Dimensions of Ecosystem Complexity: Heterogeneity, Connectivity, and History
ecological complexity 3 (2006) 1–12 available at www.sciencedirect.com journal homepage: http://www.elsevier.com/locate/ecocom Viewpoint Dimensions of ecosystem complexity: Heterogeneity, connectivity, and history M.L. Cadenasso a,*, S.T.A. Pickett b, J.M. Grove c a Hixon Center for Urban Ecology, School of Forestry and Environmental Studies, Yale University, 205 Prospect Street, New Haven, CT 06511, United States b Institute of Ecosystem Studies, Box AB, Millbrook, NY 12545, United States c USDA Forest Service, Northeastern Research Station, 705 Spear Street, P.O. Box 968, Burlington, VT 05401, United States article info abstract Article history: Biocomplexity was introduced to most ecologists through the National Science Foundation’s Received 2 June 2005 grant program, and the literature intended to introduce that program. The generalities of that Received in revised form literature contrast with the abstract and mathematical sophistication of literature from 30 June 2005 physics, systems theory, and indeed even of pioneering ecologists who have translated the Accepted 2 July 2005 conceptintoecology. Thissituation leaves a middle ground, that isboth accessibletoecologists Published on line 23 January 2006 in general, and cognizant of the fundamentals of complexity, to be more completely explored. To help scope this middle ground, and to promote empirical explorations that may be located Keywords: there, we propose a non-exclusive framework for the conceptual territory. While recognizing Biocomplexity the deep foundations in the studies of complex behavior, we take ecological structure as the Framework entry point for framework development. This framework is based on a definition of biocom- Coupled systems plexity as the degree to which ecological systems comprising biological, social and physical Spatial heterogeneity components incorporate spatially explicit heterogeneity, organizational connectivity, and Legacies historical contingency through time.
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