ARQUIVOS DO MUSEU BOCAGE

Nova Série, VoLID, N.o 20, pp. 529-550 30 de Agosto de 2003

TWENTY FIVE NEW RECORDS ON PORTUGUESE FUNGUS GNATS (DIPTERA: )

EUGÉNIA RIBEIRO (1.2)

INTRODUCTION

The mycetophilids or fungus gnats are dipterous whose larvae show some degree af assodation with fungi, mainly agarics, using them as feeding or breeding sites. The adults have very conspicuous characters as a highly arched thora<, prominent coxae and long legs with tibial spurs, being usually found in shady and humid areas. Sixty five mycetophilids species were known for Portugal in Ribeiro (1991 c; see also Ribeiro 1990 a, 1990 b, 1991 a, 1991 b, 1992). Chandler (1993) showed that the known fUl1gorum (De Geer) in more southern countries was another species, Mycetophila perpaLlida Chandler. After checking Qur material we realised lhat ali material -:identified as M. fUl1gorul1l was in fact M. perpallida. Also Qur material previously identified as Mycetophila ruficollis Meigen is Mycetophila britamlica Lastovka & Kidd. Meanwhile, nine more species were added to the Portuguese entomofauna [Carles-Tolrá, 2001; Carles-Tolrá (ed.) (2002)]. The material of this study was catched with Malaise traps setted in Herdade da Ribeira Abaixo (Grândola, Portugal) (Fig. I). Twenty five species of mycetophilids are new records to Qur country and twenty others are shawn as new

(I) Departamento de Biologia , Faculdade de Ciênci~s da Universidade de Lisboa" Campo Grande, Bloco C2, 1749-016 Lisboa - PORTUGAL. (2) Centro de Biologia Ambiental, Faculdade de Ciências da Universidade de Lisboa, Campo Grande, Bloco C2, 1749-016 Li sboa - PORTUGAL. 530 Eugénia Ribeiro for that area. One thousand , >Cour hundred and seventy six specimens distributed by forty five speeies of fungus gnats were eoll eeted·(Tab. I). For eaeh genus some generie data will be given and for eaeh myeetophilid speeies data wi ll be listed aeeording to the type-Ioeality, date(s) of eolleetion and number of males and females eolleeted. A sign (*) is given to the new reeords and their distribution is referred according to the catalogue af Palaearctic Diptera, S60s and Papp (1988). The site for eaeh speeies is not indieated beeause ali the material belong to Herdade da Ribeira Abaixo (Grândola).

STUDIED MATERIAL

Acnemia Winnertz Aellemia Winnertz, 1863: 798. Type species: Leia Ilitidieollis Meigen, 1818: 255 (designation by Johannsen, 1909: 63).

Revised by Zaitzev (I 982a, 1982b), it is mainly a Holaretie genus, not showing many species. From Qur material we found two species, new records to IHortugal: Acnemia Qmoena Winnertz, a yellowish species presenting macro and microtrichia OTI lhe wings and Acnemia nitidicollis (Meigen), a clark species with only macrotrichia present on lhe wings.

* AC{lemia amoena Winnertz, 1863 Acnemia amoena. Verh. zool.-bot. Ges. Wien, 13: 800 . . Type-Ioeality: "Sammlung des Herrn Seantors von Heyden in Frankfurt am Main" (?distriet ofFrankfurt) [D]. Material examined: l-V to 20-V-1997, 2 33 + 2 'j''j'; 20-V to 2-VI-1997, I 3 ; 2-VI to 27-VI-1997, 7 33 + I 'j' ; 5-VIII to 4-IX-1997,'2 33 + 3 'j''j'; 2-XI to 27-XI-1997,2 'j''j'; ll-V 10 ll-VI-1998, 1 'j' ; ll-X to 3-XII-1998, 1 'j' ; 25-I1I to 24-IV-2000,3 33 + 1 'j' ;22-VIO 18-VI-2000, I 'j' ;9-VIIto 15-VIII-2000, 1 'j' . Distribution: Widespread in Europe and also known from Japan.

* Acnemia nitidicollis (Meigen, 1818) Leia Ilitidieollis. Syst. Besehr., 1: 255. Type-Ioeality: not given (?Stolberg) [D] Material examined: 27-VI to 4-VII-1997, 1 3 + 2 'j''j' ; IS-III to 8-IV-1998, 1 3 ; 8-IV to 13-V-1998, 1 3 ; 13-V to 11-VI-1998, 1 3 + 2 'j''j'; 25-I1I to 24-IV-2000, 1 'j'. Distribution: Widely di~tributed in Europe and known from Japan . . Twenty Five Records on Portuguese Fungus Gnats 531

Allodia Winnertz Allodia Winnertz, 1863: 826. Type species: Mycetophila omaticollis: \Vinnertz, 1863: 830 (designation Johannsen, 1909: 104) [=Mycetophila lllgells (Wiedemann, 1817: 68), mi sidentification] .

Adult fungus gnats from this genus can be found in wet woods and larvae breed in fungi , mainly agaries. Tuomikoski (1966) aeeept two subgenera in Allodia' genus: ALLodia sensu stricto and Brachycampta. Qur material incorporate ALLodia ornaticollis (Meigen) and Allodia barbala (Lundstrom), this one as a new reeord to Portugal. Eaeh speeies belongs, respeetively, to eaeh subgenus indieated abave .

• Allodia barbata (Lundstrom, 1909) Brachycampta barbala. Acta Soe. Falllla Flora!ellll. , 32 (2): 26. Type-Iocality: Karislojo (Finland). Material examined: 13-IV to I-V-1997, I rJ. Distribution: Known in many countries ofEurope.

Allodia ornaticollis (Meigen, 1818) Mycetophila ornaticollis. Syst. Besehr., 1: 269. Type· ldeality: nol given (Germany). \ .. Matehil"il"X amitÍéd: 15-I1I to 8-IV-1998, I rJ ; 8-IV to 13-V-1998, I rJ ; 21-1 to 23- II-2000, I rJ ; 25-I1I to 24-IV-2000, I rJ + I

Allodia sp. Material examined: 27-XI to 18-XII·1997, I

Boletina Staeger Boletilla Staeger, 1840: 233. Typé- ilpeeies, . Leia· Irivittata Meigen, 1818: 258 (designation Johannsen, 1909: 739;:

It is quite a widespreal!h egenus . with a large number of Palaearctic species. However, the biology of most speeies is still fairly unknown.

Boletina gripha Dziedzieki, 1885 Boletilla griplza. Pam. fi zyogr., 5: 9. Type-Iocality: "Zaezernie v Bialorussii" (USSR). Material examined: 2-XI to 27-XI·1997, 8 rJrJ + 2

4 r3r3 ; 20-XII-1999 to 22-1-2000, 3 r3r3 ; 28-X to 26-XI-2ooo, 5 r3r3 + 2 ~~ ; 21-1 to 19-11-2001, I ~ .

Bolitophila Meigen" Bolilophila Meigen, 1818: 220. Type speeies: Bolilophila ci/lerea Meigen, 1818: 221 (designation Westwood, 1840: 127).

Larvae inhabit fungi , mainly agaric species and adults showing medium size and a slender body shape ean be found along streams in woods.

Bolitophila pseudohybrida Landrock, 1912 Bolilophila pseudohybrida. Berl. e/ll. Z., 57: 45. Type-Iocality: Strzygi (Poland). Material examined: 2-XI to 27-XI-1997, 2 ~~; 19-V to 26-VI-1999, I ~ ; 17-X to 20-XI-1999, I ~; 20-XII-1999 to 22-1-2000, I ~; 22-1 to 23-11-2000, I r3 + 2 ~~ ; 23-11 to 25-I1I-2ooo, 2 ~~; 28-X to 26-XI-2ooo, 3 r3r3 + 2 ~~ ; 26-XI to 22-XII- 2000,2 r3r3 + 5 ~~ .

Brevicornu Marshall Brevicomu Marshall, 1896: 306. Type species: Brevicomu fiavum Marshall, 1896: 307 (designation Tuomikoski, 1966: 185).

The genus is well represented in the Palearetie region and Tuomikoski (1966) paid attention to this genus establishing Stigmatomeria as a second subgenus after lhe existing 8revicornu sensu stricto. When lhe femates do not appear associated with males, we consider them in this work as Brevicornu sp ..

* Brevicornufuscipenne (Staeger, 1840) Mycelophilafuscipe/llle. Nalurl •. Tidsskr., 3: 259. Type-Iocality: not given (Germany). Material examined: 27-XI to 18-XII-1997, I r3; 20-XII-1999 to 22-1-2000, I r3; 25-I1I to 24-IV-2oo0, I r3, Distribution: Known fram many European countries.

* Brevicornu griseicoUe (Staeger, 1840) Mycelophila griseicolle. Naturl •. Tidsskr., 3: 258. Type-Iocality: not given (Denmark). Twellty Five Records 011 Porlllguese FUIIgus GI/ ats 533

Material examined: 2-VI to 27-VI- 1997, 3 aa;1 8-XII-1997 to 6-1- 1998, I a; 6-1 to 6-11-1998, 4 aa; 15-111 to 8-1V-1 998, I a + 2 'i''i' ; 8-IV to 13-V- 1998, 2 aa + I I 'i''i' ; 20-XII- 1999 to 22-1-2000, I a; 25-TIl to 24-IV-2000, I a. Distribution: Most countries in Europe.

* 8revicorllll Iligrofllscllm (Lundstrom, 1909) Brachycampta lligrofuscllIll. Acta Soco Fauna Florafenn., 32 (2): 27. Type-Ioeality: Karislojo (Finland). Material examined: 17-X to 2 1-XI-1999, I a + I 'i' . Distribution: Many countri es in Europe.

Brevicornlt sericoma (Meigen, 1830) Mycetophila sericoma. Syst. Beschr., 6: 302. Type·locality: not given ("aus hiesiger Gegend") (Aachen). Material examined: 24-Il to 15-111-1 998, 3 aa; 15-111 to 8-IV- 1998, I a; 8-IV to J3 -V-1998, 3 aa + 3 'i''i' ; 3 1-1 to 7-11-1999, I a + I 'i' ; 21-XI to 20-XIl-1999, 3 aa;23-Il to 25-1II-2oo0, 2 aa;25-111 to 24-IV-2ooo, 4 aa + I 'i' ; 25-111 to 24- IV-2000, 2 aa;22-V to 18-VI-2000, I a + 4 'i''i' ; 15-VIII to 24-IX-2000, I a; 2 1-1 to 19-11-2001,5 aa;1 9-11 to 25-111-200 1, I a.

8revicorllll sp. Material examined: 2-VI to 27-VI-1997, 2 'i''i' ; 27-XI to 18-XII-1997, 2 'i''i'; 18- XIl-1997 to 6-1-1 998, 3 'i''i' ; 6-1 to 6-11 -1998,2 'i''i'; 6-1I to 24- Il-1998, I 'i' ; 24-1I to 15-111- 1998, 4 'i''i' ; 15-111 to 8- IV- 1998, 6 'i''i' ; 8-IV to 13-V-1998, 4 'i''i' ; 13-V to II-VI-1998, 3 'i''i' ; 31-1 to 7-1I- 1999, I 'i' ; 21-XI to 20-XII-1999, I 'i' ; 20-XII- 1999 to 22-1-2000, 5 'i''i' ; 22-1 to 23-11-2000, 2 'i''i' ; 23-Il to 25-III-2oo0, I 'i' ; 25- m to 24-IV-2ooo, 3 'i''i' ; 25-1IJ to 24-IV-2oo0, 6 'i''i' ; 22-V to 18-VI-2000, 2 'i''i' ; 9-VIl to 15-VIII-2ooo, I 'i' ; 15-VIII to 24-IX-2ooo, I 'i' ; 2 1-1 to 19-Il-2ool, 2 'i''i'.

Cordyla Mei gen Cordyla Meigen, 1803: 263. Type speeies: Cordy/ajusea Meigen, 1804: 93 (slIbsqllent mon.).

T he genus is formed by tiny fungus gnals. The adults can be found in woods and larvae ha ve been found breeding in fungi or in rotten \Voad . Some females were nOl po ss ible to identify at lhe spec ies levei, bei ng named in thi s work as Cordyla sp ..

Cordyla brevicornis (Staeger, 1840) Pachypalpus brevicomis. Na lllrh. Tidssk r., 3: 269. Type-Iocality: "Fredriksbergs Sloth ave" (Denmark). 534 Eugénia Ribeiro

Material examined: 20-V to 2-VI-1997, 4 1313 + 4 'jl'jl ; 8-IV to 13-V-1998, 3 1313 ; 13-V to II-VI-1998, 21313 ; II-VI to 14-VII-1998, 7 1313 .

* Cordyla crassicomis Meigen, 1818 Cordyla erassieomis. Sys/. Bese"r., 1: 275. Type-I ocali ty: Austria. Material examined: 13-IV to I-V- 1997, 2 1313 ; l -V to 20-V- 1997, I 13 ; 6-1 to 6- Il- 1998, I 13 ; 8-IV to 13-V-1998, 3 1313 ; 13-V to II-VI- 1998, 2 1313 ; 16-IV to 19- V- 1999, 3 1313 ; 19-V to 26-VI-1999, I 13 ; 21-XI to 20-XII-1999, 113 ; 22-1 to 23- II-2000, I 13 ; 25-III to 24-IV-2000, 2 1313 ; 22-XII-2000 to 21-1-2001, I 13 ; 22-IV to 19-V-2ool , 113 ; 18-VI to 15-VII-2ool , 113 . Distribution: Most parts of Europe.

* Cordylafasciata Meigen, 1830 Cordyla faseia/a. Syst. Bese"r., 6: 304. Type-Iocality: "Gegend von Berlin". Material examined: 27-XI to 18-XII-1997, 5 1313 ; 18-XII-1 997 to 6-1-1998, 113 + 2 'jl'jl ; 6-1 to 6-II-1998, I 13 ; 24-1I to 15-III-1998, I 13 ; 8-IV to 13-V- 1998, 2 'jl'f ; 25-lll to 24-IV-2ooo, 113 . Distribution: Widely distributed in Europe.

Cordylafusca Meigen, 1804 Cordylafllsca. Klass. Bese"r. , 1: 93. Type-Iocality: Aachen (D). Material examined: 2-VI to 27-VI-1 997, 3 1313 ; 27-VI to 4-VII-1 997, I 13 ; 18-X to 2-XI-1997, I 13 ; 2-XI to 27-XI-1997, 6 1313 ; 27-XI to 18-XII-1997, I 13 ; 18-XII- 1997 to 6-1-1998, 53 1313 + 24 'jl'jl ; 6-1 to 6-1I- 1998, 113 ; IS-III to 8-IV- 1998, 113 ; 8-IV to 13-V-1998, I 13 + I 'jl ; 3-XII-1998 to 31-1-1 999, I 13 ; 25-lll to 24-IV- 2000,2 1313 ; 28-X to 26-XI- 2ooo, I 13 ; 18- VI to 15-VII-2ool, I 13 .

* Cordyla pusilla Edwards, 1925 Cordyla pusilla. Tralls. R. em. Soe. Lond., 1924: 615. Type-Iocality: Shefford, Beds. (Great Britain). Material examined: 13-I V to I-V-1 997, 2 1313 ; l -V to 20-V-1997, I 13 ; 2-VI to 27-VI-1997,3 1313 ; 24-1I to 15-III-1998, 5 1313 ; 15-IIl to 8-IV- 1998, 2 1313 ; 8-IV to 13-V-1998, 4 1313 ; 13-V to II-VI-1998, I 13 ; II-VI to 14-VII-1998, 3 1313 ; 3- XII-1998 to 31-1-1999, I 13 ; 20-VIII to 9-IX-1999, I 13 ; 19-II to 25-1II-2ool , I 13 . Distribution: Some countries in Europe.

Cordyla sp. Material examined: 13-IV to I-V- 1997, I 'jl ; l-V to 20-V- 1997, 2 'jl'jl ; 2-VI to 27- VI-1 997,45 'jl'jl ; 27-VI to 4-VIl- 1997, 1 'jl ; 18-X to 2-XI- 1997, 1 'jl ; 2-XI to 27- Twellry Five Records 011 Portuguese FllIlgUS Gnats 535

XI-1997 , 30 'j''j' ; 27-Xl to 18-XlI-1997, 3 'j''j' ; 6-1 to 6-11-1998, I 'j'; 24-11 to 15- III-1998,4 'j''j'; 15-1II to 8-IV-1998, 2 'j''j' ; 8-IV to 13-V-1998, 16 'j''j'; 13-V to ll-VI-1 998,2 'j''j'; II-VI to 14-VII-1998, I 'j' ; 31-1 to 7- 11-1999, 6 'j''j' ; 19-V to 26-VI-1999,4 'j''j'; 21-XI to 20-XII-1999, I 'j' ; 20-XII-199910 22-1-2000, I 'j' ;22- I to 23- 11-2000, 5 'j''j' ; 23- 11 to 25-I11-2000, I 'j' ; 25-I1I to 24-IV-2ooo, 2 'j''j' ; 22-V to 18-VI-2ooo, 2 'j''j' ; 19-11 to 25-I1I-2ool, I 'j' ; 22-IV to 19-V-2ool, I 'j' ; 19-V to 18- VI-2ool , I 'j'; 18-VI to 15-VII-2ool, 5 'j''j' .

Ectrepestholleura Enderlein Ectrepestholle/lra Enderlein, 19l1 : ll5. Type species: Tetragonellra hirta Winnertz, 1846: 19 (origin al designation).

Vockeroth (1980), as well as other authors eonsidered that Eetrepesthollellra was so related to the Telragoll eura genus, that it should not be co nsidered an autonomous genus. However, lhrough severa I eharaeters e handl er (1980) showed that in the Palaearctic region lhe two genera can be distinguished, being independent from eaeh other.

• EctrepeSlllOnellra hirta (Wi nnertz, 1846) Tetragoneura hirta. Stettill. enl. Ztg., 7: 19. Type-Ioeality: not given (?Krefeld, lee totype designation). Material examined: l-V to 20-V- 1997, 4 'j''j'; 20-V to 2-VI-1997, 2 éSéS + I 'j'; 2-VI to 27-VI-1997, 4 éSéS + I 'j' ; 27-VI to 4-VII-1997, I éS + I 'j' ; 13-V to 11- VI-1998, 6 éSéS + 2 'j''j' ; li-VI to 14-VII -1998, I 'j' ; 14-VII to 14-VIII-1998, I éS; li-X to 3-XII-1998, I éS + I 'j'; 19-V to 26-VI-1999, I éS ; 26-VI to 23-VII- 1999, I 'j' ; 20-XIl-1999 to 22-1-2000, I 'j'; 25-JIl to 24-IV-2000, I 'j' ; 24-IX to 28-X-2000, I 'j' ; 18-VI to 15-VII-2ool, I 'j'. Distribution: Widely distributed in Europe.

Exechia Winnertz Exechia Winnertz, 1863: 879. Type species: Tip/l/a fimgorum: Winnertz, 1863: 886, misidentified as Exeehia fimgarum (De Geer) (designated Joh annsen, 1909: 106) [= fi/sea (Meigen, 1804)].

It is a widespread genus in ali zoogeographieal regions belonging from the Exechiini tribe. Tuomikoski (1966) separated th e genus in several genera. Our material belongs to eleven different species, fi ve of lhem being new to Portugal.

Exechin bicincla (Staeger, 1840) Mycetophila bicillcta. Naturh. Tidsskr., 3: 263. Type-Iocality: not given (Denmark). 536 Eugénia Ribeiro

Material examined: 13-V to ll-VI-1998, I Ô; 20-XIl-1999 to 22-[-2000, I 'j' ; 22-1 to 23-Il-2000, I Ô + I 'j' ; 23-Il to 25-IlI-2000, I 'j' ; 25-lII to 24-IV-2000, I Ô.

* Exechia conflnis Winnertz, 1863 Exechia cOllfinis. Verll. zoo/. -bot. Ges. Wi ell , 13: 892. Type-Iocality: not given (?Krefeld) [O). Material examined: 27-XI to 18-XIl-1997, I Ô; 18-XIl-1997 to 6-1- 1998, 2 ÔÔ; 28-X to 26-XI-2000, 4 ÔÔ; 26-XI to 22-XIl-2000, 4 ÔÔ. Distributi on: MOSl countries in Europe.

* Exechia deli/ata Lundstrom, 1916 Exechia dewafa. AI111ls !tist.-nat. Mus. nam. lumg., 14: 75. Type-Iocality: Kovácspatak (Czechoslovakia). Material examined: 3 1-1 to 7-[[-1 999, I Ô; 9-IX to 17-X-1999, I Ô; 17-X to 20- X[-1999, I Ô; 20-XI to 20-XIl- 1999, I Ô; 20-XII-1999 to 22-1-2000, I Ô; 22-1 to 23-11-2000, I Ô + I 'j' ; 23-Il to 25-IlI-20DO, I 'j' ; 25-I11 to 24-IV-2000, I Ô; 28- X to 26-XI-2oo0, I Ô. Distribution: Known in some European countri es.

Exechia dorsalis (Staeger, 1840) Mycetophila dorsalis. Naturll. Tidsskr. , 3: 262. Typc-Iocality: not given (Oenmark). Material examined: 18-XII-1997 to 6-[- 1998, I Ô; ll-VI to 14-VII-1998, I Ô; 21- XI to 20-XII-1999, I Ô; 22-1 to 23-11-2000, I Ô + I 'j'; 23-Il to 25-III-2000, I 'j' ; 25- I1I to 24-IV-20oo, 3 ÔÔ + 3 'j''j' .

Exechiaflllva Santos Abreu, 1920 Exechiaflllva. Mems. R. Acad. eienc. Artes Barcelona, 16: 78. Type-Iocality: Fuente Bermeja, La Palma (Canary Islands). Material examined: 2-XI to 27-XI-1997, I Ô + I 'j' ; 18-XIl-1997 to 6-1-1998, 2 ÔÔ; 6-1 to 6-11-1998, 3 ÔÔ; 15-III to 8-IV- 1998, 4 ÔÔ + I 'j' ; II-V[ to 14-VIl- 1998, I Ô; 31-1 to 7-Il- 1999, 3 ÔÔ + 2 'j''j'; 21-XI to 20-XII-1999, 5 ÔÔ; 20-XII- 1999 to 22-1-2000, 2 ÔÔ.

Exechia jilSca (Meigen, 1804) Myceropllila jllsca. Kla ss. Bescll,.. , 1: 9 \. Type-Iocality: not given (?Germany) . Material examincd: 20-V to 2-VI-1997, I Ô; 27-XI to 18- XIl-1997, 4 ÔÔ + 7 'j''j' ; 18-XIl-1997 to 6-1-1998, 9 ÔÔ + 14 'j''j' ; 6-1 to 6- Il-1 998, I Ô; 8-IV to 13-V- 1998, 5 ÔÔ + 2 'j''j' ; 21-XI to 20-XIl-1999, 5 ÔÔ; 20-XII- 1999 to 22-1-2000, 2 ÔÔ; 22-1 to 23-Il-2oo0, 3 ÔÔ + 2 'j''j' ; 23- 11 to 25-I1I-2oo0, 3 ÔÔ + 2 'j''j' ; 25-III to 24-IV-2000, 3 ÔÔ; 28-X to 26-XI-2000, 2 ÔÔ + 8 'j''j' ; 26-XI to 22-XII- 2000,3 ÔÔ; 21-lto 19-[[-2001,2 ÔÔ. Twenty Five Records 011 Porruguese FUf/ gus Gf/ats 537

Exechia fusca group Material exam ined: 2-VI to 27-VI- 1997, I

* Exechia Iligrosclllella/a Landrock, 1912 Exechia lIigrosclltellata. Wi ell. Ellt. Ztg. , 31: 309. Type-Iocality: "Mahren, Josefstal bei Adamstal" (= Adamov ValJ ey) (Czechoslovakia). Material examined: 18-XII- 1997 to 6-1-1998, 3 ~~ . Distributi on: Widely distributed in Europe and also known from Mongolia in Asia .

• Exechia repallda Johannsen, 1912 Exechia repallda. BulI. Me. Agric. Exp. SIII. , (2) 200: 73. Type-Iocalit y: Boston, Mass. (USA). Material examined: 27-IX to 18-X-1997, I ~ . Distribution: Most European countries and fram North America.

* Exechia repalldoides Caspers, 1984 Exechia repandoides. Ze irsch. fllr Ent. , 5: 173-205. Type-Iocality: Teichbach (Lunz) Material examined: 8-IV to 13-V-1998, I ~ . Distribution: Known for many countri es in Eurape.

Exechia r epa Ilda group Material examined: 22-1 to 23-11-2000, I

Exechia seria/a (Meigen, 1830) Mycelophila seriala. Sysl. Beschr. , 6: 302 . Type-Iocality: "Berliner Gegend". Material examined: 2-VI to 27-VI-1997, I ~ + 2

Exechia spÍllllligera Lundstrom, 1912 Exechia spimdigera. Acta Soco Fauna Florafenll., 36 (1): 33. Type-Iocalities: Kuusto, Karislojo, Vasa (Finl and) and Sorta vala (USSR). Material examined: 6-1 to 6- I1-1998, 2 ~~ . 538 Eugénia Ribeiro

MOlloe/olla Mik Monoe/ol/a Mik, 1886: 279. Type speeies: Seiophila hallerata Staeger, 1840: 275 (a u!.) (=nifilalera (Walker, 1837».

Monoe/ona Mik is represented by a small number of species. Aeeording to Hutson el aI. (1980) the four speeies described from Europe belong to a single one, Monoelona rufilalera (Walker).

* Monoe/olla rufllatera (Walker, 1837) Sciophila rufilatera. Em. Mag., 4: 115. Type-Ioeality: "Near London" (Great Britain). Material exami ned: 2-VI to 27-VI- 1997, I <3' ; 2-XI to 27-XI-1997, I <3' ; 24-11 to 15-I1I-1998, I '1' ; 8-IV to 13-V-1998, 2 <3'<3' ; 13-V to II-VI-1998, I '1' ; 25-I1I to 24-IV-2000, I '1' . Distribution: Widely distributed in Europe.

Mycetophi/a Meigen Mycelophila Meigen, 1803: 263. Type speeies: Tipula agarici Vi ll ers, 1789: 393 (designation Johannsen, 1909: 116).

It is a widespread genus which had th e attenti on of researehers as Laffoon (1957) in the Nearetie region and Lastovka (1972) and Lastovka & Kidd (1975) contributions in the Palearctic regian. Specimens from eleven spec ies were collected, seven of them being new records for Portugal. Some females bel ong to a eomplex of species named as Mycelophila nificollis Meigen, 1818 (group). Some other femates are designated as Mycetoplzila sp. because it was not possible to reac h lhe specific leveI.

Mycetophi/a alea Laffoon, -1965 Mycetophila alea. , in Stone & aI.: A catalog of lhe Diplera of America NorrhofMexico: 210. Type-Iocality: not given (Europe). Material examined: 20-V to 2-VI-1997, I '1' ; 2-VI to 27-VI-197, 20 <3'<3' + 8 '1''1' ; 27-VI to 4-V II -1997, I <3' ; 2-XI to 27-XI- 1997, I '1' ; 27-XI to 18-XII-1997, I <3' + 2 '1''1' ; 13-V to II-VI-1998, I <3' + I '1' ; li-X to 3-XII-1998, I '1' ; 25-I1I to 24-IV- 2000, 1 <3' + I '1' ; 9-VII to 15-VIII-2000, 1 '1' .

Mycetophila britannica Lastovka & Kidd, 1975 MycelOphila britclIlllica. Ellfol11ologist's 111011. Mag., 110: 203. Twenty Five Records 011 POrluguese FUflgllS Gflats 539

Type-Iocalities: Gerrard's Cross, Bucks, and many other localities in England (Great Britain). Material examined: l-V to 20-V- 1997, I <3 ; 2-VI to 27-VI- 1997, 3 <3<3 + I <;! ;'2-XI to 27-XI-1997, 3 <3<3 + I <;! ; 27-XI to 18-XII-1997, 98 <3<3 + 76 <;!<;! ; 18-XII to 6-1- 1998,39 <3<3 + 48 <;!<;! ; 6-1 to 6-11-1998, 4 <30 + 3 <;!<;! ; 24-11 to 15-111- 1998, I <3 ; 13-V to ll-VI- 1998, 1 <3 ; II-X to 3-XII-1998, I <3 ; 18-XII to 6-1-1998, 79 <3<3 + 83 <;!<;! ; 10-111 to 16-IV-1999, I <3 ; 19-V to 26-VI-1999, 2 <3<3 ; 21-XI to 20-XlI- 1999,2 <3<3 ; 20-XII-1999 to 22-1-2000, I <3 ; 22-1 to 23-1I-2ooo, I <3 + 2 <;!<;! ; 23-11 to 25-I1I-2ooo, 3 <3<3 ; 26-XI to 22-XII-2ooo, l o .

* Mycetophila caudata Staeger, 1840 Myce/ophila caudara. Na/urh. Tidsskr. , 3: 243. Type-Iocality: not given (Denmark). Material examined: l-V to 20-V-1997, I <;! ; 2-VI to 27-VI-1997, 4 <;!<;! ; 2-XI to 27-XI-1997, I <3 + 3 <;!<;! ; 18-XlI-1997 to 6-1-1998, I <3 ; II-X to 3-XII-1998, I <;! Distribution: Widespread in Europe and also known from the Nearctic region.

* Mycetophila deflexa Chandler, 2001 Mycetophila deflexa. Br. l.Em. Nat. Hist., 13: 235. Type-Iocalities: Chobham Common, Gracious Pond, Surrey (England). Material examined: l-V to 20-V- 1997, 3 <3<3 + 2 <;!<;! ; 20-V to 2-VI- 1997, 1 <3 + 2 <;!<;! ; 2-VI to 27-VI- 1997, I <;! ; 27-IX to 18-X-1997, I <3 ; 2-XI to 27-XI-1997, 10 <3<3 + 2 <;!<;! ; 27-XI to 18-XII-1997, 2 <3<3 + 2 <;!<;! ; 18-XII to 6-1-1998, 2 <3<3 + I <;! ; 24-11 to 15-I1I-1998, I <3 ; 15-III to 8-IV-1998, 1 <3 + I <;! ; ; 8-IV to 13-V- 1998, I <;! ; 13-V to II-VI-1998, 2 <3<3 ; 3-XII-1998 to 31-1-1999, I <3 : 31-1 to 7- 11-1999, 3 <30 + I <;! ; 17-X to 21 -Xl-1999, 1 <;! ; 21-XI to 20-XII-1999, 3 <3<3 + 2 <;!<;! ; 20-XII- 1999 to 22-1-2000, 2 <3<3 + 2 <;'<;! ; 23-11 to 25-111-2000, I <3 ; 25-I1I to 24-IV-2ooo, 5 <3<3 + 7 <;!<;! ; 9-VII to 15-VIII-2ooo, I <3 ; 21 -1 to 19-11-2001 , 1 <3; 18-VI to 15-VII-2ool, 1 <3 + I <;! . Distribution: known in some countries in Europe.

* Mycetophilaformosa Lundstrom, 1911 Mycetophilaformosa. Annls hist.-nat. Mu s. Nam. Hllllg., 9: 409. Type-Iocality: Jasenak (Yugosloavia). Material examined: 22-1 to 23-I1-2oo0, 1 <;! . Di str ibution: Widely di stributed in Europe and known from Iran in Asia.

Mycetophila luctuosa Meigen, 1830 Mycetophila luctllosa. Syst. Beschr., 6: 299. Type-Iocality: not given (Europe). Material examined: II-VI to 14-VII-1 998, 3 <30 + 1 <;! . 540 Eugénia Ribeiro

* Mycetophila mitis (Johannsen, 19 12) Myco/hera lIIitis. BI/l/. Me agric. Exp. S/II, 200: 82. Type-Iocality: Winconsin (USA). Material examined: 27-XI to 18-XIl-1 997, 6 rJrJ + 6 'j''j' ; 18-XIl- 1997 to 6-1 1998,7 rJrJ + 3 'j''j' ; 21-XI to 20-XII-1999, 4 rJrJ; 22-1 to 23-11- 2000, I rJ; 23-11 to 25-I1I-2ooo, I rJ ; 28-X to 26-XI-2000, I rJ + I 'j' . Distributi on: Known for some European countries and for lhe Nearctic regian.

* Mycetophilapllmila Winnertz, 1863 Mycelophila pl/lllila. Verl!. Zool.-bOI. Ges. Wiell , 13: 922. Type- Iocality: not given (" ln der Samm lung des Herrn Baron vo n Osten-Sacken in St. Petersburg") (USSR). Material examined: 13-V to ll-VI-1998, I 'j' . Di stribution: Widely di stributed in Europe and known in Iran from Asia.

Mycetophila "'ficollis Meigen, 1818 (group) Material examined: 2-XI to 27-XI- 1997, 2 'j''j' ; 8-IV to 13-V-1998, I 'j' ; l3-V to II-VI-1998,2 'j''j' ; 31-/ to 7-11- 1999, I 'j' ; 2 1-XI to 20-XII- 1999, I 'j' ; 24-IX to 28-X-2000, I 'j' .

Mycetophila sigllatoides Dziedzicki, 1884 M ycetophila signafoides. Pam. Fizyogr., 4: 310. Type-Iocality: not given (probably district of Krefeld, D). Materi al examined: 18-XII-1997 to 6-1-1998, I 'j' ; 13-V to 11-VI-1 998, I rJ.

* Mycetophila stolida Walker, 1856 Mycetophila slOlida. Il1s. Brit., Dipt., 3: 15. Type-Iocality: England (Oreat Britain). Material examined: l-V to 20-V-1997, I rJ; 2-XI to 27-XI- 1997, I rJ + I 'j' ; 27- VI to 4-V11- 1997, I rJ; 18-XII-1997 to 6-1-1998, 4 rJrJ + 7 'j''j' ; 6-1 to 6- 11-1998, 2 rJrJ; 15-I1I to 8-IV-1988, I rJ; 13-V to ll-VI- 1998, I rJ + I 'j' ; 19-V to 26-VI- 1999, I rJ ; 25-111 to 24-1V-2ooo, I rJ ; 25-1II to 8- VII-2ooo, 4 rJrJ. Dist ri but ion: Widespread in Europe; Known from Iran and from lhe Nearctic regian.

* Mycetophila trillotata Staeger, 1840 Mycerophila trillorata. Nawrll. Tidsskr. , 3: 242. Type-Iocality: not give n (Denmark). Materi al examined: l -V to 20-V-1997, I 'j' ; 15-111 to 8- IV- 1998, I rJ . Distributi on: Widely distr ibuted in Europe; known from lran and fTOm the Nearc li c region. Twenty Five Records OH Porluguese Fungus Gnats 541

Mycetophila sp. Material examined: 2-XI to 27-XI-1997, I 'j' ; 27-XI to 28-XIl-1997, 1 'j' ; 18-XIl- 1997 to 6-1-1998,1 'j' ; IS-III to 8-IV-1998, 1 'j' ; 13-V to 1 I-VI-1998, 1 'j' ; II-VI to 14-VIl- 1998, 1 'j' ; 20-XIl- 1999 to 22-1-2000, I 'j' ; 25-1Il to 24-IV-2000, 1 'j' ; 28-X to 26-XI-2000, 2 'j''j'.

Mycomya Rondani Ronda ni , 1856: 194. Type species: Scioplúla lIIargillata Meigen, 1818: 249 (orig. designation).

It is an enormous genus of fu ngus gnats appearing in a great number of habitats. The group was especiall y studied by Vaisanen (l984),'who revised the genus in the Holarctic region. Our specimens belong to five species, two of them new to Portugal: Mycomya cinerascens (Macquart) and Mycomya tumida (W innertz). Four fema les are designated as Mycomya sp. because it was not possible to reach the specific leveI.

* Mycomya cillerascells (Macquart, 1826) Scioplzila cilleraseells. Mélll. Soe. Sei. Agrie. Lille, 1823-1824: 10 1. Type-Iocali ty: not given (? Northern France). Material examined: lO-III to 16-IV- 1999, 1 r3 . Distribution: Largely wi despread in Europe and also known from Japan and Mongoli a.

Mycolllya lIIargillata (Meigen, 1818) Sciophila marginara. Syst. Beschr., 1: 249. Type-Iocali ty: "Osterreich" (Austri a). Material examined: 5-VIII t04-IX-1997, 1 'j' .

Mycomya moura (Walker, 1856) Sciophila maura. / IIS. Brit., Dipt., 3: 42. Type-Iocal ity: England. Material examined: 27-XJ to 18-XII-1997, 1 r3 ; 6-1 to 6-Il-1998, 1 r3 + 1 'j' ; 24-I1 to 15-I1I- 1998, I r3 ; 15-1lI to 8-IV- 1998, 2 r3r3 ; 16-IV to 19-V-1999, 1 r3 ; 19-V to 26-VI- 1999, 1 r3 + 2 'j''j' ; 20-XIl-1999 to 22-1-2000, 1 r3 + 1 'j' ; 22-1 to 23-I1- 2000,1 'j' .

Mycomya promillells (Lundstrom, 1913) Sciopltilaprolll ill ens. Annls hist.-nat. Mus. Il(lfll. 11I/I1g., 11: 306. Type- Iocality: "Hungari a: Plitvica" (Y ugoslav ia). Material examined: 2-XI to 27-XI-1 997, 1 r3 ; 20-XII -1999 to 22-1-2000, 1 r3 ; 22-1 to 23-I1-2oo0, 1 r3 . 542 Eugénia Ribeiro

* Mycomya tumida (Winnertz, 1863) Sciophila tumida. Verh. zool. -bot. Ces. Wien, 13: 727. Type-Iocality: not given (? Germany). Material examined: 6-1 to 6-11-1998,1 éS + I 'j' . Distribution: Known for some European countries.

Mycomya sp. Material examined: 2-VI to 27-VI-1997, 1 'j' ; 31-1 to 7-11-1999, I 'j' ; 22-1 to 23-11- 2000,1 'j' ; 23-11 to 25-I1I-2000, I 'j'.

ACKNOWLEDGEMENTS

The author would Iike to thank Fernando Sousa for his continuous help, especially in field work and to André Sousa, Gonçalo Duarte, José Miguel Simões and Ana Cristina Farinha by their help separating the material from other groups. I also would like to express my gratitude to my colleague Margarida Matos who review lhe engl ish in this paper and to Dr. Peter Chandler for his continuous advice and help. This work was partially financed by C.B.A. (Centro de Biologia Ambiental).

ABSTRACT

ln Herdade da Ribeira Abaixo (Grândola, south west of Portugal) specimens of fungus gnats were collected usi ng Malaise traps. Data for forty five species are given. Twenty five af these species are new records to lhe entomofauna af Portugal. For twenty species, a new locality is presented. The new mycetophilid species for Portugal are: Acnemia amoena Winnertz, 1863; ACllemia nitidicollis (Meigen, 1818); Allodia barbata (Lundstrom, 1909); 8revicomu fuscipenne (Staeger, 1840); 8revicoTl1u griseicolle (Staeger. 1840); Brevicomu Iligrofuscum (Lundstrom, 1909); Cordyla erassieomis Meigen, 1818; Cordyla faseiata Meigen, 1830; Cordyla pusilla Edwards, 1925; Eetrepesthollellra hirta (Wi nnertz, 1846); Exechia confinü Winnertz, 1863; Exechia dentara Lundstrom, 191 6; Exechia nigroscutellata Landrock, 1912; Exechia repanda Johannsen, 1912; Exechia repandoides Caspers, 1984; Monodona rufilarera (Walker, 1837); Myeetophila caudara Staeger, 1840; Myeetophila defiexa Chandler, 200 1; Myeetophila formosa Lundstrom, 1911 ; Myeetophila mitis (Johannsen, 1912); Myeetophila pumila Winnertz, 1863; Myeetophila stolida Walker, 1856; Myeetophila trinotata Staeger, 1840; Mycomya cinerascens (Macquart, 1826) and Mycomya tumida (Winnertz, 1863). Twenty Five Records on Portuguese Fungus Gnats 543

RESUMO

Na Herdade da Ribeira Abaixo (Grândola, sudoeste de Portugal) foram capturados espécimens de micetofilídeos utilizando armadilhas Malaise. Neste trabalho são apresentados os dados para quarenta e cinco espécies. Vinte e cinco destas espécies são novi dades faunÍsticas para a entomofauna de Portugal e para vinte outras é dada uma nova .área de di stribui ção. As espécies novas para Portugal são: Acnemia amoena Winnertz, 1863; Acnemia nitidicollis (Meigen, 1818); AlIodia barbata (Lundstrom, 1909); Brevicomu fuscipel1ne (Staeger, 1840); Brevicornu griseicolle (Staeger, 1840); Brevicornu nigrofuscum (Lundstrom, 1909); Cordyla crassicomis Meigen, 1818; Cordyla fasciata Meigen, 1830; Cordyla pusilla Edwards, 1925; Ectrepesthol1eura hirta (Winnertz, 1846); Exechia confinis Winnertz, 1863; Exechia dentata Lundstrom, 1916; Exechia nigroscutellata Landrock, 1912; Exechia repanda Johannsen, 1912; Exechia repandoides Caspers, 1984; Monoc/ona rufilatera (Walker, 1837); Mycetophila caudata Staeger, 1840; Mycetophila deflexa Chandler, 2001; Mycetophila formosa Lundstrom, 1911 ; Mycetophila mitis (Johannsen, 1912); Mycetophila pumila Winnertz, 1863; Mycetophila stolida Walker, 1856; Mycetophila trinotata Staeger, 1840; Mycomya cinerascens (Macquart, 1826) e Mycomya tumida (Winnertz, 1863).

REFERENCES

CARLES-ToLRÁ, M. 2001 - Algunos dipteros nuevos para Portugal (Insecta, Diptera). Zoo/. Baetica, 12: 91-100. CARLES-TOLRÁ, M. 2002 - Catalogo de los Dipteros de Espana, Portugal y Andorra (Insecta). Monografias Sociedad Elltolllologica Aragonesa, 8: 1-313. CASPERS , N. 1984 - Mycetophiliden aus Lunz, Niederosterreich (Diptera, Nematocera, Mycetophilidae). Zeitschriftfur Elllomologie 5: 173-205. CHANDLER, P. J. 1980 - The European and eastern Nearctic fungus-gnats in the genus Ectrepesthoneura (Mycetophi li dae). Systematic Entomology, 5: 27-41. CHANDLER, P. J. 1993 - The Holarctic species of the Mycetophila fUllgorulll (De Geer) Group (Diptera, Mycetophilidae). British Joum. Ent. & Nat. Hist., 6: 5- II. CHANDLER, P. J. 2001 - Fungus gnats (Diptera: Sciaroidea) new to Britain. Br. J. En!. Na!. His!., 13: 215-243. DZIEDZICKI, H. 1884 - Przyczynek do fauny owadów dwuskryzydlych. l. Teil. (Mycothera, Mycetophila, Staegeria). Pamietnik Fizyjograficzny 4: 298-324. DZIEDZICKI, H. 1885 - Przyczynek do fauny owadów dwuskryzydlych. Rodzaje nowe: Hertwigia novo gen., Ewycera no vo gen. i. gatunki rodzajów; Boletina, Sciophila. Pal1lietnik FizyjograficZI1Y 5: 164-194. 544 Eugénia Ribeiro

EDWA RDS, F. W. 1925 - British fu ngus-gnats (Diptera, Myeetophilidae) with a revised generic c1assification af the fam ily. Transactiolls of lhe Royal EllIomological Society of London 1924: 505-670. ENDERLEI N, G. 1911 - Neue Gattungen und Arten au Bereuropaiseher Fliegen. Sleuill elll. Zlg., 72: 135-209. HUTSON, A. M.; ACKLAND, D. M. and KIDD, L. N. 1980 - M yeelOphi lidae (Bolitophilinae, Ditomyiinae, Diadoeidiinae, Keropltinae, Seiophilinae and Manotinae) Diptera, Nematocera. Handbooks for Idelllification of British Insecls, 9 (3): 1-109. JOHANNSEN, O. A. 1909 - Diptera. Fam. Myeetoph ilidae. ln WYTSMAN, P. (ed.): Genera Insectorwn, Fase. 93: 1-14l. JOHANNSEN, O. A. 1912 - Fungus-gnats of Nonh Ameriea. IV. Bul/. Me agric. Exp. Sln, (2) No. 200: 57-146. LAFFOON, J. L. 1957 - A revision of the Nearetie speeies of Fungivora (Diptera, Myeetophilidae). Iowa SI. Col/. J. Sci., 31(1956): 141-340. LAFFOON, J. L. 1965 - Family Myeetophilidae (Fungivoridae). ln STONE, A., et ai. (eds): A cararog of lhe Diplera of America 1I0rlh of Mexico. Washington: 169-229. . '. LANDROCK, K. 1912 - Zur Monographie der Gattung Bolitophila Meig .. Ber/. Enl. Z., 57: 33-51. L ASTOVKA, P. 1972 - Holaretie speeies of Mycelophila ru[icollis-group (Diptera, Myeetophilidae). ACla elll. bohemoslov., 69: 275-294. LASTOVKA, P. and KIDD, L. N. 1975 - Review of the British and notes on other speeies of the Mycelophila ru[ieollis-group, with the deseription of a new speeies (Dipt., Myeetophi li dae). Enlomologisl 's mal/. Mag., 110: 203-214. LUNDSTROM , C. 1909 - Beitrage zur Kenntnis der Dipteren Finnlands. IV. Supplement. Myeetophili dae. Acla Soco Fauna Florafelln. , 32 (2): 1-63. LUNDSTROM, C. 19II - Neue oder weni g bekannte europaisehe Myeetophiliden. Al1llls hist.-nat. Mus. Na11l. Hung., 9: 390-4 19. LUNDSTROM , C.1912 - Beitrage zur Kenntnis der Dipteren Finnlands. VIII, Supplement 2. Myeetophilidae etc. Acla Soco Faulla Florafellll., 36 (1): 1-39. L UN DSTROM, C. 191 3 - Neue oder wenig bekannte europaische Mycetophili den. III. Alll1ls /tis/. -nat. Mus. nClll1. lumg., 11 : 305-322. LUNDSTROM, C. 19 16 - Neue oder wenig bekannte europaische MycelOphiliden. IV. Annls !tisl.-nat. Mus. 1/alll. hung. , 14: 72-80. MACQUART, 1. 1826 - Inseetes Dipteres du nord de la France. Tipulaires. Mém. Soco Sci. Agric. Lille, 1823-1824: 59-224. MARSHALL, P. 1896 - New Zealand Diptera, No. 2. Mycetophilidae. Tran s. ProC. N. Z. Insl., 28: 250-309. ME1 GEN, J. W. 1803 - Versuch einer neuen GattungsEintheilung der europai schen zweiflugeligen Insekten. Mag. Illseklenk., 2: 259-28 1. Twenty Five Records Oll Portuguese Fungus Gllats 545

MEIGEN, J. W. 1804 - Klassifikazioll ll1ul Beschreibung der europaischen zweiflugeligell lllsekten (Diptem Lilln.), Braunschweig (=Brunswick): 1(1): 1- 152. MEIGEN, 1. W. 1818 - Systematische BeschreibulIg der bekannten europaischell zweiflugeligell lnsektell. Aachen, I:XXXVI+I-333. MEIGEN, 1. W. 1830 - Systematische BeschreibuJ/.g der bekannten eu ropaischen zweiflugeligelllllsektell. Hamm, 6: XI+ 1-401. MIK, J. 1886 - Dipterologische Miscellen. II., Wiell. em. Ztg., 5: 276-279 . . RIBEIRO, E. 1990 a - Contribution to the study of fungus-gnats (Diptera: Mycetophiloidea) of Portugal. II - Seven new records. Bolm. Soe. porto Em. 118 (IV-16): 173-196. RIBEIRO, E. 1990 b - Contribution to the study of fungus-gnats (Diptera: Mycetophiloidea) of Portugal. 1II - Four new records. Bolm. Soe. porto En/. 127 (IV-25): 301-308. RIBEIRO, E. 1991 a - Contribution to the study of fungus-gnats (Diptera: Mycetophiloidea) of Portugal. IV - Three new records. Bolm. Soe. porto EnI. 134 (V-2): 21-28. RIBEIRO, E. 1991 b - Contribution to the study of fungus-gnats (Diptera: Mycetophiloidea) of Portugal. V - Eighteen new records. Bolm. Soe. porto Em. 135 (V-3): 29-40 . . RIBEIRO, E. 1991 c - Contribution to the study of fungus-gnats (Diptera: Myeetophiloidea) of Portugal. VI - Six new records. Bolm. Soe. porto Ellt. 136 (V-4): 41-52. RIBEIRO, E. 1992 - Contribution to lhe study of fungus-gnats (Diptera: Myeetophiloidea) of Portugal. I - Twenty-three new records. Bolm. Soe. porto ciências naturais (2a Série), 25: 2 1-30. RONDANI, C. 1856 - Dipterologiae italicae prodomus. Genera ítali ca ordini s dipterorum ordinatim dísposita et dístincta et in famílias et stirpes aggregata. Parmae [=Parma]: 1: 1-228. SANTOS ABREU, E. 1920 - Monografia de los Fungivoridos de las Islas Canárias. Memorias de la Real Academia de Ciências y Artes de Barcelona, 16: 1-152. SÓOS, A. and PA PP, L. (ed.) 1988 - Catalogue of Palaearetie Diptem, 3: 1-448. STAEGER, R. C. 1840 - Systematisk fOrlegnelse over de i Danmark hidtil funde Diptera. Na/url!. Tidsskr., 3: 1-58,228-288. TUOM IKOSK I, R. 1966 Generic laxonomy of the Exeehilli (Diplera, Mycetophilidae). AIIII/s. Ellt. Felln. , 32: 159-194. V AISANEN, R. 1984 - A monograph of the genus Myeomya Rondani in the Holarc!ic region (Diptera, Mycetophilidae). Ae/a Zoo/. Fel/nica 177: 1-346. V ILLERS. C . 1789 - Cm'oli Linllaei ewomologia, fauna sueciae descriptioniblls alicIa. Lugd uni [=Lyon], 3: 1-657. VOCKEROTH, J. R. 1980 - New genera and species of Mycetophilidae (Diptera) from the Holarctic reg ion, with notes on other spec ies. Ca flad. Ewomol., 112: 529-544. 546 Eugénia Ribeiro

W ALKER, F. 1837 - Notes on Diptera. Em. Mag., 4: 226-230. WALKER , F. 1856 - Illsecta Britallllica. Diptera. London, 3: XXIV+I-352. WESTWOOD, J. O. 1840 - Synopsis of the gellera of British illsects: 1-158 (separately paginated). ln : An introdueti on to the modero classifieation of insects; founded on lhe natural habits and corresponding organization af the different families. London, 2: XI+I-587. WIEDEMANN, C. R. W. 1817 - Neuer Zweiflugler (Diptera Linn.) aus der Gegend um Kiel. Zoo/. Mag., Kiel, 1: 61-86. WINNERTZ, J. 1846 - Besehreibung einiger neuen Gattungen aus der Ordnung der Zweiflugler. Steuill. ent. Ztg., 7(1): 11-20. WI NNERTZ, 1. 1863 Beitrag zu einer Monographie der Pilzmueken (Myeetophilidae). Verll. zoo/. -bot. Ces. Wien , 13: 637-964. ZA ITEV, A. I. 1982 a -(Dipterans of the genus ACllemia Winn. (Myeetophilidae) of the Holaretie fauna. I). Zoologicheskyi Zhllmal6l: 707-715 (in Russian). ZA ITEV, A. I. 1982 b -(Dipterans of the genus ACIlemia Winn. (Myeetophilidae) of the Holaretie fauna. 2). Zoologicheskyi Zhllrnal61: 867-874 (in Russian). Twenty Five Records 012 Portuguese Fungus Cnats 547

8 7 6 5 Me p G 4 3 2 1 o 9 8 7 6 5 4 MF F P 3 2 1 o 9 8 7 6 5 ME 4 3 1 o 9 8 7 6 5 MO o ao 4 3 2 1 o 9 8 7 6 5 MC c .aç 4 NC 3 2 1 o 9 8 7 6 5 B B as 4 3 2 1 o 9 ANOJA 84 567 8 9 o 1 23 4 56789 01234 567 89 o 1 234 5

Figure 1 - Distribution of new records: Grândola (Herdade da Ribeira Abaixo) (29SNC 32). 548 Eugénia Ribeiro

Table 1 . Mycetophilid species, number of males and fe males and their respective total nu mber

Species Number of males Number of females Total ACI/emia amoeno 15 I3 28 Ao/emia Ilitidicollis 4 5 9 Allodia barbata O I Allodia omaticollis 5 6 Allodia sp. O 5 5 Bolellno g ripha 40 19 59 Bolitophifa pseudohybrida 6 16 22 B revicomu fllscipelllle 3 O 3 Brevicorllu griseicolle 13 13 26 Brevicomu nigrofuscum I I 2 Brevicomll sericoma 27 9 36 8revicornu sp. O 52 52 Cordyla brevicornis 16 4 20 CordyLa crassicomis 20 O 20 Cordyla faseia/a 9 4 13 Cordyla fusca 73 25 98 Cordyla pusilla 24 O 24 Cordyla sp. O 139 139 EctrepeSfhoneura hirta 16 16 32 Exechia bicillCla 3 3 6 Exechia conflllis II O II Exechia dentola 8 2 10 Exechia dorso/is 7 5 12 Exechia fi/Ivo 21 4 25 Exechia fusca 43 35 78 Exechiafusca grollp O 37 37 Exechia nigrosclltellata 3 O 3 Exechia repanda O Exech ia repandoides O Exechia reponda grollp O I I Exechia seriata 3 4 7 Exechia spilluligera 2 O 2 Twenty Five Records 011 Porluguese FUÍlgus Gnals 549

Table 1 ~ Mycetophilid species, number of males and females and their respective total number (continuation)

Se!!:ies Number of males Number of females Tolal MonocJona rufilatera 4 3 7 Mycetophila a/ea 24 16 40 Mycetophila britallllica 241 214 455 Mycetophila caudata 2 9 II Mycetophila deflexa 41 26 67 Mycetophila formosa O 3 4 Mycetophila mitis 20 10 30 Mycetophila pumila O I I MycetophiJa ruficollis group O 8 8 MycetophiJa sigllatoides I 2 MycetophiJa stotida 17 9 26 Mycetophila trinotata I 2 Mycetophila sp. O 10 10 Mycomya cillerascellS O Mycomya margillata O I I Mycomya maura 8 5 13 Mycomya promillells 3 O 3 Mycomya tumida 2 Mycomya sp. O 4 4 TOTAL 743 733 1476