Dendrology Series

WSFNR15-14 March 2015 NativeNative PinePine RangesRanges inin GeorgiaGeorgia by Dr. Kim D. Coder, Professor of Tree Biology & Health Care, Warnell School of Forestry & Natural Resources, University of Georgia

Georgia pines are more than filters for romantic moonlight. These pines have been and continue to be the most important trees for our quality of life. Georgia pines are the cardboard boxes, paper products, wood of our houses, and renewable resources which drove the industrial revolution in the old South and which drives the future of the new South. Pines are considered by many to be the lowest of the trees because they are so common. Yet they are both the lead actors in an ecological play in rural lands, as well as all the diverse commu- nities of Georgia. Pines are trees which smell, sound, and look like home to many people. Pines are also the most impor- tant industrial tree group in the Northern Hemisphere. Georgia has eleven native and naturalized pines. Each is akin to the others, but unique in where and how it grows. This publication helps define where our pines grow in Georgia.

The PINES! Pines are in one of six families within the conifers (Pinales). The conifers are divided into roughly 50 genera and more than 500 species. The conifer famillies include pine (Pinaceae) and cypress (Cupressaceae) of the Northern Hemisphere, and podocarp (Podocarpaceae) and araucaria (Araucariaceae) of the Southern Hemisphere. Plum-yew (Cephalotaxaceae) and umbrella-pine (Sciadopityaceae) families are less common. The pine family (Pinaceae) has many genera (~9) and many species (~211). The most common of the genera are fir (Abies), cedars (Cedrus), larch (Larix), spruce (Picea), pine (Pinus), Douglas-fir (Pseudotsuga), and hemlock (Tsuga).

Genus Trees generically considered pines were first placed in the Pinus genus in 1753. The name of the genus is the Latin word for pine. The pine genus (Pinus) contains ~95 species (range from 66-120 species) from around the Northern Hemisphere. One pine species does range across the equator into the Southern Hemi- sphere in Sumatra. Worldwide, pines are divided into 15 sub-sections of the genera. Of the worldwide total of ~95 pines, ~60 are in the Western Hemisphere. Mexico has ~35 different species, the United States has ~36 species, and Canada has 9 species. Pines can be found as far South in

In compliance with federal law, including the provisions of Title IX of the Education Amendments of 1972, Title VI of the Civil Rights Act of 1964, Sections 503 and 504 of the Rehabilitation Act of 1973, and the Americans with Disabilities Act of 1990, the University of Georgia does not discriminate on the basis of race, sex, religion, color, national or ethnic origin, age, disability, or military service in its administration of educational policies, programs, or activities; its admissions policies; scholarship and loan programs; athletic or other University-administered programs; or employment. In addition, the University does not discriminate on the basis of sexual orientation consistent with the University non-discrimination policy. Inquiries or complaints should be directed to the director of the Equal Opportunity Office, Peabody Hall, 290 South Jackson Street, University of Georgia, Athens, GA 30602. Telephone 706-542-7912 (V/TDD). Fax 706-542-2822. AN EQUAL OPPORTUNITY / AFFIRMATIVE ACTION INSTITUTION. Central America as Nicaragua. Pines can also be found across the Carribean islands (4 species). The Eastern United States has 13 native pines, of which 11 are native or naturalized in Georgia. Georgia’s eleven native and naturalized pines include:

Pinus clausa -- sand pine (naturalized) Pinus echinata -- shortleaf pine Pinus elliottii -- slash pine Pinus glabra -- spruce pine Pinus palustris -- longleaf pine Pinus pungens -- table mountain pine Pinus rigida -- pitch pine Pinus serotina -- pond pine Pinus strobus -- Eastern white pine Pinus taeda -- loblolly pine Pinus virginiana -- Virginia pine

Figure 1 shows the genetic relationships among these pines in Georgia.

Local Genetics The ecological distribution of pines can be immense or minute. Within each pine’s native growth range are genetic differences which allow for more efficient survival and growth under a variety of conditions. Many pines have regional races which do not perform well outside their native area. A pine species may appear to be the same, but regional races within one species may have widely variable attributes based upon climate, soil, and pests differences. Pines also form hybrids, where one species will interbreed with another species. The hybrid off-spring may pose identification problems and regeneration opportunities with their mix of growth attributes.

Native Range Knowing the native ranges of Georgia pines can assist in identification. Knowing a species’ native range is also helpful in judging potential tree stress from being out-of-range. Maps in this publication (i.e. map number #1 through #11) provide a native range for each Georgia pine. These native or naturalized range maps were developed over time through referencing federal species maps, state species maps, herbarium specimens, and personal observations by the author. Outlying and disjunct small populations are not shown. Without other supporting information for within-county distributions, mapping lines follow county political boundaries. For example, if a species is found in the Southern end of a county, the whole county could be included in the native species range.

Species Distributions Figure 2 demonstrates the North / South distribution of our native and naturalized pine species. Gener- ally, Georgia has mountain pines, statewide pines and Coastal plain pines. Figure 3 provides an estimated number of native or naturalized pines for each Georgia county by species number class. Note the relatively low numbers of native pine species in the Peidmont of Georgia.

Summary Georgia’s native pines have been, and continue to be, a great asset for the State. Our pines are “green gold” for Georgia’s future. Urban, rural, or interface sites can all make use of pines, whether for recreation, plywood, pine straw, lumber, resin products, habitat, watershed protection, carbon-dioxide capture, or aesthet- ics. Pines represent both Georgia’s glorious past and its wonderful future.

Dr. Kim D. Coder, Warnell School, University of Georgia 2 Pinus Sub-Genera Groups

1) white pine group = Pinus strobus – Eastern white pine

2) Southern yellow pine group = Pinus echinata -- shortleaf pine Pinus elliottii – slash pine Pinus glabra – spruce pine Pinus palustris -- longleaf pine Pinus pungens -- table mountain pine Pinus rigida -- pitch pine Pinus serotina -- pond pine Pinus taeda -- loblolly pine

3) jack pine group = Pinus clausa -- sand pine Pinus virginiana – Virginia pine

Figure 1: The sub-genera groups of Georgia pines.

Dr. Kim D. Coder, Warnell School, University of Georgia 3 Naturalized Growth Range For Pinus clausa: sand pine Naturalized range derived from federal and state maps, herbarium samples and personal observations.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM

DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW MAP Y S R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE #1 NEWTON COLUMBIA

CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

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Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 4 Native Growth Range For Pinus echinata: shortleaf pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. Native range includes all areas North and West of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM

DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW Y S MAP R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE NEWTON COLUMBIA#2 CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

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Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 5 Native Growth Range For Pinus elliottii: slash pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. Native range includes all areas South of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM

DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW Y S MAP R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE NEWTON COLUMBIA#3 CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

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Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 6 Native Growth Range For Pinus glabra: spruce pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. Native range includes all areas South of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM

DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW Y MAP S R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE NEWTON COLUMBIA#4 CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

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Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 7 Native Growth Range For Pinus palustris: longleaf pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. Native range includes all areas South and West of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM

DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW MAP Y S R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE #5 NEWTON COLUMBIA

CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

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Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 8 Native Growth Range For Pinus pungens: table mountain pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. The native range includes all areas North and East of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM

DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW Y S MAP R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE NEWTON COLUMBIA#6 CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

SEMINOLE BROOKS

ECHOLS

Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 9 Native Growth Range For Pinus rigida: pitch pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. Native range includes all areas North and East of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM

DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW Y S MAP R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE NEWTON COLUMBIA#7 CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

SEMINOLE BROOKS

ECHOLS

Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 10 Native Growth Range For Pinus serotina: pond pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. Native range includes all areas South and East of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM

DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW MAP Y S R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE #8 NEWTON COLUMBIA

CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

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Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 11 Native Growth Range For Pinus strobus: Eastern white pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. Native range includes all areas North of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM

DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW Y S MAP R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE NEWTON COLUMBIA#9 CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

SEMINOLE BROOKS

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Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 12 Native Growth Range For Pinus taeda: loblolly pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. Native range includes all areas South and West of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM MAP DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW Y S R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN#10 DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE NEWTON COLUMBIA

CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

SEMINOLE BROOKS

ECHOLS

Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 13 Native Growth Range For Pinus virginiana: Virginia pine Native contiguous range derived from federal and state maps, herbarium samples and personal observations. Native range includes all areas North and West of the lines.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION WALKER WHITFIELD GILMER WHITE CHATTOOGA LUMPKIN GORDON STEPHENS PICKENS HABERSHAM MAP DAWSON FRANKLIN HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW Y S R O F JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN#11 DOUGLAS FULTON

CARROLL ROCKDALE MORGAN GREENE NEWTON COLUMBIA

CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK

R E GLASCOCK H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN

SEMINOLE BROOKS

ECHOLS

Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 14 Figure 2: Native range centroids for Georgia’s pines. Numbers represent common names of pines and are placed at the idealized center of their native or naturalized range within Georgia.

DADE CATOOSA TOWNS FANNIN TOWNS pitch pine = 1 RABUN MURRAY UNION table mountain pine = 2 WALKER WHITFIELD GILMER 1 Eastern white pine = 3 WHITE Virginia pine = 4 CHATTOOGA 3LUMPKIN GORDON STEPHENS HABERSHAM shortleaf pine = 5 PICKENS 2 HABERSHAM

DAWSON FRANKLIN loblolly pine = 6 HALL BANKS HART FLOYD CHEROKEE H GEORGIAlongleaf pine = 7 T GEORGIA BARTOW Y 4 S R O pond pine = 8 FFORSYTH JACKSON MADISON ELBERT POLK spruce pine = 9 BARROW COBB GWINNETT CLARKE OGLETHORPE slash pine = 10 more NorthernPAULDING OCONEE HARALSON sand pine = 11 WALTON DEKALB WILKES LINCOLN DOUGLAS LE FULTON A D K C CARROLL O ROCKDALER MORGAN GREENE O NEWTON RR COLUMBIA FE LIA CLAYTON HENRY TALIAFERROTA

McDUFFIE COWETA WARREN McDUFFIE HEARD FAYETTEFAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK K OC SC LA R GLASCOCKG E 5 H T PIKE E BALDWIN JEFFERSON BURKE moreTROUP StatewideLAMAR IW LAMAR MONROE R JONES E WASHINGTON MMERIWETHER WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN 6 TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH

LAURENS HOUSTON CHATTA- MARION BLECKLEY TREUTLEN BULLOCH HOOCHEE MARION MACON CANDLER EFFINGHAM

SCHLEY more Southern PULASKI DODGE DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM MONTGOMERY WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH 7 BEN HILL DAVIS APPLING BEN HILL8 DAVIS TURNER LONG

CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH 9 COFFEE BACON McINTOSH TIFT EARLY BAKER 10 PIERCE BERRIEN ATKINSON MITCHELL GLYNN MILLER COLQUITT BRANTLEY COOK WARE

LANIER DECATUR GRADY CLINCH CAMDEN GRADY THOMAS LOWNDES CHARLTON CAMDEN

SEMINOLE BROOKS 11 ECHOLS BackgroundBackground mapmap fromfrom CarlCarl VinsonVinson InstituteInstitute ofof Government,Government, UniversityUniversity ofof Georgia.Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 15 Figure 3: Number of Georgia pine species in each county. Native range counts derived from federal and state maps, herbarium samples and personal observations.

DADE CATOOSA FANNIN TOWNS RABUN MURRAY UNION B KEY: WALKER WHITFIELD GILMER A = 2 to 3 species. B WHITE CHATTOOGA LUMPKIN GORDON STEPHENS B = 4 to 5 species. C HABERSHAM B A PICKENS DAWSON FRANKLIN C = 6 to 7 species. HALL BANKS HART H FLOYD CHEROKEE T GEORGIA BARTOW Y S R O F B JACKSON MADISON ELBERT POLK BARROW COBB GWINNETT CLARKE OGLETHORPE PAULDING OCONEE HARALSON WALTON DEKALB WILKES LINCOLN DOUGLAS FULTON A CARROLL ROCKDALE MORGAN GREENE NEWTON COLUMBIA

CLAYTON HENRY TALIAFERRO

COWETA WARREN McDUFFIE FAYETTE HEARD JASPER PUTNAM RICHMOND SPALDING BUTTS HANCOCK B R GLASCOCK AE H T PIKE JEFFERSON TROUP E BALDWIN BURKE IW LAMAR MONROE R JONES E M WASHINGTON UPSON

BIBB WILKINSON JENKINS HARRIS TALBOT CRAWFORD JOHNSON SCREVEN TWIGGS EMANUEL MUSCOGEE TAYLOR PEACH B LAURENS HOUSTON CHATTA- MARION BLECKLEY TREUTLEN BULLOCH HOOCHEE MACON CANDLER CEFFINGHAM BSCHLEY PULASKI DOOLY DODGE EVANS STEWART WHEELER TOOMBS SUMTER BRYAN CHATHAM WEBSTER WILCOX MONTGOMERY TATTNALL CRISP TELFAIR LIBERTY QUITMAN TERRELL LEE JEFF RANDOLPH BEN HILL DAVIS APPLING TURNER LONGB CLAY IRWIN CALHOUN DOUGHERTY WAYNE WORTH COFFEE BACON McINTOSH C TIFT EARLY BAKER PIERCE BERRIEN ATKINSON MITCHELL GLYNN COLQUITT B MILLER BRANTLEY COOK WARE

LANIER DECATUR CLINCH GRADY THOMAS LOWNDES CHARLTON CAMDEN BROOKS SEMINOLE C

ECHOLS

Background map from Carl Vinson Institute of Government, University of Georgia.

Dr. Kim D. Coder, Warnell School, University of Georgia 16