G. Thomas WATTERS 1315 Kinnear Road, Ohio State University, Columbus, Ohio 43212 USA
Total Page:16
File Type:pdf, Size:1020Kb
G. T. - WATTERS & G. DUFFY NOVAPI-X 11(1): 1 1 2, 20 mars 20 1 New species of Annulariidae (Gastropoda) from the Bahamas and Dominican Republic G. Thomas WATTERS 1315 Kinnear Road, Ohio State University, Columbus, Ohio 43212 USA Watters. 1 @osu.edu Glenn DUFFY 5679 Old Ranch Road, Sarasota, Florida 34241 USA KEYWORDS. Gastropoda, Annulariidae, Bahamas, Dominican Republic. ABSTRACT. Ten new taxa of Annulariidae are described from the Bahamas and the Dominican Republic: Abbottella (Abbottella) harpeza n. sp.; A. (A.) mellosa n. sp.; A. {A.) milleaccmtha n. sp.; Chondropoma (Chondropoma) marmoreum n. sp.; C. {Wetmorepoma) ociileum n. sp.; Chondropomiiim hooksi n. sp.; C. alyshae n. sp.; C. piimiliim n. sp.; Chondropomella elegtms n. sp. from the Dominican Republic; and Opisthosiphon (Opisthosiphona) simpsoni williamsae n. subsp. from the Bahamas. Chondropomella is removed from the Chondropominae to the Tudorinae. INTRODUCTION Abbottella (Abbottella) harpeza n. sp. Figs 1-3 The family Annulariidae is one of the largest land snail familles in the world despite its relatively narrow Type material. Holotype UF 420731 (5.6 mm distribution in the Caribbean région. Many species maximum length, including peristome x 7.1 mm hâve very narrow ranges. Hispaniola, and to a lesser maximum width, including peristome). extent the Bahamas, are areas of particularly high diversity of thèse snails. The annulariids of thèse Type locality, Dominican Republic, Isla Beata. islands were covered in détail by Bartsch (1946) and it Distribution. the locality. seemed that there was little left to be done. Known only from type Nevertheless, récent collections there reveal no Material Examined. UF 420731, Dominican apparent end to the diversity of thèse snails - ten new Republic, Isla Beata, under leaf mold (holotype), May, taxa are described hère, most from a fairly narrow 1993,G. Duffy! région of the Barahona Peninsula, an area known to harbor numerous endémies and peculiar taxa. Description. Shell small (holotype 5.6 mm maximum length, including peristome x 7.1 mm maximum Repositories width, including peristome), helicoid, umbilicus wide (holotype 26% of maximum width), circular, and open BMNH: The Natural History Muséum, London, UK. to apex. Nuclear whorls 1 Va, scarcely demarcated GTW: Collection of G. Thomas Watters, Columbus, from the teleoconch, smooth, prominent. Teleoconch Ohio, USA. of 2 Va whorls, adnate except for immediately behind OSUM: The Ohio State University Muséum of the peristome. Suture deeply impressed. Peristome Biological Diversity, Columbus, Ohio, USA. double (holotype 2.1 mm diameter maximum inner UF: Florida Muséum of Natural History, Gainesville, aperture height x 2.0 mm diameter maximum inner Florida, USA. aperture width; holotype 3.6 mm diameter maximum outer peristome height x 3.5 mm diameter maximum SYSTEMATICS outer peristome width, but broken). Outer lip fairly thick, widely expanded, tluted, perpendicular to the Superfamily LITTORINOIDEA whori except posteriorly where it forms a wide Family ANNULARIIDAE Henderson & Bartsch, concave auricle adhèrent to the previous whorl, 1920 ventrally covered with numerous erect, concentric Subfamily ANNULARIINAE Henderson & Bartsch, lamellae. inner lip exserted, very short. Spiral 1920 sculpture of low, squarish threads (~26 on the final Genus Abhotfella Henderson & Bartsch, 1920 whorl), every flfth one or so stronger than intervening Type species Chondropomum niorcletiainim Crosse, threads, becoming stronger and more widely separated 1873 (by original désignation) Récent, Hispaniola, towards the umbilicus. Axial sculpture of closely Cuba. spaced, tlnc lamellae belwecn which are numerous 1 G. T. Wattkrs «S: G. Duffy New spccics t)!" Amiulaiiidac microscopic lamellae. Intersections ol" spiral and axial immediately bclmul the peristome. Suture impressed. sculptures produccd into sliarp. ercct denticles, cxcept Peristome double (holotype 2.6 mm diameter in the umbilicus. Suture bounded by nunierous maximum inner aperture height x 2.7 mm diameter denticles. I^ackground color tan with darker spiral maximum inner aperture width; holotype 3.9 mm bands on top orwliorls, breaking up into linear spots diameter maximum outer peristome height x 3.7 mm on the base and umbilicus. Both sides ol'outer lip with diameter maximum outer peristome width). Outer lip bold. radiating brown bands. Operculuni nuillispiral fairly thick, Huted, perpendicular to the whori at its with obliqucK projecting laniella. base, concave at the apex, widely expanded, barely adnate to the previous whorl. Inner lip exserted, very Discussion. Abbottella harpeza inost closely short. Spiral sculpture of feeble threads (--30 on the resembles .-(. crossci (Piisbry, 1933). Pilsbry gave llnal whorl), widely separated, becoming stronger onl\ "Sanlo Domingo" as the type local ily for A. towards the umbilicus. Axial sculpture of numerous crossei. referring to the island rather than the city. feeble threads between which are microscopic Bartsch (1946) identitled Pilsbry 's species with lamellae. Spiral and axial sculptures intersect forming spécimens from Samanà Bay in the iiorthcastern square pits, the junctures produced into very weak Dominican Republic. Ahhottclla harpeza is more nodules. At the suture the axial sculpture forms darkl\ colored than .(. crossci and the ihorny sculpture minute cusps. On the holotype every tlfth axial or so is more pronounced; the denticles are more aligned forms a peculiar enlarged scale at the suture; this is with the spiral sculpture than the axial sculpture in A. variable on other spécimens examined. Shell honey- harpeza. the opposite is truc of .1. crossci. Species of colored, waxy in appearance, with pale tan spots Abbottella. including A. crossei. occiir along the arranged in a barely discernable spiral pattern. Both northern coast of Hispaniola and eastemmost Cuba, sides of outer lip with radiating brown bands. although a few species hâve not been locali/ed since Operculum multispiral with obliquely projecting their descriptions. This species from Isia Beata and A. lamella. milleacantha n. sp. from Isla Saona are the only species known from the southern coast of Hispaniola. Discussion. Abbottella mellosa is characterized by its Although known from a single spécimen, this species subdued sculpture and waxy appearance; it is probably is sufficiently distinct and geographically isolated to the least sculptured of ail the Abbottella. Only A. warrant description. adolfi (Pfeiffer, 1852) approaches it in this regard. Abbottella adolfi has not been localized although Etymology. Greek harpeza. a thorny hedge, in Bartsch (1946) described the subspecies peninsularis référence to the prickly sculpture of the shell. from the Samanâ Peninsula. Abbottella mellosa differs in having even flner, almost obsolète sculpture, the Abbottella (Abbottella) mellosa n. sp. peculiar suturai tufts, and a lighter color. Figs 4-6 Etymology. Latin mellosa. honey-colored, in Type material. Holotype UF 420729 (6.9 mm référence to the color of the shell. maximum length, including peristome x 7.9 mm maximum width, including peristome). Paratype Abbottella (Abbottella) milleacantha n. sp. OSUM 32477, from type locality (5.6 mm maximum Figs 7-9 length. including peristome x 6.6 mm maximum width, including peristome). Type material. Holotype UF 420728 (5.7 mm maximum length, including peristome x 7.5 mm Type locality. Dominican Republic, Los Brazos, near width, including peristome). Paratype OSUM 32478, Sosùa. from type locality (5.5 mm maximum length, including peristome x 7.8 mm width, including Distribution. Known only from the type locality. peristome). Material Examined. Dominican Republic, Los Type locality. Dominican Republic, northeast Isla Brazos, near Sosùa, on limestone after rain, April, Saona. 2004, A. Bodart & M. Coltro! (holotype, paratype, GTW 14055a [4 dd]) Distribution. Known only from the type locality. Description. Shell small (holotype 6.9 mm maximum Material examined. Dominican Republic, northeast length, including peristome x 7.9 mm maximum Isla Saona, on limestone rocks. May. 1997, G. Duffy! width, including peristome), helicoid, umbilicus wide (holotype & paratype). (holotype 26% of maximum width), elliptical, and open to apex. Nuclear whoris 1 Va. scarcely Description. Shell small (holotype 5.7 mm maximum demarcated from the teleoconch, smooth, prominent. length, including peristome x 7.5 mm width, including Teleoconch of 3 I/2 whoris, adnate except for peristome), helicoid, umbilicus wide (holotype 27% of G.T. WATT1£RS&G. DUFFY NOVAPI.X 11(1): 1-12, 20 mars 2010 maximum width), circular, and open to apex. Nuclear decollate x 5.2 mm maximum width. including whorls 1 1/4, scarcely demarcated from the teleoconch. peristome). Paratype OSUM 32479, Berry Islands, smooth, prominent. Teleoconch of 2 % whorls, adnate Chub Cay, east end of airport runway, under rubble except for immediately behind the peristome. Suture (11.5 mm maximum lenglh. including peristome, impressed. Peristome double (holotype 2.8 mm decollate x 6.0 mm maximum width, including diameter maximum inner aperture height x 2.5 mm peristome). diameter maximum inner aperture width; holotype 3.9 mm diameter maximum outer peristome height x 3.3 Type locaiity. Bahamas, Andros Island, Twin Lakes mm diameter maximum outer peristome width). Outer Farm along Fresh Creek. lip thin, tluted, perpendicular to the whorl, narrow except where the posterior edge is produced into a Material Examined. Bahamas. OSUM 6214. Andros triangular auricle adhèrent with