo, Democratic Republic of Congo. Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

Neurellipes ducarmei ducarmei Libert, 2010

Neurellipes ducarmei ducarmei Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 279.

Type locality: Democratic Republic of Congo: Mangina, April 1994 (R. Ducarme); genitalia Libert 107-674. Holotype (male) in M.R.A.C., Tervuren. Distribution: Democratic Republic of Congo. Specific localities: Democratic Republic of Congo – Mangina (TL); Beni (Libert, 2010); Mambasa (Libert, 2010); Irangi (Libert, 2010); Bafwasende (Libert, 2010); Kisangani (Libert, 2010); Molegbwe (Libert, 2010); Embangobango near Lusambo (Libert, 2010).

Neurellipes ducarmei occidentalis Libert, 2010

Neurellipes ducarmei occidentalis Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 280.

Type locality: Cameroon: Ebogo, February 1985 (M. Libert). Holotype (male) in M.N.H.N., Paris. Distribution: Cameroon, Gabon, Congo. Specific localities: Cameroon – Ebogo (TL); Mbankomo (Libert, 2010); Meyila (Libert, 2010); Nkoloveng (Libert, 2010); Yaounde (Libert, 2010); Sangmelima (Libert, 2010); Mintom II (Libert, 2010); Moloundou (Libert, 2010). Gabon – Talaguga (Libert, 2010). Congo – Kelle (Libert, 2010); Ketta Forest (Libert, 2010); Ouesso (Libert, 2010).

*Neurellipes georgiadisi (Larsen, 2009)

Anthene georgiadisi Larsen, 2009. Entomologist’s Record and Journal of Variation 121 (1): 49 (47-51). Anthene georgiadisi Larsen, 2009. d’Abrera, 2009: 794. Neurellipes georgiadisi (Larsen, 2009). Libert, 2010: 280 n. comb.

Nurellipes georgiadisi holotype male. Images courtesy of Torben Larsen.

Type locality: Liberia: “Sapo National Park”. Holotype (male), 5°24'40"N, 8°24'53"W, February 2008 (S. Georgiadis); genitalia Libert 108-079; A.B.R.I., Nairobi. Distribution: Liberia. Specific localities: Liberia – Sapo National Park (TL). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes rhoko Safian, 2014

Anthene rhoko Safian, 2014. Zootaxa 3860: 196 (195-200).

Type locality: Nigeria: “Cercopan Research and Conservation Area, Rhoko Forest, Iko Esai, Cross

20 River Loop. 18.V.2012. Leg. Szabolcs Safian. Coordinates: 5°39'27.20"N, 8°15'42.16"E. Holotype male in A.B.R.I., Nairobi. Distribution: Nigeria, Cameroon. Specific localities: Nigeria – Rhoko Forest (TL). Cameroon – Ebogo (Safian, 2014). Habitat: Evergreen Forest (Safian, 2014). Habits: Found on forest edges, perching low down. When disturbed they fly quite weakly for a few metres, before resettling. It is quite probably a canopy species (Safian, 2014). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes pyroptera (Aurivillius, 1895)

Lycaenesthes pyroptera Aurivillius, 1895. Entomologisches Nachrichten. Berlin 21: 382 (379-382). Anthene pyroptera (Aurivillius, 1895). Ackery et al., 1995: 626. Anthene pyroptera Aurivillius, 1895. d’Abrera, 2009: 795. Neurellipes pyroptera (Aurivillius, 1895). Libert, 2010: 282 n. comb.

Type locality: [Democratic Republic of Congo]: “Banana (Congo)” [False locality (Libert, 2010)]. Holotype (male), Lualabourg, Democratic Republic of Congo; M.R.A.C., Tervuren. Neallotype (female), Etoumbi, Congo, November 1962 (T.H.E. Jackson); M.N.H.N., Paris (Libert, 2010). Distribution: Cameroon, Gabon, Congo, Angola, Central African Republic, Democratic Republic of Congo, Uganda, Recorded, in error, from Tanzania (as leptala = pyroptera). Specific localities: Cameroon – Ebodje (Libert, 2010); Kribi (Libert, 2010); Lena (Libert, 2010); Adamaooua (Libert, 2010); Yaounde (Libert, 2010); Sangmelima (Libert, 2010); Maan (Libert, 2010); Goyoum (Libert, 2010); Bertoua (Libert, 2010). Gabon – Bateke Plateau (Libert, 2010); Ipassa (Libert, 2010); Oyem (Libert, 2010); camp PPG (Vande weghe, 2010); Leke (Vande weghe, 2010); Franceville (Vande weghe, 2010). Congo – Etoumbi (Libert, 2010); Fort Rousset = Owando (Libert, 2010); Kelle (Libert, 2010); Ketta (Libert, 2010); Mambili (Libert, 2010); Ngo (Libert, 2010). Angola – Quango (Libert, 2010). Central African Republic – Dzanga (Noss, 1998); Bangui (Libert, 2010). Democratic Republic of Congo – Lualabourg (TL); Isangila (Libert, 2010); Albert National Park (Libert, 2010); Abumonbazi (Libert, 2010); Kapanga (Libert, 2010). Uganda – Bunyoro (Libert, 2010). Habitat: Forest. Early stages: Nothing published. Larval food: Nothing published.

leptala Courvoisier, 1914 (as sp. of Lycaenesthes). In: Strand, 1914. Lepidoptera Niepeltiana [1]: 43 (1-64). Type locality: [Tanzania]: “D.-W.-Afrika”. Kielland (1990d) does not include this species in his book on the butterflies of Tanzania, nor do Congdon & Collins (1998) list it in the supplement to Kielland’s book. D’Abrera (2009: 796) treats leptala as a good species but avers that it may be a synonym of pyroptera (Aurivillius, 1895). Libert (2010: 282) treats it as a synonym of pyroptera (Aurivillius, 1895).

*Neurellipes erythropoecilus (Holland, 1893) Curious Hairtail

Lycaenesthes erythropoecilus Holland, 1893. Entomological News 4: 26 (22-28). Anthene erythropoecilus (Holland, 1893). Ackery et al., 1995: 622. Anthene erythropoecilus Holland, 1893. d’Abrera, 2009: 795. Neurellipes erythropoecilus (Holland, 1893). Libert, 2010: 284 n. comb.

Alternative common name: Curious Ciliate Blue. Type locality: Gabon: “Talaguga, Ogové Valley”. Holotype (female); genitalia Libert 106-935; C.M.N.H., Pittsburgh. Neallotype (male), Benito, Equatorial Guinea, January 1897; C.M.N.H., Pittsburgh (Libert, 2010). Distribution: Ghana, Togo, Nigeria, Cameroon, Equatorial Guinea, Gabon, Congo, Central African Republic, Democratic Republic of Congo. Habitat: Nothing published. Habits: A scarce butterfly, usually found in ones and twos while mud-puddling or while defending territories on the edges of forest or along forest paths (Larsen, 2005a). The flight is quite weak (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

Note: D’Abrera (2009: 794) avers that erythropoecilus is a synonym of scintillula but makes no formal taxonomic changes.

Neurellipes erythropoecilus erythropoecilus (Holland, 1893)

Type locality: Gabon: “Talaguga, Ogové Valley”. Holotype (female); genitalia Libert 106-935; C.M.N.H., Pittsburgh. Neallotype (male), Benito, Equatorial Guinea, January 1897; C.M.N.H., Pittsburgh (Libert, 2010). Distribution: Cameroon, Equatorial Guinea, Gabon, Congo. Specific localities: Cameroon – Campo (Libert, 2010); Londji (Libert, 2010); Batanga (Libert, 2010); Bule Country (Libert, 2010); Efulen (Libert, 2010); Maan (Libert, 2010). Equatorial Guinea – Benito (Libert, 2010). Gabon – Talaguga, Ogove Valley (TL); Libreville (Libert, 2010); Ogooue (Libert, 2010); Kinguele (Vande weghe, 2010). Congo – Kuilu (Libert, 2010).

Neurellipes erythropoecilus locuples (Grose-Smith, 1898)

Lycaenesthes locuples Grose-Smith, 1898. Novitates Zoologicae 5: 357 (350-358). Anthene locuples (Grose-Smith, 1898). Ackery et al., 1995: 625. Anthene locuples Grose-Smith, 1898. d’Abrera, 2009: 795. Neurellipes erythropoecilus locuples (Gr.-Sm., 1898). Libert, 2010: 285 comb. n.

Type locality: Nigeria: “Warri”. Holotype (male), February 1896; N.H.M., London. Neallotype (female), Bitje, Ja River, Cameroon, April-June 1910 (G.L. Bates); N.H.M., London (Libert, 2010). Distribution: Ghana, Togo, Nigeria (western Nigeria, Cross River loop), Cameroon, Congo, Central African Republic, Democratic Republic of Congo (Kinshasa). Specific localities: Ghana – Kukurantumi (Larsen, 2005a); Likpe (Larsen, 2005a); Wli Falls (Larsen, 2005a); Bobiri Butterfly Sanctuary (Larsen et al., 2007); Bia (Libert, 2010); Anfoega (Libert, 2010); Ankasa (Libert, 2010); Mparaeso (Libert, 2010); Mpasaso (Libert, 2010). Togo – Kloto (Libert, 2010). Nigeria – Warri (TL); Agege (Larsen, 2005a); Ikom (Larsen, 2005a); Oban Hills (Larsen, 2005a); Okomu (Libert, 2010); Sapoba (Libert, 2010). Cameroon – Korup (Larsen, 2005a); Mamfe (Libert, 2010); Abong Mbang (Libert, 2010); Akom (Libert, 2010); Maan (Libert, 2010). 22 Congo – Kelle (Libert, 2010); Odzala (Libert, 2010). Central African Republic – Moloukou (Libert, 2010).; Boukoko (Libert, 2010); Botambi near Bangui (Libert, 2010). Democratic Republic of Congo – Kinshasa (Ackery et al., 1995).

*Neurellipes juba (Fabricius, 1787) Anomalous Hairtail

Papilio juba Fabricius, 1787. Mantissa Insectorum 2: 82 (382 pp.). Hafniae. Anthene juba (Fabricius, 1787). Ackery et al., 1995: 623. Anthene juba Fabricius, 1787. d’Abrera, 2009: 796. Neurellipes juba (Fabricius, 1787). Libert, 2010: 286 n. comb.

Alternative common name: Anomalous Ciliate Blue. Type locality: Sierra Leone: “Sierra Leon Africae”. Holotype (male); Zoological Museum, University of Copenhagen. Neallotype (female), Sierra Leone (Preuss); Z.M.H.U., Berlin (Libert, 2010). Distribution: Guinea, Sierra Leone, Liberia, Ivory Coast (Warren-Gash, pers. comm., 2002), Ghana, Togo, Nigeria (east), Cameroon (extreme west). Misattributed to the Togo fauna by Carcasson (1981) (Larsen, 2005a). Specific localities: Guinea – Fouta Djalon (Larsen, 2005a); Nimbas (Larsen, 2005a); Dubreka near Conakry (Libert, 2010); Mamou (Libert, 2010); Labe (Libert, 2010); Macenta (Libert, 2010). Sierra Leone – Freetown (Libert, 2010); Baranga (Libert, 2010); Bumbuna (Libert, 2010); Gola (Libert, 2010). Liberia – Kpaine (Libert, 2010); Wanau (Libert, 2010). Ivory Coast – Alepe (Larsen, 2005a). Ghana – Atewa Range (Maessen, vide Larsen, 2005a; single male); Ankasa (Libert, 2010); Kakum (Libert, 2010); Kibi (Libert, 2010); Tano Ofin (Libert, 2010). Togo – no locality (Libert, 2010). Nigeria – Bonny (Larsen, 2005a). Cameroon – no locality (Libert, 2010). Habitat: Primary, good quality, forest (Larsen, 2005a). Habits: A scarce butterfly, with the usual slow flight of the ‘red’ anthenes (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

dewitzi Staudinger, 1891 (as sp. of Lycaenesthes). Deutsche Entomologische Zeitschrift, Iris 4: 155 (61- 157). Sierra Leone.

*Neurellipes zenkeri (Karsch, 1895)

Lycaenesthes zenkeri Karsch, 1895. Entomologische Nachrichten. Berlin 21: 293 (289-322). Anthene zenkeri (Karsch, 1895). Ackery et al., 1995: 628. Anthene zenkeri Karsch, 1895. d’Abrera, 2009: 796. Neurellipes zenkeri (Karsch, 1895). Libert, 2010: 288 n. comb.

Neurellipes zenkeri zenkeri (Karsch, 1895)

Lycaenesthes zenkeri Karsch, 1895. Entomologische Nachrichten. Berlin 21: 293 (289-322). Anthene zenkeri Karsch, 1895. d’Abrera, 2009: 796. Anthene zenkeri (Karsch, 1895). Ackery et al., 1995: 628. Neurellipes zenkeri zenkeri (Karsch, 1895). Libert, 2010: 288.

Type locality: Cameroon: “Yaúnde-Station”. Holotype (male), (S. Zenker); Z.M.H.U., Berlin. Neallotype (female), Ebogo, Nyong River, Cameroon, July 1995 (S. Collins); A.B.R.I., Nairobi (Libert, 2010). Distribution: Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo (Uele, Tshopo), Uganda (west). Specific localities: Cameroon – Yaounde Station (TL); Yakaduma Forest (Schultze & Aurivillius, 1923); Maan (Libert, 2010); Mebassa (Libert, 2010). Gabon – Nouna near Belinga (Libert, 2010); Waka (Vande weghe, 2010); Langoue (Vande weghe, 2010). Congo – Etoumbi (Libert, 2010); Ketta (Libert, 2010); Sembe (Libert, 2010). Central African Republic – Moloukou (Libert, 2010); Mbata (Libert, 2010); Bangui (Libert, 2010). Democratic Republic of Congo – Lukolela (Libert, 2010); Mbandaka (Libert, 2010); Budjala (Libert, 2010); Kuma (Libert, 2010); Kisangani (Libert, 2010); Bafwasende (Libert, 2010); Paulis (Libert, 2010); Beni (Libert, 2010); Sankuru (Libert, 2010); Bena Dibele (Libert, 2010). Uganda – Bwamba (Libert, 2010).

connexa Aurivillius, 1923. In: Schultze & Aurivillius, 1923 (as ab. of Lycaenesthes zenkeri). Ergebnisse der Zweiten Deutschen Zentral-Afrika Expedition 1(17): 1237 (1113-1242). Cameroon: “Yakaduma-Urwald”.

Neurellipes zenkeri vandeweghei Libert, 2010

Neurellipes zenkeri vandeweghei Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 289.

Type locality: Gabon: Langoue, P.N. Ivindo, 20 November 2005 (G. Vande weghe). Holotype (female) in the M.R.A.C., Tervuren. Distribution: Gabon. Specific localities: Gabon – Langoue (TL).

*Neurellipes lachares (Hewitson, [1878]) Silky Hairtail

Lycaenesthes lachares Hewitson, [1878]. In: Hewitson, [1862-78]. Illustrations of diurnal lepidoptera. Lycaenidae : 225 (1: 1-228; 2: 95 pp.; Supplement: 1-48.). London. Anthene lachares (Hewitson, 1878). Ackery et al., 1995: 623. Anthene lachares Hewitson, 1878. d’Abrera, 2009: 794. Neurellipes lachares (Hewitson, 1878). Libert, 2010: 291 n. comb.

Alternative common name: Silky Ciliate Blue. Type locality: Cameroon: “Cameroons”. Holotype (female) in the N.H.M., London. Neallotype (male), Bitje, Ja River, 2000 ft, Cameroon, 1898 (G.L. Bates); N.H.M., London (Libert, 2010). Distribution: Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria, Cameroon, Equatorial Guinea, Gabon, Congo, Angola, Central African Republic, Democratic Republic of Congo, Uganda. Habitat: Primary forest and dense secondary forest (Larsen, 2005a). Habits: A fairly scarce species, which almost never leaves the confines of the forests it inhabits 24 (Larsen, 2005a). It appears not to be very active and specimens of both sexes are usually encountered while perched on green leaves inside forest (Larsen, 2005a). Males occasionally mud-puddle (Larsen, 2005a). Early stages:

Lamborn, 1914: 478. Larva green and onisciform, becoming reddish with oblique lighter stripes just before pupation. Both DNO and TO’s are present. The larvae occur under leaves, usually in groups of two to eight and are always attended by ants. They tend to pull leaves together, especially when about to pupate. Pupal period 9 to 10 days (n = 19).

Larval food: Nothing published. Associated ant: Pheidole aurivillii Mayr. race kasaiensis For. (Formcidae) [Lamborn, 1914: 478; Nigeria]. Pheidole rotundata For. (Formcidae) [Lamborn, 1914: 478; Nigeria].

pulcher Grose-Smith & Kirby, 1893. In: Grose-Smith & Kirby, 1892-7 (as sp. of Lycaenesthes). Rhopalocera exotica, being illustrations of new, rare and unfigured species of butterflies 2: 105 (261 pp.). London. Gabon: “North Gaboon”.

liparis Grose-Smith, 1898 (as sp. of Lycaenesthes). Novitates Zoologicae 5: 357 (350-358). Nigeria: “Warri”.

obsolescens Bethune-Baker, 1910 (as var. of Lycaenesthes lachares). Transactions of the Entomological Society of London 43: 48 (1-84). No locality.

Neurellipes lachares lachares (Hewitson, [1878])

Lycaenesthes lachares Hewitson, [1878]. In: Hewitson, [1862-78]. Illustrations of diurnal lepidoptera. Lycaenidae : 225 (1: 1-228; 2: 95 pp.; Supplement: 1-48.). London. Anthene lachares lachares (Hewitson, 1878). Ackery et al., 1995: 623. Anthene lachares Hewitson, 1878. d’Abrera, 2009: 794. Neurellipes lachares lachares (Hewitson, 1878). Libert, 2010: 292.

Type locality: Cameroon: “Cameroons”. Holotype (female) in the N.H.M., London. Neallotype (male), Bitje, Ja River, 2000 ft, Cameroon, 1898 (G.L. Bates); N.H.M., London (Libert, 2010). Distribution: Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria (south and Cross River loop), Cameroon, Equatorial Guinea, Gabon, Congo, Angola, Democratic Republic of Congo. Specific localities: Guinea – Massadou near Macenta (Libert, 2010). Sierra Leone – Bafodia (Libert, 2010); Kasewe Forest (Libert, 2010); Moyamba (Libert, 2010); Pujehun (Libert, 2010); Bumbuna (Libert, 2010); Gola (Libert, 2010). Liberia – Monrovia (Libert, 2010); Ganta (Libert, 2010); Kpain (Libert, 2010); Siabli (Libert, 2010); Wanau (Libert, 2010). Ivory Coast – Touba (Libert, 2010); Abengourou (Libert, 2010). Ghana – Bobiri Butterfly Sanctuary (Larsen et al., 2007); Bia (Libert, 2010); Nakpanduri (Libert, 2010). Nigeria – Warri (Grose-Smith, 1898); Lagos (Libert, 2010); Ikom (Libert, 2010); Kagoro (Libert, 2010). Cameroon – Bitje, Ja River (Stempffer, 1946); Victoria (Libert, 2010); Mamfe (Libert, 2010); Ako (Libert, 2010); Goyoum (Libert, 2010); Batouri district (Libert, 2010); Maan (Libert, 2010). Equatorial Guinea – Alen (Libert, 2010); Benito (Libert, 2010). Gabon – Oyem (Libert, 2010); Ougooue (Libert, 2010); Ivindo (Libert, 2010); Waka (Libert, 2010); Tchibanga (Libert, 2010); Kinguele (Vande weghe, 2010); Ipassa (Vande weghe, 2010); Langoue (Vande weghe, 2010); Kongou (Vande weghe, 2010); camp Nouna (Vande weghe, 2010); Ekouyi (Vande weghe, 2010). Congo – Sembe (Libert, 2010); Odzala (Libert, 2010); Etoumbi (Libert, 2010); Fort Rousset (Libert, 2010); Kuilu (Libert, 2010); Brazzaville (Libert, 2010).

25 Angola – Bihe (Libert, 2010) Democratic Republic of Congo – Bombo Lumene (Libert, 2010); Kimuenza (Libert, 2010); Maluku near Kinshasa (Libert, 2010); Kapalumba in Kasai (Libert, 2010); Ikela (Libert, 2010); Bafwasende (Libert, 2010).

Neurellipes lachares toroensis (Stempffer, 1947)

Anthene lachares toroensis Stempffer, 1947. Bulletin de la Société Entomologique de France 52: 40 (35-41). Anthene lachares toroensis Stempffer, 1947. Ackery et al., 1995: 624. Anthene lachares toroensis Stempffer, 1947. Larsen & Collins, 1998 [see note below] Anthene toroensis Stempffer, 1947. d’Abrera, 2009: 794. Neurellipes lachares toroensis (Stempffer, 1947). Libert, 2010: 294 n. comb.

Type locality: Uganda: “Bwamba, Toro, Ouganda occidental”. Holotype (male), December 1942 (T.H.E. Jackson); N.H.M., London. Allotype (female), same data (Libert, 2010). Distribution: Central African Republic, Democratic Republic of Congo (east – Ituri), Uganda (west). Specific localities: Central African Republic – Dzanga (Noss, 1998; as lachares); Bangui (Libert, 2010); Moloukou (Libert, 2010). Democratic Republic of Congo – Kuma (Libert, 2010); Bamania (Libert, 2010).; Yakoma (Libert, 2010); Bambesa (Libert, 2010); Paulis (Libert, 2010); Kisangani (Libert, 2010); Semliki (Libert, 2010); Walikali (Libert, 2010). Uganda – Bwamba (TL).

Note: Larsen & Collins (1998) averred that subspecies toroensis may be specifically distinct but did not formally emend its status. It was duly raised to species level by d’Abrera (2009: 794). However, Libert (2010: 294) treats toroensis as a subspecies of lachares.

*Neurellipes emkopoti (Larsen & Collins, 1998) Mkpot Hairtail

Anthene emkopoti Larsen & Collins, 1998. Lambillionea 98 (3) (Tome I): 373 (372-376). Anthene emkopoti Larsen & Collins, 1998. d’Abrera, 2009: 796. Neurellipes emkopoti (Larsen & Coll., 1878). Libert, 2010: 295 n. comb.

Neurellipes emkopoti. Male, Nigeria. Left – upperside; right – underside. Images courtesy Torben Larsen.

Alternative common name: Mkpot Ciliate Blue. Type locality: Nigeria: “Butatong, Okwangwo, xi.1995. (T.B. Larsen leg.).” Holotype (male) in the N.H.M., London (Libert, 2010).

26 Distribution: Nigeria (south and Cross River loop), Cameroon (west). Specific localities: Nigeria – Butatong, Okwangwo (TL); Okomu Nature Sanctuary, near Benin City (Larsen, 2005a); Oban Hills (Larsen, 2005a); Calabar (Libert, 2010). Cameroon – Korup (Larsen, 2005a); Victoria (Larsen, 2005a); Douala/Edea road (Libert, vide Larsen, 2005a); Buea (Larsen, 2005a); Ebogo (Larsen, 2005a); Kumba (Libert, 2010); Bonepoupa (Libert, 2010); Telep near Edea (Libert, 2010); Kribi (Libert, 2010); Mbebe (Libert, 2010); Ndoupe (Libert, 2010); Campo (Libert, 2010); Lolodorf (Libert, 2010). Habitat: Forest (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes ramnika (d’Abrera, 1980)

Anthene ramnika d'Abrera, 1980. Butterflies of the Afrotropical region: 532 (593 pp.). Melbourne. Anthene ramnika d’Abrera, 1980. Ackery et al., 1995: 627. Anthene ramnika d’Abrera, 1980. d’Abrera, 2009: 794. Neurellipes ramnika (d’Abrera, 1980). Libert, 2010: 296 n. comb.

Type locality: [Democratic Republic of Congo]: “Yopole Dist., Buta, B. Congo, 3500 ft”. Holotype (male), October 1946 (T.H.E. Jackson); N.H.M., London (Libert, 2010). Distribution: Cameroon, Democratic Republic of Congo (Uele). Specific localities: Cameroon – Meyo Mela (Libert, 2010); Bitje (Libert, 2010); Ebogo (Libert, 2010). Democratic Republic of Congo – Buta, Yopole District (TL); Beni (Libert, 2010); Mount Hoyo (Libert, 2010); Walikali (Libert, 2010); Paulis (Libert, 2010); Opala (Libert, 2010). Habitat: Forest. Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes atewa (Larsen & Collins, 1998) Atewa Hairtail

Anthene atewa Larsen & Collins, 1998. Lambillionea 98 (3) (Tome I): 373 (372-376). Anthene atewa Larsen & Collins, 1998. d’Abrera, 2009: 796. Neurellipes atewa (Larsen & Collins, 1998). Libert, 2010: 297 n. comb.

Neurellipes atewa. Male, Ghana. Left – upperside; right – underside. Images courtesy Torben Larsen.

Neurellipes atewa. Female, Ghana. Upperside. Image courtesy Torben Larsen.

Alternative common name: Atewa Ciliate Blue. Type locality: Ghana: “Atewa Range, Kibi, viii.1996 (T.B. Larsen leg.).” Holotype (male) in the Natural History Museum, London (Libert, 2010). Distribution: Guinea, Sierra Leone, Ivory Coast (east), Ghana. Specific localities: Guinea – Ziama Forest near Seredou (Libert, 2010). Sierra Leone – Gola (Libert, 2010). Ivory Coast – Yapo (H. Warren-Gash, vide Larsen, 2005a); Alepe (Libert, 2010); Banco (Libert, 2010); Danane-Nimba (Libert, 2010). Ghana – Kibi, Atewa Range (TL); Hotopo (Larsen, 2005a); Subri (Maessen, vide Larsen, 2005a); Ankasa (Libert, 2010). Habitat: Primary forest (Larsen, 2005a). Habits: A rare species (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes xanthopoecilus (Holland, 1893)

Lycaenesthes xanthopoecilus Holland, 1893. Entomological News 4: 27 (22-28). Anthene xanthopoecilus Holland, 1893. d’Abrera, 2009: 794. Neurellipes xanthopoecilus (Holl., 1893). Libert, 2010: 300 n. comb.

Neurellipes xanthopoecilus xanthopoecilus. Male. Left – upperside; right – underside. Ipassa, Ivindo N.P., Gabon. 17 June 2016. J. Dobson. Images M.C. Williams ex Dobson collection.

Type locality: Gabon: “Talaguga”. Holotype (female), (A.C. Good); C.M.N.H., Pittsburgh. Neallotype (male), Efulen, Cameroon, 23 April 1925 (H.L. Weber); C.M.N.H., Pittsburgh (Libert, 2010).

28 Distribution: Nigeria, Cameroon, Equatorial Guinea, Gabon, Congo, Central African Republic, Democratic Republic of Congo. Habitat: Forest. Early stages: Nothing published. Larval food: Nothing published.

Neurellipes xanthopoecilus xanthopoecilus (Holland, 1893)

Lycaenesthes xanthopoecilus Holland, 1893. Entomological News 4: 27 (22-28). Anthene xanthopoecilus Holland, 1893. d’Abrera, 2009: 794. Neurellipes xanthopoecilus xanthopoecilus (Holl., 1893). Libert, 2010: 300.

Neurellipes xanthopoecilus xanthopoecilus. Male. Left – upperside; right – underside. Ipassa, Ivindo N.P., Gabon. 17 June 2016. J. Dobson. Images M.C. Williams ex Dobson collection.

Type locality: Gabon: “Talaguga”. Holotype (female), (A.C. Good); C.M.N.H., Pittsburgh. Neallotype (male), Efulen, Cameroon, 23 April 1925 (H.L. Weber); C.M.N.H., Pittsburgh (Libert, 2010). Distribution: Cameroon, Equatorial Guinea, Gabon, Congo, Central African Republic, Democratic Republic of Congo. Specific localities: Cameroon – Bitje, Ja River (Bethune-Baker, 1910 – as bitje); Nyong (Libert, 2010); Maan (Libert, 2010); Djoum (Libert, 2010). Equatorial Guinea – Benito (Libert, 2010). Gabon – Talaguga (TL); Ipassa (Vande weghe, 2010); Langoue (Vande weghe, 2010); Kongou (Vande weghe, 2010); camp Nouna (Vande weghe, 2010); Bakouaka (Vande weghe, 2010); Kebiri (Vande weghe, 2010). Congo – Souanke (Libert, 2010); Ouesso (Libert, 2010); Odzala (Libert, 2010); Kelle (Libert, 2010); Fort Rousset (Libert, 2010); Mah (Libert, 2010). Central African Republic – Moloukou (Libert, 2010); Bangui (Libert, 2010); Fort Sibut (Libert, 2010). Democratic Republic of Congo – Lukolela (Libert, 2010).

bitje Druce, 1910 (as sp. of Lycaenesthes). Proceedings of the Zoological Society of London 1910: 373 (356-378). Type locality: Cameroon: “Bitje, Ja River, Camerouns, 2000 ft.”. Treated as Anthene versatilis bitje (Druce, 1910) by Larsen, 2005a: 239, stat. rev. D’Abrera (2009: 794) formally sank bitje as a junior synonym of versatilis. Libert (2010: 300) treated bitje as a synonym of xanthopoecilus (Holland, 1893) n. syn.

Neurellipes xanthopoecilus lukokescha (Karsch, 1895)

Lycaenesthes lukokescha Karsch, 1895. Entomologische Nachrichten. Berlin 21: 295 (289-322). Synonym of Anthene xanthopoecilus (Holland, 1893). Ackery et al., 1995. Neurellipes xanthopoecilus lukokescha (Karsch, 1895). Libert, 2010: 302 n. comb. 29

Type locality: [Democratic Republic of Congo]: “Mukenge”. Holotype (female), (P. Pogge); Z.M.H.U., Berlin. Neallotype (male), Katako-Kombe, Sankuru, Democratic Republic of Congo, 2 April 1952 (Dr M. Fontaine); M.R.A.C., Tervuren (Libert, 2010). Distribution: Democratic Republic of Congo. Specific localities: Democratic Republic of Congo – Mukenge (TL); Katako-Kombe (Libert, 2010); Kinshasa (Libert, 2010); Beni (Libert, 2010); Epulu (Libert, 2010); Kapanga (Libert, 2010).

Neurellipes xanthopoecilus peteri Libert, 2010

Neurellipes xanthopoecilus peteri Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 302.

Type locality: Nigeria: Mkpot, Calabar, 1-7 December 1996 (P. Walwanda). Holotype (female) in A.B.R.I., Nairobi. Allotype (male), same data and place of deposition. Distribution: Nigeria, Cameroon. Specific localities: Nigeria – Mkpot, Calabar (TL); Ikom (Libert, 2010); Oban Hills (Libert, 2010). Cameroon – Mamfe (Libert, 2010).

*Neurellipes versatilis (Bethune-Baker, 1910)

Lycaenesthes versatilis Bethune-Baker, 1910. Transactions of the Entomological Society of London 43: 58 (1-84). Anthene versatilis (Bethune-Baker, 1910). Ackery et al., 1995: 628. Anthene versatilis Bethune-Baker, 1910. Larsen, 2005a. Anthene versatilis Bethune-Baker, 1910. d’Abrera, 2009: 794. Neurellipes versatilis (Bethune-B., 1910). Libert, 2010: 303.

Type locality: Gabon: “Gaboon”. Holotype (male) in the N.H.M., London. Neallotype (female), Gabon (Mocquerys); Z.M.H.U., Berlin (Libert, 2010). Distribution: Cameroon, Gabon. Misattributed to the Liberia, Ivory Coast and Ghana fauna by Clench (1965) and Stempffer (1966) for the then undescribed Anthene atewa (Larsen, 2005a). Has also been recorded, in error, from Nigeria and the Democratic Republic of Congo (Libert, 2010). Specific localities: Cameroon – Douala (Libert, 2010); Batanga (Libert, 2010); Ngambe (Ndokon); Djoum (Libert, 2010); Avobengou (Libert, 2010). Gabon – Libreville (Libert, 2010); Fernan Vaz (Libert, 2010); Ogowe (Libert, 2010); Pongara (Vande weghe, 2010); Nyonie (Vande weghe, 2010); Rabi (Vande weghe, 2010); Langoue (Vande weghe, 2010); Kongou (Vande weghe, 2010). Habitat: Habits: Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes lycotas (Grose-Smith, 1898)

Lycaenesthes lycotas Grose-Smith, 1898. Novitates Zoologicae 5: 356 (350-358). Synonym of Anthene xanthopoecilus (Holland, 1893). Ackery et al., 1995. Neurellipes lycotas (Grose-Smith, 1898). Libert, 2010: 305 n. comb.

Type locality: [Democratic Republic of Congo]: “Bopoto, Upper Congo”. Holotype (female), (Balfern); N.H.M., London. Neallotype (male), Beni, Ituri, 4000 ft, March 1947 (T.H.E. Jackson);

30 genitalia GET 1965-503; N.H.M., London (Libert, 2010). Distribution: Democratic Republic of Congo. Specific localities: Democratic Republic of Congo – Bopoto (TL); Lukolela (Libert, 2010); Eala (Libert, 2010)Mount Hoyo (Libert, 2010); Lubero (Libert, 2010); Beni (Libert, 2010); Yakoma (Libert, 2010); Bambesa (Libert, 2010). Habitat: Habits: Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes lysicles (Hewitson, 1874) Violet-spotted Hairtail

Male (left) and female (right) of the Violet-spotted Hairtail. Ghana. Images courtesy Pamela Sai.

Lycaenesthes lysicles Hewitson, 1874. Transactions of the Entomological Society of London 1874: 348 (343-355). Anthene lysicles (Hewitson, 1874). Ackery et al., 1995: 625. Anthene lysicles Hewitson, 1874. d’Abrera, 2009: 794. Neurellipes lysicles (Hewitson, 1874). Libert, 2010: 305 n. comb.

Alternative common name: Violet-spotted Ciliate Blue. Type locality: Nigeria: “Old Calabar”. Holotype (male) in the N.H.M., London. Neallotype (female), Warri, Nigeria, April 1897 (Dr Roth); N.H.M., London (Libert, 2010). Distribution: Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, Nigeria, Cameroon, Equatorial Guinea, Gabon, Congo, Central African Republic, Democratic Republic of Congo, Uganda. Habitat: Forest (Larsen, 2005a). Habits: An uncommon species met with occasionally in closed forest (Larsen, 2005a). Both sexes perch on green leaves and do not appear to be very active (Larsen, 2005a). Males rarely mud-puddle (Larsen, 2005a). Early stages: Nothing published. Larval food: Albizia species (Fabaceae) [Fontaine, 1988; Democratic Republic of Congo].

Note: Berger (1981) recorded it from Shaba (DRC) but Larsen (2005a) thinks that these populations may represent a distinct species.

Neurellipes lysicles lysicles (Hewitson, 1874)

31 Type locality: Nigeria: “Old Calabar”. Holotype (male) in the N.H.M., London. Neallotype (female), Warri, Nigeria, April 1897 (Dr Roth); N.H.M., London (Libert, 2010). Distribution: Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, Nigeria (south and Cross River loop), Cameroon, Equatorial Guinea, Gabon, Congo, Central African Republic, Democratic Republic of Congo (Shaba). Specific localities: Guinea – Labe (Libert, 2010). Sierra Leone – Freetown (Libert, 2010); Kafogo (Libert, 2010); Moyamba (Libert, 2010). Liberia – Fisabu (Libert, 2010); Fisherman Lake (Libert, 2010); Harbel (Libert, 2010); Wanau (Libert, 2010). Ivory Coast – Danane (Libert, 2010); Abidjan (Libert, 2010). Ghana – Bobiri Butterfly Sanctuary (Larsen et al., 2007); Ankasa (Libert, 2010); Mamiri (Libert, 2010); Tano Ofin (Libert, 2010); Kumasi (Libert, 2010); Kwahu Scarp (Libert, 2010). Togo – Badou (Libert, 2010); Kpalime (Libert, 2010). Nigeria – Old Calabar (TL); Warri (Libert, 2010); Lagos (Libert, 2010); Ubiaja (Libert, 2010); Nsukka (Libert, 2010). Cameroon – Korup (Larsen, 2005a); Rumpi Hills (Libert, 2010); Mamfe (Libert, 2010); Bitje (Libert, 2010); Goyoum (Libert, 2010). Equatorial Guinea – Benito (Libert, 2010). Gabon – Ipassa (Libert, 2010); Ivindo (Libert, 2010); Ogooue (Libert, 2010); Libreville (Libert, 2010); Mayumbe (Libert, 2010); Pongara (Vande weghe, 2010); Kinguele (Vande weghe, 2010); Lake Evaro (Vande weghe, 2010); Mboumie (Vande weghe, 2010); Ndjole (Vande weghe, 2010); Waka (Vande weghe, 2010); Massouna 2000 (Vande weghe, 2010); Langoue (Vande weghe, 2010); camp Nouna (Vande weghe, 2010); Kebiri (Vande weghe, 2010). Congo – Souanke (Libert, 2010); Ngo (Libert, 2010); Brazzaville (Libert, 2010). Central African Republic – Moloukou (Libert, 2010); Boukoko (Libert, 2010); Bangui (Libert, 2010). Democratic Republic of Congo – Mbata Siala (Libert, 2010); Mayumbe (Libert, 2010); Paulis (Libert, 2010); Kafakumba (Libert, 2010).

Neurellipes lysicles clarior Libert, 2010

Neurellipes lysicles clarior Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 307.

Type locality: Uganda: Kalinzu Forest, West Ankole, November 1937 (T.H.E. Jackson). Holotype (male) in the N.H.M., London. Allotype (female) same data and deposition, except collected in October. Distribution: Uganda. Specific localities: Uganda – Kalinzu Forest (TL).

*Neurellipes buchholzi (Plötz, 1880)

Lycaenesthes buchholzi Plötz, 1880. Stettiner Entomologische Zeitung 41: 202 (189-206). Anthene buchholzi (Plötz, 1880). Ackery et al., 1995: 621. Anthene buchholzi Plötz, 1880. d’Abrera, 2009: 794. Neurellipes buchholzi (Plötz, 1880). Libert, 2010: 307 n. comb.

Type locality: Cameroon: “Victoria”. Distribution: Cameroon. Known only from the type series. Specific localities: Cameroon – Victoria (= Limbe) (TL). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes lyzanius (Hewitson, 1874) Black-patches Hairtail

Lycaenesthes lyzanius Hewitson, 1874. Entomologist’s Monthly Magazine 11: 36 (36). Neurypexina lyzanius (Hewitson, 1874). Ackery et al., 1995: 629. Neurypexina lyzanius Hewitson, 1874. d’Abrera, 2009: 801. Neurellipes lyzanius (Hewitson, 1874). Libert, 2010: 309 n. comb.

Alternative common name: Black-patches. Type locality: Nigeria: “Old Calabar”. Holotype (female) in the Natural History Museum, London. Neallotype (male), Uwet, southern Nigeria, March 1921 (D. Cator); N.H.M., London (Libert, 2010). Distribution: Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, Nigeria (south and Cross River loop), Cameroon, Equatorial Guinea, Gabon, Congo, Angola (north), Central African Republic, Democratic Republic of Congo. Specific localities: Guinea – Ratoma (Libert, 2010); Dubreka near Conakry (Libert, 2010); Mamou (Libert, 2010); Nzo near the Nimba Mountains (Libert, 2010). Sierra Leone – Banta (Libert, 2010); Guma near Freetown (Libert, 2010); Panguma (Libert, 2010); Bumbuna (Libert, 2010); Gola (Libert, 2010). Liberia – Boma Hills (Libert, 2010); Harbel (Libert, 2010); Kpaine (Libert, 2010); Monrovia (Libert, 2010); Wanau (Libert, 2010). Ivory Coast – Danane (Libert, 2010); Bereby (Libert, 2010); Banco near Abidjan (Libert, 2010); Alepe (Libert, 2010); Bouna (Libert, 2010). Ghana – Bobiri Butterfly Sanctuary (Larsen et al., 2007); Bia (Libert, 2010); Cape Three Points (Libert, 2010); Kumasi (Libert, 2010); Kyabobo (Libert, 2010). Togo – Danyi (Libert, 2010). Nigeria – Old Calabar (TL); Uwet (Libert, 2010); Lagos (Libert, 2010); Oban Hills (Libert, 2010); Omuo (Libert, 2010). Cameroon – Mamfe (Libert, 2010); Victoria (Libert, 2010); Bafia (Libert, 2010); Yaounde (Libert, 2010); Wak (Libert, 2010); Ngaoundere (Libert, 2010); Maan (Libert, 2010); Lomie (Libert, 2010); Moloundou (Libert, 2010). Equatorial Guinea – Benito (Libert, 2010). Gabon – Upper Ogove River (Holland, 1891); Libreville (Libert, 2010); Fernan Vaz (Libert, 2010); Ogooue (Libert, 2010); Tchibanga (Libert, 2010); Nyonie (Vande weghe, 2010); Safala (Vande weghe, 2010); Mondah (Vande weghe, 2010); Iguela (Vande weghe, 2010); Rabi (Vande weghe, 2010); Waka (Vande weghe, 2010); Mikongo (Vande weghe, 2010); Langoue (Vande weghe, 2010); Ipassa (Vande weghe, 2010); Nouna (Vande weghe, 2010). Congo – Ouesso (Libert, 2010); Odzala (Libert, 2010); Etoumbi (Libert, 2010); Kelle (Libert, 2010); Kuilu (Libert, 2010). Angola – Bihe (Libert, 2010). Central African Republic – Bangui (Libert, 2010). Democratic Republic of Congo – Lukolela (Libert, 2010); Eala (Libert, 2010); Elizabetha (Libert, 2010); Embangobango (Libert, 2010); Opala (Libert, 2010); Beni (Libert, 2010). Habitat: Nothing published. Habits: A relatively common butterfly that is often met with inside forests, perched on broad green leaves (Larsen, 2005a). Males do not appear to mud-puddle and are somewhat shy (Larsen, 2005a). Early stages:

Lamborn, 1914: 484. Six larvae were found near each other on new shoots, but were not gregarious. Pupal period 9 to 10 days (n = 6).

Larval food: Nothing published. Associated ant: Pheidole rotundata For. (Formcidae) [Lamborn, 1914: 484; Nigeria].

33 regillus Holland, 1891 (as ?). Psyche, a Journal of Entomology. Cambridge, Mass.6: 51 (50-53). Gabon: “upper waters of the Ogové River”.

turbatus Grose-Smith & Kirby, 1893. In: Grose-Smith & Kirby, 1892-7 (as ?). Rhopalocera exotica, being illustrations of new, rare and unfigured species of butterflies 2: 105 (261 pp.). London. Gabon: “Gaboon”.

*Neurellipes ferenczi Libert, 2010

Neurellipes ferenczi Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 311.

Type locality: Guinea: Coyah, 19 March 1964 (Dr K. Ferencz); genitalia Libert 109-163. Holotype (male) in the Hungarian Natural History Museum, Budapest. Distribution: Guinea. Specific localities: Guinea – Coyah (TL). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

*Neurellipes flavomaculatus (Grose-Smith & Kirby, [1893]) Yellow-spot Hairtail

Lycaenesthes flavomaculatus Grose-Smith & Kirby, [1893]. In: Grose-Smith & Kirby, [1892-7]. Rhopalocera exotica, being illustrations of new, rare and unfigured species of butterflies 2: 104 (261 pp.). London. Anthene flavomaculatus (Grose-Smith & Kirby, 1893). Ackery et al., 1995: 622. Anthene flavomaculatus Grose-Smith & Kirby, 1893. d’Abrera, 2009: 796. Neurellipes flavomaculatus (Grose-Smith & Kirby, 1893). Libert, 2010: 312 n. comb.

Alternative common name: Yellow-spot Ciliate Blue. Type locality: “West Africa”. Holotype (male) in the N.H.M., London. Neallotype (female), Afikpo, northern Nigeria (Reddick); N.H.M., London (Libert, 2010). Distribution: Nigeria, Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo, Angola. Probably misattributed to the Ivory Coast fauna by Ackery et al. (1995) (Larsen, 2005a). Habitat: Forest (Larsen, 2005a). Habits: A rather rare species, mostly encountered while perched on green leaves, in forest (Larsen, 2005a). Early stages:

Lamborn, 1914: 483. Three larvae were found side by side on the underside of a leaf. DNO present; TO’s marked by small white spots. Larvae drop to the ground if disturbed. Pupal period 9 to 10 days (n = 3).

Larval food: Nothing published. Associated ants: Crematogaster buchneri For. race alligatrix For. (Formcidae) [Lamborn, 1914: 483; Nigeria]. Crematogaster buchneri For. race winkleri For. (Formcidae) [Lamborn, 1914: 483; Nigeria]. Odontomachus haematodes Linn. (Formcidae) [Lamborn, 1914: 483; Nigeria]. Pheidole rotundata For. (Formcidae) [Lamborn, 1914: 483; Nigeria].

34 Neurellipes flavomaculatus flavomaculatus (Grose-Smith & Kirby, [1893])

Lycaenesthes flavomaculatus Grose-Smith & Kirby, [1893]. In: Grose-Smith & Kirby, [1892-7]. Rhopalocera exotica, being illustrations of new, rare and unfigured species of butterflies 2: 104 (261 pp.). London. Anthene flavomaculatus (Grose-Smith & Kirby, 1893). Ackery et al., 1995: 622. Anthene flavomaculatus Grose-Smith & Kirby, 1893. d’Abrera, 2009: 796. Neurellipes flavomaculatus flavomaculatus (Grose-Smith & Kirby, 1893). Libert, 2010: 312.

Type locality: “West Africa”. Holotype (male) in the N.H.M., London. Neallotype (female), Afikpo, northern Nigeria (Reddick); N.H.M., London (Libert, 2010). Distribution: Nigeria (south and Cross River loop), Gabon. Probably misattributed to the Ivory Coast fauna by Ackery et al. (1995) (Larsen, 2005a). Specific localities: Nigeria – Agege (Larsen,2005a); Bassa (Larsen, 2005a); Okomu (Larsen, 2005a); Kagoro (Larsen, 2005a); Oban Hills (Larsen, 2005a); Lagos (Libert, 2010); Calabar (Libert, 2010); Benoue (Libert, 2010). Gabon – Camp Nouna (Vande weghe, 2010).

Neurellipes flavomaculatus bipuncta (Joicey & Talbot, 1921)

Lycaenesthes bipuncta Joicey & Talbot, 1921. Bulletin of the Hill Museum, Witley 1: 96 (40-166). Anthene bipuncta (Joicey & Talbot, 1921). Ackery et al., 1995: 621. Anthene bipuncta Joicey & Talbot, 1921. d’Abrera, 2009: 796. Neurellipes flavomaculatus bipuncta (Joicey & Talbot, 1921). Libert, 2010: n. comb.

Type locality: [Democratic Republic of Congo]: “Cartouche, near Lesse, W. Semliki River”. Holotype (male), January 1920 (T.A. Barns); N.H.M., London. Allotype (female), between Lindi and Lubila Rivers, north of Batama, Democratic Republic of Congo, April 1920 (T.A. Barns); N.H.M., London (Libert, 2010). Distribution: Nigeria, Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo, Angola. Specific localities: Nigeria – Kagoro Forest (Libert, 2010); Nabarawa = ?Nassarawa (Libert, 2010). Cameroon – Ebogo (Libert, 2010); Akoafim (Libert, 2010). Gabon – Fougamou (Libert, 2010); camp PPG (Vande weghe, 2010). Congo – Etoumbi (Libert, 2010); Mambili (Libert, 2010). Central African Republic – Bangui (Libert, 2010); Mbaiki (Libert, 2010). Democratic Republic of Congo – Cartouch, near Lesse (TL); Katako-Kombe (Libert, 2010); Mwene Ditu (Libert, 2010); Lukolela (Libert, 2010). Angola – Bihe (Libert, 2010).

*Neurellipes boormani Libert, 2010

Neurellipes boormani Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 314.

Type locality: Nigeria: Lagos, January 1960 (J. Boorman). Holotype (female) in A.B.R.I., Nairobi. Allotype (male), Lagos, Nigeria, January 1963 (J. Boorman); genitalia Libert 109-333; in A.B.R.I., Nairobi. Distribution: Nigeria. Specific localities: Nigeria – Lagos (TL). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. 35 Associated ants: Nothing published.

*Neurellipes lamprocles (Hewitson, [1878]) Lesser Black-patches Hairtail

Lycaenesthes lamprocles Hewitson, [1878]. In: Hewitson, [1862-78]. Illustrations of diurnal lepidoptera. Lycaenidae: 225 (1: 1-228; 2: 95 pp.; Supplement: 1-48.). London. Neurypexina lamprocles (Hewitson, 1878). Ackery et., 1995: 629. Neurypexina lamprocles Hewitson, 1878. d’Abrera, 2009: 801. Neurellipes lamprocles (Hewitson, 1878). Libert, 2010: 315 n. comb.

Alternative common name: Lesser Black-patches. Type locality: “West Africa”. Probably Cameroon or Gabon (Larsen, 2005a). Holotype (male) in the N.H.M., London. Neallotype (female), Uwet, southern Nigeria, March 1921 (D. Cator); N.H.M., London (Libert, 2010). Distribution: Nigeria, Cameroon, Equatorial Guinea, Gabon, Congo, Central African Republic, Democratic Republic of Congo. Misattributed to the Ivory Coast fauna by Stempffer (1966) (Larsen, 2005a). Habitat: Forest. Habits: An uncommon species, usually found perched on large green leaves, conspicuous because of the white underside (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

Neurellipes lamprocles lamprocles (Hewitson, [1878])

Type locality: “West Africa”. Probably Cameroon or Gabon (Larsen, 2005a). Holotype (male) in the N.H.M., London. Neallotype (female), Uwet, southern Nigeria, March 1921 (D. Cator); N.H.M., London (Libert, 2010). Distribution: Nigeria (Cross River loop), Cameroon, Equatorial Guinea, Gabon, Congo. Misattributed to the Ivory Coast fauna by Stempffer (1966) (Larsen, 2005a). Specific localities: Nigeria – Uwet (Libert, 2010); Kabbo (Libert, 2010); Omuo (Libert, 2010); Calabar (Libert, 2010); Ikom (Libert, 2010). Cameroon – Bitje, Ja River (Stempffer, 1946 – for bitje); Korup (Larsen, 2005a); Mamfe (Libert, 2010); Maan (Libert, 2010); Deng Deng (Libert, 2010); Bertoua (Libert, 2010); Lomie (Libert, 2010). Equatorial Guinea – Balengue (Libert, 2010). Gabon – Libreville (Libert, 2010); Ogooue (Libert, 2010); Ivindo (Libert, 2010); Rabi (Libert, 2010); Mayoumbe (Libert, 2010); Waka (Vande weghe, 2010); Lope N.P. (Vande weghe, 2010); Langoue (Vande weghe, 2010); Ipassa (Vande weghe, 2010); camp Nouna (Vande weghe, 2010). Congo – Etoumbi (Libert, 2010); Menengue (Libert, 2010); Koubotchi (Libert, 2010); Kouilou (Libert, 2010).

bitje Stempffer, 1946 (as sp. of Pseudoliptena). Revue Française d’Entomologie 13: 9 (8-19). Cameroon: “Bitje, Ja River”. Synonymized with Neurellipes lamprocles lamprocles (Hewitson, 1878) by Libert, 2010: 315 n. syn.

36 Neurellipes lamprocles reducta Libert, 2010

Neurellipes lamprocles reducta Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 316.

Type locality: Democratic Republic of Congo: Blukwa. Holotype (male) in M.R.A.C., Tervuren. Allotype (female), Paulis, Democratic Republic of Congo, 16 August 1959 (Dr M. Fontaine); in M.R.A.C., Tervuren. Distribution: Central African Republic, Democratic Republic of Congo (north-east). Specific localities: Central African Republic – Dzanga (Noss, 1998); Bangui (Libert, 2010). Democratic Republic of Congo – Blukwa (TL); Paulis (Libert, 2010); Bangupa (Libert, 2010); Kisangani (Libert, 2010); Makala (Libert, 2010); Mambasa (Libert, 2010); Beni (Libert, 2010).

*Neurellipes mirellae Belcastro & Libert, 2010

Neurellipes mirellae Belcastro & Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 318.

Type locality: Sierra Leone: Gola Forest south, near Gorahun, 28 April 2006 (C. Belcastro). Holotype (male) in Museo Civico di Zoologia di Roma. Distribution: Sierra Leone. Specific localities: Sierra Leone – Gola Forest (TL). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

*Neurellipes inferna (Bethune-Baker, 1926)

Triclema inferna Bethune-Baker, 1926. Annals and Magazine of Natural History (9) 17: 399 (384-402). Triclema inferna Bethune-Baker, 1926. d’Abrera, 2009: 802. Neurellipes inferna (Bethune-Baker, 1926). Libert, 2010: 318 n. comb.

Type locality: Cameroon: “Bitje, 2000 ft.”. Holotype (male), (G.L. Bates); N.H.M., London (Libert, 2010). Distribution: Cameroon. Specific localities: Cameroon – Bitje (TL); Koukoue south-east of Edea (Libert, 2010). Early stages: Nothing published. Larval food: Nothing published.

*Neurellipes darwini Libert, 2010

Neurellipes darwini Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 318.

Type locality: Cameroon: Ebondi, 22 May 1989 (M. Libert); genitalia Libert 106-255. Holotype (male) in M.N.H.N., Paris. Distribution: Cameroon (south). Specific localities:

37 Cameroon – Ebondi (TL). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.

*Neurellipes dubia Libert, 2010

Neurellipes dubia Libert, 2010. Libert, 2010. Révision des Anthene africains. A.B.R.I., Nairobi, Kenya and Lambillionea, Chênée, Belgique: 319.

Type locality: Cameroon: Ekona, 19 May; genitalia Libert 107-304. Holotype (male) N.H.M., London. Distribution: Cameroon. Specific localities: Cameroon – Ekona (TL). Habitat: . Habits: . Early stages: Nothing published. Larval food: Nothing published. Associated ants: Nothing published.