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Introducción a Los Escarabajos Carabidae (Coleoptera) De Colombia

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Introducción a Los Escarabajos Carabidae (Coleoptera) De Colombia Introducción a los escarabajos Carabidae (Coleoptera) de Colombia Claudia Martínez Instituto de Investigación de Recursos Biológicos Alexander von Humboldt COLOMBIA, DIVERSA POR NATURALEZA INSTITUTO DE INVESTIGACIÓN DE RECURSOS BIOLÓGICOS ALEXANDER VON HUMBOLDT © Instituto de Investigación de Recursos Biológicos Alexander von Humboldt 2005 Los textos pueden ser utilizados total o parcialmente citando la fuente CONTRIBUCIÓN IAvH # 328 REVISIÓN TÉCNICA Claudia Moreno COORDINACIÓN EDITORIAL Claudia María Villa G. María Margarita Gaitán U. ILUSTRACIÓN Gilberto Mendoza Nora Mendoza DISEÑO Y DIAGRAMACIÓN Liliana Patricia Aguilar G. IMPRESIÓN Ramos López editorial Primera edición Impreso en Bogotá D. C., Colombia Mayo 2005 REPÚBLICA DE COLOMBIA Ministerio de Ambiente, Vivienda y Desarrollo Territorial CÍTESE COMO: Martínez C. 2005. Introducción a los escarabajos Carabidae (Coleoptera) de Colombia. Instituto de Investigación de Recursos Biológicos Alexander von Humboldt. Bogotá D. C., Colombia. 546 p. G E F ISBN 958-8151-46-5 PALABRAS CLAVE: 1. Claves 2. Diversidad 3. Insectos Banco Mundial 4. Neotrópico 5. Suramérica 6. Taxonomía Esta obra contribuye Embajada Real de los al Inventario Nacional de la Biodiversidad Paises Bajos FERNANDO GAST HARDERS Director General Sobre la autora SOBRE LA AUTORA Claudia Martínez [email protected] Bióloga con Maestría en Biología - línea sistemática de Universidad Nacional de Colombia. Su campo de interés es la entomología, en particular la taxonomía de insectos del orden Coleoptera. Ha publicado trabajos sobre el tema y espera seguir contribuyendo en el conoci- miento de esta área para el país y fuera de él. Introducción a los escarabajos Carabidae de Colombia (Coleoptera) 4 INSTITUTO DE INVESTIGACIÓN DE RECURSOS BIOLÓGICOS ALEXANDER VON HUMBOLDT Tabla de contenido TABLA DE CONTENIDO PRESENTACIÓN 19 PRÓLOGO 21 AGRADECIMIENTOS 23 INTRODUCCIÓN 25 CAPÍTULO 1. GENERALIDADES DE CARABIDAE 29 Aspectos de biología ................................................................................................................ 31 Morfología y glosario ............................................................................................................... 34 Sistemática de Carabidae ........................................................................................................ 55 Evolución ............................................................................................................................... 61 Estudios de Carabidae en Colombia ......................................................................................... 65 Materiales y métodos en campo y laboratorio ........................................................................... 66 CAPÍTULO 2. CLAVE PARA TRIBUS Y GÉNEROS DE CARABIDAE 73 Clave para las familias y tribus suramericanas de Geadephaga ................................................... 75 Key to adults of families and tribes of Geadephaga of South America ........................................ 91 Clave para subtribus y géneros de Carabidae en Colombia ......................................................... 100 CAPÍTULO 3. SINOPSIS DE LOS GÉNEROS DE CARABIDAE EN COLOMBIA 137 CAPÍTULO 3.1 SUBFAMILIA CARABINAE 139 Tribu Carabini ................................................................................................................... 139 Calosoma Weber, 1801 ............................................................................................. 139 Tribu Cicindelini ............................................................................................................... 141 CAPÍTULO 3.2 SUBFAMILIA SCARITINAE 141 Tribu Clivinini ................................................................................................................... 141 Ardistomis Putzeys, 1846 .......................................................................................... 141 Aspidoglossa Putzeys, 1846 ....................................................................................... 145 Clivina Latreille, 1802 .............................................................................................. 147 Oxydrepanus Putzeys, 1866 ...................................................................................... 149 Schizogenius Putzeys, 1846 ....................................................................................... 151 Semiardistomis Kult, 1950 ....................................................................................... 153 Tribu Scaritini ................................................................................................................... 155 Subtribu Forcipatorina ................................................................................................. 155 Camptodontus Dejean, 1826 ..................................................................................... 157 Forcipator Maindron, 1904 ....................................................................................... 157 Stratiotes Putzeys, 1846 ........................................................................................... 159 Subtribu Scaritina ........................................................................................................ 161 Scarites Fabricius, 1775 ............................................................................................ 161 Tribu Siagonini ................................................................................................................. 163 Enceladus Bonelli, 1803 ........................................................................................... 163 Introducción a los escarabajos Carabidae de Colombia (Coleoptera) INSTITUTO DE INVESTIGACIÓN DE RECURSOS BIOLÓGICOS ALEXANDER VON HUMBOLDT 5 Tabla de contenido CAPÍTULO 3.3 SUBFAMILIA PAUSSINAE 165 Tribu Brachinini ............................................................................................................... 165 Brachinus Weber, 1861 ............................................................................................ 166 Pheropsophus Solier, 1833 ......................................................................................... 169 Tribu Ozaenini .................................................................................................................. 172 Ozaena Olivier, 1811 ................................................................................................ 172 Pachyteles Perty, 1830 .............................................................................................. 174 Physea Brullé, 1834 .................................................................................................. 177 Platycerozaena Bänninger, 1927 ............................................................................... 179 Tribu Paussini ................................................................................................................... 181 Homopterus Westwood, 1841 .................................................................................... 181 CAPÍTULO 3.4 SUBFAMILIA BROSCINAE 183 Tribu Cymbionotini ........................................................................................................... 183 Cymbionotum Baudi di Selve, 1864 ........................................................................... 183 CAPÍTULO 3.5 SUBFAMILIA PSYDRINAE 185 Tribu Bembidiini ............................................................................................................... 185 Subtribu Bembidiina ..................................................................................................... 185 Bembidion Latreille, 1802 ......................................................................................... 185 Subtribu Tachyina ........................................................................................................ 187 Elaphropus Motschulsky, 1839 .................................................................................. 187 Meotachys Erwin, 1974 ............................................................................................. 189 Micratopus Casey, 1914 ............................................................................................ 191 Paratachys Casey, 1918 ............................................................................................ 193 Pericompsus LeConte, 1851 ....................................................................................... 195 Polyderis Motschulsky, 1862 ..................................................................................... 197 Tachys Dejean, 1821 ................................................................................................ 199 Subtribu Xystosomina .................................................................................................... 201 Batesiana Erwin, 1994 .............................................................................................. 201 Geballusa Erwin, 1994 .............................................................................................. 203 Gouleta Erwin, 1994 ................................................................................................. 205 Mioptachys Bates, 1882 ............................................................................................ 207 Tribu Pogonini .................................................................................................................. 209 Diplochaetus Chaudoir, 1871 ...................................................................................
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