Indian Journal of Geo Marine Sciences Vol. 49 (03), March 2020, pp. 398-405
Morphometric and meristic traits of three ambassid fish species (Chanda nama, Parambassis lala and Parambassis ranga)
Md. Alomgir Hossen, Md. Yeamin Hossain*, Dalia Khatun, Md. Nasir Uddin Pramanik, Most Farida Parvin, Julia Jasmin, Suraiya Sharmin, Obaidur Rahman, Zannatul Mawa, Md. Ashekur Rahman & Md. Rabiul Hasan
Department of Fisheries, Faculty of Agriculture, University of Rajshahi, Rajshahi, Bangladesh *[E-mail: [email protected]]
Received 08 August 2018; revised 13 October 2018
This research describes the first complete and inclusive information on morphometric and meristic characteristics of three Ambassid species (Chanda nama, Parambassis lala and Parambassis ranga) from the Mathabhanga River collected during August 2017 to July 2018. In our study, range of the total length was 2.0–7.2 cm for C. nama; 1.7-3.9 cm for P. lala and 1.9-6.4 cm for P. ranga. All length-weight relationships were highly significant (p < 0.001) with r2 values ≥ 0.900 (except W vs. PrDL relationship in P. lala, where, r2 = 0.861). Also, all length-length relationships were highly correlated with r2 values being ≥ 0.900. Based on r2 value, W vs. FL and TL vs. FL was best fitted model for all the three species among the 11 equations. The calculated fin formula is: dorsal, D. 21–22 (VIII/13–14); pectoral, P1. 8–10 (2/6–8); pelvic, P2. 6 (I/5); anal, A. 17-18 (III/14–15); and caudal, C. 20–22 (4/16–18) for C. nama; D. 20–21 (VIII/12–13); P1. 8–9 (1-2/7); P2. 6 (I/5); A. 16-18 (III/13–15); C. 18–20 (4-6/14) for P. lala and D. 20–21 (VIII/12–13); P1. 9–10 (2/7–8); P2. 6 (I/5); A. 18-20 (III/15–17); C. 22–24 (4-6/18) for P. ranga. These outcomes will be a functional tool for identification of Ambassid fishes up to species level in any water-bodies of Bangladesh as well as in the adjacent countries.
[Keywords: Ambassid species, Bangladesh, Mathabhanga River, Meristic traits, Morphometric characters] Introduction Morphometric and meristic traits i.e., L-W The fishes Chanda nama (Hamilton, 1822), relationships18-20, L-L relationships21,22 of several Parambassis lala (Hamilton, 1822) and Parambassis threatened species from Bangladesh are well ranga (Hamilton, 1822) are important part of the recognized. But there are no previous works on family Ambassidae. They are found in Bangladesh, morphometric and meristic traits of three Ambassid Combodia, India, Japan, Myanmar, Malaysia, Nepal fish species coverture a large number of linear and Pakistan1. They have high market value and mainly dimensions. Therefore, the goal of our research is to used for aquarium purposes2,3. These fishes inhabit describe the morphometric and meristic clear streams, rivers, canals, ditches, inundated paddy characteristics of 3 ambassid species (C. nama, fields, ponds, and beels4,5 and feed on invertebrates, P. lala and P. ranga) from the Mathabhanga River, worms and crustaceans4,6. The natural populations of south western (SW) Bangladesh. these species are waning for various causes counting habitat destruction and modification7,8, use of illegal Materials and Methods fishing gears and reckless fishing, water pollution and This research was conducted in the Mathabhanga other ecological changes to their habitat9,10. Worldwide, River (Lat. 23°21' – 23°51' N; Long. 88°38' – 88°62' the C. nama and P. ranga are considered as least E) located in Chuadanga district, SW, Bangladesh. concern and P. lala is as near threatened11. This River is supposed to be an essential feeding and Morphometric and meristic features can be breeding ground the fishes of southwestern considered as a functional tool for accurate Bangladesh18,21. Samples were occasionally collected recognition of any species and its classification12-14. from different parts of the Mathabhanga River in Additionally, in fisheries research, morphometric Chuadanga region, during August 2017 to July 2018 characters play a vital role for assessing the welfare of from the commercial fishers’ catch. The gears used by individuals as well as appraising the morphological commercial fishers including gill net, seine net, cast traits and the life history of populations from various net, conical and box trap. Fishes were instantly geographic region15-17. preserved in ice and fixed in 10 % formalin on HOSSEN et al.: MORPHOMETRIC AND MERISTIC TRAITS OF THREE AMBASSID FISHES 399
entrance in the research laboratory. The identification and a digital slide calipers (0.1 cm) was used to of fishes was done13,23, and the scientific names were measure various lengths (Table 1 and Fig. 1). The LWR confirmed with FishBase1. was estimated using the equation: W = a×Lb. The Magnifying glass was used to count the numbers of regression parameters (a and b) were calculated by: ln fin rays and the absence/presence of scale was (W) = ln(a) + b ln(L). Besides, 95 % confidence determined using an electric microscope (Model: BS- intervals (CI) of a and b and the co-efficient of 6022 trf). An electronic balance (0.1 g accuracy) was determination (r2) were estimated. All the extremes 24 used to measure the body weight (BW) of each individuals outliers were removed from the regress analyses . To
Table 1 ─ Morphometric measurements of three Ambassid fish species captured from the Mathabhanga River, southwestern Bangladesh Species n Measurements Min Max Mean ± SD 95 % CI % TL Chanda nama 128 TL (Total length) 2.0 7.2 4.26 ± 0.95 4.10 - 4.43 (Hamilton, 1822) FL (Fork length) 1.8 6.2 3.70 ± 0.82 3.56 - 3.84 86.8 SL (Standard length) 1.5 5.5 3.25 ± 0.75 3.12 - 3.38 76.2 HL (Head length) 0.4 1.8 0.96 ± 0.25 0.92 - 1.00 22.6 PrDL (Pre-dorsal length) 0.6 2.1 1.20 ± 0.30 1.14 - 1.24 27.9 PoDL (Post-dorsal length) 1.3 4.8 2.78 ± 0.65 2.67 - 2.89 65.2 DFBL (Dorsal fin base length) 0.7 2.7 1.59 ± 0.36 1.53 - 1.65 37.3 PcL (Pectoral length) 0.5 1.8 0.99 ± 0.26 0.94 - 1.03 23.2 PvL (Pelvic length) 0.5 1.8 1.08 ± 0.25 1.03 - 1.12 25.2 PrAnL (Pre-anal length) 0.9 3.0 1.78 ± 0.39 1.71 - 1.85 41.8 PoAnL (Post-anal length) 1.3 5.0 2.82 ± 0.66 2.70 - 2.94 66.2 AFBL (Anal fin base length) 0.4 2.0 1.04 ± 0.29 0.99 - 1.09 24.4 W (Body weight) 0.1 3.2 0.75 ± 0.57 0.65 - 0.85 Parambassis lala 119 TL (Total length) 1.7 3.9 2.91 ± 0.35 2.85 - 2.97 (Hamilton, 1822) FL (Fork length) 1.5 3.4 2.56 ± 0.31 2.50 - 2.61 87.8 SL (Standard length) 1.3 2.8 2.10 ± 0.24 2.06 - 2.15 72.3 HL (Head length) 0.3 0.9 0.65 ± 0.10 0.64 - 0.67 22.5 PrDL (Pre-dorsal length) 0.5 1.2 0.83 ± 0.11 0.81 - 0.85 28.6 PoDL (Post-dorsal length) 1.1 2.7 1.90 ± 0.27 1.85 - 1.95 65.4 DFBL (Dorsal fin base length) 0.6 1.5 1.07 ± 0.17 1.04 - 1.10 36.8 PcL (Pectoral length) 0.4 1.1 0.78 ± 0.12 0.76 - 0.80 26.7 PvL (Pelvic length) 0.5 1.1 0.78 ± 0.12 0.76 - 0.80 26.8 PrAnL (Pre-anal length) 0.5 1.5 1.02 ± 0.17 0.99 - 1.05 35.0 PoAnL (Post-anal length) 0.9 3.0 1.99 ± 0.35 1.92 - 2.05 68.3 AFBL (Anal fin base length) 0.4 1.5 0.97 ± 0.19 0.93 - 1.0 33.3 W (Body weight) 0.1 0.9 0.39 ± 0.14 0.37 - 0.42 Parambassis ranga 118 TL (Total length) 1.9 6.4 3.50 ± 1.06 3.31 - 3.70 (Hamilton, 1822) FL (Fork length) 1.6 5.5 3.02 ± 0.95 2.84 - 3.19 86.2 SL (Standard length) 1.4 5.0 2.62 ± 0.86 2.47 - 2.78 75.0 HL (Head length) 0.4 1.7 0.82 ± 0.30 0.76 - 0.87 23.3 PrDL (Pre-dorsal length) 0.7 2.2 1.13 ± 0.37 1.06 - 1.20 32.2 PoDL (Post-dorsal length) 1.2 4.4 2.27 ± 0.75 2.13 - 2.41 64.8 DFBL (Dorsal fin base length) 0.5 2.2 1.14 ± 0.39 1.07 - 1.21 32.5 PcL (Pectoral length) 0.5 1.7 0.87 ± 0.28 0.82 - 0.93 25.0 PvL (Pelvic length) 0.5 1.7 0.93 ± 0.30 0.87 - 0.98 26.5 PrAnL (Pre-anal length) 0.8 2.5 1.38 ± 0.44 1.30 - 1.46 39.4 PoAnL (Post-anal length) 1.1 4.4 2.33 ± 0.77 2.19 - 2.47 66.5 AFBL (Anal fin base length) 0.3 1.9 0.95 ± 0.34 0.89 - 1.01 27.1 W (Body weight) 0.1 4.1 0.82 ± 0.88 0.66 - 0.98
400 INDIAN J. GEO-MAR. SCI., VOL. 49, NO. 03, MARCH 2020
Fig. 1 — Showing the morphometric characters of three Ambassid fish species, (a) Chanda nama, (b) Parambasis lala and (c) Parambassis ranga confirm, whether the b values obtained in the linear Linear regression analyses were used to estimate a regressions were significantly different or not from total of 11 LLRs18. Best model for both L-W and L-L the isometric value (b = 3)25 the t-test was used. relationships was chosen based on the maximum HOSSEN et al.: MORPHOMETRIC AND MERISTIC TRAITS OF THREE AMBASSID FISHES 401
value of r2. Graph Pad Prism 6.5 software was used are given in Table 3. All L-W relationships were for statistical analyses. All statistical analyses were highly significant (p < 0.001) with r2 values ≥ 0.900 envisaged at 5 % significant level (p < 0.05). (except W vs. PrDL relationship in P. lala where, r2 = 0.861). Based on r2 value, W vs. FL was the best Results fitted model among 11 equations. The LLR Among the three Ambassid species the body shape measurements are presented in Table 4 and all LLRs of C. nama is strongly compressed laterally almost were also highly correlated with r2 values being flat, P. lala is small and almost rounded and P. ranga ≥ 0.900. In keeping with maximum r2 value, TL vs. is comparatively short and deeply compressed. Scales FL relationship was the best fitted model for all the are absent, body color is transparent and there are three ambassid fish species among 11 equations, in some black spot in P. ranga. The fins are yellowish in the Mathabhanga River, SW Bangladesh. C. nama; reddish in P. lala and transparent in P. ranga. The fin formula is: dorsal, D. 21–22 Discussion (VIII/13–14); pectoral, P1. 8–10 (2/6–8); pelvic, P2. 6 Information on morphometric and meristic (I/5); anal, A. 17-18 (III/14–15); and caudal, C. 20–22 characters were missing for these Ambassid species 23 (4/16–18) for C. nama; D. 20–21 (VIII/12–13); P1. except meristic information for C. nama . But our 8–9 (1-2/7); P2.6 (I/5); A. 16-18 (III/13–15); C. 18–20 calculated fin formula is dissimilar with that of (4-6/14) for P. lala and D. 20–21 (VIII/12–13); Talwar and jhingran23, because in our study we have P1. 9–10 (2/7–8); P2. 6 (I/5); A. 18-20 (III/15–17); identified a total of 8-10 pectoral fin rays, whereas the C. 22–24 (4-6/18) for P. ranga. In our study TL referenced study estimated total 13-15 pectoral fin ranged from 2.0–7.2 cm (mean ± SD = 4.26 ± 0.95) rays. In addition we have not found any spiny rays in and the BW ranged from 0.1–3.2 g (0.75 ± 0.57) for pectoral fins, which is in accordance with that study. C. nama; 1.7-3.9 cm (2.91 ± 0.35) and 0.1-0.9 g Also, there is difference with scale, because in our (0.39 ± 0.14) for P. lala and 1.9-6.4 cm (3.50 ± 1.06) study we have not found any scale for these species. and 0.1-4.1 g (0.82 ± 0.88) for P. ranga, respectively During this study, a total of 365 individuals of in the Mathabhanga River. The fork length (86.8 % three Ambassid species were collected. However, for C. nama; 87.8 % for P. lala and 86.2 % for C. nama smaller than 2.0 cm TL; P. lala smaller P. ranga) contains the high proportion of TL. All the than 1.7 cm TL and P. ranga smaller than 1.9 cm morphometric and meristic measurements are TL were not harvested because of inappropriate presented in Table 1 and 2. selection of fishing gears26-29. The maximum length The regression parameters (a and b) with their recorded in present study was 7.2 cm TL for C. nama, 95 % confidence intervals (Cl) of LWRs, coefficients that was lower than 11 cm in TL reported by of determination (r2) of three Ambassid fish species Menon30; 3.9 cm TL for P. lala, which was near
Table 2 — Meristic characters of three Ambassid fish species captured from the Mathabhanga River, southwestern Bangladesh Species n Characters Number of fin rays Spiny/ Unbranched/ Branched Chanda nama (Hamilton, 1822) 128 Dorsal fin rays 21 - 22 VIII/-/13 - 14 Pectoral fin rays 8 - 10 -/2/6 - 8 Pelvic fin rays 6 I/ - /5 Anal fin rays 17 - 18 III/-/14 - 15 Caudal fin rays 20 - 22 -/4/16 - 18 Parambassis lala (Hamilton, 1822) 119 Dorsal fin rays 20 - 21 VIII/-/12 - 13 Pectoral fin rays 8 - 9 -/1 - 2/7 Pelvic fin rays 6 I/ - /5 Anal fin rays 16 - 18 III/-/13 - 15 Caudal fin rays 18 - 20 -/4 - 6/14 Parambassis ranga (Hamilton, 1822) 118 Dorsal fin rays 20 - 21 VIII/-/12 - 13 Pectoral fin rays 9 - 10 -/2/7 - 8 Pelvic fin rays 6 I/ - /5 Anal fin rays 18 - 20 III/-/15 - 17 Caudal fin rays 22 - 24 -/4 - 6/18 402 INDIAN J. GEO-MAR. SCI., VOL. 49, NO. 03, MARCH 2020
Table 3 — Descriptive statistics and estimated parameters of the length-weight relationships of three Ambassid fish species captured from the Mathabhanga River, southwestern Bangladesh Species n Equation Regression 95 % CL of a 95 % CL of b r2 parameters a b Chanda nama (Hamilton, 1822) 128 W = a* TLb 0.0082 3.00 0.0071 - 0.0096 2.89 - 3.10 0.962 W = a* FLb 0.0124 3.01 0.0108 - 0.0143 2.90 - 3.12 0.964 W = a* SLb 0.0205 2.92 0.0182 - 0.0230 2.82 - 3.02 0.963 W = a* HLb 0.7105 2.55 0.6889 - 0.7327 2.44 - 2.67 0.938 W = a* PrDLb 0.4035 2.70 0.3912 - 0.4162 2.59 - 2.80 0.950 W = a* PoDLb 0.0336 2.88 0.0302 - 0.0372 2.78 - 2.99 0.961 W = a* DFBLb 0.1671 2.88 0.1551 - 0.1799 2.73 - 3.03 0.920 W = a* PcLb 0.6620 2.58 0.6413 - 0.6832 2.46 - 2.70 0.932 W = a* PvLb 0.5185 2.85 0.4995 - 0.5382 2.69 - 3.01 0.907 W = a* PrAnLb 0.1064 3.10 0.0982 - 0.1154 2.97 - 3.24 0.942 W = a* PoAnLb 0.0318 2.89 0.0290 - 0.0349 2.80 - 2.98 0.960 Parambassis lala (Hamilton, 1822) 119 W = a* TLb 0.0138 3.10 0.0120 - 0.0158 2.97 - 3.23 0.951 W = a* FLb 0.0219 3.03 0.0195 - 0.0245 2.91 - 3.15 0.960 W = a* SLb 0.0327 3.28 0.0295 - 0.0363 3.15 - 3.42 0.950 W = a* HLb 1.0186 2.33 0.9666 - 1.0734 2.22 - 2.44 0.935 W = a* PrDLb 0.6168 2.66 0.5885 - 0.6463 2.46 - 2.85 0.861 W = a* PoDLb 0.0709 2.60 0.0654 - 0.0768 2.48 - 2.73 0.938 W = a* DFBLb 0.3222 2.34 0.3147 - 0.3298 2.20 - 2.46 0.909 W = a* PcLb 0.6937 2.40 0.6728 - 0.7153 2.30 - 2.49 0.952 W = a* PvLb 0.6997 2.47 0.6801 - 0.7199 2.38 - 2.57 0.958 W = a* PrAnLb 0.3662 2.12 0.3583 - 0.3742 2.00 - 2.24 0.913 W = a* PoAnLb 0.0978 2.00 0.0917 - 0.1043 1.89 - 2.07 0.940 Parambassis ranga (Hamilton, 1822) 118 W = a* TLb 0.0105 3.22 0.0091 - 0.0121 3.10 - 3.34 0.962 W = a* FLb 0.0194 3.10 0.0171 - 0.0219 2.99 - 3.22 0.963 W = a* SLb 0.0325 3.02 0.0290 - 0.0365 2.90 - 3.14 0.957 W = a* HLb 1.0501 2.68 0.9830 - 1.1217 2.53 - 2.84 0.911 W = a* PrDLb 0.4099 3.05 0.3926 - 0.4281 2.91 - 3.19 0.942 W = a* PoDLb 0.0508 3.01 0.0455 - 0.0568 2.88 - 3.15 0.945 W = a* DFBLb 0.4181 2.81 0.3950 - 0.4425 2.64 - 2.99 0.900 W = a* PcLb 0.8929 3.01 0.8405 - 0.9487 2.84 - 3.18 0.912 W = a* PvLb 0.7422 2.95 0.7037 - 0.7829 2.79 - 3.11 0.919 W = a* PrAnLb 0.2299 2.98 0.2161 - 0.2446 2.83 - 3.13 0.930 W = a* PoAnLb 0.0500 2.94 0.0454 - 0.0552 2.83 - 3.06 0.957 See Table 1 for abbreviations in details; n, sample size; a and b are regression parameters of length-weight relationship; CI, confidence intervals; r2, co-efficient of determination similar with 3.9 cm in SL by Geetakumari31 and essential for fisheries resource planning and 6.4 cm TL for P. ranga which was lower than the management34,39. maximum value of 8.0 cm in TL32. It was quite hard The allometric co-efficient (b) of length-length to catch larger specimens from the Mathabhanga relationships may vary between 2.0 to 4.040 and River because of lack of larger-sized fishes 2.5-3.524. In our findings, maximum b values were in the fishing grounds33-35 or may be where the between the limit of 2.5–3.5, however, some were larger sized fishes exist the fishermen did not go also observed between the limit 2.0–4.0. The b values there36-38. The information on maximum length is may be fluctuated even in case of same species which needed to calculate the growth parameters i.e., may have happened because of several factors such as growth coefficient, asymptotic length, that are physiology, sex, preservation technique and variation HOSSEN et al.: MORPHOMETRIC AND MERISTIC TRAITS OF THREE AMBASSID FISHES 403
Table 4 ─ The estimated parameters of the length-length relationships (y = a +b * x) of three Ambassid fish species captured from the Mathabhanga River, southwestern Bangladesh Species n Equation Regression parameters 95 % CL of a 95 % CL of b r2 a b Chanda nama 128 TL = a + b× FL -0.0014 1.154 -0.0875 to 0.0848 1.131 - 1.176 0.988 (Hamilton, 1822) TL = a + b× SL 0.1549 1.264 0.0589 - 0.2509 1.235 - 1.293 0.984 TL = a + b× HL 0.7201 3.683 0.5841 - 0.8560 3.546 - 3.820 0.958 TL = a + b × PrDL 0.5294 3.144 0.4016 - 0.6572 3.039 - 3.248 0.966 TL = a + b × PoDL 0.2142 1.458 0.1263 - 0.3021 1.427 - 1.488 0.986 TL = a + b × DFBL 0.1597 2.581 -0.0058 to 0.3253 2.479 - 2.682 0.953 TL = a + b × PcL 0.8042 3.496 0.6561 - 0.9524 3.351 - 3.641 0.948 TL = a + b × PvL 0.2298 3.754 0.0589 - 0.4006 3.599 - 3.909 0.948 TL = a + b × PrAnL -0.0040 2.400 -0.1521 to 0.1442 2.318 - 2.481 0.964 TL = a + b × PoAnL 0.2485 1.424 0.1656 - 0.3314 1.395 - 1.453 0.987 TL = a + b × AFBL 0.9019 3.237 0.7555 - 1.0483 3.101 - 3.372 0.946 Parambassis lala 119 TL = a + b × FL 0.1272 1.089 0.0511 - 0.2033 1.059 - 1.118 0.978 (Hamilton, 1822) TL = a + b × SL -0.0991 1.430 -0.2158 to 0.0177 1.375 - 1.485 0.958 TL = a + b × HL 0.7118 3.357 0.5796 - 0.8439 3.157 - 3.557 0.904 TL = a + b × PrDL 0.4328 2.980 0.2812 - 0.5844 2.799 - 3.161 0.901 TL = a + b × PoDL 0.5294 1.250 0.4365 - 0.6223 1.202 - 1.299 0.957 TL = a + b× DFBL 0.8179 1.950 0.6912 - 0.9445 1.834 - 2.067 0.904 TL = a + b × PcL 0.7409 2.790 0.6212 - 0.8607 2.637 - 2.942 0.918 TL = a + b × PvL 0.7311 2.790 0.6011 - 0.8610 2.626 - 2.955 0.906 TL = a + b × PrAnL 0.8940 1.980 0.7829 - 1.0051 1.873 - 2.088 0.919 TL = a + b × PoAnL 0.9882 0.967 0.9059 - 1.0705 0.926 - 1.008 0.950 TL = a + b × AFBL 1.2256 1.738 1.1210 - 1.3302 1.632 - 1.844 0.900 Parambassis 118 TL = a + b × FL 0.1582 1.108 0.1005 - 0.2160 1.090 - 1.126 0.992 ranga (Hamilton, TL = a + b × SL 0.3013 1.220 0.2303 - 0.3723 1.194 - 1.246 0.987 1822) TL = a + b × HL 0.7225 3.405 0.6001 - 0.8448 3.265 - 3.546 0.952 TL = a + b × PrDL 0.3067 2.822 0.1950 - 0.4185 2.728 - 2.916 0.968 TL = a + b × PoDL 0.3183 1.403 0.2317 - 0.4048 1.366 - 1.439 0.981 TL = a + b × DFBL 0.4435 2.689 0.3180 - 0.5689 2.585 - 2.794 0.957 TL = a + b × PcL 0.2932 3.669 0.1569 - 0.4296 3.520 - 3.817 0.954 TL = a + b × PvL 0.2654 3.484 0.1347 - 0.3961 3.350 - 3.618 0.959 TL = a + b × PrAnL 0.2313 2.373 0.1294 - 0.3331 2.303 - 2.444 0.975 TL = a + b × PoAnL 0.3241 1.365 0.2602 - 0.3880 1.339 - 1.391 0.989 TL = a + b × AFBL 0.5965 3.058 0.4818 - 0.7112 2.944 - 3.172 0.961 in length sizes of the specimens33,41,42. Besides, all LLRs Acknowledgement were highly correlated but comparison was not possible The authors would like to extend their sincere due to lack of previous literature. However, the current appreciation to (i) University of Rajshahi, Bangladesh work estimates the best model among the 11 equations for funding of this project (No. 375/5/52/RU/Agri- (using different lengths) on the basis of co–efficient of 25/17-18; 01/03/2018) and (ii) PIU-BARC, NATP-2, determination that is very helpful for future studies. Our CRG sub-Project: 484 for technical supports to collect research will be supportive for the fisheries scientist to morphological and meristic data. the assessment of the remaining stocks and to identify the Ambassid species in the Mathabhanga River, SW, References Bangladesh and contiguous ecosystems. 1 Froese, R. & Pauly, D. (Eds.). FishBase 2018: World Wide Web electronic publication. Available 404 INDIAN J. GEO-MAR. SCI., VOL. 49, NO. 03, MARCH 2020
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