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Palaeoenvironmental Interpretation of the Triassic Sandstones of Scrabo
PALAEOENVIRONMENTAL INTERPRETATION OF THE TRIASSIC SANDSTONES OF SCRABO, COUNTY DOWN, NORTHERN IRELAND: ICHNOLOGICAL AND SEDIMENTOLOGICAL STUDIES INDICATING A MIXED FLUVIATILE-AEOLIAN SUCCESSION JAMES O. BUCKMAN, PHILIPS. DOUGHTY,MICHAEL J. BENTON and ANDREW J. JERAM (Received 19 May 1997) Abstract The Sherwood Sandstone Group at Scrabo, Co. Down, NorthernIreland, has been variously interpretedas aeolian or fluviatile. The sedimentological and ichnological data show that these sandstone-dominatedfacies were deposited within a mixed fluviatile-aeolian regime, in which fluviatile processes were responsible for the majorityof the preservedsedimentary sequence. The sequence contains a moderately diverse non-marine ichnofauna comprising the invertebratetrace fossils Biformites, Cruziana/Rusophycus(Isopodichnus), Herpystezoum(Unisulcus), Planolites, 'small arthropodtrackways', 'large arthropodtrackways' (cf. in part Paleohelcura), and the vertebrate trace fossil Chirotherium? (as well as a number of other, presently unnamed, vertebrate footprints). The recorded ichnofauna greatly increases that previously known from the Irish Triassic, comparesfavourably with that from the rest of Europe, and representsthe only known Irish locality for reptiliantrackways. Introduction north-east, are exposed at Scrabo Hill (Charlesworth 1963). Several Palaeogene The Sherwood Sandstone Group (Triassic) dolerite dykes and sills are exposed at the in north-eastIreland comprises part of a thick southern end of the hill, and these form a sequence of non-marine sediments, with a resistant cap that has shielded the sandstones maximum thickness of 1850m (Parnell et al. against removal by ice during the last 1992), which are rarely exposed and known glaciations. mostly from borehole data. At Scrabo (Fig. 1) Various environmental interpretationshave the Sherwood SandstoneGroup occurs within a been made of the Scrabo succession. The deep trough of Carboniferousto Triassic age succession has been interpretedas representing (Parnell et al. -
Sequence of Post-Moult Exoskeleton Hardening Preserved in a Trilobite Mass Moult Assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco
Editors' choice Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco HARRIET B. DRAGE, THIJS R.A. VANDENBROUCKE, PETER VAN ROY, and ALLISON C. DALEY Drage, H.B., Vandenbroucke, T.R.A., Van Roy, P., and Daley, A.C. 2019. Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco. Acta Palaeontologica Polonica 64 (2): 261–273. Euarthropods have a tough exoskeleton that provides crucial protection from predation and parasitism. However, this is restrictive to growth and must be periodically moulted. The moulting sequence is well-known from extant arthropods, consisting of: (i) the long inter-moult stage, in which no changes occur to the hardened exoskeleton; (ii) the pre-moult stage where the old exoskeleton is detached and the new one secreted; (iii) exuviation, when the old exoskeleton is moulted; and (iv) the post-moult stage during which the new exoskeleton starts as soft, thin, and partially compressed and gradually hardens to the robust exoskeleton of the inter-moult stage. Trilobite fossils typically consist of inter-moult carcasses or moulted exuviae, but specimens preserving the post-moult stage are rare. Here we describe nine specimens assigned to Symphysurus ebbestadi representing the first group of contemporaneous fossils collected that preserve all key stages of the moulting process in one taxon, including the post-moult stage. They were collected from a single lens in the Tremadocian part of the Fezouata Shale Formation, Morocco. Based on cephalic displacement and comparison to other trilobite moults, one specimen appears to represent a moulted exoskeleton. -
Copertina Guida Ai TRILOBITI V3 Esterno
Enrico Bonino nato in provincia di Bergamo nel 1966, Enrico si è laureato in Geologia presso il Dipartimento di Scienze della Terra dell'Università di Genova. Attualmente risiede in Belgio dove svolge attività come specialista nel settore dei Sistemi di Informazione Geografica e analisi di immagini digitali. Curatore scientifico del Museo Back to the Past, ha pubblicato numerosi volumi di paleontologia in lingua italiana e inglese, collaborando inoltre all’elaborazione di testi e pubblicazioni scientifiche a livello nazonale e internazionale. Oltre alla passione per questa classe di artropodi, i suoi interessi sono orientati alle forme di vita vissute nel Precambriano, stromatoliti, e fossilizzazioni tipo konservat-lagerstätte. Carlo Kier nato a Milano nel 1961, Carlo si è laureato in Legge, ed è attualmente presidente della catena di alberghi Azul Hotel. Risiede a Cancun, Messico, dove si dedica ad attività legate all'ambiente marino. All'età di 16 anni, ha iniziato una lunga collaborazione con il Museo di Storia Naturale di Milano, ed è a partire dal 1970 che prese inizio la vera passione per i trilobiti, dando avvio a quella che oggi è diventata una delle collezioni paleontologiche più importanti al mondo. La sua instancabile attività di ricerca sul terreno in varie parti del globo e la collaborazione con professionisti del settore, ha permesso la descrizione di nuove specie di trilobiti ed artropodi. Una forte determinazione e la costruzione di un nuovo complesso alberghiero (AZUL Sensatori) hanno infine concretizzzato la realizzazione -
Reassignment of Vertebrate Ichnotaxa from the Upper Carboniferous 'Fern
Reassignment of vertebrate ichnotaxa from the Upper Carboniferous ‘Fern Ledges’, Lancaster Formation, Saint John, New Brunswick, Canada MATTHEW R. STIMSON1,2*, RANDALL F. MILLER1, AND SPENCER G. LUCAS3 1. Steinhammer Palaeontology Laboratory, New Brunswick Museum, Saint John, New Brunswick E2K 1E5, Canada 2. New Brunswick Geological Surveys Branch, Department of Energy and Mines, Fredericton, New Brunswick E3B 5H1, Canada 3. New Mexico Museum of Natural History and Science, Albuquerque, New Mexico 87104, USA *Corresponding author <[email protected]> Date received: 14 April 2015 ¶ Date accepted: 04 September 2015 ABSTRACT Vertebrate ichnotaxa described by George Frederic Matthew in 1910 from the Upper Carboniferous (Lower Pennsylvanian) ‘Fern Ledges’ of Saint John, New Brunswick, were dismissed as dubious trackways by previous authors. Thus, three new ichnospecies Matthew described appeared in the 1975 Treatise on Invertebrate Paleontology as “unrecognized or unrecognizable” and were mostly forgotten by vertebrate ichnologists. These traces include Hylopus (?) variabilis, Nanopus (?) vetustus and Bipezia bilobata. One ichnospecies, Hylopus (?) variabilis, here is retained as a valid tetrapod footprint ichnotaxon and reassigned to the ichnogenus Limnopus as a new combination, together with other poorly preserved specimens Matthew labeled, but never described. Nanopus (?) vetustus and Bipezia bilobata named by Matthew in the same paper, have been reexamined and remain as nomina dubia. RÉSUMÉ Les ichnotaxons vertébrés de Fern Ledges du Carbonifère supérieur (Pennsylvanien inférieur) de Saint John, au Nouveau-Brunswick, décrits par George Frederic Matthew en 1910 avaient été rejetés par des auteurs précédents les considérant comme des séries de traces douteuses. Trois nouvelles ichnoespèces décrites par Matthew sont ainsi appa- rues dans le Treatise on Invertebrate Paleontology (Traité sur la paléontologie des invertébrés) de 1975 à titre d’espèces « non identifiées ou non identifiables » et ont principalement été laissées aux ichnologues des vertébrés. -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More By Michelle M. Casey1, Perry Kennard2, and Bruce S. Lieberman1, 3 1Biodiversity Institute, University of Kansas, Lawrence, KS, 66045, 2Earth Science Teacher, Southwest Middle School, USD497, and 3Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Middle level laboratory exercise for Earth or General Science; supported provided by National Science Foundation (NSF) grants DEB-1256993 and EF-1206757. Learning Goals and Pedagogy This lab is designed for middle level General Science or Earth Science classes. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens 3) to highlight major events or trends in the evolutionary history and ecology of the Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. They only lived in the oceans but occurred in all types of marine environments, and ranged in size from less than a centimeter to almost a meter across. They were once one of the most successful of all animal groups and in certain fossil deposits, especially in the Cambrian, Ordovician, and Devonian periods, they are extremely abundant. -
The Evolution of Trilobite Body Patterning
ANRV309-EA35-14 ARI 20 March 2007 15:54 The Evolution of Trilobite Body Patterning Nigel C. Hughes Department of Earth Sciences, University of California, Riverside, California 92521; email: [email protected] Annu. Rev. Earth Planet. Sci. 2007. 35:401–34 Key Words First published online as a Review in Advance on Trilobita, trilobitomorph, segmentation, Cambrian, Ordovician, January 29, 2007 diversification, body plan The Annual Review of Earth and Planetary Sciences is online at earth.annualreviews.org Abstract This article’s doi: The good fossil record of trilobite exoskeletal anatomy and on- 10.1146/annurev.earth.35.031306.140258 togeny, coupled with information on their nonbiomineralized tis- Copyright c 2007 by Annual Reviews. sues, permits analysis of how the trilobite body was organized and All rights reserved developed, and the various evolutionary modifications of such pat- 0084-6597/07/0530-0401$20.00 terning within the group. In several respects trilobite development and form appears comparable with that which may have charac- terized the ancestor of most or all euarthropods, giving studies of trilobite body organization special relevance in the light of recent advances in the understanding of arthropod evolution and devel- opment. The Cambrian diversification of trilobites displayed mod- Annu. Rev. Earth Planet. Sci. 2007.35:401-434. Downloaded from arjournals.annualreviews.org ifications in the patterning of the trunk region comparable with by UNIVERSITY OF CALIFORNIA - RIVERSIDE LIBRARY on 05/02/07. For personal use only. those seen among the closest relatives of Trilobita. In contrast, the Ordovician diversification of trilobites, although contributing greatly to the overall diversity within the clade, did so within a nar- rower range of trunk conditions. -
Innovations in Animal-Substrate Interactions Through Geologic Time
The other biodiversity record: Innovations in animal-substrate interactions through geologic time Luis A. Buatois, Dept. of Geological Sciences, University of Saskatchewan, 114 Science Place, Saskatoon SK S7N 5E2, Canada, luis. [email protected]; and M. Gabriela Mángano, Dept. of Geological Sciences, University of Saskatchewan, 114 Science Place, Saskatoon SK S7N 5E2, Canada, [email protected] ABSTRACT 1979; Bambach, 1977; Sepkoski, 1978, 1979, and Droser, 2004; Mángano and Buatois, Tracking biodiversity changes based on 1984, 1997). However, this has been marked 2014; Buatois et al., 2016a), rather than on body fossils through geologic time became by controversies regarding the nature of the whole Phanerozoic. In this study we diversity trajectories and their potential one of the main objectives of paleontology tackle this issue based on a systematic and biases (e.g., Sepkoski et al., 1981; Alroy, in the 1980s. Trace fossils represent an alter- global compilation of trace-fossil data 2010; Crampton et al., 2003; Holland, 2010; native record to evaluate secular changes in in the stratigraphic record. We show that Bush and Bambach, 2015). In these studies, diversity. A quantitative ichnologic analysis, quantitative ichnologic analysis indicates diversity has been invariably assessed based based on a comprehensive and global data that the three main marine evolutionary on body fossils. set, has been undertaken in order to evaluate radiations inferred from body fossils, namely Trace fossils represent an alternative temporal trends in diversity of bioturbation the Cambrian Explosion, Great Ordovician record to assess secular changes in bio- and bioerosion structures. The results of this Biodiversification Event, and Mesozoic diversity. Trace-fossil data were given less study indicate that the three main marine Marine Revolution, are also expressed in the attention and were considered briefly in evolutionary radiations (Cambrian Explo- trace-fossil record. -
Th TRILO the Back to the Past Museum Guide to TRILO BITES
With regard to human interest in fossils, trilobites may rank second only to dinosaurs. Having studied trilobites most of my life, the English version of The Back to the Past Museum Guide to TRILOBITES by Enrico Bonino and Carlo Kier is a pleasant treat. I am captivated by the abundant color images of more than 600 diverse species of trilobites, mostly from the authors’ own collections. Carlo Kier The Back to the Past Museum Guide to Specimens amply represent famous trilobite localities around the world and typify forms from most of the Enrico Bonino Enrico 250-million-year history of trilobites. Numerous specimens are masterpieces of modern professional preparation. Richard A. Robison Professor Emeritus University of Kansas TRILOBITES Enrico Bonino was born in the Province of Bergamo in 1966 and received his degree in Geology from the Depart- ment of Earth Sciences at the University of Genoa. He currently lives in Belgium where he works as a cartographer specialized in the use of satellite imaging and geographic information systems (GIS). His proficiency in the use of digital-image processing, a healthy dose of artistic talent, and a good knowledge of desktop publishing software have provided him with the skills he needed to create graphics, including dozens of posters and illustrations, for all of the displays at the Back to the Past Museum in Cancún. In addition to his passion for trilobites, Enrico is particularly inter- TRILOBITES ested in the life forms that developed during the Precambrian. Carlo Kier was born in Milan in 1961. He holds a degree in law and is currently the director of the Azul Hotel chain. -
An Inventory of Trilobites from National Park Service Areas
Sullivan, R.M. and Lucas, S.G., eds., 2016, Fossil Record 5. New Mexico Museum of Natural History and Science Bulletin 74. 179 AN INVENTORY OF TRILOBITES FROM NATIONAL PARK SERVICE AREAS MEGAN R. NORR¹, VINCENT L. SANTUCCI1 and JUSTIN S. TWEET2 1National Park Service. 1201 Eye Street NW, Washington, D.C. 20005; -email: [email protected]; 2Tweet Paleo-Consulting. 9149 79th St. S. Cottage Grove. MN 55016; Abstract—Trilobites represent an extinct group of Paleozoic marine invertebrate fossils that have great scientific interest and public appeal. Trilobites exhibit wide taxonomic diversity and are contained within nine orders of the Class Trilobita. A wealth of scientific literature exists regarding trilobites, their morphology, biostratigraphy, indicators of paleoenvironments, behavior, and other research themes. An inventory of National Park Service areas reveals that fossilized remains of trilobites are documented from within at least 33 NPS units, including Death Valley National Park, Grand Canyon National Park, Yellowstone National Park, and Yukon-Charley Rivers National Preserve. More than 120 trilobite hototype specimens are known from National Park Service areas. INTRODUCTION Of the 262 National Park Service areas identified with paleontological resources, 33 of those units have documented trilobite fossils (Fig. 1). More than 120 holotype specimens of trilobites have been found within National Park Service (NPS) units. Once thriving during the Paleozoic Era (between ~520 and 250 million years ago) and becoming extinct at the end of the Permian Period, trilobites were prone to fossilization due to their hard exoskeletons and the sedimentary marine environments they inhabited. While parks such as Death Valley National Park and Yukon-Charley Rivers National Preserve have reported a great abundance of fossilized trilobites, many other national parks also contain a diverse trilobite fauna. -
A Giant Rusophycus from the Arisaig Group (Siluro
A G I A N T RUSOPHYCUS FROM THE ARISAIG GROUP (SILURO-DEVONIAN) OF NOVA SCOTIA y. Tansathien and R.K. P ic k e r ill Department of Geology, U niversity of New Brunswick Fredericton, New Brunswick, E3B 5A3 Date Received October 16, 1986 Date Accepted December 24, 1986 A unique specimen of the trace fossil Rusophycus in the Moydart Formation (Ludlow) of the Siluro-Devonian Arisaig Group of northeastern Nova Scotia is at least 35 cm long, 18 cm wide and 8.5 cm deep. The specimen represents the largest Rusophycus yet described and can truly be regarded as a giant. It is speculated that the most likely producer of the trace fossil was a homalonotid trilobite. La Formation de Moydart (Ludlow) du Groupe siluro-dAvonien d'Arisaig, au nord-est de la Nouvelle-Ecosse, a livrA un exemplaire unique de l'ichnite Rusophycus mesurant au moins 35 cm an de long. 18 cm de large et 8.5 cm de profond. II s'agit 14 du plus grand Rusophycus jamais dAcrit et l'on est done en prAsence d'un vAritable gAant. Nous entrevoyons que 1'auteur de cette trace a pu etre un trilobite homalonotidA. [Traduit par le Journal] INTRODUCTION is exposed in the Arisaig area of northwestern Nova Scotia on the southern shores of Northumberland The trace fossil Rusophycus (Hall, 1852) is an Strait (Fig. 1). The succession consists of 1400- ichnogenus commonly, if not universally, attributed 1500 m of black to greenish grey mudstone with to arthropods, in particular to trilobites (in coarse-grained siltstone and fine-grained sandstone marine environments) or isopods or notostracan interbeds. -
Ordovician Stratigraphy and Benthic Community Replacements in the Eastern Anti-Atlas, Morocco J
Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco J. Javier Alvaro, Mohammed Benharref, Jacques Destombes, Juan Carlos Gutiérrez-Marco, Aaron Hunter, Bertrand Lefebvre, Peter van Roy, Samuel Zamora To cite this version: J. Javier Alvaro, Mohammed Benharref, Jacques Destombes, Juan Carlos Gutiérrez-Marco, Aaron Hunter, et al.. Ordovician stratigraphy and benthic community replacements in the eastern Anti- Atlas, Morocco. The Great Ordovician Biodiversification Event: Insights from the Tafilalt Biota, Morocco, 485, The Geological Society of London, pp.SP485.20, In press, Geological Society, London, Special Publication, 10.1144/SP485.20. hal-02405970 HAL Id: hal-02405970 https://hal.archives-ouvertes.fr/hal-02405970 Submitted on 13 Nov 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. The Geological Society Special Publications Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco --Manuscript Draft-- Manuscript Number: GSLSpecPub2019-17R1 Article Type: Research article Full Title: Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco Short Title: Ordovician stratigraphy of the Anti-Atlas Corresponding Author: Javier Alvaro Instituto de Geociencias SPAIN Corresponding Author E-Mail: [email protected] Other Authors: MOHAMMED BENHARREF JACQUES DESTOMBES JUAN CARLOS GUTIÉRREZ-MARCO AARON W.