Vol. 7, No.1 - 2, March - June, 1993 27

A Review of tt Ie RI lopalopl 101 i1li (eoleoplel a. eelal Ilbycidae) af Nartll alld eelIb al AI Ilei ica

Edmund F. Glesbert 9780 Dlake Lalle Beverly Hills, CA 90210

and

John A. Chemsal< Entomology & Parasitology University of California 201 Wellman Hall Berkeley, CA 94720

Abstract The tribe Rhoplllophorini is reviewed tor North andC-entral America Keys to genera and species are provided, and known species are redescribed. The genus Rhopalophora Audinet-Serville is revised to include 16 species, WIth the followmg additIons: R. sernpenmsn. sp. and R. yucatana n. sp. are descnbed from MeXICO; R. rUgLcolhs hOl1Orei n ssp is described from Baja California; and R meeskei Casey, 1891 is restored to species status New synonymIes are: R. mcrustata Chevrolat, 1859 R. tenuis (Chevrolat, 1855); R. laevicollis LeConte, 1873 R. cltpricollis Gnerin-MlmeviJle, 1844 Distribution maps are includedfor the more common species. The subgenus Ischwnodonta Chevrolat IS raIsed to genenc status, and includes 3 species: 1. serripes (Bates, 1872) n. comb.; I. versicolor (Chevrolat, 1859) n. comb.; and 1. mexicana n. sp. from Mexico. Cosmisoma Audinet-Servilleis revised to mclude 6 species, with 2 newly described: C. militaris n. sp. from Panama and Costa Rica, and C. ,haptos n. sp from Panama and O>lombia. New synonymies are: C. nudicornis Bates, 1892 C. martyr Thomson, 1860, and C. alboscutellata linsley, 1935 =C. reticulata Bates, 1885. Cycnoderus Audinet-Serville is newly recorded from Mexico and centralAmerica with 6 species: C. virginiae n. sp.. C. copei n. sp., C. liuidus n. sp., C. breuicolle n. sp. from Mexico, and C. guatemalicus n. sp. from Guatemala. Cycnoderus barbatus Gounelle is recorded from O>sta Rica and Panama. Anew genus, Muxbalia, is proposed and tentatively placedin the Rhopalophotini, and M. monzoni n. sp. is describedfrom Guatemala. Ozodes Audinet-Serville, andLissozodes Bates are removedfrom the RhopalophOlini and placed pruvisiOllally in the NeC}'dalopsilli. Rhopalophola cup, icoUis, R. longipes, R. serripennis, Cycnoderus virginiae, Cosmisoma rhaptos, and Muxbalia monzoni are illustrated.

Introduction fauna nOl th of South America has been largely ig- nored. Since theefforts ofBates(1879-1885), new taxa The tribe Rhop_alo~horiniis found ~rimarilv in have been proposed only by Linsley (1935, 1942) theWestern Hemisphe~(atpresentcon;stingoft 19 species in 13 genera) with only 3 genera occurring Adult specimens ofRhopalophorini are often en- elsewhere: Ampherlwe Newman from Australia, countered in large numbers, uswlIly on the flowers of Pelossus Thomson from Mrica, and Dymorpho various plants, 01 on dead twigs and blanches. Mem- bers ofthe tribe appear to be diurnal. cosmisoma Picfrom China ~Aurivillius1912; Gressitt This revision deals wit.h members of t.he t.ribe 1951). The group is predrnrinIDltly NeotIOpEal with found from North America through Panama, these :::~~~~~er:~g~~~~rt't;~~t~~~i:;~~:~~:t::~ limits being strictlya matterOf convenience. CladiStic These species were treated by Linsley (1964), but the analysis has not been considered due to these geo- graphicallimitations. PortionsoftheSouthAmerican 28 Insecta Mundi fauna havebeen studiedLy Zajciw (1962), MOlIne and diffCI fwm othcIs in that bibe by the fOlln of the Magno (1988), and by Monne (1989) pronotum, different style ofprosternaI sexual modifi- For the geographIcal area under conSIderatIon, catIOns, the exsertedpro- and mesocoxae, andfrom all the tribe Rhopalophoriniconsists ofsix genera and 33 butllluxhalian. gen. bythe graduallycla'latefemora. species. In the preparation of this work, over 3000 Martins (1975), in attempting to clarify the status of specimens 'tvere examined. Data on the material several genera in the Rhopalophorini and examined for the less common species is listed indi- Tillomogphini, pointed out the close relationship of vidually, disbibutiOllal maps have been providedfor Ozodes, lvTeozodes Zajciw, Parozodes Autivillius, species collected in a large numbers of localities Argyrodines Bates, Aphysotes Bates, and Epropetes DepOSItIon oftypes and speCImens ill illstItutIons and Bates, but did not choose to reassign these genera private collections are indicated by the following tribally at that time. Of these genera,Orodes and abbreviations: [AMNH] = American MuseumofNatu­ Lissozodes (of which an as yet undescribed species talHistmy, IBMNHl TheNatutalHistmy Museum, OCCutS in Costa Rica) OCCut in the the area of this London; [CAS] - California Academy of Sciences; study, and appeartobe closely related to Euclwra<;sw; [EG] - Edmund GIesbert; [FH] - Frank Hovore; Audinet-Serville whIch IS presently aSSIgned to the [FMNH] - Field Museum of Natural History, Chi Necydalopsini.Untilthetribalstatusofthisgroupcan cago; [GNI - Gale NelSon; [llliJ - Henry Howden; be definitely resolved, OZodes and Lissozodes are [lIS] Henry Stockwell; [JAG] John A. Chemsak; placed into the Nee)' dalopsini. [JB] = Joseph Beier!; [JC] = James Cope; [JH] = Jeff The six genera of Rhopalophorini occurring in 'Huether, [JM] JoseMonwn; [tJ""W) JamesWappes; North and Central America may be separated as [RB] - Keve Ribardo; (I ACM] - Los Angeles County follows' Museum of Natural HIstory; [MNS] - NatIonal Museum of NaturalSciences, Ottawa; [RP] Richard Key to the Genera of Rhopalophorini Penrose; [RT] =RobertTurnbow; [RW] =RickWestcott; in North and Central America [SU] Stanford University; crAM] Texas A&M University; [VA] = University of Arizona; [VCB] = 1. Meso and metafemora abruptly clavate beyond Umversity of CalifornIa, Berkeley; [UCD] - UnIVer­ middle (See fig. 1) 2. sity of California, Davis; [UDV] Universidad del Meso- and metatemora graduallY clavate (See fig Valle de Guatemala; [UK] = University of Kansas; 12) Muxbalia [UNAM] UnivetsidadNacionalAut6nomadeMexico, [VU] =University of Utah. 2. Elytral apices truncate or sinuate-truncate. An- tenna I fourth segment. much shorter t.han third Tribe RHOPALOPHORINI Lacordaire (See fig 1) 3 Lacordaire, 1869:109 (as Rhopalophorides); LeConte, Elytral apices acuminate or rounded. Antennal 1873.307; LeConte and Hom, 1883.292; Leng, 1884.114; Knull, 1946:187; Gressitt, 1951:315; Linsley, 1964:187. 3.(2) Elytra bearing distinct transverse pubescent fas- DESCRIPTION: Formparallel-sided. Headwithfrons ciae at base and middle Rho;alor;horella large, Oblique, concave; eyes finely faCetted, deeply Elytra lacking transverse pubescent [~ci& ...... emarginate;mandibles acute; antennaeslender, elon ...... Rlwpaloplwra gate, withouta poriferoussystem,segmentsunarmed; palpi short, equal, last segment oval, not dilated. 4.(2) iilltennal fifth segment bearing a tuft of hail' Wee Anterior coxal cavities small, not angulate, open setae, other segments sometimes tufted. Hind behind Legslong, slender,femorausuallypedllnclIlate legs laeking tufts of setae Cosmisema and abruptly clavate; tibial spurs small. Stridulatory Antennalfifth segmentusuallylackingtuftsofhair­ like setae [exeept Cycooderus harbatus, which surface of mesonotum large, undIVIded; scutellum has hind tibiae and tarsi also tufted] 5. small, obtuse. Elytral disk flattened, especially near base. 5.(4) Antennae ofmales 11 segmented. Elytradull,punc­ The Rhopalophorini may be recognized by the tation fine, dense, uniform. Pronotum at most slender, somewhatflattenedbody form, slender, elon­ slightly longer than wide. Femora at least feebly gate antennae, and usually pedUnculate femora. asperate at base lscf'uonodonta Ozodes Audinet-Serville and Lissozodes Bates have Antennae of males 12 segmented. Elytra shining, been previously included in the Rhopalophorini but punctatIonmoderatelycoarse,Irregular. Pronotum Vol. 7, No.1· 2, March· June, 1993 29

at least 1 li3 times as long as wide. Femora transversely, somewhat narrowed behind apex, smooth, not asperate at base Cycrwderus pubescence, if evident, moderately dense, trans- verselyappressed, usuallyinterruptedat middle; Genlls RhopaLopbora Alldinet-Serville punctation absent or mostly obscured. Length 8- 15.5mm 2 Elytra subdepressed or not; margins often at least 18~: Rhopatoplwffi l'.udinet Serville, 100; Chevrolat, somewhat asperate; punctures usually coarse, ;ot ~hsc'lr:d (See fi~ 3) Pron~tum lac;ng :­ mc sculp unng; pu escence, i presen ,usu Gounelle, 1911:99; Knull, 1946:227; Linsley, 1964:188. y for;i~ n~w vittae; punctation usually evi- Rb6polilpborlls, I.e! :Onte, 18'1:3'311/; I.e! :onte and Horn, den. eng -J:lmm ~6 1883:292. D:n.0PWl I.e! :Onte, 1850'19 (Type species' Stenocorus 2. (1) Integument with strong metallic greenish reflec­ longipes Say, monobasic). tions a enda es metallic bluish. Pronotal u- a ora a ora' uriv' 'us (Hidalgo, Guerretu) R. eximia Bates DESCRIPTION: Formslender, somewhatdepressed; Integument without distinct metallic reflections 3 integument usually blackish, often with pronotum red. Head moderately prolonged; antennae filiform, 3. (2) Pronotal pubescence indistinct or obscure. lntegu­ at least as long as body in male, shorter in female, mentbrightly, distinctly I eddish 01 angeandblack. punctulate and sericeus, nearly always eleven seg­ Pubescence of elytra and underside obscure. mented, basal segments slightly swollen apically, Length 10 llmm. Guatemala ...... R. nigriventris Bates Pronotal pubescence distinct, whitish to bright or ronotum unarme ,usual yonger an wi e. ute­ ange. Elytra and underside densely golden or Hum small, triangular; apex obtuse, notched. Elytra whitish pubescent 4. somewhat depressed near base and suture, with apices tr uncate, smface punctate. Abdomen nOImally 4 (3) Pronotal dIsk pICeous, WIth a paw of broad, golden segmented. Legs with mesoand metafemoraabruptly pubescent vittae; prosternum broadly orange at clavatenear apIces, fIrst segmentofmetatarsIatleast apex. Length 8-13mm. MeXICO (Chlapas) to COsta twice as long as second. Rica R wheel/In Bates Pronotum dark red to reddish-orange, often WIth a TYPE SPECIES. RhopalophOl a sanguinicoUis pic?~u: :~~l~ at middle ~f disk, clot~~d w.ith WhItiS 0 ng t orange pu escence no ormmg Audinet-Serville (fhomson designation, 1864). distinctvittae, middleirregularlyglabrous, sparse punctules often visible 5. A complex ofclosely-related and variable pheno- typesin this genus, mcIudirigR.l.Ong£pes, R. tenUls, R. 5. (4) Pr-onotum and elytra uniformly dark reddish, ap­ bicoloreUa, R. rugicoUis, R. serripennis, R. pendagesdal ker. PlOnotal pubescence fine, whit- punctatipennis, and R. miniaWcollis, has been diffi­ ish, broadly interrupted at middle, with a wide cult to separate in the past In several cases, pheno- glabrous latel al "itta abcwe a nailow pubescent types which are sympatric and clearly separable at vittaon each side; sculpturing reduced. Length 8­ one IOCatiOn intergrade at another. Treating these 1O.5mm. Mexico (Yueat{m, Quintalia Roo) ...... R. yucatana n. sp. forms as separate species is, we believe, preferable to Pronotum not coneolorous with elytIa, often with II wholesale synonymy, but examples from a few popu­ dark spot at middle ofdisk; pubescence variable, lations will not fit neatly illtu the parameters given in 'llhitish to bright orange, narrowlyinterruptedat the following key. middle, without lateral vittae; surface strongly sculPtured. Length 10 15.5mm. Southern Texas Key to the Species of Rhopalophora to C<>sta Rica .... R. cupricollis GUl3rin-Meneville in North and Central America 6 (1) Head red, or mostly red 7 Head blackIsh, somehmes WIth OCCIput partly red 1. Elytrabroadly, shallowly, mediallydepressed; punc- 9 tUles small, oftenpartiall) obseuted, malgilis not asperate (See fig. 1). Pronotum with disk usually 7 (6) Pmnotum nearly twice as long as wide, nearly dOl sally seulptuted, with a btoadobtuse swelling parallel-SIded, only slightly WIder at mIddle; disk near base on each side ofa broad median impres- nearly impunctate, with a pair ofwhitish pubes- sion; sides swollen somewhat behind middle, cent vittae. Length 6-8.5mm. Lower Rio Grande 30 Insecta Mundi

Valley,Texas toMexico (Nuevo Leon, Talllaulipas) 14.(13) PlOnotulll about as wide as long, sides moderately ...... R. angustata Schaeffer widely rounded, obtusely tuberculateoneach side Pronotum slightly less than 1 1/2 times as long as of disk near base, base constricted; pubescent wide, sidesslightlyrounded, widestbehindmiddle vittae orange or golden. Length 6-lOmm. Utah, ...... 8. Colorado, Arizona and Texas to '''lexico (wide spread) R. meeskei Casey 8. (1) Pronotal disk moderately coarsely, densely punc Pronotum distinctly longer than wide 15. tate, pubescentvittae (sometimesindistinct)com posedofpaIegolden pubescence. Length6-8.6mm. 15.(14) Pronotum feebly subangulate at Sides, WIdest Mexico (Sinaloa) slightly behind middle; punctures indistinct ...... R. mimawcolhs Chevrolat (m part) Length 5-9mm. Eastern and rmdwestern North Pronotal disk nearly impunctate, lacking vittae. America to central Texas (see fig. 3) Length5-8mm. SouthernArizona, Mexico (Sonora) ...... R. longipes (Say) ...... R. prorubra Knull Pronotum subcylindrical or slightly tapering, punc- tures distinct 16. 9.(8) Pronotal disk not plicate nor rugulose 10. Pronotal disk transversely rugose. Length 7-lOmm. 16.(15) Metafemorashort, at most barely attainingelytral Texas, New Mexico, Arizona and Baja Califomia apices in female, barely surpassing apices in south to Guatemala ...... R. rugicollis (leConte) male. Antennae with terminal segment shorter thantenth, apexstrongly curved, blunt. Pronotum 10.(9) Elytral punctation subseriate; epipleural margins subcylindrical, pubescence evenly distributed on strongly asper ate. Plonotum I ed, subeylindrieal, each side of middle, usually not forming distinct pubescentvittae absentor feeble. Length 5-9mm. vittae. Length 4.5-9mm. Texas (Rio Grande Val- Southwestern Mexico (see fig. 8) . ley)tosouther n ArizonaandMexiw (Talllaulipas, ...... R. serripennis n. sp. Coahuila, Sonora, Baja California) . EIYtra with punctation irregular, not subseriate; ...... R. bioolorella Knull epipleuralmarginsfeebly to moderately asperate. Metafemora surpassin~ elytral apices in female, Pronotum red or black 11. surpassing apices by -lOre than halfthe length of club in male. Antennae with terminal segment at II.(IO) Pronotum black 12. least subequal to tenth, apex subacute, feebly Pronotum red (sometimes piceous in R bicolorella) appendiculate (sometimes forming short t\velfth ...... 13. segment at appendiculation). Pronotum slightly tapering, sides slightly rounded, pubescence usu 12.(11) Pronotum WIth SIdes feebly rounded, tapenng to ally forming distinct vittae. Elytra piceous, shin­ apex; disk with a wide golden vilta on each side ing, pubescence moderately indistinct. Length 6 ofa narrow, glabrous, sparsely punctate median lOmm. MeXICO to GuatemaIa . vitta; prosternum of males without erect setae. R punctatlpennrs Linsley Length 8-11mm. Mexico to EI Salvador ...... R. lineicollis Chevrolat Pronotum subcylindrical, slightly wider behind middle, disk moder Htely densely punctate, vittae Rhopalophora angustata Schaeffer varyingfrom moderatelywide toindistinct, white to pale golden, prosteruum of males with short, RhopalophonJ angustata Schaeffer, 1905'162 (TYPE LO pale, erect setae present. Length 6-9mm. Mexico CALI'I'Y: Esperanza Ranch, BrownsVille, Texas); to Costa Rica R. tennis (Che'lolat) Linsley andMartin,1933'181 (habits); Vogt, 1949'175 (habits); Linsley, 1964:188; Hovore and Giesbert, 13.(11)Male pl'ostemalpunctatepatchessmaH, indistinct, 1976'357 (babJts); Manley and hench, 1976'49; elongate. Pronotal punctures deep, moderately Turnbow and Wappes, 1978:369 (habits); Hovore, dense on each side of median impunctate area, Penrose, and Neck, 1987'308 (habitS) usually separated by about one diameter; pubes- Rhopalophora (Rhopalophora) angustata; Aurivillius, cent vlttae broad, usually dlstmct. Elytra 1912:431. castaneous to piceous; pubescence fine, pale Length 6-IOmm. MeXICO . DESCRIPTION: MALE: Formmoderatelysmall, elon­ R miniatocollis Chevrolat (in part) gate, slender; integument piceous, head and protho rax red, elytra rufopiceous. Head finely punctate above, mOle coarsely so on {IOns; antennae extending 3 segments at most beyond elytral apices, eleventh segment filiform, nearly as long as tenth. Pronotum piceous 14. Vol. 7, No.1- 2, March - June, 1993 31

distinctly narrowet than elytral humeri, about 1 1/2 width of the base and apex subequal, and by the times as long as broad, subcylindrical. feebly wider at entirely red head. Adults have been taken from flow- middle; disk with a shallow, median, longitudinal ersofB

appearance of the pronotal pubescence, and by the shorter Ie s and antennae. Rho alo hora bicolorella

(Figs. 1 & 2)

Rhopalophora cupricollis Guerln-M€meville, 1844:235

White, 1855:205; Bates, 1872:188; Bates, 1874:224; a es, ; a es, Rho a10 hora Rho a10 h 1912:431.

AbTY. "Texas"). NE'vV SYN f'

\ y

~ ~ -jJ I":u. T? \ ~ r) Ie .; ~ 1 I

Figure 2. Known geographic distribution ofRhopalophora cupricollis Guerin-Meneville. elytral apices by about length of club. Length 10­ southern part ofthe range. In the north, from Texas 155mm to northern Mexico, this pubescence IS whitiSh, wIth a dark pronotal macula usually present and some­ }<'EMALE: Form similar to maIe.•A.ntennae exceeding times quite Luge. These charaeters appear to inter elytmlaprees by about 2 segments. Presternum lack grade elinally and indWiduals with pale golden or ing glabrous punctate patches. Metafemora exceed­ light orange pubescence occur in central Mexico. ing elytral apices by less than length of club Length Elytral pubescence appears to be denser and more lO-15mm. coppery in the southern populations and in Costa Rica, and elytraI punctation is also somewhat fmer RANGE: Lower Rio Grande Valley, Texas to Costa andpartiallyobscuredby thepubescence. Theform of Rica. (See fIg. 2) the pronotum is also somewhat variable, with indi­ vidualsfmm centralMexicooften showinga tendency REMARKS: Distinguished by the larger size, non­ toward more pronounced sculpturing of the discal metalliC integument, and dense, silky, appressedpu- surface. bescence of the pronotum, this widespread species Host plants are recorded as Pithecellobium, appears to be quite variable. Typreal R. cupricoUis, Pr6sopis, (]()ndalia, Citrus, Di6spyros, and with the pubescence oftheheadandpronotum dense, ZanthoxYlum. Specimens have been collected on a bright golden or coppery, occur over most of the variety offlowers. 34 Insecta Mundi

R17opalop17018 eximia Bates R17opaloph0l8 Iineico/lis CtJevlolat (Fig.9) Rhopalophora eximia Bates, 1892:166 p1.7, fig. 1 (TYPE .. . ~ va CALITY: "Mexico"); Bates, 1880:62; Bates, 1885:309. Rhvpaloplwra (Rlwpalophor a) lineicollis, Aurivillius, 1912:431. DESCRIPTION'MAl ;p;. Form moderate-sized, elon- gate, slender; integument black with metallic green reflections, antennae and legs '....ith metallic blue er, e ongae;mtegumen pIceous ac ; ea mer­ reflections. Head with frons deeply transversely im- ately sparsely punctate, densely clothed with recum plessed, modetately coaIsely punctate between an- bent golden pubescence; antennae long, exceeding ~:~:~~~l;b::~~::Il~:~;;t:~I~::n~;a~~:;g~II~:;: ehttal apices by about 4 segments, scape ments, scapefinelY punctate, thirdandfifth segments each nearly twice as long as fourth. Pronotum longer than wide, sides feebly rounded, tapering slightly to than wide, sides somewhat swollen behind middle, apex, WIdest at base, with subbasalimpression feeble; basewiderthan apex; disk with anobtuse tubercle on disk 'Nith an inconspicuous tubercle on each side near each side of shallow, median, longitudinal, subbasal base, surfacesparselypunctateandglabrous atmiddle; impression, surface sparsely punctate, transversely, disk with a wide, dense, golden pubescent villa on . shallowly rugose, clothed with moderately dense, each side from base to apex, bordered lateraJly by a tranS"v'ersely appressed, yellowish pubescence inter glabrous vitfu on each SIde; prostemum WIth basal 314 rupted at middle by longitudinal glabrous vitta; densely clothed with whitish appressed pubescence, erect setae lacking, with an indistinct, triangular, glablOus, punctatepatchon each side infrontofcoxae; metasternum m erate ydense y Cothe WIth fme, meso- and metasternum clothed with whitish pubes- short, golden pubescence, episterna more densely cence. Elytra 3 tImes as long as WIdth across humen, pubescent. Elytra slightly more than 3 times as long sides slightly tapering, outer margins feebly serrate; as width actOss humeli, sides subpatallel, slightly apices truncate with hind angles dentate; diskfeebly, . . . narrower at mid e ic run u ur , en a e; is su epresse, sur ace ense y bordered by a vague carina on basal half; surface :icropl~nctat~, with larger pu~tur~s moderately moderately coarsely, densely punctate, pubescence ense, rrregu ar; pubescence ob que y transverse, short, fine, appressed, pale golden, partially obscur- appressed, yellowish, moderatery dense toward su­ mgsurlace. Abdomen densely clothedWIth appressed ture moderatel s arse toward mar . s. Abdomen . . e e y ense y c 0 e wi I e, s or ,go en elytral apices by at least length of club. Length 8­ pubescence which is more dense laterally. Legs long, llmm slender; metafemora surpassing elytra.J. apices by entire length ofelub. Length 11 14.5mm. FEM ALE; Form similar to male. ,A.ntennae exceeding elytral apices by about 2 1/2 segments. Presternum FEMALE. Unknown lacking glabrous punctate patches. Metafemora ex­ ceeding ebttal apices by about half length of club. RANGE; Central MeXIcan hIghlands (Guerrero, Length 8-1O.5mm Hidalgo). RANGE: Central Mexico to EI Salvador. (See fig. 9) MATERIALEXAMINED. MEXICO. 1male, Hidalgo, 1km N Tlachinol, Hwy 105, 30-31 V, 1984 (D L REMARKS: This species may be distinguished by the Caldwell) [JW]. pair ofwide, distinct, golden pubescent vittae on the REMARKS: This species appears to be rare in collec blackprothorax From R tenuis, which a1somayhave tions. Itmay be recognized immediately by the metal­ a vittate pronotum, it may be separated by the larger lic greenish integument, yellowish pubescence, and average size, less coarsely, indistinctly punctate rugose pronotal disk. pronotum with the punctures less evident at the Vol. 7, No.1- 2, March· June, 1993 35 middle of the disk bet\\' een the \' ittae, and by the absence oferect setae on the rosternum.

(Figs. 3&4)

76; Fattig, 1947:27 (habits); Linsley, 1964:191, fig.60; os 'n . os in a: a is' os'n 1984b:80(habits); Hovore, Penrose, andNeck, 1987:308

Rhopalophora (Rhopalophora) longipes; Aurivillius,

RhopalophoraporosaWhite, 1855:206 (TYPE LOCALITY: " • tl

vittae; prosternum WIt a very arge, su va, granu- tar and punctate pateh in front of each coxa. Elytra more than 3 times as long as width across humeri. Figure 3 RhopalophoUl longip€s (Say), maill.

REMARKS: Rhopalophora rongipes is the northern- most component in a complex of closely related and superficially similar species It is sometimes difficult irregularly punctate, finely, indistinctly pubescent. to distinguish structurally, but is the only member of Abdmnen densely whitish pubescent, fIfth stemite the genus found m the midwestern and eastern :~Z:~;o~iX:~:~~~:;~:~~~;~~er~;~i7:I~~~ regions of the United States. This species may most readilybe separatedfrom othermembersofthis group Length 4. 5 9mm. of species by the fmm of the plOlloturn, whieh is subangulately widened behind the middle. In addi- FEMALE. Similar to male. Antennae at most barely twn, R long£pes may be separated from R. surpassing elytral apices Prosternum lacking oval punctatipClUbis and R. miniatoooUis by the indistinct granular patches. Metafemora barely surpassmg pronotal punctation, and by the shorter legs and ap~es. Len~ w~. shorter tenninalantennalsegment,fromR. bicofmella = by the entirely red integumentandnarrow pubescent vittae or the pronotum, and by the somewhat lOnger legs and antennae; from R. rugicollis by the lack of pronotal rugosities; from R. meeskei by the more 36 Insecta Mundi

I \ 1/If ___d ) I \ )Jf\

f'l I '\ \ / /L /1.1. / I r I

\ \ J p 'I \I .'"• \ \rr I J\ Y ~ \.) i1r- --J\ • • .. --­• '} \

o o

Figure 4. Know n geogr aphic distribution ofRlwpqlophoTU longipes (Say) [solid cit des]; and R. bicolotella Knull [open circles].

elongate form of the pronotum, and pale, not golden Rhopalophora longip€s m€€sk€i; Linsley, Knull, and pubescence; from R. serripennis b; the less coarse Statham, 1961:19, hg. 12 (habits); Linsley, 1964:192; elytral punctUles, and nmt-asperat epipleural mat­ Rice, Turnbow & Bovoro, 1985'20 ginS~ ~ ~~ Rhopalophora (Rhopalophora) longipes var. Meeskei; and from ant::tata the large, round AunVi1hus, 1912-431

DESCRIPTION: MALE: FormmOderatelysmall, slen der, elongate; integument black, prothorax red with baseandapex narrowly infuscated. Head moderately coarsely punctate, moderately sparsely pale pubes- Rhopaiophora meeskei Casey cent; antennae exceeding elytral apices by about 2 to (Fig. 5) 3 segments, scape scabrous punctate, with base dis tinctly impressed, eleventh segment curved, slightly RhopalOphora m€esk€l Casey, 1891:30 (TYPE LOCALITY: shorb~I than tenth. PIOnotum subquadra~ or very Las Vegas, New Mexico); Hamilton, 1892:159;Leng . h lin r h n i i r un nl an am' ton, 1896:168. rounded, widest behind middle, nearly as wide as Vol. 7, No.1 - 2, March· June, 1993 37

• •• \ { .r/ " • "- \ "' V..---- l • • \ 'i .---.-- '\ ) • • / \ • ...... -. • ~.--..., J \ • ~. \ ! ~ .. ~ • II ...... " --. \ • ( \j " ••• ( \ ,J A

"'\

Figure 5. Known geographic distribution ofRhopubJphoTU meeskei Casey.

lar; pubescencefme, short,indistinct. Abdomenclothed surface slightly uneven, with a small, obtuse tubercle with short, fine, pale pubescence. Legs with on eaeh side near base; punctures round, moderately metafemOla exceeding elyttal apices by at most half coarse. moderately dense; each side with a more or length ofclub. Length 6-1Omm less indistinct vitta composed of short, fine, pale orange or golden, appressed pubescence; prostemum FEMALE: Fonn similar to male. Antennae barely wIth a large, round, shghtly swollen, granulate patch exceeding elytrar apices. Prosternum lacking large, of moderately fine punctures on each side in front of roundpunctatepatches. Metaremora barely reaching coxae; meso· and metasternum clothed with very elytral apices. Length 6-1Omm. ShOI t, fale, whitish pubescence. Ely tIa 8 timesas lung as width across humeri, sides subparallel, margins RANGE- South Dakota and Colorado to western feebly serrate; apIces smuate-truncate, WIth hmd Texas and Arizona, south to Guatemala (See fIg. 5) angles dentate; disk convex, with an indistinct costa at the middle of each side from humerus to about REMARKS: This species may be distinguished from middle, punctUles dense, modemtely comse, inegu- othms in this study by the broader form of the 38 Insecta Mundi

pronoturn, which is aboutaswide aslongwithbroadly elytral apices by about 1/2 length of club. Length 6­ rounded sides and indistinct pubescent vittae com- tOrnm. prised of short orange pubescence; by the large prosternal punctate patches ofthe males; and by the RANGE: Pacific slope and West coast ofMexico. (See densely punctate, black elytra. IndivIdualS from the fIg. 6). southern part of the range generally appear to have slightly longer legs and antennae. REMARKS: This species may be distinguished from Adults have been collected on a wide variety of R. longipes, R. pU1'Wtatipennis, and R. sern'pennis by blossoms No larval hosts have been recorded thecombination ofround, moderately dense pronotal punctures, oblong prosternalpunctate patches of the Rhopalophora miniatooollis Ghevrolat male, and by the longer legs and antennae. The (Fig. 6) specimen identified as the type [BMNH] exhibits somewhat less dense than average prmlOtal puncta- Rlwpalophora miniawcollis Ghe.rolat, 1859.61 (pyPE tion. Further variation within the species appears in LOCALITY: "Mexico"); Bates, 1880:62; Bates 1885:310. the color and pubescence of the pronotum, WhICh IS Rhopalophora (Rhopalophora) miniawmllis; Auri.illius, darkerand with widerpubescent'1ittae in thecoastal 1912:431. Jalisco specimens

DESCRIPTION: MALE: Formmoderatelysmall, slen­ der, elongate; integumentpiceous, with headpiceous Rhopalophora nigriventris Bates tored,prothoraxredtodarkred, elytra dullcastaneous Rhopalophora nigriventris Bates, 1885:310 (TYPE LOGALITY."Purula", Guatemala. [Pur ulhlt, Baja Verapaz]). RlwpalophoFa (Rhopalophora) nigriventris; Aurivillius, scabrouslypunctate, thirdsegmentoften flattened or 1912:431. feebly grooved, eleventh segment subequat to tenth. PmnDtum about 1 2/5 as long as wide, sides feebly DESCRIPTION: MALE: Form moderate-sized, elon- rounded, WIdest behmd mIddle; diSk WIth surface gate, slender, integumentblack: with vertex andocci­ moderately densely, moderately coarsely, irregularly putofhead, entire prothorax, mesosternum, pro- and punctate, punctures round, deep; each side usually mesocoxae reddiSh-orange, elytraslightlytmgedWIth with a distinct, moderately wide pubescentvittaeOln- reddish at base. Head with frons and vertex nearly posed of short, pale pubescence, erect setae very impunctate, pubescencevery fme, obscure; antennae sparse, ifpresent; prosternum clothed WIth fme, pale exceeding ely trat apices by about :3 segments, scape recumbent pubescence, interrupted in front of each flat.tened dorsally, eleventh segmentsubequal totenth coxa by an mdiStmct, oblong, subtriangular, glabrous Pronotum longer than WIde, SIdes unevenly rounded, pateh of fine punctures; meso- and metastemum slightly swollen on each side behindmiddle, shallowly clothed with fine, pale pubescence. Elytra slightly impressed in middle before base, base slightly wider more than 3 times as long as width across humeri, than apex, disk with surface opaque, impunctate, sides subparallel, sometimes slightly tapering, or obscurely clothed with very fme, short, recumbent., slightly narrowedm mIddle, epipleuraImarginsmod­ pale reddish pubescence; prosternum WIth a moder­ erately asperate; apices truncate, with apical angles a;e~T small, oval, punctate patch on e~ side in front dentate; disk slightly depressed on each side at base, smfacemoderatelydensely, moderatelycoarselypJmc- tate, pubescenceveryfine, short, appressed, partially tures on metasternum, pubescence very fme, short, obscurmg surface, but not obscunng punctures. Ab­ silky, not obscuring surface. Elytra about 3 tImes as domen clothed with fme, pale pubescence. Legs long; longaswidth across humeri, sidesslightlyincuntedin metafemora exceedingelytralapices by atleastlength mIddle; apIces subtruncate, WIth mner angle feebly ofclub. Length 6-lOmm. dentate; disk flattened, subdepressed, with moder ately densely scattered, moderate-sized punctures, FEMAI,F> Form similar to male Antennae exceeding becoming slightly smaller toward apices; pubeseence elytralapices by about 3 segments. Prosternum lack- indistinct, very fine, short, not obscuring surface. ing distinctpunctate patches. Metafemora exceeding Abdomen mOderately clothed WIth fme, short, silky Vol. 7, No. 1 - 2, March - June, 1993 39

\ ,..- "( "'\ V \J1V \ \ l.J --- \ I \

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FiguIe 6. Kno>,n geoglaphic distIibution ofR,'wpaloph()ffl serripennis, new species fsolid squaIes], R. punctatipennis Linsley [solid circles); and R. miniatocollis Chevrolat [open circles). pubescence Legs moderately long; metafemora ex- REMARKS' This species may be easily separated ceeding elytral apIces by abOut length ofclub. Length from other RlWpaliJphora by the obscure dorsal and 10 12mm. ventral pubescence, '.vhich allows the integument to appear brightly and distinctly black and red-orange. FEMALE. Form similar to male. Antennae at most The species is quite rare in collections. barely surpassingelytral apices Prosternum lacking sman punctate patches. Metafemora at most barely QXCQQding Qlytral apicQs. LQngth 1O.5mm. Rhopalophora prorubra Knull RANGE. Guatemala. RMpaWph()fflprorubla Knull, 1944.91, pI. 1, fig. 7 (fYPE MATERIAL EXAMINED. GUATEMALA. 1 male, 1 LOCALITY: Sabino Canyon, Santa Catalina Mts., female, Baja Verapaz, 6-9km E Pumilia, 15-24 IV, Arizona); Linsley, 1964:189, fig. 59; lIo,ole and 1990 (E. GIesbert)[EG). Giesbert, 1976:357 (habits). 40 Insecta Mundi

DESCRIPTION. MALE. Fmmmodewtely small, elon­ male, SonUla, Alamos, 15-20 VII, 1958 (R.L. gate; integument black with bead, protborax, Westcott)[I,ACM] mesosternum, procoxae and mesocoxae red. Head with frons sligh~T concav~, d@s~ly punctat~; ,,~rt~x REMARKS: This sp~ci~s may b~ s~parat~d by th~ transversely rugose, pubescence inconspICUOUS; an­ comblllatlOn of the red head, prothorax, and tennae extending 8 segments at most beyond elytral mesosternum, the nearly glabrous pronotal disk, and apices, eleventhsegmentsubegualtotenth. Pronotum reduced elytral pubescence. Adults have been taken longer than wide, widest behindmiddle, sides moder­ on flowers ofCondalia, Baccharis, andMimosa. Lar­ ately rounded; base wider than apex; disk feebly, val host associations have not been recorded obtusely tuberculate on each SIde of mIddle and near base; surface sparsely, moderately fmely punctate; Rhopa,'ophora punctatipennis Linsley pubescencealmostentirelyabsent;prosternumfmely (Fig. 6) rugose andpunetatebehindapicalthird, with a small, Rlwpalophora punctatjpennis Linsley, 1935.92 (TYPE LOCALITY: Bejucos, Temescaltepec, Mexico. [Stateof across humeri, widestathumeri, subparallelto apical Mexico]). 1/6, then broadly rounded to apex, margins feebly asperate, apices tlnncate, sometimesfeebly spiculate, DESCRIPTION. MALE. Form moderately small, slen- with hind angles usually dentate; disk with surface der, elongate; integumentpiceous black, with protho- ~oderatelyshining,coarsely, densely, punctate, punc­ raxred. HeadmOderatelycoarsely, mOderately densely tures vaguely subseriate; pubescence inconspicuous, punctate, pubescence short, fme, moderately dense; shghtlymore distmctnearscutellum. Abdomen fmely antennae usually n-segmented, exceeding elytral punctate; pubescence short, dense, not obscuring apices by about 4 1/2 segments. eleventh segment surface. Legs slender; metafemora exceeding elytral longer than tenth, feebly appendiculate, rarely di­ apices by about 112 length ofclub Lengtb 5-7mm VIded to fOrm a Short twelfth segment. Pronotum slightly longer than wide, sides rounded, tapering FEMALE: Form SImilar to male. Aritennae and slightly toapex, widestbehind middle, nearly as wide metafemora at most barely attaining elytral apices. as elytmlhumeIi; disk even, surface with modemtely Prosternum lacking small punctate patches. Length sparse, often transverse, shallow punctures; pubes­ 5.5-8mm. cence sparse, fine, sllberect, pale, with a narrow vit.ta of short, appressed, pale pubescence on each side; RANGE- Sonthern AriZona to northwestern Mexico prosternum with a large, subovril, granular, glabrous (Sonora). punetate pateh on eaeh side in front of coxae; meso- and metasternum clothed with fine, appressed, sil­ MATERIAL EXAMINED. ARIZONA. 1 male, 1 fe- verypllbescence Elytra about3 timesaslongaswidth male, Pima Co., Madera Canyon, 6 VIII, 1971 across humeri, sides subparallel, epipleural margins (E Giesbert)[EG]; I male,8females, Pima Q) ,Sm! W at most feebly asperate; apices truncate, "vith hind Box Canyon, 14 VII, 1975 (Giesbert)[EG]; 1male, Box mar~inS dentate; disk with surface shining; ~unc­ Canyon, Santa Rita Mts., 19 24 VII, 1978 (A.E. tUllY modemtely coalse, modetately dense; p bes- Lewis)[J\V); 1male, PimaCo., SabinoCanyon, 25'111, eenee vely fUle, short, appressed, not obscuring sur- 1974 (N. Rulien)[JW); 1 female, Santa Cruz Co., face. Legs with metafemora exceeding elytral apices Patagonia. 10 VIII, 1958 (F.G. Werner) [UA]; 1 fe- by nearly length OfclJlb Abdomen clothed with fine, male, Santa Cruz Co., 3 mi. NE Nogales, 22 VII, 1959 short, appressed, silverypubescence. Length 6-9mm. (RadfOrd, Patblrson & Samulllson)[OA] . NEW MEXICO: 2 females, Hidalgo Co., Guadalupe Can- FEMALE: Form similar to male. Antennae 11 seg- yon, 7 VIII, 1967 (J.W. Smith)[TAM). MEXICO: 32 mented, exceedingelytralapicesbyabout2segments, males, 35 fcmales, SonOIa, Minas Nuevas, 7 VIII, terminal segment s1Jbeq1J aI to tenth Prosternum 1952 (C.&P. Vaurie)[AMNH]; 1male, lfemale,Sonora, lackingglabrouspubescentpatches. Metatibiaebarely GUll"OCoba, I I- 19 VII, 1933 [LACM]; 1female, Sonora, exeeeding elytral apiees. Length 6-8.5mm. 10mi. W Alamos, 21 VII, 1954(M. Cazier, W. Gertsch, Bradts)[UCB); 1 female, Sonora, 7 mi. Vi Alamos, 8 RANGE. Mexico to Guatemala. (See fig. 6) VIII, 1964 (JA Chemsak & J. Powell)[UCB); 1 fe- Vol. 7, NO.1 - 2, March - June, 1993 41

REMARKS. This species is very similar to R. with small, round granulate patchesofdistinct punc­ miniatocollis and R. longipes. It may be separated tures in front of coxae, basal 2/3 clothed with ap- from the former by the slightly shorter and wider pressed, silky, pale pubescence; meso- and pronotum with sparser, shallower, usually trans- meta:;;ternum clothed with fme, appressed, whitish verse punctures, and the larger, subOval prostemal pubescence. Elytra slightly more than 3 times as long punctate patches and longer terminal antennal seg as '.vidth acrosshumeri, sidesnearly straight, slightly ments ofthe males. From R. longipes it differs by the tapering; apices subtruncate, with hind angles feebly slightly tapering,subcylindIicalfoIlnoftheplOnotum, dentate,disksubdeplessed,punctUles modelate-sized, the longer appendages, the elongate, appendiculate slightlyfmer towardapices, moderatelydense; pubes- eleventh segment of male antennae (except in those cence short, dense, appressed, dull golden, somewhat cases where the appendiculationhasformed a twelfth obscuring surface. Abdomen clothed with fine, ap segment), and the male mefiifemora surpassing the pressed, whitish pubescence. Legs moderately long; elytralapicesbyaboutthefull club length. Occasional metafemora exceeding elytIal apices by about length specimens are found with the pronotal disk slightly of club. Length 8-13.5 mm. rugose. These may be separatedfrom R. rugicoltts by the absence of shallow longitudinal grooves on the FEMALE: Form :;;imilar to male.•Antennae exceeding thIrd and fourth antennal segments, by havmg the elytral apIces by less than 3 segments. Prostemum terminal segment ofthe male antennae acute, feebly lacking small punctate patches. Metafemora exceed appendiculate, and longer than the tenth, andby the ingelytral apices by about halflength ofclub. Length slightly longer metafemOia. In a small pelcentage of 8-12nlln. the males examined the antennal appendiculation is developed as a short but distmct twelfth segment. No RANGE: Southern MeXICO (Chiapas) to Costa RIca. other

DESCRIPI10N: MALE: Form mooorate-siZed, €lIOn REMARKS: This species may be distinguishedby the gate, slender; integument piceous, with elytra dull piceous integument with the apical margin of the brownish piceous, a moderately wide band across plostemuIll widely mange, the golden pwnotal pu- apex ofprosternum orange, and sometimes extreme bescent vittae, and the dull brown elytra. base and apex ofpronotum tmged with orange. Head WIth frons moderately densely p1lDctate, densely to moderately densely clothed on frons and vertex with Rhopalophora rugicollis rugicollis (LeConte) appressed golden pubescence; antennaelong, exeeed Rhopalophorus rugicollis LeConte, 1858:83 (TYPE 1,( )( :AI ,ITY ·"Texas") Rhopalophora rlIglCOUIS Leng, eleventh segment slightly shorter than t.enth 1886:31;Casey, 1891:30; LinsleyandMartin, 1933:181 (habIts); Linsley, 1964:190; Turnbow and Wappes, Pronotum longer than wide, sides unevenly rounded, ~::~=(habits);Hovore,PenmseandNeck 1987-308 often slightly swollen behind middle, base slightlY wider than apex; disk with an obtuse tubercleoneach RhopaloDhora (RhoDalophora) rugicollis; AllrivillillS side neal base, and a shallow, median, longitudinal 1912:431. RhopafJJphora rugulosus; Knull, 1934:336 depression near base; surface sparsely punctate, (error), clothedon each SIde ofa WIde, median, glabrous VItla withdense, appressed, goldenpubescence;prosternum 42 Insecta Mundi

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FigUIe ..,. Known geogt aphic disttibutioll ofRlwpalophUla 1 agicullis 1 agicullis (LeConte) [solid circles], and R 1 agicullis hovorei. new subspecies [solid squaresl.

DESCRIPTION.MALE. POI m mudelatelysmall, elun- cent. Elytra about 3 times as long as width aeross gate, slender; int.egument piceous, protborax often bumeri, sides subparallel, feebly narrowed in middle, reddiSh. Head densely punctate, pubescence whIte; outer margms feebly serrate on apICal 172; apIces antennae exceeding elytral apices by about 4 seg sinuate truncate, hind angles dentiform; disk -Nith ments, segments 3 to 5 with dorsal surface shallowly surface moderatelycoarsely, moderatelydenselypunc­ sulcate longitudinally. Plonotum longel than wide, tate, pubescence moderately dense, fIne, shOl t, ap- slightly wider at base, sides feebly rounded; disk pressed, pale. Abdomen densely white pubescent, transversely plicate and rugose from base to apex at WIth scattered suberect setae; fifth sternite rounded middle, puncturei'i i'icattered, moderately coari'ie; pu- at apex. Legi'i elongate; metafemoraexce@ding @lytral bescence on each side fine, appressed, pale, often apices by at least half the length of club. Length 7­ forming mOle 01 less distinct, moderately wide vittae; lOmill. prosternum with basal 3/4 clothed with pale pubes­ cence except. for a large, opaque, subquadrafe or FEMAI oK Form similartomale Antennae with basal rounded, granulate patch on each side in front of segments at most slightly flattened, lacking distinct coxae; meso- and metasternum densely white pubes- longitudinal sulci. Metafemora at most slightly ex- Vol. 7, No.1· 2, March - June, 1993 43

ceeding elytral apices. Plostemum lacking opaque ~ short, with metafemOIa I earning about apical 1/10 glabrous patches. Length 7-9.5mm. of elytra. Length 6-8mm.

RANGE' Southwestern United States (Texas, New RANGE- SouthernBajaCalifomia, Mexico (See fig 7). MeXICo and ArIZona) to MexIco and Guatemala. (See fig. 7) TYPES: Holotype male, allotype [CAS], and 14 paratypes (8 males, 6 females) from MEXICO, Baja REMARKS. This species differs from others in this California Sur, 5 mi. N Santa Anita, 2-3 X, 1981 (F. study except R exim;a, which bas metallic integu. Hovore, R Penrose)[FH, RPJ 9 additional paratypes, ment, by thedistmctlyplIcateorrugose pronotaldisk. all from Baja CalifornIa Sur as follows: I male, 9 mI. Howwer, specimens have been seen ofboth R. tenuis N, 2-5 mi. W SantaAnita, 20VIII, 1982 (Hovore)[FH]; and R. punctatipennis in which the pronotum is 1 male, San Pedro, 5 VII, 1919 (G. F. Ferris)[SU]; 1 slightly rugose. From these indi\'iduals R. rug;,collis male, 1 female, San Pedlo, 23 VIII, 1976 (E. may be separated by the shallowly grooved third and Giesbert)[EG]; 1 female, 10 mi. N EI Triunfo, on fourth antennal segments, and from R. tenuis by the Asclepias sp., X, 1966; 10, 5km NEITriunfo, 24 VIII, large male prosternal patches 1976 (Giesbert)(EG); 2 females, 6 mi. N San Jose del Vanation m elytralpunctatIOn andpronotalcolor CabO, lUIX, 1967 (J. Chemsak,A&M. Michelbacher) in this species is notable. Individuals resembling R. [UGB); 1 male, 1 mi. 8'11 Punta Palmilla, 13 IX, 1967 lineicollis, with a piceous pronotum bearing golden (Chemsak, Michelbachers)[UCB]. pubescentvittae, occurfrequentlyinthemoresouthern populations R rugicolliR may be separated from the REMARKS' This isolated and subspecific population latter by the broader, rugose pronotum, and m the differs from R. ruglCollu; ruglCollu; by the shorter males by thelarge, distinct, ovalprosternalgranulate appendages and piceous pronotum, and is sympatric patches, and more distinct grooves on the third and withR. bicolorellaintheCapeRegion ofBaja California. fourth antennal segments. A number of inm\'iduals It may be easily separated from that species by the seen from Arizona havesomewhatfiner elytralpunc- rugose pronotum and sulcate third antennal seg- tation. Adults have been reared from Celtis and ments of the male. In additIOn, the pronotum m the Pitlutcellobium in the northern partofthe range, and Cape populations of R. lJicolorella is nearly ahvays collected at blossoms, including those of Mimosa, red, with the disk slightly impressed behind the Acacia, Lupinus, and Cearwthus. middle and considelably less densely pubescent. We take pleasure in naming this subspecies for Frank T. Hovore, who first brought it to our attention Rhopalophorn rugicollis hovorei, new subspecies (Figure 7) Rhopalophofa serripennis, new species (Figures 6, 8 ) Rhopalophorarugicollis; Hom, 1894:338; Linsley, 1942:58. DESCRIPTION: MALE: (See fig 8) Form small, DESCRIPTION: MALE: Form similartoR. rugicollis elongate, slender; integument black, prothorax red, lugiwllis. Integument piceous tllloughout~Anten­ WIth base and apex often narrowly intuscated. Head nae exceeding elytral apices by 2 to 3 segments; third with scattered, moderately coarse punctures; pubes- segment longitudinany sulcate. Pronotum trans­ cence moderately sparse,pale, applessed,vel texfinely, versely, irregularly rugose; disk with whitish pubes transversely rugose; antennae exceeding elytral api- cence narrowly interrupted in middle from base to ces by about 4 segments, scape moderately slender, apex,pIOstenmmwithopaqueglabIOuspateheslalge, canaliculate Pronotum about 1 1/2 times as long as oval. Legs slender. short, with metafemora not or wide, subcylindrical, slightly tapering, or with sides barely attaining elytral apices. Length 6·8mm. slightly rounded, 'Nidest behind middle; disk with a pair ofvery feeble tubercles on each side near base; FEMALE: Form similar to male. Antennae reaching punctationmoderatelysparse, pubescence,ifpresent, to about apical 1/8 of elytra, basal segments lacking ~=i~t~:~:::~~;:::~;~~::e:~~~n;t~:x;;: sulci. Prosternum lacking opaque glabrous patches. punctate patches small, indistinct; meso and 44 Insecta Mundi

[KR); 1 female, 5km. E La Ventos~, Oaxaca, 25 VI, 1989 (Cope}(JCl; 1 female, 3mi. VI Tehuantepec. Oaxaca, on Prosopis juli/lora, 20 VII, 1965 (G.H. Figure 8. Rhopawphorn serripennis, new species, male. Ne1son)[GN] metasternum densely clothed with silky, white pu­ REMARKS: This species may be separated from R. bescence. Elytra 3 times as long as width across punctatipennis and R. miniatocollis by the strongly asperateepipleuralmarginsoftheelytra, thecoarsely, humeri somewhat narrowed before middle mar~ns stIUngG aspemte, apices sinuate-truncate, ill er subseriately cribrate-punctate elytral disk, and the small, indistinct prosternal punctate patches of the ~n~les dentate; disk r~arsebY' densely cribrate.p~lDc. ate, punctures usua y su senate; pubescence me, male. Variation occurs mainIY in the wm, pubes short, appressed, notobscuring punctures. Abdomen cence and punctation of the pronotum. A number of specimens have been seen from clothed with silk;, white ;ubescence. Le~ with metafemOlasUlpa singelytl 1apices by abou ength westem Mexico which exhibit plffilOtal punctatiffil ofclub Length 5-9mm typical of R. miniatocollis, and can be placed only dO'lbtfuDy ill th~ pr~s~nt SP~Cl~S: 2 males, 2 f~mal~s, Guerrero, Hwy 95, 6 mi. S Rio Mexcala, 5 VIII, 1965 (Nelson)[GN]; 4 males, Oaxaca, 3mi. E Juchitan, July 8, 1988 (Cope)[JC]; 5 males, Chiapas, 9-12km N Arriaga, .26 VI, 1~87 (Wappes, Giesbert)[JW, EG); 1

(Turnbow)[RT). RANGE: Central and southwestern Mexico from Morelos and Nayarit to Chiapas (See Fig. 6). Vol. 7, No.1· 2, March· June, 1993 45

Rhopat'ophora tenuis (Chevrolat) (Figure 9)

"Mexico"); White 1855:210. prono urn, erect setae on t e prosternum, t e ac 0 RhoparophOla tenuis, Chevtulat, 1859.61, Bates, 1880.62, distinct punctate patches on the male prostemum, Bates, 1885:310. and throughout most of its range, by the all black RlwpakJplwra(Rlwparoplwra) tenuis, Autio villius, 1912.431. integumental cohn, often with more-or-less golden Rhopalophora incrustata Chevrolat, 1859:62 (TYPE LO­ pubescent pronotal vittae CALITY: Oaxaca, Mexico), Bates, 1880.62, Bates, SOme confUSIOn eXISts regarding the Identlty of 1885:310; Linsley, 1935:91. NEW SYNONYMY Rhopalophora tAAuis, since Ghevrolat's original de Rhopalophorn (Rhopalophorn) ineruslala; i\urivillius, 1912:431. scription was of an individual with a red pronotum. Subsequently the same authm in 1859(b), keys the DESGRPTION: MALE: Form small, slender, elon species as having the pronotum the same color as the gate; integumentpiceousblack, with pronotum rarely body. that is, black. Bates (1885) calls the species dark red at sides. Head moderately densely, coarsely "drn '~fl I)"aassuming t?e Orig~l de~c:iptio~~ohave punctate, vertex moderately finely, transversely been ase on a speClIDen 0 . m£nwtoco {/,S. The rugoso-punctate; antennae exceeding elytral apices problem is compounded by the fact that this single by about 4 segments, scape moderately finely sca- specimen from the Mniszech collection has been im- brous-punctate, fourth segment halfas long as fifth, possible to locate, but later specimens labeled as R. terminal segment longer than fourth. PlOnotumless ten:.i.<: p~sentlY in ~e BMNH:nd MC~;t.lectin than 11/2 times aslong aswide, sides slightlyrounded, widest behind middle; diSk often narrowly, longltudl- nally impressed at middle of basal half, with incon­ author'slatertreatment. We arefollowing Chevrolat's spicuous obtuse tubercles on each side near base· subsequent charaeterization ofthis species based wt surface opaque, moderately densely, distinctly, mod~ available material. erately coarsely punctate except for a narrow, me­ The name incrustata was given by Chevrolat to a di~nct p~enotype dian, irregular, glabrous area; pubescence whitish fjj afR. tenuis with longerlegs, more golden- forming a pair of moderately wide vittae whiCh vary from dense and distinct to nearlY absent or indistinct; ;rosternum with ~unctate ~atches in­ individual, superficially similar to R. lineicollis. Only distinct, pube cence shmt, mu er atelyense, ap- about.] 0% ottbeMexican material examined exhibits pressed, whitish, with short, erect, pale setaeevident; meso- and metasternum moderately densely whitish pubescent. Elytra slightly more than 3 times as lOng as width across humeri, sides subparallel, epipleural of this phenotype would appear to be distinct from R. margins slightly asperate; apices sinuate truncate tennis. in large series of specimens taken at several hind an les dentate· disk moderatel densel unc~ locations in Mexico bothphenotypesoccur, alongwith ,, ppres e ,pa e 0 en, varying from sparse to dense AbdOmen clOthed with tion. In the material examined from throughout the moderately dense, whitish, appressed pubescence. Mexican highlands, intergradepopulations appear to MetafCmora exceeding elytral apices by at least 1/3 be abundant, and a gr eat deal ofstluctural variation length of club. Length 6-9mm. is found. VariatiOns in integumental colOr apparently oc- FEMAI.Eo Form similar to male Antennae exceeding cur only at the northwestern extreme of the range, elytral apices by about 3 segments, eleventh segment with abouthalftheindividualsseen from the Mexican subequal to fourth. Metafemora shorter, at most state of Sonor a having the plOllotum partially dark barely exceeding elytral apices. Length 6-9mm. redat thesides. These Sonoran individuals aresuper­ ficially similar to R bicolnrelln, hut the males may be RAN~E· Mexico (widespread) and Costa Rica (See easily separated from males of that species by the fig. 9 indiStinct prosternal punctate patches. Occasional specimens have been seen with a slightly rugose pronotal disk, but these are easily distinguishedfrom 46 Insecta Mundi

~ I r I ( \ ..., I" ... - 'I ) \f~ "'vJ\ '\ y~ }V '"\...'lJ\,l'"",, 1- 'I \ '-, ? \ 'P\..~ ~ 7~ ~ '\.t ~~ ~ -'"'""' ~ /!1 7" \.) '"~~. f·/, \~

.'

Figure 9. Known geographic distribution ofRlropaloph01a tennis (Chevrulat) [solid circles], alldR. lineicollis Chevrulat [open circles].

R. I ugicollisby the more slenderform, andby thelaek widest behind middle; disk slight.ly uneven, with a oflarge, munded granulate punctate patches on the shallowmedian impression nearbase; slutacesparsely. male prosternum. mdiStmctIy punctate, WIth transversely laid, tme, v,hitish pubeseenee forming an indistinet, '.vide vitta on each side ofa wide, median, glabrous space, later­ RhopakJpfJm8 yllcafana, new spedes ally with a wide, glabIOuS vitta above a narrow pubescentvittaoneachsideaboveprocoxa; prosternum DESCRIPTION: MALE: Form moderate-sized, slen­ shallowly lffipressed, transversely plicate, WIth an der, elongate; integument datk reddish, with flOns, indistinct, round, granulate patch of punctures on legs, and antennae piceous. Head with frons trans­ each side before coxae; base and middle between verselyimpressed, densely whitISh pubescent; anten- granulate patches whitish pubescent; meso- and nae exceeding elytral apices by slightly more than 4 metasternum moderately densely clothed with fine, segments, seape shallowly, longitudinallyimpressed, short, appressed, pale pubescence Elytra abOut 3 segments 3 to 5 flattened or shallowly grooved dor- times as long as width across humeri, widest at sally. Pronotumlongerthanwide, sidesfeebly rounded, humeri, tapering slightly; apices slightly sinuately Vol. 7, No.1· 2, March· June, 1993 47

tIUncate, with hind angles dentate, disk broadly TYPE SPECIES. RhopaJoplwra bidncta Hom (origi­ subdepressed in middle, slightly elevated and nal designation). subcostate near sides; surface moderately coarsely, Rhopalophorella differs from Rhopalophora by irregularly punctate; pubescence short, moderately the fasciate elytra, carinate metepisterna, and dis- dense, appressed, notobseurmgpunctures. Abdomen tallyflattened, subserratefemale antennae. Thesmgle moderately densely clothed with short, pale pubes speciesknO'Nn alsodiffersbyhayingpaleappendages, cence. Legs elongate; metafemora exceeding elytral and is apparently limited in distribution. apices by about length ofelub. Length S-lO.5nnn.

FEMALE: Form sImilarW male. Antennae exceeding Rhopalophorella rase/ala (LeConte) elytral apices by slightly more than 2 segments. Prosternum lacking punctate patches. Metafemora Gracilia fasciata Leconte 1873:171 (TYPE LOCALITY: exceeding ely tralapicesby slightly less than length of "LowerCalifornia" [Baja California]); Leng, 1884:119; club. Length 8-1Omm. Linsley, 1942:50. RlUJpalvphol ellafasciata, Chemsak and Linsley, 1982:48; TYPES' Holotype male [CAS] from Piste, Yucatan, Hovore, 1988:16. MEXICO, 12 VIII, 1967 (KC. Welling). 24paratypes Rlwpalophorabicincta Hom, 1895.245 (TYPEbeCALITY. from MEXICO as follO'tvs: 1 female, GhichCn !tzu, San Jose del Cabo, Mexico); Leng and Hamilton, 1896:168. YucatBn, 18 VIII, 1965 (N.J. Dickey)[RW]; 1 male, Rhopalophora (Rhopalophora) bicincta; Aurivillius, 20km N Calillo Puerto, Quintana Roo, 12-14 Til, 1983 1912:431. (J Wappes) &IW]; ] male, same locality and dates (E Rlwpawphorella hicincta; Linsley, 1942:58 (pi. 4, fig. 6). Giesbert)[EG]; 1 female, 18km N Carillo PuertO, QuintanaRoo,28V, 1984(R. Tumbow)[RT); 1female, DESCRIPTION: MALE. Form small, elongate, slen- 17km NW Felipe CarilloPuerto, QuintanaRoo, 18VI, der, sides subparallel; integument dull black, anten­ 1990 (Fnrnbow)1RT1; 4 males, 6 females, same local- naeand legs tesf.aceolls, femora dark testaceOIlS Head ity, 1VI, 1992 (F. Hovore)[FH); 1male, LeonaVicario, narrowerthan pronotum; antennae about 11/4times as Qumtana Roo, 26 V, 1992 (Hovore)[FH); 1 female, long. as body, thirdsegmentlonger than. seape, nearly , fIfth, segments 5 to 11 subequal; eyessmall, lowerlobe wider than long, UPpel lobe small. narrow. PnIDotum with sidesrounded, widestbehind middle; diskfinely, mOderatelypunctate, WIthoutpubesc€lnc€l; prost€lmum REMARKS' Thisspecies differs from R cltpricollis by elongate, with a large granulate pat:eh on each side in the dark reddIsh mtegumental color, and the some­ front ofcoxae; meso- and metasternum with patches' .....hatmore even disealsurface andless distinctpubes " of silvely white pubescence latetally, metasternum cence of the pronotum. moderately convex, metepisterna narrow, slightly arcuate, with a longitudinal carma over about 273 Of the length. Elytra widest near apex; disk flat, without Genus RhopalOPhorelfa Linsley lateral carinae; apices obtuse, tri or quadtidentate, diskdensely cribrate-punctate, with fasciae ofsilveIY Rhopalophorella Linsley, 1942:57. white pubescence across base and obliquely behind middle Abdomen witb a silvery wbite lateral pubes- DESCRIPTION: Form small, slender, somewhat de­ cent patch on each side offrrst sternite; fifth sternite pressed. Elytra WIth pub€lsc€lnt fasciae. Head with shorter than rourth. Legs with metafemoFa surpass frons oblique;vertexevenlyconvex; antennrefIliform, ing elytral apices. Length 3.5-5mm. eleven-segmented, lacking setae, female with outer segmentsflattened, subsellate; scave robust, davate FEMALE- Form similar m male Antennae shorter, Pronotum unarmed, elongate, constrictedatbase and about 2/3 as long as body, with distal segments flat­ apex Elytra WIdest near ap€lX; disk flat, without tened,subserrate,eleventhsegmentIOngerthan tenth. lateral carina; apices obtuse. Underside with Metafemora not reaching elytral apices. Abdomen metepistema longitudinally calinate. Abdomennol­ with fifth stemite lungel than four tho Length 4-5mrn. mally segmented. Legs with frrst segment of meta- tarsi nearly 3 times as long as second segment. 48 Insecta Mundi

RANGE. Soathem Baja Califomia, Mexico. Rica to BI asil. (subgenus Ulododerus) . .. G. (u.) barbatus Gounelle MATERIAL EXAMINED: MEXICO, BAJA CALI­ Antennae andlegs"ithout setalblushes (subgenus FORNIASUR: 2 mahls, 1fQmalQ, Triunfo, 7 VII, 1938 Cycnoderus) 2. (Michelbacher & Ross) [CAS]; 2 males, 11 females, 6 2.(1) Elytraldiskwith wide, shallow longitudinaldepres­ km N EI Triunfo, 21 VIII, 1976 (E.Giesbert) [EG]; 1 sion near suture densely punctate and clothed female, same locality, 15 VII, 1938 (Michelbacher & with pale, transverse pubescence, punctation and Russ) [UCB], 1male, 25mi. 'vVLaPaz,4IX, 1959 (K.'vV. pubescence sparsertowardmargms; metafemora Radford & FG Werner) [IICB]; 1male, 8 mi NW La abruptly clavate 3 Paz, 18 IX, 1967 (J. Chemsak, A&M. Michelbacher) Elytral disk lacking depressIOn, punctatIOn and [UCB]. pubescence nearly uniform; metafemora gradu- ally, moderately clavate 5 REMARKS. This species may be easily recognized by the small size, black integument with pale append- 3.(2) Antennae of females with third segment distinctly longer than fourth. densely fringed along inner ages, and the distinct silvery white fasciae of the side with black hair-like setae; length at most ~Iytr~ Spe~;enshavQ ~n ta~QI~ on gowe~:.tclud- barely exceeding elytral apices. Length 8-9mm. Guatemala (male unknown) ...... 0. (G.) guatemalicus n.sp. Antennae with third segment subequal to fourth, ~ellus CycnodelUs Audillet-Ser ville lightly fringed along inner side; length exceeding elytral apices by at least 3 segments 4. Cycnoderus Audinet-Serville, 1834:101; White, 1855:213; 4. (3) Elytralpunctation nearbase andsutureverydense, Thomson, 1860:202; Thomson, 1864:218; Lacordaire, fine; scutellum densely pubescent; pronotum of 1869: 112; Gounelle, 1911:99; Martins and Napp, males bearing long, sinuate grooves at sides; 1989:65. femora with patches of fine, recumbent, silky puboscence.umgth9.5-14mm. (See fig. 10). Mexico DESCRIPTION. Form moderately small, (Chlapas) C. (C.) mrgmwe n. sp. subdepressed, elongate, sidessubparallel; integument Elytralpunctation nearbase andsuture moderately shmmg, often WIth metallic reflectIOns. Antennae 12­ dense andfine; scutellum indistinctly pubescent; segmented in malQs, II-segmented in females, ocell pronotnmofmales lackingsinnategrooves; femora sIOnally WIth tufts of harr-like setae. Pronotum un­ WIthOut recumbent, silkypubescence. Length '7.5­ armed, elongate, disk somev.'hat flattened, lightly 11 5mm Mexico (.Ialisco, Guerrero) sculptured. Elytrasomewhatflattened, apices acumi- ...... C. (C.) cope£ n.sp. nate. Abdmnen nOllnally segmented. Legs slender, 5.(2) Integument metallic blue-black; pronotum 1 1/2 femora pedunculate-clavate, posterior femora and times as long as wide; antennae with third seg- tarSI occaSIOnally tufted. ment fringed on underside only. Length 9.5-11 mm. Mexico (Guerrero) ...... e eel lividus n.sp. TYPE SPECIES: Cycnoderus tenuatus Audinet­ Integument metallic dark reddish-brown with legs Serville (monobasic). CyclwdefUs was listed by and prunutum often reddish; prunutum about 1 1/3 Blackwelder (1946) as including six species Martins times longer than wide; antennae with third & Napp ( 1989) reaSSIgned an additIOnal speCIes from segmentfringed above andbeluw. Length 8-91111n. Rhopalophora. TheSQ sevenspecies are reportedfrom Mexico (Jalisco) G. (G.) brevicolle n. sp. COlombIa, Brasil, and Argentina Five new species from Mexico andGuatemala, and a northward range extension to Costa Rica of a South American species Subgenus Ulododerus Gounelle are recorded here. Cycnoderus (TUododems) Gounelle,191 1 '101 Key to MeXican and Central Amencan Species of Cycnoderus DESCRIPTION: Antennae 'Nith tufts of hair like setae on fifth segment in males, sixth segment in

1. Antennal segments 5 (males) 01' 6 (females), females. Legs with metatibiae and metatarsi beating metatibiae, andmetatarsibearingdistinctbrushes dense tufts oflonghair-like setae; metatarsiwithfIrst oflong hair-like setae. Length 7-1O.5mm. Costa Vol. 7, No.1 - 2, March - June, 1993 49 segment 1 1/2 times longel than following segments PanamapIO\l., Ceno Campana, 800m., 25V, 1990 (F. together. Hovore) [FH].

TYPE SPECIES' Cycnoderus (Wododerus) barbatus REMARKS This speciesis distinguishedby theblack GOunelIe (present desIgnatIOn). tufts of harr-like setae on the antennae and hind legs. Field obselVations by the authors indieate that these This subgenus is distinguished primarily by the distinctly tuftedhindlegs andantennae are extended tufted antennae, metatibiae, and metatalsi. A single laterally dUIing flight. species extends its range northward into Costa Rica

Subgenus Gycnoderus Audinet Serville Cycnoderus (Ulododerus) barbatus Gounelle Antennae lacking tufts of setae. Legs with metatibiae and metatarsi lacking tufts ofsetae; meta- Cycnoderu.s (Wododerlls) barbat/lS GOl! n aJla 191 1·1 OJ tarsi with first segm~mt subequal to following seg (TIPE LOCALITI: Jatahy, Goyaz, Brasil); Hovore ments together. 1989·252 TYPE SPECIES: Cycnoderus tenuatus Audinet· DESCRIPTION: MALE: Formmod@rat@lysmall, @lon Serville, 1834 (monobasic) gate, slender; integumentshining, metallicblue-black, Five new species of this subgenus are here reo el:yna somewhat more greenish metallic, bases of corded from MeXICO and Guatemala. metafemora, andvague longitudinal discal markings 011 elytra sometimes testaceous. Head short, moder­ Cycnoderus (Cycnoderus) brevicolle, ately densely punctate; antennae about 1 1/2 times as long as bOdy, apicaI 172 of fifth segment 6earrng a new species brush of long black hair like setae on inside, distal segments short. Pronotum more than 11/2 times as DESCRIPTION: FEMALE: Form moderately small, long as wide, disk glabIOus, witlI a medianlongitudi- mocleratelyslender, depressed, parallel-sided; integu- nal callus and 4 indistinct rounded calluses, 2subapi- ment shining, piceous-black with violet-metallic re- calandaless dlStrnct subbasalparr; puncturesIrregu­ flections, legs usually reddish, pronotum and head lar; presternum moderately densely, coarsely punc often reddish-brown. Head short, with punctures tate and rugose, pubescence moderately sparse, frne, moderately coarse and dense, irregular; antennae elect; meso- and metasternum shining, pubescence moderatel s arse erect. EI traabout 3 times as Ion s wi s u ri, ic ir s·g y e is- cent; apices with outer margins rounded, sutural with dark hair-like setae above and below, scape and angles acute; disk WIth moderately dense punctures otlIel basal segments fl inged below. PIOIlOtuIIl about nearsuture, becoming sparser andfiner towardmar 1 1/3 times as long as wide, sides slightly rounded, gins. Abdomen shining, pubescence moderatelysparse, tapenng feebly to apex; disk feebly sculptured, WIth erect Legsslender;femora ahruptlyclavate; metatihiae m@man longitudinal callus obscure, punctures mod with a distinct brush oflong black setae on each side erately dense, impressedandcoarse, long, erect setae of apicru 172; metatarSI WIth hrst segment 6earmg a numeIOus, pIO-, meso- and metasteIIlUm shining, brush of long black setae for its entire length. Length ;::~a:::r~;7::~~1@~:~::::lra~~~:::~:)z:~e~ 5.5-8.5mm. width across humeri; apices rounded, sutural angles FEMALE: Form similar to male. Antennae about 1 1/3 acute; diSk flattened, mOderatelydensely andcoarsely, tImes as long as body, bearrng a brush ofblack setae subunifOI'mly punctate; pubescence tranS\' erse, dalk, on inside of almost entire sixth segment; pronotum inconspicuous. Abdomen moderately coarsely punc- less distinctly sculptured. Length 7-10.5mm. tate, sparsely pubescent. Legs with femora shining, moderately gradually clavate Length 8-9mm MATERIAL EXAMINED: 1 male, COSTA RICA, Cartagoprov ,CATTE, 3Km SE Turrialba, 60Qm , 16 MALE: Unknown. V, 1985 (J. Chemsak) [UCB]; 1 female, same locality, 9·14 VI, 1988 E. (Giesbert) [EG]; 1female, PANAMA, 50 Insecta Mundi

TYPES. IIolotype female [Institutode Biologia, UNAM)] verse, subrecumbent pubescence, outer portion of and2 female paratypesfrom MEXICO, Jalisco, Est. BioL each elytron more sparsely punctate. Abdomen with Chamela, 1-8 XII, 1988 (J. Chemsak) [UCB]. terminalsternitesubequaltofourth, apexsubtruncate: pubescence fine, pale, erect, with frne, silky, recum- REMARKS: ThIS speCIes, slffiilar m punctatlOn to C. bentpubescence at sides. Legs long, slender, shining; lividus, may be separated by the shorter pronotum, femora abruptly clwv'ate, clavae nearly impunctate, reddish legs, and the moderately densely fringed pubescence sparse,pale, suberect. Length 7.5 - 11. 5mm. thiId antennalsegment. Thefringed thirdsegrnent is also characteristic orc gllnfemalieus, from which C FEMAlE' FoOD similar to male Antennae less than brevlColle differs by the somewhat more robust bOdy tWIce as long as bOdy. Pronotum WIth SIdes lackmg and the style of eJ)'tral punctation and pubescence. definedcalluses, erectpubescence evidentlaterally as The small type senes was taken from flowers. wellasdorsally. Prosternumlacking micropunctures. Abdomen with terminal stemite longer than fourth, apex rounded. Length 7.5-11mm. Cycnoderus (Cycnoderus) copei, new species

DESCRlP'I10N: MALE: FormmOderatelysmall, elOri gate, nearlyparallel-sided; integumentshining, black- additional paratypes from MEXICO as follows: 2 ish with metallic reflections; pronotum sometimes males, 7 females, Guerrero, 6km N Chilpancingo, 15- parbally reddish, ely traldiskdarktestaceous, heavily infuscated with metallic blackish near suture, mar­ gins, and apices, punctures darker HeadmOderateIy males, Guerrero, 6km N Ghilpancingo, 16 18 VII, short; puncturesmoderatelycoarse andsparse, denser 1984(Ribardo) [KR); 2females, lOkm N Chilpancingo, and coarser on vertex and occiput; pubescence erect, 22 VII, 1987 (R.Turnbow) IRT1; 1 male, 1 female, moderatelylong, fme, sparse; antennaeslender, more lalisco, 9km S Chamela, 15-19 VII, 1987 (Cope) [JC); than3timesas longasbody, scapelessthan 1/2aslong 1 female, Jalisco, 27.4km S Chamela, 16 VII, 1987 as tbird segment, wbicb is subequal tofourth, under- (Turnbow) [RT]. sideofsegments 2and3moderatelyfringedwith long, REMARKS. Qvcnoderus copei appears robe structur- ally closely related to the SouthAmerican C. tenuatus Audinet-Senrille We are pleased to dedicate this specie~ to James Cope, who first brought it to our base, with a longitudinal, shining, glabrous, shallow attentIon. callus at middle surrounded by coarse, deep punc- tures, sides slightly swollen, with punctures slightly more coarse, sparse, and irregular, and often with CYGnodems (Cycnodems) guatemaliclIs, irregularlateralcallusesdefinedbyindistinctareasof new species finer punctures, pubescence erect, fme, pale, longer than second antennal segment, and angled slightly DESCRlPI10N: FEMALE: Form small, slender, de forward, sparse on sides; prosternum very finely, pressed, parallel-sided; integument shining, dark densely micropunctate with deep, moderately sparse brown with strong aeneous retleetions, head and punctures interspersed; meso- and metasternum antennae darker, legs reddish brown. Head moder- alutaceous, moderatelysparselypunctate;pubescence ately short, punctation moderately dense, Irregular, fine, pale, erect, with fine, recumbent, silky pubes- denser on vertex and occiput; pubescence sparse, cence on prosternalprocess, mesosternum, and epis­ antennae slender, about as long as body, scape less terna. Scutellum transverse, sparsely pubescent. than 1/2 as long as third segment, 'tvhich is distinctly Elytra nearly 3 times as long as width across humeri, longer than fourth, with underside of third segment feebly tapeting, apices rounded, saturalangles acate, distinctly fl inged with black setae, fourth segment disk flattened, slightly depressed near suture; base lightly fringed on underside, distal segments short, and sutural depreSSIon densely, dIStmctly punctate, eleventhsegmentshorterthanscape. Pronotumnearly bearingmoderatelysparse, fine, pale, obliquely trans 1 1/2 times as long as wide, sides slightly rounded; Vol. 7, No.1- 2, March - June, 1993 51

baseandapexwidely, shallowlyimpressedandrugose; with a median, longitudinal raisedcallus sunounded disk depressed near base, with an indistinct longitu- by coarse, close punctures, sides of disk sparsely dinalcallus, punctation moderately com se anddense, punctate, with several raised swellings, a pair of irregular; puhescence sparse, fine, pale, erect to which are more distinct on each side of apical 1/3; suberect; prostemum coarsely, moderately densely pubescence sparse, long, dark, erect, nearly absent punctate; metasternum moderatelypunctate andfee laterally; presternum '.vith surface densely bly rugose at sides. Scutellum small, semicircular; micropunctate and sparsely, coarsely, deeply punc- pubescence shmt, pale, modeIately dense. ElytIa tate, meso- and metastemum moderately densely about 3 times as long as width across humeri, feebly punctate;puhescencepale,fine, erect Scutellum wider tapenng; apIces rounded to acute suturalangles; disk than long, mdlstmctlypunctateandpubescent. Elytra flattened, slightly depressed near suture; sutural slightly less than 3 times as long as width across depression densely punctate, bearing moderately humeri,feebly tapering; apices rounded, suturalangles sparse, fme, pale, obliquely transverse, subrecumbent acute; disk flattened, densely, eribIately punctate, pubescence, each elytron more sparsely punctate pubescence dark, sparse, obscure. Abdomen with luwmd mmgin. Abdomen with terminal sternite tenninal stemite about as long as fourth, apex trun­ suhtnJDcate at apex; surface nearly impunctate, with cate Tlegs with femora moderately gradually clavate; scatteredfme, pale, short setae. Legs slender; femora surface slightIy opaque, pubescence sparse, suberect. abruptly clavate, clavae nearly impunctate, pubes Length 9.5 1O.5mm. cence sparse, pale, suberect. Length 8-9mm. FEMALE. FalIn similar lu male. Antennae shorter, MALE· Unknown about 1 1/3 times as long as body, third segment 1 1/2 !lmes as long as fourth. Pronotum less dIstmctIy TYPES: Holotypefemale [CAS] and2female paratypes sculptured, erect pubescence evident laterally. from GUATEMALA, Tacat6n, Lago Amatitlan, near Prostemum lacking micropunctation. Abdomen with Villa Canales, 10 I, 1989 (E. C. Kondratieft) [FH, EO]. terminal sternite slightly longer than fourth, apex rounded. Length 9.5-11mm. REMARKS. This species shares characterswith both C copei and C brel1icolle, differing from the former TYPES' Holotype male, allotype [C t\S] and 12 prmcIpally by the antennal measurements and the paratypes (3 males, 9 females) from MEXICO, distinctfringe en the third segment..",hieh gives it the Guerrero, 10. 5km. N Iguala, 4OOOft., 19 21 IX, 1989 appearanceofbeingsomewhatheavierthanthefourth. (J. Wappes, R.Turnbow & E.Giesbert) [JW, RT, EG). From C. brevicolle it differs by the sculpture and 2additionalparatypes, 1male, 1female,fmmMEXTCO, pnnctation of the elytra, the somewhat more slender Colima, EI Terrero rd 4500£1 (vicinity orEI SaHz), 30 form, and the longer pronotum. IX-4X, 1991 (J. Wappes, R. Morns, E. Glesbert) [JW, EO].

CyGmxiezas (Cycmx'iezas) Iividas, REMARKS. Thisspeciesis distinguishedby its metal- new species lic bluish-black integumentalcolor. Itdiffers from the two preceding specie!> by the more or Ie!>!> unifOrm DESCRIPTION- MALE' FormmoderatelylOman, !>Ien- elytral punctation. Several specimens in the type der, depressed, parallel-sided; integument shining series were found clinging to narrow leaves ofgrass. metallic blue black, elytraless metallic, neady black. Headmoderatel short unctationmoderatel coarse esse ,ense vcr ex a ipu, u escence sparse, erect; antennae nearly 3 times as long as body, new species scape less than 1/2 as long as third segment, which is (Figure 10) subequal to fourth, tv"elfth segment cUlVed at tip, undersideofsegments2 and3fringed withlong, black DESCBTJYITON- MAl IE- (See fig 10) Form moderate setae. PlOnotum 1 1/2 times as long as wide, sides SIzed, elongate, nearly parallel-SIded; mtegument sligbtlyrounded, widestbefore base, taperingsligbtly shiningmetallie greenish black, prothorax reddish at to apex; base and apex WIdely, shallowly lII1pressed sides and beneath, bases offemora testaceous, elytra and transversely rugose; disk heavily sculptured, dark testaceous, heavily infuscated with metallic 52 Insecta Mundi

, " cence at sides; terminal sternite subequal to fourth, apex su rune e. en - mm.

lacking sinuate groove, erect pubescence evident lat-

sternitelongerthanfourth, apexsubtruncate. Length - mm.

TYPES: Holotype male, allotype [CAS], and 1 male

Tuxtla Gutierrez, 3300ft., 27 VI-8 VII, 1986 Figure 10 Cycnodi?rus (CycniXkrus) lJIrginiae, new spe (E Giesbert) 8additional paratypesfrom MEXICO as follows: 1 male, 3 females, Chiapas, Sumidero Can yon, 6-8 VII. 1986 (J.Wappes. Giesbert) [JW, EG]; 1 greenish·black. Head moderately short, punctation male, 1female, Chiapas, EI Surnidero Pargue, 23·24 moderately coarse, dense on vertex and occiput, pu­ VI, 1990 (J. Huether)[JH]; 1 male, Chiapas, EI bescence rille, erect, scattered; antennae slender, Sumidero, Miradorla Coyota, 24VJ, 1890(R TurnbOw) more than 3 times as long as body, scape less than 113 as [RT]; 1male, Oaxaca, 39.1kmSETeotitIan delCamino, longas thirdsegment, .....hieh is subequaltoorslightly 518m, 21 VII, 1987 (R. Anderson) [TAM). shorter than fourth, underside of segments 2 and 3 moderately fringed with dark hair·likesetae, twelfth REMARKS: CycrwderusvirginiaRappearstobeclosely segment curved Pronoturn nearly twice as long as related to the Brasilian C. tenuatus, and although wide, sides slightly rounded, widest before base, tao differing in integumental color, shares with that pering somewhat to apex; base and apex ',videly, species the diStillctlve grooved pronotal disk Of the shallowly impressed; disk transversely rugose, de· male. This species is similar to C. copei, but may be pressednearbase, with a longitudinal raisedcallus at easily separated by details of the male pronotal disk middle surrounded by moderately coarsely, densely such as the sinuate "waves and distinctly shmtet punctate, shallowimpressions, sidesmoderatelyswol· pubescence, and by the silky recumbent pubescence len and coarsely, irregularly punctate, with a fine, on the ~mora, the diStinct pubescence on the scutel sinuate, internally pubescent groove on each side lum, and the somewhat finer, less distinct elytral cunmgupwardfrom procoxalcavity, then fOIwaldto punctation. In addition, the integumental color is apical 1/6; pubescence fme, pale, erect, shorter than somewhatdifferent, with thebasiccolor morestrongly second antennal segment, sparse on lateral surface; metallic, and the femoral bases pale. Vol. 7, No.1- 2, March - June, 1993 53

Adults of this and C. copei fly in the summel Erytta with black area behind middle, not apical, months in southwestern Mexico. Specimens were with 2 pairs of pale oblique lines. Length 11- encountered by beatmg hvmg foliage m an area of 15mm. Honduras to Colombia . rncently cut brush...... C. plumicornis (Drury) Itis a pleasureto dedicate this species toVirginia 3.(1) Integumentmetallicblue-green. Pronotaldisksculp­ Ciesbertinappreciationforhermanyyearsofsupport tured Length 14 17mm. Guatemala to Panama and patience. C martyr Thomson Integument blackish, or mostly blackish. PronotaI disk convex 4 Genus Cosmlsoma Audinet-Serville 4 (3) Elytra hlack, with hurneralarea orange C.osta Rica, Panama C. mtlHiins n. sp. CosmisomaAudinet-Serville, 1834:19; Castelnau, 1840:434; Elytra Wack, or with white fasciae; humeral area White, 1855:214; Thomson, 1860.177, Thomson, never orange 5. 1864:169, 418; Burmeister,1865:170; Lacordaire, 1869.112, Bates, 1870.404, Bates, 1880.63, Zajciw, 5.(4) Elytra with a pair of short, oblique fasciae behind 1962:285; Monne & Magno, 1988: 1. Cosmosoma Gemminger & Halold, 1872.2984, GouneHe, mid'He consisting of whitish pubescence. 1911:103 (emend.) Pronotum cribrate-punctate, dull. Length 8- 10nlln. Panama and Colombia ...... C. rhaptos n. sp. DESCRIPTION: Form small to moderate-sized, Elytra ·,..,ithout fasciae. Pronotum reticulate-punc- subparallel or slightly tapering. Antennae ll-seg­ tate, somewhat shining. Length 6-9mm. Mexico mented, with tufts (lfJong hair-like setae on the fifth, to Panama G. retieuktta Bates andsometimesothersegments. Scutellumrounded at apex, notnotehed. Pronotumunarmed, slightlylonger thanwide, sidesrounded. Elytrasubparallelorslightly Cosmisoma madyr Thomson tapeIing, apices acuminate OI IOunded. Abdomen nonnallysegmented I .egssJender, clavate, with meso- Cosmis()ntamal tyl Thomson 1860.180 (FYPELOCALITY. and metafemora pedunculate clavate. Costa Rica). Cosmisoma martyra Bates 1872:189; Bates, 1880:63 (pI. 5, fig. 16), Bates,1885.311. TYPE SPECIES: Saperda scopulicornis Kirby (sub- Cosmisomanudicome Bates 1892:166 (pI. 7, fig. 2) (TYPE sequent designation by Castelnau, 1840). LOCALITY: Chiriqui, Panama) NEVl gYNOl'NMY.

COsmisoma IS prunarily South American with a total Of over 40 speciQs, six Of which occur ill the area DESCRIPTION. MALE. fum moderate-sized, eloo- ofthis study. The South American components ofthe gate, slightly tapering; integument shining dark me­ genus ha\'e been re\l iew ed and added to by Zajciw talliC green, appendages metalliC blUe-black, eaCh (1962) and Monne & Magno (1988). elytron with a moderately broad longitudinalvitta of Specimens of Cosmisoma have been taken diur­ golden pubescence. Head moderately elongate, shin· nally at flowers and on recently dead trunks and ing, moderately sparsely, distinctly punctate, pubes- branches of trees. cence fine, sparse, suberect; frons narrow WIth a distinct longitudinal median groove; antennae 2 to 2 1/2 times as long as body, scape subcylindrical, fmely Key to Ute Mexican and Central American scabrose punctate, base ,...,ith a small, distinctimpl'es species of Cosmisoma black hair-like setae on out.side at apical 2/5, fifth and sixth segmentswith a distinct, curved, apical spineon 1. underside, tenninal segment lOng, subequal to third. Elytra, pronotum, and ant@nnal tufts without or Pronotum longer than wide, sides unevenly rounded; ange pubescence 3. subapicalandsubbasalimPI essionsmodelately deep, disk uneven, wit.h a pair of large, feeble, sparsely 2.(1) Elytra WIth aplCaI area black andWIth a smgle pmr punctate swellings oneach SIde; surface shmmg,very ofpale oblique lines Length 8-9 5mm Nicaragua finely, densely punctate, '.'lith a narrow, impunctate, to Panama C. titania Bates 54 Insecta Mundi

median vitta, pubescence applessed, fIne, shOt t, in­ Cosmisoma mililatis, 'lew species distinct, intermixed with longer, coarser, erect setae; prosternum broadly impressed, moderately plicate DESCRIPTION: MALE: Formsmall, moderatelyslen­ ~ht~ ap~~eso- and metasternum moderately del; integument piceous to black, basal1f4oEely tIa at humeri, first 2 pairs oflegs, and bases ofmetafemora orange; tibiae and segments 3 W 5 ofantennae some- tapering; apices subacuminate; disk slightly, longitu- times reddish-brown Head coarsely, closely reticu- dinally deplessed on each side of middle, smface late-punctateonvertex; antennae abOut 1173 tImes as finely, denselypunctate:pubescencemoderatelydense, longas body, fifth segmentcurved, bearinga brush of black hair-like setae on apical half, segments 6 to 11 shOt t, modelately stlongly sellate. PlOnotumsliglltly longer than wide, sides broadly rounded at middle; pale pubescence, Legs moderately elongate; femora disk moderafuly opaque, lacking pubescence, shal- moderatelygraduallyclavate; metatarsiclothedwith lowly, coarselyreticulate-punctate,with an impunctate coarse, white hair-like setae, Length 14-17mm. band across base and apex; pro-, meso- and metastemum clothed with silky white pubescence FEMALE: Form SImiliU" to maIe. Aritennae abOut 1 1/3 which is more dense laterally on mesepisternum and times as long as body, segments 6 to 11 shorter, apical halfofmetepisternum Scutellum clothed with terminalsegmentmuch shorter than third. Abdomen subrecumbent white pubescence. Elytra about 2 1/2 somewhat mOte lObust, tellninal segment slightly tImes as long as Width across humen; apices nar­ 'longer than fourth. Length 15-16mm rowly, separately rounded; disk lacking pubescence, denselymicrosculptured,scabrous,puncturesobscure. RP-.NGE: Guatemala to Panama. Abdomen with tellninal stemite subequal to fom th, pubescence silky, white, denser laterally on first MATERIAL EXAMINED, COSTA RICA. 1 female, sternite Legs with femora abruptly clavate; Cartago prov., CATIE grounds, Turrialba, 28-31 V, metafemora slightly exceeding elytralapices. Length 1987 [EGI; 1 male, same locality, 14 V, 1985, 5,5-7.5mm, (J.ltChemsak) [DCB); 1female, same locality, 6 VI, 1986(Chemsak,Katsura,A&M, Michelbacher) [UCB]; FEMALE: Form somewhat more robust than male, PANAMA: 1 male, 1female, Canal Zone, Barre Colo- Antennae sligbtly longer tban body Abdomen more radoI., 20-23 VII, 1977 (R.B.& LB. KinIsey) [UCB]; 1 strongly convex, with terminal sternite longer than male, tremale, Bayano dist., vicinity Ipeti, 16'1, 1982 fourth, MetafemoranotattainingeIYtralapices. Length (E Giesbert) [EG]; 1 male, Bayano dist, 55km E 67.5mm. Chepo, 20 V, 1981 (Glesbert) LEG]; 1 male, 56km E Chepo, 20 V, 1981 g. Wappes) [JW); 2 males, 4 TYPES. Holotype male, allotype, [CAS] and 16 females, PanamaProv., lOkm NElLIano 1400',16-20 paratypes (7 males, 9 females)[EG] from PANAMA, V, 1984(Giesbet t) [EG] ,2 males, PanaImiplOV" CellO Panamaprov., CerroCampana, 2200ft., 18-19 V, 1984 Campana 2200', 18-19 V, 1984 (Giesbert) [EG). (E. Giesbert). 45 additional paratypes as follows: PANAMA: 1 male, CacM pIOv., El Valle de Anton REMARKS: This species differs from others in Cen 2500ft., 5 VI, 1945 (C.D. Michener) [UK]; 2 males. 1 tralAmericaby the metallicgreenintegument. Bates, female, Cocle prov., 4km SEIValle, 220Oft., 16V-2VI, in hm miginaldescIiption ore. nudimme, pointedout 1981 (Giesbert) [EG]; I male, CerroCampana, 2'700ft , that he was describing the species from a single 5 V, 1981 (Giesbert) [EG]; 2 males, 1 female, same individual, possibly a speeimen of Co martyr, from locality, on novlCrs of Glethra sp" 18 V, 1984 (H. which the antennal tufts had been abraded, since he Stockwell) S; 1 male, 3 females, same localit , 13- o serve no ot er erences. eave examme a 14 V, 1985 (F. Hovore) [FH]; 1 female, same locality, photographofthe type, and this appears tohavebeen 17-19V, 1987 (Giesbert) [EG]; 1female, same localit.y, the case. No other examples matching the description 2100ft., 29IV, 1992 (Giesbert)[EG]; lfemale, Panama of C. nudicome have been seen prov., Cerro Azul, 220Oft., 9V, 1984 (Giesbert) [EG); 3 males, 3 females, same locality, 24V, 1985 (Hovore) [PH], 8 males, same locality, 25 V, 1988 (Giesbet t) [EG); 4males, 2 females, same locality, 17-]8V, ]987 (Hovore) [FH]; 1 male, 1 female, Panama prov., 8- Vol. 7, No.1- 2, March - June, 1993 55

lOkm N EI Llano, 1200ft., 26 IV-4 V, 1992 and transversely plicate behind apex, base beming (Giesbert)[EG]; 1 female, Col6n prov.. Santa Rita orange pubescence; mesosternum finely silvery pu- RIdge, 12VI, 1976(D. Engleman) [u"W]. COSTARlCA: bescent; metasternurn mostlyorangepubescent. Elytra 1male, 1female, GuanacastepI"()V., Hacienda Comelco, less than 3times as long as width across humeri, sides 24km NW Canas, 50m., on blossoms of A1LOphyllus subparallel, feebly tapering; apices jointly rounded, sp., 20 V, 1971 (E.R. Heithaus) [UCB); 1 male, 1 sutural angles obtuse;

diskmodel8telyshining,pubescencelacking, COOl sely , 24km N Salama 4500', 25-31 V, 1989 (Giesbert) [EG], shallowly reticulate-punctate, with an impunctate 1 male, same data (Wappes) [JW]. ELSALVADOR: 2 bandacrossbaseand apex; prosternum unimpressed, males,Dept, Ahuachapan,BosqueElImposible, Cerro somewhat rugose near apex, fmely, densely punctate deLomadePaja,49Om,onflowersofAreliabarbinerius, and moderately densely pale pubescent on basal 374; 19 VI, 1979 (R.D, Cave) [UCB). PANAMA: 1 male, metasternum somewhat s.....ollen; pubescenee silky Panama prov" Gel'l'O Azul Summit, 15 22 Y, 1987 whitish at middle, concentratedlaterally into distinct (Giesbert) lEG). patches on mesepistemum and apical half of metepisternum Scutellum densely clothed with REMARKS' This small, obscure species is character- subrecumbent whitiSh pubeSCence. Elytra abOut 2 I/'2 Ized by the all-black color WIth the scutellum densely times as long as width across humeri; sides slightly white pubescent and parts of the antennae and legs curvedinwardat middle; apices narrowly, separately often partially dull orange. Specimens with paler rounded; disk laeking pubescence, densely appendages occur throughout the range. microsculptured, scabrous, puncturesobscure. Abdo- men with terminal sternite subequal W fourth, apex broadly subtruncate; basal sternites bearing patches Cosrnisoma rhapfos, new species ofsilkyWhItISh pubescence. LegswIthfemora abruptly (Fig, 11) clavate; metafemoraslightlyexeeedingelytralapiees. Length 6-9mm. DESCRIPTION: FEMALE: (See fig. 11) Form moder- ately small, parallel-sided; integument piceous to black, each elytron with a short, obliquely transverse, linear patch of white pubescence slightly behind mid

4000',20 VI, 1987 (Giesbert) [EG]; 1 male, Chiapas, I6km W Ocozocoautla, 20 VI, 1987 (Giesbert) [EG]; 1 male, Chiapas, 17kmWTuxtlaGutierrez3300', 21-25 VI, 1987 (Giesbert) [EG). GUATEMALA: 3 males, 19- Vol. 7, NO.1· 2, March· June, 1993 57

, long, erect setae. Length 8-9.5mm.

CATIE groun~, Turrialba, 28-31 V, 1987 (Giesbert) IEG1.

REMARKS: ThIS attractIve little speCIes IS fairly close in color andstructure to C. plumicomis, butdiffers by REMARKS: This species is easily separated by the the smaller size, elytral pattern, and more clavate blackintegumentwith anobliquely tIansverse, white, antennal scape. post-median pubescent mark on each elytron.

Cosmisoma titania Bates Genus Ischionodonta Chevrolat, new status .

Cosmlsoma t1tanla Bates 1870'407 (TYPE [J ICALI'l'Y' Rhopalilphora(Tschiiiiiildilnlo) ( $evroJat 1859·58, ( \ounelle Chontales, Mcaragua); Bates, 1880:63. 1911:98. ', BrochyrhopalaBurmeister, 1865:171;Lacordaire. 1869:41; Berg, 1886:239(Type species: Brochyrlwpalaaenescens DESCRIPTION: MALE: Form moderatelY small, elon Burmeister, 1865, present designation), gate; integumentpiceousblack, with diskofpronotum (except base andapex), apical 1/3 offifth andbasal 415 of DESCRIPTION: Form moderately elongate, body sixth antennaI segments, and basal 3/5 of elyfra subdepressed, sides nearly parallel; integumentdark orange with orange pubescence, the apical 215 black with metallic reflections. Head moderately short; pubeseent ',vith an oblique, curved white pubeseent antennal tubercles acute; antennae elongate, fourth line near middle ofblack area Headslightly elongate, segment more or less subequal to third. Pronotum moderately sparsely pubescent, with numerous erect se11le; antennae about 1 1/2 times as long as body, scape abruptlyclavate, segments3 to 5 long, carinate, fifth segment longest, curved, remaining segments shorter, third segment bearing a moderately large meso- andmetafemora abruptlyclavate, metafemoral blUsh of long black hair-like setae, fifth segment peduneles and metatibiaefeebly W strongly asperate bearing a large brush over apical 7/8, with basal half black, apicalhalfcomposedoforangesetae, scapewith 58 Insecta Mundi

TYPE SPECIES. RhupuluphOlu bmsiliensis pubescence, meso- and metasternum fInely, densely Chevrolat, 1859 (by present designation) punctate, pubescence pale Elytra slightly more than A number of conSIstent structural characters tWICe as long as WIdth across humeri; apices narrowly separate !schionodorlta from Rhopolaphoffi and ap- rounded; dil>k feebly granulate near base; punctures pear to warrant full generiC status. lschi01wdOnta IS hne, very dense; pubescence goWen, very hne and primarily South l'.merican in distribution with three short, subrecumbent. Abdomen ...... ith terminal stern species occuring in Mexico and Central America. ite subequal to fourth, apex widely subtruncate; pu- bescence pale. Legs with aspetiteson posteriOl edge of Key to the Mexican and Central American metatibiae well developed, spinose Length 1O-12mm species of Ischionodonta FEl\L\LE: Form similar to male. hltennae exceeding 1. Underside and femoral clubs black with metallic elytralapices by about 4 segments, eleventh segment blue reflections; metatibiae with spinose asperi shOlter than third. Abdomen with terminal sternite ti@s on post@rior @dgl:l. Nicaragua to Panama.... longer than fmIrth, apex narrowly rounded Length ... 1. sernpes (Bates) 12mm. IInderside and femoral clubs piceous or dull metal- lic; metatlblae WIth aspentes on postenor edge RANGE: Nicaragua to Panama. not prominent .... . 2 MATERIAL EXAMINED: PANAMA: 1female, Canal 2.(1) Head and pronotum immaculate reddish orange; Zone, Barro Colorado I., 20 VII, 1977 elytradarkmetallicgreenish. NicaraguatoSouth America 1. versicolor (Chevrolat) Head black, pronotum reddish orange, often with a median black vitta; elytra dull blackish with metallic reflections. Mexico (Jabsco, Chiapas) ...... 1. mexiCUflU n. sp. REMARKS: This speeies may be easily distinguished by the well developed asperites on the posterior edge of the metatIbIae, by the quadrate reddish-orange Ischionodonta serripes (Bates), pronotum with a wide black median vitta, the rela new combination tively short elytra, andthe metallic blue reflections of themetasternum, metepisterna, abdomen, andfemo- Rhopalophoraserripes Bates 1872:188 (TYPELOCALITY: ral clubs. Chontales, Niearag\la); Bates, 1330:63. Rh~;~;\t:;a (lschiouodouta) serripes; Aurivillius, Jschionodonta versicolor (Chevrolat), new combination

Rhopalophora versicolor Chevrolat, 1859:62 (TYPE LO- DESCRIPTION. MALE. Form moderately ShOlt, CALITY. "Nou'I.'tllle Grenade" [Colombia)); Bates, moderately slender; integument blackish, prothorax 1872: 188, Bates 1880:63; Bates, 1885:310. reddish orangewith a wide, dorsal, median hlackvitfa Rhopalophora (!~chionodonta) versicolor, A\lrivilli\ls, from base to apex, elytra blackish with dull metallic 1912:432 reflectIOns, underSIde andfemoraIclubsdark metallIc blue. Head finely, densely punctate; frons sparsely, finely pubescent; antennaeelongate, exceeding elytral DESCRIPTION: MALE: Form moderate sized; in­ apiees hy ahout 6 segments, scape asperate-punctate, tegument black with dull metallic reflections, head short, about twice as long as wide, third segment and prothorax bright reddish orange, elytra dark feebly asperate, slightly shorterthanfourth, eleventh metanic greenISh. Head mOderately finely, densely segmentlonger than third. Pronotumas wide as long, punctate; frons with a fme, longitudinal, median sides strongly rounded, widest near middle, trans­ impression, pubescence sparse, recumbent; antennae versely impressed near base and apex; disk with 4 slender, elongate, exceeding ely tr al apices by about 5 feeble swellings; surface opaque, inconspicuously 1/2 to 6 segments, scape short, densely asperate­ asperate-punctate; pubescence absent orverysparse; punctilfe, segments3 fii 5suhequal, eleventh segment prosternumwith apical 1/3 transverselyrugose, basal subequal to third. Pronotum somewhat longer than 2/3 densely punctate, clothed with pale, appressed WIde, SIdes broadly rounded, WIdest behmd mIddle; Vol. 7, No.1 - 2, March - June, 1993 59

baseandapexbroadly, shallow ly impressed; diskwith REMARKS: This species may be recognized by the 4feeble swellings: surface opaque, moderately, incon- immaculate bright red-orange head and prothorax, spicuously scabrose-punctate, very inconspicuously and dull metallic greenish elytra. Itappears to be the pub@scent; meso· and metasternum finely, densely most commonly collected speci@s of Ischionodonta in punctate and pubescent. Elytra abOut 2 172 times as Central AIrierica long as','/idth acrosshumeri, slightly elevated atbase; Blackwelder( 1946) JistedRJwpaloplwraaeneseens apices narrowly rounded; disk finely, densely punc- Burmeister,R. semirubraBurmeisterandR. platensis tate, pubescence fIne, shm t, golden. Abdomen with Chevrornt as synonyms ofR. Vet sicolUl. Subsequent terminal sternite subequal to fourth. apex slightly work by Zajciw and Ruffinelli (1962) recognized flexed downward, tufted, moderately WIdely plalensisandsemirubraasvalidspeCIes WIth aenescens subtruncate; fmely, densely punctate andpubescent. a synonym ofplatensis. Legs with metatibial asperities feebly developed. Length 1O-12.5mm. Iscbionodonta mexicana, new species FEMALE: Form similar to male. Antennae exceeding @lytral apices by about 4 segments, eleventh segment DESCRIPTION. MALE. Forni mooerate sized, in- much shorter than thrrd. Abdomen WIth termmal tegument black with dullmetallic reflections, protho­ sternite longer than fourth, apex narrowly rounded. lax reddish-mange, usually marked at mid

and elytra, and by the presence of sparse, golden,

Muxbalia, new genus

DESCRIPTION: Form linear, subdepressed, slender.

, third. Scutellum small, oblong, apically rounded. 1 with surface uneven. Elytrasubdepressed, morethan 3 112 times as lOng as basal width; apiCes obliquely emarginate-truncate. Abdomen normallysegmented. Figure 12. Muxbalia monzoni, new species, male. Legs very slender, elongate, with meso- and metafemOI a gmdually clavate. Muxbalia monzoni, new species (Fig. 12) TYPE SPECIES· MuXbalio monzoni Giesbert and Chemsak. DESCRIPTION: MALE: (See fig. 12) Form moder- MuxbaliaeroselyresemblesRhopalopJwrain size, ately small, subparallel, slender; integument blaek, form and color, but may be immediately separated by with a vague metallic sheen on elytral margins and the gradually clavate meso- and metafemora, which legs, elytrapiceous, oftenwith avaguemetallicpurple in all Rhnpalnpbora are abmptJy pedunculate-clav- sheen, pronotuID occasionally red with hase and apex ate. Although the femoral form is anomalous, in all black. Head moderately short, nearly glabrous; frons other morphological eharaeteristiesMuxbalia is typi wide, moderatelY punctate; vertex opaque; antennae calofthe tribe, andis therefore provisionally assigned slender, exceeding elytral apices by about 2 1/2 seg- to the RhopalophOIini. A single species is presently ments, basal segments VeIy sparsely ciliate, scape known subcylindrical, basal half shallowly, longitudinally canaliculate, thirdsegmentlongerthanfourth, shorter Vol. 7, No.1· 2, March - June, 1993 61 than ftfth, segments 10 and 11 eaeh feebly areuate, Ackilowledgeilielits slightly longer than third. Pronotum longer than We would like to thank the authorities of the British Museum (Natural History), London for the loan oftypes in theircare Additionally, the following institutions and individuals are gratefully acknowl· edgedfor makingmaterialavailablefor studY: .Ameri

Museum, University of California, Davis; California Academy of Sciences, San FraneiS(~(); Canadian Na- tional Collection, Ottawa; Estaci6n de Biologia Chame1a, Mexico; COrnell University, Ithaca; ESSig Museum of Entomology, University of California, Berkeley; FieldMuseum of NaturalHistory, Chicago; Los Angeles County Museum of Natural History; Museum of Comparative Zoology, Cambridge; Na­ tional MuseumofNaturalSeienees, Ottawa; Stanford University, PaloAlto; TexasA&M University, College dull, microalutaceous, punctures moderately coarse, StatiOn; UnitQd StatQs MusQum Of Natural History; moderately denre, irregular; very sparse pubescence Snow Entomological Museum, Lawrence; Joseph near base and apex fine, pale, incffilspicuous. AbOO· Beierl, James Cope, Frank Hovore, Henry Howden, Jeff Huether, Gale Nelson, Richard Penrose, Keve concavewhenvIewe ram eSIewinpexrourFe ; Ribardo, HenryStockwell,RobertTurnbow,andJames su~acQ finQI~ oonsQl)'punc~atQ; ~bQ8CQnCQ :YQry Wappes ally clavate; metatarsi with first segmentlonger than second and third together. Length 9-12.5mm. Literature Cited Audinet..senille, J. G. 1834. Nouvelle dassifiea- FEMALE: Form SImilar to male. Aritennae nearly tion de la famille des longicornes. Ann. Soc. reaehing elytral apices, segments 10 and 11 slightly Entom 01 France 3'5-110 shorter than third. Prosternum somewhat rugose, without punctate patches Abdomen with terminal Aurivillius, C. 1912. Goleopterorum catalogus, pars fe;:ite convex when viewed from the side I .ength 8- 39, Cerambycidae:Cerambyeinae, pp. 1574. VI: Junk, Berlin.

TYPES: Holotype male, allotype [CASl, and 55 Bates, H. W. 1870. Contributions to an insect fauna paratypes(30 males, 25 females) flOm GUATEMALA, Of the AmazonValley (COleoptera, CerambyCIdae) Dept. de Guatemala, Santa Catarina de Pinula, San Trans. Entomol. Soc. London, 1870:243-335,391- .Jorge MllxbaJ, 3 X, 1990 (E Giesbetf) [lICB, EG); 62 444. additional paratypQs (34 rnalQs, 28 {QrnaIQs), sarnQ locality, 1-8 XI, 1990 (J.Monz6n) [JM, UDV]. Bates, H. W. 1872. On the longicorn Coleoptera of CbontaJes, Nicaragua Trans Entomol SOC LOn- REMARKS: This species has been collected from the don, 1872: 163·238. flowers of several different Compositae at a single location. In the type series of119 specimens, 10 males Bates, H. W. 1874. Supplement to the longicorn and 5 females are bIcolored. It IS a pleasure to name Coleoptera of Chontalia, Nicaragua. Trans. thisspeciesfor JoseMonz6n, a friend andenthusiastic Entomol Soc London, 1874'219-235 studentofentomology atthe UniversidaddelValle in Guatemala, in appreciation fm his assistance in col- Bates, HW 1879·1885. BiOlOgm CQfitrali-ArnQri- lecting the type series. cana, Insecta, Coleoptera, vol. 5, Longicornia, 436 pp. 62 Insecta Mundi

Bates, II. VI. 1892. Additions to the Longieomia of Craighead, F. C. 1923. NOI th Ameliean eetambyeid Mexico and Central America with remarks on laIVae. Can. Dep. Agric. Tech. Bull. 27 (n.s.), 239 some ofthe previously recorded species. With an pp., figs 1-8. introduction byFredrick DuCaneGodman,FRS Trans. Entomol. SOC. LOndon, Part II, 1892: 143­ Urury, U. 1782. lllustatlOns of Natural 183. History...Exotic Insects... vo1.3, 76 pp. 50 pIs.

Berg, F. G. C. 1886. Notas sinonimicas acelca de Fattig,P.W.1947. TheCerambycidaeOllong-horned algunoscerambicidosdelafauna Argentina Anal beetles of Georgia Emory IIniv Mus Bull no 5 SOC. Clent. Argentma, 21:234-240. 48PP-

Blackwelder, R. E. 1946. Checklistofthe coleopter­ Gemminger,M.,& E. von Harold. 1872. Catalogus ous insects ofMexico, CentIal Amelica, the 'Nest coleoptelolum hucusgue descIiptolum Indies. and South America. PtA. U.S. Nat. Mus. synonymicus et systematicus, 9:2669-2988. Bull. 185.551-763. Gosling, D. C. L. 1973. An annotatgd list of thg Burmeister, H. C. C. 1865. LOngIcornla Argentma­ Cerambycidae of Michigan (Coleoptera) Part I, Systematische Ubersicht der Bockkafer, der Ia introduction and the subfamilies Parandrinae, Plata -Staaten. Stett. Entomol. Zeitung, 26: 156­ Prioninae, Spondylinae, Aseminae and 181. Cerambycinae. Great Lakes EnWmol. 6.65-84.

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