植物研究雑誌 J､Jpn､Bot. Originals 71:305-309(1996)

Therqpogonpallidus)NewtotheFloraofThailand

NoriyukiTANAKA

DepartmentofEducation,SchoolofLiberalArts,TeikyoUniverslty,

3590tsuka,Haci句i-Shi,Tokyo,192-03JAPAN

(ReceivedonMay7,1996)

771erqPOgOn,amOnOtyPICgenuSOfIliliaceaeknown丘omtheHimalayanrangeanditsadjacentreglOnS,

wasconfirmedtoexistalsoinnorthernThailand.ThisnewThailocalitybecomesthenewsoutheastemmost

localityinthegeographicalrangeofthegenus･ThespecimenfromThailandhassomedifferencesfrom

Therqpogon(T.pallidus)aspresentlyknown,buttheoveralldiffbrenceseemsnotsogreat,andtherefbreIassign

thisspecimentothesamespecies･ThisreportcontainsmorphologlCalandotherdataonthisspecimen･

InhisNumericalList,Wal1ich(1832)listedthe 丘omOクゐg呼OgO托. nameOphippogon?pallidusWall.fbraplantfrom However,thisspecimenhassomedifferencesfrom Nepalwithoutanydescription.Kunth(1850)studied Therqpogon(T.pallidus),aSPreSentlyknown.First, thisplantandgaveitadetaileddescrlPtlOnunderthe itisrelativelysmallinsizeasawhole･Forexample,

Same name.Later,having studied this species, itspedicelsareOnly3to4mmlong,equalingthe

Maximowicz(1871)established the new genus Shortestlengtheverrecordedinthisspecies(Noltie T%erqpogon.ThisgenushasbeenmonotypICSinceits 1994),anditsstyles2.1to2.6mmlong,beingnearly establishmentcomprlSlngOnlyonespecies,T･Pal- halfoftheshortestlength(4.3mm:Noltie1994)ever

1idus(Wall.exKunth)Maxim. reported.Second,there exists a differencein the ForthepurposeofstudyingthegenusOphi甲OgOn flowercolour,aSfollows.Flowersofthisspecimen taxonomically,Ihavehadaloanofsomespecimens arewhite,andtepalshaveapaleyellowcentralline

Ofthatgenus丘･OmtheNaturalHistoryMuseumin andyellowtlP,aCCOrdingtothecollector'srecord Paris,andIfoundthatitincludedaspecimenwhich (citedbelow),WhiletheflowersofT.pallidussofar

CanbeidentifiedasTherppogon(Figs.1&2).This reportedarewhiteortingedpinkishorpurplish(Baker SPeCimenwasformerlyidenti丘edasOphiqpogonby 1875,Hooker1875,1894,Krause1930,Hara1971, theorlglnalcollectors.However,inmanyrespectsthe Liang1978,Utech1979,PoluninandStainton1984,

SPeCimenaccordswellwiththeknowncharacteristics Chen1987,Noltie1994).Intheserespects,theplants

Of771erqPOgOn･Forexample,thisspecimenhasa Ofthisspecimenappeartobedifferentiatedtosome

SuPeriorovarywith9to120Vulesineachloculus degreefromT･Pallidussofarknownl)･However,the

(Fig.2B),filamentsofnearlythesamelengthasthe OVeralldeviationofthisspecimenseemsnotsogreat anthers(Fig.2A),andleaveswithaprominentmidrib astowarrantestablishinganewspecies.Therefbre,I beneath(Fig.2A).ManyofthesecharaCterSdiffer asslgnthisspecimentoT.pallidus,andthespeciesis

ー305- 306 植物研究雑誌 第71巻 第6号 平成8年12月 henceregardedashavingabroaderrangeOfvariation thatitwascollectedfromalimestonepeak･However, thanbefore. ageneralftatureofthehabitatandthealtitudewhere

Asforthehabitatofthisspecimen,itisnoteworthy thisspecimenwascollectedalsocoincidewiththe

Fig･1･AspecimenidentifiedasT･PallidusfromThailand･ December1996 JournalofJapaneseBotanyVol.71No.6 307

isolated,andsouthernmOStinitstotalrange･Thereis PreViousrecordsforT.pallidus(Forexample,Noltie (1994)reportsthatitshabitatisdryrockyhillsides, agoodpossibilitythatinfuturethisspeciswillalso anditisfoundataltitudesbetween1830-3970m). beR)undinthereglOnSbetweenthisnewlocalityand Therppogonpallidusis known to existin the thefbrmerlyknownlocalities(Forexample,itmay HimalayanrangeandreglOnSadjacenttoit,including be丘)undinMyanmar,etC.). Uttar Pradesh(Kumaon),,,, ThehabitatofthisverylntereStlngSPeCiesinthe

Assam,Khasia,,andN.andN.W.ofYunnan SOuthernmOSt,isolatedlocalityinThailandisimpor-

(Maximowicz1871,Baker1875,Hooker1875,1894, tantinvariousrespects,andIhopeitwillnotbelost Engler1888,Krause1930,Kitamura1955,Haraetal. infutureduetohumanactivities.

1978,Liang1978,PoluninandStainton1984,Wu

1984,Chen1987,Noltie1994).Whereas,aSPeCimen Theropogonpallidus(Wall.exKunth)Maxim., hereconcernedwas collectedinnorthernThailand. Bull.Acad.Imp.Sci.St.-P6tersb.15:90(1871). AsfarasIamaware,thisreportisthefirsttorecord qphiqpogon?pallidus[Wa11.,Numer.List:179, theoccurrenceofT.pallidusinThailand.Thisnew n.5138(1832),nOm.nud.】exKunth,Enum.Pl.5:300 localityinThailandisfarawayeven丘■Omthenearest (1850). knownlocality(N.W.Yunnan)ofT.pallidus,hence DescrlPtionbasedonthespecimenfromThailand:

Fig･2･Close-uPSOfsomepartsofthespecimeninFig･1･A･Aninflorescenceandleaves･Stamen,PIStil,and midribareindicatedbyus",りpり,and=m‖,reSPeCtively･B･Lowerpartofapistil,Showinganalignmentofovules inaloculus･Theupperpartofthestylehasbeenlost.Scaleinmm. 308 植物研究雑誌 第71巻 第6号 平成8年12月

Tuftedherb.Totallyglabrous.Rootstockthickand habitat.Accordingly,athoroughexaminationofaset

Very Short with many thick fibrous roots.Leaves Ofthesespecimensmayhelpconsiderablyourunder- linear,attenuatedtotheapex,Sheathedatbase,uPtO Standingofthevariationofthisspeciesatthelocality. 27cmlong,2.5-5.7mmwide,midribelevatedbe- 2)specimenswiththesameCO11ectionnumberkeptat

neathandprominent.Scapeslender,COmPreSSedes- AAH,BKF,C,andLwereidentiBedbyCharoenphol

PeCiallyinupperpart,angledandnarrowlywlnged, (1974)asOphiqpogonmalcolmsoniiRoyleexHook.f. 14･5-17cmlong.Inflorescenceraceme,1.7-1.8cm Accordingtohispaper,thesespecimenswerecol-

long,4-flowered.FlowersborneSlngly,Subtendedby 1ectedin the samelocality as the specimen here

bothabractandabracteole.Pedicelscurveddown- examined,butatadifferentaltitude,1200m.

Ward,3-lmmlong.Bractslanceolatetonarrowly Retもrences lanceolate,WithmembranousmarglnS,0･35-2cm BakerJ.G.1875.Revision ofthe Genera and Species of long･Flowerscampanulate,drooplng.Tepals6,free .(771erqPOgOn:P,562･)Journ･Linn･Soc.,Bot. 14:508-632,pl.17-20. tobase,imbricated,OVate,5-6mmlong,2.5-3mm CharoenphoIC.C.1974.StudiesinThaiLiliaceae.ThaiFor. Wide,1-Veined,Slightlyrecurvedoutwardinupper Bull.,Bot.$:88-94. ChenS.-C.1987.TherqpogonMaxim.p.562,Pl.304(ト2).In: Part･Stamens6.AntherslanCeOlate,minutelyapICulate WuC.-Y.(ed.),FloraXizangica.Vol.5.SciencePress, at apex,2mmlong,1ntrOrSe.Pollen grains pale Be毎1ng. EnglerA･1888･Liliaceae･P･10-9-1･(77wqpogon:P･81)･In:A･ yellow･Filamentsc.1.5mmlong,broadandflat- EnglerundK.Prantl(ed.),DieNattirlichenPflanZenfamilien tened,C･0.9mmwide,basevery shortlyconnate II(5).Verlagv.WilhelmEngelmann,Leipzig. HaraH.1971.Liliaceae,P.165-175.(771erPPOgOn:P･174).In: laterally betweenneighbourlng丘1aments.Ovaries HaraH.(ed.),FloraofEasternHimalaya.2ndReport.Univ. SuPerior,Subglobose with3longitudinalshallow OfTokyoPress,Tokyo. grooves(i･e･,trilobedincross-SeCtion),2mmlong,3- +,SternW.T.andWi11iamsL.H.J.1978.AnEnumera- tionoftheFloweringPlantsofNepal.Vol.1.(771erPPOgOn locular.Ovules9-12ineachlochlus,arrangedin2 Maxim,P.80.).BritishMuseum,London. HookerJ.D.1875.771erqPOgOnPallidus･Curtis'Bot･Mag.101: rowslongitudinally on centralmarglnalplacenta･ t.6154. Styleveryslender,Slightlyattenuated,2.1-2.6mm +1894.FloraofBritishIndia.Vol･6.77wqpogon:P･ 324.L.Reeve,London. long･Stigmaveryminute. KrauseK･1930.Liliaceae,P.227-386･(771erPPOgOn:P.370- Specimenexamined:Thailand,ChiangMai,Doi 371).In:A.EnglerundK.Prantl(ed.),DieNatむ1ichen Pflanzenfamilien.15a.Verlagv.WilhelmEngelmann,Leip- Chiengdao.Limestonepeak.Alt.c.2075m.Between Zig. rocks.FIs.white,butlobeswithapaleyellowcentral KitamuraS.1955.FlowerlngPlants andFernS.P.73-290. (771erqPOgOn:P.96).In:KiharaH.(ed.),FaunaandFloraof lineandye1lowtip･20June1968･C･F･VanBeusekom NepalHimalaya.Vol.I.FaunaandFloraResearchSociety. &C.Phengkhlai,n.13522)(P,eXHerb.Lugd.Batav.). KyotoUniv.,Kyoto. KunthC.S.1850.Enumerati6plantarum.Tomus5.Qphiqpogon ?pallidusWall･:P.300-301.StutgardiaeetTubingae. Ithank the curator ofthe NationalMuseum of LiangS･-Y･1978.771erqPOgOnMaxim.:P.2&pl.1.In:Wang F･-T･etTangT.(ed.),Liliaceae(2).FloraReipublicae NaturalHistoryin Paris(P)fbr theloan ofthis PopularisSinicae15.SciencePress,Beijing. SPeCimen. Maximowicz C.187l.Ophiopogonis speciesin herbariis Petropolitanis.(771erqPOgOnMaxim.p.89-90.)Bu11.Acad. Imp.Sci.St.-Pitersb.15:83-90. Endnotes NoltieH･J･1994･FloraofBhutan･Vol･3(1)･771erPPOgOn:P･ l)Aspecimenhereexaminedisoneofthosedistrib- 192.RoyalBotanicGardenEdinburgh,Edinburgh. PoluninO･andStaintonA.1984･FlowersoftheHimalaya. utedfromtheherbariuminLeiden(L),Netherlands. 771erqPOgOn:P.428,Pl.123.OxfbrdUniv.Press,Oxford. UtechF.H.1978.FloralvascularanatomyoftheHimalayan Thismeansthatpluralnumberofspecimenswiththe T71erqPOgOnPallidusMaxim.(Liliaceae-Conva11arieae).An- Same COllection number were madefromthe same nalsofCamegieMuseum48:25-41. December1996 JournalofJapaneseBotanyVol.71No.6 309

WallichN.1832.AnumericallistofdriedspecimensintheEast Wu.C.-Y.1984.Index Florae Yunnanensis.TomusII. IndiaCompany'sMuseum.Ophiqpogon?pallidusWall･:P･ (Therqpogon:P.1939).ThePeople'sPublishingHouse, 170,n.5138.London. Yunnan,China.

田中教之:撒e叩聯〝pαJJf血ざタイに産す ジャノヒゲ属((わゐ∫呼曙0乃)の研究のためにパ の多くの個体についてさらに詳しく研究する必要 リの自然誌博物館から標本を借用していたが,そ もあるが,現時点においては,この標本は,r

｡の中の一点に,採集者によって(わ鋸呼曙0乃と同 pα招血∫であると同定する. 定されているが同属とは明らかに異なるものがあっ rpαJ〃血∫はこれまでヒマラヤとそれに続く山 た.この標本は,多くの性質において乃eJ叩曙0乃 地,ないしはその近傍の山地に分布することが知 pαJJ励ぶ(ユリ科)と一致する.しかし,全体的 られていたが,この標本の採集地はタイの北部で に小形であり,花梗は長さ3-4mmで,花柱も あり,これまで知られているもっともそこに近い 長さ2.ト2.6mmしかない.また,採集者の記録に 生育地(中国雲両省北西部)からもかなり離れて よれば,花は白色であり,花被片の端部は黄色で, いる.タイにおける本種の分布はこれまで知られ 花被片の中央には淡黄色の筋があるという.これ ていないと思われるので,ここに報告することに まで知られているrクα肪血ぶの花色は,白色か, した.この地が同種の分布域の南限となる. ピンクないし紫色を帯びる.このような点におい 乃eJ叩卿乃属はrpαJJ誠一∫一種からなる単型属で てこの標本の個体は,これまで知られているr ある.その分類学的位置づけについては,今後の pαJJ肋ぶとは僅かながら分化しているのかもしれ 研究に委ねられるところが多い.この極めて興味 ない.しかし,仮にこれらの性質が原産地におい 深い種の,しかもその分布域の最東南部の隔離的 て安定しているものだとしても,その差異は種を 分布地であるこのタイの生育地が,いっまでも無 分かっはど大きいものではないと判断する.生育 事であることを望みたい. 地や海抜も既知のものとあまり変わりはない(た (帝京大学文学部教育学科) だし,この標本の産地は石灰岩地である).現地