Belgian Arachnological Society ARABEL

Spiders of Cyprus (Araneae)

A catalogue of all currently known species from Cypr us.

R OBERT B OSMANS , J OHAN V AN K EER , A NTHONY R USSELL - S MITH , M ICHAEL H ADJICONSTANTIS , M ARJAN K OMNENOV , J AN B OSSELAERS , S IEGFRIED H UBER , D UNCAN M C C OWAN , R OWLEY S NAZELL , A RTHUR D ECAE , C HRISTOS Z OUMIDES , K ARL - H INRICH K IELHORN AND P IERRE O GER .

Arachnological Contributions

Newsletter Belgian Arachnological Society, volume 34 (suppl.). 15 November 2019 ISSN 0774 - 7225

2

R OBERT B OSMANS 1,14 , J OHAN V AN K EER ², A NTHONY R USSELL - S MITH ³, M ICHAEL H ADJICONSTANTIS 4 , M ARJAN K OMNENOV 5 , J AN B OSSELAERS 6 , S IEGFRIED H UBER 7 , R OWLEY S NAZELL 8 , D UNCAN M C C OWAN 9, A RTHUR D ECAE 10 , C HRISTOS Z OUMIDES 11 , K ARL - H INRICH K IELHORN 12 & P IERRE O GER 13

(1) Robert Bosmans, Terrestrial Ecology Unit, Ledeganckstraat 35, B - 9000 Gent, Belgium (2) Johan Van Keer, Bormstraat 204, bus 3, B - 1880 Kapellen op den Bos, Belgium (3) Anthony Russell - Smith, 1 Bailiffs Cottage, Doddington, Sittingbourne, Kent ME9 0JU, United Kingdom (4) Michael Hadjiconstantis, Association for the Protection of Natural Heritage and Biodiversity of Cyprus, Kastellorizou 4, Lakatamia, 2314 Nicosia, Cyprus (5) Marjan Komnenov, Blvd. Kuzman Josifovski Pitu, 19/5/3, 1000 S kopje, North Macedonia (6) Jan Bosselaers, Rerumnovarumlaan 2, B - 2340 Beerse, Belgium (7) Siegfried Huber, Ottenbohlstraße 12, 88690 Mühlhofen, Germany (8) Rowley Snazell, Four Winds, 10, Bon Accord Road, Swanage, Dorset BH19 2DS, United Kingdom (9) Duncan McCowan, 13, Agios Andronikos, Dali 2540, Nicosia, Cyprus (10) Arthur Decae, Natural History Museum, Rotterdam, the Netherlands (11) Christos Zoumides, Energy, Environment & Water Research Center, The Cyprus Institute, 2121 Nicosia, Cyprus (12 ) Ka rl - Hinrich Kielhorn, Albertstra ß e 10, 10827 Berlin, Germany (13) Pierre Oger, Rue du Grand Vivier 14, 4217 Warêt l’Evêque, Belgium (14) Corresponding author: [email protected]

Arachnologica l Contributions. Newsletter of the Belgian arachnological Society 34 (suppl.). 1 5 November 2019

ARABEL v.z.w. / a.s.b.l.

Bestuur/Bureau

V OORZITTER /P RÉSIDENT : Robert Bosmans, Ooievaarstraat 98, 9000 Ge nt O NDERVOORZITTER /V ICE - P RÉ SIDENT : Jan Bosselaers, Rerum Noovarumlaaan 2, 90 Melle S ECRETARIS /S ÉCRÉTAIRE : Rudy Jocqué,Veeweidestraat, 3040 Huldenberg P ENNINGMEESTER /T RÉSORIER : Johan Van Keer, Bormstraat 204 bus 3, 1880 Kapelle - op - den - Bos W EBMASTER Arnauld Henrard, Ru e de Namur 69/2, 1300 Wavre R EDACTEUR N IEUWSBRIEF Léon Baert, Leopold I - plein 3A, 8400 Oostende E DUCATIEVE ACTIVITEIT EN : Mark Alderweireldt, Begoniastraat 5, 9090 Melle B EH E ERDER DATABANK Frederik Hendrickx, Hoekskenstraat 11, 9070 Heusden L IDGELD /C ÔTISATION : 20 Euro R EKENING /C OMPTE : IBAN: BE65 0014b4419 4196 BIC: GEBABEBB Vermelding: “Lidgeld/côtisation ARABEL”

Spiders of Cyprus 3

Table of contents SUMMARY ...... 5 I. INTRODUCTION ...... 6 II. HISTORY OF THE STUDY OF THE SPIDERS OF CY PRUS ...... 6 III. METHODS AND MATERIAL ...... 7 IV. TAXONOMIC PART ...... 9

F AMILY A RANEIDAE ...... 9 F AMILY C TENIDAE ...... 10 F AMILY D ICTYNIDAE ...... 11 F AMILY G NAPHOSIDAE ...... 12 F AMILY L INYPHIIDAE ...... 16 F AMILY L YCOSIDAE ...... 24 F AMILY N EMESIIDAE ...... 25 F AMILY P HYXELIDIDAE ...... 26 F AMILY T ETRAGNATHIDAE ...... 28 F AMILY T HERIDIIDAE ...... 30 F AMILY Z ODARIIDAE ...... 32 V. LIST OF SPECIES AND RECORDS ...... 33

F AMILY A GELENIDAE ...... 34 F AMILY A NYPHAENIDAE ...... 35 F AMILY C HEIRACANTHIIDAE ...... 43 F AMILY C ITHAERONIDAE ...... 44 F AMILY C LUBIONIDAE ...... 44 F AMILY C TENIDAE ...... 45 F AMILY C YBAEIDAE ...... 45 F AMILY D ICTYNIDAE ...... 45 F AMILY D YSDERIDAE ...... 48 F AMILY F ILISTATIDAE ...... 51 F AMILY G NAPHOSIDAE ...... 52 F AMILY H AHNIIDAE ...... 68 F AMILY H ERSILIIDAE ...... 68 F AMILY L INYPHIIDAE ...... 69 F AMILY L IOCRANIDAE ...... 82 F AMILY L YCOSIDAE ...... 84 F AMILY M IMETIDAE ...... 90 F AMILY M ITURGIDAE ...... 91 F AMILY M YSMENIDAE ...... 91 F AMILY N EMESIIDAE ...... 91 F AMILY O ECOBIIDAE ...... 91 F AMILY O ONOPIDAE ...... 93 F AMILY O XYOPIDAE ...... 94 F AMILY P ALPIMANIDAE ...... 96 F AMILY P HILODROMIDAE ...... 96 F AMILY P HOLCIDAE ...... 100 F AMILY P HYXELIDIDAE ...... 101 F AMILY P ISAURIDAE ...... 102 F AMILY S ALTICIDAE ...... 102 F AMILY S CYTODIDAE ...... 117 F AMILY S EGESTRIIDAE ...... 119 F AMILY S ICARIIDAE ...... 119 F AMILY S PARASSIDAE ...... 120 F AMILY T ETRAGNATHIDAE ...... 121 F AMILY T HERAPHOSIDAE ...... 122 Spiders of Cyprus 4

F AMILY T HERIDIIDAE ...... 123 F AMILY T HOMISIDAE ...... 138 F AMILY T ITANOECIDAE ...... 144 F AMILY T RACHELIDAE ...... 145 F AMILY U LOBORIDAE ...... 145 F AMILY Z ODARIIDAE ...... 146 F AMILY Z OROPSIDAE ...... 149 VI. DISCUSSION ...... 151 ACKNOWLEDGEMENTS ...... 152 REFERENCES ...... 153 ADDENDUM. LIST OF SP IDERS OF CYPRUS ...... 160

Male of Evarcha nepos (O. Pickard - Cambridge, 1872)

Spiders of Cyprus 5

Summary The a rachnofauna of Cyprus currently includes 449 species, of which all known and all new localities are presented. Nineteen species are considered endemic, but several new species will be described in the future. The unknown male of Dipoena galilaea Levy & Ami tai and the unknown females of Improphantes cypriot Tanasevitch, I. turok Tanasevitch, Palliduphantes minimus Deeleman - Reinhold and Steatoda trianguloides Levy are described for the first time. Male and female of Phaeocedus hebraeus Levy, 1999 have been in correctly matched and the female is described for the first time. Lepthyphantes lagunculus Denis from Algeria is redescribed and transferred to the genus Theonina n. comb. The species is also cited for the first time in Spain, Morocco and Cyprus. Lathys ar abs Simon, 1911 is not a synonym of L. stigmatisata (Menge, 1869) but a distinct species, contra Lethinen, 1967. Ctenidae and Anahita syriaca is a new family, genus and species for Europe. The following species are new to Europe: Improphantes turok Tanasev itch, Mermessus fradeorum (Berland), Mogrus logunovi Prószyński, Neoscona sodom Levy, Phaeocedus hebraeus Levy , Pterotricha schaefferi (Audouin), Silhouettella tomer Saaristo, Talanites dunini Platnick & Ovtsharenko, Tetragnatha demissa (C. L. Koch), Zygiometella perlongipes (O. Pickard - Cambridge, 1872), Ox yopes sobrinus O. Pickard - Cambridge and Theridion ochreolum Levy & Amitai. Not less than 325 species are cited here for the first time in Cyprus. A number of species records were critically evaluated and were apparently misidentific a tions.

Spiders of Cyprus 6

I. Introduc tion Cyprus is an island in the Eastern Basin of the Mediterranean with a total land area of 9251 km². For a detailed description of the geography, the climate, the geology and the habitats, we refer to Sparrow & John (2016) ‘An introduction to the wildl ife of Cyprus’. Cyprus has two distinct mountain ranges, the large, central, gradually sloping Troodos Massif, up to 1951m, and the narrow, steep, Pentadaktylos Range ( Keryneia mountains), reaching 1024m and fringing the northern coast for 160 km. Cyprus has an extreme Mediterranean climate, with hot and dry summers, and mild and rainy winters. Natural and semi - natural habitats in Cyprus include woodlands (coniferous, oak and riparian), scrubland (phrygana) and some grasslands, dunes and small wet lands. Woodlands are preferred habitats for spiders and several endemic species can be found here. Coniferous forests occur at all altitudes in Cyprus. At lower levels, they are dominated by Calabrian pine Pinus brutia . Black pine ( Pinus nigra pallasiana ) forests occur on the higher peaks of the Troodos Mountain, at somewhat lower altitudes mixed with Pinus brutia . Cypress forests can mainly be found on the slopes of the Pentadaktylos range, and Juniperus forests on the slopes of the Akamas Peninsula and Karpasia. Finally, cedar forest with the endemic Cedrus brevifolia are mainly found in the ‘Cedar Valley’ in the Paphos forest. Oak forests ( Quercus infectoria veneris , Q. alnifolia ) occur as small stands, mainly on the Troodos mountain range. Riparian for ests with Platanus orientalis , Salix alba and Alnus orientalis occur along the upper parts of the major rivers of the island, while in the lower parts of the rivers which are often dry they are replaced by riparian galleries with Nereum oleander and Tamari x spp. Scrublands of phrygana are very common in Cyprus, but difficult to explore for spiders. The main type is a short, pillow - forming, thorny vegetation, grading into more arborescent vegetation with Olive ( Olea europaea ), Pistachio ( Pistacia lentisca ) and Kermes oak ( Quercus coccifera ). Wetland habitats are important for spiders including the marshy areas surrounding them. Some natural or brackish lakes include the Akrotiri, Famagusta, Larnaka, and Paralimni lakes. Many reservoirs and dams have been created by man, some of them with interesting marshy areas, such as the Achna Lake. Agricultural environments such as olive, carob and almond groves, vineyards, patches of low intensity cereals and grazing land may also offer suitable habitats for spider species.

II. History of the study of the spiders of Cyprus Very little arachnological research has been carried out on Cyprus and the spider fauna is very poorly known. The first spider mentioned in the literature was also one of largest and most str iking species found on the island. In an important paper on Mediterranean Mygalomorpha, the German arachnologist, Ausserer (1871) described Ischnocolus gracilis , now considered a junior synonym of Chaetopelma olivaceum (C. L. Koch, 1841). In the same paper , he described another mygalomorph spider, Brachythele incerta which has never again been reported in the literature. In the early years, Simon (1874) described Nurscia flavipes from Cyprus, but this appears to have been a synonym of N. albosignata Simo n 1874. Similarly, he described Micaria todilla from Cyprus (Simon, 1878), but this appears to have been a junior synonym of M. ignea O. Pickard - Cambridge, 1872. Strand (1907) added 3 species to the fauna of Cyprus in a small contribution. The Polish ara chnologist, Kulczyński (1908) was the first to publish a paper dealing entirely with spiders of Cyprus. Nineteen species were listed for the first time in Cyprus, of which seven were new to science. In general, researchers contributing to the arachnofaun a of Cyprus concerned themselves with revisions of families or genera. Deeleman - Reinhold (1988) contributed with the genus Dysdera (four species), Bosmans & Van Keer (1999) with the theridiid genus Enoplognatha (two species), Knoflach (1999) with the theri diid genus Neottiura (one species), Bosmans (2009) with the zodariid genus Zodarion (one new and three revised species), Jäger et al. (2011) with the sparassid genus Olios (one species) and finally Zonstein & Marusik (2016 , 2019 ) contributed to the Filist atidae of Cyprus. Wunderlich (1980, 1995a - c, 2011) published many contributions to the Mediterranean arachnofauna, including the fauna of Cyprus: Walckenaeria cyprusensis in 1995, Theridion cyprusense in 2011 and Ariadna cyprusensis in 2011. A few au thors described single species from Cyprus. Deeleman - Reinhold (1985) discovered a new linyphiid spider, Palliduphantes minimus . Bosselaers (2002) described a new genus, Akamasia (named after

Spiders of Cyprus 7

the Akamas Peninsula in Cyprus) which includes only one known spe cies at present; the Cyprus endemic Akamasia cyprogenia . Vollmer (1997) described a second Chaetopelma species from Cyprus: C. karlamani . Azarkina (2006) described the salticid species Aelurillus cypriotus , Panayiotou et al. (2010) described Berinda cypria . In a paper on European Linyphiidae, Tanasevitch (2011) described several new species from the Mediterranean, among them two new species ( Araeoncus cypriacus and Improphantes cypriot ) and nine species cited for the first time in Cyprus. Logunov (2013, 2015) is an important researcher of the spider fauna of Cyprus. In 2013, he added 15 species to the fauna of the island and a further 13 species of Salticidae in 2015, as a result of which, salticids are among the best known of Cypriot spiders. Further recent contributions are those of Chatzaki & Russell - Smith (2017) adding 7 species of Gnaphosidae, Aharon et al. (2017) clarifying the identity of the Artema species occurring in Cyprus, Azarkina et al 2018 ) , describing the unknown female of Aelurillus cy priotus , Bosmans et al. (2018) revising the genus Haplodrassu s, Gücel et al. (2018, 2019) describing new Dysderocrates and Harpactea species from Cypriot caves, Özkütük et al . (2019), citing a Cheiracanthium species for the first time and finally Chatzaki & Van Keer (2019) citing Echemus levyi for the first time. Finally, we should mention two recent books dealing with the spider fauna of Cyprus. McCowan’s book (2014), “60 Cypriot Spiders” is a popular introdu ction to the spiders of Cyprus. Bosmans, in Sp arrow & John (2016 , Wildlife of Cyprus ), wrote a contribution on spiders which adds no new species for the island . Summarizing, Nentwig et al. (2019) mention 107 species reported from Cyprus in the literature. Compared with other Mediterranean islands t his number is relatively low, as can be seen in Table 1:

Surface area Distance from Number of Author (km²) mainland (km) known species Sardinia 24.090 200 459 Pantini et al ., 2013 Crete 8.336 96 430 Bosmans et al. , 2013 Lesbos 1632 12 296 Bosmans et al. , 2009 Chios 904 8 315 Russell - Smith et al. , 2011 Korfu 607 7 243 Russell - Smith 2014 Lefkada 334 2 117 Russell - Smith 2014 Cyprus 9.251 70 107 Nentwig et al. , 2019

Table 1: Number of spider species on, and surface area of, the Mediterranean islands Sardinia, Crete, Lesbos, Chios, Korfu, Lefkada and Cyprus.

Many more spider species are thus to be expected in Cyprus. Along with a team of European arachnologists, several collecting trips have been organised, and some local arachnologists started to cooperate with the project. III. Methods and Material The material studied was collected by different arachnologists. RB, A.R - S, MK, JVK and KHK identified their materia l themselves. They are not listed as collectors in the text, only the deposition in their collection is referred to. Material of Michael Hadjiconstantis and Christos Zoumides was identified by RB and material of John Murphy was identified by AR - S. AR - S h ad some Salticidae and Thomisidae identified by Dmitri Logunov, the Agelenidae by Angelo Bolzern, some Pholcidae by Bernhard Huber and some Gnaphosidae by Maria Chatzaki . Material of the Natural History Museum was identified by Marjan Komnenov.

An overview of all examined collections is given in table 2.

Collector Year Location Seen by Spiders of Cyprus 8

M. Askins 1998, 2007 Personal collection A.R - S R. Bosmans 2010, 2011 Personal collection RB J. Bosselaers 1996, 1997 Personal collection JB C. Felton 1997 Liverpool Museum A.R - S M. Hadjiconstantis 2013 - 201 8 Personal collection RB and colleagues S. Huber 2017 Personal collection RB K. - H. & U. Kielhorn 2011, 2013, 2014, 2018 Perso nal collection KHK, RB and friends M. Komnenov 1986 NHMW MK D. McCowan 2004 - 201 8 Personal collection A.R - S J. Murphy 1983, 2002 Manchester Museum A.R - S T. Russell - Smith 2007, 2013, 2017 Personal collection A.R - S R. Snazell 2007, 2013 Personal collection A.R - S J. Van Keer 2007, 2010, 2011 Personal collection JVK Different collectors Different years British Museum RB

Table 2. List of collections examined and number of records

Abbreviations BMNH: British Musuem of Natural History CAR - S: Collection Tony Russell - Smith CDM: Collection Duncan McCowan CJB: Collection Jan Bosselaers CJVK: Collection Johan Van Keer CKHK: Collection Karl - Hinrich Kielhorn CMA: Collection Martin Askins CMH: Collection Michael Hadjiconstantis (and including Christos Zoumides) CRB: Collection Robert Bosmans CRS: Collection Rowley Snazell CSH: Collection Siegfried Huber MMUE: Manchester University Museum NHML: Naturalis History Museum, Leiden NH MW: Collection of the Natural History Museum in Vienna WML: World Museum, Liverpool ZMUT: Zoological Museum of the University of Turku

Spiders of Cyprus 9

IV. Taxonomic part

In this part, s ome poorly known or rarely illu strated species are discussed .

Family Aranei dae

Neoscona sodom Levy, 1998 (Figs 1 a - e) Neoscona sodom Levy, 1998a: 340, figs 117 - 126.

Figs 1a - e. Neoscona sodom Levy. a. Male, dorsal view; b. Male palp, retrolateral view; c. Idem, ventral view; d. Idem, prolateral view; e. Idem, ventro - prolateral view.

Diagnosis Neoscona sodom Levy differs clearly from Neoscona adianta (Walckenaer, 1802) and N. subfusca (C. L. Koch), the other Neoscona species occurring in Cyprus, by its general colour, abdominal pattern and further by the sh ape of the median apophysis with a black spike, and the fan - shaped terminal apophysis.

Records See part V, page 4 1 .

Spiders of Cyprus 10

Comments This species is only known from two localities in Israel, the male from the Dead Sea region, the female from the Yizre ’el Valley. It is new to Cyprus.

Family Ctenidae Anahita syriaca (O. Pickard - Cambridge, 1872) (Figs 2 a - f) Anahita syriaca ; Levy, 2003: 11, figs 16 - 22 (descr. ♂, ♀).

Diagnosis Ctenidae are recognised by the peculiar eye disposition in rows of 2 - 4 - 2 (Fig. 1c). Since Anahita syriaca is the only European species of the family, it cannot be confused with any other. The male is further characterised by the absenc e of a tibial apophysis and the semi - circular embolus (Figs 1d - f), the female by the shape of the square, epigynal plate.

Comments This is the first record of both the family and this genus for Europe. The female was first described by O. Pickard - Camb ridge (1872) from the plains of the Jordan, the male was described much later by Levy (2003), from Israel.

Records See part V, page 45 .

Figs 2 a - f. Anahita syriaca (O. Pickard - Cambridge). a. Male, dorsal view; b. Male, dorsal view, living specimen; c. Male prosoma, dorsal view; d. Male palp, retrolateral view; e. Idem, ventral view; f. Idem, proventral view.

Spiders of Cyprus 11

Family Dictynidae

Lathys arabs Simon, 1911 (Figs 3 a - e) Lathys arabs Simon, 1911: 278. Lathys puta ; Lehtinen 1967: 243, figs. 257, 266 (synonymy, here rejected).

Figs 3a - e. Lathys arabs Simon. a. Male palp, ventral view; b. Idem, retrolateral view; c. Part of male palpal cymbium and tibia, detail, dorso - prolateral view; d. Epigyne, ventral view; e. Vu lva, ventral view.

Type material Lectotye ♂, 1 ♀ 2 ♀♀ paralectotypes of Lathys arabs Simon from Algeria, S. Bou Saada, Aïn Oghrab, MNHN AR 403; 3 paratypes from Algeria, Biskra, MNHN AR 396; ♀♀ paralectotypes from Tunisia, Nefzaoua, Vibert leg., MNHN AR 420; examined; all by present designation, and all mentioned in the original description by Simon 1910.

Comparative material examined Lectotye ♂, 4 ♀♀ paralectotypes of Lathys maura Simon from Algeria, Annaba, Forêt de l’Edough, on bushes togeth er with Lat h ys humilis , VI.1884, MNHN AR 413.

Diagnosis Lathys arabs was described from Algeria by Simon (1910). This species was synonymized with Lathys stigmatisata by Lehtinen (1967), although he did not mention examining type material. The first author examined the type material of Lathy arabs and figures of the species were published in Bosmans et al. (2009). Males differ from the related Lathys stigmatisata (Menge) by the absence of a patellar apophysis and the shorter distal part of the conduc tor, directed in posteo - dorsal direction. In females of L. arabs, the

Spiders of Cyprus 12

epigynal openings are widely separated and there are two pairs of vertical loops in the fertilisation duct (fig. 3e), but only one pair in Lathys stigmatisata . Lathys arabs has to be con sidered a distinct species. The paper of Marusik et al. (2006) was of great help to describe the different parts of male palps and epigynes. Description Measurements: Male (n = 7): Total length 2.2 - 3.2; prosoma 0.81 - 1.05 long. Female (n = 15): Total le ngth 2.2 - 3.4; prosoma 0.88 - 1.38 long. Colour: Prosoma yellowish brown, with foveal region grey, often some pale grey radiating stripes; legs pale yellowish; abdomen dorsally with large pale grey spot, filled with 5 - 6 grey chevrons, laterally with a patte rn of oblique, alternate pale grey and darker grey stripes, the paler stripes sometimes interrupted by darker spots or completely dark grey with whitish spots. Male palp (Figs 3 a - c): Patella without apophysis; tibia with short, pointed retrodorsal and l arger prodorsal apophyses; base of conductor trapezoid in ventral view, distal part cork - shrew - like, with 3.5 loops, terminally bluntly pointed; bulbus circular, flattened, embolar base not visible, arising at ventral side of bulbus, embolus whip - like, imb edded in lateral grooves of bulbus, describing 3.5 circles. Epigyne (Figs 3 d - e): Median septum as wide as the diameter of each atrium; copulation ducts with 3 horizontal and two vertical loops

Records See part V, page 47 .

Family Gnaphosida e

Berinda idae Lissner, 2016 (Figs 4 a - e) Berinda idae Lissner, in Lissner & Chatzaki, 2016a : 9, figs 1a - c, 2a - b, 3a - b, 4a - b, 5a - b, 6 - 7 ( descr. ♂♀).

Figs 4 a - e. Berinda idae Lissner. a. Male, dorsal view; b. Male palp, retrolateral view; c. Idem, ventral view; d. Idem, prolateral view; e. Female, dorsal view; f. Epigyn e, ventral view; g. Vulva, dors al view.

Spiders of Cyprus 13

Comments According to the World Spider Catalog (2018), the genus Berinda i ncludes seven species , all from the eastern Mediterranean region or Central Asia. Three of them occur in Cyprus. Males of Berinda idae differ from all other Berinda species by the short, triangular retrolateral tibial apophysis (Figs 4b - c), females by the short, rectangular hood (Figs 4 f). This is the first record of Berinda idae outside of Greece. Previously, it was known from the Dodekanese (Kalymnos, Nisyros) and the Cyclades (Santorini, Christiani Islet) (Lissner & Chatzaki, 2012). In the specimens from Cyprus, the tibial apophysis of the male looks shorter (probably due to a different angle of view) and the epigynal hood is more incised posteriorly, but these are considered minor variations.

Records See part V, page 53 .

Drassyllus cypri us Chatzaki & Russell - Smith, 2017 (Figs 5 a - k) Drassyllus cyprius Chatzaki & Russell - Smith 2017: 24 (descr. ♂♀).

Figs 5 a - k. Drassyllus cyprius Chatzaki & Russell - Smith. a. Male prosoma, dorsal view; b. Female, dorsal view; c. Idem, ventral view; d., Male palp, retrolateral view; e. Idem, ventral view; f. Idem, prolateral view; g. Male palpal tibia, lateral view; h.Tip of bulbus, ventral view; i. Epiyne, ventral view; j. Vulva, ventral view; k. Vulva, dorsal view.

Spiders of Cyprus 14

Comments This recently described species is apparently common and widespread in Cyprus.

Records See part V, page 56 .

Phaeocedus hebraeus Levy, 1999 (Figs 6 a - i) Phaeocedus hebraeus Levy, 1999: 429, figs 1 - 5 (descr. ♂, non ♀).

Figs 6 a - i. P haeocedus hebrae us Levy a. Male, dorsal view; b. Male palp, retrolateral view; c. Idem, ventral view; d. Idem, proventral view; e. Female, do rsal view; f. Idem, ventral view; g. Epigyne, ventral view; h. Vulva, vent ral view; i. Vulva, dorsal view.

Spiders of Cyprus 15

Type material Holotype male from Israel, Mount Scopus,14.VI.1944, paratype female (misidentified, see below) from Israel, Poriya, 20.IV.1940, A. Shulov leg; examined by Chat z aki & Van Kee r (2019).

Diagnosis Males of this species are easily recognised by the large, elongate rectangular conductor (fig. 6 c), females by the small, rounded postero - median depression in the epigyne (fig. 6 g).

Comments Phaeocedus hebraeus was described from Israel by Levy (1999), the male from Mount Scopus in Jerusalem, the female from Poriya in Galilea, about 100 km to the North. Levy (1999) considered them to belong to the same species. However, in Cyprus we collected males of the species together with females that are d ifferent from Levy’s female specimen. It is clear that Levy (1999) mismatched both sexes, the female belonging to another genus . Chatzaki & Van Keer (2019) came to the same conclusion and attributed the female to a juvenile Drassodes .

Records See par t V, page 65 .

Other material examined GREECE: Attiki: Alepohori camping site, 1 ♂, 22.V.1998, R. Bosmans leg. (CRB).

Disribution Until now only known from Israel, cited here for the first time in Greece and Cyprus.

Synaphosus shirin Ovt sharenko, Levy & Platnick, 1994 (Figs 7 a - e) Synaphosus shirin Ovtsharenko, Levy & Platnick, 1994 : 23, figs 82 - 83 (descr. ♂); Chatzaki & R ussell - Smith, 2017 : 248, figs 29 - 34 (descr. ♂♀).

Figs 7 a - e. Synaphosus shirin Ovtsharenko, Levy & Platnick. a. Male, dorsal view; b. Male palp, retrolateral view; c. Idem, ventral view; d. Idem, prolateral view; e. Idem, dorsal view.

Comment s Synaphosus shirin was originally described from Iran by Ovtsharenko et al. (1994 ) . Only the female was known. Recently, Chatzaki & Russell - Smith (2017 ) found females together with males which they described for the first time. More Synaphosus species are cited further down in th is paper.

Records See part V, page 65 .

Spiders of Cyprus 16

Talanites dunini Platnick & Ovtsharenko, 1991 (Figs 8 a - e) Talanites dunini Platnick & Ovtsharenko, 1991 : 117, figs 5 - 8 (descr. ♂♀). Talanites dunini Levy, 2009 : 23, figs 51 - 52.

Comments According to the World Spider Catalog (2018), the genus Talanites comprises 15 species: one from Tunisia, 1 from Egypt, 8 from the Near and Middle East, and 5 from the United States and Mexico. Talanites dunini was described from Azerbaijan and Turkmenistan (Platnick & Ovtsharenko, 1991), and Levy (2009) cited it also in Israel. Here it is mentioned for the first time from Europe in Cyprus.

Records: See part V, page 66 .

Figs 8 a - e. Talanites dunini Platnick & Ovtsharenko. a. Male, dorsal view; b. Male palp, retrolateral view; c. Idem, ventral view; d. Idem, prolateral view; e. Idem, dorsa l view.

Family Linyphiidae

Canariphantes tenerrimus (Simon, 1929) Lepthyphantes tenerrimus Simon, 1929: 611, 737 (descr. ♀). Lepthyphantes homonymus Denis, 1934: 76, figs 3 (descr. ♂).

The specimens cited below correspond best with the drawings of Tanasevitch (2011) of Canariphantes homonymus Denis (now a junior synonym of C. tenerrimus (Simon), based on ma terial from Greece. Typical for C. tenerrimus is the L - shaped, terminally pointed lamella, clearly differing from the terminally bifurcate lamella of C. nanus . Except for the citation of Tanasevitch from Greece, C. tenerrimus is only known from the w ester n part of the Mediterrranean (Spain, Portugal, France, Morocco and Algeria). Further studies are necessary to confirm the present identification.

Records: See part V, page 71 .

Spiders of Cyprus 17

Figs 9 a - k. a - f. Improphantes cypriot Tanasevitch. a. Male, dor sal view; b. Female, dorsal view; c. Male palp, retrolateral view; d. Idem, prolateral view; e. Epigyne, ventral view; f. Idem, lateral view. g - k. Improphantes turok Tanasevitch. g. Male, dorsal view; h. Female, dorsal view; i. Male palp, retrolateral view ; j. Idem, prolateral view; k. Epigyne, ventral view.

Spiders of Cyprus 18

Improphantes cypriot Tanasevitch, 2011 (Figs 9 a - f, 10a - b) Improphantes cypriot Tanasevitch, 2011: 67 (descr. ♂).

Diagnosis Closely related to I. turok , males differing by the numerous teeth on the dorsal margin of the lamella in I. cypriot (Fig 10a), compared to only small, terminal teeth in I. turok (Fig. 10c). However, some of the teeth of the lamella can be broken off. Females differ in the scape being wider than long in I. cypriot , as long as wide in I. turok (Fig. 10b versus fig. 10e).

Description Measurements: Male: Total length 0.61 - 0.67; prosoma 0.55 - 0.59 long, 0 .43 - 0.49 wide. Female: Total length 0.73 - 0.90; prosoma 0.53 - 0.62 long, 0.39 - 0.49 wide. Spination: Fe I with 1 prolateral spine, Fe II - IV spineless; Ti with 2 dorsal spines, Ti I with additional pro - and retrolateral spine, Ti II with additional retrolate ral spine.

Records See part V, page 74 .

Improphantes turok Tanasevitch, 2011 (Figs 9 g - k, 10 c - g) Improphantes turok Tanasevitch, 2011: 69, figs 65 - 69 (descr. ♂, ♀).

Description Male: Total length 0.59 - 0.70; prosoma 0.41 - 0.66 long, 0. 31 - 0.38 wide. Female: Total length 0.63 - 0.77; prosoma 0.41 - 0.55 long, 0.31 - 0.52 wide. Colour and spination as in Improphantes cypriot .

Diagnosis See Improphantes cypriot .

Records See part V, page 75 .

Spiders of Cyprus 19

Figs 10a - g. a - b. I mprophantes cypriot Tanasevitch. a. Male palp, retrolateral view; b. Epigyne. c - g. Improphantes turok Tanasevitch c. Male palp, retrolateral view; d. Embolic division, prolateral view; e. Epigyne, ventral view; f. Epigyne, dorsal view. g. Vulva, ventral vi ew, distal part dissected.

Spiders of Cyprus 20

Megalepthyphantes turkeyensis Tanasevitch, Kunt & Seyyar, 2004 (Figs 11 a - f) Lepthyphantes congener Kulczyński, 1908 : 68, pl. 2, fig. 17 (descr. ♂ , misidentified). Megalepthyphantes turkeyensis Tanasevitch, Kunt & Seyyar, 2005 : 279, figs 1 - 9 (descr. ♂ ). Megalepthyphantes turkeyensis Tanasevitch , 2011 : 74, figs 82 - 83 ( misidentification as L congener confirmed ). Palliduphantes montanus ; Topçu et al. 2013: 91 (misiden tification).

Comments Kulczyński (1908 ) described a male collected in Cyprus as the unknown male of Lepthyphantes congener (O. Pickard - Cambridge, 1872), now considered a synonym of Frontinellina frutetorum (C. L. Koch, 1834). Tanasevitch (2011) poin ted out that this is not correct and that the specimen described by Kulczyński is in reality Megalepthyphantes turkeyensis Tanasevitch, Kunt & Seyyar, described in 2005 from Turkey. Tanasevitch ( op. ci t .) cites the species also from Ikaria, an island of the Cyclades, and this record has been overlooked by Nentwig et al. (2019) and Chatzaki et al. (2015). Tanasevitch, Kunt & Seyyar ( op. cit. ) mention examining females of Megalepthyphantes turkeyensis but these were never figured or described. The first author was able to examine specimens cited as Palliduphantes montanus ( Kulczyński , 1898) from Turkey and these appeared to belong to Megalepthyphantes turkeyensis as well. Palliduphantes montanus must be deleted from the Turkish list of spiders.

Figs 11 a - f. Megalepthyphantes turkeyensis Tanasevitch, Kunt & Seyyar . a. Female, dorsal view; b. Male palp, retrolateral view; c. Idem, ventral view; d. Idem, prolateral view; e. Epigyne, ventral view; f. Vulva, ventral view .

Diagnosis The species is closely related to Megalepthyphantes lydiae Wunderlich, 1994 from Gr eece. Males differ in the terminally pointed lamella (fig. 10b), rounded in M. lydiae , and the absence of a dorsal tooth in the embolus of M. turkeyensis . Females differ in the scape being more rectangular and longer than wide (fig. 10e), while it is more rounded and somewhat wider than long in M. lydiae .

Comparative material examined Greece, Lesbos, Mandamanos, 2 ♂♂ 2 ♀♀ of Megalepthyphantes lydiae (CRB).

Records: See part V, page 7 6 .

Spiders of Cyprus 21

Palliduphantes minimus (Deeleman - Reinhold, 1986) (Figs 12 a - h)

Lepthyphantes minimus Deeleman - Reinhold, 1986: 46, figs 35a - b (descr. ♂).

Figs 12 a - h. Palliduphantes minimus (Deeleman - Reinhold). a. Male, dorsal view; b. Female, dorsal view; c. Idem, ventral view; d. Male palp, retrolateral view; e. Idem, prolateral view; f. Idem, ventral view; g. Lamella, ventral view; h. Epigyne, ven t ral view.

Spiders of Cyprus 22

Diagnosis Palliduphantes minimus is the only Palliduphantes species in Cyprus. It is a small, pale species of which males are recognised by the strongly pointed distal tooth of the lamella and the presence of three teeth along its dorsal margin (fig. 11g), females by the short, strongly curved scape (fig. 11h).

Description Measurements: Male: Total length 1.5; prosoma 0.65 long, 0.51 wide. Female: Total length 1.6 ; prosoma 0.71 long, 0.56 wide. Colour: Pale yellowish, only eyes with narrow black rings. Prosoma: Anterior median eyes separated by half their diameter, from the anterior laterals by their diameter; posterior eyes separated by their diameter. Chelice rae with 3 large promarginal teeth and 4 smaller, closely set retromarginal teeth, laterally with 18 stridulating ridges in te male. Spination: Legs with long spines, proximal dorsal spine on tibia I 2.7 as long as the tibias diameter; femur I with 1 pr olateral spine; tibia I with 2 dorsal, 1 prolateral and 1 retrolateral spine, Tibia II with 2 dorsal and 1 retrolateral spine, Ti III - IV with 2 dorsal spines; metatarsi I - IV with 1 dorsal spine. Male palp (Figs d - g): Dorsal spine on patella slightly long er than its diameter, that on tibia 1.5 times as long as its diameter; proximal part of paracymbium with 4 hairs and small basal tooth, distal part small and rounded; lamella elongated, with 2 basal tooth, terminally split into 3 sharp teeth. Epigyne (Fi g. 12 h): Scape sigmoid, short and strongly curved.

Records See part V, page 78 .

Theonina laguncula (Denis, 1937) n. comb. (Figs 13 a - k)

Lepthyphantes laguncula Denis, 1937: 1045 (descr. ♂, ♀); Bosmans, 1985: 172 (citation); Bosmans 2006: 178 (citation).

Type material Type series of one ♂ and ♀ from Algeria, For êt de Zouagha, Djebel Daya, 13.V.1936; depositon unknown, not in MNHNP.

Etymology The etym ological meaning of the name laguncula was not given by Denis (1937). In Latin, laguncula means ‘a small flask’ and this obviously refers to the shape of the apophysis on the palpal patella.

Diagnosis Both sexes are at once recognised by their contr asting colour pattern. Males are further easily recognised by the shape of the chelicerae and the bottle - shaped apophysis on the palpal patella, females by the rectangular scape.

Comments The shape of the male palp (large basal tubercles on cymbium, C - shaped embolus), and of the female epigyne (flattened S - shaped scape with broad distal part) resemble closely those of Theonina cornix and T. linyphioides (see Bosmans, 2006) and Lepthyphantes laguncula is therefore best placed in this genus.

Description Measurements: Male (n=1): Total length 2.1; carapace 0.83 long, 0.68 wide. Female (n=6): Total length 1.8 - 2.3; carapace 0.65 - 0.86 long, 0.58 - 0.71 wide. Colour: Carapace speckled olive brown, region behind the eyes and thoracic region yellowish orange; chelicerae and sternum yellowish orange; legs yellowish white, tibiae, metatarsi and tarsi with median and distal grey annulations, often poorly marked; abdomen grey, with two oblique white chevrons and 5 transverse white stripes, venter pale yellow with small scattered white spots.

Spiders of Cyprus 23

Figs 13 a - k. Theonina lag uncula (Denis) n. comb. a. Male, dorsal view; b. Female, dorsal view; c, d. Male palp, retrolateral view; e. Idem, prolateral view; f. Embolic division, ventral view; g, h. Cymbium, dorsal view; i. Epigyne, ventral view; j. Vulva, ventral view; k. Idem, do rsal view.

Spiders of Cyprus 24

Chelicerae: In male tapering at the edge, diverging, laterally with 10 - 11 stridulating ridges, fang groove with one anterior and one posterior tooth, chelicerae of female normal, fang groove with 3 lar ge anterior and 4 smaller posterior teeth. Legs relatively short, Femur I only slightly longer than length of prosoma. Spinulation: Femora spineless; Tibiae I - II with 2 dorsal, 1 prolateral and 1 retrolateral spine, tibiae III - IV wih 2 dorsal spines, metarsi spineless. Male palp (Figs 13 c - h): Patella with strong cylindrical apophysis bearing a long hair; cymbium with curved, posterior horn, at retrolateral side accompanied by a smaller tubercle; paracymbium U - shaped, basal part with median tooth, distal part gradually n arrowing, terminally rounded; radix with a strong distal concavity at prolateral side and two median tubercles on retrolateral side; lamella elongated, gently curved, with parallel margins, terminally rounded; embolus large, sickle - shaped,with transverse c arina (fig. 13f). Epigyne (Figs 13 i - k): With flattened and S - shaped scape, anterior part rectangular, distal part much wider with strong lateral lobes protruding from the epigynal cavity and with small stretcher.

Material examined MOROCCO: Tanger - Tétouan: 10 km E. Chefchaouen, 500m, 1 ♀, stones in Quercus suber forest, 15.V.1985, R. Bosmans leg. (CRB). SPAIN: Alicante: Sierra de Aitana, 1 ♂, pitfalls in Pinus forest, 19.IV.1997, M. Perez leg. (CRB). Baleares: Ibiza: Sa Talaia, 200m, 1 ♀, pitfall in Pinus forest, 8 - 18.IV.2018, R. Bosmans leg. (CRB). Malaga: Sierra de Mijas, 1 ♀, low and open dry woods, IV.1095, P. Hillyard leg. (BM). CYPRUS: Paphos: Akamas Peninsula Agios Minas, 4 ♀♀, in dense Pinus litter, 2.V.2007 (CAR - S, C RB, CRS); Smigies, W. Neo Chorio, 300m, 1 ♀, litter and stones in Pinus forest, 30.IV.2010 (CRB).

Distribution Theonina laguncula was until now only known from its type locality in Algeria but appears to have a large distribution area in the Mediter ranean, from Morocco and Spain in the West to Cyprus in the East.

Family Lycosidae Pardosa roscai (Roewer, 1951) (Figs 14 a - i)

Lycosa maculata Rosca, 1939 : 95, figs 7 - 9 (descr. ♂♀; preoc cupied by Hahn, 1822). Lycosa roscai Roewer, 1951 : 438 (replacement name). Pardosa bulgarica Buchar, 1968 : 124, figs 4D - G (synonymy). Pardosa crib rata roscai ; Fuhn & Niculescu - Burlacu, 1971 : 90, figs 37a - e (transfer from Lycosa , reduced to subspecies). Pardosa roscai ; Bayram, Efil & Deltshev, 2009 : 465, figs 1 - 5 (elevated to species).

Comments Pardosa roscai belongs to the proxima group of Pardosa , a group of species that are not easy to identify. It is closely related to Pardosa cribrata Simon and P. hortensis (Thorell). Males can be r ecognised by the relatively short cymbium, apically densely covered with black setae. Females are recognised by the relatively wide anterior cavity of the epigyne. See also Bayram et al. (2009).

Distribution The species is common in Bulgaria (Buchar 1 968), Romania (Fedoriak et al. 2016), Turkey (Bayram et al. 2009) and Iran ( Zamani & Mozaffarian (2017) . Fedoriak et al. ( op. cit. ) mention also material examined from Italy, but this is not retained in the WSC (2019).

Records See part V, page 88 .

Spiders of Cyprus 25

Figs 14 a - i. Pardosa roscai (Roewer). a. Male, dorsal view; b. Female, dorsal view; c. Idem, ventral view; d. Male palp, retrolateral view; e. Idem, ventral view; f. Idem, prolateral view; g. Epigyne, ventral view; h. Vulva, ventral view; i. Id em, dorsal view.

Family Nemesiidae

Brachythele incerta Ausserer, 1871 (Figs 15 a - d) Brachythele incerta Ausserer, 1871: 175, descr. ♀.

Spiders of Cyprus 26

Fig 15 a - d. Brachythele incerta Ausserer a. Labelled vial containing the type specimen; b. Chelicerae showing presence of strong distal bristles and absence of rastellar teeth; c. maxillae with spicules in groups; d. Strongly damaged abdomen missing all diagnostic characters.

Type material Holotype ♀ from Cyprus, without further locality , labelled “Arach. 62, Könshing leg. 1862”; NHMW, examined by Arthur Decae. The specimen was found to be in very poor condition. Absence of strong rastellar teeth (15b, arrow) and presence of spicules grouped on maxillae (15c, arrow) indicate the genus lev el identity is correct.

Comments Although badly damaged, the morphology of the preserved specimen clearly indicate that both the family level (Nemesiidae) and the genus level ( Brachythele ) are correct. As such, it remains the only record of Brachyt hele from Cyprus.

Records See part V, pag 91 .

Family Phyxelididae

Phyxelida anatolica Griswold, 1990 (Figs 16 a - j) Diagnosis See Griswold, 1990.

Spiders of Cyprus 27

Figs 16 a - i. Phyxelida anatolica Griswold. a. Male, dorsal view; b. Fe male, dorsal view; c. Idem, ventral view; d. Male palp, retrolateral view; e. Idem, ventral view; f. Idem, prolateral view; g. Male palpal tibia, dorsal view; h. Male metarsus with clasping spine; i. Epigyne, ventral view; j. Vulva, ventral view.

Spiders of Cyprus 28

Diagno sis See Griswold, 1990.

Comments This family with only one genus and one species in Europe occurs also in Cyprus. Identification of this species for European arachnologists not familiar with the family is not easy, and therefore photos of the spe cies are provided. The disposition of the eyes is very typical (fig. 16b), the metatarsi have a typical spine (fig. 16h) and male palps have very peculiar tibial apophyses (fig. 16d - e), females on the contrary have no special characters in the epigyne.

Records See part V, page 101 .

Family Tetragnathidae

Tetragnatha demissa L. Koch, 1872 (Figs 17 a - v ) Diagnosis Males of this species are recognised by the sligh t S - shape of the conductor when viewed ventrally, females by the epigyne w hich is wider than long, and by the bilobed spermathecae.

Comments In April 2010, males and females of an unknown Tetragnatha species were collected in the marshy vegetation at the border of Asprokremmos Dam . They were collected again in April 2011 at the same site and in Meneou in a salt marsh and neighbouring fields. None of the European species appeared to correspond with the species. Earlier descriptions of Tetragnatha demissa (e. g. Saaris to, 2010 ) correspond well with the species and it is cited here as such.

Records See part V, page 121 .

Spiders of Cyprus 29

Figs 17 a - v. Tetragnatha demissa L. Koch. a. Male, dorsal view; b. Left chelicera of male, anterior view; c. Idem, interior vie w; d. Idem, posterior view; e. Idem, exterior view; f. Male palp, retrolateral view; g. Idem, ventral view; h. Idem, prolateral view; i - n. Embolus and conductor, different views; o. Female, dorsal view; p. Eye region of female, dorsal view; q. Left chelice ra of female, anterior view; r. Idem, interior view; s. Idem, posterior view; t. Idem, exterior view; u. Epigyne, ventral view; v. Vulva, ventral view.

Spiders of Cyprus 30

Family Theridiidae

Dipoena galilaea Levy & Amitai, 1981 (Figs 18 a - g)

Dipoena galilaea Levy & Amitai 1981: 186, figs 37 - 42 (descr. ♀).

Diagnosis The male of Dipoena galilaea can be easily recognised by the abdominal pattern (figs 18 a - c) and the large, elongated palp (figs 18d - g).

7

Figs 18 a - g. Dipoena galilaea Levy & Amitai, 1981. a. Male, dorsal view; b. Idem, late ral view; c. Female, dorsal view; d. Male palp, retrolateral view; e. Idem, ventral view; f. Idem, prolateral view; g. Detail, ventral view.

Comments The female of Dipoena galilaea was described from Israel by Levy & Amitai (1981). Females were furt her recorded in Greece on three occasions: Russell - Smith et al (2011) in Chios, Bosmans et al. in Lesbos (2009) and Van Keer et al . (2010) in Macedonia. The male has not been described previously and resembles most to the one of Dipoena. punctisparsa Yagin uma, 1967 from Korea and Japan.

Records See part V, page 125 .

Spiders of Cyprus 31

Steatoda trianguloides Levy, 1991 (Figs 19 a - e) Steatoda trianguloides Levy, 1991: 228, figs 3 - 6.

Figs 19 a - e. Steatoda trianguloides Levy, 1991. a. Fem ale, dorsal view; b. Idem, ventral view; c. Epigyne, ventral view; d. Vulva, ventral view; e. Idem, dorsal view. ell Comments Levy (1991) described a new male theridiid species from Israel, naming it Steatoda trianguloides , because of its close relat ions to Steatoda triangularis . The colour and the densely punctated prosoma were very alike. It was collected at 1700m on the rocky slopes of Mt Hermon in Israel. Later, the same species was found in Corsica by Lissner (2016), but no details of the habitat were given. A female collected on the top of Mount Olympos is here tentatively described as its mate.

Records: See part V, page 138 .

Spiders of Cyprus 32

Family Zodariidae Zodarion barbarae Bosmans, 2009 (Figs 20 a - g) Zodarion barbarae Bosmans, 2009: 248, figs 80 - 83, 138 - 139.

Figs 20 a - g . Zodarion barbarae Bosmans a. Male, dorsal view; b. Male palp, retrolateral view; c. Idem, ventral view; d. Idem, detail; e. Epigyne, ventral view; f. Vulva, ventral view; g. Vulva, dorsal view.

This species w as so far only known from the Taigetos and Parnonas Mountains in the Greek Peloponnisos (Bosmans, 2009). The specimens from Cyprus are slightly larger than the Greek specimens, but the palp and epigyne are identical.

Description Measu rements: Male (n=9 ): Total length 2.2 - 2.8; prosoma 1.08 - 1.37 long, 0.82 - 0.99 wide. Female (n = 3): Total length 3.1 - 4.2; prosoma 1.51 - 1.61 long, 1.11 - 1.20 wide. Male palp and epigyne: See Bosmans (2009) and figs 20 a - g.

Records See part V, page 1 47 .

Spiders of Cyprus 33

V. List of species and records The following is a checklist of all spider species currently known from Cyprus. Earlier published records and new observations are grouped separately. For each species, data are given by the 6 districts of Cyprus: Famagusta (Ammoc hostos) , Keryneia , Larnaca, Nicosia (Lefkosia), Limassol and Paphos (Fig. 21).

Fig. 21. Districts of Cyprus.

Habitats and microhabitats of the localities are briefly described, and dates, collectors, sexes and number of individuals ar e given. We also provide references to where illustrations can be found. If species are figured in European Spiders ( N entwig et al. 2019, http://www.araneae.unibe.ch), these references are given first as they are available on the internet. Finally, the kno wn distribution and general comments are provided.

Spiders of Cyprus 34

Family Agelenidae Citations Without precise locality (Strand, 1907). Agelena orientalis C. L. Koch, 1837 New records Identification : https://araneae.nmbe.ch/ . LARNACA: Achna Reservoir, 50m, 1 ♀, under stones and litter bordering lake, 16.V.2011 Citations (CRB); Meneou, 1 ♀, in fields, 8.IV. - 15.IX.2016 None. (CMH). LIMASSOL: Alassa SE, 1 ♀, near river Limnatis, New records under stones at base of limestone rock face of FAMAG USTA: Cape Greco, Ayia Napa, 3 ♂♂ 1 gorge, 5.V.2007 (CAR - S); Amathus, 1 ♀, under ♀, 35m, under stones in phrygana, 27.IV.2010 stones in dry valley, 6.IV.1986 (NHMW); Fasoula, 2 ♀♀, olive garden with dry st one walls, (CRB); Prastio S., Agios Andronikos, 30m, 1 ♂, 16.IV.1986 (NHMW); Kolossi, N of the castle, 2 on herbs in Eucalyptus forest, 27.V.2011 (CRB); ♀♀, under stones, 11.IV.1986, 1 ♀, outer wall of Vokolida S., Cape Elaia, 10m, 1 ♀, on shrubs near the castle, 7.IV.1986 (NHMW); road Pano the beach, 26.V.2011 (CRB). Platres - Troodos, 2 ♀♀, under stones in Pinus LARNAC A : Achna Reservoir, 50m, 1 ♀, on nigra forest, 1.V.2007 (CAR - S); Pissouri Beach, herbs bordering lake, 27.V.2011 (CRB); Kornos 1 ♀, 27.IV.1982, coll. A. Stubbs (MMUE); Pissouri, 1 ♀, under stone, 24.III.1998, coll. J. Picnic site, 350m, 1 ♀, on shrubs in Pinus forest, Murphy (MMUE); Pano Platres, track near 28.V.2011 (CRB). Caledonia Waterfall, 1 ♀, under stones in rocky NICOSIA: Agridi, near Dali, 1 ♂, in grass in valley, 26.IV.2007 (CAR - S). house yard, VII.2009 (CDM). NICOSIA: Agios Sozomenos , near Dali, 1 ♂, in dry grassland , 30. IV.2007 (CAR - S), 1 ♀ , Distribution: Italy to Centra l Asia. New to 29.IV.2013 (CAR - S); idem, 1 ♀, in house, Cyprus. 1.V.2012 (CDM); idem, 1 ♀, under stones in pine plantation, 2.V.2013 (CAR - S); Agridi, near Dali, Benoitia lepida (O. P - Cambridge, 1876) 1 ♂ 2 ♀♀, in garden, IV. 2011; idem, 1 ♀, in Agelena lepida ; De Blauwe, 1980b: 19. house, 1.V.2012 (CDM); Athalassa National Forest Pa rk, Nicosia, 1 ♀, by stream, 13.IV.2008 Identification (CDM), 1 ♀, under stones in pine plantation, Levy, 1996: 94, figs 36 - 39. 2.V.2013 (CAR - S); Athalassa, 4 ♀♀, without date, G. P. Georghiou leg. (BMNH); Dali, Citations Lymbourena Hill, in field, V.2004 (CDM), 1 ♀ NICOSIA: Kythrea (De Blauwe, 1980b). under stones in riverbed, 12.V.2007 (CAR - S ); Kapedes, 4 ♀♀, pitfalls in Pinus brutia forest, New Records 17.I. - 23.IV.2016 (CMH); Lakatamia, 1 ♀, in None. olive grove, 1.I. - 1.X. 2016 (CMH); Oikos, 700m, 1 ♀, limey slope, 23.IV.2017, S. Huber leg. Comments (CSH); Pallouriotissa, Nicosia, 2 ♂♂ 1 ♀, in Levy (1996) considers the citation of De Blauwe garden, V.2012 (CDM). (1980b) as mistaken, but the material was re - PAPHOS : Avakas gorge and surroundings, 3 ♀♀, examined by us (in KBIN) and was correctly 22.III.2018 (CKHK); Evretou Reservoir, 350m, 1 identified. ♀, under stones in grassland around lake,

Distribution: North Africa, Israel, Cyprus, 1.V.2010 (CRB); Konia, 1 ♂, on herbs, Yemen. 20.IV.2016 (CMH); Neo Chorio, 1 ♀, under Lycosoides coarctata (Dufour, 1831) stone, 17.III.1998, coll. J. Murphy (MMUE ); Textrix coarctat a ; Strand, 1907: 397. Kannaviou, 1 ♀, under stone, 18.XII.1985, coll. J. Murphy (MMUE); Kato Akourdaleia, 1 ♀, in Identification garden, VIII.2005 (CDM). Levy, 1996: 107, figs 96 - 100.

Spiders of Cyprus 35

Distribution: Mediterranean. Distribution: Lebanon, Israel, Turkey, and new to Cyprus. Tegenaria dalmatica Kulczyński, 1906 Tegenaria zinzulensis ; Brignoli, 1976: 568. Tegenaria pagana C. L. Koch, 1840

Identific ation Identification : https://araneae.nmbe.ch/ . Levy, 1996: 104, figs 65 - 69. Citations Citations No ne. KERYNEIA : Keryneia (Brignoli, 1976). New records New records NICOSIA: Dali, 13, Agios Andronikos, 1 ♀, in PAPHOS: Avakas Gorge, 2 ♂♂ 1 ♀, hand catch, garden, IV.2007, det. A. Bolzern (CDM). XI.1996 (CJB). Distribution: Europe to Central Asia, USA to Distribution: Central and Eastern Mediterranean. Chile, New Zealand.

Tegenaria dentifera Kulc zyński, 1908 Tegenaria parietina (Fourcroy, 1785) Tegenaria dentifera Kulczyński, 1908: 78 Identification Identification Bolzern et al. 2013: 806, figs 1E, F, 21N – R. Kulczyński, 1908: 78, pl. 2, fig. 18. Citations Citation None. Without precise locality (Kulczyński, 1908). New records New records NICOSIA: Pedoulas, 1750m, 1 ♀, in web against Distribution: Cyprus. wall, 19.V.2010 (CRB).

Tegenaria domestica (Clerck, 1757) Distribution: Mediterranean to Central Asia. New to Cypru s. Identification : https://araneae.nmbe.ch/ . Tegenaria sp. Citation Without precise locality (Kulczyński, 1908). A number of Tegenaria specimens could not yet be i dentified to species level and will be treated in a further paper. New records NICOSIA: Kakopetria SE., Platania Camping site, Textrix caudata L. Koch, 1872 1 ♀, pitfalls in Pinus brutia fore st, 6.IV. - Textrix caudata ; De Blauwe, 1980a: 12. 8.VI.2014, M. Hadjiconstantis leg. (CMH). Identification : https://araneae.nmbe.ch/ . Distribution: Cosmopolitan. Citations Tegenaria maronita Simon, 1873 Wit hout precise locality (De Blauwe, 1980a).

Identification New records Levy, 1996: 101, figs 59 - 64. None.

Citations Distribution: Mediterranean, expanding to None. Central Europe.

New records NICOSIA: Cedar Valley Forest, 1 ♂, under Family Anyphaenidae s tones, 29.IV.2007, det. A. Bolzern. (CAR - S). Anyphaena sabina L. Koch, 1866

Spiders of Cyprus 36

Identification PAPHOS: Akamas Peninsula, Avakas Gorge, 1 ♀, Simon 1932: 895, 960, figs 1345 - 1346; Urones, under bushe s, 27.III.1998, coll. J. Murph y Barrientos & Espuny, 1995: 123, figs 18 - 22. (MMUE); Drouseia, 539m, 1 ♀, in p hrygana, 23.III.2018 (CKHK); Evretou Reservoir, Stavros Citations tis Psokas river, 2 ♀♀, 20.III.2018 (CKHK); None. Kelefos bridge, dam on Diarizos River, 1 ♀, 550m, beaten from shrubs, 21.V.2011 (CRB); New records Kour daka, Ezousa river, 1 ♀, 14.III.2018 LIMASSOL: Agios Georgios E., Palaiomylos, (CKHK); Mavrokolympos Reservoir, 1 ♂, 1250m, 5 ♂♂ 2 ♀♀, beaten in mixed Pinus and 13.III.2018 (CKHK); near Kritou Terra, 1 ♀, Quercus forest, 4.V.2010 (CJVK); Pano Platres, swept from grass in dry fallow, 14.IV.2017; 1050m, 1 ♂ 2 ♀♀, beaten near Mesa Po tamos Paphos NW, 1 ♀, under stones at the edge of a Waterfall s, 23.V.2011 (CJVK, CRB); Troodos S., large sand pit, 24.IV.1986 (NHMW). 1350m, 1 ♂ 4 ♀♀, beaten in mixed Quercus - Pinus forest, 3.V.2010 (CJVK, CRB). Distrib ution: Palaearctic. New to Cyprus. NICOSIA: Cedar Valley Forest, 1180m, 1 ♂, under stone by track, 29.IV.2007 (CAR - S), 3 ♀♀, Araneus diadematus Clerck, 1757 beaten in Cedrus forest, 19.V.2011 (CRB); Kakopetria S., Platania forest station, 1100m, 1 ♀, Identification : https://araneae.nmbe.ch/ . beaten in dense Pinus forest, 4.V.2010 (CHVK); road Pedoulas - Lemithou, 1 ♀, pitfalls in Pinus Citations nigra and Pinus brutia forest, 6.IV. - 8.VI.2014 None. (CMH); Prodromos S., Trooditissa picnic site, 1450m, 2 ♂♂, under stones in Pinus forest, New records 21.V.2011 (CRB). LARNACA: Ora, 1 ♂, on its web in maquis, 28.XII. 2016 (CMH). Distribution: Southern Europe, Russia, Georgia, LIMASSOL: Pelendri, 4 ♀♀, on bushes in Pinus Azerbaijan. New to Cyprus. brutia forest, 11. XII. 2018 (CMH). NICOSIA: Klirou, 1 ♀, on its web in house Family Araneidae garden, 5. VI. 2018 (CMH). PAPHOS : Stroumpi, 1 ♂ 2 ♀♀, on its web in Aculepeira armida (Audouin, 1826) house garden, 16. VI. 2018 (CMH).

Identification Distri bution: Holarctic. First confirmed record in Levy, 1998a: 329, figs 68 - 79. Cyprus.

Citations . Araneus grossus (C. L. Koch, 1844) None. Identification : https://araneae.nmbe.ch/ . New records FAMAGUSTA: Frenaros, 1 ♂, 26.IV.1957, S. Citations Pieris leg, (BMNH); Liopetri, 1 ♀, 9.IV.1957, D. None. Pieris leg. (BMNH). New records Distribution: Palaearctic. New to Cyprus. KERYNEIA : road Chalefka - S ourp Magar Monastery (Armenomonastiro), 660m, 1 ♀ in Agalenatea redii (Scopoli, 1763) open Pinus forest, 23.XI.2010 (CRB); Klepini, 300m, 2 ♀♀ in open Pinus forest, 23.XI.2010 Identificatio n : https://araneae.nmbe.ch/ . (CRB). PAPHOS: Vretsia W., near Kelefos Bridge, 1 ♀, Citations . 650m, beaten from shrubs, 19.V.2011 (CRB). None. Distributio n: Europe to Central Asia. New to New records Cyprus. LIMASSOL: Agios Tychonas, 1 ♀, under stones near house walls, 19.IV.1986 (NHMW).

Spiders of Cyprus 37

Kapedes S., 600 m, 1 juvenile, herbs along rivulet, Araniella alpica (L. Koch, 1869) 5.V.2010 (CRB). PAPHOS: Akamas Peninsula, Cape Geronnisos Identification : https://araneae.nmbe.ch/ . S., 1 juvenile, under stones in dry phrygana, 3.V.2007 (CAR - S); Akamas Peninsula, Lara Bay, Citations 1 ♀, C. Saure leg., 1 9.VI.2013 (CKHK); None. Asprokremmos Dam, N. Kouklia, 150m, New records juveniles, herbs along lake, 29.IV.2010 (CRB), LIMASSOL: Pano Platres, Caledonia Waterfall , 1 and 1 subadult ♂, 23.V.2011 (CRB); Kampos NE, ♂, swept from herbs and shrubs in rocky stream Xeros River, 550m, 2 ♀♀, beaten from shrubs, valley, 26.IV.2007 (CAR - S). 20.V.2011 (CRB).

PAPHOS : Vretsia W., near Kelefos bridge, 650m, Distribution: Old world. New to Cyprus. 1 ♀, beaten from shrubs, 19.V.2011 (CRB). Argiope trifasciata (Forsskål, 1775) Distribution: Europe to Azerbaijan. New to Cyprus. Identification : https://araneae.nmbe.ch/ .

Araniella inconspicua (Si mon, 1874) Citations None. Identification : https://araneae.nmbe.ch/ . New records Citations FAMAGUSTA: Prastio S ., Agios Andronikos, None. 30m, 1 ♀, under stones and herbs in Eucal yptus forest, 27.V.2011 (CRB) New records LIMASSOL: Akrotiri Peninsula, 1 ♂ 1 ♀, margin NICOSIA: Cedar Valley, 1120m, 1 ♀, Cedrus of central salt lake, 7.V.1972, coll. A. Stubbs forest, 19.V.2011 (CKHK). (MMUE); Akrotiri Peninsula, Fasouri reedbeds, 1 ♀, swept from grass and reeds, 17.IV.2017 (CAR - PAPHOS: Koudraka, Ezousa river, 1 ♂, S). 14.I II.2018 (CKHK). NICOSIA: Agios Sozomenos , n ear Dali, 1 ♀, in dry grassland, IV.2011 (CDM) ; A gridi, near Dali, Distribution: Europe to China. New to Cyprus. 1 ♂, in field, IV.2007 (CDM), 1 ♂, on wall inside house, XI.2010 (CDM). Argiope lobata (Pallas, 1772) Distribution: Cosmopolitan. New to Cyprus. Identification : https://araneae.nmbe.ch/ . Cyclosa groppalii Pesarini, 1998 Citations None. Identification

New records Citat ions : FAMAGUSTA: Flamoudi W., 25m, 2 ♂♂ 2 ♀♀ 1 None. subadult ♀, phrygana near the beach, 25.V.2011 (CJVK, CRB); Karpasia Peninsula, Chelones, New records juveniles, herbs in salt marsh, 16.V.2011 (CRB); LIMASSOL: Limassol, A mathus hotel, 1 ♀, front Vokolida S ., Cape Elaia, 10m, 5 ♂♂ 1 subadult yard, 8.IV.1986 (NHMW). ♀, on bushes near t he beach, 26.V.2011 (CJVK, PAPHOS: near Avakas Gorge, 1 ♂, beaten from CRB) . junipers on rocky hillside, 14.IV.2017 (CAR - S). LARNACA: Kornos picnic site, 350m, 1 subadult ♀, on branches in Pinus forest, 28.V.2011 (CRB); Distribution: Spain, Balearic Is., Crete. New to Larnaca SW, Hala Sultan Tekke Mosque, 5m, Cypr us juveniles, on herbs along salt lake, 16.V.2011 (CRB). Cyclosa sierrae Simon, 1870 NICOSIA: Agridi, near Dali, 1 ♂, on house wall, Cyclosa sierrae ; Logunov, 2013: 5. VII.2009; i n garden, 1 ♂, VIII.2012 (CDM);

Spiders of Cyprus 38

Identification plantation, 23.III.2018 (CKHK); Smigies, W. Neo Levy, 1998a: 318, figs 24 - 32. Chorio, 300m, 4 ♀♀, bushes in Pinus forest, 30.IV.2010 (CJVK, CRB). Citations : PAPHOS : Polis, Akamas forest (Logunov, 2013). Distribution: Europe, Caucasus.

New records Cyrtarachne ixoides (Simon, 1870) FAMAGUSTA: Kantara E., 610m, 1 ♀, beaten from s hrubs and stones in open Pinus forest, Identi fication 25.V.2011 (CJVK); Salamis ruins, 50m, 1 ♀, Levy, 1998a: 314, figs 1 - 11. beating in ruins, 15.V.2011 (CRB). KERYNEIA: Keryneia , near the town, 1 ♀, Citations Juniperus phoenicea forest, 30.IV.2016 (CMH); None. Pentadactylos R ange , Saint Hilarion Castle W, 600m, 4 ♀♀, beating in Pinus forest 17.V.2011 New records (CJVK, CRB); Vouni ruins, 250m, 1 subadult ♂, LIMASSOL: Akrotiri Peninsula, 1 ♀, margin of on herbs in ruins, 20.III.2011 (CRB). central salt lake, 7.V.1972, coll. A. Stubbs LARNACA: Agia Anna, 1 ♀, in gulley with (MMUE). pines, 26.IV.1982, coll. A. Stubbs (MMUE); Delikipos, near Kornos, 2 ♀, in grass in open pine Distribution: Mediterranean to Georgia, wood, 9.V.2013 (CAR - S); Stavrovouni Madagascar. New to Cyprus. Monastery, 1 ♂, in phrygana, 26. IV.1982, coll. A. Stubbs (MMUE). Cyrtophora citricola (Forsskål, 1775) LIMASSOL: Agios Tychonas, 1 ♀, on bushes near house walls, 19.IV.1986 (NHMW); Fasoula, Identification: https://araneae.nmbe.ch/ . 1 ♀, olive grove, 16.IV.1986 (NHMW); Platres, 1 ♀, on walls, 22.IV.1986 (NHMW). Citations NICOSIA: 13, Agios Andronikos, Dali, 1 ♀, in None. garden, IV. - V.2008 (CDM); Agridi near Dali, 1 ♀, in garden, IV.2004 (CDM); Athalassa National New records Forest Park, 1 ♀, Pinus brutia forest, 21.II.2016 LARNACA: Mari (N 34°44'32.4" E 33°18'19.8"), (CMH); Oikos, Xerargaka picnic site, 350m, 1 ♂ 2 ♂♂, 5 ♀♀, on Pistacia lentiscus bush , on their 2 ♀♀, beaten from shrubs in Pinus forest, web. 25. XII. 2018, R. D. Symeou leg. (CMH). 20.V.2011 (CJVK, CRB); Kapedes S., 600m, 1 ♀, herbs and stones along rivulet, 5.V.2010 (CRB); Distribution: Limnitis River, Kato Pyrgos SE, 200m, 1 ♂, in Europe, Turkey, New to Cyprus. Pinus forest, 2.V.2010 (CRB). PAPHOS : Akamas Peninsula, Cape Geronnisos Gibbaranea bituberculata (Walckenaer, 1802) S., 1 ♀, under stones in dry phrygana, 3.V.2007 (CAR - S), 1 ♂, (CRS); Akamas Peninsula, Kioni, Identification : https://araneae.nmbe.ch/ . 10m, 2 ♀♀, in Juniperus phrygana, 30.IV.2010 (CJVK, CRB); Avakas gorge, 50m, 2 ♀♀, herbs Citations in citrus orchard at entrance of gorge, 30.IV.2010 None. (CRB); near Avakas Go rge; 1 ♂, beaten from junipers on rocky hillside, 14.IV.2017 (CAR - S) ; 9 New records km N. Pano Panagia, 1050m, 1 ♂, in litter near FAMAGUSTA : Cape Apostolos Andreas, 1 ♂, spring, 19.V.2011 (CJVK); Gialia, 1 ♂ 1 ♀, on Juniperus phoenicea forest 30.III.2016 (CMH); bush in pines, 25.III.1998, coll. J. Murphy Gialousa E ., Agios Thyrsos, 20m, 1 ♀ , beaten (MMUE); Kinousa SE, 400m, 4 ♀♀, on low from shrubs, 16.V.2011 (CJVK). bushes i n Pinus forest, 1.V.2010 (CJVK, CRB); LARNACA: Agioi Vavatsinias, 700m, 1 ♀, litter Neo Chorio, 3 ♂♂ 3 ♀♀, swept from phrygrana in Pinus forest, 28.IV.2010 (CJVK); Pergamos, 1 and grass, 2.V.2007 (CAR - S); same site, 1 ♂ ♀, 28.II.1957, S. Pieris leg. (BM NH). (CRS); idem, 1 ♀, beaten from shrubs in gorge, LIMASSOL: Pano Platres, 1050m, 1 ♀, by 22.V.2011 (CRB); Pegeia, Pikni Forest, 366m, 2 woodland stream, 4.V.1982, coll. A. Stubbs ♀♀, in pine forest, 11.III.2018 (CKH K); Pegeia, (MMUE). Pikn i Forest, 346m, 1 ♀, in former c arob

Spiders of Cyprus 39

NIKOSIA: Cedar Valley, 1150m, 1 ♂, 5.V.1982, coll. A. Stubbs (MMUE); Neo Chorio, 21.III.2018, forest road, beaten from shrubs 1 ♀, in grass, 30.III.1998, coll. J. Murphy (CKHK). (MMUE); PAPHOS: Pegeia, Pikn i Forest, 346m, 1 ♀, in former c arob plantation , 11.III.2018, same site, 1 Distribution: Europe, North Africa, Russia, ♀, 23.III.2018 (CKHK). Ukraine. New to Cyprus.

Distribution: Palearctic. New to Cyprus. Hypsosinga pyg maea (Sundevall, 1831)

Gibbaranea gibbosa (Walckenaer, 1802) Identification : https://araneae.nmbe.ch/ .

Identification : https://araneae.nmbe.ch/ . Citations None. Citations None. New records LIMASSOL: Akrotiri Peninsula, 1 ♂ 3 ♀♀, New records margin of central salt lake, 7.V.1972, coll. A. FAMAGUSTA : Cape Greco, 1 ♂, under stones in Stubbs (MMU E). phrygana, 3.V.2013 (CDM); Kantara E., 610m, 1 ♀, beaten from shrubs in open Pinus forest, Distribution : Holarctic. New to Cyprus. 25.V.2011 (CRB). Larinia chloris (Audouin, 1826) Distribution: Europe to Azerbaijan. New to Cyprus. Identification Levy, 1986 : 5, figs 18 - 27. Glyptogona sextuberculata (Keyserling, 1863) Citations None. Identification Levy, 1998a : 315, figs 12 - 23. New records LARNACA : Achna Reservo ir, 50m, 1 ♂ 1 ♀, Citations under stones and litter bordering lake, 27.V.2011 None. (CJVK); Oroklini, 4 ♀♀, Phragmites bordering lake, 3.VIII.2016 (CMH). New records FAMAGUSTA : Kantara E., 610m, 1 ♂, stones in Distribution: Europe, North Africa, Russia, open Pinus forest, 25.V.2011 (CJVK). Ukraine. New to Cyprus. LA RNACA : Achna Reservoir, 50m, 1 ♀, stones and litter bordering lake, 16.V.2011 (CJVK). Larinioides suspicax (O. Pickard - Cambrid ge, 1876) Distribution: Italy to Israel. New to Cyprus. Identification : https://araneae.nmbe.ch/ . Hypsosinga albovittata (Westring, 1851) Citations Identification : https://a raneae.nmbe.ch/ . None.

Citations New records None. LIMASSOL: Akrotiri Peninsula, Fasouri reed beds, 1 ♂ 3 ♀♀, on their web on Juncus sp. , 29. New records IV. 2018, C. Makris leg. (CMH). PAPHOS : Evretou Reservoir, 350m, 2 ♀♀, grassland around lake, 1.V.2010 (CJVK, CRB); Distribution: Mediterranean to Central Asia. Evretou Reservoir, Stavros tis Psokas river, 1 ♂, 20.III.2018 (CKHK); Kelefos bridge, dam on Leviellus kochi (Thorell, 1870) Diarizos River, 550 m, beaten from shrubs, 21.V.2011 (CJVK); Mandria, 1 ♂, in scrub,

Spiders of Cyprus 40

Identification LIMASSOL: Agios Tychonas, 1 ♀, under stones, Levy, 1974a : 283, figs 85 - 91. 19.IV.1986 (NHMW); Akrotiri Peninsula, 1 ♀, margin of central salt lake, 7.V.1972, coll. A. Citations Stubbs (MMUE), 1 ♀, 2.IV.1997 (CJB); Akrotiri None. Peninsula N., 5m, 1 ♂ 10 ♀♀, litter and beating in Euc alyptus forest, 29.IV.2010 (CJVK, CRB); N ew records Fasoula, 1 ♀, in olive grove, 16.IV.1986 LARNACA : Dip otamos Reservoir, 150m, 1 ♀, (NHMW); Kourion, Apollo Hylates Temple, 1 ♀, herbs and stones bordering lake, 28.IV.2010 under stones in ruins, 23.IV.1986 (NHMW); Pano (CJVK). Platres, 1050m, 3 ♂♂ 3 ♀♀, on bushes near Mesa PAPHOS : Smigies, W. Neo Chorio, 300m, 1 ♀, Potamos Waterfall s, 23.V.2011 (CJVK, CRB); litter and stones in Pinus forest, 30.IV.2010 Pano Platres, track near Caledonia Waterfall, 1 ♀, (CJVK). swept from herbs and shrubs, 26.IV.2007 (CAR - S); Pano Platres, near road junction to Troodos, 1 Distribution: Europe, North Africa, Russia, ♂, 28.IV.2007 (CAR - S); Pano Platres, near top of Ukraine. New to Cyprus. road to Millomeris Waterfalls, 5 ♂♂ 1 ♀, swept Lipocrea epeiroides (O. Pickard - Cambridge, from gr ass and herbs in abandoned vineyard, 1872) 28.IV.2007 (CRS); Pissouri Beach, 1 ♂ 1 ♀, 27.IV.1982, coll. A. Stubbs (MMUE); Platres, 1 Identification ♀, on walls, 22.IV.1986 (NHMW); Troodos S., Levy, 1986: 7, figs 28 - 39. 1350m, 1 ♂, under stones in mixed Quercus - Pinus forest, 3.V.2010 (CRB); Troodos, Moun t Citations : Olympos (Chionistra), 1900m, 1 ♂, stones in PAPHOS : Polis (Wunderlich, 2011). Pinus forest, 23.V.2011 (CJVK). PAPHOS : Akamas Peninsula, 1 ♂, in coastal New records grassland, 22.III.1998, coll. J. Murphy (MMUE); NICOSIA: Agridi near Dali, 1 ♂, in grass field, Akamas Peninsula, Mandria, 4 ♂♂ 1 ♀, in fields, D. McCowan leg. (CDM). 6.V.1982, coll. A. Stubbs (MMUE); Aspr okremmos Dam, N. Kouklia, 150m, ♀♀, Dist ribution: Italy (Sardinia, Sicily), Greece, herbs and stones along lake, 23.V.2011 (CJVK); Cyprus, Turkey, Israel, Yemen, India . Evretou Reservoir, 350m, 1 ♂ 6 ♀♀, shrubs and grassland around lake, 1.V.2010 (CJVK, CRB); Mangora acalypha (Walckenaer, 1802) Evretou Reservoir, 1 ♀, under stones and in grass Mangora acalypha ; Logunov, 2013: 5 tussocks, 13.IV.2017 (CAR - S); Ke lefos bridge, dam on Diarizos River, 550m, 7 ♀♀, under stones Identification : https://araneae.nmbe.ch/ . and beaten from shrubs, 21.V.2011 (CJVK, CRB) ; Gialia, 3 ♂♂, on bush in pine forest , Citations 25.III.1998, coll. J. Murphy (MMUE); Kato LIMASSOL: Pano Platres (Logunov, 2013). Akourdaleia, 2 ♀♀, in garden, VIII.2005 (CDM); PAPHOS : Polis W., Akamas forest (Logunov, Kato Paphos, 1 ♂, un der stones near hotel, 2013); Avakas (Logunov, 2013). 4.IV.1997 (CJB); Kelefos bridge, dam on Diarizos River, 550m, 7 ♀♀, under stones and beaten from New records shrubs, 21.V.2011 (CJVK, CRB); near Kritou FAMAGUSTA : Agios Sergios, 30m, 1 ♀, beating Terra, 2 ♀♀, swept from grass in dry fallow, in Pinus plantation, 15.V.2011, CRB); Gialousa 14.IV.2017; Mavrokolympos Reservoir, 1 ♀, E., Agios Thyrsos, 20m, 1 ♀, beaten from shrubs, 13.III .2018 (CKHK); Pegeia, Pikn i Forest, 346m, 16.V.2011 (CRB); Gialousa E., Agia Triada, 50m, 1 ♀, in former c arob plantation, 11.III.2018 1 ♀, under stones in ruins, 18.III.2011 (CRB). (CKHK); Sm igies, W. Neo Chorio, 300m, 1 ♀, KERYNEIA : Pentadactylos Ra nge, Saint Hilarion beating in Pinus forest, 30.IV.2010 (CRB), 3 ♂♂ Castle W., 600m, 1 ♂ 1 ♀, beaten from shrubs in 5 ♀♀, swept from phrygrana and gras s, 2.V.2007 Pinus forest, 17.V.2011 (CJVK). (CAR - S), 1 ♂, (CRS); Vretsia W. , near Kelefos LARNACA: Agioi Vavatsinias, 700m, 2 ♂♂ 2 bridge, 650m, 2 ♀♀, beaten from shrubs, ♀♀, beating in Pinus forest, 28.IV.2010 (CJVK, 19.V.2011 (CJVK). CRB); Stavrovouni Monastery, 1 ♂, in phrygana, 26.IV.1982, coll. A . Stubbs (MMUE). Distribution: Palaearctic.

Spiders of Cyprus 41

Neosc ona adianta (Walckenaer, 1802) (CAR - S); Masari S., Merika river, 120m, 1 ♂ 1 Neoscona adianta ; Logunov, 2013: 5. ♀, under stones in wasteland, 24.V.2011 (CRB). PAPHOS: Akrotiri Peninsula, 2 ♂♂, 27.IX.2007 Identification : https:// araneae.nmbe.ch/ . (CDM); Akrotiri Peninsula E., 5m, 2 ♂♂ 2 ♀♀, on herbs in salt marsh, 29.IV.2010 (CRB); Citations Akrotir i Peninsula, near British Airbase, 3 ♂♂ 2 LIMASSOL: Akrotiri Salt Lake (Logunov, 2013); ♀♀, swept in pine wood, 4.V.2007 (CAR - S), 1 ♂ Plano Platres (Logunov, 2013). (CRS); Alassa SE, near river Limnatis, 1 ♂, swept PAPHOS : Amargeti (Logunov, 2013). from lush grass, 5.V.2007 (CAR - S), 3 ♂♂ (CRS); 2 km S. of Kouklia, 1 ♂ 1 ♀, in riverine marsh, New records 30.IV.1982, coll. A. St ubbs (MMUE); FAMAGUSTA: Cape Greko, 1 ♀, in Juniperus Asprokremmos Dam, N. Kouklia, 150m, 2 ♂♂ 2 phoenicea forest, 1.V.2016 (CMH); Flamoudi W., ♀♀, herbs along lake, 29.IV.2010 (CJVK, CRB); 25m, 2 ♀♀, phrygana near the beach, 25.V.2011 Evretou Reservoir, 350m, 4 ♂♂ 2 ♀♀, grassland (CRB); Gialousa E., Agios Thyrsos, 20m, 2 ♀♀, around lake, 1.V.2010 (CJVK, CRB) and 2 ♂♂ 1 beaten from shrubs, 16.V.20111 (CJVK); ♀, 23.V.2011 (CRB); Kelefos bridge, dam on Karpasia Peninsula, Apostolos Andreas Diarizos River, 550m , 1 ♂ 1 ♀, under stones and Monastery, 50m, 1 ♂ 2 ♀♀, herbs in wasteland, beaten from shrubs, 21.V.2011 (CRB); Kidasi N., 16.V.2011 (CRB); Prastio S., Agios Andron ikos, along Diarizos River, 300m, 3 ♂♂ 1 ♀; beaten 30m, 1 ♀, herbs in Eucalyptus forest, 27.V.2011 from shrubs, 21.V.2011 (CRB); near Souskiou, 1 (CRB); Salamis ruins, 50m, 2 ♂♂ 4 ♀♀; beating ♀, on bush in dry grassland, 1.V.2013 (CAR - S); in ruins, 15.V.2011 (CJVK, CRB); Vokolida S., Vretsia W., near Kelefos bridge, 650m, 2 ♂♂ 3 Cape Elaia, 10m, 1 ♂ 1 ♀, bushes near the beach, ♀♀, beaten from shrubs, 19.V.2011 (CJVK, 26.V.2011 (CJVK, CRB); CRB). LARNACA : Achna Reservoir, 50m, 1 ♂, on herbs bordering lake, 27.V.2011 (CJVK); Agios Distribution: Palaearctic. Theodoros, 2 ♂♂ 1 ♀, on herbs, 23.IV.2016 (CMH). Neoscona sodom Levy, 1998 LIMASSOL: Agios Tychonas, 4 ♀♀ 3 juveniles, under stones near house walls, 19.IV.1986 Identification (NHMW); Akrotiri Peninsula, 1 ♀, margin of Levy, 1998a: 340, figs 117 - 126. central salt lake, 7.V.1972 , coll. A. Stubbs (MMUE); Akrotiri Peninsula, Fasouri reedbeds, 3 Citations ♂♂, swept from grass and reeds, 17.IV.2017, None. (CAR - S); Amathus, 2 ♂♂ 5 ♀♀ 4 juveniles, on herbs in dry valley, 25.IV.1986, 1 ♀ 5 juveniles, New records 6.IV.1986,1 ♀ 1 juvenile, in the excavation area, FAMAGUSTA: Gialousa E., Agios Thyrsos, 9. IV.1986 , 4 juveniles, on house walls, 9.IV.1986 2 0m, 1 subadult ♀, marshy area near spring near (NHMW); Armenochori, 2 ♀♀ 5 juveniles, on the sea, 18.III.2011 (CRB). bushes, 14.IV.1986 (NHMW); Erimi, 1 juvenile, under stones in dry riverbed, 18.IV.1986 LARNACA: Achna Reservoir, 50m, 1 ♂, on (NHMW); Fasoula, 2 juveniles, olive garden with herbs bordering lake, 16.V.2011 (CRB), and 3 dry stone walls, 16.IV.1986 (NH MW). ♂♂, 27.V.2011 (CJVK, CRB). Parekklisia, 4 juveniles, under stones in dry riverbed, 13.IV.1987 (NHMW). Distribution: Israel. New to Cyprus. NICOSIA: Agios Sozomenos , near Dali, 1 ♂, in dry grassland, 30.IV.2007 (CRS), 5 ♂♂ 4 ♀♀ Neoscona subfusca (C. L. Koch, 1837) (CAR - S); Agridi, near Dali, 1 ♂, in field, V.2007 (CDM); Kato Koutrafas - Astromeritis road, 120m, Identification 1 ♂ 1 ♀, beaten from shrubs in dry riverbed, Levy, 1998a: 335, figs 96 - 107. 18.V.2011 (CJVK); Lefka NW, 120m, 4 ♂♂ 4 ♀♀, beaten from shrubs in fruit plantation, Citations 18.V.2011 (CJVK, CRB); Limnitis, 1 ♀, Pinus None. brutia forest, 13.V.2016 (CMH); Lympia, Lympia Reservoir, 1 ♀, in gra ss tussocks, 9.V.2013 New records

Spiders of Cyprus 42

FAMAGUSTA: Gialousa E., Agios Thyrsos, 20m, Distribution: Old World. New to Cyprus. 7 ♂♂ 7 ♀♀, beaten from shrubs, 16.V.20111 (CJVK, CRB); Gialousa E., Agia Triada, 50m, 2 Singa neta (O. Pickard - Cambridge, 1872) ♂♂ 3 ♀♀, herb s in ruins, 18.III.2011 (CRB); Kantara E., 610m, 1 ♀, beaten from shrubs in open Identification Pinus forest, 25.V.2011 (CRB); Karpasia Levy, 1984: 124, figs 26 - 41. Peninsula, Apostolos Andreas Monastery, 50m, 2 ♀♀, herbs in wasteland, 16.V.2011 (CRB); Komi Citations Kebir, 150m, 2 ♂♂ 1 ♀, beaten from shrubs, None. 26.V.2011 (CJVK); Salamis ruins, 50m, 1 ♀, beaten from low trees, 15.V.2011 (CJVK); New records LARNACA: Larnaca salt lake, 5m, 4 ♂♂ 1 ♀, trees FAMAGUSTA: Frenar os, 1 ♂, 6.IV.1957, S. bordering lake, 27.IV.2010 (CJVK, CRB); Pieris leg. (BMNH); Karpasia Peninsula, Cape Meneou, 1 ♀, salt marsh and fields, 8.IV. - Apostolos Andreas, 70m, 1 ♀, stones in grassland, 15.IX.2016 (CMH); Stavrovouni Monaster y, 16.V.2011 (CJVK). 350m, 1 ♀, litter in Pinus forest, 28.IV.2010 LARNACA: Achna Reservoir, 50m, 2 ♀♀, herbs (CJVK). in marshy area near lake, 16.V.2011 (CRB). LIMASSOL: Akrotiri Peninsula, 1 ♂ 1 ♀, LIMASSOL: Akrotiri Peninsula, edge of central 27.IX.2007 (CDM); Akrotiri Peninsula, near salt pan, in shrubs, 2 ♀♀, 27.IV.2007 (CRS); British Airbase, 1 ♀, swept in pine wood, Alassa SE, near river Limnatis, 1 ♀, in dry 4.V.2007 (CAR - S); Akrotiri Peninsula, Turtle grassland, 4.V.2007 (CRS). Beach, 1 ♀, sandy beach, bea ten from vegetation, NICOSIA: Agios Sozomenos , near Da li , 1 ♀, in 15.III.2018 (CKHK); Erimi, 1 ♀, under stones, dry grassland, 30.IV.2007 (CRS ) . 18.IV.1986 (NHMW); Kolossi , ♀, outer wall of PAPHOS: Asprokremmos Dam, N. Kouklia, the castle, 7.IV.1986 (NHMW); Pissouri Beach, 1 15 0m, 1 ♀, herbs along lake, 29.IV.2010 (CRB); ♂ 1 ♀, 27.IV.1982, coll. A. Stubbs (MMUE); Kidasi N., along Diarizos River, 300m, 1 ♀, Pissouri, 1 ♀, in bushes, 24.III.1998, coll. J. beaten from shrubs, 21.V.2011 (CRB). Murph y (MMUE). NICOSIA: 13, Agios Andronikos, Dali, 1 ♂ 1 ♀, Distribution: Mediterranean to Caucasus. New to in garden, IV.2007 (CDM); Ayia Koroni, Agios Cyprus. Epifanios, 1 ♀, in house, IV.2005 (CDM); Agridi, near Dal i, 2 ♂♂, in garden, 1.V.2010, 1 ♀, IV. Zilla diodia (Walckenaer, 1802) 2011, 1 ♂, in house 30.IV.2010, 1 ♀, in house, X.2010; 2 ♂♂, in house, V.2011; 1 ♂ 1 ♀, in Identification : https://araneae.nmbe.ch/ . house, VII.2012 (all CDM); Geri, 1 ♀, in olive grove, 11.V.2016 (CMH); Kampos NE, Xeros Citations Ri ver, 550m, 2 ♀♀, beaten from shrubs, None. 20.V.2011 (CJVK); Lefka NW, 120m, 2 ♂♂ 7 ♀♀, beaten from shrubs in fruit plantation, New records 18.V.2011 (CJVK, CRB); Pallouriotissa, Nicosia, LARNACA: Agioi Vavatsinias, 700m, 1 ♀, litter 1 ♂ 1 ♀, in garden, 15.VI.2008 (CDM); in Pinus forest, 28.IV.2010 (CJVK). Xerargaka S, Orkontas, 1 ♂, riverside vegetation, LIMASSOL : Agros SW, 950m, 1 ♀, beaten in 26.III.2016 (CMH). Pinus forest, 5.V.2010 (CJVK , CRB); Fasoula, 1 PAPHOS: Asprokremmos Dam, N. Kouklia, ♀, olive garden with dry stone walls, 16.IV.1986 150m, 11 ♂♂ 11 ♀♀, herbs along lake, (NHMW); Pano Platres, 1050m, 2 ♂♂ 3 ♀♀, 23.V.2011 (CJVK, CRB); Avakas gor ge, 50m, 1 beating in forest near Mesa Potamos Waterfall , ♀, in citrus yard at entrance of gorge, 30.IV.2010 23.V.2011 (CJVK, CRB); Pano Platres, below Caledonia Waterfall, 3 ♂♂, swept from herbs and (CRB); Evretou Reservoir, 350m, 1 ♀, on shrubs shrubs in rocky stream valley, 26.IV. and 1.V.2007 around lake, 1.V.2010 (CRB); Kato Akourdaleia, (CAR - S); Pano Platres, Millomeris Waterfall s, 1 1 ♀, in garden, VIII.2005 (CDM); Paphos, 5 ♀♀, ♂, swept from grass and herbs in abandoned in garden, 2.XI.1996 (CJB); Vretsia W., near vineyard, 28.IV.2007 (CAR - S), 1 ♂ (CRS). Kelefos bridge, 650m, 1 ♀, beaten from shrubs, NICOSIA: Agridi, near Dali, 2 ♀♀, in garden, 19.V.2011 (CRB). V.2011 (CDM); Cedar Valley Forest, 1 180m, 1 ♂, swept from herbs, 29.IV.2007 (CAR - S); Cedar Valley Forest, 1180m, 1 ♀, beating in Cedrus

Spiders of Cyprus 43

forest, 19.V.2011 (CRB); Chrysoroyiatissa Distribution: Europe to Central Asia. Monastery, 1 ♂ 2 ♀♀, 2.V.1982, coll. A. Stubbs (MMUE); N. Oikos, Xerargaka picnic site, 350m, Cheiracanthium erraticum (Walckenaer, 1802) 1 subadult ♀, beaten from shrubs in Pinus forest, Identification : https://araneae.nmbe.ch / . 20.V.2011 (CRB). PAPHOS: Agios Georgios E., Palaiomylos, Citations 1250m, 1 ♂, beaten in mixed Pinus and Quercus None. forest, 4.V.2010 (CJVK); Asprokremmos Dam, New records N. Kouklia, 150m, 1 ♀, herbs and stones along LARNACA: Mosfiloti, 1 ♀, under stone in open lake, 23.V.2011 (CJVK); Evretou Reservoir, pine woodland, 12.IV.2017 (CAR - S). 350m, 1 ♀, beating shrubs around lake, 1.V.2010 NICOSIA: Agridi, near Dali, 1 ♂, in yard, V.2009 (CRB); Kelefos bridge, dam on Diarizos River, (CDM). 550m, 3 ♀♀, beaten from shrubs, 21.V.2011 PAPHOS: Evretou Reservoir, 1 ♀, under stone in (CJVK, CRB); Kelefos bridge, 465m, deciduous grass tussocks, 13.IV.2017 (CAR - S). wood, 1 ♀, 16.III.2018 (CKHK); Gialia, 2 ♀♀, on bush in pine fo rest , 25.III.1998, coll. J. Murphy Distribution: Palaearctic. (MMUE); Kinousa, 1 ♂ 1 ♀, in shrubs, 19.III.1998, coll. J. Murphy; s ame site, 1 ♀, Cheiracanthium mildei L. Koch, 1864 28.III.1998 (MMUE); Mandria, 2 ♂♂ 1 ♀, in scrub, 2.V.1982, coll. A. Stubbs (MMUE). Identification : https://araneae.nmbe.ch/ .

Distribution: Western and Mediterranean Europe Citations to Azerbaijan. New to Cyprus. NICOSIA: Trapeza Mountains ( Özkütük et al. 2013); Kalyvakia , Chalefka (Özkütük et al. 2013). Zygiella atrica (C. L. Koch, 1845) New records Identification : https://araneae.nmbe.ch/ . KERYNEIA : Saint Hilarion Castle, 610m, 1 ♀, under stones in Pinus forest, 22.XI.2010 (CRB). Citations LIMASSOL: Amathus, 1 ♀, under stones in dry None. valley, 6.IV.1986 (N HMW),1 ♂, front yard of

New records hotel, 8.IV.1986 (NHMW); Armenochori, 1 ♀, LIMASSOL: Troodos Square, 1 ♀, beaten from under stones of a dry stone wall, 14.IV.1 986 shrub in Pi nus nigra forest, 11.V.2016 (CMH). (NHMW); Troodos Square, 1 ♀, Pinus nigra NICOSIA: Agridi, near Dali, 1 ♀, in garden, forest, 11.V.2016 (CMH). 26.II.2011; 1 ♂, in house, XI.2012 (CDM). NICOSIA : Agios Sozomenos , near Dali, 1 ♀, under stones and in dry grassland, 29.IV.2013; Distribution: Europe, Russia. New to Cyprus. Agridi, near Dali, 1 ♀, under eucalyptus bark in

garden, 29.IV.2013, 1 ♀, in garden, V.2004, 1 ♀, Family Cheiracanthiidae in field, IV.2007, 1 ♀, in garden, V.2009; Athalassa National Forest Park, Nicosia, 1 ♀, Cheiracanthium elegans Thorell, 1875 under stones in pine plantation, 8.V.2013 (all CDM), 1 ♀ (CAR - S); Dali, 13, Agios Identification : https://araneae.nmbe.ch/ . Andronikos, 1 ♀, in garden, V.2007(CDM); Dali, Lymbourena Hill, 1 ♂ 1 ♀, in field, IV.2004 Citations (CDM); Kapedes, 1 ♀, pitfalls in Pinus brutia None. forest, 17.I. - 23.IV.2016 (CMH); Lakatamia, 2 New records ♂♂, in olive grove, 1.I. - 1.X. 2016 (CMH); LIMASSOL: Akrotiri Peninsula, edge of central Pallouriotissa, Nicosia, 1 ♂, in garden, salt pan, 1 ♂, Salicornia marsh, 27.IV.2007 15.VI.2008 (CDM); Kampos NE, Xeros River, (CDM), 1 ♀ (CAR - S). 550m, 1 ♀, stones in riverbed , 20.V.2011 (CRB); NICOSIA: Agrid i, near Dali, 1 ♂, inside house, Limnitis River, Kato Pyrgos SE, 200m, 2 ♂♂, 28.IV.2009; 1 ♂, in garden, 18.IV.2012 (CDM). beaten in Pinus forest, 2.V.2010 (CRB); Masari

Spiders of Cyprus 44

S., Merika River, 120m, 1 ♂, stones in wasteland, None. 24.V.2011 (CRB). PAPHOS : Akamas Peninsula, Neo Chorio picnic New records NICOSIA : Agridi, near Dali, 1 ♂, in grass in site, 1 ♀, swept from phrygana an d grass, yard, V.2009 (CDM); Kapedes, 1 subadult ♂, 2.V.2007 (CRS); Kato Pyrgos, 700m, 1 ♀, under pitfalls in Pinus brutia forest, 17.I. - 23.IV.2016 stones in Pinus forest, 2.V.2010 (CRB); Kinousa (CMH); Lakatamia, 2 ♀♀, in olive grove, 1.I. - E., Agios Merkourios, 350m, 1 ♀, under stones 1.X. 2016 (CMH); Xerargaka S, Orkontas, 1 ♂, and litter in Pinus forest, 1.V.2010 (CRB). riverside vegetation, 26.III.2016 (CMH).

Distribution: Holarctic, in Europe originally Distribution: Greece, Libya to Malaysia, limited to the Mediterranean region but recently Australia, USA, Cuba, Brazil. New to Cyprus, and expanding into Western Europe. only found in the Nicosia district.

Cheiracanthium seidlitzi L. Koch, 1864 Family Clubionidae Identification Borges & Wunderlich, 2008 : 264, figs 17a - b. Clubiona pseudoneglecta Wunderlich, 1994

Citations Identification N one. Merrett, 2001: 32: fig s 1 - 3, 7 - 9.

New records Citations NICOSIA : Kapedes, 1 ♀, pitfalls in Pinus brutia None. forest, 17.I. - 23.IV.2016 (CMH); Lakatamia, 1 ♂, in olive grove, 1.I. - 1.X. 2016 (CMH); Lefka New records NW, 120m, 3 ♂♂ 1 LIMASSOL: SW Limassol, Lady’s Mile Beach, 1 subadult ♀, beaten from shrubs in fruit plantation, ♀, in salt marsh, 19.II.2002, D. W. Wrase leg. 18.V.2011 (CRB). (CKHK). PAPHOS: Mavrokolympos Reser voir, 1 ♂, 13.III.2018 (CKHK). Distribution: Europe, Algeria, Morocco, Caucasus. New to Cyprus. Distribution: Mediterranean to Central Asia. New to Cyprus. Porrhoclubiona vegeta Simon, 1918

Cheiracanthium sp. Identification Van Helsdingen: 1979: 299, figs 5, 8 - 9 . Identification Several Cheiracanthium specimens were captured Citations that cannot be identified to species level. The None. genus is in need of revision, because species are New records insuffiently described and specific variation is not FAMAGUSTA : Cape Greko, 1 ♀, beaten from well understood. shrubs in Juniperus phoenicea forest, 1.V.2016 (CMH); Gialousa E., Agios Thyrsos, 20m, 1 ♀, beaten from shrubs, 16.V.20111 (CRB); Salamis Family Cithaeronidae ruins, 50m, 1 ♀, beaten from low tree s, 15.V.2011 (CRB). Cithaeron praedonius O. Pickard - Cambridge, LARNACA: Agios Minas, 1 ♀, under stone in 1872 pine wood, 20.III.1998, coll. J. Murphy (MMUE); Mosfiloti, 1 ♀, under stones in dry eroded Identification grassland on hillside, 12.IV.2017 (CAR - S). Platnick, 1991: 6, figs 9 - 13. LIMASSOL : Pissouri, 2 ♀♀, under stone, 24.III.1998, coll . J. Murphy (MMUE); Agros SW, Citations 950m, 1 ♀, under stones in Pinus forest, 5.V.2010

Spiders of Cyprus 45

(CJVK); Pano Platres, near top of road to Family Cybaeidae Millomeris Waterfalls, 1 ♀, under stones in abandoned vineyard, 28.IV.2007 (CAR - S). Cedicus flavipes Simon, 1875 NICOSIA : 13, Agios Andronikos, Dali, 1 ♀, in Cedicus flavipes ; Kulczyński, 1908: 77. gar den, IV.2007 (CDM); Agridi, near Dali, 1 ♂, in garden, 14.III.2010; 1 ♂ in garden, 20.VI.2011 Identification (CDM); Masari S., Merika River, 120m, 1 ♀, Levy, 1996: 118, figs 129 - 134. stones in wasteland, 24.V.2011 (CRB). Citations PAPHOS: Akamas Peninsula, 1 ♂, in coastal Without precise locality (Kulczyński, 1908). grassland, 22.III.1998, coll. J. Murp hy (MMUE); KERYNEIA : Mount Kyparissovoun (Özkütük et Gialia, 1 ♀, under bush in pines, 25.III.1998, coll. al . 2013); Trap eza Mountain, Pentadactylos J. Murphy (MMUE). Kinousa, 1 ♂ 1 ♀, under Range (Özkütük et al . 2013). stone in shrubs, 19.III.1998; 1 ♀, under stones in pine wood, 23.III.1998, coll. J. Murphy (MMUE), New records 1 ♂ 3 ♀♀, under stones in pine wood, 28.III.1998, FAMAGUSTA : Prastio S., Lake Kouklion, 1 ♀, fallow land with Salicornia/Suaeda , 20.II.2011, c oll. J. Murphy (MMUE); Kourdaka, Ezousa D. W. Wrase leg. (CKHK). river, 3 ♂♂ 11 ♀♀, under bark of Eucalyptus , KERYNEIA : Larnakas tis Lapithou, 1 ♀, stones 14.III.2018 (CKHK); Makounta, 4 ♀♀, under in grassland, 17.I.2016 (CMH); Saint Hilarion stones in scrub, 19.III.1998, coll. J. Murphy Castle, 2 km W., 610m, 1 ♀, under stones in open (MMUE); Neo Chorio, 2 ♀♀, under stone in low Pinus forest, 20.III.2011 (CRB). scrub, 30.III.1998, coll. J. Murphy (M MUE). LIMASSOL: Limassol, 1 ♂ 5 ♀♀, G. Mavromoustakis leg., II. 1933 (BMNH); Mount Distribution: Europe, Central Asia, North Africa, Olympos (Chionistra), S. slope, 1900m, 4 ♀♀, Canary Is. New to Cyprus. under stones in o pen Pinus forest, 4.V.2010 (CRB); Mount Olympos (Chionistra), just below summit, 4 ♀♀, under large rocks, Pinus nigra Family Ctenidae forest, 28.IV.2007 (CAR - S), 2 ♀♀, 28.IV.2007 (CRS); Mount Olympos (Chionistra), 1830m, 1 Anahita syriaca (O. Pickard - Cambridge, 1872) ♀, pine forest with open places, 23.II.2011, D. W .

Identification Wrase leg. (CKHK); Troodos Square, 1 ♀, Pinus Anahita syriaca ; Levy, 2003: 11, figs 16 - 22 nigra forest, 11.V.2016 (CMH). (descr. ♂, ♀). NICOSIA: road Pedoulas - Prodromos, 1 ♂, pitfalls in Pinus nigra forest, 6.IV. - 8.VI.2014 (CMH); Citations near Platania Scout camping site, 1 ♂, pitfalls in N one. Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH). PAPHOS: Akamas Peninsula, 1 ♀, in coastal New records grassland, 22.III.1998, coll. J. Murphy (MMUE); NICOSIA: Marathasa Dam, 2 ♂♂, pitfalls in Kathikas W., S Arodes, Aspros Valley, 527m, 1 riverside vegetation, 7.V. - 10.VII.2016 (CMH); ♀, open pasture on clay, 24.II.2011, D. W. Wrase road Pedoulas - Lemithou, 2 ♂♂, pitfalls in Pinus leg (CKHK); Kidasi S., Dhiarizos river, 260m, 1 nigra and Pinus brutia forest, IV. - VII.2017 ♀, field edges, 21.II.2011, D. W. Wrase leg (CRB). (CKHK); Kinousa SE, 400m, 1 ♀, under stones KERYNEIA: Larnakas tis Lapithou, 1 subadult ♀ , and litter in Pinus forest, 1.V.2010 (CRB). stones in grassland, 17.I.2016. PAPHOS: Avakas Gorge area, 1 su badult ♀, hand Distribution: Eastern Mediterranean. collected under stones, 23.II.2014 (CDM) ; Souskiou, 2 subadult ♀, hand collected under stones , 5.II.2017 (CMH). Family Dictynidae

Distribution: Israel. New to Cyprus . Argenna patula (Simon, 1874)

Spiders of Cyprus 46

Identification : https://araneae.nmbe.ch/ . 4.V.2007 (CAR - S), 1 ♂, (CRS); Akrotiri Peninsula, Fasouri, 1 ♂, swept from flowering Citations herbs and grass, 27.IV.2007 (CAR - S); same site, None. 1 ♂ 3 ♀♀, in reed litter, 27 .IV.2007, 2 ♀♀ (CAR - S), 1 ♂ 3 ♀♀, 4.V.2007 (CRS); Fasouri reedbeds, New records Akrotiri Peninsula, 4 ♂♂ 5 ♀♀, on muddy LIMASSOL: Akrotiri Peninsula, Fasouri reed ground, edge of reedbed, 17.IV.2017 (CAR - S); beds, 1 ♂ in reed litter, 27.IV.2007 (CAR - S). Caledonia Waterfall, Pano Platres, 2 ♂♂ 5 ♀♀, PAPHOS: Kinousa, 2 ♀♀, under stones in pine under stones and in litter, edge of P. nigra forest, wood, 28.II I.1998, coll. J. Murphy (MMUE). 26.I V.2007 (CAR - S); Pano Platres, 1 ♀, in small web on shrubs, 6.V.2007 (CAR - S); Troodos S., Distribution: Palaearctic. New to Cyprus. road to Pano Platres, 2 ♂♂ 3 ♀♀, under stones in Pinus nigra forest, 28.IV.2007 (CRS), 2 ♂♂ 5 ♀♀ (CAR - S), 2 ♂♂ 3 ♀♀, 26.IV.2007 (CAS). Brigittea civica (Lucas, 1850) NICOSIA: Agridi, near Dali, 1 ♂, in yard, 14.III.2010 (CDM), 3 ♀♀, on and under stones, Identification : https://araneae.nmbe.ch/ . river bank, 29.IV.2013 (CDM), 1 ♂, in eucalyptus Citations litter, 29.IV.2013 (CAR - S); Athalassa National LIMASSOL: Akrotiri Salt Lake (Logu nov , 2013). Forest Park, Nicosia, 3 ♂♂ 10 ♀♀, in Eucalyptus New records plantation, 2.V.2013 (CRS), 1 ♂ 3 ♀♀, under LIMASSOL: Pissouri, 1 ♀, under stone, stones in pine plantation, 8.V.2013, 1 ♂ 1 ♀, in 24.III.1998, coll. J. Murphy, (MMUE). deep eucalyptus litter (CAR - S), 10 ♀♀ (CRS); Marathassa Dam, 3 ♀♀, beaten from shrubs Distribution: Circum - Mediterranean, N. around lake, 20.V.2011 (CRB); Lympia America. Reservoir, Lympia, 1 ♀, under stones below dam

Brigittea innocens (O. Pickard - Cambridge, wall, 9.V.2013 (CAR - S), 3 ♀♀ (CDM), 1 ♀ 1872) (CRS). PAPHOS: Akamas Peninsula, ca. 2 km. S. of Identification Cape Geronnisos, 1 ♂ 3 ♀♀, under stones in dry Lehtinen, 1967: 219, figs 309, 323. phrygana, 3.V.2007 (CRS); Avakas gorge and surroundings, 1 ♀, 22.III.2018 (CKHK); Evretou Citations Reservoir, 1 ♀, under stones and in grass None. tussocks, 13.IV.2017 (CAR - S); Kel efos bridge, Diarizos River Dam, 550m, 1 ♀, beaten from New records shrubs, 21.V.2011 (CRB); Lefka NW, 120m, 1 ♀, FAMAGUSTA: Salamis ruins, 50m, 3 ♂♂, beaten beaten from bushes in fruit plantation, 18.V.2011 from low trees, 15.V.2011 (CRB); Agios Sergios , (CRB); Oikos N., Xerargaka picnic site, 350m, 2 30m, 1 ♀, beaten in Pinus plantation, 15.V.2011, ♀♀, beating shrubs and stones in Pinus forest, CRB). 20.V.2011 (CRB); KERYNEIA : Panagia Apsinthiotissa Monastery ruins , 350m, 2 ♂♂ 7 ♀♀, beaten in clearing in Distribution: Italy, Eastern Mediterranean, Pinus forest, 24.V.2011 (CRB). Kazakhstan. New to Cyprus. LARNACA: Kornos picnic site, 350m, 2 ♀♀, beaten in Pinus forest, 28.V.2011 (CRB); Larnaca Devade tenella (Tyschchenko, 1965) SW, Hala Sultan Tekke Mosque, 5m, 1 ♀, under stones and litter bordering salt lake, 16.V.201 1 Identification (CRB); Mosfiloti, 1 ♀, under stones in dry eroded Esyunin & Efimik, 2000: 680, figs 1.7 - 8, 2.10, grassland on hillside, 12.IV.2017 (CAR - S); 3.5. Stavrovouni Monastery, 350m, 1 ♀, on bark of Pinus trees, 28.IV. 2010 (CRB). Comments LIMASSOL: Akrotiri Peninsula, 2 ♂♂ 8 ♀♀, There has been much confusion in the past a bout margin of central salt lake, 7.V.1972, co ll. A. Devade species. The first Devade species was Stubbs (MMUE); Akrotiri Peninsula, near British described in the genus Amaurobius by O. Pickard - Airbase, 2 ♂♂ 3 ♀♀, swept in pine wood, Cambridge (1872) as A. indistinctus and later

Spiders of Cyprus 47

described again under different names and in LIMASSOL: Mount Olympos (Chionistra), different genera: Diotima hirsitussima Simon, 1820m, 1 ♀, under stones in Pinus forest, 1880, Altella lybica Denis, 1947 and Strinatella 25.XI.2010 (CRB). spinosa Denis, 1957. This bears out the doubts PAPHOS: Kelefos br idge, 465m, deciduous arachnologists had in placing the species in a wood, 1 ♀, 16.III.2018 (CKHK). correct genus. Devade tenella (Tyschchenko, 1965) from Kazakhstan, with its synonyms Comments Momius hispidus Andreeva & Tyschchenko, 1969 In Cyprus, only two females were collected, their and Devade uiensis Esyuin1994 is a second, epigyne with two vertical ducts not differing from closely related Devade species. Esyunin (1994) the type material of L. arabs described above. The considered it a synonym of D. indistincta whereas captures of males should confirm th is Esyunin & Marusik (2000) and Esyunin & Efimik identification. (2000) considered it an independent species. The holotype female of Devade indisticta (O. Pickard - Distribution: Algeria, Greece. New to Cyprus. Cambridge, 1872) and the type materal of D. tenella should be re - examined to clarify the status Lathys humilis (Blackwall, 1855) of the two species. Our specimens resemble most the careful drawings of Esyunin & Efimik (2000) Identification : https://araneae.nmbe.ch/ . of Devade tenella and are therefore iden tified as such. Citations None. Citations None. New records LIMASSOL: Troodos, near summit of Mount New records Olympos (Chionistra), 1 ♀, under stones in Pinus FAMAGUSTA : Salamis S., Silver Beach, 5m, 1 brutia forest, 19.IV.2017 (CAR - S). ♂, in pitfalls in salt marsh, 15 - 27.V.2011 (CRB). PAPHOS: Akamas Peninsula, 1 ♀, in coastal LARNACA : Larnaca salt lake, 5m, 2 ♂♂ 11 ♀♀, scrub, 22.III.1998, coll. J. Murphy (MMUE). stones and litter in salt marsh, 27.IV.2010 (CJVK, Distribution: Palaearctic. New to Cyprus. CRB); Larnaca W., Hala Sultan Tekke mosque, 5m, 2 ♂♂ 2 ♀♀, under stones and in litter in salt M arilynia bicolor (Simon, 1870) marsh, 20.XI.2010 (CRB), 1 ♂ 2 ♀♀, pitfalls at same site. Identification : https://araneae.nmbe.ch/ . LIMASSOL: Akrotiri peninsula, edge of central salt pan, 1 ♀, under stones & debris in Salicornia Citations None. etc., 30.IV.2013 (CAR - S); Salt Lake at Akrotiri village SW Limassol, 1 ♂, in salt marsh, New records 28.II.2011, D. W. Wrase leg. (CKHK); Akrotiri LIMASSOL : Alassa SE, near river Limnatis, 1 ♀, Peninsula, salt lake, 2 ♂♂ 3 ♀♀, under stones in dry grassland, 5.V.2007 (CAR - S). Salicornia , 15.III.2018 (CKHK). PAPHOS : Kelefos bridge, dam on Diarizos river, PAPHOS : Mavrokolympos Reservoir, NE 550m, 1 ♂, under stones along rivulet, 21.V.2011 Potima, 150m, 1 ♀, under stones along lake, (CRB).

22.V.2011 (CRB). Distribution: Mediterranean region to Central Distribution: Mediterran ean. New to Cyprus. Asia. New to Cyprus.

Lathys arabs Simon, 1911 Nigma flavescens (Walckenaer, 1830)

Identification Identification : https://araneae.nmbe.ch/ . Bosmans et al. 2009: 32. Citations Citations None. None. New records New records

Spiders of Cyprus 48

LIMASSOL: Pano Platres, 1050m, 2 ♂♂ 1 ♀, Kinousa, 1 ♀, on bush in pine wood, 28.III.1998, litter and beaten near Mesa Pota mos Waterfall s, coll. J. Murphy (MMUE); Kourdaka, Ezousa 23.V.2011 (CRB); Pano Platres, 1 ♀, in small web river, 6 ♂♂ 2 ♀♀, 14.III.2018 (CKHK); in shrub, 6.V.2007 (CAR - S ). Mavrokolympos Reservoir, 1 ♂, 13.III.2018 NICOSIA : 13, Agios Andronikos, Dali 1 ♂ 2 ♀♀, ( CKHK); Pegeia, Pikni Forest, 370 m, 1 ♂, pine in garden, V.2007 (CDM); Agridi, near Dali, 1 ♀, forest, 11.III.2018 (CKHK). in field, IV.2007, 2 ♀♀, V.2008; 1 ♀, V.2012 (CDM); Agios Sozomenos , near Dali, 1 ♀, in dry Distribution: Europe, Azores, Madeira, Canary grassland, 29.IV.2013 (CAR - S); Lympia, Lympia Islands. New to Cyprus. Reservoir, 1 ♀, i n rushes below dam wall, 9.V.2013 (CAR - S); Pallouriotissa, Nicosia, 2 ♀♀, Nigma walckenaeri (Roewer, 1951) in building, V.2006 (CDM); Prodromos S., Trooditissa picnic site, 1450m, 4 ♂♂, beating in Identification : https://araneae.nmbe.ch/ . Pinus forest, 21.V.2011 (CRB). PAPHOS: Kelefos bridge, 465m, deciduous Citations wood, 1 ♂, 16.III.2 018 (CKHK); Kourdaka, None. Ezousa river, 1 ♂, 14.III.2018 (CKHK). New records

LARNACA: Larnaca SW., Hala Sultan Tekke Distribution: Palearctic. New to Cyprus. mosque, 5m, 2 ♀♀, beaten in garden, 15.V.2011

(CRB). Nigma puella (Simon, 1870) LIMASSOL : Agios Georgios E., Palaiomylos,

1250m, 2 ♀♀, beaten in mixed Pinus and Identification : https://araneae.nmbe.ch/ . Quercus forest, 4.V.2010 (CRB); Akrotiri

Peninsula, 1 ♂, swept from grass and reeds, Citations Fasouri reedbeds, 17.IV.2017 (CAR - S). None. NICOSIA: Cedar Valley Forest, 1180m, 1 ♂, beaten in Cedrus forest, 19.V.2011 (CRB); New records PAPHOS : Gialia, 2 ♂♂ 1 ♀, on bush in pines, FAMAGUSTA: Lysi, 2 ♀♀, beaten from shrubs 25.III.1998, coll. J. Murphy (MMUE); Vretsia W., in garden, 15.IV.2016 (CMH). near Kelefos bridge, 650m, 1 ♀, beaten from KERYNEIA : Larnakas tis Lapithou N. , 950m, 1 shrubs, 19.V.2011 (CRB). ♀, under stones in Pinus forest, 25.XI.2010

(CRB). Distribution: Palearctic. New to Cyprus. LIMASSOL : Pano Platres, 1050m, 2 ♂♂, beaten near Mesa Potamos Wate rfall s, 23.V.2011 (CRB);

Troodos S., 1350m, 1 ♂, beating in mixed Quercus - Pinus forest, 3.V.2010 (CRB); Troodos Fa mily Dysderidae Square, 1 ♂ 12 ♀♀, beaten from shrubs in Pinus nigra forest, 11.V.2016 (CMH). Dysdera gigas Roewer, 1928 NICOSIA : Cedar Valley Forest, 1180m, 1 ♂, beaten in Cedrus fores t, 19.V.2011 (CRB), 1 ♀, Identification swept from shrubs, 29.IV.2007 (CAR - S), 1 ♂ Deeleman - Reinhold & Deeleman, 1988: 224, figs (CRS); Kakopetria S., Platania forest station, 271 - 276. 1100m, 1 ♂, beating in dense Pinus forest, 4.V.2010 (CRB); Pedoulas, 1 ♂ 1 ♀, by woodland Citations stream, 3.V.1982, coll. A. Stubbs (MMUE); road None. Pedo ulas - Prodromos, 1 ♀, pitfalls in Pinus nigra forest, 6.IV. - 8.VI.2014 (CMH). New records PAPHOS : Akamas Peninsula, 1 ♂, woodland near LIMASSOL: Pano Platres, 1050m, 1 ♀, litter near Agios Minas, 3.V.2007 (CRS); Avakas Gorge, 2 Mesa Potamos Waterfall s, 23.V.2011 (CRB). ♂♂ 1 ♀, stones in gorge, 5.IV.1997 (CJB); Evretou Reservoir, 350m, 1 ♀, on shrubs and Distribution: Crete and new to Cyprus. grassland around lake, 1.V.2010 (CRB); Evretou Reservoir, Stavros tis Psokas river, 2 ♂♂ 2 ♀♀, 20.III.2018 (CKHK); Gialia, 1 ♂, on bush in pines, 25.III.1998, coll. J. Murphy (MMUE); Dysdera lata Reuss, 1834 Spiders of Cyprus 49

Dysdera lata ; Deeleman - Reinhold & Deeleman, Dysdera spinicrus ; Deeleman - Reinhold & 1988: 200. Deeleman, 1988: 204.

Identification Identification Deeleman - Reinhold & Deeleman, 1988: 200, figs Deeleman, 1988: 204, figs 206, 207, 211 - 214, 201 - 205, 210. 219, 220.

Citations PAPHOS : Timi (Deel eman, 1988). Citations PAPHOS : Petra tou Romiou, Baths of Aphrodite New records (Deeleman - Reinhold & De eleman, 1988); LIMASSOL: near Prodromos reservoir, pitfalls in Episkopi Game Reserve (Deeleman - Reinhold & Pinus nigra forest, 6.IV - 8.VI.2014, M. Deeleman, 1988). Hadjiconstantis leg. NICOSIA: Laka matia, 1 ♂, stones in olive New records groove, 2016 (CMH); Marathassa Dam, 1 ♂, FAMAGUSTA : Agios Sergios , 30m, 1 ♂, under pitfalls in riverside vegetation, 7.V - 10.VII.2016 stones in phrygana, 17.III.2011 (CRB), 1 ♀, (CM); Pano Amiantos, 2 ♂♂, pitfalls in Pinus 11.V.2011 (CRB); Rizokarpas, 1 ♀, Juniperus brutia forest, 6.IV - 8.VII.2014 (CMH); near Platania Scout camp, 2 ♂♂, pitfalls in Pinus forest, 12.III.2016 (CMH). bruti a forest, 6.IV - 8.VI.2014 (CMH). LARNACA: Meneou, 1 ♀, fields, 8.IV.2016 (CMH); Mosfiloti, 1 ♀, under stones in open pine Distribution: Algeria, Greece, Turkey, Crimea woodland, 12.IV.2017 (CRS). and Cyprus. LI MASSOL: Alassa SE, near river Limnatis, 2 ♀♀, under large rocks in open Pinus nigra forest, Dysdera longimandibularis Nosek, 1905 27.IV.2007 (CRS); Mount Olympos (Chionistra), Dysdera longimandibularis ; Deeleman - Reinhold just below summit, 1 ♂ 1 ♀ , 28.IV.2007 (CAR - & Deeleman, 1988: 163. S) ; road Chionistra - Prodromos, 1 ♂, pitfalls in Pinus nigra forest, 6.IV. - 8.VI.2014 (CMH); Identification Pano Platres, Millomeris Waterfalls, 1 ♀, under Deeleman - Reinhold & Deelem an, 1988: 163, figs stones in abandoned vineyard, 28.IV.2007 (CAR - 39 - 43. S); Pano Platres, near top of road to Millomeris at base of limestone rock face of gorge, 5.V.2007 Citations (CAR - S); Prodromos E., near Prodromos dam, PAPHOS : Episkopi game reserve (Deeleman - 1 550m, 4 ♀♀, pitfalls in Pinus forest, XI.2010 - Reinhold & Deeleman, 1988). III.2011 (CRB); Prodromos S., Trooditissa picnic place, 1300m, 1 ♂ 1 ♀, pitfalls in open Pinus New records forest, XI.2010 - III.2011 (CRB); Troodos, near LIMASSOL: Prodromos S., Trooditissa picnic summit of Mount Olympos (Chionistra), 2 ♂♂ 2 place, 1300m, 2 ♂♂, pitfalls in open Pinus forest, ♀♀, under stones in Pinus brutia forest, 19.IV.2017 (CAR - S); 10 km S. of Troodos, 1 ♀, XI.2010 - III.2011 (CRB). under stones in Pinus brutia forest, 19.IV.2017 NICOSIA : Cedar Valley Forest, 1180m, 1 ♂, (CAR - S). stones in Cedrus forest, 19.V.2011 (CRB); NICOSIA : Agios Sozomenos , near Dali, 1 ♀, Prodromos E., near Prodomos dam, 1550m, 1 ♀, stones in disturbed phrygana, 30.IV.2007 (CAR - under stones a nd litter in Pinus forest, 2.V.2010, 2 S); Agridi, near Dali, 1 ♀, in yard, V.200 9, 1 ♂, ♂♂, pitfalls at same site, XI.2010 - III.2011 in garden, V.2010 (CDM); Athalassa National Forest Park, Nicosia, 2 ♀♀, under stones in pine (CRB), 1 ♂ 1 ♀, III. - V.2011 (CRB). plantation, 2.V.2013 (CAR - S); Avakas Gorge, PAPHOS : Avakas Gorge, 1 ♂ 1 ♀, litter along Akamas Peninsula, 2 ♂♂ 1 ♀, in rocky gorge, stream, 22.II.2011, D. W. Wrase leg. (CKHK). 23.II.2014 (CDM); Strovolos, 2 ♀♀; around houses, 1.V.2016 (CMH). Distribution: Turkey and Cyprus. PAPHOS : Akamas Peninsula, Agios Minas, 2 ♂♂, under stones in meadow with tall grass, 1.V.2007 (CAR - S); Akamas Peninsula, 1 ♀, in Dysdera spinicrus Simon, 1882 hotel grounds , 3.V.2007 (CAR - S); Asprokremmos

Spiders of Cyprus 50

Dam, N. Kouklia, 150m, 1 ♂, herbs and stones along lake, 29.IV.2010 (CRB); Evret ou Reservoir. At least 2 new species of Dysdera were collected. 1 ♀, under stone in grass tussocks. 13.IV.2017 They will be described in a separate paper. ( CAR - S); Kathikas W. Aspros valley, 527m, 1 ♀, under stones in open pasture, 24.II.2011, D. W. Dysderocrates kibrisensis Gücel, Wrase leg. (CKHK); Neo Chorio, 1 ♀, under Charalambidou, Göçmen & Kunt, 2019 stone in field, 20.III.1998, coll. J. Murphy Dysderocrates kibrisensis Gücel et al ., 2019: 46, (MMUE). figs 2 - 6 (descr. ♀).

D istribution: The Balkans to Syria, Cyprus. Citations NICOSIA: Trapeza Mountains, Trape za cave Dysdera westringi O. Pickard - Cambridge, 1872 ( Gücel et al., 2018; type locality ). Dysdera westringi ; Strand 1907: 393; Deeleman - New records Reinhold & Deeleman, 1988: 202. None.

Identification Distribution: An endemic species of the Trapeza Deeleman - Reinhold & Deeleman, 1988: 202, figs Mountains (= Pentadactylos range). 209, 216 - 218. Harpactea cecconii (Kulczyński, 1908) Citations Harpactes cecconii Kulczyński, 1908: 51 (descr. Without precise locality (Strand, 1907). ♂, ♀); Brignoli, 1980: 77. LIMASSOL : Mount Troodos, 1600m (Deeleman, 1988). Identification PAPHOS : Episkopi game reserve (Deeleman - Kulczyński, 1908: 51, pl. 2, figs 1 - 2. Reinhold & Deeleman, 1988); Petra tou Romiou (Baths of Aphrodite) (Deeleman - Reinhold & Citations Deeleman, 1988); T imi (Deeleman - Reinhold & Without precise locality (Kulczyński, 1908; Deeleman, 1988). Brignoli, 1980).

New records New records LIMASSOL : Agios Tychonas, 1 ♀, under stones, LARNACA : Achna Reservoir, 50m, 1 ♂ , in 19.IV.1986 (NHMW); Amathus, 1 ♀, on house pitfalls in marshy area around lake, 15 - walls, 9.IV.1986 (NHMW); Pano Platres, 1 ♂ 1 27.V.2011 (CRB); Larnaca salt lake, 10m, 1 ♂, in ♀, under stones in gorge below Caledonia pitfalls around lake, III. - V.2011 (CRB). Waterfall, 19.IV.2017 ( CAR - S); LIMASSOL: Mount Olympos (Chionistra), Troodos S., road to Pano Platres, 1 ♂, under 1900m, 1 ♂ 3 ♀♀, in pitfalls in Pinus forest, III. - stones in Pinus nigra forest, 1.V.2007 (CAR - S). V.2011 (CRB); Mount Olym pos (Chionistra), S. PAPHOS : Akamas Peninsula, S. Neo Chorio, slope, 1900m, 1 ♂ 1 ♀, under stones in open 270m, 1 ♀, stones in karst landscape, 22.IV.2017, Pinus forest, 4.V.2010 (CRB); Pano Platres, track S. Huber leg. (CSH); near Avakas Gorge, 1 ♂ 1 near Caledonia Waterfall, 2 ♂♂, under stones by ♀, under ston es in field with tall grass, edge of P. nigra forest, 26.IV.2007 (CAR - S); 14.IV.2017 (CAR - S); Drouseia S., Panagia Pedoulas (N34°58'17'' E32°49'48''), 960m, 1 ♂, Chrysospiliotissa, 580m, 1 ♀, night catch in small n ight catch in orange garden, 20.IV.2017, S. cave, 22.IV.2017, S. Huber leg. (CSH); Neo Huber leg. (CSH); ca. 5 km S. of Troodos, road to Chorio, 1 ♀, under stone in scrub, 30.III.1998, Pano Platres, 1 ♂, under stones in Pinus nigra coll. J. Murphy (MMUE); Polis W., Latsi, forest, 28.IV.2007 (CAR - S), 2 ♂♂ 1 ♀, 1.V.2007 Aph rodite Beach, 60m, 1 ♀, night catch, (CAR - S); Troodos, near summit of Mount 22.IV.2017, S. Huber leg. (CH); Polis W., Latsi Olympos (Chionistra), 1 ♂, un der stones in Pinus Botanical Garden, 60m, 1 ♂, night catch, brutia forest, 19.IV.2017 (CAR - S). 22.IV.2017, S. Huber leg. (CSH ). NICOSIA: Cedar Valley Forest, 1 ♀, under stones, 29.IV.2007 (CAR - S). Distribution: Turkey and Cyprus. PAPHOS: 3 km SE Baths of Aphrodite, 1 ♂ 1 ♀, Eucalyptus forest along fields, 5.V.2007 (CRS); Akamas Peninsula, 1 ♀, in litter of coastal scrub, 22.III.1998, coll. J. Murphy (MMUE); Avakas Dysdera sp. Gorge, 2 ♂♂ 1 ♀, in rocky gorge, 23.II.2014

Spiders of Cyprus 51

(CDM), 1 ♂ 1 ♀, in litter in gorge, 5.IV.1997 (CJB); Evretou Reservoir, 1 ♂, under stones in Identification : https://araneae.nmbe.ch/ . grass tussocks, 13.IV.2017 (CAR - S); Kinousa, 1 ♂, under sto ne in pine wood, 28.III.1998, coll. J. Citations Murphy (MMUE); Neo Chorio, 1 ♀, under stone LARNACA: Larnaca (Marusik et al ., 2014). in field, 17.III.1998, coll. J. Murphy (MMUE). KERYNEIA: Kerynaeia ( Z onstein & Marusik , 2019). Distribution: Cyprus. Also cited from Epidavros PAPHOS : Gialia ( Z onstein & Marusik , 2019) ; in continental Greece (Deltsjev & Paraschi, 1990) Kinousa ( Z onstein & Marusik , 2019 ) but this record is stro ngly doubted. New records Harpactea gunselorum Gücel, Fuller, Göçmen FAMAGUSTA : Cape Greco, 1 ♂, under stones in & Kunt, 2018 phrygana, 3.V.2013 (CRS); Davlos W ., 25m, 1 subadult ♀, under stones in phrygana near the Identification : Gücel et al., 2018: 81, figs 1A - D, beach, 25.V.2011 (CRB); Paralimni Lake, 65m, 1 2A - H, 3A - B. ♂ 1 juvenile, herbs and litter around dried lake, Citations 27.IV.2010 (CRB); Salamis ruins, 30m, 1 KERYNEIA : St Hilarion castle ( Gücel et al ., juvenile, under stones in ruins, 17.III.2011 (CRB). 2018). KERYNEIA : Karavas , 660m, 1 ♂ 1 ♀, wall NICOSIA: between Chalefka and Keryneias Cliff around garden of hotel, 22.XI.2010 (CRB); ( Gücel et al., 2018); Trapeza Mountains, near Kazafani , 150m, 1 juvenile, under stones in olive Trapeza cave ( Gü cel et al., 2018; type locality ); grove, 23.XI.2010 (CRB). Keryneias Rock ( Gücel et al ., 2018); Kalyvakia , Chalefka ( Gücel et al., 2018). LARNACA : Larnaca salt lake, 5m, 6 juveniles, under stones and litter in salt marsh, 27.IV.2010 New records (CRB); Larnaca SW, Hala Sultan Tekke Mosque, None. 5m, 1 ♀, under stones and litter bordering salt lake, 16.V.2011 (CRB). Distribution: An endemic species of the Trapeza LIMASSOL : Alassa SE, near River Limnatis, 1 Mountains (= Pentadactylos range). juvenile, under stones in dry g rassland, 5.V.2007

Harpactea kalavachiana Gücel, (CAR - S). Charalambidou, Göçmen & Kunt, 2019 NICOSIA : Agios Tychonas, 3 ♀♀, under stones Harpactea kalavachiana Gücel et al., 2019, figs near houses, 19.IV.1986 (NHMW); Amathus, 1 9 - 11, 13 - 14 (descr. ♀). ♂, under stones in dry va lley, 25.IV.1986 (NHMW), 2 ♀♀, 9.IV.1986, 1 ♀, 9.IV.1986 Citations (NHMW); Armenochori, 1 juvenile, under stones NICOSIA: Kalyvakia , Kalyvakia Cave (Gücel et al., 2019; type locality ). of a dry s t one wall, 14.IV .1986 (NHMW); Erimi, 1 ♀, under stones in dry riverbed, 18.IV.1986 New records (NHMW); Gialia, 1 ♀, on pine trunk, 25.III.1998, None. coll. J. Murphy (MMUE); Kinousa, 1 ♀, under stone in pine wood, 28.III.1998, coll. J. Murphy Distribution: An endemic species of Cyprus . (MMUE); Lymbourena Hill, Dali, 1 ♀, in field,

IV.2004 (CDM); Kato Koutrafas - Astromeritis Harpactea sp. road, 120m, 1 ♂, herbs on pond shore, 18.V.2011 Several new species of Harapactea were (CRB); Parekklisha, 1 juvenile, under stones in collected. They will be described in a separate riverbed, 13.IV.1987 (NHMW); Pedoulas, 960m, paper. 1 ♀, road embankment, 18.IV.2017, S. Huber leg. (CSH); Pla tres, 1 juvenile, on walls, 22.IV.1986 Family Filistatidae (NHMW). PAPHOS : N. Pano Panagia, 1050m, 1 ♀, in litt er near spring, 19.V.2011 (CRB). Filistata insidiatrix (Forsskål, 1775)

Spiders of Cyprus 52

Zaitunia kunti Zonstein & Marusik 2016: 23, figs Distribution : Mediterranean to Turkmenistan, 10 - 11, 43C (descr. ♂♀). Cape Verde Islands. Identification Pritha nana (Simon, 1868) Zonstein & Marusik 2016: 23, figs 10 - 11, 43C.

Identification : https://araneae.nmbe.ch/ . Citations LIMASSOL: Troodos Mountains, 1 km. S. of Citations Troodos (Zonstein & Marusik 2016). None. New records New records FAMAGUSTA : Komi Kebir , 1 ♂, 150m, in NICOSIA: A gridi, near Dali, 1 ♀, in yard, V.2009 pitfalls in Pinus forest, III. - V.2011 (CRB). (CDM). PAPHOS : Evretou Reservoir, 350m, 1 ♀, shrubs PAPHOS : Kato Paphos archaeological site, 1 ♂, and grassland around lake, 1.V.2010 (CRB); under stones in grass, 10.V.2007 (CAR - S); nea r Kidasi N., along Diarizos River, 300m, 1 ♀, u nder Souskiou, 2 ♀♀, under stones in dry grassland, stones, 21.V.2011 (CRB); Kinousa SE, 400m, 1 1.V.2013 (CAR - S). ♀, under stones and litter in Pinus forest, Distribution: Mediterranean. 1.V.2010 (CRB).

Zaitunia annulipes (Kulczyński, 1908) Distribution: Turkey and Cyprus. Filistata annulipes (Kulczyński, 1908): 49 (descr. ♀). Zaitunia annulipes ; Brignoli, 1982 (transfer not Family Gnaphosidae fol lowed by Platnick); Wunderlich, 2011: 171; Zonstein & Marusik 2016: 20, figs 8 - 9, 43B Anagraphis pallens Simon, 1893 (descr. ♂♀). Identification Identification. Anagraphis pallens Chatzaki, Thaler & Mylonas, Zonstein & Marusik 2016,: 20, figs 8 - 9, 43B. 2002a : 567, figs 2 - 7.

Citations Citations Without precise locality (Kulczyński, 1908; type None. locality). LARNACA: Vyzakia, Larnaca salt lake (Zonstein New records & Marusik 2016). KERYNEIA : near Agios Georgios, 1 ♀, under LIMASSOL: Germasogeia (Zonstein & Marusik stones, 27.II.2011, H. Henderickx leg. (CJB). 2016); Paramali (Zonstein & Marusik 2016). LARNACA : Achna Rese rvoir, 50m, 1 ♂, pitfalls in marshy area around lake, 15 - 27.V.2011 New records (CRB). LARNACA: Mosfiloti, 1 ♂ 1 ♀, under stones in NICOSIA: Agridi , near Dali, 1 ♀, in grass in dry eroded grassland on hillside, 12.IV.2017 field, III.2014, det. M. Chatzaki (CDM); near (CAR - S ). Platania Scout camping site, 2 ♂♂, pitfalls in NICOSIA: Agios Sozomenos , near Dali, 1 ♂ 2 ♀♀, under stones in dry, rocky phrygana, Pinus brutia forest, IV. - VII.2017 (CMH ). 20.IV.2017 (CAR - S). PAPHOS: Agios Minas picnic site, 1 ♂, under Distribution: South Africa, Libya, Malta, Turkey, stone in open pine woodland, 13.IV.2017 (CAR - Syria, Israel, Iran, Crete, Chios. New to Cyprus. S). Aphantaulax cincta (L. Koch, 1866) Distribution Only known from Cyprus. Identification : https://araneae.nmbe.ch/ .

Zai tunia kunti Zonstein & Marusik, 2016 Citations

Spiders of Cyprus 53

None. LIMASSOL: Armenochori, 1 ♂, under stones of a dry stone wall, 14.IV.1986 (NHMW); Erimi, 1 ♀, New records under s tones along dry riverbed, 21.IV.1986 FAMAGUSTA : Kantara E., 610m, 1 ♀, beaten (NHMW); Fasoula, 2 ♂♂ 1 subadult ♂, olive from shrubs and stones in open Pinus forest, garden with dry stone walls, 16.IV.1986 25.V.2011 (CRB). (NHMW). KERYNEIA : Pentadactylos Range, Saint Hilarion PAPHOS : Avakas gorge, 50m, 1 ♂, litter in citrus Castle W, 600m, 2 ♂♂, under stones in Pinus yard at entrance of gorge, 30.IV.2010 (CRB); forest 17.V.2011 (CRB). Kinousa SE, 400m, 2 ♂♂ 1 ♀, under stones and LIMASSOL: Akrotiri Peninsula, Fasouri reed litter in Pinus forest, 1.V.2010 (CRB); Kinousa beds, 1 ♀, in reed litter, 27.IV.2007 (CRS). E., Agios Merkourios, 350m, 1 ♂, under stones PAPHOS : Vretsia W., near Kelefos Bridge, and in litter in Pinus forest, 1.V.2010 (CRB); n ear 650m, 1 ♂, beaten from shrubs, 19.V.2011 Souskiou, 1 ♀, under stones in grassland on (CRB). eroded hillside, 17.IV.2017 (CAR - S).

Distribution: S. and S. E. Europe, N. Africa, Israel. New to Cyprus. Comments Unt il now, Berinda cypria was only known from Aphantaulax trifasciata (O. Pickard - the type locality near Limassol. The species appears to be widespread in Cyprus and was most Camb ridge, 1872) frequently found in pine forest, but also in Identification : https://araneae.nmbe.ch/ . phrygana and in a citrus yard.

Citations Distribution : Endemic species to Cyprus . None. Berinda ensigera (O. Pickard - Cambridge, New records 1874) FAMAGUSTA: Protaras, Profitis Elias, 1 ♂, 2005 (CDM). Identification LARNACA: Hala Sultan Tekke mosque, 1 ♀, Chatzaki, Thaler & Mylonas, 2002a: 573, figs 14 - stones in su rrounding park, 20.IV.1986 (NHMW). 15, 18 - 19.

LIMASSOL: Akrotiri Peninsula, 27.IX.2007 (CDM). Citations PAPHOS: Kato Akourdaleia, in garden, VIII.2008 None. (CDM). New records Distribution: Palaearctic. New to Cyprus. FAMAGUSTA : Salamis ruins, 50m, 5 ♂♂, 2 ♀♀, under stones, 15.V.2011 (CRB). Berinda cypria Chatzaki & Panayiotou, 2010 LIMASSOL: Kourion archaeological site near Episkopi, 1 ♀, under stones, 9.V.2007 (CAR - S). Identification Chatzaki& Panayiotou, 2010: 50, figs 13 - 14, 17, Distribution : Greece, Turkey. New to Cyprus. 20 - 21. Berinda idae Lissner, 2016 Citations LIMASSOL: Kourion (Chatzaki & Panyiotou, Identif ication 2010; type locality). Lissner & Chatzaki 2016: 2016, figs 1 - 7 (descr. ♂♀). New records FAMAGUSTA : Cape Greco, Ayia Napa, 35m, 1 Citations ♂, under stones in phrygana , 27.IV.2010 (CRB); None. Kantara E., 650m, 1 ♂, pitfalls in Pinus forest, XI.2010 - III.2011 (CRB).

Spiders of Cyprus 54

New records New records LARNACA: Hala Sultan Tekke, 50m, stones and LARNACA : Dekhelia military camp, 1 ♂, litter bordering salt lake, 1 ♂, 15.V.2011 (CRB). 3.VIII.1958, M. Connelly leg. (BMNH). NICOSIA: Elia - Aggolemi road, 100m, 1 ♂ ♀, in LIMASSOL : road Chionistra - Prodromos, 1 ♂, rubbish i n dry river bed, 18.V.2011 (CRB). pitfalls in Pinus nigra forest, 6.IV. - 8.VI.2014 (CMH). Distribution : Only known from two Greek islands in the Dodekanese (Kalymnos, Nisyros) Distribution: Bulgaria, Greece, Israel, Turkey. and the Cyclades (Santorini, Christiani Islet). New New to Cyprus. to Cyprus.

Cryptodrassus helvolus (O. Pickard - Cambridge, Berlandina plumalis (O. Pickard - Cambridge, 1872) 1872) Cryptodrassus helvolus (Chatzaki & Russell -

Smith 2017: 238, figs 1 - 6. Identifi cation Chatzaki, Thaler & Mylonas, 2002b: 609, figs 13 - Identification 16. Chatzaki & Russell - Smith, 2017: 238, figs 1 - 6).

Citations Citations None. NICOSIA: Agridi, Dali, (Chatzaki & Russell -

New records Smith 2017) LARNACA : Larnaca SW, Hala Sultan Tekke mosque, 50m, 3 ♂♂, pitfalls in salt marsh, 15 - New records NICOSIA: Agios Sozomenos, near Dali, 1 ♂, 28.V.2011 (CRB). under stones in dry grassland, 30.IV.2007 (CAR - S); Dali, 1 ♂ 1 ♀, under stones around old water Distribution: Mediterranean region to China. mill, 12.V.2007 (CAR - S). New to Cyprus. Distribution: Cyprus, Israel Cesonia aspida Chatzaki, 2002 Drassodes cupreus (Blackwall, 1834) Identification Drassodes cupreus ; Logunov, 2013: 5 Cesonia aspida Chatzaki, in Chatzaki et al ., 2002b: 624, figs 63 - 68. Identification Roberts 1985: 66 , figs 22b,e, 23b - e . Citations None. Citations LIMASSOL: Episkopi, Kourion Beach (Logunov, 2013). New records PAPHOS : Mavrokolympos Reservoir (Logunov, LIMASSOL: road Kaminaria - Lemithou, 2 ♂♂, 2013); Xeropotamos River (Logunov, 2013). pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH). New records None. Distribution Greece (Crete), Turkey. New to Cyprus. Distribution: Palaearctic.

Civizelotes solsti tialis (Levy, 1998) Drassodes lacertosu s (O. Pickard - Cambridge, 1872) Identification Levy, 1998c : 139, figs 97 - 101. Identification Chatzaki, Thaler & Mylonas, 2002b: 618, figs 40 - Citations 43. None.

Spiders of Cyprus 55

Citations with dry stone walls, 16.IV.1986 (NHMW); None. Kourion, Apollo Hylates Temple, 1 ♀, under stones in the area of the archaeological site, New records 23.IV.1986 (NHMW); Panagia Glossa Monastery LARNACA : Dipotamos Reservoir, 150m, 1 ♀, near Kellaki, 1 ♂, under stones in cultivated land, herbs and stones bordering lake, 28.IV. 2010 12.IV.1986 (NHMW); Mount Olympos (CRB). (Chionistra), just below summit, 1 ♂ 2 ♀♀, under large rocks in open Pinus nigra forest, 2 7.IV.2007 LIMASSOL: Erimi, 3 ♂♂ 3 ♀♀, under stones (CRS); Pissouri, 2 ♂♂, under stones, 24.III.1998, along dry riverbed, 21.IV.1986 (NHMW); coll. J. Murphy (MMUE). Fasoula, 5 ♂♂ olive garden with dry stone walls, NICOSIA : Agridi, near Da li, 4 ♂♂, inside house, 16.IV.1986 (NHMW); Parekklisia, 2 ♂♂ 2 ♀♀, under stones in riverbed, 13.IV.1986 (NHMW). IV.2012, 1 ♂ , IV.2009, 1 ♀, V.2009, on veranda, 1 ♀, 28.IV.2009 (CDM); Cedar Valley Forest, 1 Distribution : Greece, Turkey, Israel, Syria. New ♂, under stones by track, 29 .IV.2007 (CRB); to Cyprus. Idalion archaeological Site, Dali, 1 ♂, under stones in phrygana, 1.V.2013 (CAR - S); Drassodes lapidosus (Walckenaer, 1802) Kakopetria S., Platania forest station, 1100m, 1 ♂

Identification : https://araneae.nmbe.ch/ . 1 ♀, under stones in dense Pinus forest, 4.V.2010 (CRB). Citations PAPHOS : Akamas Peninsula, Agios Minas None. Church, 1 ♂ 1 ♀, stones in grassland, 3.IV.1997 (CJB); Akamas Peninsula, Avakas Gorge, 1 ♂, New records under stone, 27.III.1998, coll. J. Murphy FAMAGUSTA : Agios Sergios , 1 ♂ 1 ♀, pitfalls (MMUE); Akamas Peninsula, near Agios Minas in Pinus plantation, II. - V.2011 (CRB); Cape church, 1 ♀, under sto nes in meadow with tall Greco, Ayia Napa, 35m, 1 ♂, under stones in grass, 2.V.2007 (CRB); Akamas Peninsula, Baths phrygana, 27.IV.2010 (CRB); Karpasia Peninsula, of Aphrodite, 1 ♂ 1 ♀, stones in rock strewn Apostolos Andreas Monastery, 50m, 2 ♀♀, under grassland, 3.IV.1997 (CJB); Akamas Peninsula, stones in wasteland, 16.V.2011 (CRB); Salamis between Cape Drepanum and Lara Bay, 1 ♂, ruins, 30m, 1 ♂, under stones in ruins, 17.III.2011 stones in rock strewn maquis near beach, (CRB); 2.IV.1997 (CJB); Akamas Peninsula, Lara Bay, 2 KERYNEIA : Agios Georgios, 1 ♀, under stone in ♀♀, stones in rock strewn beac h, 3.IV.1997 coastal scrub, 21.III.1998, coll. J. Murphy (CJB); Akamas Peninsula, Smigies, 352m, 1 (MMUE); Panagia Apsinthiotissa Monastery ♂,m aquis, 12.III.2018 (CKHK); Asprokremmos ruin s, 350m, 1 ♀, stones in clearing in Pinus Dam, N. Kouklia, 150m, 1 ♀, herbs and stones forest, 24.V.2011 (CRB); Pentadactylos Range, along lake, 29.IV.2010 (CRB); Drouseia, 539m, 1 Lapithos S ., 800m, 1 ♀, under stones in Pinus ♂, in p hrygana, 23.III.2018 (CKHK); Kakopetria forest, 17.V.2011 (CRB); Saint Hilarion Castle, 2 S., Platania fores t station, 1100m, 1 ♂ 1 ♀, under km W., 610m, 2 ♂♂, under stones in open Pinus stones in dense Pinus forest, 4.V.2010 (CRB); forest, 20.III.2011 (CRB); N. Vouni; 250m, 1 ♂, Kato Akourdaleia, 1 ♂, in guest house swimming under stones in ruins, 20.III.2011 (CRB). pool, 3.V.2007 (CRS); Kouklia - Pissouri road, LARNACA : Larnaca SW, Hala Sultan Tekke 150m, 3 ♂♂, stony slope near sea, 22.III.2011 Mosque, 5m, 1 ♀, under stones and litter (CRB); Makounta, 4 ♀♀, under stones in scrub, bordering salt lake, 16.V.2010 (CRB); Mosfiloti, 19.III.1998, coll. J. Murphy (MMUE); Kinousa, 1 1 ♀, under stone in open pine woodland, ♂, under stone in pine wood, 19.III.1998, coll. J. 12.IV.2017 (CAR - S). Murphy (MMUE); Neo Chorio, 2 ♂♂, under LIMASSOL : Agros SW, 950m, 1 ♂, under stones stone in field, 17.III.1998, 2 ♂♂, 30.III.1998, coll. in Pinus forest, 5.V.2010 (CRB); Alassa SE, near J. Murphy (MMUE); Paphos, Bay at hotel river Limnatis, 1 ♀, under stones in dry grassland, “Paphian Bay”, 1 ♂ 1 ♀, sieving compost heap 5.V.2007 (CAR - S), 3 ♂♂, under pebbles and in near hotel, 4.IV.1997 (CJB); Paphos NW, 2 ♀♀, grass, 5.V.2007 (CRS); Amathus, 1 ♂, under under stones at the edge of a large sand pit, stones in dry valley, 6.IV.1986, 1 ♂, 7.IV.1 986, 1 24.IV.1986 (NHMW); Polis W., Latsi Botanical ♂ 3 ♀♀, on house walls, 9.IV.1986 (NHMW); Garden, 60m, 1 ♂, night catch, 22.IV.2017, S. Erimi, 1 ♀, under stones along dry riverbed, Huber leg. (CSH); Pyrgos, ♀, stones in dry 18.IV.1986 (NHMW); Fasoula, 1 ♀, olive garden gra ssland, 15.IV.1986 (NHMW); Smigies, 2 ♂♂ Spiders of Cyprus 56

1 ♀, under stones in hillside scrub, 26.III.1998, Iden tification coll. J. Murphy (MMUE); Vretsia W., near Levy, 2004: 8, figs 20 - 26. The species is closely Kelefos bridge, 650m, 1 ♀, under stones, related to D rassodes lapidosus but differs in the 19.V.2011 (CRB). less darkened chelicerae with three, closely set teeth in the promargin, widely separated in D. Distribution: Palaearctic. New to Cyprus. lapidosus .

Drassodes lute scens (C. L. Koch, 1839) Citations None. Identification Drassodes lutescens; Chatzopoulou & Chatzaki, New reco rds 2009: 352, figs 3 - 24. FAMAGUSTA: Rizokarpaso , 1 ♂, Juniperus phoenicea forest, 12.III.2016 (CMH). Citations KERYNEIA : Pentadactylos Range, Ftericha S ., None. 800m, 1 ♂, under stones in grassland, 17.V.2011 (CRB); Pentadactylos Range, Lapithos S ., 800m, 1 New records ♂, under stones in Pinus forest, 17.V.2011 (CRB). KERYNEIA : Pentadactylos Range, Lapithos S ., LIMASS OL: Agros, 1 ♀, Pinus brutia forest, 800m, 1 ♀, under stones in Pinus forest, 20.V.2016 (CMH). 17.V.2011 (C RB). NICOSIA: Pano Amiantos, 3 ♂♂ 1 ♀, pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH); LIMASSOL : Mount Olympos (Chionistra), road Pedoulas - Lemithou, pitfalls in Pinus nigra 1900m, 2 ♂♂ 1 ♀, pitfalls in Pinus forest, and P. brutia forest, 6.IV. - 8.VI.2014 (CMH). XI.2010 - III.2011 (CRB). NICOSIA: Idalion Archaeological Site, Dali, 1 ♀, Distribu tion : Israel, Lebanon. New to Cyprus. under stones in phrygana, 1.V.2013 (CAR - S). Drassyllus cyprius Chatzaki & Russell - Smith, Distribution : Mediterranean to Central Asi a. 2017 New to Cyprus. Drassyllus cyprius Chatzaki & Russell - Smith 2017: 244. Drassodes serratichelis (Roewer, 1928) Identificaion Identification Chatzaki & Russell - Smith 2017: 244, figs 16, 18 – Chatzaki, Thaler & Mylonas, 2002b: 618, figs 40 - 20, 23, 25 −26. 43. Citations Citations PA PHOS: Kinousa (Chatzaki & Russell - Smith None. 2017).

New records New records FAMAGUSTA : Agios Sergios , 30m, 1 ♀, in KERYNEIA : Agios Georgios, 1 ♀ , under stones pitfalls in Pinus plantation, 15 - 27.V .2011 in coastal scrub, 21.III.1998 (CJM). (CRB). LARNACA: Agios Minas, 1 ♂, in forest. PAPHOS : Mavrokolympos Reservoir, NE 20.III.1998 (CMA), 4 ♂♂ 3 ♀♀ , under stones on mountain pass, 26.II I.1 998 (CMA); Delikipos, 1 Potima, 150m, 1 ♀, under stones along lake, 22.V. ♀, u nder stones & in grass around small dam, 2011 (CRB). 18.IV.2017 (CAR - S). LIMASSOL: Near river S.E. of Alassa, 1 ♀, on Distribution : Crete, Kos, Karpathos, Israel, and at base of limestone rockface of gorge, Ukraine, USA. New to Cyprus. 5.V.2007 (CAR - S); Pano Platres, track near Caledonia Waterf alls, 1 ♀, under stones and in Drassodes unicolor (O. Pickard - Cambridge, litter, edge of P. nigra forest. 26.IV.2007 (CAR - 1872) S). NICOSIA: Cedar Valley Forest, 1050m, 1 ♀, stones in Cedrus forest, 19.V.2010 (CRB), 1 ♂ 2 ♀♀, under stone by track in forest,

Spiders of Cyprus 57

29.IV.2007 (CRS), 1 ♀ (CMA); between ♂, pitfalls in Pinus nigra and P. brutia f orest, IV. Cedar Valley and Stavros tis Psokas, 1 ♀, - VII.2017 (CMH). under stones by track in P. nigra forest LIMASSOL: Amathus, 1 ♂, under stones in dry 29.IV.2007 (CRS); below Machairas valley, 6.IV.1986 (NHMW); Mount Olympos Monastry, 3 ♀♀, under stones beside track in (Chionistra), just below summit, 1 ♂, under large phrygana. 4.V.2013 (CAR - S); road Pedoulas - rocks in open Pinus nigra forest, 27.IV.2007 Lemithou, 1 ♂, pitfalls in Pinus nigra and P. brutia forest, 8.VI.2014, 3 ♂♂ 2 ♀♀, IV. - (CAR - S). VII.2017 (CMH). PAPHOS: near Souskiou, 1 ♂, under s tone on PAPHOS: Agio s Minos picnic site, 2 ♂♂ 1 ♀, rocky eroded grassland on hill, 17.IV.2017 (CRS). u nder stones in open pine woodland, 13.IV.2017 (CAR - S); Akamas Peninsula, 2 ♀♀ , under stones Distribution: Turkey, Israel. New to Cyprus. in coastal scrub, 22.III.1998 (CJM); Akamas Peninsula, ca. 2 km S of Point Geranisos, 1 ♀, Drassyllus pusillus (C. L. Koch, 1833) un der stones in dry phrygana, 3.V.2007 (CAR - S); Akamas Peninsula, Avakas Gorge, 1 ♀, under Identification stone in gorge, 27.III.1998 (CJM); Akamas Roberts, 1985: 72, figs 26c . Peninsula, Smigies, 352m, 4 ♂♂ 1 ♀, in m aquis, 12.III.2018 (CKHK); Cape Drepanon, 1 ♀, under Citations stone, 29.III.1998 (CJM); Evre tou Reservoir, 1 ♀, None. under stones and in grass tussocks, 13.IV.2017 (CA R - S); Fontana Amorosa, 1 ♂ 3 ♀♀, New Records 22.III.1998 (CMA); Gialia, 2 ♀♀, under stones in NICOSIA: Pi cnic site 5 km from Machairas pine woods. 25.III.1998 (CJM); Kinousa, 1 ♀, Monastery, 1 ♀, under stone in open grassy pine under stones in pine woods, 19.III.1998 (CJM), 2 woods, 4.V.2013 (CAR - S). ♂♂ 1 ♀, 23 .III.1998 (CMA), 4 ♂♂ 9 ♀♀, 28.III.1998 (CMA), 1 ♀ , 29.III.1998 (CMA); N. Distribution: Palaearctic. New to Cyprus. Kouklia, Lake Asprogeminou, 1 ♀, 100m, stones around lake, 29.IV.2010 (CRB); Neo Chorio, 1 ♂ Echemus levyi Kovblyuk & Seyar, 2009 1 ♀, under stones in low scrub, 30.III.1998 (CJM); Panagia, 1 ♂, in litter, 29.III.1998 ( CMA); Identification Pegeia, Pikni Forest, 374m, 5 ♂♂, pitfalls in open Chatzaki et al., 2002a: 582, figs 39 - 42 ruderal site, 14.III. - 23.III.2018 (CKHK); Smigies, 2 ♂♂, under stones in hillside scrub, Cita tions 26.III.1998 (CJM), 2 ♂♂, 29.IV.2007 (CAR - S). LIMASSOL: Pano Platres, abov e Caledonia Waterfall (Chatzaki & Van Keer 2019). Distribution: An endemic species of Cyprus. New records Drassyllu s jubatopalpis Levy, 1998 None.

Identification Distribution: Turkey, Cyprus. Levy, 1998c: 154, figs 133 - 137 figs 17 - 19. Gnaphosa bithynica Kulczyński, 1903 Citations None. Identification Chatzaki et al., 2002a: 582, figs 39 - 42 New records FAMAGUSTA : Komi Kebir , 150m, 1 ♂, pitfalls Citations None. in Pinus forest, III. - V.2011 (CRB).

LARNACA : Larnaca salt lake, 10m, 2 ♀♀, New records p itfalls around lake, III. - V.2011 (CRB); LIMASSOL : road Chionistra - Prodromos, 2 ♂♂, Stavrovouni Monastery W., 1 ♀, pitfalls in Pinus pitfalls in Pinus nigra forest, 6.IV. - 8.VI.2014 forest, III. - V.2010 (CRB). (CMH); Mount Olympos (Chionistra), just below NICOSIA: Dali, 1 ♀, under stones in riverbed, summit, 1 ♀, under large rocks in open Pinus 30.IV.2007 (CAR - S); road Pedoulas - Lemithou, 1 nigra forest, 27.IV.2007 (CAR - S); M ount Spiders of Cyprus 58

Olympos (Chionistra),1 ♂, pitfalls in open Citations Juniperus foetidissima forest, 6.IV. - 8.VI.2014 FAMAGUSTA : Karpasia Peninsula, Apostolos (CMH); road Pedoulas - Prodromos, 5 ♂♂, pitfalls Andreas Monastery (Bosmans et al. 2018); in Pinus nigra forest, 6.IV. - 8.VI.2014 (CMH); Flamoudi W . (Bosmans et al. 2018). near Prodromos reservoir, 3 ♂♂, pitfalls in Pinus LARNACA : Kornos picnic site (Bosmans et al. nigra forest, 6.IV. - 8.VI.2014 (CMH); Troodos 2018). Square, 3 ♂♂, pitfalls in open, mixed P. nigra and PAPHOS : Akamas Peninsu la, Agios Minas ; J. foetidissima forest, 6.IV. - 8.VI.2014 (CMH). Akamas Peninsula, Cape Geronnisos S., Mavrokolymnos Dam, Potima NE (Bosmans et al. Distribution: Crete, Turkey. New to Cyprus, 2018). where it w as only found in the forests on Mount Troodos. New records NICOSIA: A gridi, nea r Dali, 1 ♂, on veranda, Gnaphosa dol osa Herman, 1879 V.2009, 1 ♂, in house, 2.II.2010, 1 ♂, in house, Gnaphosa barroissi ; Kulczyński, 1908: 53. 21.II.2011, 1 ♀, under stone, river bank, 29.IV.2013 (CDM). Identification : https://araneae.nmbe.ch/ . Distribution: Egypt, Israel, Azerbaijan, Cyprus. Citations CYPRUS Haplodrassus omissus (O. Pickard - Cambridge, Without precise locality (Kulczyński, 1908). 1872)

New records Identification LARNACA : Lar naca salt lake, 2 ♂♂, pitfalls in Bosmans et al. 2018: 25, figs 87 - 97, 108 - 112. salt marsh, III. - V.2011 (CRB). Citations Distribution: Palaearctic. FAMAGUSTA: Cape Greko (Bosmans et al. 2018). Haplodrassus dalmatensis (L. Koch, 1866) NICOSIA: Lakatamia (Bosmans et al. 2018).

Identification : https://araneae.nmbe.ch/ . New records: None. Citations None. Distribution: Circum - Mediterranean. New to Cyprus. New records LIMASSOL: Panagia Glossa Monastery near Haplodrassus pseu dosignifer Marusik, Hipppa Kellaki, 1 ♀, under stones in cultivated land, & Kopponen, 1996 12.IV.1986 (NHMW); Pissouri, 1 ♂ 1 ♀, under stones, 24.III.1998, coll. J. Murphy (MMUE). Identification PAPHOS: Makounta, 1 ♂ 1 ♀, under stones in Marusik et al. 1996, figs 63 - 70. scrub, 19. III.1998, coll. J. Murphy (MMUE); Pegeia, Pikni Forest, 374m, 2 ♂♂, pitfalls in open ruderal site, 14.III. - 23.III.2018 (CKHK). Citations None. Distribution: Palearctic, Ethiopia. New to Cyprus. New records LIMASSOL: road Chionistra - Prodromos, 2 ♂♂, Haplodrassus invalidus (O. Pickard - pitfalls in Pinus nigra forest, 6.IV. - 8.VI.2014 Cambridge, 1872) (CMH); Pano Amiantos, 1 ♀, pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH); Troodos Identificat ion Square, 1 ♀, pitfalls in open, mixed P. nigra and Bosmans et al. 2018: 54, figs 223 - 231. J. foetidissima fore st, 6.IV. - 8.VI.2014 (CMH). PAPHOS: Mavrokolympos Reservoir, 1 ♀, 13.III.2018CKHK (CKHK); Pegeia, Pikni Forest, 366m, 1 ♂ 1 ♀, pitfalls in pine forest, 11.III. -

Spiders of Cyprus 59

23.III.2018 (CKHK); Pegeia, Pikni Forest, 374m, Identification 10 ♂♂, pitfalls in open ruderal site, 14.II I. - Chatzaki, Thaler & Mylonas, 2003: 73, figs 82 - 23.III.2018 (CKHK). 86.

Comments Citations Our specimens fit completely the description of NICOSIA: Pissouri (Platnick & Shadab 1983). H. pseudosignifer by Marusik et al. (1996). Distribution: Ukraine, Russian, Iran. New to New records Cyprus. FAMAGUSTA : Gialousa E ., Agios Thyr sos, 20m, 1 ♂, beaten from shrubs, 16.V.2011 (CRB). Haplodrassus signifer (C. L. Koch, 1839) NICOSIA : Agios Sozomenos , near Dali, 1 ♀, under stones in dry grassland, 30.IV.2007 (CAR - Identification : https://araneae.nmbe.ch/ . S), 1 ♂ (CRS); Agridi, ne ar Dali, 1 ♀, in garden, V.2011, 1 ♀, in garden, IV.2012 (CDM); Elia - Citations Angolemi road, 100m , 2 ♂♂, under stones in FAMAGUSTA: Komi Kebir ( Bosmans et al. grassland, rubbish in dry riverbed, 18.V.2011 2018); Agios Sergios ( Bosmans et al. 2018). (CRB); Marathasa Dam, 1 ♂ 2 ♀♀, pitfalls in KERYNEIA : Kalo Chorio (Bosmans et al. 2018); lake side vegetation, 7.V. - 10.VII.2016 (CMH); Vouni ruins (Bosmans et al. 2018). Masari S., Merika r iver, 120m, 1 ♀, under stones PAPHOS: Kouklia - Pissoura road (Bosmans et al. in wasteland, 24.V.2011 (CRB); Xerargaka S, 2018). Or kontas, 1 ♂, riversid e vegetation, 26.III.2016 NICOSIA:Prodromos E., near Prodromos dam (CMH). (Bosmans et al. 2018). LIMASSOL: Pissouri Beach, 1 ♂ 1 ♀, 27.IV.1982. coll. A. Stubbs (MMUE). New records PAPHOS : Kato Paphos archaeological site, 1 ♂, LIMASSOL: Alassa SE, near river Limnatis, 1 ♀, under stones in grass, 10.V.2007 (CAR - S). under stones in dry grassland, 5.V.2007 (CRS); Pano Platres, valley be low Caledonia Waterfall, 1 ♀, under stones, 26.IV.2007 (CRS); Pissouri, 1 ♀, Distribution under stones, 24.III.1998, coll. J. Murphy Mediterranean, Cana ry Is., Burkino Faso, USA. (MMUE); ca. 5 km S. of Troodos, road to Pano Platres, 1 ♀, under stones in P. nigra forest, Lasophorus zografae Chatzaki, 2018 1.V.2007 (CRS). NICOSIA : Kapouti , 100m, 1 ♀, under stones in Identification Chatzaki 2018: 532, figs 27 - 28, 31 - 40. grassland, 20.III.2011 (CRB).

PAPHOS : Akamas Peninsula, 2 ♀♀, under stones Citations in coastal scrub, 22.III.1998, coll. J. Murphy None. (MMUE); Baths of Aphrodite, 1 ♀, under stone in coastal scrub, 18.III.1998, coll. J. Murphy New records (MMUE); Akamas Peni n sula, Smigies, 352m, 1 ♂ LIMASSOL: road Pedoulas - Lemithou, 1 ♂ 2 ♀♀, 1 ♀, m aquis, 12.III.2018 (CKHK); Drouseia S., pitfalls in Pinus brutia forest, IV. - VII. 2017 Panagia Chrysospiliotissa, 580m, 1 ♂, night catch (CRB). in small cave, 22.IV.2017, S. Huber leg. (CSH); Makounta, 1 ♂ 2 ♀♀, under stones in scrub, Distribution: Northern Gr eece, Dodekanese. 19.III.1998, coll. J. Murphy (MMUE); Kinousa , 1 New to Cyprus ♂ 1 ♀, under stone in pine wood, 23.III.1998, 4 ♂♂ 4 ♀♀, 28.III.1998, coll. J. Murphy (MMUE); Leptodrassus femineus (Simon, 1873) Neo Chorio, 1 ♂ 2 ♀♀, under stone s in field, 20.III.1998, 1 ♀, 30.III.1998, coll. J. Murphy Identification (MMUE). Chatzaki, Thaler & Mylonas, 2002a: 590, figs 66 - 67, 73. Distribution: Holarctic. Citations Heser nilicola (O. Pic kard - Cambridge, 1874) None. Zelotes nilicola ; Platnick & Shadab, 1983: 186.

Spiders of Cyprus 60

New records Micaria guttulata (C. L. Koch, 1839) KERYNEIA : Panagia Ap sinthiotissa Monastery ruins, 350m, 1 ♀, under stones in clearing in Identification : https://araneae.nmbe.ch/ . Pinus forest, 24.V.2011 (CRB). Citations LARNACA : Achna Reservoir, 50m, 1 ♀, under None. stones bordering lake, 16.V.2011 (CRB), 1 ♀, 27.V.2011 (CRB). New records NICOSIA: Masari S., Merika River, 120m, 1 ♀, PAPHOS: Akamas Peninsula, Avakas Gorge, 1 und er stones in wasteland, 24.V.2011 (CRB). ♀, under stone, 27.III.1998, coll. J. Murp hy PAPHOS: Mavrokolympos Reservoir, 1 ♀, C. (MMUE). Saure leg., 20.VI.2013 (CKHK). Distribution: Palaearctic. New to Cyprus.

Distribution: Mediterranean. New to Cyprus. Micaria ignea O. Pickard - Cambridge, 1872 Micaria todilla Simon, 1878: 28. Micaria coarctata (Lucas, 1846) Identification Identification : https://araneae.nmbe.ch/ . Bosmans & Blick, 2000: 477, figs 5 - 8.

Citations Citations None. Without precise locality (Simon, 1878).

New records New records FAMAGUSTA : Flamoudi W ., 25m, 1 ♂, under NICOSIA: Lakatamia, 1 ♀, on a stone in olive stones and phrygana near the beach, 25.V.2011 grove, 1.I. - 1.X. 2016 (CMH). (CRB); Karpasia Peninsula, Cape Apostolos Andreas, 70m, 1 ♀, under stones in grassland, Distribution: Mediterranean to Uzbekistan. 16. V.2011 (CRB). LARNACA : Dipotamos Reservoir, 150m, 1 ♀, Micaria pallipes (Lucas, 1846) herbs and stones bordering lake, 28.IV. 2010 Micaria pallipes ; Chatzaki & Russell - Smith 2012 . (CRB). NICOSIA: near Platania Scout camping site, 1 ♂, pitfalls in Pinus brutia forest, IV. - VII.2017 Identification (CMH). Bosmans & Blick, 2000: 4 49, figs 9 - 12.

PAPHOS: Akamas Peninsula, Avakas Gorge , 1 Citations ♀, under stone, 27.III.1998, coll. J. Murphy LIMASSOL: Fassouri reed beds (Chatzaki & (MMUE); Kinousa, 1 ♂ 1 ♀, under stone in pine Russell - Smith 2012). wood, 28.III.1998, coll. J. Murphy (MMUE).

New records Distribution : Mediterranean to Central Asia. LIMASSOL: road Kaminaria - Lemithou, 1 ♀, New to Cyprus. pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014

(CMH). Micaria dives (Lucas, 1846)

Distribution: Identification : https://araneae.nmbe.ch/ . Mediterranean to Centr al Asia.

Citations Nomisia excerpta (O. Pickard - Cambridge, 1872 None. Nomisia excerpta ; Chatzaki et al ., 2010: 6.

New records Identification LIMASSOL : Alassa SE, near river Limnatis, 1 ♂, Chatzaki 2010: 6, figs 9 - 23. under stones in dry grassland, 5.V.2007 (CAR - S). Citations Distribution: Palaearctic. New to Cyprus. Without precise locality (Chatzaki et al., 2010).

Spiders of Cyprus 61

New records Citations FAMAGUSTA: Protaras, Profitis Elias, 1 ♀, 2005 None. (CDM). KERYNEIA: Agios Georgios, 1 ♀, under stone in Ne w records coastal scrub, 21.III.1998, coll. J. Murphy FAMAGUSTA : Gialousa E ., Agios Thyrsos, (MMUE). 20m, 2 ♂♂, under stones, 16.V.20111 (CRB); LARNACA: Mosfiloti, 1 ♂, under stone in dry Kantara E., 610m, 1 ♀, under stones in open eroded grassland on hillside, 12.IV.2017 (CAR - Pinus forest, 25.V.2011 (CRB); Rizokarpaso , 5 S). ♂♂, Juniperus phoenicea forest, 12.III.2016 NICOSIA: Agios Sozomenos , near Dali, 1 ♀, (CMH). under stones in disturbed phrygana, 30.IV.2007 KERYNEIA : Pentadact ylos Range, Saint Hilarion (CAR - S); Agridi, near Dali, 1 ♀, in field IV.2007 Castle, 600m, 1 ♀, under stones, 17.V.2011 (CDM); Dali, Idalion Archaeological Site, 2 ♂♂, (CRB). under stones in phrygana, 1.V.2013 (CAR - S), 1 LARNACA : Kornos picnic site, 350m, 2 ♀♀, ♂, same site, 1.IV.2012, 1 ♂, under st ones in under stones in Pinus forest, 28.V.2011 (CRB). phrygana, 30.IV.2013 (CDM); Lympia Reservoir, LIMASSOL: road Chionistra - Prodromos, 16 ♂♂, Lympia, 1 ♂, in clump of rushes below dam wall, pitfalls in Pinus nigra forest, 6.IV. - 8.V I.2014 9.V.2013 (CDM); below Machairas Monastery, 1 (CMH); road Kaminaria - Lemithou, 2 ♂♂ 1 ♀, ♂, under stones beside track in phrygana, pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 4.V.2013 (CAR - S). (CMH); Kourion archaeological site near PAPHOS : Akamas Peninsula, Agios Minas, 1 ♀, Episkopi, 2 ♀♀, under stones, 9.V.2007 (CAR - S); in thick pine litter, 2.V.2007 (CAR - S); Dhiarizos Troodos Square, 1 ♀, pitfalls in open, mixed P. river near Kelefos bridge, 444m, 1 ♀, open river nigra and J. foetidi ssima forest, 6.IV. - 8.VI.2014 bank, 16.III.2018 (CKHK). (CMH). NICOSIA: Agios Sozomenos, near Dali, 3 ♀♀, Distribution: Canary Islands, Tunisia, Greece, under stones in dry, rocky phrygana, 20.IV.2017 Israel and Cyprus. (CAR - S); Cedar Valley Forest, 1180m, 1 ♂, under stones in in Cedrus forest, 19.V.2011 (CRB); Nomisia exornata (C. L. Koch, 1839) between Cedar Valley and Stavr os tis Psokas, 1 ♂, under stones in P. nigra forest, 29.IV.2007 Identification : https://araneae.nmbe.ch/ . (CAR - S); Chrysoroyiatissa Monastery, 1 ♂, 2.V.1982. coll. A. Stubbs (MMUE); Dali, 1 ♀, Citations under stones in grassland, 30.IV.2007 (CAR - S); None. Elia - Angolemi road, 100m, 2 ♀♀, under stones in grassland, 18.V.2011 (CRB); Limnitis, 1 ♂, Pinus New records brutia forest, 13.V.2016 (CMH); Marathasa Dam, LARNACA: Cape Kiti, 1 ♀, under stones below 5 ♂♂ 1 ♀, pitfalls in riverside vegetation, 7.V. - coastal cliffs, 25.IV.1982, coll. A. Stubbs 10.VII.2016 (CMH); road Pedoulas - Lemithou, 3 (MMUE). ♂♂, pitfalls in Pinus nigra and P. brutia forest, LIMASSOL : Mouttagiaka, 20m, 1 ♂, in 6.IV. - 8.VI.2014 (CMH); road Pedoulas - swimming p ool of Caravel hotel, 29.IV.2010 Prodromos, 2 ♂♂, pitfalls in Pinus nigra forest, (CRB). 6.IV. - 8.VI.2014 (CMH); near Platania Scout camping site, 3 ♂♂, pitfalls in Pinus brutia forest, Distribution: Europe to Central Asia. New to 6.IV. - 8.VI.2014 (CMH); Prodromos E., near Cyprus. Prodromos Dam, 1550m, 1 ♂, under stones in Pinus f orest, 23.V.2011 (CRB); near Prodromos Nomisia recepta (Pavesi, 1880) reservoir, 5 ♂♂, pitfalls in Pinus nigra forest, Nomisia recepta ; Chatzaki, 2010: 153. 6.IV. - 8.VI.2014 (CMH); Xerargaka S, Orkontas, 2 ♀♀, riverside vegetation, 26.III.2016 (CMH); Identification PAPHOS : Kato Paphos archaeological site, 3 ♂♂ Chatzaki 2010: 15, figs 40 - 46. 2 ♀♀, under stones in gra ssland, 10.V.2007 (CAR - S); Mavrokolympos Reservoir, 1 ♀, C. Saure leg., 20.VI.2013 (CKHK). Spiders of Cyprus 62

Distribution: Algeria, Tunisia, Sicily, Malta, Phaeocedus hebraeus Levy, 1999 Greece. New to Cyprus. Identification Nomisia ripariensis (O. Pickard - Cambridge, Levy, 1999: 429, figs 1 - 5. 1872) Citations Identification None. Chatzaki 2010: 21, figs 47 - 50. New records Citations LARNACA: Cape Gkreko, Ayia Napa, 35m, 1 ♂ None. 1 ♀ , under bark of Pinus , 2 7.IV.2010 (CRB) 5.V.2010 (CRB); Marathassa Dam, 550m, 1 ♂, New records under stones around lake, 20.V.2011 (CRB). FAMAGUSTA: Karpasia Peninsula, Apostolos Andreas Monastery, 50m, 2 ♂♂ 1 ♀, under stones Distribution: Israel, Cyprus. in wasteland, 16.V.2011 (CRB). Poecilochroa senilis (O. Pickard - Cambridge, LARNACA : Larnaca SW, Hala Sultan Tekke 1872) Mosque, 5m, 1 ♂, litter bordering salt la ke, 16.V.2011 (CRB). Identification NICOSIA: Agios Sozomenos , near Dali, 2 ♂♂ 1 Chatzaki, Thaler & Mylonas, 2002a: 56 9, figs 8 - ♀, under stones in dry grassland, 30.IV.2007 11; Kovblyuk & Tuneva, 2009: 157, figs 1 - 9. (CAR - S); Dali, 1 ♀, under stones in grassland, Citations 12.V.2007 (CAR - S); Elia - Angolemi road, 100m, None. 1 ♀, under stones in grassland, 18.V.2011 (CRB); Eri mi, 1 ♂, under stones along dry riverbed, New records 21.IV.1986 (NHMW); Masari S., Merika river, LARNACA : Cape Kiti, 1 ♂ 2 ♀♀, under stones 120m, 1 ♀, under stones in wasteland, 24.V.2011 below coastal cliffs, 25.IV.1982, coll. A. Stubbs (CRB). (MMUE); Larnaca salt lake, 5m, 1 ♂, in litter in PAPHOS : Akamas Peninsula, Neo Chorio, 1 ♂, at salt marsh, 27.IV.201 0 (CRB). picnic site , under stone in phrygana, 2.V. 2007 LIMASSOL: Akrotiri Peninsula, edge of central (CRS); Kidasi N., al ong Diarizos River, 300m, 1 salt pan, 1 ♂, under stones and debris in ♂, under stones, 21.V.2011 (CRB); Salicornia , 27.IV.2007 (CRS).

Mavrokolympos Reservoir, NE Potima, 150m, 1 Distribution: East Mediterranean, from Corsica ♀, under stones along lake, 22.V.2011 (CRB). to Turkmenistan. New to Cyprus.

Distribution : Greece to Azerbaijan. New to Poecilochroa taborensis Levy, 1999 Cyprus. Identification Phaeocedus braccatus (L. Koch, 1866) Lissner & Chatzaki 2016: 17, figs 1, 2A - C, 3A - D, 4A - E, 5A - C, 6A - D. Ident ification : https://araneae.nmbe.ch/ . Citations Citations NICOSIA: Dali, Idalion Archaeological Site None. (Lissner & Chatzaki 2016).

New records New records NICOSIA: Kapedes S., 600m, 1 ♂, under stones NICOSIA: Alona, 1 ♀, mixed forest, 27.II.2016 along rivulet, 5.V.2010 (CRB); Marathassa Dam, (CMH). 550m, 1 ♂, under stones around lake, 20. V.2011 (CRB). Distr ibution: Portugal, Greece, Israel, Cyprus.

Distribution: Palaearctic. New to Cyprus.

Spiders of Cyprus 63

Poecilochroa sp. Identification Chatzaki et al., 2002a: 580, figs 43 - 46 Poecilochroa sp.; Chatzaki & Russell - Smith 2017: 242, figs 13 - 15. Citations PAPHOS : Avakas Go rge (Logunov, 2013). Citations LIMASSOL: Akrotiri, Fassouli reed beds New records (Chatzaki & Russell - Smith 2017). FAMAGUSTA : Flamoudi W., 25m, 1 subadult ♀, under stones in phrygana near the beach, Distribution: Cyprus. 25.V.2011 (CRB); Karpasia Peninsula, Apostolos Andreas Monastery, 50m, 2 ♀♀, under stones in Prodidomus amaranthinus (Lucas, 1846) wasteland, 16.V.2011; Karpasia Peninsul a, Cape Apostolos Andreas, 70m, 1 ♂, 2 ♀♀, under stones Identification in grassland, 16.V.2011 (CRB); Paralimni Lake, Pérez & Blasco, 1986: 156, figs 1 - 6. 65m, 1 ♀, litter along dry lake, 21.XI.2010 (CRB); Protaras, Profitis Elias, 1 ♂, 2005 (CDM); Citations Salamis ruins, 30m, 1 subadult ♀, under stones in None. ruins, 17 .III.2011 (CRB). KERYNEIA : Panagia Apsinthiotissa Monastery New records ruins, 350m, 1 ♀, under stones in clearing in FAMAGUSTA: Cape Greco, 1 ♀, under stones in Pinus forest, 24.V.2011 (CRB); Saint Hilarion phrygana, 3.V.2013 (CAR - S). Castle W, 600m, 1 ♂, under stones in Pinus forest NICOSIA : Agios Sozomenos, near Dali, 1 17.V.2011 (CRB). juvenile, under stones in dry grassland, LARNACA : Achna Reservoir, 50m, 2 ♀♀, under 30.IV.2007 (CAR - S), 1 ♂ (CRS). stones and litter bordering lake, 16.V.2011 PAPHOS : Akamas Peninsula, Agios Minas, 1 ♂, (CRB); Dipotamos Reservoir, 150m, 1 ♀, under in thick pine litter, 2.V.2 007 (CAR - S); Baths of stones bordering lake, 28.IV.2010 (CRB); Aphrodite , 1 ♂, under stones in tall grass in Larnaca SW, Hala Sultan Tekke mosque, 50m, 1 meadow, 15.IV.2017 (CAR - S); ♂, pitfalls in salt marsh, 15 - 28.V.2011 (CRB). near Avakas Gorge, 1 ♂, under stone in field with LIMASSOL: Alassa S. E., near river Limnatis, 1 ♂ 1 ♀, under stones in dry grassland, 4.V.2007 tall grass, 14.IV.2017 (CAR - S); Kato Paphos (CRS); Pano Platres, valley below Caledonia archaeological site, 1 ♂, under stones in grass, Waterfall, 1 ♀, under stone, 26.IV.2007 (CRS); 10.V.2007 (CAR - S). Troodos, Mount Olympos (Chionistra), 1900m, 1 ♀, under stones in Pinus forest, 23.V.2011 Distribution: Mediterranean. New to Cyprus. (C RB); Mount Olympos (Chionistra), 1830m, 1 ♀, pine forest with open places, 23.II.2011, D. W. Pterotricha kochi (O. Pickard - Cambridge, Wrase leg. (CKHK); Troodos, 1 ♀, under rocks, 1872) 30.XI.2008 (CDM); Fasoula, 1 ♀, olive garden with dry stone walls, 16.IV.1986 (NHMW). Identification NICOSIA: Agios Sozomenos , nea r Dali, 2 ♂♂ 3 Levy, 1995: 967, figs 121 - 125. ♀♀, under stones in dry grassland and in disturbed phrygana, 30.IV.2007 (CAR - S); Agridi, Citations near Dali, 3 ♂♂ 1 ♀, in house, V.2009, 1 ♂, in None. grass in lawn, 1.VII.2010, 1 ♂ 1 ♀, in garden, New records V.2011 (all CDM); Cedar Valley Forest, 1 ♂, NICOSIA: Agridi, near Dali, 1 ♂, in grass in yard, under stones b y track, 29.IV.2007 (CAR - S); Dali, IV.2009 (CDM ). 13, Agios Andronikos, 1 ♂ 1 ♀, in garden, V.2008 (CDM); Dali, 2 ♀♀, under stones in Distribution: Turkey, Lebanon, Syria, Israel. riverbed, 12.V.2007 (CRS); Elia - Angolemi road, New to Cyprus. 100m, 1 ♀, under stones in grassland, 18.V.2011 (CRB); Kakopetria S., Platania forest stat ion, Pterotricha lentiginosa (L. Koch, 1837) 1100m, 1 ♀, under stones in dense Pinus forest, Pterotricha lentiginosa ; Logunov, 2013: 5 . 4.V.2010 (CRB); Lympia Reservoir, Lympia, 1 ♀, in clump of rushes b elow dam wall, 9.V.2013

Spiders of Cyprus 64

(CAR - S), 1 ♂, (CDM); Panagia Glossa Monastery Citations near Kellaki, 1 ♀, under stones in cultivated land, None. 12.IV.1986 (NH MW); Oikos, 700m, 1 ♂, limy slope, 23.IV.2017, S. Huber leg. (CSH); Pyrgos, 1 New records ♀, d ry grassland, 15.IV.1986 (NHMW). LARNACA : Stavrovouni Monastery, 350m, 1 ♀, PAPHOS : Akamas Peninsula, 1 ♀, under stones under bark of Pinus trees, 28.IV.2010 (CRB). in coastal scrub, 22.III.1998, coll. J. Murphy (MMUE); Akamas Peninsula, Avakas Gorge, 1 ♀, Distribution : Greece, Israel. New to Cyprus. un der stone, 27.III.1998, coll. J. Murphy (MMUE); Akamas Peninsula, Avakas gorge, Scotophaeus scutulatus (L. Koc h, 1866) 50m, 1 ♀, litter in citrus orchard at entrance of gorge, 30.IV.2010 (CRB); Akamas Peninsula, Identification : https://araneae.nmbe.ch/ . track along the sea, 10m, 1 ♂ 1 ♀, under stones in Juniperus phrygana, 30.IV.2010 ( CRB); Akamas Citations Peninsula, Cape Drepanum, 1 ♀, stones in rock None. strewn slope at sea front, 31.III.1997 (CJB); Akamas Peninsula, Cape Geronnisos S., 1 ♂ 1 ♀, N ew records under stones in dry phrygana, 3.V.2007 (CAR - S); LIMASSOL: Troodos, Persephone Trail, 1732m, Akamas Peninsula, at Neo Chorio picnic site, 2 1 ♀, in Pinus nigra forest, 19.III.2018 (CKHK). ♀♀, under stone in phrygana, 2.V.2007 (CRS); Kinousa, 1 ♂, under stone in pine wood, NICOSIA : Agridi, nea r Dali, 1 ♀, under 28.III.1998, coll. J. Murphy (MMUE); Gialia, 1 eucalyptus bark in garden, 29.IV.2013 (CAR - S); ♀, under stones in pine wood, 25.III.1998, coll. J. Cedar Valley Forest, 1180m, 1 ♂, beaten in Murphy (MMUE); Makounta, 1 ♀, under stones Cedrus forest, 19.V.2011 (CRB). in scrub, 19.III.1998, coll. J. Murphy (MMUE); PAPHOS: Gialia, 1 ♀, under stones in pine wood, Pedoulas, 960m, 1 ♂ 1 ♀, night catch in orange 25.III.1998, coll. J. Murphy (MMUE); Kelefos garden, 20.IV.2017, S. Huber leg. (CSH); Paphos bridge, 465m, decid uous wood, 1 ♀, under bark, NW, 1 ♂, under stones at the edge of a large sand 16.III.2018 (CKHK). pit, 24.IV.1986 (NHMW); Smigies, 2 ♀♀, under stones in hillside scrub, 26.III.1998, coll. J. Distribution : Europe to Central Asia, North Murphy (M MUE). Africa. New to Cyprus.

Distribution: Mediterranean to Ukraine. Setaphis carmeli (O. Pickard - Cambridge, 1872) Identification Pterotricha schaefferi (Audouin, 1826) Chatzaki, Thaler & Mylonas, 2003: 55, figs 40 - 41. Identification Levy, 1995: 964, figs 108 - 112. Citation s None. Citations New records None. FAMAGUSTA : Gialousa E., Agios Thyrsos, 20m, 1 ♀, in litter, 16.V.20111 (CRB). New records LIMASSOL : Troodos S., road to Pano Platres, 1 LIMASSOL : Akrotiri Peninsula, Ladies Mile ♂, under stones in P. nigra forest, 28.IV.2007 Beach, 1 ♂, in sand dune s, 27.IV.2007 (CAR - S). (CAR - S), 1 ♀, 1.V.2007 (CRS). PAPHOS: Akamas Peninsula, Lara Bay, 1 ♂, C. NICOSIA: Masari S., Mer ika River, 120m, 1 ♀, Saure leg., 18.VI.2013 (CKHK). under stones in wasteland, 24.V.2011 (CRB).

Distribution: Libya, Egypt, Sudan, Israel. New to PAPHOS : Akamas Peninsula, 1 ♂, under stones Cyprus. in coastal scrub, 22.III.1998 coll. J. Murphy (MMUE); Akamas Peninsula, track along the sea, Scotophaeus peninsularis Roewer, 1928 10m, 1 ♀, under stones in Juniperus phrygana, 30.IV .2010 (CRB); Neo Chorio, 1 ♂, under stone Identification : https://araneae.nmbe.ch/ . in field, 20.II I.1998, 1 ♀, 30.III.1998, coll. J.

Murphy (MMUE).

Spiders of Cyprus 65

Distribution: Circum - Mediterranean. New to Distribution: Greece. New to Cyprus. Cyprus. Talanites dunini Platnick & Ovtsharenko, 19 91 Synaphosus palearcticus Ovtsharenko, Levy & Identification Platnick, 1994 Platnick & Ovtsharenko, 1991: 117, figs 5 - 8.

Identification Citations Ch atzaki et al. , 2002b: 623, figs 50 - 52, 57 - 58. None.

Citations New records None. LIMASSOL: Akrotiri Peninsula N., 5m, 2 ♂♂, litter in Eucalyptus forest, 29.IV.2010 (CRB). New records PAPHOS : Akamas Peninsula, Agios Minas, 1 ♂, Distribution : Israel, Central Asia. New to in thick pine litter, 2.V.2007 (CAR - S); Kato Europe. Paphos archaeological site, 1 ♂, under stones in grass, 10.V.2007 (CAR - S). Trachyzelotes jaxartensis (Kroneberg, 1875)

Distribution: Central Palaearctic. New to Cyprus. Identification Platnick & Murphy, 1984: 10, figs 19 - 22. Synaphosus shirin Ovtsharenko, Levy & Platnick, 1994 Citations Chatzaki & Russell - Smith 2017: 248. None.

Identification: New records Chatzaki & Russell - Smith , 2017: 248, figs 29 - 34. NICOSIA: Marathasa Dam, 1 ♂ 2 ♀♀, pitfalls in riverside vegetation, 7.V. - 10.VII.2016. PAPHOS : Akamas Peninsula, Agios Minas, 1 ♀,

wet area by stream, 2.V.2007 (CAR - S). Citations LARNACA: Lympia R eservoir (Chatzaki & Russell - Smith 2017). Distribution : Holarctic, expanding to tropical NICOSIA: Agios Sozomenos, near Dali (Chatzaki Africa and Hawai. New to Cyprus. & Russell - Smith 2017); Agridi, near Dali (Chatzaki & Russell - Smith 2017). Trachyzelotes lyonneti (Audouin, 1826)

New records Identification LARNACA: Kornos picknick site, 350m, 1 ♂, Chatzaki et al. , 2003: 54, figs 24 - 25, 29. stones in Pinus forest, 28.V.2011 (CRB). Citations Distribution : Iran, Cyprus. None.

Synaphosus trichopus (Roewer, 1928) New records FAMAGUSTA : Prastio S., Agios Andronikos, Identification 30m, 1 ♂ 1 ♀, under stones and herbs in Chatzaki et al. , 2002b: 623, figs 53 - 54, 59 - 60. Eucalyptus forest, 27.V.2011 (CRB); Salamis S., Silver Beach, 5m, 3 ♂♂, pitfalls in salt marsh, 15 Citations - 27.V.2011 (CRB). None. LIMASSO L: Akrotiri Peninsula , 1 ♀, 27.IX.2007

(CDM). New records PAPHOS : Kato Akourdaleia, 1 ♂, in swimming PAPHOS : Kato Paphos, archaeological site, 1 ♂, pool of guest house, 3.V.2007 (CAR - S); Evretou under stones in grass, 10.V.2007 (CAR - S). Reservoir, 1 ♂, under stones in sparse grass, 13.IV.2017 (CRS).

Spiders of Cyprus 66

Distribution : Mediterranean region to Central NICOSIA: Agios Sozomenos , near Dali, 1 ♂, Asia. New to Cyprus. under stones in dry grassland, 29.IV.2013 (CAR - S); near Platania Scout camping site, 1 ♀, pitfalls Trachyzelotes malkini Platnick & Murphy, in Pinus brutia forest, IV. - VII.2017 (CMH). 1 984 PAPHOS : Avakas Gorge, 50m, 2 ♂♂, litter in Identification citrus yard at e ntrance of gorge, 30.IV.2010 Platnick & Murphy, 1984: 22, figs 51 - 54. (CRB).

Citations Distribution: Israel, Cyprus. None. Turkozelotes mccowani (Chatzaki & Russell - New records Smith, 2017) NICOSIA: Agridi, near Dali, 2 ♂♂, in yard, V.2009 (CDM). Identification Chatzaki & Russell - Smith 2017: 247, figs 27 - 28. Distribution: Romania, North Macedonia, Crete, Turkey, Ukraine, Kazakhstan. New to Cypr us. Citations NICOSIA: Agridi , near Dali (Chatzaki & Russell - Trachyzelotes stubbsi Platnick & Murphy, 1984 Smith 2017). Trachyzelotes stubbsi Platnick & Murphy, 1984: 9; Chatzaki, 2010: 48. New records Trachyzelotes cfr. stubbsi ; Logunov, 2013: 5. None.

Identification Distribution: Cyprus, Northern Greece. Platnick & Murphy, 1984: 9, figs 15 - 16. Zelotes harmeron Levy, 2009 Citations LA RNACA : Cape Kiti, coastal cliffs near Larnaca Identification airport (Platnick & Murphy, 1984). Levy, 1998c: 148, figs 120 - 122. LIMASSOL : Kourio (Chatzaki, 2010). PAPHOS: W. Polis, Akamas forest (Logunov, Citations 2013). None.

New records New records FAMAGUSTA : Gialousa E., Agios Thyrsos, FAMAGUSTA : Prastio S., Agios 20m, 3 ♂♂ 2 ♀♀, litter under shrubs, 16.V.20111 Andro nikos, 30m, 1 ♂ 1 ♀, under stones and (CRB); Prastio S., Agios Andronikos, 30m, 1 ♂, herbs in Eucalyptus forest, 27.V.2011 (CRB). under stones and herbs in Eucalyptus forest, LIMASSOL: Armenochori, 1 ♂, under stones of a 27.V.2011 (CRB); Vokolida S., Cape Elaia, 10m, dry stone wall, 14.IV.1986 (NHMW); road 1 ♀, under stones and on shrubs near the beach, Kaminaria - Lemithou, 2 ♂♂, pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH). 26 .V.2011 (CRB). NICOSIA: Marathasa Dam, 2 ♂♂ 10 ♀♀, pitfalls LARNACA: Cape Kiti, 1 ♂, under stones below in riverside vegetation, 7.V. - 10.VII.2016 coastal cliffs, 25.IV.1982, coll. A. Stubbs (MMUE); Delikipos, near Kornos, 1 ♂, in grass in (CMH). open pine wood, 9.V.2013 (CAR - S); Mosfiloti, 1 PAPHOS: N. Kouklia, Lake Asprogeminou, 1 ♀, ♀, under stones in field with tall grass, 100m, stones around lake, 29.IV.2010 (CRB). 12.IV.2017 ( C AR - S); Stavrovouni Monastery, 350m, 1 ♂, pitfalls in open Pinus forest, III - Distribution: Greece (Thrakia, Rhodos and V.2011 (CRB). Chios), Israel, Turkey . New to Cyprus. LIMASSOL: A krotiri Peninsula, edge of central salt pan, 1 ♂, under stones and debris in Zelotes limnatis Chatzaki & Russell - Smith, Salicornia , 27.IV.2007 (CDM), 1 ♂, 30.IV.2013 2017 (CAR - S), 1 ♂, 30.IV.2013 (CD M). Zelotes limnatis Chatzaki & Russell - Smith, 2017: 250.

Spiders of Cyprus 67

Id entification Identification Senglet, 2011: 523, figs 40 - 49, 72. Chatzaki & Russell - Smith, 2017: 250, figs 35, 37 – 40. Citations None. Citations LIMASSOL: Valley of river Limnatis ne ar Kouris New records Dam (Chatzaki & Russell - Smith, 2017). LARNACA : Achna Reservoir, 50m, 2 ♂♂ 2 ♀♀, under stones and in litter bordering lake, New records 16.V.2011 (CRB), 1 ♂ 1 ♀, 27.V.2011 and 6 ♂♂ LARNACA: Achna Reservoir, 1 ♂, litter along 3 ♀♀, pitfalls at same site, 15 - 27.V.2011 (CRB); the lake, 27.V.2011 (CRB). Dipotamos Reservoir, 150m, 1 ♂, herbs and Distribution: Cyprus. stones bordering lake, 28.IV.2010 (CRB); Larnaca SW, Hala Sultan Tekke mosque, 50m, 11 Zelotes metellus Roewer, 1928 ♂♂ 10 ♀♀, pitfalls in salt marsh, 15 - 28.V.2011 (CRB). Identification LIMASSOL : Akrotiri Peninsula, edge of central Senglet 2011: 555, figs 165 - 170 salt p an,1 ♂ 1 ♀, under stones and debris in Salicornia , 27.IV.2007 (CAR - S); Alassa SE, near Citations river Limnatis, under stones in dry grassland, None. 5.V.2007 (CAR - S), 2 ♂♂ 1 ♀ (CRS). NICOSIA: Athalassa National Forest Park, 2 ♂♂, New records under stones in pine plantation, 2.V.20 13; same LARNACA: Achna Reservoir, 50 m, 1 ♂, pitfalls bordering lake, 15 - 27.V.2011 (CRB). site, 4 ♂♂ 4 ♀♀, under stones and in deep eucalyptus litter, 2.V.2013; same site, 3 ♂♂ 1 ♀, Distribution: Greece, Turkey, Ukraine, S. Russia, 8.V.2013 (CAR - S); Dali, 2 ♂♂, under stones in Israel, Iran. New to Cyprus. riverbed, 12.V.2007 (CAR - S), 1 ♀, in garden, Zelotes scrutatus (O. Pickard - Cambridge, 1872) VII.2012 (CDM); Lympia, Lympia Reservoir, 2 ♀♀, in rushes below dam wall, 9.V.2013 (CAR - Identification S), same site, 2 ♂♂ (CDM); Marathasa Dam, 19 Chatzaki et al. , 2003: 69, figs 72 - 76. ♂♂ 10 ♀♀, pitfalls in riverside vegetation, 7.V. - Citations 10.VII.2016 (CMH); Pano Amiantos, 1 ♀, pitfalls None. in Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH). PAPHOS : Avakas gorge, 50m, 1 ♂, litter in citrus New records yard at entrance of gorge, 30.IV.2010 (CRB); NICOSIA : Agios Sozomeno s , near Dali, 2 ♀♀, Kato Akourdaleia, 1 ♀, in garden, VIII.2005 under stones in dry grassla nd, 30.IV.2007, 1 ♂, (CDM). 29.IV.2013 (CAR - S), 1 ♂ (CRS); Dali, 2 ♂♂, under stones in riverbed, 12.V.2007 (CAR - S); Distribution: Circum - Mediterranean, USA. New road Pedoulas - Lemithou, 3 ♂♂ 1 ♀, pitfa lls in to Cyprus. Pinus nigra and P. brutia forest, 6.IV. - 8.VI.2014 (CMH). Zelotes zekharya Levy, 2009 PAPHOS : Kidasi N., along Diarizos River, 300m, 1 ♂ 1 ♀, under stones, 21.V.2011 (CRB). Identification : Levy, 2009: 29, figs 63 - 64 ( ♂); Chatzaki & Russell - Smith, 2017: 251, figs 41 - 42 ( ♀). Distribution: Circum - Mediterranean. New to

Cyprus. Citations NICOSIA: Agridi, near Dali (Chatzaki & Russell - Smith, 2017).

New records: Zelotes tenuis (L. Koch, 1866)

Spiders of Cyprus 68

NICOSIA: Platania, near Scout camping site, 8 Idalion Archaeological Site, 1 ♀, under stones in ♂♂ 2 ♀♀, pitfalls in Pinus brutia forest, IV - phrygana, 30.IV.2013 (CDM). VII.2017, M. Hadjiconstantis leg. (CRB, CMH). PAPHOS: Akamas Peninsula, Avakas Gorge, 1 ♀, under stone, 27.III.1998, coll. J. Murphy Distribution: Iran, Israel, Cyprus. (MMUE); Kinousa, 1 ♀, under stone in pine wood, 28.III.1998, coll. J. Murphy (MMUE); Neo Zelotes sp. Chorio, 2 ♀♀, under stones in field, 20.III.1998, At least two Zelotes species were captured that coll. J. Murphy (MMUE); Smigies, 1 ♀, under could not be identified, even the most abundantly stones in hillside scrub, 26.III.1998, coll. J. collected one. Also, it is often difficult to match Murphy (MMUE); males and femal es, so some species will eventually be described in a future paper, some Distribution: Mediterranean, Mali, Yemen. New will remain undescribed. to Cyprus.

Hersiliola cfr. simoni (O. Pickard - Cambridge, Family Hahniidae 1872)

Iberina candida (Simon, 1875) Identification Levy, 2003: 25, figs 47 - 56; Marusik & Fet, 2009: Identification : https://araneae.nmbe.ch/ . 82, figs 4.2, 5.2, 6.2 - 3.

Citatio ns Citations None. None.

New records New records LIMASSOL : Fasouri, Akrotiri Peninsula, 1 ♀, in FAMAGUSTA : Cape Greco, Ayia Napa, 35m, 1 reed litter, 4.V.2007 (CAR - S); Pano Platres, ♀, stones in phrygana, 27.IV.2010 (CJVK); Caledonia Waterfall , 3 ♀♀, in litter in rocky Davlos W., 25m, 2 ♀♀, stones in phrygana near stream valley, 4.V.2007 (CAR - S); Pano Platres, the beach, 25.V.2011 (CRB); Flamoudi W., 25m, 2 ♂♂ 3 ♀♀, stones and phrygana near the beach, 1050m, 1 ♀, litter near Mesa Potamos Wat erfall s, 25.V.2011 (CJVK); Karpasia Peninsula, Cape 23.V.2011 (CRB). Apostolos Andreas, 70m, 1 ♀, stones in gra ssland, Distribution: Europe, North Africa. New to 16.V.2011 (CRB). Cyprus. KERYNEIA: Panagia Apsinthiotissa Monastery ruins, 350m, 1 ♂, stones in clearing in Pinus forest, 24.V.2011 (CJVK). Family Hersiliidae LIMASSOL: 10 km S. of Troodos, 1 ♀, under stone in Pinus brutia forest, 19.IV.2017 (CAR - S). Hersiliola macullulata (Dufour, 1831) NICOSIA : Agios Soz omenos, near Dali, 1 ♀, under stones in dry grassland, 20.IV.2017 (CAR - Identification S); Cedar Valley Forest, 2 ♀♀, under stones by Levy, 2003: 28, figs 57 - 60; Marusik & Fet, 2009: track, 29.IV.2007 (CAR - S); between Cedar 81, figs 4.1, 5.1, 6.1. Valley and Stavros tis Psokas, 4 ♀♀, under stones in P inus nigra forest, 29.IV.2007 (CRS); Elia - Citatio ns A ngolemi road, 100m, 1 ♀, stones in grassland, None. rubbish in dry riverbed, 18.V.2011 (CJVK); Kampos NE, Xeros River, 550m, 1 ♀, stones in New records riverbed , 20.V.2011 (CJVK); Machairas Forest FAMAGUSTA: Cape Greco, 1 ♀, under stones in near Kionia, 1300m, 2 ♀♀, stony phrygana, phrygana, 3.V.2013 (CAR - S) . 5.V.2010 (CJVK). NICOSIA : Agios Sozomenos , near Dali, 1 ♂ 1 ♀, PAPHOS : Akamas P eninsula Cape Geronnisos S., under stones in dry grassland, 30.IV.2007, 1 ♀ , on 1 ♂ 1 ♀, under stones in dry phrygana, 3.V.2007 (CAR - S), 1 ♂ 2 ♀♀ (CRS); Akamas Peninsula, limestone ridge in disturbed phrygana, 30.IV.2007 Kioni, track along the sea, 10m, ♂♂ 5 ♀♀, stones (CAR - S), 1 ♂ (CRS); Cedar Valley Forest, 3 ♀♀, in Juniperus phrygana, 30.IV.2010 (CJVK); under stones by track, 29.IV.2007 (CRS); Dali, Akamas Peninsula, Smigies, 352m, 2 ♀♀, maquis ,

Spiders of Cyprus 69

12.III.2018 (CKHK); Kinousa SE, 400m, 1 ♂ 5 marsh, 27.IV.2010 (CRB), 1 ♀, pitfalls at same ♀♀, under stones and litter in Pinus forest, site, III; - V.2011 (CRB); Larnaca SW, Hala 1.V.2010 (CJVK);1 km N.E. of Pegeia, 1 ♀, in pine Sultan Tekke mosque, 50m, 5 ♂♂ 4 ♀♀, pitf alls wood, 30.IV.1982, coll. A. Stubbs (MMUE); in salt marsh, 15 - 28.V.2011 (CRB); Kornos Pegeia, Agios Georgios, rocky shore, 1 ♀, Picnic site, 350m, 2 ♀♀, beating in Pinus forest, 13.III.2018 (CKHK). 28.V.2011 (CRB); Stavrovouni Monastery, 350m, 1 ♂, on bark of Pinus trees, 28.IV.2010 (CRB). Commen ts LIMASSOL : Akrotiri Peninsula E., 5m, 1 ♀, The specimens cited above are closely related to under stones and litt er in salt marsh, 29.IV.2010 Hersiliola simoni but there are some differences. (CRB); Akrotiri Peninsula, near British airbase, 1 Further research is necessary to establish their ♀, under stone in phrygana, 4.V.2007 (CRS); exact identity. Alassa SE, 2 ♀♀, in grass at edge of river Limnatis, 5.V.2007 (CAR - S), 1 ♀ (CRS); Fasouri Distribution: Mediterranean, Nigeria. New to reedbeds, 1 ♂, in reed litter, 27.IV.2007 (CRS); Cyprus. Pano Platres, track near Caledonia Waterfall, 2 ♂♂ 6 ♀♀, under stones at edge of Pinus nigra forest, 26.IV.2007 (CAR - S); Pano Platres, near Family Linyphiidae top of road to Millomeris Waterfalls, 1 ♀, under stones in abandoned vineyard, 28.IV.2007 (CAR - Agyneta fuscipalpa (C. L. Koch, 1836) S); Troodos, n ear top of Mount Olympos (Chionistra), 4 ♂♂ 2 ♀♀, beaten from Pinus , Identification 28.IV.2007 (CRS); Troodos S., 1350m, 1 ♀, under stones in mixed Quercus - Pinus forest, Citations 3.V.2010 (CRB); Troodos, Mount Olympos None. (Chionistra), 1900m, 2 ♂♂, under stones in Pinus forest, 23.V. 2011 (CRB); Troodos S., road to New records Pano Platres, 1 ♀, under stones in Pinus nigra LIMASSOL: Amathus, dry valley along the forest, 28.IV.2007 (CAR - S), 2 ♀♀ (CAR - S); highway, 1 ♂ 2 ♀♀, beating from vegetation, Troodos Square, 1 ♂, pitfalls in open, mixed P. 25.IV.1986 (NHMW). nigra and J. foetidissima forest, 6.IV. - 8.VI.2014 (CMH). Distribution: Palearctic. New to Cyprus. NICOSIA: Agios Sozomenos , near Dali, 1 ♂, under stones in dry grassland, 29.IV.2013 (CAR - Agyneta pseudorurestris (Wunderlich, 1980) S); Agridi, near Dali, 1 ♀, in house, V.2009; Agyneta pseudorurestris Wunderlich, 1980: 319. idem, 1 ♀, in house, IV.2012 (CDM); Athalassa National Forest Park, Nicosia, 1 ♀, under stones Ident ification in pine plantation, 8.V.2013 (CAR - S); between Bosmans, 2006: 140, figs 49 - 54. Cedar valley and Stavros tis Psokas, 1 ♀, under stones in Pinus nigra forest, 29.IV.2007 (CAR - Citations S); Kapedes S., 600m, 1 ♀, herbs and stones Cyprus, without precise locality (Wunderlich, along rivulet, 5.V.2010 (CRB); Lakatamia, 1 ♀, in 1980). olive grove, 1.I. - 1.X. 2016 (CMH); Lefka NW, 120m, 1 ♀, beaten from shrub s in fruit plantation, New records 18.V.2011 (CRB); Limnitis River, Kato Pyrgos FAMAGUSTA: Salamis S., Silver Beach, 5m, 2 SE, 200m, 1 ♀, under stones in Pinus forest, ♀♀, pitfalls in salt marsh, 15 - 27.V.2011 (CRB). 2.V.2010 (CRB); Lympia Reservoir, Lympia, 1 KERYNEIA : Pentadactylos Ran ge, Fterich S., ♂, in clump of rushes below dam wall, 9.V.2013 800m, 1 ♀, under stones in grassland, 17.V.2011 (CAR - S), 1 ♀, 9.V.2013 (CDM). (CRB). PAPHOS : Akamas Penin sula, wood near Agios LARNACA: Achna Reservoir, 50m, 2 ♂♂ 1 ♀, Minas church, 1 ♀, in thick pine litter, 2.V.2007 pitfalls in marshy area around lake, 15 - (CRS); Aphrodites Bath, 1 ♀, under stones in 27.V.2011 (CRB); Delikipos, 1 ♀, in moist grass meadow, 15.IV.2017 (CAR - S); Evretou around small dam, 18.IV.2017 (CAR - S); Reservoir, 350m, 1 ♀, grassland around lake, Dipotamos Dam, 150m, 1 ♀, herbs and stones 1.V.2010 (CRB); Evretou Reservoir, Stavros tis bordering lake, 28.IV.2010 (CRB); Larnaca salt Psokas rive r, 2 ♂♂ 1 ♀, 20.III.2018 (CKHK); lake, 5m, 4 ♀♀, under stones and litter in salt Kato Koutrafas - Astromeritis road, 120m, 1 ♂ 1

Spiders of Cyprus 70

♀, herbs on pond shore, beaten from shrubs in dry Amathus, 1 ♂, dry valley, beaten from vegetation, riverbed, 18.V.2011 (CRB); Kato Paphos, 1 ♂ 1 25.IV.1986 (NHMW); Mount Olympos ♀, beating tall herbs near hotel, 4.IV.1997 (CJB); (Chionistra), S. slope, 1900m, 4 ♀♀, under stones Pegeia, Pikni Forest, 346m, 1 ♀, pitfalls i n former in open Pinus forest, 4.V.2010 (CRB); Pano c arob plantation, 11.III. - 23.III.2018 (CKHK); Platres, Caledonia Waterfall , 1 ♂ 3 ♀♀, litter by Pegeia, Pikni Forest, 374m, 3 ♂♂ 2 ♀♀, pitfalls rocky track, 26.IV.2007 (CAR - S), 1 ♂ (CRS); 1 in open ruderal site, 14.III. - 23.III.2018 (CKHK). ♂ 10 ♀♀, in P. nigra litter by track, 26.IV.2007; 1 ♀, 7.V.2013 (CAR - S); Pano Platres, 1050m, 2 Distribution: Spain, Cyprus, Sardinia, Algeria, ♀♀, litter near Mesa Potamos Waterfall , Tunisia. 23.V.2011 ( CRB); Troodos S., road to Pano Platres, 1 ♂, under stones in Pinus nigra forest, Agyneta simplicita rsis (Simon, 1884) 28.IV.2007 (CAR - S); Troodos, near top Mount Olympos (Chionistra), 1 ♂ 7 ♀♀, beating Cyprus Identification : https://araneae.nmbe.ch/ . and Pinus , 28.IV.2007 (CAR - S), 3 ♂♂ (CRS). NICOSIA: Cedar Valley Forest, 3 ♀♀, und er Citations stones, 29.IV.2007 (CAR - S) , 1 ♀ (CRS); Kapedes None S., 600m, 1 ♀, herbs and stones along rivulet, 5.V.2010 (CRB); Kato Koutrafas - Astromeritis New records road, 120m, 1 ♂ 1 ♀, under stones in dry LIMASSOL : Akrotiri Peninsula, 3 ♀♀, in reed riverbed, 18.V.2011 (CRB); Lakatamia, 1 ♀, in litter, 4.V.2007 (CRS). olive grove, 1.I. - 1.X. 201 6 (CMH). PAPHOS: Akamas Peninsula, between Cape Distribution: Europe, Russia, K azakhstan. New Drepano and Lara Bay, 1 ♀, beating bushes, to Cyprus. 2.IV.1997 (CJB); near Avakas Gorge, 1 ♂ 1 ♀, under stones in field with tall grass, 14.IV.2017 Alioranus pastoralis (O. Pickard - Cambridge, (CAR - S), 1 ♂, beaten from junipers on rocky 1872) hillside (CAR - S); Kinous a, 1 ♀, under stone in Hubertinus planiceps Wunderlich, 1980: 124. pine wood, 28.III.1998, coll. J. Murphy (MMUE); Alioranus pastoralis ; Tanasevitch, 2011: 51. Kelefos bridge, dam on Diarizos River, 550m, 1 Identification ♀, under stones, 21.V.2011 (CRB); Pegeia, Pikni Wunderlich, 1980: figs 8 - 16. Forest, 346m, 6 ♂♂ 3 ♀♀, pitfalls in former carob plantation, 11.III. - 23.III.2018 (CKHK) ; Citations Pegeia, Pikni Forest, 366m, 1 ♂, pitfalls in pine FAMAGUSTA : Agia M arina Kalograias E. forest, 11.III. - 23.III.2018 (CKHK); Pegeia, Pikni (Tanasevitch, 2011). Forest, 374m, 10 ♂♂ 3 ♀♀, pitfalls in open NICOSIA : Cedar valley (Tanasevitch, 2011). ruderal site, 14.III. - 23.III.2018 (CKHK). LIMASSOL : near Platres, 1300m (Wunderlich, 1980). Distribution: Crete, Cyprus, Israel, Tajikistan.

New records Araeon cus cypriacus Tanasevitch, 2011 FAMAGUSTA: Gialousa E ., Agios Thyrsos, Araeoncus cypriacus Tanasevitch, 2011: 31, figs 20m, 2 ♀♀, litter under fig tree, 18.III.2011 1 - 5. (CRB); Paralimni lake, 65m, 1 ♀, litter around dried lake, 27.IV.2010 (CRB). Identification Tanasevitch, 2011: 31, figs 1 - 5. KERYNEIA: Komi Kebir , 150m, 1 ♀, pitfalls in Pinus forest, XI.2010 - III.2011 (CRB); Saint Citations Hilarion castle, 610m, 1 ♀, under stones in Pinus PAPHOS : Polis W., Baths of Aphrodite forest, 22.XI.2010 (CRB); Vouni ruins, 250m, 1 (Tanasevitch, 2011; type locality). ♀, under stones, 20.III.2011 (CRB). LARNACA : Larnaca, 9.XI.1932, 1 ♀, A. Ball leg. New records (KBIN). LIMAS SOL: Pano Platres, Millomeris Waterfalls, LIMASSOL: Akrotiri Peninsula, Fasouri reed 1 ♂, under rocks and in litter, 28.IV.2007 (CRS). beds, 1 ♂, in reed litter, 27.IV.2007 (CRS) ; Alassa S.E., near river Limnatis, 1 ♂ 2 ♀♀, under Distribution: An edemic species of Cyprus. stones in dry grassland, 5.V.2007 (CRS);

Spiders of Cyprus 71

Araeoncus humilis (Blackwall, 1841) Distribution: Mediterranaen. New to Cyprus.

Identification : https://arane ae.nmbe.ch/ . Cresmatoneta mutinensis (Canestrini, 1868) Cresmatoneta mutinensis ; Tanasevitch, 2011: 58. Citations Identification None. Pesarini, 1997: 253, fig 102. Citations New records PAPHOS : Polis ( Tanasevitch, 2011). LIMASSOL: Alassa SE, 2 ♂♂ 1 ♀, under pebbles and in grass at edge of river Limnatis, 5.V.2007 New records (CAR - S). FAMAGUSTA: Gialousa E ., Agios Thyrsos, 20m, 1 subadult ♂ 1 subadult ♀, under stones,

Distribution: Palaearctic, New Zealand. New to 16.V.2011 (CRB); Gialousa E . Agia Triada, 50m, 1 subdult ♂ 1 sub a dult ♀, under stones in ruins, Cyprus. 18.III.2011 (CRB).

NICOSIA: Kato Koutrafas, Astromeritis road, Bathyphantes gracilis (Bl ackwall, 1841) 120m, 3 ♀♀, beaten from shrubs in dry riverbed,

18.V.2011 (CRB); Nicosia, Athalassa National Identification : https://araneae.nmbe.ch/ Forest Park, 1 ♂ 1 ♀, under stones in pine

plantation, 8.V.2013 (CAR - S). Citations PAPHOS: Baths of Aphrodite, 1 ♀, in damp grass None near small stream, 15.IV.2017 (CAR - S); Evretou

Reservoir, 350m, 1 ♀, und er stones in grassland New records around lake, 1.V.2010 (CJVK). LIMASSOL: Akrotiri Peninsula, Fassouli reed beds, 1 ♂ 2 ♀♀, in reed litter, 27.IV.2007, 1 ♀, in Distribution: Mediterranean. reed litter, 30.IV.2013 (CAR - S); 1 ♂, same site, 4.V.2007 (CAR - S). Dactylopisthes digiticeps (Simon, 1881)

Distribution: Holarctic. New to Cyprus. Identification Bosmans et al. 2009: 63, figs 1 - 6. Canariphantes tenerrimus (Simon, 1929)

Citations Identification None. Bosmans & Bouragba, 1992 : 259, figs 28 - 34 (sub

Lepthyphantes homonymus ); Tanasevitch, 2011 : New records . 57, figs 22 - 26 (sub Canariphantes homonymus ). LARNACA: Achna Reservoir , 50m, 4 ♂♂ 4 ♀♀, Citations pitfalls in marshy area bordering lake, 15 - None. 27.V.2011 (CRB). LIMASSOL: Fasouri, Akrotiri Peninsula, 1 ♂ 18 New records ♀♀ in reed litter, 27.IV.2007, 8 ♀♀, 27.IV.2007 FAMAGUSTA : Agios Sergios , 30m, 5 ♀♀, in (CRS), 3 ♂♂ 36 ♀♀, 4.V.2007 (CAR - S). pitfalls in Pinus plantation, 15 - 27.V.2011 PAPHOS: Diarizos river, 1 ♀, 445m, open r iver (CRB); Kantara E. 650m, 1 ♀, in pitfalls in Pinus bank, 16.III.2018, K. - H. & U. Kielhorn leg. forest, XI.2010 - III.2011 (CRB). (CKHK); Evretou Reservoir, Stavros tis Psokas KERYNEIA: Chaleuka Forest Station, road to river, 1 ♂, 20.III.2018 (CKHK); Kidasi village, herbarium, 1 ♀, sweeping in mixed Pinus, Diarizos Valley, 440m, 1 ♀, 16.III.2018 (CKHK). Arbutus and Cystus forest, 12.V.2001 (CRS). LIMASSOL: Akrotiri Peninsula, Fasouri reed Distribution: Mediterranean. New to Cyprus. beds, 1 ♂ 2 ♀♀, in reed litter, 27.IV.2007, 1 ♀, in reed litter, 30.IV.2013 (CAR - S), 1 ♂, 4.V.2007 Diploc ephalus graecus (O. P - Cambridge, 1873) (CRS); Mount Olympos (Chionistra), S. slope, Identification 1900m, 1 ♀, under stones in open Pinus forest, Bosmans, 1996: 136, figs 43 - 50. 4.V.2010 (CRB). NICOSIA: Lympia, Lympia Reservoir, 2 ♀♀, in Citations grass tussocks, 9.V.2013 ( CAR - S). None. Spiders of Cyprus 72

New records . bordering lake, 28.IV.2010 (CRB); Evretou KERYNEIA: Road to Buffavento, 1 ♂, under Reservoir, 350m, 1 ♀, under s tones around lake, stones in woods, 15.IV.2001 (CAR - S). 1.V.2010 (CRB); Larnaca salt lake, 5m, 2 ♂♂ 1 LIMASSOL: Pano Platres, below Kalidonia Falls, ♀, under stones and litter in salt marsh, 1 ♀, in herbs, 1.V.2007 (CAR - S); Pano Platres, 27.IV.2010 (CRB); Larnaca SW, Hala Sultan Myomeris Falls, 2 ♀♀, litter near falls, Tekke mosque, 50m, 2 ♂♂ 1 ♀, pitfalls in salt 28.IV.2007 (CAR - S); Troodos, near top of Mount marsh, 15 - 28.VI.2011 (CRB). Olympos (Chionistra), 1 ♀, beaten from Cyprus LIMASSOL: Alassa SE, 5 ♂♂ 5 ♀♀ , under and Pinus , 28.IV.2007 (CAR - S). pebbles and in grass at edge of river Limnatis, PAPHOS: Evretou Reservoir, Stavros tis Ps okas 5.V.2007 (CAR - S), 1 ♂, (CRS); Erimi W., 1 ♀, river, 1 ♀, 20.III.2018 (CKHK). grass heathland with scattered stones, 21.IV.1986

Distribution: Mediterranean. New to Cyprus. (NHMW). NICOSIA: Agridi, near Dali, 1 ♂ 1 ♀, in house, Diplocephalus turcicus Brignoli, 1972 V.2011, in garden 1 ♀ (CDM); Akrotiri Peninsula, 1 ♂, edge of central salt pan, in Identification Salicornia marsh, 17.IV.2017 (CAR - S); Agios Bosmans et al. 2009: 63, figs 7 - 12. Sozomenos , near Dali, 6 ♂♂ 7 ♀♀, wet area by stream, 30.IV.2007 (CAR - S); Kapedes S., 600m, Citations 2 ♂♂ 2 ♀♀, herbs and stones along rivulet, None. 5.V.2010 (CRB); Kato Koutrafas - Astromeriti s road, 120m, 4 ♀♀, under stones in dry riverbed, New records . 18.V.2011 (CRB); Lympia Reservoir, Lympia, 1 PAPHOS: Avakas Gorge, 50m, 1 ♀, litter in ♂, under stones below dam wall, 9.V.2013 (CAR - citrus grove, 30.IV.2010 (CRB). S) ), 1 ♂ (CDM); Strovolos, 1 ♂, around houses, LIMASSOL: Pano Platres, Caledonian Waterfalls, 1.V.2016 (CMH). 1 ♀, forest litter, 26.IV.2007 (CRS). PAPHOS: A kamas Peninsula, Agios Minas, 1 ♀, wet a rea by stream, 2.V.2007 (CAR - S); Akamas, Distribution: Greece, Turkey. New to Cyprus. entrance to Avakas Gorge, 50m, 1 ♂ 2 ♀♀, litter in citrus yard, 30.IV.2010 (CRB); Akamas Diplocephalus sp. Peninsula, Lara Bay, 1 ♂, under stones, 3.IV.1997 (CJB); Avakas Gorge, 150m, under stones in An unidentified Diplocephalus species will be gorge, 22.V.2011 (CRB); Evretou R eservoir, described in a further paper. Stavros tis Psokas river, 1 ♀, 20.III.2018 (CKHK); Kelefos bridge, dam on Diarizos River, Erigone dentipalpis (Wider, 1834) 550m, 1 ♀, under stones, 21.V.2011 (CRB); Erigone dentipalpis ; Tanasevitch, 2011: 64. Mavrokolympos Reservoir, NE Potima, 150m, 1 ♀, under stones along lake, 22.V.2011 (CRB); Identification : https://araneae.nmbe.ch/ . Pegeia, Coral Bay, 1 ♀, u nder stones, 31.III.1997 (CJB); Pegeia, Pikni Forest, 374m, 1 ♂, pitfalls in Citations open ruderal site, 14.III. - 23.III.2018 (CKHK); LIMASSOL : Moniatis ( Tanasevit ch, 2011). Polis, 5m, 1 ♀, floating in swimming pool of hotel, 1.V.2010 (CRB). New records FAMAGUSTA: Paralimni Lake, 65m, 6 ♂♂ 5 Distribution: Holarctic. ♀♀, herbs and litter around dried up lake, Erigonoplus spinifemuralis Dimitrov, 2003 27.IV.2010 (CRB); Prastio S ., Agios Andronikos,

30m, 1 ♀, under stones in Eucalyptus forest, Identification 27.V.2011 (CRB). Bosmans et al. 2013: 13, figs 5 - 10. LARNACA: Achna Reservoir, 50m, 15 ♂♂ 6 ♀♀, pitfalls in marshy area borde ring lake, 15 - Citations 27.V.2011 (CRB), 4 ♀♀, 15.V.2011 (CRB); None. Delikipos, 2 ♂♂ 5 ♀♀ in moist grass around small dam, 18.IV.2017 (CAR - S); Dipotamos New records Reservoir, 150m, 2 ♂♂ 5 ♀♀, herbs and stones NICOSIA: Dali, 1 ♂, in garden, IV. - V.2008, D.

Spiders of Cyprus 73

McCowan leg., IV. - V.2010 (CDM). shrubs, 1.V.2007 (CAR - S); Pano Platres S., along PAPHOS: Neo Chorio, 1 ♀, on bushes, Kryos River, 1050m, 1 ♀, he rbs in Pinus forest, 30.III.1998, coll. J. Murphy (MMUE). 3.V.2010 (CJVK); Pissouri Beach, 1 ♀, 27.IV.1982, coll. A. Stubbs (MMUE); Pissouri, 1 Distribution: Bulgaria, Greece, Ukraine, Turkey, ♀, on bush, 24.III.1998, coll. J. Murphy Kazakstan. New to Cyprus (MMUE); Platres, 3 ♀♀, on walls, 22.IV.1986 (NHMW); Trooditissa Monastery, Trooditissa Frontinellina frutetorum (C. L. Koch, 1835) river, 1360m, 1 ♂, be aten from shrubs, 19.III.2018 (CKHK). Frontinellina frutetorum ; Bosmans, 1994: 235; NICOSIA: Ayia Koroni, Agios Epifanios, 1 ♂, in Logunov, 2013: 5. house, IV.2005 (CDM); Agios Sozomenos, near Dali , 2 ♀♀, in disturbed phrygana, 30.IV.2007 Identification : https://araneae.nmbe.ch/ (CAR - S); Agridi, near Cedar Valley Forest, 1180m, 1 ♂ 1 ♀, beating in Cedrus fo rest, Citations 19.V.2010 (CRB); D ali, 1 ♂ 1 ♀, in field, Without precise locality (Kulczyński, 1908, sub. IV.2007, 1 ♂ 1 ♀, in garden 14.III.2010, 1 ♀, on Lepthyphantes congener , misidentifiation). wall inside house, I.2011, 1 ♂, on wall inside LIMASSOL: Akrotiri Salt Lake (Logunov, 2013); house, IV.2012 (CDM); Dali, Lymbourena Hill, 1 Pano Platres (Logunov, 2013). ♂, in field, IV.2004 (CDM); Kapedes S., 600m, 2 PAPHOS: Avakas Gorge (Logunov, 2013). ♀♀, herbs and sto nes along rivulet, 5.V.2010 (CRB); Lakatamia, 1 ♀, in olive grove, 1.I. - 1.X. New records 2016 (CMH); Nicosia, Pallouriotissa, 1 ♂, in KERYNEIA: Panagia Apsinthio tissa Monastery building, V.2006. (CDM); Lympia Reservoir, ruins, 350m, 1 ♀, herbs in clearing in Pinus Lympia, 1 ♀, in grass tussocks, 9.V.2013 (CAR - forest, 24.V.2011 (CRB); Pentadactylos Range, S); Oikos N., Xerargaka picnic site, 350m, 1 ♀, Ftericha S ., 800m, 2 ♀♀, in grassland, 17.V.2011 beaten from shrubs in Pinus forest, 20.V.2010 (CRB); Saint Hilarion Castle W, 600m, 1 ♀, (CRB); Pano Amiantos, 1 ♂, pitfalls in Pinus beaten from shrubs, 17.V.2011 (CRB). brutia forest, 6.IV. - 8.VI.2014 (CMH); Pedoulas, LARNACA: Delikipos , near Kornos, 1 ♀, under 1 ♀, by woodland stream, 3.V.1982, coll. A. stones in grass of open pine wood, 9.V.2013 Stubbs (MMUE); near Prodromos reservoir, 1 ♂, (CAR - S); Larnaca SW., Hala Sultan Tekke pitfalls in Pinus nigr a forest, 6.IV. - 8.VI.2014 mosque, 5m, 2 ♀♀, herbs in salt marsh, (CMH). 16.III.2011 (CRB); Mosfiloti, 1 ♀, under stone in PAPHOS: Avakas Gorge, 150m, 1 ♀, in herbs in dry eroded grassland on hillside, 12.IV.2017 gorge, 22.V.2010 (CRB); Evretou Reservoir, (CAR - S). 350m, 3 ♀♀, on shrubs around lake, 1.V.2010 L IMASSOL: Akrotiri Peninsula, near British (CRB); Evretou Reservoir, Stavros tis Psokas Airbase, 3 ♀♀, swept in pine wood, 4.V.2007 river, 1 ♂ 4 ♀♀, 20.III.2018 (CKHK); Giali a, 1 ♂ (CAR - S); Akrotiri Peninsula N., 5m, 1 ♀, herbs in 1 ♀, on bush in pine wood, 25.III.1998, coll. J. Eucalyptus forest, 29.IV.2010 (CRB); Akrotiri Murphy (MMUE); Kato Akourdaleia, 1 ♂ 1 ♀, in Peninsula, 1 ♀, edge of central salt pan, in garden, VIII.2005 (CDM); Kelefos bridge, dam Salicornia marsh, 27.IV.2007 (CAR - S), 1 ♀, on Diarizos River, 550m, 1 ♀, beaten from 17.IV.2017 (CAR - S); Amathus, 1 ♂ 1 ♀, under shrubs, 21.V.2011 (CRB); Kidasi N., along s tones on plants in dry valley, 6.IV.1986 Diarizos River, 300m, 1 ♀, beaten from shrubs, (NHMW); Armenochori, 1 ♀, on bushes, 21.V.2011 (CRB); Kinousa, 1 ♂ 1 ♀, on shrub in 14.IV.1986 (NHMW); Germasogeia, 4 ♀♀, along pine wood, 19.III.1998, coll. J. Murphy (MMUE); the stream below the dam on plants, 10.IV.1986 Kinousa SE, 400m, 5 ♀♀, shrubs in Pinus forest, (NHMW); Kolossi castle, 1 ♀, walls in the castle 1.V.2010 (CJVK, CRB); Kourdaka, Ezousa river, area, 11.IV.1986 (NHMW); Nikitari S., Panagia 1 ♂ 1 ♀, 14.III.2018 (CKHK); near Kritou Terra , tou Asinou, 1 ♀, under stones in phrygana, 1 ♀, swept from grass in dry fallow, 14.IV.2017 9.V.2005 (CAR - S); gorge below Pano Platres, 1 (CAR - S); Mandria, 1 ♂, in scrub, 6.V.1982. coll. ♂, in webs on shrubs, 5.V.2007 (CAR - S); Pano A. Stubbs (MMUE); Mavrokolympos Reservoir, Platres, 1050m, 1 ♀, litter near Mesa Potamos NE Potima, 150m, 1 ♂, herbs along lake, water fal l, 23.V.2010 (CRB); Pano Platres, near 22.V.2010 (CRB); Mavrokolympos Reservoir, 1 top of road to Millomeris Waterfalls, 1 ♀, in litter, ♀, 13.III.2018 (CKHK); Neo Cho rio, 1 ♂, in 28.IV.2007 (CAR - S); Pano Platres, below grass in field, 30.III.1998, coll. J. Murphy Caledonia Waterfall , 1 ♀, swept from herbs and (MMUE); Paphos, NW of the city, 2 ♂♂ 2 ♀♀,

Spiders of Cyprus 74

under stones at the edge of a large sand pit, in rocky valley, 26.IV.2007 (CAR - S), 4 ♂♂ 24.IV.1986 (NHMW); Paphos, near Tombs of the (CRS). Kings, 25m, 2 ♀♀, on herbs, 23.III.2011 (CRB); PAPHOS: Kelefos bridge, 465m, deciduous Pegeia, Pikni For est, 346m, 3 ♀♀, in former wood, 2 ♀♀, 16.III.2018 (CKHK). c arob plantation, 11.III.2018, same site, 1 ♂ 1 ♀, 23.III.2018 (CKHK); near Souskiou, 1 ♀, on bush Distributi on: Palaearctic. in dry grassland, 1.V.2013 (CAR - S); Vretsia W., near Kelefos bridge, 650m, 1 ♀, beaten from Improphantes cypriot Tanasevitch, 2011 shrubs, 19.V.2010 (CRB). Improphantes cypriot Tanasevitch, 2011: 67 (descr. ♂). Distrib ution: Palaearctic. Identification Gnathonarium dentatum (Wider, 1834) Tanasevitch, 2011: 67, figs 59 - 64 ; present paper, page 20 . Identification : https://araneae.nmbe.ch/ . Citations Citations NICOSIA : Nicosia ( Tanasevitch, 2011). FAMAGUSTA : Koma tou Gialou (Tanasevitch, 2011). New records LARNACA : N. Larnaca airport (Tanasevitch, LIMASSOL: Akrotiri Peninsula, 1 ♂, margin of 2011; type locality). central salt lake, 7.V.1972, coll. A. Stubbs (MMUE); Akrotiri Peninsula, Fasouri reed beds, 2 New records ♂♂ 6 ♀♀ in reed litter, 27.IV.2007, 3 ♂♂ 17 ♀♀, FAMAGUSTA: Agios Sergios , 30m, 5 ♂♂ 5 ♀♀, pitfalls in Pinus plantation, 15 - 27.V.2011 30.IV.2013, 10 ♂♂ 7 ♀♀, 4.V.2007 (CAR - S), 4 (CRB); Cape Greco, 1 ♀, in deep pine litter, ♀♀, 30.IV.2013 (CDM). 3.V.2013 (CAR - S). NICOSIA: L YMPIA R ESERVOIR , Lymp ia, 2 ♀♀, LARNACA: Ac hna Reservoir, 50m, 1 ♀, under under stones below dam wall, 9.V.2013 (CDM). stones and litter bordering lake, 27.V.2011 PAPHOS : Avakas Gorge, 50m, 1 ♀, litter in (CRB); Larnaca salt lake, 10m, 4 ♂♂ 5 ♀♀, gorge, 30.IV.2010 (CJVK), 1 ♀, stones in gorge, pitfalls around lake, XI.2010 - III.2010 (CRB); 5.IV.1997 (CJB). Larnaca SW, Hala Sultan Tekke mosque, 50m, 1 ♀, in pitfalls in salt marsh, 15 - 28.V.2011 (CRB) ; Distribution: Mosfiloti, 1 ♂, under stones in dry eroded Palaearctic. grassland on hillside, 12.IV.2017 (CAR - S); Stavrovouni Monastery W., 350m, 1 ♀, pitfalls in Gongylidiellum murcidum Simon, 1884 open Pinus forest, XI.2010 - III.2011 (CRB). Gongyldiellum murcidum ; Tanasevitch, 2011: 66. LIMASSOL: Akrotiri Peninsula, Fasouri reed beds, 3 ♀♀ in reed litter, 4. V.2007 (CRS); Pano Identification : https://araneae.nmbe.ch/ . Platres, Caledonia Waterfall, 1 ♀, in litter near track, 26.IV.2007 (CAR - S); Troodos, near top of Citations Mount Olympos (Chionistra), 1 ♀, under stones in LARNACA: Larnaca 6 km E., hotel Lordos open P inus nigra forest, 28.IV.2007 (CRS); (Tanasevitch, 2011). Troodo s E., Livadi Tou Passia, 1750m, 1 ♂, LIMASSOL : Agios Dimitrios (Tanasevitch, pitf alls in peat marsh, III. - V.2011 (CRB); 2011); Pano Platres ( Tanasevitch, 2011); Phini - Troodos S., 1350m, 3 ♀♀, stones in mixed Agios Dimitrios road ( Tanasevitch, 2011); Quercus - Pinus forest, 3.V.2010 (CRB). Prodromos, 3 km S. ( Tanasevitch, 2011 ). NICOSIA : Athalassa National Forest Park, NICOSIA: Cedar valley ( Tanasevitch, 2011). Nicosia S., 3 ♂♂ 4 ♀♀, Eucalyptus litter, 2 - PAPHOS : Alonoudhi junction ( Tanasevitch, 8.V.2013 (CRS), 3 ♀♀, under stones in pine 2011); Maa - Palaiocastro, Coral Bay ( Tanasevitch, planta tion, 8.V.2013 (CAR - S); Kato Koutrafas - 20 11). Astromeritis road, 120m, 1 ♂ 3 ♀♀, herbs on pond shore, 18.V.2011 (CRB); Mosfiloti, 1 ♂, New records under stone in dry eroded grassland on hillside, LIMASSOL : Pano Platres, track near Caledonia 12.IV.2017 (CAR - S). Waterfall , 20 ♂♂ 19 ♀♀, in patch of moist litter

Spiders of Cyprus 75

PAPHOS : Aphrodites Bath, 1 ♂, under stones in meadow, 1 5.IV.2017 (CAR - S); Avakas gorge, Distribution: Turkey. New to Cyprus. 50m, 1 ♂ 2 ♀♀, litter in citrus yard at entrance of gorge, 30.IV.2010 (CRB); near Kidasi village, Jacksonella bidens Tanasevitch, 2011 Diarizos Valley, 1 ♂, in flood debris by stream, Jacksonella bidens Tanasevitch, 2011: 71. 1.V.2013 (CAR - S); Evretou Reservoir, 350m, 1 ♀, shrubs and grassland around lake, 1.V.2010 Identification (CRB); Smigies, W. Neo Chorio, 300m, 1 ♀, litter Tanasevitc h, 2011: 71, figs 70 - 76. and stones in Pinus forest, 30.IV.2010 (CRB). Citations Distribution: Only known from Cyprus. It can LIMASSOL : Prodromos S. (Tanasevitch, 2011; co - occur with Improphantes turok , but this type locality). species mostly occurs at higher altitudes. NICOSIA : Above Ayia (Tanasevitch, 2011). PAPHOS : Mamonia, bank of Diarizos Impro phantes turok Tanasevitch, 2011 (Tanasevitch, 2011). Improphantes turok Tanasevitch, 2011: 69, figs 65 - 69 (descr. ♂, ♀). New records LARNACA: Livadia, 1 ♂, riverbed, 20.IV .2016 Identification (CMH). Tanasevitch, 2011: 69, figs 65 - 69 ; present paper, LIMASSOL : Fasouri, Akrotiri Peninsula, 1 ♀, page 20 . reed beds, 4.V.2007 (CAR - S).

Citations Distribution: Cyprus, Samos. None. Linyphia maura Thorell, 1875 New records LIMASSOL: Kapedes, 1 ♀, Pinus bru tia forest, Identification pitfalls, 17.I. - 23.IV.2016 (CMH); Mount Van Helsdingen, 1969: 41, figs 31 - 39; Bosmans, Olympos (Chionistra), 1900m, 1 ♀, under stones 2006: 136, figs 33 - 36. in Pinus forest, 28.IV.201 (CRB), 1 ♂ 1 ♀, in Citat ions pitfalls in Pinus forest, III. - V.2011 (CRB); Pano None. Platres E., Mesa Potamos Waterfall s, 950m, 1 ♀, in litter in Pinus forest, 4.V.2010 (CRB); road Pedoulas - Prodromos, 2 ♀♀, pitfalls in Pinus New records nigra forest, 6.IV. - 8.VI.2014 (CMH); Prodromos PAPHOS : Akamas Peninsula, Neo Chorio, 1 ♀, S., Trooditissa picnic site, 1300m, 4 ♂♂ 1 ♀, swept from grasses, 2.V.2005 (CAR - S). pitfalls in open Pinus forest, XI.2010 - III.2011 Distribution: Western Mediterranean. New to (CRB); Prodomos E., near Pro dromos dam, 1 ♀, Cyprus. in pitfalls in Pinus forest, XI.2010 - III.2010, 1 ♀, III. - V.2011 (CRB); Troodos E., Livadi Tou Linyphia mimonti Simon, 1884 Passia, 1750m, 1 ♂, pitfalls in peat marsh, XI.2010 - III.2011 (CRB); Troodos S., 1350m, 3 Identification ♀♀, stones in mixed Quercus - Pinus forest, Van Helsdingen, 1969: 68, fi gs 71 - 78.

3.V.2010 (CRB). Citations NICOSIA: Cedar Valley Forest, 1180m, 1 ♀, None. under stone in forest, 28.IV.2017 (CAR - S). PAPHOS : Agios Georgios E., Palaiomylos, New records 1250m, 1 ♀, beaten in mixed Pinus and Quercus NICOSIA: Pedoulas, 1 ♀, by woodland stream, forest, 4.V.2010 (CRB); Kelefos bridge, 465m, 3.V.1982, coll. A. Stubbs (MMUE). LIMASSOL : Agios Georgios E., Palaiomylos, deciduous wood, 4 ♀♀, 16.III .2018 (CKHK); 1250m, 9 ♀♀, beating in mixed Pinus and Pegeia, Pikni Forest, 346m, 2 ♀♀, in former Quercus forest, 4.V.2010 (CJVK, CRB); Ceda r c arob plantation, 23.III.2018 (CKHK). Valley Forest, 1180m, 4 ♀♀, beating in Cedrus

Spiders of Cyprus 76

forest, 19.V.2010 (CRB), 4 ♀♀, 29.IV.2007 Lepthyphantes albuloides ; Kulczyński, 1908: 67, (CAR - S), 1 ♀ (CRS); Cedar Valley, 1150m, 5 pl. 2, fig. 19 (misidentific ation). ♀♀, 21.III.2018, forest road, beaten from shrubs Megalepthyphantes turkeyensis ; Tanasevitch , (CKHK); Pano Platres, 2 ♀♀, in webs on shrubs, 2011: 74. 6.V.2007 (CAR - S); Pa no Plat res, below Caledonia Waterfall , 1 ♂ 3 ♀♀, swept from herbs Identification and shrubs, 1.V.2007 (CAR - S); Pano Platres S., Tanasevitch et al. , 2005 : 279 , 2011: 74, figs 1 - 9 . along Kryos River, 1050m, 1 ♀, beating in Pinus 82 - 83. forest, 3.V.2010 (CJVK); Pano Platres E., Mesa Potamos Waterfall s, 950m, 3 ♀♀, beating in Comments Pinus forest , 4.V.2010 (CJVK); Troodos S., Although Tanasevitch (2011) claimed to hav e 1350m, 3 ♀♀, beating in mixed Quercus - Pinus examined females, the epigyne of forest, 3.V.2010 (CJVK); Trooditissa Monastery, Megalepthyphantes t urkeyensi s was never Trooditissa river, 1360m, 1 ♂ 6 ♀♀, beaten from illustrated. shrubs, 19.III.2018 (CKHK). . Citations Distribution: Palearctic. New to Cyprus. Without precise locality (Kulczyński, 1908, sub Lepthyphantes albuloides ( ♀) and L. congener Liny phia triangularis (Clerck, 1757). ( ♂)). NICOSIA: C edar Valley Forest (Tanasevitch, Identification : https://araneae.nmbe.ch/ . 2011).

Citations New records None. KERYNEIA: road Chalefka - Sourp Magar Monastery (Armenomonastiro), 660m, 1 subadult New records ♀, under stones in open Pinus forest, 23.XI.2010 KERYNEIA: Klepini , 300m, 2 ♀♀, on low (CRB); Klepini , 300m, 1 subadult ♀, under stones bushes in open Pinus forest, 23.XI. 2010 (CRB). in open Pinus forest, 23.XI.2010 (CRB); Keryneia , 1 ♀, 24.X.1933, G. Mavromoustakis Distribution: Palearctic. New to Cyprus. leg. (BMNH); Saint Hilarion castle, 610m, 1 ♂ 1 ♀, under stones in Pinus f orest, 22 .XI. 2010 Maculoncus parvipalpus Wunderlich, 1995 (CRB). LIMASSOL : Pedoulas, 960m, 1 ♀, road Identification embankment, 20.IV.2017, S. Huber leg. (CSH); Wunderlich, 1995c: 647, figs 12 - 19. Platres, 1400m, 1 ♀, in pine forest, 28.X.1981, P. R. Deeleman leg. (NHML); Troodos Square, Citations 1600m, 2 ♂♂ 1 ♀, under stones in pine forest, None. 31.X.1989 , P. R. Deeleman leg. (NHML). NICOSIA: Agios Sozomenos, near Dali, 1 ♀, New records under stone in dry, rocky phrygana, 20.IV.2017 LIMASSOL : Agros SW, 950m, 1 ♀, under stones (CAR - S); Kapedes, 2 ♀♀, pitfalls in Pinus brutia forest, 17.I. - 23.IV.2016 (CMH). in Pinus forest, 5.V.2010 (CJVK). PAPHOS: Kato Akourdaleia, 1 ♀, VIII.2008 PAPHOS: Limnitis River, Kato Pyrgos SE, 200m, (CDM). 1 ♀, litter in Pinus forest, 2.V.2010 (CRB); Pegeia, Pikni Forest, 346m, 1 ♂, in former c arob Distribution: Turkey, Cyprus. plantation, 23.III.2018 (CKHK). Mermessus denticulatus (Banks, 1898) Distribution: Greece, Peloponnisos and the islands of Chios, Crete a nd Naxos. New to Cyprus. Identificaton Van Helsdingen, 2009: 618, figs 1 - 10. Megalepthyphantes turkeyensis Tanasevitch, Kunt & Seyyar, 2005 Citations None. Lepthyphantes congener ; Kulczyński, 1908: 68, pl. 2, fig. 17 (misidentification).

Spiders of Cyprus 77

New records Distribution : Hola rctic. FAMAGUSTA: Paralimni lake, 65m, 1 ♂ 2 ♀♀, herbs and litter around dried lake, 27. IV.2010 Microlinyphia pusilla (Sundevall, 1830) (CRB). Microlinyphia pusilla ; Van Helsdingen, 1970: 9. LARNACA: Achna Reservoir, 50m, 1 ♂ 1 ♀, under stones and litter bordering lake, 16.V.2011 Identification : https://araneae.nmbe.ch/ . (CRB), 1 ♀, 27.V.2022, 4 ♀♀, pitfalls at same site, 15 - 27.V.2011 (CRB); Larnaca salt lake, 5m, Citations 1 ♂ 2 ♀♀, under stones and litter in salt marsh, None. 27. IV.2010 (CRB). LIMASSOL: Alassa, in grass by edge of river, New records 5.V.2007, 2 ♂♂ 5 ♀♀ (CRS). FAMAGUSTA: Vokolida S ., Cape Elaia, 1 ♀, 10m, under stones and on bushes near the beach, Distribution: USA to Colombia, introduced in 26.V.2011 (CRB). Europe and North Africa. New to Cyprus. LARNACA: Achna Reservoir, 50m, 1 ♀, 15.V.2010, 1 ♀, herbs in marshy area bordering Mermessus fradeorum (Berland, 1932) lake, 19.V.2011 (CRB); Kelefos bridge, dam on Diarizos River, 550m, 1 ♀, beaten from s hrubs, Identification 21.V.2011 (CRB); Dipotamos Reservoir, 150m, 1 Millidge, 1988 : 67, figs 333 - 339. ♀, herbs and stones bordering lake, 28.IV.2010 (CRB). Citations NICOSIA: Agridi, near Dali, 1 ♂, in garden, None. 14.III.2010 (CDM); Alassa, by river Limnatis, 1 ♂, in dry grassland, 4.V.2007 (CRS); Kato New records Koutrafas - Ast romeritis road, 120m, 1 ♂ 1 ♀, FAMAGUSTA: Paralimni Lake, 65m, 1 ♀, herbs beaten from shrubs in dry riverbed, 18.V.2011 and litter around dried lake, 27.IV.2010 (CRB). (CRB); Machairas forest near Kionia, 1300m, 1 ♂, stony phrygana, 5.V.2010 (CRB); Cedar Distribution: Cosmopolitan. New to Cyprus. Valley Forest, 1180m, 1 ♂, beating in Cedrus forest, 19.V.2010 (CRB); Troodos S., 1350m, 1 Microctenonyx subitaneus (O. Pickard - ♀ , herbs in mixed Quercus - Pinus forest, 3.V.2010 Cambridge, 18 75) (CJVK). Microctenonyx subitaneus ; Tanasevitch, 2011: 75. PAPHOS: Asprokremmos dam, N. Kouklia, 150m, 1 ♂, herbs along lake, 29.IV.2010 (CRB); Identification : https://araneae.nmbe.ch/ . Kinousa, 1 ♀, in litter in pine wood, 19.III.1998, coll. J. Murphy (MMUE); Kykkos Monastery, 1150m, 1 ♀, on hedg es around monastery, Citations 19.V.2010 (CRB); Troodos S., 1350m, 1 ♂, on LARNACA : Xylofagou 4 km E. (Tanasevitch , herbs in mixed Quercus - Pinus forest, 3.V.2010 2011). (CRB).

New records Distribution : Holarctic. New to Cyprus. FAMAGUSTA: Paralimni lak e, 65m, 1 ♀, herbs and litter around dried lake, 27.IV.2010. Neriene clathrata (Sundevall, 1830) LARNACA: Larnaca salt lake, 5m, 1 ♀, under stones and litter in salt marsh, 27.IV.2010 (CRB), Identification : https://araneae.nmbe.ch/ . 1 ♀, pitfalls at same site, III. - V.2011 (CRB). NICOSIA: Agridi, near Dali, 1 ♀, in eucalyptus Citations l itter in garden, 29.IV.2013 (CAR - S). None. PAPHOS: Avakas Gorge, Akamas Peninsula , 50m, 4 ♀♀, litter in citrus yard at entrance of New records LARNACA: Agios Minas, 1 ♀, in litter in pine gorge, 30.IV.2010 (CRB); Pegeia, Pikni Forest, wood, 20.III.1998, coll. J. Murphy (MMUE). 346m, 1 ♀, in former carob plantation, 23.III.2018 (CKHK). Distribution: Holarctic. New to Cyprus.

Spiders of Cyprus 78

Neriene furtiva (O. Pickard - Cambri dge, 1871). Distribution: An endemic species of Cyprus.

Identification : https://araneae.nmbe.ch/ . Pelecopsis elongata (Wider, 1834) Pelecopsis elongata ; Tanasevitch, 2011: 77. Citations None. Identification : https://araneae.nmbe.ch/ .

New records Citations PAPHOS: Smigies, W. Neo Chorio, 300m, 1 ♀, LIMASSOL : Doros ( Tanasevitch, 2011); herbs in Pinus forest, 30.IV.2010 (CRB). Limassol, salt lake ( Tanasevitch, 2011). PAPHOS : Polis W., Baths of Aphrodite Distribution: Europe, North Africa, Turkey. New ( Tanasevitch, 2011); Pro dromi - Androlikou road to Cyprus. ( Tanasevitch, 2011).

Ostearius melanopygius (O. Pickard - New records Cambridge, 1879) FAMAGUSTA: Gialousa E., Agios Thyrsos, 20m, 2 ♀♀, litter under fig tree, 18.III.2011 (CRB); Identification: https://araneae.nmbe.ch/ . Kantara castle, 650m, 1 ♀, under stones in ruins, 19.III.2011 (CRB). Citations LIMASSOL : Fasouri reedbeds, Akrotiri None. Peninsula, 1 ♀, in reed litter, 4.V.2007 (CRS); Pano Platres, Caled onia Waterfall , 1 ♀, in moist New records litter, 26.IV.2007 (CA R - S); Pano Platres, LIMASSOL: Erimi, 1 ♂, under stones in dry Millomeris Waterfall s, 1 ♀, litter near Waterfall , riverbed, 18.IV.1986 (NHMW); Germasogeia, 1 28.IV.2007 (CAR - S); Troodos, near top of Mount ♂, on herbs along the stream below the dam, Olympos (Chionistra), 1 ♀, under stone in open P. 10.IV.1986 (NHMW); Limassol, 1 ♀, hotel nigra forest, 28.IV.2007 (CRS). Sheraton, 5.X.1988 (NHMW). NI COSIA: Dali, 13, Agios Andronikos, 1 ♂, in garden, IV.2007 (CDM); Agridi, near Dali, 1 ♀, Distribution: Cosmopolitan. New to Cyprus. in grass in lawn, 20.VI.2011; 1 ♂, on wall in house, 13.XII.2011 (CDM). Palliduphantes minimus (Deeleman - Reinhold, PAPHOS: Dhiarizos river near Kelefos bridge, 1986) 444m, 1 ♀, open river bank, 16.III.2018 (CKHK); Lepthyphantes minimus Deeleman - Reinhold, Gialia, 1 ♀, bushes in pine wood, 25.III.1998, 1986: 46, figs 35a - b. coll. J. Murphy (MMUE); Mavrokolympos Reservoir, 4 ♀♀, 13.III.2018 (CKHK). Identification Deeleman - Reinhold, 1986: 46, figs 35a - b; present Distribution: Europe, Russia. paper, page 23. Prinerigone vagans (Audouin, 1826) Citations Prinerigone vagans ; Tanasevitch, 2011: 77. LIMASSOL : Platres, south slope of Mount Troodos, 1400m (type local ity; Deeleman - Identifi cation : https://araneae.nmbe.ch/ . Reinhold, 1985). Citations New records PAPHOS : Mamonia, bank of Diarizos river KERYNEIA: Chaleuka Forest Station, road to ( Tanasevitch, 2011). herbarium, 1 ♀, sweeping in mixed Pinus, Arbutus and Cystus forest, 12.V.2001 (CRS). New records LARNACA: Larnaca salt lake, 10m, 1 ♀, pitfalls LARNACA: Achna Reservoir, 50m, 3 ♂♂, along salt lake, XI.2010 - III.2011 (CRB). pitfalls in marshy area around lake, 15 - LIMASSOL: Prodromos S., Trooditissa picnic 27.V.2011 (CRB); Larnaca salt lake, 5m, 1 ♀, site, 1300m, 1 ♂ 1 ♀, pitfalls in open Pinus forest, under stones and litter in salt marsh, 27.IV.2010 XI.2010 - III.2011 (CRB). (CRB). NICOSIA: Lympia Reservoir, 1 ♀, in Arundo litter, 15.V.2013 (CRS).

Spiders of Cyprus 79

LIMASSOL: Alassa SE, 12 ♂♂ 14 ♀♀, under E., Livadi Tou P assia, 1750m, 2 ♂♂, pitfalls in pebbles and in grass at edge of River Limnatis, peat marsh, XI.2010 - III.2011 (CRB). 5.V.2007 (CAR - S), 2 ♀♀ (CAR - S). NICOSIA: Kapedes, 1 ♀, pitfalls in Pinus brutia NICOSIA: Kato Koutrafas - Astro meritis road, forest, 17.I. - 23.IV.2016 (CMH); Prodromos E., 120m, 1 ♀, beaten from shrubs in dry riverbed, near Prodromos dam, 1550m, 1 ♂, pitfalls in 18.V.2011 (CRB). Pinus forest, XI.2010 - III.2011 (CRB). PAPHOS: Akamas Peninsula, Agios Minas, 1 ♂ 1 PA PHOS: Kato Akourdaleia, 1 ♂, in garden, ♀, wet area by stream, 2.V.2007 (CAR - S); VIII.2005 (CDM); Pegeia, Pikni Forest, 366m, 1 Dhiarizos river near Kelefos bridge, 444m, 2 ♀♀, ♀, pitfalls in pine forest, 11.III. - 23.III.2018 open river bank, 16.III.2018 (CKHK); Evre tou (CKHK). Reservoir, Stavros tis Psokas river, 1 ♂ 1 ♀, 20.III.2018 (CKHK). Distribution : S. France, Italy, the Balkans, Caucasus, Cyprus. Distribution: Old World. Styloctetor romanus (O. Pickard - Cambri dge, Silometopus reussi (Thorell, 1871) 1873)

Identification : https://araneae.nmbe.ch/ . Identification : https://araneae.nmbe.ch/ .

Citations Citations None. None.

New records New records FAMAGUSTA: Paralimni Lake, 65m, 1 ♀, litter FAMAGUSTA: Agios Sergios , 30m, 2 ♂♂ 1 ♀, around dried lake, 27.IV.2010 (CRB). in pitfalls in Pinus plantation, 15. - 27.V.2011 (CRB). Distribution : Palaearctic. New to Cyprus. PAPHOS: Gial ia, 1 ♀, bushes in pine wood, 25.III.1998, coll. J. Murphy (MMUE).

Sintula retroversus (O. Pickard - Cambridge, Distribution : Palaearctic. New to Cyprus. 1875)

Sintula retroversus ; Tanasevitch, 2011: 78. Tallusia vindobonensis (Kulczyński 1898)

Id entification Identification Tanasevitch, 1990: 108, figs 14.9 - 10. Polenec & Thaler, 1980: 103, figs 1 - 7.

Citations Citations PAPHOS : Polis W., Baths of Aphrodite None. ( Tanasevitch, 2011); between Baths of Aphrodite and Fontana Amorosa ( Tanasevitch, 2011). New re cords

LIMASSOL: Livadi Tou Passia, E. Troodos, New records 1750m, 1 ♀, pitfalls in peat marsh, XI.2010 - KERYNEIA : Saint Hilarion Castle, 610m, 1 subadult ♂ 1 ♀, under stones in Pinus forest, III.2011, 1 ♀, III. - V.2011 (CRB); Pano Platres, 22.XI.2010 (CRB), 2 ♀♀, pitfalls at same site, gorge below Caledonia Waterfall , 1 ♀, under XI.2010 - III.2011 (CRB). stones, 26.IV.2004 (CAR - S). LARNACA: Achna Reservoir, 50m, 1 ♂ 1 ♀, PAPHOS: Kelefos bridge, 465m, deciduous under stones and litter bordering lake, 27.V.2011 wood, 1 ♀, under bark, 16.III.2018 (CKHK). (CRB). LIMASSOL : Malia - Dora road, 650m, 1 ♀ , under Distribution : Central & Eastern Europe. New to stones in grassland, 21.III.2011 (CRB); Pano Cyprus, but only found at high altitude. Platres E., Mesa Potamos Waterfall s, 950m, 1 ♀, litter in Pinus forest, 4.V.2010 ( CRB); Pano Platres, Millomeris Waterfall s, 1 ♀, litter near Tapinocyba n. sp. Waterfall , 28.IV.2007 (CAR - S); Troodos range A new Tapinocyba species will be described in a further paper. Spiders of Cyprus 80

Tenuiphantes floriana (Van He lsdingen, 1977) and litter in Pinus forest, 3.V.2010 (CRB); Pano Platres E., Mesa Potamos Waterfall s, 950m, 3 ♂♂ Identification : https://araneae.h/ . 7 ♀♀, litter in Pinus forest, 4.V.2010, and 1 ♂, 23.V.2011 ( CRB); Pano Platres, Millomeris Citations Waterfall s, 1 ♀, in litter, 28.IV.2007 (CA R - S); None. Troodos E., Livadi tou Passia, 1170m, 39 ♂♂ 39 ♀♀, in mosses in peat marsh, 3.V.2010 (CJVK, New records CRB), 1 ♀ in pitfalls, XI.2010 - III.2011 (CRB); NICOSIA: Prodromos S., Trooditissa picnic Troodos S., 1350m, 2 ♂♂, under stones in mixed place, 1300m, 2 ♂♂ 1 ♀, pitfalls in open Pinus Quercus - Pinus forest, 3.V.2010 (CRB). forest, XI.2010 - I II.2011 (CRB). NICOSIA: Agridi, near Dali, 1 ♂, in garden, IV.2010 (CDM); Athalassa National Forest Park, Comments Nicosia, 1 ♀, under stones in pine plantation, Males can be distinguished from the commo n 8.V.2013 (CAR - S); Cedar Valley Forest, 1180m, Tenuiphantes tenuis by the presence of 2 teeth on 4 ♂♂ 10 ♀♀, beating in Cedrus forest, 19.V.2010 the posterior margin of the paracymbium. The (CRB); Chrysoroyiatissa Monastery, 1 ♀, in pi ne disposition of the teeth of the specimens from and oak woodland, 2.V.1982. coll. A. Stubbs Cyprus differs slightly from material fr om (MMUE); Dali, 1 ♀, on outside wall of house, continental Greece . 5.V.2013 (CAR - S); Kapedes S., 600m, 1 ♀, herbs and stones along rivulet, 5.V.2010 (CRB); Kato Distribution Koutrafas, Astromeritis road, 120m, 3 ♂♂ 4 ♀♀, Bulgaria, Albania, Romania, Serbia, North under stones in dry riv erbed, 18.V.2011 (CRB); Macedonia. New to Cyprus. Lympia Reservoir, Lympia, 3 ♂, in clump of rushes below dam wall, 9.V.2013 (CAR - S); Tenuiphantes tenuis (Blackwall, 1852) Machairas forest near Kionia, 1300m, 2 ♂♂ 6 Tenuiphantes tenuis ; Tanasevitch, 2011: 83. ♀♀, stony phrygana, 5.V.2010 (CJVK); below Machairas Monastery, 2 ♂♂ 1 ♀, under stones Identification : https://araneae.nmbe.ch/ . beside track in p hrygana, 4.V.2013 (CAR - S); N. Oikos, Xerargaka picnic site, 350m, 3 ♂♂ 4 ♀♀, Citations beating bushes and under stones in Pinus forest, LIMASSOL : Agios Dimitrios ( Tanasevitch, 20.V.2010 (CRB); Prodromos E., near Prodomos 2011). dam, 1550m, 1 ♀, under stones and litter in Pinus NICOSIA : Cedar Valley Forest ( Tanasevitch, forest, 2.V.2010, 1 ♀ in pitfalls, XI.2010 - III.2011; 2011). Prodromos S., Trooditissa picnic site, 1450m, 1 PAPHOS: Al o noudhi junction ( Tanasevitch, ♂, under stones in Pinus forest, 21.V.2011 (CRB). 2011). PAPHOS: Akamas Peninsula, Agios Minas, 1 ♂ 1 ♀, in thick pine litter, in wet area by stream, New records 2.V.2007 (CAR - S); Aphrodites Bath, 1 ♀, under LARNACA: Agioi Vavatsinias, 700m, 4 ♂♂ 5 stones in meadow, 15.IV.2017 (CAR - S); Avakas ♀♀, litter in Pinus forest, 28.IV.2010 (CRB). gorge, 50m, 3 ♂♂ 4 ♀♀, litter in citrus yard at LIMASSOL: Alassa SE, 7 ♂♂ 6 ♀♀, under entrance of gorge, 30.IV.2010 (CJVK, CRB), 1 ♀, pebbles and in grass at edge of river Limnatis, at under stones in gorge, 22.V.2010 (CRB); Cedar base of limestone rock, 5.V.2007 (CAR - S), 2 ♂♂ Valley Forest, 1180m, 4 ♂♂ 10 ♀♀, beating in (CRS); Amathus, excav ation area, 1 ♀, under Cedrus forest, 19.V.201 0 (CRB), 1 ♂ 2 ♀♀, stones, 9.IV.1986 (NHMW ); Malia - Dora road, under stones, 29.IV.2007 (CAR - S), 2 ♂♂ 2 ♀♀ 650m, 1 ♀, under stones in grassland, 21.III.2011 (CAR - S) ; between Cedar Valley and Stavros tis (CRB); Mount Olympos (Chionistra), S. slope, Psokas, 1 ♀, under stones in P. nigra forest, 1900m, 9 ♂♂ 9 ♀♀, under stones in open Pinus 29.IV.2007 (CRS); 9 km N. Pano Panagia, forest, 4.V.2010 and 7 ♂♂ 1 ♀, pitfalls at same 1050m, in litter near spring, 6 ♂♂ 3 ♀♀, site, XI.2010 - III.2011 (CRB); Pano Platres, 1 ♀, 19.V.2011 (CRB); Evr etou Reservoir, 350m, 4 on stone wall, 6.V.2007 (CAR - S); Pano Platres, ♂♂ 3 ♀♀, shrubs and grassland around lake, below Caledonia Waterfall , 9 ♂♂, 11 ♀♀, swept 1.V.2010 (CJVK, CRB); Kinousa E., Agios from herbs and shrubs, 1.V.2007 (CAR - S), 14 ♂♂ Merkourios, 350m, 2 ♂♂, under stones and, litter 6 ♀♀, under stones and in grass, rocky stream in Pinus forest, 1.V.2010 (CRB). valley, 7.V.2013 (CAR - S); Pano Platres S., along Kryos River, 1050m , 3 ♂♂ 5 ♀♀, under stones Distribution: Cosmopolitan.

Spiders of Cyprus 81

Theonina laguncula (Denis, 1937) N. Comb. Citations None. Identification Present paper, p. 25 . New records LIMASSOL: Pano Platres, below Caledonia Citations Waterfall , 1 ♂ 5 ♀♀, under stones and in grass, None. 26.IV.2007 (CAR - S).

New records Distribution: Palaearctic. New to Cyprus. PAPHOS : Akamas Peninsula Agios Minas, 4 ♀♀, . in dense Pinus litter, 2.V.2007 (CAR - S, CRB, Walckenaeria cyprusensis Wunderlich, 1995 CRS); Smigies, W. Neo Chorio, 300m, 1 ♀, litter Walckenaeria cyprusensis Wunderlich, 1995b: and stones in Pinus for est, 30.IV.2010 (CRB). 674; Tanasevitch, 2011: 89.

Distribution: North Algeria, Morocco, Spain, Identification Greece and new to Cyprus. Wunderlich, 199b5: 674, figs 110 - 115.

Typhochrestus digitatus (O. Pickard - Citations Cambridge, 1873) LIMASSOL : Agios Dimitrios ( Tanasevitch,

2011); Troodos Mts, 1500m (type locality; Identification: https://araneae.nmbe.ch/ . Wunderlich, 1995b).

Ci tations New records None. LIMASSOL: Livadi Tou Passia, E. Troodos,

1750m, 1 ♂, pitfalls in peat marsh, XI.2010 - New records III.2011, 1 ♀, III. - V.2011 (CRB); Mount LIMASSOL: Platres, Kalidonia falls track, 1 ♀, in Olympos (Chionistra), 1850 m , 1 ♂ 9 ♀♀, pitfalls woods along track, 26.IV.2007 (CAR - S). in Pinus forest, XI.2010 - III.2011 (CRB), 3 ♂♂ 2

♀♀, III. – V.2011 (CRB); near top of Mount Distribution: Europe, N. Africa. New to Cyprus. Olympos (Chionistra), 1 ♀, stones in Pinus nigra

forest, 28.IV.2007 (CAR - S); Troodos, Persephone Walckenaeria alticeps (Denis, 1952) Trail, 1732m, 2 ♀♀, in Pinus nigra forest, Walckenaeria alticeps ; Tanasevitch, 2011: 88. 19.III.2018 (CKHK).

NICOSIA: Cedar Valley Forest, 1050m, 1 ♀, Identification : https://araneae.nmbe.ch/ . litter and beating in Cedar forest, 19 .V.2010

(CRB); Cedar valley, river valley in Cedrus Citations forest, 1120 m, 1 ♀, 21.III.2018 (CKHK); LIMASSOL : Agios Dimitrios (Tanasevitch, Prodromos E., near Prodromos dam, 1550m, 3 2011). ♂♂ 2 ♀♀, pitfalls in Pinus forest, XI.2010 - NICOSIA : Cedar Valley Forest ( Tanasevitch, III.2011, 2 ♀♀, III. – V.2011 (CRB). 2011). PAPHOS: Akamas Peninsula, Agios Mina s, 1 ♀, PAPHOS: Al o noudhi junction ( Tanasevitch, in dense Pinus litter, 2.V.2007 (CAR - S). 2011).

New records Distribution: An endemic species of Cyprus. LIMASSO L : Pano Platres, 1 ♀, in grass at base of wall, 6.V.2007 (CAR - S); Pano Platr es, below Walckenaeria stylifrons (O. Pickard. - Caledonia Waterfall , 1 ♀, under stones at river, Cambridge, 1875) 1.V.2013 (CAR - S). Walckenaeria stylifrons ; Wunderlich, 1995b: 674.

Distribution: Palaearctic. Identification : https://araneae.nmbe.ch/ .

Walckenaeria antica (Wider, 1834) Citations

Without precise locality (Wunderlich, 1995b). Identification : https://araneae.nmbe.ch/ .

New records

None Spiders of Cyprus 82

(NHMW); Livadi Tou Passia, E. Troodos, 1750m, Distribution: Europe. 1 ♂, pitfalls in peat marsh, XI.2010 - III.2011 (CRB); Mount Olympos (Chionistra), 2 ♀♀, Walckenaeria n. sp. pitfalls in open Junip erus foetidissima forest, 6.IV. - 8.VI.2014 (CMH); Mount Olympos A new Walckenaeria species closely related to W. (Chionistra), 1 ♀, litter in Pinus forest, cyprusensis will be described in a separate paper. 14.IV.2017 (CAR - S); Mount Olympos (Chionistra), S. slope, 1900m, 2 ♀♀, under stones in open Pinus forest, 4.V.2010 (CRB); Prodromos Family Liocranidae S., Trooditissa picnic site, 1300m, 9 ♂♂ 2 ♀♀, pitfalls in open Pinus forest, XI.2010 - III.2011 Agroeca inopina O. Pickard - Cambridge, 1886 (CRB), 5 ♂♂ 3 ♀♀, in litter, 25.XI.2010 (CRB); Trooditissa Monastery, 5 ♂♂ 3 ♀♀, litter and Identification : https://araneae.nmbe.ch/ . stones in Quercus forest, 25.XI.2010 (CRB). NICOSIA: Ayia Koroni, Agios Epifanios ., 1 ♀, in Citations house, IV.2005 (CDM); Agridi, near Dali, 1 ♀, on None. stones, river bank, 29.IV.2013 (CDM); Athalassa National Forest Park, Nicosia, 1 ♀, under stones New records in pine plantation,8.V.2013 (CAR - S); Cedar LIMA S SOL: Trood os, near summit of Mount valley, river valley in Cedrus forest, 1120 m, 1 ♀, Olympos (Chionistra), 1 ♀, under stone in Pinus 21.I II.2018 (CKHK); Kapedes, 2 ♂♂ 1 ♀, pitfalls brutia forest, 19.IV.2017 (CAR - S); 10 km S. of in Pinus brutia forest, 17.I. - 23.IV.2016 (CMH); Troodos, 1 ♀, under stones in Pinus brutia forest, Pano Amiantos, 5 ♀♀, pitfalls in Pinus brutia 19.IV.2017 (CAR - S). forest, 6.IV. - 8.VI.2014 (CMH); road Pedoulas - NICOSIA : between Cedar Valley and Stavros tis Lemithou, 3 ♀♀, pitfalls in Pinus nigra and P. Psokas, 1 ♀, unde r stones in P. nigra forest, brutia forest, 6.IV. - 8.VI.2014 (CMH); road 29.IV.2007 (CAR - S); Troodos S., road to Pano Pedoulas - Prodromos, 2 ♀♀, pitfalls in Pinus Platres, 1 ♀, under stones in Pinus nigra forest, nigra forest, 6.IV. - 8.VI.2014 (CMH); near 1.V.2007 (CRS). Platania Scout camping site, 10 ♀♀, pitfalls in PAPHOS: Smigies, 1 ♀, under stone in hillside Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH). scrub, 26.III.1998, coll. J. Murphy (MMUE). PAPHOS : Pegeia, Pikni Forest, 346m, 1 ♀, pitfalls in former carob plantation, 11.III. - Distribution: Euro pe, Algeria. New to Cyprus. 23.III.2018 (CKHK); same site , 366m, 1 ♀, pitfalls in pine forest plantation, 11.III. - Agroeca parva Bosmans, 2011 23.III.2018 (CKHK).

Identification Distribution: Greece (Lesbos), Turkey, Israel. Agroeca parva Bosmans, 2011: 19, figs 11 - 14. New to Cyprus.

Citations Arabelia pheidoleicomes Bosselaers, 2 009 None. Arabelia pheidoleicomes Bosselaers, 2009: 51.

New records Identification FAMAGUSTA: Rizokarpaso , 2 ♀♀, Juniperus Bosselaers, 2009: 51, figs 2 K; 7 A - G; 8 h; 9. phoenicea forest, 12.III.2016 (CMH). Citations KERYNEI A: Larnakas tis Lapithou N. , 950m, 1 PAPHOS : Neo Chorio (Bosselaers, 2009); Polemi ♀, under stones in Pinus forest, 25.XI.2010 (Bosselaers, 2009). (CRB). LARNACA : Agioi Vavatsinias, 700m, ♀, litter in New records Pinus forest, 28.IV.2010 (CRB). None.

LIMASSOL: road Chionistra - Prodromos, 3 ♀♀, Distribution: Greece ( Rhodos, Lesbos) and pitfalls in Pinus nigra forest, 6.IV. - 8. VI.2014 Cyprus. (CMH); Kolossi castle, ca. 12 km W . of Limassol,

1 ♀, walls in the castle area, 11.IV.1986

Spiders of Cyprus 83

Mesiotelus cyprius Kulczyński, 1908 Akamas Peninsula, Smigies, 1 ♀, stony slope, Mesiotelus cyprius Kulczyński, 1908: 76. 25.IV.2007 (CJVK); Ak amas P eninsula, Smigies, 352m, 8 ♀♀, m aquis, 12.III.2018 (CKHK); near Identification Avakas Gorge, 1 ♀, under stone in field with tall Kulczyński, 1908: 76, pl. 2, figs 13. grass, 14.IV.2017 (CAR - S); Avakas gorge and Citations surroundings, 1 ♀, 22.III.2018 (CKHK); Evretou Without precise locality (Kulczyński, 1908). Reservoir, 1 ♀, under stone in grass tussocks , LIMASSOL: Troodos Mountain (Bosselaers & 13.IV.2017 (CAR - S); Kinousa SE, 400m, 2 ♀♀, Jocqué, 2002). under stones and litter in Pinus forest, 1.V.2010 (CRB); near Kritou Terra, 2 ♀♀, under stones in

dry fallow, 14.IV.2017 (CAR - S); Peristerona, New recor ds FAMAGUSTA : Kantara castle, 650m, 1 ♀, under Atichoullis gorge, 425m, 2 ♀♀, stones in maquis, stones in ruins, 19.III.2011 (CRB); Salamis ruins, 2.V.2007 (CJVK); S migies, W. Neo Chorio, 30m, 1 ♀, under stones in ruins, 17.III.2011 300m, 1 ♀, litter in Pinus forest, 30.IV.2010 (CRB); Komi Kebir , 150m, 1 ♀, under stones in (CRB); Souskiou, 1 ♀, 16.III.2013 (CDM). Pinus forest, 18.III.2011 (CRB). Distribution: An endemic species of Cyprus. KERYNEIA: Klepini , 300m, 1 ♂ 1 1 ♀♀, under stones in open Pinus forest, 23.XI.2010 (CRB); Mesiotelus tenuissimus (L. Koch, 1866) Larnakas tis Lapithou E ., 950m, 6 ♀♀, under stones in Pinus forest, 24.XI.2010 (CRB), 1 ♀, Identification under stones in Pinus forest, 25.XI.2010 (CRB); Bosmans et al., 2009: 35, figs 24 - 28. Palaiosofos , 1 ♀, under stones in forest, 24.XI.2010 (CRB) ; Pentadactylos Range, Ftericha Citations S ., 800m, 1 ♀, under stones in grassland, None. 17.V.2011 (CRB); Saint Hilarion Castle, 2 km W., 610m, 1 ♀, under stones in open Pinus forest, New records 20.III.2011 (CRB), 1 ♀, 17.V.2011 (CRB). LIMASSOL: Amathus, 1 ♀, on house walls, LARNACA: Delikipos, 1 ♀, in grass around 9.IV.1986 (NHMW); Pissouri, 1 ♀, under stone, sma ll dam, 18.IV.2017 (CAR - S); Mosfiloti, 1 ♀, 24.III.1998, coll. J. Murphy (MMUE). under stone in dry eroded grassland on hillside, NICOSIA: Pyrgos, 1 ♀, dry grassland, 12.IV.2017 (CAR - S). 15.IV.1986 (NHMW); Strovolos, 1 ♀, around LIMASSOL: Pano Platres, track near Caledonia hous es, 1.V.2016 (CMH). PAPHOS: Akamas Peninsula, 7 ♀♀, in coastal Waterfall , 1 ♀, under stones and in litter, grassland, 22.III.1998, coll. J. Murphy (MMUE); 26.IV.2007 (CRS); Prodromos S., Trooditissa Kinousa, 1 ♀, under stone in pine wood, picnic p lace, 1300m, 3 ♀♀, pitfalls in open Pinus 28.III.1998, coll. J. Murphy (MMUE); Smigies, 4 forest, XI.2010 - III.2011 (CRB); Troodos, near ♀♀, under stone in hillside scrub, 26.III.1998, summit of Mount Olympos (Chionistra), 1 ♀, coll. J . Murphy (MMUE). under stone in Pinus brutia forest., 9.IV.2017 (CAR - S); 10 km S. of Troodos, 1 ♀, under stone Distribution: Europe, North Africa, Ukraine, Turkmenistan. New to Cyprus. in Pinus brutia forest, 19.IV .2017 (CAR - S).

NICOSIA : Machairas, 1 ♀, Pinus brutia forest, Mesiotelus sp. 1.XI.2015 (CMH); Prodromos E., near Prodromos dam, 1550m, 1 ♀, pitfalls in Pinus forest, XI.2010 A new Mesiotelus species will be described in a - III.2011 (CRB). separate paper. PAPHOS : Agios Minas picnic site, 2 ♀♀, under stones in open pine woodland, 1 3.IV.2017 (CAR - Scotina celans Blackwall, 1841

S); Akamas Peninsula, track along the sea, 10m, 1 Identification : https://araneae.nmbe.ch/ . ♀, under stones in Juniperus phrygana, 30.IV.2010 (CRB); Akamas Peninsula, Neo Citations Chorio, 1 ♀, under stones, 25.IV.2007 (CJVK); None.

Spiders of Cyprus 84

New records orange orchard, 30.IV.2013 (CAR - S); Kolossi FAMAGUSTA : Agios Sergios , 30m, 1 ♀, pitfalls castle, 1 ♀, walls in the castle area, 11.IV.1986 in Pinus plantation, 15. - 27.V.2011 (CRB); Komi (NHMW); Pano Platres, track near Caledonia Kebir , 150m, 1 ♀, pitfalls in Pinus forest, III. - Waterfall , 2 ♂♂, under stones and in litter at edge V.2011 (CRB). of Pinus nigra forest, 26.IV.2007 (CAR - S); LIMASSOL: Mount Olympos (Chionistra), Pissouri, 1 ♀, in grass, 24.III.1998, coll. J. 1900m, 1 ♀, pitfalls in Pinus forest, III. - V.2011 Murphy (MMUE); Troodos S., road to Pano (CRB); Troodos Range, Livadi t ou Passia, 1750m, Platres, 2 ♂♂, 2 ♀♀, under stones in P. nigra 1 ♀, pitfalls in peat marsh, III. - V.2011 (CRB). forest, 28.IV.2007 (CAR - S), 1 ♂ (CRS); same site NICOSIA : Oikos N., Xerargaka picnic site, 350m, 4 ♂♂ 5 ♀♀, 1.V.2007 (CRS); 10 km S. of 4 ♂♂, under stones in Pinus forest, 20.V.2011 Troodos, 1 ♂ 1 ♀, in Pinus brutia forest, (CRB); Kato Koutrafas - Astromeritis road, 120m, 19 .IV.2017 (CAR - S). 1 ♀, herbs on pond shore, 18.V.2011 (CRB). NICOSIA: Agia Koroni, Agios Epifanios, 1 ♂, in PAPHOS : 9 km N. Pano Panagia, 1050m, 1 ♀, in house, IV.2005 (CDM); Agridi, near Dali, 1 ♀, in litter near spring, 19.V.2011 (CRB). garden, IV.2004, 2 ♂♂ 1 ♀, in field, V.2008, 1 ♀, on veranda, 12.IV.2008, 1 ♂, on veranda, Distribution: Europe, Algeria, Russia, Georgia. 12.IV.2009, 1 ♂ 1 ♀, in garden, 14.III.2010, 1 ♀ , New to Cyprus. in garden, 20.VI.2011 (all CDM); Athalassa National Forest Park, Nicosia, 1 ♀, under stone in pine plantation, 2.V.2013 (CDM); Cedar Valley Family Lycosidae Forest, 1 ♀, under stones by track, 29.IV.2007 (CRS); Cedar Valley, 1150m, 2 ♂♂ 2 ♀♀, Alopecosa albofasciata (Brullé, 1832) 21.III.2018, for est road, in gravel (CKHK); same Alopecosa albofasciata ; Bristowe, 19 35: 768. site , 1 ♂, 19.V.2011 (CJVK); between Cedar Valley and Stavros tis Psokas, 1 ♂, under stones Identification : https://araneae.nmbe.ch/ . in P. nigra forest, 29.IV.2007 (CAR - S); Dali, Lymbourena Hill, 2 ♂♂, in field, V.2004 (CDM); Citations Chrysoroyiatissa Monastery, 2 ♂♂ 1 ♀, in pine Without precise locality (Bristowe, 1935). and oak woodland, 2.V.1982 , coll. A. Stubbs (MMUE); Dali , 13, Agios Andronikos, 1 ♀, in New records garden, V.2008 (CDM); Ikos N., Xerargaka picnic FAMAGUSTA : Agios Sergios , 30m, 3 ♂♂ 1 ♀, site, 350m, 1 ♀, stones in Pinus forest, 20.V.2011 in pitfalls in Pinus plantatio n, 15 . - 27.V.2011 (CJVK); Kapouti , 100m, 1 ♂, under stones in (CRB); Gialousa E ., Agios Thyrsos, 20m, 1 ♀, in grassland, 20.III.2011 (CRB). litter under fig tree, 18.III.2011 (CRB); Gialousa PAPHOS : Agia, 1 ♂, stones at picnic site, E., Agia Triada, 50m, 1 ♂ 1 subadult ♀, under 30.IV.2007 (CJVK); Agios Georgios, Potamos stones in ruins, 18.III.2011 (CRB) ; Kantara castle, Diarizos, 2 ♂♂ 1 ♀, pitfalls at border or reed bed, 650m, 1 subadult ♂, under stones in ru ins, 26.V – 5.V.2007 (CJVK); Akamas Peninsula, 19.III.2011 (CRB). Smigies, 352m, 1 ♂, maquis, 12.III.2018 KERYNEIA: Keryneia , 1 ♂, 28.IV.1936, E. (CKHK); Drouseia S., Panagia Chry sospiliotissa, Green leg. (BMNH); Vouni ruins N., 250m, 1 ♂, 580m, 1 ♂, night catch in small cave, 22.IV.2017, under stones in ruins, 20.III.2011 (CRB). S. Huber leg. (CSH), Gialia, 1 ♀, under stones in LARNACA : Achnas Dam, 50m, 4 ♀♀, under pine wood, 25.III.1998, coll. J. Murphy (MMUE); stones and litter bordering lake, 27.V.2011 Kato Akourdaleia, 2 ♂♂, in guest house (CJVK); Delikipo s, near Kornos, 1 ♀, under swimming pool, 3.V.2007 (CRS); Kato Paphos stones in grass of open pine wood, 9.V.2013 archaeologi cal site, under stones in grass, 1 ♀, (CAR - S); Larnaca SW., Hala Sultan Teke 10.V.2007 (CAR - S); Kinousa E., Agios mosque, 5m, 1 ♀, salt marsh, 15.V.2011 (CJVK); Merkourios, 350m,1 ♀, under stones and litter in Stavrovouni Monastery, 350m, 1 ♀, in pitfalls in Pinus forest, 1.V.2010 (CJVK); Lysos E., 550m, 1 open Pinus forest, III. - V.2011 (CRB). ♀, while digging for Nemesia , 27.IV.2017, S. LIMASSOL :); Agios Georgios E., Palaiomylos, Huber leg. (CSH); Mavrokolympos Reserv oir, 1 1250m, 1 ♀, under stones in mixed Pinus and ♀, 13.III.2018 (CKHK); Neo Chorio, 1 ♂, in Quercus forest, 4.V.2010 (CRB); Akrotiri grassy scrub, 17.III.1998, coll. J. Murphy Peninsula, near Agios Nikolaos Monastery, 25m, (MMUE); Oikos, 700m, 1 ♂, limy slope, 2 ♀♀, in grassy ditch, 23.III.2011 (CRB); Akrotiri 23.IV.2017, S. Huber leg. (CSH); Oikos N., Peninsula, near Fasouri reedbeds, 1 ♀ , in irrigated Xerargaka picnic site, 350m, 1 ♀, under stones in

Spiders of Cyprus 85

Pinus forest, 20.V.2011 (CRB); P egeia, Pikni Troodos, Persephone Trail, 1732m, 1 ♀, in Pinus Forest, 345m, 13 ♂♂, pitfalls in former carob nigra forest, 19.III.2018 (CKHK). plantation, 11.III. - 23.III.2018 (CKHK); same NICOSIA: Lakatamia, 1 ♀, in olive grove, 1.I. - site , 365m, 4 ♂♂, pitfalls in pine forest, 11.III. - 1.X. 2016 (CMH); Pano Amiantos, 1 ♀, pitfalls in 23.III.2018 ( CKHK); same site, 375 m, 5 ♂♂, Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH); pitfalls in open ruderal site, 14.III. - 23.III.2018 near Platania Scout camp ing site, 1 ♂ 2 ♀♀, ( CKHK); Souskiou, 1 ♀, 16.III.2013 (CDM); pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 Stavros - Lysos road, 900m, 1 ♂, leaf litter, (CMH); near Prodromos reservoir, 2 ♀♀, pitfalls 21.IV.2017, S. Huber leg. (CSH). in Pinus nigra forest, 6.IV. - 8.VI.2014 (CMH). P APHOS: Akamas Peninsula, Avgas E stuary, 2 Distribution: Mediterranean to Central Asia. ♂♂, stones in rock strewn s lope near beach, 31.III.1997 (CJB); Paphos, Bay at hotel “Paphian Alopecosa sp. Bay”, 6 ♂♂ 1 ♀, pitfall traps on rocky beach, 1 - ? Tarentula simoni ; Kulczyński, 1908: 83 6.IV.1997 (CJB), 4 ♂♂ 4 ♀♀, stones in rock (misidentification) . strewn terrain near beach, 5.IV.1997 (CJB); Paphos, Tombs of the Kings, 1 ♂ 1 ♀, stones in Citations rock strewn terrain, 1.IV .1997 (CJB). Without precise locality (Kulczyński, 1908, sub Tarentula simoni ). Distribution: Holarctic. New to Cyprus.

Comments Arctosa cinerea (Fabricius, 1777) Two species of Alopecosa oc cur on Cyprus, the very common A . albofasciata and a nother Identification : https://araneae.nmbe.ch/ . common species , currently under study. Most probably this is the spe cies that Kulczyński ( 1908 ) Citations mentioned as Tarentula simon . It is closely related None to that species but it is perhaps a new species. New records Records LIMASSOL : Alassa SE, near river Limnatis, 1 ♀, FAMAGUSTA: Cape Greco, Ayia Napa, 35m, 1 under large stone in riverine shingle, 5.V.2007 ♀, under stones in phrygana, 20.XI.2010 (CRB). (CAR - S). LARNACA : Mosfiloti, 1 ♀, under stone in dry eroded grassland on hillside, 12.IV.2017 (CAR - Arctosa leopardus (Sundevall, 1833) S). Tarentula brevispina Kulczyński, 1908: 80 (descr. LIMASSOL : road Chionistra - Prodromos, 2 ♀♀, ♀). pitfalls in Pinus nigra forest, 6.IV. - 8.VI.2014 (CMH ); road Kakopetria - Pedoulas , Marathos Identification : https://araneae.n mbe.ch/ . picnic site, 2 ♂♂ 1 ♀, 25.XI.2010 (CRB); Livadi T ou Passia, E. Troodos, 1750m, 1 ♂ 1 ♀, pitfalls Citations in peat marsh, XI.2010 - III.2011(CRB); Mount Without precise locality (Kulczyński, 1908, sub Olympos (Chionistra), 1900m, 3 ♂♂ 3 ♀♀, Tarentula brevispina ). pitfalls in Pinus f orest, XI.2010 - III.2011 (CRB), 3 ♀♀, pitfalls in open Juniperus foetidissima New records forest, 6.IV. - 8.VI.2014 ( CMH) , 2 ♂♂ 1 ♀, FAMAGUSTA : Gialousa E., Agios Therissos , 1820m, under stones in Pinus forest , 25.XI.2010 20m, 4 ♂♂ 2 ♀♀, stones around spring, (CRB), 1 ♂, pitfalls in Pinus forest, XI.2010 - 16.V.20111 (CRB). III.2011 (CRB); Pano Platres E., Mesa Potamos LARNACA: Achna Reservoir, 50m, 11 ♂♂ 2 Waterfall s, 1 ♂, 950m, in litter in Pinus forest, ♀♀, pitfalls in marshy area around lake, 15 - 4.V.2010 (CRB); Troodos Square, 2 ♀♀, pitfalls 27.V.2011 (CRB), 2 ♂♂ 1 ♀, hand catch at same in open, mixed Pinus nigra and Juniperus site, 15.V.2011 (CJVK, CRB); Livadia,1 ♀, river foetidissima forest, 6.IV. - 8.VI.2014 (CMH); bed, 20.IV.2016 (CMH). Troodos Square, 1 ♀, Pinus nigra forest, LIMASSOL : Akrotiri Peninsula, Fasouri reed 11.V.2016 (CMH); Troodos, near summit of beds, 3 ♂♂ 1 ♀, in reed litter, 27.IV.2007; sa me Mount Olympos (Chionistra), 2 ♀♀, under stones site, 3 ♂♂, 4.V.2007, 2 ♀♀, 30.IV.2013 (all in Pinus brutia forest, 19.IV.2017 (CAR - S); CAR - S); idem, Fasouri marsh, 1 ♀, in low grass, 17.VI.2016 (CMH).

Spiders of Cyprus 86

NICOSIA: Agridi, near Dali, 3 ♂♂, edge of river, C itations 22.IV.2012 (CDM); Athalassa National Forest None Park, Nicosia, 2 ♂♂, by stream, 13.IV.2008 (CDM) ; Kapedes S., 600m, 2 ♂♂ 1 ♀, herbs and New records stones along rivulet, 5.V.2010 (CJVK); Lympia PAPHOS: Evretou Reservoir, 350m, 2 ♂♂, stones Reservoir, 6 ♂♂ 5 ♀♀, under stones below dam around lake, 1.V.2007 (CJVK). wall, 9.V.2013 (CAR - S); Marathassa Dam, 550m, 1 ♂, stones around lake, 20.V.2011 (CJVK). Distribution: Canary Is., Mediterranean. New to PAPHOS : Akamas Peninsula, Agios Mi nas, 2 Cyprus. ♀♀, wet area by stream, 2.V.2007 (CAR - S); Avakas Gorge, 50m, 3 ♂♂ 3 ♀♀, litter in citrus Aulonia kratochvili Dunin, Buchar & Absolon, yard at entrance of gorge, 30.IV.2010 (CJVK, 1986 CRB); Evretou Reservoir, 250m, 4 ♂ 1 ♀, 1.V.2007 (CJVK); Kelefos bridge, 550m, 1 ♂♂ 2 Identification ♀♀, dam on Diarizos River, un der stones, Dunin, Buchar & Absolon, 1986: 28, figs 1 - 7; 21.V.2011 (CJVK, CRB); Stavros tis Psokas, 2 Kronestedt, 1997: 77, figs 1 - 2. ♂♂ 1 ♀, along small pond, 30.IV.2007 (CJVK). Citations Distribution: Palearctic. None.

Arctosa similis Schenkel, 1938 New records LARNACA : Larnaca SW, Hala Sultan Tekke Identification mosque, 50m, 1 ♀, in pitfalls in salt marsh, 15 - Buchar, Knoflach & Thaler, 2006 : 334, figs 3 - 4, 28.V.2011 (CRB). 10, 12, 19 - 20, 27 - 30. LIMASSOL: Akrotiri Peninsula N., 5m, 6 ♀♀, litter in Eucalyptus forest, 29.IV.2010 (CJVK, Citations CRB); Fasouri, Akrotiri Peninsula, 4 ♀♀, in reed None litter, 27.IV. and 4.V.2007 ; same site , 1 ♂ 2 ♀♀ (CRS), 1 ♀, 30.IV.2013 (CAR - S). New records LARNACA : Achna Reservoir, 50m, 2 ♂♂, stones Distribution: Greece to Central Asia. New to and litter bordering lake, 16.V.2011 (CJVK, Cyprus. CRB); same site, 2 ♀♀, in pitfalls, 15 – 27.V.2011 (CJVK). Hogna ferox (Lucas, 1838) NIC OSIA: Kapedes S., 600m, 1 ♂, herbs and stones along rivulet, 5.V.2010 (CJVK). Identification PAPHOS : Evretou Reservoir, 250m, 1 ♂ 5 ♀♀, Hepner & Paulus, 2009 : 341, figs 14 - 16. along lake, 1.V.2007 (CJVK); Evretou Reservoir, Stavros tis Psokas river, 1 ♂, 20.III.2018 Citations (CKHK); Kelefos bridge, 1 ♂, dam on Diariz os None. River, 550m, 2 ♀♀, under stones, 21.V.2011 (CJVK, CRB); Mavrokolympos Reservoir, NE New records Potima, 150m, 1 ♂ 1 ♀, under stones along lake, LARNACA: Achna Reservoir, 50m, 1 ♂ 2 ♀♀, 22.V.2011 (CRB); Mavrokolympos Reservoir, 1 under stones and litter bordering lake, 29.V.2011 ♂ 4 ♀♀, 13.III.2018 (CKHK); same site , 1 ♀, (CJVK). stones around lake, 22.V.2011 (CJ VK). LIMASSOL : Am athus, 1 ♂, under stones in dry valley, 6.IV.1986 (NHMW); Amathus, 1 ♀ 1 Distribution: Canary Is., Mediterranean. New to juvenile, front yard in hotel, 8.IV.1986 (NHMW); Cyprus. Mouttagiaka, 20m, 2 ♂ 1 ♀ 2 subadult ♀♀, drown in swimming pool of Caravel hotel, Arctosa variana C. L. Koch, 1847 29.IV.2010 (CRB). PAPHOS: Evretou Reservoir, 350m, 3 ♀♀, stones Identification around lake, 1.V.2007 (CJVK). Buchar, Knoflach & Thaler, 2006 : 329, figs 6 - 7, 13 - 14, 21 - 22, 31. Distribution: Canary Islands, Mediterranean. New to Cyprus.

Spiders of Cyprus 87

Hogna radiata (Latreille, 1817) Citations None. Identification Loksa,1972: 49, figs 44 - 45. New records LIMASSOL : Liva di tou Passia, E. Troodos, Citations 1170m, 8 ♂♂ 6 ♀♀, in peat marsh, 3.V.2010 None. (CJVK, CRB) ; Troodos S., 1350m, 1 ♂, litter in mixed Pinus - Quercus forest, 3.V.2010 (CJVK). New records LIMASSOL: Akroti ri Peninsula, 1 ♂, margin of Distribution: Greece to Central Asia. New to central salt lake, 7.V.1972, coll. A. Stubbs Cyprus. (MMUE); Τ rooditissa Monastery, 1300m, 1 ♂, under stones in Quercus forest, 25.XI.2010 Pardosa atomaria (C. L. Koch, 1847) (CRB). Lycosa atomaria ; Kulczyński, 1908: 80. NICOSIA: Agridi, near Dali, 1 ♀, in garden, Pardosa atomaria ; Buchar & Thaler, 2002: 446, IV.2004, 2 ♂♂, on veranda, XI.2009, 1 ♂, in figs 11 - 18. lawn, 1.X.2010, 1 ♂, in garden, VIII.2012 (all CDM); Dali 13, Agios Andronikos, 1 ♂, in Identification garden, IV. - V.2008 (CDM). Buchar & Thaler, 2002: 446, figs 11 - 18. PAPHOS: Kato Akourdaleia, 1 ♀, in garden, VIII.2005 (CDM) Citations Without precise locality (Kulczyński, 1908). Distribution: Central Europe to Central Asia, LIMASSOL : Troodos (Buchar & Thaler, 2002). Iran, Central Africa. New to Cyprus. NICOSIA: Cedar valley Forest, 1100m (Buchar & Thaler, 2002). Lycosa praegrandis C. L. Koch, 1836 PAPHOS : Akamas Peninsula, Avakas Gorge (Buchar & Thaler, 2002). Identification Zyuzin & Logunov , 2000: 306, figs 1 - 2. New records FAMAGUSTA: Paralimni Lake, 65m, 2 ♂♂ 2 Citations ♀♀, herbs and litter around dried lake, None. 27.IV.2010 (CRB). LARNACA : Delikipos, 2 ♀♀, in moist grass New records around small dam, 18.IV.2017 (CAR - S). FAMAGUSTA: Agia Triada, 1 ♂, under stones in LIMASSOL : Akrotiri Peninsula, 1 ♂ 3 ♀♀, on carob grove, 19.VI.2016 (CMH); Protaras, 1 ♂, in muddy ground, edge of reedbed, 17.IV.2017 hotel, 6.VII I.2016 (CMH). (CAR - S); Fasouri, Akrotiri Peninsula, 1 ♀, swept KERYNEIA: Saint Hilarion Castle, 2 km W., from grass, 4.V.2007 (CAR - S); G ermasogeia 610m, 1 subadult ♀, stones in open Pinus forest, Dam, 1 ♀, 20.II.2011, D. W. Wrase leg. (CKHK); 20.III.2011 (CRB). Pano Platres, below Caledonia W aterfall, 1 ♂, LIMASSOL : Agros, 1 ♀, night catch in Pinus under stones and in grass in rocky valley, brutia forest, 20.V.2016 (CMH); Mount Olympos 1.V.2007 (CAR - S); Pano Platres, Millomeris (Chionistra), 1820m, 1 subadult ♀, unde r stones Waterfalls, 1 ♀, in litter, 28.IV.2007 (CAR - S); in Pinus forest, 25.XI.2010 (CRB), 1 ♀, 4.V.2010 Parekklisia, 1 ♀, under stones in riverbed, (CJVK), 1 ♀, 23.V.2011 (CJVK). 13.IV.1986 (NHMW); Troodos S., road to Pano NICOSIA: Anthoupolis, 1 ♀, night catch in Platres, 2 ♀♀, under stones in P. nigra forest, gardens, 13.III.2016 (CMH); road Pedoulas - 28.IV.2007 (CRS); Prodromos S., Trooditissa Lemithou, 1 subadult ♀, pitfalls in Pinus nigra picnic site, 1450m, 1 ♂, under stones in Pinus and Pinus brutia forest, 6.IV. - 8. VI.2014 (CMH). forest, 21.V.2011 (CRB). NICOSIA: Agridi, ne ar Dali, 2 ♂♂ 4 ♀♀, by Distribution: Greece to to Kazakhstan. New to edge of river, 22.IV.2012; in garden, 1 ♀, Cyprus. 1.V.2012 (CDM); Athalassa National Forest Park, 1 ♀, by stream, 13.IV.2008 (CDM), 1 ♂ 3 ♀♀, in Pardosa albatula (Roewer, 1951) eucalyptus litter by edge of lake, 2.V.2013 (CAR - Identification : https://araneae.nmbe.ch/ . S); Cedar Valley Forest, 1180m, 1 ♂ 3 ♀♀, litt er in Cedrus forest, 19.V.2011 (CJVK); same site , 2

Spiders of Cyprus 88

♂♂ 1 ♀, under stones by track, 29.IV.2007 Citations (CRS); Kapedes S., 600m, 3 ♂ 6 ♀♀, herbs and None. stones along rivulet, 5.V.2010 (CJVK, CRB); Lympia Reservoir, Lympia, 6 ♂♂ 2 ♀♀, under New records stones below dam wall, 9.V.201 3 (CAR - S); NICOSIA: Agridi, near Dali, 1 ♂, in garden, Prodromos E., near Prodromos Dam, 1550m, 1 ♀, 20.VI.2011 (CDM); Cedar Valley Forest, 2 ♂♂ 2 stones in Pinus forest, 2.V.2010 (CRB); ♀♀, edge of track in cedar forest, 2.V.1982, coll. Prodromos S., Trooditissa picnic site, 1450m, 1 ♂ A. Stubbs (MMUE); Kythrea, 1 ♂, no date, G. P. 3 ♀♀, under stones in Pinus forest, 21.V.2011 (CJVK). Georghiou leg. (BMNH). PAPHOS: Agia, 1 ♀, under stones at picnic site, LIMASSOL : Fasouri, Akrotiri Peninsula, in reed 30.IV .2007 (CJVK); Agios Georgios, Dhiarizos litter, 27.IV.2007 1 ♂ 4 ♀♀, 1 ♂, 4.V.2007 river, 1 ♂, pitfalls at edge of reed bed, 26.IV – (CRS), 4 ♂♂ 1 ♀, 4.V.2007 (CAR - S). 5.V.2007 (CJVK); Agios Merkouris, 4 ♂♂ 5 ♀♀, banks of river in forest, 27.IV.2007 (CJVK); Distribution: Palaearctic, Canary Islands, Azores. Akamas Peninsula, Agios Minas, 1 ♂ 1 ♀, wet New to Cyprus. area by stream, 2.V.2007 (CAR - S) ; Avakas Gorge, 150m, 5 ♂ 8 ♀♀, litter in citrus yard at Pardosa roscai (Roewer, 1951) entrance of gorge, 30.IV.2010 (CKVK, CRB), 2 ♂♂, stones in gorge, 22.V.2011 (CRB), 1 ♂ 2 Identification ♀♀, 5.IV.1997 (CJB); Dhiarizos river near Bayram et al. 2009: 464, figs 1 - 5. Kelefos bridge, 444m, 4 ♂♂ 5 ♀♀, open river bank, 16.III.2018 (CKHK) ; Evretou Reservoir, Citations 350m, 4 ♀♀, in grassland around lake, 1.V.2010 None. (CRB); Kelefos bridge, 4 ♂♂ 4 ♀♀, dam on Diarizos River, 550m, under stones 21.V.2011 New records (CRB); Kinousa E., Agios Merkourios, 350m, 2 FAMAGUSTA : Gialousa E ., Agios Thyrsos, ♂♂ 5 ♀♀, under stones and litter in Pinus forest, 20m, 2 ♂♂ 2 ♀♀, stones in ruins, 18.III.2011 1.V.201 0 (CJVK, CRB); Livadi, 400m, 4 ♂♂ 10 (CRB); Salamis S., Silver Beach, 5m, 8 ♂♂ 1 ♀, ♀♀, under stones along rivulet, 28.IV.2007 (CJVK); Marathassa Dam, 550m, 2 ♂♂ 3 ♀♀, in pitfalls in salt marsh, 15 - 27.V.2011 (CJVK, stones around lake, 20.V.2011 (CJVK); CRB). Mavrokolympos Reservoir, 2 ♀♀, 13.III.2018 LARNACA : Achna Reservoir, 50m, 5 ♂ 5 ♀♀, (CKHK); 9 km N. Pano Panagia, 1050m, 4 ♀♀, in under stones and litter bordering lake, 21.XI.2010, litter n ear spring, 19.V.2011 (CJVK, CRB); 1 ♂ 2 ♀♀, 16.V.2011, 347 ♂♂ 56 ♀♀, 15 - Stavros tis Psokas S., 1 ♂ 1 ♀, banks of small 27.V.2011, pitfalls at same site (all CJVK, CRB); river, 30.IV.2007 (CJVK). Larnaca SW, Hala Sul tan Tekke mosque, 50m, 23 Distribution: Balkans, Cyprus, Rhodes, Aegean ♂♂ 7 ♀♀, in pitfalls in salt marsh, 15 - 28.V.2011 Is. (CRB), 1 ♂ 2 ♀♀, hand catch at same site, 15.V.2011 (CJVK); Paralimni Lake, 65m, 1 ♀, Pardosa proxima (C. L. Koch, 1847) litter along dry lake, 27.IV.2010 (CJVK). Pardosa tenuipes L. Koch, 1882 LIMASSOL: Akrotiri Peninsula, salt lake, 1 ♀,

unde r Salicornia , 15.III.2018 (CKHK). Identification Pardosa proxima : Isaia et al. 2018: 7, figs 1 - 2, 5 - PAPHOS: Agios Georgios, Dhiarizos river, 2 ♂♂ 10, 19, 21 - 22, 26. 1 ♀♀, pitfalls along reed bed, 26.IV – 5.V.2007 Pardosa tenuipes Isaia et al. , 2018 : 12, figs 3 - 4, (CKVK); Dhiarizos river near Kelefos bridge, 11 - 16, 20, 23. 444m, 1 ♀, open river bank, 16.III.2018 (CKHK); Evretou Reservoir, Stavros tis Ps okas river, 3 ♀♀, Comments 20.III.2018 (CKHK); Evretou Reservoir, 350m, 1 Recently, Isaia et al. (2018) pointed out the ♀, shrubs and grassland around lake, 1.V.2010 differences between Pardosa proxima and (CRB); same site , along lake, 2 ♂♂ 12 ♀♀, P. tenuipes . When our specimens were identified, 1.V.2007 (CJVK); Mavrokolympos Reservoir, NE this study was not yet available and these species Potima, 150m, 1 ♂ 1 ♀, under stones along lake, c ould not be distinguished in the present paper. 22.V.2011 (CJVK).

Spiders of Cyprus 89

Distribution: Bulgaria, Romania, Turkey, Iran. Citations New to Cyprus, where it is locally very abundant None. in wetlands around reservoirs. New records Pirata piraticus (Clerck, 1757) LIMASSOL : Akrotiri Peninsula, Fasouri reed beds, in reed litter, 1 ♂ 1 subadult ♀, 27.IV.2007, Identification : https://araneae. nmbe.ch/ . 1 ♀, 4.V.2007, 1 ♀, 30.IV.2013 (CAR - S), 2 ♀♀

(CDM). Citations None. NICOSIA: Agridi, near Dali, 1 ♂, in garden, V.2010 (CDM); near Prodromos reservoir, 1 ♂ 3 New records ♀♀, pitfalls in Pinus nigra forest, 6.IV. - PAPHOS: Agios Georgios, Potamos Diarizos, 2 8.VI.2014 (CMH); Xerargaka S, Orkontas, 3 ♀♀, ♂♂, pitfalls in reed bed, 26.IV – 5.V.2007 riverside vegetation, 26.III.2016 (CMH). (CJVK); Agios Mamas, 215m, 2 ♂♂ 4 ♀♀, near PAPHOS : Akamas Peninsula, Agios Minas, 1 ♀, spring, 28.IV.2007 (CJVK). wet area by stream, 2.V.2007 (CAR - S); Kato Paphos, 1 subadult ♂, seevin g hay, 4.IV.1997 Distribution: North America, Europe, Turkey, (CJB). Caucasus, Russia, Central Asia, China, Japan . Distribution: Circum - mediterranean. New to Trabea paradoxa Simon 1876 Cyprus.

Identification Wadicosa fidelis (O. Pickard - Cambridge, 1872) Russell - Smith, 1982: 73, figs 1A - D, 2A - E, 11A. Identification Citations Kronestedt & Zyuzin 2009: 813, figs 1 - 17. None. Citations New records None. FAMAGUSTA : Kantara E., 610m, 1 ♂, stones in New records open Pinus forest, 25.V.2011 (CRB); Paralimni LARNACA : Achna Reservoir, 50m, 1 ♂ 2 ♀♀, Lake, 65m, 1 subadult ♀, h erbs and litter around under stones and litter bordering lake, 21.XI.2010, dried lake, 27.IV.2010 (CRB). 2 ♂ 8 ♀♀, 16.V.2011, 14 ♂♂ 9 ♀♀, pitfalls at LARNACA : Agios Minas, 1 ♀, in litter in pine wood, 20.III.1998, coll. J. Murphy (MMUE). same site, 15 . - 27.V.2011 (CJVK, CRB). LIMASSOL: Akrotiri Peninsula N., 5m, 1 ♂, LIMASSOL : Alass a SE, 1 ♂, under pebbles and litter in Eucalyptus forest, 29.IV.2010 (CRB); in grass at edge of river Limnatis, 5.V.2007 Pano Platres E., Mesa Potamos Waterfall s, 950m, (CAR - S). 2 ♂♂ 3 ♀♀, litter in Pinus forest, 4.V.2010 NICOSIA : Cataldere dam, Kythrea N ., 105m, 1 (CJVK, CRB); road Pano Platres - Troodos, 1 juvenile, under stones bordering lake, 23.XI.2010 juvenile, under stones in Pinus nigra forest, (CRB); Lympia Reservoir, Lympia, 1 ♀, under 1.V.2007 (CAR - S). stone below dam wall, 9.V.2013 (C AR - S). NICOSIA: Athalassa National Forest Park, PAPHOS : Evretou Reservoir, 350m, 3 ♂♂ 7 ♀♀, Nicosia, 1 ♂ 3 ♀♀, in deep eucalyptus litter, border of lake, 1.V.2007 (CJVK); 8.V.2013 (CAR - S); near Platania Scout camping Gefira Kelefou, along Diarizos Potamos, 1 ♂, site, 1 ♀, pitfalls in Pinus brutia forest, 6.IV. - 21.V. 2001 (CJVK); Mavrokolympos Reservoir, 8.VI.2014 (CMH); near Prodromos reservoir, 2 NE Potima, 150m, 2 ♂♂ 2 ♀♀, under stones ♂♂, pitfalls in Pinus nigra forest, 6.IV. - 8.VI.2014 (CMH). along lake, 22.V.2011 (CJVK, CRB); Ma vrokolympos Reservoir, 4 ♂♂ 1 ♀, 13.III.2018 Trochosa hispanica Simon, 1870 (CKHK).

Ide ntification Distribution: Palearctic, Canary Is. New to Thaler, Buchar & Knoflach, 2000: 1079, figs 37 - Cyprus. Common along lakes and rivers. 39, 41.

Spiders of Cyprus 90

Family Mimetidae LIMASSOL: Trooditissa Monastery, Trooditissa river, 1360m, 1 ♀, beaten from shrubs, Ero aphana (Walckenaer, 1802) 19.III.2018 (CKHK). PAP HOS : Kykkos Monastery, 1150m, 1 ♀, on Identification : https://araneae.nmbe.ch/ . walls in monastery, 19.V.2011 (CRB).

Citations Distribution: Palaearctic. New to Cyprus. None. Mimetus laevigatus (Keyserling, 1863) New records Identification : https://araneae.nmbe.ch/ LIMASSOL: Akrotiri Peninsula, 1 ♂, margin of central salt lake, 7.V.1972, coll. A. Stubbs Citations (MMUE). None. NICOSIA : Lympia Reservoir, Lympia, 1 ♀, in rushes below dam wall, 9.V.2013 (C AR - S); Marathassa Dam, 550m, 1 ♀, beaten from shrubs New records around lake, 20.V.2011 (CRB); Oikos N., FAMAGUSTA : Gialousa E ., Agios Thyrsos, Xerargaka picnic site, 350m, 1 ♀, beaten from 20m,1 subadult ♀, beaten from shrubs, shrubs and stones in Pinus forest, 20.V.2011 16.V.20111 (CRB); Gialousa E ., Agia Triada, (CRB). 50m, 1 subadult ♀, on herbs in ruins, 18.III.2011 PAPHOS: ca. 2 km S of Cape Geronnisos, (CRB). Akamas Peninsula, 1 ♀, u nder stones in dry KERYNEIA: Pentadactylos Range, Sa int Hilarion phrygana, 3.V.2007 (CRS); Evretou Reservoir, Castle W, 600m, 1 subadult ♂ 1 subadult ♀, Stavros tis Psokas river, 1 ♂ 1 ♀, 20.III.2018 beaten in Pinus forest 17.V.2011 (CRB). (CKHK); near Souskiou, 1 ♀, on bush in dry LIMASSOL : Akrotiri Peninsula, 1 ♂, on muddy grassland, 1.V.2013 (CDM). ground, edge of Fasouri reedbeds, 17.IV.2017

(CAR - S); Pano Platres, track near Caledonia Distribution : Palaearctic. New to Cyprus. Waterfall, 1 juvenile, sw ept from herbs and shrubs Ero flammeola Simon, 18 81 in rocky stream valley, 26.IV.2007 (CAR - S). NICOSIA : Cedar Valley Forest, 1180m, 1 Identification subadult ♀, beaten in Cedrus forest, 19.V.2011 Ero flammeola ; Thaler et al. 2004: 362. (CRB); Kampos NE, Xeros River, 550m, 1 subadult ♂, beaten from shrubs, 20.V.2011 Citations (CRB); Marathassa Dam, 550m, 1 subadult ♀, None. beaten from shrubs around lake, 20.V.2011 New records (CRB); Oikos N., Xerargaka picnic site, 350m, 1 NICOSIA : Kato Koutrafas - Astromeritis road, subadult ♀, beaten from shrubs in Pinus forest, 120m, 1 ♀, herbs on pond shore, 18.V.2011 20.V.2011 (CRB). (CRB). NICOSIA: Cedar Valley Forest, 1180m, 1 subadult ♀, beaten in Cedrus fores t, 19.V.2011 Distribution: Palaearctic. New to Cyprus. (CRB). PAPHOS : Akamas Peninsula, Neo Chorio, 1 Ero t uberculata (De Geer, 1778) subadult ♂, swept from phrygrana and grass,

Identification : https://araneae.ch/ . 2.V.2007 (CAR - S).

Citations Distribution : Mediterranean to Central Asia. None. New to Cyprus.

New records

Spiders of Cyprus 91

Family Miturgidae Identification: https://araneae.nmbe.ch/ .

Zora manicata Simon, 1878 Citations Identifi cation : https://araneae.nmbe.ch/ . None.

Citations None. New records FAMAGUSTA : Gialousa , Agios Thyrsos, 1 ♀, New records 16.V.2011 (CRB). FAMAGUSTA : Gialousa E ., Agia Triada, 50m, 1 PAPHOS: Kelefos bridge, 465m, deciduous wood, 1 ♀, i n litter, 16.III.2018 (CKHK). ♂, stones in ruins, 18.III.2011 (CRB).

KERYNEIA: Saint Hilarion Castle, 610m, 1 ♀, in Distribution: Cyprus, Syria, Israel. New to pitfa lls in Pinus forest, XI.2010 - III.2011 (CRB). Cyprus. LARNACA : Delikipos, 1 ♀, under stones in grass around small dam, 8.IV.2017 (CAR - S); Dipotamos Reservoir, 150m, 1 ♀, herbs and Family Nemesiidae stones bordering lake, 28.IV.2010 (CRB). LIMASSOL : Pano Platres, track near C aledonia Brachythele incerta Ausserer, 1871 Waterfall, 2 ♂♂ 5 ♀♀, under stones and in grass Brachythele incerta Ausserer, 1871: 175, descr. in rocky valley, 26.IV.2007 (CAR - S), 4 ♂♂ ♂. (CRS); road Pano Platres - Troodos, 1 ♂, under stones in Pinus nigra forest, 28.IV.2007 (CAR - S). Identification NICOSIA : Athalassa National Forest Park, See page 28. Nicosia, 2 ♀♀, unde r stones in pine plantation, Citations 8.V.2013 (CAR - S); between Cedar Valley and Without precise locality (Ausserer, 1871). Stavros tis Psokas, 1 ♀, under stones in P. nigra forest, 28.IV.2007 (CAR - S); Lympia Reservoir, New records Lympia, 6 ♀♀, in rushes below dam wall, None. 9.V.2013 (CAR - S); Oikos N., Xerargaka picnic sit e, 350m, 2 ♀♀, under stones in Pinus forest, Distribution 20.V.2011 (CRB); Pano Amiantos, 2 ♂♂, pitfalls Only known from Cyprus. No Brachythele has been recorded from the island since the original in Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH); description. Prodromos S., Trooditissa picnic site, 1450m, 1 ♀, stones in Pinus forest, 21.V.2011 (CRB) . Nemesia sp. PAPHOS : Akamas Penins ula, 1 ♀, in coastal At a few locations in Cyprus, Nemesia spe cimens grassland, 22.III.1998, coll. J. Murphy (MMUE); were discovered. The genus was not previously Akamas Peninsula, Agios Minas, 1 ♂, wet area by known from Cyprus and the species will be stream, 2.V.2007 (CAR - S); Makounta, 1 ♂, under described in a separate paper. stones in scrub, 19.III.1998, coll. J. Murphy (MMUE); Pegeia, Pikni Forest, 366m, 1 ♂, Family Oecobiidae pitfa lls in pine forest, 11.III. - 23.III.2018 (CKHK); same site , 374m, 2 ♂♂, pitfalls in open Oecobius cellariorum (Dugès, 1836) ruderal site, 14.III. - 23.III.2018 (CKHK). Identification : https://araneae.nmbe.ch/ . Distribution: Europe, Ukraine, Israel. New to Citations Cyprus. None.

New records Family Mysmenidae LIMASSOL: Agios Tychonas, 1 ♂ 1 ♀, under stones, bushes near house walls, 19.IV.1986

Mysmena leucoplagiata (Simon, 1880)

Spiders of Cyprus 92

(NHMW); Kolossi, N . of the castle, 1 ♂ 4 ♀♀, Citations under stones, 11.IV.1986 (NHMW). None. NICOSIA : Kato Koutrafas - Astromeritis road, 120m, 1 ♂, herbs on pond shore, 18.V.2011 New records (CRB); Pedoulas, 1100m, 1 ♀, on wall of hotel, LIMASSOL : Mouttagiaka, 20m, 1 ♀, on wall of 4.V.2010 (CRB). Caravel hotel, 29.IV.2010 (CRB).

Distribution: Cosmopolitan. New to Cyprus. Distribution: Cosmopolitan. New to Cyprus.

Oecobius maculatus Simon, 1870 Oecobius putus O. Pickard - Cambridge, 1876

Identification: https://araneae.nmbe.ch/ . Identification Kritscher (1966): 290, figs 11 - 12. Wunderlich, 1995a: 594, figs 29 - 30. Citations Citations None. None. New records New records NICOSIA: Dali, 13, Agios Andronikos, 1 ♂, in FAMAGUSTA : Paralimni Lake, 65m, 2 ♀♀, house, V.2008 (CD M). herbs and litter around dried lake, 27.IV.2010 Distribution: Egypt, Sudan to Azerbaijan, India. (CRB); Salamis ruins, 30m, 1 ♀, stones in ruins, New to Cyprus. 17.III.2011 (CRB). LARNACA : Larnaca salt lake, 5m, 3 ♀♀, stones Oecobius teliger (O. Pickard - Cambridge, 1872) in salt marsh, 27.IV.2010 (CRB). LIMASSOL : Akrotiri Peninsula, 2 ♀♀, edge of Identification central salt pan, under stones and debris in Wunderlich, 1995a: 599, figs 54 - 57.

Salicornia , 27.IV.2007 (CAR - S); Armenochori, 1 Citations ♀, in garag e, 14.IV.1986 (NHMW); Erimi, 1 ♀, None. under stones in dry riverbed, 18.IV.1986 (NHMW); Kolossi castle, 2 ♂♂ 1 ♀, walls in the New records castle area, 11.IV.1986 (NHMW). FAMAGUSTA : Prastio S ., Agios Andro nikos, NICOSIA : Agios Sozomenos , near Dali, 2 ♀♀, 30m, 3 ♀♀, under stones in Eucalyptus forest, stones in disturbed phrygana, 30.IV.2007 (CAR - 27.V.2011 (CRB); Karpasia Peninsula, Cape S). Apostolos Andreas, 70m, 2 ♀♀, stones in PAPHO S: Evretou Reservoir, 1 ♀, under stone in grassland, 16.V.2011 (CRB). grass tussocks, 13.IV.2017 (CAR - S); Kato LIMASSOL : gorge below Caledonia Paphos, 2 ♂♂ 4 ♀♀, under stones near hotel, Waterfall, 1 ♀, under stone, 26.IV.2007 (CRS); 4.IV.1997 (CJB); near Souskiou, 3 ♀♀, under Pano Platr es, 1 ♀; on stone wall, 26.IV.2007 stones on rocky eroded grassland on hill, (CAR - S). 17.IV.2017 (CAR - S). NICOSIA: Agios Sozomenos , near Dali, 1 ♂ 1 ♀, under stones in dry grassland, 30.IV.2007, on Distribution: Med iterranean to Azerbaijan. New limestone ridge behind village, 2 ♀♀, under to Cyprus. stone, 30.IV.2007 (CRS); Agridi, near Dali, 1 ♀, in house, IV.2009, 1 ♂, on stones, river bank, Oecobius navus Blackwall, 1859 29.IV.2013 (CDM); Dali, 13, Agios Andronikos, 2 ♀♀, in garden, IV.2008 (CDM).

PAPHOS: Near Souskiou, 1 ♀, under stones in Identification Wunderlich , 1995c: 595, figs 31 - 35. dry grassland, 1.V.2013 (CAR - S), 1 ♀ (CDM).

Spiders of Cyprus 93

Distribution: Lebanon, Turkey, and the Greek Paphos, 1 ♀, under stones near hotel, 1.IV.1997 Island of Chios . New to Cyprus. (CJB).

Uroctea durandi (Latreille, 1809) Distribution: Turkey, Lebanon, Israel. New to Cyprus. Identification: https://araneae.nmbe.ch/ . Opopaea saaristoi Wunderlich, 2011 Citations Opopaea saaristoi Wunderlich, 2011: 199, figs 1 - None. 5.

New records Identification LARNACA : Agioi Vavatsinias, 700m, 1 juvenile, Wunderlich, 2011: 199, figs 1 - 5. stones in Pinus for est, 28.IV.2010 (CRB). LIMASSOL: Pano Platres, below Caledonia Citations North Cyprus (Wunderlich, 2011; type locality). Wa terfall, 1 juvenile, under stone in gorge,

19.IV.2017 (CAR - S); 10 km S. of Troodos, 1 ♀, New records under stone in Pinus brutia forest, 19.IV.2017 None. (CAR - S). NICOSIA : Dali, Idalion Archaeological Site, 1 Distribution: Only known from Cyprus. juvenile, under stones in phrygana, 30. IV .2013 (CDM); Prodromos S., Trooditissa picnic site, Opopaea santschii Brignoli, 1974 1450m, 1 ♀, stones in Pinus forest, 21.V.2011 Opopaea santschii ; Saaristo, 2007: 135, figs 79 - (CRB) ; Profitis Elias, Machairas Forest,1 85. juvenile, 2005 (CDM). PAPHOS: Kykkos Monastery, road to Vigilios Ident ification church, 950m, 1 subault ♀, 20.IV.2017, S. Huber Saaristo, 2007: 135, figs 79 - 85. leg. (CSH). Citations

LIMASSOL : Limassol (Saaristo, 2006). Distrib ution: Mediterranean. New to Cyprus. New records None. Family Oonopidae Distribution: Tunisia, Cyprus, Egypt, Israel. Opopaea punctata (O. Pickard - Cambridge, 1872) Orchestina sp. Identification Identification Saaristo & Marusik, 2008: 29, figs 64 - 72, 155 - Citations 165, 196, 202, 227. None.

Citations New records None. PA PHOS: Akamas Peninsula, Agios Minas church; 1 ♂ 1 ♀, 2.V.2007 (CAR - S). New records FAMAGUSTA : Agio s Sergios , 30m, 1 ♂1 ♀, in Comments : These specimens could not be pitfalls in Pinus plantation, 15 . - 27.V.2011 identified . (CRB). NICOSIA : Prodromos S., Trooditissa picnic site, Silhouettella tomer Saaristo, 2007

1450m, 1 ♂, stones in Pinus forest, 21.V.2011 Identification (CRB). Saaristo, 2007: 137, figs 99 - 105. PAPHOS: near Avakas Gorge, 1 ♀, under stones in field with tall grass, 14.IV.2017 (CAR - S); Kato Citations None.

Spiders of Cyprus 94

New reco rds (MMUE); Platres, 1 ♀, on walls, 22.IV.1986 FAMAGUSTA :Rizokarpaso W ., Panagia Eleousa (NHMW). Monastery, 30m, 1 ♂, stones in Pinus forest, NICOSIA : Agios Sozomenos , near Dali, 1 ♀, 18.III.2011 (CRB); Salamis S., Silver Beach, 5m, under stone in dry grassland, 29.IV. 2013 (CAR - S); Agios Tychonas, 1 ♂, under stones near house 1 ♂, in pitfalls in salt marsh, 15 - 27.V.2011 walls, 19.IV.1986 (NHMW); Agridi, near Dali, 2 (CRB). ♂♂, in field, IV.2007; same site, 2 ♂♂, V.2008, 2 KERYNEIA: Pentadactylos Range, Ftericha S ., ♂♂, in garden, V.2011 (all CDM); Lefka NW, 800m, 1 ♀, st ones in grassland, 17.V.2011 (CRB). 120m, 6 ♀♀, beaten from bushes in fruit plantation, 18.V. 2011 (CJVK, CRB); Machairas Distribution: Israel, and new to Cyprus. forest near Kionia, 1300m, 1 ♂ 1 ♀, stony phrygana, 5.V.2010 (CJVK); Pyrgos, 1 ♂, in dry grass steppe, 15.IV.1986 (NHMW). Family Oxyopidae PAPHOS : Agios Merkourios, 250m, 1 ♂, sweeping grassland, 27.IV.2007 (CJVK); Oxyopes globifer Simon, 1876 Asprokremmos Dam, N. Koukli a, 150m, 1 ♀, herbs and stones along lake, 23.V.2011 (CRB); Identification Avakas gorge and surroundings, 1 ♀, 22.III.2018 Levy, 1999: 41, figs 10A - E, 11A - D, 12A - C. (CKHK); Drouseia S., Panagia Chrysospiliotissa, 580m, 1 ♂, night catch in small cave, 22.IV.2017, Citations S. Huber leg. (CSH), Evretou Reservoir, 350m, 1 None. ♀, shrubs and grassland around lake, 1.V.2010 New records (CJVK); Kelefos bridge, dam on Diarizos River, LIMASSOL : Avd imou, 1 ♀, on coast, 550m, 3 ♂♂ 1 ♀, beaten from shrubs, 21.V.2011 14.VII.1992, coll. P. Selden (BMNH). (CJVK, CRB); Gialia, 1 ♂, under stones in pine wood, 25.III.1998, coll. J. Murphy (MMUE), 2 Distribution: Mediterranean to central Asia. New ♂♂ 1 ♀, sweeping Cystus , 28 .IV.2007 (CJVK); to Cyprus. near Kritou Terra, 1 ♀, swept from grass in dry fallow, 14.IV.2017 (CAR - S); Loukrounou E., Oxyopes heterophthalmus (Latreille, 1804) 140m, 1 ♀, sweeping herbs bordering cultivated Oxyopes heterophthalmus ; Logunov, 2013: 5 fields, 1.V.2007 (CJVK); Mavrokolympos Reservoir, 1 ♂, 13.III.2018 (CKHK); Pelathousa Identification : https://araneae.nmbe.ch/ . SE., 245m, 1 ♂ 4 ♀♀, sweeping grassland, 30.IV.2007 (CJVK); Peristerona, Atichoullis Citations gorge, 425m, 1 ♀, beating in maquis, 2.V.2007 PAPHOS : Avakas Gorge (Logunov, 2013); (CJVK); Pomos S., 530m, 2 ♂♂, beating Cystus , Xeropotamos River (Logunov, 2013). 28.IV.2007 (CJVK); Souskiou, 1 ♂, 16.III.2013 (CDM). New records KERYNEIA: Pentadactylos Range, Saint Hilarion Distribution: Palaearctic. Castle W., 600m, 1 ♀, beaten from shrubs in Pinus forest, 17.V.2011 (CJVK). Oxy opes lineatus Latreille, 1806 LARNACA: Agia Anna, 1 ♂, in gulley with Identification : https://araneae.nmbe.ch/ . pines, 26.IV.1982, coll. A. Stubbs (MMUE). LIMASSOL: Agios Tychonas, 2 ♂♂ 1 ♀, under Citations stones near house walls, 19.IV.1986 (NHMW); None. Akrotiri, Fasouri reedbeds, 1 subadult ♀, swept from gra ss and reeds, 17.IV.2017 (CAR - S); New records Alassa SE, near river Limnatis, 1 ♂ 1 ♀, FAMAGUSTA : Agios Sergios , 30m, 1 ♂ 3 ♀♀, sweeping lush grassland, 5.V.2007 (CAR - S) ; beaten in Pinus plantation, 15.V.2011, CRB); Amathus, 1 ♀, on house wall, 9.IV.1986 Salamis ruins, 50m, 1 ♂, beaten from low trees, (NHMW); Kannavia, 780m, 1 ♂, sweeping 15.V.2011 (CRB). grassland, 4.V.2007 (CJVK); Pano Platres E., LARNACA : Kornos Picnic site, 350m, 3 ♂♂, Mesa Potamos Wat erfall s, 950m, 1 ♂, beaten in under stones in Pinus forest, 28.V.2011 (CJVK). Pinus forest, 4.V.2010 (CJVK); Pissouri beach, 1 ♀, under stone, 24.III.1998, coll. A. Stubbs

Spiders of Cyprus 95

LIMASSOL : Alassa SE, 1 ♂ 1 ♀, swept from Oxyopes nigripalpis Kulczyński, 1891 grass at edge of river Limnatis, 5.V.2007 (CAR - S); Pano Platres, near top of road to Millomeris Identification : https://araneae.nmbe.ch/ . Waterfalls, 1 juvenile, swept from grass and herbs in abandoned vineyard, 28.IV.2007 (CAR - S); Citations Mount Troodos, 1 ♀ (BMNH). None. NICOSIA: Agridi, near Dali, 1 ♂, in garden, V.2011; 1 ♀, IV.2012 (CDM); Cedar Valley, 1 New records juvenile, swep t from herbs, 29.IV.2007 (CAR - S); Without precise locality, 1 ♂, VI.1999. coll. J. Kampos NE, Xeros River, 550m, 19 ♂♂ 4 ♀♀, Murphy (MMUE). under stones in riverbed and beaten from shrubs, 20.V.2011 (CJVK, CRB); Kapedes S., 600m, 3 Distribution: Mediterranean. New to Cyprus. ♂♂, herbs along rivulet, 5.V.2010 (CJVK, CRB); Kato Koutrafas - Astromeritis road, 120m, 2 ♂♂, Oxyopes ramosus (Martini & Goeze, 17 78) herbs on pond shore, beaten from shrubs in dry riverbed, 18.V.2011 (CJVK); Lefka NW, 120m, Identification : https://araneae.nmbe.ch/ . 12 ♂♂ 1 ♀, beaten from shrubs in fruit plantation, 18.V.2011 (CRB); Machairas forest near Kionia, Citations 1300m, 2 ♂♂, stony phrygana, 5.V.2010 (CRB); None. Limnitis River, Kato Pyrgos SE, 200m, 1 ♂, stones and litter in Pinus forest, 2.V.2010 New records (CJVK); Marathassa Dam, 550m, 1 ♂, beaten NICOSIA: Agridi, near Dali, 1 ♂, in house, from shrubs around lake, 20.V.2011 (CRB); V.2011 (CDM). Oikos N., Xerargaka picnic site, 350m, 1 ♂ 1 ♀, beaten from shrubs and stones in Pinus forest, Distribution: Palaearctic. New to Cyprus. 20.V.2 011 (CJVK). PAPHOS : Akamas Peninsula, Neo Chorio, 1 Oxyopes sobrinus Latreille, 1806 subadult ♂, swept from phrygrana, 2.V.2007 (CAR - S); Akamas Peninsula, Agios Minas, 1 Identification subadult ♂, on tall grasses, 2.V.2007 (CAR - S); Levy, 1999: 37, figs 7 - 8. Asprokremmos Dam, N. Kouklia, 150m, 3 ♂♂ 4 ♀♀, herbs and stones along lake, 29.IV.2010 Citations (CJVK); Evretou Reservoir, 350m, 2 ♂♂ 4 ♀♀, None. on shrubs around lake, 1.V.2011 (CJVK, CRB); Mavrokolympos Reservoir, NE Potima, 150m, 1 New records ♂ 1 ♀, on shrubs, 22.V.2011 (CRB); Pela NICOSIA : Agridi, near Dali, 1 ♂, in yard, V.2009 thousa SE., 245m, 2 ♂♂, sweeping grassland, (CDM); Agios Sozomenos , near Dali , 1 ♂, under 30.IV.2007 (CJVK); P olis, 15m, 1 ♂, herbs and stones in disturbed phrygana, 30.IV.2007 (CAR - stones around building, 27.IV.2007 (CJVK). S).

Distribution: Palaearctic. New to Cyprus. Distribution: Libya, Israel. New for Cyprus. Oxyopes mediterraneus Levy, 1999 Peucetia arabica Simon, 1882 Identification Levy, 1999 : 43, fi gs 14 - 15. Identification Levy, 1999: 51, figs 24A - B, 25A - C. Citations None. Citations None New records NICOSIA : Elia - Angolemi road, 100m, 1 ♀, stones in grassland, 18.V.2011 (CJVK). New records KERYNEIA: Keryneia , 1 ♀, E. Green leg. Distribution: Mediterranean. New to Cyprus. (BMNH).

Spiders of Cyprus 96

LARNACA : Agioi Vavatsinias, 700m, 1 ♂, Platres, 1 ♀, under stones, in P. nigra forest, beaten in Pinus forest, 28.IV.2010 (CJVK). 1.V.2007 (CRS); 10 km S. Troodos, 1 ♀, under LIMASSOL : Agros SW, 950m, 1 ♂, beating in stones in Pinus brutia forest, 19.IV.2017 (CAR - S). Pinus forest, 5.V.2010 (CJVK). NICOSIA : Cedar Valley Forest, 1 ♀, under stones NICOSIA : Kampos NE, Xeros River, 550m, 1 ♀, beaten from shrubs, 20.V.2011 (CRB) ; by track, 29.IV.2007 (CRS); Dali, Idalion Lefka NW, 120m, 1 ♂ 3 ♀♀, beaten from shrubs Archaeological Site, 1 ♀, under stones in in fruit plantation, 18.V.2011 (CJVK); Marathassa phrygana , 30.IV.2013 (CDM); Kakopetria S., Dam, 550m, 1 ♀, b eaten from shrubs around lake, Platania forest station, 1100m, 1 ♀, under stones 20.V.2011 (CJVK); Oikos N., Xerargaka picnic in dense Pinus forest, 4.V.2010 (CRB); below site, 350m, 1 ♂, beaten from shrubs in P inus Machairas Monastery, 1 ♀, under stones beside forest, 20.V.2011 (CJVK). track in phrygana, 4.V.2013 (CAR - S); Nikitari S., PAPHOS : Agios Mamas ruins, Gialia E., 215m, 1 ♀, beating Cystus , 28.IV.2007 (CJVK); Kinousa Panagia Asinou, 1 ♀, under sto nes in disturbed SE, 400m, 1 ♂, under stones in Pinus forest, phrygana, 9.V.2007 (CAR - S); Pyrgos, 1 ♂ 1 ♀, 1.V.2010 (CRB); Limnitis River, Kato Pyrgos SE, steppe with individual stones and bushes and dry 200m, 5 ♂♂ 2 ♀♀, stones and litter in Pinus grass, 15.IV.1986 (NHMW). forest, 2.V.2010 (CJVK); Pelathousa SE., 245m, 1 PAPHOS: Akamas Peninsula, Cape Drepanum, 1 ♂, sweeping grassland, 30.IV.2007 (CJVK). ♀, 31.III.1997 (CJB); Akamas Peninsula, Cape Geronnisos S., 1 ♂, under st ones in dry phrygana, Distribution: Spain, Greek Islands, Africa, Middle 3.V.2007 (CAR - S), 1 ♀ (CRS); Akamas East. New to Cyprus. Peninsula, S. Neo Chorio, 270m, 1 ♀, stones in karst landscape, 22.IV.2017, S. Huber leg. (CSH); Family Palpimanidae Kinousa, 1 ♂, under stone in pine wood, 28.III.1998, coll. J. Murphy (MMUE); Kato Palpimanus cyprius Kulczyński, 1909 Akourdaleia, 1 ♂, i n garden, VIII.2005 (CDM); Palpimanus cypri us Kulczyński, 1909: 671; Paphos NW, 1 ♂, under stones at the edge of a Platnick, 1981: 172. large sand pit, 24.IV.1986 (NHMW); Smigies, 1 ♀, under stone in hillside scrub, 26.III.1998, coll. Identification J. Murphy (MMUE). Platnick, 1981: 172, figs 7, 16. Distribution: Cyprus, Syria, Israel. Citations Without precise locality (Kulczyński, 1909; Platnick, 1981). Family Phi lodromidae

New records Philodromus aureolus (Clerck, 1757) FAMAGUSTA : Komi Kebir , 150m, 1 ♀, pitfalls in Pinus forest, III. - V.2011 (CRB). Identification : https://araneae.nmbe.ch/ KERYNEIA: near Agios Georgios, 2 ♀♀, under stones, 27.II.2011, H. Henderickx leg. (CJB); Citations Keryneia , 1 ♀, E. Green leg., 28.IV. 1936 None. (BMNH); Pentadactylos Range, Saint Hilarion Castle W, 600m, 1 ♀, under stones in Pinus forest New records 17.V. 2011 (CRB); Pentadactylos Range, Ftericha NICOSIA : Kato Koutrafas - Astromeritis road, S ., 800m, 1 juvenile, under stones in grassland, 120m, 1 ♂, beaten from shru bs in dry riverbed, 17.V.2011 (CRB). 18.V.2011 (CRB). LIMASSOL : Agros SW, 950m, 1 ♀, under stones in Pinus forest, 5.V.2010 (CRB); Alassa SE, river Distribution: Europe. New to Cyprus. Limnatis, 1 ♀, under stones at base of limestone roc k face of gorge, 5.V.2007 (CAR - S); Panagia Philodromus cespitum (Walckenaer, 1802) Glossa Monastery near Kellaki, 1 ♂, stones in cultivated land, 12.IV.1986 (NHMW); Pedoulas, Identification : https://araneae.nmbe.ch/ . 960m, 1 ♀, road embankment, 18.IV.2017, S. Huber leg. (CSH); 5 km S. Troodos, road to Pano

Spiders of Cyprus 97

Citations NICOSIA : Agr idi, near Dali, 1 ♂, in garden, None. V.2009 (CDM); Lefka NW, 120m, 1 ♀, beaten from shrubs in fruit plantation, 18.V.2011 (CRB); New records below Machairas Monastery, 1 ♂, under stones NICOSIA : Marathassa Dam, 550m, 1 ♀, beaten beside track in phrygana, 4.V.2013 (CAR - S); from shrubs around lake, 20.V.2011 (CRB). Marathassa Dam, 550m, 1 ♀, beaten from shru bs LIMASSOL: Akrotiri Peninsula E., 5m, 2 ♀♀, around lake, 20.V.2011 (CRB); Masari S., Merika under stones and in litter in salt marsh, River, 120m, 1 ♂ 1 ♀, beaten from shrubs in 29.IV.2010 (CRB). wasteland, 24.V.2011 (CRB); Oikos N., Xerargaka picnic site, 350m, 1 ♂ 4 ♀♀, beaten D istribution: Europe. New to Cyprus. from shrubs in Pinus forest, 20.V.2011 (CRB). PAPHOS : Vretsia W., near Kele fos Bridge, Philodromus longipalp is Simon, 1870 650m, 1 ♀, beaten from shrubs, 19.V.2011 (CRB). Identification : https://araneae.nmbe.ch/ Distribution: Eastern Mediterranean. New to Citations Cyprus. None. Philodromus margaritatus (Clerck, 1757) New records NICOSIA : Agridi, near Dali, 1 ♀, in hedge in Identification : https://araneae.nmbe.ch/ . garden, 20.V.2011 , 1 ♂ in garden, V.2011, 2 ♂♂, in garden, V.2 012 (all CDM). Citations None. PAPHOS : Evretou Reservoir, 350m, 1 ♀, beaten from shrubs around lake, 1.V.2010 (CRB). New records NICOSIA: Pano Amiantos, 1 ♂, pitfalls in Pinus Distribution: Europe. New to Cyprus. brutia forest, 6.IV. - 8.VI.2014 (CMH). LIMASSOL: Troodos Square, 1 ♂, Pinus nigra Philodromus lunatus Muster & Thaler, 2004 forest, 11.V.2016 (CMH). 1 Identification Distribution: Palaearctic. New to Cyprus. Muster & Thaler, 2004: 319, figs 4, 6, 19, 26, 27. Philodr omus marmoratus Kulczyński, 1891 Citations None. Identification Kubcová, 2004: 292, figs 11a - d. New records FAMAGUSTA : Agios Sergios , 30m, 1 ♀, beaten Citations in Pinus plantation, 15.V.2011 CRB); Kantara E., None. 610m, 1 ♂ 5 ♀♀, beaten from shrubs in open Pinus forest, 25.V.2011 (CRB); Prastio S ., Agios New records Andronikos, 30m, 1 ♀ , swept from herbs in FAMAGUSTA : Salamis ruins, 50m, 2 ♂♂, beaten from low trees, 15.V.2011 (CRB). Eucalyptus forest, 27.V.2011 (CRB). LARNACA : Achna Reservoir, 50m, 1 ♂, beating KERYNEIA: Panagia Apsinthiotissa Monastery bushes bordering lake, 15.V.2011 (CRB). ruins, 350m, 1 ♂ 1 ♀, beaten in clearing in Pinus forest, 24.V.2011 (CRB); Pentadactylos Range, Distribution: Central Europe, Bulgaria, Ukraine. Saint Hilarion Castle, 600m, 5 ♂♂ 1 ♀, beaten New to Cyprus. from shrubs , 17.V.2011 (CRB). LIMASSOL: Armenochori, 1 ♀, city area, Philodromus poecilus (Thorell, 1872) 14.IV.1986 (NHMW); Episkopi game reserve, 1 ♂, C. & P. R. Deeleman leg., 31.I.1988, moulted Identification : https://araneae.nmbe.ch/ . to adult in VI.1988 (CRB); Parekklisia, 1 ♀, under stones in riverbed, 13.IV.1986 (NHMW). Citations None.

Spiders of Cyprus 98

NW, 120m, 2 ♀♀; beaten f rom shrubs in fruit New records plantation, 18.V.2011 (CRB); Kato Koutrafas - NICOSIA: Xerargaka S, Orkontas, 1 ♀, riverside Astromeritis road, 120m, 1 ♀, swept from herbs vegetation, 26.III.2016, M. Hadjiconstantis leg. on pond shore, 18.V.2011 (CRB); Oikos N., (CRB). Xerargaka picnic site, 350m, 1 ♀, beaten from Distribution: Palaearctic. New to Cyprus. shrubs in Pinus forest, 20.V.2011 (CRB). PAPHOS : Asprok remmos Dam, N. Kouklia, Philodromus rufus Walckenaer, 1826 150m, 2 ♀♀, swept from herbs along lake, 23.V.2011 (CRB); Evretou Reservoir, 350m, 2 Identification : h ttps://araneae.nmbe.ch/ . ♀♀, on shrubs around lake, 1.V.2010 (CRB);

Kelefos bridge, dam on Diarizos River, 550m, 1 Citations None. ♂ 1 ♀, beaten from shrubs, 21.V.2011 (CRB); Kidasi N., al ong Diarizos River, 300m, 2 ♀♀, New records beaten from shrubs, 21.V.2011 (CRB); Vretsia NICOSIA: Agridi, near Dali, 1 ♂, in yard, W., near Kelefos bridge, 650m, 1 ♀, beaten from IV.2009 (CDM). shrubs, 19.V.2011 (CRB). PAPHOS: Stavros tis Psokas, 850m, under litter, 21.IV.2017, S. Huber leg. (CSH). Distribution: Greece, Cyprus, Israel, Azerbaijan.

Distribution: Holarctic. New to Cyprus . Pulchellodromus pulchellus (Lucas, 1846) Phi lodromus pulchellus ; Logunov, 2013: 5. Pulchellodromus medius (O. Pickard - Cambridge, 1872) Identification Philodromus medius ; Kulczyński, 1908: 75; Muster et al ., 2007: 60, figs 5, 7, 21, 32, 43, 54, Muster, Bosmans & Thaler, 2007: 57. 63 - 64.

Identification Citations Muster, Bosmans & Thaler, 2007: 57, figs 9, 23, PAPHOS : Avakas Gorge (Logunov, 2013). 34, 45, 56, 67 - 68. New records NICOSIA: Athalassa National Forest Park, Citations Nicosia, 1 ♀, under ston es in pine plantation, Without preci se localiy (Kulczyński, 1908; 2.V.2013 (CDM); Dali, Idalion Archaeological Muster et al., 2007). Site, 1 ♀, under stones in phrygana, 30.IV.2013 (CDM); Nikitari S., Panagia Asinou, 1 ♂, under New records stones in disturbed phrygana, 9.V.2007 (CAR - S). FAMAGUSTA : Cape Greco, Ayia Napa, 35m, 1 LIMASSOL : Akrotiri Peninsula N., 5m, 1 ♀, ♀, stones in phrygana, 27.IV.2010 (CRB); l itter and beating in Eucalyptus forest, 29.IV.2010 Salamis ruins, 50m, 2 ♀♀, beaten from low trees, (CRB); Akrotiri Peninsula, near British airbase, 1 15.V.2011, CRB); ♂, swept from grass in pine wood. 4.V.2007 KERYNEIA: Pentadactylos Range, Sai nt Hilarion (CAR - S); Akrotiri Peninsula, Fassouri reed beds, Castle, 600m, 1 ♂, beaten from shrubs, 17.V.2011 3 ♂♂, swept from grass, 17.IV.2017 (CAR - S); Alassa SE, near ri ver Limnatis, 1 ♂, stones in dry (CRB). grassland, 5.V.2007 (CAR - S); Pissouri beach, 1 LARNACA : Dipotamos Reservoir, 150m, 1 ♀, ♀, under stone, 24.III.1998, coll. A. Stubbs herbs and stones bordering lake, 28.IV.2010 (MMUE). (CRB); Larnaca salt lake, 5m, 1 ♀, on herbs in PAPHOS: Akamas Peninsula, Avakas Gorge, 1 salt marsh, 27.IV.2010 (CRB). ♂, under stone, 27.III.1998, coll. J. Murphy LIMASSOL: Agios Tych o na s, 1 ♂, under stones (MMUE); Akamas Peninsula, Agios Mi nas, 1 ♂ 2 near house walls (NHMW) . ♀♀, under stones in phrygana , 2.V.2007 (CAR - S); Akamas Peninsula, Neo Chorio, 1 ♂ 1 ♀, NICOSIA: Agridi, near Dali, 1 ♂, in garden, swept from phrygana, 2.V.2007 (CAR - S); Gialia, V.2011 (CDM); Amathus, 1 ♀, under stones in 1 ♀, in pine wood, 25.III.1998, coll. J. Murphy dry valley, 6.IV.1986 (NHMW); Amat hus, 1 ♂, (MMUE); near Kidasi village, Diarizos Valley, 1 front yard of hotel, 8.IV.1986 (NHMW); Lefka ♂, in flood deb ris by stream, 1.V.2013 (CAR - S); Spiders of Cyprus 99

Kinousa, 2 ♂♂, in pine wood, 19.III.1998, coll. J. 25.V.2011 (CRB); Kantara E., 610m, 1 ♂ 1 ♀, Murphy (MMUE). stones in open Pinus forest, 25.V.2011 (CRB); Karpasia Peninsula, Apostolos Andreas Distribution: Mediterranean. Monastery, 50m, 1 ♂, stones in wasteland,

16.V.2011 (CRB); Salamis ruins, 50m, 1 ♀, under Thanatus fabricii (Audouin, 1866) stones, 15.V.2011 (CRB); Salamis S., Silver Identification Beach, 5m, 1 ♀, in pitfalls in salt marsh, 15. - Crespo et al., 2010: 97, figs 2A - C. 27.V.2011 (CRB). LARNACA : Achna Reservoir, 50m, 1 ♂ 1 Citations subadult ♀, stones and litt er bordering lake, None. 16.V.2011 (CRB), 3 ♂♂, pitfalls at same site, 15. - 27.V.2011 (CRB); Cape Kiti, 1 ♀, under stones New records below coastal cliffs, 25.IV.1982, coll. A. Stubbs NICOSIA: Agios Sozomenos , near Dali, 1 ♂ 1 ♀, (MMUE); Larnaca salt lake, 5m, 2 ♀♀, stones under stones in dry grassland, 30.IV.2007 (CAR - and litter in salt marsh, 27.IV.2010 (CRB); S), 1 ♂ (CRS); Agridi, near Dali, 1 ♂, in garden, S tavrovouni Monastery, 350m, 1 ♂, on bark of IV.2011 (CDM); Dali, 1 ♀, under stones and in Pinus trees, 28.IV.2010 (CRB). LIMASSOL: Agios Tychonas, 1 ♂, under stones grass in riverbed, 12.V.2007 (CAR - S). near house walls, 19.IV.1986 (NHMW); Akrotiri PAPHOS: Ak amas Peninsula, between Cape Peninsula E., 5m, 1 ♀, under stones and in litter in Drepano and Lara Bay, 1 ♀, 2.IV.1997 (CJB). salt marsh, 29.IV.2010 (CRB); Akroti ri Peninsula, edge of central salt pan, 1 ♀, under stones and Distribution: Canary Islands to Central Asia. debris in Salicornia , 27.IV.2007 (CAR - S); New to Cyprus. Akrotiri Peninsula, Ladies Mile Beach, 1 ♂, in sand dunes, 27.IV.2007 (CAR - S); Alassa SE, near Thanatus imbecillus L. Koch, 1878 river Limnatis, 2 ♂♂, under stones in dry grassland, 5.V. 2007 (CAR - S), 1 ♂ 3 ♀♀ (CRS); Identification Kourion archaeological site near Episkopi, 1 ♀, Logunov & Huseynov, 2008: 124, figs 19 - 22. under stones, 9.V.2007 (CAR - S). NICOSIA : Agridi, near Dali, 3 ♂♂, in garden, Citati ons VI.2010, in lawn, 4 ♂♂ 5 ♀♀, 1.VII.2010, in None. lawn, 6 ♂♂ 10 ♀♀, 20.VI.2011, 2 ♂♂, in garden, V.2011, 1 ♂, in house, VI.2012 (all CDM); Agios New records Sozomenos , near Dali, 1 ♂ 1 ♀, under stones in LIMASSOL: Pissouri, 1 ♀, in grass, 24.III.1998, dry grssland, 30.IV.2007 (CAR - S) , 1 ♂, coll. J. Murphy (MMUE). 1.VI.2012 (CDM); Armenochori, 1 ♂, city area, 14.IV.1986 (NHMW); Dali, 13, Agios Distribution: North Macedonia to central Asia. Andronikos, 1 ♀, in garden, V.2008 (CDM); Dali, New to Cyprus. 1 ♀, under stone near river, 12.V.2007; Dali, Idalion archaeological site, 1 ♀, 1.V.2013 (CAR - Thanatus vulgaris Simon, 1870 S); Elia - Angolemi road, 100m, 5 ♀♀, under Thanatus vulgaris ; Logunov, 2013: 5. rubbish in dry riverbed, 18.V.2011 (CRB); Erimi, 1 ♂, under stones along dry riverbed, 21.IV.1986 Identif ication : https://araneae.nmbe.ch/ . (NHMW); Kato Koutrafas - Astromeritis road, 120m, 1 ♀, herbs on pond shore, 18.V.2011 Citations (CRB); Kourion, Apollo Hylates Temple, 1 ♀, LIMASSOL: Episkopi, Kourion Beach (Logunov, under stones in the area of the antique site, 2013). 23.IV.1986 (NHMW); Machairas forest near PAPHOS : Amargeti (Logunov, 2013); Kionia, 1300m, 1 ♂, stony phrygana, 5.V.2010 Xeropotamos River (Logunov, 2013). (CRB); Masari S., Merika River, 120m, 1 ♀, stones in wasteland, 24.V.2011 (CRB); New records Pallouriotissa, Nicosia, 1 ♂, in building, IV.2006 FAMAGUSTA : Cape G reco, 1 ♀, under stones in (CDM). phrygana, 3.V.2013 (CAR - S); Flamoudi W., 25m, PAPHOS : Akamas Peninsula, track along the sea, 1 ♂ 1 ♀, stones and phrygana near the beach, 10m, 1 ♀, stones in Juniperus phrygana,

Spiders of Cyprus 100

30.IV.2010 (CRB); Akamas Peninsula, Lara Bay, Citations 1 ♀, C. Saure leg., 18.VI.2013 (CKHK); FAMAGUSTA: Lisi (Aharon et al. 2017); Asprokremmos Dam, N. Kouklia, 150m, 1 ♀, Panagia Lysis church (Aharon et al . 2017). herbs and stones along lake, 29.IV.2010 (CRB); NICOSIA: Nicosia (A haron et al. 2017); near Evretou Reservoir, 350m, 1 ♀, under stones in Agios Sozomenos (Aharon et al. 2017); Geri grassland around lake, 1.V.2010 (CRB); Kato (Aharon et al., 2017). Paphos, 1 ♀, under stones near hotel, 5.IV.1997 (CJB); Kato Paphos archaeological site, 1 ♀, New records under stones in grass, 10.V.2007 (CAR - S); near NICOSIA: Dali, 13, Agios Andronikos, 1 ♂, Souskiou, 1 ♀, under stones in dry grassland, in house, XII.2009 (CDM). 1.V.2013 (CAR - S). Distribution: Israel, Greece, Turkey, Cyprus. Distribution: Holarctic. Holocnemus pluchei (Scopoli, 1763) Tibellus macellus Simon, 1875 Identification : https://araneae.nmbe.ch/ . Identification Ef imik, 1999: 113, figs 3, 8, 15 - 16, 25, 33, 41, 49, Citations 66 - 70. None.

Citations New records None FAMAGUSTA : Cape Greco, Ayia Napa, 35m, 1 ♀, under stones in phrygana, 27.IV.2010 (CRB), 1 New records ♀, under stones in phrygana, 3.V.2013 (CAR - S); LIMASSOL: Akrotiri Peninsula, Fasouri, 1 ♂ 1 Kantara E., 650m, 3 ♂♂, in toilets, 19.III.2011 ♀, swept from grass and herbs, 27.IV.2007 (CAR - (CRB); Karpasia Peninsula, Aposto los Andreas S). Monastery, 50m, 1 ♂ 1 ♀, under stones in wasteland, 16.V.2011 (CRB); Paralimni lake, Distribution: Europe to Central Asia. New to 65m, 1 ♀, litter along dry lake, 21.XI.2010 Cyprus. (CRB); Rizokarpaso W ., Panagia Eleousa Monastery, 30m, 1 subadult ♀, under stones in Tibellus oblongus (Walckenaer, 1802) Pinus forest, 18.III.2011 (CRB ); Salamis ruins, 30m, 1 subadult ♀, under stones in ruins, Identification : https://araneae.nmbe.ch/ . 17.III.2011 (CRB); Tula (Engomi) E., Silver beach, 10m, 1 ♀, under stones in salt marsh, Citations 15.V.201 (CRB). None KERYNEIA: Karavas , 660m, 1 ♀, garden of hotel, 22.XI.2010 (CRB); Pentadactylos Range, New records Fte richa S ., 800m, 1 juvenile, under stones in LIMASSOL: Akrotiri Peninsula, 1 ♀, margin of grassland, 17.V.2011 (CRB). central salt lake, 7.V.1972, coll. A. S tubbs LARNACA : Larnaca salt lake, 5m, 3 ♀q, under (MMUE); Akrotiri Peninsula, near British air stones in salt marsh, 27.IV.2010 (CJVK, CRB); base, 1 ♀, swept from marshy area, 4.V.2007 Larnaca SW., Hala Sultan Tekke mosque, 5m, 1 (CAR - S), same site 1 ♀ (CRS). ♂, salt marsh, 16.III.2011 (CRB); Stavrovouni M onastery, 350m, 2 ♀♀, on bark of Pinus trees, Distribution: Holarctic. New to Cyprus. 28.IV.2010 (CRB). LIMASSOL: Akrotiri Peninsula, Fasouri reedbeds, 1 juvenile, swept from grass and reeds, Family Pholcidae 17.IV.2017 (CAR - S); Akrotiri Peninsula, central salt pan, 2 ♀♀, under stones, 17.IV.2017 (CAR - Artema nephilit Aharon, Huber & Gavish - S), 1 ♀ 2 juveniles, 2.IV.1997 (CJB); Alassa SE, Regev, 2017 1 ♂, near river Limnatis, under stones at base of Identifi cation limestone rock face of gorge, 5.V.2007 (CAR - S). Artema nephilit Aharon, Huber & Gavish - Regev, NICOSIA : Agios Sozomenos, near Dali , 1 ♂, in 2017 : 15, figs 4 - 6, 9 - 14, 35 - 75, 203, 210 disturbed phrygana, 30.IV.2007 (CRS); Apliki S., 800m, 1 juv., under s tone, 24.IV.2017, S. Huber

Spiders of Cyprus 101

leg., det. B. Huber (CSH); Dali, 13, Agios FAMAGUSTA: Karpasia Peninsula, 1 ♀, in cave Andronikos, 1 ♀, in garden, IV.2008 (CDM); near the ruins of Aphendrika church, 10. VII. Elia - Angolemi road, 100m, under stones in 2016 (CMH); Agios Andronikos, 1 ♂ 2 ♀♀, in grassland, rubbish in dry riverbed, 18.V.2011 the cave of Agia Fotou, 19. VI. 2016 (CMH); (CRB); Limnitis River, Kato Pyrgos SE, 200m, 2 Agia Trias, 1 ♀, cave of Ayios Therisso. ♀♀, und er stones and litter in Pinus forest, PAPHOS: Tsada, 1 ♂ 1 ♀, in fruit tre e orchard, 2.V.2010 (CRB) ; Nikitari S., Panagia Asinou, 1 ♂ 23.VII.2017 (CMH). 1 ♀, on stone walls, 9.V.2007 (CAR - S); Masari S., Merika River, 120m, 1 ♂, under stones in Distribution: Cosmopolitan. New to Cyprus. wasteland, 24.V.2011 (CRB); Pedoulas, 960m, 1 ♂, 1 ♀, road embankment, 18.IV.2 017, S. Huber Spermophora senoculata (Dugès, 1836) leg., det. B. Huber (CSH); Pissouri beach, 1 ♂ 1 ♀, under stones, 27.IV.1982, coll. A. Stubbs Identification : https://araneae.nmbe.ch/ . (MMUE); PAPHOS : Akamas Peninsula, Avgas E stuary, 2 Citations ♀♀, under stones, 31.III.1997 (CJB); Drouseia, None. 539m, 1 ♂ 1 ♀, in p hrygana, 23.III.2018 (CKHK) ; Kinousa SE, 400m, 2 ♀♀, under stones New records and litter in Pinus forest, 1.V.2010 (CRB). NICOSIA: Agridi, near Dali, 1 ♂, in yard, V.2009 (CDM). Distribution PAPHOS : Akamas Peninsula, Cape Drepanos, 1 Mediterranean, expanding to temperate Europe. ♀, in plastic bottle, 4.VIII.1998 (CJB); Kinousa New to Cyprus. SE, 400m, 2 ♂♂ 1 ♀, under stones and litter in Pinus forest, 1.V.2010 (CRB). Hoplopholcus sp. Distribution : Holarctic. New to C yprus. Records LIMASSOL: near Kidasi, 1 ♀, in flood litter by Family Phrurolithidae stream, 1.V.2103 (CAR - S); Pano Platres E., Mesa Potamos Waerfall s, 950m, 1 ♀, litter in Pinus Phrurolithus sp. forest, 4.V.2012 (CRB); Troodos S, 1350m, 1 ♀ 1 The only Phrurolithus species captured on the subadult ♀, litter in mixed Pinus - Quercus forest, island appears to be a new species and will be 3.V.2010 (CRB). described in a separate paper. NICOSIA: Kampos, 500m, 1 ♂ 1 ♀, abandoned village, 20 .IV.2017, S. Huber leg. (CSH).

PAPHOS: Agios Georgios E., Palaiomylos, 1250m, 1 ♂ 2 ♀♀, stones in mixed Pinus and Family Phyxelididae Quercus forest, 4.V.2010 (CRB); Avakas Gorge, 1 ♀, XI.1998 (CJB); Kinousa SE, 400m, 1 ♂ 2 Phyxelida anatolica Griswold, 1990 ♀♀, under stones and litter in Pinus forest, Ph yxelida anatolica Griswold, 1990: 167; Thaler 1.V.2 010 (CRB); Oikos, 700m, 2 juveniles, small & Knoflach, 1998: 36; Özkütük et al. 2013: 74. creek, 23.IV.2017, S. Huber leg. Identification Comments Thaler & Knoflach, 1998: 36, figs 1 - 16, 18 - 23. The species is probably new and is currently studied by B. Huber. Citations LIMASSOL: Troodos Range, Mount Olympos Pholcus phalangioides (Fuesslin, 1775) (Thaler & Knoflach, 1998).

Identification : https://araneae.nmbe.ch/ . New r ecords Citations FAMAGUSTA: Agios Andronikos, 1 ♀, hand None. catch in cave of Agia Fotou church, 19.VI.2016 (CMH); Sygkrasi, 1 ♂, cave under church ruins, New records 19.VI.2016 (CMH).

Spiders of Cyprus 102

LIMASSOL : Mount Olympos (Chionistra), S. under stones in dry valley, 25.IV.1986, 2 ♀♀, slope, 1900m, 1 ♀, under stones in open Pinus 6.IV.1986 (NHMW); Armenochori, 1 ♀, on forest, 4.V.2010 (CRB). bushes, 14.IV.1986 (NHMW); Germasogeia, 1 ♀, NICOSIA: Agridi, near Dali, 1 ♂, in field, on plants along stream below the d am, 10.IV.1986 IV.2008 (CDM); Marathasa Dam, 275m, 1 ♂, pitfalls in riverside vegetation, 7.V. - 10.VII.2016 (NHMW). (CMH). NICOSIA : Lefka NW, 120m, 1 ♀, beaten from shrubs in fruit plantation, 18.V.2011 (CRB). Distribution: Greece (Crete), Cyprus, Turkey. PAPHOS : Baths of Aphrodite, 1 ♀, in tall grass in meadow, 15.IV.2017 (CAR - S); Avakas gorge, 50m, 1 ♀, litter in citrus yard at entrance of gor ge, Family Pisauridae 30.IV.2010 (CRB); Drouseia, 539m, 1 ♀, in p hrygana, 23.III.2018 (CKHK); Paphos NW, 2 Pisaura conso cia (O. Pickard - Cambridge, ♀♀, under stones at the edge of a large sand pit, 1872) Pisaura consocia ; Logunov, 2013: 5. 24.IV.1986 (NHMW).

Identification Distribution: Palaearctic. New to Cyprus. Levy, 1999: 58, figs 31 - 32. Pisaura novicia (L. Koch, 1878) Citations LIMASSOL: Episkopi, Kourion beach (Logunov, Identific ation 2013). Nadolny et al ., 2012: 262, figs 2, 4, 24 - 25, 28 - 30, 33 - 34, 350, fig. 911. New records LARNACA : Delikipos, near Kornos, 1 ♀, in grass Citations of open pine wood, 9.V.2013 (CAR - S); None.

Stavrovouni Monastery, 350m, 1 ♂, on wall in New records entrance hall of monastery, 28.IV.2010 (CRB). LARNACA : Larnaca, 1 ♀, without date, C. LIMASSOL: Limassol, Curium stadium, 1 ♀, Glasyznet leg. (BMNH). stones in rock strewn grassland, 2.IV.1997 (CJB). NICOSIA: Agridi, near Dali, 1 ♂, i n garden, Distribution: Mediterranean to Central Asia. IV.2012 (CDM). New to Cyprus. PAPHOS: Pegeia, Pikni Forest, 345m, 1 ♂, in former carob plantation, 23.III.2018 (CKHK). Family Salticidae Distribution: Turkey, Israel, Lebanon, Syria. New to Cyprus. Aelurillus aeruginosus (Simon, 1871)

Pisaura mirabilis (Clerck, 1757) Identification: https://www.araneae.nmbe.ch/

Identification : https://araneae.nmbe.ch/ . Citations None. Citations None. New records LIMASSOL: Troodos, Mount Olympos, 1 ♂, New records 9.IX.1997, Coll. Felton, det. G. Azarkina LARNACA : Larnaca SW, Hala Sultan Tekke (WML). Mosque, 5m, 1 ♂ 1 ♀, on herbs bordering salt lake, 16.V.2011 (CRB). Distribution: Mediterranean. New to Cyprus.

LIMASSOL : Akrotiri Peninsula, near Fasouri Aelurillus blandus (Simon, 1871) r eed beds, 1 ♂ 1 ♀, swept from flowering herbs and grass, 27.IV.2007 (CAR - S); Amathus, 2 ♀♀,

Spiders of Cyprus 103

Identification ♂♂ 1 ♀, pitfalls in open, mixed Pinus nigra and Azarkina, 2002; https://www.jumping - J uniperus foetidissima forest, 6.IV. - 8.VI.2014 spiders.com/ (CMH). NICOSIA: Cedar Valley Forest, 7 ♂♂, swept Citations from herbs, 29.IV.2007 (CRS), 1 ♂ (CAR - S); None. Cedar Valley, 1150m, 1 ♂, 21.III.2018, forest road, in gravel (CKHK); between Cedar Valley New records and Stavros tis Psokas, 2 ♀♀, under stones in PAPHOS : Akam as Peninsula, Lara Bay, 1 ♂, C. P inus nigra forest, 29.IV.2007 (CAR - S), same Saure leg., 18.VI.2013 (CKHK). site 1 ♂ 1 ♀ (CRS); Kampos, 1 ♂, Pinus brutia forest, 21.V.2016 (CMH); Pano Amiantos, 3 ♂♂, pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 Distribution: Greece, Crete. New to Cyprus. (CMH).

PAPHOS: Akamas Peninsula, Cape Geron nisos Aelurillus cypriotus Azarkina, 2006 Aelurillus cypriotus Azarkina, 2006: 68, figs 28 - S., 1 ♀, under stones in dry phrygana, 3.V.2007 36; Azarkina et al ., 2018: 49, figs 1 - 12, 15 - 19 , 24 - (CAR - S); Akamas Peninsula, between Cape 25. Drepanum and Lara Bay, 1 ♂ 1 ♀ rock strewn maquis near beach, 2.IV.1997, turning stones ù , Identification (CJB); Akamas Peninsula, Lara Bay, 1 ♂, C. Azarkina, 2006: 68, figs 28 - 36; Azarkina et al. Saure leg., 18.VI.20 13, same site, 1 ♂, 17.III.2018 2018: 49, figs 1 - 12, 15 - 19, 24 - 25. In the male, the longitudinal, white band on the abdomen is (CKHK); Akamas Peninsla, between Cape composed of white scales. In some specimens, Drepanou and Lara Bay, 1 ♂, 2.IV.1997 (CJB); these are completely rubbed off, the abdomen Avakas Gorge, 50m, 1 ♀, litter in citrus yard at then being completely black. entrance of gorge, 30.IV.2010 (CRB).

Citations Distribution: An endemic species of Cyprus. Without precise locality (Azarkina, 2006). LIMASSOL: Foini, Pano Platres; Trooditissa ; Mt. A ttulus distinguendus (Simon, 1868) Olympus, near Ski resort ( all Azarkina et al., 2018) . Identification NICOSIA: Mt Troodos, road Pedoulas - Lemithou Metzner, 1999: 522, fig. 1390. (Azarkina et al., 2018). PAPHOS: Akamas Peninsulae, Kioni; Kinousa; Citations Pano Arkhimandrita ( all Azarkina et al., 2018). None.

New records New records FAMAGUSTA: Cape Greco, 2 ♂♂, under stones LIMASSOL : Pano Platres, Gorge below in phrygana, 3.V.2013 (CAR - S), 1 ♀ (CDM); Caledonia Waterfall, 1 ♂, under stone, Rizokarpaso , 1 ♂, Juniperus phoenicea forest , 26.IV.2007 (CRS). 12.III.2016 (CMH). LIMASSOL : Akrotiri Peninsula, edge of central salt pan, 2 ♂♂, under stones and debris in Distribution: Palaearctic. Ne w to Cyprus . Salicornia , 27.IV.2007 (CAR - S) 1 ♀, (CDM); Akrotiri Peninsula E., 5m, 1 ♂, under stones and Ballus chalybeius (Walckenaer, 1802) in litter in salt marsh, 29.IV.2010 (CRB); Pano Plat res, valley near Caledonia Waterfall, 1 ♂, Identification under stones and in dried out stream bed, Metzner, 1999: 40, figs 6a - g. 26.IV.2007 (CAR - S); Troodos, Mount Olympos (Chionistra), 1900m, 2 ♂♂, stones in Pinus forest, Citations 23.V.2011 (CRB); idem, 2 ♂♂, open Juniperus None. phoenicea forest, 13.VI.2016 (CMH ); same site , 3 ♂♂, pitfalls in open Juniperus foetidissima forest, New records 6.IV. - 8.VI.2014 (CMH); Troodos Mountain, NICOSIA : Kakopetria S., Platania forest station, Agios Nikolaos tis Stegis, 1 ♂, on river bank at 1100m, 1 ♂, beating in dense Pinus forest, trout farm, 9.IX.1997 (WML); Troodos Square, 6 4.V.2010 (CRB). Spiders of Cyprus 104

forest, 28.IV.2007 (CRS); Troodos, near top of Distribution : Europe and N. Africa to Central Mount Olympos (Chionistra), 1 ♂, under stone in Asia. New to Cyprus. open P inus nigra forest, 28.IV.2007 (CRS)

NICOSIA: Agios Sozomenos , near Dali, under Ballus rufipes (Simon, 1868) Ballus rufipes ; Logunov, 2015: 53. stones in dry grassland, 30.IV.200 7 (CAR - S); Agridi, near Dali , 3 ♂♂, in house, 14.III.2010 Identification (CDM); Cedar Valley Forest, 1 ♂, under stones, Metzner, 1999: 40, figs 6a - g. 29.IV.2007 (CRS), 1 ♂ (CAR - S); Dali, 1 ♀, under stones and in grass in riverbed, 12.V.2007 (CAR - Citations S). NICOSIA: Dali (Logunov, 2015). PAPHOS : Akamas Peninsula, Smigies, 352m, 3

New reco rds ♂♂, ma quis, 12.III.2018 (CKHK); Evretou NICOSIA: Agridi, near Dali, 1 ♂, in garden, Reservoir, 350m, 1 ♀, in grassland around lake, IV.2004 (CDM). 1.V.2010 (CRB); Dhiarizos river near Kelefos bridge, 444m, 1 ♂, open river bank, 16.III.2018 Distribution : Europe and N. Africa to Central (CKHK); Gialia, 1 ♂, in pine wood, 25.III.1998, Asia. coll. J. Murphy (MMUE); Kelefos b ridge, dam on

Diarizos River, 550m, 1 ♀, under stones, Chalcoscirtus infimus (Simon, 1868) 21.V.2011 (CRB); Kinousa, 1 ♂, in pine wood, Identification 23.III.1998, 1 ♂, coll. J. Murphy (MMUE); Metzner, 1999: 45, fig. 11a. Mandria, 1 ♂, in scrub, 6.V.1982, coll. A. Stubbs (MMUE); Mavrokolympos Reservoir, 1 ♂, Citations 13.III.2018 (CKHK); Smigie s, 1 ♂, under stone in LARNACA: Tochni (Logunov, 2015). hillside scrub, 26.III.1998, coll. J. Murphy PAPHOS : Gialia (Logunov, 2015). (MMUE). New records FAMAGUSTA : Agios Sergios , 30m, 1 ♀, pitfalls Distribution : Southern Europe to Central Asia. in Pinus plantation, 15 - 27.V.2011 (CRB); Komi New to Cyprus. Kebir , 150m, 1 ♂, stones in Pinus forest, Cyrba algerina (Lucas, 1846) 18.III.2011 (CRB); Rizokarpaso, 1 ♂, in Cyrba algerina ; Logunov, 2013: 5; Logunov, Juniperus phoenicea forest, 12.III.2016 (CMH). 2015: 5. KERYNEIA: Pentadactylos Range, Saint Hilarion Castle, 600m, 1 ♀, stones in grassland, 17.V.2011 Identification (CRB). Me tzner, 1999: 35, figs 1a - h. LARNACA : Kornos picnic site, 350m, 1 ♀, stones in Pinus forest, 28.V.2011 (CRB); Larnaca Citations FAMAGUSTA: Protaras, Agios Ilias (Logunov, salt lake, 5m, 1 ♀, litter in s alt marsh, 27.IV.2010 2015). (CRB). LIMASSOL: Episkopi, Kourion Beach (Logunov, LIMASSOL : Alassa SE, 1 ♂ 2 ♀♀, under pebbles 2013). and in grass at edge of river Limnatis, 5.V.2007 PAPHOS : Avakas Gorge (Logunov, 2013). (CAR - S), 1 ♂ (CRS); Amathus, 1 ♀, under stones in dry valley, 6.IV.1986 (NHMW); Germasogeia, New records along stream below the dam, 1 ♀, under stones, FAMAGUSTA : Agios Sergios , 30m, 1 subadult 10.IV.1986 (NHMW); Pano Platres, Caledonia ♂, st ones in phrygana, 17.III.2011 (CRB); Cape Waterfall, 1 ♂, under stones in dried out stream Greco, Ayia Napa, 35m, 1 ♂ 1 ♀, under stones in bed, 26.IV.2007 (CAR - S); Pissouri, 1 ♂, in grass, phrygana, 27.IV.2010 (CJVK, CRB); Cape Greco, 24.III.1998, coll. J. Murphy (MMUE); Platres, 1 1 ♀, under stones in phrygana, 3.V.2013 (CAR - ♂, on walls, 22.IV.1986 (NHMW); Troodos S., S), 1 ♂ 2 ♀♀, (CDM); Flamoudi W ., 25m, 2 ♂♂ road to Pano Platres, 1 ♂, under stones in P. nigra 1 ♀, stones in phrygana near th e beach, 25.V.2011

Spiders of Cyprus 105

(CRB); Kantara castle, 650m, 1 subadult ♂, NICOSIA : Agridi, near Dali, 1 ♀, V.2004, 1 ♂, in stones in ruins, 19.III.2011 (CRB); Kantara E., yard, V.2009, 1 ♂, in house, 30.IV.2010, 1 ♂, in 610m, 1 ♀, stones in open Pinus forest, house, 5.XI.2010 (CDM); between Cedar Valley and Stavros tis Psokas, 1 ♂ 3 juveniles, under 25.V.2011 (CRB); Karpasia Peninsula, Cape stones in P inus nigra forest, 29.IV.2007 (CAR - S); Apostolos Andreas, 70m, 2 ♀♀, stones in Elia - Angolemi road, 100m, 1 ♂, stones in grassland, 16.V.2011 (CR B); Prastio S ., Agios grassland, 18.V.2011 (CRB); Kampos NE, Xeros Andronikos, 30m, 1 ♂, under stones in Eucalyptus River, 550m, 1 ♀, stones in riverbed, 20.V.2011 forest, 27.V.2011 (CRB); Profitis Elias, 1 ♂, 2005 (CRB); Kampos, 1 ♀, Pinus br utia forest, (CDM); Rizokarpaso , 1 ♀, Juniperus phoenicea 21.V.2016 (CMH); Kapedes S., 600m, 1 ♂, under forest, 7.V.2016, 1 ♀, 12.III.2016 (CMH); stones along rivulet, 5.V.2010 (CJVK); Kato Dali, Idalion Archaeological Site, 1 ♂, under stones in Salamis ruins, 30m, 2 subadult ♀♀, stones in phrygana, 30.IV. 2013 (CDM); Limnitis, 1 ♂ 1 ♀, ru ins, 17.III.2011 (CRB); Tula (Engomi) E., Pinus brutia forest, 13.V.2016 (CMH). Silver beach, 10m, 1 ♀, stones in salt marsh, PAPHOS : A kamas Peninsula, track along the sea, 15.V.2011 CRB); Vokolida S ., Cape Elaia, 10m, 1 10m, 1 ♂, stones in Juniperus phrygana, ♂ 1 ♀, under stones near the beach, 26.V.2011 30.IV.2010 (CJVK); Akamas Peninsula, Agios (CRB). Minas, 2 ♂♂, under stones in meadow with tall KERYNEIA: Larnakas tis Lapithou N. , 950m, 1 grass, 2.V.2007 (CAR - S); Akamas Peninsula, subadult ♂, under sto nes in Pinus forest, Cape Geronnisos S., 2 ♂♂ 2 juveniles, under 25.XI.2010 (CRB); Saint Hilarion Castle, 2 km st ones in dry phrygana, 3.V.2007 (CAR - S); W., 610m, 1 subadult ♀, stones in open Pinus Akourdaleia, 1 ♂, in garden, VIII.2005 (CDM); forest, 20.III.2011 (CRB), 1 ♀, beaten from Asprokremmos Dam, N. Kouklia, 150m, 1 ♂, shrubs in Pinus forest, 17.V.2011 (CRB). stones along lake, 29.IV.2010 (CJVK); Avakas LARNACA : Achna Reservoir, 50m, 1 ♀, stones Gorge, 50m, 1 ♂, litter in citrus orchard at bordering lake , 16.V.2011 (CRB); Cape Kiti, 2 entrance of gorge, 30.IV.2010 (CRB); 1 km N.E. ♂♂ 2 immatures, under stones below coastal of Pegeia, 1 ♂ 2 ♀♀, in pine wood, 30.IV.1982, cliffs, 25.IV.1982, coll. A. Stubbs (MMUE); Larnaca salt lake, 5m, 1 ♀, under stones and litter coll. A. Stubbs (MMUE); Souskiou, 3 ♂♂ 1 ♀, in salt marsh, 27.IV.2010 (CHVK), 1 ♀, pitfalls at 16.III.2013 (CDM). same site, III. - V.2011 (CRB). LIMASSOL : Akrotiri Peninsula, edge of central Distribution: Canary Islands to Central Asia. salt pan, 3 ♂♂ 2 ♀♀, under stones and debris in Salicornia , 27.IV.2007 (CAR - S), 1 ♀, 30.IV.2013 Euophrys herbigrada (Simon, 1871) (CAR - S), 2 ♂♂ (CDM); Amathus, 1 ♀, under Identification stones in dry valley, 1 ♂ 1 ♀, 6.IV .1986, 1 ♀, on Metzner, 1999: 50, figs 15 a - i. house on walls, 9.IV.19 86 (NHMW); Armenochori, 1 ♂, under stones of a dry stone Citations wall, 14.IV.1986 (NHMW); Erimi, 2 ♂♂, under None. stones along dry riverbed, 18.IV.1986 (NHMW); Germasogeia, along stream below the dam, 1 ♂, New records under stones, 10.IV.1986 (NHMW); Pano Platres, PAPHOS: Paphos, Bay at hotel “Paphian Bay”, 2 ♂♂ 1 ♀, sieving compost heap near hotel, valley near Cale donia Waterfall, 1 immature, 4.IV.1997 (CJB). under stones in dried out stream bed, 26.IV.2007 (CAR - S); Parekklisia, 1 ♂ 1 ♀, under stones in Distribution: Europe. New to Cyprus. riverbed, 13.IV.1986 (NHMW); Troodos, near top of Mount Olympos (Chionistra), 3 ♂♂, under Euophrys rufibarbis (Simon, 1868) stone in open P inus nigra forest, 28.IV.2 007 Euophrys rufibarb is ; Logunov, 2015: 56.

(CAR - S); Troodos S., road to Pano Platres, 1 ♂, Identification under stones in P inus nigra forest, 28.IV.2007 Metzner, 1999: 51, figs 16a - k. (CAR - S).

Spiders of Cyprus 106

Citations Com ments LIMASSOL : Troodos, near Kannoures (Logunov , The male of this species differs clearly from E. 2015). rufibarbis by the white palps, the orange basal fringe and the yellowish white subocular hairs, New records separated by a hairless area. FAMAGUSTA : Cape Greco, Ayia Napa, 35m, 1 ♂, under stones in phrygana, 20.XI.2010 (CRB); Distribution: Southern Europe, Turkey, Syria. Salam is ruins, 30m, 1 ♀, stones in ruins, First citation in Cyp rus.

17.III.2011 (CRB); Vokolida S , Cape Elaia, 10m, Evarcha insularis (Metzner, 1999) 1 ♀, under stones nea r the beach, 26.V.2011 (CRB). Identification KERYNEIA: Saint Hilarion Castle, 2 km W., Logunov , 2001: 60, figs 4 - 7. 610m, 2 ♀♀, stones in open Pinus forest, 20.III.2011 (CRB). Citations LARNACA : Kor nos picnic site, 350m, 1 ♀, under None. stones, 28.V.2011 (CRB); Stavrovouni New records Monastery, 350m, 2 ♀♀, on bark of Pinus trees, LARNACA: Agios Theodoros, 1 ♂, on herbs, 23. 28.IV.2010 (CRB). IV. 2016 (CMH). LIMASSOL : Fasouri marsh, 1 ♂, 17.VI.2016 NICOSIA: Kapedes, 520m, 1 ♂, pitfalls in Pinus (CMH); Troodos Range, Livadi tou Passia, 1750m, brutia f oret, 17.I. - 23.IV.2016, M. Hadjiconstantis 1 ♀, in pitfalls in peat mars h, III. - V.2011. NICOSIA : Cataldere dam, Kythrea N ., 105m, 1 leg. (CRB). ♂, under stones bordering lake, 23.XI.2010 PAPHOS: Stavros tis Psokas, 1 ♂, on the ground (CRB); Marathassa Dam, 550m, 1 ♀, under stones near the camp site, Pinus brutia forest, 12.V. around lake, 20.V.2011 (CRB); Prodromos S., 2018, M. Smith leg. (CMH). Trooditissa picnic site, 1450m, 1 ♀, stones in Pinus fore st, 21.V.2011 (CRB); Prodromos E., Distribution : Greece (Lesbos and Chios), Turkey, Iran. New to Cyprus. near Prodromos Dam, 1550m, 1 ♀, stones in

Pinus forest, 23.V.2011 (CRB). Evarcha jucunda (Lucas, 1846) PAPHOS : Akamas Peninsula, Cape Geronnisos S., 1 ♂ 1 ♀, under stones in dry phrygana, Identification 3.V.2007 (CRS); Evretou Reservoir, 350m, 1 ♀, Metzner, 1999: 148, fig. 113. beaten from shr ubs around lake, 1.V.2010 (CRB); Kato Akourdaleia, 1 ♂, in garden, VIII.2005 Citations (CDM). None.

Distribution : Mediterranean. New records NICOSIA: near Platania Scout camping site, 1 Euophrys sulphurea (L. Koch, 1867) ♀ pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH). Identification Metzner, 1999: 31, figs 17a - m. Distribution : Medi terranean.

Evarcha nepos (O. Pickard - Cambridge, 1872) N ew records Evarcha nepos ; Logunov, 2015: 61. FAMAGUSTA: Paralimn i Lake, 65m, 1 ♂, herbs and litter around dried lake, 27.IV.2010 (CRB). Identification LARNACA: Delikipos, 1 ♂, under stones and in Prószyn'ski, 2003: 58, figs 201, 217 - 219, 222 - 223. grass around small dam, 18.IV.2017 (CAR - S). NICOSIA: Marathassa Dam, 550m, 2 ♀♀, Citations beating bushes near water, 20.V.2011 (CRB). PAPHOS: Paphos (Logunov, 2015).

Spiders of Cyprus 107

New records deciduous wo od, 1 ♀, 16.III.2018 (CKHK); near FAMAGUSTA: Cape Gre co, Ayia Napa, 35m, 1 Kidasi, 1 ♂, in flood litter by stream, 1.V.2103 ♀, stones in phrygana, 27.IV.2010 (CRB). (CAR - S).

LARNACA : Dipotamos Reservoir, 150m, 4 ♂♂, Distribution: Greece, Syria, Cyprus, Israel, Syria. swept from herbs bordering lake, 28.IV.2010 (CRB). Heliophanillus fulgens (O. Pickard - Cambridge, LIMASSOL: Agios Tychonas, 1 ♂, under stones 1872) near house walls, 19.IV.1986 (NHMW). Identification NICOSIA: Ath alassa National Forest Park, 1 ♂, Metzner, 1999: 97, figs 62a - i. Pinus brutia forest, 21.II.2016 (CMH). Citations PAPHOS : Mavrokolympos Reservoir, NE None. Potima, 150m, 1 ♀, under stones along lake, 22.V.2011 (CRB). New records LIMASSOL : Alassa SE, near river Limnatis, 2 Distribution : Turkey, Israel, Cyprus. ♂♂; under stones in dry grassland and at edge of river, 5.V.2007 (CAR - S), 1 ♀ (CRS); Kolossi Evarcha patagiata (O. Pickard - Cambri dge, castle, 1 ♂, walls in the castle area, 11.IV.1986 1872) (NHMW). Evarcha patagiata ; Logunov, 2013: 5; Logunov, 2015: 61. NICOSIA : Agridi, near Dali, 2 ♂♂ 3 ♀♀, in house, V.2010, 3 ♀♀ in garden, 1 ♂ 2 ♀♀, in Identification : https://araneae.nmbe.ch/ . house, IV.2011, (all CDM); 13, Dali, Agios Andronikos, 1 ♂, in garden, 2007 CDM); Lefka Citations NW, 120m, 1 ♀, beaten from shrubs in fruit LIMASSOL: Troodos, Agios Nikolaos (Logunov, plantation, 18.V.2011 (CRB); Lympia Reservoir, 2015). 2 ♂♂, in grass tussocks and on concrete duct, PAPHOS : Paphos forest (Lo gunov, 2015); Polis W., Akamas Forest (Logunov, 2013). 9.V.2013 (CAR - S); 1 ♀ (CDM); Marathassa Dam, 550m, 3 ♂♂, swept from herbs around lake, New records 20.V.2011 (CRB) . KERYNEIA: Pentadactylos Range, Saint Hilarion PAPHOS : Evretou Reservoir, 350m, 3 ♂♂ 1 ♀, Castle W., 600m, 1 ♂, beaten from shrubs in wet grassland around lake, 1.V.2010 (CRB); Pinus forest, 17.V.2011 (CRB). Kelefos bridge, dam on Diar izos River, 550m, 2 LIMASSOL: Amathus, 1 ♂, under stones in dry ♂♂, under stones near lake, 21.V.2011 (CRB); valley, 25.IV.1 986 (NHMW). Mavrokolympos Reservoir, NE . Potima, 150m, 1 NICOSIA : Cedar Valley Forest, 1180m, 1 ♂, ♀, under stones along lake, 22.V.2011 (CRB); beaten in Cedrus forest, 19.V.2011 (CRB); Lefka Mavrokolympos Reservoir, 1 ♀, C. Saure leg., NW, 120m, 1 ♀, beaten from shrubs in fruit 20.VI.2013 (CKHK). plantation, 18.V.2011 (CRB); Marathassa Dam, 550m, 1 ♀, beaten from shrubs around lake, Distribution: Middle East, Greec e, North Africa. 20.V.2011 (CRB); Oik os N., Xerargaka picnic New to Cyprus. site, 350m, 1 ♂, beaten from shrubs and stones in Pinus forest, 20.V.2011 (CRB). Heliophanus auratus C. L. Koch, 1835 Heliophanus auratus ; Wesolowska, 1986: 212. PAPHOS: Akamas Peninsula, Avakas Gorge, 2

♂♂, under stone, 27.III.1998, coll. J. Murphy (MMUE); Avakas gorge and surroundings, 1 ♂, Identification : https://araneae.nmbe.ch/ . 22.III.2018 (CKHK ); Akamas Peninsula, Agios Minas, 2 ♂♂ 1 ♀, under stones in meadow with Citations tall grass, 2.V.2007 (CRS); Evretou Reservoir, NICOSIA : Pendayia (Wesolows ka, 1986).

Stavros tis Psokas river, 2 ♂♂, 20.III.2018 (CKHK); Kato Akourdaleia, 1 ♂, in garden, New records None. VIII.2010 (CDM); Kelefos bridge, 465m,

Spiders of Cyprus 108

Distribution: Palearctic. Citations None. Heliophanus decoratus L. Koch, 1875 Ne w records Identification LARNACA: Stavrovouni Monastery, 3 ♀♀, in Wesolowska, 1986: 208, figs 549 - 557, 559 - 566, phrygana, 26.IV.1982, coll. A. Stubbs (MMUE). 570 - 584. LIMASSOL : Akrotiri Peninsula, 6 ♂♂ 14 ♀♀, Citations margin of central salt lake, 7.V.1972, coll. A. None. Stubbs (MMUE); Akrotiri Peninsula, near Fasouri reed beds, 1 ♂ 4 ♀♀, swept from flowering herbs New records and grass, 27.IV. and 4.V.2007 (CAR - S), 1 ♀, in FAMAGUSTA: Paralimni Lake, 65m, 2 ♂♂ , reed litter, 27.IV.2007 (CAR - S), same site, 3 ♂♂ herbs and litter around dried lake, 27.IV.2010 (CRS). (CRB); Rizokarpaso , 1 ♀, Juniperus phoenicea NICOSIA: Dali, 13, Agios Andronikos, 1 ♂, in forest, 12.III.2016 (CMH). garden, IV.2007 (CDM); Lympia Reservoir, LARNACA : Larnaca salt lake, 5m, 1 ♂ 2 ♀♀, Lympia, 2 ♂♂ 2 ♀♀, in phrygana b elow dam stones and litter in salt marsh, 27.IV.2010 (CRB); wall, 9.V.2013 (CDM). Larnaca SW, Hala Sultan Tekke Mosque, 5m , 1 ♂ PAPHOS: Near Paphos, 1 ♂, in marshy area, 1 ♀ stones and litter bordering salt lake, 1.V.1982, coll. A. Stubbs (MMUE).

16.V.2011; (CRB); Meneou, 2 ♂♂ 2 ♀♀, salt Distribution: Corsica, Greece, Asia Minor. New marsh and fields, 8.IV. - 15.IX.2016 (CMH). to Cyprus. LIMASSOL : Akrotiri Peninsula, 1 ♀, margin of central salt lake, 7.V.1972, coll. A. Stubbs Heliophanus encifer Simon, 1871 (MMUE); Akrotiri Peninsula, 2m 1 ♀, 27.IX.2007 Heliophanus encifer ; Wesolowska, 1986: 40; (CDM); Akrotiri Peninsula, edge of central salt L ogunov, 2015: 65. pan, 1 ♂, under stones and debris in Salicornia , Identification 27.IV.2007; 1 ♀, 30.IV.2013 (CAR - S); Akrotiri Wesolowska, 1986: 40, figs 456 - 466. Peninsula, salt lake, 2 ♂♂, under Salicornia , 15.III.2018 (CKHK); Akrotiri Peninsula, near Citations Britis h Airbase, 1 ♀, swept in wet area, 4.V.2007 Without precise locality (Wesolowska, 1986). (CAR - S); Alassa SE, near river Limnatis, 2 ♂♂, NICOSIA: Dali (Logunov, 2015); under stones in dry grassland and at edge of river, PAPHOS: Gialia (Logunov, 2015)

5.V.2007 (CRS); 5 km S. of Troodos, road to New records Pano Platres, 1 ♂, under stones, Pinus nigra FAMAGUSTA : Cape Greco, 1 ♀, under stones in forest, 1.V.2007 (CRS); Troodos Square, 2 ♂♂, phrygana, 3.V.2013 (CAR - S); Kantara E., 610m, Pinus nigra forest, 11.V.2016 (CMH). 1 ♂ 4 ♀♀, beaten from shrubs and stones in open NICOSIA: Agridi, near Dali, 1 ♂, in field, Pinus forest, 25.V.2011 (CRB); Lysi, 1 ♂, in IV.2007, 1 ♂, in house, V.2009, 1 ♂, in house, garden, 15.IV.2016 (CMH). 7.V.2010 (all CDM); Dali, 13, Agios Andronikos, KERYNEIA: Pentadactylos Range, Lapithos S ., 3 ♂ 1 ♀, in garden, IV.2007 (CDM); Dali, 1 ♂, 800m, 1 ♀, under stones in Pinus f orest, under stone s and in grass in riverbed, 12.V.2007 17.V.2011 (CRB); Pentadactylos Range, Saint (CAR - S). Hilarion Castle, 600m, 2 ♂♂ 1 ♀, beaten from shrubs in Pinus forest, 17.V.2011 (CRB). Distribution : Morocco, Algeria, Tunisia, Lybia, LARNACA : Larnaca SW, Hala Sultan Tekke Egypt, Syria, Iran. New to Cyprus. Mosque, 5m, 1 ♀, swept from herbs bordering salt

Heliophanus edentulus Simon, 1871 lake, 16.V.2011 (CRB); Meneou , 1 ♀, salt marsh and fields, 8.IV. - 15.IX.2016 (CMH). Identification LIMASSOL : Alassa SE, near river Limnatis, 2 Metzner, 1999: 98, figs 63a - h. ♂♂, under stones in dry grassland, 5.V.2007 (CAR - S); Kouklia - Pissouri road, 150m, 1 ♀,

Spiders of Cyprus 109

stony slope near sea, 22.III.2011 (CRB); Pano Citations Platres, below Caledoni a Waterfall, 1 ♂, under None. stones and in grass in rocky valley, 26.IV.2007 (CAR - S); Pano Platres, Millomeris Waterfalls, 1 New records LARNACA : Achna Reservoir, 50m, 2 ♂♂, 2 ♀♀, ♂, under rocks and litter, 28.IV.2007 (CAR - S); 5 swept from low herbs bordering lake 27.V.2011 km S. of Troodos, road to Pano Platres, 1 ♂, (CRB); Pano Lefkara, 200m, 1 ♀, very dry karst, under stones, P. nigra forest, 28.IV.2007 (CRS). 29.IV.2017, S. Huber leg. (CSH); Stavrovouni NICOSIA : Agios Sozomenos , near Dali, 1 ♂, Monastery, 2 ♂♂ 1 ♀, in phrygana, 26.IV.1982, under stones in dry grassland, 30.IV.2007 (CAR - coll. A. Stubbs (MMUE). S); Agridi, near Dali, 1 ♀, in garden, IV.2004 LIMASSOL : Akrotiri Peninsula, 1 ♀, margin of (CDM); Chrysoroyiatissa Monastery, 1 ♂, in pine central salt lake, 7.V.1972, coll. A . Stubbs wood 2.V.1982, coll. A. Stubbs (MMUE); (MMUE); Akrotiri Peninsula, near Fasouri reed Lymbourena Hill, Dali, 1 ♂, IV.2004 (CDM); beds, 1 ♂, swept from flowering herbs and grass, Cedar Valley Forest, 1180m, 3 ♀♀, beaten in Cedrus forest, 19.V.2011 (CAR - S), 1 ♂ (CRS); 27.IV.2007 (CRS); Alassa SE, near river between Cedar Valley and Stavros tis Psokas, 1 Limnatis, 1 ♀, under stones in dry grassland, ♂, under stones in P. nigra forest, 29.IV.2007 5.V.2007 (CRS); Amathus, 1 ♀, under stones in (CAR - S); Dali, 13, Agios Andronikos, 1 ♂, in dry valley, 6.IV.198 6. NICOSIA : Lefka NW, garden, 1 0.III.2009 (CDM); Kampos NE, Xeros 120m, 2 ♂♂, 2 ♀♀, beaten from shrubs in fruit River, 550m, 1 ♂ 6 ♀♀, beaten from shrubs, plantation, 18.V.2011 (CRB); Lympia Reservoir, 20.V.2011 (CRB); Lefka NW, 120m, beaten from Lympia, 1 ♂, in grass tussocks and on concrete shrubs in fruit planta tion, 3 ♂♂ 1 ♀, 18.V.2011 (CRB) , Lympia Reservoir, Lympia, 1 ♀, under duct, 9.V.2013 (CAR - S). stones below dam wall, 9.V.2013 (CAR - S) ; PAPHOS : Akamas Peninsula, Avakas Gorge, 1 Marathassa Dam, 550m, 1 ♀, beaten from shrubs ♂, under stone, 27.III.199 8, coll. J. Murphy around lake, 20.V.2011 (CRB); Prodromos S., (MMUE); Akrotiri peninsula, 1 ♀, swept from Trooditissa picnic site, 1450m, 2 ♂♂ 2 ♀♀, grass and reeds, Fassouri reedbeds, 17.IV.2017 beaten from shrubs in Pinus forest, 21.V.2011 (CAR - S); Souskiou, 1 ♂, under stones on rocky (CRB); Strovolos, 1 ♀, around houses, 1.V.2016 eroded grassland on hill, 17.IV.2017 (CAR - (CMH). S).Drouseia, 539m, 1 ♂, in p hrygana, 23.III.2018 PAPHOS : Akamas Peni nsula, 2 ♀♀, in coastal (CKHK); M avrokolympos Reservoir, 1 ♀, C. grassland, 22.III.1998, coll. J. Murphy (MMUE); Saure leg., 20.VI.2013 (CKHK); Paphos, Bay at Akamas Peninsula, Smigies, 352m, 2 ♂♂, hotel “Paphian Bay”, 1 ♂, stones on rock strewn maquis, 12.III.2018 (CKHK); Baths of Aphrodite, terrain near beach, 31.III.1997 (CJB); Pegeia, 1 ♂, under stone in coastal scrub, 18.III.1998, Pikni Forest, 374m, 1 ♂, pitfalls in open ruderal coll. J. Murphy (MMUE); Evretou Reservoir, site, 14.III. - 23.III.2018 (CKHK ); Vretsia W., Stavro s tis Psokas river, 1 ♀, 20.III.2018 near Kelefos Bridge, 650m, 2 ♀♀, beaten from (CKHK); Gialia, 3 ♂♂ 4 ♀♀, in pine wood, shrubs, 19.V.2011 (CRB). 25.III.1998, coll. J. Murphy (MMUE); Kato Paphos archaeological site, 1 ♀, under stones in Distribution : Italy, the Balkans, the Caucasus, grass, 10.V.2007 (CAR - S); Akamas Peninsula, Asia Minor. New to Cyprus. Lara Bay, 1 ♂, stones on rock strewn beach, 3.IV.1997 (CJB); Smigies, 1 ♂, under stone in Heliophanus flavipes Hahn, 1832 hillside scrub, 26.III.1998, coll. J. Murphy (MMUE); Mavrokolympos Reservoir, 2 ♂♂ 4 Identification : https://araneae.nmbe.ch/ . ♀♀, 13.III.2018 (CKHK); Pegeia, Pikni Forest, 346m, 3 ♂♂ 1 ♀, in former carob plantation, Citations 11.III.2018 (CKHK). None.

Distribut ion: Corsica, Italy, Israel, Syria, Cyprus. New records LARNACA : Achna Reservoir, 50m, 1 ♀, under Heliophanus equester L. Koch, 1867 stones and litter bordering lake, 27.V.2011 (CRB). Identification Metzner, 1999: 102, figs 68a - i. Distribution: Palearctic. New to Cyprus.

Spiders of Cyprus 110

Citations Heliophanus melinus L. Koch, 186 7 LIMASSOL: Agridia (Logunov, 2015). Heliophanus melinus ; Kulczyński, 1903: 35; NICOSIA: Dali (Logunov, 2015). Wesołowska, 1986: 220; Metzner, 1999: 108; PAPHOS: Souskiou (Logunov, 2015). Dobroruka, 2003: 14; New records Identification FAMAGUSTA : Kantara E., 610m, 4 ♀♀, beaten Metzner, 1999: 108, figs 74a - g. from shrubs in open Pinus forest, 25.V .2011 (CRB). Citations KERYNEIA: Pentadactylos Range, Ftericha S ., None. 800m, 1 ♂, stones in grassland, 17.V.2011 (CRB); New records Pentadactylos Range, Lapithos S ., 800m, 2 ♂♂ 1 LARNACA : Agios Theodoros, 1 ♂ 1 ♀, on herbs , ♀, beaten in Pinus forest, 24.V.2011 (CRB). 23.IV.2016 (CMH); Larnaca SW, Hala Sultan LIMASSOL : Alassa SE, near river Limnatis, 2 Tekke Mosque, 5m, 1 ♀, in litter bordering salt ♂♂, under s tones in dry grassland, 5.V.2007 lake, 16.V.2011 (CRB). (CAR - S); Amathus, 1 ♀, under stones in dry KERYNEIA: Panagia Apsinthiotissa Monastery valley, 25.IV.1986 (NHMW); Fasoula, 1 ♂, olive ruins, 350m, 1 ♀, stones in clearing in Pinus garden with dry stone walls, 16.IV.1986 forest, 24.V.2011 (CRB). (NHMW); Pissouri beach, 1 ♂, under stones, LIMASSOL: Akrotiri Peni nsula, 2 ♀♀, 27.IV.1982, coll. A. Stubbs (MMUE); Platres, 1 27.IX.2007 (CDM); Alassa SE, near river ♂, on walls, 22.IV.1986 (NHMW). Limnatis, 1 ♀, under stones in dry grassland, NICOSIA: Agridi, near Dali, 1 ♂, in garden, 5.V.2007 (CRS). IV.2004, 1 ♀, in garden, V.2011, 1 ♂, in garden, NICOSIA : Agridi, near Dali 1 ♀, grass in lawn, V.2012 (CDM); Cedar Valley Forest, 1180m, 3 1.VII.2010, 2 ♀♀, in house, V.2011 (CDM); ♀♀, beaten in Cedrus forest, 19.V.2011 (CRB); Athalassa National Forest Park, Nicosia. 1 ♀, Kakopetria S., Platania forest station, 11 00m, 1 ♂, unde r stones in pine plantation, 2.V.2013 (CDM); under stones in dense Pinus forest, 4.V.2010 Dali, under stones near old water mill, 1 ♀, (CRB). 12.V.2007 (CAR - S); Kampos NE, Xeros River, 550m, 1 ♀, beaten from shrubs, 20.V.2011 PAPHOS: Akamas Peninsula, Avakas Gorge, 1 (CRB); Lympia Reservoir, Lympia, 2 ♀♀, in ♂, under stone, 27.III.1998, coll. J. Murphy clump of rushes below dam wall, 9.V.2013 . (MMUE); 1 km N.E. of Pegeia, 1 ♂, in pine PAPHOS : Evretou Reservoir, 350m, 1 ♂ 2 ♀♀, wood, 30.IV.1982, coll. A. Stubbs (MMUE); near Souskiou, 1 ♂, u nder stones in dry grassland, beaten from shrubs, 1.V.2010 (CRB); Kidasi N., 1.V.2013 (CAR - S). along Diarizos River, 300m, 3 ♀♀, beaten from shrubs, 21.V.2011 (CRB); Mavrokolympos Distribution : Greece to Central Asia. Reservoir, NE Potima, 150m, 1 ♀, under stones along lake, 22.V.2011 (CRB); ne ar Souskiou, 1 ♀, Macaroeris nidicolens (Walckenaer, 1802) under stones in dry grassland, 1.V.2013 (CAR - S); Macaroeris nidicolens ; Logunov, 2015: 69. 1 ♀ (CDM). Identification

Metzner, 1999: 43, figs 9d - e, 10a - l. Distribution : South Europe to Mediterranean.

New to Cyprus. Citations

PAPHOS: Pano Panagia (Logunov, 2015). Heliophanus mordax (O. Pickard - Cambridge,

1872) New records Heliophanus mordax ; Logunov, 2015: 66. FAMAGUSTA : Agios Sergios , 30m, 1 ♀, beaten

Identification Salamis ruins, 50m, 2 ♂♂ 2 ♀♀, beaten from low Metzner, 1999: 110, figs 75a - h. trees, 15.V.2011 (CRB); in Pinus plantation, 15.V.2011 (CRB).

Spiders of Cyprus 111

KERYNEIA: Pentadactylos Range, Saint Hilario n beds, 1 subadult ♂, 1 immature, swept from Castle, 600m, 1 ♂ 1 ♀, beaten from shrubs, flowering herbs and grass, 27.IV.2007 (CAR - S). 17.V.2011 (CRB). Distribution : Palaearctic and N. Africa. New to LIMASSOL: Pano Platres, 1050m, 1 ♂, beaten Cypr us. near Mesa Potamos Waterfall s, 23.V.2011 (CRB). NICOSIA : Agridi, near Dali, 1 ♀, in garden, Menemerus bivittatus (Dufour, 1831) V.2011 (CDM); Chrysoroyiatissa Monastery, 3 ♂♂, in pine wood 2.V.1982, coll. A. Stubbs Identification : https://araneae.nmbe.ch/ (MMUE); Marathassa Dam, 550m, 1 ♀, beaten from shrubs around lake, 20.V.2011 (CRB); Citations Oikos N., Xerargaka picnic site, 350m, 1 ♂ 1 ♀, None. beaten from shrubs in Pinus forest, 20.V.2011 (CRB). New records PAPHOS : A kamas Peninsula, Agios Minas, 1 ♀, PAPHOS: Kato Akourdaleia, 1 ♂, in garden, beaten from climbers on pines, 2.V.2007 (CAR - VII.2008 (CDM). S), 1 ♀ (CRS); near Avakas Gorge, 1 ♂, beaten from junipers on rocky hillside, 14.IV.2017 Distribution : P antropical. Introduced to Europe. (CAR - S); Evretou Reservoir, 350m, 1 ♀, beaten New to Cyprus. from shrubs and grassland around lake, 1.V.2010 (CRB); Kelefos bridge , 465m, deciduous wood, 1 Menemerus fagei Berland & Millot, 1941 ♀, 16.III.2018 (CKHK); Kourdaka, Ezousa river, 1 ♂, under bark of Eucalyptus , 14.III.2018 Identification : https://araneae.nmbe.ch/ (CKHK); Pegeia, Pikni Forest, 366m, 1 ♂, in pine forest under bark, 11.III.2018 (CKHK). Citations None. Distribution : Mediterranean to Central Asia. New records Marpissa nivoyi (Lucas, 1846) NICOSIA: Agridi, near Dali, 1 ♀, in field, Marpissa nivoyi ; Metzner, 1999: 139. IV.2007, 1 ♀, on bark of Eucalyptus , 8.II.2010, det. D. Logunov (CAR - S). Identification : https://araneae.nmbe.ch/ . Distribution : West Africa to Yemen, Malta. New Citations to Cyprus. None. Menemerus semilimbatus (Hahn, 1829) New records LIMASSOL : Akrotiri Peninsula, Fasouri reed Identification : h ttps://araneae.nmbe.ch/ beds, 1 ♀, in reed litter, 4.V.2007 (CRS); Lady’s Mile Beach, SW Limassol, 1 ♀, in salt marsh, 19.II.2011, D. W. Wrase leg. (CKHK). Citations None. Distribution : Europe to Central Asia. New to Cyprus. New records LARNACA : Lakatamia, 1 ♂, in olive grove, 1.I - Mendoza canestrinii (Ninni, 1868) 1.X. 2016 (CMH); Larnaca SW, Hala Sultan

Tekke Mosque, 5m, 1 ♂, on stone wall, Identification Metzner, 1999 : 144, figs 109a - h. 16.V.2011 (CRB); Meneou, 1 ♂ 1 ♀, salt marsh and fields, 8.IV. - 1 5.IX.2016 (CMH). Citations NICOSIA: Agridi, near Dali, 1 ♀, in garden, None. V.2004, 1 ♂, in garden, VIII.2010 (CDM); Dali, Lymbourena Hill, 1 ♀, IV.2004 (CDM). New records PAPHOS: Akamas Peninsula, 1 ♂, 8.VII.2009 LARNACA: Akrotiri Peninsula, near Fasouri reed (CDM); Kato Akourdaleia, 1 ♂ 1 ♀, in garden,

Spiders of Cyprus 112

VIII.2005, 2 ♀♀, in garden, VIII.2010, 1 ♂, on NICOSIA : Kampos NE, Xeros River, 550m, 2 building, 10.VIII.2012 (CDM). ♀♀, beaten from shrubs, 20.V.2011 (CRB); Kapedes, 1 ♂, pitfalls in Pinus brutia forest, 17.I. Distribution : Canary Islands to Central Asia. - 23.IV.2016 (CMH); Lefka NW, 120m, 1 ♂, New to Cyprus. beaten from shrubs in fruit plantation, 18.V.2011 Mogrus logunovi Prószyński, 2000 (CRB) ; Profi tis Ilias, Machairas Forest, 1 ♀, 2005 (CDM). Identification : https://araneae.nmbe.ch/ Distribution: Greece, North Macedonia, Turkey, Citations Israel, Azerbaijan. None. Neaetha absheronica Logunov & Guseinov, 2002 New Records LARNACA: Akrotiri Peninsula, edge of central Identification: https://araneae.nmbe.ch / . salt pan, 2 ♀♀, in silken cells in Salicornia , 30.IV.2013 , det. D. Logunov (CAR - S), 2 ♀♀, Citations 17.IX.2007 (CDM). None. NICOSIA: Agridi, near Dali, 1 ♀, in field, V.2007 (CDM). New records LIMASSOL: Akrotiri Peninsula, Turtle Beach, 1 Distribution : Egypt, Israel, Jordan, Yemen, ♂, 15.III.2018, sandy beach (CKHK). United Arab Emirates. New to Cyprus. NICOSIA: Agios Sozomenos, near Dali, 1 ♂,

under stones in dry rocky phrygana, 20.IV.2017, Mogrus neglectus (Simon, 1868) Mogrus neglectus ; Logunov, 2015: 70. det. D. Logunov (CAR - S). PAPHOS: A kamas Peninsula, Lara Bay, 1 ♂ 1 ♀, Identification C. Saure leg., 18.VI.2013 (CKHK); Paphos, Bay Metzner, 1999: 70, figs 35a - l. at hotel “Paphian Bay”, 1 ♂, pitfalls traps in rock

Citations strewn terrain near beach, 6.IV.1997 (CJB). FAMAGUSTA : Profitis Ilias (Log unov, 2015). LIMASSOL: Akrotiri Peninsula (Logunov, 2015). Distribution: North Macedonia, Greece, Turkey, Azerbaijan. New to Cyprus. New records FAMAGUSTA : Lysi, 1 ♂, in garden, 15.IV.2016 Neon rayi (Simon, 1875) (CMH); Paralimni Lake, 65m, 1 ♀, on herbs around dried lake, 27.IV.2010 (CRB); Prastio S ., Identification Agios Andronikos, 30m, 1 subadult ♂ 4 ♀♀, on Metzner, 1999: 66, figs 31a - g. herbs in Eucalyptus plantation, 27.V.2011 (CRB); Vokolida S ., Cape Elaia, 10m, 4 ♀♀, under stones Citations None. and on shrubs near the beach, 26.V.2011 (CRB).

LARNACA : Larnaca SW, Hala Sultan Tekke New records mosque, 5m, 2 ♀♀, surrounding park, 20.IV.1986 FAMAGUSTA : Agios Sergios , 30m, 1 ♂, stones (NHMW); same site, 2 ♀♀, on herbs in salt in phrygana, 17.III.2011 (CRB). marsh, 27.IV.2010 (CRB); same site, 1 ♂ 1 ♀, on PAPHOS : Akamas Peninsula, Agios Minas, 1 ♀, herbs bordering salt lake, 16.V.2011 (CRB); in thick p ine litter, 2.V.2007 (CAR - S); Akamas Meneou, 1 ♀, salt marsh and fields, 8.IV. - Peninsula, Cape Geronnisos S., 1 ♀, under stones 15.IX.2016 (CMH). in dry phrygana, 3.V.2007 (CAR - S); Gialia, 1 ♀, LIMASSOL: Akrotiri Peninsula, near British in pine wood, 25.III.1998, coll. J. Murphy (MMUE). Airbase, 1 ♀, swept in pine wood, 4.V .2007

(CAR - S). Distribution : Europe to southern regions of Central Asia, Algeria. New to Cyprus.

Spiders of Cyprus 113

Pellenes brevis (Simon, 1868) Identification Pellenes brevis ; Logunov, 2015: 72. Metzner, 1999: 12 8 , figs 94 a - j.

Identification Citations Metzner, 1999: 125, figs 90 a - k . PAPHOS: Troodos, Agio s Nikolado (Logunov, 2015). Citations PAPHOS: N. Paphos, Lara (Logunov, 2015); New records Stavros Tis Psokas, Paphos forest (Logunov, None. 2015). Distribution: Southern Europe, Africa, Turkey, N ew records Ukraine, Caucasus to Central Asia . None.

Distribution: France, Spain, Bulgaria, Greece. Philaeus chrysops (Poda, 1761)

Pellenes diagonalis (Simon, 1868) Identification Metzner, 1999: 140, figs 105a - h, 106a - l. Identification Metzner, 1999: 128, figs 91a - i ( ♀, sub P. Citatio ns ostrinus ); Metzner, 1999: 121, figs 86a - k ( ♂, sub None. P. diagonalis ). New records Citations KERYNEIA: Keryneia , 1 ♀, near the town in None. Juniperus phoenicea forest, 30.IV.2016 (CMH); Pentadactylos Range, Saint Hilarion Castle W., New records 600m, 1 ♀, beaten from shrubs in Pinus forest, LIMASSOL : Agios Georgios E., Palaiomylos, 17.V.2011 (CRB). 1250m, 1 ♀, beaten in mixed Pinus and Quercus LARNACA : Agioi Vavatsinia s, 700m, 2 ♀♀, forest, 4.V.2010 (CRB). litter in Pinus forest, 28.IV.2010 (CRB); Meneou, 1 ♂, salt marsh and fields, 8.IV. - 15.IX.2016 Distribution: Greece (Corfu), Macedonia, (CMH). Turkey, Israel. New to Cyprus. NICOSIA : Dali, 13, Agios Andronikos, 1 ♀, in garden, IV.2007 (CDM); Machairas forest near Pellenes flavipalpis (Lucas, 1853) Kionia, 1300m, 3 ♂♂ 2 ♀♀, stony phrygana, 5.V. 2010 (CJVK, CRB); Peristerona, 1 ♀, Identification riverbed, 7.V.2016 (CMH). Azarkina, 2006: 68, figs 28 - 36 LIMASSOL: Akrotiri Peninsula N., 5m, 1 ♀,

Citations beating in Eucalyptus forest, 29.IV.2010 (CRB); None. In giving the general distribution area of Erimi, 1 ♂, under stones along dry riverbed, this species, Drensky (1936) and Hadjissarantos 18.IV.1986 (NHMW); Platres, 1 ♂, on walls, (1940) mention Cyprus, but we did not find any 22.IV.1986 (NH MW). precise citation or information. PAPHOS : Evretou Reservoir, 1 ♂, under stones and in grass tussocks, 13.IV.2017 (CAR - S); New rec ords Kelefos bridge, dam on Diarizos River, 550m, 1 None. ♀, beaten from shrubs, 21.V.2011 (CRB); near

Souskiou, 1 ♀, under stones in dry grassland, Distribution: Greece, Crete. The presence in Cyprus must be confirmed. 1.V.2013 (CDM).

Pellenes geniculatus (Simon, 1868) Di stribution : Southern Palaearctic. New to P ellenes geniculatus ; Logunov, 2015: 72. Cyprus.

Spiders of Cyprus 114

Phlegra bresnieri (Lucas, 1846) Plexippoides flavescens (O . Pickard - Cambridge, 1872) Identification Metzner, 1999: 70, figs 35a - l Identification: https://araneae.nmbe.ch/ .

Citations Citations None. None.

New records New records FAMAGUSTA : Prastio S ., Agios Andronikos, LIMASSOL: Amathus, 1 ♀, under stones in dry 30m, 1 ♀, under stones and h erbs in Eucalyptus valley, 6.IV.1986 (NHMW). forest, 27.V.2011 (CRB); Rizokarpaso , 1 ♂, Distribution : Greece to Central Asia, Pakistan, Juniperus phoenicea forest, 12.III.2016 (CMH). Sudan. LIMASSOL : Pano Platres, below Caledonia Waterfall, 1 ♂, under stones and in grass in rocky Plexippoides gestroi (Dalmas, 1920) valley, 26.IV.2007 (CAR - S). Plexippoides gestroi ; Logunov, 2015: 73. NICOSIA : Agios Sozomenos , near Dali, 2 ♀♀, under stones in dry grassland, 30.IV.2007 (CAR - Identification S). Metzner, 1999: 135, figs 99a - i.

Distribution : Mediterranean, Syria. New to Citations Cyprus. LIMASSOL: E. Limassol, Mount Amathus

(Logunov, 2015). Phlegra cinereofasciata (Simon, 1868) New r ecords Phlegra cinereofasciata ; Logunov, 2015: 73. LIMASSOL: Pissouri, 1 ♀, in grass, 24.III.1998,

coll. J. Murphy (MMUE); Platres, 1 ♀, on walls, Identification 22.IV.1986 (NHMW). Metzner, 1999: 69, fig s 34a - k.

Distribution : Mediterranean. Citations

LIMASSOL : Troodos, Mt Olympos (Logunov, Plexippus devorans (O. Pickard - Cambridge, 2015). 1872)

New records Identification LIMASSOL: road Chionistra - Prodromos, 1 ♂, Metzner, 1999: 136, fig s 100a - j. pitfalls in Pinus nigra forest, 6.IV. - 8.VI.2014

(CMH). Citations

None. Distribution : Portugal to Central Asia.

New records Phlegra fasciata (Hahn, 1826) NICOSIA: Lympia Reservoir, Lympia, 1 ♂, under Phlegra fasciata ; Logunov, 2015: 74. stones below dam wall, 9.V.2013 (CDM).

Identification Distribution : Greece to Central Asia, Algeria. Metzner, 1999: 68, figs 33a - i. New to Cyprus.

Citations Plexippus paykulli (Audouin, 1826) Cyprus, without further data (Logunov , 2015).

Identif ication: https://araneae.nmbe.ch/ . New records

None. Citations

None. Distribution : Palaearctic.

Spiders of Cyprus 115

New records New records KERYNEIA: Karavas , 660m, 1 ♂, on stone walls KERYNEIA: Pe ntadactylos Range, Saint Hilarion in garden of hotel, 22.XI.2010 (CRB). Castle, 600m, 1 ♀, beaten from shrubs, 17.V.2011 LARNACA : Larnaca SW, Hala Sultan Tekke (CRB). Mosque, 5m , 1 ♀, on stone walls near mosque, 16.V.2011 (CRB); same site , 1 ♀, 20.IV.1986 Distribution : Mediterranean to Azerbaijan. New (NHMW). to Cyprus.

LIMASSOL: Ama thus, hotel, 2 ♀♀, front yard, Pseudicius vankeeri Metzner, 1999 8.IV.1986 (NHMW); Amathus , 1 ♀, in the Pseudicius vankeeri ; Logunov, 2015: 76. excavation area, 9.IV.1986 (NHMW); Limassol, Identification Sheraton hotel, 1 ♀, 5.X.1988 (NHMW). M etzner, 1999: 95, figs 60a - h. NICOSIA: Agridi, near Dali, 1 ♂, in house, IV.2009, 1 ♂, in garden, V.2011 (all CDM); Dali, Citations Lymbourena Hill, 1 ♀, in field, IV.2004 (CDM); NICOSIA: Agridi , near Dali (Logunov, 2015).

Dali, 13, Agios Andronikos, 1 ♀, in garden, N ew records V.2007, 1 ♂ 1 ♀, in garden, IV.2008, 1 ♂ 4 ♀♀, in garden, V.2008 (all CD M); Geri, 1 ♀, olive NICOSIA: Agridi, near Dali, 1 ♀, in garden, grove and fields, 11.V.2016 (CMH); Lakatamia, 1 IV.2010, 1 ♂, in yard, V.2009, 1 ♂, stones on ♀, in olive grove, 1.I. - 1.X. 2016 (CMH); river bank, 29.IV.2013, 1 ♂, in garden, VIII.2010, Pallouriotissa, 1 ♀, in building, V.2006 (CDM). 1 ♂, VII.2012 (all CDM). PAPHOS : Akamas Peninsula, Lara Bay, 1 ♂, C. LARNACA : Stavrovouni Monastery, 350m, 1 ♂, on bark of Pinus trees, 28.IV.2010 (CRB). Saure leg., 19.VI.2013 (CKHK); Kidasi N., along

Dia rizos River, 300m, 1 ♀, beaten from shrubs, Distribution : Greece, Turkey. New to Cyprus. 21.V.2011 (CRB). Pseudomogrus gavdos (Logunov & Marusik, Distribution : Cosmopolitan. New to Cyprus. 2003) Identification Pseudeuophrys obsoleta (Simon, 1868) Logunov & Marusik, 2003: 43, figs 147 - 152. Pseudeuophrys obsoleta ; Logunov, 2015: 75. Identification Citations Metzner, 1999: 54, figs 19a - h. None.

Cit ations New records NICOSIA: Near Cedar Valley (Logunov, 2015). LARNACA: Larnaca salt lake, 1 ♀, litter in salt marsh, 27.IV.2010 (CRB). New records LIMASSOL: Akrotiri Peninsula, Turtle Beach, 2 NICOSIA: between Cedar Valley and Stavros tis ♀♀, beaten from vegetation on sandy beach, Psokas, 3 ♂♂, under stones in P. nigra forest, 15.III.2018 (CKHK). 29.IV.2007 (CAR - S), 1 ♀ (CRS); Cedar Valley Forest, 1 ♀, under stones, 29.IV.2007 (CAR - S) 1 ♂ (CRS); Cedar Valley, 1150m, 3 ♂♂, 21.III.2018, Distributio n : Canary Islands, Sardinia, Algeria, forest road, in gravel (CKHK). Crete. New to Cyprus.

Distribution : Palaearctic. Salticus noordami Metzner, 1999 Salticus noordami ; Logunov, 2009: 915; Pseudicius picaceus (Simon, 1868) Logunov, 2015: 78. Identification Metzner, 1999: 93, figs 58a - g. Identification Metzner, 1999: 115, figs 79a - f; Logunov, 2009: Citations 915, figs 36 - 42. None. Citations NICOSIA: Lympia Reservoir (Logunov, 2015).

Spiders of Cyprus 116

LIMASSOL: Troodos Mts., Panagia (Logunov, (MMUE); Akrotiri Peninsula, salt lake, 1 ♀, 2015); Pano Platres (Logunov, 2015). beaten from Salicornia , 15.III.2018 (CKHK). PAPHOS : Akamas Peninsula, near Agios Minas PAPH OS: Akamas Peninsula, Smigies, 352m, 1 (Logunov, 2015); Coral bay (Logunov, 2009). ♀, Maquis, 12.III.2018 (CKHK); Mavrokolympos Reservoir, 1 ♀, 13.III.2018 (CKHK); Paphos, Bay New records at hotel “Paphian Bay”, 1 ♀, beating vegetation FAMAGUSTA : Kantara E., 610m, 3 ♀♀, beaten near beach, 4.IV.1987 (CJB). from shrubs and stones in open Pinus forest, 25.V.2011 (CRB); Gialousa E ., Agios Thyrsos, Distribution : Circum - mediterranean. 20m, 1 ♀, beaten from shrubs, 16.V.20111 S ibianor aurocinctus (Ohlert, 1865) (CRB). Identification: https://araneae.nmbe.ch/ . LARNACA: Meneou, 1 ♂, salt marsh & fields, 8.IV. - 15.IX.2016 (CMH). Citations LIMASSOL: Alassa SE, near r iver Limnatis, 3 None. ♂♂, stones at base of limestone rock face of gorge (CAR - S); Armenochori, 1 ♀, under stones New records of a dry stone wall, 14.IV.1986 (NHMW); Mount NICOSIA: Lympia Reservoir, Lympia, 1 ♂, under Olympos (Chionistra), just below summit, 1 ♂, in stones below dam wall, 9.V.2013 (CDM). open Pinus nigra forest, 27.IV.2007 (CRS); Pano Platr es, 1 ♂, on stone wall, 6.V.2007 (CAR - S). Distribution : Palaearctic. New to Cyprus. NICOSIA: Agridi, near Dali, 1 ♂, on stone in garden, 4.IV.2012 (CDM); Dali, 13, Agios Synageles dalmaticus (Keyserling, 1863) Andronikos, 1 ♂, in garden, IV.2007 (CDM); Germasogeia, 1 ♀, on house at the dam, Identification 10.IV.1986 (NHMW); Lympia Reservoir, Metzner, 1999: 38, figs 5a - g. Lympia , 1 ♀, on concrete duct, 9.V.2013 (CAR - S); Nikitari S., Panagia Asinou, 1 ♀, on stone Citations wall, 9.V.2007 (CAR - S); Xerargaka S, Orkontas, None. 1 ♂, riverside vegetation, 26.III.2016 (CMH). PAPHOS: Akamas Peninsula, Agios Minas, 1 ♀, New records in dense Pinus litter, 2.V.200 7 (CRS); near FAMAGUSTA : Agios Sergios , 30m, 1 ♂ 2 ♀♀, Avakas Gorge, 1 ♀, beaten from junipers on beaten in Pinus plantation, 1 5.V.2011 CRB); rocky hillside, 14.IV.2017 (CAR - S); Cape Greco, Ayia Napa, 35m, 1 ♂, stones in Mavrokolympos Reservoir, 1 ♂, 13.III.2018 (CKHK). phrygana, 27.IV.2010 (CRB), 1 ♀, under stone in phrygana, 3.V.2013 (CDM); Gialousa E ., Agios Distribution Thyrsos, 20m, 1 ♀, beaten from shrubs, Lesbos, Turkey, Caucasus, Israel, Iran, Cyprus. 16.V.20111 (CRB); Gialousa E ., Agia Triada, 50m, 1 ♀, herbs in r uins, 18.III.2011 (CRB); Salticus propinquus Lucas, 1846 Kantara E., 610m, 2 ♀♀, beaten from shrubs in Saltic us propinquus ; Logunov, 2015: 78. open Pinus forest, 25.V.2011 (CRB); Salamis Identification ruins, 50m,1 ♀, beaten from low trees, 15.V.2011 Metzner, 1999: 115, figs 81a - i. (CRB). KERYNEIA : near Agios Georgios, under stones, Citations 1 subadult ♂, 27.II.2011, H. Henderick x leg. LIMASSOL: Pissouri (Logunov, 2015). (CJB).

LIMASSOL: Akrotiri Peninsula, near British New records FAMAGUSTA : Salamis ruins, 30m, 1 ♂, stones Airbase, 1 ♀, swept in pine wood, 4.V.2007 in ruins, 17.III.2011 (CRB). (CRS); Mount Olympos (Chionistra), just below LARNACA: Agios Minas, 1 ♂, under stone in summit, 1 ♂, in open Pinus nigra forest, pine wood, 20.III.1998, coll. J. Murphy (MMUE). 28.IV.2007 (CRS). LIMASSOL: Akrotiri Peninsula, 1 ♀, margin of NICOSIA : Agridi, near Dali, 1 ♀, in garden, central salt lake, 7.V.1972, coll. A. Stubbs IV .2004, 2 ♂♂, in house, IV.2009, 1 ♂, in garden,

Spiders of Cyprus 117

V.2011, 1 ♂, in garden, V.2012 (CDM); LARNACA : Larnaca salt lake, 5m, 1 ♂, stones Athalassa National Forest Park, Nicosia, 1 ♂, and litter in salt marsh, 27.IV.2010 (CRB). under stones in pine plantation, 2.V.2013; Dali, 1 ♂, on outside wall of house, 5.V.2013 (CAR - S); Distribution : Palearctic. New to Cyprus. Dali, 13, Agios Andronikos , 1 ♂, in garden, V.2007, 1 ♀, in garden, III.2008 (CDM); Kampos Thyene imperialis (Rossi, 1846)

NE, Xeros River, 550m, 1 ♂, beaten from shrubs, Identification 20.V.2011 (CRB); Kapedes S., 600m, 1 ♂, herbs Metzner, 1999: 132, figs 97a - l. along rivulet, 5.V.2010 (CRB); Lefka NW, 120m, 1 ♂, beaten from shrubs in fruit plantation, Citations 18.V. 2011 (CRB); Lympia Reservoir, Lympia, 1 Non e. ♂, on concrete duct, 9.V.2013 (CAR - S), 1 ♀, New records (CDM); Nikitari S., Panagia Asinou, 1 juvenile, FAMAGUSTA : Salamis ruins, 50m, 1 ♂, beaten under stones in disturbed phrygana, 9.V.2007 from low trees, 15.V.2011 (CRB). (CAR - S); Strovolos, 1 ♂, around houses, LARNACA : Kornos Picnic site, 350m, 1 ♀, 1.V.2016 (CMH). beaten in Pinus forest, 28.V.2011 (CRB). PAPHOS : Evretou Reser voir, 1 ♂, under stones NICOSIA: Agridi, near Dali, 1 ♂, in garden, and in grass tussocks, 13.IV.2017 (CAR - S); IV.2012 (CDM); Dali, 13, Agio s Andronikos, 2 Kelefos bridge, dam on Diarizos River, 550m, 1 ♂♂, in garden, IV.2007 (CDM). subadult ♀, beaten from shrubs, 21.V.2011 LIMASSOL: Akrotiri Peninsula, near British (CRB). Airbase, 1 ♂, swept in pine wood, 4.V.2007

(CAR - S); Akrotiri Peninsula, Fassouri reedbeds, 1 Distribution : Mediterranean. New to Cyprus. ♂, swept from grass and reeds, 17.IV.2017 (CAR - Synageles hilarulus (C. L. Koch, 1846) S). Synageles hilarulus ; Logunov, 2015: 81. Distribution : Ol d World. New to Cyprus. Identification Metzner, 1999: 38, figs 5a - g. Citations Family Scytodidae LIMASSOL: Akrotiri Peninsula, Fasouri reed beds (Logunov, 2015). Scytodes thoracica (Latreille, 1802)

New records Identification : https://araneae.nmbe.ch/ . LIMASSOL: Akrotiri Peninsula, Fasouri reed beds, 1 ♀, in reed l itter, 27.IV.2007 (CAR - S); 2 Citations ♀♀ (CRS). None.

Distribution : Palaearctic. New records FAMAGUSTA : Kantara E., 650m, 1 ♀, pitfalls in Talavera aequipes (O. Pickard - Cambridge, Pinus forest, XI.2010 - III.2011 (CRB); Komi 1871) Kebir , 150m, 1 ♂ 2 ♀♀, stones in Pinus forest, 18.III.2011 (CRB); Prastio S ., Agios Andronikos, Identification: h ttps://araneae.nmbe.ch/ . 30m, 2 ♀♀, under stones in Eucalyptus forest, 27.V.2011 (CRB). Citations KERYNEIA: Klepini , 300m, 1 ♂, und er stones in None. open Pinus forest, 23.XI.2010 (CRB); Larnakas tis Lapithou, 2 ♀♀, stones in grassland, 17.I.2016 (CMH); Pentadactylos Range, Ftericha S ., 800m, 1 ♀, under stones in grassland, 17.V.2011 (CRB). LARNACA : Hala Sultan Tekke mosque, 1 ♂ 2 New reco rds ♀♀, fro m the area of the mosque and the surrounding park, 20.IV.1986 (NHMW); Spiders of Cyprus 118

Stavrovouni Monastery, 350m, 1 subdult ♀, under New records bark of Pinus trees, 28.IV.2010, 28.IV.2010 NICOSIA: Agridi, near Dali, 1 ♂ 1 ♀, in house, (CRB). 28.VIII.2010 (CDM); Strovolos, 1 subadult ♀, LIMASSOL: Akrotiri Peninsula N., 5m, 1 ♀, around houses, 1.V.2016 (CRB). litter and beating in Eucalyptus forest, 29 .IV.2010 (CJVK); Alassa SE, 1 ♂, near river, under stones Distribution: Canary Isles, Spain to Myanmar. New to Cyprus. at base of limestone rock face of gorge (CAR - S);

Amathus, 1 ♂, under stones in dry valley, Scytodes velutina Heineken & Lowe, 1832 25.IV.1986 , 1 ♀, under stones on plants, 6.IV.1986, 1 ♀ 2 juveniles, 9.IV.1986 (NHMW); Identification : https://araneae.nmbe.ch/ Armenochori, 1 ♀, un der stones of a dry stone wall, 14.IV.1986 (NHMW); Fasoula, 1 ♂ 1 ♀ 1 Citations juvenile, olive garden with dry stone walls, None.

16.IV.1986 (NHMW); Germasogeia, 1 ♂ 2 ♀♀ 4 New records juveniles, on house at the dam, 10.IV.1986 FAMAGUSTA : Agios Sergios , 30m, 1 ♂ 3 ♀♀, (NHMW); Kolossi, N of the castle, 5 ♂♂, under pitfalls in Pinus plantation, 15 - 27.V.2011 stones, 1 1.IV.1986 (NHMW); Mount Olympos (CRB); Gialousa, Agios Thyrsos, 20m, 1 ♂ 1 ♀, (Chionistra), just below summit, 1 ♀, 2 juveniles, litter under fig tree, 18.III.2011 (CRB); Kantara under large rocks in open Pinus nigra forest, E., 610m, 1 ♂, under stones in open Pinus fore st, 28.IV.2007 (CRS); Platres, 1 ♀, on walls, 25.V.2011 (CRB), 1 ♂, pitfalls at same site, 22.IV.1986 (NHMW). XI.2010 - III.2011 (CRB); Paralimni lake, 65m, 7 NICOSIA: Limnitis, 1 ♀, Pinus brutia forest, ♀♀, litter along dry lake, 21.XI.2010 (CRB); 13.V.2016 (CMH); Ma rathasa Dam, 3 ♂♂ 2 ♀♀, Protaras, Profitis Elias, 1 ♀, 2005 (CDM); pitfalls in riverside vegetation, 7.V. - 10.VII.2016 Salamis S., Silver Beach, 5m, 1 ♂ 1 ♀, pitfalls in (CMH); Nikitari S., Panagia Asinou, 1juvenile, salt marsh, 15 - 27 .V.2011 (CRB). under stones in disturbed phrygana, 9.V.2007 KERYNEIA : Klepini , 300m, 1 ♂, under stones in (CAR - S). open Pinus forest, 23.XI.2010 (CRB); Saint PAPHOS : Akamas Peninsula, 1 ♀, in coastal Hilarion Castle, 610m, 1 ♂ 2 ♀♀, pitfalls in Pinus grassland, 22.III.1998, coll. J. M urphy (MMUE); forest, XI.2010 - III.2011 (CRB). Akamas Peninsula, Agios Minas, 2 juveniles, under stones in meadow, 2.V.2007 (CAR - S); Kato LARNACA : Larnaca salt lake, 10m, 1 ♂, pitfalls Paphos, 1 ♀, seevin hay, 4.IV.1997 (CJB); around lake, III . - V.2011 (CRB); road to Kelefos bridge, 465m, deciduous wood, 1 ♂, Stavrovouni Monastery, 1 ♂, pitfalls in open 16.III.2018 (CKHK); Kinousa SE, 400m, 1 ♀, Pinus forest, III. - V.2011 (CRB). under stones and litter in Pinus forest, 1.V.2010 NICOSIA: Agioi Trimithias, 2 ♀♀, olive grove (CRB); Makounta, 1 ♀, under stones in scrub, and fields, 22.I.2016 (CMH); Agridi, near Dali, 1 19.III.1998, coll. J. Murphy (MMUE); Paphos ♂, in house, VIII.2009, 1 ♀, in house, IV.2012, 1 NW, 1 ♀, under stones at the edge of a large sand pit, 24.IV.1986 (NHMW); Souskiou,1 ♂ 3 ♀♀, ♀, in house, VII.2012 (CDM); Athalassa National 16.III.2013 (CDM). Forest Park, 1 ♀, Pinus brutia forest, 21.II.2016 (CMH); Kapedes, 1 ♀, pitfalls in Pinus brutia Distribution: Holarctic , Pacific. New to Cyprus. forest, 17.I. - 23.IV.2016 (CMH); Lymbourena Hill, Dali, 1 ♀, in field, IV.2004 (CDM); Lympia Scytodes univittata Simon, 1882 Reservoir, Lympia, 1 ♀, in rushes below dam

wall, 9.V.2013 (CDM); Marathasa Dam, 1 ♂, Identification Carrillo & Ferrández, 2014: 54, figs 1 - 6. pitfalls in riverside vegetation, 7.V. - 10.VII.2016 (CMH). PAPHOS : Akamas Peninsula, Cape Drepanum, 2 Citations ♀♀, 31.III.1997 (CJB); Akamas Peninsula, Avgas None. E stuary, 2 ♀♀, 31.III.1997 (CJB); Kato Paphos archaeological site, 1 ♀, under stones in grass,

Spiders of Cyprus 119

10.V.2007 (CAR - S); Mavrokolympos Reservoir, None. Pano Archimandrita, 1 subadult ♀, in cave, 29.VII.1998, H. Henderickx leg. (CJB); NE New records Potima, 150m, 1 ♀, under stones along lake, 22.V. NICOSIA: Xerargaka S, Orkontas, 1 ♀, riverside 2011 (CRB); Pegeia, P ikni Forest, 366m, 1 ♀, vegetation, 26.III.2016 (CMH). pine forest under bark, 11.III.2018 (CKHK). LIMASSOL: Pedoulas, 960m, 1 ♂, 1 ♀, road embankm ent, 18.IV.2017, S. Huber leg. (CSH). Distribution: Mediterranean, Cape Verde Is., PAPHOS : Drouseia S., Panagia Chrysospiliotissa, Seychelles. New to Cyprus. 580m, 1 ♀, night catch in small cave, 22.IV.2017, S. Huber leg. (CSH); Neo Chorio, 1 ♂, under stone, 17.III.1998, coll. J. Murphy (MMUE); Kato Family Segestriidae Akourdaleia, 1 ♀, VII.200 8 (CDM); Kelefos bridge, 465m, deciduous wood, 2 ♀♀ 3 juveniles, under bark, 16.III.2018 (CKHK); Paphos, 1 ♀, on Ariadna cyprusensis Wunderlich, 2011 Ariadna cyprusensis Wunderlich, 2011: 1 87. terrace, VI.1998 (CJB); Pegeia, 100m, 1 ♀, under bark of fruit tree, 22.V.2011 (CRB), Pegeia, 268m, 1 ♂, on terrace of building, 21.III.2018 Identification Wunderlich, 2011:187, figs 51. (C KHK); Pomos, 1 ♀, 31.VIII.1998, in sea cave, H. Henderickx leg. (CJB). Citations LIMASSOL : Pissouri beach (Wunderlich, 2011). Distribution: Europe to Georgia, N. Africa. New PAPHOS : Agios Georgios, Cape Drepano to Cyprus. (Wunderlich, 2011); Akamas Peninsula, Fontana Amorosa (Wunderlich, 2011). Segestria senoculata (Linnaeus, 1758 )

New records Identification : https://arane ae.nmbe.ch/ . PAPH OS : Pegeia, Agios Georgios, rocky shore, 1 ♀ 1 juvenile, 13.III.2018 (CKHK). Citations None.

Distribution: Cyprus, probably Kos. Only known from coastal localities. New records NICOSIA: Agridi, near Dali, 1 ♀, in garden, IV.2011 (CDM). Segestria bavarica C. L. Koch, 1843

Identification : https://araneae.nmbe.ch/ . Distribution: Palaearctic. New to Cyprus.

Citations None. Family Sicariidae New records L IMASSOL : Troodos, Mount Olympos Loxosceles rufescens (Dufour, 1820) (Chionistra), 1900m, 1 juvenile, under stones in Loxosceles rufescens ; Log unov , 2013: 5. Pinus forest, 23.V.2011 (CRB). NICOSIA: Xerargaka S, Orkontas, 1 ♀, riverside Identification vegetation, 26.III.2016 (CMH). Simon 1904: 75, fig. 126. PAPHOS: Near Kidasi village, Diarizos Valley, 1 ♀, under bark on large plane tree, 1.V.2013 Citations (CAR - S); Neo Chorio, 1 ♀, stones in grassland PAPHOS : Avakas Gorge (Logunov, 2013). (CMH). New records Distribution: Europe to Caucasus. New to FAMAGUSTA : Cape Apostolos Andreas, 1 ♀, Cyprus. Juniperus phoenicea forest 30.III.2016 (CMH); Salamis ruins, 30m, 1 subadult ♀, under stones in Segestria florentina (Rossi, 1790) ruins, 17.III.2011 (CRB), 2 ♀♀, 15.V.2011 (CRB); Davlos W., 25m, under stones in Identification : https://araneae.nmbe.ch/ . phrygana near the bea ch, 25.V.2011 (CRB); Citations Rizokarpaso , 1 ♀, Juniperus phoenicea forest, Spiders of Cyprus 120

7.V.2016 (CMH); Agios Sergios , 30m, 1 subadult PAPHOS : Evretou Reservoir, 350m, 1 ♀, under ♂, pitfalls in Pinus pl antation, 15 - 27.V.2011 stones in grassland around lake, 1.V.2010 (CRB); (CRB); Cape Greco, 35m, 1 subadult ♀, under Kato Akourdaleia, 1 ♂, in garden, VIII.2005 stones in phrygana, 21.XI.2010 (CRB); Paralimni (CDM); Kato Paphos, ar chaeological site, 1 lake, 65 m,1 subadult ♀, in litter along dry lake, juvenile, under stones in grass, 10.V.2007 (CAR - 21.XI.2010 (CRB). S); Kinousa E., Agios Merkourios, 350m, 1 ♂, KERYNEIA: near Agios Georgios, 1 ♀, under under stones in Pinus forest, 1.V.2010 (CRB); stones, 27.II.2011, H. Hen derickx leg. (CJB); Mavrokolympos Reservoir, 1 ♀, 13.III.2 018 Kazafani , 150m, 1 ♀, under stones in olive grove, (CKHK); Pano Archimandrita E stuary, 1 ♀, 23.XI.2010 (CRB); Klepini , 300m, 2 ♀♀, under 9.III.1999, H . Henderickx leg. (CJB); Paphos stones in open Pinus forest, 23.XI.2010 (CRB); NW, 1 ♂ 1 ♀ 1 juvenile, under stones, 24.IV.1986 Pentadactylos Range, Saint Hilarion Castle W, (NHMW); Pegeia, Coral Bay, 1 ♀, under stones, 600m, 2 ♀♀, under stones in Pinus forest 31.III.1997 (CJB); Pegeia, Pikni Forest, 346m, 1 17.V.20 11 (CRB); Vasileia S ., 200m, 2 juveniles, ♀, in former carob plantation, 23.III.2018 dry riverbed in olive grove, 25.XI.2010 (CRB). (CKHK); Pomos E., 140m, 1 ♀, road to reservoir, LARNACA : Achna Reservoir, 50m, 1 ♀, under 23.IV.2017, S. Huber leg. (CSH); Pomos, 1 ♀, in stones bordering lake, 16.V.2011 (CRB); sea cave, H. Henderickx leg. (CJB); 31.VII.1998; Delikipos, 1 ♀, under stone in grass around small near Souskiou, 1 ♂, under stones in dry grassland, dam, 18.IV.2017 (CAR - S); L arnaca salt lake, 5m, 1.V.2013 (CAR - S). 2 juveniles, under stones in salt marsh, Distribution: Cosmopolitan. 20.XI.2010 (CRB), 1 ♀, pitfalls at same site (CRB); Ora, 1 ♀, in maquis, 28.XII.2016 (CMH); Stavrovouni Monastery, pitfalls in open Pinus Family Sparassidae forest, III. - V.2011 (CRB). LIMASSOL : Agios Tychonas, 1 ♀, 19.IV.1986 Eusparassus wa lckenaeri (Audouin, 1826) (NHMW); Akrotiri Peninsula, near Agios Eusparassus tersa ; Strand 1907: 397. Nikolaos Monastery, 25m, 1 ♀, in grassy ditch, Eusparassus walckenaeri ; Moradmand & Jäger, 23.III.2011 (CRB); Akrotiri Peninsula E., 5m, 1 2012: 2455. ♀, under stones and litter in salt marsh, 29.IV.2010 (CRB); Akrotiri Peninsula, edge of Identification central salt pan, 1 ♀, under stones and debris in Levy, 1989: 132, figs 3 - 18; Moradmand & Jäger Salicornia , 30.IV.2013 (CDM); Amathus, 1 ♀ 2 2012: 2455, figs 5 A - G. subadult ♂♂, under stones in dry valley, 25.IV.1986, 1 ♀ 8 juvenile s, 6.IV.1986, 1 ♀ 1 Citations juvenile, 9.IV.1986 (NHMW); Armenochori, 2 Without precise locality (Strand 1907). ♀♀ 6 juveniles, under stones of a dry stone wall, FAMAGUSTA: Protaras; Agios Ilias Village 14.I V.1986 (NHMW); Erimi, 1 ♀, under stones in (both Moradmand & Jäger, 2012 ). dry riverbed, 18.IV.1986 (NHMW); Fasoula, 3 juveniles, olive garden with dry stone walls, New records 16.IV.1986 (NHMW); Malia - Dora road, 650m, 2 FAMAGUSTA : Vokolida S., Cape Elaia, 10m, 1 ♀♀, under stones in grassland, 21.III.2011 subadult ♀, under stones and on shrubs near the (CRB); Mount Olympos (Chionistra), j ust below summit, 1 ♀, under large rocks in open Pinus beach, 26.V.2011 (CRB). nigra forest, 28.IV.2007 (CRS); Pissouri, 2 ♀♀, LARNACA: Dekeleia Military Ba se, 1 ♂, in cave, 3.VII.2016 (CMH). 10.VI.1958, M. Connelly leg. (BMNH); Dekeleia, NICOSIA: Agia Marina, 1 ♂, Pinus brutia forest, Royal Army Medical College, Department of 7.III.2016 (CMH); 13, Agios Andronikos, Dali 1 Tropical Medicine, 1 ♀, 18.VII.1959, Baldry leg. ♂ 2 ♀♀, in garden, V.2008 (CD M); A gridi, near (BMNH). Dali, 1 ♀, in eucalyptus litter in garden, NICOSIA : Kampos NE, Xeros River, 550m, 1 ♀, 29.IV.2013; 1 ♂, in garden, IV.2011 (CDM); Athalassa National Forest Park, 1 ♂ 2 ♀♀, under stones in riverbed, 20.V.2011 (CRB); Limnitis, 1 stones in pine plantation, 8.V.2013 (CAR - S); ♂, Pin us brutia forest, 13.V.2016 (CMH). Marathasa Dam, 4 ♂ 1 ♀, pitfalls in riverside vegetation, 7. V. - 10.VII.2016 (CMH); Pedoulas, Distribution: Eastern M editerranean to 960m, 1 ♀, road embankment, 18.IV.2017, S. Afghanistan. Huber leg. (CSH).

Spiders of Cyprus 121

Micrommata formosa Pavesi, 1878 Citatio ns Agridi near Dali (Jäger et al. 2011). Identification Levy, 1989: 165, figs 100 - 114. New records NICOSIA: Agridi, near Dali, 1 ♂, in house, 2014 Citations (CDM). None. Distribution: Israel (Levy, 1989), Egypt (O. New records Pickard - Cambridge, 1876; Denis, 1947), Cyprus. FAMAGUSTA : Prastio S., Lake Kouk lion, 1 ♀, fallow land with Salicornia/Suaeda , 20.II.2011, D. W. Wrase leg. (CKHK). Family Tetragnathidae LARNACA: Ora, 1 ♀, in maquis, 28.XII. 2016 (CMH). Metelli na mengei (Blackwall, 1869) NICOSIA: Agridi, near Dali, 6 ♂♂, in house, 2014 (CDM); Geri, 1 ♂, in olive grove, Identification : https://araneae.nmbe.ch/ . 11.V.2016 (CMH). Citations Distribution: Algeria to Israel. New to Cyprus. None.

Micrommata ligurina (C. L. Koch, 1845) New records LIMASSOL : Pano Platres, track near Caledonia Identification Waterfall, 1 ♂ 1 ♀, swept from herbs and shrubs Levy, 1989: 168, figs 115 - 125. i n rocky stream valley, 26.IV.2007 (CAR - S). Citations None. Distribution: Europe to Georgia. New to Cyprus.

New records Tetragnatha demissa L. Koch, 1872 LARNAKA: Meneou, 4 ♀♀, 8.II.1957, G. P. Georgiou (BMNH). Identification NICOSIA: Makedonitissa , 1 ♂, 13.II.1957, G. P. Tetragnatha demissa ; Saaristo, 2010 : 236 , figs 36: Georgiou (BMNH). 29, 33.

Distribution: Mediterranean to Central Asia. New records New to Cyprus. LARNACA: Meneou, 3 ♂♂ 3 ♀♀, salt marsh and fields, 8.IV. - 15.IX.2016 (CMH). Micrommata virescens (Clerck, 1757) PAPHOS : Asprokremmos Dam, N. Kouklia, 150m, 2 ♂♂ 2 ♀♀, on herbs along lake, Identification : https://araneae.nmbe.ch/ . 29.IV.2010 (CRB); idem, 5 ♂♂ 11 ♀♀, herbs along lake, 23.V.2011 ( CRB). Citations None. Distribution: South Africa, Tanzania, Seychelles, Australia. New to Cyprus. New records NICOSIA: Dali, Lymbourena Hill, 2 ♀♀, in field, V.2004 (CDM). Tetragnatha intermedia Kulczyński, 1891

Distribution: Palaearctic. New to Cyprus. Identification : https://araneae.nmbe.ch/ .

Olios suavis (O. Pickard - Cambridge, 1876) Citations None.

Identification Ne w records Levy, 1989: 152, figs 71 - 81, 115 - 125. NICOSIA: Dali, Idalion Archaeological Site, 1 ♂, in phrygana, 30.IV.2013 (CDM).

Spiders of Cyprus 12 2

PAPHOS : Evretou Reservoir, 350m, 2 ♂♂ 2 ♀♀, Wunderlich, 1995: 640, figs 1 - 10. shrubs and grassland around lake, 1.V.2010 (CRB). Citations None. Distribution Palaearctic. New to Cyprus. New records LIMASSOL: Episkopi, Fassouri reed beds, 1 ♂, Tetragnatha ni tens (Audouin, 1826) on vege tation at the edge of the reed bed, 30.IV.2013 (CRS). Identification : https://araneae.nmbe.ch/ Comments Citations A male of this species was collected by Rowley None. Snazell in 2013. It will be redescribed by him in a separate paper. New records LARNACA: Meneou, 2 ♂♂ 1 ♀, salt marsh and Distribution: Until now, this species was only fields, 8.IV. - 15.IX.2016 (CMH). known from Israel. It is new to Europe and NICOSIA: Geri, 2 ♂♂ 1 ♀, i n olive grove, Cyprus.

11.V.2016 (CMH).

PAPHOS : Avakas Gorge, 150m, 1 ♂, on rocks in Family Theraphosidae gorge, 22.V.2011 (CRB); Ktima, 1 ♂, G.

Mavromoustakis leg., without date (BMNH). Chaetopelma karlamani Vollmer, 1997

Distribution : Cosmotropical, Mediterranean. Chaetopelma karlamani Vollmer, 1997: 5; Peters, New to Cyprus. 1998: 24; Peters, 2000: 87; Guadanucci & Gallon, 2008: 39, figs 11 - 18; Gallon et al . 2012: 138. Tetragnatha obtusa C. L. K och, 1837 Identification Identification : https://araneae.nmbe.ch/ . Vollmer, 1997: 5, figs 3 - 13.

Citations Citations None. KERYNEIA : Lapithos (Vollmer, 1997; type locality); Trapeza Mountain (Vollmer, 1997); New records Karmi (Vollmer, 1997). NICOSIA: Marathassa Dam, 550m, 1 ♂ 1 ♀, LARNACA : Without precise locality stones and beaten from shrubs around lake, (Guadanucci & Gallon, 2008). 20.V.2011 (CRB); Nikitari S., Panagia Asinou, 1 NICOSIA: Withou t precise locality (Guadanucci ♀, in disturbed phrygana, 9.V.2007 (CAR - S). & Gallon, 2008). PAPHOS : Akamas Peninsula, Neo Chorio, 2 ♂♂, swept from phrygrana and grass, 2.V.2007 (CAR - New records S); Asprokremmos Dam, N. Kouklia, 150m, 3 ♂♂ FAMAGUSTA: Kantara E., 1 ♂, 650m, pitfalls in 1 ♀, herbs along lake, 23.V.2011 (CRB); Avakas Pinus forest, III. - V.2011 (CRB) idem, 1 juv., gorge, 1 ♂ 1 ♀, 22.III.2018 (CKHK); Evretou stones in ruins, 19.III.2011 (CRB). Reservoir, Stavros tis Psokas river, 1 ♂, KERYNEIA : Vasileia S., 200m, 2 juveniles, 20.III.2018 (CKHK); Kelefos bridge, dam on stones in dry r iverbed in olive grove, 25.XI.2010 Diarizos River, 550m, 1 ♂ 2 ♀♀, beaten from (CRB); Saint Hilarion Castle, 2 km W., 610m, 1 shrubs, 21.V.2011 (CRB); Kelefos bridge, 465m, juv., stones in open Pinus forest, 20.III.2011 deciduous wood, 1 ♂ 1 ♀, 16.III.2018 (CKHK). (CRB). NICOSIA: Kampos N. (35°04'01"N 32°44'15"E), Distribution : Palaeartic. New to Cyprus. 500m, 2 juv., in leaf litter, 23.IV.2017 (CSH); Kampos N. (35°04'19"N 32°4 4'36"E), 470m, 2 Zygiometella perlongipes (O. Pickard - juv., unde r stones, 20.IV.2017 (CSH); Oikos Cambridge, 1872) (N34°59’58” E 32°59’50”), 660m, 1 ♂, sitting at bottom of road emban kment, night catch, Identification 19.IV.2017 (CSH); Oikos (35°00'02"N

Spiders of Cyprus 123

32°49'58"E), 700m, 1 juv., near small creek, 1 ♂, some juv., in rock shelter, 21.IV.2017 19.IV.2017 (CSH). (CSH); Latsi W., Baths of Aphrodite, 4 juv., under large stones near the beach, 15.IV.2017 Dis tribution: Only known from Cyprus, where it (CAR - S). occurs at higher altitudes. Distribution: Egypt, Turkey, Cypr us, Lebanon, Chaetopelma olivaceum (C. L. Koch, 1841) Palestine, Israel, Jordan, Syria and Saudi Arabia. In Cyprus found mainly in the Paphos province. Mygale olivacea C. L. Koch, 1841: 34, fig. 712 (descr. imm. ♀). Ischnocolus gracilis Ausserer, 1871: 187 (descr. Family Theridiidae imm. ♀); Smit h 1990: 127. Chaetopelma olivacea ; Strand 1907: 393. Anelosimus pulchellus (Walckenaer, 1802) Chaetopelma anatolica Schmidt & Smith, 1995: 1, figs 1 - 9. Identification : https://a raneae.nmbe.ch/ . Chaetopelma aegypt ac i um (sic) ; Smith, 1990: 110 (citation). Chaetopelma gracile ; Vollmer, 1997: 5; Peters, Citations 1998: 20; Peters, 2000: 86. None. Chaetopelma olivaceum ; Guadanucci & Gallon, 2008: 39, figs 11 - 18; Gallon et al ., 2012: 136. New records FAMAGUSTA: Gialousa E., Agios Thyrsos, Identification 20m, 1 subadult ♀, beaten from shrubs, Vollmer, 1997: 5, figs 3 - 13. 16.V.20111 (CRB). Citations KERYNEIA: Panagia Apsinthiotissa Monastery Without precise locality (Ausserer, 1871, type ruins, 350m, 1 ♀, on low herbs in clearing in locality of C. gracile ; Strand, 1907). Pinus fo rest, 24.V.2011 (CRB); Pentadactylos KERYNEIA : Lapithos (Vollmer, 1997); Saint Range, Saint Hilarion Castle, 600m, 1 subadult ♀, Hilarion (Gallon et al., 2012). beaten from shrubs, 17.V.2011 (CRB). LARNACA : Ayia Napa (Schmidt & Smith, 1995, LIMASSOL: Pano Platres, 1050m, 1 ♀, beaten sub C. karlamani , misidentification; Guadanucci & Gallon, 2008); Larnaca (Gallon et al ., 2012). near Mesa Potamos Waterfall s, 23.V.2011 (CRB). NICOSIA : Dali (Gallon et al., 2012); NICOSIA : Marathassa Dam, 550m, 1 subad ult ♀, Asprokremmos (Guadanucci & Gallon, 2008); beaten from shrubs around lake, 20.V.2011 Nicosia (Gallon et al ., 2012). (CRB); Oikos N., Xerargaka picnic site, 350m, 2 PAPHOS : Asprokremmos, Solea Valley (Gallon subadult ♀♀, beaten from shrubs in Pinus forest, et al ., 2012); Paphos (Smith, 1990, sub C. 20.V.2011 (CRB); aegyptiacum ).

New records Distribution : Europe to Russia, North Africa. NICOSIA: Dali, Agios Andronikos, house New to Cyprus. veranda and garden, surrounding ga rdens, 5 ♂♂ 8 ♀♀, VI, 1995 - 2009 (CDM); Kythrea, 1 ♀, G. P. Anelosimus vittatus ( C. L. Koch, 1836) Gheorgiou leg., no date (BMNH). PAPHOS : Akamas Peninsula, Cape Drepanum, 1 Identification : https://araneae.nmbe.ch/ . ♂ 1 ♀ - , rock strewn slope at sea front, 15.V.1998, H. Henderickx leg. (CJB); idem, 1 ♂ 1 Citations juv. ♀, raised in captivity (C JB); Avakas Gorge, 1 None. ♀, 16.VIII.2014, N. &. L. Wendlandt leg. (CKHK); W. Kathikas, Aspros valley, 527m, 1 ♂ New records 2 ♀♀, under stones in open pasture, 24.II.2011, NICOSIA : Kakopetria S., Platania forest station, D. W. Wrase leg. (CKHK); Latsi W., Blue 1100m, 1 ♂, beating in dense Pinus forest, Lagoon (35°03'42"N 32°20'21"E), 50m, 1 juv., 4.V.2010 (CJ VK). rock shelter, 2 1.IV.2017 (CSH); Latsi W., Aphrodite’s Bath (35°03'22"N 32°20'40"E), 60m, Distribution: Palaearctic. New to Cyprus.

Spiders of Cyprus 124

litter, 2 ♀♀, under bark, 16.III.2018 (CKHK); Argyrodes argyrodes (Walckenaer, 1841) Kinousa SE, 400m, 2 ♀♀, in litter in Pinus forest, 1.V.2010 (CJVK, CRB); Kinousa E., Agios Identification Merkou rios, 350m, 2 ♂♂ 1 ♀, under stones in Levy, 1998b: 121, figs 224 - 237. Pinus forest, 1.V.2010 (CJVK, CRB).

Citations Distribution: Mediterranean. New to Cyprus. None. Dipoena galilaea Levy & Amitai, 1981 New records FAMAGUSTA : Gialousa E., Agios Thyrsos, Identification 20m, 1 ♀, beaten from shr ubs, 16.V.20111 Dipoena galilaea Levy & Amitai , 1981: 186, figs (CRB). 37 - 42; present paper figs 18a - g .

Citations Distribution: Mediterranean to West Africa, None. Seychelles. New to Cyprus.

New records Crustulina scabripes Simon, 1881 PAPHOS : Makounta, Argaka Dam, 85m, 1 ♂ 1

♀, beating bushes, 27.IV. - 5.V.2007 (CJVK). Identification

Knoflach, 1994: 322, figs 21, 22, 25, 30, 33, 43, Distribution: Greece, Israel. New to Cyprus. 44.

Dipoena melanogaster (C. L. Koch, 1837) Citations None. Identificatio n : https://araneae.nmbe.ch/ .

New records Citations F AMAGUSTA: Cape Greco, Ayia Napa, 35m, 1 None. ♂ 4 ♀♀, stones in phrygana, 27.IV.2010 (CJVK,

CRB). New records KERYNEIA: Larnakas tis Lapithou, 1 ♂, stones NICOSIA : Kakopetria S., Platania forest station, in grassland, 17.I.2016 (CMH). LARNACA : Cape Kiti, 1 ♂ 1 ♀, under stones 1100m, 1 ♂, under stones in dense Pinus forest, below coastal cliffs, 25.IV.1982, coll. A. Stubbs 4.V.2010 (CJVK); Machairas forest near Kion ia, (MMUE); Larnaca SW., Hala Sultan Tekke 1300m, 2 ♂♂ 10 ♀♀, beaten in stony phrygana, mosque, 5m, 1 ♂, salt marsh, 15.V.2011 (CJVK); 5.V.2010 (CJVK); Marathassa Dam, 550m, 1 ♀, Stavrovouni Monastery, 350m, 1 ♀, under bark of beaten from shrubs around lake, 20.V.2011 Pinus trees, 28.IV.2010 (CRB). (CRB); Prodromos S., Trooditissa picnic site, LIMASSOL: Akrotiri Peninsula E., 5m, 2 ♂♂ 1 1450m, 1 ♂, beaten in Pinus forest, 21.V.2011 ♀, under stones and in litter in salt marsh, (CRB); Prodromos S., Trooditi ssa picnic site, 29.IV.2010 (CJVK); Armenochori, 2 ♂♂, on 1450m, 1 ♂, stones in Pinus forest, 21.V.2011 bushes, 14.IV.1986 (NHMW); Kourion (CJVK). archaeological site near Episkopi, 1 ♀, under PAPHOS : Akamas Peninsula, 1 ♀, in coastal stones, 9.V.2007 (CAR - S); Pano Platres, 1 ♂, on grassland, 22.III.1998, coll. J. Murphy (MMUE); stone wall, 6.V.2007 (CAR - S); Pano Platres E., near Avakas Gorge, 2 ♀♀, beaten from junipers Mesa Potamos Waterfall s, 950m, 1 ♀, in litter in on rocky hillside, 14.IV.2017 (CAR - S); Baths of Pinus forest, 4.V.2010 (CJVK). Aphrodite, 1 ♀, under stone in coastal scrub, NICOSIA: Cedar valley, river valley in Cedrus 18.III.1998, coll. J. Murphy (MMUE); Kelefos forest, 1120 m, 1 ♀, 21.III.2018 (CKHK); bridge, dam on Diarizos River, 550m, 2 ♀♀, beaten from shrubs, 21.V.2011 (CRB); Makounta, Limnitis River, Kato Pyrgos SE, 200m, 1 ♂, Argaka Dam, 85m, 1 ♂, beating bushes, 27.IV. - under stones in Pinus forest, 2.V.2010 (CRB) . 5.V.2007 (CJVK). PAPHOS: Akamas Peninsula, A gios Minas, 1 ♀, under stones in wet grassland, 2.V.2007 (CAR - S); Kelefos bridge, 465m, deciduous wood, 2 ♀♀, in Spiders of Cyprus 125

Distrib ution: Europe, North Africa to Azerbaijan. New to Cyprus. Identification Knoflch & Thaler, 2000: 412, figs 1 - 5, 8, 54 - 57. Enoplognatha gemina Bosmans & Van Keer, 1999 Citations None. Identification Bosmans & Van Keer, 1999: 235, figs 103 - 107. New records FAMAGUSTA : Kantara E., 650m, 1 ♂ 2 ♀♀, in Citations pitfal ls in Pinus forest, XI.2010 - III.2011 (CRB); None. Gialousa E., Agia Triada, 50m, 1 ♂, stones in

ruins, 18.III.2011 (CRB). New records FAMAGUSTA : Cape Apostolos Andreas, 2 ♀♀ , Distribution: Rhodes, Israel, Az er baijan. New to Juniperus phoenicea forest 30.III.2016 (CMH); Cyprus. Karpasia Peninsula, Cape Apostolos Andreas, 70m, 1 ♀, stones in grassland, 16.V.2011 (CJVK); Enoplognatha macrochelis Levy & Amitai, RizokarpasoW., Panagia Eleousa Monastery, 1981 30m, 1 ♀, stones in Pinus forest, 18.III.2011 Enoplognatha macrochelis ; Bosmans & Van (CRB); idem, 1 ♂, Juniperus phoenicea forest, Keer, 1999: 229, figs 83 - 87.

12.III.2016 (CMH). Identification KERYNEIA: Pa nagia Apsinthiotissa Monastery Bosmans & Van Keer, 1999: 229, figs 83 - 87. ruins, 350m, 2 ♀♀, stones in clearing in Pinus forest, 24.V.2011 (CJVK). Citations LIMASSOL : Agros, 2 ♀♀, Pinus brutia forest, LIMASSOL : Limassol (Bosmans & Van Keer, 20.V.2016 (CMH); Akrotiri Peninsula, edge of 1999); Pissouri (Bosmans & Van Keer, 1999). central s alt pan, 1 ♀, under stones and debris in PAPHOS : Akamas Peninsula (Bosmans & Van Salicornia , 27.IV.2007 (CAR - S); Kapouti , 100m, Kee r, 1999); Kouklia (Bosmans & Van Keer, 2 ♀♀, stones in grassland, 20.III.2011 (CRB); 1999); Lara (Bosmans & Van Keer, 1999); Malia - Dora road, 650m, 2 ♀♀, stones in Paphos (Bosmans & Van Keer, 1999); Pano, grassland, 21.III.2011 (CRB). Archimandrita road (Bosmans & Van Keer, NICOSIA: Lakatamia, 1 ♂ ♀, in olive grove, 1.I. - 1999); Petra tou Romiou (Bosmans & Van Keer, 1.X . 2016 (CMH). 1999). PAPHOS : Akamas Peninsula, Agios Minas, 1 ♀, in thick pine litter, 2.V.2007 (CAR - S), 1 ♀, New records 3.IV.1997 (CJB); Akamas Peninsula, Lara Bay, 7 FAMAGUSTA: Cape Greco, Ay ia Napa, 35m, 1 ♀♀, under stones, 3.IV.1997 (CJB); Evretou E., ♀, stones in phrygana, 27.IV.2010 (CRB); 200m, 2 ♀♀, stones in maquis, 1.V.2007 (CJVK); Karpasia Peninsula, Apostolos Andreas Evretou Re servoir, 180m, 6 ♀♀, stones on slope Monastery, 50m, 6 ♀♀, stones in wasteland, to lake, 1.V.2007 (CJVK); Kato Paphos, 1 ♀, under stones near hotel, 31.III.1997 (CJB); 16.V.2011 (CJVK); Komi Kebir , 150m, 1 ♀, Kelefos bridge, dam on Diarizos River, 550m, 1 stones in Pinus forest, 18.III.2011 (CRB); ♀, und er stones , 21.V.2011 (CJVK); Kouklia - Rizokarpaso W., Panagia Eleousa Mo nastery, Pissouri road, 150m, 1 ♀, stony slope nea r sea, 30m, 1 ♂, stones in Pinus forest, 18.III.2011 22.III.2011 (CRB); Mavrokolympos Reservoir, 1 (CRB); Salamis ruins, 30m, 1 ♀, stones in ruins, ♀, 13.III.2018 (CKHK); Neo Chorio, 200m, 2 17.III.2011 (CRB). ♀♀, under stones, 25.IV.2007 (CJVK); Stavros tis KERYNEIA : Panagia Apsinthiotissa Monastery Psokas S., 180m, 1 ♀, stones along rivulet, 30.IV.2007 (CJVK). ruins, 350m, 10 ♀♀, stones in clearing in Pinus forest, 24.V.2011 (CJVK, CRB). Distribution: France, Italy, the Balkans, I srael, LARNACA : Larnaca salt lake, 5m, 2 ♀♀, stones Turkey, Syria, Egypt. New to Cyprus. and litter in salt marsh, 27.IV.2010 (CJVK), 1 ♂, pitfalls at same site, III. - V.2011 (CRB); Larnaca Enoplognatha giladensis (Levy & Amitai, 1982) SW., Hala Sultan Tekke mosque, 5m, 1 ♀, stones

Spiders of Cyprus 126

in salt marsh, 16.III.2011 (CRB); Stavrovouni (CJVK); Mavrokolympos Reservoir, 1 ♀, Monastery, 350m, 2 ♀♀, under stones in Pinus 13.III.2018 (CKHK); Pegeia, Pikni Forest, 346m, forest, 28.IV.2010 (CJVK, CRB). 1 ♀, in former car ob plantation, 23.III.2018 (CKHK); Smigies, 400m, 5 ♀♀, stones in maquis, LIMASSOL: Akrotiri Peninsula, central salt pan, 25.IV.2007 (CJVK); Souskiou, 2 ♀♀, 16.III.2013 2 ♀♀, under stones, 17.IV.2017 (CAR - S), 1 ♀, (CDM). 2.IV.1997 (CJB); Akrotiri Peninsula E., 5m, 2 ♀♀, under stones in salt marsh, 29.IV.2010 Distribution (CRB); Alassa SE, near river Limnatis, 1 ♀, Greece, Turkey, Israel, Caucasus and Cyprus. stones in dry grassland, 5.V.2007 (CAR - S), 9 ♀♀, (CRS); Malia - Dora road, 650m, 2 ♀♀, stones in Enoplognatha mariae Bosmans & Van Keer, 1999 grassland, 21.III.2011 (CRB); Panagia Glossa

Monastery near Kellaki, 4 ♀♀, under stones in Identif ication nearly bare cultivated land, 12.IV.1986 (NHM W), Bosmans & Van Keer, 1999: 215, figs 24 - 29. 1 ♀, outer wall of the castle, 7.IV.1986 (NHMW); Pano Platres, near top of road to Millomeris Citations Waterfalls, 1 ♂, under stones in abandoned None. vineyard, 28.IV.2007 (CRS). New records NICOSIA: Agios Sozomenos , near Dali, 2 ♀♀, KERYNEIA : Pentadactylos Range, Lapithos S., under stones in dry, rocky phrygana, 20.IV .2017 800m, 1 ♀, stones in Pinus forest, 17.V.2011 (CAR - S); Agridi, near Dali, 1 ♀, in garden, (CJVK). 1V.2010 (CDM); Amathus, 1 ♀, on house walls, LARNACA : Achna Reservoir, 50m, 1 ♀, under 9.IV.1986 (NHMW); Armenochori, 1 ♀, on stones and litter bo rdering lake, 27.V.2011 bushes, 14.IV.1986 (NHM W); Dali, 1 ♀, in grass (CJVK). in riverbed, 12.V.2007 (CAR - S) 1 ♀, 1.IV.2012 PAPHOS : Evretou Reservoir, 350m, 1 ♀, shrubs (CDM); Dali, 13, Agios Andro n ikos, 1 ♂ 2 ♀♀, and grassland around lake, 1.V.2010 (CRB); in garden, IV. 2007 (CDM); Pallouriotissa, Evretou E., 200m, 3 ♀♀, sweeping low herbs, Nicosia. 1 ♂, in garden, V.2012 (CDM). 1.V.2007 (CJVK); Pelathousa SE., 245m, 1 ♂ 1 PAPHOS : Akamas Peninsula, between Cape ♀, sweeping grassland, 30.IV.2007 (CJVK). Drepano and Lara Bay, 2 ♀♀, 31.III. and 2.IV.1997 (CJB); Akamas Peninsula, Cape Geronnisos, 25m, 4 ♀♀, stones in maqui s, Distribution: Crete, Rhodes. New to Cyprus. 29.IV.2007 (CJVK); Akamas Peninsula, 2 ♀♀, 3.V.2007 (CRS); Akamas Peninsula, 3 ♀♀, in Enoplognatha mediterranea Levy & Amitai, coastal shrub, 22.III.1998, coll. J. Murphy 1981 (MMUE); Asprokremmos Dam, N. Kouklia, Enoplognatha mediterranea ; Bosmans & Van 150m, 1 ♀, herbs and stones along lake, Keer, 1999: 223, 29.IV.2010 (CJVK); Avakas Gorge, 50m, 2 ♀♀, l itter in citrus yard at entrance of gorge, Identification 30.IV.2010 (CRB); Avakas Gorge, 55m, 2 ♀♀, Bosmans & Van Keer, 1999: 223, figs 58 - 62. stones along rivulet, 29.IV.2007 (CJVK); Drouseia, 539m, 1 ♀, in p hrygana, 23.III.2018 Citations (CKHK); Evretou Reservoir, 180m, 1 ♀, stones PAPHOS : Akamas P eninsula (Bosmans & Van on slope to lake, 1.V.2007 (CJVK); Evr etou E., Keer, 1999). 200m, 1 ♂, stones in maquis, 1.V.2007 (CJVK); Kato Paphos, 2 ♀♀, under stones near hotel, New records 31.III.1997 (CJB); Kouklia, Sanctuary of LIMASSOL: Amathus, 2 ♀♀, under s tones on Aphrodite, 1 ♀, 8.IV.1997 (CJB); Kouklia - plants in dry valley, 6.IV.1986, 1 ♀, 7.IV.1986 Pissouri road, 150m, 3 ♀♀, stony slope near sea, (NHMW); NW . Erimi, 6 ♀♀, grass heathland 22.III.2011 (CRB); Lara, 2 5m, 10 ♀♀, stony with individual stones, 21.IV.1986 (NHMW); Juniperus maquis near the beach, 29.IV.2007 idem, 1 ♀, under stones in dry riverbed, (CJVK); Mavrokolympos Reservoir, NE Potima, 18.IV.1986 (NHMW); Fasoula, 1 ♀, olive garden 150m, 4 ♀♀, under stones along lake, 22.V.2011 with dry stone walls, 16.IV.1986 (NHMW).

Spiders of Cyprus 127

FAMAGUSTA : Agios Sergios, 30m, 1 ♂, stones Distribution in phrygana, 17.III.2011 (CRB). Turkey, Cyprus, Israel, Caucasus. LARNACA : Stavrovouni Monastery W., 350m, 1

♂ , pitfalls in open Pinus forest, XI.2010 - III.2011 Enoplognatha parathoracica Levy & Amitai, 1981 (CRB). LIMASSOL : Agios Georgios E., Palaiomylos, Identification 1250m, 2 ♀♀, stones in mixed Pinus and Quercus Bosmans & Van Keer, 1999: 220 , figs 42 - 47. forest, 4.V.2010 (CRB); Agros SW, 950m, 1 ♀, under stones in Pinus forest, 5.V.2010 (CJVK); Citations Akrotiri Penin sula, Kalathas Bay, 1 ♀, 9.IV.1997 None. (CJB); Pano Platres S., along Kryos River, New records 1050m, 1 ♀, litter in Pinus forest, 3.V.2010 LIMASSOL: Troodos square, in mixed Pinus (CRB); Pano Platres E., Mesa Potamos nigra - Juniperus foetidisssima forest, 1 ♂, Waterfall s, 950m, 4 ♀♀, in litter in Pinus forest, 23.VI.2017 (CMH). 4.V.2010 (CJVK); Troodos 3 km S. , 3 ♂♂, NI COSIA: road Pedoulas - Lemithou, 2 ♂♂ , pitfa lls in maquis, 26.IV. - 5.V.2007 (CJVK). pitfalls in Pinus nigra and P. brutia fores t, 6.IV. - NICOSIA : Limnitis River, Kato Pyrgos SE, 8.VI.2 014 (CMH ); near Platania camping site, 4 200m, 1 ♀, stones and litter in Pinus forest, ♂♂ 2 ♀♀, pitfalls in Pinus brutia fores t, 6.IV. - 2.V.2010 (CRB); Oikos N., Xerargaka picnic site, 8.VI. 2014 (CRB ). 350m, 1 ♀, beaten from shrubs and stones in Pinus forest, 20.V.2011 (CJVK). Distribution: Greece, Turkey, Israel, Azerbaijan. PAPHOS : Agia, Stavros tis Psokas S., 590m, 2 New to Cyprus. ♀♀, stones in picnic site, 30.IV.2007 (CJVK); Agios Merkourios, 250m, 1 ♀, stones in Pinus Enoplognatha penelope Hippa & Oksala, 1982 forest, 27.IV.2007 (CJVK); Akamas Peninsula, Cape Geronnisos, 25m, 1 ♀, stones in maquis, Identification 29.IV.007 (CJVK); Livadi pic nic site, 350m, 1 ♀, Hippa & Oksala, 1982: 221, figs 7 - 10. under stones in forest, 28.IV.2007 (CJVK); 9 km N. Pano Panagia, 1050m, 1 ♀, in litter near Citations spring, 19.V.2011 (CJVK); Peristerona, None. Atichoullis gorge, 425m, 1 ♀, stones in maquis, 2.V.2007 (CJVK); Smigies, 400m, 1 ♀, stones in New records maquis, 2 5.IV.2007 (CJVK). NICOSIA : Lefka NW, 120m, 2 ♂♂ 4 ♀♀; beaten from shrubs in fruit plantation, 18.V.2011 (CJVK, Distribution: Mediterranean. New to Cyprus.

CRB); Marathassa Dam, 550m, 1 ♂ 1 ♀, beaten Enoplognatha thoracica (Hahn, 1833) from shrubs around lake, 20.V.2011 (CR B). PAPHOS : Vretsia W., near Kelefos Bridge, Identification 650m, 3 ♂♂ 1 ♀, beaten from shrubs, 19.V.2011 Bosmans & Van Keer, 1999: 218, figs 36 - 41. (CJVK, CRB). Citations Distribution: Greece, Bulgaria. New to Cyprus. None.

Enoplognatha quadripunctata Simon, 1884 New records Enoplognatha quadripunctata ; Bosmans & Van FAMAGUSTA : Agios S ergios , 30m, 3 ♂♂, in Keer, 1 999: 218; Knoflach, 2004: 237. pit falls in Pinus plantation, 15 - 27.V.2011 (CRB). Identification KERYNEIA: Pentadactylos Range, Saint Hilarion Bosmans & Van Keer, 1999: 218, figs 36 - 41. Castle W., 600m, 3 ♂♂ 1 ♀, stones in Pinus Citations forest, 17.V.2011 (CRB), 3 ♂♂ 1 ♀, pitfalls at None. same site, III. - V.2010 (CRB). LARNACA : Delikipos, near Kornos, 1 ♀ , under New records stones in grass of open pine wood, 9.V.2013

Spiders of Cyprus 128

(CAR - S); Stavrovouni Monastery, 350m, 2 ♂♂ 1 Levy & Amitai, 1981: 178, figs 1 - 10. ♀, in pitfalls in open Pinus forest, III. - V.2011 (CRB). Citations PAPHOS : Mavrokol ympos Reservoir (Logunov, LIMASSOL : NW Erimi, 1 ♀, grass heathland 2013). with individual stones, 21.IV.1986 (NHMW); Pano Platres, t rack near Caledonia Waterfall, 1 ♀, New records under stones and in litter at edge of P inus nigra FAMAGUSTA : Davlos W., 25m, 1 subadult ♂, forest, 26.IV.2007 (CAR - S); road Pano Platres - stones in phrygana near the beach, 25.V.2011 Troodos, 1 ♀, under stones in Pinus nigra forest, (CRB); Kantara E., 610m, 1 subadult ♂, under 1.V.2007 (CAR - S). stones in open Pinus forest, 25.V.2011 (CRB); Protaras, Profitis Elias, 1 ♂, 2005 (CDM); NICOSIA: Athalassa National Forest Park, Salamis ruins, 50m, 1 ♀, under stones, 15.V.2011, Nicosia, 1 ♀, under stones in pine plantation, CRB) 2.V.2013 (CDM), 1 ♀, (CAR - S); Dali, 1 ♀, under KERYNEIA : near Agios Georgios, 1 ♀, under stones and in grass in riverbed, 12.V.2007 (CAR - stones, 27.II.2011, H. Henderickx leg. (CJB); S); Pallouriotissa, Nicosia 1 ♀, in garden, Saint Hilarion castle, 610m, 1 subadult ♂ 1 ♀, 15.VI.2008 (CDM). under stones in Pinus forest, 22.XI.2010 (CRB) . LARNACA : Achna Reservoir, 50m, 1 ♀, 1 Distribution: Holarctic. New to Cyprus. subadult ♀, stones and litter bordering lake, 16.V.2011 (CRB), 1 subadult ♂ 2 ♀♀, 27.V.2011 Episinus maculi pes Cavanna, 1876 (CJVK, CRB); Larnaca salt lake, 5m, 1 ♀, stones and litter in salt marsh, 27.IV.2010 (CRB); Identification Larnaca SW., Hala Sultan Tekk e mosque, 5m, 1 Hillyard, 1983: 88, figs 1 - 7. ♂ 1 ♀, litter in salt marsh, 15.V.2011 (CJVK). LIMASSOL : Pallouriotissa, Nicosia, 1 ♂, in Citations garden, IV.2012 (CDM); Pano Platres, near top of None. road to Millomeris Waterfalls, 1 ♂, under stones in abandoned vineyard, 28.IV.2007 (CAR - S); New records Pissour i beach, 1 ♂, under stones, 27.IV.1982, NICOSIA: Agridi, near Dali, 1 ♀, in garden, coll. A. Stubbs (MMUE). IV.2004 (CDM). NICOSIA : Agios Sozomenos , near Dali, 1 ♀, under stone in dry, rocky phrygana, 20.IV.2017 Distribution: Europe, Ukraine, Russia. New to (CAR - S); Agios Sozomenos , near Dali, 1 ♀, Cyprus. under stones and in dry grassland, 30.IV. 2007 (CRS), 1 ♂ , 29.IV.2013 (CAR - S); Agridi, near Episinus truncatus L atreille, 1809 Dali, 1 ♂, in garden, IV.2004, 1 ♂, in field, V.2007, 1 ♀, grass in lawn, 1.VII.2010 (CDM); Identification : https://araneae.nmbe.ch/ . Dali, 13, Agios Andronikos, 1 ♂, in garden, V.2007; 1 ♂, in garden, IV.2008 (CDM); Kampos Citations NE, Xeros River, 550m, 1 ♂, stones in r iverbed, None. 20.V.2011 (CRB); Kidasi N., along Diarizos River,300m, 1 subadult ♀, under stones, 21.V.2011 (CRB); Lakatamia, 1 ♂ 2 ♀♀, in olive New records PAPHOS : Kinousa SE . , 400m, 3 ♂♂, under grove, 1.I. - 1.X. 2016 (CMH); Limnitis, 1 ♀, stones and litter in Pinus forest, 1.V.2010 (CRB); Pinus brutia forest, 13.V.2016 (CMH); Oikos N., Xerarga ka picnic site, 350m, 1 ♂, Marathassa Dam, 550m, 1 ♂, under st ones around stones in Pinus forest, 20.V.2011 (CJVK). lake, 20.V.2011 (CRB); Masari S., Merika River, 120m, 1 ♂ 2 ♀♀, stones in wasteland, 24.V.2011 (CJVK, CRB); Nicosia, 1 ♀, under stone in grassland, 1.IV.1997 (CJB); Strovolos, 1 ♀, Distribution: Palaearctic. New to Cyprus. around houses, 1.V.2016 (CMH). PAPHOS : Evretou Reservoir, 1 ♀ , under stone in Euryopis episinoides (Walckenaer, 1847) grass tussocks, 13.IV.2017 (CAR - S); Evretou Reservoir, 350m, 1 ♀, under stones in grassland Identification around lake, 1.V.2010 (CRB); Kato Akourdaleia, Spiders of Cyprus 129

1 ♂, in guest house swimming pool, 3.V.2007 None. (CRS), 1 ♀, in garden, VIII.2005, 3 ♀♀, VII.2008 (CDM); Kato Paphos archaeological site, 1 New records subadult ♀, under stones in grass, 10.V.2007 FAMAGUSTA : Karpasia Peninsula, Cape (CAR - S); Kinousa, 1 ♂, in pine wood, Apostolos Andreas, 70m, 1 ♂, stones in grassland, 23.III.1998, coll. J. Murphy (MMUE); 16.V.2011 (CJVK). Mavrokolympos Reservoir, 1 ♂, C. Saure leg., NICOSIA : Dali, Idalion Archaeological Site, 1 ♀, 20.VI.2013 (CKHK); Mesana, 1 subadult ♂, in under stones in p hrygana, 1.V.2013 (CDM); garden, 10.XI .2015 (CMH); Pelathousa SE., Kapedes S., 600m, 1 ♂, under stones along 245m, 1 ♂, sweeping grassland, 30.IV.2007 rivulet, 5.V.2010 (CJVK); Nikitari S., Panagia (CJVK). Asinou, 1 ♂, under stones in disturbed phrygana, 9.V.2007 (CAR - S); Oikos N., Xerargaka picnic Distribution: Mediterranean, China. New to site, 350m, 1 ♀, stones in Pinus forest, 20.V.20 11 Cyprus. (CRB); near Platania Scout camping site, 2 ♂♂, pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 Euryopis laeta (Westring, 1861) (CMH). LIMASSOL: Parekklisia, 1 ♂, under stones in Identification : https://araneae.nm be.ch/ . riverbed, 13.IV.1987 (NHMW); Pissouri, 1 ♂, in grass, 24.III.1998, coll. J. Murphy (MMUE). Citations None. Distri bution: Algeria, Tunisia, Libya, Greece, Israel. New to Cyprus. New records LIMASSOL: Armenochori, 1 ♀, on bushes, Kochiura aulica (C. L. Koch, 1838) 14.IV.1986 (NHMW). Identification : https://araneae.nmbe.ch/ . Distribution: Europe, Russia, Tunisia to Tajikistan. New to Cyprus. Citations None. Euryopis quinqueguttata Thorell, 1875 New records FAMAGUSTA : Agio s Sergios, 30m, 1 ♂, beaten Identification : https://araneae.nmbe.ch/ . in Pinus plantation, 15.V.2011 (CRB); Kantara E., 610m, 1 ♂ 1 ♀, beaten from shrubs in open Pinus Citations fore st, 25.V.2011 (CRB); Rizokarpaso, 3 ♀♀, in None. Juniperus phoenicea forest, 12.III.2016 (CMH); New records KERYNEIA : Saint Hilarion Castle, 600m, 2 ♂♂ LARNACA : Achna Reservoir, 50m, 1 ♂, under 2 ♀♀, beaten from shrubs, 17.V.2011 (CJVK, stones and litter bordering lake, 27.V.2011 CRB). (CJVK); Larnaca SW . , Hala Sultan Tekke LARNACA : Achna Reservoir, 50m, 1 ♂, on mosque, 50m, 1 ♀, i n pitfalls in salt marsh, 15 - herbs bordering lake, 27.V.2011 (CRB); Hala 28.V.2011 (CJVK). Sultan Tekke Mosque, 1 ♀, herbs in the NICOSIA : Masari S., Merika River, 120m, 1 ♂, surrounding park, 20.IV.1986 (NHMW); Larnaca stones in wasteland, 24.V.2011 (CJVK). salt lake, 5m, 2 ♀♀, on her bs in salt marsh, 27.IV.2010 (CJVK, CRB); Kornos Picnic site, Distribution: Europe, Egypt to Turkmenistan. 350m, 1 ♂ 1 ♀, beaten in Pinus forest, 28.V.2011 New to Cyprus. (CJVK); Stavrovouni Monastery, 350m, 1 ♂ 2 ♀♀, swept in Pinus forest, 28.IV.2010 (CJVK, CRB). LIMASSOL: Akrotiri Peninsula, near British Euryopis sexalbomaculata (Lucas, 1846) Airbas e, 1 ♀, swept in pine wood, 4.V.2007

(CAR - S); Akrotiri Peninsula, near Fasouri reed Identif ication Levy, 1998b: 146, figs 275 - 283. beds, 1 ♀, swept from flowering herbs and grass, 27.IV.2007 (CAR - S); Amathus , 1 ♀, on herbs in Citations dry valley, 6.IV.1986 (NHMW); Fasoula, 2 ♀♀,

Spiders of Cyprus 130

olive garden with dry stone wall s, 16.IV.1986 Distribution: Mediterranean. New to Cypru s. (NHMW); Kannavia, 780m, 1 ♂ 1 ♀, sweeping grassland, 4.V.2007 (CJVK); Kolossi, N of the Lasaeola coracina (C. L. Koch, 1837) castle, 1 ♀, under stones, 11.IV.1986 (NHMW). Identification : https://araneae.nmbe.ch/ . NICOSIA : Agridi, near Dali, 1 ♂, in field, V. 2007, 1 ♂, in field, IV. 2008, 1 ♂, in garden, Citations 1V.2010, 1 ♂, in garden, V.2011 (all CDM); Dali, None. 13, Agios Andronikos, 1 ♂ 1 ♀, in garden, IV.2007 (CDM); Kampos NE, Xeros River, 550m, 2 ♂♂ 1 ♀, beaten from shrubs, 20.V.2011 New records (CRB); Kapedes, 1 ♀, pitfalls in Pinus brutia KERYNEIA : Larnakas tis Lapithou, stones in forest, 17.I. - 23.IV.2016 (CMH); Lakatamia, 1 ♂ grassland, 1 ♂, 17.I.2016 (CMH). 3 ♀♀, in olive grove, 1.I. - 1.X. 2016 (CMH); Lefka NW, 120m, 4 ♀♀, beaten from shrubs in NICOSIA : Prodromos S., Trooditissa picnic site, fruit plantation, 18.V.2011 (CJVK, CRB); Masari 1450m, 1 ♂, stones in Pinus forest, 21.V.2011 S., Merika River, 120m, 1 ♂, beaten from shrubs (CRB). in wasteland, 24.V.2011 (CRB); Xerargaka S, Orkontas, 2 ♀♀, riverside vegetation, 26.III.2016 Distribution: Europe. New to Cyprus. (CMH). PAPHOS : Agios Georgios, 180m, 1 ♀, pitfalls in Lasaeola testaceomarginata Simon, 1881 reed bed of river Diarizos, 26.IV. - 5.V.2007 (CJVK); Asprokremmos Dam, N. Kouklia, 150m, Identification 1 ♂ 2 ♀♀, on herbs along lake, 23.V.2011 (CRB); Brignoli, 1968: 97, figs 7, 8, 11. Avakas gorge and surroundings, 1 ♂, 22 .III.2018 (CKHK); Evretou Reservoir, 350m, 1 ♀, swept Cita tions from shrubs and grassland around lake, 1.V.2010 None. (CRB); Kato Paphos, 1 ♂ 1 ♀, beating tall herbs near hotel, 4.IV.1997 (CJB); Kidasi N., along New records Diarizos River, 300m, 1 ♂ 4 ♀♀, beaten from LARNACA : Larnaca salt lake, 5m, 1 ♀, beaten shrubs, 21.V.201 1 (CRB); Kourdaka, Ezousa from shrubs along lake, 27.IV.2010 (CRB). river, 1 ♂, 14.III.2018 (CKHK); Makounta, Argaka Dam, 85m, 1 ♂, beating in maquis, Distribution: Mediterranean: the south of France, 27.IV.2007 (CJVK); Mavrokolympos Reservoir, 2 Sicilia, Portugal, Morocco, Greece. New to ♂♂, 20.VI.2013 , C. Saure leg., (CKHK) ; same Cyprus. site, 1 ♂, 13.III.2018 (CKHK); Pegeia, Pikni Forest, 346m , 1 ♂, in former carob plantation, Latrodectus dahli Lev y, 1959 11.III.2018 (CKHK); Vretsia W., near Kelefos Bridge, 650m, 2 ♂♂ 9 ♀♀, beaten from shrubs, Identification 19.V.2011 (CJVK). Levy, 1998b: 108, figs 198 - 202; Knoflach & van Harten, 2002 : 333, figs 3, 12 - 19. Distribution: Macaronesia to Azerbaijan. Citations Lasaeola convexa (Blackwall, 1870) None.

Identification New recor ds Levy, 1998b : 154, figs 287 - 293. LIMASSOL: Limassol, 1 ♂ 4 ♀♀, in hotel Sheraton, 5.X.1988 (NHMW). NICOSIA: North Dali, 1 ♀, collected from field Citations under stones, IV.2004 (CDM). None. Distribution: Morocco to Central Asia. New to New records Cyprus. PAPHOS : Asprokremmos Dam, N. Kouklia, 150m, 1 ♂, herbs along lake, 29.IV.2010 (CRB). Latrodectus geometricus C. L. Koch, 1841

Spiders of Cyprus 131

Identification PAPHOS : Akamas Peninsula, Agios Minas Knoflach & van Harten, 2002: 339, figs 2, 20 - 28. (Knoflach et al., 2008); Makounta, Argaka dam Citations (Knoflach et al ., 2008). PAPHOS : Paphos (Knoflach & Van Harten, 2002). New records, FAMAGUSTA : Gialousa E., Agia Triada, 50m, 1 New records ♀, on shrubs in ru ins, 18.III.2011 (CRB); LARNACA : Larnaca SW, Hala Sultan Tekke Gialousa E., Agios Thyrsos, 20m, 1 ♀, beaten Mosque, 5m, 2 ♂♂ 4 ♀♀, on walls in mosque, from shrubs, 16.V.20111 (CRB). 20.XI.2010 (CRB), 1 ♂ 8 ♀♀ , 15.V.2011 (CJVK, KERYNEIA : Saint Hilarion castle, 610m, 1 CRB). subadult ♂, swept from herbs in Pi nus forest, NICOSIA: Agridi near Dali, 1 ♀, collected on 22.XI.2010 (CRB). house veranda, VII.2014 (CDM); Dali, 1 ♀ under LARNACA : Agioi Vavatsinias, 700m, 1 ♀, swept bicycle saddle, VIII. 2010 (CDM); Lakatamia, 1 ♂ in Pinus forest, 28.IV.2010 (CRB). ♀♀, in olive grove, 1.I - 1.X. 2016 (CMH). LIMASSOL : Alassa SE, near river Limnatis, 1 ♂, PAPHOS: Latchi beach, 2 ♀♀, collected from swept from lush grass, 5.V.2007 (CAR - S). sig n post, VII.2008 (CDM); Pegeia, 1 ♂, under NICOSIA : Marathassa Dam, 550m, 4 ♀♀, beaten stone, 15.XI.2016 (CMH). from shr ubs around lake, 20.V.2011 (CJVK). PAPHOS: Akamas Peninsula, Neo Chorio, 1 Distribution: Cosmopolitan. subadult, swept from phrygrana and grass, 2.V.2007 (CAR - S); Akamas Peninsula, Agios Latrodectus tredecimguttatus (Rossi, 1790) Minas, 1 ♂ 1 ♀, in thick pine litter in wood, 2.V.2007 (CRS); Avakas Gorge, 150m, 3 ♂♂ 4 Identification ♀♀, on shrubs in gorge, 28.IV.2010 (CJVK), 1 ♂ Levy, 1998b: 90, figs 161 - 177. 14 ♀♀, 22.V.2011 (CJVK, CRB); Evretou Reservoir, 350m, 1 ♀, swept from shrubs around Citations lake, 1.V.2010 (CRB); Makounta, Argaka Dam, None. 85m, 1 ♂, beating in maquis, 27.IV.2007 (CJVK); 9 km N. Pano Panagia, 1050m, 10 ♂♂ 11 ♀♀, on New records shrubs near spring, 19.V.2011 (CJVK, CRB); FAMAGUS TA : Karpasia Peninsula, Cape Smigies, W. Neo Chorio, 300m, 1 subadult ♀, Apostolos Andreas, 70m, 1 ♀, stones in grassland, swept in forest, 30.IV.2010 (CRB). 16.V.2011 (CJVK). LARNACA : Achna Reservoir, 50m, 1 ♂ 1 ♀, in Distribution: Lebanon, Israel, Crete, Rhodos, litter bordering lake, 27.V.2011 (CJVK, CRB); Cyprus. Larnaca SW., Hala Sultan Tekke mosque, 5m, 2 subadult ♂♂, under ston es in salt marsh, Neottiura herbigrada (Simon, 1873) 16.III.2011 (CRB). Neottiura herbigrada ; Knof lach, 1999: 353; NICOSIA: North Dali, 2 subadult ♀♀, under Logunov, 2013: 5. stones in field, IV, 2004 (CDM); Pera Chorio, 1 ♀, in laundry room, V . 2017 (CDM). Identification Knoflach, 1999: 353, figs 6, 9, 20 - 22, 36 - 37, 45, Distribution: Mediterranean to China. New to 54, 58, 61, 67. Cyprus.

Neospintharus syriacus (O . Pickard - Citations Cambridge, 1872) LIMASSOL : Kourion (Knoflach, 1999). PAPHOS : Akamas Peninsula, Fontana Amorosa Identification (Knoflach, 1999); W. Polis, Akamas Forest Levy, 1985a: 103, figs 29 - 43. (Logunov, 2013). New records LARNACA : Agioi Vavatsinias, 700m, 1 ♂, litter Citations in Pinus forest, 28.IV.2010 (CJVK). LIMASSOL : Alassa SE (Knoflach et al. , 2008). LIMASSOL: Akrotiri Peninsula, near British Airbase, 1 ♀, swept in pine wood, 4.V.2007

Spiders of Cyprus 132

(CAR - S); road Kaminaria - Lemithou, 1 ♂, pitfall s Kouklia, 150m, 1 ♀, herbs and stones along lake, in Pinus brutia forest, 6.IV. - 8.VI.2014 (CMH). 29.IV.2010 (CJVK); Evretou E., 200m, 1 ♂, NICOSIA : Marathassa Dam, 550m, 1 ♀, beaten sweeping low herbs, 1.V.2007 (CJVK); Evretou from shrubs around lake, 20.V.2011 (CRB); Reservoir, 350m, 1 ♀ , shrubs and grassland Masari S., Merika River, 120m, 1 ♀, on bushes in around lake, 1.V.2010 (CJVK); Gialia, 110m, 3 wasteland, 24.V.2011 (CRB). ♀♀, sweeping Cystus , 28.IV.2007 (CJVK); PAPHOS : Kato Akourdaleia, 1 ♂, in guest house Pelathousa SE., 245m, 2 ♀♀, sweeping grassland, swimming pool, 3.V.2007 (CRS); Kelefos bridge, 30.IV.2007 (CJVK). dam on Diarizos River, 550m, 1 ♀, beaten from shrubs, 21.V.2011 (CJVK); 9 km N. Pano Distribution: Mediterranean. Panagia, 1050m, 1 ♀, in litter near spring, 19.V.2011 (CJVK); Pegeia, Agios Georgios, 1 ♂, Paidiscura dromedaria (Simon, 1880 ) rocky shore, 13.III.2018 ( CKHK); Smigies, W. Neo Chorio, 300m, 4 ♀♀, swept in Pinus forest, Identification 30.IV.2010 (CRB). Knoflach & Thaler, 2000: 429, figs 30, 33, 36, 41 - 42, 58. Distribution: European part of the Mediterranean, from France to Israel and Cyprus. Citations None. Neottiura suaveolens (Simon, 1880) New records Identification LARNACA : Larnaca salt lake, 5m, 3 ♀♀, swept Knoflach, 1999: 357, figs 15, 16, 23, 40, 41, 46, from shrubs around lake, 27.IV.2010 (CRB); 51, 59, 62, 66, 70. Larnaca SW, Hala Sultan Tekke mosque, 50m, 1 ♀, in pitfalls in salt marsh, 15 - 28.V.2011 (CRB). Citations LIMASSOL: Akrotiri Peninsula, salt lake, 4 ♂♂ 2 None. ♀♀, beaten from Salicornia , 15.III.2018 (CKHK); Akrotiri Peninsula, Turtle Beach, 1 ♂, sandy New records beach, beaten from vegetation, 15.III.2018 KE RYNEIA:Keryneia , dunes near Topset Hotel, (CKHK). 1 ♂ 1 subadult ♂, 28.II.2011, H. Henderickx leg. PAPHOS : As prokremmos Dam, N. Kouklia, (CJB). 150m, 22 ♀♀, herbs along lake, 29.IV.2010 (CRB), 2 ♂♂ 22 ♀♀, 23.V.2011 (CJVK, CRB); Distribution: South and Centr a l Europe. New to Paphos,3 ♀♀, bay at hotel “Paphian Bay”, beating Cypru s. herbs near beach, 4.IV.1997 (CJB).

Neottiura uncinata (Lucas, 1846) Distribution: Spain, North Africa to Middle East, Neottiura uncinata ; Logunov, 2013: 5. Cape Verde Is., Greece. New to Cyprus.

Identification Pholcomma gibbum (Westring, 1851) Knoflach, 1999: 358, figs 19, 42 - 43, 48, 53, 57, 60, 63, 68. Identification : https://araneae.nmbe.ch/ .

Citations Citations PAPHOS : Avakas Gorge (Logunov, 2013). None. LIMASSOL: Akrotiri Salt Lake ( Logunov, 2013). New records New records LARNACA: Agios Minas, 1 ♀, under stone in NICOSIA: Agridi, near Dali, 1 ♂, in garden, pine wo od, 20.III.1998, coll. J. Murphy (MMUE). III.2014 (CDM). LIMASSOL: Akrotiri Peninsula, near Fasouri PAPHOS : Agios Mamas ruins, Gialia E., 215m, 1 reed beds, 8 ♀♀, swept from flowering herbs and ♂, beating Cystus , 28.IV.2007 (CJVK); Agios grass, 4.IV.2007 (CRS). Merkourios, 250m, 2 ♂♂, sweeping grassland, PAPHOS : Akamas Peninsula, Agios Minas, 2 27.IV.2007 (CJVK); Akamas Peninsula, Neo Chorio, 2 ♀♀, swept from phrygana and grass, ♀♀, in thick pine litter, 2.V.2007 (CAR - S); 2.V.2007 (CAR - S); Asprokremmos Dam, N.

Spiders of Cyprus 133

Kelefos br idge, 465m, 3 ♀♀, deciduous wood, NICOSIA : Kampos NE, Xeros River, 550m, 3 16.III.2018 (CKHK). ♂♂ 1 ♀, beaten from shrubs, 20.V.2011 (CJVK, CRB); Lefka NW, 120m, 1 ♂ 1 ♀, beaten from Distribution: Europe, Russia, Ukraine, North shrubs in fruit plantation, 18.V.2011 (CRB); Africa. New to Cyprus. Oikos N., Xerargaka picni c site, 350m, 5 ♀♀, beaten from shru bs in Pinus forest, 20.V.2011 Phycosoma inornatum (O. Pickard - Cambridge, (CJVK, CRB); Prodromos E., near Prodomos 1861) dam, 1510m, 1 ♀, swept in Pinus forest, 2.V.2010 (CJVK); Xerargaka S, Orkontas, 1 ♀, riverside Identification : https://ar aneae.nmbe.ch/ . vegetation, 26.III.2016 (CMH). LIMASSOL: Akrotiri Pe ninsula, near British Citations Airbase, 1 ♂, swept in pine wood, 4.V.2007 None. (CRS); Troodos Square, 1 ♂, Pinus nigra forest, 11.V.2016 (CMH); Amathus, 3 ♂♂ 2 ♀♀, herbs New records in dry valley, 25.IV.1986 (NHMW). PAPHOS: Akamas Peninsula, Neo Chorio, 1 ♂ 1 PAPHOS : Agios Mamas ruins, Gialia E., 215m, 1 ♀, swept from phrygrana and grass, 2.V.2007 ♂ 1 ♀, beating Cys tus , 28.IV.2007 (CJVK); (CRS). Akamas Peninsula, Neo Chorio, 1 ♀, swept from phrygrana and grass, 2.V.2007 (CAR - S); Distribution: Europe to Azerbaijan. New to Asprokremmos Dam, N. Kouklia, 150m, 3 ♀♀, Cyprus. herbs along lake, 29.IV.2010 (CJV, CRB); near Avakas Gorge, 1 ♀, beaten from junipers on Phylloneta impressa (L. Koch, 1881) rocky hillside, 14.IV.2017 (CAR - S); Evretou Reservoir, 350m, 1 ♂ 1 ♀, swept from shrubs Identification : https://araneae.nmbe.ch/ . around lake, 1.V.2010 (CRB); Evretou Reservoir, Stavros tis Psokas river, 2 ♂♂, 20.III.2018 Citations (CKHK); Kelefos bridge, dam on Diarizos River, None. 550m, 2 ♂♂ 2 ♀♀, under stones and beaten from shrubs, 21.V.2011 (CJVK); Gialia, 110m, 1 ♂, New records sweeping Cystus , 28.IV.2007 (CJVK); Kinousa Without precise locality, 1 ♂ 2 imm atures , SE, 400m, 2 ♂♂ 4 ♀♀, herbs in Pinus forest, VI.1999, coll. J. Murphy (MMUE). 1.V.2010 (CJVK, CRB); near Kritou Terra, 2 ♂♂ 1 ♀, swept from grass in dry fallow, 14.IV.2017 Distribution: Holarctic. New to Cyprus. (CAR - S); Mavrokolympos Reser voir, 1 ♂ 1 ♀, 13.III.2018 (CKHK); Pelathousa SE., 245m, 3 Platnickina nigropunctata (Lucas, 1846) ♂♂, sweeping grassland, 30.IV.2007 (CJVK); Peristerona, Atichoullis gorge, 425m, beating in Identification maquis, 2.V.2007 (CJVK); Pegeia, Pikni Forest, Levy, 1998b: 189, figs 357 - 363. 346m, 1 ♂, in former carob plantation, 23.III.2018 (CKHK); Polis, 10m, 1 ♂, herbs and stones Citations around building, 27.IV.2007 (CJVK); Vretsia W., None. near Kelefos Bridge, 650m, 1 ♂ 1 ♀, beaten from shrubs, 19.V.2011 (CRB). New records FAMAGUSTA : Gialousa E., Agios Thyrsos, Distribution: Mediterranean. New to Cyprus.

20m, 1 ♀, beaten from shrubs, 16.V.20111 Rhomphaea longicaudata O. Pickard - (CRB); Gialousa E., Agia Triada, 50m, 4 ♀♀, Cambridge, 1872 herbs in ruins, 18.III.2011 (CJVK, CRB); Kantara E., 610m, 2 ♂♂ 1 ♀, beaten from shrubs in open Pinus forest, 25.V.2011 (CJVK, CRB); Karpasia Identification Peninsula, Cape Apostolos Andreas, 70m, 1 ♀, Lissner , 2017 : 288, figs 1A - C, 2A - C, 3A - B, 4A - swept in grassland, 16.V.2011 (CRB). B, 5 - 8.

LARNACA : Kornos Picnic sit e, 350m, 1 ♀, Citations beaten in Pinus forest, 28.V.2011 (CRB). None.

Spiders of Cyprus 134

New records New records LIMAS SOL: Pissouri beach, 2 ♀♀ 1 immature , FAMAGUSTA : Kantara E., 610m, 1 ♂ 15 ♀♀ , on shrubs, 27.IV.1982, coll. A. Stu bbs (MMUE). beaten from shrubs in open Pinus forest, 25.V.2011 (CJVK, CRB). Distribution: Greece, Lebanon. New to Cyprus. KERYNEIA : Pentadactylos Range, Saint Hilarion Castle, 600m, 2 ♂♂ 4 ♀♀, beaten from shrubs, Simitidion agaricographum (Levy & Amitai, 17.V.2011 (CRB). 1982) LARNACA : Agioi Vavatsinias, 7 00m, 1 ♂ 7 ♀♀, Simithidion agaricographum ; Knoflach, 1996: litter in Pinus forest, 28.IV.2010 (CJVK); 149. Stavrovouni Monastery, 350m,1 ♂ 2 ♀♀, on bark of Pinus trees, 28.IV.2010 (CJVK). Identification LIMASSOL: Akrotiri Peninsula, near British Knoflach, 1996, figs 1 - 3, 7, 9, 11 - 12. Airbase, 1 ♀, swept in pine wood, 4.V.2007 (CRS); Alassa SE, near river Limnatis, 1 ♂, swept Citations from lush grass, 5.V.2007 (CAR - S), 5 ♀♀ (CRS); PAPHOS : Akama s Peninsula, Fontana Amorosa Pano Platres, 1050m, 2 ♂♂ 1 ♀, beaten near Mesa (Knoflach, 1996). Potamos Waterfall s, 23.V.2011 (CRB); Pano Platres, near top of road to Millomeris Waterfalls, New records 4 ♂♂ 4 ♀♀, swept from grass and herbs in FAMAGUSTA : Kantara E., 610m, 1 ♂, beaten aband oned vineyard, 28.IV.2007 (CAR - S); Pano from shrubs and stones in open Pinus forest, Platres, below Caledonia Waterfall, 2 ♂♂, swept 25.V.2011 (CJVK); Karpasia Peninsula, Cape from herbs and shrubs in rocky stream valley, Apostolos Andreas, 70m, 1 ♀, stones in grassland, 26.IV. and 1.V.2007 (CAR - S), Pano Platres E., 16.V.2011 ( CJVK). Mesa Potamos Waterfall s, 950m, 1 ♂ 10 ♀♀, in LARNACA : Kornos Picnic site, 350m, 1 ♀, under litter in Pinus forest, 4.V.201 0 (CJVK); Troodos stones and beaten in Pinus forest, 28.V.2011 S., 1350m, 1 ♂, on herbs in mixed Quercus - Pinus (CRB). forest, 3.V.2010 (CRB); same site ; 5 ♂♂ 7 ♀♀, LIMASSOL : Parekklisia, 3 ♂♂ 1 ♀, under stones 28.IV.2007 (CRS). in riverbed, 13.IV.1987 (NHMW). NICOSIA : Cedar Valley Forest, 1180m, 2 ♂♂ 4 NICOSIA: Agridi , near Dali, 1 ♂, in garden, ♀♀, beaten in Cedrus forest, 19.V.2011 (CRB); Cedar Valley, 1150m, 1 ♂, 21.III.2 018, forest IV. 2004 (CDM). road, beaten from shrubs (CKHK); Dali, 13, PAPHOS : Agios Mamas ruins, Gialia E., 215m, 1 Agios Andronikos, 1 ♀, in garden, IV.2007 ♀, beating Cystus , 28.IV.2007 (CJVK); near (CDM); Kakopetria S., Platania forest station, Avakas Gorge, 2 ♂♂ 3 ♀♀, beaten from junipers 1100m, 1 ♂ 7 ♀♀, beating in dense Pinus forest, on rocky hillside, 14.IV.2017 (CAR - S); Lara, 4.V.2010 (CJVK); Kapedes, 1 ♂, pitfalls in Pinus 25m, 1 ♀, stony Juniperus maquis near the beach, brutia forest, 17.I. - 23.IV.2016 (CMH); Lefka 29.IV.2007 (CJVK); Makounta, Arga ka Dam, NW, 120m, 3 ♀♀, beaten from shrubs in fruit plantation, 18.V.2011 (CRB); Kampos NE, Xeros 85m, 1 ♂ 1 ♀, beating bushes, 27.IV. - 5.V.2007 River, 550m, 3 ♀♀, beaten from shrubs, (CJVK); Mavrokolympos Reservoir, 1 ♂ 1 ♀, 20.V.2011 (CRB); Limnitis River, Kato Pyrgos 13.III.2018 (CKHK); Neo Chorio, 1 ♀, swept SE, 200m, 2 ♀♀, beaten in Pinus forest, 2.V.2010 from phrygana and grass, 2.V.2007 (CAR - S); Neo ( CRB); Oikos N., Xerargaka picnic site, 350m, 3 Chorio, 200m, 1 ♀, under stones, 25.IV.2007 ♂ 28 ♀♀, beaten from shrubs and stones in Pinus (CJVK); Pelathousa SE., 24 5m, 1 ♂ 1 ♀, forest, 20.V.2011 (CJVK, CRB); Pedoulas, sweeping grassland, 30.IV.2007 (CJVK). 1100m, 1 ♀, in garden of hotel, 4.V.2010 (CRB). PAPHOS : Agios Mamas ruins, Gialia E., 1 ♂ 9

♀♀, beating Cystus , 28.IV. 2007 (CJVK); Agios Distribution: Israel, Cyprus, Lesbos. Merkourios, 250m, 2 ♂♂ 2 ♀♀, sweeping Simitidion simile (C. L. Koch, 1836) grassland, 27.IV.2007 (CJVK); Akamas

Peninsula, Neo Chorio, 1 ♀, swept from Identification: https://araneae.nmbe.ch/ . phrygrana and grass, 2.V.2007 (CAR - S) near

Agios Minas, 1 ♂ 1 ♀, 3.V.2007 (CRS); Argaka Citations SE, 160m, 3 ♂♂ 2 ♀♀, beating in maq uis, None. 27.IV.2007 (CJVK); Evretou Reservoir, Stavros

Spiders of Cyprus 135

tis Psokas river, 2 ♂♂ 1 ♀, 20.III.2018 (CKHK); Gialia, 110m, 4 ♀♀, sweeping Cystus , 28.IV.2007 Distribution: Cosmopolitan. New to Cyprus. (CJVK); Kelefos bridge, dam on Diarizos River, 550m, 2 ♀♀, beaten from shrubs, 21.V.2011 Steatoda grossa (C. L. Koch, 1838) (CRB); Kidasi N., al ong Diarizos River, 300m, 3 ♀♀, beaten from shrubs, 21.V.2011 (CRB); Identification : https://araneae.nmbe.ch/ . Makounta, Argaka Dam, 85m, 2 ♂♂ 2 ♀♀, beating in maquis, 27.IV.2007 (CJVK); Citations Pelathousa SE., 245m, 11 ♂♂ 23 ♀♀, sweeping None. grassland, 30.IV.2007 (CJVK); Peristerona, Atichoullis gorge, 425m, 11 ♀♀, beating in New records maquis, 2.V.2007 (CJVK); Vretsia W., near NICOSIA: Agridi, near Dali, 1 ♀, in garden, Kelefos Bridge, 650m, 1 ♂ 4 ♀♀, beaten from V.2004, 1 ♂, in house,V.2011, 1 ♂, in house, shrubs, 19.V.2011 (CJVK). IV.2012 (all CDM).

Steatoda albomaculata (Degeer, 1778) Distribution: Cosmopolitan. New to Cyprus.

Identification : https://araneae.nmb e.ch/ . Steatoda paykulliana (Walckenaer, 1806) Citations Lithyphantes paykullianus ; Strand, 1907: 29 3. None. Steatoda paykulliana ; Logunov, 2013: 5.

New records Identification LARNACA : Dekeleia Military Base, 1 ♂, Levy, 1998b: 63, figs 116 - 122. 11.VI.1958, M. Connelly leg. (BMNH); Kornos picnic site, 350m, 1 ♂ 2 ♀♀, under stones in Citations Pinus forest, 28.V.2011 (CRB); Larnaca SW., Between Larnaca and Limassol (Strand, 1907). Hala Sultan Tekke mosque, 5m, 1 ♀, s alt marsh, LIMASSOL: Episkopi, Kourion Beach (Logunov, 15.V.2011 (CJVK). 2013). NICOSIA: Agridi, near Dali, 1 ♂, in house, V.2009 (CDM); 13, Agios Andronikos, Dali, 1 ♀, New records in garden, IV.2008 (CDM); Dali, 1 ♀, under FAMAGUSTA : Agios Sergios , 30m, 2 ♀♀, under stones in riverbed, 12.V.2007 (CAR - S); stones in phrygana, 17.III.2011 (CRB); Cape Peristerona, 1 ♀, riverbed, 7.V.2016 (CMH). Greco, 1 ♀, under stones in phrygana, 3.V.2013 PAPHOS : Akamas Peninsula, Cape Geronnisos, (CAR - S); Cape Greco, Ayia Napa, 35m, 2 ♀♀, 25m, 1 ♀, stones in maquis, 29.IV.007 (CJVK); stones in phrygana, 27.IV.2010 (CJVK); Drouseia, 539m, 1 ♀, in p hrygana, 23.III.2018 Flamoudi W ., 25m, 2 ♀♀, stones in phrygana (CKHK). near the beach, 25.V.2 011 (CRB); Rizokarpaso Distribution: Cosmopolitan. New to Cyprus. W ., Panagia Eleousa Monastery, 30m, 1 ♀, under stones in Pinus forest, 18.III.2011 (CRB); Salamis Steatoda erigoniformis (O. Pickard - Cambridge, ruins, 50m, 1 ♀, under stones, 15.V.2011 (CJVK). 1872) KERYNEIA : Karavas , 660m, 1 juvenile , garden of hotel, 22.XI.2010 (CRB); Larnakas t is Identificatio n Lapithou, 1 ♀, stones in grassland, 17.I.2016 Levy, 1998b : 81, figs 152 - 160. (CMH); Saint Hilarion castle, 610m, 1 juvenile, under stones in Pinus forest, 22.XI.2010 (CRB). Citations LARNACA : Achna Reservoir, 50m, 1 juvenile, None. under stones and litter bordering lake, 21.XI.2010 (CRB); Cape Kit i, 2 ♂♂ 1 ♀, under stones below coastal cliffs, 25.IV.1982, coll. A. Stubbs (MMUE); Evretou Reservoir, 350m, 2 ♀♀, under New records stones in grassland around lake, 1.V.2010 (CRB); LARNACA: Larnaca SW., Hala Sultan Tekke Larnaca, 2 ♀♀, C. Glasynet leg. (CMNH); mosque, 5m, 1 ♂, litter in salt marsh, 15.V.2011 Larnaca salt lake, 5m, 2 ♀♀, stones and litter in (CJVK). salt marsh, 27.IV.2010 (CJVK, CRB); idem, 1 ♀, LIMASSOL: Amathus, 1 ♂, herbs in dry valley, 20.XI.2010 (CRB). 25.IV.1986 (NHMW).

Spiders of Cyprus 136

LIMASSOL : Agios Tychonas, 2 ♀♀, under stones near house walls, 19.IV.1986 (NHMW); New records Akrotiri Peninsula, edge of central salt pan, 1 ♀, LIMASSOL : Mt Olympos, 1850m, 1 ♀, stones in under stones and debris in Salicornia , 27.IV.2007 Pinus nigra f orest, 26. IV.2007 (CJVK); Polis, (C AR - S); Alassa SE, near river Limnatis, 1 ♀, 15m, 1 ♀, herbs and stones around building, under stones in dry grassland, 5.V.2007 (CAR - S); 27.IV.2007 (CJVK). Amathus, 5 ♀♀, under stones, in dry va lley, 25.IV.1986 (NHMW), 7 ♀♀, 6.IV.1986, 4 ♀♀, Distribution: Cosmopolitan. New to Cyprus. 9.IV.1986 (NHMW); Armenochori, 1 ♀, under stones of a dry stone wall, 14.IV.1 986 (NHMW); Theridion cinereum Thorell, 1875 Erimi NW, 1 ♀, grass heathland with individual stones, 21.IV.1986 (NHMW); Erimi, 1 ♀, under Identification stones along dry riverbed, 18.IV.1986 (NHMW); Knoflach, 1998: 589, figs 46, 52, 55d. Fasoula, 2 ♀♀, olive garden with dry stone walls, 16.IV.1986 (NHMW); Germasogeia, 1 ♀, under Citations stones along the stream below the dam, None. 10.IV.1986 (NHMW); Limassol, 2 ♂♂ 1 juv. ♀, A. Mavromoustakis leg., 2.X.1933 (BMNH); New records Panagia Glossa Monastery near Kellaki, 1 ♀, FAMAGUSTA : Gialousa E., Agios Thyrsos, 1 ♀, stones in cultivated land, 12.IV.1986 (NHMW). 20m, litter under fig tree, 18.III.2011 (CRB). NICOSIA: Agios Sozomenos , near Dali, 1 ♀, in disturbe d phrygana, 30.IV.2007 (CRS); Agridi, Distribution: Switzerland, Austria, Balkans, near Dali, 2 ♂♂, on veranda, 10.III.2010 (CDM); Bulgaria, Greece, Turkey, Russia. New to Cyprus.

Geri, 1 subadult ♀, in olive grove, 11.V.2016 Theridion cyprusense Wunderlich, 20 11 (CMH). Theridion cyprusense Wunderlich, 2011: 240, figs PAPHOS : Akamas Peninsula, Agios Minas, 1 ♀, 3 - 5. under stones in meadow with tall grass, 2.V.2007 (C AR - S); Akamas Penins ula, Avgas E stuary, 1 ♀, Identification under stones, 31.III.1997 (CJB); Asprokremmos Wunderlich, 2011: 240, figs 3 - 5. Dam, N. Kouklia, 150m, 2 ♀♀, herbs and stones along lake, 29.IV.2010 (CJVK); Evretou Citations Reservoir, 180m, 3 ♀♀, stones on slope to lake, PAPHOS : beach near Polis (Wunderlich, 2011; 1.V.2007 (CJVK); Kato Paphos, 1 ♂ 2 ♀♀, under type locality). stones near hotel, 31.III.1997 (CJB); Kelefos bridge, dam on Diarizos River, 550m, 1 ♀, under New records stones and beaten from shrubs, 21.V.2011 PAPHOS : 9 km N. Pano Panagia, 1050m, 1 ♂, in (CJVK); Makounta, 1 ♀, under stones in scrub, l itter near spring, 19.V.2011 (CJVK); Evretou 19.III.1998, coll. J. Murphy (MMUE); Kathikas Reservoir, Stavros tis Psokas river, 1 ♂, W., S Arodes, Aspros Valley, 527m, 1 ♂ 1 ♀, 20.III.2018 (CKHK). open pasture on clay, 24.II.2011, D. W. Wrase leg (CKHK); Mavrokolympos Reservoir, 1 ♀, Distribution: Only known from Cyprus. 13.III.2018 (CKHK); Neo Chorio, 1 ♀, under stone, 17.III.1998, coll. J. Murphy (MMUE); Theridion helena Wunderlich, 2011 Polis, 15m, 1 ♂ 1 ♀, stones around building, 27.IV.2007 (CJVK). Identification Wunderlich, 2011 : 250, figs 99 - 101. Dis tribution: Mediterranean to Central Asia. Citations N one.

Steatoda trianguloides (Levy, 1991) New records FAMAGUSTA : Cape Greco, Ayia Napa, 35m, 1 Identification ♀, stones in phrygana, 27.IV.2010 (CRB); Levy, 1991: 228, figs 3 - 6 ( ♂); present paper , f igs Kantara E., 610m, 2 ♀♀, beaten from shrubs and 19 a - e ( ♀). stones in open Pinus forest, 25.V.2011 (CRB).

Spiders of Cyprus 137

KERYNEIA : Panagia Apsinthiotissa Monastery ruins, 350m , 1 ♀, herbs in clearing in Pinus forest, 24.V.2011 (CRB); Pentadactylos Range, Ftericha S., 800m, 1 ♀, stones in grassland, New records 17.V.2011 (CRB). LARNACA : Larnaca SW, Hala Sultan Tekke LARNACA : Agioi Vavatsinias, 700m, 1 ♂ 1 ♀ mosque, 50m, 1 ♂, in pitfalls in salt marsh, 15 - litter in Pinus forest, 28.IV.2010 (C RB); 28.V.2011 (CJVK). Dipo po tamos Reservoir, 150m, 1 ♀, herbs and stones bordering lake, 28.IV.2010 (CRB); Kornos Distribution: A rare species, actually known Picnic site, 350m, 2 ♀♀, beating in Pinus forest, from Italy, North Macedon ia and Romania. New 28.V.2011 (CRB); Stavrovouni Monastery, 350m, to Cyprus. 2 ♀♀, on bark of Pinus trees, 28.IV.2010 (CRB). LIMASSOL: Livadi tou Passia, E. Troodos, Theridion melanostictum O. Pickard - 1170m , 1 ♂, in peat marsh, 3.V.2010 (CRB); Cambridge, 1876 Prodromos E., near Prodomos dam, 1510m, 1 ♀, Theridion melanostictum ; Knoflach, 1998: 546. stones and litter in Pinus forest, 2.V.2010 (CRB); Xyliatos Dam, 2 ♀♀, stones in Pinus forest, Identification 4.V.2007 (CJVK). Levy & Amitai, 1982: 99, figs 32 - 37. NICOSIA: Lefka, 120m, 1 ♀, on walls of building, 18.V.2011 (CRB); Prodromos E., near Citations Prodomos dam, 1510m, 1 ♀, stones and litter in PAPHOS : Paphos (Knoflach, 1996). Pinus forest, 2.V.2010 (CRB); Strovolos, 1 ♀, in building, 1 - V.2016 (CMH). New rec ords PAPHOS : Agia, 2 ♀♀, on walls of small building None. (CJVK); Akamas, Avakas Gorge, 1 ♀, on rocky slope, 29.IV.20 07 (CJVK), 2 ♀♀, under stones in Distribution: North Africa, Israel, Canary gorge, 22.V.2011 (CRB); Akamas, Smigies, Islands, Cyprus, North America. 300m, 1 ♂ 12 ♀♀, handcatch on small building, 5.V.2207 (CJVK); Kelefos bridge, dam on Theridion ochreolum Levy & Amitai, 1982 Diarizos River, 550m, 1 ♀, under stones and beaten from shrubs, 21.V.2011 (CRB); Kinousa Identification E., Agi os Merkourios, 350m, 2 ♀♀, under stones Levy & Amitai, 1982: 99, figs 32 - 37. and litter in Pinus forest, 1.V.2010 (CRB); Kykkos Monastery, 1150m, 1 ♂ 7 ♀♀, on walls Citations in monastery, 19.V.2011 ( CRB); Kinousa SE, None. 400m, 2 ♀♀, under stones and litter in Pinus forest, 1.V.2010 (CRB); Livadi, 1 ♂ 2 ♀♀ , walls New records of small building, 28.IV.2007 (CJVK); Makounta, LIMASSOL: Agios Georgios E., Palaiomylos, Argaka Dam, 85m, 1 ♀, beating bushes, 1250m, 3 ♂♂, beaten in mixed Pinus and 27.IV.2007 (CJVK); Pano Platres E., Mesa Quercus forest, 4.V.2010 (CJVK); Akrotiri Potamos Waterfall s, 950m, 1 ♀, in litter in Pinus Peninsula N., 5m, 1 ♂, beaten in Eucalyptus forest, 4.V.2010 (CRB); Peristerona, Atichoullis forest, 29.IV.2010 (CJVK, CRB); Erimi, 2 ♂♂, gorge, 425m, 2 ♀♀, bea ting in maquis, 2.V.2007 under stones along dry riverbed, 18.IV.198 6 (CJVK); Polis, 2 ♂♂ 14 ♀♀, on walls of hotel, (NHMW); near river Limnatis SE. Alassa, 2 ♀♀, 25.IV.2007 (CJVK); Smigies, W. Neo Chorio, under stones in dry grassland, 5.V.2007 (C RS); 300m, 2 ♀♀, litter and stones in Pinus forest, Pano Platres, 1050m, 6 ♂♂, beaten near Mesa 30.IV.2010 (CRB). Potamos Waterfall s, 23.V.2011 (CJVK, CRB). NICOSIA : Agridi, near Dali, 1 ♀, in garden, Distribution: Until now only known from Crete . V.2012 (CDM); Cedar Valley Forest, 1180m, 2 New to Cyprus , where it app ears to be very ♂♂ 1 ♀, beaten in Cedrus forest, 19.V.2011 common. (CRB); Kakopetria S., Platania forest station, Theridion italiense Wunderlich, 1995 1100m, 14 ♂♂ 8 ♀♀, under stones in dense Pinus forest, 4.V.2010 (CJVK). Identification : https://araneae.nmbe.ch/ . PAPHOS : Agia, Stavros tis Psokas S., 590m, 1 ♂, Citations on walls in picnic site, 30.IV.2007 (CJVK); Agios None. Georgios E., Palaiomylos, 1250m, 13 ♂♂, beaten

Spiders of Cyprus 138

in mixed Pinus and Qu ercus forest, 4.V.2010 Wunderlich, 1995d: 764, figs 52 - 53 ( ♂); Demir, (CJVK); Akamas Peninsula, Neo Chorio, 200m, 2 Aktas & Seyyar, 2008: 65, figs 1 - 7 ( ♀). ♂♂ 5 ♀♀, beating bushes, 25.IV.2007, 1 ♀, 30.IV.2010, (CJVK); Akamas Peninsula, wood Citations near Agios Minas church, 5 ♀♀, beaten from None. climbers on pines, 2.V.2007 (CRS), 2 ♀♀, (CAR - S); Avakas Go rge, 55m, 1 ♂, stones along rivulet, New records 29.IV.2007, 1 ♀, 22.V . 2011 (CJVK); ca. 2 km. S FAMAGUSTA: R izokarpaso, 1 ♀, in Juniperus of Cape Geronnisos, 1 ♀, under stones in dry phoenicea forest, 12.III.2016 (CMH); Cape phrygana, 3.V.2007 (CRS); Evretou Reservoir, Greco, Ayia Napa, 35m, 1 ♀, stones in phrygana, Stavros tis Psokas river, 1 ♂, 20.III.2018 27.IV.2010 (CRB). (CKHK); Kourdaka, Ezousa river, 1 ♂, KERYNEIA : Agios Georgios, 1 ♀, under stone in 14 .III.2018 (CKHK); Pelathousa SE., 245m, 1 ♂, coastal scrub, 21.III.1998, coll. J. Murphy sweeping grassland, 30.IV.2007 (CJVK); Polis, (MMUE); Pentadactylos Rang e, Saint Hilarion 15m, 1 ♂, beating herbs, 27.IV.2007 (CJVK). Castle, 600m, 2 ♀♀, under stones, 17.V.2011 (CJVK). Distribution: Israel. New to Cyprus. NICOSIA: Alona, 1 ♀, mixed forest, 27.II.2016 (CMH). LIMASSOL: Limassol, 2 ♀♀, 24.II.1933, G. A. Family Thomisidae Mavromoustakis leg. (BMNH); Pissouri, 1 ♀, in grass, 24.III.1998, coll. J. Murphy (MM UE); Bassaniodes Pocock, 1903 Millomeris Waterfalls, near Pano Platres, 1 ♀, Recently, the large crab spider genus Xysticus was under rock, 28.IV.2007 (CRS). split up (Breitling, 2019), and the Cypriot species PAPHOS : Akamas Peninsula, track along the sea, Xysticus caperatus , X. pseudorectolineus , X. 10m, 2 ♀♀, stones in Juniperus phrygana, tenebrosus , and X. tristrami are no longer known 30.IV.2010 (CJVK, CRB); Akamas Peninsula, under their familiar name Xysticus , but as Avgas E stuary, 1 ♀, under stones, 31.III.1 997 Bassinioides . The other speci es occurring in (CJB); Akamas Peninsula, Cape Geronnisos, 25m, Cyprus ( Xysticus cor, X. edax , X. laetus , X. 1 ♀, stones in maquis, 29.IV.2007 (CJVK); pseudoluctuosus and X. xerodermus ) remain into Akamas Peninsula, Lara Bay, 25m, stony Xysticus . Juniperus maquis near the beach, 29.IV.2007 (CJVK); Akamas Peninsula, 1 ♀, in coastal Bassaniodes caperatus ( Simon, 1875 )ba grassland, 22.III.1998, coll. J. Murph y (MMUE); Baths of Aphrodite, 1 ♀, under stone in coastal Identification scrub, 18.III.1998, coll. J. Murphy (MMUE); Levy, 1976: 21, figs 33 - 40. Kinousa, 2 ♀♀, in pine wood, 25.III.1998, coll. J. Murphy (MMUE); Kato Paphos, 1 ♀, under Citations stones near hotel, 1.IV.1997 (CJB); near Kidasi None. village, Diarizo s River, 1 ♀, under bark of plane tree, 1.V.2013 (CAR - S); New records Kouklia, Sanctuary of Aphrodite, 1 ♀, 8.IV.1997 KERYNEIA : Pentadactylos Range, Lapithos S. , (CJB); Makounta, 1 ♀, under stones in scrub, 800m, 1 ♂, stones in Pinus forest, 17.V.2011 19.III.1998, coll. J. Murphy (MMUE); (CJVK). Mavrokolympos Reservoir, NE . Potima, 150m, 1 FAMAGUSTA : Cape Greco, Ayia Napa, 35m, 1 ♀, under stones alon g lake, 22.V.2011 (CJVK); ♂, stones in phrygana, 27.IV.2010 (CJVK); Cape Mavrokolympos Reservoir, 3 ♀♀, 13.III.2018 Greco, 1 ♀, under stones in phrygana, 3.V.2013 (CKHK). (CAR - S). Distribution: Turkey, Greece (Lesbos and Distr ibution Chios). New to Cyprus. Israel, Turkey. New to Cyprus. Bassaniodes pseudorectilineus (Wunderlich, 1995) Bassaniodes tenebrosus (Silhavy, 1944)

Identification Identification Wunderlich, 1995d: 762, figs 43 - 44.

Spiders of Cyprus 139

C itations Identification None. Marusik et al. 2005: 156 , figs 1 - 4, 15 - 16, 19, 25 - 26, 30, 33 - 34, 40, 45. New records LARNACA : Stavrovouni Monastery, 350m, 1 ♀, Citations on bark of Pinus tree , 28.IV.2010 (CJVK). None. LIMASSOL: Troodos Mountain, 1 ♂ 1 ♀, G. Mavromoustakis leg., without date (BMNH). New records LIMASSOL : Mount Olympos (Chionistra), Distribution: Bulgaria, North Macedonia, Gree ce 1850m, 2 ♂♂, pitfalls in Pinus nigra forest, (Lesbos and Chios), Israel. New to Cyprus. 26.IV. - 5.V.2007 (CJVK); Troodos 3 km S, 1600m, 3 ♂♂, pitfalls in maquis, 26.IV. - Bassaniodes tristrami (O. Pickard - Cambridge, 5.V.2007 (CJVK). 1872) Distribution: Albania, Bulgaria, Ukraine, Identification Turkey. New to Cyprus. Wunderlich, 1995d: 762, figs 43 - 44. Diaea livens Simon, 1876 Citations Without precise locality (Kulczyński, 1908, sub X. Identification : https://araneae.nmbe.ch/ . tristrami ). Citations Comments None. The identification of the female s pecimens mentioned below is a bit doubtful, as no adult New records male ss were collected. Identification based on NICO SIA : Kakopetria S., Platania forest station, females, are difficult to distinguish them from 1100m, 1 ♀, beating in dense Pinus forest, Bassanioides pseudorectolineus . The capture of 4.V.2010 (CJVK). males should confirm the presenc e of the species PAPHOS : Agios Georgios E., Palaiomylos, in the island. 1250m, 3 ♂♂, beaten in mixed Pinus and Levy (1976), without examining the material, Quercus forest, 4.V.2010 (CJVK). considered the record of Bassanioides tristrami of Kulczyńksi (1908, sub Xysticus ) as a Distribution: USA, Central Europ e, Greece to misidentification and attributed it to Bassanioides Azerbaijan. New to Cyprus. ferus . Kulczyński’s excell ent drawing (fig. 14, epigyne longer than wide) does not look like Heriaeus graminicola (Doleschall, 1852) Levy ’s figures at all (figs 13, epigyne wider than long). They look on the contrary very well like Identification those of Bassanioides pseudorectolineus (see Loerbroks, 1983 : 107, figs 15, 29 - 36. Demir et al., 2008: 65, figs 1 - 7, sub Xysticu s p. ), the commonest Bassanioides species in Cyprus according to our collections. So, Kulczyński’s Citations citation (1908) remains doubtfull. None.

New records New records LIMASSOL : Panagia Glossa Monastery near NICOSIA : Ced ar Valley Forest, 1050m, 2 ♂♂, Kellaki, 1 ♀, under stones in cultivated land, beating in Cedar forest, 19.V.2010 (CRB). 12.IV.1986 (NHMW) ; Troodos S., 1350m, 1 ♀, stones in mixed Quercus - Pinus forest, 3.V.2010 (CJVK). Distribution: Central and South - East Europe, Balkans, Russia, Turkey, Caucasus. New to Distribution: Crete, Turkey, Saudi Arabia to Cyprus. Central Asia. New to Cyprus. Heriaeus simoni Kulczyński, 1903 Cozyptila thaleri Marusik & Kovblyuk, 2005

Spiders of Cyprus 140

Identification Kelefos bridge, dam on Diarizos River, 550m, 1 Loerbroks, 1983 : 107, figs 15, 29 - 36. ♀, beate n from shrubs, 21.V.2011 (CRB); Kykkos Monastery, 1150m, 2 ♀♀, on bushes around Citations monastery, 19.V.2011 (CJVK); Pelathousa E., None. 245m, sweeping grassland, 30.IV.2007 (CJVK); Peristerona, Atichoullis gorge, 425m, 1 ♂, beating New records in maquis, 2.V.2007 (CJVK). KERYNEIA : Pentadactylos Range, Saint Hilarion Castle W., 600m, 1 ♂, beaten from shrubs in Dis tribution: Greece, Turkey, Israel, Lebanon, Pinus forest, 17.V.2011 (CJVK). Central Asia. New to Cyprus.

Ozyptila rigida (O. Pickard - Cambridge, 1872) Distribution: Palearctic. New to Cyprus.

Identification Misumena vatia (Clerck, 1757) Levy, 1985b: 65, figs 96 - 97, (sub. O. rigida ); 66,

Identification : https://araneae.nmbe.ch/ . figs 98 - 99 (sub O. complicata ).

Citations

None Citations

None. N ew records

LIMASSOL : Fasouri, Akrotiri Peninsula, 1 ♀, in New records reed litter, 27.IV.2007, 3 ♂♂ 2 ♀♀, 4.V.2007, PAPHOS : Evretou Reservoir, 350m, 1 ♀, shrubs det. D. Logunov (CAR - S). and grassland around lake, 1.V.2010 (CJVK).

Distribution: Israel, Saudi Arabia, Azerbaijan. Distribution: Holarctic. New to Cyprus.

Ozyptila tricoloripes Strand, 1913 Monaeses israeliensis Levy, 1973 Oxyptila rigida ; Kulczyński, 1908: 70, pl. 2, figs

15 (misidentification according to Levy, 1975). Identification

Levy, 1973: 111, figs 8 - 11. Identification

Levy, 1975: 167, figs 22 - 23 ( ♂, sub O. Citations tricoloripes ); Levy, 1975: 165, figs 19 - 21 ( ♀, sub None. O. pickardi ).

New records KERYNEIA : Saint Hilarion castle, 610m, 1 ♀, 1 Citations Without precise locality (Kulczyński, 1908) . suba dult ♀, beaten in Pinus forest, 22.XI.2010 (CJVK, CRB). New records LIMASSOL: Fasoula, 1 ♀, olive garden with dry FAMAGUSTA : Salamis S., Silver Beach, 5m, 4 stone walls, 16.IV.1986 (NHMW). ♀♀, in pitfalls in salt marsh, 15 - 27.V.2011 NICOSIA : Agridi, near Dali, 1 ♀, in yard, (CRB). VIII.2009, 1 ♂, in garden, 1V.2010, 1 ♂, in LARNACA: Dekeleia Military Base, 1 ♀, garden, IV.2012 (CDM); Elia - Ango lemi road, VI.1958, M. Connelly leg. (BMNH); Larnaca 100m, 1 ♀, rubbish in dry riverbed, 18.V.2011 SW . , Hala Sultan Tekke mosque, 50m, 1 ♀, (CJVK); Masari S., Merika River, 120m, 1 ♂, on pitfalls in sal t marsh, 15. - 28.V.2011 (CJVK). shrubs in wasteland, 24.V.2011 (CRB). LIMASSOL: Amathus, 1 ♀, on house walls, PAPHOS : Akamas Peninsula, Lara Bay, 1 ♀, C. 9.IV.1986 (NHMW); road Kaminaria - Lemithou, 1 Saure leg., 18.VI.2013 (CKHK); Evretou ♀, pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 Reservoir, 350m, 1 ♂, beaten from shrubs and (CMH). grassland around lake, 1.V.2010 (CRB); Gialia, NICOSIA : Dali, 13, Agios Andronikos, 2 ♂♂, in 110m, 1 ♂, sweeping Cystus , 28.IV.2007 (CJVK); garden, IV.2007 (CDM); Elia - Angolemi road, Loukrounou E., 140m, 1 ♂ 1 ♀, sweeping herbs bordering cultivated fields, 1.V.2007 (CJVK);

Spiders of Cyprus 141

100m, 1 ♀, under rubbish in dry riverbed, NICOSIA : Lefka NW, 120m, 1 subadult ♀, 18.V.2011 (CRB). beaten from shrubs in fruit plantation, 18.V.2011 PAPHOS: Baths of Aphrodite, 5m, 1 ♀, pitfalls (CRB). under Eucalyptus trees bordering fields, 25.IV. - LIMASSOL : Akrotiri Peninsula, near Fasouri 5.V.2007 (CJVK); Gialia, 1 ♀, in pine wood, reed beds, 3 ♂♂, swept from flowering herbs and 25.III.1998, coll. J. Murphy (MMUE); Kato grass, 27.IV.2007 (CAR - S), 1 ♂ (CDM), 1 ♂ , 4.V.2007 (CAR - S). Paphos, 1 ♀, seevin hay, 4.IV.1997 (CJB); PAPHOS : Asprokremmos Dam, N. Kouklia, Kinousa, 1 ♀, in pine wood, 23.III.1998, coll. J. 150m, 8 ♂♂ 1 su badult ♀, herbs and stones along Murphy (MMUE); near Kritou Terra, 1 ♀, under lake, 23.V.2011 (CJVK, CRB); Evretou stone in dry fallow field, 17.IV.2017 (CAR - S); Reservoir, 350m, 2 ♂♂ 1 ♀, beaten from shrubs Neo Chorio, 1 ♀, under stone, 30.III.1998, coll. J. around lake, 1.V.2010 (CJVK, CRB). Murphy (MMUE); Pege ia, Pikni Forest, 366m, 2

♀♀, pitfalls in pine forest, 11.III. - 23.III.2018 Distribution: Palaearctic, St Helena, South Africa. (CKHK). New to Cyprus.

Distribution: Israel, Cyprus, Saudi Arabia, Synema globosum (Fabricius, 1775) Azerbaijan. Identification : https://araneae.nmbe.ch/ . Ozyptila sp. Citations A new Ozyptila species will be described in a None. separate paper. New records Pistius truncatus ( Pallas, 1772) FAMAGUSTA : Flamoudi W., 25m, 1 ♀, beaten in phrygana near the beach, 25.V.2011 (CRB); Identification : https://araneae.nmbe.ch/ . Kantara E., 610m, 1 ♂ 1 ♀, beaten from shrubs

and stones in open Pinus forest, 25.V.2011 Citations None. (CRB); Karpasia Peninsula, Apostolos Andreas Monastery, 50m, 1 ♀, stones in wasteland, New records 16.V.2011 (CJVK). NICOSIA : Cedar Valley Forest, 1 ♂, swept from KERYNEIA : Keryneia , near the town, 1 ♀, herbs, 29.IV.2007 (CAR - S); Dali, 13, Agios Juniperus phoenicea forest, 30.IV.2016 (CM H); Andronikos, 1 sub ad ult ♂, in garden, V.2007 Lapithos S., 800m, beating in clearing in Pinus (CDM). forest, 17.V.2011 (CJVK); Panagia Apsinthiotissa LIMASSOL : Pano Platres, track near Caledonia Monastery ruins, 350m, 1 ♀, stones in clearing in Waterfall, 1 ♂, beaten from shrub at edge of Pinus forest, 24.V.2011 (CRB). P inus nigra forest, 26.IV.2007 (CAR - S). LARNACA: Dekeleia Military Base, 2 ♀♀, 11.VII.1958, M. Connelly leg. (BM NH); Distribution: Europe. New to Cyprus. Stavrovouni Monastery, 1 ♂ 1 ♀, 3 immatures, in phrygana, 26.IV.1982, coll. A. Stubbs (MMUE). Runcinia grammica (C. L. Koch, 1837) LIMASSOL : Agios Tychonas, 1 ♂, under stones

near house walls, 19.IV.1986 (NHMW); Pano Ide ntification : https://araneae.nmbe.ch/ . Platres, near top of road to Millomeris Waterfalls, Citations 1 ♂, swept from grass and herbs in abandoned None. vineyard, 28.IV.2007 (CAR - S). NICOSIA : Agridi, near Dali, 3 ♂♂, in garden, New records V.2004, 1 ♀, in garden, IV.2011 (CDM); between LARNACA : Larnaca salt lake, 5m, 1 ♂, on herbs Cedar Valley and Stavros tis Psokas, 1 ♂, under in salt marsh, 27.IV.2010 (CRB). stones in Pinus nigra forest, 29.IV.2007 (CAR - S); Evre tou Kampos NE . , Xeros River, 550m, 2 ♂♂ 1

Spiders of Cyprus 142

♀, beaten from shrubs, 20.V.2011 (CJVK); Lefka None. NW, 120m, 4 ♂♂ 5 ♀♀, beaten from shrubs in fruit plantation, 18.V.2011 (CJVK, CRB); New records LIMASSOL: Akrotiri Peninsula N., 5m, 1 ♂, Limnitis River, Kato Pyrgos SE . , 200m, 1 ♂, beating in Eucalyptus forest, 29.IV.2010 (CRB). stones and litter in Pinus forest, 2.V. 2010 NICOSIA: Agridi, near Dali, 1 ♂, in ga rden, (CJVK); Masari S., Merika River, 120m, 1 ♀, VI.2010 (CDM). stones in wasteland, 24.V.2011 (CRB).

PAPHOS : Akamas Peninsula, Avakas Gorge, 1 Distribution: Mediterranean, Africa, Yemen, ♀, under stone, 27.III.1998, coll. J. Murphy Socotra, Seychelles. New to Cyprus. (MMUE); Asprokremmos Dam, N. Kouklia, 150m, 2 ♂♂ , beaten from herbs along lake, 23.V.2011 (CRB); Evretou E., 200m, 2 ♂♂, Thomisus onustus Walckenaer, 1805 Thomisus onustus ; Logunov, 2013: 5. sweeping low herbs, 1.V.2007 (CJVK); Evretou

Reservoir, 350m, 1 ♂ 1 ♀, shrubs and grassland Identification : https:// araneae.nmbe.ch/ . around lake, 1.V.2010 (CJVK); Kelefos bridge, dam on Diarizos River, 550m, 1 ♀, beaten from Citations shrubs, 21.V.2011 (CRB); 2 km S . of Kouklia, 1 LIMASSOL: Akrotiri Phasouri (Logunov, 2013); ♀ 1 immature, in riverine marsh, 30.IV.1982, coll. Plano Platres (Logunov, 2013). A. Stubbs (MMUE); Ktima, 3 ♂♂ 1 ♀, G.

Mavromoustakis leg., without date (BMNH); New records Vretsia W., near Kelefos Bridge, 650m, 1 ♂, FAMAGUSTA : Salamis ruins, 50m, 4 ♂♂ 3 beaten from shrubs, 19.V.2011 (CRB). ♀♀, beaten from low trees, 15.V.2011 (CRB).

Distribution: Palaearct ic. New to Cyprus. LARNACA : Agioi Vavatsinias, 700m, 1 ♀, b eaten in Pinus forest, 28.IV.2010 (CRB); Kato Synema plorator (O. Pickard - Cambridge, 1872) Drys, 1 ♂, Quercus litter, 26.III.2016 (CMH); Larnaca, 1 ♀, C. Glasynet leg., without date Identification: https://araneae.nmbe.ch/ . (BMNH).

LIMASSOL: Akrotiri Peninsula, 1 ♂, 27.IX.2007 Citations (CDM); Armenochori, 1 ♀, city area, 14.IV.1986 None. (NHMW). New records NICOSI A : 13, Agios Andronikos, Dali, 1 ♀, in LIMASSOL: Akrotiri Peninsula, near British garden, V.2008 (CDM); Agios Sozomenos , near Airbase , 1 ♀, swept in pine wood, 4.V.2007 Dali, 1 ♂, swept in dry grassland, 30.IV.2007 (CRS); Troodos S., road to Pano Platres, 1 ♂, (CRS); Agridi, near Dali, 1 ♀, in garden, IV.2004, under stones in P. nigra forest, 1.V.2007 (CRS). 1 ♂, in field, IV.2007, 1 ♀, on house wall, NICOSIA : Dali, 13 Agios Andronikos, 1 ♀, in 5.V.2009, 1 ♂, in garden, III. 2010, 1 ♂, in garden, garden, IV.2007 (CDM); Cedar Valley Forest, VI.2010, 2 ♂♂, in garden, V.2011, 1 ♂, in garden, V.2012, 1 ♂, in garden, 12.IX.2012 1180m, 1 ♂, beaten in Cedrus forest , 19.V.2011 (CDM); Elia - Angolemi road, 100m, 4 ♂♂ 1 ♀, (CRB); Evretou E., 200m, 1 ♂, sweeping low stones in grassland, rubbish in dry riverbed, herbs, 1.V.2007 (CJVK); Pelathousa SE, 245m, 1 18.V.2011 (CJVK); Kampos NE, Xeros River, ♂ 1 ♀, beating Cystus in Pinus forest, 28.IV.2007 550m, 2 ♀♀, beaten from shrubs, 20.V.2011 (CJVK). (CRB); Kapedes S., 600m, 1 ♂, on herbs along rivulet, 5.V.2010 (CRB). Distribution: Greece, Bulgaria, Israel, Lebanon, PAPHOS : Agios Mamas ruins, Gialia E., 215m, 1 Russia, Central Asia. New to Cyprus. ♂, beating Cystus , 28.IV.2007 (CJVK); Asprokremmos Dam, N. Kouklia, 150m, 3 ♂♂, Thomisus citrinellus Simon, 1875 herbs along lake, 29.IV.2010 (CJVK , CRB); Avakas gorge and surroundings, 1 ♂ 1 ♀, Identification 22.III.2018 (CKHK); Evretou Reservoir, 350m, 2 Levy, 1973: 128, figs 43 - 46. ♂♂, shrubs and grassland around lake, 1.V.2010 Citations (CJVK, CRB); Gialia, 1 ♂, in pine wood,

Spiders of Cyprus 143

25.III.1998, coll. J. Murphy (MMUE); Gialia, LIMASSOL: Akamas Peninsula, Agios Minas, 1 110m,1 ♂, sweeping Cystus , 28 .IV.2007 (CJVK); ♂, in meadow with tall grass, 2.V.2007 (CAR - S); Kato Paphos, 1 ♂, beating tall herbs near hotel, Akrotiri Peninsula, near British Airbase, 1 ♀, 4.IV.1997 (CJB); Kourdaka, Ezousa river, 1 ♂, swept in pine wood, 4.V.2007 (CAR - S). 14.III.2018 (CKHK); near Kritou Terra, 1 ♀, NICOSIA : Kapedes, 1 ♂, pitfalls in Pinus brutia under stone in dry fallow field, 17.IV.2017 (CAR - forest, 17.I. - 23.IV.2016 (CMH); Lefka NW, S); Ktima, 6 ♀♀, G. Mavromoustakis leg., 120m, 1 ♂ 1 ♀, beaten from shrubs in fruit without da te (BMNH); Kykkos Monastery, plantation, 18.V.2011 (CJVK, CRB); Oikos N., 1150m, 4 ♂♂ 2 ♂♂, on bushes around monastery, Marathassa Dam, 550m, 1 ♂, stones and beat en 19.V.2011 (CJVK); Pelathousa SE., 245m, 1 ♂, from shrubs around lake, 20.V.2011 (CRB); sweeping grassland, 30.IV.2007 (CJVK); Polis, Xerargaka picnic site, 350m, 2 ♂♂, beaten from 15m, 1 ♀, herbs and stones around building, shrubs and stones in Pinus forest, 20.V.2011 27.IV.2007 (CJVK). (CRB). PAPHOS : near Avakas Gorge; 1 ♀, beaten from Distribution: Palearct ic. junipers on rocky hillside, 14.IV.2017 (CAR - S); Kato Akourdaleia, 1 ♀, in garden, VIIII.2005 Thomisus zyuzini Marusik & Logunov, 1990 (CDM); Makounta, Argaka Dam, 85m, beating in

Identification maquis, 27.IV.2007 (CJVK); Smigies, W. Neo Marusik & Logunov, 1990: 34, figs 6 - 8 ( ♀); Chorio, 300m, 1 ♀, litter and stones in Pinus Marusik & Logunov, 1995: 144, figs 19 - 20 ( ♂). forest, 30.IV.2010 (CRB). Distribution: Mediterranean. New to Cyprus. Citations None. Xysticus cor Cane strini, 1873

New records Identification FAMAGUSTA : Karpasia Peninsula, Apostolos Thaler & Notflatscher, 1990: 177, figs 42 - 45. Andreas Monast ery, 50m, 1 ♂, stones in wasteland, 16.V.2011 (CRB). Citations None. Distribution: Turkey, Saudi Arabia to Central New records Asia. New to Cyprus. NICOSIA: Cedar Valley Forest, 1 ♂, swept from

herbs, 29.IV.2007 (CRS). Tmarus piochardi (Simon, 1866)

Distribution: Southern Europe, Iran. New to Identification Cyprus. Levy, 2007: 19, figs 51 - 52.

Xysticus edax (O. Pickard - Cambridge, 1872) Citations

None. Identificication

Levy, 1976: 21, figs 33 - 40. New records

FA MAGUSTA : Kantara E., 610m, 1 ♂ 1 ♀, Citations beaten from shrubs and stones in open Pinus None. forest, 25.V.2011 (CRB); Komi Kebir , 150m, 1 ♂, beaten from shrubs, 26.V.2011 (CJVK). KERYNEIA: Panagia Apsinthiotissa Monastery ruins, 350m,1 ♀, stones in clearing in Pinu s New records NICOSIA: Agridi, near Dali, 1 ♀, in field, forest, 24.V.2011 (CRB). V.2008, 1 ♀, V.2009 (CDM); Dali, 13, Agios LARNACA : Kornos Picnic site, 350m, 1 ♂, Andronikos, 1 ♀, in garden, I V. 2007 (CDM); beaten in Pinus forest, 28.V.2011 (CRB). Lymbourena Hill, Dali, 1 ♀, in field, V.2004 (CDM); Kapedes S., 600m, 1 ♀, herbs and stones

Spiders of Cyprus 144

along rivulet, 5.V.2010 (CRB). Nicosia, 1 ♀, Citations 30.IV.1957, G. P. Georghiou leg. (BMNH). None. PAPHOS : Agios Merkourios, 250m, 1 ♀, sweeping grassland, 27.I V.2007 (CJVK); Akamas New records Peninsula, track along the sea, 10m, 1 ♀, stones in NICOSIA: Agridi, near Dali, 1 ♂, in grass in Juniperus phrygana, 30.IV.2010 (CRB); garden, III.2014 . leg. D. McCowan (MMUE). Drouseia, 539m, 1 ♂, in p hrygana, 23.III.2018 (CKHK); Limnitis River, Kato Pyrgos SE, 200m, Distribution: Turkey, Tajikistan. New to Cyprus. 1 ♀, stones and litter in Pinus forest, 2.V.2010 (CJ VK); Mavrokolympos Reservoir, 1 ♂, Xysticus xerodermus Strand, 1913 13.III.2018 (CKHK); Peristerona, Atichoullis gorge, 425m, 1 ♂, beating in maquis, 2.V.2007 Identification (CJVK); Pegeia, Pikni Forest, 346m, 2 ♂♂, Levy, 1976 : 27, figs 49 - 52 pitfalls in former carob plantation, 11.III. - 23.III.2018 (CKHK); Pegeia, Pikni Forest, 374m, Citations 1 ♂, pitfalls in open ruderal site, 14.III. - None. 23.III.2018 (CKHK); near Souskiou, 1 ♀, under stones in dry grassland, 1.V.2013 (CDM). New records KERYNEIA : Kazafani , 150m, 1 ♀, under stones Distribution: Israel, Turkey, New to Cyprus. in olive grove, 23.XI.2010 (CRB). NICOSIA: Athalassa National Forest Park, 1 ♂, Xysticus laetus Thorell, 1875 Pinus brutia forest, 21.II.2016, M.

Identification Hadjiconstantis leg. (CRB); Lakatamia, 1 ♂, in Logunov , 2006: 63, figs 5, 6, 13 - 20. olive grove, 1.I. - 1.X. 2016, M. Ha djiconstantis leg. (CRB); Kapedes S., 600m, 2 ♀♀, herbs and Citations stones along rivulet, 5.V.2010 (CRB). None. LIMASSOL: Fasouri marsh, 1 ♀, in low grass, 17.VI.2016 (CMH). New records NICOSIA : Agridi, near Dali, 1 ♀, in garden, Distribution: Greece, Turkey, Israel. New to III.2010 (CDM); Oikos N., Xerargaka picnic site, Cyprus. 350m, 1 ♀, beaten from shrubs and stones in Pinus forest, 20.V.2011 (CJVK). LIMASSOL : Akrotiri Peninsula, Fasouri reed Family Titanoecidae beds, 1 ♂ 5 ♀♀, in reed litter, 27.IV.2007 (CAR - S), det. D. Logunov; same site, 5 ♀♀ (CRS), 4 Nurs cia albomaculata (Lucas, 1846)

♀♀, 4.V.2007 (CAR - S). Identification : https://araneae.nmbe.ch/ . PAPHOS : Agios Mamas ruins, Gialia E., 215m, 1 ♀, beating Cystus , 28.IV.2007 (CJVK); Evretou Citations Reservoir, 35 0m, 1 ♀, stones in grassland around None. lake, 1.V.2010 (CJVK); Evretou Reservoir, Stavros tis Psokas river, 1 ♂, 20.III.2018 New records (CKHK); Pelathousa SE., 245m, 1 ♀, sweeping LARNACA : Achna Reservoir, 50m, 3 ♂♂ 1 ♀, in grassland, 30.IV.2007 (CJVK). pitfalls in marshy area around lake, 15 -

27.V. 2011 (CRB); Larnaca SW, Hala Sultan Distribution: Eastern Mediterranean region, from Italy t o the Caucasus. New to Cyprus. Tekke mosque, 50m, 4 ♂♂, in pitfalls in salt marsh, 15 - 28.V.2011 (CRB). Xysticus pseudoluctuosus Marusik & Logunov, NICOSIA: Dali, 13, Agios Andronikos, 1 ♀, in 1995 garden, V.2007 (CDM); Limnitis, 1 ♀, Pinus brutia forest, 13.V.2016 (CMH); Lympia Identification Reservoir, Lympi a, 1 ♂ 4 ♀♀, under stones below Marusik & Logunov, 1995: 147, figs 26 - 27. dam wall, 9.V.2013 (CDM); Marathasa Dam, 1 ♂,

Spiders of Cyprus 145

pitfalls in riverside vegetation, 7.V. - 10.VII.2016 NICOSIA: Lympia Reservoir, Lympia, 1 ♀, under (CMH). stones below dam wall, 9.V.2013 (CDM).

Distribution: Mediterranean region to Central Comments Asia. New to Cyprus. In th e absence of males, these females cannot be identified to species level. Species occurring in Nurscia albosignata Simon, 1874 Turkey and possible candidates are Titanoeca Nurscia flavipes Simon, 1874: 235. caucasius Dunin 1985, T. incerta Nosek, 1905 and T. schineri L. Koch, 1872. Identification Lehtinen, 1967: 253, figs 430 - 431, 433. Kovblyuk et al . 2016: 129, figs 315 - 317, 319, 321, 324 - 325, Family Trachelidae 329 - 331. Trachelas mi nor O. Pickard - Cambridge, 1872 Citations Without precise locality (Simon, 1874). Identification Simon, 1932: 958, 977, figs 1498 - 1499; New records Bosselaers et al., 2009: 18, figs 4, 9 - 15. FAMAGUSTA : Flamoudi W. , 25m, 3 ♂♂ 1 ♀, stones and phrygana near the beach, 25.V.2011 Citations (CRB). None.

LARNACA : Achna Reservoir, 50m, 2 ♂♂, stones and litter bordering lake, 16.V.2011, 1 female, 21.XI.2010, (CRB). New records LIMASSOL: Agios Tychonas, 3 subadult ♂♂ 2 LARNACA: Achna Reservoir, 50m, 2 ♀♀, under juveniles, under stones near house walls, stones and litter bordering lake, 27.V.2011 19.IV.1986 (NHMW); Amathus, 2 ♂♂ 6 subadult (CRB), 1 ♂, pitfalls at same site, 15 - 27.V.2011 ♂♂ 4 juvenile s, under stones in dry valley, (CRB).

6.IV.1986, 2 juveniles, 6.IV.1986, 1 subadult ♂ , Distribution: Mediterranean to Central Asia, 7.IV.198 6, 1 juvenile, on house walls, 9.IV.1986 West Africa. New to Cy prus. (NHMW); Armenochori, 3 subadult ♂♂ 2 juveniles, under sto nes of a dry stone wall, 14.IV.1986 (NHMW); Fasoula, 1 juvenile, olive Family Uloboridae garden with dry stone walls, 16.IV.1986 (NHMW); Germasogeia, along the stream below Hyptiotes paradoxus (C. L. Koch, 1834) the dam, 2 juveniles, under stones, 10.IV.1986 Identification : https://araneae.nmbe.ch/ . (NHMW); Parekklisia, 3 subadult ♂♂ 6 juveniles, under stones in riverbed, 13.IV. 1986 (NHMW). Citations NICOSIA: Pyrgos, 1 subadult ♂ 2 juveniles, dry None. grassland, 15.IV.1986 (NHMW). PAPHOS : Mavrokolympos Reservoir, NE New records Potima, 150m, 1 ♀, under stones along lake, NICOSIA: Agia Marina, 1 ♀, Pinus brutia forest, 22.V.2011 (CRB). 7.III.2016 (CMH). LIMASSOL: Pelendri, 1 ♂ 2 ♀♀, on herbs in

Pinus brutia forest, 11. XII. 2018 (CMH). Distribution: Bulgaria, Cyprus to Central Asia. PAPHOS: Kinousa, 1 ♀, in pine wood,

23.III.1998, coll. J. Murphy (MMUE). Titanoeca sp.

Distribution : Palaearctic. New to Cyprus. Records

LARNACA : Achna Reservoir, 50m, 1 ♀, under Uloborus plumipes Lucas, 1846 stones and litter bordering lake, 21.XI.2010 (CRB).

Spiders of Cyprus 146

Identification Mandria, 1 ♂, in scrub, 5.V.1982, coll. A. Stubbs (MMUE). Identification : https://araneae.nmbe.ch/ . Distribution : Palaearctic. New to Cyprus. Citations None. Family Zodariidae New records NICOSIA : Agridi, near Dali, 2 ♀♀, in garden, Lachesana blackwalli (O. Pi ckard - Cambridge, IV.2004 (CDM). 1872) LIMASSOL: Akrotiri Peninsula, near British Laches blackwallii ; Kulczyński, 1908: 59. Airbase, 1 ♀, swept in pine wood, 4.V.2007 Lachesana blackwalli ; Thaler & Knoflach, 2004: (CAR - S); Kolossi, N . of the castle, 2 ♀♀, under 41. Identification stones, 11.IV.19 86 (NHMW). Thaler & Knoflach, 2004: 41, figs 4, 12, 14, 17, PAPHOS : Akamas Peninsula, Neo Chorio, 1 ♀, 19, 22, 25; swept from phrygrana and grass, 2.V.2007 (CAR - S); Oikos, 700m, 1 subadult ♀, small creek, Citations 23.IV.2017, S. Huber leg. (CSH); 9 km N. Pano Without further locality (Kulczyński, 1908). Panagia, 1050m, 1 subadu l t ♂, 1 ♀, 1 subadult ♀, PAPHOS: Paphos, lowland near coast (Thaler & in litter near spring, 19.V.2011 (CRB); Pomos Knoflach, 2004); Agios Georgios (Thaler & Dam, 3 ♀♀, Pinus brutia forest, 27.VII.2016 Knoflach, 2004). (CMH). New records Distribution : Old world. New to Cyprus. FAMAGUSTA : Karpasia Peninsula, Apostolos Andreas Monastery, 50m, 1 ♂, stones in Uloborus walckenaerius Latreille, 1806 wasteland, 16.V.2011 (CJVK). KERYNEIA : Larn akas tis Lapithou, 1 subadult ♀, Identification : https://araneae.nmbe.c h/ . stones in grassland, 17.I.2016 (CMH).

Citations LIMASSOL: Mount Olympos (Chionistra), 1 None. subadult ♀, pitfalls in open Juniperus foetidissima forest, 6.IV. - 8.VI.2014 (CMH). New records NICOSIA : Agios Sozomenos , near Dali, 1 FAMAGUSTA : Prastio S., Agios Andronikos, juvenile, under s tones in dry grassland, 30m, 3 ♀♀, under stones and herbs in Eucalyptus 30.IV.2007 (CAR - S). forest, 27.V.2011 (CRB); Protaras, Profitis Elias, PAPHOS: Lysos E., 550m, 1 subadult ♀, while 1 ♂, 2005 (CDM); Salamis ruins, 30m, 1 ♀, digging for Nemesia , 27.IV.2017, S. Huber leg. (CSH). stones in ruins, 17.III.2011 (CRB).

LARNACA : Pervolia, 1 ♀, VII.2005 (CDM). Distribution: Crete, Cyprus, Turkey, Israel, LIMASSOL : Akrotiri Peninsula, near Agios Lebanon. Nikolaos Monastery, 25m, 1 ♀, in grassy ditch, 23.III.2011 (CRB); Akrotiri Peninsula, near Palaestina expolita O. Pickard - Cambridge, British Airbase, 1 ♀, swept in pine wood, 1872 4.V.2007 (CRS); Mount Olympos ( Chionistra), Identification Levy, 1992: 69, figs 1 - 12. just below summit, 1 ♂, on shrub in Pinus nigra forest. 28.IV.2007 (CRS). NICOSIA : Agridi, near Dali, 1 ♂, in garden, Citations V.2011 (CDM); Lakatamia, in olive grove, 1 ♀, None. 1.I. Pervoli - 1.X. 2016 (CMH). PAPHOS : Avakas gorge, 50m, 1 ♀, on rocks in New records g orge, 30.IV.2010 (CJVK); near Paphos, 1 ♀, in FAMAGUSTA : Kantara E. 650m, 1 ♂, in pitfalls marshy area, 1.V.1982, coll. A. Stubbs (MMUE); in Pinus forest, XI.2010 - III.2011 (CRB);

Spiders of Cyprus 147

Salamis S., Silver Beach, 5m, 1 ♂, in pitfalls in salt marsh, 15 - 27.V.2011 (CR B). Distribution LARNACA : Achna Reservoir, 50m, 1 ♀, under Greek Peloponnisos; first citation in Cyprus. stones and litter bordering lake, 27.V.2011 (CJVK); Kornos picnic site, 350m, 1 ♂, under Zodarion granulatum Kulczyński, 1908 stones, 28.V.2011 (CRB); Larnaca SW, Hala Zodarium granulatum Kulczyński, 1908: 59, pl. 2, Sultan Tekke mosque, 50m, 66 ♂♂ 7 ♀♀, in figs 4, 10 (descr. ♂); Bosmans, 2009: 274. pitfalls in salt marsh, 15 - 28.V.201 1 (CJVK, CRB). Identification LIMASSOL: Akrotiri Peninsula, edge of central Bosmans 2009: 275, figs 170 - 171, 194 - 195. salt pan, 2 ♂♂, Salicornia marsh, 27.IV.2007 (CRS); Akrotiri Peninsula, Fasouri reedbeds, 1 ♂, Citations 4.V.2007 (CRS). Without further locality (Kulczyński, 1908). NICOSIA: Dali, 1 ♀, under stones near river, LIMASSOL: Kourion, Apollo Hylates Temple (Bosmans, 2009). 18.V.2007 (CAR - S); Elia - Angolemi ro ad, 100m, PAPHOS: Saranta Kolones (Bosmans, 2009). 1 ♀, rubbish in dry riverbed, 18.V.2011 (CRB); Kato Koutrafas - Astromeritis road, 120m, 1 ♂, New records herbs on pond shore, 18.V.2011 (CJVK); Lympia NICOSIA : Agios Sozomenos , near Dali, 1 ♂, Reservoir, Lympia, 1 ♂ 1 ♀, in grass tussocks and under stones in dry grassland, 29.IV.2007 (CAR - on concrete duct, 9.V.2013 (CAR - S). S), 1 ♂, 29.IV.2013 (CAR - S); Kampos NE, Xeros PAPHOS : Evretou Res ervoir, 350m, 1 ♂, shrubs River, 550m, 1 ♂, stones in riverbed, 20.V.2011 and grassland around lake, 1.V.2010 (CJVK); (CRB); Marathasa Dam, 10 ♂♂ 3 ♀♀, pitfalls in Kidasi N., along Diarizos River, 300m, 2 ♂♂, riverside vegetation, 7.V. - 10.VII.2016 (CMH). under stones, 21.V.2011 (CRB). PAPHOS : Kinousa SE, 400m, 1 ♀, under stones Distribution: Crete, Turkey, Israel, Lebanon. and litter in Pinus forest, 1.V.2010 (CJVK). New to Cyprus. Distribution: Continental Greece and islands, Zodarion barbarae Bosmans, 2009 Turkey, Cyprus, Israel and Lebanon.

Id entification Zodarion lutipes (O. Pickard - Cambridge, 1872) Bosmans, 2009: 248, figs 80 - 83, 138 - 139. Zodarion lutipes ; Bosmans, 2009: 2 81.

Citations Identification None. Bosmans 2009: 281, figs 184 - 185, 178 - 179.

New records Citations FAMAGUSTA: Rizokarpas, 1 ♂, Juniperus PAPHOS: Agios Georgios, Cape Drepanum phoenicea forest, 12.III.2016 (CMH). (Bosmans, 2009); Akrotiri Peninsula (Bosmans, LARNACA : Larnaca SW, Hala Sultan Tekke 2009). mosque, 50m, 1 ♀, in pitfalls in salt mars h, 15 - 28.V.2011 (CJVK). New records LIMASSOL: Pano Platres, 1050m, 1 ♂, litter near LIMASSOL : Akrotiri Peninsula, edge of central Mesopotamos Falls, 23.V.2011 (CRB), 3 ♀♀, salt pan, 1 ♂, Salicornia marsh, 27.IV.2007 4.V.2010 (CJVK). (CAR - S). NICOSIA: Lefka Dam, 13 ♂♂, pitfalls in PAPHOS : Evretou Reservoir, 350m, 4 ♂♂ 1 ♀, riverside vegetation, 7.V. - 10.VII.2016 (CMH, stones in grassland around lake, 1.V.2010 (CJVK, CRB); near Platania Scout camping si te, 2 ♂♂, CRB); near Kidasi village, Diarizos Valley, 2 ♀♀, pitfalls in Pinus brutia forest, 6.IV. - 8. VI.2014, 1 under bark of plane tree near stream, 1.V.2013 ♀, IV. - VII.2017 (CRB ). (CAR - S); near Souskiou, 1 ♂, under stones in dry PAPHOS : Kelefos bridge, 465m, deciduous grassland, 1.V.2013 (CDM). wood, 1 ♂ 3 ♀♀, in litter, 5 ♂♂ 10 ♀♀, under bark, 16.III.2018 (CKHK); Kidasi S., Dhiarizos Distribution: Israel, Lebanon, Jordan and river, 244m, 2 ♀♀, 21.II.2011, Alnus forest, D. Cyprus. W. Wrase leg. (CRB).

Spiders of Cyprus 148

Zodarion reticulatum Kulczyński, 1908 stones in dry riverbed, 18.IV.1986 (NHMW); Zodarion reticulatum Kulczyński, 1908: 61, pl. 2, Kampos NE, Xeros River, 550m, 1 ♀, stones in figs 16 (descr. ♀); Bosmans, 2009: 280. rive rbed and beaten from shrubs, 20.V.2011 (CJVK); Kapedes, 1 ♂, pitfalls in Pinus brutia Identificat ion forest, 17.I. - 23.IV.2016 (CMH); Machairas Bosmans, 2009: 280, figs 174 - 175, 182 - 183. forest near Kioni, 1300m, 1 ♂ 1 ♀, stony phrygana, 5.V.2010 (CJVK); Marathasa Dam, 2 Citations ♂♂ 1 ♀, pitfalls in riverside vegetation, 7.V. - Cyprus, without further locality (type locality; 10.VII.2016 (CMH); Panagia Glossa Monastery Kulczyński, 1908). near Kellaki, 1 ♀, stones in cultivated land, LIMASSOL: Kourion, Apollo Hylates Temple 12.IV.1986 (NHMW); Lakatamia, 1 ♀, in olive (Bosmans, 2009); Kourion (Bosmans, 2009); grove, 1.I. - 1.X. 2016 (CMH); Pano Amiantos, Episkopi (Bosmans, 2009); Avdimou coa st pitfalls in Pinus brutia forest, 6.IV. - 8.VI.2014 (Bosmans, 2009); between Platres and Caledonia (CMH); road Pedoula s - Lemithou, 8 ♂♂ 3 ♀♀, Waterfall (Bosmans, 2009); Troodos mountains, pitfalls in Pinus nigra and Pinus brutia forest, road Platres - Trodoos (Bosmans, 2009). 6.IV. - 8.VI.2014 (CMH). PAPHOS: Panagia (Bosmans, 2009); Pegia PAPHOS : Agios Minas picnic site, 1 ♀, under (Bosmans, 2009). stone in open pine woodland, 13.IV.2017 (CAR - S); Akamas Peninsula, Agios Mamas, 250m, 1 ♀, New records road embankment, 22.IV .2017, S. Huber leg. FAMAGUSTA: Rizokarpaso , 8 ♂♂ 1 ♀, Juniperus p hoenicea forest, 12.III.2016 (CMH). (CSH); Akamas Peninsula, Avgas E stuary, 1 ♀, KERYNEIA : Agios Georgios, 1 ♂ 3 ♀♀, under under stones, 31.III.1997 (CJB); Asprokremmos stone in coastal scrub, 21.III.1998, coll. J. Murphy Dam, N. Kouklia, 150m, 2 ♂♂, herbs and stones (MMUE); Chalefka, Sourp Magar Monastery along lake, 29.IV.2010 (CJVK); near Avakas (Armenomonastiro) road, 660m, 1 ♂, under Gorge, 1 ♂ 2 ♀♀, under stones in field with tall stones in open Pinus forest, 23.XI.2010 (C RB); grass, 14.IV.2017 (CAR - S); Cape Drepanum, 1 Kazafani , 150m, 1 ♂, under stones in olive grove, ♀, under stone , 29.III.1998, coll. J. Murphy 23.XI.2010 (CRB); Klepini , 300m, 1 ♀, under stones in open Pinus forest, 23.XI.2010 (CRB); (MMUE); Kelefos bridge, dam on Diarizos River, Vasileia S ., 200m, 1 ♀, in dry riverbed in olive 550m, 1 ♂, under stones, 21.V.2011 (CJVK); Neo grove, 25.XI.2010 (CRB). Chorio, 1 ♂ 2 ♀♀, under stone, 30.III.1998, coll. LARNACA : Kornos Picnic site, 350m, 1 ♂, 2 J. Murphy (MMUE); Vretsia W., near Kelefos ♀♀, under stones in Pinus forest, 28.V.2011 bridge, 650m, 1 ♀, under stones, 19.V.2011 (CJVK, CRB); Stavrovouni Monastery, 350m, 1 (CJVK); Pegeia, Pikni Forest, 346m, 1 ♂, pitfalls ♀, stones in Pinus forest , 28.IV.2010 (CJVK). in former carob plantation, 11.III. - 23.III.2018 LIMASSOL: Agros SW, 950m, 1 ♀, under stones (CKHK); Pegeia, Pikni Forest, 366m, 2 ♀♀, in in Pinus forest, 5.V.2010 (CJVK); Akrotiri pine forest, 11.III. - 23.III.2018 (CKHK). Peninsula E., 5m, 1 ♀, un der stones and in litter in salt marsh, 29.IV.2010 (CJVK); road Kaminaria - Distr ibution: Cyprus , and further only known Lemithou, 7 ♂♂ 4 ♀♀, pitfalls in Pinus brutia from Chios ( Russell - Smith et al. 2011 ) . forest, 6.IV. - 8.VI.2014 (CMH); Pano Platres, Zodarion thoni Nosek, 1905 below Caledonia Waterfall, 1 ♂, under stones in Zodarium thoni var. cypria Kulczyński, 1908: 57, rocky valley, 26.IV.2007 (CRS); Pedoulas, 960m, pl. 2, fig. 39. 1 ♂ 1 ♀, night catch in orange garden, Zodarion thoni; Bosmans, 2009: 272. 18.IV.2017, S. Huber leg. (CSH); Troodos S., road to Pano Platres, 2 ♀♀, under stones in Pinus Identification nigra forest, 28.IV.2007 (CRS); Troodos village, Bosmans, 2009: 272, figs 168 - 169, 176 - 177. 1 ♂, stones in Pinus forest, 22.IV.86 (NHMW). Citations NICOSIA : Agios Soz omenos , near Dali, 1 ♀, in Without prec ise locality (Kulczyński, 1908, sub Z. dry grassland, 1.IV.2012 (CDM); Armenochori, 1 thoni cyprium ). ♂ 1 ♀ 3 juveniles, under stones of a dry stone LARNACA: Larnaca (Bosmans, 2009). wall, 14.IV.1986 (NHMW); Erimi, 1 ♀, under LIMASSOL: Avdimou coast (Bosmans, 2009).

Spiders of Cyprus 149

NICOSIA: Agios Sozomenos , near Dali, sto nes in wasteland, 24.V.2011 (CJVK, CRB); (Bosmans, 2009). Panagia Glossa Monastery near Kellaki, 1 ♀, PAPHOS: Pa phos, Coral Bay (Bosmans, 2009). agricultural land, stones, rarely plants, 12.IV.1986

(NHMW); Xerargaka S, Orkontas, 2 ♂♂, New records FAMAGUSTA : Cape Greco, Ayia Napa, 35m, 1 riverside vegetation, 26.III.2016 (CMH). ♂ 1 ♀, stones in phrygana, 27.IV.2010 (CJVK, PAPHOS : Akamas Peninsula, Avakas Gorg e, 3 ♀♀, under stone, 27.III.1998, coll. J. Murphy CRB); Gialousa E., Agios Thyrsos, 20m, 1 ♂ 1 ♀ (MMUE); Akamas Peninsula, track along the sea, 2 subadult ♀♀, litter under fig tree, 18.III.2011 10m, 1 ♀, stones in Juniperus phrygana, (CJVK, CRB); Kantara castle, 650m, 1 ♀, stones 30.IV.2010 (CRB); Akamas Peninsula, Agios in ruins, 19.III.2011 (CRB); Karpasia Peninsula, Minas, 2 ♂♂, under stones in meadow with tall Apostolos Andreas Monastery, 50m, 1 ♀, stones grass, 2.V.2007 (CAR - S); Akama s Peninsula in wasteland, 16.V.2011 (CRB); Karpasia Cape Geronnisos S., 2 ♀♀, under stones in dry phrygana, 3.V.2007 (CAR - S); Kinousa, 1 ♂, in Peninsula, Cape Apostolos Andreas, 70m, 1 ♀, pine wood, 19.III.1998, coll. J. Murphy (MMUE); stones in grassland, 16.V.2011 (CJVK); Paralimni Smigies, 1 ♀, under stone in hillside scrub, L ake, 65m, 2 ♂♂, herbs and litter around dried 26.III.1998, coll. J. Murphy (MMUE). lake, 27.IV.2010 (CJVK, CRB); Prastio S ., Agios Andronikos, 30m, 2 ♂♂, under stones and herbs Distribution: Gree ce, Bulgaria, Romania, in Eucalyptus forest, 27.V.2011 (CRB); Salamis Macedonia, Russia, Ukraine, Azerbaijan, ruins, 50m, 1 ♀, under stones, 15.V.2011 (CJVK). Armenia, Georgia, Turkey, Cyprus, Lebanon.

KERYNEIA: Panagia Apsinthiotissa Monastery ruins, 350m, 1 ♀, stones in clearing in Pinus Family Zoropsidae forest, 24.V.2011 (CJVK ).

LARNACA : Achna Reservoir, 50m, under stones Akamasia cyprogenia (Bosselaers, 1997) and litter bordering lake, 4 ♂♂ 1 ♀, 27.V.2011 Zoropsis cyprogenia Bosselaers, 1997: 164, figs (CJVK, CRB), 20 ♂♂ 1 ♀, pitfalls at same site, 2 - 10. 15 . - 27.V.2011 (C JVK, CRB); Larnaca SW, Hala Akamasia cyprogenia ; Bo sselaers, 2002: 149, figs Sultan Tekke mosque, 50m, 3 ♂♂ 3 ♀♀, in 3C, 4A - B. pitfalls in salt marsh, 15 - 28.V.2011 (CJVK, Identification CRB), 1 ♀, hand collected at same site, Bosselaers, 2002: 148, figs 4A - C. 15.V.2010 (CJVK). LIMASSOL: Akrotiri Peninsula, near British Citations Airbase, 1 ♀, under stones in phrygana, 4 .V.2007 PAPHOS : Akamas Peninsula, Avakas Gorge (CAR - S); Kourion archaeological site near (Bosselaers, 1997; type locality); Akamas Episkopi, 3 ♂♂, under stones, 9.V.2007 (CAR - S); Peninsula, Neo Chorio (Bosselaers 2002). Pissouri, 1 ♀, in grass, 24.III.1998, coll. J. Murphy (MMUE). New records NICOSIA : Agioi Trimithias, 1 ♂, in olive grove, NICOSIA: Kampos, 500m, 1 ♂ 1 ♀, old holiday 22.I.2016 (CMH); Agios Sozomenos , near Dali, 2 village, 23.IV.2017, S. Huber leg. (CSH); near ♂♂, under stones in dry grassland, 30.IV.2007 Platania Scout camping site, 1 ♀, pitfalls in Pinus (CAR - S), 4 ♂♂, under stones in phrygana, brutia forest, 6.IV. - 8.VI.2014 (CMH). PAPHOS: Akamas Peninsula, Agios Mamas, 30.IV.2007 (CAR - S), 1 ♂, 29.IV.2013 (CRS), 1 250m, 1 ♂, road embankment, 22.IV.2017, S. ♂, IV.2011 (CDM); Agridi, near Dali, 1 ♀, in Huber leg. (CSH). yard, IV.2009, 1 ♀, on veranda, 1.II.2010, 1 ♀, in house, II.2010, 1 ♂, in gar den; Dali, 13, Agios Distribution: A rare endemic species of Cyprus. Andronikos, 1 ♂, in garden, V.2008 (all CDM); Zoropsis lutea (Thorell, 1875) Lakatamia, 3 ♀♀, in olive grove, 1.I. - 1.X. 2016 Zoropsis lutea asiatica Kulczyński, 1911: 12, pl. (CMH); Marathasa Dam, 26 ♂♂ 5 ♀♀, pitfalls in 1, figs 1. riverside vegetation, 7.V. - 10.VII.2016 (CMH); Identification Masari S., Merika River, 120m, 5 ♂♂ 6 ♀♀, Levy, 1990: 141, figs 1 - 16.

Spiders of Cyprus 150

area of the archaeological site, 23.IV.1986 Citations (NHMW); Oikos, 700m, 1 ♀, limy slope, Without precise loc ality Kulczyński, 1911, as 23.IV.2017, S. Huber leg. (CSH); Pano Platres, Zoropsis lutea asiatica , now considered a track near Caledonia Waterfall, 1 juvenile; under synonym of the nominal species. ). stones and in litter at edge of Pinus nigra forest, 26.IV.2007 (CAR - S); road Pano Platres - Troodos, New records 1 juvenile, under stones in Pinus nigra forest, FAMAGUSTA : Agios Sergios , 30m, 1 ♀, stones 1.V.2007 (CAR - S); Pissouri, 1 ♀, in grass, in phrygana, 17.III.2011 (CRB); Cape Apostolos 24.III.1998, coll. J. Murphy (MMUE). Andreas, 1 ♀, Juniperus phoenicea forest PAPHOS : Akamas Peninsula, 2 ♀♀, in coastal grass land, 22.III.1998, coll. J. Murphy (MMUE); 30 .III.2016 (CMH); N. Vouni ruins, 250m, 1 ♀, Baths of Aphrodite, 1 ♀, under large stone near stones in ruins, 20.III.2011 (CRB); Rizokarpaso beach, 15.IV.2017 (CAR - S); Evretou Reservoir, W ., Panagia Eleousa Monastery, 30m, 2 ♀♀, Stavros tis Psokas river, 1 ♀, 20.III.2018 stones in Pinus forest, 18.III.2011. (CKHK); Gialia, 1 ♀, in pine wood, 25.III.1998, KERYNEIA : Agios Georgios, 1 ♀, under stone in coll. J. Murphy (MMUE); Kino usa E., Agios coastal scrub, 21.III.1998, coll. J . Murphy Merkourios, 350m, 1 ♀, under stones and litter in (MMUE). Pinus forest, 1.V.2010 (CRB); Kouklia - Pissouri NICOSIA: Ayia Koroni, Agios Epifanios, 1 ♀, in road, 150m, 1 ♀, stony slope near sea, 22.III.2011 house, XI.2004, 1 ♀, in house, IV.2005 (CDM); (CRB); Makounta, 1 ♂, under stones in scrub, Agridi, near Dali, 1 ♀, under bark of pine tree, 19.III.1998, coll. J. Murphy (MMUE); Neo I.2011, 1 ♀, in yard, 7.I.2010 (CDM); 1 ♀, in Chorio, 1 ♀, under stone, 30.III.1998, coll. J. Eucalyptus litter in garden, 29.IV.2013 (CAR - S); Murphy (MMUE); Paphos NW, 1 ♀, under stones Dali, Lymbourena Hill, 1 ♀, 2005 (CDM). at the edge of a large sand pit, 24.IV.1986 LIMASSOL : Akrotiri Peninsula, Turtle Beach, 1 (NHMW); Polis W., Latsi, path to Blue Lagoon, ♀, in phrygana, 15.III.2018 (CKHK); Amathus, 1 50m, 1 ♂, 22.IV.2017, S. Huber leg. (CSH); ♀, under stones in dry valley, 6.IV.1986 Smigies, 1 ♀, under stone in hillside s crub, (NHMW); Armenochori, 1 ♀, under stones of a 26.III.1998, coll. J. Murphy (MMUE). dry stone wall, 14.IV.1986 (NHMW); K ourion, Apollo Hylates Temple, 1 ♀, under stones in the Distribution: Eastern Mediterranean, Ukraine.

Spiders of Cyprus 151

VI. Discussion

Prior to this publication, only 103 spider species had been recorded from Cyprus (Nentwig et al ., 2019). The 44 9 species recorded here more than quadruples that number, a testimony to the efficacy of focused surveys by teams of arachnologists in documenting biodiversity. Figure 1 shows the increase in the number of species per year. Interest in the spider of Cyprus appears to have been ve ry low and grew very gradually until 2012, rising quickly from then on.

Increasing number of spiders 500 450 400 350 R

R 300 E B 250 M

U 200 N 150 100 50 0 2 2 2 6 0 4 8 2 6 0 4 8 2 6 0 4 8 2 6 0 4 8 2 6 0 4 8 2 6 0 4 8 7 8 0 0 1 1 1 2 2 3 3 3 4 4 5 5 5 6 6 7 7 7 8 8 9 9 9 0 0 1 1 1 8 8 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 0 0 0 0 0 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 2 2 2 2 2 YEAR

Figure 1: The number of species recorded per year in Cyprus between 1872 and 201 9 .

A consideration of Table 1 above s hows that the total number of spider species in Cyprus (44 9 ) i s close to that for the island of Crete (430 species), a similarly sized eastern Mediterranean island (Bosmans et al ., 2013). Furthermore, a comparison of the family composition of the spider faunas of these two islands indicates that they are closely simi lar in this respect as well. For the seven most species - rich families (Araneidae, Gnaphosidae, Linyphiidae, Lycosidae, Salticidae, Theridiidae, Thomisidae) the proportion of each family represented in the total lies within 1.5% of each other in the two isl ands. This is not to suggest that there are no significant differences at all. The fauna of Cyprus includes two families, Ctenidae and Phyxelididae (each represented by a single species) not represented in Crete or elsewhere in Europe and a number of gener a that also are not known to occur in Crete (e.g. Hersiliola (Hersiliidae) , Cithaeron (Cithaeronidae) , Zaituna (Filistatidae) and Akamasia (Zoropsidae)).

Another clear difference between the spider faunas of Cyprus and Crete lies in the number of end emic species represented in each island. In Crete there are 58 species recorded as endemic to the island, representing 13.5% of all spiders (Bosmans et al ., op. cit.) . By contrast, in Cyprus only 17 species endemic to the island are currently known, repres enting 4.3% of all spiders.The reasons for this difference are not entirely clear but a significant factor could be the close proximity of Cyprus to Turkey on the one hand (only 70 km at the nearest point) and to Syria on the other (only 105 km at the near est point). It seems likely that there has been frequent movement of spider species from Turkey and the Middle East to Cyprus. This suggestion is strongly borne out by the distribution data provided in this catalogue which includes a relatively high propor tion of species which are also represented in Turkey and/or the Middle East.

How complete is our knowledge of the Cyprus spider fauna? Although the present work provides a useful starting point, it is almost certain that the species list will expand significantly over the coming decades. A paper in preparation includes 23 species new to science from the island (Bosmans et al ., in prep.) and further collecting will very likely reveal many more, both new to Cyprus and to science. Given the island’s posi tion at a crossroads between Asia , Africa and Europe and our limited knowledge of the fauna of the eastern Mediterranean as a whole, we can confidently expect to see a rapid growth in the number of species from Cyprus.

Spiders of Cyprus 152

Table 3 lists the 1 7 currently know n endemic species of Cyprus. The species that will be described as new in a future paper are not taken into account here. Dysderidae and Linyphiidae are the most abundant species.

Six of the species listed show no special habitat preferences and have larg e distribution areas, occuring in all or almost all island districts. These species perhaps also occur in nearby Turkey. Four species prefer litter in forest or phrygana. two species are restricted to caves and one species seems to prefer wetland habitats. Several species have insufficiently known habitats and more research is necessary to establish the preferences of these species.

Family Species Districts Habitat Dysderidae Dysderocrates kibrisensi Ni Caves Dysderidae Harpactea cecconii La Li Ni Pa Various Dysderidae Harpactea gunselorum Ke Ni Forests Dysderidae Harpactea kalavachiana Ni Caves Gnaphosidae Berinda cypria Fa Li Pa Various Gnaphosidae Drassyllus cyprius Ke La Li Ni Pa Forest, phrygana Linyphiidae Araeoncus cypriacus Li Pa Litter Linyphiidae Improphantes cypriot Fa La Ni Pa Various Linyphiidae Palliduphantes minimus Ke La Li Ni Forests, grass tussocks Linyphiidae Walckenaera cypresensis Li Ni Pa Forests, mostly at high altitudes Liocranidae Mesiotelus cyprius Fa Ke La Li Ni Pa Various Nemesiidae Brachythele incerta Unknown Unknown Oonopidae Opopaea saaristoi ‘North Cyprus’ ‘Plants on sandy ground’ Salticidae Ael urillus cypriotus Fa Li Ni Pa Various Theridiidae Theridion cyprusense Pa Bushes along rivers, springs and near beach Theraphosidae Chaetopelma karlamani Fa Ke La Ni Forests at higher altitude Zoropsidae Akamasia cyprogenia Ni Pa Forest litter

Table 3: Currently known endemic species of Cyprus, their distribution in districts and recorded habita t preferences (Fa: Famagusta; Ke : Keryneia ; La: Larnaca; Li:Limassol; Ni: Nicosia; Pa: Paphos).

Acknowledgements B. Hub er is thanked for the identification of Pholcidae, D. Logunov and G. Azarkina for the identification of Salticidae and Thomisidae, M. Chatzaki for the identification of Gnaphosidae and A. Bolzern for the identification of Agelenidae. Without their willing help this catalogue would have been impossible to complete. Many thanks also to Martin Askins , Maria Chatzaki , George Constantinou, Herman Dekoninck, M arios Giannapas, Evan Jones , Christodoulos Makris, Eleni Panayiotou , Matthew Smith, Robin D. Symeou and Lut Vannieuwenhuyse for their valuable help during sampling. Theo Blick and Aydin Topçu are thanked for examining material of M egalepthyphantes . turkeyensis . We thank Christo f Hörweg (NHMV) for making available for study the holotype of Brachythele incert a . Finally, we would like to thank Theo Blic k, Wo l f gang Nentwig and some ARABEL members for interesting discussions.

Spiders of Cyprus 153

References A HARON , S., H UBER , B. A. & G AVISH - R EGEV , E. ( 2017). Daddy - long - leg giants: revision of the spider genus Artema Walcken aer, 1837 (Araneae, Pholcidae). European Journal of Taxonomy 376 : 1 - 57. A USSERER , A . (1871). Beiträge zur Kenntniss der Arachniden - Familie der Territelariae Thorell (Mygalidae Autor). Verhandlungen der kaiserlich - königlischen zoologisch - botanischen Gesel lschaft in Wien 21 : 117 - 224. A ZARKINA , G. N. (2002). New and poorly known species of the genus Aelurillus Simon, 1884 from Central Asia, Asia Minor and the eastern Mediterranean (Araneae: Salticidae). Bulletin of the British Arachnological Society 12 (6) : 249 - 263. A ZARKINA , G. N. (2006). Four new species of the genus Aelurillus Simon, 1884 (Araneae: Salticidae). In C. Deltshev & P. Stoev (eds.), European Arachnology 2005. Acta zoologica bulgarica Supplement 1: 63 - 72. A ZARKINA , G. N., Z OUMIDES , C. & H A DJICONSTANTIS , M. (2018). First description of the female of Aelurillus cypriotus Azarkina 2006 (Araneae: Salticidae). Acta Arachnologica 67 (1): 49 - 54. B AYRAM , A., E FIL , L. & D ELTSHEV , C. (2009). Pardosa roscai (Roewer, 1951), a spider new to the fauna o f Turkey (Araneae: Lycosidae). Turkish Journal of Zoology 33 : 463 - 467. B LAUWE , R. de. (1980a). Revision de la famille des Agelenidae (Araneae) habitant la region méditerranéene (3e partie). Bulletin de l’ Institut royal des Sciences naturelles de Belgiqu e 52 (11): 1 - 28. B LAUWE , R. de. (1980b). Revision de la famille des Agelenidae (Araneae) de la region mediterraneene (2e partie). Bulletin de l’Institut royal de Sciences naturelles de Belgique 52 : 1 - 54. B ORGES , P. A. V. & W UNDERLICH , J. (2008). Spider biodiversity patterns and their conservation in the Azorean archipelago, with descriptions of new species. Systematics and Biodiversity 6 : 249 - 282. B OSMANS , R. (1996). The genera Araeoncus Simon, Delorripis Simon and Diplocephalus Bertkau in northern Afr ica (Araneae: Linyphiidae: Erigoninae): Studies on North African Linyphiidae VII. Belgian Journal of Zoology 126 : 123 - 151. B OSMANS , R. (2006). Contribution to the knowledge of the Linyphiidae of the Maghreb. Part XI. Miscellaneous linyphiid genera and ad ditions (Araneae: Linyphiidae: Linyphiinae). Bulletin et Annales de la Société royal belge d’Entomologie 141: 125 - 161. B OSMANS , R. (2009). Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae, Zodariidae). Cont ributions to natural History 12 : 211 - 295. B OSMANS , R. (2011). On some new or rare spider species from Lesbos, Greece (Araneae: Agelenidae, Amaurobiidae, Corinnidae, Gnaphosidae, Liocranidae). Arachnologische Mitteilungen 40 : 15 - 22. B OSMANS , R., B AERT , L., B OSSELAERS , J., D EONINCK , H., M AELFAIT , J. - P. & V AN K EER , J. (2009). Spiders of Lesbos (Greece). Nieuwsbrief van de Belgische Arachnologische Vereniging 24 (SUPPL.): 1 - 70. B OSMANS , R. & B LICK , T. (2000). Contribution to the knowledge of the genus Mi caria in the West - palaearctic region, with description of the new genus Arboricaria and three new species (Araneae Gnaphosidae). Memorie della Società Entomologica Italiana, Genova 78 : 443 - 476. B OSMANS , R. & B OURAGBA , N. (1992). Trois nouvelles Linyphiida e de l'Atlas Algérien, avec la description du mâle de Lepthyphantes djazairi Bosmans, et la redescription de Lepthyphantes homonymus Denis (Araneae). Bulletin & Annales de la Société Entomologique de Belgique 128 : 245 - 262. B OSMANS , R. & V AN K EER , J. (199 9). The genus Enoplognatha Pavesi, 1880 in the Mediterranean region (Araneae: Theridiidae). Bulletin of the British Arachnological Society 11 : 209 - 241. B OSMANS , R., K ALTSAS , D., K URT , K., Y AGMUR , E. A., B IRD L. A. & H ARVEY M. S. (2016). Chapter 19: Arach nidae, 467 - 531. In: S PARROW , D. J. & J OHN E. An introduction to the wildlife of Cyprus. Terra Cypria, Cyprus, 895 B OSMANS , R., K HERBOUCHE - A BROUS , O., B ENHALIMA , S. & H ERVÉ , C. (2018). The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the M aghreb in particular (Araneae: Gnaphosidae). Zootaxa 4451 (1): 1 - 67. pp. B OSSELAERS , J. (1997). Zoropsis cyprogenia sp. n., a new, probably endemic spider species from Cyprus (Araneae, Zoropsidae). Phegea 25 : 162 - 168. B OSSELAERS , J. (2002). A cladistic a nalysis of Zoropsidae (Araneae), with the description of a new genus. Belgian Journal of Zoology 132 : 141 - 154. B OSSELAERS , J. (2009). Studies in Liocranidae (Araneae): redescriptions and transfers in Apostenus Westring and Brachyanillus Simon, as well as description of a new genus. Zootaxa 2141: 37 - 55. B OSSELAERS , J. & J OCQUÉ , R. (2002). Studies in Corinnidae: cladistic analysis of 38 corinnid and liocranid genera, and transfer of Phrurolithinae. Zoologica Scripta 31 : 241 - 270. B REITLING , R. (2019). A barcode - based phylogenetic scaffold for Xysticus and its relatives (Araneae: Thomisidae: Coriarachnini). Ecologica Montenegrina 20 : 198 - 206. B RIGNOLI , P. M. (1968). Su alcuni Araneidae e Theridiidae di Sicilia (Araneae). Atti dell' Accademia Gioenia di S cienze Naturali in Catania (6) 20 : 85 - 104. B RIGNOLI , P. M. (1976) . Ragni di Grecia IX. Specie nuove o interessanti delle famiglie Leptonetidae, Dysderidae, Pholcidae ed Agelenidae (Araneae). Revue Suisse de Zoologie 83 (3): 539 - 578. B RIGNOLI , P. M . (19 80) . Some new or interesting eastern Mediterranean Dysderidae and Agelenidae (Araneae). Annales Zoologici, Warszawa 35 : 75 - 82.

Spiders of Cyprus 154

B RIGNOLI , P. M. (1982) . Contribution à la connaissance des Filistatidae paléarctiques (Araneae). Revue Arachnologique 4 : 65 - 75. B UCHAR , J. (1968). Zur Lycosidenfauna Bulgariens (Arachn., Araneae). Věstník Československé Zoologické Společnosti v Praze 32 : 116 - 130. B UCHAR , J. & T HALER , K. (2002). Über Pardosa atomaria (C.L. Koch) und andere Pardosa - Arten an Geröllufern in Süd - und Mitteleuropa (Araneae, Lycosidae). Linzer Biologische Beiträge 34 : 445 - 465. B UCHAR , J., K NOFLACH , B. & T HALER , K. (2006) . On the identity of Arctosa varians C. L. Koch and Artosa similis Schenkel, with notes on related species. Bulletin of the Britis h Arachnological Society 13 : 329 - 336. C ARILLO , J. & F ERR Á NDEZ , M .A. (2014). Primeros registros de Scytodes univittata Simon 1882 (Araneae: Scytodidae) en la Península Ibérica. Revista Ibérica Aracnologia 24 : 53 - 57. C HATZAKI , M. (2010). A revision of th e genus Nomisia in Greece and neighboring regions with the description of two new species. Zootaxa 2501: 1 - 22. C HATZAKI , M. (2018). On the ground spider genera Marjanus gen. n., Lasophorus gen. n. and Turkozelotes Kovblyuk & Seyyar, 2009 (Araneae: Gnapho sidae) from Greece. Zootaxa 4392 (3): 521 - 545. C HATZAKI , M., T HALER , K. & M YLONAS , M. M. (2002a). Ground spiders (Gnaphosidae; Araneae) of Crete (Greece). Taxonomy and distribution. I. Revue Suisse de Zoologie 109 : 559 - 601. C HATZAKI , M., T HALER , K. & M Y LONAS , M. (2002b). Ground spiders (Gnaphosidae, Araneae) of Crete and adjacent areas of Greece. Taxonomy and distribution. II. Revue Suisse de Zoologie 109 : 603 - 633. C HATZAKI , M., T HALER , K. & M YLONAS , M. (2003). Ground spiders (Gnaphosidae; Araneae) fro m Crete and adjacent areas of Greece. Taxonomy and distribution. III. Zelotes and allied genera. Revue Suisse de Zoologie 1109 : 45 - 89. C HATZAKI , M. & R USSELL - S MITH , A. (2017). New species and new records of ground spiders (Araneae: Gnaphosidae) from Cypr us. Zootaxa 4329 (3): 237 - 255. C HATZAKI , M. & V AN K EER , J. (2019). Ground spiders (Araneae: Gnaphosidae, Liocranidae, Prodidomidae) from the Greek islands Rodos, Symi and Karpathos, with the description of new species. Zootaxa 4646 (3): 434 - 460 C HATZOPOULO U , E. & C HATZAKI , M. (2009). Taxonomic review of some Drassodes species from Greece and other east Medit erranean countries (Araneae: Gnaphosidae). Contributions to Natural History 12 : 349 - 359. C RESPO , L. C., M ENDES , S. & P RADO E. C ASTRO , C. (2010). Two additions to the portuguese and iberian spider (Arachnida, Araneae) fauna. Graellsia 66 : 97 - 100. D EELEMAN - R EINHOLD , C. L. (1985). Contribution à la connaissance des Lepthyphantes du groupe pallidus (Araneae, Linyphiidae) de Yougoslavie, Grece et Chypre. Mémoires de Biospéléologie 12: 37 - 50. D EELEMAN - R EINHOLD , C. L. & D EELEMAN , P. R. (1988). Revision des Dysderinae (Araneae, Dysderidae), les espèces mediterranéennes occidentales exceptées. Tijdschrift voor Entomologie 131 : 141 - 269 D ELTSHEV , C. D. & P A RASCHI , L. (1990). A contribution to the study of spiders (Araneae: Dysderidae, Salticidae, Agelenidae) in Greece, with a description of a new species ( Malthonica spinipalpis Deltshev, sp. n. Agelenidae). Biologia Gallo - Hellenica 17 : 3 - 12. D EMIR , H., A KT AŞ , M. & S EYYAR , O . (2008) . The female of Xysticus pseudorectilineus (Wunderlich, 1995) (Araneae: Thomisidae) from Turkey. Zootaxa 1674 : 65 - 68. D ENIS , J. (1937). On a collection of spiders from Algeria. Proceedings of the Zoological Society of London 19 36 : 1027 - 1060. D UNIN , P. M., B UCHAR , J. & A BSOLON , K. (1986). Die Dritte Paläarktische Aulonia - Art: Aulonia kratochvili sp. n. (Araneida, Lycosidae). Věstník Československé Zoologické Společnosti v Praze 50 : 28 - 32. E FIMIK , V. E. (1999). A review of the spider genus Tibellus Simon, 1875 of the East Palearctic (Aranei: Philodromidae). Arthropoda Selecta 8 : 103 - 124. E SYUNIN , S. L. (1994). Remarks on the spider fauna of the Urals, 3. Devade Simon, 1884, a genus new to the Urals, with notes on Devade indi stincta (O. P. - Cambridge, 1872) (Arachnida Aranei Dictynidae). Arthropoda Selecta 3 (1 - 2): 39 - 47 E SYUNIN , S. L. & E FIMIK , V. E. (2000). A review of the genus Devade (Aranei, Dictynidae) from fauna of central Asia and southern Russia. Zoologicheski ĭ Zhurna l 79 : 679 - 685. E SYUNIN , S. L. & M ARUSIK , Y. M. (2001). A new species of the genus Devade Simon, 1884 from Mongolia, with notes on D. tenella (Tyschchenko, 1965) (Aranei: Dictynidae). Arthropoda Selecta 9 : 129 - 131. F EDORIAK , M., V OLOSHYN , V. L. & M OSCAL IUC , L. A. (2016). Scientific heritage of Alexandru Roşca: publications, spider collection, described species. Arachnologische Mitteilungen 51 : 85 - 91. G UADANUCCI , J. P. L. & G ALLON , R. C. (2008). A revision of the spider genera Chaetopelma Ausserer 1871 and Nesiergus Simon 1903 (Araneae, Theraphosidae, Ischnocolinae). Zootaxa 1753: 34 - 48. G ÜCEL , S., F ULLER , Ö. Ö., G ÖÇMEN , B. & K UNT , K. B . (2018). Harpactea gunselorum sp. n., a new spider species from northern Cyprus (Araneae: Dysderidae). Serket 16 (2): 80 - 86. G ÜCEL , S., C HARALAMBIDOU , I., G ÖÇMEN , B. & K UNT , K. B. (2019). New data of spiders (Arachnida, Araneae) of Cyprus. 1. Dysderidae found in caves. ZooKeys 825 : 43 - 53. H EPNER , M. & P AULUS , H. F. ( 2009) . Contributions on the wolf spider fauna (Ara neae, Lycosidae) of Gran Canaria (Spain). Bulletin of the British Arachnological Society 14 : 339 - 346.

Spiders of Cyprus 155

H ILLYARD P. D. (1983). Episinus maculipes Cavanna (Araneae, Theridiidae): rediscovery in Britain. Bulletin of the British Arachnological Society 6 : 88 - 9 2. H IPPA , H. & O KSALA , I. (1982). Definition and revision of the Enoplognatha ovata (Clerck) group (Araneae: Theridiidae). Entomologica Scandinavica 13 : 213 - 222. I SAIA , M., K RONESTEDT , T., B ALLARIN , F. & C HIARLE , A . (2018). On the morphological separat ion of two sibling species: Pardosa proxima ( P. vlijmi syn. nov.) and P. tenuipes (Araneae: Lycosidae). Arachnologische Mitteilungen 56 : 6 - 16 J ÄGER , P., M C C OWAN , D. & R USSELL - S MITH , A . (2011 ) . First record of Olios suavis for southern Europe in Cyprus (A raneae: Sparassidae: Sparassinae). Bulletin of the British Arachnological Society 15 : 127 - 129. K NOFLACH , B. (1994). Zur Genitalmorphologie und Biologie der Crustulina - Arten Europas (Arachnida: Araneae, Theridiidae). Mitteilungen der Schweizerischen Entom ologischen Gesellschaft 67 : 327 - 346. K NOFLACH , B. (1996). Das Männchen von Simitidion agaricographum (Levy & Amitai) (Arachnida: Araneae, Theridiidae). Bericht des naturwissenschaftlich medizinischen Vereins in Innsbruck 83: 149 - 156. K NOFLACH , B. (1998 ). Mating in Theridion varians Hahn and related species (Araneae: Theridiidae). Journal for natural History 32: 545 - 604. K NOFLACH , B. (1999). The comb - footed spider genera Neottiura and Coleosoma in Europe (Araneaem Theridiidae). Mitteilungen der schweiz erischen entomologischen Geselschaft 72: 341 - 371. K NOFLACH , B. & VAN H ARTEN , A. (2002). The genus Latrodectus (Araneae: Theridiidae) from mainland Yemen, the Socotra Archipelago and adjacent countries. Fauna of Arabia 19: 321 - 361. K NOFLACH , B. & T HALER , K. (2000). Notes on Mediterranean Theridiidae (Araneae) - I. Memorie della Società Entomologica Italiana, Genova 78 : 411 - 442. K NOFLACH - T HALER , B., J. V AN K EER , A. A SKINS & R USSELL - S MITH , A. (2008). Neospintharus syriacus – a new widespread species in the Eastern Mediterranean? Newsletter of the British Arachnological Society 113: 9. K OVBLYUK , M. M. & T UNEVA , T. K. (2009). Three interesting species of Gnaphosidae from Crimea (Arachnida: Aranei). Arthropoda Selecta 17 : 157 - 164. K OVBLYUK , M. M., K ASTR YGINA , Z. A. & O MELKO , M. M. (2012). A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae). ZooKeys 205 : 59 - 89. K OVBLYUK , M. M., G NELITSA V. A., N ADOLNY A. A., K ASTRYGINA , Z. A., K UKUSHKI N O. V. (2016). Spiders (Arachnida: Aranei) of the Karadag Nature Reserve (Crimea). Ekosystem y 3 (33): 3 – 288. K RITSCHER , E. (1966). Die paläarktischen Arten der Gattung Oecobius (Aran., Oecobiidae). Annalen des Naturhistorischen Museums in Wien 69 : 285 - 2 95. K RONESTEDT , T. (1997) . First record of Aulonia kratochvili (Araneae, Lycosidae) from Europe. Arachnologische Mitteilungen 14 : 77 - 80. K RONESTEDT , T. & Z YUZIN , A. A. (2009). Fixation of Lycosa fidelis O. Pickard - Cambridge, 1872 as the type species f or the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescripton of the species. Contributions to Natural History 12 : 813 - 828. [incl. corrigendum: vol. 13/2010: 89]. K UBCOVÁ , L. (2004). Separation of the females of Philodromus praedatus O.P. - Cambridge and Philodromus aureolus (Clerck) (Philodromidae, Araneae). In: F. Samu & C. Szinetár (eds.) European Arachnology 2002. Plant Protection Institute & Berzsenyi College Budapest, pp. 57 - 62 K ULCZYŃSK , W. (1908). Fragmenta arachnologica VI. X. Aran eae nonnuli in Cyprio insula et in Palaestina a Cel. Profigs Dre G. Cecconi lectae. Bulletin international de l’Académie des Sciences de Cracovie 1908 : 49 - 85. K ULCZYŃSK , W. (1909). Fragmenta arachnologica VIII. Palpimanorum species Mediterranorum. Bullet in international de l’Académie des Sciences de Cracovie 1909 : 667 - 676. K ULCZYŃSK , W. (1911). Fragmenta Arachnologica XVI, XVII. Bulletin international de l’Académie des Sciences de Cracovie 1911: 12 - 75. L EHTINEN , P. T. (1967). Classification of the cr ibellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4 : 199 - 468. L EVY , G. (1973). Crab - spiders of six genera from Israel (Araneae: Thomisidae). Israel Journal of Zoology 22 : 107 - 14 1. L EVY , H. W. (1974). The orb - weaver genus Zygiella (Araneae: Araneidae). Bulletin of the Museum of Comparative Zoology at Harvard College 146 : 267 - 290. L EVY , G. (1976). The spider genus Xysticus (Araneae: Thomisidae) in Israel. Israel Journal of Zool ogy 25: 1 - 37. L EVY , G. & A MITAI , P. (1981). spiders of the genera Euryopis and Dipoena (Araneae: Theridiidae) from Israel. Bull Br arachnol Soc 5 : 177 - 188. L EVY , G. & P. A MITAI (1982). The comb - footed spider genera Theridion , Achaearanea and Anelosimus of Israel (Araneae: Theridiidae). Journal for Zoology, London 196: 81 - 131. L EVY , G. (1984). The spider genera Singa and Hypsosinga (Araneae, Araneidae) in Israel. Zoologica Scripta 13 : 121 - 133. L EVY , G. (1985a). Spiders of the genera Episinus , Argyro des and Coscinida from Israel, with additional notes on Theridion (Araneae: Theridiidae). Journal of Zoology, London (A) 207 : 87 - 123. L EVY , G. (1985b). Araneae: Thomisidae. In: Fauna Palaestina, Arachnida II. Israel Academy of Sciences and Humanities, J erusalem, 115 pp

Spiders of Cyprus 156

L EVY , G. (1986). Spiders of the genera Siwa , Larinia , Lipocrea and Drexelia (Araneae: Araneidae) from Israel. Bulletin of the British Arachnological Society 7 : 1 - 10. L EVY , G. (1989). The family of huntsman spiders in Israel with annota tions on species of the Middle East (Araneae: Sparassidae). Journal of Zoology, London 217 : 127 - 176. L EVY , G. (1990). On the cribellate spider Zoropsis lutea in Israel (Araneae, Zoropsidae). Bulletin of the British Arachnological Society 8 : 139 - 143. Le vy, G. ( 1991) . On some new and uncommon spiders from Israel (Araneae). Bulletin of the British Arachnological Society 8 : 227 - 232. L EVY , G. (1992). The spider genera Palaestina , Trygetus , Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotatio ns on species of the Middle East. Israel Journal of Zoology 38 : 67 - 110. L EVY , G. (1995). Revision of the spider subfamily Gnaphosinae in Israel (Araneae: Gnaphosidae). Journal of Natural History 29 : 919 - 981. L EVY , G. (1996). The agelenid funnel - weaver family and the spider genus Cedicus in Israel (Araneae, Agelenidae and Cybaeidae). Zoologia Scripta 25: 85 - 122. L EVY , G. (1998a). Twelve genera of orb - weaver spiders (Araneae, Araneidae) from Israel. Israeli Journl of Zoology 43 : 311 - 365. L EVY , G. (199 8b). Araneae: Theridiidae. In: Fauna Palaestina, Arachnida III. Israel Academy of Sciences and Humanities, Jerusalem, 228 pp. L EVY , G. (1998c). The ground - spider genera Setaphis , Trachyzelotes , Zelotes , and Drassyllus (Araneae: Gnaphosidae) in Israel. Is rael Journal of Zoology 44 : 93 - 158. L EVY , G. (1999). The lynx and nursery - web spider families in Israel (Araneae, Oxyopidae and Pisauridae). Zoosystema 21 : 29 - 64. L EVY , G. (2003). Spiders of the families Anyphaenidae, Hahniidae, Ctenidae, Zoridae, and Hersiliidae (Araneae) from Israel. Israel Journal of Zoology 49 : 1 - 31. L EVY , G. (2004). Spiders of the genera Drassodes and Haplodrassus (Araneae, Gnaphosidae) from Israel. Israeli Journal of Zoology 50 : 1 - 37. L EVY , G. (2007). Calommata (Atypidae) and new spider species (Araneae) from Israel. Zootaxa 1551 : 1 - 30. L EVY , G . (2009). New ground - spider genera and species with annexed checklist of the Gnaphosidae (Araneae) of Israel. Zootaxa 2066 : 1 - 49. L ISSNER , J. (2017c) . A contribution to the knowledge of Rhomphaea L. Koch, 1872 (Araneae: Theridiidae) in the Mediterranean and Macaronesian regions. Arachnology 17 (6): 287 - 293 L ISSNER , J. & C HATZAKI , M. (2016a). First records of Poecilochroa taborensis Levy, 1999 (Araneae: Gnaphosidae) from Greece and Cy prus with notes on some closely related species. Nieuwsbrief SPINED 36 : 16 - 21 L ISSNER , J. & C HATZAKI , M. (2016b). A new species of Berinda (Araneae, Gnaphosidae) from the eastern Aegean Islands, Greece. Arachnologische Mitteilungen 51 : 9 - 11. L ISSNER , J . (2016 ). A small study of the Corsican spider and pseudoscorpion fauna (Araneae, Pseudoscorpiones). Nieuwsbrief SPINED 36 : 5 - 15 L OERBROKS , A. (1983). Revision der Krabbenspinnen - Gattung Heriaeus Simon (Arachnida: Araneae: Thomisidae). Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg 26 : 85 - 139. L OGUNOV , D. V. (1979). New and poorly known species of Salticidae (Araneae) from Turkey and Iran. Contributions to natural History 12 : 899 - 919. L OGUNOV , D. V. (2006). Notes on Xysticus kempeleni Th orell, 1872 and two closely related spider species (Araneae: Thomisidae). Acta Arachnologica, Tokyo 55 : 59 - 66. L OGUNOV , D. V. (2013). On a small spider collection from Cyprus (Arachnida: Araneae). Newsletter of the British Arachnological Society 127 : 4 - 5. L OGUNOV , D. V. (2015) . Taxonomic - faunistic notes on the jumping spiders of the Mediterranean (Aranei: Salticidae). Arthropoda Selecta 24 (1): 33 - 85. L OGUNOV , D. V. & H USEYNOV , E. F. (2008 ) . A faunistic review of the spider family Philodromidae (Arane i) of Azerbaijan. Arthropoda Selecta 17 : 117 - 131. L OGUNOV , D. V. & M ARUSIK , Y. M. (2003). A revision of the genus Yllenus Simon, 1868 (Arachnida, Araneae, Salticidae). KMK Scientific Press, Moscow, 168 pp. L OKSA , I. (1972). Araneae II. Fauna Hungariae 109 : 1 - 112. L OKSA , I. (1973). On the morphology and systematical position of Mysmena leucoplagiata (Simon, 1879) (Araneae: Symphytognathidae). Acta Zoologica Academiae Scientiarum Hungaricae 19 : 283 - 287. M ARUSIK , Y. M. & L OGUNOV , D. V. (1990). The crab spiders of middle Asia, USSR (Aranei, Thomisidae). 1. Descriptions and notes on distribution of some species. Korean Arachnology 6 : 31 - 62. M ARUSIK , Y. M. & L OGUNOV , D. V. (1995). The crab spiders of Middle Asia (Aranei, Thomisidae), 2. Beiträge zur Aran eologie 4 : 133 - 175. M ARUSIK , Y. M., L EHTINEN , P. T. & K OVBLYUK , M. M. (2005). Cozyptila , a new genus of crab spiders (Aranei: Thomisidae: Thomisinae : Coriarachnini) from the western Palaearctic. Arthropoda Selecta 13 : 151 - 163. M ARUSIK , Y. M. & F ET , V. (2009). A survey of east Palearctic Hersiliola Thorell, 1870 (Araneae, Hersiliidae), with a description of three new genera. ZooKeys 16 : 75 - 114 . Spiders of Cyprus 157

M ARUSIK , Y. M. & Z AMANI , A. (2015) . The spider family Filistatidae (Araneae) in Iran. ZooKeys 516 : 123 - 135. M C C OWAN , D . (2014). Sixty Cypriot Spiders. Mikrokyklos Books, 1 - 155 M ERRETT , P. (2001). Clubiona pseudoneglecta Wunderlich, 1994, a clubionid spider new to Britain (Araneae: Clubionidae). Bulletin of the British Arachnological Society 12 : 32 - 34. M E TZNER , H. (1999). Die Springspinnen (Araneae, Salticidae) Griechenlands. Andrias 14 : 1 - 279. M ILLIDGE , A. F. (1988). The spiders of New Zealand: Part VI. Family Linyphiidae. Otago Museum Bulletin 6 : 35 - 67. M ORADMAND , M. & J ÄGER , P. (2012). Taxonomic rev ision of the huntsman spider genus Eusparassus Simon, 1903 (Araneae: Sparassidae) in Eurasia. Journal of narural History 46: 2439 - 2496. M URPHY , J. (2007). Gnaphosid genera of the world . British Arachnological Society, St Neots, Cambs 1, i - xii, 1 - 92; 2:i - 11, 93 - 605 M USTER , C., R., T HALER , K. (2004). New species and records of Mediterranean Philodromidae (Arachnida, Araneae): I. Philodromus aureolus group. In: Thaler, K. (ed.) Diversität und Biologie von Webspinnen, Skorpionen und anderen Spinnentieren. D enisia 12 , 305 - 326. M USTER , C., B OSMANS , R. & T HALER , K. (2007). The Philodromus pulchellus - group in the Mediterranean: taxonomic revision, phylogenetic analysis and biogeography (Araneae: Philodromidae). Invertebrate Systematics 21: 39 - 72. N ADOLNY , A . A., P ONOMAREV , A. V., K OVBLUYK , M. M. & D VADNENKO , K. V. (2012). New data on Pisaura novicia (Aranei: Pisauridae) from eastern Europe. Arthropoda Selecta 21 : 255 - 267. N ENTWIG , W., B LICK T., G LOOR D., H ÄNGGI A. & K ROPF C . (2018). Araneae. Spiders of Eu rope. Online at http://wsc.nmbe.c h, version 19.0, last accessed on 15 September 2019. doi: 10.24436/2. O VTSHARENKO , V. I., P LATNICK , N. I. & S ONG , D. X. (1992). A review of the North Asian ground spiders of the genus Gnaphosa (Araneae, Gnaphosidae). Bul letin of the American Museum of Natural History 212 : 1 - 88. O VTSHARENKO , V. I., L EVY , G. & P LATNICK , N. I. ( 1994) . A review of the ground spider genus Synaphosus (Araneae, Gnaphosidae). American Museum Novitates 3095 : 1 - 27 Ö ZKÜTÜK R. S., E LVERICI , M., K UNT , K. B. & Y AĞMUR , E. A. (2013). Faunistic notes on the cybaeid spiders of Turkey (Araneae: Cybaeidae). Journal of Applied Biological Sciences 7 (2) : 71 - 77. Ö ZKÜTÜK , R. S., G ÜCEL , S., F ULLER , Ö. Ö. & K UNT , K. B . (2019). Cheiracanthium mildei L. Koch, 1864 (Araneae: Cheiracanthiidae), a new faunistic record for Cyprus. Serket 16 (4): 184 - 187. P ANAYIOTOU , E., K ALTSAS , D., S EYYAR , O. & C HATZAKI , M. (2010). Revision of the genus Berinda (Araneae, Gnaphosidae) in the east Mediterranean with the descriptio n of two new species. Zootaxa 2362 : 44 - 54. P ÉREZ , P., J. A. & B LASCO F., A. (1986). Nota sobre los Prodidominae (Araneae: Gnaphosidae) de la Peninsula Iberica. Mémoires de la Société Royal Belge d'Entomologie 33 : 155 - 164. P ESARINI C. (1997). I ragni ( Arachnida Araneae) del Monte Barro (Italia, Lombardia, Lecco). Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 27 : 251 - 263. P ETERS , H. - J. (1998). Tarantulas of the world: Afrika's Vogelspinnen. Published by the author, 122 pp. P ETERS , H. - J. (2000). Tarantulas of the world: Kleiner Atlas der Vogelspinnen - Band 1. Published by the author, 148 pp. P ICKARD - C AMBRIDGE , O. (1872). General list of the spiders of Palestine and Syria, with descriptions of nume rous new species, and characters of two new genera. Proceedings of the Zoological Society of London 40 (1): 212 - 354, pl. XIII - XVI. P LATNICK , N. I. (1981) . A review of the spider subfamily Palpimaninae (Araneae, Palpimanidae), I. Bulletin of the British Ar achnological Society 5 : 169 - 173. P LATNICK , N. N. I. (1991). A revision of the ground spider family Cithaeronidae (Araneae, Gnaphosoidea). American Museum Novitates 3018 : 1 - 13. P LATNICK , N. N. I. & M URPHY , J. A. (1984). A revision of the spider genera T rachyzelotes and Urozelotes (Araneae, Gnaphosidae). American Museum Novitates 2792 : 1 - 30. P LATNICK , N. N. I. & O VTSHARENKO , V. I. (1991). On Eurasian and American Talanites (Araneae, Gnaphosidae). Journal of Arachnology 19 : 115 - 121. P LATNICK , N. I. & S HADAB , M. U. (1983) . A revision of the American spiders of the genus Zelotes (Araneae, Gnaphosidae). Bulletin of the American Museum of Natural History 174 : 97 - 192. P OLENEC , A. & T HALER , K. (1980). Zwei wenig bekannte Deckennetzspinnen Südost - Europas: C entromerus vindobonensis Kulczynski und Leptyphantes istrianus Kulczynski (Arachnida: Araneae: Linyphiidae). Senckenbergiana Biologica 61 : 103 - 111. R OBERTS , M. J. (1985). The spiders of Great Britain and Ireland, Volume 1: Atypidae to Theridiosomatidae. H arley Books, Colchester, England. R USSELL - S MITH , A. (1982). A revision of the genus Trabaea Simon (Araneae: Lycosidae). Zoological Journal of the Linnean Society 74 : 69 - 91. R USSELL - S MITH , A. , A LLISON , R., A SKINS , M, B LUMSOM , W., S NAZELL R. & S PILLIN , C. (2011). A provisional checklist and gazetteer of the spiders of Chios, Greece (Arachnida: Araneae). Bulletin of the British Arachnological Society 8 : 38 - 40. R USSELL - S MITH , A. Spiders from the Ionian islands of Kerkyra (Corfu) and Lefkada, Greece (Ara chnida: Aranei). Arthropoda Selecta 23 (3): 285 – 300.

Spiders of Cyprus 158

S AARISTO , M. I. (2007). The oonopid spiders (Aranei: Oonopidae) of Israel. Arthropoda Selecta 15: 119 - 140. S AARISTO , M. I. (2010). Araneae. In: Gerlach, J. & Y. Marusik (eds.) Arachnida and Myriapoda of the Seychelles islands. Siri Scientific Press, Manchester UK, pp. 8 - 306. S AARISTO , M.I. & M ARUSIK , Y.M. (2008): A survey of African Opopaea Simon, 1891 (Arachnida, Aranei, Oonopidae). Arthropoda Selecta 17 : 17 - 53. S ENGLET , A. (2011). New species in the Zelotes tenuis - group and new or little known species in other Zelotes groups (Gnaphosidae, Araneae). Revue Suisse de Zoologie 118 : 513 - 559. S IMON , E. (1874). Simon, E. (1874a). Les arachnides de France . Paris 1, 1 - 272. S IMON , E. (1878). Les arachni des de France . Paris 4, 1 - 334. S IMON , E. (1911). Catalogue raisonné des arachnides du nord de l'Afrique (1re partie). Annales de la Société Entomologique de France 79 (3, for 1910): 265 - 332. S IMON , E. (1932). Les arachnides de France. Tome VI. Synopsis générale et catalogue des espèces françaises de l'ordre des Araneae; 4e partie . Paris 6, 773 - 978. S MITH , A. M. (1990). Baboon spiders: Tarantulas of Africa and the Middle East . Fitzgerald Publishing, London, pp. 1 - 142 S PARROW , D. J. J OHN E. (Eds) (20 16). An Introduction to the Wildlife of Cyprus. 870 pp. Limassol. S TRAND , E. (1907). Spinnen des zoologischen Instituts in Tübingen. Zoologische Jahrbücher. II. Abteilung für Systematik 24 : 391 - 468. T ANASEVITCH , A. V. (2011). On linyphiid spiders from the eastern and central Mediterranean kept at the Muséum d'histoire naturelle, Geneva. Revue Suisse de Zoologie 118 : 49 - 91. T ANASEVITCH , A. V ., K UNT , K. B. & S EYYAR , O. (2005). A new species of the genus Megalepthyphantes Wunderlich from Turkey (Aranei: Linyphiidae: Micronetinae). Arthropoda Selecta 13 : 279 - 280. T HALER , K. & K NOFLACH , B. ( 1998). Phyxelida anatolica Griswold, new to Cyprus (Arachnida Araneae: Amaurobiidae, Phyxelidinae). Bulletin of the British Arachnological Society 11: 36 - 40. T HALER , K. & N OFLATSCHER , M. - T. (1990). Neue und bemerkenswerte Spinnenfunde in Südtirol (Arachnida: Aranei). Veröffentlichungen des Museum Ferdinandeum in Innsbruck 69 : 169 - 190. T HALER , K., B UCHAR , J. & K NOFLACH , B. (2000). Notes on wolf spiders from Greece (A raneae, Lycosidae). Linzer Biologische Beiträge 32 : 1071 - 1091. T HALER , K. & K NOFLACH , B. (2004). A new Lachesana species from Greece and additional records from the Near East (Arachnida, Araneae, Zodariidae). Bulletin of the British Arachnological Societ y 13 : 41 - 46. T HALER , K., H ARTEN , A. V AN & K NOFLACH , B. (2004). Pirate spiders of the genus Ero C.L. Koch from southern Europe, Yemen, and Ivory Coast, with two new species (Arachnida, Araneae, Mimetidae). Denisia 13 : 359 - 368. T OPÇU , A., S EYYAR O. & EMI RCAN N. (2013). A contribution to the cave spider fauna of Turkey. Zoology in the Middle East 59 : 91 - 92. U RONES , C., B ARRIENTOS , J. A. & E SPUNY , A. (1995). El género Anyphaena Sundevall, 1833 (Araneae: Anyphaenidae) el la Península Ibérica. Boletín de la Asociación Española de Entomología 19 : 109 - 131. V AN H ELSDINGEN , P. J. (1969). A reclassification of the species of Linyphia Latreille based on the functioning of the genitalia (Araneida, Linyphiidae), I. Zoologische Verhandelingen 105 : 1 - 303. V AN H ELSD INGEN , P. J. (1970). A reclassification of the species of Linyphia based on the functioning of the genitalia (Araneida, Linyphiidae), II. Zoologische Verhandelingen 111 : 1 - 86. V AN H ELSDINGEN , P. J. (1979). Remarks concerning Clubionidae. Bulletin of the British Arachnological Society 4 : 298 - 302. V AN H ELSDINGEN , P. J. (2009). Mermessus denticulatus (Banks, 1898) and Mermessus trilobatus (Emerton, 1882), adventive species in the Netherlands (Araneae, Linyphiidae). Contributions to Natural History 12 : 617 - 626. V AN K EER , J., V AN K EER K., D E K ONINCK H. A R AMEL G. (2010). The Arachnofauna (Araneae) of Wetland Kerkini (Macedonia - Northern Central Greece). Acta zoologica bulgarica 62 (2): 141 - 160. V OLLMER , P. (1997). Chaetopelma karlamani sp. n., (Araneida: T heraphosidae: Ischnocolinae) eine Vogelspinne aus Nord - Zypern. Tarantulas of the World , Ausgabe 16, Dec. 1997: 4 - 18. W ESOLOWSKA , W. (1986). A revision of the genus Heliophanus C. L. Koch, 1833 (Aranei: Salticidae). Annales Zoologici, Warszawa 40 : 1 - 254. W ORLD S PIDER C ATALOG (2019). World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 19.0, last accessed on 15 September 2019. doi: 10.24436/2. W UNDERLICH , J. (1980). Linyphiidae aus Süd - Europa und Nord - Afrika (Arachn. : Araneae). Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (N.F.) 23 : 319 - 337. W UNDERLICH , J. (1995a). Zu Taxonomie und Biogeographie der Arten der Gattung Oecobius Lucas 1846, mit Neubeschreibungen aus der Mediterraneis und von der Arabisc hen Halbinsel (Arachnida: Araneae: Oecobiidae). Beiträge zur Araneologie 4 : 585 - 608. W UNDERLICH , J. (1995b). Beschreibung bisher unbekannter Arten der Baldachinspinnen aus der Östlichen Mediterraneis (Arachnida: Araneae: Linyphiidae). Beiträge zur Arane ologie 4: 655 - 686. W UNDERLICH , J. (1995c). Zur Taxonomie europäischer Gattungen der Zwergspinnen (Arachnida: Araneae: Linyphiidae: Erigoninae). Beiträge zur Araneologie 4 : 643 - 654. Spiders of Cyprus 159

W UNDERLICH , J. (1995d). Zur Kenntnis west - paläarktischer Arten der Gatt ungen Psammitis Menge 1875, Xysticus C. L. Koch 1835 und Ozyptila Simon 1864 (Arachnida: Araneae: Thomisidae). Beiträge zur Araneologie 4 : 749 - 774. W UNDERLICH , J. (2011). Extant and fossil spiders (Araneae). Beitrage zur Araneologie wept 1 - 640. Z AMANI , A. & M OZAFFARIAN , F . (2017). Further spider (Arachnida: Araneae) material deposited in the Agricultural Zoology Museum of Iran (AZMI), Iranian Research Institute of Plant Protection. Arachnologische Mitteilungen 54 : 8 - 20. Z ONSTEIN , S. & M ARUSIK , Y.M. (2 009). A revision of the spider genus Zaitunia (Araneae, Filistatidae). European Journal of Taxonomy 214: 1 – 97. Z ONSTEIN , S. & M ARUSIK , Y. M. (2016). A revision of the spider genus Zaitunia (Araneae, Filistatidae). European Journal of Taxonomy 214 : 1 - 97. doi:10.5852/ejt.2016.214 Z ONSTEIN , S. & M ARUSIK , Y . M. (2019). A revision of the spider genus Filistata (Araneae: Filistatidae). Arachnology 18 (2): 53 - 93. Z YUZIN , A. & L OGUNOV , D. V. (2000). New a nd little - known species of the Lycosidae from Azerbaijan, the Caucasus (Araneae, Lycosidae). Bulletin of the British Arachnological Society 11 : 305 - 319.

Spiders of Cyprus 160

Addendum. List of spiders of Cyprus

Family Species First cited in 1 Agelenidae Agelena orien talis C. L. Koch, 1837 Present paper

2 Agelenidae Benoitia lepida (O. P - Cambridge, 1876) De Blauwe, 1980

3 Agelenidae Lycosoides coarctata (Dufour, 1831) Strand, 1907

4 Agelenidae Tegenaria dalmatica (Kulczyński, 1906) Brignoli, 1976

5 Agelenidae Tegen aria dentifera Kulczyński, 1908 Kulczyński,1908

6 Agelenidae Tegenaria domestica (Clerck, 1757) Present paper

7 Agelenidae Tegenaria maronita (Simon, 1873) Present paper

8 Agelenidae Tegenaria pagana C. L. Koch, 1840 Present paper

9 Agelenidae Tegen aria parietina (Fourcroy, 1785) Present paper

10 Agelenidae Textrix caudata L. Koch, 1872 De Blauwe, 1980

11 Anyphaenidae Anyphaena sabina L. Koch, 1866 Present paper

12 Araneidae Aculepeira armida (Audouin, 1826) Present paper

13 Araneidae Agalena tea redii (Scopoli, 1763) Present paper

14 Araneidae Araneus diadematus Clerck, 1757 Present paper

15 Araneidae Araneus grossus (C. L. Koch, 1844) Present paper

16 Araneidae Araniella alpica (L. Koch, 1869) Present paper

17 Araneidae Araniella inconspicua (Simon, 1874) Present paper

18 Araneidae Argiope lobata (Pallas, 1772) Present paper

19 Araneidae Argiope trifasciata (Forskål, 1775) Present paper

20 Araneidae Cyclosa groppal ii Pesarini, 1998 Present paper

21 Araneidae Cyclosa sierrae Simon, 1870 Logunov, 2013

22 Araneidae Cyrtarachne ixoides (Simon, 1870) Present paper

23 Araneidae Cyrtophora citricola (Forsskål, 1775) Present paper

24 Araneidae Gibbaranea bituberculata (Walckenaer, 1802) Present paper

25 Araneidae Gibbaranea gi bbosa (Walckenaer, 1802) Present paper

26 Araneidae Glyptogona sextuberculata (Keyserling, 1863) Present paper

27 Araneidae Hypsosinga albovittata (Westring, 1851) Present paper

28 Araneidae Hypsosinga pygmaea (Sundevall, 1831) Present paper

29 Aran eidae Larinia chloris (Audouin, 1826) Present paper

30 Araneidae Larinioides suspicax (O. Pickard - Cambridge, 1876) Present paper

31 Araneidae Leviellus kochi (Thorell, 1870) Present paper

32 Araneidae Lipocrea epeiroides (O. Pickard - Cambridge, 1870) Wunderlich, 2011

Spiders of Cyprus 161

33 Araneidae Mangora acalypha (Walckenaer,1802) Logunov, 2013

34 Araneidae Neoscona adianta (Walckenaer,1802) Logunov, 2013

35 Araneidae Neoscona sodom Levy, 1998 Present paper

36 Araneidae Neoscona subfusca (C. L. Koch, 1837) Present paper

37 Araneidae Singa neta (O. Pickard - Cambridge, 1872) Present paper

38 Araneidae Zilla diodia (Walckenaer, 1802) Present paper

39 Araneidae Zygiella atrica (C. L. Koch, 1845) Present paper

40 Cheiracanthidae Cheiracanthium elegans Thorell, 18 75 Present paper

41 Cheiracanthidae Cheiracanthium erraticum (Walckenaer, 1802) Present paper

42 Cheiracanthidae Cheiracanthium mildei L. Koch, 1864 Present paper

43 Cheiracanthidae Cheiracanthium seidlitzi (L. Koch, 1864) Present paper

44 Cheiraca nthidae Cheiracanthium sp. Present paper

45 Cithaeronidae Cithaeron praedonius O. Pickard - Cambridge, 1872 Present paper

46 Clubiona Clubiona pseudoneglecta Wunderlich, 1994 Present paper

47 Clubiona Porrhoclubiona vegeta (Simon, 1918) Present paper

48 Ctenidae Anahita syriaca (O. Pickard - Cambridge, 1872) Present paper

49 Cybaeidae Cedicus flavipes Simon, 1875 Kulczyński,1908

50 Dictynidae Argenna patula (Simon, 1874) Present paper

51 Dictynidae Brigittea civica (Lucas, 1850) Logunov, 2013

52 Di ctynidae Brigittea innocens (O. Pickard - Cambridge, 1872) Present paper

53 Dictynidae Devade tenella (Tyschchenko, 1965 ) Present paper

54 Dictynidae Lathys arabs Simon, 1910 Present paper

55 Dictynidae Lathys humilis (Blackwall, 1855) Present paper

56 Dictynidae Mariyinia bicolor (Simon, 1870) Present paper

57 Dictynidae Nigma flavescens (Walckenaer, 1830) Present paper

58 Dictynidae Nigma puella (Simon, 1870) Present paper

59 Dictynidae Nigma walckenaeri (Roewer, 1951) Present paper

60 Dysd eridae Dysdera gigas Roewer, 1928 Present paper

61 Dysderidae Dysdera lata Reuss, 1834 Deeleman - Reinhold, 1988

62 Dysderidae Dysdera longimandibularis Nosek, 1905 Deeleman - Reinhold, 1988

63 Dysderidae Dysdera sp. Present paper

64 Dysderidae Dysdera sp . Present paper

65 Dysderidae Dysdera spinicrus Simon, 1882 Deeleman - Reinhold, 1988

66 Dysderidae Dysdera westringi (O. Pickard - Cambridge, 1872) Strand, 1907

Spiders of Cyprus 162

Dysderocrates kibrisensis Gücel, Charalambidou, 67 Dysderidae Gücel et al ., 2019 Göçmen & Kunt, 2019 68 Dysderidae Harpactea cecconii (Kulczyński, 1908) Kulczyński,1908 Harpactea gunselorum Gücel, Fuller, Göçmen & Kunt, 69 Dysderidae Gücel et al ., 2018 2018 Harpactea kalavachiana Gücel, Charalambidou, Göçmen 70 Dysderidae Gücel et al ., 2019 & Kunt, 2019 71 Dysderidae Harpactea sp. Present paper

72 Dysderidae Harpactea sp. Present paper

73 Dysderidae Harpactea sp. Present paper

74 Dysderidae Harpactea sp. Present paper

75 Filistatidae Filistata insidiatrix (Forsskål, 1775) Brignoli, 1982

76 Filistatidae Pritha nana (Simon, 1868) Present paper

77 Filistatidae Zaitunia annulipes (Kulczyński, 1908) Kulczyński,1908

78 Filistatidae Zaitunia kunti Zonstein & Marusik Zonstein & Marusik 2016

79 Gnaphosidae Anagraphis pallens Simon, 1893 Presen t paper

80 Gnaphosidae Aphantaulax cincta (L. Koch, 1866) Present paper

81 Gnaphosidae Aphantaulax trifasciata (O. Pickard - Cambridge, 1872) Present paper

82 Gnaphosidae Berinda cypria Chatzaki & Panayiotou, 2010 Chatzaki & Panayiotou, 2010

83 Gna phosidae Berinda ensigera (O. Pickard - Cambridge, 1872) Present paper

84 Gnaphosidae Berinda idae Lissner, 2016 Present paper

85 Gnaphosidae Berlandina plumalis (O. Pickard - Cambridge, 1872) Present paper

86 Gnaphosidae Cesonia aspida Chatzaki, 2002 Pr esent paper

87 Gnaphosidae Civizelotes solstitialis Levy, 1998 Present paper Chatzaki & Russell - Smith, 88 Gnaphosidae Cryptodrassus helvolus (O. Pickard - Cambridge, 1872) 2017 89 Gnaphosidae Drassodes cupreus (Blackwall, 1834) Logunov, 2013

90 Gnapho sidae Drassodes lacertosus (O. Pickard - Cambridge, 1872) Present paper

91 Gnaphosidae Drassodes lapidosus (Walckenaer, 1802) Present paper

92 Gnaphosidae Drassodes lutescens (C. L. Koch, 1839) Present paper

93 Gnaphosidae Drassodes serratichelis (Ro ewer, 1928) Present paper

94 Gnaphosidae Drassodes unicolor (O. Pickard - Cambridge, 1872) Present paper Chatzaki & Russell - Smith, 95 Gnaphosidae Drassyllus cyprius Chatzaki & Russell - Smith, 2017 2017 96 Gnaphosidae Drassyllus jubatopalpis Levy, 1998 Pr esent paper

97 Gnaphosidae Drassyllus pusillus (C. L. Koch, 1833) Present paper

98 Gnaphosidae Echemus levyi (Kovblyuk & Seyyar, 2009) Chatzaki & Van Keer, 2019

99 Gnaphosidae Gnaphosa bithynica Kulczyński, 1903 Present paper

100 Gnaphosidae Gnaphos a dolosa Herman, 1879 Kulczyński, 1908

Spiders of Cyprus 163

101 Gnaphosidae Haplodrassus dalmatensis (L. Koch, 1866) Present paper

102 Gnaphosidae Haplodrassus invalidus (O. Pickard - Cambridge, 1872) Present paper

103 Gnaphosidae Ha p lodrassus omissus (O. P ickard - C ambr idge , 1872 Present paper Haplodrassus pseudosignifer Marusik, Hippa & 104 Gnaphosidae Present paper Kopponen, 1996 105 Gnaphosidae Haplodrassus signifer (C. L. Koch, 1839) Present paper

106 Gnaphosidae Heser nilicola (O. Pickard. - Cambridge, 1874) Platnick & Shadab, 1983

107 Gnaphosidae Lasophorus zografae Chatzaki, 2018 Chatzaki, 2018

108 Gnaphosidae Leptodrassus femineus (Simon, 1873) Present paper

109 Gnaphosidae Micaria coarctata (Lucas, 1846) Present paper

110 Gnaphosidae Micaria dives (Lucas, 1846) Present paper

111 Gnaphosidae Micaria guttulata (C. L. Koch, 1839) Present paper

112 Gnaphosidae Micaria ignea O. Pickard - Cambridge, 1872 Simon, 1876

113 Gnaphosidae Micaria pallipes (Lucas, 1846) Present paper

114 Gnaphosidae Nomisia excerpt a (O. Pickard. - Cambridge, 1872) Chatzaki & Panayiotou, 2010

115 Gnaphosidae Nomisia exornata (C. L. Koch, 1839) Present paper

116 Gnaphosidae Nomisia recepta (Pavesi, 1880) Present paper

117 Gnaphosidae Nomisia ripariensis (O. Pickard. - Cambridge, 187 2) Present paper

118 Gnaphosidae Phaeocedus braccatus (L. Koch, 1866) Present paper

119 Gnaphosidae Phaeocedus hebraeus Levy, 1999 Present paper

120 Gnaphosidae Poecilochroa senilis (O. Pickard - Cambridge, 1872) Present paper Chatzaki & Russell - Smith, 121 Gnaphosidae Poecilo chroa sp. 2017 122 Gnaphosidae Poecilochroa taborensis Levy, 1999 Lissner & Chatzaki, 2016

123 Gnaphosidae Prodidomus amaranthinus (Lucas, 1846) Present paper

124 Gnaphosidae Pterotricha kochi (O. Pickard - Cambridge, 1872) Pres ent paper

125 Gnaphosidae Pterotricha lentiginosa (L. Koch, 1837) Logunov, 2013

126 Gnaphosidae Pterotricha schaefferi (Audouin, 1826) Present paper

127 Gnaphosidae Scotophaeus peninsularis Roewer, 1928 Present paper

128 Gnaphosidae Scotophaeus scutu latus (L. Koch, 1866) Present paper

129 Gnaphosidae Setaphis carmeli (O. Pickard - Cambridge, 1872) Present paper Synaphosus palearcticus Ovtsharenko, Levy & Platnick, 130 Gnaphosidae Present paper 1994 Chatzaki & Russell - Smith, 131 Gnaphosidae Synaphosus shirin Ovtsharenko, Levy & Platnick , 1994 2017 132 Gnaphosidae Synaphosus trichopus (Roewer, 1928) Present paper

133 Gnaphosidae Talanites dunini Platnick & Ovtsharenko, 1991 Present paper

134 Gnaphosidae Trachyzelotes jaxartensis (Kroneberg, 1 875) Present paper

Spiders of Cyprus 164

135 Gnaphosidae Trachyzelotes lyonneti (Audouin, 1826) Present paper

136 Gnaphosidae Trachyzelotes malkini Platnick & Murphy, 1984 Present paper

137 Gnaphosidae Trachyzelotes stubbsi Platnick & Murphy, 1984 Platnick & Shadab, 1983 Chatzaki & Russell - Smith, 138 Gnaphosidae Turkozelotes mccowani (Chatzaki & Russell - Smith, 2017) 2017 139 Gnaphosidae Zelotes harmeron Levy, 2009 Present paper Chatzaki & Russell - Smith, 140 Gnaphosidae Zelotes limnatis Chatzaki & Russell - Smith, 2017 20 17 141 Gnaphosidae Zelotes metellus Roewer, 1928 Present paper

142 Gnaphosidae Zelotes scrutatus (O. Pickard. - Cambridge, 1872) Present paper

143 Gnaphosidae Zelotes sp. Present paper

144 Gnaphosidae Zelotes tenuis (L. Koch, 1866) Present paper Chatzaki & Russell - Smith, 145 G naphosidae Zelotes zekharya Levy, 2009 2017 146 Hahniidae Iberina candida (Simon, 1875) Present paper

147 Hersiliidae Hersiliola macullulata (Dufour, 1831) Present paper

148 Hersiliidae Hersiliola simoni (O. Pickard - Cambridge , 1872) Present paper

149 Linyphiidae Agyneta fuscipalpa (C. L. Koch, 1836) Present paper

150 Linyphiidae Agyneta pseudorurestris (Wunderlich, 1980) Wunderlich, 1980

151 Linyphiidae Agyneta simplicitarsis (Simon, 1884) Present paper

152 Linyphiidae Alioranus pastoralis (O. Pickard - Cambridge, 1872) Wunderlich, 1980

153 Linyphiidae Araeoncus cypriacus Tanasevitch, 2011 Tanasevitch, 2011

154 Linyphiidae Araeoncus humilis (Blackwall, 1841) Present paper

155 Linyphiidae Bathyphantes gracilis (Blackwal l, 1841) Present paper

156 Linyphiidae Canariphantes tenerrimus (Simon, 1929) Present paper

157 Linyphiidae Cresmatoneta mutinensis (Canestrini, 1868) Tanasevitch, 2011

158 Linyphiidae Dactylopisthes digiticeps (Simon, 1881 Present paper

159 Linyphii dae Diplocephalus graecus (O. P - Cambridge, 1873) Present paper

160 Linyphiidae Diplocephalus sp. Present paper

161 Linyphiidae Diplocephalus turcicus Brignoli, 1972 Present paper

162 Linyphiidae Erigone dentipalpis (Wider, 1834) Tanasevitch, 2011

16 3 Linyphiidae Erigonoplus spinifemuralis (Dimitrov, 2003) Present paper

164 Linyphiidae Frontinellina frutetorum (C. L. Koch, 1835) Kulczyński,1908

165 Linyphiidae Gnathonarium dentatum (Wider, 1834) Tanasevitch, 2011

166 Linyphiidae Gongylidiellum murc idum Simon, 1884 Tanasevitch, 2011

167 Linyphiidae Improphantes cypriot Tanasevitch, 2011 Tanasevitch, 2011

168 Linyphiidae Improphantes turok Tanasevitch, 2011 Present paper

Spiders of Cyprus 165

169 Linyphiidae Jacksonella bidens Tanasevitch, 2011 Tanasevitch, 2011

170 L inyphiidae Linyphia maura Thorell, 1875 Present paper

171 Linyphiidae Linyphia mimonti Simon, 1884) Present paper

172 Linyphiidae Linyphia triangularis (Clerck, 1757). Present paper

173 Linyphiidae Maculoncus parvipalpus Wunderlich, 1995. Present pap er

174 Linyphiidae Mecopisthes sp. Present paper Megalepthyphantes turkeyensis Tanasevitch, Boğaç, 175 Linyphiidae Tanasevitch et al ., 2004 Kunt & Seyyar, 2005 176 Linyphiidae Mermessus fradeorum (Berland, 1932) Present paper

177 Linyphiidae Mermessu s denticulatus (Banks, 1898) Present paper

178 Linyphiidae Microctenonyx subitaneus (O. Pickard - Cambridge, 1875) Tanasevitch, 2011

179 Linyphiidae Microlinyphia pusilla (Sundevall, 1830) Present paper

180 Linyphiidae Neriene clathrata (Sundevall, 1830 ) Present paper

181 Linyphiidae Neriene furtiva (O. Pickard. - Cambridge, 1871). Present paper

182 Linyphiidae Ostearius melanopygius (O. Pickard - Cambridge, 1879) Present paper

183 Linyphiidae Palliduphantes minimus (Deeleman - Reinhold, 1985) Deeleman - Reinhold, 1985

184 Linyphiidae Pelecopsis elongata (Wider, 1834) Tanasevitch, 2011

185 Linyphiidae Prinerigone vagans (Audouin, 1826) Tanasevitch, 2011

186 Linyphiidae Silometopus reussi (Thorell, 1871) Present paper

187 Linyphiidae Sintula retroversu s (O. Pickard - Cambridge, 1875) Tanasevitch, 2011

188 Linyphiidae Styloctetor romanus (O. Pickard - Cambridge, 1873) Present paper

189 Linyphiidae Tallusia vindobonensis ( Kulczyński, 1898) Present paper

190 Linyphiidae Tapinocyba sp. Present paper

191 Linyphiidae Tenuiphantes floriana (van Helsdingen, 1977) Present paper

192 Linyphiidae Tenuiphantes tenuis (Blackwall, 1852) Tanasevitch, 2011

193 Linyphiidae Theonina laguncula (Denis, 1937) N. Comb. Present paper

194 Linyphiidae Typhochrestus digitatu s (O. Pickard - Cambridge, 1873) Present paper

195 Linyphiidae Walckenaeria alticeps (Denis, 1952) Present paper

196 Linyphiidae Walckenaeria antica (Wider, 1834) Present paper

197 Linyphiidae Walckenaeria cyprusensis Wunderlich, 1995 Wunderlich, 1995

198 Linyphiidae Walckenaeria sp. Present paper

199 Linyphiidae Walckenaeria stylifrons (O. Pickard - Cambridge, 1875) Wunderlich, 1995

200 Liocranidae Agroeca inopina O. Pickard - Cambridge, 1886 Present paper

201 Liocranidae Agroeca parva Bosmans, 2011 P resent paper

202 Liocranidae Arabelia pheidoleicomes Bosselaers, 2009 Bosselaers, 2009

Spiders of Cyprus 166

203 Liocranidae Mesiotelus cyprius Kulczyński, 1908 Kulczyński,1908

204 Liocranidae Mesiotelus sp. Present paper

205 Liocranidae Mesiotelus tenuissimus (L. Koch, 1 866) Present paper

206 Liocranidae Scotina celans Blackwall, 1841 Present paper

207 Lycosidae Alopecosa sp.(sub A. simoni ) Kulczyński,1908

208 Lycosidae Alopecosa albofasciata (Brullé, 1832) Bristowe, 1934

209 Lycosidae Arctosa cinerea (Fabricius, 17 77) Present paper

210 Lycosidae Arctosa leopardus (Sundevall, 1833) Kulczyński,1908

211 Lycosidae Arctosa similis Schenkel, 1938 Present paper

212 Lycosidae Arctosa variana C. L. Koch, 1847 Present paper

213 Lycosidae Aulonia kratochvili Dunin, Bucha r & Absolon, 1986 Present paper

214 Lycosidae Hogna ferox (Lucas, 1838) Present paper

215 Lycosidae Hogna radiata (Latreille, 1817) Present paper

216 Lycosidae Lycosa praegrandis C. L. Koch, 1836 Present paper

217 Lycosidae Pardosa albatula (Roewer, 1951) Present paper

218 Lycosidae Pardosa atomaria (C. L. Koch, 1847) Buchar & Thaler, 2002 Pardosa proxima (C. L. Koch, 1847) , P.tenuipes L. Koch, 219 Lycosidae Present paper 1882 220 Lycosidae Pardosa roscai (Roewer, 1951) Present paper

221 Lycosi dae Pirata piraticus (Clerck, 1757) Present paper

222 Lycosidae Trabea paradoxa Simon 1876 Present paper

223 Lycosidae Trochosa hispanica Simon, 1870 Present paper

224 Lycosidae Wadicosa fidelis (O. Pickard - Cambridge, 1872) Present paper

225 Mimeti dae Ero aphana (Walckenaer, 1802) Present paper

226 Mimetidae Ero flammeola Simon, 1881 Present paper

227 Mimetidae Ero tuberculata (De Geer, 1778) Present paper

228 Mimetidae Mimetus laevigatus (Keyserling, 1863) Present paper

229 Miturgidae Zora manicata Simon, 1878 Present paper

230 Mysmenidae Mysmena leucoplagiata (Simon, 1880) Present paper

231 Nemesiidae Brachythele incerta Ausserer, 1871 Ausserer, 1871

232 Nemesiidae Nemesia sp. Present paper

233 Oecobiidae Oecobius cellariorum (Dugès , 1836) Present paper

234 Oecobiidae Oecobius maculatus Simon, 1870 Present paper

235 Oecobiidae Oecobius navus Blackwall, 1859 Present paper

236 Oecobiidae Oecobius putus O. Pickard - Cambridge, 1876 Present paper

Spiders of Cyprus 167

237 Oecobiidae Oecobius teliger (O. Pickard - Cambridge, 1872) Present paper

238 Oecobiidae Uroctea durandi (Latreille, 1809) Present paper

239 Oonopidae Opopaea punctata (O. Pickard - Cambridge, 1872) Present paper

240 Oonopidae Opopaea saaristoi Wunderlich, 2011 Wunderlich, 2011

241 Oo nopidae Opopaea santschii Brignoli, 1974 Saaristo, 2006

242 Oonopidae Orchestina sp. Present paper

243 Oonopidae Silhouettella tomer Saaristo, 2007 Present paper

244 Oxyopidae Oxyopes globifer Simon, 1876 Present paper

245 Oxyopidae Oxyopes heterop hthalmus (Latreille, 1804) Logunov, 2013

246 Oxyopidae Oxyopes lineatus Latreille, 1806 Present paper

247 Oxyopidae Oxyopes mediterraneus Levy, 1999 Present paper

248 Oxyopidae Oxyopes nigripalpis Kulczyński, 1891 Present paper

249 Oxyopidae Oxyopes ramosus (Martini & Goeze, 1778) Present paper

250 Oxyopidae Oxyopes sobrinus Latreille, 1806 Present paper

251 Oxyopidae Peucetia arabica Simon, 1882 Present paper

252 Palpimanidae Palpimanus cyprius Kulczyński, 1909 Kulczyński, 1909

253 Philodromi dae Philodromus aureolus (Clerck, 1757) Present paper

254 Philodromidae Philodromus cespitum (Walckenaer, 1802) Present paper

255 Philodromidae Philodromus longipalpis Simon, 1870 Present paper

256 Philodromidae Philodromus lunatus Muster & Thaler, 2 004 Present paper

257 Philodromidae Philodromus margaritatus (Clerck, 1757) Present paper

258 Philodromidae Philodromus marmoratus Kulczyński, 1891 Present paper

259 Philodromidae Philodromus poecilus (Thorell, 1872) Present paper

260 Philodromida e Philodromus rufus Walckenaer, 1826 Present paper

261 Philodromidae Pulchellodromus medius ( O. Pickard - Cambridge, 1872) Kulczyński,1908

262 Philodromidae Pulchellodromus pulchellus ( Lucas, 1846) Logunov, 2013

263 Philodromidae Thanatus fabricii ( Audou in, 1866) Present paper

264 Philodromidae Thanatus imbecillus (C. L. Koch, 1878) Present paper

265 Philodromidae Thanatus vulgaris Simon, 1870 Logunov, 2013

266 Philodromidae Tibellus macellus Simon, 1875 Present paper

267 Philodromidae Tibellus obl ongus (Walckenaer, 1802) Present paper

268 Pholcidae Artema nephilit Aharon, Huber & Gavish - Regev, 2017 Aharon et al. , 2017

269 Pholcidae Holocnemus pluchei (Scopoli, 1763) Present paper

270 Pholcidae Hoplopholcus sp. Present paper

Spiders of Cyprus 168

271 Pholcidae Pho lcus phalangioides (Fuesslin, 1775) Present paper

272 Pholcidae Spermophora senoculata (Dugès, 1836) Present paper

273 Phrurolithidae Phrurolithus sp. Present paper

274 Phyxelididae Phyxelida anatolica Griswold, 1990 Thaler & Knoflach, 1998

275 Pisa uridae Pisaura consocia (O. Pickard - Cambridge, 1872) Logunov, 2013

276 Pisauridae Pisaura mirabilis (Clerck, 1757) Present paper

277 Pisauridae Pisaura novicia (L. Koch, 1878) Present paper

278 Salticidae Aelurillus aeruginosus (Simon, 1871) Present p aper

279 Salticidae Aelurillus blandus (Simon, 1871) Present paper

280 Salticidae Aelurillus cypriotus Azarkina, 2006 Azarkina, 2006

281 Salticidae Attulus distinguendus (Simon, 1868) Present paper

282 Salticidae Ballus chalybeius (Walckenaer, 1802) Present paper

283 Salticidae Ballus rufipes (Simon, 1868) Logunov, 2015

284 Salticidae Chalcoscirtus infimus (Simon, 1868) Logunov, 2015

285 Salticidae Cyrba algerina (Lucas, 1846) Logunov, 2013

286 Salticidae Euophrys herbigrada (Simon, 1871) Present paper

287 Salticidae Euophrys rufibarbis (Simon, 1868) Logunov, 2015

288 Salticidae Euophrys sulphurea (L . Koch, 1867) Present paper

289 Salticidae Evarcha jucunda (Lucas, 1846) Present paper

290 Salticidae Evarcha nepos (O. Pickard - Cambridge, 1872) Present paper

291 Salticidae Evarcha patagiata (O. Pickard - Cambridge, 1872) Logunov, 2013

292 Salticidae Evarcha insularis (Metzner, 1999) Present paper

293 Salticidae Heliophanillus fulgens (O. Pickard - Cambridge, 1872) Present paper

294 Salticidae H eliophanus auratus C. L. Koch, 1835 Wesolowska, 1986

295 Salticidae Heliophanus decoratus L. Koch, 1875 Present paper

296 Salticidae Heliophanus edentulus Simon, 1871 Present paper

297 Salticidae Heliophanus encifer Simon, 1871 Wesolowska, 1986

298 S alticidae Heliophanus equester L. Koch, 1867 Present paper

299 Salticidae Heliophanus flavipes (Hahn, 1832) Present paper

300 Salticidae Heliophanus melinus L. Koch, 1867 Present paper

301 Salticidae Heliophanus mordax (O. Pickard - Cambridge, 1872) Lo gunov, 2015

302 Salticidae Macaroeris nidicolens (Walckenaer, 1802) Logunov, 2015

303 Salticidae Marpissa nivoyi (Lucas, 1846) Present paper

304 Salticidae Mendoza canestrinii (Ninni, 1868) Present paper

Spiders of Cyprus 169

305 Salticidae Menemerus bivittatus (Dufour, 1 831) Present paper

306 Salticidae Menemerus fagei Berland & Millot, 1941 Present paper

307 Salticidae Menemerus semilimbatus (Hahn, 1829) Present paper

308 Salticidae Mogrus logunovi Prószyński, 2000 Present paper

309 Salticidae Mogrus neglectus (S imon, 1868) Logunov, 2015

310 Salticidae Neaetha absheronica Logunov & Guseinov, 2002 Present paper

311 Salticidae Neon rayi (Simon, 1875) Present paper

312 Salticidae Pellenes brevis (Simon, 1868) Logunov, 2015

313 Salticidae Pellenes diagonalis (Sim on, 1868) Present paper

314 Salticidae Pellenes flavipalpis (Lucas, 1853) Drensky, 1936

315 Salticidae Pellenes geniculatus (Simon, 1866) Logunov, 2015

316 Salticidae Philaeus chrysops (Poda, 1761) Present paper

317 Salticidae Phlegra bresnieri (Luca s, 1846) Present paper

318 Salticidae Phlegra cinereofasciata (Simon, 1868) Logunov, 2015

319 Salticidae Phlegra fasciata (Hahn, 1826) Logunov, 2015

320 Salticidae Plexippoides flavescens (O. Pickard - Cambridge, 1872) Present paper

321 Salticidae Plexi ppoides gestroi (Dalmas, 1920) Logunov, 2015

322 Salticidae Plexippus devorans (O. Pickard - Cambridge, 1872) Present paper

323 Salticidae Plexippus paykulli (Audouin, 1826) Present paper

324 Salticidae Pseudeuophrys obsoleta (Simon, 1868) Logunov, 2015

325 Salticidae Pseudicius picaceus (Simon, 1868) Present paper

326 Salticidae Pseudicius vankeeri Metzner, 1999 Logunov, 2015

327 Salticidae Pseudomogrus gavdos (Logunov & Marusik, 2003) Present paper

328 Salticidae Salticus noordami Metzner, 1999 Lo gunov, 2009

329 Salticidae Salticus propinquus Lucas, 1846 Logunov, 2015

330 Salticidae Sibianor aurocinctus (Ohlert, 1865) Present paper

331 Salticidae Synageles dalmaticus (Keyserling, 1863) Present paper

332 Salticidae Synageles hilarulus (C. L. K och, 1846) Logunov, 2015

333 Salticidae Talavera aequipes (O. Pickard - Cambridge, 1871) Present paper

334 Salticidae Thyene imperialis (Rossi, 1846) Present paper

335 Scytodidae Scytodes thoracica (Latreille, 1802) Present paper

336 Scytodidae Scytode s univittat a Simon, 1882 Present paper

337 Scytodidae Scytodes velutina (Heineken & Lowe, 1832) Present paper

338 Segestriidae Ariadna cyprusensis (Wunderlich, 2011) Wunderlich, 2011

Spiders of Cyprus 170

339 Segestriidae Segestria bavarica (C. L. Koch, 1843) Present paper

340 Segestriidae Segestria florentina (Rossi, 1790) Present paper

341 Segestriidae Segestria senoculata (Linnaeus, 1758) Present paper

342 Sicariidae Loxosceles rufescens (Dufour, 1820) Logunov, 2013

343 Sparassidae Eusparassus walckenaeri (Audouin, 1 826) Strand, 1907

344 Sparassidae Micrommata formosa Pavesi, 1878 Present paper

345 Sparassidae Micrommata ligurina (C. L. Koch, 1845) Present paper

346 Sparassidae Micrommata virescens (Clerck, 1757) Present paper

347 Sparassidae Olios suavis (O. Pickard - Cambridge, 1876) Jäger et al. 2011 348 Tetragnathidae Metellina mengei (Blackwall, 1869) Present paper

349 Tetragnathidae Tetragnatha demissa L. Koch, 1872 Present paper

350 Tetragnathidae Tetragnatha intermedia Kulczyński, 1891 Present paper

351 Tetragnathidae Tetragnatha nitens (Audouin, 1826) Present paper

352 Tetragnathidae Tetragnatha obtusa C. L. Koch, 1837 Present paper

353 Tetragnathidae Zygiometella perlongipes (O. Pickard - Cambridge, 1872) Present paper

354 Theraphosidae Chaetop elma karlamani Vollmer, 1997 Vollmer, 1997

355 Theraphosidae Chaetopelma olivaceum (C. L. Koch, 1841) Ausserer, 1871

356 Theridiidae Anelosimus pulchellus (Walckenaer, 1802) Present paper

357 Theridiidae Anelosimus vittatus (C. L. Koch, 1836) Present pa per

358 Theridiidae Argyrodes argyrodes (Walckenaer, 1841) Present paper

359 Theridiidae Crustulina scabripes Simon, 1881 Present paper

360 Theridiidae Dipoena galilaea Levy & Amitai, 1981 Present paper

361 Theridiidae Dipoena melanogaster (C. L. K och, 1837) Present paper

362 Theridiidae Enoplognatha gemina Bosmans & Van Keer, 1999 Present paper

363 Theridiidae Enoplognatha giladensis (Levy & Amitai, 1982) Present paper

364 Theridiidae Enoplognatha macrochelis Levy & Amitai, 1981 Bosmans & Van Keer, 1999

365 Theridiidae Enoplognatha mariae Bosmans & Van Keer, 1999 Present paper

366 Theridiidae Enoplognatha mediterranea Levy & Amitai, 1981 Bosmans & Van Keer, 1999

367 Theridiidae Enoplognatha parathoracica Levy & Amitai, 1981 Present paper

368 Theridiidae Enoplognatha penelope Hippa & Oksala, 1982 Present paper

369 Theridiidae Enoplognatha quadripunctata Simon, 1884 Present paper

370 Theridiidae Enoplognatha thoracica (Hahn, 1833) Present paper

371 Theridiidae Episinus maculipes Cav anna, 1876 Present paper

372 Theridiidae Episinus truncatus Latreille, 1809 Present paper

Spiders of Cyprus 171

373 Theridiidae Euryopis episinoides (Walckenaer, 1847) Logunov, 2013

374 Theridiidae Euryopis laeta (Westring, 1861) Present paper

375 Theridiidae Euryopis q uinqueguttata Thorell, 1875 Present paper

376 Theridiidae Euryopis sexalbomaculata (Lucas, 1846) Present paper

377 Theridiidae Kochiura aulica (C. L. Koch, 1838) Present paper

378 Theridiidae Lasaeola convexa (Blackwall, 1870) Present paper

379 Ther idiidae Lasaeola coracina (C. L. Koch, 1837) Present paper

380 Theridiidae Lasaeola testaceomarginata Simon, 1881 Present paper

381 Theridiidae Latrodectus dahli Levi, 1959 Present paper

382 Theridiidae Latrodectus geometricus C. L. Koch, 1841 Knofla ch & Van Harten, 2002

383 Theridiidae Latrodectus tredecimguttatus (Rossi, 1790) Present paper

384 Theridiidae Neospintharus syriacus (O. Pickard - Cambridge, 1872) Knoflach et al. 2008

385 Theridiidae Neottiura herbigrada (Simon, 1873) Knoflach, 1999

3 86 Theridiidae Neottiura suaveolens (Simon, 1880) Present paper

387 Theridiidae Neottiura uncinata (Lucas, 1846) Logunov, 2013

388 Theridiidae Paidiscura dromedaria (Simon, 1880) Present paper

389 Theridiidae Pholcomma gibbum (Westring, 1851) Present p aper

390 Theridiidae Phycosoma inornatum (O. Pickard - Cambridge, 1861) Present paper

391 Theridiidae Phylloneta impressa (L. Koch, 1881) Present paper

392 Theridiidae Platnickina nigropunctata (Lucas, 1846) Present paper

393 Theridiidae Rhomphaea l ongicaudata O. Pickard - Cambridge, 1872 Present paper

394 Theridiidae Simitidion agaricographum (Levy & Amitai, 1982) Knoflach, 1996

395 Theridiidae Simitidion simile (C. L. Koch, 1836) Present paper

396 Theridiidae Steatoda albomaculata (Degeer, 177 8) Present paper

397 Theridiidae Steatoda erigoniformis (O. Pickard - Cambridge, 1872) Present paper

398 Theridiidae Steatoda grossa (C. L. Koch, 1838) Present paper

399 Theridiidae Steatoda paykulliana (Walckenaer, 1806) Logunov, 2013

400 Theridiida e Steatoda trianguloides Levy, 1991 Present paper

401 Theridiidae Steatoda triangulosa (Walckenaer, 1802) Present paper

402 Theridiidae Theridion cinereum Thorell, 1875 Present paper

403 Theridiidae Theridion cyprusense Wunderlich, 2011 Wunderlich, 201 1

404 Theridiidae Theridion helena Wunderlich, 2011 Present paper

405 Theridiidae Theridion italiense Wunderlich, 1995 Present paper

406 Theridiidae Theridion melanostictum O. Pickard. - Cambridge, 1876 Knoflach, 1998

Spiders of Cyprus 172

407 Theridiidae Theridion ochreolum Levy & Amitai, 1982 Present paper

408 Thomisidae Bassaniodes caperatus ( Simon, 1875 ) Present paper

409 Thomisidae Bassaniodes pseudorectilineus (Wunderlich, 1995) Present paper

410 Thomisidae Bassaniodes tenebrosus (Silhavy, 1944) Present paper

411 Thomisidae Bassaniodes tristrami (O. Pickard - Cambridge, 1872) Present paper

412 Thomisidae Cozyptila thaleri Marusik & Kovbyuk, 2005 Present paper

413 Thomisidae Diaea livens Simon, 1876 Present paper

414 Thomisidae Heriaeus graminicola (Doleschall, 1852) Present paper

415 Thomisidae Heriaeus simoni Kulczyński, 1903 Present paper

416 Thomisidae Misumena vatia (Clerck, 1757) Present paper

417 Thomisidae Monaeses israeliensis Levy, 1973 Present paper

418 Thomisidae Ozyptila rigida (O. Pickard - Ca mbridge, 1872) Present paper

419 Thomisidae Ozyptila tricoloripes Strand, 1913 Kulczyński,1908

420 Thomisidae Ozyptila sp. Present paper

421 Thomisidae Pistius truncatus (Pallas, 1772) Present paper

422 Thomisidae Runcinia grammica (C. L. Koch, 1837) Present paper

423 Thomisidae Synema globosum (Fabricius, 1775) Present paper

424 Thomisidae Synema plorator (O. P - Cambridge, 1872) Present paper

425 Thomisidae Thomisus citrinellus Simon, 1875 Present paper

426 Thomisidae Thomisus onustus Walckena er,1805 Logunov, 2013

427 Thomisidae Thomisus zyuzini Marusik & Logunov, 1990 Present paper

428 Thomisidae Tmarus piochardi (Simon, 1866) Present paper

429 Thomisidae Xysticus cor Canestrini, 1873 Present paper

430 Thomisidae Xysticus edax (O. Picka rd - Cambridge, 1872) Present paper

431 Thomisidae Xysticus laetus Thorell, 1875 Present paper

432 Thomisidae Xysticus pseudoluctuosus Marusik & Logunov, 1995 Present paper

433 Thomisidae Xysticus xerodermus Strand, 1913 Present paper

434 Titanoecida e Nurscia albomaculata (Lucas, 1846) Present paper

435 Titanoecidae Nurscia albosignata Simon, 1874 Present paper

436 Titanoecidae Titanoeca sp. Present paper

437 Trachelidae Trachelas minor O. Pickard - Cambridge, 1872 Present paper

438 Uloboridae H yptiotes paradoxus (C. L. Koch, 1834) Present paper

439 Uloboridae Uloborus plumipes Lucas, 1846 Present paper

440 Uloboridae Uloborus walckenaerius Latreille, 1806 Present paper

Spiders of Cyprus 173

441 Zodariidae Lachesana blackwalli (O. Pickard - Cambridge, 1872) Kulczy ński,1908

442 Zodariidae Palaestina expolita O. Pickard - Cambridge, 1872 Present paper

443 Zodariidae Zodarion barbarae Bosmans, 2009 Present paper

444 Zodariidae Zodarion granulatum Kulczyński, 1908 Kulczyński,1908

445 Zodariidae Zodarion lutipes (O. Pickard - Cambridge, 1872) Bosmans, 2009

446 Zodariidae Zodarion reticulatum Kulczyński, 1908 Kulczyński,1908

447 Zodariidae Zodarion thoni Nosek, 1905 Kulczyński,1908

448 Zoropsidae Akamasia cyprogenia Bosselaers, 1997 Bosselaers, 1997

449 Zoropsidae Z oropsis lutea (Thorell, 1875) Kulczyński,1911

Spiders of Cyprus