Tree Squirrels: Managing Habitat and Controlling Damage
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Squirrels and Allies (Sciuridae Spp.)
Squirrels and Allies (Sciuridae Spp.) There are about 275 species of tree, ground, and flying squirrels throughout the world, ranging from the five-inch pygmy squirrel (Myosciurus pumilio) of Africa to the nearly three-foot giant squirrel (Ratufa indica) of Asia. Currently, there are five species of squirrel found in Rhode Island: The Eastern gray squirrel, the Southern flying squirrel, the red squirrel, the chipmunk, and the woodchuck. Tree squirrels are those species that nest in trees, while ground squirrels nest in underground burrows. This publication is intended to provide information on the above species found in Rhode Island. Eastern Gray Squirrel (Sciurus carolinensis) Gray squirrels are the most common species of squirrel in Rhode Island. They are a tree squirrel, however they spend much of their time on the ground, and have adapted well to human-altered environments. They are commonly found in urban and suburban habitats. Gray squirrels can be grayish-brown, gray or black, with a white or light brown underside. Adults range in size from 15 to 20 inches, and between 1 to 1 ½ pounds. Their bushy tail accounts for almost half of their length. Gray squirrels may live between four and eight years in the wild, but they have been known to live up to ten years in captivity. They are active mainly during the day and do not hibernate. In the winter they spend days at a time in their nests, leaving only to gather food. Their habitat is mainly any wooded area with mast producing trees, where they live in tree cavities or leaf nests. -
Symposium on the Gray Squirrel
SYMPOSIUM ON THE GRAY SQUIRREL INTRODUCTION This symposium is an innovation in the regional meetings of professional game and fish personnel. When I was asked to serve as chairman of the Technical Game Sessions of the 13th Annual Conference of the Southeastern Association of Game and Fish Commissioners this seemed to be an excellent opportunity to collect most of the people who have done some research on the gray squirrel to exchange information and ideas and to summarize some of this work for the benefit of game managers and other biologists. Many of these people were not from the southeast and surprisingly not one of the panel mem bers is presenting a general resume of one aspect of squirrel biology with which he is most familiar. The gray squirrel is also important in Great Britain but because it causes extensive damage to forests. Much work has been done over there by Monica Shorten (Mrs. Vizoso) and a symposium on the gray squirrel would not be complete without her presence. A grant from the National Science Foundation through the American Institute of Biological Sciences made it possible to bring Mrs. Vizoso here. It is hoped that this symposium will set a precedent for other symposia at future wildlife conferences. VAGN FLYGER. THE RELATIONSHIPS OF THE GRAY SQUIRREL, SCIURUS CAROLINENSIS, TO ITS NEAREST RELATIVES By DR. ]. C. MOORE INTRODUCTION It seems at least slightly more probable at this point in our knowledge of the living Sciuridae, that the northeastern American gray squirrel's oldest known ancestors came from the Old \Vorld rather than evolved in the New. -
Tree Squirrels
Tree Squirrels When the public is polled regarding suburban and urban wildlife, tree squirrels generally rank first as problem makers. Residents complain about them nesting in homes and exploiting bird feeders. Interestingly, squirrels almost always rank first among preferred urban/suburban wildlife species. Such is the Figure 1. The Eastern gray squirrel is from the deciduous paradox they present: We want them and we don’t and mixed coniferous-deciduous forests of eastern North want them, depending on what they are doing at any America, and was introduced into city parks, campuses, given moment. and estates in Washington in the early 1900s. (Drawing by Elva Paulson) Although tree squirrels spend a considerable amount of time on the ground, unlike the related ground squirrels, they are more at home in trees. Washington is home to four species of native tree squirrels and two species of introduced tree squirrels. Native Washington Tree Squirrels The Douglas squirrel, or chickaree (Tamiasciurus douglasii) measures 10 to 14 inches in length, including its tail. Its upper parts are reddish-or brownish-gray, and its underparts are orange to yellowish. The Douglas squirrel is found in stands of fir, pine, cedar, and other conifers in the Cascade Mountains and western parts of Washington. The red squirrel (Tamiasciurus hudsonicus, Fig. 4) is about the same size as the Douglas squirrel and lives in coniferous forests and semi-open woods in northeast Washington. It is rusty-red on the upper part and white or grayish white on its underside. The Western gray squirrel (Sciurus griseus, Fig. 2) is the largest tree squirrel in Washington, ranging from 18 to 24 inches in length. -
Breeding Ecology of Barred Owls in the Central Appalachians
BREEDING ECOLOGY OF BARRED OWLS IN THE CENTRAL APPALACHIANS ABSTRACT- Eight pairs of breedingBarred Owls (Strix varia) in westernMaryland were studied. Nest site habitat was sampledand quantifiedusing a modificationof theJames and Shugart(1970) technique (see Titus and Mosher1981). Statisticalcomparison to 76 randomhabitat plots showed nest sites werb in moremature forest stands and closer to forest openings.There was no apparent association of nestsites with water. Cavity dimensions were compared statistically with 41 randomlyselected cavities. Except for cavityheight, there were no statistically significant differences between them. Smallmammals comprised 65.9% of the totalnumber of prey itemsrecorded, of which81.5% were members of the familiesCricetidae and Soricidae. Birds accounted for 14.6%of theprey items and crayfish and insects 19.5%. We also recordedan apparentinstance of juvenile cannibalism. Thirteennestlings were produced in 7 nests,averaging 1.9 young per nest.Only 2 of 5 nests,where the outcome was known,fledged young. The Barred Owl (Strix varia) is a common noc- STUDY AREA AND METHODS turnal raptor in forestsof the easternUnited States, The studywas conducted in Green Ridge State Forest (GRSF), though few detailedstudies of it havebeen pub- Allegany County, Maryland. It is within the Ridge and Valley lished.Most reportsare of singlenesting occurr- physiographicregion (Stoneand Matthews 1977), characterized by narrowmountain ridges oriented northeast to southwestsepa- encesand general observations(Bolles 1890; Carter rated by steepnarrow valleys(see Titus 1980). 1925; Henderson 1933; Robertson 1959; Brown About 74% of the countyand nearly all of GRSF is forested 1962; Caldwell 1972; Hamerstrom1973; Appel- Major foresttypes were describedby Brushet al. (1980).Predom- gate 1975; Soucy 1976; Bird and Wright 1977; inant tree speciesinclude white oak (Quercusalba), red oak (Q. -
Phylogenies of Flying Squirrels (Pteromyinae)
Journal of Mammalian Evolution, Vol. 9, No. 1/2, June 2002 (© 2002) Phylogenies of Flying Squirrels (Pteromyinae) Richard W. Thorington, Jr.,'''* Diane Pitassy,^ and Sharon A. Jansa^ The phylogeny of flying squirrels was assessed, based on analyses of 80 morphological characters. Three published hypotheses were tested with constraint trees and compared with trees based on heuristic searches, all using PAUP*. Analyses were conducted on unordered data, on ordered data (Wagner), and on ordered data using Dollo parsimony. Compared with trees based on heuristic searches, the McKenna (1962) constraint trees were consistently the longsst, requiring 8-11 more steps. The Mein (1970) constraint trees were shorter, requiring five to seven steps more than the unconstrained trees, and the Thorington and Darrow (20(X)) constraint trees were shorter yet, zero to one step longer than the corresponding unconstrained tree. In each of the constraint trees, some of the constrained nodes had poor character support. The heuristic trees provided best character support for three groups, but they did not resolve the basal trichotomy between a Glaucomys group of six genera, a Petaurista group of four genera, and a Trogopterus group of four genera. The inclusion of the small northern Eurasian flying squirrel, Pteminys, in the Peiaiiiisia group of giant South Asian flying squirrels is an unexpected hypothesis. Another novel hypothesis is the inclusion of the genus Aemmys, large animals from the Sunda Shelf, with the Trogopterus group of smaller "complex-toothed flying squirrels" from mainland Malaysia and southeast Asia. We explore the implications of this study for future analysis of molecular data and for past and future interpretations of the fossil record. -
Habitat Fragmentation Provides a Competitive Advantage to an Invasive Tree Squirrel, Sciurus Carolinensis
Biol Invasions DOI 10.1007/s10530-017-1560-8 ORIGINAL PAPER Habitat fragmentation provides a competitive advantage to an invasive tree squirrel, Sciurus carolinensis Tyler Jessen . Yiwei Wang . Christopher C. Wilmers Received: 13 April 2017 / Accepted: 2 September 2017 Ó Springer International Publishing AG 2017 Abstract Changes in the composition of biological (Sciurus griseus) by non-native eastern gray tree communities can be elicited by competitive exclusion, squirrels (Sciurus carolinensis). We tested this wherein a species is excluded from viable habitat by a hypothesis along a continuum of invasion across three superior competitor. Yet less is known about the role study sites in central California. We found that within of environmental change in facilitating or mitigating the developed areas of the University of California at exclusion in the context of invasive species. In these Santa Cruz campus and city of Santa Cruz, S. situations, decline in a native species can be due to the carolinensis excluded S. griseus from viable habitat. effects of habitat change, or due to direct effects from The competitive advantage of S. carolinensis may be invasive species themselves. This is summarized by due to morphological and/or behavioral adaptation to the ‘‘driver-passenger’’ concept of native species loss. terrestrial life in fragmented hardwood forests. We We present a multi-year study of tree squirrels that classify S. carolinensis as a ‘‘driver’’ of the decline of tested the hypothesis that tree canopy fragmentation, native S. griseus in areas with high tree canopy often a result of human development, influenced the fragmentation. Future habitat fragmentation in west- replacement of native western gray tree squirrels ern North America may result in similar invasion dynamics between these species. -
Phylogenetic Relationships Among Six Flying Squirrel Gener,A Inferred from Mitochondrial Cytochrome B Gene Sequences
Phylogenetic Relationships among Six Flying Squirrel Gener,a Inferred from Mitochondrial Cytochrome b Gene Sequences 著者(英) Oshida Tatsuo, Lin Liang-Kong, Yanagisawa Hisashi, Endo Hideki, Masuda Ryuichi journal or Zoological Science publication title volume 17 number 4 page range 485-489 year 2000-05 URL http://id.nii.ac.jp/1588/00004215/ ZOOLOGICAL SCIENCE 17: 485–489 (2000) © 2000 Zoological Society of Japan Phylogenetic Relationships among Six Flying Squirrel Genera, Inferred from Mitochondrial Cytochrome b Gene Sequences Tatsuo Oshida1*, Liang-Kong Lin2, Hisashi Yanagawa3, Hideki Endo4 and Ryuichi Masuda1 1Chromosome Research Unit, Faculty of Science, Hokkaido University, Sapporo 060-0810, Japan 2Laboratory of Wildlife Ecology, Department of Biology, Tunghai University, Taichung 407, Taiwan 3Laboratory of Wildlife Ecology, Obihiro University and Veterinary Medicine, Obihiro 080-0843, Japan 4Department of Zoology, National Science Museum, Shinjuku-ku, Tokyo, 169-0073, Japan ABSTRACT—Petauristinae (flying squirrels) consists of 44 extant species in 14 recent genera, and their phylogenetic relationships and taxonomy are unsettled questions. We analyzed partial mitochondrial cyto- chrome b gene sequences (1,068 base pairs) to investigate the phylogenetic relationships among six flying squirrel genera (Belomys, Hylopetes, Petaurista, Petinomys, and Pteromys from Asia and Glaucomys from North America). Molecular phylogenetic trees, constructed by neighbor-joining and maximum likelihood methods, strongly indicated the closer relationship between Hylopetes and Petinomys with 100% bootstrap values. Belomys early split from other flying squirrels. Petaurista was closely related to Pteromys, and Glaucomys was most closely related to the cluster consisting of Hylopetes and Petinomys. The bootstrap values supporting branching at the deeper nodes were not always so high, suggesting the early radiation in the evolution of flying squirrels. -
Nest Box Utility for Arboreal Small Mammals in Vietnam's Tropical Forest Использование Дуплянок Для И
Russian J. Theriol. 10(2): 5964 © RUSSIAN JOURNAL OF THERIOLOGY, 2011 Nest box utility for arboreal small mammals in Vietnams tropical forest Ami Kato, Tatsuo Oshida, Son Truong Nguyen, Nghia Xuan Nguyen, Hao Van Luon, Tue Van Ha, Bich Quang Truong, Hideki Endo & Dang Xuan Nguyen ABSTRACT. To test nest box utility in a Southeast Asian tropical forest, we set 30 wooden nest boxes on trees in the Cuc Phuong National Park, Vietnam for a year. During the rainy season, we checked each nest box each month in the daytime. We expected that arboreal rodents might be more likely to use the nest boxes as shelter from the heavy rain. During dry season, we additionally checked each nest box every two months. We expected that nest boxes would be used as a shelter from the rain by small arboreal mammals, such as rats and flying squirrels in the rainy season more than in the dry season. During the rainy season, we found ants, bees, and birds mainly nested the nest boxes for reproduction: bees in April; ants from May to August; and birds from April to June. From the late rainy season to the dry season, arboreal small mammals mainly used nest boxes: rats from August to February and flying squirrel in December. Nest resource competition between birds and rodents may be minimal since they use cavities in different seasons. Also, unlike our expectation, it was preliminary suggested that arboreal small rodents would use more frequently nest box in the dry season than in the rainy season. KEY WORDS: flying squirrel, rat, rainy season, dry season, competition, nest resource, nest material. -
Periodic Status Review for the Western Gray Squirrel
STATE OF WASHINGTON February 2016 Periodic Status Review for the Western Gray Squirrel Gary J. Wiles Washington Department of FISH AND WILDLIFE Wildlife Program The Washington Department of Fish and Wildlife maintains a list of endangered, threatened, and sensitive species (Washington Administrative Codes 232-12-014 and 232-12-011). In 1990, the Washington Wildlife Commission adopted listing procedures developed by a group of citizens, interest groups, and state and federal agencies (Washington Administrative Code 232-12-297). The procedures include how species list- ings will be initiated, criteria for listing and delisting, a requirement for public review, the development of recovery or management plans, and the periodic review of listed species. The Washington Department of Fish and Wildlife is directed to conduct reviews of each endangered, threat- ened, or sensitive wildlife species at least every five years after the date of its listing by the Washington Fish and Wildlife Commission. In addition, as was the case with this western gray squirrel periodic status review, the Department may initiate a review of a species if a petition is received from an interested person setting forth specific evidence and scientific data to suggest that a species may be in need of reclassification. The periodic status reviews are designed to include an update of the species status report to determine whether the status of the species warrants its current listing status or deserves reclassification. The agency notifies the general public and specific parties who have expressed their interest to the Department of the periodic status review at least one year prior to the five-year period so that they may submit new scientific data to be included in the review. -
Tree Squirrel Habitat Selection and Predispersal Seed Predation in a Declining Subalpine Conifer
Oecologia DOI 10.1007/s00442-009-1496-5 PLANT-ANIMAL INTERACTIONS - ORIGINAL PAPER Tree squirrel habitat selection and predispersal seed predation in a declining subalpine conifer Shawn T. McKinney · Carl E. Fiedler Received: 11 February 2008 / Accepted: 22 October 2009 © Springer-Verlag 2009 Abstract DiVerential responses by species to modern variables were the least important in squirrel habitat selec- perturbations in forest ecosystems may have undesirable tion. Squirrel habitat use was lower and varied more in impacts on plant-animal interactions. If such disruptions P. albicaulis-dominant forests, and predispersal cone cause declines in a plant species without corresponding predation decreased linearly with increasing P. albicaulis declines in a primary seed predator, the eVects on the plant relative abundance. In Northern Rocky Mountain sites, could be exacerbated. We examined one such interaction where P. albicaulis mortality was higher and abundance between Pinus albicaulis (whitebark pine), a bird-dispersed, lower, squirrel predation was greater than in Central Rocky subalpine forest species experiencing severe population Mountain sites. In ecosystems with reduced P. albicaulis declines in the northern part of its range, and Tamiasciurus abundance, altered interactions between the squirrel and hudsonicus (red squirrel), an eYcient conifer seed predator, pine may lead to a lower proportion of P. albicaulis con- at 20 sites in two distinct ecosystems. Hypotheses about tributing to population recruitment because of reduced seed squirrel habitat preferences were tested to determine how availability. Reducing the abundance of competing conifers changes in forest conditions inXuence habitat use and will create suboptimal squirrel habitat, thus lowering cone subsequent levels of predispersal cone predation. -
Controlling Tree Squirrels in Urban Areas
L-1914 5-98 Controlling Tree Squirrels in Urban Areas Tree squirrels inhabit many parts of Texas. The Tree squirrels feed on a variety of material; nuts, three types of tree squirrels found in the state are fruit, seeds, berries, insects and bark make up the the fox squirrel, the gray squirrel and the flying bulk of their diet. Nesting sites include tree cavities, squirrel. Tree squirrels are interesting animals, but old woodpecker holes, attics, etc. If such places can- in urban and suburban areas they sometimes can not be found, tree squirrels will construct leafy be pests. nests in the branches of trees. Fox squirrels, the most common of the tree squir- rels, are found in east and central Texas. Gray squir- Biology and Reproduction rels, also known as “cat” squirrels, are found in the eastern part of Texas. Flying squirrels are the small- Fox Squirrels est of the tree squirrels, the least common, and the Adult weight: 11/2 to 21/2 pounds. only nocturnal tree squirrel. They are found in the eastern part of the state. Total length: 18 to 27 inches. Color: Brownish-gray with an orange underside. Tail: 7 to 14 inches, bushy. Gestation period: 42 to 45 days. Litter size: Average of three. Litter number: One to two per year, usually born in March or April and again in August or September. Weaning: 10 to 12 weeks. Life span: 4 to 7 years. Gray Squirrels Adult weight: 1 to 2 pounds. Total length: 16 to 20 inches. Color: Grayish. Tail: 7 to 10 inches, bushy. -
Northern Flying Squirrel (Glaucomys Sabrinus) Species Guidance Family: Sciuridae – the Squirrels
Northern Flying Squirrel (Glaucomys sabrinus) Species Guidance Family: Sciuridae – the squirrels Species of Greatest Conservation Need (SGCN) State Status: SC/P (Special Concern/Fully Protected) State Rank: S3S4 Federal Status: None Global Rank: G5 Wildlife Action Plan Counties with documented locations Mean Risk Score: 3.2 of northern flying squirrels in Wildlife Action Plan Area of Wisconsin. Source: Natural Heritage Photo by Ryan Stephens Importance Score: 2 Inventory Database, April 2013. Species Information General Description: The northern flying squirrel and its sister species, the southern flying squirrel (Glaucomys volans), both occur in Wisconsin. The northern flying squirrel is slightly larger than the southern flying squirrel, but is small compared to other tree squirrels. Adult northern flying squirrels in the Great Lakes region weigh 70-130 g (2.5-4.6 oz) (Kurta 1995). Total length (including tail) ranges from 245-315 mm (10-12 in), tail length 110-150 mm (4.3-5.9 in), hindfoot length 35-40 mm (1.4-1.6 in), and ear height 18-26 mm (0.7-1.0 in) (Jackson 1961, Kurta 1995). Pelage (fur) is silky and usually cinnamon–colored, but can range from dark brown to red. Belly hair is white at the tips and gray at the base. The northern flying squirrel uses its patagium (loose flap of skin between the front and hind legs; Fig. 1) for gliding between trees. A cartilaginous projection called a styliform process extends from the wrist (Fig. 1) to widen the patagium and enhance its effect (Wells-Gosling and Heaney 1989). Similar Species: The flying squirrels can easily be distinguished from the other eight Wisconsin squirrels by the presence of a furry patagium (flap of skin) that runs from the wrist to the ankle, and also by a cartilaginous projection on the wrist called a styliform process.