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Home , African Invertebrates, Ananteris, Bothriuridae, Brachistosternus, Chactidae, Charinidae

Curriculum Vitae: Lorenzo Prendini, Ph.D.

Curator of Arachnida & Myriapoda Division of Invertebrate American Museum of Natural History Central Park West at 79th Street New York, NY 10024-5192, USA

tel: +1-212-769-5843; fax: +1-212-769-5277; email: [email protected] @ AMNH: http://scorpion.amnh.org Division of Invertebrate Zoology: http://www.amnh.org/our-research/staff-directory/lorenzo-prendini

Education Ph.D., Zoology, University of Cape Town, South (2001) B.Sc. (Hons), Zoology, University of Cape Town, South Africa (1995) B.Sc., Botany & Zoology, University of the Witwatersrand, Johannesburg, South Africa (1994)

Appointments Curator, Division of Invertebrate Zoology, AMNH (2012–) Professor, Richard Gilder Graduate School of Comparative Biology, AMNH (2008–) Associate Curator, Division of Invertebrate Zoology, AMNH (2007–2012) Assistant Curator, Division of Invertebrate Zoology, AMNH (2002–2007)

Adjunct Appointments Adjunct Professor, Department of Entomology, Colorado State University (2008–) Research Associate, National Museum of Namibia (2006–) Research Associate, Ditsong National Museum of Natural History, Pretoria, South Africa (2004–) Adjunct Professor, Department of Entomology, Cornell University (2004–) Adjunct Professor, Department of Biology (, and Behavior), CUNY (2003–)

Associate Editor/Editorial Board Member Iranian Journal of Biosystematics (2018–) European Journal of (2012–) and Phylogeny (, Pantopoda) (2010–) ISRN Zoology (2010–2014) American Museum Novitates and Bulletin of the American Museum of Natural History (2008–) ZOOTAXA (, , Scorpiones, , ) (2005–) Cladistics (2004–) Journal of Afrotropical Zoology (2004–2011)

Professional Affiliations African Arachnological Society; American Arachnological Society; Entomological Society of Southern Africa; The Explorer’s Club (Fellow); International Society of Arachnology (Executive Committee, 2007–2010); Society of Systematic Biologists; South African Society of Systematic Biology; Willi Hennig Society (Fellow; Council, 2008–2011); Zoological Society of Southern Africa.

Publications González-Santillán, E. and Prendini, L. 2018. Systematic revision of the North American syntropine vaejovid genera Balsateres, Kuarapu, and Thorellius with descriptions of three new . Bulletin of the American Museum of Natural History 420: 1–81. Esposito, L.A., Yamaguti, H.Y., Pinto-da-Rocha, R. and Prendini, L. 2018. Plucking with the plectrum: Phylogeny of the New World buthid scorpion subfamily Centruroidinae Kraus, 1955 (Scorpiones: ) reveals evolution of three pecten-sternite stridulation organs. Arthropod Systematics and Phylogeny 76(1): 87–122. Schaffrath, S., Prendini, L., and Predel, R. 2018. Intraspecific variation in southern African scorpion species of the genera , and (Scorpiones: Buthidae, ). Toxicon 144: 83–90.

1 Curriculum Vitae: Lorenzo Prendini, Ph.D.

Barrales Alcalá, D.A., Francke, O.F., and Prendini, L. 2018. Systematic revision of the giant North American vinegaroons, Mastigoproctus (Thelyphonida: ). Bulletin of the American Museum of Natural History 418: 1–62. Loria, S.F., and Prendini, L. 2018. Ultrastructural comparison of the eyespot and ocelli of , and implications for the systematics of Chaerilidae Pocock, 1893. Zoologischer Anzeiger 273: 183–191. DOI: 10.1016/j.jcz.2017.12.011. Silva de Miranda, G., Giupponi, A.P.L., Prendini, L., and Scharff, N. 2018. Weygoldtia, a new genus of Charinidae Quintero, 1986 (Arachnida, Amblypygi) with a reappraisal of the genera in the family. Zoologischer Anzeiger 273: 23–32. DOI: 10.1016/j.jcz.2018.02.003. Rubin, M., Lamsdell, J.C., Prendini, L. and Hopkins, M.J. 2017. Exocuticular hyaline layer of sea scorpions and horseshoe crabs suggests cuticular fluorescence is plesiomorphic and non- adaptive in scorpions. Journal of Zoology (London) 303: 245–353. DOI: 10.1111/jzo.12493. Esposito, L.A., Yamaguti, H.Y., Souza, C.A., Pinto-da-Rocha, R. and Prendini, L. 2017. Systematic revision of the Neotropical club-tailed scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, revalidation of Heteroctenus, and description of two new genera and three new species (Buthidae: Rhopalurusinae). Bulletin of the American Museum of Natural History 415: 1–134. Rogers, D.C., et al. [list of authors including Prendini, L.] 2017. Images are not and should not ever be type specimens: A rebuttal to Garraffoni & Freitas. Zootaxa 4269(4): 455–459. Wheeler W.C., Coddington, J.A., Crowley, L.M., Dimitrov, D., Goloboff, P.A., Griswold, C.E., Hormiga, G., Prendini, L., Ramírez, M.J., Sierwald, P., Almeida-Silva, L., Alvarez-Padilla, F., Arnedo, M.A., Benavides Silva, L.R., Benjamin, S.P., Bond, J.E., Grismado, C.J., Hasan, E., Hedin, M., Izquierdo, M.A., Labarque, F.M., Ledford, J., Lopardo, L., Maddison, W.P., Miller, J.A., Piacentini, L.N., Platnick, N.I., Polotow, D., Silva-Dávila, D., Scharff, N., Szuts, T., Ubick, D., Vink, C.J., Wood, H.M., and Zhang, J. 2016. The tree of life: Phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 1–43 DOI: 10.1111/cla.12182. Ceriaco, L.M.P., Gutiérrez, E.E., Dubois, A., et al. [493 coauthors including Prendini, L.] 2016. Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa 4196(3): 435–445. Franz-Guess, S., Klußman-Fricke, B.-J., Wirkner, C.S., Prendini, L., and Starck, J.M. 2016. Morphology of the tracheal system of camel (Chelicerata: Solifugae) based on micro- CT and 3D-reconstruction in exemplar species from three families. Arthropod Structure & Development 45: 440–451. Amorim, D.S., Santos, C.M.D., Krell, F.-T., Dubois, A., Nihei, S.S., Oliveira, O.M.P., Pont, A., Song, H., Verdade, V.K., Fachin, D.A., Klassa, B., Lamas, C.J.E., Oliveira, S.S., De Carvalho, C.J.B., Mello-Patiu, C.A., Hajdu, E., Couri, M.S., Silva, V.C., Capellari, R.S., Falaschi, R.L., Feitosa, R.M., Prendini, L., Pombal Jr., J.P., Fernández, F., Rocha, R.M., Lattke, J.E., Caramaschi, U., Duarte, M., Marques, A.C., Reis, R.E., Kurina, O., Takiya, D.M., Tavares, M., Fernandes, D.S., Franco, F.L., Cuezzo, F., Paulson, D., Guénard, B., Schlick- Steiner, B.C., Arthofer, W., Steiner, F.M., Fisher, B.L., Johnson, R.A., Delsinne, T.D., Donoso, D.A., Mulieri, P.R., Patitucci, L.D., Carpenter, J.M., Herman, L., and Grimaldi, D. 2016. Timeless standards for species delimitation. Zootaxa 4137(1): 121–128. Prendini, L. 2016. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of , sensu lato. Bulletin of the American Museum of Natural History 407: 1–66. González-Santillán, E. and Prendini, L. 2016. Systematic revision of the North American syntropine vaejovid scorpion genera Maaykuyak, Syntropis, and Vizcaino, with description of the adults of Syntropis williamsi. Bulletin of the American Museum of Natural History 405: 1– 67.

2 Curriculum Vitae: Lorenzo Prendini, Ph.D.

Adams, A., Marais, E., Turner, S., Prendini, L., and Pinshow, B. 2016. Similar burrow architecture of three arid-zone scorpion species implies similar ecological function. Natuurwissenschaften (The Science of Nature) 103: 56. DOI: 10.1007/s00114-016-1374-z. Ceccarelli, F.S., Ojanguren-Affilastro, A.A., Ramírez, M.J., Ochoa, J.A., Mattoni, C.I., and Prendini, L. 2016. Andean uplift drives diversification of the bothriurid scorpion genus . Journal of . [Early Edition: DOI: 10.1111/jbi.12760] Janion-Scheepers, C., Measey, J., Braschler, B., Chown, S.L., Coetzee, L., Colville, J., Dames, J., Davies, A.B., Davis, A., Davies, S., Dippenaar-Schoeman, A., Duffy, G., Fourie, D., Griffiths, C., Haddad, C., Hamer, M., Herbert, D., Hugo-Coetzee, L.E.A., Jacobs, A., Jansen van Rensburg, C., Lamani, S., Lotz, L.N., Louw, S., Lyle, R., Malan, A., Marais, M., Neethling, J.-A., Nxele, T., Plisko, D., Prendini, L., Swart, A., Theron, P., Truter, M., Ueckermann, E., Uys, V.M., Villet, M.H., Willows-Munrow, S., and Wilson, J.R.U. 2016. Soil biota in a megadiverse country: Current knowledge and future directions in South Africa. Pedobiologia – J. Soil Ecol. 59: 129–174. Ojanguren-Affilastro, A.A., Mattoni, C.I., Ochoa, J.A. Ramírez, M.J., Ceccarelli, F.S., and Prendini, L. 2016. Phylogeny, species delimitation and convergence in the South American bothriurid scorpion genus Brachistosternus Pocock 1893: Integrating morphology, nuclear and mitochondrial DNA. Molecular Phylogenetics and Evolution 94: 159–170. Maquart, P.O., Réveillion, F., Prendini, L., Burger, M., Fisher, B.L., and Van Noort, S. 2016. New distribution records for African Amblypygi. African Entomology 24(1): 245–246. Amr, Z., Abed, O.A., Al Share, T., Hamidan, N., and Prendini, L. 2015. New records of Jordanian scorpions. Jordan Journal of Natural History 2: 30–38. Prendini, L. 2015. Three new Uroplectes (Scorpiones: Buthidae) with punctate metasomal segments from tropical central Africa. American Museum Novitates 3840: 1–32. González-Santillán, E. and Prendini, L. 2015. Systematic revision of the North American syntropine vaejovid scorpions with a subaculear tubercle, Konetontli González-Santillán and Prendini, 2013. Bulletin of the American Museum of Natural History 397: 1–78. Prendini, L. 2015. A remarkably small species of Uroplectes Peters, 1861 (Scorpiones: Buthidae) endemic to the Succulent Karoo of the Northern Cape Province, South Africa. African Invertebrates 56(2): 499–513. Cushing, P.E., Graham, M.R., Prendini, L., and Brookhart, J.O. 2015. A multilocus molecular phylogeny of the endemic North American camel spider family Eremobatidae (Arachnida: Solifugae). Molecular Phylogenetics and Evolution 92: 280–293. Bird, T.L., Wharton, R.A., and Prendini, L. 2015. Cheliceral morphology in Solifugae (Arachnida): Primary homology, terminology, and character survey. Bulletin of the American Museum of Natural History 394: 1–355. Talal, S., Tesler, I., Sivan, J., Ben-Shlomo, R., Tahir, H.M., Prendini, L., Snir, S., and Gefen, E. 2015. Scorpion speciation in the Holy Land: Multilocus phylogeography corroborates diagnostic differences in morphology and burrowing behavior among subspecies and justifies recognition as phylogenetic, ecological and biological species. Molecular Phylogenetics and Evolution 91: 226–237. González-Santillán, E. and Prendini, L. 2015. Phylogeny of the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, based on morphology, mitochondrial and nuclear DNA. Cladistics 31(4): 341–405. Mattoni, C.I., García-Hernández, S., Botero-Trujillo, R., Ochoa, J.A., Ojanguren-Affilastro, A.A., Pinto-da-Rocha, R., and Prendini, L. 2015. Scorpion sheds ‘tail’ to escape: Consequences and implications of autotomy in scorpions (Buthidae: Ananteris). PLoS One 10(1): e0116639 (15 pp.). DOI: 10.1371/journal.pone.0116639. Santibáñez-López, C.E., Francke, O.F. and Prendini, L. 2014. Shining a light into the world’s deepest caves: Phylogenetic systematics of the troglobiotic scorpion genus Alacran Francke, 1982 (Typhlochactidae: Alacraninae). Invertebrate Systematics 28: 643–664.

3 Curriculum Vitae: Lorenzo Prendini, Ph.D.

Santibáñez-López, C.E., Francke, O.F. and Prendini, L. 2014. Phylogeny of the North American scorpion genus Peters, 1861 (Scorpiones: ) based on morphology, nuclear and mitochondrial DNA. Arthropod Systematics and Phylogeny 72(3): 257–279. (cover article) Loria, S.F. and Prendini, L. 2014. Homology of the lateral eyes of Scorpiones: A six-ocellus model. PLoS One 9(12): e112913 (30 pp.). DOI: 10.1371/journal.pone.0112913. Santibáñez-López, C.E., Francke, O.F. and Prendini, L. 2014. Kolotl n. gen. (Scorpiones: Diplocentridae) a new scorpion genus from Mexico. American Museum Novitates 3815: 1–14. Foord, S.H., Gelebe, V., and Prendini, L. 2014. Effects of aspect and altitude on scorpion diversity along an environmental gradient in the Soutpansberg, South Africa. Journal of Arid Environments 113: 114–120. Tahir, H.M., Navidpour, S., and Prendini, L. 2014. First reports of Razianus Farzanpay, 1987 (Scorpiones: Buthidae) from Iraq and Pakistan, descriptions of two new species, and redescription of Razianus zarudnyi (Birula, 1903). American Museum Novitates 3806: 1–26. Tahir, H.M. and Prendini, L. 2014. Redescription of latimanus and confirmation of the genus Heterometrus (Scorpiones: Scorpionidae) in Pakistan. American Museum Novitates 3805: 1–23. Cushing, P.E., Casto, P., Knowlton, E.D., Royer, S., Laudier, D., Gaffin, D.D., Prendini, L., and Brookhart, J.O. 2014. Comparative morphology and functional significance of setae called papillae on the of male camel spiders (Arachnida, Solifugae). Annals of the Entomological Society of America 107(2): 510–520. Monod, L. and Prendini, L. 2014. Evidence for Eurogondwana: The roles of dispersal, extinction and vicariance in the evolution and biogeography of Indo-Pacific (Scorpiones: Scorpionoidea). Cladistics 31(1): 71–111. Bryson, R.W., Jr., Prendini, L., Savary, W.E., and Pearman, P.B. 2014. Caves as microrefugia: Pleistocene phylogeography of the troglophilic North American scorpion Pseudouroctonus reddelli. BMC Evolutionary Biology 14(9): 1–16. Miller, A.L., Makowsky, R.A., Formanowicz, D.R., Prendini, L., and Cox, C.L. 2014. Cryptic genetic diversity and complex phylogeography of the boreal North American scorpion, boreus (). Molecular Phylogenetics and Evolution 71: 298–307. González-Santillán, E. and Prendini, L. 2013. Redefinition and generic revision of the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, with descriptions of six new genera. Bulletin of the American Museum of Natural History 382: 1–71. Monod, L., Harvey, M.S., and Prendini, L. 2013. Stenotopic Hormurus Thorell, 1876 scorpions from the monsoon ecosystems of northern , with a discussion on the evolution of burrowing behaviour in Hormuridae Laurie, 1896. Revue suisse de Zoologie 120(2): 281– 346. Graham, M.R., Jaeger, J.R., Prendini, L., and Riddle, B.R. 2013. Phylogeography of Beck's desert scorpion, Paruroctonus becki, reveals diversification in the Eastern California Shear Zone and postglacial expansion in the Great Basin Desert. Molecular Phylogenetics and Evolution 69: 502–513. Bryson, R.W., Jr., Savary, W.E., and Prendini, L. 2013. Biogeography of scorpions in the Pseudouroctonus minimus complex (Vaejovidae) from south-western : Implications of ecological specialization for pre-Quaternary diversification. Journal of Biogeography 40(10): 1850–1860. DOI: 10.1111/jbi.12134. Santibáñez-López, C.E., Francke, O.F. and Prendini, L. 2013. Systematics of the keyserlingii group of Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae), with descriptions of three new species from Oaxaca, Mexico. American Museum Novitates 3777: 1–47. Ochoa, J.A., Rojas-Runjaic, F., Pinto-da-Rocha, R., and Prendini, L. 2013. Systematic revision of the Neotropical genus Chactopsis Kraepelin, 1912 (Scorpiones: ), with

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descriptions of two new genera and four new species. Bulletin of the American Museum of Natural History 378: 1–121. Bryson, R.W., Jr., Riddle, B.R., Graham, M.R., Tilston Smith, B., and Prendini, L. 2013. As old as the hills: Montane scorpions in southwestern North America reveal an ancient association between biotic diversification and landscape history. PLoS One 8(1): e52822 (11 pp.). Murienne, J., Benavides, L.R., Prendini, L., Hormiga, G., and Giribet, G. 2013. Forest refugia in Western and Central Africa as ‘museums’ of Mesozoic . Biology Letters 9: 20120932 (8 pp.). Graham, M.R., Jaeger, J.R., Prendini, L., and Riddle, B.R. 2013. Phylogeography of the Arizona hairy scorpion ( arizonensis) supports a model of biotic assembly in the Mojave Desert and adds a new Pleistocene refugium. Journal of Biogeography 40(7): 1298–1312. DOI: 10.1111/jbi.12079. Klußman-Fricke, B.-J., Prendini, L., and Wirkner, C.S. 2012. Evolutionary morphology of the hemolymph vascular system in scorpions: A character analysis. Arthropod Structure & Development 41: 545–560. Volschenk, E.S., Harvey, M.S. and Prendini, L. 2012. A new species of Urodacus (Scorpiones: Urodacidae) from Western Australia. American Museum Novitates 3748: 1–18. Sissom, W.D., Hughes, G.B., Bryson, R.W., Jr., and Prendini, L. 2012. The vorhiesi group of C.L. Koch, 1836 (Scorpiones: Vaejovidae) in Arizona, with description of a new species from the Hualapai Mountains. American Museum Novitates 3742: 1–19. Prendini, L. 2012. The scorpion king. Among the most long-lived terrestrial . Asian Geographic 87(2): 86–89. Ojanguren-Affilastro, A.A., Mattoni, C.I., Ochoa, J.A., and Prendini, L. 2012. Rumikiru, n. gen. (Scorpiones: ), a new scorpion genus from the Atacama Desert. American Museum Novitates 3731: 1–43. Mattoni, C.I., Ochoa, J.A., Ojanguren-Affilastro, A.A., and Prendini, L. 2012. Orobothriurus (Scorpiones: Bothriuridae) phylogeny, Andean biogeography, and the relative importance of genitalic and somatic characters. Zoologica Scripta 41(2): 160–176. Engelbrecht, I. and Prendini, L. 2012. Cryptic diversity of South African trapdoor spiders: Three new species of Stasimopus Simon, 1892 (Mygalomorphae, Ctenizidae), and redescription of Stasimopus robertsi Hewitt, 1910. American Museum Novitates 3732: 1–42. Prendini, L. 2011. Order Scorpiones C.L. Koch, 1850. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 115–117. Prendini, L. 2011. Order Solifugae Sundevall, 1833. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 118. Prendini, L. 2011. Order Thorell, 1888. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 121. Prendini, L. 2011. Order Thorell, 1876. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 122. Prendini, L. 2011. Order Amblypygi Thorell, 1883. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 154. Prendini, L. 2011. Order Thelyphonida Latreille, 1804. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 155. Prendini, L. 2011. Order Schizomida Petrunkevitch, 1945. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 156. Prendini, L. 2011. Scorpions (Scorpiones). In: Grzimek’s Animal Life (N. Simmons, ed.). Gale Cengage. http://animals.galegroup.com Last modified: December 22, 2011.

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Pizarro-Araya, J., Ojanguren-Affilastro, A.A., and Prendini, L. 2011. First report of an arboreal scorpion (Bothriuridae: Urophonius) from the temperate forests of southern . Gayana 75(2): 166–170. Ojanguren-Affilastro, A.A., Pizarro-Araya, J., and Prendini, L. 2011. New data on Chilean Urophonius Pocock, 1893 (Scorpiones, Bothriuridae), with description of a new species. American Museum Novitates 3725: 1–44. Ochoa, J.A., Ojanguren-Affilastro, A.A., Mattoni, C.I., and Prendini, L. 2011. Systematic revision of the Andean scorpion genus Orobothriurus Maury, 1976 (Bothriuridae), with discussion of the altitude record for scorpions. Bulletin of the American Museum of Natural History 359: 1– 90. Engelbrecht, I. and Prendini, L. 2011. An assessment of the taxonomic resolution of southern African trapdoor spiders (Araneae: Ctenizidae; Cyrtaucheniidae; Idiopidae) and implications for their conservation. Biodiversity and Conservation 20: 3101–3116. Prendini, L., Francke, O.F. and Vignoli, V. 2010. Troglomorphism, trichobothriotaxy and typhlochactid phylogeny (Scorpiones, Chactoidea): More evidence that troglobitism is not an evolutionary dead-end. Cladistics 26: 117–142. (cover article) Prendini, L. and Esposito, L.A. 2010. A reanalysis of Parabuthus (Scorpiones: Buthidae) phylogeny with descriptions of two new Parabuthus species endemic to the Central Namib gravel plains, Namibia. Zoological Journal of the Linnean Society 159: 673–710. Prendini, L. 2010. Order Scorpiones C.L. Koch, 1837. Pp. 321–330 In: Gerlach, J. and Marusik, Y. (Eds.) Arachnida and Myriapoda of the Seychelles Islands. Siri Scientific Press, Manchester, UK. Ojanguren-Affilastro, A.A., Ochoa, J.A., Mattoni, C.I., and Prendini, L. 2010. Systematic revision of the granulatus group of Urophonius Pocock 1893 (Scorpiones, Bothriuridae) with description of a new species from central Chile. American Museum Novitates 3695: 1–40. Ochoa, J.A. and Prendini, L. 2010. The genus Hadruroides Pocock, 1893 (Scorpiones: Iuridae) in : New records and descriptions of six new species. American Museum Novitates 3687: 1–56. Ochoa, J.A., Botero-Trujillo, R., and Prendini, L. 2010. On the troglomorphic scorpion Troglotayosicus humiculum (Scorpiones, Troglotayosicidae), with first description of the adults. American Museum Novitates 3691: 1–19. Mattoni, C., Ochoa, J., Ojanguren-Affilastro, A., and Prendini, L. 2010. Towards an all-species phylogeny of the scorpion family Bothriuridae. Cladistics 26: 216–217. [Abstract] Vignoli, V. and Prendini, L. 2009. Systematic revision of the troglomorphic scorpion family Typhlochactidae (Scorpiones: Chactoidea). Bulletin of the American Museum of Natural History 326: 1–94. Prendini, L., Esposito, L.A., Huff, J.C. and Volschenk, E.S. 2009. Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from Hispaniola. Journal of Arachnology 37: 206–244. Prendini, L. 2009. Assembling the Scorpion Tree of Life: ‘High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni)’ reanalysed. In: Arrivillaga, J., El Souki, M. & Herrera, B. (Eds.) Monografía Digital Sociedad Venezolana de Entomología 2: 1–41. ISBN 978-980-12-3750-1 Huff, J.H. and Prendini, L. 2009. On the African whip scorpion, Etienneus africanus (Hentschel, 1899) (Thelyphonida: Thelyphonidae), with a redescription based on new material from Guinea-Bissau and . American Museum Novitates 3658: 1–17. Francke, O.F., Vignoli, V. and Prendini, L. 2009. A new species of Typhlochactas (Scorpiones, Typhlochactinae) from eastern Mexico. American Museum Novitates 3647: 1–11.

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Dippenaar-Schoeman, A.S., Van den Berg, A. and Prendini, L. 2009. Spiders and scorpions (Arachnida: Araneae, Scorpiones) of the Nylsvley Nature Reserve, South Africa. Koedoe 51(1): 1–9. DOI: 10.4102/koedoe.2009.161 Volschenk, E.S. and Prendini, L. 2008. Aops oncodactylus gen. et sp. nov., the first troglobitic urodacid (Urodacidae: Scorpiones), with a re-assessment of cavernicolous, troglobitic and troglomorphic scorpions. Invertebrate Systematics 22: 235–257. Volschenk, E.S., Mattoni, C.I. and Prendini, L. 2008. Comparative anatomy of the mesosomal organs of scorpions (Chelicerata, Scorpiones), with implications for the phylogeny of the order. Zoological Journal of the Linnean Society 154: 651–675. Vignoli, V. and Prendini, L. 2008. A new species of Akentrobuthus Lamoral, 1976 (Scorpiones: Buthidae) from the Republic of . Journal of Afrotropical Zoology 4: 61–70. Prendini, L. and Bird, T.L. 2008. Scorpions of the Brandberg Massif, Namibia: Species richness inversely correlated with altitude. African Invertebrates 49(2): 77–107. Kamenz, C. and Prendini, L. 2008. An atlas of book lung ultrastructure in the Order Scorpiones (Arachnida). Bulletin of the American Museum of Natural History 316: 1–259. Huff, J.C., Viquez, C., and Prendini, L. 2008. Redescription of Mimoscorpius pugnator (Butler, 1872) (Arachnida: Thelyphonida), with first description of the female. American Museum Novitates 3633: 1–9. Francke, O.F. and Prendini, L. 2008. Phylogeny and classification of the giant hairy scorpions, Hadrurus Thorell (Iuridae Thorell): A reappraisal. Systematics and Biodiversity 6(2): 205–223. Dunlop, J.A., Tetlie, O.E. and Prendini, L. 2008. Reinterpretation of the scorpion Proscorpius osborni (Whitfield): Integrating data from Palaeozoic and Recent scorpions. Palaeontology 51(2): 303–320. Cushing, P.E., Prendini, L., Savary, W.E., Brookhart, J.O., Wharton, R.A., Bird, T., and Gromov, A.V. 2008. Global survey and inventory of Solifugae. American Arachnology 76: 11. [Abstract] Wirkner, C.S. and Prendini, L. 2007. Comparative morphology of the hemolymph vascular system in scorpions – a survey using corrosion casting, MicroCT and 3D-reconstruction. Journal of Morphology 268: 401–413. Ramírez, M.J., Coddington, J.A., Maddison, W.P., Midford, P., Prendini, L., Miller, J., Griswold, C.E., Hormiga, G., Sierwald, P., Scharff, N., Benjamin, S.P. and Wheeler, W.C. 2007. Linking of digital images to phylogenetic data matrices using a morphological ontology. Systematic Biology 56(2): 283–294. Ojanguren Affilastro, A.A., Mattoni, C.I. and Prendini, L. 2007. The genus Brachistosternus (Scorpiones: Bothriuridae) in Chile, with descriptions of two new species. American Museum Novitates 3564: 1–44. Mattoni, C.I. and Prendini, L. 2007. Phylogeny and biogeography of the family Bothriuridae (Scorpiones). Darwiniana 45(supl.): 96–98. Baldo, L., Prendini, L., Corthals, A. and Werren, J. 2007. Wolbachia are present in southern African scorpions and cluster with supergroup F. Current Microbiology 55: 367–373. Prendini, L., Volschenk, E.S., Maaliki, S. and Gromov, A.V. 2006. A ‘living ’ from Central Asia: the morphology of Pseudochactas ovchinnikovi Gromov, 1998 (Scorpiones: Pseudochactidae), with comments on its phylogenetic position. Zoologischer Anzeiger 245: 211–248. (cover article) Prendini, L. 2006. Scorpiones. In: McGraw-Hill Encyclopedia of Science and Technology. AccessScience@McGraw-Hill. DOI 10.1036/1097–8542.607800. Available at: http://www.accessscience.com/email_pdf/2dbdce746a7af4696920181c26717f67.pdf Last modified: April 7, 2006. Prendini, L. 2006. New South African flat rock scorpions (Liochelidae: ). American Museum Novitates 3502: 1–32.

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Prendini, L. and Wheeler, W.C. 2005. Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for in online publishing. Cladistics 21(5): 446–494. Prendini, L., Weygoldt, P. and Wheeler, W.C. 2005. Systematics of the Damon variegatus group of African whip spiders (Chelicerata: Amblypygi): Evidence from behaviour, morphology and DNA. Organisms, Diversity and Evolution 5: 203–236. (cover article) Prendini, L. 2005. New records and observations on the natural history of Lisposoma elegans and Lisposoma josehermana (Scorpiones: Bothriuridae). American Museum Novitates 3487: 1–11. Prendini, L. 2005. Scorpion diversity and distribution in southern Africa: Pattern and process. Pp. 25–68 In: Huber, B.A., Sinclair, B.J. & Lampe, K.-H. (Eds.) African Biodiversity: Molecules, Organisms, Ecosystems. Proceedings of the 5th International Symposium on Tropical Biology, Museum Alexander Koenig, Bonn. Springer Verlag, New York. Prendini, L. 2005. On Hadogenes angolensis Lourenço, 1999 syn. n. (Scorpiones, Liochelidae), with a redescription of H. taeniurus (Thorell, 1876). Revue suisse de Zoologie 112(2): 1–28. Prendini, L. 2005. Comment on “Identifying spiders through DNA barcodes”. Canadian Journal of Zoology 83(3): 481–491. Prendini, L. 2005. Revision of Karasbergia Hewitt (Scorpiones: Buthidae), a monotypic genus endemic to southern Africa. Journal of Afrotropical Zoology 1: 77–93. Prendini, L., and Wheeler, W.C. 2004. Assembling the scorpion Tree of Life: Phylogeny of extant scorpions based on molecules, morphology and exemplars. Cladistics 20: 602–603. [Abstract] Prendini, L., Crowe, T.M., and Wheeler, W.C. 2004. Burrowing scorpion phylogeny and the evolution of paleodrainage systems in southern Africa. Cladistics 20: 93. [Abstract] Prendini, L. 2004. The systematics of southern African Parabuthus Pocock (Scorpiones, Buthidae): Revisions to the taxonomy and key to the species. Journal of Arachnology 32(1): 109–186. Prendini, L. 2004. Systematics of the genus Pseudolychas Kraepelin (Scorpiones: Buthidae). Annals of the Entomological Society of America 97(1): 37–63. Prendini, L. 2004. On the scorpions of Gabon and neighbouring countries, with a reassessment of the synonyms attributed to Babycurus buettneri Karsch and a redescription of Babycurus melanicus Kovařík. In: Fisher, B.L. (Ed.) Monts Doudou, Gabon. A Floral and Faunal Inventory with Reference to Elevational Variation. Memoirs of the California Academy of Sciences 28: 235–267. Druce, D., Hamer, M., Slotow, R. and Prendini, L. 2004. Checklist of (Diplopoda), centipedes (Chilopoda) and scorpions (Arachnida: Scorpionida) from a savanna ecosystem, Limpopo Province, South Africa. African Invertebrates 45: 315–322. Coddington, J.A., Giribet, G., Harvey, M.S., Prendini, L., and Walter, D.E. 2004. Arachnida. Pp. 296–318 In: Cracraft, J. and Donoghue, M. (Eds.) Assembling the Tree of Life. Oxford University Press, Oxford. Prendini, L., Crowe, T.M. and Wheeler, W.C. 2003. Systematics and biogeography of the family Scorpionidae Latreille, with a discussion of phylogenetic methods. Invertebrate Systematics 17(2): 185–259. (cover article) Prendini, L. 2003. Revision of the genus Lisposoma Lawrence, 1928 (Scorpiones: Bothriuridae). Systematics and Evolution 34: 241–264. Prendini, L. 2003. Discovery of the male of Parabuthus muelleri, and implications for the phylogeny of Parabuthus (Scorpiones: Buthidae). American Museum Novitates 3408: 1–24. Prendini, L. 2003. A new genus and species of bothriurid scorpion from the Brandberg Massif, Namibia, with a reanalysis of bothriurid phylogeny and a discussion of the phylogenetic position of Lisposoma Lawrence. Systematic Entomology 28(2): 149–172.

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Prendini, L. 2003. Threats facing southern Africa’s unique scorpion fauna. Colophon, Newsletter of the IUCN/SSC Southern African Invertebrates Specialist Group (SAISG) Newsletter No. 3: 7–11. Prendini, L., Hanner, R. and DeSalle, R. 2002. Obtaining, storing and archiving specimens and tissue samples for use in molecular studies. Pp. 176–248 In: DeSalle, R., Giribet, G. and Wheeler, W.C. (Eds.) Methods and Tools in Biosciences and Medicine. Techniques in Molecular Evolution and Systematics. Birkhäuser Verlag AG, Basel. Nishiguchi, M.K., Doukakis, P., Egan, M., Goldstein, P.Z., Kizirian, D., Phillips, A., Prendini, L., Rosenbaum, H.C., Torres, E., Wyner, Y., DeSalle, R. and Giribet, G. 2002. DNA isolation procedures. Pp. 249–287 In: DeSalle, R., Giribet, G. and Wheeler, W.C. (Eds.) Methods and Tools in Biosciences and Medicine. Techniques in Molecular Evolution and Systematics. Birkhäuser Verlag AG, Basel. Dyason, K., Brandt, W., Prendini, L., Verdonck, F., Tytgat, J., Du Plessis, J., Müller, G. and Van der Walt, J. 2002. Determination of species-specific components in the venom of Parabuthus scorpions from southern Africa using matrix-assisted laser desorption time-of-flight mass spectrometry. Rapid Communications in Mass Spectrometry 16: 768–773. Prendini, L. 2001. Book reviews: New volumes on the “minor” orders. Journal of Arachnology 29(3): 234–237. Prendini, L. 2001. Substratum specialization and speciation in southern African scorpions: the Effect Hypothesis revisited. Pp. 113–138 In: Fet, V. and Selden, P.A. (Eds.) Scorpions 2001. In Memoriam Gary A. Polis. British Arachnological Society, Burnham Beeches, Bucks, UK. Prendini, L. 2001. Further additions to the scorpion fauna of Trinidad and Tobago. Journal of Arachnology 29(2): 173–188. Prendini, L. 2001. Two new species of Hadogenes (Scorpiones, Ischnuridae) from South Africa, with a redescription of Hadogenes bicolor and a discussion on the phylogenetic position of Hadogenes. Journal of Arachnology 29(2): 146–172. Prendini, L. 2001. Species or supraspecific taxa as terminals in cladistic analysis? Groundplans versus exemplars revisited. Systematic Biology 50(2): 290–300. Prendini, L. 2001. Phylogeny of Parabuthus (Scorpiones, Buthidae). Zoologica Scripta 30(1): 13–35. Prendini, L. 2001. A review of synonyms and subspecies in the genus Opistophthalmus C.L. Koch (Scorpiones: Scorpionidae). African Entomology 9(1): 17–48. Prendini, L. 2000. Chelicerata (Scorpiones). In: Kirk-Spriggs, A.H. and Marais, E. (Eds.) Dâures – Biodiversity of the Brandberg Massif, Namibia. Cimbebasia Memoir 9: 109–120. Prendini, L. 2000. A new species of Parabuthus Pocock (Scorpiones, Buthidae), and new records of Parabuthus capensis (Ehrenberg), from Namibia and South Africa. Cimbebasia 16: 201–214. Prendini, L. 2000. Phylogeny and classification of the superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): An exemplar approach. Cladistics 16(1): 1–78. (cover article) Prendini, L., Crowe, T.M., Linder, H.P. and Wheeler, W.C. 1999. Taking the sting out of scorpion systematics: An outline of progress in understanding the systematics of Opistophthalmus C.L. Koch. American Arachnology 60: 6–7. [Abstract] Prendini, L. and Linder, H.P. 1998. Phylogeny of the South African species of restioid leafhoppers (Cicadellidae, Cephalelini). Entomologica scandinavica 29: 11–18. Prendini, L., Linder, H.P. and Picker, M.D. 1997. Parallel cladogenesis, coevolution and sequential evolution in insect-plant associations: A test case. Cladistics 13: 178–179. [Abstract] Prendini, L., Crowe, T.M. and Linder, H.P. 1997. Challenges to scorpion cladistics: Effects of sex, segmentation and psammophily. Cladistics 13: 177–178. [Abstract] Prendini, L. 1997. Two new host-restricted restioid leafhoppers in the genus Cephalelus

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(Cicadellidae: Cephalelini), with descriptions of the females of C. brevipilus Davies, C. daviesi Davies and C. rawsonia Davies. African Entomology 5(2): 273–281. Newlands, G. and Prendini, L. 1997. Redescription of Hadogenes zumpti (Scorpiones: Ischnuridae), an unusual rock scorpion from the Richtersveld, South Africa. South African Journal of Zoology 32(3): 76–81. Prendini, L., Theron, L.-J., Van der Merwe, K. and Owen-Smith, N. 1996. Abundance and guild structure of grasshoppers (Orthoptera: Acridoidea) in communally grazed and protected savanna. South African Journal of Zoology 31(3): 120–130.

Interviews: National Public Radio (http://www.npr.org/templates/story/story.php?storyId=16493817): ‘Fossil shows sea scorpions stretched to 8 feet.’ by J. Hamilton, 21 November 2007. Scholastic SuperScience ‘Scorpions: Friend or Foe?’ (Grades 3–6) by E. Carney, January 2008, pp. 10, 11 (http://www2.scholastic.com/browse/article.jsp?id=3751918). Scholastic Science World ‘Scorpions: Friend or Foe?’ (Grades 6–10) by E. Carney, January 2008, pp. 10, 11 (http://www2.scholastic.com/browse/article.jsp?id=3751917) ‘Scorpions: friend or foe? Do scorpions deserve their bad reputation? One scientist finds out.’ by E. Carney, January 2008(http://findarticles.com/p/articles/mi_hb5019/is_4_19/ai_n29399348/) The Forum Magazine of the American Tarantula Society: ‘Interview with Dr. Lorenzo Prendini by P. MacDonald 2008/2009’ 18(1), 2010: 6–15. Practical Reptile Keeping: Feature – Scorpions. ‘Sting in the Tail: Professor Lorenzo Prendini discusses the world’s scorpions’ by V. Neblik, August 2011: 55–59. CBS News (http://www.cbsnews.com/video/watch/?id=7383853n): ‘In Search of Scorpions’, 7 October, 2011. Also: AMNH Website (http://www.amnh.org/explore/news- blogs/news/scorpion-expert-featured-in-cbs-video2) ‘Scorpion Expert Featured in CBS Video.’ 17 October 2011. AMNH.tv (http://www.amnh.org/explore/amnh.tv/(watch)/meet-the-scientists/profile-lorenzo- prendini) and YouTube (https://www.youtube.com/watch?v=PgsN6dSd3T0), 27 July 2012. Arachne, Deutsche Arachnologische Gesellschaft ‘Interview mit Dr. Lorenzo Prendini, Kurator für Arachniden und Myriapoden im American Museum of Natural History’ by C. Turiel, 6: 34–35, September 2012. Facebook Live – Inverse (https://www.facebook.com/naturalhistory/posts/10153458341886991), 22 June 2016.

Selected Media: Rotunda, AMNH Member’s Newsletter ‘2004 Scientific Expeditions’ December 2004, 29(6): 4–5. Martha Stewart Kids Magazine ‘Beware Scorpions’ Number 19, Fall 2005, pp. 70, 76–81, 143. ‘Fathomless Skin’ (http://www.asianart.com/articles/lacquer/index.html#3): online article comparing the of scorpions and centipedes with Chinese lacquer art by J. Rauer, 13 January, 2006. AMNH Website (http://www.amnh.org/science/papers/scorpion_images.php): ‘Microscopic morphology adds detail to scorpion family tree. Atlas of scanning electron microscope images of the book lung published by the American Museum of Natural History.’ 13 January 2009. Physorg.com (http://www.physorg.com/news150984224.html) and e! Science News (http://esciencenews.com/sources/physorg/2009/01/12/microscopic.morphology.adds.scorpio n.family.tree): ‘Microscopic morphology adds to the scorpion family tree’. Seed Magazine (http://seedmagazine.com/portfolio/22_library-of-lungs.html): ‘Portfolio: A library of lungs.’ June 2009. AMNH Website (http://www.amnh.org/science/papers/species_names.php): ‘The names within the names. AMNH scientists honored with an array of scientific names.’ 27 March, 2009. AMNH Website (http://www.amnh.org/science/papers/prendini_clad_2010.php) and Physorg.com (http://www.physorg.com/news187613062.html): ‘Phylogenetic analysis of Mexican cave scorpions suggests adaptation to caves is reversible. New evidence that specialized adaptations are not evolutionary dead ends.’ 12 March 2010.

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Rotunda, AMNH Member’s Newsletter ‘Spotting New Species’ Winter 2011, 36(1): 10. Wired Science: ‘Scorpions, Spiders and Sharks: Electron-Microscope Images’ by B. Mason, 20 April 2011 (http://www.wired.com/wiredscience/2011/04/sem-gallery/?pid=1232&viewall=true) Scientific American (http://www.scientificamerican.com/blog/post.cfm?id=art-and-science-meet- in-images-of-m-2011-06-18): ‘Art and Science Meet in Images of Museum Specimens and Artifacts’ by M. Bradley, 18 June 2011. Live Science (http://www.livescience.com/14759-amazing-images-science-art-imaging- technologies.html): ‘Amazing Images Reveal the Art of Science’ by W. Parry, 24 June 2011. Live Science (http://www.livescience.com/14758-science-art-gallery-imaging.html): ‘Science as Art: A Gallery’, 24 June 2011. Wired Science (http://www.wired.com/wiredscience/2011/06/picturing-science-exhibit/): ‘Hidden Blades, Glowing Scorpions and Bug Genitalia: Great Science Images from the American Museum of Natural History’ by D. Mosher, 24 June 2011. AMNH Website (http://www.flickr.com//photos/amnh/sets/72157626899911609/show/): ‘Picturing Science: Museum Scientists and Imaging Technologies Photo Slideshow’, 25 June 2011–24 June 2012. Rotunda, AMNH Member’s Newsletter ‘Now on View: Picturing Science’ Summer 2011, 36(3): 3. The New York Times (http://www.nytimes.com/2011/07/12/science/12museum.html?_r=1): ‘Specialized Windows on the Natural World’ by H. Fountain, 7 July 2011. Also: http://www.nytimes.com/slideshow/2011/07/11/science/12Museum-12.html http://www.timesofoman.com/featuredetail.asp?fid=1068 http://m.post-gazette.com/news/science/specialized-windows-on-the-natural-world-1159868 Folha.com (http://www1.folha.uol.com.br/ciencia/943051-microscopia-produz-obras-de-arte-com- estetica-cientifica.shtml): ‘Microscopia produz obras de arte com estética científica’, 13 July, 2011. Time Out New York (http://newyork.timeout.com/arts-culture/museums/1594001/picturing- science-museum-scientists-and-imaging-technologies): ‘Picturing Science: Museum Scientists and Imaging Technologies’, July 2011. The New York Times (http://www.nytimes.com/2012/05/29/science/would-eradicating-deer-- hurt-the-ecosystem.html?_r=0): ‘If deer ticks were eradicated would it hurt the ecosystem?’ by C.C. Ray, 28 May 2012. AMNH Website (http://www.amnh.org/explore/news-blogs/news/training-colleagues-around-the- world-to-collect-scorpions) ‘Training Colleagues Around the World to Collect Scorpions.’ 17 June 2012. Live Science (http://www.livescience.com/21818-spiders-scoprions-exhibit.html): ‘Beware, Arachnophobes: Half of Spiders Are Undiscovered’ by W. Parry, 25 July 2012. AMNH Website (http://www.amnh.org/explore/news-blogs/research/the-arachnid-collection- behind-spiders-alive): ‘The Arachnid Collection Behind Spiders Alive!’. 30 July 2012. National Geographic Omnivore blog (http://newswatch.nationalgeographic.com/2013/03/27/why- did-the-bushman-put-a-scorpion-in-his-mouth-on-the-amazing-race/): ‘Why did the Bushman put a scorpion in his mouth on ‘The Amazing Race’?’ by L. Poon, 27 March 2013. LeMauricien.com (http://www.lemauricien.com/article/considere-eteint-sur-coin-mire- redecouverte-du-scorpion-lychas-serratus): ‘Considere eteint sur le coin de mire: Redecouverte du scorpion “Lychas serratus”’, 13 April 2013. Also: http://www.lemauricien.com/topics/Lorenzo%20Prendini); Indian Ocean Times: ‘An endemic species of scorpion reappears after 150 years of disappearance’ 17 April 2013 (http://en.indian-ocean-times.com/Mauritius-An-endemic-species-of-scorpion-reappears-after- 150-years-of-disappearance_a1162.html]; Zinfos974.com: ‘Maurice: Un scorpion disparu depuis 150 ans refait son apparition’ 18 April 2013 [http://www.zinfos974.com/Maurice-Un- scorpion-disparu-depuis-150-ans-refait-son-apparition_a55870.html). National Geographic (http://phenomena.nationalgeographic.com/2015/01/29/how-the-scorpion- lost-its-tail-and-its-anus/): ‘How the Scorpion Lost Its Tail and Its Anus’ by E. Yong, 29 Januray 2015. AMNH Website (http://www.amnh.org/explore/news-blogs/research- posts/scorpions-detachable-tail-shakes-predators): ‘Scorpion’s Detachable Tail Shakes Predators’ by I. Chant, 5 February 2015.

11 Curriculum Vitae: Lorenzo Prendini, Ph.D.

AMNH press center (http://www.amnh.org/about-us/press-center/a-close-look-at-formidable- camel-spider-jaws) and Eurekalert (http://www.eurekalert.org/pub_releases/2015-06/amon- stc062215.php): ‘Study Takes Close Look at Formidable Camel Spider Jaws’ by K. Snyder, 22 June 2015, and (http://www.amnh.org/explore/news-blogs/research-posts/close-up-on- the-fearsome-jaws-of-camel-spiders): ‘Close Up on the Fearsome Jaws of Camel Spiders’; Live Science (http://www.livescience.com/51377-camel-spider-jaw-study.html): ‘Camel Spider’s Fierce Jaw is Focus of New Creepy Crawly ‘Dictionary’’ by E. Palermo, 29 June 2015; io9 (http://io9.com/to-gaze-into-the-jaws-of-camel-spiders-is-to-gaze-into-1713133140): ‘To Gaze into the Jaws of Camel Spiders is to Gaze into the Jaws of Hell’ by E. Inglis-Arkell, 23 June 2015; International Business Times (http://www.ibtimes.co.uk/fearsome-jaw-camel- spider-unveiled-visual-atlas-graphic-images-1507411): ‘Warning potentially scary images: Fearsome jaw of camel spider unveiled in 'visual atlas'’ by H. Osbourne, 22 June 2015; BBC Earth (http://www.bbc.com/earth/story/20150630-inside-the-jaws-of-camel-spiders): ‘Inside the Jaws of Camel Spiders’ by M. Hogenboom, 30 June 2015; Natural History Magazine (http://www.naturalhistorymag.com/features/122921/the-naming-of-parts): ‘The Naming of Parts’ by T. Bird, November 2015. LabRoots (http://labroots.com/trending/plants-and- /1291/miss-muffet-s-worst-nightmare): ‘Miss Muffet’s Worst Nightmare’ by B.K. Kim, 25 June, 2015. LiveScience (http://www.livescience.com/53701-snake-loses-battle-against-spider-photo.html): ‘Reptile Dysfunction: Snake Loses Wild Battle Against Spider’ by J. Bryner, 12 February 2016. UnDark (http://undark.org/article/zika-and-the-folly-of-mosquito-specicide/): ‘The Allure and Folly of Specicide’ by L. Peeples, 13 May 2016. AMNH News & Blogs (http://www.amnh.org/explore/news-blogs/research-posts/dna-study- suggests-scorpions-diversified-as-andes-formed): ‘DNA Study Suggests Scorpions Diversified as Andes Formed’ by K. Snyder, 16 May 2016. AMNH press center (http://www.amnh.org/about-the-museum/press-center/scorpions-have- similar-tastes-in-burrow-architecture) : ‘Scorpions Have Similar Tastes in Burrow Architecture’ by K. Snyder, 21 June 2016. Also: https://www.sciencedaily.com/releases/2016/06/160621143632.htm; https://phys.org/news/2016-06-scorpions-similar-burrow-architecture.html; http://esciencenews.com/sources/newswise.scinews/2016/06/21/scorpions.have.similar.taste s.burrow.architecture; Science News (http://www.upi.com/Science_News/2016/06/21/Burrow- building-scorpions-have-consistent-tastes/5281466538123/): ‘Burrow-building scorpions have consistent tastes’ by B. Hays, 21 June 2016. IFL Science (http://www.iflscience.com/plants- and-animals/scorpions-similar-tastes-interior-burrow-design/page-2/): ‘Scorpions Have Similar Tastes In Interior Burrow Design’, 22 June 2016. PopSci (http://www.popsci.com/aluminum-casts-scorpion-burrows): ‘Different scorpion species share similar taste in interior design’ by C. McDonald, 23 June 2016. Live Science (http://www.livescience.com/55236-scorpions-burrows-revealed-in-3d.html): ‘Scorpion Architects Build Lairs with Porches and Mating Rooms’ by M. Weisberger, 29 June 2016. The Site (http://blog.therainforestsite.com/cs-scorpion-burrows): ‘Scorpions Everywhere Love to Build Burrows’, 20 July 2016. AMNH News & Blogs (http://www.amnh.org/explore/news-blogs/research-posts/camel-spider- respiration-makes-them-potent-predators): ‘Respiration Makes Camel Spiders Potent Predators’ by A. Chant, 15 September 2016. Also: https://article.wn.com/view/2016/09/15/Respiration_Makes_Camel_Spiders_Potent_Predator s_American_Mu/ California Academy of Sciences (http://www.calacademy.org/press/releases/three-new- %E2%80%9Cclub-tailed%E2%80%9D-scorpions-join-the-tree-of-life): ‘Three New 'Club- Tailed' Scorpions Join the Tree of Life’, 24 July 2017. Also: EurekAlert! (https://www.eurekalert.org/pub_releases/2017-07/caos-tn072117.php); PhysOrg (https://phys.org/news/2017-07-club-tailed-scorpions-tree-life.html); ScienceDaily (https://www.sciencedaily.com/releases/2017/07/170724090753.htm); Jersey Tribune (http://jerseytribune.com/2017/07/24/three-new-club-tailed-scorpions-join-the- tree-of-life); and Science News (http://www.sci-news.com/biology/three-new-species-club-

12 Curriculum Vitae: Lorenzo Prendini, Ph.D.

tailed-scorpions-05083.html): ‘Arachnologists Discover Three New Species of 'Club-Tailed' Scorpions’, 28 July 2017. New York Times (https://www.nytimes.com/2017/08/04/science/new-scorpions-species.html): ‘Scorpions Are Ancient, but Some Species Are New to Science’ by J. Gorman, 4 August 2017. Live Science (https://www.livescience.com/61999-tarantula-spider-rescued-from-flood.html): ‘Faced with Drowning, This Giant Tarantula Goes Out on a Limb’ by L. Geggel, 12 March 2018.

Oral Presentations Plenary: Inaugural Conference, South African Society for Systematic Biology (1999); 5th International Symposium on Tropical Biology, Museum Alexander Koenig, Bonn (2004); XXI Congreso Venezolano de Entomología, Universidad Simón Bolívar, Caracas (2009); 26th European Congress of Arachnology, Ben-Gurion University of the Negev, Israel (2011); University of Sargodha, Pakistan (2012). Invited: University of Cape Town (1995, 1998, 2000); University of Pretoria (2000); Symposium, Scorpion Toxins from southern Africa, Katolieke Universiteit, Leuven (1998); AMNH, New York (2000, 2002, 2005, 2007, 2008, 2012); New Mexico State University, Las Cruces (2002); Cornell University, Ithaca (2003); Major Systematic Entomology Facilities Meeting, Plant Protection Research Institute, Pretoria (2003); Zoologisches Museum der Universität Humboldt, Berlin (2004); Friedrich-Schiller-Universität, Jena (2004); Villanova University, PA (2005); Natural History Museum of Zimbabwe, Bulawayo (2013); Rutgers University, Newark (2013); Columbia University Club, New York (2013); Dana Biosphere Reserve, Jordan (2013); Systematics Association Biennial Meeting, Oxford University, UK (2015); South African National Biodiversity Institute, Kirstenbosch, Cape Town (2018). Contributed: 5th International Colloquium, Research Group for the Study of African (1996); 15th Meeting, Willi Hennig Society (1996); Symposia, Zoological Society of Southern Africa (1996, 1997); Joint congress: Entomological Society of Southern Africa (11th Congress) and African Association of Insect Scientists (12th Congress) (1997); XIV International Congress of Arachnology and 22nd Annual Meeting, American Arachnological Society (1998); 23rd Annual Meeting, American Arachnological Society (1999); 6th International Colloquium, African Arachnological Society (1999); 15th International Congress of Arachnology (2001), convened symposium on Scorpiology; 23rd Meeting, Willi Hennig Society (2004); 16th International Congress of Arachnology (2004); 17th International Congress of Arachnology (2007); 19th International Congress of Arachnology (2013). Coauthored: 15th International Congress of Arachnology (2001); Symposium, Assembling the Tree of Life: Science, Relevance, and Challenges, AMNH, New York (2002); III Encuentro de Aracnólogos del Cono Sur (2003); IV Encuentro de Aracnólogos del Cono Sur (2003); 19th Annual Meeting of the Society for the Protection of Natural History Collections (2004); 16th International Congress of Arachnology (2004); Assembling the Tree of Life Principal Investigators’ Meeting, National Science Foundation, Arlington, VA (2004); Primer Congreso Latinoamericano de Aracnología (2005); 17th International Congress of Arachnology (2007); 32nd Annual Meeting, American Arachnological Society (2008); 24th European Colloquium of Arachnology (2008); 27th Meeting of the Willi Hennig Society and VIII Reunión de Cladística y Biogeografía (2008); 2 Congreso Latinoamericano de Aracnología (2008); 25th European Colloquium of Arachnology (2009); 29th Meeting, Willi Hennig Society (2010); 18th International Congress of Arachnology (2010); 2nd International Congress on Invertebrate Morphology (2011); 35th Annual Meeting of the American Arachnological Society (2011); 26th European Congress of Arachnology (2011); 3rd Latin American Congress of Arachnology (2011); X Reunion Argentina de Cladística y Biogeografía, Mendoza, Argentina (2012); 27th European Congress of Arachnology (2012); Young Systematists’ Forum, London (2012); 19th International Congress of Arachnology (2013); 39th Annual Meeting of the American Arachnological Society (2015); 20th International Congress of Arachnology (2016); 4th Conference of the Asian Society of Arachnology (2017).

Posters 16th International Congress of Arachnology (2004); Expanding the Ark: The Emerging Science

13 Curriculum Vitae: Lorenzo Prendini, Ph.D. and Practice of Invertebrate Conservation. Spring Symposium, Center for Biodiversity and Conservation, American Museum of Natural History, New York (2004); Assembling the Tree of Life Principal Investigators’ Meeting, National Science Foundation, Arlington, VA (2004); 22nd European Colloquium of Arachnology (2005); 23rd European Colloquium of Arachnology (2006); 31st Annual Meeting, American Arachnological Society (2007); 17th International Congress of Arachnology (2007); 2 Congreso Latinoamericano de Aracnología (2008); 14th Natural History Symposium, Gerace Research Center, San Salvador Island, Bahamas (2011); 35th Annual Meeting of the American Arachnological Society (2011); 26th European Congress of Arachnology (2011); 3rd Latin American Congress of Arachnology (2011); Society for Integrative and Comparative Biology, Annual Meeting (2014); Evolution Meetings (2015); 20th International Congress of Arachnology (2016); Annual Meeting of the Geological Society of America (2016); 12th Colloquium of the African Arachnological Society (2017).

Grants American Arachnological Society: Arachnology Research Fund: USD 400 (1998), USD 800 (2000). AMNH Explore21: 2015: ‘Biodiversity Surveys in Poorly Explored Regions of Cuba: Developing New Research Collections and Collaborations’ (USD 85,000), 2016: Part 2 (USD 26,716), Co-PI with S. Perkins et al. Australian Biological Resources Study: Co-PI (with E.S. Volschenk and M. Harvey), on ‘Revision and Phylogeny of the Australian scorpion genus Urodacus Peters (Scorpiones)’: AUD 181,135 (2006–2009). Ben Gurion University of the Negev, Jacob Blaustein Center for Scientific Cooperation: Award from Foreign Visiting Lecturers’ Program: Salary and travel expenses for fieldwork and intensive course on ‘Systematics and Diversity of Scorpions, with Introduction to the Israeli Fauna’ at Ben-Gurion University of the Negev, Sede Boqer, in August 2013. Botswana International University of Science and Technology Research Initiation Grant: Collaborator (with PI Tharina Bird) on ‘Identification and characterization of novel molecules with potential biomedical applications from scorpion (Arachnida) and centipede (Chilopoda) venom’: BWP 200,000 (2018–2019). Higher Education Commission, Pakistan: PI on ‘Molecular systematics of medically important scorpions (Family Buthidae) from Pakistan’: stipend and subsistence for 12 months in New York for postdoctoral fellow, Muhammad Tahir; USD 10,000 bench fees for Prendini (2012–2013); Visiting Scholar Program: airfare and expenses to lecture and conduct workshop at University of Sargodha, Punjab, and visit collections at Pakistan Museum of Natural History, Islamabad (2012). Israel Taxonomy Initiative: Funds for fieldwork and intensive course on ‘Systematics and Diversity of Scorpions, with Introduction to the Israeli Fauna’ at Ben-Gurion University of the Negev, Sede Boqer, in August 2013 (USD 3,020). JRS Biodiversity Foundation: Co-PI (with Bridgit Davis) on ‘Improvement and Integration of Arachnid Biodiversity Information in South Africa’ [http://jrsbiodiversity.org/grant/national- museum-bloemfontein/] (USD 180,000), 2012–2014. Mohammad bin Zayed Species Conservation Fund: PI on grant 11252329 ‘Assessing the Conservation Status of Endangered Scorpions (Scorpionidae: Opistophthalmus) in the Western Cape Province, South Africa’ [http://www.speciesconservation.org/case-studies-projects/black- legged-burrowing-scorpion/2329]: USD 20,000 (2011–2012). National Geographic Society Expedition Grants: Co-PI (with Pio Colmenares) on WW-214R- 17 ‘Exploring Unknown Arachnid Diversity in the Caves of an Ancient Tepui’: USD 18,100 (2017– 2018). National Science Foundation, U.S.A.: Co-PI (with W.C. Wheeler, J.A. Coddington, G. Hormiga and P. Sierwald) on EAR 0228699 ‘Assembling the Tree of Life: Phylogeny of Spiders’: USD 2,738,000 (2002–2006); PI (with W.D. Sissom) on DEB 0413453 ‘REVSYS: Revisionary Systematics of the North American Scorpion Family Vaejovidae’ [http://www.vaejovidae.com/]: USD 414,578 (2004–2008); Major User (PI: A. Klaus) on DBI 0619559 ‘Acquisition of a Variable Pressure SEM at the American Museum of Natural History’: USD 386,563 (2006); Co-PI (with P.E. Cushing) on DEB 0640219 ‘Collaborative Research: Global Survey and Inventory of Solifugae’ [http://www.solpugid.com/]: USD 649,984 (2007–2011); PI (with Lauren Esposito) on

14 Curriculum Vitae: Lorenzo Prendini, Ph.D.

DEB 0910147 ‘Dissertation Research: Revision of the New World scorpion genus Centruroides Marx, 1890: Systematics and Biogeography’: USD 14,847 (2009–2010); PI (with Lionel Monod) on DEB 0910091 ‘Dissertation Research: The Liochelidae of the Indo-Pacific Region: Systematics and Biogeography’: USD 14,750 (2009–2010); PI (with Stephanie Loria) on DEB 1310855 ‘Dissertation Research: Patterns of Diversity and Endemism in South-East Asian scorpions: Systematics and Biogeography of Chaerilidae’: USD 20,766 (2013–2014); PI on ‘Living : Integrating Phylogenomics and Comparative Morphology to Assemble the Scorpion Tree of Life’: USD 808,499 (2017–2020). Richard Lounsbery Foundation: Co-PI (with M.E. Siddall and R. DeSalle) on ‘DNA barcoding of Leeches and Scorpions’: USD 325,000 (2004–2007). South African National Research Foundation (NRF), Foundational Biodiversity Information Programme (FBIP): ‘BioGaps: Filling Biodiversity Information Gaps to Support Development Decision Making in the Karoo’ [http://www.sanbi.org/biogaps] (ZAR 972,000), Collaborator with co-PIs Domitilla Raimondo and Krystal Tolley from the South African National Biodiversity Institute and ca. 30 participants including ca. 10 taxon specialists; AMNH budget: ZAR 89,000 for fieldwork in South Africa and identification of materials collected plus ZAR 140,000 to support South African student to digitize 8,000 scorpion records in AMNH collection. Stavros Niarchos Foundation: Constantine S. Niarchos Expedition grant: USD 7,000 (2003), USD 7,000 (2007), USD 7,500 (2008), USD 6,000 (2009), USD 5,490 (2011), USD 17,788 (2016). United States-Israel Binational Science Foundation: Co-PI (with E. Gefen) on BSF 2014046 ‘Diversity, Origins and Affinities of the Israeli Scorpion Fauna’: USD 135,000 (2015–2018). U.S. Academic Institutions: Theodore Roosevelt Memorial Fund, American Museum of Natural History: USD 1,500 (1997), USD 1,500 (1998); Ernst Mayr Grant, Museum of Comparative Zoology, Harvard University: USD 1,500 (1998); Collections Study Grant, American Museum of Natural History: USD 1,000 (1999); Visiting Researcher, Program for the Promotion of International Partnerships, University of Michigan: airfare from Cape Town, board, lodging and research expenses in Ann Arbor for 3 weeks (1999).

Scholarships and Bursaries Corporate Sector, South Africa: Standard Bank of South Africa: ZAR 7,000 (1992–1994); The Premier Group: 85 % of tuition fees p. a. (1992–1994). Private Organisations: Skye Foundation and Charitable Trust: ZAR 60,000 (2000). Foundation for Research Development, South Africa: B.Sc. Bursary: ZAR 5,000 p.a. (1992– 1994); B.Sc. (Hons) Bursary: ZAR 7,000 (1995); M.Sc. Bursary: ZAR 10,000 (1996); Prestigious Bursary, M.Sc.: ZAR 40,000 (1997); Prestigious Bursary, Ph.D.: ZAR 50,000 p.a. (1998, 1999). University of Cape Town: Myer Levinson (EMDIN) Scholarship: ZAR 6,000 p.a. (1996, 1997); S.A. College Croll Scholarship: ZAR 4,900 p.a. (1996, 1997).

Awards Anglo-American Corporation Open Scholarship Book Prize (1992); Certificates of Merit (Botany, Zoology), University of the Witwatersrand (1992, 1993, 1994); South African Breweries Bronze Medal (1992), Silver Medal (1993), Gold Medal (1994); Tree Society of Southern Africa Prize (1993); Peter Thompson Prize (1993, 1994); John Twinch Memorial Prize (1994); Zoological Society of Southern Africa Award (1994, 1995); Wayne Rendell Memorial Award (1995); Purcell Memorial Prize (1995, 2002); 5th International Colloquium of the Research Group for the Study of African Arachnids (1996); 6th International Conference of the African Arachnological Society (1999); University of Sargodha, Pakistan (2012).

Reviewer Books: Catalog of the Scorpions of the World (1758–1998) (Fet et al., 2000); Methods and Tools in Biosciences and Medicine: Techniques in Molecular Evolution and Systematics (DeSalle et al., 2001); Proceedings of the 19th European Colloquium of Arachnology (Scharff & Toft, 2001); The Biology of (Pinto-da-Rocha, Machado & Giribet, 2007); The Evolution of ‘Primary’ Sexual Characters in Animals (Leonard & Cordoba-Aguilar, 2008); Arachnids of the World (Beccaloni, 2008); Scorpions: On the Hunt (2008); Grzimek’s Animal Life Encyclopedia (2009);

15 Curriculum Vitae: Lorenzo Prendini, Ph.D.

Springer Handbook of Toxinology (Gopalakrishnakone, 2015); book proposal for the University of Arizona Press. Grants: Australian Biological Resources Study; British Ecological Society; Czech Science Foundation; Fondo para la Investigación Científica y Tecnológica, Argentina; Fonds de la Recherche Scientifique, Belgium; National Geographic Society (Expedition Grants); National Research Foundation, South Africa; National Science Foundation, U.S.A. (panelist); Netherlands Organization for Scientific Research (Innovational Research Incentives Scheme); U.S.-Israel Binational Science Foundation (panelist); U.S. Fish and Wildlife Service. Evaluator: Rating of South African scientists for National Research Foundation, South Africa (2008); referee for South African Council for Natural Scientific Professions (2012). Journals: Acta Amazonica; Acta Zoologica; African Entomology; African Invertebrates; Amazoniana; American Museum Novitates; Animal Biology; Arachnologische Mitteilungen; Arachnology; Arnoldia Zimbabwe; Arthropod Structure and Development; BioInvasions Records; Biologia; Biota Colombiana; Biota Neotropica; BMC Evolutionary Biology; Canadian Journal of Zoology; Caribbean Journal of Science; Cimbebasia; Cladistics; Comptes Rendus Biologie; ; Current Opinion in Insect Science; Entomological News; Environmental Entomology; Fauna of Arabia; Frontiers in Zoology; Genetica; Insect Systematics and Evolution; International Journal of Modern Biological Research; International Journal of Tropical Insect Science; Invertebrate Systematics; Israel Journal of Zoology; Journal of Arachnology; Journal of Arid Environments; Journal of Biogeography; Journal of Ethology; Journal of Genetics; Journal of Insect Conservation; Journal of Insect Science; Journal of Morphology; Journal of Natural History; Journal of Threatened Taxa; Living World: Journal of the Trinidad and Tobago Field Naturalists Club; Molecular Ecology; Molecular Phylogenetics and Evolution; Natural History Magazine; North-Western Journal of Zoology; Organisms, Diversity and Evolution; Pakistan Journal of Zoology; Palaeontology; Palaeontologia Electronica; Pan-Pacific Entomologist; PLoS One; Proceedings of the National Academy of Sciences, USA; Revista Ibérica Aracnología; Revista Mexicana de Biodiversidad; Rivista Italiana di Paleontologia e Stratigrafia; Revue Suisse de Zoologie; Scientifica; Scientific Reports, Nature; Senckenbergiana biol.; Studies on Neotropical Fauna and Environment; Systematic Biology; Systematic Entomology; Terrestrial Arthropod Reviews; Texas Memorial Museum, Speleological Monographs; Texas Tech University, Occasional Papers of the Museum; Transactions of the Royal Society of Edinburgh: Earth and Environmental Science; Transactions of the Royal Society of South Africa; Turkish Journal of Zoology; ZooKeys; Zoologica Scripta; Zoological Journal of the Linnean Society; Zoologischer Anzeiger; Zoology; Zoosystema; ZOOTAXA.

Teaching and Mentoring Postgraduate Teaching: New Mexico State University, Las Cruces: Arthropod Diversity and Evolution (2002); AMNH, Richard Gilder Graduate School: Core Course in Systematic Biology (2008–2010): lectures covering Classification, Taxonomy, Nomenclature, PhyloCode, DNA Barcoding, Species Concepts, Reticulation, Phylogeography and Coalescence; Entomology Course, guest lecture (2012): ‘Insect Relatives with Introduction to Chelicerata’. Ben-Gurion University of the Negev, Sede Boqer, Israel: Israel Taxonomy Initiative: Systematics and Diversity of Scorpions, with Introduction to the Israeli Fauna (2013), seven lectures (‘The Fascinating World of Scorpions’; ‘Collecting, Preserving and Documenting Scorpions for Systematics Research’; two lectures on ‘Phylogeny and Classification of Scorpions’; ‘Identification of Biological Taxa’; two lectures on ‘Israeli Scorpion Fauna’; two afternoon practicals on how to identify scorpions and construct dichotomous identification keys, two nights of fieldwork (rocky and sandy desert habitats); and a final examination. Postdoctoral Fellows: Erich S. Volschenk (2003–2006); Camilo I. Mattoni (2004–2006); Jose A. Ochoa Camara (2007–2009); Carsten Kamenz (2009–2011); H. Muhammad Tahir (2013–2014). Patricia Carrera, Cordoba University, Argentina (four months, 2009); Ricardo Botero-Trujillo (2018–). Major Advisor: Ph.D.: Lauren Esposito, City University of New York (2004–2011); Edmundo González Santillan, City University of New York (2005–2012); Lionel Monod, City University of New York (2005–2011); Stephanie Loria, Richard Gilder Graduate School, AMNH (2011–2015). M.S.: Jayson Slovak, City University of New York (2016–).

16 Curriculum Vitae: Lorenzo Prendini, Ph.D.

External Examiner: Ph.D.: Erich S. Volschenk, Curtin University of Technology, Perth, Australia (2002); Rabia Yaqoob, University of Sargodha, Pakistan (2015); Imtiaz Alam, University of the Punjab, Lahore, Pakistan (2017). Committees: Ph.D.: Joyce Fassbender, City University of New York (2004–2008); Tharina Bird, Colorado State University (2008–2014); Ed Stanley, Richard Gilder Graduate School (RGGS), AMNH (2010–2011); Ansel Payne, RGGS (2011–2012); Bernardo Santos, RGGS (2013–2016); Jeremy Frank, RGGS (2015–). M.S.: Sonia Maartens, University of Pretoria, South Africa (2007– 2008); Samuel Mwangi, Africa Nazarene University, Nairobi, Kenya (2007–2008), West Texas A&M University (2015–2016); Jessica Azzinari, West Texas A&M University (2015–2016). Visiting Students: Undergraduate: Samuel Mwangi, National Museums of Kenya, Nairobi (8 weeks, 2005). M.S.: Lionel Monod, University of Geneva, Switzerland (4 weeks, 2002); Christina Bisulca, University of Delaware (6 months, 2003–2004); Alfonso Ballesteros Chavez, UNAM, Mexico (8 weeks, 2008); Stephan Schaffrath, University of Cologne, Germany (3 months, 2013– 2014); Samuel Mwangi, West Texas A&M University (4 weeks, 2016). Ph.D.: Valerio Vignoli, University of Siena, Italy (8 weeks, 2004; 8 weeks, 2006); Christian Wirkner, Humboldt University, Berlin, Germany (12 weeks, 2004); Andres Ojanguren Affilastro, Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’ (MACN), Buenos Aires (6 weeks, 2005; 6 weeks, 2007); Alexander V. Gromov, Institute of Zoology, Almaty, Kazakhstan (3 months, 2007); Carsten Kamenz, Humboldt University, (12 weeks, 2005–2006; 8 weeks, 2008); Humberto Yamaguti, Universidade São Paulo, (5 months, 2009; 1 month, 2010); Carlos Santibañez-López, UNAM, Mexico (2 weeks, 2009; 4 months, 2011, 1 month, 2012; 1 month, 2014); Tharina Bird, Colorado State University, Fort Collins (3 months, 2009; 4 months, 2010–2011; 6 months, 2012); Rene Barba, Instituto de Ecología y Sistemática, Havana, Cuba (3 weeks, 2012); Roberta Engel, University of Connecticut, Storrs (several days, 2012); David Vrech, Universidad Nacional de Córdoba, Argentina (1 month, 2012); Rodrigo Monjaraz-Ruedas, UNAM, Mexico (1 month, 2014, 1 month, 2016, 2 months, 2017); Daniela Ramírez, UNAM, Mexico (2 weeks, 2014); Gustavo de Miranda, Copenhagen Museum, Denmark (1 week, 2014; 3 months, 2015); Ingrid Catalina-Ortíz, Universidad Nacional de , Bogotá (10 days, 2015); Bastian Klussmann-Fricke, Rostock University, Germany (2 months, 2015); Gerardo Contreras-Felix, UNAM, Mexico (1 month, 2015, 3 months, 2016, 2 months, 2017); Rebecca Godwin, Auburn University, AL (1 week, 2016); Diego Barrales Alcalá, UNAM, Mexico (2 months, 2016); Ricardo Botero-Trujillo, MACN, Buenos Aires (2.5 months, 2016); Francisco J. Salgueiro Sepulveda, Facultad de Ciencias, UNAM, Mexico (2 weeks, 2016); Dulce F. Piedra, UNAM, Mexico (2 weeks, 2016); Fabio Akashi Hernandes, São Paulo State University (UNESP) (2008, 2016); Jesus A. Cruz López, UNAM, Mexico (1 week, 2017); Shlomo Cain, University of Haifa, Oranim, Israel (2.5 months, 2018). Undergraduate Interns: NSF-funded summer Research Experiences for Undergraduates (REU) program, AMNH: Lauren Esposito, University of Texas, El Paso (2002); Samara Maaliki, City University of New York (2003) [joint UMEB (Undergraduate Mentoring in Evolutionary Biology) program]; Michelle McCoy, University of North Carolina (2004); Kanvaly Ben Bamba, Yale University (2004); Steve Webb, Muhlenberg College (2005); Sarah Schoenbrun, Brown University (2007); Margaret Rubin, Oberlin College (2016). Other: Gena Esposito, University of Texas, El Paso (2007); Angela Holuba, Barnard College, New York (2008); Sylvia Johnson, Barnard College, New York (2008); Michelle Yun, York College, New York (2010); Diogo Castellato, Federal University of Rio de Janeiro and Baruch College, Brazil Scientific Mobility Program of the Institute for International Education (2015–2016); Deborah Chin, Fairleigh Dickinson University (2016, 2017). High-school Interns: NSF-funded After-School Centers in Exploration and New Discovery (ASCEND) program, AMNH: Adrian Tung (1998); Brian Webster (1999); Bernard Laszczower (2003–2004); Boitumelo McCallum (2003–2004). AMNH High School Science Research Program in Biodiversity: Connie Cai (2004–2005); Melanie Ng (2004–2005); JianHua Lin (2006–2007); Qiao Rong Huang (2006–2007); Eleanor Goetz and Sasha Reiter (2013–2014). AMNH Lang Science Program: Sofia Castro and Geeta Sharma (2017). Other: Michelle Bayefsky-Anand (2008); Adam Getzler (2011); Massimiliano Roppo (2013); Billy Conlan (2015); Chelsea Silva (2015, 2016); Tamar Cohen (2017).

17 Curriculum Vitae: Lorenzo Prendini, Ph.D.

Molecular Systematics Laboratory University of Cape Town (1996); American Museum of Natural History (1997–2000, 2002–); University of Michigan, Ann Arbor (1999). NGS (Anchored Hybrid Enrichment), collaboration with Florida State University (2016–).

Fieldwork Angola (2015), Argentina (2003), Botswana (2011), Chile (2003), Djibouti (2017), Ethiopia (2018), Israel (2011, 2013), Jordan (2013), Kazakhstan (2003), (2012, twice), Malawi (2007, 2018), Malaysia (2013), Mexico (2002, 2005), Mozambique (2007), Namibia (1996/1997, 1999, 2003/2004, 2006–2010 annually, 2015), Oman (2017), Pakistan (2012), Puerto Rico (2009), South Africa (1989–2000 annually, 2003–2017 annually), Swaziland (2001), (2015), Trinidad and Tobago (1999), Tunisia (2017), U.S.A. (AZ, CA, NM, NV, TX) (1997, 2002, 2005, 2007, 2012), U.S. Virgin Islands (2009), Uzbekistan (2003), Venezuela (2009), (2012, twice), Zimbabwe (2013). Twenty-nine years (1989–) in southern Africa. Field trips (e.g. the Namib, Kalahari, Kaokoveld, Richtersveld and Sperrgebiet) required proficiency in off-road driving with a 4x4, and complete self-sufficiency (all necessary fuel and water had to be brought, in addition to standard supplies). Two 5-day hikes to survey scorpion fauna of Brandberg Massif, which involved carrying all supplies and water in a 40 kg backpack (2004, 2006). Hiked up El Tucuche, Trinidad (1999) and Auyan Tepui, Venezuela (2009). 4-day kayak trip through Xe Bang Fai River Cave, Laos (2012). Six-day hike to summit of Mount Mulanje, Malawi (2018). A collection of ca. 8,000 scorpion specimens, accumulated during six years’ collecting in southern Africa, was disseminated to various southern African, North American, and European museums; collections enlarged annually at the AMNH since 2002.

Collections Since 1995, the following collections visited, besides the AMNH: Albany Museum, Grahamstown; California Academy of Sciences, San Francisco; Field Museum of Natural History, Chicago; Forschungsinstitut und Natur-Museum Senckenberg, Frankfurt; Institute of Fundamental Studies, Kandy; Manuel A. González-Sponga Collection, Caracas; McGregor Museum, Kimberley; Musée Royal de l’Afrique Centrale, Tervuren; Museum National d’Histoire Naturelle, Paris; Museum of Comparative Zoology, Harvard University, Cambridge, MA; Natal Museum, Pietermaritzburg; National Collection of and Arachnids, Pretoria; National Museum of Namibia, Windhoek; National Museum of Natural History, Smithsonian Institution, Washington, DC; National Museum, Bloemfontein; Natural History Museum of Zimbabwe, Bulawayo; South African Museum, Cape Town; Natural History Museum, London; Oxford University Museum of Natural History, U.K.; Pakistan Museum of Natural History, Islamabad; Transvaal Museum, Pretoria; Zoologisch Museum, Universiteit van Amsterdam; University of Puerto Rico, Rio Piedras; Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn; Zoologisches Institut und Zoologisches Museum, Universität Hamburg; Zoologisches Museum, Universität Humboldt, Berlin. Material loaned from a further 16 collections. Curated, sorted, identified and partially digitized African scorpion collections (1995–2001, 2004, 2009, 2010, 2012–2015): McGregor Museum, Kimberley; Natal Museum, Pietermaritzburg; National Collection of Arachnida, Plant Protection Research Institute, Pretoria; National Museum, Bloemfontein; National Museum of Namibia, Windhoek; South African Museum, Cape Town; Transvaal Museum, Pretoria. Arranged purchase of Alexis Harington Scorpion Collection (ca. 6,500 specimens) by the AMNH (2002) and currently oversee curation of arachnid and myriapod collections in the AMNH (2002–). Started and continue to enlarge the world’s only synoptic collection of scorpion tissues at the Ambrose Monell Collection for Molecular and Microbial Research of the AMNH (1999–): ca. 20,000 tissue samples and associated vouchers, acquired by personal collections and donations from colleagues.

Exhibitions At AMNH, contributed images and reviewed copy on UV fluorescence, SEM and CT to Picturing Science: Museum Scientists and Imaging Technologies (2011); provided images and reviewed copy for Creatures of Light (2011) and Spiders Alive! (2012); co-curated Spiders Alive! (2015).

Interests

18 Curriculum Vitae: Lorenzo Prendini, Ph.D.

Lorenzo Prendini’s research program emphasises systematics, collections-based research and fieldwork. His primary research interests lie in the empirical study of phylogeny, macroevolution, speciation, biogeography, and comparative evolutionary biology, using scorpions and minor arachnid orders (specifically Amblypygi, Solifugae, Schizomida and Thelyphonida) as model organisms. Major research interests are: higher phylogeny and classification of scorpions; revisionary systematics of scorpions; phylogeny and classification of minor arachnid orders; systematic theory and practice; adaptation and biogeography of arachnids; distribution and conservation; evolution of insect-plant associations.

Biography Dr Lorenzo Prendini curates the collections of Arachnida and Myriapoda at the AMNH, where he has worked for fifteen years. His research addresses the systematics, biogeography, and evolution of scorpions and lesser known arachnid orders (whip spiders, camel spiders and whip scorpions) using a combination of morphological, molecular, and distributional data, and diverse analytical tools. Current research on scorpions is focused in three areas: reconstructing the phylogeny and revising the contentious suprageneric classification of scorpions; systematic revisions of Afrotropical, New World, and Australasian scorpion families; testing adaptational and biogeographical hypotheses in sub-Saharan Africa. The search for interesting arachnids has taken Prendini and his research group to more than 30 countries on all continents except Antarctica. Besides arachnids, Prendini is interested in the evolution of insect-plant associations and in theoretical issues concerning the phylogenetic analysis of morphological data.

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