DOCSLIB.ORG

Rove Beetles of the World, Staphylinidae (Insecta: Coleoptera: Staphylinidae)1 J

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Rove Beetles of the World, Staphylinidae (Insecta: Coleoptera: Staphylinidae)1 J EENY114 Rove Beetles of the World, Staphylinidae (Insecta: Coleoptera: Staphylinidae)1 J. Howard Frank and Michael C. Thomas2 Introduction Classification The family Staphylinidae belongs to the suborder Polyphaga Phylogeny of the order Coleoptera (beetles). The superfamily Staphy- Four phyletic lines are now included in Staphylinidae linoidea includes the small families Hydraenidae, Ptiliidae, (Lawrence and Newton 1995): Agyrtidae, Leiodidae, and Silphidae, and the huge family Staphylinidae. • subfamilies Glypholomatinae, Microsilphinae, Omaliinae, Empelinae, Proteininae, Micropeplinae, Neophoninae, SUBORDER Polyphaga—SUPERFAMILY Staphylinoidea— Dasycerinae, Protopselaphinae, Pselaphinae, and FAMILY Staphylinidae Scydmaeninae; At present, more species of beetles are known than of any • subfamilies Phloeocharinae, Olisthaerinae, Tachyporinae, other insect order. As now constituted, Staphylinidae are Trichophyinae, Habrocerinae, and Aleocharinae; the largest family of beetles, with over 63,000 species known • subfamilies Trigonurinae, Apateticinae, Scaphidiinae, worldwide and probably over 75% of tropical species still Piestinae, Osoriinae, and Oxytelinae; undescribed. It is the largest family in the British Isles and • subfamilies Oxyporinae, Megalopsidiinae, Steninae, in America north of Mexico, and it may prove to be so in Euaesthetinae, Solieriinae, Leptotyphlinae, Pseudopsinae, other regions when huge numbers of now-unknown species Paederinae, and Staphylininae. are described (if they are described before they become extinct by habitat destruction). In the future, systematists The former taxa Brathinidae (now just part of subfamily may, however, choose to split the family into four phyletic Omaliinae), Dasyceridae (now subfamily Dasycerinae), lines to form four families. This document takes a world- Empelidae (now subfamily Empelinae), Glypholomatini wide perspective, emphasizing the role of the extremely few (formerly a tribe of Silphidae), Microsilphinae (formerly species that have been studied; identification is far beyond a subfamily of Silphidae), Pselaphidae (now subfamily the scope of this document because of the size of the family. Pselaphinae), Scydmaenidae (now subfamily Scydmaeni- nae), and Scaphidiidae (now subfamily Scaphidiinae) of earlier authors are here in Staphylinidae. 1. This document is EENY114, one of a series of the Entomology and Nematology Department, UF/IFAS Extension. Original publication date September 1999. Revised April 2016 and October 2019. Visit the EDIS website at https://edis.ifas.ufl.edu for the currently supported version of this publication. This document is also available on the Featured Creatures website at http://entnemdept.ifas.ufl.edu/creatures/. 2. J. Howard Frank, retired professor, Entomology and Nematology Department; and Michael C. Thomas, retired, Florida Department of Agriculture and Consumer Services, Division of Plant Industry; UF/IFAS Extension, Gainesville, FL 32611. The Institute of Food and Agricultural Sciences (IFAS) is an Equal Opportunity Institution authorized to provide research, educational information and other services only to individuals and institutions that function with non-discrimination with respect to race, creed, color, religion, age, disability, sex, sexual orientation, marital status, national origin, political opinions or affiliations. For more information on obtaining other UF/IFAS Extension publications, contact your county’s UF/IFAS Extension office. U.S. Department of Agriculture, UF/IFAS Extension Service, University of Florida, IFAS, Florida A & M University Cooperative Extension Program, and Boards of County Commissioners Cooperating. Nick T. Place, dean for UF/IFAS Extension. Fossils Pleistocene fossil Staphylinidae have been reported from peat bogs in northeastern North America and from Europe. They seem to be of extant species and help to show former distribution of some of these species. Oligocene fossils are from amber from the Baltic, the Dominican Republic, and elsewhere, and from shales in the United States (Colorado), France, Germany, and elsewhere. Most of these are recognizably members of modern genera. Deposits from Figure 1. Representative adults of four subfamilies (left to right): the mid-Cretaceous to lower Jurassic in Eurasia have also Proteinus thomasi Frank (Proteininae) 1.5 mm; Coproporus rutilus yielded fossils. Most of these resemble members of modern (Erichson) (Tachyporinae) 3.8 mm; Scaphidium quadriguttatum (Say) subfamilies. The oldest recorded staphylinid, more than 200 (Scaphidiinae) 4.3 mm; Neobisnius ludicrus (Erichson) (Staphylininae) 4.1 mm. million years old, is from the upper Triassic of the United Credits: M. C. Thomas, Division of Plant Industry States (Virginia). Species-level identification of present-day Non-entomologists sometimes confuse Staphylinidae staphylinids normally requires dissection, at least of the with the order Dermaptera. However, the non-opposable genitalia; when this cannot be done with fossil specimens, valvulae (appendages of the ninth abdominal segment) they have limited value. are not the opposable forcipes of Dermaptera, and the radial wing-folding pattern of Dermaptera is unlike that of Appearance Staphylinidae. Adults The length of adult Staphylinidae ranges from less than Immature Stages 1 mm to 40 mm. Most are under 7 mm long. Most have Staphylinid eggs typically are white, spherical, spheroidal, short elytra, exposing several abdominal segments, but or pyriform (pear-shaped). Eggs of some genera in the it would be an error to imagine that all have short elytra, subfamily Staphylininae (e.g., Philonthus) have pronounced or that all beetles with short elytra are Staphylinidae. surface sculpture, allowing identification at least to the Typically, they are slender with short elytra and powerful species-group level. Larvae are campodeiform (sometimes abdominal musculature that renders them very flexible called staphyliniform), meaning they are elongated and and thus able to enter narrow crevices. Those that have flattened). In some subfamilies (Paederinae, Staphylininae, short elytra trade flexibility for exposure, rendering them and to a lesser extent in their immediate relatives) the subject to desiccation and dependent upon humid habitats. head is relatively more heavily sclerotized and there is a Abdominal segments are surrounded by sclerotized plates distinct “neck” (nuchal constriction of the head). Prepupae (a large dorsal plate called a tergite, a large ventral plate of the subfamily Steninae, most species within subfamily called a sternite, and up to two pairs of dorso-lateral plates Aleocharinae, and Astenus (Paederinae) spin a silken called paratergites) with membranous connections. In some cocoon in which they pupate. Pupae are obtect, pigmented, genera (e.g., Paederinae, Euaesthetinae, and Osoriinae, and and sclerotized in the subfamily Staphylininae, but exarate, partially so in Steninae) the tergites-paratergites-sternites white, and unsclerotized in all the other subfamilies. In plates are fused into rings around each segment, probably general, the immature stages develop rapidly, in a few days limiting water loss. The length of adults (Figure 1), which is to a few weeks, and the adults are long-lived. nevertheless normally used to express size, is an inaccurate determinant of size because abdominal segments typically Habitat and Food can be telescoped, making the body appear longer when under moist conditions and alive, but shorter when dried Habitat and dead. A lack of eyes has evolved in some soil-inhabiting Staphylinidae occupy almost all moist environments (Leptotyphlinae) and cave-inhabiting species, and wing- throughout the world. Because none of them is truly lessness has evolved in species occupying environments aquatic, they do not live in open waters; although winged including mountains, the soil, caves, and seashores. adults may be skimmed from the sea surface far from land, their presence is due to misadventure but attests to their dispersive ability. They live in leaf litter of woodland and forest floors and grasslands. They concentrate in fallen decomposing fruits; the space under loose bark of fallen, Rove Beetles of the World, Staphylinidae (Insecta: Coleoptera: Staphylinidae) 2 decaying trees; drifted plant materials on banks of rivers Eulissus (Staphylininae) on adult dung-inhabiting scarab and lakes; and dung, carrion, and nests of vertebrate beetles. Aleochara (Aleocharinae) has evolved to become animals. Several hundred species live only on seashores. parasitoidal in fly puparia. Many are specialized to exist in nests of social insects. Many inhabit caves, underground burrows of vertebrate animals, Behavior and smaller soil cavities, even of burrows that they (a few of them) excavate. Many live in mushrooms. Adults and even Presocial or Subsocial Behavior larvae of a few are associated with living flowers. Others Such behavior is known in Bledius and Platystethus (Oxy- climb on plants, especially at night, and hunt for prey. A few telinae) and Eumicrota (Aleocharinae). Adults construct seem to live with terrestrial snails. Their distribution in arid chambers—Bledius in sandy or muddy shores, Platystethus environments is restricted to moist microhabitats. in the dung of ungulates, and Eumicrota in mushrooms—in which they deposit and guard their eggs and young. Food A little about the feeding habits of Staphylinidae has been Relationships with Social and Communal deduced from casual observations by many observers, Insects from dissections
Load more

Recommended publications
  • A Catalogue of Coleoptera Specimens with Potential Forensic Interest in the Goulandris Natural History Museum Collection
    ENTOMOLOGIA HELLENICA Vol. 25, 2016 A catalogue of Coleoptera specimens with potential forensic interest in the Goulandris Natural History Museum collection Dimaki Maria Goulandris Natural History Museum, 100 Othonos St. 14562 Kifissia, Greece Anagnou-Veroniki Maria Makariou 13, 15343 Aghia Paraskevi (Athens), Greece Tylianakis Jason Zoology Department, University of Canterbury, Private Bag 4800, Christchurch, New Zealand http://dx.doi.org/10.12681/eh.11549 Copyright © 2017 Maria Dimaki, Maria Anagnou- Veroniki, Jason Tylianakis To cite this article: Dimaki, M., Anagnou-Veroniki, M., & Tylianakis, J. (2016). A catalogue of Coleoptera specimens with potential forensic interest in the Goulandris Natural History Museum collection. ENTOMOLOGIA HELLENICA, 25(2), 31-38. doi:http://dx.doi.org/10.12681/eh.11549 http://epublishing.ekt.gr | e-Publisher: EKT | Downloaded at 27/12/2018 06:22:38 | ENTOMOLOGIA HELLENICA 25 (2016): 31-38 Received 15 March 2016 Accepted 12 December 2016 Available online 3 February 2017 A catalogue of Coleoptera specimens with potential forensic interest in the Goulandris Natural History Museum collection MARIA DIMAKI1’*, MARIA ANAGNOU-VERONIKI2 AND JASON TYLIANAKIS3 1Goulandris Natural History Museum, 100 Othonos St. 14562 Kifissia, Greece 2Makariou 13, 15343 Aghia Paraskevi (Athens), Greece 3Zoology Department, University of Canterbury, Private Bag 4800, Christchurch, New Zealand ABSTRACT This paper presents a catalogue of the Coleoptera specimens in the Goulandris Natural History Museum collection that have potential forensic interest. Forensic entomology can help to estimate the time elapsed since death by studying the necrophagous insects collected on a cadaver and its surroundings. In this paper forty eight species (369 specimens) are listed that belong to seven families: Silphidae (3 species), Staphylinidae (6 species), Histeridae (11 species), Anobiidae (4 species), Cleridae (6 species), Dermestidae (14 species), and Nitidulidae (4 species).
    [Show full text]
  • Local and Landscape Effects on Carrion-Associated Rove Beetle (Coleoptera: Staphylinidae) Communities in German Forests
    insects Article Local and Landscape Effects on Carrion-Associated Rove Beetle (Coleoptera: Staphylinidae) Communities in German Forests Sandra Weithmann 1,* , Jonas Kuppler 1 , Gregor Degasperi 2, Sandra Steiger 3 , Manfred Ayasse 1 and Christian von Hoermann 4 1 Institute of Evolutionary Ecology and Conservation Genomics, University of Ulm, 89069 Ulm, Germany; [email protected] (J.K.); [email protected] (M.A.) 2 Richard-Wagnerstraße 9, 6020 Innsbruck, Austria; [email protected] 3 Department of Evolutionary Animal Ecology, University of Bayreuth, 95447 Bayreuth, Germany; [email protected] 4 Department of Conservation and Research, Bavarian Forest National Park, 94481 Grafenau, Germany; [email protected] * Correspondence: [email protected] Received: 15 October 2020; Accepted: 21 November 2020; Published: 24 November 2020 Simple Summary: Increasing forest management practices by humans are threatening inherent insect biodiversity and thus important ecosystem services provided by them. One insect group which reacts sensitively to habitat changes are the rove beetles contributing to the maintenance of an undisturbed insect succession during decomposition by mainly hunting fly maggots. However, little is known about carrion-associated rove beetles due to poor taxonomic knowledge. In our study, we unveiled the human-induced and environmental drivers that modify rove beetle communities on vertebrate cadavers. At German forest sites selected by a gradient of management intensity, we contributed to the understanding of the rove beetle-mediated decomposition process. One main result is that an increasing human impact in forests changes rove beetle communities by promoting generalist and more open-habitat species coping with low structural heterogeneity, whereas species like Philonthus decorus get lost.
    [Show full text]
  • Asymmetric Hindwing Foldings in Rove Beetles
    Asymmetric hindwing foldings in rove beetles Kazuya Saitoa,1, Shuhei Yamamotob, Munetoshi Maruyamac, and Yoji Okabea aInstitute of Industrial Science, University of Tokyo, Tokyo 153-8505, Japan; bGraduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 812-8581, Japan; and cThe Kyushu University Museum, Fukuoka 812-8581, Japan Edited by May R. Berenbaum, University of Illinois at Urbana–Champaign, Urbana, IL, and approved October 13, 2014 (received for review May 23, 2014) Foldable wings of insects are the ultimate deployable structures storage space (with some minor exceptions in soil- and cave- and have attracted the interest of aerospace engineering scientists dwelling species). The strategy is achieved by their extraordinary as well as entomologists. Rove beetles are known to fold their right–left asymmetric wing folding. As a result, rove beetles be- wings in the most sophisticated ways that have right–left asym- came highly diverse group, such that they account for 15% (i.e., metric patterns. However, the specific folding process and the rea- nearly 60,000 species) of all known species of Coleoptera. son for this asymmetry remain unclear. This study reveals how Despite the great potential of the process for engineering these asymmetric patterns emerge as a result of the folding pro- applications, few studies have been undertaken revealing the cess of rove beetles. A high-speed camera was used to reveal the details of this asymmetric wing folding. The wings of a rove beetle details of the wing-folding movement. The results show that these have two different crease patterns, but previous studies have characteristic asymmetrical patterns emerge as a result of simulta- described only one side.
    [Show full text]
  • The Beetle Fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and Distribution
    INSECTA MUNDI, Vol. 20, No. 3-4, September-December, 2006 165 The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and distribution Stewart B. Peck Department of Biology, Carleton University, 1125 Colonel By Drive, Ottawa, Ontario K1S 5B6, Canada stewart_peck@carleton. ca Abstract. The beetle fauna of the island of Dominica is summarized. It is presently known to contain 269 genera, and 361 species (in 42 families), of which 347 are named at a species level. Of these, 62 species are endemic to the island. The other naturally occurring species number 262, and another 23 species are of such wide distribution that they have probably been accidentally introduced and distributed, at least in part, by human activities. Undoubtedly, the actual numbers of species on Dominica are many times higher than now reported. This highlights the poor level of knowledge of the beetles of Dominica and the Lesser Antilles in general. Of the species known to occur elsewhere, the largest numbers are shared with neighboring Guadeloupe (201), and then with South America (126), Puerto Rico (113), Cuba (107), and Mexico-Central America (108). The Antillean island chain probably represents the main avenue of natural overwater dispersal via intermediate stepping-stone islands. The distributional patterns of the species shared with Dominica and elsewhere in the Caribbean suggest stages in a dynamic taxon cycle of species origin, range expansion, distribution contraction, and re-speciation. Introduction windward (eastern) side (with an average of 250 mm of rain annually). Rainfall is heavy and varies season- The islands of the West Indies are increasingly ally, with the dry season from mid-January to mid- recognized as a hotspot for species biodiversity June and the rainy season from mid-June to mid- (Myers et al.
    [Show full text]
  • The Evolution and Genomic Basis of Beetle Diversity
    The evolution and genomic basis of beetle diversity Duane D. McKennaa,b,1,2, Seunggwan Shina,b,2, Dirk Ahrensc, Michael Balked, Cristian Beza-Bezaa,b, Dave J. Clarkea,b, Alexander Donathe, Hermes E. Escalonae,f,g, Frank Friedrichh, Harald Letschi, Shanlin Liuj, David Maddisonk, Christoph Mayere, Bernhard Misofe, Peyton J. Murina, Oliver Niehuisg, Ralph S. Petersc, Lars Podsiadlowskie, l m l,n o f l Hans Pohl , Erin D. Scully , Evgeny V. Yan , Xin Zhou , Adam Slipinski , and Rolf G. Beutel aDepartment of Biological Sciences, University of Memphis, Memphis, TN 38152; bCenter for Biodiversity Research, University of Memphis, Memphis, TN 38152; cCenter for Taxonomy and Evolutionary Research, Arthropoda Department, Zoologisches Forschungsmuseum Alexander Koenig, 53113 Bonn, Germany; dBavarian State Collection of Zoology, Bavarian Natural History Collections, 81247 Munich, Germany; eCenter for Molecular Biodiversity Research, Zoological Research Museum Alexander Koenig, 53113 Bonn, Germany; fAustralian National Insect Collection, Commonwealth Scientific and Industrial Research Organisation, Canberra, ACT 2601, Australia; gDepartment of Evolutionary Biology and Ecology, Institute for Biology I (Zoology), University of Freiburg, 79104 Freiburg, Germany; hInstitute of Zoology, University of Hamburg, D-20146 Hamburg, Germany; iDepartment of Botany and Biodiversity Research, University of Wien, Wien 1030, Austria; jChina National GeneBank, BGI-Shenzhen, 518083 Guangdong, People’s Republic of China; kDepartment of Integrative Biology, Oregon State
    [Show full text]
  • On the Aleocharini of Turkey, with Notes on Some Species from Adjacent Regions
    View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Beiträge zur Entomologie = Contributions to Entomology (E-Journal) Beitr. Ent. Keltern ISSN 0005 - 805X 57 (2007) 1 S. 177 - 209 30.06.2007 On the Aleocharini of Turkey, with notes on some species from adjacent regions (Coleoptera: Staphylinidae, Aleocharinae) With 102 figures and 1 table VOLKER ASSING Summary Types and additional material of Aleocharini from Turkey and adjacent regions are revised. 6 species are (re-)described and illustrated: Megalogastria alata sp. n. (Bitlis, Erzurum), Aleochara caloderoides sp.sp. n. (Mer sin), A. (Ceranota) membranosa spsp.. n. (Antalya), A. (C.) bodemeyeri BERNHAUER, 1914, A. (C.) bitu- berculata BERNHAUER, 1900, and A. (C.) erythroptera GRAVENHORST, 1806. External and sexual characters of A. (C.) caucasica EPPELSHEIM, 1889, A. (Rheochara) leptocera EPPELSHEIM, 1889, and A. (R.) spalacis SCHEERPELTZ, 1969 are figured. The following synonymy is proposed: Aleochara moesta GRAVENHORST, 1802 = A. ebneri SCHEERPELTZ, 1929, syn. n. A lectotype is designated for A. spalacis SCHEERPELTZ. Additional records of Aleocharini from Turkey are reported, among them 7 first records. A checklist of the Aleocharini known from Turkish territory is compiled. Keywords Coleoptera, Staphylinidae, Aleocharinae, Aleocharini, Aleochara, Megalogastria, Pseudocalea, Tinotus, Palaearctic region, Turkey, taxonomy, new species, new records, distribution, checklist Zusammenfassung Typen und weiteres Material von Arten der Tribus Aleocharini aus der Türkei und angrenzenden Gebieten werden revidiert. 6 Arten werden beschrieben bzw. redeskribiert und abgebildet: Megalogastria alata sp. n. (Bitlis, Erzurum), Aleochara caloderoides sp.sp. n. (Mersin),(Mersin), A. (Ceranota) membranosa sp.sp. n. (Antalya),(Antalya), A. (C.) bodemeyeri BERNHAUER, 1914, A.
    [Show full text]
  • Coleoptera, Staphylinidae, Oxytelinae) Living in Floral Bracts of Columnea Medicinalis L
    Zootaxa 4394 (4): 559–566 ISSN 1175-5326 (print edition) http://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2018 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4394.4.6 http://zoobank.org/urn:lsid:zoobank.org:pub:83AA97F4-AC3F-41AD-A339-5A67A64AF656 Description and notes on natural history of a new species of Parosus Sharp, 1887 (Coleoptera, Staphylinidae, Oxytelinae) living in floral bracts of Columnea medicinalis L. (Gesneriaceae) MARGARITA M. LÓPEZ-GARCÍA1,2 & OSCAR H. MARÍN-GÓMEZ1,2 1Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, Colombia. E-mail: [email protected], [email protected] 2 Instituto de Ecología A.C., Carretera Antigua a Coatepec 351, El Haya, 91070 Xalapa, Veracruz, Mexico Abstract A new species of the recently revised genus Parosus is described, P. amayae López-García & Marín-Gómez sp. nov., from adult and larval specimens collected in bracts of Columnea medicinalis in the Natural Reserve Río Ñambí (Southwestern Colombia). Observations on the interaction with the plant, subsocial behavior, and population density are presented and discussed. Adults and larvae apparently live together and feed on eggs and larvae of flies that develop inside the decom- posing fruits of C. medicinalis. The new species is illustrated by color habitus photos, as well as its L1 and L3 larvae, male and female genitalia are depicted by line drawings. Key words: Oxytelinae, new species, decomposing fruits, gesneriads, subsocial behavior, cloud forest, Colombia, larvae Introduction Staphylinidae is the largest animal family with more than 61,300 described species (Newton 2015) and is dominant in a great variety of ecosystems, showing several ecological interactions (Thayer 2005).
    [Show full text]
  • Relative and Seasonal Abundance of Beneficial Arthropods in Centipedegrass As Influenced by Management Practices
    HORTICULTURAL ENTOMOLOGY Relative and Seasonal Abundance of Beneficial Arthropods in Centipedegrass as Influenced by Management Practices S. KRISTINE BRAMAN AND ANDREW F. PENDLEY Department of Entomology, University of Georgia, College of Agriculture Experiment Stations, Georgia Station, Griffin, GA 30223 J. Econ. Entomol. 86(2): 494-504 (1993) ABSTRACT Pitfall traps were used to monitor the seasonal activity of arthropod preda­ tors, parasitoids, and decomposers in replicated plots of centipedegrass turf for 3 yr (1989-1991) at two locations. During 1990 and 1991, the influence of single or combined herbicide, insecticide, and fertilizer applications on these beneficials was assessed. In total, 21 species of carabids in 13 genera and 17 species of staphylinids in 14 genera were represented in pitfall-trap collections. Nonsminthurid collembolans, ants, spiders, and parasitic Hymenoptera were adversely affected in the short term by insecticide applica­ tions targeting the twolined spittlebug, Prosapia bicincta (Say). Other taxa, notably orib­ atid Acari, increased over time in response to pesticide or fertilizer applications. Although various taxa were reduced by pesticide application during three of four sample intervals, a lack ofoverall differences in season totals suggests that the disruptive influence ofcertain chemical management practices may be less severe than expected in the landscape. KEY WORDS Arthropoda, centipedegrass, nontarget effects CENTIPEDEGRASS, Eremochloa ophiuroides Potter 1983, Arnold & Potter 1987, Potter et al. (Munro) Hack, a native of China and Southeast 1990b, Vavrek & Niemczyk 1990). Asia introduced into the United States in 1916, Studies characterizing the beneficial arthropod has become widely grown from South Carolina community and assessing effects of management to Florida and westward along the Gulf Coast practices on those invertebrates are especially states to Texas (DubIe 1989).
    [Show full text]
  • Remarks on Some European Aleocharinae, with Description of a New Rhopaletes Species from Croatia (Coleoptera: Staphylinidae)
    Travaux du Muséum National d’Histoire Naturelle © Décembre Vol. LIII pp. 191–215 «Grigore Antipa» 2010 DOI: 10.2478/v10191-010-0015-6 REMARKS ON SOME EUROPEAN ALEOCHARINAE, WITH DESCRIPTION OF A NEW RHOPALETES SPECIES FROM CROATIA (COLEOPTERA: STAPHYLINIDAE) LÁSZLÓ ÁDÁM Abstract. Based on an examination of type and non-type material, ten species-group names are synonymised: Atheta mediterranea G. Benick, 1941, Aloconota carpathica Jeannel et Jarrige, 1949 and Atheta carpatensis Tichomirova, 1973 with Aloconota mihoki (Bernhauer, 1913); Amischa jugorum Scheerpeltz, 1956 with Amischa analis (Gravenhorst, 1802); Amischa strupii Scheerpeltz, 1967 with Amischa bifoveolata (Mannerheim, 1830); Atheta tricholomatobia V. B. Semenov, 2002 with Atheta boehmei Linke, 1934; Atheta palatina G. Benick, 1974 and Atheta palatina G. Benick, 1975 with Atheta dilaticornis (Kraatz, 1856); Atheta degenerata G. Benick, 1974 and Atheta degenerata G. Benick, 1975 with Atheta testaceipes (Heer, 1839). A new name, Atheta velebitica nom. nov. is proposed for Atheta serotina Ádám, 2008, a junior primary homonym of Atheta serotina Blackwelder, 1944. A revised key for the Central European species of the Aloconota sulcifrons group is provided. Comments on the separation of the males of Amischa bifoveolata and A. analis are given. A key for the identification of Amischa species occurring in Hungary and its close surroundings is presented. Remarks are presented about the relationships of Alevonota Thomson, 1858 and Enalodroma Thomson, 1859. The taxonomic status of Oxypodera Bernhauer, 1915 and Mycetota Ádám, 1987 is discussed. The specific status of Pella hampei (Kraatz, 1862) is debated. Remarks are presented about the relationships of Alevonota Thomson, 1858, as well as Mycetota Ádám, 1987, Oxypodera Bernhauer, 1915 and Rhopaletes Cameron, 1939.
    [Show full text]
  • Phylogenetic Relationships and Distribution of the Rhizotrogini (Coleoptera, Scarabaeidae, Melolonthinae) in the West Mediterranean
    Graellsia, 59(2-3): 443-455 (2003) PHYLOGENETIC RELATIONSHIPS AND DISTRIBUTION OF THE RHIZOTROGINI (COLEOPTERA, SCARABAEIDAE, MELOLONTHINAE) IN THE WEST MEDITERRANEAN Mª. M. Coca-Abia* ABSTRACT In this paper, the West Mediterranean genera of Rhizotrogini are reviewed. Two kinds of character sets are discussed: those relative to the external morphology of the adult and those of the male and female genitalia. Genera Amadotrogus Reitter, 1902; Amphimallina Reitter, 1905; Amphimallon Berthold, 1827; Geotrogus Guérin-Méneville, 1842; Monotropus Erichson, 1847; Pseudoapeterogyna Escalera, 1914 and Rhizotrogus Berthold, 1827 are analysed: to demonstrate the monophyly of this group of genera; to asses the realtionships of these taxa; to test species transferred from Rhizotrogus to Geotrogus and Monotropus, and to describe external morphological and male and female genitalic cha- racters which distinguish each genus. Phylogenetic analysis leads to the conclusion that this group of genera is monophyletic. However, nothing can be said about internal relationships of the genera, which remain in a basal polytomy. Some of the species tranferred from Rhizotrogus are considered to be a new genus Firminus. The genera Amphimallina and Pseudoapterogyna are synonymized with Amphimallon and Geotrogus respectively. Key words: Taxonomy, nomenclature, review, Coleoptera, Scarabaeidae, Melolonthinae, Rhizotrogini, Amadotrogus, Amphimallon, Rhizotrogus, Geotrogus, Pseudoapterogyna, Firminus, Mediterranean basin. RESUMEN Relaciones filogenéticas y distribución de
    [Show full text]
  • An Electronic Checklist of the New World Chafers (Coleoptera: Scarabaeidae: Melolonthinae)
    AN ELECTRONIC CHECKLIST OF THE NEW WORLD CHAFERS (COLEOPTERA: SCARABAEIDAE: MELOLONTHINAE) Version 3 ARTHUR V. EVANS Research Associate, Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC; Department of Recent Invertebrates, Virginia Museum of Natural History, Martinsville, VA; Department of Biology, Virginia Commonwealth University, Richmond, VA; c/o1600 Nottoway Ave., Richmond, VA 23227, USA; [email protected] and ANDREW B. T. SMITH Canadian Museum of Nature, P.O. Box 3443, Station D, Ottawa, ON, K1P 6P4, Canada; [email protected] INTRODUCTION The following is a checklist of all Melolonthinae (Coleoptera: Scarabaeidae) found in the New World. It has been modified from Evans (2003), Evans and Smith (2005), and Smith and Evans (2005) and has been updated to 13 March 2009. Included in this checklist are all of the available names given for New World Melolonthinae (both valid and invalid). Tribes are listed in traditional order (pseudo-phylogenetically) with genera, species, and subspecies listed alphabetically within. Under each valid generic name the subgenera and synonymies are listed as are type species and, in some cases, citations for keys, checklists, and bibliographies. Listed under each valid species are synonymies, distributional data by country, and citations for new combinations and spellings. A complete bibliography is included in the “References” section of all papers mentioned in the checklist. The purpose of this checklist is to present accurate and complete information for all the names of Melolonthinae in the New World. The taxonomy herein is based on the current literature (even if we have unpublished data contradicting what has been published) and the nomenclature carefully follows the International Code of Zoological Nomenclature.
    [Show full text]
  • The Little Things That Run the City How Do Melbourne’S Green Spaces Support Insect Biodiversity and Promote Ecosystem Health?
    The Little Things that Run the City How do Melbourne’s green spaces support insect biodiversity and promote ecosystem health? Luis Mata, Christopher D. Ives, Georgia E. Garrard, Ascelin Gordon, Anna Backstrom, Kate Cranney, Tessa R. Smith, Laura Stark, Daniel J. Bickel, Saul Cunningham, Amy K. Hahs, Dieter Hochuli, Mallik Malipatil, Melinda L Moir, Michaela Plein, Nick Porch, Linda Semeraro, Rachel Standish, Ken Walker, Peter A. Vesk, Kirsten Parris and Sarah A. Bekessy The Little Things that Run the City – How do Melbourne’s green spaces support insect biodiversity and promote ecosystem health? Report prepared for the City of Melbourne, November 2015 Coordinating authors Luis Mata Christopher D. Ives Georgia E. Garrard Ascelin Gordon Sarah Bekessy Interdisciplinary Conservation Science Research Group Centre for Urban Research School of Global, Urban and Social Studies RMIT University 124 La Trobe Street Melbourne 3000 Contributing authors Anna Backstrom, Kate Cranney, Tessa R. Smith, Laura Stark, Daniel J. Bickel, Saul Cunningham, Amy K. Hahs, Dieter Hochuli, Mallik Malipatil, Melinda L Moir, Michaela Plein, Nick Porch, Linda Semeraro, Rachel Standish, Ken Walker, Peter A. Vesk and Kirsten Parris. Cover artwork by Kate Cranney ‘Melbourne in a Minute Scavenger’ (Ink and paper on paper, 2015) This artwork is a little tribute to a minute beetle. We found the brown minute scavenger beetle (Corticaria sp.) at so many survey plots for the Little Things that Run the City project that we dubbed the species ‘Old Faithful’. I’ve recreated the map of the City of Melbourne within the beetle’s body. Can you trace the outline of Port Phillip Bay? Can you recognise the shape of your suburb? Next time you’re walking in a park or garden in the City of Melbourne, keep a keen eye out for this ubiquitous little beetle.
    [Show full text]