CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM.
III. OXYTELINE GROUP
LEE H. HERMAN
BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY NUMBER 265 NEW YORK : 2001 Recent issues of the Bulletin may be purchased from the Museum. Lists of back issues of the Bulletin, Novitates, and Anthropological Papers published during the last five years are available at World Wide Web site http://nimidi.amnh.org. Or address mail orders to: American Museum of Natural History Library, Central Park West at 79th St., New York, N.Y. 10024. TEL: (212) 769- 5545. FAX: (212) 769-5009. E-MAIL: [email protected] CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM.
III. OXYTELINE GROUP
LEE H. HERMAN Curator, Division of Invertebrate Zoology American Museum of Natural History
BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Number 265, pages 1067–1806 Issued July 18, 2001 (Parts I–VII)
Copyright © American Museum of Natural History 2001 ISSN 0003-0090
CATALOG OF THE STAPHYLINIDAE, PARTS I–VII
PART I, pages 1–650 Introduction Brief History of Taxonomic Studies of the Staphylinidae Research History • Biographical Sketches Overview of Classification Omaliine Group Dasycerinae • Empelinae • Glypholomatinae • Micropeplinae • Microsilphinae • Neophoninae • Omaliinae • Proteininae • Protopselaphinae
PART II, pages 651–1066 Tachyporine Group Habrocerinae • Olisthaerinae • Phloeocharinae • Tachyporinae • Trichophyinae
PART III, pages 1067–1806 Oxyteline Group Apateticinae • Osoriinae • Oxytelinae • Piestinae • Trigonurinae
PART IV, pages 1807–2440 Staphylinine Group Euaesthetinae • Leptotyphlinae • Megalopsidiinae • Oxyporinae • Pseudopsinae • Solieriinae • Steninae
PART V, pages 2441–3020 Staphylinine Group, continued Staphylininae Diochini • Maorothiini • Othiini • Platyprosopini • Staphylinini (Amblyopinina • Hyptiomina • Anisolinina • Philonthina)
PART VI, pages 3021–3840 Staphylinine Group, continued Staphylininae, continued Staphylinini (Quediina • Staphylinina • Tanygnathinina • Xanthopygina • Incertae Sedis) • Xantholinini Staphylinidae: Subfamily Incertae Sedis Protactinae†
PART VII, pages 3841–4218 Bibliography Index
iii CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM
ABSTRACT This catalog (published in seven parts, all released on the same day) is based on only the published lit- erature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylin- inae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or descrip- tion are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or syn- onyms are listed for each valid name. Annotated subsequent references are presented for all names. Dis- tributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Alesˇ Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies.
Address correspondence to: Lee H. Herman, Curator, Division of Invertebrates, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA ([email protected]). Please send comments, corrections, or updates to either the postal or e-mail address. Reprints of new publications would be helpful and appreciated.
iv CONTENTS
PART III OXYTELINE GROUP
Oxyteline Group ...... 1067 Apateticinae ...... 1067 Osoriinae ...... 1071 Eleusinini ...... 1072 Leptochirini ...... 1094 Osoriini ...... 1145 Thoracophorini ...... 1230 Clavilispinina ...... 1231 Glyptomina ...... 1239 Lispinina ...... 1250 Thoracophorina ...... 1293 Osoriinae: Tribe Incertae Sedis ...... 1305 Oxytelinae ...... 1307 Coprophilini ...... 1308 Deleasterini ...... 1321 Oxytelini ...... 1332 Thinobiini ...... 1491 Oxytelinae: Tribe Incertae Sedis ...... 1778 Piestinae ...... 1781 Trigonurinae ...... 1803
PARTS I, II, IV–VII Introduction, History, and Biographical Sketches ...... Part I Omaliine Group ...... Part I Tachyporine Group ...... Part II Staphylinine Group ...... Parts IV–VI Staphylinidae: Subfamily Incertae Sedis ...... Part VI Protactinae† ...... Part VI Bibliography ...... Part VII Index ...... Part VII
v
APATETICINAE [2 genera; 24 species; Oriental and Palaearctic Regions] Apateticinae Fauvel, 1895. — Newton and Thayer, 1992: 55 (summary of current classification). — Lawrence and Newton, 1995: 827 (discussion). — M. Hansen, 1996: 33, 36 (relatively basal position within superfamily; listed in Oxyteline group of subfamilies but should probably be omitted from family). Apateticae Fauvel, 1895: 190 (genera included: Apatetica, Trygaeus). TYPE GENUS: Apa- tetica (see Newton and Thayer, 1992: 55). — M. Hansen, 1997: 78, 79, 80, 135, 166, 180 (characters supporting clade; autapomorphic features; description; notes). Apateticini Fauvel, 1895. — Eichelbaum, 1909: 115 (list of genera with some references; generic synonyms; number of genera; number of species in each genus; general distribution of genera). — Newton, 1982a: 14 (larval characters).
Apatetica [19 species; Oriental and Palaearctic Regions] Apatetica Westwood, 1848: 86 (species included: lebioides). TYPE SPECIES: Apatetica lebioides Westwood, fixed by monotypy. — Fauvel, 1895: 190 (characters; notes). — Bernhauer and Schubert, 1910: 4 (world catalog; 12 species). — Winkler, 1925: 323 (catalog for Palaearctic region). — Cameron, 1925: 1 (catalog of Indian species). — Cameron, 1930: 26 (characters; key to species of British India). — Scheerpeltz, 1933: 992 (world catalog supplement). — Jansson, 1947: 1 (characters; discussion). — Blackwelder, 1952: 411 (type species: lebioides). — Abdullah and Qadri, 1970: 115 (cited as Apatelica; key to species of East Pakistan). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Shibata, 1976: 76 (checklist of species of Japan). — Lawrence and Newton, 1995: 827 (Apateticinae). Trygaeus Sharp, 1874a: 420 (species included: princeps). TYPE SPECIES: Trygaeus princeps Sharp, fixed by monotypy. — Duvivier, 1883: 203 (catalog; cited as Trigaeus). — Lewis, 1893a: 248 (synonym of Apatetica). — Fauvel, 1895: 190 (synonym of Apatetica). — Bernhauer and Schubert, 1910: 4 (synonym of Apatetica). — Cameron, 1930: 26 (synonym of Apatetica). — Jansson, 1947: 5 (characters; may be valid genus or synonym of Nodynus). — Blackwelder, 1952: 414 (type species: princeps). Idiocheila Frivaldszky, 1883: 135 (species included: spinipennis). TYPE SPECIES: Idiocheila spinipennis Frivaldszky, fixed by monotypy. — Fauvel, 1895: 190 (synonym of Apatetica). — Bernhauer and Schubert, 1910: 4 (synonym of Apatetica). — Cameron, 1930: 26 (synonym of Apatetica). — Blackwelder, 1952: 412 (type species: spinipennis). birmana Cameron, 1930: 33 (Apatetica; Type locality: Upper Burma: Seinghku Valley, alt. 5000 feet). — Scheerpeltz, 1933: 992 (Apatetica; catalog). DISTRIBUTION: Myanmar.
1067 1068 APATETICINAE: APATETICA Bulletin AMNH birmanica Jansson, 1947: 7 (Apatetica; Type locality: Kambaiti at 7000 feet). DISTRIBUTION: Myanmar. brunnipes Ritsema, 1879: 46 (Apatetica; Type locality: Sumatra). — Fauvel, 1895: 192 (Apatetica; characters; Sumatra; Borneo). — Bernhauer and Schubert, 1910: 4 (Apatetica; catalog; Sumatra, Borneo). — Cameron, 1928d: 90 (Apatetica; Sumatra). DISTRIBUTION: Sumatra, Borneo. caeruleipennis Cameron, 1930: 32 (Apatetica; Type locality: Manipur). — Scheerpeltz, 1933: 992 (Apatetica; catalog). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). DISTRIBUTION: India, Bangladesh. callosa Jansson, 1947: 10 (Apatetica; Type locality: Kambaiti, 6000 feet). DISTRIBUTION: Myanmar. indica Cameron, 1930: 29 (Apatetica; Type locality: Bhutan). — Scheerpeltz, 1933: 992 (Apatetica; catalog). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). DISTRIBUTION: Bhutan, Bangladesh. intermedia Cameron, 1930: 30 (Apatetica; Type locality: Burma: Ruby Mines; Bengal). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). DISTRIBUTION: Myanmar, India, Bangladesh. javanica Sharp, 1892: 61 (Trygaeus; Type locality: Mount Poentjak, West Java). — Fauvel, 1895: 194 (Apatetica; characters; Bengal; Burma; Java). — Bernhauer and Schubert, 1910: 4 (Apatetica; catalog; Java; East Indies). — Cameron, 1930: 29 (Apatetica; characters; Assam; Bengal). — Scheerpeltz, 1933: 993 (Apatetica; catalog). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). DISTRIBUTION: India, Bangladesh, Java, Sumatra. sumatrana Cameron, 1930d: 326 (Apatetica; variety of javanica; Type locality: Gunung Singgalang). — Scheerpeltz, 1933: 993 (Apatetica; variety of javanica). laevicollis Fauvel, 1904b: 86 (Apatetica; Type locality: Tonkin central: Chiem-Hoa). — Bernhauer and Schubert, 1910: 4 (Apatetica; catalog; Tonkin). DISTRIBUTION: Vietnam. lebioides Westwood, 1848: 86 (Apatetica; Type locality: Himalayan region). — Fauvel, 1895: 193 (Apatetica; characters; Himalaya; Sikkim). — Bernhauer and Schubert, 1910: 4 (Apatetica; catalog; East Indies). — Cameron, 1930: 31 (Apatetica; characters; Sikkim). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). DISTRIBUTION: India, Bangladesh. nitiduloides Westwood, 1864: 11 (Apatetica; Type locality: Java). — Fauvel, 1895: 192 (Apatetica; characters; Java). — Bernhauer and Schubert, 1910: 5 (Apatetica; catalog; Java). DISTRIBUTION: Java. princeps Sharp, 1874a: 421 (Trygaeus; Type locality: Maiyasan Temple, Hiogo). — Sharp, 1889: 467 (Trygaeus; notes; Japan). — Lewis, 1893a: 248 (Trygaeus; collecting notes). — Fauvel, 1895: 194 (Apatetica; characters; Japan). — Bernhauer and Schubert, 1910: 5 (Apatetica; catalog; Japan). — Nakane, 1963a: 81 (Apatetica; characters; habitus photograph; Japan). DISTRIBUTION: Japan. Bulletin AMNH APATETICINAE: APATETICA 1069 rotundicollis Fauvel, 1904b: 85 (Apatetica; Type locality: Khasia Hills). — Bernhauer and Schubert, 1910: 5 (Apatetica; catalog; East Indies). — Cameron, 1930: 28 (Apatetica; characters; Khasia Hills). — Scheerpeltz, 1933: 993 (Apatetica; catalog). — Abdullah and Qadri, 1970: 115 (Apatetica; cited as rotondicollis; characters; East Pakistan). DISTRIBUTION: Bangladesh. semiviolacea Fauvel, 1904b: 85 (Apatetica; Type locality: Java: Mont Ardjuno, 1000 m). — Bernhauer and Schubert, 1910: 5 (Apatetica; catalog; Java). DISTRIBUTION: Java. siamensis Lewis, 1893a: 248 (Apatetica; Type locality: Siam). — Fauvel, 1895: 192 (Apatetica; characters; Siam). — Bernhauer and Schubert, 1910: 5 (Apatetica; catalog; Siam). DISTRIBUTION: Thailand. sikkimi Fauvel, 1895: 193 (Apatetica; Type locality: Sikkim, Darjeeling). — Bernhauer and Schubert, 1910: 5 (Apatetica; catalog; East Indies). — Cameron, 1930: 28 (Apatetica; characters; Sikkim). — Scheerpeltz, 1933: 993 (Apatetica; catalog). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). DISTRIBUTION: India, Bangladesh. sparsicollis Fauvel, 1904b: 86 (Apatetica; Type locality: Lombock: Sapit, 700 m). — Bernhauer and Schubert, 1910: 5 (Apatetica; catalog; East Indies). DISTRIBUTION: Indonesia. spinipennis Frivaldszky, 1883: 137 (Idiocheila; Type locality: Borneo: m. Matang). — Fauvel, 1895: 192 (Apatetica; synonym of brunnipes). — Bernhauer and Schubert, 1910: 4 (Apatetica; synonym of brunneipes). — Hammond, 1984: 196 (Apatetica; valid species; Borneo). DISTRIBUTION: Borneo. sumatrana Cameron, 1930, see: javanica Sharp, 1892. viridipennis Fauvel, 1895: 192 (Apatetica; Type locality: Sikkim, Darjeeling). — Bernhauer and Schubert, 1910: 5 (Apatetica; catalog; East Indies). — Cameron, 1930: 31 (Apatetica; characters; Sikkim; Assam; Burma). — Scheerpeltz, 1933: 993 (Apatetica; catalog). — Jansson, 1947: 6 (Apatetica; characters; Burma). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). DISTRIBUTION: India, Bangladesh, Myanmar.
Nodynus [5 species; Oriental and Palaearctic Regions] Nodynus C. Waterhouse, 1876: 12 (species included: nitidus). TYPE SPECIES: Nodynus nitidus C. Waterhouse, fixed by monotypy. — Bernhauer and Schubert, 1910: 5 (world catalog; 2 species). — Winkler, 1925: 323 (catalog for Palaearctic region). — Cameron, 1925: 1 (catalog of Indian species). — Cameron, 1930: 33 (characters). — Scheerpeltz, 1933: 993 (world catalog supplement). — Jansson, 1947: 1 (characters; discussion). — Blackwelder, 1952: 413 (type species: nitidus). — Crowson and Ellis, 1969: 168 (notes on classification). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Shibata, 1976: 77 (checklist of species of Japan). — Lawrence and Newton, 1995: 827 (Apateticinae). 1070 APATETICINAE: NODYNUS Bulletin AMNH leucofasciatus Lewis, 1879: 460 (Nodynus; Type locality: Yamato). — Sharp, 1889: 467 (Nodynus; notes; Japan). — Bernhauer and Schubert, 1910: 5 (Nodynus; catalog; Japan). — Sharp and Muir, 1912: 501 (Nodynus; characters of male genitalia). — Nakane, 1963a: 81 (Nodynus; characters; habitus photograph; Japan). DISTRIBUTION: Japan. nitidus C. Waterhouse, 1876: 13 (Nodynus; Type locality: E. Indies). — Bernhauer and Schubert, 1910: 5 (Nodynus; catalog; East Indies). — Cameron, 1930: 34 (Nodynus; characters; East Indies). — Scheerpeltz, 1933: 993 (Nodynus; catalog). DISTRIBUTION: “East Indies”. rubrofasciatus Jansson, 1947: 11 (Nodynus; Type locality: Kambaiti, 7000 feet). DISTRIBUTION: Myanmar. silphoides Jansson, 1947: 14 (Nodynus; Type locality: Kambaiti, 7000 feet). DISTRIBUTION: Myanmar. unicolor Jansson, 1947: 13 (Nodynus; Type locality: Kambaiti, 7000 feet). DISTRIBUTION: Myanmar. OSORIINAE [105 genera; 2058 species; All Regions except the Antarctic] Osoriinae Erichson, 1839. — Portevin, 1929: 408 (characters). — Scheerpeltz, 1933: 1127 (synonym of Osoriini). — Blackwelder, 1942: 76, 78 (characters; key to tribes). — Blackwelder, 1943: 119 (key to genera of West Indies). — Blackwelder, 1944: 107 (checklist of species from Mexico, Central and South America, and the West Indies). — Fagel, 1955: 7 (key to genera of Ethiopian region). — Crowson, 1955: 33, 38 (adult and larval characters in key; relationships). — Hatch, 1957: 49, 243 (characters; key to subtribes and genera of Pacific Northwest). — Fagel, 1959: 5 (key to African genera). — Coiffait, 1960a: 515 (biogeography of eyeless species). — Arnett, 1963: 242, 262 (key to North American genera; characters; list of North American genera). — Lohse, 1964: 19 (characters; key to central European species). — Moore, 1964a: 88 (characters; notes). — Kasule, 1966: 265 (larval characters). — Coiffait and Saiz, 1968: 414 (list of Chilean species). — Abdullah and Qadri, 1970: 114 (cited as Piestinae; key to species of West Pakistan). — Abdullah and Qadri, 1970: 116 (key to species of East and West Pakistan). — Moore and Legner, 1974c: 115 (characters; key to genera of America north of Mexico). — Moore and Legner, 1974g: 555 (key to genera of America north of Mexico). — Moore and Legner, 1975: 172 (catalog; North America north of Mexico). — Topp, 1978: 306 (larval characters). — Coiffait, 1979: 7 (systematic position of subfamily; biological and biogeographic notes; key to genera of Madagascar species; species of Madagascar). — Burakowski, Mroczkowski, and Stefan´ska, 1979: 7 (catalog; Poland). — Moore and Legner, 1979: 152 (characters; tabular key to genera of North America north of Mexico). — McColl, 1982 (checklist of taxa of New Zealand; historical review; phylogenetic discussion; ecology; notes on larvae; characters; keys to genera and species of New Zealand). — Watt and McColl, 1982: 13, 19 (characters; notes on New Zealand taxa). — Newton, 1982a: 14 (larval characters). — Outerelo and Gamarra, 1985: 77 (key to genera of Iberian Peninsula). — Naomi, 1986: 33 (characters; key to genera of Japan). — Newton, 1990: 91 (key to genera of Australia). — Frank, 1991: 345 (larval characters). — Newton and Thayer, 1992: 13, 59 (summary of current classification; subfamily of Staphy- linidae; discussion). — Lawrence and Newton, 1995: 828 (discussion). — Sparacio, 1995: 135 (characters; species of Sicily). — Klimaszewski, Newton, and Thayer, 1996: 150 (review of subfamily for New Zealand; notes; list of New Zealand genera; number of New Zealand species). — M. Hansen, 1996: 36 (listed in Oxyteline group of subfamilies). — Downie and Arnett, 1996: 428 (characters; key to species of Northeastern North America). — M. Hansen, 1997: 125, 176, 177, 179 (autapomorphic features; key adult and larval characters; notes). — Drugmand, 1998a (list of types in l’Institut royal des Sciences naturelles de Belgique). — Cho and Ahn, 1999: 175 (review of subfamily for Korea; key to Korean tribes, genera, and species). — Newton, Thayer, Ashe, and Chandler, 2000: 320, 376 (key to Nearctic tribes, subtribes, and genera; characters; notes). — Klimaszewski, 2000: 21, 55 (characters; notes on classification, natural history, distribution, and key to tribes in Canada). 1071 1072 OSORIINAE: ELEUSININI Bulletin AMNH
Tribe Eleusinini Eleusinini Sharp, 1887. — Eichelbaum, 1909: 111 (list of genera with some references; generic synonyms; number of genera; number of species in each genus; general distribution of genera). — Cameron, 1930: 24 (characters). — Newton and Thayer, 1992: 59 (tribe of Osoriinae). — Klimaszewski, Newton, and Thayer, 1996: 150 (list of New Zealand genera; number of New Zealand species). — Newton, Thayer, Ashe, and Chandler, 2000: 376 (characters; notes). Eleusinina Sharp, 1887: 728 (genera included: Eleusis, Eumalus). TYPE GENUS: Eleusis (see Newton and Thayer, 1992: 59). — Jakobson, 1908: 446, 466 (characters; list of species of Russia and western Europe). Eleusii Bernhauer and Schubert, 1910: 10 (world catalog; Eumalus, Triga, Eleusis). — Blackwelder, 1942: 79 (characters). — Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies). — Newton and Thayer, 1992: 59 (synonym of Eleusinini). Eleusiinae: — Watt and McColl, 1982: 18, 19 (characters; valid subfamily; notes on New Zealand taxa).
Eleusis [200 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions] Eleusis Laporte, 1835: 131 (species included: tibialis). TYPE SPECIES: Eleusis tibialis Laporte, fixed by monotypy. — Laporte, 1840: 186 (characters). — Lacordaire, 1854: 127 (characters; notes; list of species). — Fauvel, 1865: 35 (synonym of Isomalus). — Gemminger and Harold, 1868: 675 (catalog). — LeConte, 1877: 249 (characters; notes). — Fauvel, 1878d: 180 [= 1878e: 16] (key to North American species). — Fauvel, 1878f: 206 (characters; key to New Guinea species). — Duvivier, 1883: 203 (catalog). — Sharp, 1887: 728 (characters; Mexican and Central American species). — Bernhauer and Schubert, 1910: 10 (world catalog; 75 species). — Leng, 1920: 93 (catalog of North American species). — Cameron, 1921a: 350, 398 (characters; key to Singapore species; catalog). — Winkler, 1925: 323 (catalog for Palaearctic region). — Cameron, 1925: 2 (catalog of Indian species). — Bernhauer, 1926d: 246 (key to species of Philippines). — Cameron, 1930: 74, 453 (characters; key to British Indian species). — Scheerpeltz, 1933: 996 (world catalog supplement). — Blackwelder, 1943: 155 (characters; type species: tibialis). — Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 145 (type species: tibialis). — Steel, 1950a: 215 (characters in key). — Scheerpeltz, 1956a: 488 (key to African species). — Scheerpeltz, 1961g: 239 (key to species of Madagascar). — Abdullah and Qadri, 1970: 116, 117 (key to species of East and West Pakistan). — Shibata, 1973c: 24 (checklist of species of Taiwan). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Moore and Legner, 1974b: 80 (characters; notes). — Moore and Legner, 1975: 167 (catalog; North America north of Mexico). Bulletin AMNH OSORIINAE: ELEUSIS 1073
— Shibata, 1976: 85 (checklist of species of Japan). — Moore and Legner, 1979: 145 (characters; notes). — Hammond, 1984: 196 (checklist of species of Borneo). — Newton, 1990: 91 (characters in key; one described Australian species). — Naomi, 1996: 49 (key to species of Japan). — Downie and Arnett, 1996: 428 (characters). — Drugmand, 1998a: 207 (list of types in l’Institut royal des Sciences naturelles de Belgique). — Newton, Thayer, Ashe, and Chandler, 2000: 376 (2 Nearctic species; notes; characters in key). — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 57 (necrophilous species in Mexico). Chasolium Laporte, 1835: 132 (species included: ernestini). TYPE SPECIES: Chasolium ernestini Laporte, fixed by monotypy. — Laporte, 1840: 190 (characters). — Lacordaire, 1854: 129 (characters; notes; list of species). — Fauvel, 1865: 35 (synonym of Isomalus). — Gemminger and Harold, 1868: 676 (catalog). — Fauvel, 1869: 494 (synonym of Eleusis). — Duvivier, 1883: 204 (catalog). — Bernhauer and Schubert, 1910: 10 (synonym of Eleusis). — Cameron, 1930: 74 (synonym of Eleusis). — Blackwelder, 1943: 155 (synonym of Eleusis; type species ernestini). — Blackwelder, 1952: 99 (synonym of Eleusis; type species: ernestini). Isomalus Erichson, 1839a: 31 (without included species, for first included species see Erichson, 1840). TYPE SPECIES: Isomalus humilis Erichson, fixed by subsequent designation by Duponchel, 1841: 57. — Erichson, 1840: 838 (first included species: complanatus, humilis, interruptus, bicolor, testaceus; characters). — Lacordaire, 1854: 128 (characters; notes; list of species). — LeConte, 1861a: 72 (characters). — LeConte, 1863: 27] (list of North American species). — Fauvel, 1864b: 31 [= 1865: 35 (characters; list of and key to species). — Gemminger and Harold, 1868: 675, 676 (synonym of Eleusis and Chasolium). — Horn, 1871: 297 (characters). — Sharp, 1876: 410 (notes). — Bernhauer and Schubert, 1910: 10 (synonym of Eleusis). — Cameron, 1930: 74 (synonym of Eleusis). — Blackwelder, 1943: 155 (subgenus of Eleusis; type species humilis). — Steel, 1950a: 215 (subgenus of Eleusis; characters in key). — Blackwelder, 1952: 205 (synonym of Eleusis; type species). Leiosoma Chevrolat, 1846: 279 (species included: none; [Note: Nomen nudum; it was published without description, without species and as synonym of Isomalus (Articles 11.6, 12.1); Blackwelder, 1952: 213 cited the genus as an isogenotypic synonym of Isomalus]). Liosoma Agassiz, 1847: 204 (emendation of Leiosoma; [Note: Nomen nudum; it was proposed to replace an unavailable name (Article 12.2.3)]). acuticollis Bernhauer, 1902: 171 (Eleusis; Type locality: Port Daufin, Madagaskar). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). — Scheerpeltz, 1961g: 240 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. adusta Fauvel, 1864b: 33 [= 1865: 37] (Isomalus; Type locality: Brésil: Nuovo Friburgo, prov. de Rio Janeiro). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). DISTRIBUTION: Brazil. 1074 OSORIINAE: ELEUSIS Bulletin AMNH agilis Sharp, 1876: 410 (Isomalus; Type locality: Ega). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). DISTRIBUTION: Brazil. alternans Bernhauer, 1926d: 252 (Eleusis; Type locality: Luzon, Provinz Mountain, Balbalasang). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. alutella Fauvel, 1903e: 237 (Eleusis; Type locality: Cameroon; Gabon: Libreville; Mada- gascar: Diego-Suarez, Tananarivo, Fort Dauphin). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). — Eichelbaum, 1913: 115 (Eleusis; Tanzania). — Scheerpeltz, 1956a: 481, 496 (Eleusis; characters; Nigeria; Cameroon; French Congo; Belgian Congo; Kenya; Uganda; Tanganyika; Mozambique; Madagascar). — Scheerpeltz, 1961g: 245 (Eleusis; characters; Madagascar). — Levasseur, 1968a: 49 (Eleusis; Central African Republic). — Frank, 1982: 6 (Eleusis; list of parasites). DISTRIBUTION: Gabon, French Equatorial Africa, Cameroon, Central African Republic, Congo, Uganda, Kenya, Tanzania, Mozambique, Madagascar. alutiventris Bernhauer, 1926d: 248 (Eleusis; Type locality: Luzon). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. andrewesi Cameron, 1930: 79 (Eleusis; Type locality: Niligiri Hills). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: India. angustae Bernhauer, 1926, see: Zeoleusis. angusticeps Bernhauer, 1926d: 249 (Eleusis; Type locality: Luzon, Provinz Camarinas Sur, Mount Isarog: Provinz Bataan, Limay. Mindanao, Provinz Lanao, Mumungan. Palawan, Binaluan. Basilan. Mindoro, San Teodoro. Masbate, Aroroy). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. angusticollis Bernhauer, 1937b: 577 (Eleusis; Type locality: W. Usambara). — Scheerpeltz, 1956a: 495 (Eleusis; characters; Uganda; Kenya; Tanganyika). DISTRIBUTION: Tanzania, Kenya, Uganda. apicalis Fauvel, 1904e: 298 (Eleusis; Type locality: Tananarive; Fort Dauphin et forêts au nord de ce dernier). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). — Scheerpeltz, 1961g: 245 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. apicipennis Fairmaire, 1849: 291 (Isomalus; Type locality: Taïti). — Fauvel, 1877: 186 (Lispinus; characters; Tahiti). — Fauvel, 1878g: 482 (Lispinus; characters). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). — Bernhauer, 1920: 3 (Eleusis; Australia). — Cameron, 1927c: 259 (Eleusis; Fiji). — Coiffait, 1976: 219 (Eleusis; Tahiti). DISTRIBUTION: Society Islands, Fiji, Australia. apicipennis Kraatz, 1859, see: terminata Fauvel, 1869. armuellensis Bierig, 1931: 426 (Eleusis; Type locality: Panama: Puerto Armuelles, Paz. Küste nahe Kostarika). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Panama. Bulletin AMNH OSORIINAE: ELEUSIS 1075 atra Scheerpeltz, 1956a: 480, 493, 502 (Eleusis; Type locality: Belgisch-Kongo: Hembe Bitale. Also cited from French Congo). DISTRIBUTION: French Equatorial Africa, Congo. atriceps Bernhauer, 1926d: 250 (Eleusis; Type locality: Luzon, Provinz Mountain, Balbalasang). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. augustae Bernhauer, 1926d: 247 (Eleusis; Type locality: Luzon, Provinz Mountain, Pauai, Haight’s Place; Mount Polis). — Bierig, 1932: 516 (Eleusis; characters; Philippines). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. australis Fauvel, 1878, see: planicollis W. J. MacLeay, 1873. banghaasi Bernhauer, 1926, see: Zeoleusis. banosana Bernhauer, 1926d: 251 (Eleusis; Type locality: Luzon, Provinz Laguna, Los Baños). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. basilewskyi Cameron, 1956: 177 (Eleusis; Type locality: Urundi: Bururi, 1800–2100 m). DISTRIBUTION: Burundi. basilewskyi Scheerpeltz, 1961, see: pierrei Herman, 2001. beesoni Cameron, 1930: 81 (Eleusis; Type locality: Mussoorie District: Kaligad). — Scheerpeltz, 1933: 996 (Eleusis; catalog). — Abdullah and Qadri, 1970: 116 (Eleusis; characters; West Pakistan). DISTRIBUTION: Pakistan, India. belua Bierig, 1932: 516 (Eleusis; Type locality: Philippinen: Mount Data, 7000 Fuss). DISTRIBUTION: Philippines. bernhaueri Wendeler, 1928, see: quadraticeps Bernhauer, 1926. bicarinata Bernhauer, 1904: 7 (Eleusis; Type locality: Madagaskar). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). — Scheerpeltz, 1961g: 244 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. bicolor Erichson, 1840: 839 (Isomalus; Type locality: Mexico). — Fauvel, 1864b: 34 [= 1865: 38] (Isomalus; characters; Mexico). — Solsky, 1868a: 144 (Isomalus; Mexico). — Sharp, 1887: 729 (Eleusis; characters; Mexico; Guatemala). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). DISTRIBUTION: Mexico, Guatemala. divisa Sharp, 1887: 729 (Eleusis; variety of bicolor; Type locality: Guatemala, Cerro Zunil, Zapote, Senahu, San Gerónimo). biimpressa Scheerpeltz, 1965: 131 (Eleusis; Type locality: British Sikkim, 200 m, Valley at Tista Bridge). DISTRIBUTION: India. bipunctata Eichelbaum, 1913, see: picta Eppelsheim, 1895. birmana Cameron, 1930: 453 (Eleusis; Type locality: Burma: Ruby Mines). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Myanmar. bisulcata Bernhauer, 1904: 8 (Eleusis; Type locality: Madagaskar). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). — Scheerpeltz, 1961g: 242 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. 1076 OSORIINAE: ELEUSIS Bulletin AMNH bitalensis Fagel, 1957b: 200 (Eleusis; Type locality: Congo Belge: Kivu: territoire de Kalehe, Bitale, N.O. Kahuzi, 1800 m). DISTRIBUTION: Congo. boops Fauvel, 1904e: 299 (Eleusis; Type locality: forêts au nord de Fort Dauphin). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). — Scheerpeltz, 1961g: 245 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. brachycephala Bernhauer, 1934j: 171 (Eleusis; Type locality: Ost-Java, Nongkodjadjar, 4000 f.). DISTRIBUTION: Indonesia. brachyptera Bernhauer, 1904: 6 (Eleusis; Type locality: Madagaskar). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). — Scheerpeltz, 1961g: 241 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. brasiliana Wendeler, 1955: 188 (Eleusis; Type locality: Brasilien: Espirito Santo, Affonso Claudio, Laranja da Terra). DISTRIBUTION: Brazil. breviceps Sharp, 1887: 731 (Eleusis; Type locality: Guatemala, Senahu 2500 feet). — Bernhauer and Schubert, 1910: 10 (Eleusis; catalog). DISTRIBUTION: Guatemala. brevipennis Fauvel, 1889b: 242 (Eleusis; Type locality: Nouvelle-Calédonie). — Fauvel, 1903d: 256 (Eleusis; New Caledonia; New Hebrides). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: New Caledonia, Vanuatu. brunnea Bernhauer, 1926d: 249 (Eleusis; Type locality: Mindanao, Provinz Lanao, Mumungan). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. burgeoni Bernhauer, 1934, see: capitata Schubert, 1911. canadensis Horn, 1871, see: Renardia. capitata Schubert, 1911: 2 (Eleusis; Type locality: Brit. Uganda). — Scheerpeltz, 1933: 996 (Eleusis; catalog). — Scheerpeltz, 1956a: 479, 489 (Eleusis; characters; Belgian Congo; Uganda; Kenya; Tanganyika; French Congo). DISTRIBUTION: Tanzania, Kenya, Uganda, French Equatorial Africa, Congo. burgeoni Bernhauer, 1934g: 228 (Eleusis; Type locality: Mombasa, 36 km. südlich von Lubero; Ende). — Scheerpeltz, 1956a: 489 (Eleusis; synonym of capitata). cephalotes Bernhauer, 1904a: 218 (Eleusis; Type locality: Mexiko). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Mexico. circuliceps Bernhauer, 1926d: 252 (Eleusis; Type locality: Luzon, Provinz Laguna, Los Baños). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. coarctata Sharp, 1889: 466 (Eleusis; Type locality: Yuyama; Sapporo; Nikko). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Nakane, 1963a: 81 (Eleusis; characters; habitus photograph; Japan). — Frank, 1982: 6 (Eleusis; list of parasites). — Naomi, 1996: 49 (Eleusis; characters; Japan). — Cho and Ahn, 1999: 176 (Eleusis; characters; Korea). DISTRIBUTION: Japan, Korea. Bulletin AMNH OSORIINAE: ELEUSIS 1077 complanata Erichson, 1840, see: ernestini Laporte, 1835. conifrons Fauvel, 1904b: 82 (Eleusis; Type locality: Sumatra: Monts Singalang). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Cameron, 1928d: 91 (Eleusis; illustrations; Sumatra). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Indonesia. conradti Fauvel, 1904d: 292 (Eleusis; Type locality: Usambara: Derema, 850 m). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Scheerpeltz, 1956a: 490 (Eleusis; characters; Uganda; Kenya; Tanganyika). — Frank, 1982: 6 (Eleusis; list of parasites). DISTRIBUTION: Tanzania, Kenya, Uganda. corvina Sharp, 1887: 730 (Eleusis; Type locality: Guatemala, Senahu). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Guatemala. crassicornis Bernhauer, 1926d: 250 (Eleusis; Type locality: Luzon, Provinz Laguna, Los Baños). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. cribriceps Cameron, 1950a: 179 (Eleusis; Type locality: Kenya Colony: Tumutumu, 5500 feet). DISTRIBUTION: Kenya. darlingtoni Blackwelder, 1943: 156 (Eleusis; Type locality: Cuba; Soledad, near Cienfuegos, province of Santa Clara). DISTRIBUTION: Cuba. decipiens Cameron, 1932, see: Zeoleusis. densiceps Bernhauer, 1926d: 254 (Eleusis; Type locality: Luzon, Provinz Mountain, Pauai, Haight’s Place). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. denticollis Cameron, 1930: 80 (Eleusis; Type locality: Niligiri Hills). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: India. derivata Bierig, 1932: 518 (Eleusis; Type locality: Philippinen: Mount Santo Tomas, 6500 Fuss). DISTRIBUTION: Philippines. discalis Fauvel, 1904b: 83 (Eleusis; Type locality: Sumatra: Monts Singalang). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Bernhauer, 1926d: 249 (Eleusis; characters; Philippines). — Cameron, 1928d: 91 (Eleusis; Sumatra). DISTRIBUTION: Indonesia. divergens Bernhauer, 1926d: 247 (Eleusis; Type locality: Luzon, Provinz Laguna, Los Baños). — Bernhauer, 1928: 1 (Eleusis; Sumatra). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Philippines. diversicollis Fauvel, 1879: 77 (Eleusis; Type locality: Mysol). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Cameron, 1931: 355 (Eleusis; New Guinea). DISTRIBUTION: New Guinea. divisa Sharp, 1887, see: bicolor Erichson, 1840. dohertyi Cameron, 1950d: 1 (Eleusis; Type locality: Perak). DISTRIBUTION: Malaysia. 1078 OSORIINAE: ELEUSIS Bulletin AMNH drescheri Cameron, 1936a: 26 (Eleusis; Type locality: Java: G. Slamat, Batoerraden). DISTRIBUTION: Indonesia. dubia Sharp, 1876: 411 (Isomalus; Type locality: Ega). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Brazil. dubiosa Bernhauer, 1929b: 81 (Eleusis; Type locality: Haut-Uele: Watsa; Ituri: La Moto, Madyu). — Scheerpeltz, 1933: 996 (Eleusis; catalog). — Scheerpeltz, 1956a: 488 (Eleusis; characters; French Congo; Belgian Congo). DISTRIBUTION: French Equatorial Africa, Congo. ellipticeps Scheerpeltz, 1956a: 480, 495, 498 (Eleusis; Type locality: Belgisch-Kongo: Yangambi. Also cited from Cameroon and French Congo). — Fagel, 1957b: 196 (Eleusis; notes; Belgian Congo). DISTRIBUTION: Cameroon, French Equatorial Africa, Congo. ernestini Laporte, 1835: 132 (Chasolium; Type locality: Madagascar). — Laporte, 1840: 190 (Chasolium; characters; Madagascar). — Fauvel, 1869: 494 (Eleusis). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Scheerpeltz, 1961g: 233, 244 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. complanata Erichson, 1840: 839 (Isomalus; Type locality: Madagascar). — Bernhauer and Schubert, 1910: 11 (Eleusis; synonym of ernestini). exigua Bernhauer, 1926, see: Zeoleusis. fageli Scheerpeltz, 1961g: 243 (Eleusis; Type locality: Not cited but presumably Madagascar). DISTRIBUTION: Madagascar. fasciata LeConte, 1863, see: humilis Erichson, 1840. fauveli Fagel, 1957b: 202 (Eleusis; Type locality: Natal). DISTRIBUTION: South Africa. feae Fauvel, 1895: 188 (Eleusis; Type locality: Bírmanie, Carin Cheba, 900–1100 m). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Cameron, 1930: 78 (Eleusis; characters; Burma; Java). — Scheerpeltz, 1933: 996 (Eleusis; catalog). DISTRIBUTION: Myanmar, Indonesia. fenestrata Sharp, 1887: 730 (Eleusis; Type locality: Guatemala, Cerro Zunil). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Guatemala. ferruginea Cameron, 1925b: 231 (Eleusis; Type locality: Selangor, Federated Malay States; The Gap; Borneo, Tutai River). — Cameron, 1928d: 91 (Eleusis; illustrations; Sumatra). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Borneo, Malaysia, Indonesia. flava Fagel, 1957b: 194 (Eleusis; Type locality: Urundi: territoire de Bururi, Nyamurembe, 900 m). DISTRIBUTION: Burundi. flaviceps Bernhauer, 1926d: 249 (Eleusis; Type locality: Luzon, Provinz Camarines Sur, Mount Isarog: Provinz Laguna, Los Baños). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Philippines. flavipennis Sharp, 1887: 731 (Eleusis; Type locality: Guatemala, Purula). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Guatemala. Bulletin AMNH OSORIINAE: ELEUSIS 1079 frater Fauvel, 1864b: 35 [= 1865: 39] (Isomalus; Type locality: Caracas). — Fauvel, 1891: 88 (Eleusis; Venezuela). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Venezuela. frontalis Schubert, 1911: 3 (Eleusis; Type locality: Antananarivo, Madagascar). — Scheerpeltz, 1933: 997 (Eleusis; catalog). — Scheerpeltz, 1961g: 240 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. fulvicornis Eppelsheim, 1895a: 140 (Eleusis; Type locality: Gabon). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Bernhauer, 1942c: 347 (Eleusis; Cameroon). — Scheerpeltz, 1956a: 480, 496 (Eleusis; characters; Liberia; Cameroon; Togo; Nigeria; French Congo; Belgian Congo). — Cameron, 1952e: 827 (Eleusis; Togo). DISTRIBUTION: Liberia, Togo, Nigeria, Cameroon, French Equatorial Africa. fusciceps Kraatz, 1859, see: Zeoleusis. ghumensis Herman, 2001: 39 (Eleusis; replacement name for inermis Cameron). DISTRIBUTION: India. inermis Cameron, 1940c: 210 (Eleusis; [preoccupied]; Type locality: Ghum District). — Herman, 2001: 39 (Eleusis; synonym of ghumensis). glabricollis Cameron, 1928d: 92 (Eleusis; Type locality: Sumatra; Fort de Kock). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Indonesia. globiceps Cameron, 1929b: 436 (Eleusis; Type locality: Selangor: The Gap. Java: Buitenzorg). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Malaysia. haddeni Bierig, 1932: 517 (Eleusis; Type locality: Philippinen: Mount Maquiling). — Nishida, 1997: 62 (Philonthus; [Note: An evident error but I am unsure whether the mistake is in the generic or specific name]; Hawaii). DISTRIBUTION: Philippines, Hawaii (?). hesperidum Wollaston, 1857a: 504 (Isomalus; Type locality: Cape Verde Islands: St. Vincent). — Wollaston, 1867: 257 (Trogophloeus; characters; notes; Cape Verde Islands). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Cape Verde Islands. heterodera Fauvel, 1904e: 299 (Eleusis; Type locality: Tananarive; Diego-Suarez; forêt Tanala). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Scheerpeltz, 1961g: 244 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. howa Bernhauer, 1902: 170 (Eleusis; Type locality: Port Daufin, Madagaskar). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Scheerpeltz, 1961g: 233, 240, 241 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. humeralis Bernhauer, 1939i: 252 (Eleusis; Type locality: Kamerun: Mt. Cameroon: Mann’s Quelle, 7400 feet). — Scheerpeltz, 1956a: 495 (Eleusis; characters; Cameroon; Fernando Po; French Congo; Belgian Congo). DISTRIBUTION: Bioko, Cameroon, French Equatorial Africa, Congo. humilis Erichson, 1840: 839 (Isomalus; Type locality: Brasilia). — Fauvel, 1904a: 44 (Eleusis; Nilghiris; Annam; China; Japan; Burma; Tenasserim; Sumatra; Bengkalis Island; Nias; Arizona; Central America; Brazil). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). 1080 OSORIINAE: ELEUSIS Bulletin AMNH
— Cameron, 1921a: 362, 398 (Eleusis; characters; Singapore). — Bernhauer, 1922c: 221 (Eleusis; Formosa). — Cameron, 1927c: 259 (Eleusis; Fiji). — Cameron, 1928d: 91 (Eleusis; illustration; Sumatra). — Bernhauer, 1928: 1 (Eleusis; Sumatra). — Cameron, 1930: 83 (Eleusis; characters; Ceylon; India; Burma; Japan). — Scheerpeltz, 1933: 997 (Eleusis; catalog). — Cameron, 1936a: 25 (Eleusis; Java). — Nakane and Sawada, 1960: 49 (Eleusis; Yakushima Island). — Watanabe, 1961: 349 (Eleusis; Izu Islands). — Nakane, 1963a: 81 (Eleusis; characters; habitus photograph; Japan). — Abdullah and Qadri, 1970: 116 (Eleusis; characters; West Pakistan). — Watanabe and Shibata, 1972: 60 (Eleusis; Yaku-shima Island). — Coiffait, 1978b: 112 (Eleusis; Bhutan). — Coiffait, 1981f: 331 (Eleusis; Andaman Islands). — Frank, 1982: 6 (Eleusis; list of parasites). — Biswas and Biswas, 1985: 141 (Eleusis; India). — Naomi, 1996: 49, 51 (Eleusis; characters; Japan). DISTRIBUTION: Brazil, Venezuela, Central America, USA, Pakistan, India, Sri Lanka, Andaman Islands, Myanmar, Singapore, Indonesia, Fiji, China, Taiwan, Japan. indica Kraatz, 1859: 183 (Isomalus; Type locality: India orientalis; Ceylan). — Fauvel, 1891: 88 (Eleusis; notes; Venezuela). — Fauvel, 1904a: 44 (Eleusis; synonym of humilis). — Bernhauer and Schubert, 1910: 11 (Eleusis; synonym of humilis). — Cameron, 1930: 83 (Eleusis; synonym of humilis). tenuis Walker, 1859: 52 (Prognathus, cited as Prognatha; Type locality: Ceylon). — Fauvel, 1891: 88 (Eleusis; synonym of indica). — Fauvel, 1904a: 44 (Eleusis; synonym of humilis). — Bernhauer and Schubert, 1910: 11 (Eleusis; synonym of humilis). — Cameron, 1930: 83 (Eleusis; synonym of humilis). fasciata LeConte, 1863a: 59 (Isomalus; Type locality: valley of Gila River, Arizona). — Horn, 1871: 297, 298 (Isomalus; characters; Arizona). — Fauvel, 1877a: xxiv (Eleusis; Brazil). — Fauvel, 1878d: 180 [= 1878e: 17] (Eleusis; characters; Arizona; Mexico; Venezuela; Brazil). — Fauvel, 1891: 88 (Eleusis; synonym of indica). — Fauvel, 1904a: 44 (Eleusis; synonym of humilis). — Bernhauer and Schubert, 1910: 11 (Eleusis; synonym of humilis). — Cameron, 1930: 83 (Eleusis; synonym of humilis). — Naomi, 1996: 51 (Eleusis; synonym of humilis). tenuis Fauvel, 1864b: 37 [= 1865: 41] (Isomalus; [preoccupied]; Type locality: Venezuela: Caracas). — Sharp, 1876: 411 (Isomalus; Brazil). — Fauvel, 1891: 88 (Eleusis; synonym of indica). — Fauvel, 1904a: 44 (Eleusis; synonym of humilis). — Bernhauer and Schubert, 1910: 11 (Eleusis; synonym of humilis). — Cameron, 1930: 83 (Eleusis; synonym of humilis). subtilis Sharp, 1889: 467 (Eleusis; Type locality: Yuyama and Oyayama, in Kiushiu). — Fauvel, 1904a: 44 (Eleusis; synonym of humilis). — Bernhauer and Schubert, 1910: 11 (Eleusis; synonym of humilis). — Cameron, 1930: 83 (Eleusis; synonym of humilis). hydrocephala Fauvel, 1904e: 300 (Eleusis; Type locality: Tananarive). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Scheerpeltz, 1961g: 246 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. Bulletin AMNH OSORIINAE: ELEUSIS 1081 impressicollis Fairmaire, 1880: 236 (Chasolium; Type locality: Nossi-Bé). — Fairmaire, 1880a: 326 (Chasolium; characters; Madagascar). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Madagascar. indica Kraatz, 1859, see: humilis Erichson, 1840. inermis Bernhauer, 1902: 171 (Eleusis; Type locality: Port Daufin, Madagaskar). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Scheerpeltz, 1961g: 242 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. inermis Cameron, 1940, see: ghumensis Herman, 2001. infima Sharp, 1887: 731 (Eleusis; Type locality: Nicaragua, Chontales). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Nicaragua. insignis Sharp, 1887: 732 (Eleusis; Type locality: Guatemala, Quiche Mountains 7000 to 9000 feet). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). DISTRIBUTION: Guatemala. insularis Cameron, 1936, see: Zeoleusis. interrupta Erichson, 1840, see: Piestus. iridescens Bernhauer, 1926d: 254 (Eleusis; Type locality: Mindanao, Provinz Zamboanga, Dapitan. Luzon, Provinz Mountain, Pauai, Haight’s Place). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Philippines. jacobsoni Cameron, 1928d: 93 (Eleusis; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Indonesia. javana Cameron, 1936a: 25 (Eleusis; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft. Sukabumi; also in the Malay Peninsula). DISTRIBUTION: Indonesia, Malaysia. kahuziensis Fagel, 1957b: 190 (Eleusis; Type locality: Congo Belge: Kivu: territoire de Kalehe, contreforts Sud du Kahuzi, 2200 m). DISTRIBUTION: Congo. kamerunensis Bernhauer, 1912f: 177 (Eleusis; Type locality: Südost Kamerun: Lolodorf). — Scheerpeltz, 1933: 997 (Eleusis; catalog). — Scheerpeltz, 1956a: 494 (Eleusis; characters; Nigeria; Cameroon; French Congo; Belgian Congo). — Fagel, 1957a: 27 (Eleusis; Ruanda). DISTRIBUTION: Nigeria, Cameroon, French Equatorial Africa, Congo, Rwanda. khumbuensis Coiffait, 1984: 137 (Eleusis; Type locality: Nepal: Khumbu, environs de Sutje, près de Lagla). DISTRIBUTION: Nepal. kolbei Fauvel, 1904d: 292 (Eleusis; Type locality: Usambara: Dereema, 850 m). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Scheerpeltz, 1956a: 490 (Eleusis; characters; Uganda; Kenya; Tanganyika). — Fagel, 1957b: 193 (Eleusis; notes). DISTRIBUTION: Uganda, Kenya, Tanzania. kraatzi Fauvel, 1878, see: terminata Fauvel, 1869. laevis Fagel, 1957b: 199 (Eleusis; Type locality: Ruanda: Parc National Albert: lac N’Gando, pied volcan Karisimbi, 2400 m). DISTRIBUTION: Rwanda. 1082 OSORIINAE: ELEUSIS Bulletin AMNH lata Coiffait, 1981f: 343 (Eleusis; Type locality: South Andaman: Rutland Island). DISTRIBUTION: Andaman Islands. laticeps Fauvel, 1895: 190 (Eleusis; Type locality: Bírmanie, Carin Cheba, 500–1100 m; Sumatra, tabacs; Ile Nias). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Cameron, 1928d: 91 (Eleusis; illustrations; Sumatra). — Cameron, 1930: 84 (Eleusis; characters; Burma; Sumatra; Nias Island). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Myanmar, Indonesia. leleupi Fagel, 1957b: 201 (Eleusis; Type locality: Congo Belge: Kivu: territoire de Kabare, contref. S.E. Kahuzi, 2000 m). DISTRIBUTION: Congo. longelytrata Bernhauer, 1926, see: philippina Bernhauer, 1914. longiceps Fauvel, 1878f: 208 (Eleusis; Type locality: Nouvelle-Guinée, Ramoi; Ile Yule). — Bernhauer, 1902b: 45 (Eleusis; Ceylon). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Cameron, 1930: 88 (Eleusis; characters; Ceylon ?). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: New Guinea, Ceylon ?. longicornis Fauvel, 1903e: 236 (Eleusis; Type locality: Cameroun; Gabon: Libreville). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1933: 997 (Eleusis; catalog). — Scheerpeltz, 1956a: 480, 488 (Eleusis; characters; Cameroon; French Congo; Belgian Congo). DISTRIBUTION: Gabon, Cameroon, French Equatorial Africa, Congo. longipennis Bernhauer, 1926, see: philippina Bernhauer, 1914. lunduensis Cameron, 1934e: 371 (Eleusis; Type locality: Mayumbe: Lundu). — Scheerpeltz, 1956a: 490 (Eleusis; characters; French Congo; Belgian Congo). DISTRIBUTION: French Equatorial Africa, Congo. lunigera Fauvel, 1904b: 84 (Eleusis; Type locality: Hue; Malacca; Singapore; Java: Tcibodas; Pengalengan, 1300 m; Sukabumi; Nouvelle-Guinee: Friedrich-Wilhelmhafen; baie Huon: Simbang; Sattelberg; Ile Deslacs). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Cameron, 1921a: 362, 398 (Eleusis; characters; Singapore). — Cameron, 1928d: 92 (Eleusis; illustrations; Sumatra). — Scheerpeltz, 1933: 997 (Eleusis; catalog). — Cameron, 1936a: 25 (Eleusis; Java). — Frank, 1982: 7 (Eleusis; list of parasites). DISTRIBUTION: New Guinea, Indonesia, Singapore, Vietnam. luvubuensis Fagel, 1957b: 197 (Eleusis; Type locality: Congo Belge: Kivu: territoire d’Uvira, Luvubu, 2650 m). DISTRIBUTION: Congo. luzonica Bernhauer, 1926d: 247 (Eleusis; Type locality: Luzon, Provinz Neuva Vizcaya, Imugan: Provinz Mountain, Mount Polis; Balaban). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Philippines. madagascariensis Bernhauer, 1917m: 554 (Eleusis; Type locality: Madagascar). — Scheerpeltz, 1933: 997 (Eleusis; catalog). — Scheerpeltz, 1961g: 242 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. magniceps Cameron, 1925b: 230 (Eleusis; Type locality: Soekaboemi, W. Preanger). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Indonesia. Bulletin AMNH OSORIINAE: ELEUSIS 1083 malaisei Scheerpeltz, 1965: 124 (Eleusis; Type locality: N.E. Burma: Kambaiti, 2000 m). DISTRIBUTION: Myanmar. marginicollis Bernhauer, 1921b: 65 (Eleusis; Type locality: Kolumbien). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Colombia. maynei Fagel, 1957b: 192 (Eleusis; Type locality: Congo Belge: Kivu: Musisi). DISTRIBUTION: Congo. megacephala Bernhauer, 1939j: 376 (Eleusis; Type locality: Belgisches Kongo: S. Kivu: N’Gweshe). — Scheerpeltz, 1956a: 481, 493 (Eleusis; characters; French Congo; Belgian Congo). DISTRIBUTION: French Equatorial Africa, Congo. microlestiformis Coiffait, 1981f: 342 (Eleusis; Type locality: South Andaman: Rutland Island). DISTRIBUTION: Andaman Islands. microphthalma Scheerpeltz, 1965: 126 (Eleusis; Type locality: N.E. Burma, Kambaiti, 7000 ft). DISTRIBUTION: Myanmar. mixta Sharp, 1887: 732 (Eleusis; Type locality: Mexico, Cordova; Guatemala, Cerro Zunil, San Geronimo, Purula). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). DISTRIBUTION: Mexico, Guatemala. mjobergi Cameron, 1928g: 424 (Eleusis; cited as mjöbergi; Type locality: Borneo: Tutau River). — Cameron, 1930b: 160 (Eleusis; cited as mjobergi; British North Borneo). — Scheerpeltz, 1933: 997 (Eleusis; cited as mjöbergi; catalog). — Hammond, 1984: 196 (Eleusis; cited as mjoebergi; Borneo). — Herman, 2001: 12 (Eleusis; spelling corrected to mjobergi). DISTRIBUTION: Borneo. montana Bernhauer, 1926d: 251 (Eleusis; Type locality: Luzon, Provinz Mountain, Mount Polis). — Bernhauer, 1928: 1 (Eleusis; Sumatra). — Scheerpeltz, 1933: 997 (Eleusis; catalog). DISTRIBUTION: Philippines, Indonesia. morstatti Scheerpeltz, 1956a: 492 (Eleusis; Type locality: Kamerun; Franz. Kongo). DISTRIBUTION: Cameroon, French Equatorial Africa. motoensis Bernhauer, 1929b: 81 (Eleusis; Type locality: Haut-Uele: Moto). — Scheerpeltz, 1933: 997 (Eleusis; catalog). — Scheerpeltz, 1956a: 494 (Eleusis; characters; French Congo; Belgian Congo). DISTRIBUTION: French Equatorial Africa, Congo. multizonata Bierig, 1932: 520 (Eleusis; Type locality: Philippines: Mount Santo Tomas, 6500 Fuss). DISTRIBUTION: Philippines. mutica Bernhauer, 1904: 9 (Eleusis; Type locality: Africa occidentalis, Gaboon, Libreville). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1956a: 480, 488 (Eleusis; characters; Cameroon; French Congo; Belgian Congo). — Levasseur, 1968a: 50 (Eleusis; Central African Republic). DISTRIBUTION: Cameroon, French Equatorial Africa, Gabon, Congo, Central African Republic. myrmidon Fairmaire and Germain, 1861, see: Zeoleusis. nigerrima Fauvel, 1864b: 33 [= 1865: 37] (Isomalus; Type locality: Brésil: Nuovo Friburgo, prov. de Rio Janeiro). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1933: 997 (Eleusis; catalog; Argentina). — Frank, 1982: 7 (Eleusis; list of parasites). DISTRIBUTION: Argentina, Brazil. 1084 OSORIINAE: ELEUSIS Bulletin AMNH nigra Cameron, 1928d: 93 (Eleusis; Type locality: Sumatra: Gunung Singgalang). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Indonesia. nigrellus LeConte, 1863, see: Renardia. nigrescens Cameron, 1925b: 231 (Eleusis; Type locality: Selangor, Federated Malay States; The Gap). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — Cameron, 1936a: 25 (Eleusis; Java). DISTRIBUTION: Malaysia, Indonesia. nigricans Scheerpeltz, 1956a: 480, 493, 501 (Eleusis; Type locality: Belgisch-Kongo: Yangambi. Also cited from French Congo). DISTRIBUTION: Congo, French Equatorial Africa. nigriventris Lea, 1910a: 140 (Eleusis; Type locality: Victoria: Portland; Tasmania: Hobart). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — E. Matthews, 1982: 8 (Eleusis; notes; Australia). — Newton, 1990: 91 (belongs in Omaliinae; [Note: No genus assigment was specified]). DISTRIBUTION: Australia. nilgiriensis Cameron, 1930: 77 (Eleusis; Type locality: Nilgiri Hills). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: India. nova Bernhauer, 1926, see: viridans Fauvel, 1895. obscura Bernhauer, 1902: 172 (Eleusis; Type locality: Port Daufin, Madagaskar). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1961g: 245 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. obscuriceps Bernhauer, 1926d: 251 (Eleusis; Type locality: Luzon, Provinz Mountain, Balbalasang). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. obscuripennis Cameron, 1928: 11 (Eleusis; Type locality: Belgian Congo: Haut Uele, Moto; Bas Uele, Koteli; Ituri: Medje). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — Scheerpeltz, 1956a: 491 (Eleusis; characters; French Congo; Belgian Congo). DISTRIBUTION: French Equatorial Africa, Congo. schoutedeni Bernhauer, 1932a: 72 (Eleusis; Type locality: Bas Uele: Koteli). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — Scheerpeltz, 1956a: 491 (Eleusis; synonym of obscuripennis). obscuripes Fagel, 1957b: 191 (Eleusis; Type locality: Congo Belge: Yangambi). DISTRIBUTION: Congo. ocellata Bernhauer, 1926d: 252 (Eleusis; Type locality: Mindanao, Provinz Lanao, Mumungan. Luzon, Provinz Nueva Vizcaya, Imugan). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. orbiceps Cameron, 1937a: 83 (Eleusis; Type locality: New Guinea: Mt. Tafa). DISTRIBUTION: New Guinea. pacifica Cameron, 1933f: 73 (Eleusis; Type locality: Marquesas Islands: Eiao: near center, altitude 1300 feet). — Coiffait, 1976: 219 (Eleusis; Tahiti). DISTRIBUTION: Marquesas Islands, Society Islands. palawanensis Bernhauer, 1926d: 249 (Eleusis; Type locality: Palawan, Binaluan). — Bierig, 1932: 518 (Eleusis; characters; Philippines). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. Bulletin AMNH OSORIINAE: ELEUSIS 1085 pallida LeConte, 1863a: 58 (Isomalus; Type locality: Pennsylvania, Lancaster County). — Horn, 1871: 297, 298 (Isomalus; characters; New York to Washington, D.C.). — Fauvel, 1878d: 181 [= 1878e: 17] (Eleusis; characters; New York to Washington; Michigan; Pennsylvania). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Downie and Arnett, 1996: 428 (Eleusis; characters; USA). DISTRIBUTION: USA. pallidipennis Fauvel, 1864b: 36 [= 1865: 40] (Isomalus; Type locality: Caracas, Venezuela; Teapa, Mexico; Amérique du Nord). — Fauvel, 1877a: xxiv (Eleusis; synonym of fasciata). — Fauvel, 1878d: 181 [= 1878e: 17] (Eleusis; synonym of fasciata). — Sharp, 1887: 730 (Eleusis; notes; Mexico; British Honduras; Guatemala). — Fauvel, 1891: 88 (Eleusis; synonym of indica). — Fauvel, 1904a: 44 (Eleusis; synonym of humilis). — Bernhauer and Schubert, 1910: 11 (Eleusis; synonym of humilis). — Cameron, 1930: 83 (Eleusis; synonym of humilis). — Scheerpeltz, 1975a: 241 (Eleusis; valid species; collecting notes; Brazil). DISTRIBUTION: Mexico, Belize, Guatemala, Venezuela, Brazil. papuana Cameron, 1937a: 83 (Eleusis; Type locality: New Guinea: Buna Bay; Mount Lamington. Kokoda, alt. 1200 ft). DISTRIBUTION: New Guinea. paralleliceps Bernhauer, 1926d: 254 (Eleusis; Type locality: Luzon, Provinz Mountain, Pauai, Haight’s Place). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. parva Blackburn, 1892, see: Zeoleusis. pauliani Cameron, 1948a: 224 (Eleusis; Type locality: Ivory Coast: Orombo Bocca, 550 m). — Cameron, 1952e: 827 (Eleusis; Togo). DISTRIBUTION: Ivory Coast, Togo. pauxilla Scheerpeltz, 1965: 133 (Eleusis; Type locality: British Sikkim, 200 m, Valley at Tista Bridge). DISTRIBUTION: India. pennata Bernhauer, 1926d: 253 (Eleusis; Type locality: Mindanao, Provinz Zamboanga, Port Banga: Provinz Lanao, Mumungan. Palawan, Binaluan). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. perrieri Fauvel, 1904e: 298 (Eleusis; Type locality: Suberbieville; Pays Androy, Ambovombe; Fort Dauphin). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1961g: 241 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. persimilis Cameron, 1945: 63 (Eleusis; Type locality: Darjeeling: Ghum district: Mangpu). DISTRIBUTION: India. phaenomenalis Bernhauer, 1926d: 246 (Eleusis; Type locality: Luzon, Provinz Neuva Vizcaya, Imugan). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. philippina Bernhauer, 1914c: 76 (Eleusis; Type locality: Philippinen: Los Banos). — Bernhauer, 1926d: 253 (Eleusis; characters; Philippines). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. longelytrata Bernhauer, 1926d: 253 (Eleusis; variety of philippina; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 998 (Eleusis; variety of philippina). 1086 OSORIINAE: ELEUSIS Bulletin AMNH
longipennis Bernhauer, 1926d: 253 (Eleusis; variety of philippina; Type locality: Not cited). — Scheerpeltz, 1933: 998 (Eleusis; variety of philippina). picta Eppelsheim, 1895a: 139 (Eleusis; Type locality: Gabon). — Fauvel, 1903e: 236 (Eleusis; Cameroon; Gabon; Togo). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Bernhauer, 1942c: 347 (Eleusis; Cameroon). — Cameron, 1950g: 3 (Eleusis; Belgian Congo). — Scheerpeltz, 1956a: 480, 491 (Eleusis; characters; Guinea; Liberia; Nigeria; Togo; Cameroon; French Congo; Belgian Congo). — Fagel, 1957b: 191 (Eleusis; notes). — Scheerpeltz, 1957: 153 (Eleusis; Tanganyika). — Scheerpeltz, 1974c: 5 (Eleusis; Sudan). — Frank, 1982: 7 (Eleusis; cited as picto; list of parasites). DISTRIBUTION: Sudan, Tanzania, Congo, Gabon, French Equatorial Africa, Cameroon, Nigeria, Togo, Liberia, Guinea. bipunctata Eichelbaum, 1913: 116 (Eleusis; variety of picta; Type locality: Amani). picturata Cameron, 1950a: 179 (Eleusis; Type locality: Uganda: Kalinzu Forest, 4000 feet). DISTRIBUTION: Uganda. pierrei Herman, 2001: 39 (Eleusis; replacement name for basilewskyi Scheerpeltz). DISTRIBUTION: Madagascar. basilewskyi Scheerpeltz, 1961g: 240 (Eleusis; [preoccupied]; Type locality: Not cited but presumably Madagascar). — Herman, 2001: 39 (Eleusis; synonym of pierrei). plagiata Fauvel, 1904b: 83 (Eleusis; Type locality: Sikkim: Darjeeling). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Cameron, 1930: 81 (Eleusis; characters; Sikkim). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — Abdullah and Qadri, 1970: 118 (Eleusis; characters; East Pakistan). DISTRIBUTION: India, Bangladesh. plagiata Schubert, 1911, see: schuberti Scheerpeltz, 1933. planicollis W. J. MacLeay, 1873: 151 (Isomalus; Type locality: Gayndah). — Fauvel, 1877: 186 (Eleusis; characters; Australia). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Bernhauer, 1920: 3 (Eleusis; Australia). — Newton, 1990: 91 (Eleusis). DISTRIBUTION: Australia. australis Fauvel, 1878g: 482 (Eleusis; Type locality: Australie, Queensland, Wide Bay). — Bernhauer and Schubert, 1910: 12 (Eleusis; synonym of planicollis). polyodonta Scheerpeltz, 1956a: 491 (Eleusis; Type locality: Liberia; Süd-Nigeria; Kamerun; Franz u. Belg. Kongo). — Scheerpeltz, 1971c: 161 (Eleusis; Cameroon). DISTRIBUTION: Liberia, Nigeria, Cameroon, French Equatorial Africa, Congo. praeustus Perroud and Montrousier, 1864 [Note: Described in Isomalus and transferred to “Clavicornes Ino” by Fauvel, 1869: 494], see: Inopeplidae. preangerana Cameron, 1945a: 140 (Eleusis; Type locality: Java: Preanger; G. Patoeha). DISTRIBUTION: Indonesia. propinqua Bernhauer, 1904: 5 (Eleusis; Type locality: Madagaskar). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1961g: 241 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. punctatella Scheerpeltz, 1956a: 492 (Eleusis; Type locality: Franz. Kongo: Umg. Brazzaville). DISTRIBUTION: Congo Republic. Bulletin AMNH OSORIINAE: ELEUSIS 1087 puncticeps Bernhauer, 1912d: 26 (Eleusis; Type locality: Columbia: Cali: Alto de las Cruces, 2200 m; Rio Vitaco, 2000 m; Rio Aquatal 1800 m). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Colombia. punctigera Fauvel, 1878f: 208 (Eleusis; Type locality: Nouvelle-Guinée, Hatam). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). DISTRIBUTION: New Guinea. pusilla Fauvel, 1895: 190 (Eleusis; Type locality: Bírmanie, Bhamao; Ile Nias). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Cameron, 1928d: 92 (Eleusis; illustration; Sumatra). — Cameron, 1930: 87 (Eleusis; characters; Burma; Nias Island). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — Cameron, 1949b: 456 (Eleusis; China). DISTRIBUTION: China, Myanmar, Indonesia. quadraticeps Bernhauer, 1926d: 248 (Eleusis; Type locality: Luzon, Provinz Mountain, Mount Polis). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. bernhaueri Wendeler, 1928b: 298 (Eleusis; replacement name for quadraticeps Bernhauer but misspelled as quadriceps; [Note: Unnecessary replacement name; Wendeler misspelled Bernhauer’s name]). — Scheerpeltz, 1933: 998 (Eleusis; synonym of quadraticeps; new name not necessary; catalog). quadriceps Cameron, 1925b: 231 (Eleusis; Type locality: Perak; Burma). — Cameron, 1930: 78 (Eleusis; characters; Burma; Perak). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Myanmar, Malaysia. quadridens Fauvel, 1904b: 84 (Eleusis; Type locality: Perak). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Naomi, 1996: 49 (Eleusis; characters; Japan). DISTRIBUTION: Malaysia, Japan. quadridentata Cameron, 1928d: 93 (Eleusis; Type locality: Sumatra: Si Rambe). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Indonesia. quadripunctata Bernhauer, 1917m: 553 (Eleusis; Type locality: Madagascar: Antananarivo). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — Scheerpeltz, 1961g: 242 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. raoultii Fairmaire, 1880: 236 (Chasolium; Type locality: Nossi-Bé). — Fairmaire, 1880a: 326 (Chasolium; characters; Madagascar). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1961g: 241 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. rectangula Bernhauer, 1904: 6 (Eleusis; Type locality: Madagaskar). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1961g: 242 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. rotundiceps Fauvel, 1904b: 83 (Eleusis; Type locality: Sikkim: Darjeeling). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Cameron, 1930: 86 (Eleusis; characters; Sikkim). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — Abdullah and Qadri, 1970: 117 (Eleusis; characters; East Pakistan). DISTRIBUTION: India, Bangladesh. 1088 OSORIINAE: ELEUSIS Bulletin AMNH rufa Cameron, 1956: 177 (Eleusis; Type locality: Ruanda: Tshuryaga, forêt du Rugege, 2400 m, terr. d’Astrida). — Fagel, 1957b: 195 (Eleusis; characters; notes). DISTRIBUTION: Rwanda, French Equatorial Africa, Congo. rufobrunnea Scheerpeltz, 1956a: 481, 496, 504 (Eleusis; Type locality: Belgisch-Kongo: Hembe Bitale. Also cited from French Congo). — Fagel, 1957b: 195 (Eleusis; synonym of rufa). ruficollis Fauvel, 1878f: 207 (Eleusis; Type locality: Nouvelle-Guinée, Hatam). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). DISTRIBUTION: New Guinea. rufiventris Fauvel, 1904b: 81 (Eleusis; Type locality: Sumatra: Monts Singalang). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Cameron, 1928d: 91 (Eleusis; Sumatra). — Bernhauer, 1928: 1 (Eleusis; Sumatra). DISTRIBUTION: Indonesia. rufobrunnea Scheerpeltz, 1956, see: rufa Cameron, 1956. rufotestacea Bernhauer, 1926d: 251 (Eleusis; Type locality: Mindanao, Provinz Lanao, Mumungan). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. rufula Sharp, 1887: 729 (Eleusis; Type locality: Panama, Bugaba, Volcan de Chiriqui 2000 to 3000 feet). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). DISTRIBUTION: Panama. saegeri Fagel, 1957b: 193 (Eleusis; Type locality: Congo Belge: Bumba). DISTRIBUTION: Congo. scapularis Bernhauer, 1902: 171 (Eleusis; Type locality: Port Daufin, Madagaskar). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1961g: 243 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. schedli Scheerpeltz, 1956a: 480, 492, 497 (Eleusis; Type locality: Belgisch-Kongo: Yanagambi. Also cited from Nigeria, Cameroon, and French Congo). DISTRIBUTION: Nigeria, Cameroon, French Equatorial Africa, Congo. schoutedeni Bernhauer, 1932, see: obscuripennis Cameron, 1928. schuberti Scheerpeltz, 1933: 998 (Eleusis; replacement name for plagiata Schubert). — Scheerpeltz, 1961g: 244 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. plagiata Schubert, 1911: 3 [preoccupied] (Eleusis; Type locality: Antananarivo, Madagascar). — Scheerpeltz, 1933: 998 (Eleusis; synonym of schuberti). scita Sharp, 1887: 731 (Eleusis; Type locality: Guatemala, near the city, Capetillo, Zapote, Cerro Zunil). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). DISTRIBUTION: Guatemala. sculpticeps Bernhauer, 1926d: 251 (Eleusis; Type locality: Luzon, Provinz Mountain Balbalan). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Philippines. secreta Cameron, 1930: 87 (Eleusis; Type locality: Siwaliks: Timli). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — Abdullah and Qadri, 1970: 116 (Eleusis; characters; West Pakistan). DISTRIBUTION: Pakistan, India. Bulletin AMNH OSORIINAE: ELEUSIS 1089 semirufa Fairmaire and Germain, 1861, see: Zeoleusis. semisplendida Bierig, 1932: 519 (Eleusis; Type locality: Philippinen: Mount Santo Tomas, 6500 Fuss). DISTRIBUTION: Philippines. sikkimensis Cameron, 1945: 63 (Eleusis; Type locality: Darjeeling: Ghum district: Mangpu). DISTRIBUTION: India. sikkimensis Scheerpeltz, 1965, see: teestaensis Herman, 2001. simillima Fagel, 1957b: 197 (Eleusis; Type locality: Congo Belge: Kivu: territoire d’Uvira, bassin Ulindi, 2650 m). DISTRIBUTION: Congo. simulans Cameron, 1930: 452 (Eleusis; Type locality: Burma: Ruby Mines). — Scheerpeltz, 1933: 998 (Eleusis; catalog). DISTRIBUTION: Myanmar. sororcula Cameron, 1936, see: Zeoleusis. soror Bernhauer, 1926d: 253 (Eleusis; Type locality: Mindanao, Provinz Lanao, Mumungan. Luzon, Provinz Camarines Sur, Mount Isarog). — Scheerpeltz, 1933: 999 (Eleusis; catalog). DISTRIBUTION: Philippines. spectabilis Bernhauer, 1904: 4 (Eleusis; Type locality: Madagaskar). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1961g: 239 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. speculiceps Bernhauer, 1926d: 250 (Eleusis; Type locality: Masbate, Aroroy). — Scheerpeltz, 1933: 999 (Eleusis; catalog). DISTRIBUTION: Philippines. speculifera Cameron, 1945: 63 (Eleusis; Type locality: Bengal: Kalimpong, Samsingh, altitude 1800 feet). DISTRIBUTION: India. speculipennis Bernhauer, 1926d: 248 (Eleusis; Type locality: Luzon, Provinz Mountain, Balbalasang). — Scheerpeltz, 1933: 999 (Eleusis; catalog). DISTRIBUTION: Philippines. splendida Bernhauer, 1926d: 252 (Eleusis; Type locality: Luzon, Burauen. Mindanao, Provinz Zamboanga, Port Banga: Provinz Lanao, Mumungan). — Scheerpeltz, 1933: 999 (Eleusis; catalog). DISTRIBUTION: Philippines. strigiceps Bernhauer, 1926d: 249 (Eleusis; Type locality: Luzon, Provinz Mountain, Lubuagan: Provinz Camarines Sur, Mount Isarog; Provinz Laguna, Los Baños: Provinz Ilocos Norte, Bangui. Mindanao, Provinz Lanao, Mumungan). — Scheerpeltz, 1933: 999 (Eleusis; catalog). DISTRIBUTION: Philippines. strigicollis Bernhauer, 1914c: 77 (Eleusis; Type locality: Sumatra: Si-Rambé). — Cameron, 1928d: 91 (Eleusis; Sumatra). — Scheerpeltz, 1933: 999 (Eleusis; catalog). DISTRIBUTION: Indonesia. subplagiata Coiffait, 1982: 134 (Eleusis; Type locality: Nepal central: Trisuli Tal, zw. Ramche und Dunche, 1800–2000 m). DISTRIBUTION: Nepal. subtilepunctata Scheerpeltz, 1965: 130 (Eleusis; Type locality: N.E. Burma, Kambaiti, 7000 ft.). DISTRIBUTION: Myanmar. 1090 OSORIINAE: ELEUSIS Bulletin AMNH subtilis Sharp, 1889, see: humilis Erichson, 1840. sulciceps Bernhauer, 1926d: 247 (Eleusis; Type locality: Luzon, Mount Banahao; Provinz Nueva Vizcaya, Imugan). — Scheerpeltz, 1933: 999 (Eleusis; catalog). DISTRIBUTION: Philippines. sulcicollis Scheerpeltz, 1956a: 480, 493, 499 (Eleusis; Type locality: Belgisch-Kongo: Hembe Bitale. Also cited from French Congo). DISTRIBUTION: French Equatorial Africa, Congo. suturalis Cameron, 1940c: 210 (Eleusis; Type locality: Ghum District: Tiger Hills, alt. 8,500–10,000 feet). DISTRIBUTION: India. teestaensis Herman, 2001: 39 (Eleusis; replacement name for sikkimensis Scheerpeltz). DISTRIBUTION: India. sikkimensis Scheerpeltz, 1965: 128 (Eleusis; [preoccupied]; Type locality: British Sikkim, 200 m, Valley at Tista Bridge). — Herman, 2001: 39 (Eleusis; synonym of teestaensis). tenuis Fauvel, 1865, see: humilis Erichson, 1840. tenuis Walker, 1859, see: humilis Erichson, 1840. terminata Fauvel, 1869: 494 (Eleusis; replacement name for apicipennis Kraatz). — Bernhauer, 1926d: 251 (Eleusis; characters in key). — Herman, 2001: 22 (Eleusis; synonymic notes). DISTRIBUTION: Seychelles, Sri Lanka, Malaysia, Singapore, Indonesia, Taiwan, Japan, China. apicipennis Kraatz, 1859: 183 (Isomalus; [preoccupied]; Type locality: Ceylan). — Bernhauer and Schubert, 1910: 11 (Eleusis; synonym of kraatzi). — Bernhauer, 1922b: 165 (Eleusis; synonym of kraatzi). — Cameron, 1930: 85 (Eleusis; synonym of kraatzi). — Scheerpeltz, 1957c: 222 (Eleusis; synonym of kraatzi). kraatzi Fauvel, 1878f: 207 (Eleusis; replacement name for apicipennis Kraatz). — Bernhauer, 1902b: 45 (Eleusis; Ceylon). — Fauvel, 1905e: 78 (Eleusis; Java; Celebes; Lombok; Engano Island; Nias Island; Sumatra; China; Ceylon; Woodlark Island). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Cameron, 1921a: 362, 398 (Eleusis; characters; Singapore). — Bernhauer, 1922a: 149 (Eleusis; Indochina). — Bernhauer, 1922b: 165 (Eleusis; Seychelles). — Bernhauer, 1922c: 221 (Eleusis; Formosa). — Cameron, 1928d: 92 (Eleusis; illustrations; Sumatra). — Bernhauer, 1928: 1 (Eleusis; Sumatra). — Cameron, 1930: 85 (Eleusis; characters; Ceylon; Malay Peninsula; Singapore; Java). — Scheerpeltz, 1933: 997 (Eleusis; catalog). — Cameron, 1936a: 25 (Eleusis; Java). — Cameron, 1949b: 456 (Eleusis; Formosa). — Scheerpeltz, 1957c: 222 (Eleusis; Sumba). — Jarrige, 1963a: 120 (Eleusis; Ceylon). — Frank, 1982: 6 (Eleusis; list of parasites). — Naomi, 1996: 49, 51 (Eleusis; characters; Japan). — Herman, 2001: 22 (Eleusis; synonym of terminata). testacea Erichson, 1840: 840 (Isomalus; Type locality: Madagascar). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1961g: 244 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. Bulletin AMNH OSORIINAE: ELEUSIS 1091 tibialis Laporte, 1835: 131 (Eleusis; Type locality: Madagascar). — Laporte, 1840: 186 (Eleusis; characters; Madagascar). — Fauvel, 1865: 36 (Isomalus; characters; Madagascar). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Scheerpeltz, 1961g: 241 (Eleusis; characters; Madagascar). DISTRIBUTION: Madagascar. tonkinensis Cameron, 1939a: 22 (Eleusis; Type locality: Tonkin: Hoa Binh). DISTRIBUTION: Vietnam. toxopei Cameron, 1952: 241 (Eleusis; Type locality: New Guinea, Snow Mountains, Moss Forest Camp, 2600 m). DISTRIBUTION: New Guinea. transversiceps Scheerpeltz, 1935a: 595 (Eleusis; Type locality: Tjisaroea, Java). DISTRIBUTION: Indonesia. turneri Bernhauer, 1940: 129 (Eleusis; Type locality: Kenya: Mt. Kinangop, bamboo zone). DISTRIBUTION: Kenya. unicolor Cameron, 1942b: 321 (Eleusis; Type locality: Chyulu Hills, altitude 5600 feet). DISTRIBUTION: Kenya. usambarae Bernhauer, 1912f: 177 (Eleusis; Type locality: Usambara). — Scheerpeltz, 1933: 999 (Eleusis; catalog). — Scheerpeltz, 1956a: 489 (Eleusis; characters; Usambara). DISTRIBUTION: Tanzania. ustulatipennis Perroud and Montrousier, 1864 [Note: Described in Isomalus and transferred to “Clavicornes Ino ?” by Fauvel, 1869: 494], see: Inopeplidae. validicornis Scheerpeltz, 1956a: 481, 496, 505 (Eleusis; Type locality: Belgisch-Kongo: Hembe Bitale. Also cited from French Congo). — Scheerpeltz, 1956a: 496 (Eleusis; characters; French Congo; Belgian Congo). DISTRIBUTION: French Equatorial Africa, Congo. vermicollis Bernhauer, 1929b: 81 (Eleusis; Type locality: Haut Uele: Watsa). — Scheerpeltz, 1933: 999 (Eleusis; catalog). — Scheerpeltz, 1956a: 490 (Eleusis; characters; French Congo; Belgian Congo). DISTRIBUTION: French Equatorial Africa, Congo. vermicularis Fauvel, 1895: 189 (Eleusis; Type locality: Bírmanie, Carin Cheba, 900–1100 m). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Cameron, 1930: 77 (Eleusis; characters; Burma). — Scheerpeltz, 1933: 999 (Eleusis; catalog). DISTRIBUTION: Myanmar. virgula Fauvel, 1889, see: Zeoleusis. viridans Fauvel, 1895: 189 (Eleusis; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Asciuii Cheba, 1200–1300 m; Tenasserim, Thagata; Java, Tcibodas; Penga- lengan, 1300 m). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Cameron, 1928d: 92 (Eleusis; illustrations; Sumatra). — Bernhauer, 1928: 1 (Eleusis; Sumatra). — Cameron, 1930: 82 (Eleusis; characters; Burma; Assam; Java; Lombok). — Scheerpeltz, 1933: 999 (Eleusis; catalog). — Abdullah and Qadri, 1970: 118 (Eleusis; characters; East Pakistan). DISTRIBUTION: India, Bangladesh, Myanmar, Indonesia. nova Bernhauer, 1926: 19 (Eleusis; Type locality: Nordost-Indien: Kobo, 400 ft.). — Cameron, 1930: 82 (Eleusis; synonym of viridans). — Scheerpeltz, 1933: 998 (Eleusis; synonym of viridans). 1092 OSORIINAE: ELEUSIS Bulletin AMNH warburgi Fauvel, 1904b: 82 (Eleusis; Type locality: Celebes: Minahassa). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). DISTRIBUTION: Indonesia. Renardia [5 species; Nearctic and Neotropical Regions] Renardia Motschulsky, 1865: 583 (species included: jubilaea). TYPE SPECIES: Renardia jubilaea Motschulsky, fixed by monotypy. — Leng, 1920: 95 (catalog of North American species). — Scheerpeltz, 1933: 1072 (world catalog supplement). — Blackwelder, 1952: 338 (type species: jubilaea). — Moore and Legner, 1974b: 82 (characters; notes). — Moore and Legner, 1975: 170 (catalog; North America north of Mexico). — Moore and Legner, 1979: 148 (characters; notes). — Downie and Arnett, 1996: 428 (characters; key to species of Northeastern North America). — Newton, Thayer, Ashe, and Chandler, 2000: 377 (2 Nearctic species; notes; characters in key). Eumalus Sharp, 1887: 732 (species included: strigosus; nigriceps). TYPE SPECIES: Eumalus nigriceps Sharp, fixed by subsequent designation by Lucas, 1920: 281. — Bernhauer and Schubert, 1910: 10 (world catalog; 3 species). — Leng, 1920: 93 (catalog of North American species). — Scheerpeltz, 1933: 996 (world catalog supplement). — Blackwelder, 1942: 88 (type species: strigosus). — Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies). — Steel, 1950a: 215 (characters in key). — Blackwelder, 1952: 156 (type species). — Hatch, 1957: 244 (characters; species of Pacific Northwest). — Hammond, 1970: 69, 70 (synonym of Renardia). canadensis Horn, 1871: 297, 299 (Isomalus; Type locality: Grimsby, Ontario). — Bernhauer and Schubert, 1910: 10 (Eumalus; synonym of nigrella). — Moore and Legner, 1975: 170 (Renardia; cited as valid species; catalog). — Downie and Arnett, 1996: 428 (Renardia; characters; USA). DISTRIBUTION: Canada. nigrella LeConte, 1863a: 59 (Isomalus; Type locality: California, San Jose and Fort Tejon). — Horn, 1871: 297, 298, 299 (Isomalus; characters; California). — Fauvel, 1878d: 182 [= 1878e: 18] (Eleusis; characters; Canada; California; Texas). — Sharp, 1887: 733 (Eumalus). — Bernhauer and Schubert, 1910: 10 (Eumalus; catalog). — Hatch, 1957: 244 (Eumalus; characters; British Columbia; Washington; Oregon; Idaho). — Legner and Moore, 1977: 174 (Renardia; found under bark). — Downie and Arnett, 1996: 428 (Renardia; characters; Canada; USA). DISTRIBUTION: Canada, USA. jubilaea Motschulsky, 1865: 584 (Renardia; Type locality: environs de New York; [Note: See lectotype designation by Hammond, 1970]). — Scheerpeltz, 1933: 1072 (Renardia; catalog). — Hammond, 1970: 69, 70 (Renardia; lectotype designation; notes; synonym of nigrella). jubilaea Motschulsky, 1865, see: nigrella LeConte, 1863. nigriceps Sharp, 1887: 733 (Eumalus; Type locality: Guatemala, Capetillo). — Bernhauer and Schubert, 1910: 10 (Eumalus; catalog). DISTRIBUTION: Guatemala. strigosus Sharp, 1887: 733 (Eumalus; Type locality: Guatemala, Totonicapam 8500 to 10,500 feet). — Bernhauer and Schubert, 1910: 10 (Eumalus; catalog). DISTRIBUTION: Guatemala. Bulletin AMNH OSORIINAE: RENARDIA 1093 tenenbaumi Bernhauer, 1929h: 186 (Eumalus; Type locality: Mexiko: Desierto de los Leones). — Scheerpeltz, 1933: 996 (Eumalus; catalog). DISTRIBUTION: Mexico. Zeoleusis [12 species; Neotropical, Oriental, and Australian Regions] Zeoleusis Steel, 1950a: 215 (species included: virgula, semirufus, insularis, sororcula, polita, annae, parva, banghaasi, helferi, exigua, fusciceps). TYPE SPECIES: Eleusis virgula Fauvel, fixed by original designation. — Steel, 1950a: 219 (key to species). — Coiffait and Saiz, 1968: 462 (characters; key to Chilean species). — Newton, 1985: 203 (notes on austral distribution). — Newton, 1990: 91 (characters in key; two described Australian species). annae Steel, 1950a: 225 (Zeoleusis; Type locality: New South Wales: Sydney). — Newton, 1990: 91 (Zeoleusis). DISTRIBUTION: Australia. banghaasi Bernhauer, 1926d: 253 (Eleusis; Type locality: Siargao, Dapa. Palawan, Binaluan. Luzon, Provinz Mountain, Balbalasang. Mindanao, Provinz Lanao, Mumungan. Masbate, Aroroy). — Scheerpeltz, 1933: 996 (Eleusis; catalog). — Steel, 1950a: 231 (Zeoleusis; characters; Philippines Islands). DISTRIBUTION: Philippines. decipiens Cameron, 1932, see: exigua Bernhauer, 1926. exigua Bernhauer, 1926d: 253 (Eleusis; variety of banghaasi; Type locality: Not cited). — Scheerpeltz, 1933: 996 (Eleusis; synonym of augustae). — Steel, 1950a: 233 (Zeoleusis; cited as valid species; characters; Philippines; Malaysia). DISTRIBUTION: Philippines, Malaysia. decipiens Cameron, 1932a: 124 (Eleusis; Type locality: Pahang: Fraser’s Hill, alt. 4000 feet). — Steel, 1950a: 232 (Zeoleusis; synonym of exigua). fusciceps Kraatz, 1859: 184 (Isomalus; Type locality: Ceylan). — Bernhauer and Schubert, 1910: 11 (Eleusis; catalog). — Cameron, 1921a: 362, 398 (Eleusis; characters; Singapore). — Bernhauer, 1926d: 253 (Eleusis; characters; Philippines). — Cameron, 1930: 88 (Eleusis; characters; Ceylon; Andamans; Siwaliks; Singapore; Sumatra). — Cameron, 1930b: 160 (Eleusis; British North Borneo). — Scheerpeltz, 1933: 997 (Eleusis; catalog). — Steel, 1950a: 233 (Zeoleusis; characters; Ceylon; India; Singapore; Sumatra; Philippines). — Jarrige, 1963a: 120 (Eleusis; Ceylon). — Abdullah and Qadri, 1970: 116 (Eleusis; characters; West Pakistan). — Coiffait, 1981f: 331 (Eleusis; cited as fuscipes; Andaman Islands). — Naomi, 1996: 51 (Zeoleusis; Japan). DISTRIBUTION: Pakistan, India, Bangladesh, Sri Lanka, Andaman Islands, Borneo, Singapore, Indonesia, Philippines, Japan. helferi Steel, 1950a: 230 (Zeoleusis; Type locality: Bosnek, Biak Island, New Guinea). DISTRIBUTION: New Guinea. insularis Cameron, 1936: 3 (Eleusis; Type locality: Rapa, Mount Vaitau, altitude 1100 feet). — Steel, 1950a: 221 (Zeoleusis; characters; Rapa Island). DISTRIBUTION: Austral Islands. myrmidon Fairmaire and Germain, 1861: 455 (Isomalus; Type locality: Santiago). — Fauvel, 1867a: 53 [= 1868: 58] (Isomalus; characters; Chile). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Coiffait and Saiz, 1968: 463 (Zeoleusis; characters; Chile). DISTRIBUTION: Chile. 1094 OSORIINAE: ZEOLEUSIS Bulletin AMNH parva Blackburn, 1892: 24 (Eleusis; Type locality: New South Wales, Blue Mountains). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Steel, 1950a: 224 (Zeoleusis; characters; Australia). DISTRIBUTION: Australia. polita Steel, 1950a: 227 (Zeoleusis; Type locality: Ceylon). DISTRIBUTION: Sri Lanka. semirufa Fairmaire and Germain, 1861: 455 (Isomalus; Type locality: Ile de Juan Fernandez). — Fauvel, 1867a: 53 [= 1868: 58] (Isomalus; characters; Chile). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Bernhauer, 1921a: 41 (Eleusis; Juan Fernandez). — Scheerpeltz, 1933: 998 (Eleusis; catalog). — Steel, 1950a: 223 (Zeoleusis; characters; Juan Fernandez). — Coiffait and Saiz, 1968: 463 (Zeoleusis; characters; Chile). DISTRIBUTION: Chile, Juan Fernandez. sororcula Cameron, 1936: 4 (Eleusis; Type locality: Rapa, Mount Vaitau, altitude, 1100 feet). — Steel, 1950a: 222 (Zeoleusis; characters; Rapa Island). DISTRIBUTION: Austral Islands. virgula Fauvel, 1889b: 243 (Eleusis; Type locality: Nouvelle-Zelande: Auckland; Greymouth). — Bernhauer and Schubert, 1910: 12 (Eleusis; catalog). — Steel, 1950a: 228 (Zeoleusis; characters; New Zealand). — Kuschel, 1990: 26 (Zeoleusis; endemic; winged; habitat; New Zealand). — Emberson, 1998: 35 (Zeoleusis; collecting notes; Chatham Islands). DISTRIBUTION: New Zealand, Chatham Islands.
Tribe Leptochirini Leptochirini Sharp, 1887. — Eichelbaum, 1909: 110 (list of genera with some references; generic synonyms; number of genera; number of species in each genus; general distribution of genera). — Cameron, 1930: 89 (characters; key to British Indian genera). — Blackwelder, 1942: 79 (characters). — Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies). — Newton and Thayer, 1992: 59 (tribe of Osoriinae). Leptochiri Sharp, 1887. — Bernhauer and Schubert, 1910: 13 (world catalog; Leptochirus, Borolinus, Priochirus, Thoracochirus). — Scheerpeltz, 1933: 999 (world catalog supplement). Leptochirina Sharp, 1887: 733 (genera included: Leptochirus, Priochirus). TYPE GENUS: Leptochirus (see Newton and Thayer, 1992: 59). — Jakobson, 1908: 446, 466 (characters; list of species of Russia and western Europe).
Borolinus [12 species; Palaearctic, Oriental, Australian, and Oceanic Regions] Borolinus Bernhauer, 1903: 116, 133 (species included: semirufus, antilope, minutus, javanicus, borneensis). TYPE SPECIES: Leptochirus javanicus Laporte, fixed by subsequent designation by R. Lucas, 1920: 144. — Bernhauer and Schubert, 1910: 15 (world catalog; 5 species). — Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog). — Cameron, 1925: 4 (catalog of Indian species). — Cameron, 1930: 92 (characters; key to British Indian species). — Scheerpeltz, 1933: 1001 (world catalog supplement). — Blackwelder, 1952: 84 (type species). Bulletin AMNH OSORIINAE: BOROLINUS 1095
— Shibata, 1973c: 25 (checklist of species of Taiwan). — Shibata, 1976: 87 (checklist of species of Japan). — Hammond, 1984: 196 (checklist of species of Borneo). — Drugmand, 1998a: 206 (list of types in l’Institut royal des Sciences naturelles de Belgique). antilope Fauvel, 1895: 181 (Leptochirus; Type locality: Bírmanie, Carin Ghécu, 1300– 1400 m; Senmijion). — Heller, 1898: 6 (Priochirus; checklist). — Bernhauer, 1903: 134 (Borolinus; characters). — Bernhauer and Schubert, 1910: 15 (Borolinus; catalog). — Cameron, 1928d: 94 (Borolinus; Sumatra). — Cameron, 1930: 93 (Borolinus; characters; Burma). — Scheerpeltz, 1933: 1001 (Borolinus; catalog; Siam). DISTRIBUTION: Myanmar, Thailand, Indonesia. nigripennis Fauvel, 1902: 10 (Leptochirus; subgenus Priochirus; variety of antilope; Type locality: Sumatra: Si-Rambé). — Bernhauer and Schubert, 1910: 15 (Borolinus; synonym of antilope). — Cameron, 1928d: 94 (Borolinus; variety of antilope; Sumatra). — Cameron, 1936a: 26 (Leptochirus; subgenus Strongylochirus; variety of antilope; Java). bispinus Erichson, 1840, see: minutus Laporte, 1840. borneensis Bernhauer, 1903: 133 (Borolinus; Type locality: Borneo). — Fauvel, 1905e: 77 (Leptochirus; synonym of minutus). — Bernhauer and Schubert, 1910: 15 (Borolinus; catalog). — Wendeler, 1928c: 117 (Borolinus; Philippines). DISTRIBUTION: Borneo, Philippines. cruentus Fauvel, 1895, see: minutus Laporte, 1840. curticollis Bernhauer, 1926b: 38 (Borolinus; Type locality: Philippinen: Mindanao, Iligan). — Wendeler, 1928c: 117 (Borolinus; Philippines). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). DISTRIBUTION: Philippines. furcatus Wendeler, 1928c: 119 (Borolinus; Type locality: Philippinen: Luzon: Imugan, Mt. Isarog). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). DISTRIBUTION: Philippines. inermis Fauvel, 1902, see: minutus Laporte, 1840. javanicus Laporte, 1835: 126 (Leptochirus; Type locality: Java). — Erichson, 1840: 827 (Leptochirus; characters; Java). — Fauvel, 1886b: 144 (Leptochirus; Philippines; Java; Sumatra). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 134 (Borolinus; characters). — Bernhauer and Schubert, 1910: 15 (Borolinus; catalog). — Cameron, 1928d: 94 (Borolinus; Sumatra). — Bernhauer, 1928: 1 (Borolinus; Sumatra). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). — Frank, 1982: 6 (Borolinus; list of parasites). DISTRIBUTION: Philippines, Indonesia. nigricollis Fauvel, 1902: 10 (Leptochirus; subgenus Priochirus; variety of javanicus; Type locality: nord de Luçon). — Bernhauer and Schubert, 1910: 15 (Borolinus; synonym of javanicus). — Scheerpeltz, 1933: 1001 (Borolinus; synonym of javanicus). minutus Laporte, 1840: 186 (Leptochirus; Type locality: Java). — Fauvel, 1878f: 186 (Leptochirus; characters; New Guinea; Java; Borneo; Philippines). — Fauvel, 1879: 72 (Leptochirus; Sumatra, Malacca; Siam; Dorei). 1096 OSORIINAE: BOROLINUS Bulletin AMNH
— Fauvel, 1886b: 144 (Leptochirus; Philippines; Siam; Malacca; Borneo; Java; Gilolo; New Guinea). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 134 (Borolinus; characters). — Fauvel, 1905e: 77 (Leptochirus; Java). — Bernhauer and Schubert, 1910: 15 (Borolinus; catalog). — Bernhauer, 1922c: 221 (Borolinus; Formosa). — Cameron, 1928d: 94 (Borolinus; Sumatra). — Bernhauer, 1928: 1 (Borolinus; Sumatra). — Wendeler, 1928c: 117 (Borolinus; Philippines). — Cameron, 1930: 94 (Borolinus; characters; Assam; Sikkim; Burma; Siam; Sumatra; Java; Lombok; Borneo). — Cameron, 1930b: 160 (Priochirus; British North Borneo). — Cameron, 1931: 355 (Borolinus; New Britain). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). — Cameron, 1936a: 26 (Leptochirus; subgenus Strongylochirus; Java). — Cameron, 1937a: 84 (Borolinus; New Guinea). — Paulian, 1941: 132 (Borolinus; larval characters; India). — Cameron, 1949b: 456 (Borolinus; China). — Scheerpeltz, 1965: 98 (Borolinus; Burma). — Abdullah and Qadri, 1970: 116 (Borolinus; characters; East Pakistan). — Coiffait, 1981f: 332 (Borolinus; Andaman Islands). — Frank, 1982: 6 (Borolinus; list of parasites). — Biswas and Biswas, 1985: 142 (Borolinus; India). DISTRIBUTION: China, Taiwan, New Guinea, New Britain, Philippines, Borneo, Indo- nesia, Singapore, Malaysia, Thailand, Myanmar, Andaman Islands, Bangladesh, India. bispinus Erichson, 1840: 827 (Leptochirus; Type locality: Java). — Fauvel, 1878f: 186 (Leptochirus; synonym of minutus). — Fauvel, 1905e: 77 (Leptochirus; synonym of minutus). — Bernhauer and Schubert, 1910: 15 (Borolinus; synonym of minutus). — Cameron, 1930: 94 (Borolinus; synonym of minutus). — Scheerpeltz, 1965: 98 (Borolinus; synonym of minutus). cruentus Fauvel, 1895: 181 (Leptochirus; variety of minutus; Type locality: Malacca; Singapore; Sumatra; Borneo). — Heller, 1898: 8 (Priochirus; variety of minutus; checklist). — Bernhauer and Schubert, 1910: 15 (Borolinus; synonym of minutus). — Cameron, 1921a: 362, 399 (Borolinus; variety of minutus; characters; Singapore). — Cameron, 1928d: 94 (Borolinus; variety of minutus; Sumatra). — Scheerpeltz, 1933: 1001 (Borolinus; variety of minutus). inermis Fauvel, 1902: 10 (Leptochirus; subgenus Priochirus; variety of minutus; Type locality: Sumatra: Palembang). — Bernhauer and Schubert, 1910: 15 (Borolinus; synonym of minutus). nigricollis Fauvel, 1902, see: javanicus Castelnau, 1835. nigripennis Fauvel, 1902, see: antilope Fauvel, 1895. oceani Bernhauer, 1919: 360 (Borolinus; Type locality: Philippinen: Magdalena, Laguna). — Wendeler, 1928c: 117 (Borolinus; Philippines). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). DISTRIBUTION: Philippines. rufus Cameron, 1928g: 424 (Borolinus; Type locality: Borneo: Mount Dulit, 3500 feet). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). DISTRIBUTION: Borneo. samarensis Bernhauer, 1928: 2 (Borolinus; Type locality: Island Samar). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). DISTRIBUTION: Philippines. Bulletin AMNH OSORIINAE: BOROLINUS 1097 semirufus Fauvel, 1895: 181 (Leptochirus; Type locality: Bírmanie, Carin Cheba, 900– 1100 m; Asciuii Cheba). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 134 (Borolinus; characters). — Bernhauer and Schubert, 1910: 15 (Borolinus; catalog). — Cameron, 1930: 93 (Borolinus; characters; Burma). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). DISTRIBUTION: Myanmar. sikkimensis Bernhauer, 1919: 359 (Borolinus; Type locality: Himalaya: Sikkim). — Cameron, 1930: 94 (Borolinus; characters; Sikkim; Burma). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). — Scheerpeltz, 1965: 98 (Borolinus; Burma). — Abdullah and Qadri, 1970: 116 (Borolinus; characters; East Pakistan). — Biswas and Biswas, 1985: 142 (Borolinus; India). DISTRIBUTION: India, Bangladesh, Myanmar. sumatrensis Bernhauer, 1928: 2 (Borolinus; Type locality: Sumatra: Brastagi). — Scheerpeltz, 1933: 1001 (Borolinus; catalog). DISTRIBUTION: Indonesia. Leptochirus [61 species; Neotropical, Palaearctic, and Oriental Regions] Leptochirus Germar, 1824: 35 (species included: scoriaceus). TYPE SPECIES: Leptochirus scoriaceus Germar, fixed by monotypy. — Latreille, 1825: 245 (cited as Leptochire; synonym of Zirophore). — Berthold, 1827: 332 (characters). — Erichson, 1840: 824 (characters). — Laporte, 1840: 186 (characters). — Lacordaire, 1854: 125 (characters; notes; list of species). — Fauvel, 1864b: 8 (characters; notes; list of species; larval characters). — Fauvel, 1865: 12 (characters; notes; list of species; larval characters). — Gemminger and Harold, 1868: 678 (catalog). — Sharp, 1876: 420 (characters; notes). — Fauvel, 1877: 184 (notes; key to species). — Duvivier, 1883: 205 (catalog). — Sharp, 1887: 733 (characters; Mexican and Central species). — Heller, 1898: 5, 6, 8 (discussion; checklist of and key to species). — Bernhauer, 1903: 116, 117 (characters; key to species). — Bernhauer and Schubert, 1910: 13 (world catalog; 35 species). — Sharp and Muir, 1912: 501 (characters of male genitalia). — Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog). — Cameron, 1925: 3 (catalog of Indian species). — Cameron, 1930: 89 (characters; key to British Indian species). — Scheerpeltz, 1933: 999 (world catalog supplement). — Paulian, 1941: 128 (larval characters). — Blackwelder, 1943: 162 (characters; type species: scoriaceus; key to species of West Indies). — Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 215 (type species: scoriaceus). — Hammond, 1984: 196 (checklist of species of Borneo). — Drugmand, 1998a: 212 (list of types in l’Institut royal des Sciences naturelles de Belgique). Mesochirus Bernhauer, 1903: 120 (subgenus of Leptochirus; species included: extimus, maxillosus, latro, brunneoniger, cayennensis, arcifer, tenuis). TYPE SPECIES: Cucujus maxillosus Fabricius, fixed by subsequent designation by R. Lucas, 1920: 405. — Bernhauer and Schubert, 1910: 14 (catalog; subgenus of Leptochirus). — Blackwelder, 1943: 162 (subgenus of Leptochirus; type species). — Blackwelder, 1952: 239 (type species). 1098 OSORIINAE: LEPTOCHIRUS Bulletin AMNH
Tropiochirus Bernhauer, 1903: 120 (subgenus of Leptochirus; species included: helleri, proteus). TYPE SPECIES: Leptochirus proteus Fauvel, fixed by subsequent designation by R. Lucas, 1920: 659. — Bernhauer and Schubert, 1910: 14 (catalog; subgenus of Leptochirus). — Scheerpeltz, 1933: 1000 (world catalog supplement; subgenus of Leptochirus). — Blackwelder, 1943: 162 (subgenus of Leptochirus; type species). — Blackwelder, 1952: 399 (subgenus of Leptochirus; type species). Strongylochirus Bernhauer, 1903: 120 (subgenus of Leptochirus; species included: quadri- dens, loriae, laevis, atkinsoni). TYPE SPECIES: Leptochirus laevis Laporte, fixed by subsequent designation by Lucas, 1920: 618. — Bernhauer and Schubert, 1910: 14 (catalog; subgenus of Leptochirus). — Scheerpeltz, 1933: 1000 (world catalog supplement; subgenus of Leptochirus). — Blackwelder, 1943: 162 (subgenus of Leptochirus; type species laevis). — Blackwelder, 1952: 364 (type species). — Biswas and Sengupta, 1977: 341 (subgenus of Leptochirus; key to species of India). albertisi Fauvel, 1878, see: Priochirus. alternus Fauvel, 1878, see: Priochirus. alticola Bernhauer, 1912d: 28 (Leptochirus; Type locality: Columbia: Cali: S. Antonio, 2000 m; Rio Aquatal, 1880 m). — Scheerpeltz, 1933: 999 (Leptochirus; catalog). DISTRIBUTION: Colombia. andinus Bernhauer, 1917f: 46 (Leptochirus; subgenus Leptochirus; Type locality: West- Kolumbien: Pacho in den Ost-Kordilleren, 1900 m). — Scheerpeltz, 1933: 999 (Leptochirus; catalog). DISTRIBUTION: Colombia. angustulus Sharp, 1887: 736 (Leptochirus; Type locality: Colombia, Venezuela). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 121 (Leptochirus; characters). — Bernhauer, 1903: 122 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Colombia, Venezuela. antennarius Fauvel, 1878, see: Priochirus. anthracinus Fauvel, 1902, see: maxillosus Fabricius, 1801. antilope Fauvel, 1895, see: Borolinus. apicalis Eppelsheim, 1895, see: Priochirus. arcifer Fauvel, 1902: 17 (Leptochirus; Type locality: Bolivie: Yuracaris, Yungas; Pérou: Marcapata). — Bernhauer, 1903: 125, 131 (Leptochirus; characters; Peru; Bolivia). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Mesochirus; catalog). DISTRIBUTION: Bolivia, Peru. ascendens Fauvel, 1903, see: Priochirus. aspericollis Fauvel, 1902, see: Thoracochirus. atkinsoni Fauvel, 1895: 182 (Leptochirus; Type locality: Bírmanie, Carin Cheba, 900–1400 m; Tenasserim, Tavoy). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 126 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Strongylochirus; catalog). — Cameron, 1930: 92 (Leptochirus; subgenus Strongylochirus; characters; Assam; Burma; Siam). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Strongylochirus; catalog). — Abdullah and Qadri, 1970: 116 (Leptochirus; subgenus Strongylochirus; characters; East Pakistan). Bulletin AMNH OSORIINAE: LEPTOCHIRUS 1099
— Biswas and Sengupta, 1977: 345 (Leptochirus; subgenus Strongylochirus; characters; Burma; India). — Biswas and Biswas, 1985: 142 (Leptochirus; subgenus Strongylochirus; India). DISTRIBUTION: India, Bangladesh, Myanmar, Thailand. beccarii Fauvel, 1878, see: Priochirus. bernhaueri Wendeler, 1957a: 47 (Leptochirus; replacement name for sharpi Bernhauer). DISTRIBUTION: Mexico. mexicanus Sharp, 1887: 739 (Leptochirus; [preoccupied]; Type locality: Mexico, Cordova). — Bernhauer and Schubert, 1910: 14 (Leptochirus; synonym of sharpi). — Wendeler, 1957a: 47 (Leptochirus; synonym of bernhaueri). sharpi Bernhauer, 1903: 123, 130 (Leptochirus; [preoccupied; replacement name for mexicanus Sharp; Note: Erroneously cited by Bernhauer and Schubert, 1910: 14 as a subsequent reference of sharpi Fauvel]). — Wendeler, 1957a: 47 (Leptochirus; synonym of bernhaueri). bicolor P. Lucas, 1857, see: maxillosus Fabricius, 1801. bicornis Fauvel, 1864, see: Priochirus. bifoveatus Eppelsheim, 1895, see: Priochirus. bifurcatus Fauvel, 1879, see: Priochirus. bipunctatus Fauvel, 1895, see: Priochirus. bispinus Erichson, 1840, see: Borolinus. bolivianus Heller, 1898, see: maxillosus Fabricius, 1801. brachycerus Kraatz, 1859, see: Priochirus. brasilianus Bernhauer, 1903: 121, 128 (Leptochirus; Type locality: Brasilia). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Brazil. brevicornis Erichson, 1840, see: Priochirus. brunneoniger Perty, 1830: 32 (Leptochirus; Type locality: Brasil: Prov. Minarum). — Erichson, 1840: 825 (Leptochirus; synonym of maxillosus). — Fauvel, 1864b: 13 (Leptochirus; characters; Mexico, French Guiana; Brazil). — Fauvel, 1865: 17 (Leptochirus; characters; Mexico; French Guiana; Brazil). — Solsky, 1868a: 144 (Leptochirus; Mexico). — Solsky, 1872: 314 (Leptochirus; Colombia). — Sharp, 1876: 421 (Leptochirus; Brazil). — Fauvel, 1877a: xxiv (Leptochirus; Brazil). — Sharp, 1887: 734 (Leptochirus; notes; Mexico). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 124 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Mesochirus; catalog). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Mesochirus; catalog). — Scheerpeltz, 1951a: 9 (Leptochirus; subgenus Mesochirus; Bolivia). DISTRIBUTION: Bolivia, Brazil, French Guiana, Colombia, Mexico. cayennensis Laporte, 1835: 125 (Leptochirus; Type locality: Cayenne). — Erichson, 1840: 825 (Leptochirus; synonym of maxillosus). — Fauvel, 1864b: 14 (Leptochirus; synonym of brunneoniger). — Fauvel, 1865: 18 (Leptochirus; synonym of brunneoniger). — Sharp, 1887: 734 (Leptochirus; synonym of brunneoniger). — Bernhauer and Schubert, 1910: 14 (Leptochirus; synonym of brunneoniger). buergeri Wendeler, 1957a: 46 (Leptochirus; subgenus Leptochirus; Type locality: Columbien: Salta di Tequendama). DISTRIBUTION: Colombia. 1100 OSORIINAE: LEPTOCHIRUS Bulletin AMNH caviceps Eppelsheim, 1895, see: Priochirus. cavifrons Fauvel, 1878, see: Priochirus. cayennensis Laporte, 1835, see: brunneoniger Perty, 1830. centralis Fauvel, 1902, see: Priochirus. cephalotes Sharp, 1887: 740 (Leptochirus; Type locality: Mexico, Guanajuato). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 123 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Mexico. ceylanensis Fauvel, 1902, see: Priochirus. collaris Fauvel, 1902, see: maxillosus Fabricius, 1801. collenettei Bernhauer, 1934e: 156 (Leptochirus; Type locality: Kolumbien: Gorgona Island). DISTRIBUTION: Colombia. columbicus Bernhauer, 1912d: 27 (Leptochirus; Type locality: Columbia: Cali: Rio Aquatal, 1600–1800 m; S. Antonio, 2000 m; Rio Vitaco, 2000 m; Villa Elvira, 1800 m). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Colombia. combustus Fauvel, 1902, see: Priochirus. comes Fauvel, 1902, see: Priochirus. condei Wendeler, 1955: 188 (Leptochirus; Type locality: Brasilien: Espirito Santo, Sta. Thereza). DISTRIBUTION: Brazil. conicicollis Fauvel, 1878, see: Priochrus. consentaneus Wendeler, 1957a: 45 (Leptochirus; subgenus Leptochirus; Type locality: Süd Peru). DISTRIBUTION: Peru. convexus Laporte, 1835, see: Priochirus. coriaceus Gray, 1832, see: scoriaceus Germar, 1824. coronatus Sachse, 1852, see: Priochirus. costaricensis Wendeler, 1927: 2 (Leptochirus; Type locality: Costa Rica: Turrialba, 800 m). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Costa Rica. costaricensis Bernhauer, 1942, see: maxi Herman, 2001. cribrellus Fauvel, 1905, see: Thoracochirus. cruentus Fauvel, 1895, see: Borolinus. davidis Fairmaire, 1889, see: Priochirus. diversus Bernhauer, 1903: 122, 129 (Leptochirus; Type locality: Guatemala; Mexiko). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Paulian, 1941: 130 (Leptochirus; larval characters; Costa Rica). DISTRIBUTION: Mexico, Guatemala, Costa Rica. dohrni Fauvel, 1902, see: Priochirus. ebeninus Erichson, 1840, see: Priochirus. edax Sharp, 1887: 736 (Leptochirus; Type locality: Mexico, Jalapa; Guatemala, El Tumbador, Cerro Zunil, San Lucas 6500 feet, Volcan de Fuego 6400 feet, Capetillo, San Geronimo, Purula, Sabo; Panama, Volcan de Chiriqui, Bugaba, Tole. Also three varieties: —Mexico, Sinaloa; —Costa Rica; —Guatemala, Dueñas). — Heller, 1898: 6 (Leptochirus; checklist). Bulletin AMNH OSORIINAE: LEPTOCHIRUS 1101
— Bernhauer, 1903: 120 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). — Paulian, 1941: 130 (Leptochirus; larval characters; Costa Rica). — Frank, 1982: 7 (Leptochirus; list of parasites). DISTRIBUTION: Mexico, Guatemala, Costa Rica, Panama. exaratus Eppelsheim, 1895, see: Priochirus. excavatus Motschulsky, 1857, see: Priochirus. eximius Wendeler, 1957a: 46 (Leptochirus; subgenus Mesochirus; Type locality: Ost-Bolivien: Prov. Sara. Also cited from Peru). DISTRIBUTION: Bolivia, Peru. extensus Fauvel, 1878, see: Priochirus. extimus Sharp, 1887: 735 (Leptochirus; Type locality: Mexico, Toxpam, Tuxtla). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 123 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Mesochirus; catalog). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Mesochirus; catalog). DISTRIBUTION: Mexico. flohri Wendeler, 1957a: 45 (Leptochirus; subgenus Leptochirus; Type locality: Mexico: Tuxtepec). DISTRIBUTION: Mexico. fontensis Sharp, 1876: 420 (Leptochirus; Type locality: Fonte Boa; Ega). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 121 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Brazil. forticornis Fauvel, 1877, see: Priochirus. galeatus Fauvel, 1902, see: Priochirus. gastralis Bernhauer, 1903: 122, 130 (Leptochirus; Type locality: Brasilien). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog; Argentina). DISTRIBUTION: Argentina, Brazil. giganteus Wendeler, 1927: 3 (Leptochirus; Type locality: Peru). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; cited as gigantulus; catalog). DISTRIBUTION: Peru. gracilis Sharp, 1887: 738 (Leptochirus; Type locality: Panama, Volcan de Chiriqui 2500 to 4000 feet). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 122 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Panama. haackei W. J. MacLeay, 1886, see: Priochirus. haeneli Bernhauer, 1910b: 351 (Leptochirus; Type locality: Mexiko). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Mexico. helleri Bernhauer, 1903: 125, 132 (Leptochirus; Type locality: Peru). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Tropiochirus; catalog). DISTRIBUTION: Peru. heterodon Fauvel, 1902, see: Priochirus. 1102 OSORIINAE: LEPTOCHIRUS Bulletin AMNH hoplites Fauvel, 1895, see: Priochirus. iconnicoffi Bernhauer, 1917f: 47 (Leptochirus; subgenus Mesochirus; Type locality: Peru: 1000 m, 11 3 südl., 75 7 ). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Mesochirus; cited as iconicoffi; catalog). DISTRIBUTION: Peru. idae Kraatz, 1859, see: laevis Laporte, 1840. incertus Bernhauer, 1903: 122, 129 (Leptochirus; Type locality: Peru). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Peru. inermis Fauvel, 1902, see: Borolinus. javanicus Laporte, 1835, see: Borolinus. kaestneri Wendeler, 1957a: 46 (Leptochirus; subgenus Leptochirus; Type locality: Mexico: Analco). DISTRIBUTION: Mexico. klimschi Bernhauer, 1907a: 281 (Leptochirus; Type locality: Brasilia, St. Catharina). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Brazil. kolbei Bernhauer, 1903: 121, 127 (Leptochirus; Type locality: Bolivien). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Bolivia. laevis Laporte, 1840: 186 (Leptochirus; Type locality: Java). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Strongylochirus; catalog). — Cameron, 1921a: 362, 399 (Leptochirus; characters; Singapore). — Cameron, 1928d: 94 (Leptochirus; subgenus Strongylochirus; Sumatra). — Bernhauer, 1928: 1 (Leptochirus; Sumatra). — Cameron, 1930: 91 (Leptochirus; subgenus Strongylochirus; characters; Assam; Siam; Singapore; Sumatra; Java; Lombok; Borneo). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Strongylochirus; catalog). — Cameron, 1936a: 26 (Leptochirus; subgenus Strongylochirus; Java). — Cameron, 1949b: 456 (Leptochirus; subgenus Strongylochirus; China). — Scheerpeltz, 1957c: 222 (Leptochirus; subgenus Strongylochirus; Sumba). — Abdullah and Qadri, 1970: 116 (Leptochirus; subgenus Strongylochirus; characters; East Pakistan). — Biswas and Sengupta, 1977: 343 (Leptochirus; subgenus Strongylochirus; characters; India). — Coiffait, 1978b: 112 (Leptochirus; subgenus Strongylochirus; Bhutan). — Biswas and Sengupta, 1982: 140 (Leptochirus; characters; India). — Biswas and Biswas, 1985: 142 (Leptochirus; subgenus Strongylochirus; India). DISTRIBUTION: India, Bhutan, Bangladesh, Thailand, Singapore, Borneo, Indonesia, China. laticeps Erichson, 1840: 826 (Leptochirus; Type locality: Java). — Bernhauer and Schubert, 1910: 14 (Leptochirus; synonym of laevis). — Cameron, 1930: 91 (Leptochirus; synonym of laevis). — Biswas and Sengupta, 1977: 343 (Leptochirus; synonym of laevis). — Biswas and Sengupta, 1982: 140 (Leptochirus; synonym of laevis). idae Kraatz, 1859: 189 note (Leptochirus; Type locality: Java). — Bernhauer and Schubert, 1910: 14 (Leptochirus; synonym of laevis). — Cameron, 1930: 91 (Leptochirus; synonym of laevis). — Biswas and Sengupta, 1977: 343 (Leptochirus; synonym of laevis). — Biswas and Sengupta, 1982: 140 (Leptochirus; synonym of laevis). laeviventris Fauvel, 1902: 16 (Leptochirus; Type locality: Bolivie: Yungas, Yuracaris, Chaco; Nouvelle Grenade; Equateur: Loja; Brésil: Santa Catharina). — Bernhauer, 1903: 121 (Leptochirus; characters; Bolivia). Bulletin AMNH OSORIINAE: LEPTOCHIRUS 1103
— Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Herman, 2001: 22 (Leptochirus; synonymic notes). DISTRIBUTION: Brazil, Bolivia, Ecuador, Colombia. laeviventris Bernhauer, 1903: 121, 126 (Leptochirus; [Note: Preoccupied]; Type locality: Bolivien). — Herman, 2001: 22 (Leptochirus; synonymic homonym of laeviventris). laeviventris Bernhauer, 1903, see: laeviventris Fauvel, 1902. laticeps Erichson, 1840, see: laevis Laporte, 1840. latro Sharp, 1876: 421 (Leptochirus; Type locality: Ega). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 124 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Mesochirus; catalog). — Scheerpeltz, 1951a: 9 (Leptochirus; subgenus Mesochirus; Bolivia). DISTRIBUTION: Brazil, Bolivia. longicornis Fauvel, 1864, see: Priochirus. loriae Heller, 1898: 14 (Leptochirus; Type locality: Penang, alt. 600–800 m). — Bernhauer, 1903: 126 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Strongylochirus; catalog). — Cameron, 1928d: 94 (Leptochirus; subgenus Strongylochirus; Sumatra). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Strongylochirus; catalog). — Cameron, 1936a: 26 (Leptochirus; subgenus Strongylochirus; Java). DISTRIBUTION: Malaysia, Indonesia. lorquini Fauvel, 1878, see: Priochirus. luzonicus Fauvel, 1886, see: Priochirus. major Bernhauer, 1903, see: quadridens Motschulsky, 1857. mandibularis Kraatz, 1859, see: Priochirus. maxi Herman, 2001: 40 (Leptochirus; replacement name for costaricensis Bernhauer). DISTRIBUTION: Costa Rica. costaricensis Bernhauer, 1942: 1 (Leptochirus; [preoccupied]; Type locality: Kostarika). — Herman, 2001: 40 (Leptochirus; synonym of maxi). maxillosus Fabricius, 1801: 93 (Cucujus; Type locality: America meridionalis). — Erichson, 1840: 825 (Leptochirus; characters; Brazil; French Guiana; Colombia; Mexico). — Fauvel, 1864b: 13 (Leptochirus; characters; Mexico, Colombia, Guyana; Brazil). — Fauvel, 1865: 18 (Leptochirus; characters; Mexico; Guyana; Colombia; Brazil). — Solsky, 1872: 314 (Leptochirus; Peru). — Sharp, 1876: 422 (Leptochirus; notes; Brazil). — Fauvel, 1877a: xxiv (Leptochirus; Brazil). — Sharp, 1887: 734 (Leptochirus; Mexico; British Honduras). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 124 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Mesochirus; catalog; Argentina). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Mesochirus; catalog). — Blackwelder, 1943: 163 (Leptochirus; characters; Trinidad). — Scheerpeltz, 1951a: 10 (Leptochirus; subgenus Mesochirus; Bolivia). DISTRIBUTION: West Indies, French Guiana, Guyana, Brazil, Argentina, Bolivia, Peru, Colombia, Nicaragua, Belize, Mexico. bicolor P. Lucas, 1857: 51 (Leptochirus; Type locality: Goyaz à Cuyaba). — Fauvel, 1877a: xxiv (Leptochirus; synonym of maxillosus). — Bernhauer, 1903: 124 (Leptochirus; aberration of maxillosus; characters). — Bernhauer and Schubert, 1910: 14 (Leptochirus; synonym of maxillosus). — Scheerpeltz, 1933: 1000 (Leptochirus; synonym of maxillosus). — Blackwelder, 1943: 163 (Leptochirus; synonym of maxillosus). 1104 OSORIINAE: LEPTOCHIRUS Bulletin AMNH
bolivianus Heller, 1898: 6, 9 (Leptochirus; variety of maxillosus; Type locality: Bolivia). — Bernhauer, 1903: 124 (Leptochirus; aberration of maxillosus; characters). — Bernhauer and Schubert, 1910: 14 (Leptochirus; synonym of maxillosus). — Blackwelder, 1943: 163 (Leptochirus; synonym of maxillosus). anthracinus Fauvel, 1902: 17 (Leptochirus; variety of maxillosus; Type locality: Mexique; Honduras; Nicaragua; Bresil; Perou). — Bernhauer and Schubert, 1910: 14 (Leptochirus; synonym of maxillosus). — Blackwelder, 1943: 163 (Leptochirus; synonym of maxillosus). collaris Fauvel, 1902: 17 (Leptochirus; variety of maxillosus; Type locality: Antilles: Trinidad; Bresil; Colombie; Bolivie; Perou). — Bernhauer and Schubert, 1910: 14 (Leptochirus; synonym of maxillosus). — Blackwelder, 1943: 163 (Leptochirus; synonym of maxillosus). mexicanus Erichson, 1840: 826 (Leptochirus; Type locality: Mexico). — Fauvel, 1864b: 15 (Leptochirus; characters; Mexico). — Fauvel, 1865: 19 (Leptochirus; characters; Mexico). — Solsky, 1868a: 144 (Leptochirus; Mexico). — Fauvel, 1891: 88 (Leptochirus; Venezuela). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 123 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). — Marcuzzi, 1977: 4 (Leptochirus; Venezuela). DISTRIBUTION: Mexico, Venezuela. quadricollis Sharp, 1887: 739 (Leptochirus; Type locality: Mexico, Jalapa). — Bernhauer, 1903: 123 (Leptochirus; synonym of mexicanus). — Bernhauer and Schubert, 1910: 13 (Leptochirus; synonym of mexicanus). mexicanus Sharp, 1887, see: bernhaueri Wendeler, 1957. micrognathus Fauvel, 1902, see: Priochirus. minutus Laporte, 1840, see: Borolinus. molossus Sharp, 1887: 735 (Leptochirus; Type locality: Mexico, Jalapa, Tuxtla, Cordova; British Honduras, R. Sarstoon, R. Hondo, Belize; Guatemala, El Tumbador, Las Mer- cedes, San Isidro, Pantaleon, Mirandilla, Capetillo, San Geronimo, Teleman, Chacoj, San Juan in Vera Paz, Cubilguitz; Salvador; Nicaragua, Chontales; Costa Rica, Irazu 6000 to 7000 feet, Cache; Panama, Bugaba, Volcan de Chiriqui 2500 to 4000 feet). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 121 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 999 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Mexico, Belize, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama. monilicornis Fauvel, 1878, see: Priochirus. montanus Bernhauer, 1912d: 28 (Leptochirus; subgenus Mesochirus; Type locality: Columbia: Cali: Rio Vitaco, 2000 m; Rio Aquatal, 1800 m; Villa Elvira, 1800 m). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Mesochirus; catalog). DISTRIBUTION: Colombia. nigricollis Fauvel, 1902, see: Borolinus. nigripennis Fauvel, 1902, see: Borolinus. nigrita Fauvel, 1902, see: Priochirus. novus Bernhauer, 1903: 121, 127 (Leptochirus; Type locality: Ecuador). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Ecuador. Bulletin AMNH OSORIINAE: LEPTOCHIRUS 1105 opacicollis Fauvel, 1878, see: Priochirus. pachoensis Bernhauer, 1917f: 47 (Leptochirus; subgenus Tropiochirus; Type locality: Westkolumbien: Pacho in den Ostkordilleren, 2000 m). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Tropochirus; catalog). DISTRIBUTION: Colombia. paradoxus Bernhauer, 1903: 121, 128 (Leptochirus; Type locality: Ecuador). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Ecuador. parcus Fauvel, 1878, see: Priochirus. parvicollis Fauvel, 1902, see: Priochirus. peruvianus Bernhauer, 1917f: 46 (Leptochirus; subgenus Leptochirus; Type locality: Peru: Chanchamayo). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Peru. piestoides Fauvel, 1902, see: Thoracochirus. politiventris Fauvel, 1902, see: Priochirus. proteus Fauvel, 1864b: 12 (Leptochirus; Type locality: Mexique; Colombie; Brésil). — Fauvel, 1865: 16 (Leptochirus; characters; Mexico; Colombia; Brazil). — Erichson, 1840: 825 (Leptochirus; [Note: Misidentification: Bernhauer and Schubert, 1910: 14 cited maxillosus Erichson as a synonym of proteus Fauvel; Erichson did not describe the species as new, he attributed it to Fabricius]; characters). — Solsky, 1875: 26 (Leptochirus; Peru). — Sharp, 1887: 735 (Leptochirus; notes; Mexico). — Fauvel, 1891: 87 (Leptochirus; Venezuela). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 125 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Tropiochirus; catalog). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Tropochirus; catalog). DISTRIBUTION: Peru, Colombia, Venezuela, Mexico. punctiger Sharp, 1887: 738 (Leptochirus; Type locality: Mexico, Oaxaca, Orizaba, Jalapa). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 123 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Mexico. pygmaeus Kraatz, 1859, see: Priochirus. quadricollis Sharp, 1887, see: mexicanus Erichson, 1840. quadridens Motschulsky, 1857a: 501 (Leptochirus; Type locality: Indes orientales). — Kraatz, 1859: 191 (Leptochirus). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 125 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 15 (Leptochirus; subgenus Strongylochirus; catalog). — Bernhauer, 1922a: 149 (Leptochirus; Indochina). — Cameron, 1928d: 94 (Leptochirus; subgenus Strongylochirus; Sumatra). — Wendeler, 1928c: 117 (Leptochirus; subgenus Strongylochirus; Philippines). — Cameron, 1930: 91 (Leptochirus; subgenus Strongylochirus; characters; Assam; Burma; Siam; Sumatra; Java; Borneo). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Strongylochirus; catalog). — Paulian, 1941: 131 (Leptochirus; larval characters; India). — Abdullah and Qadri, 1970: 116 (Leptochirus; subgenus Strongylochirus; characters; East Pakistan). — Biswas and Sengupta, 1977: 341 (Leptochirus; subgenus Strongylochirus; characters; India). 1106 OSORIINAE: LEPTOCHIRUS Bulletin AMNH
— Coiffait, 1984: 117 (Leptochirus; India). — Biswas and Biswas, 1985: 141 (Leptochirus; subgenus Strongylochirus; India). DISTRIBUTION: India, Bangladesh, Myanmar, Thailand, Borneo, Indonesia. uniformis Fauvel, 1902: 9 (Leptochirus; variety of quadridens; Type locality: Célèbes: Bonthain 1000 à 2500 m). — Cameron, 1928d: 94 (Leptochirus; subgenus Strongylochirus; variety of quadridens; Sumatra). — Scheerpeltz, 1933: 1001 (Leptochirus; variety of quadridens). major Bernhauer, 1903: 126 (Leptochirus; subspecies of quadridens; Type locality: Sumatra). — Cameron, 1928d: 94 (Leptochirus; subgenus Strongylochirus; variety of quadridens; Sumatra). quadrifidus Fauvel, 1878, see: Priochirus. raffrayi Fauvel, 1902, see: Thoracochirus. ramosus Sharp, 1887: 740 (Leptochirus; Type locality: Mexico, Jalapa). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 123 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Mexico. rugosus Fauvel, 1879, see: Thoracochirus. samoensis Blanchard, 1853, see: Priochirus. sanguinosus Motschulsky, 1857, see: Priochirus. scoriaceus Germar, 1824: 35 (Leptochirus; Type locality: Brasilia). — Blanchard, 1842: 88 (Leptochirus; Brazil). — Erichson, 1840: 825 (Leptochirus; characters; Brazil; Colombia; Mexico). — Laporte, 1840: 186 (Leptochirus; characters; Brazil; Mexico). — Guérin-Méneville, 1855: 583 (Leptochirus; Brazil). — Fauvel, 1864b: 12 (Leptochirus; characters; Mexico; Colombia; Brazil). — Fauvel, 1865: 16 (Leptochirus; characters; Mexico; Colombia; Brazil). — Solsky, 1868a: 144 (Leptochirus; Mexico). — Solsky, 1872: 314 (Leptochirus; Colombia). — Fauvel, 1877a: xxiv (Leptochirus; Brazil). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 121 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1951a: 8 (Leptochirus; Bolivia). — Scheerpeltz, 1960c: 66 (Leptochirus; subgenus Leptochirus; Bolivia). DISTRIBUTION: Paraguay, Argentina, Bolivia, Brazil, Colombia, Mexico. coriaceus Gray, 1832: 306 (Zirophorus; [Note: Attributed to Lacordaire]; Type locality: Brazil). — Erichson, 1840: 823 (Leptochirus; synonym of scoriaceus). — Bernhauer and Schubert, 1910: 13 (Leptochirus; synonym of scoriaceus). semirufus Fauvel, 1895, see: Borolinus. serriger Sharp, 1887: 738 (Leptochirus; Type locality: Mexico, Cordova, Jalapa). — Heller, 1898: 6 (Leptochirus; checklist). — Bernhauer, 1903: 121 (Leptochirus; characters). — Bernhauer and Schubert, 1910: 13 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Mexico. sharpi Fauvel, 1902: 16 (Leptochirus; Type locality: Mexique: Sierra de Durango). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Leptochirus; catalog). DISTRIBUTION: Mexico. Bulletin AMNH OSORIINAE: LEPTOCHIRUS 1107 sharpi Bernhauer, 1903, see: bernhaueri Wendeler, 1957. simoni Fauvel, 1902, see: Thoracochirus. simplex Fauvel, 1902, see: Priochirus. sinuatus Fauvel, 1902, see: Priochirus. socius Fauvel, 1902, see: Priochirus. spinosulus Fauvel, 1878, see: Priochirus. sulcifer Fauvel, 1902, see: Priochirus. sutteri Scheerpeltz, 1957c: 222, 238 (Leptochirus; subgenus Triacantochirus; Type locality: Insel Flores, Rana Mese, in 1300 m Höhe im Regenwald). DISTRIBUTION: Indonesia. taurus Fauvel, 1902, see: Priochirus. tenuicornis Bernhauer, 1903: 123, 130 (Leptochirus; Type locality: Brasilien). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Leptochirus; catalog). — Scheerpeltz, 1933: 1000 (Leptochirus; subgenus Leptochirus; catalog; Argentina). DISTRIBUTION: Argentina, Brazil. tenuis Bernhauer, 1903: 125, 132 (Leptochirus; Type locality: Chanchomayo, Bolivien). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Mesochirus; catalog). — Marcuzzi, 1977: 4 (Leptochirus; Venezuela). DISTRIBUTION: Bolivia, Venezuela. tridens Motschulsky, 1857, see: Priochirus. tridentatus Bernhauer, 1917f: 48 (Leptochirus; subgenus Tropiochirus; Type locality: West- Kolumbien: Umgebung von Cali am Rio Cauca, Cañon del Mt. Tolima, 1700 m). DISTRIBUTION: Colombia. trisinuatus Biswas and Gupta, 1977: 342 (Leptochirus; subgenus Strongylochirus; Type locality: Burma: Thingannyinaung to Sukli in Dwana Hills, 900–2100 ft.). DISTRIBUTION: Myanmar. truncatus Fauvel, 1902: 17 (Leptochirus; Type locality: Bolivie: Yuracaris). — Bernhauer and Schubert, 1910: 14 (Leptochirus; subgenus Mesochirus; catalog). DISTRIBUTION: Bolivia. unicolor Laporte, 1835, see: Priochirus. uniformis Fauvel, 1902, see: quadridens Motschulsky, 1857. variolosus Fauvel, 1895, see: Thoracochirus. venezuelanus Wendeler, 1957a: 46 (Leptochirus; subgenus Tropiochirus; Type locality: Venezuela: Merida). DISTRIBUTION: Venezuela. verrucifer Fauvel, 1895, see: Thoracochirus. vitulus Fauvel, 1878, see: Priochirus. weissi Fauvel, 1903, see: Priochirus. yunganus Scheerpeltz, 1951a: 11 (Leptochirus; subgenus Tropiochirus; Type locality: Yungas del Palmar, 2000 m). DISTRIBUTION: Bolivia. zischkai Scheerpeltz, 1951a: 8 (Leptochirus; subgenus Leptochirus; Type locality: Yungas del Palmar, 2000 m). DISTRIBUTION: Bolivia. zischkaianus Scheerpeltz, 1951a: 10 (Leptochirus; subgenus Tropiochirus; Type locality: Yungas del Palmar, 2000 m). DISTRIBUTION: Bolivia. 1108 OSORIINAE: PRIOCHIRUS Bulletin AMNH
Priochirus [270 species; Neotropical, Palaearctic, Madagascan, Oriental, Australian, and Oceanic Regions] Priochirus Sharp, 1887: 740 (species included: laxus, bicornis, haemorrhous, dilutus, diver- gens, collaris, laticornis, salvini). TYPE SPECIES: Priochirus haemorrhous Sharp, fixed by subsequent designation by Blackwelder, 1952: 321. — Fauvel, 1878f: 184 (cited as Leptochirus; characters; key to Indonesian species). — Heller, 1898: 6, 9 (checklist of and key to species). — Bernhauer, 1903: 116, 135 (characters; key to species). — Bernhauer and Schubert, 1910: 15 (world catalog; 86 species). — Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog). — Winkler, 1925: 323 (catalog for Palaearctic region). — Cameron, 1925: 4 (catalog of Indian species). — Cameron, 1930: 95, 116, 454 (characters; key to British Indian subgenera and species). — Scheerpeltz, 1933: 1001, 1008 (world catalog supplement). — Blackwelder, 1944: 110 (checklist of species from Mexico, Central and South America, and the West Indies). — P. J. M. Greenslade, 1971: 126, 185 (characters; revision of subgeneric classification; species group classification; key to subgenera; species included in subgenera). — P. J. M. Greenslade, 1972 (distribution patterns of species in Solomon Islands). — P. J. M. Greenslade, 1972a (evolution; distributional patterns and radiation). — Shibata, 1973c: 25 (checklist of species of Taiwan). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Shibata, 1976: 88 (checklist of species of Japan). — Hammond, 1984: 198 (checklist of species of Borneo). — Shibata, 1986a: 118 (supplemental list of species from Taiwan). — Newton, 1990: 91 (characters in key; one described Australian species). — Naomi, 1996a: 1 (revision of and key to species from Japan; notes). — Drugmand, 1998a: 220 (list of types in l’Institut royal des Sciences naturelles de Belgique). Triacanthus Bernhauer, 1903: 136, 158 (subgenus of Priochirus; species included: inte- grifrons, unicolor, immarginatus, apicalis, tonkinensis, poseidon, neptunus, alutaceus, subnitidus, parabolicus, caviceps, sumatrensis, subtridens, doriae, gestroi, bifurcatus, kraatzi, modiglianii, punctiventris, tridens, insularis, trifidus, depressus, coronatus, bipunctatus; preoccupied by Oken, 1817). TYPE SPECIES: Leptochirus unicolor Laporte, fixed by subsequent designation by R. Lucas, 1920: 647. — Bernhauer and Schubert, 1910: 15 (world catalog; subgenus of Priochirus). — Blackwelder, 1952: 393 (type species). — P. J. M. Greenslade, 1971: 127 (synonym of Priochirus subgenus Eutriacanthus). Cephalomerus Bernhauer, 1903: 139, 159 (subgenus of Priochirus; species included: japonicus, ducalis, fauveli, sanguinosus, exaratus, bifoveatus, pygmaeus, hoplites). TYPE SPECIES: Leptochirus pygmaeus Kraatz, fixed by subsequent designation by R. Lucas, 1920: 171. — Bernhauer and Schubert, 1910: 16 (world catalog; subgenus of Priochirus). — Cameron, 1925: 4 (subgenus of Priochirus; catalog of Indian species). — Cameron, 1930: 100 (subgenus of Priochirus; characters; key to British Indian species). — Scheerpeltz, 1933: 1003 (world catalog supplement; subgenus of Cephalomerus). — Blackwelder, 1952: 96 (type species). — P. J. M. Greenslade, 1971: 136 (subgenus of Priochirus; characters; list of species included). — P. J. M. Greenslade, 1972a: 207 (evolution; distributional patterns). Peucodontus Bernhauer, 1903: 140, 159 (subgenus of Priochirus; species included: mandibularis; [Note: The name was also cited as Pseucodontus on page 159]). TYPE SPECIES: Leptochirus mandibularis Kraatz, fixed by monotypy. — Bernhauer and Schubert, 1910: 16 (world catalog; subgenus of Priochirus). — Cameron, 1925: 4 (subgenus of Priochirus; catalog of Indian species). — Cameron, 1930: 108 (subgenus of Priochirus; characters). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1109
— Scheerpeltz, 1933: 1004 (world catalog supplement; subgenus of Priochirus). — Blackwelder, 1952: 299 (type species: mandibularis). — P. J. M. Greenslade, 1971: 130 (subgenus of Priochirus; characters; list of species included). Stigmatochirus Bernhauer, 1903: 141, 159 (subgenus of Priochirus; species included: dohrni). TYPE SPECIES: Leptochirus dohrni Fauvel, fixed by monotypy. — Bernhauer and Schubert, 1910: 16 (world catalog; subgenus of Priochirus). — Cameron, 1930: 109 (subgenus of Priochirus; characters). — Scheerpeltz, 1933: 1004 (world catalog supplement; subgenus of Priochirus). — Blackwelder, 1952: 361 (type species: dohrni). — P. J. M. Greenslade, 1971: 130 (subgenus Priochirus; characters; list of species included). — P. J. M. Greenslade, 1972a: 208 (evolution; distributional patterns). Leptarthrus Bernhauer, 1903: 141, 159 (subgenus of Priochirus; species included: longicor- nis; preoccupied by Stephens, 1829, and Dietz, 1891). TYPE SPECIES: Leptochirus longicornis Fauvel, fixed by monotypy. — Bernhauer and Schubert, 1910: 16 (world catalog; subgenus of Priochirus). — Cameron, 1925: 4 (subgenus of Priochirus; catalog of Indian species). — Cameron, 1930: 109 (subgenus of Priochirus; characters; key to British Indian species). — Scheerpeltz, 1933: 1004 (synonym of Priochirus subgenus Neoleptarthrus). — Blackwelder, 1952: 215 (type species: longicornis). — Scheerpeltz, 1965: 98 (synonym of Priochirus subgenus Neoleptarthrus). — Abdullah and Qadri, 1970: 117 (cited as Leptarthrers; subgenus of Priochrius; key to species of East Pakistan). — P. J. M. Greenslade, 1971: 132 (synonym of Priochirus subgenus Euleptarthrus). Syncampsochirus Bernhauer, 1903: 141, 159 (subgenus of Priochirus; species included: samoensis, alternus, miles, quadrifidus, lorquini, forticornis, albertisi, spinosulus, propinquus). TYPE SPECIES: Leptochirus samoensis Blanchard, fixed by subsequent designation by R. Lucas, 1920: 622. — Bernhauer and Schubert, 1910: 17 (world catalog; subgenus of Priochirus). — Cameron, 1924: 82 (subgenus of Priochirus; key to species of New Guinea). — Scheerpeltz, 1933: 1005 (world catalog supplement; subgenus of Priochirus). — Blackwelder, 1952: 367 (type species). — P. J. M. Greenslade, 1971: 155 (subgenus of Priochirus; characters; species group classifica- tion; key to some species; list of species included). — P. J. M. Greenslade, 1972a: 206 (evolution; distributional patterns). Catacamptus Bernhauer, 1903: 142, 159 (subgenus of Priochirus; species included: cilifrons, impressifrons, extensus, frater). TYPE SPECIES: Leptochirus extensus Fauvel, fixed by subsequent designation by R. Lucas, 1920: 167. — Bernhauer and Schubert, 1910: 17 (world catalog; subgenus of Priochirus). — Scheerpeltz, 1933: 1005 (world catalog supplement; subgenus of Priochirus). — Blackwelder, 1952: 95 (type species). — P. J. M. Greenslade, 1971: 174 (synonym of Priochirus subgenus Plastus). Plastus Bernhauer, 1903: 142, 160 (subgenus of Priochirus; species included: monilicornis, haakei, inaequalis, sexdentatus, excavatus, ruficrus, tetraodon, cavifrons, luzonicus, opacicollis, parcus, beccarii, antennarius, freminvillii, vitulus, conicocollis, convexus, ebeninus, brachycerus). TYPE SPECIES: Leptochirus convexus Laporte, fixed by subsequent designation by R. Lucas, 1920: 517. — Bernhauer and Schubert, 1910: 17 (world catalog; subgenus of Priochirus). — Cameron, 1924: 85 (subgenus of Priochirus; key to species of New Guinea). — Cameron, 1925: 4 (subgenus of Priochirus; catalog of Indian species). — Cameron, 1930: 111 (subgenus of Priochirus; characters; key to British Indian species). — Scheerpeltz, 1933: 1006 (world catalog supplement; subgenus of Priochirus). — Blackwelder, 1952: 311 (type species). — P. J. M. Greenslade, 1971: 174 (subgenus of Priochirus; characters; species group classifi- cation; list of species included). — P. J. M. Greenslade, 1972a: 209 (evolution; distributional patterns). 1110 OSORIINAE: PRIOCHIRUS Bulletin AMNH
Euleptarthrus Jakobson, 1908: 466 (subgenus of Priochirus; replacement name for Leptarthrus Bernhauer). TYPE SPECIES: Leptochirus longicornis Fauvel, fixed by objective synonymy with Leptarthrus. — P. J. M. Greenslade, 1971: 132 (subgenus of Priochirus; characters; list of species included). — P. J. M. Greenslade, 1972a: 208 (evolution; distributional patterns). — Shibata, 1986a: 118 (supplemental list of species from Taiwan). Eutriacanthus Jakobson, 1908: 466 (subgenus of Priochirus; replacement name for Triacanthus Bernhauer). TYPE SPECIES: Leptochirus unicolor Laporte, fixed by objective synonymy with Triacanthus. — P. J. M. Greenslade, 1971: 137 (subgenus of Priochirus; characters; list of species included). — P. J. M. Greenslade, 1972a: 207 (evolution; distributional patterns). Triacanthochirus Bernhauer, 1923c: 63 (subgenus of Priochirus; replacement name for Triacanthus Bernhauer). TYPE SPECIES: Leptochirus unicolor Laporte, fixed by objective synonymy with Triacanthus. — Cameron, 1925: 4 (subgenus of Priochirus; catalog of Indian species). — Cameron, 1930: 96 (subgenus of Priochirus; characters; key to British Indian species). — Scheerpeltz, 1933: 1001 (world catalog supplement; subgenus of Priochirus). — Blackwelder, 1952: 393 (type species). — Abdullah and Qadri, 1970: 117 (cited as Tricanthochirus; subgenus of Priochirus; key to species of East Pakistan). — P. J. M. Greenslade, 1971: 127 (synonym of Priochirus subgenus Eutriacanthus). Neoleptarthrus Scheerpeltz, 1933: 1004 (subgenus of Priochirus; replacement name for Leptarthrus Bernhauer). TYPE SPECIES: Leptochirus longicornis Fauvel, fixed by objective synonymy with Leptarthrus. — Blackwelder, 1952: 259 (type species). — P. J. M. Greenslade, 1971: 132 (synonym of Priochirus subgenus Euleptarthrus). Paraborolinus Nakane and Sawada, 1960: A121 (subgenus of Borolinus; species included: bicornis). TYPE SPECIES: Borolinus (Paraborolinus) bicornis Nakane and Sawada, fixed by monotypy. — Naomi, 1996a: 11, 13 (cited as a subgenus of Priochirus; notes; characters). Leiochirus P. J. M. Greenslade, 1971: 134 (subgenus of Priochirus; species included: latifrons). TYPE SPECIES: Priochirus (Leiochirus) latifrons P. J. M. Greenslade, fixed by original designation and monotypy. — P. J. M. Greenslade, 1972a: 208 (evolution; distributional patterns). Barychirus P. J. M. Greenslade, 1971: 135 (subgenus of Priochirus; species included: annamensis, arrowi, colossalis). TYPE SPECIES: Priochirus colossalis Bernhauer, fixed by original designation. — P. J. M. Greenslade, 1972a: 208 (evolution; distributional patterns). Exochirus P. J. M. Greenslade, 1971: 139 (subgenus of Priochirus; species included: albertisi, alces, altiusculus, aluticollis, alveatus, antennarius, aries, bacchi, beccarii, bicornutus, bisinuatus, brunneipennis, capra, cervus, cheesmani, cheesmanianus, con- fusus, conicicollis, dama, deceptor dentellicornis, divisifrons, freminvillii, germanus, heterodon, heurni, hirticornis, kaestneri, lautus, mayri, naturangai, novaebrittaniae, papuanus, parcus, parvus, popamanasiu, propinquus, quadricollis, rastri, fraterculus, reno, scyphus, sinuatus, sororculus, sparsus, spinosulus, sparsus, spinosulus, taurus, teres, tinahula, valens). TYPE SPECIES: Leptochirus beccarii Fauvel, fixed by original designation. — P. J. M. Greenslade, 1972a: 203 (evolution; distributional patterns). abori Bernhauer, 1926: 19 (Priochirus; subgenus Stigmatochirus; Type locality: Nordost- Indien: Kobo, 400 ft.; Upper Renging, 2150 ft.). — Cameron, 1930: 109 (Priochirus; subgenus Stigmatochirus; characters; India). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Stigmatochirus; catalog). — P. J. M. Greenslade, 1971: 130 (Priochirus; subgenus Stigmatochirus). DISTRIBUTION: India. Bulletin AMNH OSORIINAE: PRIOCHIRUS 1111 accessorius Cameron, 1928f: 399 (Priochirus; subgenus Triacanthochirus; Type locality: Mt. Penrissen, 4000 feet). — Scheerpeltz, 1933: 1001 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Malaysia. acietis P. J. M. Greenslade, 1971: 162 (Priochirus; subgenus Syncampsochirus; Type locality: Bismarck Archipelago: Duke of York Island, Manuan). DISTRIBUTION: Papua New Guinea. acraeus P. J. M. Greenslade, 1971: 178 (Priochirus; subgenus Plastus; Type locality: Solomon Islands: Guadalcanal, Mt. Popamanasiu, 5000 ft.). DISTRIBUTION: Solomon Islands. adjacens Cameron, 1937a: 85 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea: Mafulu, alt. 4000 ft.). — P. J. M. Greenslade, 1971: 155, 157 (Priochirus; subgenus Syncampsochirus; characters). DISTRIBUTION: New Guinea. alces P. J. M. Greenslade, 1971: 153 (Priochirus; subgenus Exochirus; Type locality: Solomon Islands: Guadalcanal, Mt. Popamanasiu, 4500 feet). DISTRIBUTION: Solomon Islands. albertisi Fauvel, 1878f: 188 (Leptochirus; Type locality: Nouvelle-Guinée, Andai; Ramoi; Amberbaki). — Fauvel, 1879: 72 (Leptochirus; Vaigheu; Salvatti; Dorei). — Heller, 1898: 6 (Priochirus; checklist). — Bernhauer, 1903: 142 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1924: 81, 82 (Priochirus; subgenus Syncampsochirus; key characters; New Guinea). — Cameron, 1931: 355 (Priochirus; New Guinea). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1937a: 84 (Priochirus; subgenus Syncampsochirus; New Guinea). — P. J. M. Greenslade, 1971: 140, 148 (Priochirus; subgenus Exochirus; New Guinea). DISTRIBUTION: New Guinea. alienus Cameron, 1945a: 141 (Priochirus; subgenus Plastus; Type locality: Java). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. alternus Fauvel, 1878f: 191 (Leptochirus; Type locality: Nouvelle-Guinée, Hatam; Amber- baki, Has). — Fauvel, 1879: 73 (Leptochirus; Amberbaki, Has). — Heller, 1898: 6 (Priochirus; checklist). — Bernhauer, 1903: 141 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1924: 82 (Priochirus; subgenus Syncampsochirus; characters in key; New Guinea). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1937a: 84 (Priochirus; subgenus Syncampsochirus; New Guinea). — P. J. M. Greenslade, 1971: 155, 157 (Priochirus; subgenus Syncampsochirus; characters). — Scheerpeltz, 1971b: 201 (Priochirus; New Guinea). DISTRIBUTION: New Guinea. altiusculus P. J. M. Greenslade, 1971: 143 (Priochirus; subgenus Exochirus; Type locality: Solomon Islands: Guadalcanal, Mt. Popamanasiu, 5000 feet). DISTRIBUTION: Solomon Islands. alutaceus Bernhauer, 1903: 138, 149 (Priochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; illustration; Sumatra). — Scheerpeltz, 1933: 1001 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. 1112 OSORIINAE: PRIOCHIRUS Bulletin AMNH aluticollis Cameron, 1924: 83, 86 (Priochirus; subgenus Plastus; Type locality: Idenburg Riv.). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 140 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. alveatus P. J. M. Greenslade, 1971: 146 (Priochirus; subgenus Exochirus; Type locality: Solomon Islands: Guadalcanal, Mt. Austen). DISTRIBUTION: Solomon Islands. analis Cameron, 1920, see: pentagonalis Bernhauer, 1914. andrewesi Cameron, 1930: 107 (Priochirus; subgenus Cephalomerus; Type locality: Niligiri Hills). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 136 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: India. angulatus Wendeler, 1928c: 123 (Priochirus; subgenus Plastus; Type locality: Philippinen: Panaon). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. annamensis Cameron, 1946a: 682 (Priochirus; subgenus Cephalomerus; Type locality: Tonkin: Chapa). — P. J. M. Greenslade, 1971: 136 (Priochirus; subgenus Barychirus). DISTRIBUTION: Vietnam. antennarius Fauvel, 1878f: 187 (Leptochirus; Type locality: Nouvelle-Guinée, Hatam, Ramoi; Amberbaki). — Heller, 1898: 6 (Priochirus; checklist). — Bernhauer, 1903: 144 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 84, 86 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 140 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. apicalis Eppelsheim, 1895c: 68 (Leptochirus; Type locality: Sikkim im Himalaya; Sumatra: Mte. Singalang). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 137 (Priochirus; characters). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1930: 97 (Priochirus; subgenus Triacanthochirus; characters; Sikkim). — Scheerpeltz, 1933: 1001 (Priochirus; subgenus Triacanthochirus; catalog). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Triacanthochirus; characters; Pakistan). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Pakistan, India, Indonesia. apiciventris Bernhauer, 1928, see: modiglianii Bernhauer, 1903. aries P. J. M. Greenslade, 1971: 154 (Priochirus; subgenus Exochirus; Type locality: Caroline Islands: Yap). DISTRIBUTION: Caroline Islands. armatus P. J. M. Greenslade, 1971: 158 (Priochirus; subgenus Syncampsochirus; Type local- ity: New Guinea: Morobe District, Edie Creek, about 6 miles S.W. Wau, Bulldog Road, 9700 feet). DISTRIBUTION: New Guinea. arrowi P. J. M. Greenslade, 1971: 136 (Priochirus; subgenus Barychirus; Type locality: Indochina: Laos: Tintoe). DISTRIBUTION: Laos. Bulletin AMNH OSORIINAE: PRIOCHIRUS 1113 arupi Saha and Vazirani, 1971: 513 (Priochirus; subgenus Plastus; Type locality: India: N.E.F.A.: Kameng Division, Jhum la). DISTRIBUTION: India. ascendens Fauvel, 1903a: 171 (Leptochirus; subgenus Priochirus; Type locality: Bírmanie; Siam [ou Sikkim]). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Cameron, 1930: 114 (Priochirus; subgenus Plastus; characters). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Plastus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: India ?, Bangladesh, Myanmar, Thailand. associatus Cameron, 1952: 242 (Priochirus; subgenus Catacamptus; Type locality: New Guinea: Snow Mountains: Moss Forest Camp, 2600–2800 m; Top Camp, 2100 m). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. astoliensis R. Dvorˇák, 1957b: 790 (Priochirus; subgenus Plastus; Type locality: India: Astoli, North Kanara). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: India. austeni P. J. M. Greenslade, 1971: 165 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Guadalcanal, Mt. Austen). DISTRIBUTION: Solomon Islands. bacchi P. J. M. Greenslade, 1971: 147 (Priochirus; subgenus Exochirus; Type locality: New Guinea: Morobe District, Edie Creek, about 6 miles S.W. Wau, Bulldog road, 9700 feet). DISTRIBUTION: New Guinea. bakeri Bernhauer, 1926b: 39 (Priochirus; subgenus Plastus; Type locality: Philippinen, Bergen von Benguet, Baguio). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. bakerianus Bernhauer, 1926b: 39 (Priochirus; subgenus Triacanthus; Type locality: Borneo: Sandakan). — Bernhauer, 1928: 2 (Priochirus; Sumatra). — Scheerpeltz, 1933: 1001 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Borneo, Indonesia. banghaasi Wendeler, 1928c: 120 (Priochirus; subgenus Catacamptus; Type locality: Philip- pinen: Luzon: Imugan). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Catacamptus; catalog). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. beabea P. J. M. Greenslade, 1971: 183 (Priochirus; subgenus Plastus; Type locality: Solomon Islands: Guadalcanal, Mt. Popamanasiu, 4500 feet). DISTRIBUTION: Solomon Islands. beccarii Fauvel, 1878f: 191 (Leptochirus; Type locality: Iles Arou, Wokan; Nouvelle-Guinée, Andai, Hatam, Ramoi). — Fauvel, 1879: 73 (Leptochirus; Mysol; Fly River; Katau). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 144 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 86 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Cameron, 1931: 355 (Priochirus; New Guinea). 1114 OSORIINAE: PRIOCHIRUS Bulletin AMNH
— Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — Cameron, 1937a: 86 (Priochirus; subgenus Plastus; New Guinea). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: Indonesia, New Guinea. bengalensis Cameron, 1945: 64 (Priochirus; subgenus Cephalomerus; Type locality: Bengal: Kalimpong, Samsingh, altitude 1800 feet). — P. J. M. Greenslade, 1971: 136 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: India. bernhaueri Wendeler, 1928c: 121 (Priochirus; subgenus Catacamptus; Type locality: Philippinen: Luzon: Imugan). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Catacamptus; catalog). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. bicolor Cameron, 1937a: 85 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft.). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. bicornis Fauvel, 1864b: 16 (Leptochirus; Type locality: Mexico). — Fauvel, 1865: 20 (Leptochirus; characters; Mexico). — Sharp, 1887: 741 (Priochirus; Mexico). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 185 (Priochirus; subgenus Priochirus). — Ito, 1997: 8 (Priochirus; subgenus Paraborolinus; Japan). DISTRIBUTION: Mexico. bicornis Nakane and Sawada, 1960, see: yakushimensis Herman, 2001. bicornutus Cameron, 1952: 242 (Priochirus; subgenus Plastus; Type locality: New Guinea: Snow Mountains: Bernhard Camp). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. bidens Cameron, 1928g: 431 (Priochirus; subgenus Catacamptus; Type locality: Borneo: Mt. Murud, 6500 feet). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Catacamptus; catalog). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Borneo. bidenticulatus Scheerpeltz, 1965: 138 (Priochirus; subgenus Cephalomerus; Type locality: N.E. Burma, Kambaiti, 2000 m). DISTRIBUTION: Myanmar. bifoveatus Eppelsheim, 1895c: 69 (Leptochirus; Type locality: Sikkim im Himalaya). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 140 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1930: 102 (Priochirus; subgenus Cephalomerus; characters; Sikkim; India). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — Paulian, 1941: 133 (Priochirus; larval characters; India). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Cephalomerus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 136 (Priochirus; subgenus Cephalomerus). — Saha and Vazirani, 1971a: 100 (Priochirus; subgenus Cephalomerus; India). DISTRIBUTION: India, Bangladesh. major Cameron, 1920a: 142 (Priochirus; subgenus Cephalomerus; Type locality: Pulney Hills, S. India). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1115
— Cameron, 1930: 102 (Priochirus; synonym of bifoveatus). — Scheerpeltz, 1933: 1003 (Priochirus; synonym of bifoveatus). bifurcatus Fauvel, 1879: 74 (Leptochirus; Type locality: Nouvelle Guinée, Dorei; Amberbaki). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 138 (Priochirus; characters). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1931: 355 (Priochirus; New Guinea). — Scheerpeltz, 1933: 1001 (Priochirus; subgenus Triacanthochirus; catalog). — Cameron, 1937a: 84 (Priochirus; subgenus Triacanthochirus; New Guinea). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: New Guinea. bilobatus Cameron, 1929b: 437 (Priochirus; subgenus Cephalomerus; Type locality: Selangor: The Gap. Perak). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Malaysia. bipunctatus Fauvel, 1895: 180 (Leptochirus; Type locality: Bírmanie, Carin Cheba, 900– 1100 m; Carin Ghécu, 1300–1400 m; Tenasserim, mont Mooleyit, 1800–1900; Plapoo; Siam, monts de La Khone). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1930: 97 (Priochirus; subgenus Triacanthochirus; characters; Sikkim; Burma). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Triacanthochirus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). — Biswas and Sengupta, 1982: 141 (Priochirus; subgenus Triacanthochirus; characters; India). — Biswas and Biswas, 1985: 143 (Priochirus; subgenus Triacanthochirus; India). DISTRIBUTION: India, Myanmar, Thailand, Indonesia. nigrita Fauvel, 1902: 13 (Leptochirus; subgenus Priochirus; variety of bipunctatus; Type locality: Malacca; Sumatra). — Bernhauer and Schubert, 1910: 15 (Priochirus; variety of bipunctatus). birmanus Cameron, 1930: 117 (Priochirus; Type locality: Burma: Ruby Mines). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Myanmar. bisinuatus Cameron, 1924: 81, 82 (Priochirus; subgenus Syncampsochirus; Type locality: Mamberamo Riv.). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. boettcheri Wendeler, 1924: 340 (Priochirus; subgenus Triacanthus; Type locality: Philippinen: Nord-Palawan, Binaluan). — Wendeler, 1928c: 118 (Priochirus; subgenus Triacanthochirus; Philippines). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Philippines. borneensis Cameron, 1928g: 425, 427 (Priochirus; subgenus Triacanthochirus; cited as corneensis [see Herman, 2001]; Type locality: Borneo: Mt. Murud, 7000–7200 feet). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; cited as borneensis; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus; cited as borneensis). — Herman, 2001: 11 (Priochrius; spelling corrected). DISTRIBUTION: Borneo. 1116 OSORIINAE: PRIOCHIRUS Bulletin AMNH borolinoides Cameron, 1928g: 427, 430 (Priochirus; subgenus Cephalomerus; Type locality: Borneo: Mt. Dulit). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Borneo. brachycerus Kraatz, 1859: 191 (Leptochirus; Type locality: Ceylan). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Cameron, 1930: 114 (Priochirus; subgenus Plastus; characters; Ceylon). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). — Saha and Vazirani, 1971: 514 (Priochirus; subgenus Plastus; India). DISTRIBUTION: Sri Lanka. brevicornis Erichson, 1840: 827 (Leptochirus; Type locality: Java). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 145 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Cameron, 1930: 117 (Priochirus; subgenus Plastus; characters; East Indies; Java). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. brevidenticulatus Scheerpeltz, 1965: 140 (Priochirus; subgenus Neoleptarthrus; Type local- ity: N.E. Burma, Kambaiti, 7000 ft.). — P. J. M. Greenslade, 1971: 132 (Priochirus; subgenus Euleptarthrus). DISTRIBUTION: Myanmar. browni P. J. M. Greenslade, 1971: 163 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Guadalcanal, Mt. Popamanasiu, 4500 feet). DISTRIBUTION: Solomon Islands. brunneipennis Cameron, 1928f: 400 (Priochirus; subgenus Syncampsochirus; Type locality: Mt. Penrissen, 4000 feet). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: Borneo. cameroni Scheerpeltz, 1933, see: difficilis Cameron, 1928. capra P. J. M. Greenslade, 1971: 151 (Priochirus; subgenus Exochirus; Type locality: New Guinea: Madang District, Finisterre Mts., Naho R. valley, Damanti, ca. 3550 feet). DISTRIBUTION: New Guinea. caviceps Eppelsheim, 1895b: 408 (Leptochirus; Type locality: Sumbawa). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 137 (Priochirus; characters). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; illustration; Sumatra). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. cavifrons Fauvel, 1878f: 194 (Leptochirus; Type locality: Iles Key). — Fauvel, 1879: 74 (Leptochirus; Fly River). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 144 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Bernhauer, 1915g: 180 (Priochirus; subgenus Syncampsochirus; Key Islands; Engano; Sumatra; Java). — Cameron, 1924: 85 (Priochirus; subgenus Plastus; characters in key; New Guinea). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1117
— Cameron, 1928d: 95 (Priochirus; subgenus Plastus; Sumatra). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — Cameron, 1936a: 27 (Priochirus; subgenus Plastus; Java). — Cameron, 1937a: 86 (Priochirus; subgenus Plastus; New Guinea). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia, New Guinea. centralis Fauvel, 1902: 18 (Leptochirus; subgenus Priochirus; Type locality: Guatemala). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 185 (Priochirus; subgenus Priochirus). DISTRIBUTION: Guatemala. cervus P. J. M. Greenslade, 1971: 153 (Priochirus; subgenus Exochirus; Type locality: Solomon Islands: S. Cristoval, 175 feet, Warahito R. camp site). DISTRIBUTION: Solomon Islands. ceylanensis Fauvel, 1902: 12 (Leptochirus; subgenus Priochirus; Type locality: Ceylan). — Fauvel, 1904a: 44 (Leptochirus; N. Kanara; Ceylon). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1930: 99 (Priochirus; subgenus Triacanthochirus; characters; Ceylon). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Sri Lanka. chapmani P. J. M. Greenslade, 1971: 131 (Priochirus; subgenus Stigmatochirus; Type local- ity: Indochina: Tonkin, Ngai-To, 4800 feet. Also cited from Siam and Laos). DISTRIBUTION: Vietnam, Laos, Thailand. cheesmani Cameron, 1934a: 20 (Priochirus; subgenus Syncampsochirus; Type locality: New Hebrides: Santo). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Hebrides. cheesmanianus Cameron, 1939b: 139 (Priochirus; subgenus Plastus; Type locality: New Guinea: Cyclops Mts.: Mt. Lina, altitude 3500 feet). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. chinensis Bernhauer, 1933: 25 (Priochirus; subgenus Leptarthrus; Type locality: Szetschwan: Tatsienlu Tjiji, Urwald-Rodungen). — P. J. M. Greenslade, 1971: 132 (Priochirus; subgenus Euleptarthrus). DISTRIBUTION: China. cilifrons Heller, 1898: 16 (Priochirus; Type locality: Celebes meridionalis, summus mons Bonthain et Maros). — Bernhauer, 1903: 142 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Catacamptus; catalog). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Catacamptus; catalog). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. circumpunctatus Heller, 1910: 12 (Priochirus; subgenus Stigmatochirus ?; Type locality: Papua). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Stigmatochirus; catalog). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. collaris Sharp, 1887: 742 (Priochirus; Type locality: Guatemala, Purula 4000 feet). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 146 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 185 (Priochirus; subgenus Priochirus). DISTRIBUTION: Guatemala. 1118 OSORIINAE: PRIOCHIRUS Bulletin AMNH colossalis Bernhauer, 1928: 3 (Priochirus; subgenus Cephalomerus; Type locality: Nordindien). — Cameron, 1930: 100 (Priochirus; subgenus Cephalomerus; characters; India). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Cephalomerus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 136 (Priochirus; subgenus Barychirus). DISTRIBUTION: India, Bangladesh. combustus Fauvel, 1902: 10 (Leptochirus; subgenus Priochirus; Type locality: Sikkim: Darjeeling). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1930: 103 (Priochirus; subgenus Cephalomerus; characters; Sikkim). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Cephalomerus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: India, Bangladesh. comes Fauvel, 1902: 11 (Leptochirus; subgenus Priochirus; Type locality: Sumatra: Mons Singalang). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Cameron, 1928d: 95 (Priochirus; Sumatra). — P. J. M. Greenslade, 1971: 186 (Priochirus; subgeneric assignment uncertain, probably not in subgenus Priochirus; notes). DISTRIBUTION: Indonesia. confusus Cameron, 1937a: 86 (Priochirus; subgenus Plastus; Type locality: New Guinea: Kokoda, alt. 1200 ft.). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. conicicollis Fauvel, 1878f: 189 (Leptochirus; Type locality: Nouvelle-Guinée, Amberbaki). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 86 (Priochirus; subgenus Plastus; cited as conicollis; characters in key; New Guinea). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. convexus Laporte, 1835: 132 (Leptochirus; Type locality: Madagascar). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 145 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — Scheerpeltz, 1961g: 234 (Priochirus; subgenus Plastus; Madagascar). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: Madagascar. coronatus Sachse, 1852, see: unicolor Laporte, 1835. crassicornis Cameron, 1930: 113 (Priochirus; subgenus Plastus; Type locality: India). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 174 (Priochirus; subgenus Plastus). DISTRIBUTION: India. currani Bernhauer, 1912g: 246 (Priochirus; subgenus Plastus; Type locality: Luzon: Benguet, Mt. Pulog). — Wendeler, 1928c: 118 (Priochirus; subgenus Plastus; Philippines). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. Bulletin AMNH OSORIINAE: PRIOCHIRUS 1119
vastus Wendeler, 1928c: 123 (Priochirus; subgenus Plastus; subspecies currani; Type locality: Philippinen: Nord-Luzon: Heightsplan). — Scheerpeltz, 1933: 1006 (Priochirus; variety of currani). dama P. J. M. Greenslade, 1971: 154 (Priochirus; subgenus Exochirus; Type locality: Moluccas: Damma Is.). DISTRIBUTION: Indonesia. davidis Fairmaire, 1889, see: japonicus Sharp, 1889. deceptor Cameron, 1937a: 84 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. delkeskampi Wendeler, 1957: 25 (Priochirus; subgenus Syncampsochirus; Type locality: Neu Guinea: Finschhafen). — P. J. M. Greenslade, 1971: 155, 157 (Priochirus; subgenus Syncampsochirus; characters). DISTRIBUTION: New Guinea. dentellicornis Wendeler, 1957: 25 (Priochirus; subgenus Syncampsochirus; Type locality: D. N. Guinea: Sattelberg). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. denticornutus Wendeler, 1928c: 121 (Priochirus; subgenus Catacamptus; Type locality: Philippinen: Mindanao: Momungan). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Catacamptus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. depressus Bernhauer, 1903, see: tridens Motschulsky, 1857. difficilis Cameron, 1928: 426, 427 (Priochirus; subgenus Triacanthochirus; [Note: This name is said to be preoccupied by Scheerpeltz, 1933: 1002, but is not]; Type locality: Borneo: Mt. Murud, 5000–6000 feet). — Cameron, 1933b: 338 (Priochirus; subgenus Triacanthochirus; British North Borneo). — Scheerpeltz, 1933: 1002 (Priochirus; synonym of cameroni). — Herman, 2001: 22 (Priochirus; valid species; not homonym). DISTRIBUTION: Borneo. difficilis Scheerpeltz, 1933: 1002 (Priochirus; [Note: Nomen nudum; Scheerpeltz cited Cameron in Entomological Monthly Magazine, 1920, 56: 142 as the reference for this name; the name is not published on that page and I cannot find it elsewhere. Scheerpeltz cited this name as the senior homonym of difficilis Cameron, 1928]). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: “East Indies”. cameroni Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; replacement name for difficilis Cameron). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). — Herman, 2001: 22 (Priochirus; synonym of difficilis). dilutus Sharp, 1887: 741 (Priochirus; Type locality: Panama, Volcan de Chiriqui 2500 to 4000 feet). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 146 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 185 (Priochirus; subgenus Priochirus). DISTRIBUTION: Panama. 1120 OSORIINAE: PRIOCHIRUS Bulletin AMNH divaricatus Cameron, 1928g: 429, 430 (Priochirus; subgenus Cephalomerus; Type locality: Borneo: Mt. Dulit, 3500 feet). — Cameron, 1928f: 400 (Priochirus; subgenus Cephalomerus; Borneo). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Borneo. divergens Sharp, 1887: 742 (Priochirus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 145 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 186 (Priochirus; subgenus Priochirus). DISTRIBUTION: Panama. divisifrons Heller, 1910: 10 (Priochirus; subgenus Cephalomerus; Type locality: Papua New Guinea). — Cameron, 1924: 86 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. dohrni Fauvel, 1902: 19 (Leptochirus; subgenus Priochirus; Type locality: Sumatra: Soekaranda; Tandjong Morawa Serdang). — Bernhauer, 1903: 141, 151 (Priochirus; characters; Sumatra). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Stigmatochirus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Stigmatochirus; Sumatra). — Bernhauer, 1928: 3 (Priochirus; Sumatra). — P. J. M. Greenslade, 1971: 130 (Priochirus; subgenus Stigmatochirus). DISTRIBUTION: Indonesia. doriae Heller, 1898: 19 (Priochirus; Type locality: Borneo septentrionalis, Sarawak). — Bernhauer, 1903: 137 (Priochirus; characters). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928f: 399 (Priochirus; Borneo). — Cameron, 1930b: 160 (Priochirus; cited as doriae f. minor Heller; [Note: Cameron cited minor as a form of doriae in a list of species but published no description; he attributed it to Heller but I find no such name by Heller so that Cameron’s form minor is an unavailable name (Article 12.1)]; British North Borneo). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Borneo. drescheri Cameron, 1936a: 27 (Priochirus; subgenus Cephalomerus; Type locality: Java: Preanger, G. Tangkoeban Prahoe, alt. 4000–5000 ft). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. ducalis Heller, 1898: 15 (Priochirus; Type locality: Sumatra, mons Singalang). — Bernhauer, 1903: 140 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Cephalomerus; Sumatra). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Indonesia. ebeninus Erichson, 1840: 826 (Leptochirus; Type locality: Madagascar). — Bernhauer and Schubert, 1910: 17 (Priochirus; synonym of convexus). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus; cited as valid species). DISTRIBUTION: Madagascar, Comoro Islands. Bulletin AMNH OSORIINAE: PRIOCHIRUS 1121
subconvexus Bernhauer, 1917m: 554 (Priochirus; Type locality: Anjouan, Comorenarchipel; Ambergebirge, N. Madagascar). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; synonym of ebeninus). eucerus Cameron, 1930: 114 (Priochirus; subgenus Plastus; Type locality: Sikkim: Gopaldhara, Rungbong Valley). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Plastus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). — Coiffait, 1978b: 113 (Priochirus; subgenus Plastus, cited as Platus; Bhutan). DISTRIBUTION: India, Bangladesh, Bhutan. exaratus Eppelsheim, 1895b: 406 (Leptochirus; Type locality: Bírmanien: Upper Burmah, Ruby Mines, 5000–7500 Fuss; Ost-Java: Montes Tengger, 4000 Fuss). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 140 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1930: 101 (Priochirus; subgenus Cephalomerus; characters; Burma). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — Scheerpeltz, 1965: 98 (Priochirus; subgenus Cephalomerus; Burma). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Myanmar, Malaysia, Indonesia. minor Bernhauer, 1928: 3 (Priochirus; subspecies of exaratus; Type locality: Perak; Sumatra: Bah Lias). — Scheerpeltz, 1933: 1003 (Priochirus; variety of exaratus). excavatus Motschulsky, 1857a: 502 (Leptochirus; Type locality: Indes orientales). — Kraatz, 1859: 191 (Leptochirus). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 144 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Plastus; Sumatra). — Wendeler, 1928c: 118 (Priochirus; subgenus Plastus; Philippines). — Cameron, 1930: 112 (Priochirus; subgenus Plastus; characters; East Indies; Sumatra; Borneo). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — Cameron, 1949b: 457 (Priochirus; subgenus Cephalomerus; Formosa). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: “East Indies”, Borneo, Indonesia, Philippines. extensus Fauvel, 1878f: 189 (Leptochirus; Type locality: Nouvelle-Guinée, Hatam). — Fauvel, 1879: 72 (Leptochirus; Ternate). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 142 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Catacamptus; catalog). — Cameron, 1924: 83 (Priochirus; subgenus Catacamptus; New Guinea). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Catacamptus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. fauveli Heller, 1898: 15 (Priochirus; Type locality: Sumatra, Si-Rambe). — Bernhauer, 1903: 140 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Cephalomerus; Sumatra). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Indonesia. 1122 OSORIINAE: PRIOCHIRUS Bulletin AMNH fletcheri Cameron, 1920a: 142 (Priochirus; subgenus Triacanthus; Type locality: Iyerpadi, S. India). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: India. formosae P. J. M. Greenslade, 1971: 133 (Priochirus; subgenus Euleptarthrus; Type locality: Formosa: Arisan). DISTRIBUTION: Taiwan. forticornis Fauvel, 1877: 185 (Leptochirus; Type locality: Iles Samoa, Upolu). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1927a: 21 (Priochirus; subgenus Syncampsochirus; figure; Samoa). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 155, 157 (Priochirus; subgenus Syncampsochirus; characters). DISTRIBUTION: Samoa. frater Heller, 1898: 17 (Priochirus; Type locality: Celebes meridionalis, mons Bonthain, 2000 m). — Bernhauer, 1903: 143 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Catacamptus; catalog). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Catacamptus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. fraterculus Cameron, 1936a: 27 (Priochirus; subgenus Plastus; Type locality: Java: G. Slamat, Batoerraden). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. fraterculus Cameron, 1937, see: rastri P. J. M. Greenslade, 1971. freminvillii Leguillou, 1841: 280 (Zirophorus; Type locality: Samoa, cited as Hamoa). — Fauvel, 1877: 185 (Leptochirus; cited as freminvillei; characters; Samoa). — Fauvel, 1878g: 480 (Leptochirus; cited as freminvillei; characters). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; cited as freminvillei; catalog). — Cameron, 1927a: 21 (Priochirus; subgenus Plastus; cited as freminvillei; figure; Samoa). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; cited as freminvillei; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus; cited as freminvillei). DISTRIBUTION: Samoa. freyi Bernhauer, 1941, see: japonicus Sharp, 1889. frontalis Wendeler, 1928c: 123 (Priochirus; subgenus Plastus; Type locality: Philippinen: Luzon: Los Baños). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. furcaticornis Scheerpeltz, 1965: 134 (Priochirus; subgenus Triacanthochirus; Type locality: N.E. Burma, Kambaiti, 200 m). DISTRIBUTION: Myanmar. galeatus Fauvel, 1902: 10 (Leptochirus; subgenus Priochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Plastus; Sumatra). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. gardneri Cameron, 1930: 454 (Priochirus; subgenus Plastus; Type locality: Bengal: Darjeeling, Rangirum, alt. 6000 feet). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Plastus; characters; East Pakistan). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1123
— P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). — Saha and Vazirani, 1971: 515 (Priochirus; subgenus Plastus; India). DISTRIBUTION: India, Bangladesh. gazella Cameron, 1928g: 428, 430 (Priochirus; subgenus Cephalomerus; Type locality: Borneo: Mt. Dulit, 3500 feet). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Borneo. germanus Cameron, 1937a: 85 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. gestroi Bernhauer, 1903: 137, 147 (Priochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; Sumatra). — Bernhauer, 1928: 2 (Priochirus; Sumatra). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. greensladei Herman, 2001: 40 (Priochirus; replacement name for minor Greenslade). DISTRIBUTION: New Guinea. minor P. J. M. Greenslade, 1971: 184 (Priochirus; subgenus Plastus; [preoccupied]; Type locality: New Guinea: Japen Is., camp 3, Central range, Mt. Ord, 3500 feet). — Herman, 2001: 40 (Priochirus; synonym of greensladei). gutturalis Bernhauer, 1915f: 135 (Priochirus; subgenus Triacanthus; Type locality: Sarawak: Mt. Matang, 1000 Fuss). — Cameron, 1930b: 160 (Priochirus; British North Borneo). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus; cited as guturalis). DISTRIBUTION: Borneo. haackei W. J. MacLeay, 1886: 142 (Leptochirus; Type locality: Fly River, New Guinea). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer and Schubert, 1910: 18 (Priochirus; synonym of monilicornis). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus; cited as haakei; cited as valid species). DISTRIBUTION: New Guinea. haemorrhous Sharp, 1887: 741 (Priochirus; Type locality: Guatemala, Cubilguitz, Chiacam). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 146 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 186 (Priochirus; subgenus Priochirus). DISTRIBUTION: Guatemala. helleri Wendeler, 1928c: 123 (Priochirus; subgenus Plastus; Type locality: Philippinen: Nord-Palawan: Binaluan). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. heterodon Fauvel, 1902: 14 (Leptochirus; subgenus Priochirus; Type locality: Nouvelle Guinée allemande). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 85, 86 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. 1124 OSORIINAE: PRIOCHIRUS Bulletin AMNH heurni Cameron, 1924: 83, 85 (Priochirus; subgenus Plastus; Type locality: Dormanpadbivak, 1410 m). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). — Scheerpeltz, 1971b: 210 (Priochirus; New Guinea). DISTRIBUTION: New Guinea. heynei Wendeler, 1927: 4 (Priochirus; Type locality: Costa Rica, Turrialba, 800 m). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 186 (Priochirus; subgenus Priochirus). DISTRIBUTION: Costa Rica. hirticornis Heller, 1910: 11 (Priochirus; Type locality: Papua, 780 m). — Cameron, 1924: 84 (Priochirus; subgenus Plastus; New Guinea). — Cameron, 1924: 86 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. hoplites Fauvel, 1895: 182 (Leptochirus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Ghécu, 1300–1400; Java). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 140 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1921a: 363, 399 (Priochirus; subgenus Cephalomerus; characters; Singapore). — Cameron, 1928d: 95 (Priochirus; subgenus Cephalomerus; Sumatra). — Cameron, 1930: 107 (Priochirus; subgenus Cephalomerus; characters; Burma; Malay Penin- sula; Singapore; Sumatra; Java). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1936a: 27 (Priochirus; subgenus Cephalomerus; Java; also listed from Burma, Sumatra, and Singapore). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Cephalomerus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). — Saha and Vazirani, 1971a: 101 (Priochirus; subgenus Cephalomerus; India). — Coiffait, 1978b: 113 (Priochirus; subgenus Cephalomerus; Bhutan). DISTRIBUTION: India, Bangladesh, Bhutan, Myanmar, Malaysia, Singapore, Indonesia. ignotus Cameron, 1943a: 336 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea). — P. J. M. Greenslade, 1971: 155, 164 (Priochirus; subgenus Syncampsochirus; characters). DISTRIBUTION: New Guinea. immarginatus Bernhauer, 1903: 137, 147 (Priochirus; Type locality: Perak). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Malaysia. impressifrons Heller, 1898: 17 (Priochirus; Type locality: Celebes meridionalis, mons Bonthain, Loka). — Bernhauer, 1903: 142 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Catacamptus; catalog). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Catacamptus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. inaequalis Bernhauer, 1903: 143, 153 (Priochirus; Type locality: Neu Guinea). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 85 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1125
— P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). — Scheerpeltz, 1971b: 201 (Priochirus; New Guinea). DISTRIBUTION: New Guinea. indicus Bernhauer, 1915h: 214 (Priochirus; subgenus Cephalomerus; Type locality: Nordost- Birma). — Cameron, 1930: 105 (Priochirus; subgenus Cephalomerus; characters; Burma). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Myanmar. infans Cameron, 1937a: 86 (Priochirus; subgenus Plastus; Type locality: New Guinea: Ishurava, alt. 3000 ft.). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. insignis Wendeler, 1928c: 124 (Priochirus; subgenus Plastus; Type locality: Philippinen: Süd-Ost-Luzon: Vivac). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. insularis Bernhauer, 1903, see: tridens Motschulsky, 1857. integrifrons Heller, 1898: 18 (Priochirus; Type locality: Celebes centralis, regio meridiana ad lacum “Poss”). — Bernhauer, 1903: 136 (Priochirus; characters). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. jacobsoni Cameron, 1945a: 140 (Priochirus; subgenus Plastus; Type locality: Java: Preanger; G. Patoeha). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. japonicus Sharp, 1889: 468 (Priochirus; Type locality: Hakone, Oyayama, island of Sado). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 139 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1949b: 456 (Priochirus; subgenus Cephalomerus; Formosa). — Nakane, 1963a: 81 (Priochirus; characters; habitus photograph; Japan). — Sawada, 1965a: 147 (Priochirus; Formosa). — P. J. M. Greenslade, 1971: 133 (Priochirus; subgenus Euleptarthrus). — Naomi, 1996a: 3 (Priochirus; subgenus Euleptarthrus; characters; Japan). — Watanabe, 1998: 313 (Priochirus; Japan). DISTRIBUTION: Taiwan, China, Japan. davidis Fairmaire, 1889: 11 (Leptochirus; Type locality: Moupin). — Heller, 1898: 7 (Priochirus; synonym of japonicus). — Bernhauer and Schubert, 1910: 16 (Priochirus; synonym of japonicus). freyi Bernhauer, 1941: 209 (Priochirus; subgenus Cephalomerus; Type locality: Japan: Kumanotaira, Kaurizawa). — P. J. M. Greenslade, 1971: 132 (Priochirus; subgenus Euleptarthrus). — Naomi, 1996a: 4 (Priochirus; synonym of japonicus). javanus Cameron, 1945a: 141 (Priochirus; subgenus Plastus; Type locality: Java: Preanger; G. Tangkoeban Prahoe, alt. 4000–5000 feet). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. 1126 OSORIINAE: PRIOCHIRUS Bulletin AMNH jhumlaensis Saha and Vazirani, 1971: 513 (Priochirus; subgenus Plastus; Type locality: India: N.E.F.A.: Kameng Division, Jhum La). DISTRIBUTION: India. jonapau P. J. M. Greenslade, 1971: 169 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Guadalcanal, Mt. Jonapau, 2600 feet). DISTRIBUTION: Solomon Islands. kaestneri Wendeler, 1957: 26 (Priochirus; subgenus Plastus; Type locality: Neu Guinea: Wandammengeb., Wasior). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. kambaitiensis Scheerpeltz, 1965: 141 (Priochirus; subgenus Plastus; Type locality: N.E. Burma, Kambaiti, 7000 ft.). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Myanmar. kamengensis Saha and Vazirani, 1971a: 99 (Priochirus; subgenus Cephalomerus; Type locality: India: NEFA: Kameng Div., Chug village, 800 m). DISTRIBUTION: India. karnyi Cameron, 1925a: 174 (Priochirus; subgenus Catacamptus; Type locality: Tjibodas). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Catacamptus; catalog). — Cameron, 1936a: 27 (Priochirus; subgenus Catacamptus; Java). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. kawamurai Bernhauer, 1943a: 170 (Priochirus; subgenus Cephalomerus; Type locality: Formosa: Horisha). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Taiwan. kimurai Naomi, 1996a: 14 (Priochirus; subgenus Plastus; Type locality: Okuma, Kunigami, Okinawa Is., Nansei Isls.). — Ito, 1997: 8 (Priochirus; subgenus Plastus; Japan). DISTRIBUTION: Japan. klimai Bernhauer, 1914c: 79 (Priochirus; subgenus Cephalomerus; Type locality: Java). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1936a: 27 (Priochirus; subgenus Cephalomerus; Java). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Indonesia. kraatzi Bernhauer, 1903: 138, 150 (Priochirus; Type locality: Celebes). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Bernhauer, 1926f: 311 (Priochirus; subgenus Triacanthochirus; Buru). — Wendeler, 1928c: 118 (Priochirus; subgenus Triacanthochirus; Philippines). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia, Philippines. kuluensis Bernhauer, 1914c: 78 (Priochirus; subgenus Plastus; Type locality: Kulu im Himalaya, 5000 Fuss). — Cameron, 1930: 113 (Priochirus; subgenus Plastus; characters; India). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: India. kuntzeni Wendeler, 1927, see: miles Bernhauer, 1903. lankcus Biswas and Sengupta, 1982: 144 (Priochirus; subgenus Plastus; Type locality: Sri Lanka: Central, Kandy, 700 m). DISTRIBUTION: Sri Lanka. Bulletin AMNH OSORIINAE: PRIOCHIRUS 1127 laticornis Sharp, 1887: 742 (Priochirus; Type locality: Mexico, Cordova). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 146 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 186 (Priochirus; subgenus Priochirus). DISTRIBUTION: Mexico. latifrons P. J. M. Greenslade, 1971: 134 (Priochirus; subgenus Leiochirus; Type locality: Sumatra: Lepong Tandai). DISTRIBUTION: Indonesia. latus P. J. M. Greenslade, 1971: 161 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea: Morobe District, Edie Creek, about 6 miles S.W. Wau, Bulldog Road, 9700 feet). DISTRIBUTION: New Guinea. lautus P. J. M. Greenslade, 1971: 145 (Priochirus; subgenus Exochirus; Type locality: Bismarck Archipelago: New Britain, Yalom, 1000 m). DISTRIBUTION: Papua New Guinea. lawrencei P. J. M. Greenslade, 1971: 185 (Priochirus; subgenus Plastus; Type locality: Solomon islands: S. Ysabel, Tatamba). DISTRIBUTION: Solomon Islands. laxus Sharp, 1887: 741 (Priochirus; Type locality: British Honduras, R. Sarstoon; Guatemala, Purula). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 145 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 186 (Priochirus; subgenus Priochirus). DISTRIBUTION: Belize, Guatemala. longicornis Fauvel, 1864b: 10 (Leptochirus; Type locality: Indes orientales). — Fauvel, 1865: 14 (Leptochirus; characters; East Indies). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 141 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Leptarthrus; catalog). — Cameron, 1930: 110 (Priochirus; subgenus Leptarthrus; characters; India). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Neoleptarthrus; catalog). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Leptarthrus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 133 (Priochirus; subgenus Euleptarthrus). — Coiffait, 1978b: 113 (Priochirus; subgenus Leptarthrus; Bhutan). — Coiffait, 1984: 117 (Priochirus; Sikkim). DISTRIBUTION: India, Bangladesh, Bhutan. opacinus Bernhauer, 1914c: 79 (Priochirus; subgenus Cephalomerus; Type locality: Ostindien, Chambagonoor). — Cameron, 1930: 110 (Priochirus; synonym of longicornis). — Scheerpeltz, 1933: 1005 (Priochrius; synonym of longicornis). longipennis Cameron, 1952: 243 (Priochirus; subgenus Plastus; Type locality: New Guinea: Snow Mountains: Lower Mist Camp, 1800 m). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. lorquini Fauvel, 1878f: 190 (Leptochirus; Type locality: Amboine; Vaigheu). — Fauvel, 1879: 73 (Leptochirus; Gilolo; Mortay). — Heller, 1898: 7 (Priochirus; checklist). — Bernhauer, 1903: 141 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). 1128 OSORIINAE: PRIOCHIRUS Bulletin AMNH
— Cameron, 1924: 82 (Priochirus; subgenus Syncampsochirus; characters in key; New Guinea). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 156, 157 (Priochirus; subgenus Syncampsochirus; characters). DISTRIBUTION: New Guinea, Indonesia. luzonicus Fauvel, 1886b: 143 (Leptochirus; Type locality: Luçon: Bilucao). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 144 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Plastus; catalog). — Wendeler, 1928c: 118 (Priochirus; subgenus Plastus; Philippines). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines, Indonesia. maitrta R. Dvorˇák, 1957b: 791 (Priochirus; subgenus Plastus; Type locality: India: Basti Karwar). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: India. major Cameron, 1920, see: bifoveatus Eppelsheim, 1895. malaisei Scheerpeltz, 1965: 136 (Priochirus; subgenus Cephalomerus; Type locality: N.E. Burma, Kambaiti, 2000 m). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus; cited as malasei). DISTRIBUTION: Myanmar. malayanus Cameron, 1936b: 40 (Priochirus; subgenus Neoleptarthrus; Type locality: Pahang: Cameron Highlands, 5000 ft). — P. J. M. Greenslade, 1971: 133 (Priochirus; subgenus Euleptarthrus). DISTRIBUTION: Malaysia. mandibularis Kraatz, 1859: 189 (Leptochirus; Type locality: Ceylan). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 140 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Peucodontus, cited as Pseucodontus; catalog). — Cameron, 1930: 108 (Priochirus; subgenus Cephalomerus; characters; Ceylon). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Peucodontus, cited as Pseucodontus; catalog). — P. J. M. Greenslade, 1971: 130 (Priochirus; subgenus Peucodontus). DISTRIBUTION: Sri Lanka. manilensis Bernhauer, 1912g: 247 (Priochirus; subgenus Plastus; Type locality: Luzon: Manila). — Wendeler, 1928c: 118 (Priochirus; subgenus Plastus; Philippines). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. mannaensis Bernhauer, 1915h: 213 (Priocharis; Type locality: Sumatra: Manna). — Cameron, 1928d: 95 (Priochirus; subgenus Plastus; Sumatra). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. masahiroi Naomi, 1996a: 6 (Priochirus; subgenus Euleptarthrus; Type locality: Mt. Kyoyo- midake, Shimogegun, Ohita Pref.). DISTRIBUTION: Japan. mayri Wendeler, 1957: 26 (Priochirus; subgenus Plastus; Type locality: Neu Guinea: Cyklopengebirge, Lager I). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. Bulletin AMNH OSORIINAE: PRIOCHIRUS 1129 melanesiae P. J. M. Greenslade, 1971: 172 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Gizo Island. Also cited from the Bismarck Archipelago, Caroline Islands, Moluccas, Philippines, and New Guinea). DISTRIBUTION: Caroline Islands, Solomon Islands, Papua New Guinea, Philippines, Indonesia. merula P. J. M. Greenslade, 1971: 180 (Priochirus; subgenus Plastus; Type locality: New Guinea: Madang District, Finisterre Mts., Naho R. Valley, Moro, 5550 feet). DISTRIBUTION: New Guinea. micrognathus Fauvel, 1902: 13 (Leptochirus; subgenus Priochirus; Type locality: Darjeeling ou Siam). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1930: 111 (Priochirus; subgenus Leptarthrus; characters; India or Siam). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Neoleptarthrus; catalog). — Paulian, 1941: 134 (Priochirus; subgenus Leptarthrus; larval characters; India). — Scheerpeltz, 1965: 99 (Priochirus; subgenus Neoleptathrus; Burma). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Leptarthrus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 133 (Priochirus; subgenus Euleptarthrus). DISTRIBUTION: Myanmar, Bangladesh, India or Thailand. micronesiae P. J. M. Greenslade, 1971: 169 (Priochirus; subgenus Syncampsochirus; Type locality: Palau Islands: Peleliu Island. Also cited from the Solomon Islands and Santa Cruz group). DISTRIBUTION: Palau, Solomon Islands. miles Bernhauer, 1903: 141, 152 (Priochirus; Type locality: Cooktown). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). — Bernhauer, 1920: 3 (Priochirus; Australia). — P. J. M. Greenslade, 1971: 156, 165 (Priochirus; subgenus Syncampsochirus; characters). — Newton, 1990: 91 (Priochirus). DISTRIBUTION: Australia. kuntzeni Wendeler, 1927: 3 (Priochirus; subgenus Syncampsochirus; Type locality: Brasilien). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 156 (Priochirus; synonym of miles; almost certainly from Australia not Brazil). mindanaoensis Wendeler, 1928c: 122 (Priochirus; subgenus Catacamptus; Type locality: Philippinen: Mindanao). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Catacamptus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. minor Bernhauer, 1928, see: exaratus Eppelsheim, 1895. minor Cameron, 1928, see: minusculus Scheerpeltz, 1933. minor P. J. M. Greenslade, 1971, see: greensladei Herman, 2001. minusculus Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; replacement name for minor Cameron). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Borneo. minor Cameron, 1928g: 430 (Priochirus; subgenus Cephalomerus; [preoccupied]; Type locality: Borneo: Mt. Dulit, 3500 feet). minutus Wendeler, 1928c: 125 (Priochirus; subgenus Plastus; Type locality: Philippinen: Luzon: Mt. Banahao, Imugan; Mindanao: Surigao). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. 1130 OSORIINAE: PRIOCHIRUS Bulletin AMNH mjobergi Bernhauer, 1928: 4 (Priochirus; subgenus Plastus; cited as mjöbergi; Type locality: Sumatra: Medan). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; cited as mjöbergi; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus; cited as mjobergi). — Herman, 2001: 12 (Priochrius; spelling corrected to mjobergi). DISTRIBUTION: Indonesia. mjobergi Cameron, 1928, see: mjobergianus Scheerpeltz, 1933. mjobergianus Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; cited as mjöbergianus; replacement name for mjobergi Cameron, cited as mjöbergi). — Hammond, 1984: 199 (Priochirus; cited as mjoebergianus; Borneo). — Herman, 2001: 12 (Priochirus; spelling corrected to mjobergianus). DISTRIBUTION: Borneo. mjobergi Cameron, 1928g: 425, 427 (Priochirus; [preoccupied]; subgenus Triacan- thochirus; cited as mjöbergi; [Note: Since this name is based on the Swedish name Mjöberg, the umlaut should be dropped without further modification and the name should be emended to mjobergi (Article 32.5.2.1)]; Type locality: Songei Tutau; Mt. Murud, 5000–7000 feet). — Scheerpeltz, 1933: 1002 (Priochirus; cited as mjöbergi; synonym of mjobergianus, cited as mjöbergianus). — Hammond, 1984: 199 (Priochirus; cited as mjobergi; synonym of mjobergianus, cited as mjoebergianus). modiglianii Bernhauer, 1903: 138, 150 (Priochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; Sumatra). — Bernhauer, 1928: 2 (Priochirus; Sumatra). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. punctiventris Bernhauer, 1903: 138, 151 (Priochirus; Type locality: Berg Singalang auf Sumatra). — Bernhauer and Schubert, 1910: 15 (Priochirus; synonym of modiglianii). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; variety of modiglianii; Sumatra). apiciventris Bernhauer, 1928: 2 (Priochirus; variety of modiglianii; Type locality: Medan, Sibajakvulkan). — Scheerpeltz, 1933: 1002 (Priochirus; variety of modiglianii). monilicornis Fauvel, 1878f: 193 (Leptochirus; Type locality: Iles Arou, Wokan; Nouvelle- Guinée, Andai, Hatam, Ramoi; Amberbaki). — Fauvel, 1879: 73 (Leptochirus; notes; Mysol; Corei; Katau; Fly River). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 143 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 85 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea, Indonesia. monstrosus Wendeler, 1957: 25 (Priochirus; subgenus Plastus; Type locality: Karolinen: Panope). — P. J. M. Greenslade, 1971: 175, 184 (Priochirus; subgenus Plastus; Caroline Islands). DISTRIBUTION: Caroline Islands. montanus Wendeler, 1928c: 122 (Priochirus; subgenus Catacamptus; Type locality: Philippinen: Luzon: Mt. Polis; Nord-Palawan: Binaluan). — Scheerpeltz, 1933: 1006 (Priochirus; subgenus Catacamptus; catalog). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1131
— P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. monticola Cameron, 1952: 242 (Priochirus; subgenus Catacamptus; Type locality: New Guinea, Snow Mountains, Moss Forest Camp, 2600–2800 m). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. moshingensis Saha and Vazirani, 1971: 511 (Priochirus; subgenus Plastus; Type locality: India: N.E.F.A.: Kameng Division, Moshing village, 750 m). DISTRIBUTION: India. moultoni Bernhauer, 1915f: 134 (Priochirus; subgenus Triacanthus; Type locality: Britisch- Nordborneo: Mt. Kinabalu, 4500 Fuss). — Cameron, 1928f: 399 (Priochirus; Borneo). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — Cameron, 1933b: 338 (Priochirus; subgenus Triacanthochirus; British North Borneo). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Borneo. mushanus Bernhauer, 1943a: 169 (Priochirus; subgenus Cephalomerus; Type locality: Formosa: Musha). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Taiwan. mutillatus Cameron, 1937a: 84 (Priochirus; subgenus Triacanthochirus; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft.). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus; cited as mutilatus). DISTRIBUTION: New Guinea. nanus Heller, 1898, see: Thoracochirus. naturangai P. J. M. Greenslade, 1971: 145 (Priochirus; subgenus Exochirus; Type locality: Solomon Islands: Russell Islands, Yandina). DISTRIBUTION: Solomon Islands. neptunus Heller, 1898: 19 (Priochirus; Type locality: Java; Sumatra, mons Singalang). — Bernhauer, 1903: 138 (Priochirus; characters). — Bernhauer and Schubert, 1910: 15 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; illustration; Sumatra). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — Cameron, 1936a: 26 (Priochirus; subgenus Triacanthochirus; Java). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Malaysia, Indonesia. pluripunctus Cameron, 1950d: 2 (Priochirus; subgenus Triancanthochirus; subspecies of neptunus; Type locality: Pahang: Cameron Highlands, Kuala Tinta, 3000 feet). nigrita Fauvel, 1902, see: bipunctatus Fauvel, 1895. novaebritanniae Bernhauer, 1915g: 180 (Priochirus; subgenus Syncampsochirus; Type locality: Not cited). — Cameron, 1924: 86 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Cameron, 1931: 355 (Priochirus; cited as novaebritaniae; New Guinea). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. novaeguineae Bernhauer, 1915g: 179 (Priochirus; subgenus Syncampsochirus; Type locality: Deutsch-Neu-Guinea: Sepik, Berg 1570 m). — Cameron, 1924: 82 (Priochirus; subgenus Syncampsochirus; characters in key; New Guinea). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. 1132 OSORIINAE: PRIOCHIRUS Bulletin AMNH olthofi Cameron, 1952: 243 (Priochirus; subgenus Plastus; Type locality: New Guinea: Snow Mountains: Bernhard Camp, 50 m). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. opacicollis Fauvel, 1878f: 193 (Leptochirus; Type locality: Nouvelle-Guinée, Ramoi). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 144 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 85 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. opacinus Bernhauer, 1914, see: longicornis Fauvel, 1864. palawanensis Wendeler, 1928c: 120 (Priochirus; subgenus Cephalomerus; Type locality: Philippinen: Palawan: Binaluan). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Philippines. pannosus P. J. M. Greenslade, 1971: 180 (Priochirus; subgenus Plastus; Type locality: New Guinea: Morobe District, Edie Creek, ca. 6 miles S.W. Wau, 7000 feet). DISTRIBUTION: New Guinea. papuanus Cameron, 1924: 84, 86 (Priochirus; subgenus Plastus; Type locality: Doorman- padbivak; Mamberamo Riv.). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: Philippines. parabolicus Heller, 1898: 18 (Priochirus; Type locality: Celebes meridionalis, mons Lompo Batang, 3000 ped.). — Bernhauer, 1903: 136 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Triacanthus; catalog). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. paradoxus Wendeler, 1928c: 125 (Priochirus; subgenus Plastus; Type locality: Philippinen: Luzon: Balbalan, Imugan, Laguna; Mindanao: Surigao). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. parcus Fauvel, 1878f: 187 (Leptochirus; Type locality: Nouvelle Guinée occidentale). — Fauvel, 1879: 72 (Leptochirus; notes; Vaigheu; Ramoi; Fly River; Amberbaki). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 144 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 86 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. parvicollis Fauvel, 1902: 18 (Leptochirus; subgenus Priochirus; Type locality: Colombie: Cachabe). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 186 (Priochirus; subgenus Priochirus). DISTRIBUTION: Colombia. Bulletin AMNH OSORIINAE: PRIOCHIRUS 1133 parvidens Cameron, 1928g: 426, 427 (Priochirus; subgenus Triacanthochirus; Type locality: Borneo: Mt. Murud, 5000–6000 feet). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — Cameron, 1933b: 338 (Priochirus; subgenus Triacanthochirus; British North Borneo). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Borneo. parvus Cameron, 1928g: 430 (Priochirus; subgenus Syncampsochirus; Type locality: Borneo: Mt. Dulit, 3500 feet). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: Borneo. pectus Wendeler, 1928c: 126 (Priochirus; subgenus Plastus; Type locality: Philippinen: Siargao: Cabuntug). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. penelope P. J. M. Greenslade, 1971: 163 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Kolombangara, Kuzi). DISTRIBUTION: Solomon Islands. peninsularis Cameron, 1929b: 436 (Priochirus; subgenus Triacanthochirus; Type locality: Selangor: The Gap). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Malaysia. pentagonalis Bernhauer, 1914c: 78 (Priochirus; subgenus Triochirus; Type locality: Chambaganoor: Madura, Südindien). — Cameron, 1930: 98 (Priochirus; subgenus Triacanthochirus; characters; India). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). — Biswas and Sengupta, 1982: 142 (Priochirus; subgenus Triacanthochirus; characters; India). DISTRIBUTION: India. analis Cameron, 1920a: 142 (Priochirus; subgenus Triacanthus; variety of fletcheri; Type locality: Kodaikanal, S. India). — Cameron, 1930: 98 (Priochirus; subgenus Triacanthochirus; synonym of pentagonalis). — Scheerpeltz, 1933: 1002 (Priochirus; synonym of pentagonalis). — Biswas and Sengupta, 1982: 142 (Priochirus; synonym of pentagonalis). peomahat P. J. M. Greenslade, 1971: 167 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Guadalcanal, Mt. Peomahat, 4650 feet). DISTRIBUTION: Solomon Islands. peregrinus Wendeler, 1928c: 126 (Priochirus; subgenus Plastus; Type locality: Philippinen: Panaon). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. persimilis Cameron, 1928a: 35 (Priochirus; subgenus Cephalomerus; Type locality: Sumatra; Java). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Indonesia. philippinus Bernhauer, 1912g: 245 (Priochirus; subgenus Plastus; Type locality: Luzon: Benguet, Baguio). — Wendeler, 1928c: 118 (Priochirus; subgenus Plastus; Philippines). 1134 OSORIINAE: PRIOCHIRUS Bulletin AMNH
— Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. pluripunctus Cameron, 1950, see: neptunus Heller, 1898. politiventris Fauvel, 1902: 15 (Leptochirus; subgenus Priochirus; Type locality: Amberbaki). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1924: 82 (Priochirus; subgenus Syncampsochirus; characters in key; New Guinea). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 156, 157 (Priochirus; subgenus Syncampsochirus; characters). DISTRIBUTION: New Guinea. popamanasiu P. J. M. Greenslade, 1971: 150 (Priochirus; subgenus Exochirus; Type locality: Solomon Islands: Guadalcanal, Mt. Popamanasiu, 6000 feet). DISTRIBUTION: Solomon Islands. poseidon Bernhauer, 1903: 138, 148 (Priochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; illustration; Sumatra). — Bernhauer, 1928: 2 (Priochirus; Sumatra). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. propinquus Bernhauer, 1903: 142, 152 (Priochirus; Type locality: Neu Guinea). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). — Bernhauer, 1915g: 180 (Priochirus; subgenus Syncampsochirus; notes). — Cameron, 1924: 82 (Priochirus; subgenus Syncampsochirus; characters in key; New Guinea). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1937a: 84 (Priochirus; subgenus Syncampsochirus; New Guinea). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. proximus Cameron, 1928g: 429, 430 (Priochirus; subgenus Cephalomerus; Type locality: Borneo: Mt. Matang). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Borneo. puncticeps Cameron, 1943c: 183 (Priochirus; subgenus Triacanthochirus; Type locality: British North Borneo: Mt. Kinabalu, Lumu Lumu, altitude 5500 feet). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Borneo. punctiventris Bernhauer, 1903, see: modiglianii Bernhauer, 1903. pygmaeoides Cameron, 1929b: 437 (Priochirus; subgenus Cephalomerus; Type locality: Singapore: Bukit Timah). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Singapore. pygmaeus Kraatz, 1859: 191 (Leptochirus; Type locality: India orientalis). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 140 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1921a: 363, 399 (Priochirus; subgenus Cephalomerus; characters; Singapore). — Cameron, 1928d: 95 (Priochirus; subgenus Cephalomerus; Sumatra). — Bernhauer, 1928: 3 (Priochirus; Sumatra). — Cameron, 1930: 106 (Priochirus; subgenus Cephalomerus; characters; East Indies). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1135
— Cameron, 1934d: 77 (Leptochirus; synonym of excavatus). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: “East Indies”, Singapore, Indonesia. quadricollis Cameron, 1924: 81, 83 (Priochrius; subgenus Syncampsochirus; Type locality: Doormanpadbivak, 1410 m). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. quadridens P. J. M. Greenslade, 1971: 168 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea: Madang District, Finisterre Mts., Moro, ca. 5550 feet). DISTRIBUTION: New Guinea. quadrifidus Fauvel, 1878f: 194 (Leptochirus; Type locality: Amboine). — Fauvel, 1879: 73 (Leptochirus; notes; Gilolo; Ceram, Wahaai; Amberbaki; Fly River). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 141 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1924: 82 (Priochirus; subgenus Syncampsochirus; characters in key; New Guinea). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — P. J. M. Greenslade, 1971: 156, 158 (Priochirus; subgenus Syncampsochirus; characters). DISTRIBUTION: New Guinea, Indonesia. quadrifoveatus P. J. M. Greenslade, 1971: 133 (Priochirus; subgenus Euleptarthrus; Type locality: Formosa: Musha). DISTRIBUTION: Taiwan. rastri P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus; replacement name for fraterculus Cameron, 1937). DISTRIBUTION: New Guinea. fraterculus Cameron, 1937a: 85 (Priochirus; subgenus Syncampsochirus; [preoccupied]; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft.). — P. J. M. Greenslade, 1971: 141 (Priochirus; synonym of rastri). rennellensis P. J. M. Greenslade, 1971: 165 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Rennell Island: Niupari). DISTRIBUTION: Solomon Islands. reno P. J. M. Greenslade, 1971: 151 (Priochirus; subgenus Exochirus; Type locality: Bismarck Archipelago: New Ireland, Lemkamin). DISTRIBUTION: Papua New Guinea. robustus Wendeler, 1928c: 127 (Priochirus; subgenus Plastus; Type locality: Philippinen: Luzon: Laguna). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. rubiginosus Cameron, 1930: 105 (Priochirus; subgenus Cephalomerus; Type locality: Sikkim: Gopaldhara, Rungbong Valley; Assam: Naga Hills). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). — Scheerpeltz, 1965: 98 (Priochirus; subgenus Cephalomerus; Sikkim). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Cephalomerus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). — Biswas and Sengupta, 1982: 143 (Priochirus; subgenus Cephalomerus; characters; India). — Biswas and Biswas, 1985: 143 (Priochirus; subgenus Cephalomerus; India). DISTRIBUTION: India, Bangladesh. ruficrus Heller, 1898: 16 (Priochirus; Type locality: Celebes septentrionalis, Tomohon). — Bernhauer, 1903: 144 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). 1136 OSORIINAE: PRIOCHIRUS Bulletin AMNH
— Wendeler, 1928c: 118 (Priochirus; subgenus Plastus; Philippines). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 175 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines, Indonesia. rufipennis Cameron, 1949, see: silvestrii Bernhauer, 1927. rufus Cameron, 1920a: 142 (Priochirus; subgenus Cephalomerus; Type locality: Santikoppa, North Coorg). — Cameron, 1930: 104 (Priochirus; subgenus Cephalomerus; characters; India). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). — Biswas and Sengupta, 1982: 143 (Priochirus; subgenus Cephalomerus; characters; India). DISTRIBUTION: India. salvini Sharp, 1887: 742 (Priochirus; Type locality: Guatemala, Volcan de Fuego 6400 feet, Quiche Mountains 7000 to 9000 feet, Cerro Zunil, Calderas 7000 feet, Volcan de Agua 8500 to 10,500 feet, San Geronimo, Sabo). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 146 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 186 (Priochirus; subgenus Priochirus). DISTRIBUTION: Guatemala. samoensis Blanchard, 1853: 54 (Leptochirus; Type locality: Samoa). — Fauvel, 1877: 184 (Leptochirus; characters; Fiji; Samoa; New Guinea; Arou; Ternate; Aus- tralia). — Fauvel, 1878f: 192 (Leptochirus; characters; New Guinea; Aru Islands; Samoa; Fiji; Australia). — Fauvel, 1878g: 480 (Leptochirus; characters; notes; Australia). — Fauvel, 1879: 73 (Leptochirus; Gilolo; Batcian; Mysol; Fly River). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 141 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). — Bernhauer, 1915g: 180 (Priochirus; subgenus Syncampsochirus; Polynesia). — Cameron, 1924: 82 (Priochirus; subgenus Syncampsochirus; characters in key; New Guinea). — Bernhauer, 1926f: 311 (Priochirus; subgenus Syncampsochirus; Buru). — Cameron, 1927a: 21 (Priochirus; subgenus Syncampsochirus; Samoa). — Wendeler, 1928c: 118 (Priochirus; subgenus Syncampsochirus; Philippines). — Cameron, 1931: 355 (Priochirus; New Guinea). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1937a: 84 (Priochirus; subgenus Syncampsochirus; New Guinea). — P. J. M. Greenslade, 1971: 156, 165, 173 (Priochirus; subgenus Syncampsochirus; characters; Samoa). DISTRIBUTION: Indonesia, Philippines, New Guinea, Australia, Fiji, Samoa. sanguinosus Motschulsky, 1857a: 501 (Leptochirus; Type locality: Indes orientales). — Kraatz, 1859: 191 (Leptochirus). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 140 (Priochirus; characters). — Fauvel, 1904a: 44 (Leptochirus; Nilghiris; Anamalai; Bírmanie; Tenasserim; Java). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1930: 103 (Priochirus; subgenus Cephalomerus; characters; Burma). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). — Cameron, 1936a: 27 (Priochirus; subgenus Cephalomerus; Java). — Paulian, 1941: 136 (Priochirus; larval characters; India). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). — Saha and Vazirani, 1971a: 100 (Priochirus; subgenus Cephalomerus; India). — Coiffait, 1978b: 113 (Priochirus; subgenus Cephalomerus; Bhutan). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1137
— Biswas and Sengupta, 1982: 144 (Priochirus; subgenus Cephalomerus; characters; India). — Coiffait, 1984: 117 (Priochirus; India). — Biswas and Biswas, 1985: 143 (Priochirus; subgenus Cephalomerus; India). DISTRIBUTION: “East Indies”, India, Bhutan, Myanmar, Malaysia, Indonesia. uniformis Cameron, 1929b: 436 (Priochirus; subgenus Cephalomerus; variety of san- guinosus; Type locality: Selangor: The Gap). — Scheerpeltz, 1933: 1004 (Priochirus; variety of sanguinosus). schuberti Wendeler, 1957: 26 (Priochirus; subgenus Priochirus; Type locality: Lombok: Segareh Anak). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Indonesia. schultzei Bernhauer, 1912g: 246 (Priochirus; subgenus Plastus; Type locality: Mindoro: Bongabon). — Wendeler, 1928c: 118 (Priochirus; subgenus Plastus; Philippines). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: Philippines. scyphus P. J. M. Greenslade, 1971: 151 (Priochirus; subgenus Exochirus; Type locality: New Guinea: Madang District, Finisterre Mts., Bodemu, about 4000 feet). DISTRIBUTION: New Guinea. sedlaceki P. J. M. Greenslade, 1971: 162 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Gizo Island). DISTRIBUTION: Solomon Islands. selangorensis Cameron, 1929b: 437 (Priochirus; subgenus Cephalomerus; Type locality: Selangor: The Gap. Perak). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). DISTRIBUTION: Malaysia. separandus Cameron, 1942c: 838 (Priochirus; subgenus Syncampsochirus; Type locality: Labuan: Batjan). — P. J. M. Greenslade, 1971: 156, 157 (Priochirus; subgenus Syncampsochirus; characters). DISTRIBUTION: Malaysia. sexdentatus Bernhauer, 1903: 143, 153 (Priochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Plastus; Sumatra). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. sikkimensis Cameron, 1930: 454 (Priochirus; subgenus Plastus; Type locality: Bengal: Darjeeling, Debrepanni, alt. 6000 feet). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — Paulian, 1941: 137 (Priochirus; larval characters; India). — Abdullah and Qadri, 1970: 117 (Priochirus; subgenus Plastus; characters; East Pakistan). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). — Saha and Vazirani, 1971: 514 (Priochirus; subgenus Plastus; India). DISTRIBUTION: India, Bangladesh. silvestrii Bernhauer, 1927f: 260 (Priochirus; subgenus Cephalomerus; Type locality: Formosa: Funkiko). — Scheerpeltz, 1933: 1004 (Priochirus; subgenus Cephalomerus; catalog). — P. J. M. Greenslade, 1971: 137 (Priochirus; subgenus Cephalomerus). DISTRIBUTION: Taiwan. rufipennis Cameron, 1949b: 457 (Priochirus; subgenus Cephalomerus; variety of silvestrii; Type locality: Formosa: Arisan). 1138 OSORIINAE: PRIOCHIRUS Bulletin AMNH similis Wendeler, 1928c: 119 (Priochirus; subgenus Triacanthochirus; Type locality: Philippinen: Basilan). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Philippines. simplex Fauvel, 1902: 15 (Leptochirus; subgenus Priochirus; Type locality: Nouvelle Guinée allemande). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 85 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 176, 178 (Priochirus; subgenus Plastus; characters; Solomon Islands; New Guinea). DISTRIBUTION: New Guinea. simus P. J. M. Greenslade, 1971: 167 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea: Buru, 1300 m). DISTRIBUTION: New Guinea. singalangensis Bernhauer, 1928: 5 (Priochirus; Type locality: Sumatra: Singalang). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Priochirus; catalog). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. sinuatus Fauvel, 1902: 14 (Leptochirus; subgenus Priochirus; Type locality: Nouvelle Bretagne). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 87 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141, 155 (Priochirus; subgenus Exochirus; characters; Bismarck Archipelago). DISTRIBUTION: Papua New Guinea. socius Fauvel, 1902: 12 (Leptochirus; subgenus Priochirus; Type locality: Sumatra: Si-Rambé). — Bernhauer, 1903: 145, 154 (Priochirus; characters; Sumatra). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Priochirus; catalog). — Cameron, 1928d: 95 (Priochirus; Sumatra). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. solomonensis P. J. M. Greenslade, 1971: 160 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Guadalcanal Kukum). DISTRIBUTION: Solomon Islands. sondaicus Bernhauer, 1928: 3 (Priochirus; subgenus Triacanthochirus; Type locality: Bandar Baroe). — Scheerpeltz, 1933: 1002 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. sororculus Cameron, 1937a: 85 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea: Orrori, alt. 3000 ft). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. sparsus Cameron, 1924: 84, 86 (Priochirus; subgenus Plastus; Type locality: Mamberamo Riv.; Idenburg Riv.). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. spinosulus Fauvel, 1878f: 190 (Leptochirus; Type locality: Nouvelle-Guinée, Hatam). — Heller, 1898: 8 (Priochirus; checklist). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1139
— Bernhauer, 1903: 142 (Priochirus; characters). — Bernhauer and Schubert, 1910: 17 (Priochirus; subgenus Syncampsochirus; catalog). — Bernhauer, 1915g: 180 (Priochirus; subgenus Syncampsochirus; New Guinea). — Cameron, 1924: 83 (Priochirus; subgenus Syncampsochirus; characters in key; New Guinea). — Cameron, 1931: 355 (Priochirus; New Guinea). — Scheerpeltz, 1933: 1005 (Priochirus; subgenus Syncampsochirus; catalog). — Cameron, 1937a: 84 (Priochirus; subgenus Syncampsochirus; New Guinea). — P. J. M. Greenslade, 1971: 141, 148 (Priochirus; subgenus Exochirus; New Guinea). DISTRIBUTION: New Guinea. staudingeri Bernhauer, 1926b: 39 (Priochirus; subgenus Plastus; Type locality: Java). — Scheerpeltz, 1933: 1007 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. steeli P. J. M. Greenslade, 1971: 158 (Priochirus; subgenus Syncampsochirus; Type locality: New Guinea: Maffin Bay). DISTRIBUTION: New Guinea. subbrevicornis Bernhauer, 1934: 1 (Priochirus; subgenus Leptarthrus; Type locality: Mittel-Szechuan: Hwa-Yin-Shan, 1600 m). — P. J. M. Greenslade, 1971: 133 (Priochirus; subgenus Euleptarthrus). DISTRIBUTION: China. subconvexus Bernhauer, 1917, see: ebeninus Erichson, 1840. subinaequalis Cameron, 1937a: 86 (Priochirus; subgenus Plastus; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft). — P. J. M. Greenslade, 1971: 176, 184 (Priochirus; subgenus Plastus; New Guinea). DISTRIBUTION: New Guinea. subnitidus Bernhauer, 1903: 138, 149 (Priochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; Sumatra). DISTRIBUTION: Indonesia. subtridens Heller, 1898: 19 (Priochirus; Type locality: Celebes meridionalis, mons Lompo Batang, 3000 ped.). — Bernhauer, 1903: 137 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Triacanthus; catalog). — Wendeler, 1928c: 118 (Priochirus; subgenus Triacanthochirus; Philippines). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia, Philippines. sulcifer Fauvel, 1902: 12 (Leptochirus; subgenus Priochirus; Type locality: Sumbawa). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Triacanthus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. sumatrensis Bernhauer, 1903: 137, 146 (Priochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; illustration; Sumatra). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Triacanthochirus; catalog). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). DISTRIBUTION: Indonesia. taprobanus Cameron, 1930: 116 (Priochirus; subgenus Plastus; Type locality: Ceylon). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Plastus; catalog). — Scheerpeltz, 1954: 110 (Priochirus; subgenus Plastus; Ceylon). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: Sri Lanka. 1140 OSORIINAE: PRIOCHIRUS Bulletin AMNH taurus Fauvel, 1902: 14 (Leptochirus; subgenus Priochirus; Type locality: Halmaheira: Patani; Batcian). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 85 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea, Indonesia. terebra P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus; Type locality: Bismarck Archipelago: New Britain, Yalom, 1000 m). DISTRIBUTION: Papua New Guinea. teres P. J. M. Greenslade, 1971: 142 (Priochirus; subgenus Exochirus; Type locality: New Guinea: Morobe District, Edie Creek, about 6 miles S.W. Wau, Bulldog Road, 9700 feet). DISTRIBUTION: New Guinea. tetraodon Bernhauer, 1903: 144, 154 (Priochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Plastus; Sumatra). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: Indonesia. tinahula P. J. M. Greenslade, 1971: 143 (Priochirus; subgenus Exochirus; Type locality: Solomon [cited as Solmon] Islands: Guadalcanal, Gold Ridge, 2600 feet). DISTRIBUTION: Solomon Islands. tonkinensis Bernhauer, 1903: 137, 148 (Priochirus; Type locality: Tonkin). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Triacanthus; catalog). — Bernhauer, 1922c: 221 (Priochirus; Formosa). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Triacanthochirus; catalog). — Cameron, 1949b: 456 (Priochirus; subgenus Triacanthochirus; China, Formosa). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). — Frank, 1982: 8 (Priochirus; list of parasites). DISTRIBUTION: Vietnam, China, Taiwan. toxopei Cameron, 1952: 243 (Priochirus; subgenus Plastus; Type locality: New Guinea: Hollandia). — P. J. M. Greenslade, 1971: 176, 178 (Priochirus; subgenus Plastus; New Guinea). DISTRIBUTION: New Guinea. tridens Motschulsky, 1857a: 502 (Leptochirus; Type locality: Indes orientales). — Kraatz, 1859: 191 (Leptochirus). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 139 (Priochirus; characters). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; Sumatra). — Bernhauer, 1928: 2 (Priochirus; Sumatra). — Cameron, 1930: 99 (Priochirus; subgenus Triacanthochirus; characters; Burma; Assam; Malay Peninsula; Java; Sumatra). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Triacanthochirus; catalog). — Scheerpeltz, 1935a: 598 (Priochirus; Java). — Cameron, 1936a: 26 (Priochirus; subgenus Triacanthochirus; Java). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). — Coiffait, 1981f: 332 (Priochirus; Andaman Islands). — Biswas and Biswas, 1985: 143 (Priochirus; subgenus Triacanthochirus; India). DISTRIBUTION: India, Andaman Islands, Myanmar, Malaysia, Indonesia, Philippines. insularis Bernhauer, 1903: 139 (Priochirus; subspecies of tridens; Type locality: Mentawei). — Bernhauer and Schubert, 1910: 16 (Priochirus; synonym of tridens). Bulletin AMNH OSORIINAE: PRIOCHIRUS 1141
— Cameron, 1921a: 363, 399 (Priochirus; subgenus Triacanthus; variety of tridens; characters; Singapore). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; variety of tridens; Sumatra). — Scheerpeltz, 1933: 1003 (Priochirus; synonym of tridens). trifidus Bernhauer, 1903: 139 (Priochirus; subspecies of tridens; Type locality: Java). — Bernhauer and Schubert, 1910: 16 (Priochirus; synonym of tridens). — Wendeler, 1928c: 118 (Priochirus; subgenus Triacanthochirus; subspecies of tridens; Philip- pines). depressus Bernhauer, 1903: 139 (Priochirus; subspecies of tridens; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 16 (Priochirus; synonym of tridens). — Cameron, 1928d: 95 (Priochirus; subgenus Triacanthochirus; variety of tridens; Sumatra). — Wendeler, 1928c: 118 (Priochirus; subgenus Triacanthochirus; subspecies of tridens; Philippines). trifidus Bernhauer, 1903, see: tridens Motschulsky, 1857. unicolor Laporte, 1835: 125 (Leptochirus; Type locality: Java). — Erichson, 1840: 826 (Leptochirus; characters; Java). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 136 (Priochirus; characters). — Fauvel, 1905e: 77 (Leptochirus; Java). — Bernhauer and Schubert, 1910: 16 (Priochirus; subgenus Triacanthus; catalog). — Cameron, 1928d: 94 (Priochirus; subgenus Triacanthochirus; illustration; Sumatra). — Scheerpeltz, 1933: 1003 (Priochirus; subgenus Triacanthochirus; catalog). — Scheerpeltz, 1935a: 598 (Priochirus; Java). — Cameron, 1936a: 26 (Priochirus; subgenus Triacanthochirus; Java). — P. J. M. Greenslade, 1971: 138 (Priochirus; subgenus Eutriacanthus). — Frank, 1982: 8 (Priochirus; list of parasites). DISTRIBUTION: Indonesia. coronatus Sachse, 1852: 146 (Leptochirus; Type locality: Java). — Heller, 1898: 8 (Priochirus; synonym of unicolor). — Fauvel, 1905e: 77 (Leptochirus; synonym of unicolor). — Bernhauer and Schubert, 1910: 16 (Priochirus; synonym of unicolor). valens Heller, 1910: 11 (Priochirus; subgenus Plastus; Type locality: Papua). — Cameron, 1924: 85, 86 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 141 (Priochirus; subgenus Exochirus). DISTRIBUTION: New Guinea. vastus Wendeler, 1928, see: currani Bernhauer, 1912. vitulus Fauvel, 1878f: 195 (Leptochirus; Type locality: Nouvelle-Guinée occidentale). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 145 (Priochirus; characters). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — Cameron, 1924: 85 (Priochirus; subgenus Plastus; characters in key; New Guinea). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. voeltzkowi Bernhauer, 1917m: 554 (Priochirus; Type locality: Anjouan, Comorenarchipel). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: Comoro Islands. volucer P. J. M. Greenslade, 1971: 167 (Priochirus; subgenus Syncampsochirus; Type locality: Solomon Islands: Bougainville, Kukugai village, 50 m). DISTRIBUTION: Solomon Islands. 1142 OSORIINAE: PRIOCHIRUS Bulletin AMNH wagneri Cameron, 1931: 355 (Priochirus; Type locality: Finsch Haven, New Guinea). — Scheerpeltz, 1933: 1008 (Priochirus; subgenus Plastus; catalog). — Cameron, 1937a: 86 (Priochirus; subgenus Plastus; New Guinea). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: New Guinea. weissi Fauvel, 1903a: 172 (Leptochirus; subgenus Priochirus; Type locality: Tuyen-Quan, Tonkin Central). — Bernhauer and Schubert, 1910: 18 (Priochirus; subgenus Plastus; catalog). — P. J. M. Greenslade, 1971: 176 (Priochirus; subgenus Plastus). DISTRIBUTION: Vietnam. yakushimensis Herman, 2001: 41 (Priochirus; replacement name for bicornis Nakane and Sawada). DISTRIBUTION: Japan. bicornis Nakane and Sawada, 1960: A121 (Borolinus; subgenus Paraborolinus; [preoccu- pied]; Type locality: Japan: Kosugidani Yakushima). — Nakane, 1963a: 83 (Borolinus; characters; habitus photograph; Japan). — Watanabe and Shibata, 1972: 60 (Borolinus; Yaku-shima Island). — Frank, 1982: 6 (Borolinus; list of parasites). — Naomi, 1996a: 11 (Priochirus; subgenus Paraborolinus; characters; Japan). — Herman, 2001: 41 (Priochirus; synonym of yakushimensis). yalomis P. J. M. Greenslade, 1971: 183 (Priochirus; subgenus Plastus; Type locality: Bismarck Archipelago: New Britain, Yalom, 1000 m). DISTRIBUTION: Papua New Guinea. yasutoshii Naomi, 1996a: 9 (Priochirus; subgenus Leiochirus; Type locality: Hatsuno, Amami Is., Nansei Isls.). DISTRIBUTION: Japan.
Thoracochirus [21 species; Palaearctic, Oriental, and Australian Regions] Thoracochirus Bernhauer, 1903: 116, 155, 160 (species included: nanus, rugosus, variolosus, verrucifer, forsteri). TYPE SPECIES: Leptochirus rugosus Fauvel, fixed by subsequent designation by Blackwelder, 1952: 388. — Bernhauer and Schubert, 1910: 18 (world catalog; 9 species). — Cameron, 1925: 6 (catalog of Indian species). — Bernhauer, 1926d: 254 (key to species of Philippines). — Cameron, 1930: 118 (characters; key to British Indian species). — Scheerpeltz, 1933: 1008 (world catalog supplement). — Shibata, 1973c: 27 (checklist of species of Taiwan). — Hammond, 1984: 199 (checklist of species of Borneo). aspericollis Fauvel, 1902: 20 (Leptochirus; subgenus Priochirus; Type locality: Java occi- dental: Pengalengan, 1300 m). — Bernhauer and Schubert, 1910: 18 (Thoracochirus; catalog). — Cameron, 1936a: 28 (Thoracochirus; Java). DISTRIBUTION: Indonesia. assamensis Cameron, 1930: 120 (Thoracochirus; Type locality: Assam: Naga Hills, alt. 9000 feet). — Scheerpeltz, 1933: 1008 (Thoracochirus; catalog). — Coiffait, 1978b: 113 (Thoracochirus; Bhutan). — Biswas and Sengupta, 1982: 136 (Thoracochirus; characters; India). DISTRIBUTION: India, Bhutan. brochus Herman, 2001: 41 (Thoracochirus; replacement name for denticollis Coiffait). DISTRIBUTION: Nepal. Bulletin AMNH OSORIINAE: THORACOCHIRUS 1143
denticollis Coiffait, 1984: 138 (Thoracochirus; [preoccupied]; Type locality: Nepal: environs de Bejepur, près de Pokhara). — Herman, 2001: 41 (Thoracochirus; synonym of brochus). cribrellus Fauvel, 1905, see: piestoides Fauvel, 1902. decanus Biswas and Sengupta, 1982: 138 (Thoracochirus; Type locality: India: Kerala: Cardamon H., Muthapatti Pass, Munnar, 1300 m). DISTRIBUTION: India. denticollis Cameron, 1945: 64 (Thoracochirus; Type locality: Bengal: Kalimpong, Samsingh, altitude 1800 feet). DISTRIBUTION: India. denticollis Coiffait, 1984, see: brochus Herman, 2001. exasperatus Cameron, 1918, see: nanus Heller, 1898. forsteri Bernhauer, 1903, see: variolosus Fauvel, 1895. formosae Cameron, 1940d: 249 (Thoracochirus; Type locality: Formosa: Kankau, Koshun). — Cameron, 1949b: 457 (Thoracochirus; Formosa). DISTRIBUTION: Taiwan. helleri Cameron, 1941b: 430 (Thoracochirus; Type locality: Philippines: Aroroy; Panaon; Subaon; Dapa). DISTRIBUTION: Philippines. intermedius Bernhauer, 1926, see: variolosus Fauvel, 1895. magnus Wendeler, 1928a: 32 (Thoracochirus; Type locality: N.O. Sumatra: Deli). — Scheerpeltz, 1933: 1008 (Thoracochirus; catalog). DISTRIBUTION: Indonesia. minutissimus Bernhauer, 1926d: 255 (Thoracochirus; Type locality: Biliran). — Scheerpeltz, 1933: 1008 (Thoracochirus; catalog). DISTRIBUTION: Philippines. minutus Cameron, 1936a: 28 (Thoracochirus; Type locality: Java: G. Slamat, Batoerraden). DISTRIBUTION: Indonesia. nanus Heller, 1898: 14 (Priochirus; Type locality: Insula Mentawei [prope Sumatram], Si Oban). — Bernhauer, 1903: 156 (Thoracochirus; characters). — Bernhauer and Schubert, 1910: 18 (Thoracochirus; catalog). — Bernhauer, 1926d: 255 (Thoracochirus; characters; Philippines; Borneo). — Cameron, 1928d: 95 (Thoracochirus; Sumatra). — Wendeler, 1928c: 119 (Thoracochirus; Philippines). — Scheerpeltz, 1933: 1008 (Thoracochirus; catalog). — Watanabe and Shibata, 1976: 328 (Thoracochirus; Malaysia). DISTRIBUTION: Philippines, Indonesia, Borneo, Malaysia. exasperatus Cameron, 1918b: 169 (Thoracochirus; Type locality: Borneo, Mt. Matang, W. Sarawak). — Cameron, 1928d: 95 (Thoracochirus; synonym of nanus). — Scheerpeltz, 1933: 1008 (Thoracochirus; synonym of nanus). papuanus Cameron, 1937a: 87 (Thoracochirus; Type locality: New Guinea: Kokoda, alt. 1200 ft.). DISTRIBUTION: New Guinea. piestoides Fauvel, 1902: 19 (Leptochirus; subgenus Priochirus; Type locality: Nord-ouest de Sumatra: Tandjong Morawa Serdang). — Fauvel, 1905e: 77 (Leptochirus; subgenus Thoracochirus; Java). — Bernhauer and Schubert, 1910: 18 (Thoracochirus; catalog). — Cameron, 1928d: 95 (Thoracochirus; Sumatra). — Wendeler, 1928c: 119 (Thoracochirus; Philippines). DISTRIBUTION: Borneo, Indonesia, Philippines. 1144 OSORIINAE: THORACOCHIRUS Bulletin AMNH
cribrellus Fauvel, 1905e: 77 (Leptochirus; subgenus Thoracochirus; variety of piestoides; Type locality: Buitenzorg, Jardin botanique). — Bernhauer and Schubert, 1910: 18 (Thoracochirus; synonym of piestoides). — Cameron, 1933b: 338 (Thoracochirus; variety of piestoides; British North Borneo). — Cameron, 1936a: 28 (Thoracochirus; variety of piestoides; Java). quadridens Wendeler, 1928c: 127 (Thoracochirus; Type locality: Philippinen: Mindanao: Surigao). — Scheerpeltz, 1933: 1008 (Thoracochirus; catalog). DISTRIBUTION: Philippines. raffrayi Fauvel, 1902: 21 (Leptochirus; subgenus Priochirus; Type locality: Penang; Sumatra, region de Deli). — Bernhauer and Schubert, 1910: 18 (Thoracochirus; catalog). — Bernhauer, 1926d: 254 (Thoracochirus; characters; Philippines). — Cameron, 1928d: 95 (Thoracochirus; Sumatra). — Wendeler, 1928c: 119 (Thoracochirus; Philippines). — Scheerpeltz, 1933: 1009 (Thoracochirus; catalog). DISTRIBUTION: Malaysia, Indonesia, Philippines. rugosus Fauvel, 1879: 74 (Leptochirus; Type locality: Nouvelle Guinée, Amberbaki). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 156 (Thoracochirus; characters). — Bernhauer and Schubert, 1910: 18 (Thoracochirus; catalog). — Scheerpeltz, 1933: 1009 (Thoracochirus; catalog). DISTRIBUTION: New Guinea. siamensis Cameron, 1940b: 181 (Thoracochirus; Type locality: Indo-China: Laos, Tintoe). DISTRIBUTION: Laos. simoni Fauvel, 1902: 21 (Leptochirus; subgenus Priochirus; Type locality: Ceylan: Kandy). — Bernhauer and Schubert, 1910: 19 (Thoracochirus; catalog). — Cameron, 1930: 120 (Thoracochirus; characters; Ceylon). — Scheerpeltz, 1933: 1009 (Thoracochirus; catalog). — Biswas and Sengupta, 1982: 135 (Thoracochirus; characters; Sri Lanka). DISTRIBUTION: Sri Lanka. sublaevicollis Bernhauer, 1926, see: variolosus Fauvel, 1895. umbilicatus Cameron, 1918b: 169 (Thoracochirus; Type locality: Borneo: Mt. Matang, W. Sarawak). — Scheerpeltz, 1933: 1009 (Thoracochirus; catalog). DISTRIBUTION: Borneo. variolosus Fauvel, 1895: 183 (Leptochirus; Type locality: Bírmanie, Carin Cheba, 500– 1100 m; Carin Asciuii Cheba, 1200–1300 m; Carin Ghécu, 1300–1400 m). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 156 (Thoracochirus; characters). — Bernhauer and Schubert, 1910: 19 (Thoracochirus; catalog). — Bernhauer, 1922c: 221 (Thoracochirus; Formosa). — Bernhauer, 1926d: 255 (Thoracochirus; characters; Philippines). — Wendeler, 1928c: 119 (Thoracochirus; Philippines). — Cameron, 1930: 119 (Thoracochirus; characters; Burma). — Scheerpeltz, 1933: 1009 (Thoracochirus; catalog). DISTRIBUTION: Myanmar, Indonesia, Borneo, Philippines, Taiwan. forsteri Bernhauer, 1903: 156 (Thoracochirus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 18 (Thoracochirus; catalog). — Bernhauer, 1926d: 255 (Thoracochirus; characters; Sumatra). — Cameron, 1928d: 95 (Thoracochirus; synonym of piestoides). — Scheerpeltz, 1933: 1009 (Thoracochirus; variety of variolosus). Bulletin AMNH OSORIINAE: THORACOCHIRUS 1145
intermedius Bernhauer, 1926d: 255 (Thoracochirus; variety of variolosus; Type locality: Biliran. Luzon, Provinz Laguna, Mount Maquiling; Los Baños. Samar. Borneo). — Wendeler, 1928c: 119 (Thoracochirus; subspecies of variolosus; Philippines). — Scheerpeltz, 1933: 1009 (Thoracochirus; variety of variolosus). sublaevicollis Bernhauer, 1926d: 255 (Thoracochirus; Type locality: Biliran). — Wendeler, 1928c: 119 (Thoracochirus; subspecies of variolosus; Philippines). — Scheerpeltz, 1933: 1009 (Thoracochirus; variety of variolosus). verrucifer Fauvel, 1895: 182 (Leptochirus; Type locality: Tenesserim). — Heller, 1898: 8 (Priochirus; checklist). — Bernhauer, 1903: 156 (Thoracochirus; characters). — Bernhauer and Schubert, 1910: 19 (Thoracochirus; catalog). — Cameron, 1930: 119 (Thoracochirus; characters; Burma). — Scheerpeltz, 1933: 1009 (Thoracochirus; catalog). — Biswas and Sengupta, 1982: 137 (Thoracochirus; characters; India; Burma). DISTRIBUTION: India, Myanmar.
Tribe Osoriini Osoriini Erichson, 1839. — Eichelbaum, 1909: 125 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera). — Blatchley, 1910: 473 (characters; key to Indiana genera). — Bernhauer and Schubert, 1911: 141 (world catalog; Osorii, Parosorii). — Cameron, 1930: 289 (characters; key to genera of British India). — Scheerpeltz, 1931: 369 (notes). — Scheerpeltz, 1933: 1127 (world catalog supplement). — Scheerpeltz, 1940: 26 (key to Palaearctic genera). — Blackwelder, 1942: 79 (characters). — Blackwelder, 1944: 110 (checklist of species from Mexico, Central and South America, and the West Indies). — Coiffait, 1955b: 262 (notes, biogeography, and key to genera with eyeless species of Mediter- ranean region). — Hatch, 1957: 243, 245 (characters). — Arnett, 1963: 242, 263 (key to North American genera; characters; list of North American genera). — Newton, 1985: 203 (notes on austral distribution). — Newton and Thayer, 1992: 59 (tribe of Osoriinae; list of subtribes). — Klimaszewski, Newton, and Thayer, 1996: 150 (list of New Zealand genera; number of New Zealand species). — Newton, Thayer, Ashe, and Chandler, 2000: 378 (characters; notes). Osorini Erichson, 1839b: 30 (genera included: Osorius, Holotrochus). TYPE GENUS: Osorius (see Newton and Thayer, 1992: 59). Osoriina Erichson, 1839. — Jakobson, 1908: 446, 476 (characters; list of species of Russia and western Europe). — Newton and Thayer, 1992: 59 (subtribe of Osoriini). Osorii Erichson, 1839. — Bernhauer and Schubert, 1911: 141 (world catalog; Cylindropsis, Paragonus, Mimogonus, Craspedus, Holotrochus, Oephronistus, Anoncosorius, Osorius). — Scheerpeltz, 1933: 1127 (world catalog supplement). Parosorii Bernhauer and Schubert, 1911: 146 (genera included: Parosorius, Mesotrochus, Teiros; catalog). TYPE GENUS: Parosorius (see Newton and Thayer, 1992: 59). Parosoriina Bernhauer and Schubert, 1911. — Newton and Thayer, 1992: 59 (subtribe of Osoriini). 1146 OSORIINAE: AFROSORIUS Bulletin AMNH
Afrosorius [20 species; Ethiopian Region] Afrosorius Fagel, 1958a: 26 (species included: assiniensis, viettei, curtipennis). TYPE SPECIES: Osorius assiniensis Fauvel, fixed by original designation. — Fagel, 1959: 71 (characters; key to African species). — Fagel, 1960a: 27 (notes). — Fagel, 1969: 72 (notes; key to species). — Drugmand, 1998a: 205 (list of types in l’Institut royal des Sciences naturelles de Belgique). adjacens Cameron, 1948, see: assiniensis Fauvel, 1903. alutipennis Bernhauer, 1939i: 253 (Osorius; Type locality: Kamerun: Mt. Cameroon, Jonga, 5000 ft; Onyanga, 5400 ft.). — Fagel, 1959: 78 (Afrosorius; characters; Cameroon). — Fagel, 1963a: 340 (Afrosorius; Guinea). DISTRIBUTION: Cameroon, Guinea. subtilior Bernhauer, 1942c: 350 (Osorius; Type locality: Mont Etinde: 1500 m. Mont Cameroun: 1800–2000 m, versant Sud Est). — Paulian, 1942: 351 (Osorius; larval characters). — Fagel, 1959: 78 (Afrosorius; synonym of alutipennis). assiniensis Fauvel, 1903e: 238 (Osorius; Type locality: Cameroun; Assinie: Ashantis). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Fagel, 1958a: 27 (Afrosorius; type species of genus). — Fagel, 1959: 75 (Afrosorius; characters; Cameroon; Ivory Coast; Guinea). DISTRIBUTION: Guinea, Ivory Coast, Cameroon. adjacens Cameron, 1948a: 226 (Osorius; Type locality: Ivory Coast: Réserve du Banco). — Fagel, 1959: 75 (Afrosorius; synonym of assiniensis). bequaerti Bernhauer, 1932, see: strigifrons H. Kolbe, 1889. bitalensis Fagel, 1959: 83 (Afrosorius; Type locality: Congo Belge: Kivu: territoire de Kaleha, Bitale, N.O. Kahuzi, 1600 m). — Fagel, 1969: 74 (Afrosorius; characters). DISTRIBUTION: Congo. curtipennis Fagel, 1958a: 30 (Afrosorius; Type locality: São Tomé: Monte Cafe, Esperença, 1300 m). — Fagel, 1959: 105 (Afrosorius; characters). DISTRIBUTION: São Tomé and Príncipe. epuluensis Fagel, 1959: 81 (Afrosorius; Type locality: Congo Belge: Kibali-Ituri: territoire d’Eupulu, Mambasa-Mungbere, 900 m. Also cited from Uganda). — Fagel, 1969: 74 (Afrosorius; characters). DISTRIBUTION: Uganda, Congo. fauveli Fagel, 1959: 86 (Afrosorius; Type locality: Gabon: Ssibange. Also cited from Dahomey, Gold Coast, and Guinea). DISTRIBUTION: Guinea, Ghana, Benin, Gabon. gigantulus Cameron, 1928: 21 (Osorius; Type locality: Belgian Congo: Arebi). — Fagel, 1959: 77 (Afrosorius; characters; Belgian Congo). DISTRIBUTION: Congo. hulstaerti Fagel, 1959: 85 (Afrosorius; Type locality: Congo Belge: Équateur: Bokuma). — Fagel, 1969: 74 (Afrosorius; characters). DISTRIBUTION: Congo. ituriensis Fagel, 1959: 100 (Afrosorius; Type locality: Congo Belge: Ituri: territoire de Bunia, mont Hoyo, 1200 m). — Fagel, 1969: 74 (Afrosorius; characters). DISTRIBUTION: Congo. Bulletin AMNH OSORIINAE: AFROSORIUS 1147 kivuensis Fagel, 1959: 93 (Afrosorius; Type locality: Congo Belge: Kivu: contref. S. Kahuzi, 2200 m. Also cited from Ruanda). DISTRIBUTION: Congo, Rwanda. marshalli Bernhauer, 1936h: 324 (Osorius; Type locality: Kenya Colony: Kaimosi). — Fagel, 1959: 79 (Afrosorius; characters; Kenya; Uganda; Belgian Congo). DISTRIBUTION: Kenya, Uganda, Congo. nitidus Fagel, 1969: 72 (Afrosorius; Type locality: Kivu: Itombwe, Kakosi, 1200–1400 m). DISTRIBUTION: Congo. overlaeti Bernhauer, 1933d: 293 (Osorius; Type locality: Lulua: Sandoa). — Fagel, 1959: 88 (Afrosorius; characters; Belgian Congo). DISTRIBUTION: Congo. pseudotruncorum Fagel, 1959: 92 (Afrosorius; Type locality: Congo Belge: Parc National de la Garamba, Morubia. Also cited from Cameroon). DISTRIBUTION: Congo, Cameroon. regularis Fauvel, 1905d: 194 (Osorius; Type locality: Congo français: Benito; Gabon; Cameroun). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Bernhauer, 1912h: 470 (Osorius; Ruwenzori). — Fagel, 1959: 98 (Afrosorius; characters; French Congo; Spanish Guinea). — Fagel, 1969: 74 (Afrosorius; characters). DISTRIBUTION: Guinea, Cameroon, Gabon, “Ruwenzori”, French Equatorial Africa. scabricollis Bernhauer, 1939i: 254 (Osorius; Type locality: Kamerun: Umg. Kamerunberg: Ekona; Tiko). — Fagel, 1959: 99 (Afrosorius; characters; Cameroon; Guinea; Sierra Leone). — Fagel, 1969: 74 (Afrosorius; characters). DISTRIBUTION: Guinea, Sierra Leone, Cameroon. secretus Cameron, 1950, see: truncorum Bernhauer, 1904. strigifrons H. Kolbe, 1889: 122 (Osorius; Type locality: Kwako bis Kimpoko). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Bernhauer, 1915o: 362 (Osorius; Fernando Poo, 2000 m). — Fagel, 1959: 87 (Afrosorius; characters; Belgian Congo; Cameroon; Uganda). — Fagel, 1959b: 5 (Afrosorius; Zaire). — Scheerpeltz, 1971c: 162 (Osorius; Cameroon). — Scheerpeltz, 1974c: 9 (Osorius; Sudan). DISTRIBUTION: Bioko, Cameroon, Congo, Rwanda, Uganda, Sudan. bequaerti Bernhauer, 1932a: 85 (Osorius; Type locality: Penghe). — Scheerpeltz, 1933: 1132 (Osorius; catalog). — Fagel, 1957a: 32 (Osorius; Ruanda). — Fagel, 1959: 87 (Afrosorius; synonym of strigifrons). — Fagel, 1959b: 5 (Afrosorius; synonym of strigifrons). subtilior Bernhauer, 1942, see: alutipennis Bernhauer, 1939. tenuis Fagel, 1959: 96 (Afrosorius; Type locality: Congo Belge: Katanga: Kundelungu, 1750 m). DISTRIBUTION: Congo. truncorum Bernhauer, 1904: 17 (Osorius; Type locality: Africa orientalis, Zanzibar). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Bernhauer, 1912h: 470 (Osorius; Central Africa). — Bernhauer, 1947: 157 (Osorius; Deutsch-Ostafrika; also Cameroon and Natal). — Cameron, 1950g: 20 (Osorius; Belgian Congo). — Fagel, 1959: 89 (Afrosorius; characters; Zanzibar; Tanganyika; Uganda; South Africa; Southern Rhodesia; Central Africa). — Fagel, 1960a: 27 (Afrosorius; Tanganyika Territory). DISTRIBUTION: Cameroon, Congo, Uganda, Tanzania, Zanzibar, Zimbabwe, South Africa. 1148 OSORIINAE: AFROSORIUS Bulletin AMNH
secretus Cameron, 1950a: 182 (Osorius; Type locality: Uganda: Bwamba Valley). — Fagel, 1959: 89 (Afrosorius; synonym of truncorum). viettei Fagel, 1958a: 27 (Afrosorius; Type locality: São Tomé: au-dessus de Lagôa Amélia, 1500 m). — Fagel, 1959: 102 (Afrosorius; characters). DISTRIBUTION: São Tomé.
Afrotyphlopsis [4 species; Ethiopian Region] Afrotyphlopsis Fagel, 1955: 86 (species included: kivuensis, punctata). TYPE SPECIES: Afro- typhlopsis punctata Fagel, fixed by original designation. — Fagel, 1959: 58 (characters). — Drugmand, 1998a: 206 (list of types in l’Institut royal des Sciences naturelles de Belgique). kivuensis Fagel, 1955: 87 (Afrotyphlopsis; Type locality: Congo Belge: Kivu: territoire de Kabare, Nyakasiba, 1900 m). DISTRIBUTION: Congo. oculata Coiffait, 1967: 272 (Afrotyphlopsis; Type locality: Bélinga). DISTRIBUTION: Gabon. pumila Fagel, 1959: 58 (Afrotyphlopsis; Type locality: Congo Belge: Kivu: territoire de Masisi, lacs Ndalaga et Lukubi, 1780 m). DISTRIBUTION: Congo. punctata Fagel, 1955: 90 (Afrotyphlopsis; Type locality: Congo Belge: Kivu: territoire de Lubero, 2200 m). — Fagel, 1959: 58 (Afrotyphlopsis; characters). DISTRIBUTION: Congo.
Allogonus [1 species; Palaearctic Region] Allogonus Coiffait, 1978b: 137 (species included: ventralis). TYPE SPECIES: Allogonus ventralis Coiffait, fixed by original designation. ventralis Coiffait, 1978b: 138 (Allogonus; Type locality: Balu Jhura, 200 m). DISTRIBUTION: Bhutan.
Allosorius [3 species; Ethiopian, Oriental, and Australian Regions] Allosorius Fagel, 1959: 106 (species included: leonensis, paradoxus). TYPE SPECIES: Osorius leonensis Cameron, fixed by original designation. leonensis Cameron, 1948b: 231 (Osorius; Type locality: Sierra Leone; also listed from Haut- Guinée). — Fagel, 1959: 106 (Allosorius; characters; Sierra Leone; Guinea). — Fagel, 1963a: 340 (Allosorius; Guinea). DISTRIBUTION: Guinea, Sierra Leone. paradoxus Bernhauer, 1927: 50 (Osorius; Type locality: Zululand: Mtunzini). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Fagel, 1959: 109 (Allosorius; characters; South Africa). DISTRIBUTION: South Africa. striola Fauvel, 1878f: 210 (Osorius; Type locality: Iles Arou, Wokan; Nouvelle-Guinée, Amberbaki). — Fauvel, 1879: 77 (Osorius; Dorei; Fly River). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Fagel, 1959: 106 (Allosorius). DISTRIBUTION: New Guinea, Indonesia. Bulletin AMNH OSORIINAE: ANANCOSORIUS 1149
Anancosorius [1 species; Neotropical Region] Anancosorius Bernhauer, 1908d: 292 (species included: klimschi). TYPE SPECIES: Ananco- sorius klimschi Bernhauer, fixed by monotypy. — Bernhauer and Schubert, 1911: 143 (world catalog; cited as Anoncosorius; 1 species). — Notman, 1925: 3 (characters in key). — Blackwelder, 1944: 111 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 50 (type species: klimschi). klimschi Bernhauer, 1908d: 293 (Anancosorius; Type locality: Brasilien: S. Catharina). — Bernhauer and Schubert, 1911: 143 (Anancosorius; catalog). DISTRIBUTION: Brazil.
Andringitrana [1 species; Madagascan Region] Andringitrana Coiffait, 1979: 85 (species included: pauliani). TYPE SPECIES: Andringitrana pauliani Coiffait, fixed by original designation. pauliani Coiffait, 1979: 86 (Andringitrana; Type locality: Madagascar Centre: massif de l’Andringitra, zone orientale, forêt Vakoana). DISTRIBUTION: Madagascar.
Arpagonus [15 species; Palaearctic, Madagascan, Oriental, and Australian Regions] Arpagonus Blackwelder, 1952: 62 (replacement name for Paragonus Fauvel). TYPE SPECIES: Paragonus birmanus Fauvel, fixed by objective synonymy with Paragonus. — Blackwelder, 1952: 62 (type species). — Newton, 1990: 92 (characters in key; one undescribed Australian species). Paragonus Fauvel, 1895: 197 (preoccupied by Gill, 1862, and Guichenot, 1869; species included: birmanus). TYPE SPECIES: Paragonus birmanus Fauvel, fixed by monotypy. — Bernhauer and Schubert, 1911: 141 (world catalog; 3 species). — Cameron, 1921a: 351, 367 (characters; key to Singapore species; catalog). — Cameron, 1925: 21 (catalog of Indian species). — Cameron, 1930: 306 (characters). — Scheerpeltz, 1933: 1129 (world catalog supplement). — Blackwelder, 1952: 290 (synonym of Arpagonus; type species: birmanus). — Fagel, 1955: 9 (characters). — Fagel, 1959: 8 (African species). — Shibata, 1973c: 35 (checklist of species of Taiwan). — Drugmand, 1998a: 220 (list of types in l’Institut royal des Sciences naturelles de Belgique). acuticollis Fauvel, 1905a: 134 (Paragonus; Type locality: Penang. Sumatra: Palembang. Java: Tcibodas). — Fauvel, 1905e: 81 (Paragonus; Java; Sumatra; Penang). — Bernhauer and Schubert, 1911: 141 (Paragonus; catalog). — Cameron, 1928d: 107 (Paragonus; Sumatra). — Scheerpeltz, 1933: 1129 (Paragonus; catalog). DISTRIBUTION: Indonesia, Malaysia. africanus Cameron, 1933, see: Euparagonus. bakeri Cameron, 1941c: 492 (Paragonus; Type locality: Philippines: Mindanao: Surigao). DISTRIBUTION: Philippines. birmanus Fauvel, 1895: 197 (Paragonus; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Carin Asciuii Ghécu, 1400–1500 m). — Bernhauer and Schubert, 1911: 141 (Paragonus; catalog). 1150 OSORIINAE: ARPAGONUS Bulletin AMNH
— Cameron, 1930: 306 (Paragonus; characters; Burma). — Scheerpeltz, 1933: 1129 (Paragonus; catalog). — Biswas, 1986: 121 (Paragonus; India). DISTRIBUTION: Myanmar, India. brevipennis Cameron, 1928d: 108 (Paragonus; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1129 (Paragonus; catalog). DISTRIBUTION: Indonesia. brunneus Coiffait, 1981d: 333 (Paragonus; Type locality: Dhumpus Kara, environs de Pokhara). DISTRIBUTION: Nepal. bryanti Cameron, 1936a: 33 (Paragonus; Type locality: Java: Batoerraden, G. Slamat. Penang). DISTRIBUTION: Indonesia, Malaysia. collarti Fagel, 1955, see: Euparagonus. congoensis Cameron, 1949, see: Euparagonus. franzi Coiffait, 1982: 143 (Paragonus; Type locality: Nepal: Nagarkotpedi, 1550 m). DISTRIBUTION: Nepal. hebridensis Cameron, 1934, see: sauteri Bernhauer, 1922. heterocerus Fauvel, 1905a: 134 (Paragonus; Type locality: Sumatra: Palembang. Java: Buitenzorg). — Fauvel, 1905e: 81 (Paragonus; Java; Sumatra). — Bernhauer and Schubert, 1911: 141 (Paragonus; catalog). — Cameron, 1921a: 367, 400 (Paragonus; characters; Singapore). — Cameron, 1928d: 107 (Paragonus; Sumatra). — Scheerpeltz, 1933: 1129 (Paragonus; catalog). DISTRIBUTION: Indonesia, Singapore. insularis Bernhauer, 1922b: 168 (Paragonus; Type locality: Seychellen: Praslin: Côtes d’Or Estate). — Scheerpeltz, 1933: 1129 (Paragonus; catalog). DISTRIBUTION: Seychelles. nepalicus Coiffait, 1982: 142 (Paragonus; Type locality: Nepal: environs de Bimal Nagar, Gorkha distr., 500 m). DISTRIBUTION: Nepal. papuanus Cameron, 1937a: 90 (Paragonus; Type locality: New Guinea: Mafulu, alt. 4000 ft.). DISTRIBUTION: New Guinea. philippinus Cameron, 1941c: 492 (Paragonus; Type locality: Philippines: Masbate: Aroroy). DISTRIBUTION: Philippines. punctatus Fagel, 1955, see: Euparagonus. sauteri Bernhauer, 1922c: 225 (Paragonus; Type locality: Kankau [Koshun]). — Scheerpeltz, 1933: 1129 (Paragonus; catalog). DISTRIBUTION: Vanuatu, Taiwan. hebridensis Cameron, 1934a: 21 (Paragonus; variety of sauteri; Type locality: New Hebrides: Banks Island: Vanua Lava. Santo; also listed from the Philippines). spectabilis Cameron, 1941c: 493 (Paragonus; Type locality: Philippines: Luzon: Imugan). DISTRIBUTION: Philippines. sumatrensis Cameron, 1928d: 107 (Paragonus; Type locality: Sumatra: Fort de Kock; Java: Soekaboemi). — Scheerpeltz, 1933: 1129 (Paragonus; catalog). DISTRIBUTION: Indonesia. Bulletin AMNH OSORIINAE: ATOPOCNEMIUS 1151
Atopocnemius [4 species; Oriental Region] Atopocnemius Bernhauer, 1914c: 92 (species included: moultoni). TYPE SPECIES: Atopocne- mius moultoni Bernhauer, fixed by monotypy. — Notman, 1925: 2 (characters in key). — Scheerpeltz, 1933: 1137 (world catalog supplement). — Blackwelder, 1952: 69 (type species: moultoni). — Naomi, 1986: 39 (characters; species of Japan). asperiventris Cameron, 1929b: 447 (Atopocnemius; Type locality: Pahang: Raub). — Scheerpeltz, 1933: 1137 (Atopocnemius; catalog). DISTRIBUTION: Malaysia. moultoni Bernhauer, 1914c: 92 (Atopocnemius; Type locality: Sarawak, Kuching). — Bernhauer, 1928: 10 (Atopocnemius; Sumatra). — Scheerpeltz, 1933: 1137 (Atopocnemius; catalog). — Cameron, 1936a: 37 (Atopocnemius; Java; Borneo). — Hammond, 1984: 196 (Atopocnemius; Borneo). DISTRIBUTION: Borneo, Indonesia. nomurai Naomi, 1986: 40 (Atopocnemius; Type locality: Mt. Yuidake, Amami-ôshima Is., Kagoshima Pref.). DISTRIBUTION: Japan. quadrifoveatus Cameron, 1929b: 447 (Atopocnemius; Type locality: Selangor: The Gap). — Scheerpeltz, 1933: 1137 (Atopocnemius; catalog). DISTRIBUTION: Malaysia. Bacillopsis [8 species; Palaearctic Region] Bacillopsis Normand, 1920: 131 (subgenus of Cylindropsis; species included: africana, cribel- lata, balearica). TYPE SPECIES: Cylindropsis africana Peyerimhoff, fixed by subse- quent designation by Blackwelder, 1952: 71. — Scheerpeltz, 1933: 1128 (world catalog supplement). — Normand, 1935: 363 (list of species of Tunisia). — Coiffait, 1955b: 263 (characters in key). — Orousset, 1985c: 173 (characters; notes). — Zanetti, 1995: 4 (list of species of Italy). africana Peyerimhoff, 1905: 187 (Cylindropsis; Type locality: Algérie: Jardins de la villa Kespi, Mustapha-Palais). — Bernhauer and Schubert, 1911: 141 (Cylindropsis; catalog). — Scheerpeltz, 1933: 1128 (Bacillopsis; catalog). — Orousset, 1985c: 175 (Bacillopsis; characters; Algeria). DISTRIBUTION: Algeria. balearica Breit, 1908: 60 (Cylindropsis; Type locality: zwischen Valldemosa und Miramar). — Bernhauer and Schubert, 1911: 141 (Cylindropsis; catalog). — Scheerpeltz, 1933: 1128 (Bacillopsis; catalog). — Español, 1945: 90 (Bacillopsis; characters; notes; Spain). — Coiffait, 1961: 82 (Bacillopsis; characters; Mallorca). — Pons and Palmer, 1996: 161 (Bacillopsis; Balearic endemic; notes; habitat; Mallorca). DISTRIBUTION: Spain. cribellata Normand, 1920: 129 (Cylindropsis; subgenus Bacillopsis; Type locality: Algérie: Tlemcen, province d’Oran). — Scheerpeltz, 1933: 1128 (Bacillopsis; catalog). — Orousset, 1985c: 178 (Bacillopsis; may be synonym of africana). DISTRIBUTION: Algeria. 1152 OSORIINAE: BACILLOPSIS Bulletin AMNH dissimilis Orousset, 1985c: 176 (Bacillopsis; Type locality: Algérie: Zaouia des Mousaïa près d’Alger). DISTRIBUTION: Algeria. marocana Coiffait, 1969c: 163 (Bacillopsis; Type locality: Berkane, Rif oriental, Maroc). — Coiffait, 1973a: 270 (Bacillopsis; Morocco). — Orousset, 1985c: 176 (Bacillopsis; characters; name emended from marocanus to marocana; Morocco). DISTRIBUTION: Morocco. mirei Orousset, 1985c: 177 (Bacillopsis; Type locality: Algérie: gorges de Kédara, au sud d’Alger). DISTRIBUTION: Algeria. normandi Orousset, 1985c: 175 (Bacillopsis; Type locality: Tunisie: Le Kef). DISTRIBUTION: Tunisia. sicula Pace, 1980: 121 (Bacillopsis; Type locality: Sicilia: Monti Iblei, Poggio Terrana, a sud di Caltagirone, 250 m). — Sparacio, 1995: 136 (Bacillopsis; characters; notes; Sicily). — Zanetti, 1995: 4 (Bacillopsis; Italy). DISTRIBUTION: Italy. vignai Pace, 1979, see: Cylindropsis. Baculopsis [1 species; Oriental Region] Baculopsis Cameron, 1928d: 106 (species included: jacobsoni). TYPE SPECIES: Baculopsis jacobsoni Cameron, fixed by monotypy. — Scheerpeltz, 1933: 1127 (world catalog supplement). — Blackwelder, 1952: 71 (type species: jacobsoni). jacobsoni Cameron, 1928d: 107 (Baculopsis; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1127 (Baculopsis; catalog). DISTRIBUTION: Indonesia. Bothrys [1 species; Oriental Region] Bothrys Fauvel, 1895: 185 (species included: personatus). TYPE SPECIES: Bothrys personatus Fauvel, fixed by original designation and monotypy. — Bernhauer and Schubert, 1910: 24 (world catalog; 1 species). — Cameron, 1925: 8 (catalog of Indian species). — Cameron, 1930: 67 (characters). — Scheerpeltz, 1933: 1020 (world catalog supplement). — Blackwelder, 1952: 84 (type species: personatus). — Drugmand, 1998a: 207 (list of types in l’Institut royal des Sciences naturelles de Belgique). minuscula Coiffait, 1982, see: Pseudespeson. personatus Fauvel, 1895: 186 (Bothrys; Type locality: Bírmanie, Bhamo, Katha; Sumatra, tabacs). — Bernhauer and Schubert, 1910: 24 (Bothrys; catalog). — Cameron, 1928d: 98 (Bothrys; Sumatra). — Cameron, 1930: 67 (Bothrys; characters; Burma; Sumatra). — Scheerpeltz, 1933: 1020 (Bothrys; catalog). DISTRIBUTION: Myanmar, Indonesia. Craspedus [1 species; Neotropical Region] Craspedus Bernhauer, 1908d: 296 (species included: iheringi). TYPE SPECIES: Craspedus iheringi Bernhauer, fixed by monotypy. — Bernhauer and Schubert, 1911: 142 (world catalog; 1 species). Bulletin AMNH OSORIINAE: CRASPEDUS 1153
— Notman, 1925: 2 (characters in key). — Blackwelder, 1944: 110 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 110 (type species: iheringi). iheringi Bernhauer, 1908d: 296 (Craspedus; Type locality: Brasilien: S. Paulo, Estaçao Raiz da Serra). — Bernhauer and Schubert, 1911: 142 (Craspedus; catalog). DISTRIBUTION: Brazil. Cylindrops [6 species; Ethiopian Region] Cylindrops Fagel, 1955: 93 (species included: congoensis, kivuensis, collarti, itombwensis, straeleni). TYPE SPECIES: Cylindrops congoensis Fagel, fixed by original designation. — Drugmand, 1998a: 207 (list of types in l’Institut royal des Sciences naturelles de Belgique). adami Paulian, 1962: 116 (Cylindrops; Type locality: République de Congo, Brazzaville, Sous-Préfecture de Kindamba, Matouridi). DISTRIBUTION: Congo Republic. angolensis Cameron, 1950f: 118 (Osorius; Type locality: Angola: Riv. Kasai). — Fagel, 1959: 60 (Cylindrops; characters). DISTRIBUTION: Angola. collarti Fagel, 1955, see: kivuensis Fagel, 1955. congoensis Fagel, 1955: 95 (Cylindrops; Type locality: Congo Belge: Kivu: territoire de Kabare, Nyakasiba, 1800 m). DISTRIBUTION: Congo. itombwensis Fagel, 1955: 101 (Cylindrops; Type locality: Congo Belge: Kivu: territoire de Fizi, Itombwe Nord, 900 m). — Fagel, 1957a: 32 (Cylindrops; Ruanda). DISTRIBUTION: Congo, Rwanda. kivuensis Fagel, 1955: 98 (Cylindrops; Type locality: Congo Belge: Kivu: territoire d’Uvira, rives basses de la Sange, 980 m). DISTRIBUTION: Congo. collarti Fagel, 1955: 100 (Cylindrops; subspecies of kivuensis; Type locality: Congo Belge: Stanleyville). — Fagel, 1959: 61 (Cylindrops; subspecies of kivuensis; characters). straeleni Fagel, 1955: 103 (Cylindrops; Type locality: Parc National de l’Upemba: Kabwe- sur-Muye, affluent Lubanga, 1320 m). DISTRIBUTION: Congo. Cylindropsis [9 species; Palaearctic Region] Cylindropsis Fauvel, 1885: 182 (replacement name for Cylindrogaster Fauvel). TYPE SPECIES: Cylindrogaster corsica Fauvel, fixed by objective synonymy with Cylindrogaster. — Heyden, Reitter, and Weise, 1906: 144 (list of species of Europe and Caucasus). — Bernhauer and Schubert, 1911: 141 (world catalog; 3 species). — Razzauti, 1914: 5 (characters). — Winkler, 1925: 347 (catalog for Palaearctic region). — Notman, 1925: 2 (characters in key). — Porta, 1926: 48 (characters; key to species of Italy). — Portevin, 1929: 408 (characters). — Scheerpeltz, 1933: 1128 (world catalog supplement). — Normand, 1935: 363 (list of species of Tunisia). — Blackwelder, 1952: 114 (type species). — Coiffait, 1955b: 263, 264 (notes; characters). — Zanetti, 1995: 4 (list of species of Italy). 1154 OSORIINAE: CYLINDROPSIS Bulletin AMNH
Cylindrogaster Fauvel, 1872: 213 [= 1873: 55] (species included: corsicus; preoccupied by Stål, 1855, Rondani, 1861, and Lioy, 1864). TYPE SPECIES: Cylindrogaster corsicus Fauvel, fixed by original designation and monotypy. — Fauvel, 1875a: XIII [= 1875b: 215] (catalog). — Duvivier, 1883: 185 (catalog). — Heyden, Reitter, and Weise, 1891: 113 (list of species of Europe and Caucasus). — Razzauti, 1914: 5 (synonym of Cylindropsis). — Blackwelder, 1952: 114 (synonym of Cylindropsis; type species: corsica). africana Peyerimhoff, 1905, see: Bacillopsis. andreinii Bordoni, 1968: 124 (Cylindropsis; Type locality: Moscona, Grosseto). — Zanetti, 1995: 4 (Cylindropsis; Italy). DISTRIBUTION: Italy. bordonii Pace, 1979: 181 (Cylindropsis; subspecies of andreinii; Type locality: Monti dell’Uccellina, presso S. Rabano). — Zanetti, 1995: 4 (Cylindropsis; subspecies of andreinii; Italy). balearica Breit, 1908, see: Bacillopsis. bordei Coiffait, 1955b: 264 (Cylindropsis; Type locality: Pont-des-Veyans, Alpes-Maritimes, Var). — Coiffait, 1965d: 290 (Cylindropsis; characters; Sardinia). DISTRIBUTION: France, Sardinia. bordonii Pace, 1979, see: andreinii Bordoni, 1968. corsica Fauvel, 1872: 213 [= 1873: 55] (Cylindrogaster; Type locality: environs de Bastia; [Note: See lectotype designation by Orousset, 1984c]). — Fauvel, 1873b: 129 [= 1873c: 22] (Cylindrogaster; Sardinia). — Saulcy, 1878: 122 (Cylindrogaster; characters; Corsica). — Sainte-Claire Deville, 1906: 78 (Cylindropsis; Corsica). — Bernhauer and Schubert, 1911: 141 (Cylindropsis; catalog). — Portevin, 1929: 408 (Cylindropsis; characters; France). — Scheerpeltz, 1933: 1128 (Cylindropsis; catalog). — Coiffait, 1965d: 290 (Cylindropsis; characters; Sardinia). — Orousset, 1984c: 210 (Cylindropsis; lectotype designation; notes; characters; Corsica). — Orousset, 1988d: 244 (Cylindropsis; larval characters; Corsica). DISTRIBUTION: Corsica, Sardinia. crassiphalla Orousset, 1984c: 214 (Cylindropsis; Type locality: forêt de Sabineto, nord de Porto, Corse du sud, 625 m). DISTRIBUTION: Corsica. cribellata Normand, 1920, see: Bacillopsis. doderoi Razzauti 1914, see: Leptotyphlopsis. exilis Mulsant and Rey, 1875, see: Leptotyphlus. insularis Orousset, 1984c: 212 (Cylindropsis; Type locality: Saint-Andréa-di-Cotone, Castagniccia, Haute-Corse, 300 m). — Orousset, 1988d: 247 (Cylindropsis; Corsica). DISTRIBUTION: Corsica. peyerimhoffi Normand, 1920, see: Leptotyphlopsis. littoralis Coiffait, 1969c: 167 (Cylindropsis; Type locality: Algoz, Algarve, Portugal). DISTRIBUTION: Portugal. loebli Orousset, 1984c: 214 (Cylindropsis; Type locality: vallée de l’Ostriconi, à l.5 km de la côte, nord-est de l’Ile Rousse, Haute-Corse). DISTRIBUTION: Corsica. lucidus Coiffait, 1958, see: Typhlosorius. sardoa Saulcy, 1878: 123 (Cylindrogaster; Type locality: Bonnari, en Sardaigne; [Note: See lectotype designation by Orousset, 1985c]). Bulletin AMNH OSORIINAE: CYLINDROPSIS 1155
— Bernhauer and Schubert, 1911: 141 (Cylindropsis; synonym of corsica). — Peyerimhoff, 1926: 119 (Cylindrogaster; identical to C. corsica). — Scheerpeltz, 1933: 1128 (Cylindropsis; catalog). — Coiffait, 1965d: 289 (Cylindropsis; valid species; characters; Sardinia). — Orousset, 1985c: 178 (Cylindropsis; lectotype designation; characters; notes; Sardinia). — Zanetti, 1995: 4 (Cylindropsis; Italy). DISTRIBUTION: Sardinia. torressalai Coiffait, 1958, see: Typhlosorius. vignai Pace, 1979e: 104 (Bacillopsis; Type locality: Sardegna presso Sassari sul M.te Tudurighe). — Orousset, 1984c: 209 (Cylindropsis). — Orousset, 1985c: 179 (Cylindropsis). — Zanetti, 1995: 4 (Bacillopsis; [Note: It is unclear whether Zanetti overlooked the change by Orousset or disagreed with it]; Sardinia). DISTRIBUTION: Sardinia. zariquieyi Dodero, 1918, see: Geomitopsis.
Edapholotrochus [4 species; Ethiopian Region] Edapholotrochus Fagel, 1958: 3 (species included: pauliani). TYPE SPECIES: Edapholotrochus pauliani Fagel, fixed by original designation. — Fagel, 1969: 59 (notes; characters; key to species). coriaceus Fagel, 1969: 63 (Edapholotrochus; Type locality: Cape Province: Swellendam district, Grootvadersbos). DISTRIBUTION: South Africa. natalensis Fagel, 1969: 60 (Edapholotrochus; Type locality: Natal: Pietermaritzburg, Town- bush Forest). DISTRIBUTION: South Africa. pauliani Fagel, 1958: 6 (Edapholotrochus; Type locality: Natal: Drakensberg, Cathedral Peak, 1500 m). — Fagel, 1969: 59 (Edapholotrochus; characters). DISTRIBUTION: South Africa. reticuliventris Fagel, 1969: 61 (Edapholotrochus; Type locality: Cape Province: Port St-John distr., Ingogo Forest Reserve). DISTRIBUTION: South Africa.
Euparagonus [10 species; Ethiopian Region] Euparagonus Fagel, 1955: 12 (subgenus of Paragonus; species included: africanus, collarti, punctatus, congoensis). TYPE SPECIES: Paragonus africanus Cameron, fixed by original designation. — Fagel, 1969: 3 (valid genus; notes; characters; key to species). africanus Cameron, 1933: 38 (Paragonus; Type locality: Belgian Congo: Faradje: Male). — Fagel, 1955: 12 (Paragonus; subgenus Euparagonus; characters; Belgian Congo). — Fagel, 1959: 8 (Paragonus; subgenus Euparagonus; notes; Zaire). — Fagel, 1960a: 15 (Paragonus; subgenus Euparagonus; Tanganyika; Angola; Congo). — Fagel, 1969: 4 (Euparagonus; characters). DISTRIBUTION: Tanzania, Congo, Angola. collarti Fagel, 1955: 13 (Paragonus subgenus Euparagonus; Type locality: Congo Belge: Stanleyville). — Fagel, 1959: 8 (Paragonus; subgenus Euparagonus; characters; notes; Zaire). — Fagel, 1969: 3 (Euparagonus; characters). DISTRIBUTION: Congo. 1156 OSORIINAE: EUPARAGONUS Bulletin AMNH congoensis Cameron, 1949a: 313 (Paragonus; Type locality: Guinée: Mt. Nimba N. E., alt. 500–700 m). — Fagel, 1955: 15 (Paragonus subgenus Euparagonus; characters; Guinea). — Fagel, 1959: 9 (Paragonus; subgenus Euparagonus; characters; notes; Zaire). — Fagel, 1969: 4 (Euparagonus; characters). DISTRIBUTION: Guinea, Congo. kwangensis Fagel, 1969: 8 (Euparagonus; Type locality: Kwango: territoire de Feshi, Kabanga, riv. Lutshina). DISTRIBUTION: Congo. machadoianus Fagel, 1969: 6 (Euparagonus; Type locality: Angola: Alto Cuilo, Na-Ipanha). DISTRIBUTION: Angola. minutus Fagel, 1969: 5 (Euparagonus; Type locality: Angola: Alto Cuilo, Na-Ipanha). DISTRIBUTION: Angola. nitidus Fagel, 1969: 7 (Euparagonus; Type locality: Kwango: territoire de Feshi, rive dr. Kwenge). DISTRIBUTION: Congo. punctatus Fagel, 1955: 13 (Paragonus subgenus Euparagonus; Type locality: Congo Belge: Ituri: territoire de Bunia, mont Hoyo, 1200 m). — Fagel, 1960a: 16 (Paragonus; subgenus Euparagonus; Uganda). — Fagel, 1969: 4 (Euparagonus; characters). DISTRIBUTION: Congo, Uganda. semirufus Fagel, 1969: 4 (Euparagonus; Type locality: Congo: Equateur: territoire de Bikoro, Mabali). DISTRIBUTION: Congo. transvaalensis Fagel, 1969: 9 (Euparagonus; Type locality: Transvaal: Pietersburg distr. dans les berges de l’Helpmekaar Rivier). DISTRIBUTION: South Africa.
Fagelia [10 species; Madagascan Region] Fagelia Coiffait, 1979: 72 (species included: andohariana, descarpentriesi, marositryana, soror, microcephala, latipes, cylindra, vicina, minuscula). TYPE SPECIES: Fagelia andohariana Coiffait, fixed by original designation. andohariana Coiffait, 1979: 77 (Fagelia; Type locality: Madagascar Centre: massif de l’Andringitra, plateau Andohariana, 2000–2100 m). DISTRIBUTION: Madagascar. cylindrica Coiffait, 1979: 81 (Fagelia; Type locality: Madagascar Centre: massif de l’Andringitra, zone sommitale, cirque Boby, 2500 m). DISTRIBUTION: Madagascar. descarpentriesi Coiffait, 1979: 75 (Fagelia; Type locality: Madagascar Centre: massif de l’Andringitra, plateau Andohariana, 2000–2100 m). DISTRIBUTION: Madagascar. grandis Dajoz, 1980: 126 (Fagelia; Type locality: Madagascar Est: station 2089, chaînes Anosyennes, massif de l’Andohahelo, massif Sud, 1850 m). DISTRIBUTION: Madagascar. latipes Coiffait, 1979: 80 (Fagelia; Type locality: Madagascar Centre: massif de l’Andringitra, zone sommitale, cirque Boby, 2500 m). DISTRIBUTION: Madagascar. marositryana Coiffait, 1979: 78 (Fagelia; Type locality: Madagascar Centre: massif de l’Andringitra, Est de la rivière Antsifotra, Marositry, 2000 m). DISTRIBUTION: Madagascar. Bulletin AMNH OSORIINAE: FAGELIA 1157 microcephala Coiffait, 1979: 80 (Fagelia; Type locality: Madagascar Centre: massif de l’Andringitra, chaîne de l’Andrianony, cirque de Manjarivolo, 1650 m). DISTRIBUTION: Madagascar. minuscula Coiffait, 1979: 83 (Fagelia; Type locality: Madagascar Centre: massif de l’Andringitra, chaîne de l’Andrianony, cirque de Manjarivolo, 1650 m). DISTRIBUTION: Madagascar. soror Coiffait, 1979: 78 (Fagelia; Type locality: Madagascar Centre: massif de l’Andringitra, forêt d’Anjavidilava, 2000 m). DISTRIBUTION: Madagascar. vicina Coiffait, 1979: 82 (Fagelia; Type locality: Madagascar Centre: massif de l’Andringitra, forêt d’Anjavidilava, 2000 m). DISTRIBUTION: Madagascar.
Gigarthrus [11 species; Ethiopian and Oriental Regions] Gigarthrus Bernhauer, 1915m: 298 (subgenus of Mimogonus; species included: bequaerti). TYPE SPECIES: Mimogonus (Gigarthus) bequaerti Bernhauer, fixed by original designation (by “nov. subg. . . . nov. spec.”) and monotypy. — Scheerpeltz, 1933: 1129 (world catalog supplement; subgenus of Mimogonus). — Blackwelder, 1952: 170 (subgenus of Mimogonus; type species bequaerti). — Fagel, 1955: 44 (characters; valid genus; type species: Gigarthrus bequaerti). — Fagel, 1959: 23 (African species). — Fagel, 1960a: 18 (characters). — Fagel, 1969: 31 (notes; key to African species). — Drugmand, 1998a: 210 (list of types in l’Institut royal des Sciences naturelles de Belgique). basilewskyi Fagel, 1960a: 19 (Gigarthrus; Type locality: Tanganyika Territory: Uluguru Mts, Bunduki, gorge de la Mungula, 1500 m). — Fagel, 1969: 31 (Gigarthrus; characters). DISTRIBUTION: Tanzania. bequaerti Bernhauer, 1915m: 298 (Mimogonus; subgenus Gigarthrus; Type locality: Katanga: Welgelegen). — Scheerpeltz, 1933: 1129 (Mimogonus; subgenus Gigarthrus; catalog). — Fagel, 1955: 53 (Gigarthrus; characters). — Fagel, 1959: 23 (Gigarthrus; characters; Mashonaland). — Fagel, 1969: 32 (Gigarthrus; characters). DISTRIBUTION: Congo, Zimbabwe. densipennis Bernhauer, 1927: 49 (Gigarthrus; Type locality: Mashonaland: Salisbury). — Scheerpeltz, 1933: 1130 (Mimogonus; subgenus Gigarthrus; catalog). — Fagel, 1959: 23 (Gigarthrus; synonym of bequaerti). densipennis Bernhauer, 1927, see: bequaerti Bernhauer, 1915. harrismithi Bernhauer, 1934, see: Mimogonellus. katangensis Fagel, 1955: 53 (Gigarthrus; Type locality: Congo Belge: Parc National de l’Upemba: Mukana, 1810 m). — Fagel, 1969: 32 (Gigarthrus; characters). DISTRIBUTION: Congo. leleupi Fagel, 1960a: 18 (Gigarthrus; Type locality: Uganda: Bugiri, 1400 m). — Fagel, 1969: 32 (Gigarthrus; characters). DISTRIBUTION: Uganda. mateui Fagel, 1955: 50 (Gigarthrus; Type locality: Congo Belge: Haut-Uele, Dili. Also cited from Spanish Guinea). — Fagel, 1969: 32 (Gigarthrus; characters). DISTRIBUTION: Congo, Equatorial Guinea. 1158 OSORIINAE: GIGARTHRUS Bulletin AMNH rhodesianus Fagel, 1969: 34 (Gigarthrus; Type locality: N. Rhodesia: Zambia, Abercorn, 1800 m, galerie forestière de la Mwengo). DISTRIBUTION: Zambia. saegeri Fagel, 1959: 25 (Gigarthrus; Type locality: Congo Belge: Parc National de la Garamba). — Fagel, 1969: 32 (Gigarthrus; characters). DISTRIBUTION: Congo. sjoestedti Bernhauer, 1928: 8 (Gigarthrus; Type locality: Sumatra: Brastagi). — Scheerpeltz, 1933: 1130 (Mimogonus; subgenus Gigarthrus; catalog). — Herman, 2001: 12 (Gigarthrus; spelling of name is correct). DISTRIBUTION: Indonesia. tshuapaensis Fagel, 1969: 32 (Gigarthrus; Type locality: Tshuapa: territoire de Bikoro, Mabali). DISTRIBUTION: Congo. turneri Bernhauer, 1927: 50 (Gigarthrus; Type locality: Pondoland: Port St. John). — Scheerpeltz, 1933: 1130 (Mimogonus; subgenus Gigarthrus; catalog). — Fagel, 1959: 24 (Gigarthrus; characters; Pondoland). — Fagel, 1969: 32 (Gigarthrus; characters). DISTRIBUTION: South Africa. wittei Fagel, 1955: 47 (Gigarthrus; Type locality: Congo Belge: Parc National de l’Upemba: Mukana, Lusinga, 1810 m). — Fagel, 1969: 32 (Gigarthrus; characters). DISTRIBUTION: Congo.
Gnatholotrochus [1 species; Ethiopian Region] Gnatholotrochus Fagel, 1955: 71 (species included: striolipennis). TYPE SPECIES: Gnatholotrochus striolipennis Fagel, fixed by original designation and monotypy. — Drugmand, 1998a: 210 (list of types in l’Institut royal des Sciences naturelles de Belgique). striolipennis Fagel, 1955: 72 (Gnatholotrochus; Type locality: Congo Belge: Kivu: territoire de Kalehe, Bitale, Nord-Ouest Kahuzi, 1600 m). DISTRIBUTION: Congo.
Heterocylindropsis [4 species; Ethiopian Region] Heterocylindropsis Fagel, 1955: 104 (species included: kahuziensis, luberensis). TYPE SPECIES: Heterocylindropsis kahuziensis Fagel, fixed by original designation. — Fagel, 1969: 68 (characters; key to species). — Drugmand, 1998a: 210 (list of types in l’Institut royal des Sciences naturelles de Belgique). kahuziensis Fagel, 1955: 106 (Heterocylindropsis; Type locality: Congo Belge: Kivu: territoire de Kabare, contreforts Sud-Est Kahuzi, 2080 m). — Fagel, 1969: 68 (Heterocylindropsis; characters). DISTRIBUTION: Congo. luberensis Fagel, 1955: 108 (Heterocylindropsis; Type locality: Congo Belge: Kivu: territoire de Lubero: Kyalamayhindi, vallée Lopo, 1400 m). — Fagel, 1969: 68 (Heterocylindropsis; characters). DISTRIBUTION: Congo. mwengensis Fagel, 1969: 68 (Heterocylindropsis; Type locality: Kivu: Mwenga, 1250 m). DISTRIBUTION: Congo. werneri Coiffait, 1977e: 662 (Heterocylindropsis; Type locality: Rwanda: Kayove, 2100 m). DISTRIBUTION: Rwanda. Bulletin AMNH OSORIINAE: HETEROSORIUS 1159
Heterosorius [2 species; Madagascan Region] Heterosorius Coiffait, 1979: 128 (species included: vadoni, clementi). TYPE SPECIES: Hetero- sorius vadoni Coiffait, fixed by original designation. clementi Coiffait, 1979: 130 (Heterosorius; Type locality: Madagascar Centre: réserve forestière à 50 km Nord-Ouest de Tananarive). DISTRIBUTION: Madagascar. vadoni Coiffait, 1979: 129 (Heterosorius; Type locality: Madagascar Est: île Sainet Marie, forêt de Kalalao). DISTRIBUTION: Madagascar. Holotrochodes [1 species; Madagascan Region] Holotrochodes Coiffait, 1979: 70 (species included: collaris). TYPE SPECIES: Holotrochodes collaris Coiffait, fixed by original designation and monotypy. collaris Coiffait, 1979: 70 (Holotrochodes; Type locality: Madagascar Est: 6 km au Nord- Ouest de Fanovana, Italaviana, 730 m). DISTRIBUTION: Madagascar. Holotrochomorphus [24 species; Ethiopian and Madagascan Regions] Holotrochomorphus Fagel, 1955: 63 (species included: straeleni, proximus, lusingensis, ger- manus, machadoi). TYPE SPECIES: Holotrochomorphus straeleni Fagel, fixed by original designation. — Fagel, 1959: 44 (notes; key to African species). — Fagel, 1969: 36 (notes; key to African species). — Coiffait, 1979: 66 (characters; key to species of Madagascar). — Drugmand, 1998a: 211 (list of types in l’Institut royal des Sciences naturelles de Belgique). angolanus Fagel, 1969: 53 (Holotrochomorphus; Type locality: Angola: riv. Camipopo, 120 km S.S.W. de Dundo). DISTRIBUTION: Angola. brunneicolor Fagel, 1969: 43 (Holotrochomorphus; Type locality: Kwango: territoire de Feshi, rive dr. Kwange). DISTRIBUTION: Congo. brunnescens Fagel, 1969: 49 (Holotrochomorphus; Type locality: Kwango: territoire de Feshi, rive g. Kwenge). DISTRIBUTION: Congo. coriaceus Fagel, 1969: 45 (Holotrochomorphus; Type locality: Angola: 120 km S.S.W. de Dundo). DISTRIBUTION: Angola. crassus Fagel, 1969: 51 (Holotrochomorphus; Type locality: Tshuapa: territoire d’Ikela, riv. Lusosa. Also cited from Angola). DISTRIBUTION: Angola, Congo. curtipennis Fagel, 1959: 48 (Holotrochomorphus; Type locality: Congo Belge: Kivu: territoire de Lubero, réserve forestière de Biena, 1800 m). — Fagel, 1969: 36 (Holotrochomorphus; characters). DISTRIBUTION: Congo. dubius Fagel, 1959: 49 (Holotrochomorphus; Type locality: Congo Belge: Stanleyville). — Fagel, 1969: 37 (Holotrochomorphus; characters). DISTRIBUTION: Congo. fortepunctatus Fagel, 1969: 46 (Holotrochomorphus; Type locality: Angola: 25 km au N. de Cazombo). DISTRIBUTION: Angola. 1160 OSORIINAE: HOLOTROCHOMORPHUS Bulletin AMNH germanus Fagel, 1955: 68 (Holotrochomorphus; Type locality: Congo Belge: Kivu: territoire de Kabare, Nyakasiba, 1900 m). — Fagel, 1969: 37 (Holotrochomorphus; characters). DISTRIBUTION: Congo. griveaudi Coiffait, 1979: 69 (Holotrochomorphus; Type locality: archipel des Comores, Mohéli, Bandamale, 420 m). DISTRIBUTION: Comoros. ikelaensis Fagel, 1969: 39 (Holotrochomorphus; Type locality: Tshuapa: territoire d’Ikela, ruiss. Befali). DISTRIBUTION: Congo. lusingensis Fagel, 1955: 67 (Holotrochomorphus; Type locality: Congo Belge: Parc National de l’Upemba: Lusinga, 1760 m). — Fagel, 1969: 37 (Holotrochomorphus; characters). DISTRIBUTION: Congo. machadoi Cameron, 1950f: 116 (Holotrochus; Type locality: Angola: Dundo: Luachimo River). — Fagel, 1955: 70 (Holotrochomorphus; characters). — Fagel, 1959: 51 (Holotrochomorphus; characters; Angola). DISTRIBUTION: Angola. madecassus Coiffait, 1979: 68 (Holotrochomorphus; Type locality: Madagascar Centre: massif de l’Ankaratra, Manjakatompo). DISTRIBUTION: Madagascar. nigrescens Fagel, 1969: 41 (Holotrochomorphus; Type locality: Kwango: territoire de Feshi, rive droite de la Kwenge). DISTRIBUTION: Congo. opacus Bernhauer, 1934h: 497 (Holotrochus; Type locality: Mashonaland: Salisbury). — Fagel, 1959: 52 (Holotrochomorphus; characters; Mashonaland). — Fagel, 1969: 37 (Holotrochomorphus; characters). DISTRIBUTION: Zimbabwe. proximus Fagel, 1955: 67 (Holotrochomorphus; Type locality: Congo Belge: Parc National de l’Upemba: Kabwe-sur-Muye, affl. Lubanga, 1320 m). — Fagel, 1969: 37 (Holotrochomorphus; characters). DISTRIBUTION: Congo. rhodesianus Fagel, 1969: 42 (Holotrochomorphus; Type locality: N. Rhodesia: Zambia, Abercorn, galerie forestière de la Mwengo, 1800 m). DISTRIBUTION: Zambia. saegeri Fagel, 1959: 47 (Holotrochomorphus; Type locality: Congo Belge: Parc National de la Garamba). — Fagel, 1969: 36 (Holotrochomorphus; characters). DISTRIBUTION: Congo. serrulatus Fagel, 1969: 52 (Holotrochomorphus; Type locality: Côte d’Ivoire: Bingerville). DISTRIBUTION: Ivory Coast. straeleni Fagel, 1955: 65 (Holotrochomorphus; Type locality: Congo Belge: Parc National de l’Upemba: Mukana-Lusinga, 1810 m). — Fagel, 1969: 36 (Holotrochomorphus; characters). DISTRIBUTION: Congo. strictus Fagel, 1969: 48 (Holotrochomorphus; Type locality: Angola: Tcha-Pemba, affl. Luangue, Xa-Cassau). DISTRIBUTION: Angola. tshuapaensis Fagel, 1959: 51 (Holotrochomorphus; Type locality: Congo Belge: Tshuapa: Flandria). — Fagel, 1969: 38 (Holotrochomorphus; characters). DISTRIBUTION: Congo. Bulletin AMNH OSORIINAE: HOLOTROCHOMORPHUS 1161 vadoni Coiffait, 1979: 69 (Holotrochomorphus; Type locality: Madagascar Est: région de Mananara-Nord, Ivontaka). DISTRIBUTION: Madagascar.
Holotrochopsis [3 species; Ethiopian Region] Holotrochopsis Fagel, 1955: 58 (species included: hulstaerti, kivuensis, muelleri). TYPE SPECIES: Holotrochus hulstaerti Bernhauer, fixed by orginal designation. — Drugmand, 1998a: 211 (list of types in l’Institut royal des Sciences naturelles de Belgique). hulstaerti Bernhauer, 1935d: 98 (Holotrochus; Type locality: Equateur: Flandria). — Fagel, 1955: 60 (Holotrochopsis; characters; Belgian Congo). DISTRIBUTION: Congo. kivuensis Fagel, 1955: 60 (Holotrochopsis; Type locality: Congo Belge: Kivu: territoire de Masisi, Walikale, 700 m). DISTRIBUTION: Congo. muelleri Bernhauer, 1927c: 123 (Holotrochus; cited as mülleri; Type locality: Lagos). — Scheerpeltz, 1933: 1130 (Holotrochus; cited as mülleri; catalog). — Fagel, 1955: 63 (Holotrochopsis; cited as mülleri). — Fagel, 1959: 42 (Holotrochopsis; cited as mülleri; characters; Nigeria). DISTRIBUTION: Nigeria.
Holotrochus [153 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions] Holotrochus Erichson, 1839b: 30 (without included species, for first included species see Erichson, 1840). TYPE SPECIES: Holotrochus volvulus Erichson, fixed by subsequent designation by Duponchel, 1841: 57, who cited the generic name as Olotrochus. — Erichson, 1840: 757 (first included species: volvulus, cylindrus, crassicollis; characters). — Lacordaire, 1854: 113 (characters; notes; list of species). — Kraatz, 1857d: 807 (notes). — Gemminger and Harold, 1868: 643 (catalog). — Sharp, 1876: 387 (notes). — Fauvel, 1879: 78 (characters; notes). — Duvivier, 1883: 185 (catalog). — Sharp, 1887: 682 (notes; Mexican and Central American species). — Blatchley, 1910: 474 (characters). — Bernhauer and Schubert, 1911: 142 (world catalog; 37 species). — Leng, 1920: 98 (catalog of North American species). — Cameron, 1921a: 351, 401 (characters; key to Singapore species; catalog). — Notman, 1925: 2 (characters in key). — Cameron, 1925: 21 (catalog of Indian species). — Bruch, 1928a: 424 (catalog; Argentina). — Cameron, 1930: 307 (characters; key to species of British India). — Scheerpeltz, 1933: 1130 (world catalog supplement). — Bernhauer, 1939f: 207 (key to New Zealand species). — Blackwelder, 1943: 169 (characters; type species volvulus; key to species of West Indies). — Blackwelder, 1944: 110 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 188 (type species volvulus). — Fagel, 1955: 56 (characters; discussion; type species volvulus). — Fagel, 1959: 32 (characters; notes; key to African species). — Coiffait, 1979: 26 (characters; key to species of Madagascar). — Irmler, 1982: 379, 380, 384 (species group classification; key to neotropical species). — Hammond, 1984: 196 (checklist of species of Borneo). — Frank, 1986a: 365 (checklist of species of Florida). 1162 OSORIINAE: HOLOTROCHUS Bulletin AMNH
— Irmler, 1987: 81 (key to Neotropical species; species group classification [scattered through article]). — Newton, 1990: 92 (characters in key; one described and one undescribed Australian species). — Lundgren, 1998: 46 (list of species from Florida; records for other States listed for each species). — Drugmand, 1998a: 211 (list of types in l’Institut royal des Sciences naturelles de Belgique). — Newton, Thayer, Ashe, and Chandler, 2000: 378 (3 Nearctic species; notes; characters in key). Neotrochus Blackwelder, 1943: 164 (species included: minor, trinitatis, cylindrus, smithi). TYPE SPECIES: Holotrochus cylindrus Erichson, fixed by original designation. — Blackwelder, 1944: 110 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 261 (type species cylindrus). — Fagel, 1955: 56 (discussion of problems of separation from Holotrochus). — Moore and Legner, 1974c: 117 (characters; notes). — Moore and Legner, 1975: 173 (catalog; North America north of Mexico). — Moore and Legner, 1979: 153 (characters; notes). — Irmler, 1982: 379 (synonym of Holotrochus). — Downie and Arnett, 1996: 428 (characters). Scapulotrochus Coiffait, 1979: 26 (subgenus of Holotrochus; species included: scapularis, aberrans). TYPE SPECIES: Holotrochus (Scapulotrochus) scapularis Coiffait, fixed by original designation. aberrans Coiffait, 1979: 34 (Holotrochus; subgenus Scapulotrochus; Type locality: Mada- gascar Est: région de Moramanga, Fanovana, 730 m). DISTRIBUTION: Madagascar. acromyrmicis Bernhauer, 1921d: 172 (Holotrochus; Type locality: Argentinien: Prov. Buenos Aires). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — Irmler, 1987: 85 (Holotrochus; characters). DISTRIBUTION: Argentina. africanus Cameron, 1935a: 372 (Holotrochus; Type locality: Stanleyville). — Fagel, 1959: 34 (Holotrochus; characters; Belgian Congo). DISTRIBUTION: Congo. amazonicus Irmler, 1982: 395 (Holotrochus; Type locality: Brazil: Manaus, Amazonas). — Irmler, 1987: 83 (Holotrochus; characters). DISTRIBUTION: Brazil. ambrensis Coiffait, 1979: 41 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Nord: montagne d’Ambre). DISTRIBUTION: Madagascar. annandalei Bernhauer, 1911: 56 (Holotrochus; Type locality: Travancore: Kalattupuzha, W. base of W. Ghats). — Cameron, 1930: 307 (Holotrochus; characters; India). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). DISTRIBUTION: India. anophthalmus Fauvel, 1900, see: Paratorchus. antennatus Wendeler, 1955: 195 (Holotrochus; Type locality: Brasilien: Espirito Santo, Sta. Theresa). — Irmler, 1982: 382 (Holotrochus; characters; Brazil). — Irmler, 1987: 84 (Holotrochus; characters). DISTRIBUTION: Brazil. setosus Wendeler, 1955: 189 (Lispinus; [preoccupied]; Type locality: Brasilien: Nova Teutonia). — Irmler, 1987: 106 (Holotrochus; synonym of antennatus). Bulletin AMNH OSORIINAE: HOLOTROCHUS 1163
condei Wendeler, 1955: 195 (Holotrochus; Type locality: Brasilien: Espirito Santo, Sta. Theresa). — Irmler, 1982: 388 (Holotrochus; synonym of antennatus). similis Wendeler, 1955: 196 (Holotrochus; Type locality: Brasilien: Espirito Santo, Sta. Theresa). — Irmler, 1982: 388 (Holotrochus; synonym of antennatus). silvaticus Wendeler, 1955: 197 (Holotrochus; Type locality: Brasilien: São Paulo, Alto da Serra, 700 m, Biol. Station). — Irmler, 1982: 388 (Holotrochus; synonym of antennatus). niger Wendeler, 1955: 197 (Holotrochus; Type locality: Brasilien: São Paulo, Alto da Serra, 700 m, Biol. Station). — Irmler, 1982: 388 (Holotrochus; synonym of antennatus). arizonicus Chapin, 1928: 66 (Holotrochus; Type locality: Santa Rita Mountains, Arizona). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — Moore and Legner, 1975: 173 (Neotrochus; catalog). DISTRIBUTION: USA. arrowi Bernhauer, 1939, see: Paratorchus. australicus Oke, 1933: 101 (Holotrochus; Type locality: Australia: Victoria: Gembrook, Belgrave). — Newton, 1990: 92 (Holotrochus). DISTRIBUTION: Australia. blackwelderi Irmler, 1982: 386 (Holotrochus; Type locality: Trinidad: Basin Hill Forest reserve E. of Flanigan Town). — Blackwelder, 1943: 167 (Neotrochus; [Note: Misidentified as cylindrus according to Irmler, 1982: 386 who cited it as cylindricus]; characters; Puerto Rico; Grenada; Trinidad). — Irmler, 1987: 84 (Holotrochus; characters). DISTRIBUTION: West Indies. bosqi Bernhauer, 1939, see: vianai Bernhauer, 1939. brachypterus Fauvel, 1905a: 136 (Holotrochus; [Note: Cited as a replacement name for minor LeConte; LeConte misidentified minor Fauvel, he did not described it as new]; Type locality: Florida: Enterprise; Cedar Keys. Caroline du Sud). — LeConte, 1877: 216 (Holotrochus; [Note: Misidentification: Bernhauer and Schubert, 1911: 142 cited minor LeConte as a synonym of brachypterus; LeConte did not describe the species as new, he attributed it to Fauvel]; characters; Florida; South Carolina). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Chapin, 1928: 65 (Holotrochus; characters in key; Florida; Louisiana; Virginia). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — Moore and Legner, 1975: 173 (Neotrochus; catalog). DISTRIBUTION: USA. brasiliensis Bernhauer and Schubert, 1911: 142 (Holotrochus; replacement name for brevi- pennis Bernhauer). — Irmler, 1982: 380 (Holotrochus; characters; Brazil). — Irmler, 1987: 82 (Holotrochus; characters). DISTRIBUTION: Brazil. brevipennis Bernhauer, 1905: 15 (Holotrochus; [preoccupied]; Type locality: Brasilia: Santos). — Irmler, 1982: 380 (Holotrochus; synonym of brasilienesis). brevipennis Bernhauer, 1905, see: brasiliensis Bernhauer and Schubert, 1911. brevipennis Broun, 1893, see: Paratorchus. brevipennis Coiffait, 1976, see: tahitiensis Herman, 2001. brouni Bernhauer, 1939, see: Nototorchus. 1164 OSORIINAE: HOLOTROCHUS Bulletin AMNH brunneipennis Coiffait, 1979: 35 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Est: forêt de Moramanga). DISTRIBUTION: Madagascar. bryanti Cameron, 1941a: 205 (Holotrochus; Type locality: W. Sarawak: Mt. Matang). — Watanabe and Shibata, 1976: 330 (Holotrochus; Malaysia). DISTRIBUTION: Malaysia. caecus Broun, 1910, see: Paratorchus. caledoniae Fauvel, 1889b: 245 (Holotrochus; Type locality: Nouvelle-Caledonie). — Fauvel, 1903d: 262 (Holotrochus; New Caledonia). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). DISTRIBUTION: New Caledonia. campbelli Irmler, 1987: 86 (Holotrochus; Type locality: Colombia: Magdalena, Campana 24 km S. St. Marta, 3000 feet). DISTRIBUTION: Colombia. celebensis Cameron, 1942c: 838 (Holotrochus; Type locality: S. Celebes: Lompa-Battau, altitude 8000 feet). DISTRIBUTION: Indonesia. centralensis Irmler, 1987: 102 (Holotrochus; Type locality: Mexico: Chiapas, 8 mi. N. Pueblo Nuevo Solistahuacan, 6000 feet). DISTRIBUTION: Mexico. centralis Sharp, 1887: 684 (Holotrochus; Type locality: Guatemala, San Juan in Vera Paz; Nicaragua, Chontales). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1987: 85 (Holotrochus; characters). DISTRIBUTION: Guatemala, Nicaragua. cerri Irmler, 1987: 101 (Holotrochus; Type locality: Panama: Cerro Campana, 2900 feet). DISTRIBUTION: Panama. chatterjeei Cameron, 1930: 308 (Holotrochus; Type locality: Dehra Dun: Golattappar). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — Abdullah and Qadri, 1970: 116 (Holotrochus; characters; West Pakistan). — Coiffait, 1977: 244 (Holotrochus; cited as chatterjei; Nepal). DISTRIBUTION: Pakistan, India, Nepal. chinensis Bernhauer, 1934: 4 (Holotrochus; Type locality: China. Umgebung Hongkongs). DISTRIBUTION: China. clavicornis Coiffait, 1979: 45 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: environs de Maroantsetra, Ambodivoangy). DISTRIBUTION: Madagascar. clavipes Sharp, 1876, see: Clavilispinus. collarti Fagel, 1959: 35 (Holotrochus; Type locality: Congo Belge: Tshela). DISTRIBUTION: Congo. columbiensis Irmler, 1987: 102 (Holotrochus; Type locality: Colombia: Valle Soladito). DISTRIBUTION: Colombia. comoni Coiffait, 1987: 497 (Holotrochus; replacement name for fauveli Coiffait). DISTRIBUTION: Madagascar. fauveli Coiffait, 1979: 56 (Holotrochus; subgenus Holotrochus; [preoccupied]; Type locality: Madagascar Est: Baie d’Antongil). condei Wendeler, 1955, see: antennatus Wendeler, 1955. conformalis Herman, 2001: 39 (Holotrochus; replacement name for similis Irmler). DISTRIBUTION: West Indies, Panama. Bulletin AMNH OSORIINAE: HOLOTROCHUS 1165
similis Irmler, 1982: 389 (Holotrochus; [preoccupied]; Type locality: Trinidad: mouth of cave, el 2600 feet, Arifro valley). — Blackwelder, 1943: 168 (Neotrochus; [Note: Misidentified as smithi Cameron according to Irmler, 1982: 389]; characters; St. Vincent; Trinidad). — Irmler, 1987: 85, 104 (Holotrochus; characters; Panama). — Herman, 2001: 39 (Holotrochus; synonym of conformalis). convertus Irmler, 1987: 97 (Holotrochus; Type locality: Panama: Canal Zone, Barro Col- orado). DISTRIBUTION: Panama. convexicollis Coiffait, 1979: 53 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Nord: 50 km au Sud-Est de Diégo-Suarez, forêt d’Analamerana, 80 m). DISTRIBUTION: Madagascar. convexus Irmler, 1987: 86 (Holotrochus; Type locality: Puerto Rico: El Yunque Sta., Luquillo Forest). DISTRIBUTION: West Indies. coriaceipennis Cameron, 1928f: 402 (Holotrochus; Type locality: Mt. Poi, 5000 feet). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). DISTRIBUTION: Borneo. coriaceus Fauvel, 1904e: 306 (Holotrochus; Type locality: Madagascar; [Note: See lectotype designation by Coiffait, 1979]. Originally cited from forêt de Moramanga). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Coiffait, 1979: 36 (Holotrochus; subgenus Holotrochus; lectotype designation: Madagascar; characters; Madagascar). DISTRIBUTION: Madagascar. courtoisi Coiffait, 1979: 54 (Holotrochus; subgenus Holotrochus; Type locality: île Maurice, Flic en Flac). DISTRIBUTION: Madagascar. crassicollis Erichson, 1840: 758 (Holotrochus; Type locality: Madagascar; [Note: See lecto- type designation by Orousset, 1984d]). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Coiffait, 1979: 50 (Holotrochus; subgenus Holotrochus; characters; Madagascar). — Orousset, 1984d: 358 (Holotrochus; lectotype designation; characters of male). DISTRIBUTION: Madagascar. curticollis Fauvel, 1898, see: Allotrochus. curtipennis Coiffait, 1979: 53 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Est: réserve naturelle intégrale n 12, Marojejy-Ouest, 1140 m). DISTRIBUTION: Madagascar. cylindricus Coiffait, 1979: 37 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: région d’Andapa, Anjanaharibe, 1600 m). DISTRIBUTION: Madagascar. cylindrus Erichson, 1840: 758 (Holotrochus; Type locality: Puerto Rico). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1982: 383 (Holotrochus; characters; Puerto Rico). — Irmler, 1987: 85 (Holotrochus; characters). DISTRIBUTION: West Indies. durus Sharp, 1876: 388 (Holotrochus; Type locality: Amazons; Ega). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1982: 380 (Holotrochus; characters; Brazil). — Irmler, 1987: 82 (Holotrochus; characters). DISTRIBUTION: Brazil. emarginatus Irmler, 1987: 88 (Holotrochus; Type locality: Panama: Panama Pr.: Cerro Campana). DISTRIBUTION: Panama. 1166 OSORIINAE: HOLOTROCHUS Bulletin AMNH enigmaticus Coiffait, 1979: 37 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar). DISTRIBUTION: Madagascar. etiennei Coiffait, 1979: 49 (Holotrochus; subgenus Holotrochus; Type locality: île de la Réunion, brûlé de Takamaka). DISTRIBUTION: Mascarene Islands. fairmairei Fauvel, 1904e: 309 (Holotrochus; Type locality: Suberbieville). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Coiffait, 1979: 48 (Holotrochus; subgenus Holotrochus; characters; Madagascar). DISTRIBUTION: Madagascar. fauveli Coiffait, 1979, see: comoni Coiffait, 1987. fauveli Sharp, 1876, see: Clavilispinus. ferrugineus Broun, 1893, see: Nototorchus. fortepunctatus Coiffait, 1979: 47 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Centre: Est d’Ambatolampy, Est de Belanitra, Ampolomita, 1400 m). DISTRIBUTION: Madagascar. fossulatus Motschulsky, 1857, see: Neolosus. foveolatus Motschulsky, 1857, see: Neolosus. franckei Wendeler, 1955: 189 (Lispinus; Type locality: Brasilien: Nova Teutonia). — Irmler, 1987: 83, 106 (Holotrochus; characters; Brazil). DISTRIBUTION: Brazil. galapagosus Coiffait, 1981g: 306 (Holotrochus; Type locality: Ile Galapagos: Ile Santa Cruz, bois au-dessus de Santa Rosa). — Hernández and Outerelo, 1994: 25 (Holotrochus; characters). DISTRIBUTION: Galapagos Islands. garambanus Fagel, 1959: 40 (Holotrochus; Type locality: Congo Belge: Parc National de la Garamba). DISTRIBUTION: Congo. geraldi Irmler, 1987: 99 (Holotrochus; Type locality: Jamaica: St. Ann. P., 4 mi. S. Monique). DISTRIBUTION: West Indies. ghanaensis Fagel, 1961: 275 (Holotrochus; Type locality: Ghana: Nr. Kumasi, Bobiri Forest Res.). DISTRIBUTION: Ghana. glaber Scriba, 1855: 301 (Holotrochus; Type locality: Venezuela). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). DISTRIBUTION: Venezuela. glaberrimus Cameron, 1928f: 402 (Holotrochus; Type locality: Mt. Poi, foot). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). DISTRIBUTION: Borneo. glabrinotus Irmler, 1987: 94 (Holotrochus; Type locality: Venezuela: Aragua, Tiara, 50 km SW Caracas, 1500 m). DISTRIBUTION: Venezuela. glabriventris Bernhauer, 1921b: 69 (Holotrochus; Type locality: Bolivien: Yuracaris). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — Irmler, 1982: 381 (Holotrochus; characters; Bolivia). — Irmler, 1987: 83 (Holotrochus; characters). DISTRIBUTION: Bolivia. gomyi Coiffait, 1979: 43 (Holotrochus; subgenus Holotrochus; Type locality: archipel des Comores, Mohéli, Convalescence, Mamoutzou). DISTRIBUTION: Comoros. Bulletin AMNH OSORIINAE: HOLOTROCHUS 1167 gracilis Broun, 1910, see: Paratorchus. grandis Bernhauer, 1928: 8 (Holotrochus; Type locality: Sumatra: Bah Lias, Medan). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). DISTRIBUTION: Indonesia. hanagarthi Irmler, 1982: 387 (Holotrochus; Type locality: Peru: Pucallpa, Loreto). — Irmler, 1987: 84 (Holotrochus; characters). DISTRIBUTION: Peru. helmsi Fauvel, 1900, see: Paratorchus. heterodontus Coiffait, 1979: 52 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Est: Baie d’Antongil). DISTRIBUTION: Madagascar. hulstaerti Bernhauer, 1935, see: Holotrochopsis. hyleae Irmler, 1987: 100 (Holotrochus; Type locality: Colombia: Amaz., 7 km N. Leticia). DISTRIBUTION: Colombia. ingae Irmler, 1982: 394 (Holotrochus; Type locality: Itajuba, Minas Gerais). — Irmler, 1987: 86 (Holotrochus; characters). DISTRIBUTION: Brazil. inpai Irmler, 1982: 392 (Holotrochus; Type locality: Brazil: Manaus, Amazonas). — Irmler, 1987: 85 (Holotrochus; characters). DISTRIBUTION: Brazil. insularis Coiffait, 1979: 51 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Sambirano, Nosy Komba). DISTRIBUTION: Madagascar. irmleri Herman, 2001: 40 (Holotrochus; replacement name for lineatocollis Irmler). DISTRIBUTION: Panama. lineatocollis Irmler, 1987: 103 (Holotrochus; [preoccupied]; Type locality: Canal Zone: Barro Colorado Is.). — Herman, 2001: 40 (Holotrochus; synonym of irmleri). isabelae Hernández and Outerelo, 1994: 22 (Holotrochus; Type locality: Isabela Island: Entrance of Sucre Cave, Sto. Thomás). DISTRIBUTION: Galapagos Islands. italavianus Coiffait, 1979: 59 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: 6 km au Nord-Ouest de Fanovana, Italaviana, 730 m). DISTRIBUTION: Madagascar. jarrigei Coiffait, 1979: 56 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: maroantsetra). DISTRIBUTION: Madagascar. javanus Cameron, 1936a: 34 (Holotrochus; Type locality: Java: G. Goentoer, Preanger). DISTRIBUTION: Indonesia. laevicauda LeConte, 1866: 376 (Lispinus; Type locality: Illinois). — LeConte, 1877: 216 (Holotrochus; characters; Illinois; New Mexico; Louisiana). — Fauvel, 1905a: 136 (Holotrochus; characters; Illinois; New Mexico; Louisiana). — Blatchley, 1910: 474 (Holotrochus; characters; Indiana). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Chapin, 1928: 65 (Holotrochus; characters in key; Michigan; Kentucky). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — Moore and Legner, 1975: 173 (Neotrochus; catalog). — Campbell and Davies, 1991: 92 (Holotrochus; checklist of species; Canada). — Downie and Arnett, 1996: 428 (Neotrochus; characters; USA). DISTRIBUTION: USA. 1168 OSORIINAE: HOLOTROCHUS Bulletin AMNH laevigatus Broun, 1910, see: Paratorchus. laticollis Bernhauer, 1908d: 295 (Holotrochus; Type locality: Brasilien: S. Catharina). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1982: 382 (Holotrochus; characters; Brazil). — Irmler, 1987: 84 (Holotrochus; characters). DISTRIBUTION: Brazil. latinotus Irmler, 1987: 94 (Holotrochus; Type locality: Colombia: Anchicaya, 1000 feet. Also cited from Panama). DISTRIBUTION: Colombia. latus Cameron, 1941, see: philippinus Cameron, 1941. leleupi Fagel, 1959: 36 (Holotrochus; Type locality: Congo Belge: Kivu: territoire de Masisi, lacs Ndalaga et Lukubi, 1780 m). DISTRIBUTION: Congo. leticiae Irmler, 1987: 97 (Holotrochus; Type locality: Colombia: Amaz., 7 km NW Leticia. Also cited from Brazil). DISTRIBUTION: Colombia, Brazil. lineatocollis Cameron, 1936, see: Heterotrochinus. lineatocollis Irmler, 1987, see: irmleri Herman, 2001. lineatus Irmler, 1982: 397 (Holotrochus; Type locality: Peru: Pucallpa, Loreto). — Irmler, 1987: 83 (Holotrochus; characters). DISTRIBUTION: Peru. longipennis Fagel, 1959: 38 (Holotrochus; Type locality: Congo Belge: Parc National de la Garamba). DISTRIBUTION: Congo. lundgreni Irmler, 1987: 82, 106 (Holotrochus; replacement name for opacus Irmler). DISTRIBUTION: Brazil. opacus Irmler, 1982: 384 (Holotrochus; [preoccupied]; Type locality: Brazil: Caixa d’agua do Grajan, Rio de Janeiro). lusitanicus Coiffait, 1978e: 179 (Holotrochus; Type locality: Portugal: Algarve, Barranco do Veilho, 600 m). DISTRIBUTION: Portugal. machadoi Cameron, 1950, see: Holotrochomorphus. maki Coiffait, 1979: 60 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: Sud de Moramanga [route d’Anosibe, km 54], Ampitameloka, 840 m). DISTRIBUTION: Madagascar. marginatus Sharp, 1887: 684 (Holotrochus; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1982: 382 (Holotrochus; characters; Guatemala). — Irmler, 1987: 84, 104 (Holotrochus; characters; Mexico). DISTRIBUTION: Guatemala, Mexico. marginicollis Fauvel, 1904e: 308 (Holotrochus; Type locality: forêt de Moramanga). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Coiffait, 1979: 43 (Holotrochus; subgenus Holotrochus; characters; Madagascar). DISTRIBUTION: Madagascar. mariannae Irmler, 1987: 92 (Holotrochus; Type locality: Panama: Chiriqui, El Mirador, Finca Collins, nr. Boquete, 6000 feet). DISTRIBUTION: Panama. maroantsetrensis Coiffait, 1979: 52 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: environs de Maroantsetra, Ambodivoangy). DISTRIBUTION: Madagascar. Bulletin AMNH OSORIINAE: HOLOTROCHUS 1169 mexicanus Irmler, 1987: 91 (Holotrochus; Type locality: Mexico: Montepio nr. Catemaco). DISTRIBUTION: Mexico. microphthalmus Fauvel, 1900, see: Paratorchus. milleri Irmler, 1982: 391 (Holotrochus; Type locality: Costa Rica: Wafer Bay, Cocos Island). — Irmler, 1987: 85, 104 (Holotrochus; characters; Panama). DISTRIBUTION: Costa Rica. minimus Coiffait, 1979: 61 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Centre: Tananarive, parc de Tsimbazaza, 1200 m). DISTRIBUTION: Madagascar. minor Fauvel, 1863: 437 (Holotrochus; Type locality: Colombie: Caracas; Cuba). — Fauvel, 1891: 91 (Holotrochus; Venezuela). — Fauvel, 1905a: 135 (Holotrochus; characters; Venezuela; Cuba; Grenada; Guyana). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Blackwelder, 1943: 165 (Neotrochus; characters; Cuba; Jamaica; Grenada; Trinidad). — Irmler, 1982: 383 (Holotrochus; characters; Argentina; Brazil; Bolivia). — Irmler, 1987: 85 (Holotrochus; characters). DISTRIBUTION: Argentina, Bolivia, Brazil, Venezuela, Guyana, West Indies. minusculus Fauvel, 1879: 79 (Holotrochus; Type locality: Mysol; Bornéo, Sarawak; Java, Tcibodas). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). DISTRIBUTION: Malaysia, Indonesia. minutus Cameron, 1928d: 108 (Holotrochus; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). DISTRIBUTION: Indonesia. monstrosus Bernhauer, 1939, see: Paratorchus. montanus Broun, 1910, see: Nototorchus. montepius Irmler, 1987: 95 (Holotrochus; Type locality: Mexico: Montepio, nr. Catemaco). DISTRIBUTION: Mexico. muelleri Bernhauer, 1927, see: Holotrochopsis. myllaenoides Cameron, 1950, see: Allotrochus. myops Coiffait, 1979: 60 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: Sud de Moramanga [route d’Anosibe, km 54], Ampitameloka, 840 m). DISTRIBUTION: Madagascar. nani Irmler, 1982: 386 (Holotrochus; Type locality: Brazil: Serra do Navio, Amapá). — Irmler, 1987: 82 (Holotrochus; characters). DISTRIBUTION: Brazil. nanus Cameron, 1940c: 210 (Holosus; Type locality: Ghum District: Mangpu). — Cameron, 1942: 108 (transferred from Holosus to Holotrochus). DISTRIBUTION: India. nationes Irmler, 1987: 95 (Holotrochus; Type locality: Brazil D.F.: Parque National, 1000 m). DISTRIBUTION: Brazil. neotropicus Irmler, 1982: 396 (Holotrochus; Type locality: Peru: Pucallpa, Loreto). — Irmler, 1987: 82 (Holotrochus; characters). DISTRIBUTION: Peru. newtoni Irmler, 1987: 96 (Holotrochus; Type locality: Mexico: Veracruz, Canyon Rio Metlac, near Fortin, 3200 feet. Also cited from Panama). DISTRIBUTION: Mexico, Panama. niger Wendeler, 1955, see: antennatus Wendeler, 1955. nigrita Fauvel, 1904e: 307 (Holotrochus; Type locality: forêt au Nord de Fort Dauphin, Mada- gascar Est; [Note: See lectotype designation by Coiffait, 1979]. Also cited from forêt de Moramanga). 1170 OSORIINAE: HOLOTROCHUS Bulletin AMNH
— Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Coiffait, 1979: 36 (Holotrochus; subgenus Holotrochus; lectotype designation: forêt au Nord de Fort Dauphin, Madagascar Est; characters; Madagascar). DISTRIBUTION: Madagascar. nitescens Fauvel, 1904e: 307 (Holotrochus; Type locality: forêt de Moramanga). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Coiffait, 1979: 40 (Holotrochus; subgenus Holotrochus; author erroneously cited as Fairmaire; characters; Madagascar). DISTRIBUTION: Madagascar. nitidipennis Coiffait, 1979: 50 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Est: environs de Maroantsetra, Ambodivoangy). DISTRIBUTION: Madagascar. nitidorsis Broun, 1910, see: Paratorchus. nitidulus Coiffait, 1979: 44 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: Sud de Moramanga, route d’Anosibe, km 54, Ampitameloka, 840 m). DISTRIBUTION: Madagascar. nitidus Cameron, 1918: 68 (Holotrochus; Type locality: Singapore: Mandai). — Cameron, 1921a: 368, 401 (Holotrochus; characters; Singapore). — Cameron, 1928d: 108 (Holotrochus; Sumatra). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). DISTRIBUTION: Singapore, Indonesia. nosykombanus Coiffait, 1979: 54 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Sambirano, Nosy Komba, 900 m). DISTRIBUTION: Madagascar. obstrusus Irmler, 1987: 103 (Holotrochus; Type locality: Panama: Cerro Campana, 3200 feet). DISTRIBUTION: Panama. ohausi Wendeler, 1955: 194 (Holotrochus; Type locality: Brasilien: São Paulo, Alto da Serra, 700 m, Biol. Station). — Irmler, 1982: 380 (Holotrochus; characters; Brazil). — Irmler, 1987: 82 (Holotrochus; characters). DISTRIBUTION: Brazil. opacus Bernhauer, 1934, see: Holotrochomorphus. opacus Irmler, 1982, see: lundgreni Irmler, 1987. osorioides Wendeler, 1955: 190 (Lispinus; Type locality: Brasilien: Nova Teutonia). — Irmler, 1987: 107 (Holotrochus; Brazil). DISTRIBUTION: Brazil. palaeotropicus Cameron, 1948a: 225 (Holotrochus; Type locality: Ivory Coast: Azaguié). — Fagel, 1959: 41 (Holotrochus; characters; Ivory Coast; Spanish Guinea; Belgian Congo). DISTRIBUTION: Ivory Coast, Equatorial Guinea, Congo. pallidus Coiffait, 1967: 277 (Holotrochus; Type locality: Bélinga). DISTRIBUTION: Gabon. parvulus Chapin, 1928: 66 (Holotrochus; Type locality: Baton Rouge, La.). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — Moore and Legner, 1975: 173 (Neotrochus; catalog). DISTRIBUTION: USA. pecki Irmler, 1989: 100 (Holotrochus; Type locality: Colombia: Amazonas, Leticia). DISTRIBUTION: Colombia. perinetensis Coiffait, 1979: 42 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Est: Périnet). DISTRIBUTION: Madagascar. Bulletin AMNH OSORIINAE: HOLOTROCHUS 1171 peruvianus Bernhauer, 1941c: 281 [= 1951: 271] (Holotrochus; Type locality: Süd-Peru: Sivia, 520 m). DISTRIBUTION: Peru. philippinus Cameron, 1941b: 435 (Holotrochus; Type locality: Philippines: Luzon; Mt. Makiling; Balbalan). DISTRIBUTION: Philippines. latus Cameron, 1941b: 435 (Holotrochus; subspecies of philippinus; Type locality: Philip- pines: Siargao; Dapa). picescens Sharp, 1887: 683 (Holotrochus; Type locality: Guatemela, Rio Naranjo 450 feet). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1982: 381 (Holotrochus; characters; Guatemala). — Irmler, 1987: 83 (Holotrochus; characters). DISTRIBUTION: Guatemala. piliventris Coiffait, 1979: 42 (Holotrochus; subgenus Holotrochus; Type locality: archipel des Comores, Mohéli, Sambia). DISTRIBUTION: Comoros. planicollis Coiffait, 1979: 45 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: région de Moramanga, Fanovana, 730 m). DISTRIBUTION: Madagascar. plaumanni Irmler, 1987: 89 (Holotrochus; Type locality: Brazil: Nova Teutonia, 300–500 m). DISTRIBUTION: Brazil. politus Sharp, 1887: 683 (Holotrochus; Type locality: Mexico). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1982: 383 (Holotrochus; characters; Mexico). — Irmler, 1987: 85, 104 (Holotrochus; characters; Mexico). DISTRIBUTION: Mexico. poundi Blackwelder, 1943: 170 (Holotrochus; Type locality: Trinidad, 5 miles west-southwest of Manzanilla). — Irmler, 1982: 382 (Holotrochus; characters; Trinidad; Peru). — Irmler, 1987: 84 (Holotrochus; characters). DISTRIBUTION: West Indies, Peru. pseudodurus Irmler, 1982: 386 (Holotrochus; Type locality: Brazil: Manaus, Amazonas). — Irmler, 1987: 82 (Holotrochus; characters). DISTRIBUTION: Brazil. pubescens Sharp, 1876: 390 (Holotrochus; Type locality: Tapajos). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1982: 381 (Holotrochus; characters; Brazil; Peru). — Irmler, 1987: 83 (Holotrochus; characters). DISTRIBUTION: Brazil, Peru. pubiventris Fauvel, 1901: 73 (Holotrochus; Type locality: Venezuela; Pernambuco). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). DISTRIBUTION: Venezuela. pumilus Irmler, 1982: 394 (Holotrochus; Type locality: Peru: Pucallpa, Loreto). — Irmler, 1987: 85 (Holotrochus; cited as pumilius; characters). DISTRIBUTION: Peru. punctatus Coiffait, 1979: 58 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: réserve naturelle intégrale n III, Andranomalaza, Antoby). DISTRIBUTION: Madagascar. puncticollis Bernhauer, 1914c: 93 (Holotrochus; Type locality: Sarawak). — Scheerpeltz, 1933: 1131 (Holotrochus; catalog). DISTRIBUTION: Malaysia. 1172 OSORIINAE: HOLOTROCHUS Bulletin AMNH punctipennis Coiffait, 1979: 39 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Est: région de Maroantsetra, Androndrona). DISTRIBUTION: Madagascar. punctulatus Scriba, 1855: 301 (Holotrochus; Type locality: Venezuela). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). DISTRIBUTION: Venezuela. pusillus Coiffait, 1979: 61 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: région de Moramanga, Fanovana, 730 m). DISTRIBUTION: Madagascar. recticollis Cameron, 1936: 5 (Holotrochus; Type locality: Society Islands: Tahiti, right fork of Papenoo Valley). — Coiffait, 1976: 221 (Holotrochus; Tahiti). DISTRIBUTION: Society Islands. regularis Coiffait, 1979: 47 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: Sud de Moramanga, route d’Anosibe, km 54, Ampitameloka, 840 m). DISTRIBUTION: Madagascar. reitteri Bernhauer, 1935: 5 (Holotrochus; Type locality: Tempelberg, 900 m, Ginyün b. Bébé, C. China). DISTRIBUTION: China. reticulipennis Coiffait, 1979: 39 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar: Analanavoro [peut-ètre Analavory au Nord-Ouest d’Anjozorobe?]). DISTRIBUTION: Madagascar. robustus Coiffait, 1979: 35 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: Nord-Ouest de Manantenina, chaînes Anosyennes, massif Nord, moyenne Ranomandry, 1050 m). DISTRIBUTION: Madagascar. rufescens Cameron, 1950, see: Lispinus. rufopygus Sharp, 1887: 683 (Holotrochus; Type locality: Guatemala, Purula 4000 feet). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1982: 381 (Holotrochus; characters; Guatemala). — Irmler, 1987: 83 (Holotrochus; characters). DISTRIBUTION: Guatemala. rufus Cameron, 1950f: 116 (Holotrochus; Type locality: Angola: Dundo: Luachimo River). — Fagel, 1959: 39 (Holotrochus; characters; Angola). DISTRIBUTION: Angola. scapularis Coiffait, 1979: 34 (Holotrochus; subgenus Scapulotrochus; Type locality: Mada- gascar Nord: montagne d’Ambre, les Roussettes, 1100 m). DISTRIBUTION: Madagascar. schubarti Irmler, 1982: 395 (Holotrochus; Type locality: Sumaré, Rio de Janeiro). — Irmler, 1987: 85, 105 (Holotrochus; characters; Colombia; Panama; Venezuela, Mexico). DISTRIBUTION: Brazil, Venezuela, Panama, Mexico. selangorensis Cameron, 1950d: 6 (Holotrochus; Type locality: Selangor: Kuala Lumpur). DISTRIBUTION: Malaysia. septentrionis Coiffait, 1979: 57 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Nord: 50 km au Sud-Est de Diégo-Suarez, forêt d’Analamerana, 80 m). DISTRIBUTION: Madagascar. setigerus Broun, 1912, see: Paratorchus. setosus Wendeler, 1955, see: antennatus Wendeler, 1955. sigridae Irmler, 1982: 396 (Holotrochus; Type locality: Brazil: Dist. Santos, São Paulo). — Irmler, 1987: 82 (Holotrochus; characters). DISTRIBUTION: Brazil. Bulletin AMNH OSORIINAE: HOLOTROCHUS 1173 sikorai Coiffait, 1979: 51 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar). DISTRIBUTION: Madagascar. silvaticus Wendeler, 1955, see: antennatus Wendeler, 1955. similis Wendeler, 1955, see: antennatus Wendeler, 1955. similis Irmler, 1982, see: conformalis Herman, 2001. simplex Sharp, 1887: 683 (Holotrochus; Type locality: Guatemala, Las Mercedes 2500 feet). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Irmler, 1982: 382 (Holotrochus; characters; Guatemala). — Irmler, 1987: 84 (Holotrochus; characters). DISTRIBUTION: Guatemala. smithi Cameron, 1913: 325 (Holotrochus; Type locality: St. Vincent). — Scheerpeltz, 1933: 1131 (Holotrochus; catalog). — Irmler, 1982: 383 (Holotrochus; characters; St. Vincent). — Irmler, 1987: 85 (Holotrochus; characters). DISTRIBUTION: West Indies. sogai Coiffait, 1979: 59 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: massif du Marojejy, réserve naturelle intégrale n XII, Masiaposa). DISTRIBUTION: Madagascar. sparsus Cameron, 1928, see: Allotrochus. specularis Fauvel, 1904e: 308 (Holotrochus; Type locality: Madagascar). — Bernhauer and Schubert, 1911: 143 (Holotrochus; catalog). — Coiffait, 1979: 44 (Holotrochus; subgenus Holotrochus; characters; Madagascar). DISTRIBUTION: Madagascar. strigipennis Bernhauer, 1921b: 68 (Holotrochus; Type locality: Bolivien: Yuracaris). — Scheerpeltz, 1933: 1131 (Holotrochus; catalog). — Irmler, 1982: 380 (Holotrochus; characters; Brazil; Bolivia). — Irmler, 1987: 83 (Holotrochus; characters). DISTRIBUTION: Brazil, Bolivia. subcrassicollis Coiffait, 1979: 40 (Holotrochus; subgenus Holotrochus; Type locality: Mada- gascar Est: Tamatave). DISTRIBUTION: Madagascar. subtilis Sharp, 1876: 391 (Holotrochus; Type locality: Ega). — Bernhauer and Schubert, 1911: 143 (Holotrochus; catalog). — Irmler, 1982: 381 (Holotrochus; characters; Brazil). — Irmler, 1987: 83 (Holotrochus; characters). DISTRIBUTION: Brazil. susannae Irmler, 1987: 89 (Holotrochus; Type locality: Colombia: Mayda, Campana, 25 km S. St. Marta, 3000 feet). DISTRIBUTION: Colombia. swezeyi Bernhauer, 1942a: 42 (Holotrochus; Type locality: Guam: Dededo). DISTRIBUTION: Marianna Islands. syntheticus Sharp, 1876: 389 (Holotrochus; Type locality: Tapajos). — Bernhauer and Schubert, 1911: 143 (Holotrochus; catalog). — Irmler, 1982: 382 (Holotrochus; characters; Brazil; Peru). — Irmler, 1987: 84 (Holotrochus; characters). DISTRIBUTION: Brazil, Peru. tahitiensis Herman, 2001: 40 (Holotrochus; replacement name for brevipennis Coiffait). DISTRIBUTION: Society Islands. brevipennis Coiffait, 1976: 235 (Holotrochus; [preoccupied]; Type locality: Tahiti: mont Aoraï). — Herman, 2001: 40 (Holotrochus; synonym of tahitiensis). 1174 OSORIINAE: HOLOTROCHUS Bulletin AMNH trinitatis Blackwelder, 1943: 166 (Neotrochus; Type locality: Trinidad: Mayaro Bay). — Irmler, 1982: 383 (Holotrochus; characters; Trinidad). — Irmler, 1987: 85, 104 (Holotrochus; characters; Mexico; Panama). DISTRIBUTION: West Indies, Panama, Mexico. uniformis Sharp, 1887: 684 (Holotrochus; Type locality: Panama, Bugaba). — Bernhauer and Schubert, 1911: 143 (Holotrochus; catalog). DISTRIBUTION: Panama. vadoni Coiffait, 1979: 58 (Holotrochus; subgenus Holotrochus; Type locality: Madagascar Est: Andapa). DISTRIBUTION: Madagascar. vagepunctus Fauvel, 1900, see: Paratorchus. vianai Bernhauer, 1939h: 234 (Holotrochus; Type locality: Argentinien: Chaco). — Irmler, 1982: 382 (Holotrochus; characters; Argentina). — Irmler, 1987: 84 (Holotrochus; characters). DISTRIBUTION: Argentina. bosqi Bernhauer, 1939h: 235 (Holotrochus; Type locality: Argentinien: Tucuman: Siambon). — Irmler, 1982: 387 (Holotrochus; synonym of vianai). villiersi Cameron, 1948, see: Lispinus. vinsoni Cameron, 1935: 35 (Holotrochus; Type locality: Mauritius: Les Mares). — Coiffait, 1979: 48 (Holotrochus; subgenus Holotrochus; characters; Mauritius). DISTRIBUTION: Mascarene Islands. volvulus Erichson, 1840: 758 (Holotrochus; Type locality: Puerto Rico). — Bernhauer and Schubert, 1911: 143 (Holotrochus; catalog). — Blackwelder, 1943: 171 (Holotrochus; characters; Puerto Rico). — Irmler, 1982: 382 (Holotrochus; cited as vovulus; characters; Puerto Rico). — Irmler, 1987: 84 (Holotrochus; characters). DISTRIBUTION: West Indies.
Idiocnemius [1 species; Australian Region] Idiocnemius Cameron, 1937a: 92 (species included: cheesmani). TYPE SPECIES: Idiocnemius cheesmani Cameron, fixed by monotypy. — Blackwelder, 1952: 197 (type species cheesmani). cheesmani Cameron, 1937a: 93 (Idiocnemius; Type locality: New Guinea: Kokoda, alt. 1200 ft.). DISTRIBUTION: New Guinea.
Indosorius [19 species; Palaearctic and Oriental Regions] Indosorius Coiffait, 1978b: 142 (species included: stricticollis, kempi, punctipennis, sikkimen- sis, nilgiriensis, peguanus, bhutanicus, gynandromorphus, spiniphallus). TYPE SPECIES: Indosorius bhutanicus Coiffait, fixed by original designation. — Coiffait, 1982: 141 (key to species of Nepal). angulifrons Coiffait, 1984e: 384 (Indosorius; Type locality: Nepal central: bois au-dessous de Fulung). DISTRIBUTION: Nepal. annandalei Bernhauer, 1911, see: nilgiriensis Fauvel, 1903. arganensis Coiffait, 1982: 138 (Indosorius; Type locality: Nepal: Col Argan). DISTRIBUTION: Nepal. armifrons Coiffait, 1982: 136 (Indosorius; Type locality: Nepal: Col Argan). DISTRIBUTION: Nepal. Bulletin AMNH OSORIINAE: INDOSORIUS 1175 begunensis Coiffait, 1984: 139 (Indosorius; Type locality: Bhutan: Beguna). DISTRIBUTION: Bhutan. bhutanicus Coiffait, 1978b: 144 (Indosorius; Type locality: Gogona, 3100 m). DISTRIBUTION: Bhutan. callifrons Bernhauer, 1926: 24 (Osorius; Type locality: Brit.-Indien: Rotung, 1400 ft.). — Cameron, 1930: 293 (Osorius; characters; India). — Scheerpeltz, 1933: 1132 (Osorius; catalog). — Paulian, 1941: 178 (Osorius; larval characters; Assam). — Scheerpeltz, 1976a: 80 (Osorius; Nepal). — Coiffait, 1982: 141 (Indosorius; characters). — Biswas and Biswas, 1985: 144 (Osorius; India). DISTRIBUTION: India, Nepal. garoensis Coiffait, 1984: 140 (Indosorius; Type locality: Meghalaya, Darugiri, Garo Hills, 450 m). DISTRIBUTION: India. gynandromorphus Coiffait, 1978b: 145 (Indosorius; Type locality: 87 km de Phuntsholing vers Thimphu). DISTRIBUTION: Bhutan. kempi Bernhauer, 1926: 24 (Osorius; Type locality: Ost-Indien: Upper Renying, 2150 ft.). — Cameron, 1930: 299 (Osorius; characters; India). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Coiffait, 1978b: 113 (Indosorius; Bhutan). — Biswas and Biswas, 1985: 144 (Osorius; India). DISTRIBUTION: India, Bhutan. parvipennis Bernhauer, 1926: 25 (Osorius; Type locality: Ost-Indien: Kobo, 400 ft.). — Cameron, 1930: 299 (Osorius; variety of kempi). — Scheerpeltz, 1933: 1134 (Osorius; synonym of kempi). mangpuensis Cameron, 1942: 110 (Osorius; Type locality: Darjeeling district: Ghum, Mangpu). — Coiffait, 1984: 118 (Indosorius; India). DISTRIBUTION: India. nilgiriensis Fauvel, 1903: 152 (Osorius; Type locality: Coonoor). — Fauvel, 1904a: 47 (Osorius; Nilghiris). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Cameron, 1930: 303 (Osorius; characters; India; Bengal). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Abdullah and Qadri, 1970: 118 (Osorius; cited as nilgriensis; characters; East Pakistan). — Coiffait, 1978b: 114 (Indosorius; cited as nilgeriensis; Bhutan). DISTRIBUTION: India, Bangladesh, Bhutan. annandalei Bernhauer, 1911: 56 (Osorius; Type locality: W. Bengalen: Paresnath, 4000 ). — Cameron, 1930: 303 (Osorius; synonym of nilgiriensis). — Scheerpeltz, 1933: 1135 (Osorius; synonym of niligiriensis). parvipennis Bernhauer, 1926, see: kempi Bernhauer, 1926. peguanus Bernhauer, 1914c: 87 (Osorius; cited as pequanus [see Herman, 2001]; Type locality: Ostindien: Pegu). — Cameron, 1930: 298 (Osorius; cited as peguanus; characters; Pegu). — Scheerpeltz, 1933: 1135 (Osorius; cited as peguanus; catalog). — Scheerpeltz, 1965: 102 (Osorius; cited as peguanus; Burma). — Coiffait, 1978b: 114 (Indosorius; cited as peguanus; Bhutan). — Biswas and Biswas, 1985: 144 (Osorius; cited as peguanus; India). — Herman, 2001: 11 (Indosorius; spelling of name corrected). DISTRIBUTION: India, Myanmar, Bhutan. 1176 OSORIINAE: INDOSORIUS Bulletin AMNH pullus Coiffait, 1982: 140 (Indosorius; Type locality: Nepal: Hitaura, Therai). — Coiffait, 1984: 118 (Indosorius; Sikkim). DISTRIBUTION: Nepal. punctipennis Fauvel, 1895: 195 (Osorius; Type locality: Bírmanie, Carin Ghécu, 1300– 1400 m; Carin Cheba, 900–1100 m; Carin Asciuii Cheba, 800–1400 m). — Cameron, 1930: 300 (Osorius; characters; Burma). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Coiffait, 1978b: 113 (Indosorius; Bhutan). — Biswas and Biswas, 1985: 144 (Osorius; India). DISTRIBUTION: India, Myanmar, Bhutan. sikkimensis Bernhauer, 1920d: 177 (Osorius; Type locality: Darjeeling, Sikkim). — Cameron, 1930: 294 (Osorius; characters; Sikkim). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Abdullah and Qadri, 1970: 118 (Osorius; characters; East Pakistan). — Coiffait, 1978b: 113 (Indosorius; Bhutan). DISTRIBUTION: India, Bangladesh, Bhutan. stricticollis Fauvel, 1895: 196 (Osorius; Type locality: Bírmanie, Carin Ghécu, 1300–1400 m; Tenasserim, mont Mooleyit, 1800–1900 m). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Cameron, 1930: 295 (Osorius; characters; Burma). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Coiffait, 1978b: 113 (Indosorius; Bhutan). — Coiffait, 1983d: 162 (Indosorius; Nepal). — Coiffait, 1984: 118 (Indosorius; Nepal). DISTRIBUTION: Myanmar, Bhutan, Nepal. spiniphallus Coiffait, 1978b: 146 (Indosorius; Type locality: Kamjee, 850 m). DISTRIBUTION: Bhutan. travei Coiffait, 1982: 138 (Indosorius; Type locality: Nepal: Col Argan). DISTRIBUTION: Nepal. truncatifrons Coiffait, 1982: 139 (Indosorius; Type locality: Nepal: Gorges de la Marsyandi, entre Tarapani et Jagath, 1650 m). DISTRIBUTION: Nepal.
Kistneria [1 species; Ethiopian Region] Kistneria Coiffait, 1967: 273 (species included: grassei). TYPE SPECIES: Kistneria grassei Coiffait, fixed by original designation and monotypy. grassei Coiffait, 1967: 273 (Kistneria; Type locality: Bélinga, piste de M’Vadi vers le km 8). DISTRIBUTION: Gabon.
Leptotyphlopsis [12 species; Palaearctic Region] Leptotyphlopsis Scheerpeltz, 1931: 376 (species included: doderoi, leucadiana, kaligoniana, ionica). TYPE SPECIES: Cylindropsis doderoi Razzauti, fixed by original designation. — Scheerpeltz, 1933: 1128 (world catalog supplement). — Normand, 1935: 363 (synonym of Cylindropsis). — Blackwelder, 1952: 217 (synonym of Cylindropsis; type species: doderoi). — Coiffait, 1955b: 263, 266 (cited as valid genus; characters in key). — Zanetti, 1995: 4 (list of species of Italy). besucheti Coiffait, 1969c: 166 (Leptotyphlopsis, cited as Leptotyphlosis; Type locality: Karatepe, Taurus, Villayet d’Adana, Turquie d’Asie). DISTRIBUTION: Turkey. Bulletin AMNH OSORIINAE: LEPTOTYPHLOPSIS 1177 bidentata Coiffait, 1969c: 166 (Leptotyphlopsis; Type locality: Urabat, Monts Amanus, Villayet d’Iskenderum, Turquie d’Asie). DISTRIBUTION: Turkey. doderoi Razzauti, 1914: 8 (Cylindropsis; Type locality: giardino dell’Istituto zoologico e dell’Orto botanico pisano). — Dodero, 1918: 105 (Cylindropsis; Tunisia; France; Catalonia). — Scheerpeltz, 1931: 381 (Leptotyphlopsis; type species of Leptotyphlopsis; characters; notes). — Scheerpeltz, 1933: 1128 (Leptotyphlopsis; catalog). — Español, 1945: 92 (Cylindropsis; notes; Spain). — Zanetti, 1995: 4 (Leptotyphlopsis; Italy). DISTRIBUTION: Italy, France, Spain, Tunisia. ionica Scheerpeltz, 1931: 386 (Leptotyphlopsis; Type locality: Kaligoni auf Levkas). — Scheerpeltz, 1933: 1128 (Leptotyphlopsis; catalog). DISTRIBUTION: Greece. iranica Coiffait, 1977e: 661 (Leptotyphlopsis; Type locality: Iran: Ilam, s/Saráb Eyván, 33 42 N, 46 25 E). DISTRIBUTION: Iran. kaligoniana Scheerpeltz, 1931: 384 (Leptotyphlopsis; Type locality: Kaligoni auf Levkas). — Scheerpeltz, 1933: 1128 (Leptotyphlopsis; catalog). DISTRIBUTION: Greece. lavagnei Coiffait, 1955b: 266 (Leptotyphlopsis; Type locality: Castelnau Hérault). DISTRIBUTION: France. libanica Coiffait, 1977e: 660 (Leptotyphlopsis; Type locality: Liban: Beit Eddine, 900 m). DISTRIBUTION: Lebanon. leucadiana Scheerpeltz, 1931: 382 (Leptotyphlopsis; Type locality: Kaligoni auf Levkas). — Scheerpeltz, 1933: 1128 (Leptotyphlopsis; catalog). DISTRIBUTION: Greece. lopadusae Bordoni, 1973j: 667 (Leptotyphlopsis; Type locality: Isola di Lampedusa). — Sparacio, 1995: 135 (Leptotyphlopsis; characters; notes; Sicily). — Zanetti, 1995: 4 (Leptotyphlopsis; Italy). DISTRIBUTION: Italy. peyerimhoffi Normand, 1920: 128 (Cylindropsis; Type locality: Tunisie: Le Kef). — Scheerpeltz, 1933: 1128 (Leptotyphlopsis; catalog). — Normand, 1947a: 6 (Leptotyphlopsis; Tunisia). DISTRIBUTION: Tunisia. zariquieyi Orousset, 1985c: 179 (Leptotyphlopsis; Type locality: Espagne: Catalogne, Arenys-de-Mar). DISTRIBUTION: Spain.
Levasseuria [10 species; Madagascan Region] Levasseuria Coiffait, 1979: 88 (species included: amplipennis, robinsoni, crassa, minima, perinetensis, monticola, italavianensis, sogai, moramangana, zahamena). TYPE SPECIES: Levasseuria minima Coiffait, fixed by original designation. amplipennis Coiffait, 1979: 91 (Levasseuria; Type locality: Madagascar Est: région de Sambava, réserve naturelle intégrale n XII, Marojejy-Ouest, 1140 m). DISTRIBUTION: Madagascar. crassa Coiffait, 1979: 92 (Levasseuria; Type locality: Madagascar Centre: Nord d’Ankazobe, lambeaux forestiers du tompoketsa d’Ambohitantely). DISTRIBUTION: Madagascar. 1178 OSORIINAE: LEVASSEURIA Bulletin AMNH italavianensis Coiffait, 1979: 95 (Levasseuria; Type locality: Madagascar Est: 6 km au Nord-Ouest de Fanovana, Italaviana, 730 m). DISTRIBUTION: Madagascar. minima Coiffait, 1979: 93 (Levasseuria; Type locality: Madagascar Centre: massif de l’Ankaratra, Manjakatompo). DISTRIBUTION: Madagascar. monticola Coiffait, 1979: 94 (Levasseuria; Type locality: Madagascar Centre: massif de l’Andringitra, zone sommitale, cirque Boby, 2500 m). DISTRIBUTION: Madagascar. moramangana Coiffait, 1979: 97 (Levasseuria; Type locality: Madagascar Est: région de Moramanga, Fanovana). DISTRIBUTION: Madagascar. perinetensis Coiffait, 1979: 94 (Levasseuria; Type locality: Madagascar Est: Périnet). DISTRIBUTION: Madagascar. robinsoni Coiffait, 1979: 91 (Levasseuria; Type locality: Madagascar Nord: environs de Diégo-Suarez, montagne des Français). DISTRIBUTION: Madagascar. sogai Coiffait, 1979: 96 (Levasseuria; Type locality: Madagascar Est: réserve naturelle intégrale n III, Andranomalaza, Ambatovositra). DISTRIBUTION: Madagascar. zahamena Coiffait, 1979: 98 (Levasseuria; Type locality: Madagascar Est: Est d’Ambaton- drazaka, Mananilaza). DISTRIBUTION: Madagascar. Lusitanopsis [7 species; Palaearctic Region] Lusitanopsis Coiffait, 1961: 79 (species included: douroensis). TYPE SPECIES: Lusitanopsis douroensis Coiffait, fixed by original designation and monotypy. — Coiffait, 1965d: 289 (key to species). algarvensis Coiffait, 1965d: 286 (Lusitanopsis; Type locality: Portugal: Algarve, environs de Loulé). DISTRIBUTION: Portugal. douroensis Coiffait, 1961: 80 (Lusitanopsis; Type locality: Portugal: Santa Marta de Penaguiao, près de Vila Réal, rive Nord du Douro). — Coiffait, 1965d: 289 (Lusitanopsis; characters). DISTRIBUTION: Portugal. herculeana Coiffait, 1965d: 287 (Lusitanopsis; Type locality: Gibraltar, sur le versant ouest du rocher). DISTRIBUTION: Gibraltar. hispanica Outerelo, 1981: 115 (Lusitanopsis; Type locality: Madrid, El Pardo, 700 m). DISTRIBUTION: Spain. monchica Coiffait, 1969c: 169 (Lusitanopsis; Type locality: Porto de Lagos, Algarve, Portu- gal). DISTRIBUTION: Portugal. salamantica Outerelo, 1977a: 26 (Lusitanopsis; Type locality: valle de Las Batuecas, Sala- manca, 590 m). DISTRIBUTION: Spain. tenerifensis Outerelo, 1990: 25 (Lusitanopsis; Type locality: Islas Canarias: Tenerife: los alrededores de Los Realejos, Valle de La Orotava). — Hernández, Outerelo, and Gamarra, 1994: 187 (Lusitanopsis; checklist of species; Canary Islands). DISTRIBUTION: Canary Islands. Bulletin AMNH OSORIINAE: MADECOSORIUS 1179
Madecosorius [19 species; Madagascan Region] Madecosorius Fagel, 1959: 69 (species included: incisicrurus). TYPE SPECIES: Osorius incisi- crurus Latreille, fixed by original designation and monotypy. — Coiffait, 1979: 99 (characters; key to species of Madagascar). alluaudi Coiffait, 1979: 112 (Madecosorius; Type locality: Madagascar Est: chaînes Anosyennes, massif Nord, moyenne Ranomandry). DISTRIBUTION: Madagascar. alutaceus Bernhauer, 1904: 17 (Osorius; Type locality: Madagaskar). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Coiffait, 1979: 123 (Madecosorius; characters; Madagascar). DISTRIBUTION: Madagascar. angulicollis Coiffait, 1979: 110 (Madecosorius; Type locality: Madagascar Est: forêt au Nord d’Anosibe [route d’Anosibe]). DISTRIBUTION: Madagascar. capuroni Coiffait, 1979: 122 (Madecosorius; Type locality: Madagascar Est: Andapa). DISTRIBUTION: Madagascar. catalai Coiffait, 1979: 111 (Madecosorius; Type locality: Madagascar Est: Vohilava, vallée du Faraony, 60 m). DISTRIBUTION: Madagascar. crenifrons Bernhauer, 1917, see: murecarius Bernhauer, 1917. descarpentriesi Coiffait, 1979: 125 (Madecosorius; Type locality: Madagascar Ouest: forêt Mahajeby, près de Morafenobe). DISTRIBUTION: Madagascar. fissifrons Coiffait, 1979: 115 (Madecosorius; Type locality: Madagascar Est: Périnet). DISTRIBUTION: Madagascar. griveaudi Coiffait, 1979: 117 (Madecosorius; Type locality: Madagascar Centre: massif de l’Andringitra, forêt d’Anjavidilava, 2000 m). DISTRIBUTION: Madagascar. impennis Fauvel, 1904e: 306 (Osorius; Type locality: Diego-Suarez). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Coiffait, 1979: 114 (Madecosorius; characters; Madagascar). DISTRIBUTION: Madagascar. incisicrurus Latreille, 1832: 85 (Osorius; Type locality: Not cited). — Klug, 1832: 139 (Osorius; characters; Madagascar). — Laporte, 1835: 131 (Osorius). — Erichson, 1840: 755 (Osorius; characters; Madagascar). — Laporte, 1840: 188 (Osorius; characters; Madagascar). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Coiffait, 1979: 110 (Madecosorius; characters; Madagascar). DISTRIBUTION: Madagascar. jarrigei Coiffait, 1979: 120 (Madecosorius; Type locality: Madagascar Est: environs de Maroantsetra, Ambodivoangy). DISTRIBUTION: Madagascar. murecarius Bernhauer, 1917m: 556 (Osorius; Type locality: Madagascar). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Coiffait, 1979: 122 (Madecosorius; characters; Madagascar). DISTRIBUTION: Madagascar. crenifrons Bernhauer, 1917m: 557 (Osorius; Type locality: Madagascar: Maroantsetra). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Coiffait, 1979: 122 (Madecosorius; synonym of murecarius). 1180 OSORIINAE: MADECOSORIUS Bulletin AMNH miles Coiffait, 1979: 122 (Madecosorius; Type locality: Madagascar Est: sous-préfecture de Brickaville, Ambadikala, rivière Ranomena). DISTRIBUTION: Madagascar. nitidus Coiffait, 1979: 107 (Madecosorius; Type locality: Madagascar Est: région de Maroantsetra). DISTRIBUTION: Madagascar. pauliani Jarrige, 1970: 61 (Madecosorius; Type locality: Madagascar: Mt. Tsaratanana, 1500–1600 m). — Coiffait, 1979: 126 (Madecosorius; characters; Madagascar). DISTRIBUTION: Madagascar. peyrierasi Coiffait, 1979: 109 (Madecosorius; Type locality: Madagascar Est: massif du Marojejy [réserve naturelle intégrale no 12], 600 m). DISTRIBUTION: Madagascar. punctifrons Bernhauer, 1917m: 555 (Osorius; Type locality: Madagascar). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Coiffait, 1979: 115 (Madecosorius; characters; Madagascar). DISTRIBUTION: Madagascar. schusteri Bernhauer, 1917m: 556 (Osorius; Type locality: Madagascar). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Coiffait, 1979: 123 (Madecosorius; characters; Madagascar). DISTRIBUTION: Madagascar. viettei Jarrige, 1970: 62 (Madecosorius; subspecies of pauliani; Type locality: Madagascar: Mt. Tsaratanana, matsabory en dessous de l’Andohanisambirano, 2050 m). — Coiffait, 1979: 126 (Madecosorius; valid species; characters; Madagascar). DISTRIBUTION: Madagascar. Madegassosorius [8 species; Madagascan Region] Madegassosorius Scheerpeltz, 1961g: 246 (species included: keiseri). TYPE SPECIES: Made- gassosorius keiseri Scheerpeltz, fixed by original designation and monotypy. — Coiffait, 1979: 14 (characters; key to species of Madagascar). caecus Coiffait, 1979: 22 (Madegassosorius; Type locality: Madagascar Est: réserve naturelle intégrale n III, Andranomalaza, Mananilaza). DISTRIBUTION: Madagascar. keiseri Scheerpeltz, 1961g: 235, 248 (Madegassosorius; Type locality: Ost-Madagaskar: Périnet). — Coiffait, 1979: 16 (Madegassosorius; characters; Madagascar). DISTRIBUTION: Madagascar. longipennis Coiffait, 1979: 18 (Madegassosorius; Type locality: Madagascar Est, réserve naturelle intégrale n 1, dite de Betampona, 50 km au Nord-Ouest de Tamatave). DISTRIBUTION: Madagascar. pauliani Coiffait, 1979: 20 (Madegassosorius; Type locality: Madagascar Centre: Nord d’Ankazobe, lambeaux forestiers du tampoketsa d’Ambohitantely). DISTRIBUTION: Madagascar. robustus Coiffait, 1979: 19 (Madegassosorius; Type locality: Madagascar Est: Périnet). DISTRIBUTION: Madagascar. scaberrinus Coiffait, 1979: 19 (Madegassosorius; Type locality: Madagascar Centre: Nord d’Ankazobe, lambeaux forestiers du tampoketsa d’Ambohitantely). DISTRIBUTION: Madagascar. sogai Coiffait, 1979: 20 (Madegassosorius; Type locality: Madagascar Est: réserve naturelle intégrale n III, Andranomalaza, vallée d’Ivelona). DISTRIBUTION: Madagascar. Bulletin AMNH OSORIINAE: MADEGASSOSORIUS 1181 sparsus Coiffait, 1979: 22 (Madegassosorius; Type locality: Madagascar Centre: massif de l’Ankaratra, Manjakatompo). DISTRIBUTION: Madagascar. Mimogonellus [22 species; Palaearctic, Ethiopian and Madagascan Regions] Mimogonellus Fagel, 1955: 26 (species included: microphthalmus, collarti, africanus, similis, leleupi, nigerrimus, methneri, curticornis, incertus). TYPE SPECIES: Mimogonus leleupi Cameron, fixed by original designation. — Fagel, 1959: 27 (notes; African species). — Fagel, 1969: 23 (notes; key to African species). — Coiffait, 1979: 11 (characters; key to species of Madagascar). — Naomi, 1996: 53 (characters; Japanese species). — Drugmand, 1998a: 215 (list of types in l’Institut royal des Sciences naturelles de Belgique). africanus Bernhauer, 1915d: 104 (Mimogonus; Type locality: Elfenbeinküste: Dimbroko). — Scheerpeltz, 1933: 1129 (Mimogonus; catalog). — Fagel, 1955: 35 (Mimogonus; characters; Belgian Congo). — Fagel, 1969: 24 (Mimogonellus; characters). DISTRIBUTION: Ivory Coast, Congo. betschi Coiffait, 1979: 13 (Mimogonellus; Type locality: Madagascar Centre: massif de l’Andringitra, forêt d’Anjavidilava, 1850–1950 m). DISTRIBUTION: Madagascar. bikoroensis Fagel, 1969: 28 (Mimogonellus; Type locality: Tshuapa: territoire de Bikoro, Mabali). DISTRIBUTION: Congo. clementi Coiffait, 1979: 14 (Mimogonellus; Type locality: Madagascar Sud, pays Mahafaly, Ouest de Betioky, plaine d’Ankazomanga). DISTRIBUTION: Madagascar. continentalis Berhauer, 1915, see: methneri Bernhauer, 1915. collarti Fagel, 1955: 32 (Mimogonellus; Type locality: Congo Belge: Likimi: Gundji). — Fagel, 1959: 27 (Mimogonellus; characters; Belgian Congo). — Coiffait, 1967: 267 (Mimogonellus; characters). — Fagel, 1969: 24 (Mimogonellus; characters). DISTRIBUTION: Congo. curticornis Fagel, 1955: 42 (Mimogonellus; Type locality: Congo Belge: Katanga: monts Kundelungu, 1750 m). — Fagel, 1969: 25 (Mimogonellus; characters). DISTRIBUTION: Congo. decelleanus Fagel, 1969: 27 (Mimogonellus; Type locality: Congo: Yangambi). DISTRIBUTION: Congo. dimorphus Coiffait, 1967: 269 (Mimogonellus; Type locality: environs de Bélinga). DISTRIBUTION: Gabon. gabonensis Coiffait, 1967: 267 (Mimogonellus; Type locality: Bélinga). DISTRIBUTION: Gabon. gracilis Coiffait, 1967: 270 (Mimogonellus; Type locality: environs de Makokou). DISTRIBUTION: Gabon. harrismithi Bernhauer, 1934, see: methneri Bernhauer, 1915. incertus Fagel, 1955: 44 (Mimogonellus; Type locality: Congo Belge: Parc National de l’Upemba: Lusinga, 1760 m). DISTRIBUTION: Congo. japonicus Naomi, 1987c: 15 (Saegerius; Type locality: Japan: Saga Pref.: Mt. Tara). — Naomi, 1996: 53 (Mimogonellus; Japan). DISTRIBUTION: Japan. 1182 OSORIINAE: MIMOGONELLUS Bulletin AMNH leleupi Cameron, 1952a: 323 (Mimogonus; Type locality: Belgian Congo: Katanga: Grotte de Kakontwé). — Fagel, 1955: 39 (Mimogonellus; characters). — Fagel, 1969: 25 (Mimogonellus; characters). DISTRIBUTION: Congo. longipennis Fagel, 1960a: 16 (Mimogonellus; Type locality: Tanganyika Territory: Uluguru Mts, sommet di Kidunda, 1800–1950 m). — Fagel, 1969: 24 (Mimogonellus; characters). DISTRIBUTION: Tanzania. meridioafricanus Scheerpeltz, 1974: 82 (Mimogonellus; Type locality: Cape Prov.: Tzitzikama Forest, Storms River Mouth). DISTRIBUTION: South Africa. methneri Bernhauer, 1915d: 105 (Mimogonus; Type locality: Deutsch-Ostafrika: Süd- Uluzuru). — Bernhauer, 1927: 50 (Mimogonus; South Africa). — Scheerpeltz, 1933: 1129 (Mimogonus; catalog). — Fagel, 1955: 39 (Mimogonellus; characters; German East Africa; Belgian Congo). — Fagel, 1959: 29 (Mimogonellus; characters; notes; German East Africa). — Fagel, 1969: 25 (Mimogonellus; characters). DISTRIBUTION: Tanzania, Congo, South Africa. continentalis Bernhauer, 1915d: 105 (Mimogonus; Type locality: Deutsch-Ostafrika: Arushu-Ju). — Bernhauer, 1927: 50 (Mimogonus; characters). — Scheerpeltz, 1933: 1129 (Mimogonus; catalog). — Fagel, 1959: 30 (Mimogonellus; subspecies of methneri; characters; German East Africa; Belgian Congo). — Fagel, 1960a: 17 (Mimogonellus; subspecies of methneri; notes; Tanganyika). — Fagel, 1969: 25 (Mimogonellus; subspecies of methneri; characters). harrismithi Bernhauer, 1934h: 496 (Gigarthrus; Type locality: Orange Free State: Harri- smith). — Fagel, 1959: 31 (Mimogonellus; subspecies of methneri; characters; South Africa). — Fagel, 1969: 25 (Mimogonellus; subspecies of methneri; characters). microphthalmus Fagel, 1955: 29 (Mimogonellus; Type locality: Ruanda: forêt du Rugege, 2400 m). — Fagel, 1969: 24 (Mimogonellus; characters). DISTRIBUTION: Rwanda. nigerrimus Cameron, 1950g: 19 (Mimogonus; Type locality: Belgian Congo: Mayumbu, volc. Nyamuragira, alt. 2100 m). — Fagel, 1955: 39 (Mimogonellus; characters; Belgian Congo). — Fagel, 1969: 25 (Mimogonellus; characters). DISTRIBUTION: Congo. nitidipennis Fagel, 1969: 30 (Mimogonellus; Type locality: Côte d’Ivoire: Bingerville). DISTRIBUTION: Ivory Coast. similis Fagel, 1955: 37 (Mimogonellus; Type locality: Congo Belge: Katanga: massif des Kundelungu, 1750 m). — Fagel, 1969: 24 (Mimogonellus; characters). DISTRIBUTION: Congo. sparsepunctatus Fagel, 1969: 25 (Mimogonellus; Type locality: Tshuapa: territoire de Bikoro, Mabali). DISTRIBUTION: Congo. yasutoshii Naomi, 1987c: 17 (Saegerius; Type locality: Japan: Kagoshima Pref.: Bozu). — Naomi, 1996: 53 (Mimogonellus; Japan). DISTRIBUTION: Japan. Bulletin AMNH OSORIINAE: MIMOGONIA 1183
Mimogonia [8 species; Neotropical and Palaearctic Regions] Mimogonia Coiffait, 1978e: 177 (species included: europaea). TYPE SPECIES: Mimogonia europaea Coiffait, fixed by monotypy. — Irmler, 1981: 144 (characters; key to Neotropical species). antennata Irmler, 1981: 150 (Mimogonia; Type locality: Peru: Pucalpa). DISTRIBUTION: Peru. brunnea Irmler, 1981: 146 (Mimogonia; Type locality: Peru: Pucalpa). DISTRIBUTION: Peru. europaea Coiffait, 1978e: 178 (Mimogonia; Type locality: Portugal: Algarve, Santa Barbara de Nexe, 100 m). DISTRIBUTION: Portugal. pumilia Irmler, 1981: 151 (Mimogonia; Type locality: Peru: Pucalpa). DISTRIBUTION: Peru. similis Irmler, 1981: 148 (Mimogonia; Type locality: Brasilien: Manaus, Amazones). DISTRIBUTION: Brazil. subopaca Irmler, 1981: 145 (Mimogonia; Type locality: Peru: Pucalpa). DISTRIBUTION: Brazil. tricolor Irmler, 1981: 149 (Mimogonia; Type locality: Manaus, Amazonas, Rio Taruma Mirîm). DISTRIBUTION: Brazil. unicolor Irmler, 1981: 149 (Mimogonia; Type locality: Brasilien: Manaus, Amazonas). DISTRIBUTION: Brazil.
Mimogonidius [1 species; Ethiopian Region] Mimogonidius Fagel, 1959: 15 (species included: rufus). TYPE SPECIES: Mimogonus rufus Cameron, fixed by original designation and monotypy. rufus Cameron, 1950f: 116 (Mimogonus; Type locality: Angola: Kasai River). — Fagel, 1955: 23 (Mimogonus; characters; Belgian Congo). — Fagel, 1959: 16 (Mimogonidius; characters; Angola; Zaire). DISTRIBUTION: Angola, Congo.
Mimogonoderus [4 species; Ethiopian Region] Mimogonoderus Fagel, 1969: 16 (species included: longipennis, microphthalmus, ineditus, angolensis). TYPE SPECIES: Mimogonoderus longipennis Fagel, fixed by original designation. angolensis Fagel, 1969: 19 (Mimogonoderus; Type locality: Angola: Alto Cuilo, Tcha- Muchito). DISTRIBUTION: Angola. ineditus Fagel, 1969: 21 (Mimogonoderus; Type locality: Kwango: territoire de Feshi, rive dr. de la Kwenge). DISTRIBUTION: Congo. longipennis Fagel, 1969: 18 (Mimogonoderus; Type locality: Tshuapa: territoire d’Ikela, tête de source de ruisseau Kilulu). DISTRIBUTION: Congo. microphthalmus Fagel, 1969: 22 (Mimogonoderus; Type locality: Angola: Tcha-Pemba, affl. Luangue, Xa-Cassau). DISTRIBUTION: Angola. 1184 OSORIINAE: MIMOGONUS Bulletin AMNH
Mimogonus [17 species; Neotropical, Palaearctic, Ethiopian, Madagascan, Oriental, and Oceanic Regions] Mimogonus Fauvel, 1903d: 261 (species included: fumator, rufipennis, microps). TYPE SPECIES: Osorius fumator Fauvel, fixed by subsequent designation by R. Lucas, 1920: 419. — Bernhauer and Schubert, 1911: 141 (world catalog; 3 species). — Cameron, 1921a: 351, 401 (characters; key to Singapore species; catalog). — Winkler, 1925: 347 (catalog for Palaearctic region). — Notman, 1925: 2, 3 (characters in key). — Cameron, 1925: 21 (catalog of Indian species). — Cameron, 1930: 304, 457 (characters; key to species of British India). — Scheerpeltz, 1933: 1129 (world catalog supplement). — Blackwelder, 1943: 172 (characters; type species fumator). — Blackwelder, 1944: 111 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 248 (type species). — Fagel, 1955: 16 (characters; type species Mimogonus fumator). — Fagel, 1959: 9 (characters; notes; African species). — Fagel, 1969: 11 (notes; key to African species). — Shibata, 1973c: 36 (checklist of species of Taiwan). — Tikhomirova, 1973a: 147 (checklist of species of USSR). — Shibata, 1976: 168 (checklist of species of Japan). — Coiffait, 1979: 10 (characters; species of Madagascar). — Hammond, 1984: 197 (checklist of species of Borneo). — Naomi, 1986: 34 (characters). — Naomi, 1987c: 12 (characters; species of Japan). — Drugmand, 1998a: 216 (list of types in l’Institut royal des Sciences naturelles de Belgique). africanus Bernhauer, 1915, see: Mimogonellus. andrewesi Cameron, 1930: 305 (Mimogonus; Type locality: Nilgiri Hills). — Scheerpeltz, 1933: 1129 (Mimogonus; catalog). DISTRIBUTION: India. angulicollis Fagel, 1959: 12 (Mimogonus; subspecies of crebrepunctatus Fagel; Type locality: Congo Belge: Kivu: territoire de Lubero, réserve forestière de Biena, 1800 m). — Fagel, 1969: 12 (Mimogonus; valid species; characters). DISTRIBUTION: Congo. bequaerti Bernhauer, 1915, see: Gigarthrus. bryanti Cameron, 1941a: 203 (Mimogonus; Type locality: W. Sarawak: Mt. Matang). DISTRIBUTION: Malaysia. congoensis Fagel, 1955, see: Saegerius. continentalis Bernhauer, 1915, see: Mimogonellus. crebrepunctatus Fagel, 1959: 10 (Mimogonus; Type locality: Congo Belge: Parc National de la Garamba, Pidigala). — Fagel, 1969: 12 (Mimogonus; characters). DISTRIBUTION: Congo. cribratus Fagel, 1961: 272 (Mimogonus; Type locality: Ghana: Nr. Kumasi, Bobiri Forest Res.). — Fagel, 1969: 11 (Mimogonus; characters). DISTRIBUTION: Ghana. curtipennis Fagel, 1959: 13 (Mimogonus; Type locality: Congo Belge: Kivu: territoire de Masisi, Mutakato, 800 m). — Fagel, 1969: 11 (Mimogonus; characters). DISTRIBUTION: Congo. Bulletin AMNH OSORIINAE: MIMOGONUS 1185 fauveli Cameron, 1913, see: fumator Fauvel, 1889. fumator Fauvel, 1889b: 246 (Osorius; Type locality: Nouvelle-Caledonie: Bourail; Sumatra). — Fauvel, 1903d: 261 (Mimogonus; New Caledonia; Sumatra; Burma; Ceylon; Madagascar; Guadeloupe). — Fauvel, 1904a: 47 (Mimogonus; Nilghiris; Ceylan; Bírmanie; Sumatra; Java; New Caledonia; Madagascar; Guadeloupe). — Bernhauer, 1910b: 359 (Mimogonus; notes; Mexico). — Bernhauer and Schubert, 1911: 141 (Mimogonus; catalog). — Eichelbaum, 1913: 122 (Mimogonus; Tanzania). — Cameron, 1921a: 368, 401 (Mimogonus; characters; Singapore). — Bernhauer, 1922b: 168 (Mimogonus; Seychelles). — Notman, 1925: 3 (Mimogonus; notes; Mexico). — Cameron, 1928d: 108 (Mimogonus; Sumatra). — Cameron, 1930: 304 (Mimogonus; characters; India). — Bernhauer, 1931d: 569 (Mimogonus; Abyssinia). — Scheerpeltz, 1933: 1129 (Mimogonus; catalog). — Blackwelder, 1943: 172 (Mimogonus; characters; Cuba; Jamaica; Haiti; Dominican Republic; Puerto Rico; Guadeloupe). — Fagel, 1955: 18 (Mimogonus; characters; New Caledonia; Sumatra; Java; Tonkin; Ceylon). — Coiffait, 1979: 11 (Mimogonus; characters; Madagascar; Mauritius). — Irmler, 1981: 144 (Mimogonus; characters). DISTRIBUTION: New Caledonia, Indonesia, Vietnam, Singapore, Malaysia, Myanmar, Sri Lanka, India, Seychelles, Mascarene Islands, Madagascar, Tanzania, Ethiopia, West Indies, Mexico. fauveli Cameron, 1913: 326 (Osorius; Type locality: Haiti). — Cameron, 1923: 398 (Osorius; synonym of Mimogonus fumator). — Cameron, 1928d: 108 (Mimogonus; synonym of fumator). — Cameron, 1930: 304 (Mimogonus; synonym of fumator). — Scheerpeltz, 1933: 1129 (Mimogonus; synonym of fumator). — Blackwelder, 1943: 172 (Mimogonus; synonym of fumator). gabonicus Coiffait, 1967: 271 (Mimogonus; Type locality: Bélinga). DISTRIBUTION: Gabon. invisus Fagel, 1955: 22 (Mimogonus; Type locality: Congo belge: Stanleyville). — Fagel, 1969: 12 (Mimogonus; characters). DISTRIBUTION: Congo. kwangoensis Fagel, 1969: 14 (Mimogonus; Type locality: Kwango: territoire de Feshi, Kabanga, riv. Lutshina). DISTRIBUTION: Congo. leleupi Cameron, 1952, see: Mimogonellus. methneri Bernhauer, 1915, see: Mimogonellus. microps Sharp, 1889: 412 (Osorius; Type locality: Nagasaki). — Fauvel, 1903d: 261 (Mimogonus). — Bernhauer and Schubert, 1911: 141 (Mimogonus; catalog). — Bernhauer, 1922c: 226 (Mimogonus; Formosa). — Scheerpeltz, 1929a: 116 (Mimogonus; notes; China). — Scheerpeltz, 1933: 1129 (Mimogonus; catalog). — Nakane, 1963a: 84 (Mimogonus; characters; habitus photograph; Japan). — Naomi, 1987c: 13 (Mimogonus; characters; Japan). — Cho and Ahn, 1999: 179 (Mimogonus; characters; Korea). DISTRIBUTION: Japan, Taiwan, China, Korea. niger Cameron, 1930: 457 (Mimogonus; Type locality: Darjeeling; Debrepanni, alt. 6000 feet). — Scheerpeltz, 1933: 1129 (Mimogonus; catalog). — Abdullah and Qadri, 1970: 118 (Mimogonus; characters; East Pakistan). DISTRIBUTION: India, Bangladesh. 1186 OSORIINAE: MIMOGONUS Bulletin AMNH nigerrimus Cameron, 1950, see: Mimogonellus. reticulatus Fagel, 1969: 12 (Mimogonus; Type locality: Congo: Albertville, Kalemie). DISTRIBUTION: Congo. rhodesianus Fagel, 1969: 13 (Mimogonus; Type locality: South Rhodesia: Chipinga, 1200–1300 m). DISTRIBUTION: Zimbabwe. rossii Bordoni, 1981: 53 (Mimogonus; Type locality: Kenia, Masai Mara Reserve, fiume Mara presso Mara Bridge). DISTRIBUTION: Kenya. rufipennis Motschulsky, 1857a: 508 (Osorius; Type locality: Indes orientales). — Kraatz, 1859: 168 (Osorius). — Fauvel, 1903d: 261 (Mimogonus). — Bernhauer and Schubert, 1911: 141 (Mimogonus; catalog). — Cameron, 1930: 305 (Mimogonus; characters; East Indies). — Scheerpeltz, 1933: 1129 (Mimogonus; catalog). — Cameron, 1934d: 78 (Mimogonus). — Cameron, 1940: 86 (Mimogonus; characters). DISTRIBUTION: “East Indies”. rufus Cameron, 1950, see: Mimogonidius. saegeri Fagel, 1955: 20 (Mimogonus; Type locality: Congo belge: Bumba). — Fagel, 1969: 11 (Mimogonus; characters). DISTRIBUTION: Congo. subterraneus Fagel, 1955, see: Saegerius. Mimotrochus [2 species; Neotropical Region] Mimotrochus Irmler, 1987: 107 (species included: pecki, columbinus). TYPE SPECIES: Mimotrochus pecki Irmler, fixed by original designation. columbinus Irmler, 1987: 108 (Mimotrochus; Type locality: Colombia: C. Amara Tequendara). DISTRIBUTION: Colombia. pecki Irmler, 1987: 107 (Mimotrochus; Type locality: Colombia: Qbda. Susamaco, 23 km W. Villavicencio). DISTRIBUTION: Colombia. Neocaledonopsis [1 species; Oceanic Region] Neocaledonopsis Pace, 1990: 15 (species included: franzi). TYPE SPECIES: Neocaledonopsis franzi Pace, fixed by original designation and monotypy. franzi Pace, 1990: 17 (Neocaledonopsis; Type locality: N. Caledonie: Col d’Amieu). DISTRIBUTION: New Caledonia. Neosorius [56 species; Ethiopian and Madagascan Regions] Neosorius Fagel, 1959: 119 (species included: schoutedeni, burgeoni, sparsiceps, mon- tishoyoanus, sparsior, upembanus, sulcicollis, kaleheanus, straeleni, biambwensis, nyakasibaensis, angolanus, humicola, paradoxifrons, kataganus, leleupi, incisifrons, kivuanus, rugegensis, opaculus, chirindanus, usambarae, arebianus, pusillus, katom- peanus, dundoensis, minimus, simillimus, rugiceps, saegeri, ituriensis, gracilis, sculp- tus, basilweskyi, uelensis, fallax, burgeoni, wittei, mutakatoensis, hoyoensis, dela- marei, fauvelianus, incertus, rugosicollis). TYPE SPECIES: Osorius burgeoni Cameron, fixed by original designation. — Fagel, 1960a: 27 (notes). — Fagel, 1969: 79 (notes; key to species). Bulletin AMNH OSORIINAE: NEOSORIUS 1187
— Coiffait, 1979: 131 (characters; species of Madagascar). — Drugmand, 1998a: 216 (list of types in l’Institut royal des Sciences naturelles de Belgique). ampliventris Fagel, 1969: 98 (Neosorius; Type locality: Kivu: territoire de Lubero, Biambwa, 950 m). DISTRIBUTION: Congo. angolanus Fagel, 1959: 181 (Neosorius; Type locality: Angola: Fl. Camissombo. Also cited from Belgian Congo). — Fagel, 1969: 81 (Neosorius; characters). DISTRIBUTION: Angola, Congo. arcuatifrons Fagel, 1969: 97 (Neosorius; Type locality: Kivu: territoire de Kalehe, riv. Luhaho). DISTRIBUTION: Congo. arebianus Cameron, 1928: 22 (Osorius; Type locality: Belgian Congo: Arebi, Bondo Mabe; Haut Uele, Abimva). — Scheerpeltz, 1933: 1131 (Osorius; catalog). — Fagel, 1957a: 32 (Osorius; Urundi). — Fagel, 1959: 153 (Neosorius; characters; Belgian Congo). — Fagel, 1959b: 6 (Neosorius; Belgian Congo). — Fagel, 1960a: 28 (Neosorius; Uganda). — Fagel, 1963a: 341 (Neosorius; Guinea). — Fagel, 1969: 83 (Neosorius; characters). DISTRIBUTION: Burundi, Congo, Guinea. basilewskyi Fagel, 1957a: 32 (Osorius; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m. Also cited from Belgian Congo). — Cameron, 1950g: 20 (Osorius; [Note: Misidenification: Cited as burgeoni but listed by Fagel, 1959: 130 as though Cameron described a new species]; Belgian Congo). — Fagel, 1959: 130 (Neosorius; characters; Belgian Congo; Urundi). — Fagel, 1961: 276 (Neosorius; Belgian Congo). — Fagel, 1969: 85 (Neosorius; characters). DISTRIBUTION: Rwanda, Burundi, Congo. biambwensis Fagel, 1959: 179 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Lubero, Biambwe, riv. Lubau, 1000 m). — Fagel, 1969: 81 (Neosorius; characters). DISTRIBUTION: Congo. bifurcatus Bernhauer, 1935, see: burgeoni Cameron, 1928. brevipennis Fagel, 1969: 92 (Neosorius; Type locality: Transvaal: Soutpansberg distr., Hanglipbos Forest). DISTRIBUTION: South Africa. bugiriensis Fagel, 1960a: 29 (Neosorius; Type locality: Uganda: Bugiri, 1400 m). — Fagel, 1969: 81 (Neosorius; characters). DISTRIBUTION: Congo. burgeoni Cameron, 1928: 22 (Osorius; Type locality: Belgian Congo: Haut Uele, Moto). — Scheerpeltz, 1933: 1132 (Osorius; catalog). — Fagel, 1959: 125 (Neosorius; characters; Belgian Congo). — Fagel, 1961: 276 (Neosorius; Belgian Congo). — Fagel, 1969: 85 (Neosorius; characters). DISTRIBUTION: Congo. bifurcatus Bernhauer, 1935d: 98 (Osorius; Type locality: Uelé: Bambesa). — Fagel, 1959: 125 (Neosorius; synonym of burgeoni; some paratypes identified as uelensis). capicola Fagel, 1969: 94 (Neosorius; Type locality: Cape Province: Humansdorp, distr., Tsisikama Forest). DISTRIBUTION: South Africa. 1188 OSORIINAE: NEOSORIUS Bulletin AMNH chirindanus Fagel, 1959: 155 (Neosorius; Type locality: Mashonaland: Chirinda Forest). — Fagel, 1960a: 28 (Neosorius; Tanganyika). — Fagel, 1969: 83 (Neosorius; characters). DISTRIBUTION: Zimbabwe. compactus Walker, 1859, see: rufipes Motschulsky, 1857. delamarei Cameron, 1948a: 226 (Osorius; Type locality: Ivory Coast: Azaguié). — Fagel, 1959: 144 (Neosorius; characters; Ivory Coast; Guinea). — Fagel, 1963a: 340 (Neosorius; Guinea; Ivory Coast). — Fagel, 1969: 86 (Neosorius; characters). DISTRIBUTION: Ivory Coast, Guinea. dolosus Fagel, 1969: 91 (Neosorius; Type locality: Angola: Alto Chicapa, riv. Tchirimbo). DISTRIBUTION: Angola. dundoensis Cameron, 1950f: 117 (Osorius; Type locality: Angola: Dundo). — Fagel, 1959: 159 (Neosorius; characters; Angola). — Fagel, 1969: 83 (Neosorius; characters). DISTRIBUTION: Angola. fallax Fagel, 1959: 140 (Neosorius; Type locality: Congo Belge: Ituri: territoire de Bunia, mont Hoyo, 1200 m). — Fagel, 1961: 279 (Neosorius; Belgian Congo). — Fagel, 1969: 85 (Neosorius; characters). DISTRIBUTION: Congo. fauvelianus Fagel, 1959: 145 (Neosorius; Type locality: Afrique occidentale: Assinie). — Fagel, 1969: 86 (Neosorius; characters). DISTRIBUTION: Ivory Coast. furcatifrons Fagel, 1969: 88 (Neosorius; Type locality: Angola: Alto Chicapa, riv. Gungo). DISTRIBUTION: Angola. ghanaensis Fagel, 1961: 277 (Neosorius; Type locality: Ghana: Nr. Kumasi, Bobiri Forest Res.). — Fagel, 1963a: 341 (Neosorius; Guinea). — Fagel, 1969: 85 (Neosorius; characters). DISTRIBUTION: Ghana, Guinea. gracilis Fagel, 1959: 137 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Masisi, Mutakato, 800 m). — Fagel, 1969: 84 (Neosorius; characters). DISTRIBUTION: Congo. hoyoensis Fagel, 1959: 127 (Neosorius; Type locality: Congo Belge: Ituri: territoire de Bunia, mont Hoyo, 1200–1300 m). — Fagel, 1969: 86 (Neosorius; characters). DISTRIBUTION: Congo. humicola Fagel, 1959: 184 (Neosorius; Type locality: Congo Belge: Bas-Congo: territoire des Cataractes, Kavuya). — Fagel, 1969: 81 (Neosorius; characters). DISTRIBUTION: Congo. incertus Fagel, 1959: 138 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Lubero, Kyalamayhindi, vallée Lopo, 1600 m). — Fagel, 1969: 86 (Neosorius; characters). DISTRIBUTION: Congo. incisifrons Fauvel, 1905d: 195 (Osorius; Type locality: Monts de Cristal). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Fagel, 1957a: 32 (Osorius; Ruanda; Urundi). — Fagel, 1959: 166 (Neosorius; characters; Gabon; Belgian Congo). Bulletin AMNH OSORIINAE: NEOSORIUS 1189
— Fagel, 1969: 82 (Neosorius; characters). DISTRIBUTION: Rwanda, Burundi, Congo, Gabon. ituriensis Fagel, 1959: 135 (Neosorius; Type locality: Congo Belge: Ituri: territoire de Bunia, mont Hoyo, 1200 m). — Fagel, 1969: 84 (Neosorius; characters). DISTRIBUTION: Congo. kaleheanus Fagel, 1959: 185 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Kalehe, Bunyakiri, 1050 m). — Fagel, 1969: 81 (Neosorius; characters). DISTRIBUTION: Congo. katanganus Fagel, 1959: 151 (Neosorius; Type locality: Congo Belge: Katanga: environs d’Elisabethville, galerie forestière de la Kisanga). — Fagel, 1969: 82 (Neosorius; characters). DISTRIBUTION: Congo. katompeanus Bernhauer, 1932a: 84 (Osorius; Type locality: Katanga: Katompe). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Fagel, 1959: 157 (Neosorius; characters; Belgian Congo: Katompe. Also cited doubtfully from Ivory Coast). — Fagel, 1969: 83 (Neosorius; characters). DISTRIBUTION: Congo, Ivory Coast ?]. kivuanus Fagel, 1959: 165 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Kabare, Lwiro, 2000–2200 m. Also cited from Ruanda; Urundi). — Fagel, 1969: 82 (Neosorius; characters). DISTRIBUTION: Rwanda, Burundi, Congo. kwangensis Fagel, 1961, see: mutakatoensis Fagel, 1959. leleupi Fagel, 1959: 168 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Masisi, Mutakato, 800 m). — Fagel, 1969: 82 (Neosorius; characters). DISTRIBUTION: Congo. lunguenaensis Fagel, 1969: 89 (Neosorius; Type locality: Angola: riv. Lunguena, affl. Luangue). DISTRIBUTION: Angola. machadonianus Fagel, 1969: 90 (Neosorius; Type locality: Angola: Alto Chicapa, riv. Tchirimbo). DISTRIBUTION: Angola. minimus Fagel, 1959: 158 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Masisi, Mutakato, 800 m). — Fagel, 1969: 83 (Neosorius; characters). DISTRIBUTION: Congo. montishoyoanus Fagel, 1959: 176 (Neosorius; Type locality: Congo Belge: Ituri: territoire de Bunia, mont Hoyo, 1100 m). — Fagel, 1969: 80 (Neosorius; characters). DISTRIBUTION: Congo. mutakatoensis Fagel, 1959: 139 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Masisi, Mutakato, 800 m). — Fagel, 1961: 278 (Neosorius; Belgian Congo). — Fagel, 1969: 85 (Neosorius; characters). DISTRIBUTION: Congo. kwangensis Fagel, 1961: 278 (Neosorius; subspecies of mutakatoensis Fagel; Type locality: Congo: Kwango: territoire de Feshi, rive gauche Kwenge). — Fagel, 1969: 85 (Neosorius; subspecies of mutakatoensis; characters). 1190 OSORIINAE: NEOSORIUS Bulletin AMNH nyakasibaensis Fagel, 1959: 180 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Kabare, Nyakasiba, 1800 m). — Fagel, 1969: 81 (Neosorius; characters). DISTRIBUTION: Congo. opaculus Eppelsheim, 1895a: 134 (Osorius; Type locality: Togoland). — Fauvel, 1903e: 239 (Osorius; Cameroon; Gabon; Togo). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Fagel, 1959: 162 (Neosorius; characters; Gabon; Cameroon; Belgian Congo). — Fagel, 1961: 279 (Neosorius; Belgian Congo). — Fagel, 1969: 83 (Neosorius; characters). — Scheerpeltz, 1971c: 162 (Osorius; Cameroon). — Frank, 1982: 8 (Neosorius; list of parasites). DISTRIBUTION: Togo, Cameroon, Gabon, Congo. paradoxifrons Bernhauer, 1932a: 84 (Osorius; Type locality: Katanga: Katompe). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Fagel, 1959: 148 (Neosorius; characters; Belgian Congo). — Fagel, 1969: 82 (Neosorius; characters). DISTRIBUTION: Congo. pusillus Fagel, 1959: 161 (Neosorius; Type locality: Congo Belge: Ituri: territoire de Bunia, mont Hoyo, 1100 m). — Fagel, 1961: 279 (Neosorius; Belgian Congo). — Fagel, 1969: 83 (Neosorius; characters). DISTRIBUTION: Congo. rufipes Motschulsky, 1857a: 508 (Osorius; Type locality: Indes orientales). — Kraatz, 1859: 168 (Osorius). — Fauvel, 1902b: 73 (Osorius; Egypt; Ceylon; India; Burma; Sumatra; Mauritius; Réunion). — Bernhauer, 1902b: 43 (Osorius; Ceylon). — Fauvel, 1904a: 47 (Osorius; Tudia; Nilghiris; Ceylan; Bírmanie; Sumatra; Maurice; Réunion; Egypt). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Cameron, 1930: 302 (Osorius; characters; Ceylon; India). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Cameron, 1949b: 462 (Osorius; Formosa). — Scheerpeltz, 1957c: 227 (Osorius; Sumba). — Abdullah and Qadri, 1970: 116 (Osorius; characters; West Pakistan). — Watanabe and Shibata, 1976: 329 (Osorius; Malaysia). — Coiffait, 1976: 222 (Osorius; Tahiti). — Coiffait, 1979: 132 (Nesorius; characters; Réunion; Mauritius; Comoros; Madagascar). — Nishida, 1994: 73 (Osorius; [Note: I assume Nishida overlooked the transfer of the species to Neosorius]; Hawaii). — Nishida, 1997: 62 (Osorius; [Note: I assume Nishida overlooked of the transfer of the species to Neosorius]; Hawaii). DISTRIBUTION: Egypt, Mascarene Islands, Pakistan, India, Sri Lanka, Myanmar, Indonesia, Taiwan, Society Islands, Hawaii. compactus Walker, 1859: 52 (Osorius; Type locality: Ceylon). — Fauvel, 1902b: 73 (Osorius; synonym of rufipes). — Fauvel, 1904a: 47 (Osorius; synonym of rufipes). — Bernhauer and Schubert, 1911: 145 (Osorius; synonym of rufipes). — Cameron, 1930: 302 (Osorius; synonym of rufipes). — Scheerpeltz, 1957c: 227 (Osorius; synonym of rufipes). syriacus Fauvel, 1875c:viii (Osorius; Type locality: Port Saïd). — Fauvel, 1878: 100 [= 1878a: 20] (Osorius; Egypt). — Fauvel, 1886: 27 [= 1886a: 19] (Osorius; Egypt). — Fauvel, 1902b: 73 (Osorius; synonym of rufipes). — Fauvel, 1904a: 47 (Osorius; synonym of rufipes). Bulletin AMNH OSORIINAE: NEOSORIUS 1191
— Bernhauer and Schubert, 1911: 145 (Osorius; synonym of rufipes). — Cameron, 1930: 302 (Osorius; synonym of rufipes). — Scheerpeltz, 1933: 1136 (Osorius; synonym of rufipes). — Scheerpeltz, 1957c: 227 (Osorius; synonym of rufipes). rugegensis Fagel, 1957a: 34 (Osorius; Type locality: Ruanda: forêt du Rugege, 2100 m). — Fagel, 1959: 164 (Neosorius; characters; Belgian Congo). — Fagel, 1961: 279 (Neosorius; Belgian Congo). — Fagel, 1969: 82 (Neosorius; characters). DISTRIBUTION: Rwanda, Congo. rugiceps Boheman, 1848: 294 (Osorius; Type locality: Caffraria meridionalis). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Fagel, 1959: 146 (Neosorius; characters). — Fagel, 1969: 84 (Neosorius; characters). DISTRIBUTION: South Africa. rugosicollis Fagel, 1959: 141 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Masisi, Mutakato, 800 m). — Fagel, 1969: 86 (Neosorius; characters). DISTRIBUTION: Congo. saegeri Fagel, 1959: 130 (Neosorius; Type locality: Congo Belge: Parc National de la Garamba: Bagbele). — Fagel, 1969: 84 (Neosorius; characters). DISTRIBUTION: Congo. schoutedeni Fagel, 1959: 169 (Neosorius; Type locality: Congo Belge: Arebi). — Fagel, 1969: 80 (Neosorius; characters). DISTRIBUTION: Congo. sculptus Fagel, 1959: 133 (Neosorius; Type locality: Congo Belge: Albertville, moyenne Kimbi, Makongu, 950 m). — Fagel, 1960a: 28 (Neosorius; Uganda). — Fagel, 1961: 277 (Neosorius; Belgian Congo). — Fagel, 1969: 85 (Neosorius; characters). DISTRIBUTION: Congo, Uganda. selindaensis Fagel, 1969: 86 (Neosorius; Type locality: South Rhodesia: Mt. Selinda, forêt de Chirinda, 1300 m). DISTRIBUTION: Zimbabwe. simillimus Fagel, 1959: 160 (Neosorius; Type locality: Congo Belge: Kivu: territoire de Masisi, Mutakato, 800 m. Also cited from Angola). — Fagel, 1969: 83 (Neosorius; characters). DISTRIBUTION: Angola, Congo. sparsiceps Bernhauer, 1938d: 327 (Osorius; Type locality: Belgisches Kongo: Rutshuru). — Fagel, 1959: 173 (Neosorius; characters; Belgian Congo). — Fagel, 1969: 80 (Neosorius; characters). DISTRIBUTION: Congo. sparsior Fauvel, 1904d: 291 (Osorius; Type locality: Usambara: Derema, 850 m). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Fagel, 1959: 171 (Neosorius; characters; German East Africa). — Fagel, 1960a: 28 (Neosorius; Tanganyika). — Fagel, 1969: 80 (Neosorius; characters). DISTRIBUTION: Tanzania. straeleni Fagel, 1959: 183 (Neosorius; Type locality: Congo Belge: Katanga: Parc National de l’Upemba: Kamitungulu, affl. g. Lusinga, sous-affl. dr. Lufwa, 1760 m). — Fagel, 1959b: 7 (Neosorius; republication of original description; Belgian Congo). — Fagel, 1969: 81 (Neosorius; characters). DISTRIBUTION: Congo. 1192 OSORIINAE: NEOSORIUS Bulletin AMNH sulcicollis Fagel, 1959: 175 (Neosorius; Type locality: Gabon). — Fagel, 1969: 80 (Neosorius; characters). DISTRIBUTION: Gabon. syriacus Fauvel, 1875, see: rufipes Motschulsky, 1857. uelensis Fagel, 1959: 132 (Neosorius; Type locality: Congo Belge: Uele: Bambesa). — Cameron, 1950g: 20 (Osorius; [Note: Misidentification: Cited as sparsiceps but listed by Fagel, 1959: 132 as though Cameron described a new species]; Belgian Congo). — Fagel, 1960a: 28 (Neosorius; Uganda). — Fagel, 1969: 85 (Neosorius; characters). DISTRIBUTION: Congo, Uganda. upembanus Fagel, 1959: 173 (Neosorius; Type locality: Parc National de l’Upemba: Masombwe, 1120 m). — Fagel, 1959b: 6 (Neosorius; republication of original description; Belgian Congo). — Fagel, 1969: 80 (Neosorius; characters). DISTRIBUTION: Congo. usambarae Bernhauer, 1915d: 103 (Osorius; Type locality: Deutsch-Ostafrika: West Usam- bara). — Scheerpeltz, 1933: 1137 (Osorius; catalog). — Fagel, 1959: 156 (Neosorius; characters; German East Africa). — Fagel, 1969: 83 (Neosorius; characters). DISTRIBUTION: Tanzania. vicarius Fagel, 1969: 96 (Neosorius; Type locality: Cape Province: Humansdorp distr., Wit Els bos). DISTRIBUTION: South Africa. wittei Fagel, 1959: 128 (Neosorius; Type locality: Congo Belge: Parc National de l’Upemba, Kamitungulu, affl. g. Lusinga, s.-affl. dr Lufwa, 1760 m). — Fagel, 1959b: 5 (Neosorius; republication of original description; Belgian Congo). — Fagel, 1969: 85 (Neosorius; characters). DISTRIBUTION: Congo.
Nepalocylindrops [1 species; Palaearctic Region] Nepalocylindrops Coiffait, 1982: 141 (species included: deharvengi). TYPE SPECIES: Nepalo- cylindrops deharvengi Coiffait, fixed by original designation. deharvengi Coiffait, 1982: 141 (Nepalocylindrops; Type locality: Nepal: forêt royale de Gokarna, 1450 m). DISTRIBUTION: Nepal.
Nototorchus [2 species; Australian Region] Nototorchus McColl, 1985: 355 (replacement name for Nototrochus McColl). TYPE SPECIES: Holotrochus ferrugineus Broun, fixed by original designation and objective syn- onymy with Nototrochus for which type had been fixed. — Klimaszewski, Newton, and Thayer, 1996: 150 (endemic to New Zealand). Nototrochus McColl, 1982: 27 (species included: ferrugineus, montanus; preoccupied by Duncan, 1884). TYPE SPECIES: Holotrochus ferrugineus Broun, fixed by original designation. — McColl, 1982: 20 (key to species of New Zealand). — McColl, 1984: 31 (characters; key to species). brouni Bernhauer, 1939, see: ferrugineus Broun, 1893. ferrugineus Broun, 1893: 1034 (Holotrochus; Type locality: Woodhill, near Kaipara Railway; [Note: See lectotype designation by McColl, 1982]). Bulletin AMNH OSORIINAE: NOTOTORCHUS 1193
— Fauvel, 1900c: 186 (Holotrochus; characters; notes; New Zealand). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — McColl, 1982: 28 (Nototrochus; lectotype. designation but stated as holotype designation; characters; New Zealand). — McColl, 1984: 31 (Nototrochus; characters). — McColl, 1985: 355 (Nototorchus). — Klimaszewski and Watt, 1997: 170 (Nototorchus; characters; collecting notes; New Zealand). DISTRIBUTION: New Zealand. brouni Bernhauer, 1939f: 208 (Holotrochus; Type locality: New Zealand: Epsom; Auckland; Te Aroha). — McColl, 1982: 28 (Nototrochus; synonym of ferrugineus). montanus Broun, 1910: 18 (Holotrochus; Type locality: Mount Cook, near the Hermitage, 2500 ft.; [Note: See lectotype designation by McColl, 1982]). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — McColl, 1982: 30 (Nototrochus; lectotype designation; characters; New Zealand). — McColl, 1984: 32 (Nototrochus; characters). — McColl, 1985: 355 (Nototorchus). DISTRIBUTION: New Zealand.
Oeophronistus [1 species; Australian Region] Oeophronistus Blackburn, 1902: 20 (species included: australicus). TYPE SPECIES: Oeophro- nistus australicus Blackburn, fixed by monotypy. — Bernhauer and Schubert, 1911: 143 (world catalog; cited as Oephronistus; 1 species). — Notman, 1925: 2 (characters in key). — Blackwelder, 1952: 270 (type species: australicus). — Newton, 1990: 92 (characters in key; one described and one undescribed Australian species). australicus Blackburn, 1902: 21 (Oeophronistus; Type locality: Victoria). — Bernhauer and Schubert, 1911: 143 (Oeophronistus; catalog). — Oke, 1933: 104 (Oeophronistus; characters; Australia). — Newton, 1990: 92 (Oeophronistus). DISTRIBUTION: Australia.
Oligotyphlopsis [1 species; Palaearctic Region] Oligotyphlopsis Scheerpeltz, 1951: 118 (species included: nevesi). TYPE SPECIES: Oligo- typhlopsis nevesi Scheerpeltz, fixed by original designation and monotypy. — Coiffait, 1955b: 263 (characters in key). nevesi Scheerpeltz, 1951: 126 (Oligotyphlopsis; Type locality: Portugal: Porto). DISTRIBUTION: Portugal.
Oryssoma [1 species; Neotropical Region] Oryssoma Notman, 1925: 2, 4 (species included: schwarzi). TYPE SPECIES: Oryssoma schwarzi Notman, fixed by original designation and monotypy. — Scheerpeltz, 1933: 1137 (world catalog supplement). — Blackwelder, 1944: 112 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 278 (type species: schwarzi). schwarzi Notman, 1925: 5 (Oryssomma; Type locality: Cacao, Trece Aguas, Alta V. Paz, Guatemala). — Scheerpeltz, 1933: 1137 (Oryssoma; catalog). DISTRIBUTION: Guatemala. 1194 OSORIINAE: OSORIELLUS Bulletin AMNH
Osoriellus [15 species; Ethiopian Region] Osoriellus Fagel, 1959: 186 (species included: linearis, zanzibaricus, kibatsiroensis, leleupi- anus, walikalensis, kabwensis, rivularis, mabaliensis). TYPE SPECIES: Osorius lin- earis Bernhauer, fixed by original designation. — Fagel, 1960a: 30 (notes). — Drugmand, 1998a: 218 (list of types in l’Institut royal des Sciences naturelles de Belgique). asperiventris Fagel, 1969: 104 (Osoriellus; Type locality: Kwango: territoire de Feshi, rive g. Kwenge). DISTRIBUTION: Congo. filiformis Fagel, 1969: 103 (Osoriellus; Type locality: Congo: Equateur: territorie de Bikoro, Mabali, esobe de M’Bolo). DISTRIBUTION: Congo. kabwensis Fagel, 1959: 196 (Osoriellus; Type locality: Congo Belge: Katanga: Parc National de l’Upemba, Kabwe-sur-Muye, affl. Lubanga, 1320 m). — Fagel, 1959b: 9 (Osoriellus; cited as new species; republication of original description; Belgian Congo). — Fagel, 1969: 101 (Osoriellus; characters). DISTRIBUTION: Congo. kibatsiroensis Fagel, 1959: 190 (Osoriellus; Type locality: Congo Belge: Kivu: territoire de Lubero, mont Kibatsiro, Visiki, 2080 m). — Fagel, 1969: 100 (Osoriellus; characters). DISTRIBUTION: Congo. leleupianus Fagel, 1959: 193 (Osoriellus; Type locality: Congo Belge: Kivu: territoire de Fizi, Bas Itombwe, 800–900 m). — Fagel, 1961: 281 (Osoriellus; Ghana). — Fagel, 1969: 101 (Osoriellus; characters). DISTRIBUTION: Congo. linearis Bernhauer, 1915d: 104 (Osorius; Type locality: Deutsch-Ostafrika: Kibosho, Arusha-Ju). — Cameron, 1950g: 20 (Osorius; Belgian Congo). — Fagel, 1959: 188 (Osoriellus; characters; East Africa). — Fagel, 1960a: 30 (Osoriellus; Tanganyika). — Fagel, 1969: 100 (Osoriellus; characters). DISTRIBUTION: Tanzania, Congo. longicollis Fagel, 1961, see: walikalensis Fagel, 1959. mabaliensis Fagel, 1959: 195 (Osoriellus; Type locality: Congo Belge: Tshuapa: lac Tumba, Mabali, 350 m). — Fagel, 1969: 101 (Osoriellus; characters). DISTRIBUTION: Congo. mariepskopensis Fagel, 1969: 101 (Osoriellus; Type locality: Transvaal: Pilgrims Rest distr., Mariepskop Forest, 1100 m). DISTRIBUTION: South Africa. mwengensis Fagel, 1969: 109 (Osoriellus; Type locality: Kivu: Mwenga, 1250 m). DISTRIBUTION: Congo. reticulifrons Fagel, 1969: 107 (Osoriellus; Type locality: Kivu: territoire de Mwenga, rive dr. de l’Elila, 650 m). DISTRIBUTION: Congo. rivularis Cameron, 1950f: 117 (Osorius; Type locality: Angola: Dundo: River Dundundu). — Fagel, 1959: 191 (Osoriellus; characters; Angola; Belgian Congo). — Fagel, 1959b: 8 (Osoriellus; Belgian Congo). — Fagel, 1969: 101 (Osoriellus; characters). DISTRIBUTION: Angola, Congo. Bulletin AMNH OSORIINAE: OSORIELLUS 1195 selindaensis Fagel, 1969: 102 (Osoriellus; Type locality: South Rhodesia: Mont Selinda, forêt de Chirinda, 1300 m). DISTRIBUTION: Zimbabwe. strictus Fagel, 1969: 105 (Osoriellus; Type locality: Congo: territoire d’Albertville, Kalemie, Bendera, 1000 m). DISTRIBUTION: Congo. walikalensis Fagel, 1959: 197 (Osoriellus; Type locality: Congo Belge: Kivu: territoire de Masisi, Walikale, 700 m). — Fagel, 1969: 101 (Osoriellus; characters). DISTRIBUTION: Ghana, Congo. longicollis Fagel, 1961: 280 (Osoriellus; subspecies of walikalensis; Type locality: Ghana: Nr. Kumasi, Bobiri Forest Res.). — Fagel, 1969: 101 (Osoriellus; subspecies walikalensis; characters). zanzibaricus Fagel, 1959: 189 (Osoriellus; Type locality: Zanzibar). — Fagel, 1969: 100 (Osoriellus; characters). DISTRIBUTION: Zanzibar.
Osoriocanthus [7 species; Ethiopian Region] Osoriocanthus Fagel, 1959: 112 (species included: alienus, nitidus, bernhaueri, leleupi). TYPE SPECIES: Osorius alienus Cameron, fixed by original designation. — Fagel, 1969: 75 (notes; key to species). — Drugmand, 1998a: 218 (list of types in l’Institut royal des Sciences naturelles de Belgique). alienus Cameron, 1950c: 93 (Osorius; Type locality: Belgian Congo: Mayidi). — Fagel, 1959: 113 (Osoriocanthus; characters; Belgian Congo). — Fagel, 1969: 75 (Osoriocanthus; characters). DISTRIBUTION: Congo. bernhaueri Fagel, 1959: 117 (Osoriocanthus; Type locality: Congo Belge: Kivu: territoire de Masisi, Mutakato, 800 m). — Fagel, 1969: 75 (Osoriocanthus; characters). DISTRIBUTION: Congo. decelleanus Fagel, 1969: 77 (Osoriocanthus; Type locality: Côte d’Ivoire: Bingerville). DISTRIBUTION: Ivory Coast. leleupi Fagel, 1959: 118 (Osoriocanthus; Type locality: Congo Belge: Kibali-Ituri: territoire d’Epulu, Mambasa-Mungbere, 900 m). — Fagel, 1969: 75 (Osoriocanthus; characters). DISTRIBUTION: Congo. nitidus Fagel, 1959: 115 (Osoriocanthus; Type locality: Congo Belge: Ituri: territoire de Bunia, mont Hoyo, 1200–1300 m). — Fagel, 1969: 75 (Osoriocanthus; characters). DISTRIBUTION: Congo. surdus Fagel, 1969: 76 (Osoriocanthus; Type locality: Angola: Alto Cuilo, Tcha-Muchito). DISTRIBUTION: Angola. trapezicollis Fagel, 1969: 78 (Osoriocanthus; Type locality: Côte d’Ivoire: Mont Nimba, base Richard Mollard, 1450 m). DISTRIBUTION: Ivory Coast.
Osoriopsis [10 species; Ethiopian Region] Osoriopsis Fagel, 1955: 119 (species included: scabriventris, ituriensis, rugegensis, crassi- cornis, urundiensis). TYPE SPECIES: Osoriopsis scabriventris Fagel, fixed by original designation. 1196 OSORIINAE: OSORIOPSIS Bulletin AMNH
— Fagel, 1959: 65 (characters). — Fagel, 1960a: 26 (key to African species). — Drugmand, 1998a: 218 (list of types in l’Institut royal des Sciences naturelles de Belgique). basilewskyi Fagel, 1960a: 23 (Osoriopsis; Type locality: Tanganyika Territory: Bunduki, Uluguru Mts., gorge de la Mungula, 1500 m). DISTRIBUTION: Tanzania. clavicornis Coiffait, 1967: 274 (Osoriopsis; Type locality: Bélinga). DISTRIBUTION: Gabon. crassicornis Fagel, 1955: 128 (Osoriopsis; Type locality: Ruanda: forêt de Rugege). DISTRIBUTION: Rwanda. ituriensis Fagel, 1955: 124 (Osoriopsis; Type locality: Congo Belge: Ituri: territoire de Bunia, mont Hoyo, 1200 m). DISTRIBUTION: Congo. kabarensis Fagel, 1959: 65 (Osoriopsis; Type locality: Congo Belge: Kivu: territoire de Kabare, riv. Nyakagera, 1600–1700 m). DISTRIBUTION: Congo. leleupi Fagel, 1960a: 23 (Osoriopsis; Type locality: Tanganyika Territory: Uluguru Mt, sommet du Kidunda, 1800–1950 m). DISTRIBUTION: Tanzania. pusilla Fagel, 1959: 66 (Osoriopsis; Type locality: Congo Belge: Kivu: territoire de Masisi, lacs Ndalaga et Lukubi, 1780 m). DISTRIBUTION: Congo. rugegensis Fagel, 1955: 125 (Osoriopsis; Type locality: Ruanda-Urundi: forêt du Rugege, 2150 m). DISTRIBUTION: Rwanda-Burundi. scabriventris Fagel, 1955: 121 (Osoriopsis; Type locality: Congo Belge: Kivu: territoire d’Uvira, Haute Sange, 2800 m). DISTRIBUTION: Congo. urundiensis Fagel, 1955: 129 (Osoriopsis; Type locality: Urundi: territoire de Bururi: rivière Sikuvyayi, 1000 m). DISTRIBUTION: Burundi. Osorius [276 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions] Osorius Guérin-Méneville, 1829: pl. 9, fig. 11 (species included: brasiliensis; [Note: Black- welder, 1952: 279, cited Latreille, 1829: 438, as the author of this name. Latreille listed no species in the genus and the date of publication for Guérin-Méneville’s work is September 12, 1829 (see note attached to the citation in the bibliography) and that of Latreille is, in lieu of other evidence, December 31, 1829]). TYPE SPECIES: Oso- rius brasiliensis Guérin-Méneville, fixed by monotypy. — Dejean, 1821: 24 (unavailable genus-group name; published without characters and with one unavailable species-group name). — Latreille, 1825: 245 (characters; genus cited with a group of genera and without characters cited that differentiate the genera). — Berthold, 1827: 332 (characters but not differentiated from other genera). — Latreille, 1829: 438 (characters; without species). — Latreille, 1832: 83 (characters; notes). — Griffith and Pidgeon, 1832: 295 (characters). — Dejean, 1833: 67 (list of species). — Erichson, 1840: 753 (characters). — Laporte, 1840: 188 (characters). — Lacordaire, 1854: 112 (characters; notes; list of species). — Kraatz, 1857d: 806 (characters; notes). Bulletin AMNH OSORIINAE: OSORIUS 1197
— LeConte, 1861a: 69 (characters). — LeConte, 1863: 25 (list of North American species). — Gemminger and Harold, 1868: 642 (catalog). — Fauvel, 1875a: XIII [= 1875b: 215] (catalog). — Sharp, 1876: 381 (notes). — Fauvel, 1878f: 209 (characters; key to New Guinea species). — Duvivier, 1883: 184 (catalog). — Lynch, 1884: 345 (characters). — Sharp, 1887: 677 (notes; Mexican and Central American species). — Bernhauer and Schubert, 1911: 143 (world catalog; 83 species). — Leng, 1920: 98 (catalog of North American species). — Cameron, 1921a: 351, 401 (characters; key to Singapore species; catalog). — Winkler, 1925: 347 (catalog for Palaearctic region). — Notman, 1925: 3, 5, 6, 25 (key to species; characters in key; catalog of New World species). — Cameron, 1925: 22, 105 (catalog of Indian species). — Cameron, 1930: 290 (characters; key to species of British India). — Scheerpeltz, 1933: 1131 (world catalog supplement). — Darlington, 1937: 285 (key to West Indian species). — Paulian, 1941: 176 (larval characters). — Bernhauer, 1942b (key to Philippine species). — Blackwelder, 1943: 174 (characters; type species: latipes; key to species of West Indies). — Blackwelder, 1944: 111 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 279 (genus attributed to Latreille, 1829; type species). — Fagel, 1958a: 27 (characters; type species: brasiliensis). — Abdullah and Qadri, 1970: 116, 117 (key to species of East and West Pakistan). — Shibata, 1973c: 36 (checklist of species of Taiwan). — Tikhomirova, 1973a: 147 (checklist of species of USSR). — Moore and Legner, 1974c: 115 (characters; notes). — Moore and Legner, 1975: 174 (catalog; North America north of Mexico). — Shibata, 1976: 168 (checklist of species of Japan). — Moore and Legner, 1979: 154 (characters; notes). — Spahr, 1981: 95 (references to specimens from amber and copal). — Hammond, 1984: 197 (checklist of species of Borneo). — Frank, 1986a: 365 (checklist of species of Florida). — Naomi, 1986: 34 (characters; species of Japan). — Newton, 1990: 92 (characters in key; two described and two undescribed Australian species). — Frank, 1991: 345 (larval characters). — Nishida, 1994: 73 (species from Hawaii). — Downie and Arnett, 1996: 428 (characters; key to species of Northeastern North America). — Lundgren, 1998: 46 (list of species from Florida; records for other States listed for each species). — Drugmand, 1998a: 219 (list of types in l’Institut royal des Sciences naturelles de Belgique). — Newton, Thayer, Ashe, and Chandler, 2000: 378 (9 Nearctic species; notes; characters in key). — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 58 (necrophilous species in Mexico). Molosoma Say, 1831: 48 [= 1834: 462] (species included: latipes). TYPE SPECIES: Oxytelus latipes Gravenhorst, fixed by monotypy. — Erichson, 1840: 753 (synonym of Osorius). — Lacordaire, 1854: 112 (synonym of Osorius). — Gemminger and Harold, 1868: 642 (synonym of Osorius). — Fauvel, 1875a: XIII [= 1875b: 215] (synonym of Osorius). — Bernhauer and Schubert, 1911: 143 (synonym of Osorius). — Cameron, 1930: 290 (synonym of Osorius). — Blackwelder, 1943: 174 (synonym of Osorius; type species latipes). — Blackwelder, 1952: 250 (type species: latipes). 1198 OSORIINAE: OSORIUS Bulletin AMNH acupunctus Bernhauer, 1933e: 332 (Osorius; Type locality: Argentinien: Buenos Aires). DISTRIBUTION: Argentina. adjacens Cameron, 1948, see: Afrosorius. affinis Sharp, 1876: 385 (Osorius; Type locality: St. Paulo). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Notman, 1925: 8 (Osorius; characters in key). — Scheerpeltz, 1933: 1131 (Osorius; catalog). DISTRIBUTION: Brazil. alienus Cameron, 1950, see: Osoriocanthus. aliiceps Cameron, 1928d: 110 (Osorius; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1131 (Osorius; catalog). DISTRIBUTION: Indonesia. almorensis Cameron, 1945: 66 (Osorius; Type locality: Almora: Kali Valley, altitude 3000 feet). DISTRIBUTION: India. alternans Bernhauer, 1920a: 137 (Osorius; Type locality: Peru: Lima; 11 3 S. 75 17 W; Bolivien: Yuracaris). — Notman, 1925: 10, 24 (Osorius; characters in key; Bolivia). — Scheerpeltz, 1933: 1131 (Osorius; catalog). DISTRIBUTION: Peru, Bolivia. alticola Cameron, 1942: 109 (Osorius; Type locality: Sikkim: Karponang, altitude 10,000 feet). — Cameron, 1928c: 559 (Osorius; [Note: Misidentified as sikkimensis according to Cameron, 1942: 109]; Sikkim). DISTRIBUTION: India. alutaceus Bernhauer, 1904, see: Madecosorius. alutipennis Bernhauer, 1939, see: Afrosorius. alutiventris Bernhauer, 1942b: 231 (Osorius; Type locality: Borneo: Sandakan). DISTRIBUTION: Borneo. amicus Bernhauer, 1942b: 232 (Osorius; Type locality: Mindanao: Surigao). DISTRIBUTION: Philippines. angolensis Cameron, 1950, see: Cylindrops. angustulus Sharp, 1874: 89 (Osorius; Type locality: Nita, near Nagasaki). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Nakane, 1963a: 84 (Osorius; characters; habitus photograph; Japan). — Naomi, 1986: 38 (Osorius; characters; Japan). — Hasegawa and Kanie, 1992: 20 (Osorius; seashore collection; Japan). — Watanabe, 1995b: 250 (Osorius; Oki-no-shima). — Watanabe, 1996a: 10 (Osorius; cave collection; Japan). DISTRIBUTION: Japan. annandalei Bernhauer, 1911, see: Indosorius. antipodum Cameron, 1937a: 91 (Osorius; Type locality: New Guinea: Kokoda, alt. 1200 ft.). DISTRIBUTION: New Guinea. arebianus Cameron, 1928, see: Neosorius. argentinus Bernhauer, 1911a: 403 (Osorius; Type locality: Argentinien: Gob. Misiones). — Notman, 1925: 6 (Osorius; characters in key). — Scheerpeltz, 1933: 1131 (Osorius; catalog). — Bernhauer, 1933e: 332 (Osorius; characters in key; Argentina). DISTRIBUTION: Argentina. Bulletin AMNH OSORIINAE: OSORIUS 1199 armifrons Cameron, 1929b: 445 (Osorius; Type locality: Pahang: Cmaeron’s Highlands, 5000 feet). — Scheerpeltz, 1933: 1131 (Osorius; catalog). DISTRIBUTION: Malaysia. aspericeps Fauvel, 1905d: 194 (Osorius; replacement name of rugiceps Kraatz). — Bernhauer, 1928: 9 (Osorius; Sumatra). — Scheerpeltz, 1933: 1131 (Osorius; catalog). DISTRIBUTION: Myanmar, Malaysia, Indonesia, Borneo, “East Indies”. rugiceps Kraatz, 1859: 166 (Osorius; [preoccupied]; Type locality: India orientalis). — Bernhauer and Schubert, 1911: 145 (Osorius; synonym of kraatzi). — Cameron, 1930: 299 (Osorius; synonym of kraatzi). kraatzi Bernhauer and Schubert, 1911: 144 (Osorius; replacement name for rugiceps Kraatz). — Cameron, 1930: 299 (Osorius; characters; Burma; Malay Peninsula; Sumatra; Borneo). — Scheerpeltz, 1933: 1131 (Osorius; synonym of aspericeps). kraatzi Cameron, 1925: 22 (Osorius; replacement name for rugiceps Kraatz; [preoccu- pied]). — Scheerpeltz, 1933: 1131 (Osorius; synonym of aspericeps). crenulatus Cameron, 1928d: 109 (Osorius; Type locality: Sumatra: Fort de Kock). — Cameron, 1930: 299 (Osorius; synonym of kraatzi). — Scheerpeltz, 1933: 1131 (Osorius; synonym of aspericeps). asperifrons Cameron, 1928g: 437, 438 (Osorius; Type locality: Borneo: Baram district). — Scheerpeltz, 1933: 1131 (Osorius; catalog). DISTRIBUTION: Borneo. assamensis Cameron, 1942: 108 (Osorius; Type locality: Assam: Shillong, altitude 6000 feet). DISTRIBUTION: India. assiniensis Fauvel, 1903, see: Afrosorius. asymmetricus Fauvel, 1901: 72 (Osorius; Type locality: Venezuela: Caracas, Puerto Cabello, San Esteban, Colonie Tovar, Sarare; Colombie; Panama; Trinidad). — Sharp, 1887: 678 (Osorius; [Note: Misidentification: Fauvel, 1891: 91 attributed canaliculatus to Sharp but Sharp credited the species to Solsky; Bernhauer and Schubert, 1911: 143 cited canaliculatus Sharp as a synonym of asymmetricus]; notes; Mexico; Guatemala; Panama). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Notman, 1925: 6 (Osorius; characters in key). — Scheerpeltz, 1933: 1131 (Osorius; catalog). DISTRIBUTION: Venezuela, Colombia, Panama, Guatemala, Mexico, West Indies. ater Perty, 1830: 30 (Osorius; Type locality: Brasil: Prov. Minarum). — Erichson, 1840: 754 (Osorius; characters; Brazil). — Laporte, 1840: 188 (Osorius; characters; Brazil). — R. Sahlberg, 1847: 803 (Osorius; Brazil). — Redtenbacher, 1867: 31 (Osorius; Brazil). — Fauvel, 1877a: xxiv (Osorius; Brazil). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Notman, 1925: 6, 21 (Osorius; characters in key; Brazil). — Scheerpeltz, 1933: 1131 (Osorius; catalog; Argentina). DISTRIBUTION: Brazil, Argentina. cornutus Laporte, 1832: 395 (Osorius; Type locality: Brésil). — Fauvel, 1877a: xxiv (Osorius; synonym of ater). — Bernhauer and Schubert, 1911: 143 (Osorius; synonym of ater). bacillus Bernhauer, 1942b: 227 (Osorius; Type locality: Luzon: Mt. Makiling). DISTRIBUTION: Philippines. 1200 OSORIINAE: OSORIUS Bulletin AMNH bakeri Bernhauer, 1915e: 117 (Osorius; Type locality: Luzon: Laguna, Mount Maquiling). — Scheerpeltz, 1933: 1131 (Osorius; catalog). — Bernhauer, 1942b: 222 (Osorius; characters; Philippines). DISTRIBUTION: Philippines. bangoensis Bernhauer, 1942b: 228 (Osorius; Type locality: Mindanao: P. Bango). DISTRIBUTION: Philippines. banksi Scheerpeltz, 1933: 1132 (Osorius; replacement name for mjobergi Cameron, cited as mjöbergi Cameron). — Hammond, 1984: 198 (Osorius; Borneo). DISTRIBUTION: Borneo. mjobergi Cameron, 1928f: 404 (Osorius; [preoccupied]; Type locality: Mt. Penrissen). — Scheerpeltz, 1933: 1132 (Osorius; cited as mjöbergi; synonym of banksi). — Hammond, 1984: 198 (Osorius; cited as mjobergi; synonym of banksi). banyosensis Herman, 2001: 40 (Osorius; replacement name for luzonicus Bernhauer, 1942). DISTRIBUTION: Philippines. luzonicus Bernhauer, 1942b: 223 (Osorius; [preoccupied]; Type locality: Luzon: Mt. Makiling, Imugan, Butac, Laguna, Los Banos, Balbalan; Siargao: Dapa). — Herman, 2001: 40 (Osorius; synonym of banyosensis). basilewskyi Fagel, 1957, see: Neosorius. basipennis Cameron, 1941c: 494 (Osorius; Type locality: Philippines: Luzon: Los Banos). — Herman, 2001: 22 (Osorius; synonymy). DISTRIBUTION: Philippines. basipennis Bernhauer, 1942b: 221 (Osorius; [preoccupied]; Type locality: Luzon: Los Banos, Mt. Makiling; Malinas, Tayabus). — Herman, 2001: 22 (Osorius; synonym of basipennis). basipennis Bernhauer, 1942, see: basipennis Cameron, 1941. beebei Bernhauer, 1914c: 86 (Osorius; Type locality: Nordost-Birma: Sansi Gorge an der chinesischen Grenze, 6000 bis 8000 Fuss). — Cameron, 1930: 293 (Osorius; characters; Burma). — Scheerpeltz, 1933: 1132 (Osorius; catalog). — Scheerpeltz, 1965: 101 (Osorius; Burma). DISTRIBUTION: Myanmar. belzebub Bernhauer, 1942b: 218 (Osorius; Type locality: Luzon: Mt. Polis). DISTRIBUTION: Philippines. bequaerti Bernhauer, 1932, see: Afrosorius. biarmatus Darlington, 1937: 287 (Osorius; Type locality: Southwestern Haiti: northeastern foothills of the Massif de la Hotte, 2000–4000 ft.). — Blackwelder, 1943: 186 (Osorius; characters; Haiti). DISTRIBUTION: West Indies. bicornis Heller, 1898: 4 (Osorius; Type locality: Celebes meridionalis, Lompo Batang, altitudine 3000 ped.). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Cameron, 1928d: 109 (Osorius; Sumatra). DISTRIBUTION: Indonesia. bicornutus Cameron, 1928g: 435, 438 (Osorius; Type locality: Borneo: Baram district). — Scheerpeltz, 1933: 1132 (Osorius; catalog). — Cameron, 1933b: 341 (Osorius; British North Borneo). DISTRIBUTION: Borneo. bidens Cameron, 1928g: 434, 438 (Osorius; Type locality: Borneo: Lio Matu). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Borneo. Bulletin AMNH OSORIINAE: OSORIUS 1201 bidentatus Cameron, 1928g: 435, 438 (Osorius; Type locality: Borneo: Baram district). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Borneo. bifurcatus Bernhauer, 1935, see: Neosorius. biimpressus Bernhauer, 1942b: 225 (Osorius; Type locality: Leyte). DISTRIBUTION: Philippines. birmanus Cameron, 1930: 301 (Osorius; Type locality: Burma: Tenasserim; Tavoy). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Myanmar. bispinosus Bernhauer, 1926f: 314 (Osorius; Type locality: Buru). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Indonesia. bistriatus Bernhauer, 1942b: 218 (Osorius; Type locality: Siargao: Dapa; Luzon: Imugan). DISTRIBUTION: Philippines. boettcheri Bernhauer, 1942b: 223 (Osorius; Type locality: Siargao: Dapa; Palawan: Binaluan; Luzon: Mt. Makiling, Los Banos; Panzon; Aroroy; Biliran). DISTRIBUTION: Philippines. boops Sharp, 1887: 679 (Osorius; Type locality: Panama, Volcan de Chiriqui 2000 to 4000 feet). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Notman, 1925: 9 (Osorius; characters in key). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Panama. borneensis Scheerpeltz, 1933: 1132 (Osorius; replacement name for gigantulus Cameron). DISTRIBUTION: Borneo. gigantulus Cameron, 1928f: 403 (Osorius; [preoccupied]; Type locality: Mt. Penrissen, 4000 feet). brachypterus Cameron, 1941c: 495 (Osorius; Type locality: Philippines: Panaon. Pinutan). DISTRIBUTION: Philippines. brasiliensis Guérin-Méneville, 1829: pl. 9, fig. 11 (Osorius; Type locality: Not cited). — Latreille, 1832: 86 (Osorius; characters; Brazil). — Laporte, 1835: 131 (Osorius; characters; notes). — Blanchard, 1842: 88 (Osorius; Bolivia). — Erichson, 1840: 755 (Osorius; characters; Brazil). — Guérin-Méneville, 1844a: 32 (Osorius; notes; Brazil). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Notman, 1925: 9, 23 (Osorius; characters in key; Paraguay). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Bolivia, Brazil, Paraguay. breviceps Notman, 1925: 7, 12 (Osorius; Type locality: San Bernardino, Paraguay). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Paraguay. brevicornis Notman, 1920b: 698 (Osorius; Type locality: Pensacola, Florida). — Notman, 1925: 8, 23 (Osorius; characters in key; Florida; Colima, Mexico). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: USA. brevipennis Notman, 1925: 7, 14 (Osorius; Type locality: Fort Grant, Arizona; also examples from Mexico and Guatemala). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: USA, Mexico, Guatemala. 1202 OSORIINAE: OSORIUS Bulletin AMNH brunneipennis Cameron, 1941c: 495 (Osorius; Type locality: Philippines: S. Luzon: Mt. Isarog. Biliran). — Herman, 2001: 22 (Osorius; synonymy). DISTRIBUTION: Philippines. brunneipennis Bernhauer, 1942b: 223 (Osorius; [preoccupied]; Type locality: Biliran; Luzon: Mt. Isarog; Mindanao: P. Bango; Siargao, Leyte). — Herman, 2001: 22 (Osorius; synonym of brunneipennis). bruneipennis Bernhauer, 1942, see: brunneipennis Cameron, 1941. brunnicornis† Hope, 1837: 52 (Osorius; copal from the Old World; Type locality: not cited). DISTRIBUTION: ?. bryanti Cameron, 1941a: 206 (Osorius; Type locality: W. Sarawak: Mt. Matang). DISTRIBUTION: Malaysia. burgeoni Cameron, 1928, see: Neosorius. buscki Notman, 1925: 9, 19 (Osorius; Type locality: Santo Domingo, W.I.; also listed from Cuba). — Scheerpeltz, 1933: 1132 (Osorius; catalog). — Darlington, 1937: 297 (Osorius; Dominican Republic; Haiti; Cuba [Note: The Cuban specimen was described as a new species, cubensis Blackwelder]). — Blackwelder, 1943: 186 (Osorius; characters). DISTRIBUTION: West Indies. callifrons Bernhauer, 1926, see: Indosorius. calvus Eppelsheim, 1895b: 406 (Osorius; Type locality: Sikkim im Himalaya). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Cameron, 1930: 295 (Osorius; characters; Sikkim). — Scheerpeltz, 1933: 1132 (Osorius; catalog). — Abdullah and Qadri, 1970: 118 (Osorius; characters; East Pakistan). DISTRIBUTION: India, Bangladesh. cameroni Scheerpeltz, 1933: 1132 (Osorius; replacement name for intermedius Cameron). DISTRIBUTION: Indonesia. intermedius Cameron, 1928d: 110 (Osorius; [preoccupied]; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1132 (Osorius; synonym of cameroni). cameronianus Scheerpeltz, 1933: 1132 (Osorius; replacement name for planifrons Cameron). DISTRIBUTION: Borneo. planifrons Cameron, 1928g: 435, 438 (Osorius; [preoccupied]; Type locality: Borneo: Baram district). — Cameron, 1933b: 341 (Osorius; British North Borneo). — Scheerpeltz, 1933: 1132 (Osorius; synonym of cameronianus). canaliculatus Solsky, 1870a: 265 (Osorius; Type locality: Mexique). — Bernhauer and Schubert, 1911: 143 (Osorius; catalog). — Notman, 1925: 10, 24 (Osorius; characters in key; Guatemala; Mexico). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Mexico, Guatemala. carinicollis Notman, 1925: 7, 15 (Osorius; Type locality: Cacao Trece Aguas, Alta V. Paz, Guatemala). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Guatemala. carinellus Fauvel, 1878f: 211 (Osorius; Type locality: Nouvelle-Guinée méridionale, Ile Yule, Roro). — Bernhauer and Schubert, 1911: 144 (Osorius; cited as cariniellus; catalog). — Cameron, 1931: 356 (Osorius; New Guinea). DISTRIBUTION: New Guinea. Bulletin AMNH OSORIINAE: OSORIUS 1203 carinifrons Cameron, 1936a: 35 (Osorius; Type locality: Java: Batoerraden, G. Slamat). DISTRIBUTION: Indonesia. cautis Darlington, 1937: 295 (Osorius; Type locality: Haiti: summit of the ridge just above Roche Croix, on a northern spur of Mt. La Hotte, 5000 ft.). — Blackwelder, 1943: 184 (Osorius; characters; Haiti). DISTRIBUTION: West Indies. celebesus Heller, 1897: 2 (Osorius; Type locality: Celebes septentrionalis, Minahassa, Tomohon et Masarang). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). DISTRIBUTION: Indonesia. cheesmani Cameron, 1937a: 90 (Osorius; Type locality: New Guinea: Mafulu, alt. 4000 ft.). DISTRIBUTION: New Guinea. chinensis Bernhauer, 1934: 4 (Osorius; Type locality: Mittel-Szechuan: Hwa-Yin-Shan, 1600 m). DISTRIBUTION: China. chinensis J. Li, 1993, see: jingkei Herman, 2001. collaris Bernhauer, 1927f: 261 (Osorius; Type locality: Sumatra). — Scheerpeltz, 1933: 1132 (Osorius; catalog). DISTRIBUTION: Indonesia. columbinus Bernhauer, 1920a: 138 (Osorius; Type locality: Kolumbien: Urwald bei Carmen, 1400 m). — Notman, 1925: 11 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Colombia. compactus Walker, 1859, see: Neosorius. confusus Notman, 1925: 10, 20 (Osorius; Type locality: Omealca, Vera Cruz, Mexico). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Mexico. congoanus Bernhauer, 1932a: 84 (Osorius; Type locality: Arebi, Bondo Mabe). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Congo. constrictus Fauvel, 1895: 196 (Osorius; Type locality: Bírmanie, Carin Ghécu, 1300–1400 m). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Cameron, 1930: 296 (Osorius; characters; Burma). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Myanmar. cordaticollis Cameron, 1942c: 839 (Osorius; Type locality: Bogor). DISTRIBUTION: Indonesia. cordicollis Fauvel, 1882: 58 (Osorius; Type locality: Sumatra: Alahan pandjang; Soengei- Boeloe). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Cameron, 1928d: 109 (Osorius; Sumatra). DISTRIBUTION: Indonesia. cordovensis Bernhauer, 1910b: 360 (Osorius; Type locality: Mexiko: Vera Cruz, Cordoba). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 7 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Mexico. cornutus Laporte, 1832, see: ater Perty, 1830. costaricensis Bernhauer, 1942: 7 (Osorius; Type locality: Kostarika: Reventazon). DISTRIBUTION: Costa Rica. 1204 OSORIINAE: OSORIUS Bulletin AMNH crassus Sharp, 1887: 681 (Osorius; Type locality: Mexico, Cordova; Guatemala, San Geronimo). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 6 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Mexico, Guatemala. crebresculpturatus Scheerpeltz, 1957c: 227, 247 (Osorius; Type locality: Sumba: Mau Marru). DISTRIBUTION: Indonesia. crenifrons Bernhauer, 1917, see: Madecosorius. crenulatus Cameron, 1928, see: aspericeps Fauvel, 1905. crenulifrons Notman, 1925: 9, 18 (Osorius; Type locality: San Diego, Cuba). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Darlington, 1937: 289, 291 (Osorius; comparative characters; Cuba). — Blackwelder, 1943: 178 (Osorius; characters; Cuba). DISTRIBUTION: West Indies. cribrum Bernhauer, 1914c: 89 (Osorius; Type locality: Sumatra: Mte. Singalang). — Bernhauer, 1928: 9 (Osorius; Sumatra). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Cameron, 1933b: 342 (Osorius; British North Borneo). DISTRIBUTION: Borneo, Indonesia. cristobalicus Coiffait, 1981g: 305 (Osorius; Type locality: Ile Galapagos: Ile San Cristobal). DISTRIBUTION: Galapagos Islands. cubensis Blackwelder, 1943: 181 (Osorius; Type locality: Cuba: San Antonio de los Banos). DISTRIBUTION: Cuba. cylindricus Latreille, 1832: 86 (Osorius; Type locality: Mexico). — Erichson, 1840: 756 (Osorius; characters; Mexico). — Sharp, 1887: 680 (Osorius; Mexico). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 8 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Mexico. daemon Darlington, 1937: 293 (Osorius; Type locality: Haiti: northeastern foothills of the Massif de la Hotte, 2000–4000 ft.). — Blackwelder, 1943: 180 (Osorius; characters; Haiti). DISTRIBUTION: West Indies. daemonicus Bernhauer, 1942b: 222 (Osorius; Type locality: Mindanao: Surigas). DISTRIBUTION: Philippines. darjeelingensis Herman, 2001: 40 (Osorius; replacement name for fraternus Cameron, 1942). fraternus Cameron, 1942: 110 (Osorius; [preoccupied]; Type locality: Darjeeling: Gopaldhara, altitude 4720–6100 feet). — Herman, 2001: 40 (Osorius; synonym of darjeelingensis). DISTRIBUTION: India. darlingtoni Blackwelder, 1943: 177 (Osorius; Type locality: Cuba, south side of Pico Turquino, elevation 3000 to 5000 feet). DISTRIBUTION: West Indies. debilis Sharp, 1887: 681 (Osorius; Type locality: Panama, Volcan de Chiriqui 2000 to 4000 feet). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 9 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Panama. Bulletin AMNH OSORIINAE: OSORIUS 1205 decipiens Cameron, 1937a: 92 (Osorius; Type locality: New Guinea: Kokoda, alt. 1300 ft.). DISTRIBUTION: New Guinea. delamarei Cameron, 1948, see: Neosorius. densicollis Bernhauer, 1919: 360 (Osorius; Type locality: Luzon: Mt. Makiling). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Bernhauer, 1942b: 224 (Osorius; characters; Philippines). DISTRIBUTION: Philippines. dentatus Bernhauer, 1908d: 294 (Osorius; Type locality: Bolivia: Mapiri). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 6 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Bolivia. dentifrons Cameron, 1933b: 342 (Osorius; Type locality: British North Borneo: Kenokok, 3300 feet). — Cameron, 1936a: 35 (Osorius; Java). DISTRIBUTION: Borneo, Indonesia. diabolicus Bernhauer, 1942b: 219 (Osorius; Type locality: Nord-Palawan: Binaluan). DISTRIBUTION: Philippines. difficilis Notman, 1925: 8, 17 (Osorius; Type locality: Chiricahua Mountains, Ariz.). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: USA. discalis Cameron, 1936a: 36 (Osorius; Type locality: Java: Tjilatjap). DISTRIBUTION: Indonesia. discicollis Fauvel, 1879: 78 (Osorius; Type locality: Nouvelle Guinée, Fly River). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). DISTRIBUTION: New Guinea. diversicollis Cameron, 1928g: 436, 438 (Osorius; Type locality: Borneo: Baram district). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Borneo. dividepunctatus Scheerpeltz, 1974: 85 (Osorius; Type locality: Cape Prov.: Tzitzikama Forest, Stormrivierpiek). DISTRIBUTION: South Africa. dohertyi Cameron, 1942c: 839 (Osorius; Type locality: Batjan: Labuan). DISTRIBUTION: Indonesia. dubius Sharp, 1887: 679 (Osorius; Type locality: Panama, Volcan de Chiriqui 2500 to 4000 feet). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 11 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Paulian, 1941: 177 (Osorius; larval characters; Costa Rica). DISTRIBUTION: Panama, Costa Rica. dundoensis Cameron, 1950, see: Neosorius. eggersi Bernhauer, 1904: 19 (Osorius; Type locality: Insula St. Thomas). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 6, 22 (Osorius; characters in key; Panama; Jamaica; Cuba). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Darlington, 1937: 298 (Osorius; notes; St. Thomas; Jamaica; Cuba; Haiti; Panama). — Blackwelder, 1943: 196 (Osorius; characters; Cuba; Jamaica; Haiti; Puerto Rico; St. Thomas). DISTRIBUTION: West Indies, Panama. 1206 OSORIINAE: OSORIUS Bulletin AMNH eppelsheimi Bernhauer, 1904: 18 (Osorius; Type locality: Sunda Inseln: Mentawei). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Cameron, 1928d: 109 (Osorius; Sumatra). — Bernhauer, 1928: 9 (Osorius; Sumatra). DISTRIBUTION: Indonesia. eucrates Fernando, 1960: 240 (Osorius; Type locality: University of Colombo, Ceylon). DISTRIBUTION: Sri Lanka. exalatus Blackwelder, 1943: 200 (Osorius; Type locality: Trinidad, Aripo Valley, elevation 2000 feet). DISTRIBUTION: West Indies. exiguus Notman, 1925: 7, 16 (Osorius; Type locality: Cuyamas, Cuba). — Notman, 1925: 7, 22 (Osorius; [Note: Misidentified as micros according to Blackwelder, 1943: 199]; characters in key; Cuba). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Darlington, 1937: 300 (Osorius; Cuba). — Blackwelder, 1943: 199 (Osorius; characters; Cuba). DISTRIBUTION: West Indies. fauveli Cameron, 1913, see: Mimogonus. fissicollis Bernhauer, 1942b: 230 (Osorius; Type locality: Luzon: Los Banos). DISTRIBUTION: Philippines. formosae Bernhauer, 1922c: 226 (Osorius; Type locality: Suisharyo, Kosempo). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Cameron, 1949b: 462 (Osorius; Formosa). — Sawada, 1965a: 147 (Osorius; Formosa). — Shibata, 1994a: 103 (Osorius; characters; Taiwan). DISTRIBUTION: Taiwan. franckei Wendeler, 1955: 197 (Osorius; Type locality: Brasilien: Nova Teutonia, 500 m). DISTRIBUTION: Brazil. fratellus Darlington, 1937: 291 (Osorius; Type locality: Haiti: Furcy, La Selle Range, probably taken at about 5000 ft.). — Blackwelder, 1943: 178 (Osorius; characters; Haiti). DISTRIBUTION: West Indies. frater Lynch, 1884: 345 (Osorius; Type locality: Buenos Aires). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 6 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Bernhauer, 1933e: 332 (Osorius; characters in key; Argentina). — Seevers, 1957: 318 (Osorius; doubtfully termitophilous). DISTRIBUTION: Argentina. fraterculus Cameron, 1936a: 36 (Osorius; Type locality: Java: Batoerraden, G. Slamat). DISTRIBUTION: Indonesia. fraternus Cameron, 1937a: 91 (Osorius; Type locality: New Guinea: Kokoda, alt. 1300 ft.). DISTRIBUTION: New Guinea. fraternus Cameron, 1942, see: darjeelingensis Herman, 2001. freyi Bernhauer, 1939k: 587 (Osorius; Type locality: N.W. China: Tienmuschan). DISTRIBUTION: China. frontalis Fauvel, 1905a: 134 (Osorius; Type locality: Sumatra: Deli; Padang. Java: Buitenzorg; Kemanglen, Tegal). — Fauvel, 1905e: 80 (Osorius; Java; Sumatra). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Eichelbaum, 1913: 122 (Osorius; Tanzania). Bulletin AMNH OSORIINAE: OSORIUS 1207
— Cameron, 1928d: 109 (Osorius; Sumatra). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Cameron, 1936a: 35 (Osorius; Java). DISTRIBUTION: Indonesia, Tanzania. fumator Fauvel, 1889, see: Mimogonus. furcifer Bernhauer, 1928: 9 (Osorius; Type locality: Sumatra: Mte. Singalang; Sibajakvulkan). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Indonesia. ganglbaueri Bernhauer, 1942b: 228 (Osorius; Type locality: Luzon: Imugan). DISTRIBUTION: Philippines. gardneri Cameron, 1930: 456 (Osorius; Type locality: Darjeeling: Ghum, alt. 7400 feet). — Scheerpeltz, 1933: 1133 (Osorius; catalog). — Abdullah and Qadri, 1970: 118 (Osorius; cited as gradneri; characters; Pakistan). DISTRIBUTION: India, Pakistan. germanus Sharp, 1887: 680 (Osorius; Type locality: Guatemala, near the city, Capetillo, Dueñas). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 8 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Guatemala. gigantulus Cameron, 1928, see: borneensis Scheerpeltz, 1933. gigantulus Cameron, 1928, see: Afrosorius. glaberrimus Bernhauer, 1926f: 314 (Osorius; Type locality: Buru). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Indonesia. granulatus Bernhauer, 1920a: 137 (Osorius; Type locality: Brasilien: Prov. Rio de Janeiro, Colonia alpina). — Notman, 1925: 10 (Osorius; characters in key). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Brazil. granulicollis Cameron, 1928f: 404 (Osorius; Type locality: Mt. Poi, 5000 feet). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Borneo. granulifrons Cameron, 1924, see: puncticollis Kraatz, 1859. guiananus Bernhauer, 1934e: 157 (Osorius; Type locality: Brit. Guiana: Essequibo R., Moraballi Creek). DISTRIBUTION: Guyana. haitiellus Darlington, 1937: 299 (Osorius; Type locality: Haiti: northeastern foothills of the Massif de la Hotte, 2000–4000 ft.). — Blackwelder, 1943: 191 (Osorius; characters; Cuba; Haiti). DISTRIBUTION: West Indies. hatamensis Fauvel, 1878f: 211 (Osorius; Type locality: Nouvelle-Guinée, Hatam). — Fauvel, 1879: 78 (Osorius; Amberbaki). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: New Guinea. hauseri Bernhauer, 1940b: 629 (Osorius; Type locality: Fokien, S. O. China). DISTRIBUTION: China. hewitti Bernhauer, 1914c: 89 (Osorius; Type locality: Sarawak: Quop.). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Malaysia. 1208 OSORIINAE: OSORIUS Bulletin AMNH hirsutus Cameron, 1928d: 109 (Osorius; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1133 (Osorius; catalog). DISTRIBUTION: Indonesia. hirtilabris Darlington, 1937: 292 (Osorius; Type locality: Haiti: northeastern foothills of the Massif de la Hotte, 2000–4000 ft.). — Blackwelder, 1943: 180 (Osorius; characters; Haiti). DISTRIBUTION: West Indies. hirtulus Fauvel, 1891: 92 (Osorius; Type locality: Colonie Tovar). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 6 (Osorius; characters in key). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Venezuela. hirtus Fauvel, 1878f: 211 (Osorius; Type locality: Nouvelle-Guinée, Ramoi). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). DISTRIBUTION: New Guinea. hubbardi Notman, 1925: 6, 11 (Osorius; Type locality: Jamaica, W.I.). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Darlington, 1937: 297 (Osorius; Jamaica). — Blackwelder, 1943: 193 (Osorius; characters; Jamaica). DISTRIBUTION: West Indies. impennis Fauvel, 1904, see: Madecosorius. impressiceps Cameron, 1941c: 493 (Osorius; Type locality: Philippines: Siargao: Dapa). — Herman, 2001: 22 (Osorius; synonymy). DISTRIBUTION: Philippines. impressiceps Bernhauer, 1942b: 222 (Osorius; [preoccupied]; Type locality: Siargao: Dapa; Mt. Banahao; Mindanao: Kolambugan). — Herman, 2001: 22 (Osorius; synonym of impressiceps). impressiceps Bernhauer, 1942, see: impressiceps Cameron, 1941. imuganensis Bernhauer, 1942b: 225 (Osorius; Type locality: Luzon: Imugan; N. Viskaya: Imugin). DISTRIBUTION: Philippines. incisicrurus Latreille, 1832, see: Madecosorius. incisifrons Fauvel, 1905, see: Neosorius. indicus Cameron, 1914: 528 (Osorius; Type locality: S. India: Nilgiri Hills). — Cameron, 1930: 304 (Osorius; characters; India). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: India. inexpectatus Filatova, 1978, see: taurus Sharp, 1889. infernus Bernhauer, 1942b: 219 (Osorius; Type locality: Luzon: Imugan). DISTRIBUTION: Philippines. insularum Bernhauer, 1942b: 231 (Osorius; Type locality: Mt. Banahao). DISTRIBUTION: Philippines. integer Sharp, 1876: 384 (Osorius; Type locality: Ega). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 10 (Osorius; characters in key). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Brazil. intermedius Cameron, 1928, see: cameroni Scheerpeltz, 1933. intermedius Erichson, 1840: 754 (Osorius; Type locality: Columbia interior). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 11, 24 (Osorius; characters in key; Costa Rica). Bulletin AMNH OSORIINAE: OSORIUS 1209
— Scheerpeltz, 1933: 1134 (Osorius; catalog). — Blackwelder, 1943: 192 (Osorius; characters; Trinidad; Venezuela; Mexico). — Frank, 1982: 8 (Osorius; list of parasites). DISTRIBUTION: West Indies, Venezuela, Colombia, Costa Rica, Mexico. interruptus Bernhauer, 1942b: 220 (Osorius; Type locality: Luzon: Mt. Makiling). DISTRIBUTION: Philippines. jamaicensis Blackwelder, 1943: 195 (Osorius; Type locality: Jamaica: Roaring River Falls, parish of St. Anns). DISTRIBUTION: West Indies. javanicus Bernhauer, 1915h: 215 (Osorius; Type locality: Java: Goenoeng Gedeh). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Indonesia. javanus Cameron, 1945a: 144 (Osorius; Type locality: W. Java: Sukabumi; E. Java: Mt. Tengger). DISTRIBUTION: Indonesia. jingkei Herman, 2001: 40 (Osorius; replacement name for chinensis J. Li). DISTRIBUTION: China. chinensis J. Li, 1993: 157 (Osorius; [preoccupied]; Type locality: Not cited but presumably China). — Herman, 2001: 40 (Osorius; synonym of jingkei). kakodanus Cameron, 1937a: 92 (Osorius; Type locality: New Guinea: Kokoda, alt. 1200 ft.). DISTRIBUTION: New Guinea. karnyi Cameron, 1925a: 176 (Osorius; Type locality: Tjibodas, 1400 m). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Indonesia. katompeanus Bernhauer, 1932, see: Neosorius. kedirianus Cameron, 1938b: 148 (Osorius; Type locality: E. Java: Res. Kediri, Pandan Aroem). DISTRIBUTION: Indonesia. kempi Bernhauer, 1926, see: Indosorius. kraatzi Bernhauer and Schubert, 1911, see: aspericeps Fauvel, 1905. kraatzi Cameron, 1925, see: aspericeps Fauvel, 1905. laeviceps Notman, 1925: 7, 15 (Osorius; Type locality: San Juan, Porto Rico). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Darlington, 1937: 301 (Osorius; Puerto Rico). — Blackwelder, 1943: 195 (Osorius; characters; Dominican Republic; Puerto Rico). DISTRIBUTION: West Indies. laevicollis Bernhauer, 1908d: 294 (Osorius; Type locality: Bolivia: Mapiri). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 10 (Osorius; characters in key). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Bolivia. laevigatulus Schubert, 1911: 4 (Osorius; Type locality: Lucena Parana, Brasil). — Notman, 1925: 9 (Osorius; characters in key). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Brazil. laevigatus Sharp, 1887: 679 (Osorius; Type locality: Guatemala, Sinanja and Sabo in Vera Paz). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 11, 24 (Osorius; characters in key; Guatemala). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Guatemala. 1210 OSORIINAE: OSORIUS Bulletin AMNH laevissimus Bernhauer, 1942b: 226 (Osorius; Type locality: Masbate: Aroroy). DISTRIBUTION: Philippines. laeviusculus Bernhauer, 1920b: 9 (Osorius; Type locality: Brasilien: Santa Catharina). — Notman, 1925: 8 (Osorius; characters in key). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Brazil. larutensis Herman, 2001: 40 (Osorius; replacement name for pendleburyi Cameron, 1950). DISTRIBUTION: Malaysia. pendleburyi Cameron, 1950d: 8 (Osorius; [preoccupied]; Type locality: Perak: Larut Hills, 3760–4500 feet). — Herman, 2001: 40 (Osorius; synonym of larutensis). latimargo Bernhauer, 1908d: 294 (Osorius; Type locality: Peru). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 9 (Osorius; characters in key). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Peru. latipes Gravenhorst, 1806: 198 (Oxytelus; Type locality: America septentrionalis). — Olivier, 1811: 616 (Oxytelus; characters; America septentrionale). — Say, 1831: 49 [= 1834: 462] (Molosoma; notes). — Erichson, 1840: 757 (Osorius; characters; America septentrionalis). — LeConte, 1877: 216 (Osorius; characters; middle and southern States; Kansas; Florida). — Blatchley, 1910: 473 (Osorius; characters; Indiana). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 8, 23 (Osorius; characters in key; South Dakota; Kansas; Iowa; Missouri; Arkansas; Texas; Alabama; Maryland; Virginia; North Carolina; Kentucky). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Darlington, 1937: 299 (Osorius; Cuba). — Blackwelder, 1943: 188 (Osorius; characters; Cuba). — Downie and Arnett, 1996: 429 (Osorius; characters; USA). DISTRIBUTION: USA, West Indies. leonensis Cameron, 1948, see: Allosorius. lewisi Blackwelder, 1943: 197 (Osorius; Type locality: Grand Cayman, west end of George- town). — Blackwelder, 1947: 121 (Osorius; Cayman Islands). DISTRIBUTION: West Indies. linearis Bernhauer, 1915, see: Osoriellus. lopchuensis Cameron, 1930: 457 (Osorius; Type locality: Darjeeling: Lopchu, alt. 5000 feet; Debrepanni; Simkona; Ranjirum, alt. 6000 feet). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Paulian, 1941: 178 (Osorius; larval characters; India). — Abdullah and Qadri, 1970: 118 (Osorius; characters; East Pakistan). DISTRIBUTION: India, Bangladesh. lucifer Bernhauer, 1942b: 217 (Osorius; Type locality: Luzon: Mt. Polis). DISTRIBUTION: Philippines. luzonicus Bernhauer, 1915e: 118 (Osorius; Type locality: Luzon: Laguna: Mount Maquiling). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Philippines, Malaysia. sarawakensis Cameron, 1941a: 206 (Osorius; subspecies of luzonicus; Type locality: W. Sarawak: Mt. Matang). luzonicus Bernhauer, 1942, see: banyosensis Herman, 2001. lynchi Bernhauer, 1933e: 333 (Osorius; Type locality: Argentinien: Prov. Buenos Aires). DISTRIBUTION: Argentina. Bulletin AMNH OSORIINAE: OSORIUS 1211 madurensis Schubert, 1911: 5 (Osorius; Type locality: Schembagamur, Madura). — Cameron, 1930: 302 (Osorius; characters; India). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: India. malabarensis Cameron, 1936a: 35 (Osorius; Type locality: Java: G. Malabar; G. Patoeha, alt. 5000 ft.; G. Tangkoeban Prahoe, alt. 4000–5000 ft.; G. Goentoer). DISTRIBUTION: Indonesia. malaisei Scheerpeltz, 1965: 164 (Osorius; Type locality: N.E. Burma, Kambaiti, 2000 m). DISTRIBUTION: Myanmar. malayanus Cameron, 1936b: 41 (Osorius; Type locality: Pahang: Cameron Highlands, 4000–5000 feet). DISTRIBUTION: Malaysia. mangpuensis Camerom, 1942, see: Indosorius. manni Notman, 1925: 9, 19 (Osorius; Type locality: Pinares Oriente, Cuba). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Darlington, 1937: 294 (Osorius; Cuba). — Blackwelder, 1943: 183 (Osorius; characters; Cuba). DISTRIBUTION: West Indies. maquilinganus Bernhauer, 1915e: 117 (Osorius; Type locality: Luzon: Laguna, Mount Maquiling). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Bernhauer, 1942b: 221 (Osorius; cited as makilinganus; characters; Philippines). DISTRIBUTION: Philippines. marshalli Bernhauer, 1936, see: Afrosorius. masuriensis Cameron, 1942: 109 (Osorius; Type locality: Mussoorie: Mossy Falls). DISTRIBUTION: India. mediocris Darlington, 1937: 296 (Osorius; Type locality: Haiti: La Visite and Vicinity, La Selle Range, 5000–7000 ft.). — Blackwelder, 1943: 185 (Osorius; characters; Haiti). DISTRIBUTION: West Indies. medius Bernhauer, 1928: 9 (Osorius; Type locality: Sumatra; Sinabong). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Indonesia. mephistopheles Cameron, 1941c: 493 (Osorius; Type locality: Philippines: Luzon: Mt. Polis). — Herman, 2001: 22 (Osorius; synonymy). DISTRIBUTION: Philippines. mephistopheles Bernhauer, 1942b: 218 (Osorius; [preoccupied]; Type locality: Luzon: Heightsplan, Mt. Polis). — Herman, 2001: 22 (Osorius; synonym of mephistopheles). mephistopheles Bernhauer, 1942, see: mephistopheles Cameron, 1941. mexicanus Bernhauer, 1910b: 360 (Osorius; Type locality: Mexiko: Oaxaca, Mitla). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 8 (Osorius; characters in key). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Mexico. microphthalmus Cameron, 1941c: 496 (Osorius; Type locality: Philippines: Pinutan). DISTRIBUTION: Philippines. microps Sharp, 1889, see: Mimogonus. micropterus Blackwelder, 1943: 201 (Osorius; Type locality: Trinidad, Ortocre River, 5 miles southeast of Rio Claro). DISTRIBUTION: West Indies. 1212 OSORIINAE: OSORIUS Bulletin AMNH micros Sharp, 1887: 682 (Osorius; Type locality: Guatemala, Pantaleon; Honduras; Panama, San Feliz). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Darlington, 1937: 301 (Osorius; Cuba). DISTRIBUTION: Guatemala, Honduras, Panama, West Indies. minor Notman, 1925: 7, 14 (Osorius; Type locality: Montserrat, Trinidad, West Indies). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Blackwelder, 1943: 190 (Osorius; characters; St. Vincent; Grenada; Trinidad). DISTRIBUTION: West Indies. minutior Bernhauer, 1942b: 226 (Osorius; Type locality: Luzon: Mt. Makiling). DISTRIBUTION: Philippines. minutissimus Bernhauer, 1914c: 90 (Osorius; Type locality: Sarawak). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: Malaysia. minutus Bernhauer, 1915e: 119 (Osorius; Type locality: Luzon: Laguna: Mount Maquiling). — Scheerpeltz, 1933: 1134 (Osorius; catalog). — Bernhauer, 1942b: 227 (Osorius; characters; Buru; Borneo; Penang). DISTRIBUTION: Philippines, Indonesia, Borneo, Malaysia. mjobergi Bernhauer, 1928: 10 (Osorius; cited as mjöbergi; [Note: Since this name is based on the Swedish name Mjöberg, the umlaut should be dropped and without further modification and the name should be emended to mjobergi]; Type locality: Sumatra: Sinabong, 2000 m., Brastagi). — Scheerpeltz, 1933: 1134 (Osorius; cited as mjöbergi; catalog). — Herman, 2001: 12 (Osorius; spelling corrected to mjobergi). DISTRIBUTION: Indonesia. mjobergi Cameron, 1928, see: banksi Scheerpeltz, 1933. montanus Cameron, 1937a: 91 (Osorius; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft.). DISTRIBUTION: New Guinea. monticola Cameron, 1914: 527 (Osorius; Type locality: S. India: Nilgiri Hills). — Cameron, 1930: 301 (Osorius; characters; India). — Scheerpeltz, 1933: 1134 (Osorius; catalog). DISTRIBUTION: India. morio Notman, 1925: 10, 21 (Osorius; Type locality: Omealca, Vera Cruz, Mexico). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Mexico. mortuorum Bernhauer, 1922c: 226 (Osorius; Type locality: Suisharyo). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Cameron, 1949b: 462 (Osorius; Formosa). DISTRIBUTION: Taiwan. mujechiensis Cho, 1998: 99 (Osorius; Type locality: Korea: Mujechi Moor, in Mt. Choˇngjoksan, Ulsan). — Cho and Ahn, 1999: 176, 179 (Osorius; characters in key; Korea). DISTRIBUTION: Korea. mundus Sharp, 1876b: 432 (Osorius; Type locality: Mexico City). — Sharp, 1887: 681 (Osorius; characters; Mexico). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 8, 22 (Osorius; characters in key; Mexico). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Mexico. murecarius Bernhauer, 1917, see: Madecosorius. Bulletin AMNH OSORIINAE: OSORIUS 1213 natalensis Scheerpeltz, 1974: 86 (Osorius; Type locality: Natal: Royal Natal National Park, Tugela Valley). DISTRIBUTION: South Africa. nefandus Bernhauer, 1942b: 221 (Osorius; Type locality: Luzon: Lamao). DISTRIBUTION: Philippines. neotropicus Bernhauer, 1908d: 295 (Osorius; Type locality: Brasilien: Rio Grande do Sul). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 9 (Osorius; characters in key). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Bernhauer, 1933e: 333 (Osorius; characters in key; Argentina). DISTRIBUTION: Argentina, Brazil. niasianus Cameron, 1945a: 144 (Osorius; Type locality: Central Nias: Lahago). DISTRIBUTION: Indonesia. nilgiriensis Fauvel, 1903, see: Indosorius. nitens Sharp, 1876: 382 (Osorius; Type locality: St. Paulo). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 10, 24 (Osorius; characters in key; Bolivia). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Brazil, Bolivia. notmani Darlington, 1937: 297 (Osorius; Type locality: Jamaica: Main Range Blue Mts., near Blue Mt. Peak, 5000–7388 ft.). — Blackwelder, 1943: 194 (Osorius; characters; Jamaica). DISTRIBUTION: West Indies. oculatus Sharp, 1876: 386 (Osorius; Type locality: Ega). — Bernhauer and Schubert, 1911: 144 (Osorius; catalog). — Notman, 1925: 7 (Osorius; characters in key). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Brazil. ogloblini Bernhauer, 1933e: 332 (Osorius; Type locality: Argentinien: Misiones, Loreto). DISTRIBUTION: Argentina. opacifrons Sharp, 1887: 680 (Osorius; Type locality: Mexico). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 9 (Osorius; characters in key). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Mexico. opacipennis Bernhauer, 1942b: 229 (Osorius; Type locality: Island Samar). DISTRIBUTION: Philippines. opaculus Eppelsheim, 1895, see: Neosorius. oriente Blackwelder, 1943: 187 (Osorius; Type locality: Cuba, mountains north of Imias, in eastern Oriente Province, elevation 3000 to 4000 feet). DISTRIBUTION: West Indies. overlaeti Bernhauer, 1933, see: Afrosorius. palawanensis Bernhauer, 1942b: 225 (Osorius; Type locality: Palawan: Binaluan). DISTRIBUTION: Philippines. paradoxifrons Bernhauer, 1932, see: Neosorius. paradoxus Bernhauer, 1927, see: Allosorius. parciventris Bernhauer, 1942b: 230 (Osorius; Type locality: Luzon: Imugan). DISTRIBUTION: Philippines. 1214 OSORIINAE: OSORIUS Bulletin AMNH parcus Sharp, 1887: 680 (Osorius; Type locality: Mexico, Cordova; Guatemala, Panzos, Cubilquitz, and San Juan in Vera Paz; Honduras; Costa Rica, Irazu 7000 feet; Panama, Volcan de Chiriqui 2500 to 4000 feet). — Fauvel, 1891: 92 (Osorius; Venezuela). — Fauvel, 1901: 72 (Osorius; Colombia; Mexico; Guatemala; Honduras; Costa Rica; Panama; Venezuela; Saint Vincent; Grenada; Ecuador; Paraguay). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 7 (Osorius; characters in key). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Moore and Legner, 1975: 174 (Osorius; catalog; Louisiana). DISTRIBUTION: USA, West Indies, Mexico, Guatemala, Honduras, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Paraguay. parviceps Notman, 1925: 6, 12 (Osorius; Type locality: Crescent City, Fla.). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: USA. parvipennis Bernhauer, 1926, see: Indosorius. parvulus Scriba, 1855: 300 (Osorius; Type locality: Venezuela). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 9 (Osorius; characters in key). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Venezuela. pectinifrons Fauvel, 1895: 196 (Osorius; Type locality: Bírmanie, Carin Ghécu, 1300– 1400 m). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Cameron, 1928c: 559 (Osorius; Darjeeling). — Cameron, 1930: 298 (Osorius; characters; Burma; Siam). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Myanmar, Thailand. pequanus Bernhauer, 1914, see: Indosorius. pendleburyi Cameron, 1945a: 145 (Osorius; Type locality: Perak: Larut Hills, altitude 3700–4500 feet). DISTRIBUTION: Malaysia. pendleburyi Cameron, 1950, see: larutensis Herman, 2001. peninsularis Cameron, 1950d: 7 (Osorius; Type locality: Pahang: Ginting Kiak, 5000 feet). DISTRIBUTION: Malaysia. peraffinis Cameron, 1942: 109 (Osorius; Type locality: Darjeeling: Gopaldhara, altitude 4720–6100 feet). DISTRIBUTION: India. peruvianus Bernhauer, 1908d: 293 (Osorius; Type locality: Peru). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 9 (Osorius; characters in key). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Peru. philippinus Bernhauer, 1915e: 118 (Osorius; Type locality: Luzon: Laguna, Mount Maquiling). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Bernhauer, 1942b: 230 (Osorius; characters; Philippines). DISTRIBUTION: Philippines. piceus Erichson, 1840: 755 (Osorius; Type locality: Brasilia). — Blanchard, 1842: 88 (Osorius; Argentina). — Lynch, 1884: 349 (Osorius; characters; Argentina). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). Bulletin AMNH OSORIINAE: OSORIUS 1215
— Notman, 1925: 10 (Osorius; characters in key). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Bernhauer, 1933e: 332 (Osorius; characters in key; Argentina). DISTRIBUTION: Brazil, Argentina. picipes Cameron, 1937a: 91 (Osorius; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft.). DISTRIBUTION: New Guinea. pilosus Fauvel, 1878f: 212 (Osorius; Type locality: Nouvelle-Guinée, Hatam; Ile Yule). — Fauvel, 1879: 78 (Osorius; New Guinea). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). DISTRIBUTION: New Guinea. planifrons LeConte, 1877: 215 (Osorius; Type locality: Southern States). — Blatchley, 1910: 473 (Osorius; characters; Indiana). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 7, 22 (Osorius; characters in key; Tennessee; Louisiana; Florida; Arizona; Jalapa, Mexico). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Smith, Lanzaro, Wheeler, and Snyder, 1978 (Osorius; habitat; feeding; natural history). — Downie and Arnett, 1996: 429 (Osorius; characters; USA). DISTRIBUTION: USA, Mexico. planifrons Cameron, 1928, see: cameronianus Scheerpeltz, 1933. politus LeConte, 1877: 215 (Osorius; Type locality: Hilsboro, Florida). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1920b: 698 (Osorius; Florida). — Notman, 1925: 6, 22 (Osorius; characters in key; Florida). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Paulian, 1941: 176, 326 (Osorius; larval and pupal characters; Costa Rica). DISTRIBUTION: USA, Costa Rica. propinquus Bernhauer, 1920a: 138 (Osorius; Type locality: Brasilien: Espirito Santo). — Notman, 1925: 11 (Osorius; characters in key). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Brazil. proximus Cameron, 1941c: 495 (Osorius; Type locality: Philippines: Luzon: Laguna). DISTRIBUTION: Philippines. puncticeps Sharp, 1887: 678 (Osorius; Type locality: Mexico, Orizaba, Jalapa, Cerro de Plumas, Tuxtla, Cordova). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 10, 24 (Osorius; characters in key; Mexico). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: Mexico. puncticollis Kraatz, 1859: 168 (Osorius; Type locality: Indiae orientalis pars borealis). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Cameron, 1930: 294 (Osorius; characters; India). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Scheerpeltz, 1965: 101 (Osorius; Burma). — Abdullah and Qadri, 1970: 116 (Osorius; characters; West Pakistan). — Scheerpeltz, 1976a: 81 (Osorius; Nepal). DISTRIBUTION: Pakistan, Nepal, India, Myanmar. granulifrons Cameron, 1924b: 178 (Osorius; Type locality: Simla Hills; Gahan, 7000 feet). — Cameron, 1930: 294 (Osorius; synonym of puncticollis). — Scheerpeltz, 1933: 1135 (Osorius; synonym of puncticollis). punctifrons Bernhauer, 1917, see: Madecosorius. punctipennis Fauvel, 1895, see: Indosorius. 1216 OSORIINAE: OSORIUS Bulletin AMNH punctiventris Cameron, 1950d: 8 (Osorius; Type locality: Selangor: The Gap). DISTRIBUTION: Malaysia. punctulatus Motschulsky, 1857a: 508 (Osorius; Type locality: Indes orientales). — Kraatz, 1859: 168 (Osorius). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Cameron, 1928d: 109 (Osorius; Sumatra). — Cameron, 1930: 302 (Osorius; characters; East Indies). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Cameron, 1934d: 77 (Osorius; types examined). — Cameron, 1940: 87 (Osorius; characters). DISTRIBUTION: “East Indies”, Indonesia. punctus Bernhauer, 1922c: 226 (Osorius; Type locality: Suisharyo, Fuhosho). — Scheerpeltz, 1933: 1135 (Osorius; catalog). — Shibata, 1994a: 103 (Osorius; characters; Taiwan). DISTRIBUTION: Taiwan. pygmaeus Laporte, 1835: 130 (Osorius; Type locality: Cayenne). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: French Guiana. regularis Fauvel, 1905, see: Afrosorius. regulus Darlington, 1937: 288 (Osorius; Type locality: Southwestern Haiti: Mt. La Hotte, 7000 ft to summit). — Blackwelder, 1943: 176 (Osorius; characters; Haiti). DISTRIBUTION: West Indies. rivularis Cameron, 1950, see: Osoriellus. robustus Cameron, 1920a: 145 (Osorius; Type locality: Sidapur, Coorg). — Cameron, 1930: 297 (Osorius; characters; India). — Scheerpeltz, 1933: 1135 (Osorius; catalog). DISTRIBUTION: India. rubripennis Bernhauer, 1942b: 224 (Osorius; Type locality: Mindanao: Butuan, Panaon, Pinutan). DISTRIBUTION: Philippines. rufipennis Motschulsky, 1857, see: Mimogonus. rufipes Motschulsky, 1857, see: Neosorius. rugegensis Fagel, 1957, see: Neosorius. rugiceps Boheman, 1848, see: Neosorius. rugiceps Kraatz, 1859, see: aspericeps Fauvel, 1905. rugicollis Kraatz, 1859: 167 (Osorius; Type locality: Ceylan). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Eichelbaum, 1913: 121 (Osorius; Tanzania). — Cameron, 1930: 292 (Osorius; characters; Ceylon, India). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Sri Lanka, India, Tanzania. rugifrons Erichson, 1840: 756 (Osorius; Type locality: Java). — Kraatz, 1859: 166 (Osorius; characters). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Cameron, 1921a: 368, 401 (Osorius; characters; Singapore). — Cameron, 1928d: 109 (Osorius; Sumatra). — Cameron, 1928f: 404 (Osorius; Borneo). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Cameron, 1936a: 35 (Osorius; Java). DISTRIBUTION: Borneo, Indonesia, Singapore. Bulletin AMNH OSORIINAE: OSORIUS 1217 rugipennis Bernhauer, 1910b: 361 (Osorius; Type locality: Mexiko: Orizaba). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 6 (Osorius; characters in key). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Mexico. rugiventris Cameron, 1945a: 145 (Osorius; Type locality: Sumatra: Fort de Kock). DISTRIBUTION: Indonesia. rugosipennis Bernhauer, 1942b: 224 (Osorius; [Note: This name was cited in a key as “rugosipennis Bernh.” but no citation to the original description was given. This is the only use of Osorius rugosipennis that I have found and the only other rugosipen- nis from the Philippines described by Bernhauer is in Lispinus]; Type locality: Luzon: Mt. Makiling). DISTRIBUTION: Philippines. sahlbergi Bernhauer, 1920a: 137 (Osorius; Type locality: Brasilien: Petropolis). — Notman, 1925: 10 (Osorius; characters in key). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Brazil. salutator Darlington, 1937: 290 (Osorius; Type locality: Haiti: La Visite and vicinity, La Selle Range, 5000–7000 ft.). — Blackwelder, 1943: 179 (Osorius; characters; Southeastern Haiti, La Viste and vicinity, La Selle Range, 5000 to 7000 feet elevation). DISTRIBUTION: West Indies. salvini Sharp, 1887: 682 (Osorius; Type locality: Guatemala, near the city 5000 feet, San Geronimo). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 7, 22 (Osorius; characters in key; Panama; Guatemala). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Guatemala, Panama. samarensis Bernhauer, 1942b: 223 (Osorius; Type locality: Island Samar). DISTRIBUTION: Philippines. samoensis Bernhauer, 1922a: 149 (Osorius; Type locality: Samoa: Apia). — Cameron, 1927a: 22 (Osorius; Samoa). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Samoa. sanguinipennis Fauvel, 1877: 192 (Osorius; Type locality: Australie, Gayndah). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Newton, 1990: 92 (Osorius). DISTRIBUTION: Australia. sarawakensis Cameron, 1941, see: luzonicus Bernhauer, 1915. satanas Bernhauer, 1942b: 220 (Osorius; Type locality: Siargao: Dapa). DISTRIBUTION: Philippines. scabricollis Bernhauer, 1939, see: Afrosorius. scheerpeltzi Bernhauer, 1942b: 229 (Osorius; Type locality: Leyte, Biliran). DISTRIBUTION: Philippines. schusteri Bernhauer, 1917, see: Madecosorius. schwarzi Notman, 1925: 7, 13 (Osorius; Type locality: Cayamas, Cuba). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Darlington, 1937: 299 (Osorius; Cuba). — Blackwelder, 1943: 191 (Osorius; characters; Cuba). DISTRIBUTION: West Indies. 1218 OSORIINAE: OSORIUS Bulletin AMNH scolytinus Fauvel, 1895: 195 (Osorius; Type locality: Bírmanie, Carin Ghécu, 1300–1400 m). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Cameron, 1930: 300 (Osorius; characters; Burma). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Watanabe and Shibata, 1976: 330 (Osorius; Malaysia). DISTRIBUTION: Myanmar, Malaysia. sechellarum H. Kolbe, 1910: 20 (Osorius; Type locality: Mahé, im gebirgigen Innern an waldigen Abhängen in 400–500 m Höhe). — Bernhauer, 1922b: 169 (Osorius; Seychelles). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Seychelles. secretus Cameron, 1950, see: Afrosorius. selangorensis Cameron, 1936b: 41 (Osorius; Type locality: Selangor: Kuala Lumpur, Ulu Klang). DISTRIBUTION: Malaysia. semicarinatus Bernhauer, 1927h: 293 (Osorius; Type locality: Neu-Guinea: Heuvel Bivak). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: New Guinea. sexpunctatus Bernhauer, 1912e: 168 (Osorius; Type locality: Argentinien: Buenos Aires). — Notman, 1925: 8 (Osorius; characters in key). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Bernhauer, 1933e: 333 (Osorius; characters in key; Argentina). — Frank, 1982: 8 (Osorius; list of parasites). DISTRIBUTION: Argentina. siamensis Cameron, 1940d: 249 (Osorius; Type locality: Indo-China: Laos; Pou Mi). DISTRIBUTION: Laos. sibuyanensis Bernhauer, 1942b: 225 (Osorius; Type locality: Island Sibuyan). DISTRIBUTION: Philippines. sikkimensis Bernhauer, 1920, see: Indosorius. silvestrii Bernhauer, 1927f: 261 (Osorius; Type locality: Yunnan: Lo-Chouli-Tong). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: China. simplex Sharp, 1876: 383 (Osorius; Type locality: Ega). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 10 (Osorius; characters in key). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Brazil. sinuatocollis Bernhauer, 1920a: 139 (Osorius; Type locality: Brasilien). — Notman, 1925: 11 (Osorius; cited as sinuaticollis; characters in key). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Brazil. smithi: [Note: Bernhauer and Schubert, 1911: 145 and Blackwelder, 1944: 112 erroneously cited smithi Bernhauer (1910b: 361) in Osorius; they also cited (Bernhauer and Schubert, 1911: 149; Blackwelder, 1944: 112) it in Oxyporus where Bernhauer described it. socors Darlington, 1937: 294 (Osorius; Type locality: Central Cuba: Central Soledad, near Cienfuegos, Santa Clara). — Blackwelder, 1943: 183 (Osorius; characters; Cuba). DISTRIBUTION: West Indies. solidus Sharp, 1876: 384 (Osorius; Type locality: St. Paulo). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 8 (Osorius; characters in key). Bulletin AMNH OSORIINAE: OSORIUS 1219
— Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Brazil. sororculus Cameron, 1937a: 91 (Osorius; Type locality: New Guinea: Mafulu, alt. 4000 ft.). DISTRIBUTION: New Guinea. sparsiceps Bernhauer, 1938, see: Neosorius. sparsifrons Cameron, 1928g: 437, 438 (Osorius; Type locality: Borneo: Baram district). — Cameron, 1928f: 401 (Osorius; Borneo). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Cameron, 1933b: 341 (Osorius; British North Borneo). DISTRIBUTION: Borneo. sparsior Fauvel, 1904, see: Neosorius. sparsus Cameron, 1928d: 110 (Osorius; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Indonesia. spectabilis Bernhauer, 1942b: 227 (Osorius; Type locality: Mindanao: Momungan). DISTRIBUTION: Philippines. stevensi Cameron, 1942: 109 (Osorius; Type locality: Nepal-Sikkim Frontier: Tonglu, altitude 10,074 feet). DISTRIBUTION: India-Nepal. stipes Sharp, 1876: 382 (Osorius; Type locality: Para; Ega). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 10, 23 (Osorius; characters in key; Bolivia). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Brazil, Bolivia. strandi Bernhauer, 1942b: 222 (Osorius; Type locality: Siargao). DISTRIBUTION: Philippines. strangulatus Fauvel, 1904a: 47 (Osorius; Type locality: N. Kanara). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Cameron, 1930: 296 (Osorius; characters; India). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: India. striaticeps Steel, 1956a: 67 (Osorius; Type locality: Huon Golf: Sattelberg). DISTRIBUTION: New Guinea. striativentris Bernhauer, 1915h: 215 (Osorius; Type locality: Sumatra: Manna). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: Indonesia. stricticollis Fauvel, 1895, see: Indosorius. strictus Blackwelder, 1943: 182 (Osorius; Type locality: Cuba: mountains north of Imias, in eastern Oriente Province, elevation 3000 to 4000 feet). DISTRIBUTION: West Indies. strigiceps Lea, 1931: 365 (Osorius; Type locality: New Guinea: Finsch Haven). — Scheerpeltz, 1933: 1136 (Osorius; catalog). DISTRIBUTION: New Guinea. strigicollis Cameron, 1941a: 205 (Osorius; Type locality: W. Sarawak: Quop). DISTRIBUTION: Malaysia. strigifrons H. Kolbe, 1889, see: Afrosorius. strigiventris Bernhauer, 1914c: 88 (Osorius; Type locality: Philippinen: Los Banos). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Bernhauer, 1942b: 232 (Osorius; characters; Philippines). DISTRIBUTION: Philippines. 1220 OSORIINAE: OSORIUS Bulletin AMNH striola Fauvel, 1878, see: Allosorius. sublaevicollis Bernhauer, 1942b: 231 (Osorius; Type locality: Luzon: Los Banos). DISTRIBUTION: Philippines. sublaevis Bernhauer, 1920b: 9 (Osorius; Type locality: Paraguay). — Notman, 1925: 8 (Osorius; characters in key). — Scheerpeltz, 1933: 1136 (Osorius; catalog). — Bernhauer, 1933e: 332 (Osorius; characters in key; Argentina). DISTRIBUTION: Paraguay. subtilior Bernhauer, 1942, see: Afrosorius. sumatranus Cameron, 1945a: 145 (Osorius; Type locality: Sumatra: Wai Lima). DISTRIBUTION: Indonesia. sumatrensis Cameron, 1945a: 145 (Osorius; Type locality: Sumatra: Fort de Kock). DISTRIBUTION: Indonesia. sutteri Scheerpeltz, 1957c: 227, 246 (Osorius; Type locality: Sumba: Waimangura). DISTRIBUTION: Indonesia. syriacus Fauvel, 1875, see: Neosorius. taurus Sharp, 1889: 411 (Osorius; Type locality: Ceylon; central and south Japan). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Watanabe, 1961: 350 (Osorius; Izu Islands). — Nakane, 1963a: 84 (Osorius; characters; habitus photograph; Japan). — Watanabe and Shibata, 1972: 61 (Osorius; Yaku-shima Island). — Yuh, Paik, Kwon, and Lee, 1985: 229 (Osorius; Korea). — Naomi, 1986: 36 (Osorius; characters; Japan). — J. Li, 1993: 23 (Osorius; cited as tanrus; China). — Watanabe, 1996a: 10 (Osorius; cave collection; Japan). — Cho and Ahn, 1999: 178 (Osorius; characters; Korea). DISTRIBUTION: Sri Lanka, Japan, Korea, China, Russia. inexpectatus Filatova, 1978: 568 [= 1979: 389] (Osorius; subspecies of taurus; Type locality: Primorskii krai Spasskii r-n, s. Gribnoe). tenuistrigosus Bernhauer, 1942b: 228 (Osorius; Type locality: Luzon: Imugan, Manila). DISTRIBUTION: Philippines. testaceus Tottenham, 1953b: 485 (Osorius; [Note: Nomen nudum; species attributed to Cameron but I find no description]). tonkinensis Bernhauer, 1914c: 87 (Osorius; Type locality: Zentral-Tonkin: Chiem-Hoa). — Bernhauer, 1922c: 227 (Osorius; Formosa). — Scheerpeltz, 1933: 1137 (Osorius; catalog). — Cameron, 1949b: 462 (Osorius; Formosa). DISTRIBUTION: Vietnam, Taiwan. toxopei Cameron, 1936a: 36 (Osorius; Type locality: Java: Tjibodas, G. Gedeh, alt. 1400 m). DISTRIBUTION: Indonesia. trinitatis Blackwelder, 1943: 199 (Osorius; Type locality: Trinidad, 10 miles north of Arima, elevation 1800 feet). DISTRIBUTION: West Indies. truncatus Cameron, 1925a: 177 (Osorius; Type locality: Verlaten Eiland; Krakatau). — Scheerpeltz, 1933: 1137 (Osorius; catalog). DISTRIBUTION: Indonesia. truncorum Bernhauer, 1904, see: Afrosorius. tuberculifrons Cameron, 1945a: 145 (Osorius; Type locality: Java: Tjibodas). DISTRIBUTION: Indonesia. tuberifrons Bernhauer, 1942b: 219 (Osorius; Type locality: Basilan, N. W. Pamay). DISTRIBUTION: Philippines. Bulletin AMNH OSORIINAE: OSORIUS 1221 tuberiventris Bernhauer, 1926f: 313 (Osorius; Type locality: Buru). — Scheerpeltz, 1933: 1137 (Osorius; catalog). DISTRIBUTION: Indonesia. turquinus Blackwelder, 1943: 188 (Osorius; Type locality: Cuba, south side of Pico Turquino, elevation 3000 to 5000 feet). DISTRIBUTION: West Indies. unicornis Cameron, 1941c: 494 (Osorius; Type locality: Philippines: Luzon: Imugan). — Herman, 2001: 22 (Osorius; synonym of unicornis). DISTRIBUTION: Philippines. unicornis Bernhauer, 1942b: 216 (Osorius; [preoccupied]; Type locality: Luzon: Imugan). — Herman, 2001: 22 (Osorius; unicornis). unicornis Bernhauer, 1942, see: unicornis Cameron, 1941. unispinosus Bernhauer, 1942b: 216 (Osorius; Type locality: Luzon: Imugan, Mt. Polis; Palikian). DISTRIBUTION: Philippines. usambarae Bernhauer, 1915, see: Neosorius. variolatus Notman, 1925: 8, 17 (Osorius; Type locality: Tucson, Ariz.). — Scheerpeltz, 1933: 1137 (Osorius; catalog). DISTRIBUTION: USA. vexator Tottenham, 1953, see: Rhabdopsidius. vicinus Sharp, 1887: 678 (Osorius; Type locality: Panama, Bugaba, Volcan de Chiriqui 2000 to 4000 feet, Caldera). — Bernhauer and Schubert, 1911: 145 (Osorius; catalog). — Notman, 1925: 10, 24 (Osorius; characters in key; Costa Rica). — Scheerpeltz, 1933: 1137 (Osorius; catalog). DISTRIBUTION: Panama, Costa Rica. victoriae Oke, 1933: 102 (Osorius; Type locality: Australia: Victoria: Carrum, Ringwood, Lorne). — Newton, 1990: 92 (Osorius). DISTRIBUTION: Australia. wasmanni Bernhauer, 1920a: 138 (Osorius; Type locality: Brasilien: Prov. Rio de Janeiro, Colonia alpina). — Notman, 1925: 9 (Osorius; characters in key). — Scheerpeltz, 1933: 1137 (Osorius; catalog). DISTRIBUTION: Brazil. weberi Blackwelder, 1943: 198 (Osorius; Type locality: Trinidad, 12 miles north of Arima). DISTRIBUTION: West Indies. wendeleri Bernhauer, 1942b: 217 (Osorius; Type locality: Nord Palawan: Binaluan). DISTRIBUTION: Philippines.
Ouloglene [1 species; Neotropical Region] Ouloglene Notman, 1925: 2, 3 (species included: barberi). TYPE SPECIES: Ouloglene barberi Notman, fixed by original designation and monotypy. — Scheerpeltz, 1933: 1137 (world catalog supplement). — Blackwelder, 1944: 112 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 280 (type species: barberi). barberi Notman, 1925: 4 (Ouloglene; Type locality: Cacao, Trece Aguas, Alta V. Paz, Guatemala). — Scheerpeltz, 1933: 1137 (Ouloglene; catalog). DISTRIBUTION: Guatemala. 1222 OSORIINAE: PARATORCHUS Bulletin AMNH
Paratorchus [40 species; Australian Region] Paratorchus McColl, 1985: 355 (replacement name for Paratrochus McColl). TYPE SPECIES: Paratrochus hamatus McColl, fixed by original designation and objective synonymy with Paratrochus for which the type had already been fixed. — Klimaszewski, Newton, and Thayer, 1996: 150 (endemic to New Zealand). Paratrochus McColl, 1982: 31 (species included: foveatus, monstrosus, helmsi, arrowi, vagepunctus, bifurcatus, caecus, curvisetis, hermes, brevisetis, fiordensis, tardus, scapulifer, microphthalmus, homerensis, humilis, brevipennis, trivialis, decipiens, hamatus, retroflexus, tubifer, alifer, pubescens, tricarinatus, flexuosus, angulatus, anophthalmus, insuetus, bucinifer, parvulus, angustus, pelorensis, aculeatus, phase- olinus; preoccupied by Pilsbry, 1893, and Kittl, 1899). TYPE SPECIES: Paratrochus hamatus McColl, fixed by original designation. — McColl, 1982: 21 (key to species of New Zealand). — McColl, 1984: 31 (characters; revised key to species). — Drugmand, 1998a: 220 (list of types in l’Institut royal des Sciences naturelles de Belgique). aculeatus McColl, 1982: 69 (Paratrochus; Type locality: New Zealand: Kirk Memorial Park, Papakura, AK, 20 m). — McColl, 1984: 34 (Paratrochus; characters). — McColl, 1985: 355 (Paratorchus). DISTRIBUTION: New Zealand. alifer McColl, 1982: 59 (Paratrochus; Type locality: New Zealand: Omahuta Kauri Reserve, ND, 135 m). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 355 (Paratorchus). DISTRIBUTION: New Zealand. angulatus McColl, 1982: 62 (Paratrochus; Type locality: New Zealand: Pinchgut Road, Kaukapakapa, AK, 91 m). — McColl, 1984: 34 (Paratrochus; characters). — McColl, 1985: 355 (Paratorchus). DISTRIBUTION: New Zealand. angustus McColl, 1982: 67 (Paratrochus; Type locality: New Zealand: Fletcher’s Creek, PB1 site, BR, 230 m). — McColl, 1984: 34 (Paratrochus; characters). — McColl, 1985: 355 (Paratorchus). DISTRIBUTION: New Zealand. anophthalmus Fauvel, 1900c: 188 (Holotrochus; Type locality: Greymouth; [Note: See lecto- type designation by McColl, 1982]). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Fagel, 1955: 75 (Typhlholotrochus). — McColl, 1982: 63 (Paratrochus; lectotype designation; characters; New Zealand). — McColl, 1984: 29, 34 (Paratrochus; characters; New Zealand). — McColl, 1985: 355 (Paratorchus). DISTRIBUTION: New Zealand. arrowi Bernhauer, 1939f: 209 (Holotrochus; Type locality: New Zealand: South Island: Mt. Owen; [Note: See lectotype designation by McColl, 1982]). — McColl, 1982: 37 (Paratrochus; lectotype designation; characters; New Zealand). — McColl, 1984: 31 (Paratrochus; characters). — McColl, 1985: 355 (Paratorchus). DISTRIBUTION: New Zealand. bifurcatus McColl, 1982: 39 (Paratrochus; Type locality: New Zealand: Canaan, NN, 650 m). — McColl, 1984: 31 (Paratrochus; characters). — McColl, 1985: 355 (Paratorchus). DISTRIBUTION: New Zealand. Bulletin AMNH OSORIINAE: PARATORCHUS 1223 brevipennis Broun, 1893: 1034 (Holotrochus; Type locality: Mokohinou). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — McColl, 1982: 52 (Paratrochus; characters; New Zealand). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 355 (Paratorchus). — Kuschel, 1990: 26 (Paratorchus; endemic; flightless; habitat; New Zealand). DISTRIBUTION: New Zealand. gracilis Broun, 1910: 17 (Holotrochus; Type locality: Waitakerei Range). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — McColl, 1982: 52 (Paratrochus; synonym of brevipennis). brevisetis McColl, 1982: 44 (Paratrochus; Type locality: New Zealand: Lake Wahapo, WD, 54 m). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. bucinifer McColl, 1982: 65 (Paratrochus; Type locality: New Zealand: Kaitoke Waterworks, Bush Walk, WN, 200 m). — McColl, 1984: 34 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. caecus Broun, 1910: 17 (Holotrochus; Type locality: Maketu, Hanua Range). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — McColl, 1982: 40 (Paratrochus; characters; New Zealand). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. curvisetis McColl, 1982: 41 (Paratrochus; Type locality: New Zealand: Pouakai Range, Ahukawakawa Track, TK, 1006 m). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. decipiens McColl, 1982: 54 (Paratrochus; Type locality: New Zealand: Kauri Grove Track, Waitakere Range, AK, 122 m). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. falcifer McColl, 1984: 27 (Paratrochus; Type locality: New Zealand: Cross Road, Ballance Bridge [WN], 213 m). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. fiordensis McColl, 1982: 45 (Paratrochus; Type locality: New Zealand: Spey River, western Manapouri, FD, 180 m). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. flexuosus McColl, 1982: 62 (Paratrochus; Type locality: New Zealand: Puketi State Forest, ND). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. foveatus McColl, 1982: 32 (Paratrochus; Type locality: New Zealand: Lake Wahapo, WD). — McColl, 1984: 31 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. 1224 OSORIINAE: PARATORCHUS Bulletin AMNH gracilis Broun, 1910, see: brevipennis Broun, 1893. grandis McColl, 1984: 23 (Paratrochus; Type locality: New Zealand: Mount Greenland [WD], 228 m). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. hamatus McColl, 1982: 55 (Paratrochus; Type locality: New Zealand: Fletcher’s Creek, BR, 230 m). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. helmsi Fauvel, 1900c: 187 (Holotrochus; Type locality: Greymouth; [Note: See lectotype designation by McColl, 1982]). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Fagel, 1955: 75 (Typhlholotrochus). — McColl, 1982: 35 (Paratrochus; lectotype designation; characters; New Zealand). — McColl, 1984: 31 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. nitidorsis Broun, 1910: 17 (Holotrochus; Type locality: Greymouth). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — McColl, 1982: 35 (Paratrochus; synonym of helmsi). setigerus Broun, 1912: 401 (Holotrochus; Type locality: Greymouth). — Scheerpeltz, 1933: 1131 (Holotrochus; catalog). — McColl, 1982: 35 (Paratrochus; synonym of helmsi). hermes McColl, 1982: 42 (Paratrochus; Type locality: New Zealand: 15 km northwest of Karamea, NN, 15 m). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. homerensis McColl, 1982: 50 (Paratrochus; Type locality: New Zealand: Mackay Creek, Eglinton Valley, FD, 400 m). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. humilis McColl, 1982: 51 (Paratrochus; Type locality: New Zealand: Arthur’s Pass, NC?, 730 m). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. insuetus McColl, 1982: 64 (Paratrochus; Type locality: New Zealand: Destruction Gulley, Waitakere Range, AK, 31 m). — McColl, 1984: 34 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. laevigatus Broun, 1910, see: vagepunctus Fauvel, 1900. maritimus McColl, 1984: 25 (Paratrochus; Type locality: New Zealand: Pahautane Bay [BR], 46 m). — McColl, 1984: 31 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. microphthalmus Fauvel, 1900c: 188 (Holotrochus; Type locality: Nouvelle-Zélande, probablement de la region d’Auckland). Bulletin AMNH OSORIINAE: PARATORCHUS 1225
— Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — McColl, 1982: 48 (Paratrochus; characters; New Zealand). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. minutus McColl, 1984: 29 (Paratrochus; Type locality: Mahoenui, Gribbons Rd. [WO]). — McColl, 1984: 34 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. monstrosus Bernhauer, 1939f: 210 (Holotrochus; Type locality: New Zealand: South Island: Hump Range, near Invercargil; [Note: See lectotype designation by McColl, 1982]). — McColl, 1982: 33 (Paratrochus; lectotype designation; characters; New Zealand). — McColl, 1984: 31 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. nitidorsis Broun, 1910, see: helmsi Fauvel, 1900. parvulus McColl, 1982: 66 (Paratrochus; Type locality: New Zealand: Karamea Bluff, Sum- mit Scenic Reserve, NN, 420 m). — McColl, 1984: 34 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. pelorensis McColl, 1982: 68 (Paratrochus; Type locality: New Zealand: Pelorus Bridge, SD, 31 m). — McColl, 1984: 34 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. phaseolinus McColl, 1982: 70 (Paratrochus; Type locality: New Zealand: Mount Pirongia, WO, 460 m). — McColl, 1984: 34 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. pubescens McColl, 1982: 60 (Paratrochus; Type locality: New Zealand: 15 km northwest of Karamea, NN, 15 m). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. relictus McColl, 1984: 28 (Paratrochus; Type locality: New Zealand: Mangaweka, Main Trunc [RI]). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. retroflexus McColl, 1982: 57 (Paratrochus; Type locality: New Zealand: Lake Aorere, Mount Domett, NN, 1260 m). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. scapulifer McColl, 1982: 47 (Paratrochus; Type locality: New Zealand: Dawson Falls, Mount Egmont, TK, 900 m). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. setigerus Broun, 1910, see: helmsi Fauvel, 1900. 1226 OSORIINAE: PARATORCHUS Bulletin AMNH tardus McColl, 1982: 46 (Paratrochus; Type locality: New Zealand: Kaitoke Waterworks, Bush Walk, WN, 200 m). — McColl, 1984: 32 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. tricarinatus McColl, 1982: 61 (Paratrochus; Type locality: New Zealand: Mount Messenger, TK, 200 m). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. trivialis McColl, 1982: 54 (Paratrochus; Type locality: New Zealand: Pinchgut Road, Kaukapakapa, AK, 91 m). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. tubifer McColl, 1982: 58 (Paratrochus; Type locality: New Zealand: Mount Messenger, TK, 200 m). — McColl, 1984: 33 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. vagepunctus Fauvel, 1900c: 187 (Holotrochus; Type locality: Nouvelle-Zelande, probable- ment de la region d’Auckland; [Note: See lectotype designation by McColl, 1982]). — Bernhauer and Schubert, 1911: 143 (Holotrochus; catalog). — Fagel, 1955: 75 (Typhlholotrochus). — McColl, 1982: 38 (Paratrochus; lectotype designation; characters; New Zealand). — McColl, 1984: 31 (Paratrochus; characters). — McColl, 1985: 356 (Paratorchus). DISTRIBUTION: New Zealand. laevigatus Broun, 1910: 16 (Holotrochus; Type locality: Mount Pirongia). — Scheerpeltz, 1933: 1130 (Holotrochus; catalog). — McColl, 1982: 38 (Paratrochus; synonym of vagepunctus). Paratrochodes [1 species; Madagascan Region] Paratrochodes Dajoz, 1980: 127 (species included: microphtalmus). TYPE SPECIES: Para- trochodes microphtalmus Dajoz, fixed by original designation. microphtalmus Dajoz, 1980: 127 (Paratrochodes; Type locality: Madagascar Est: station 2089, chaînes Anosyennes, massif de l’Andohahelo, massif Sud, 1850 m). DISTRIBUTION: Madagascar. Parosorius [1 species; Oriental Region] Parosorius Bernhauer, 1904a: 222 (species included: forsteri). TYPE SPECIES: Ancaeus forsteri Bernhauer, fixed by original designation and monotypy. — Bernhauer and Schubert, 1911: 146 (world catalog; 1 species). — Notman, 1925: 2 (characters in key). — Scheerpeltz, 1933: 1137 (world catalog supplement). — Blackwelder, 1952: 295 (type species: foersteri). forsteri Bernhauer, 1903a: 36 (Ancaeus; Type locality: Sumatra). — Bernhauer, 1904a: 222 (Parosorius). — Bernhauer and Schubert, 1911: 146 (Parosorius; catalog). — Cameron, 1928d: 110 (Parosorius; Sumatra). — Scheerpeltz, 1933: 1137 (Parosorius; catalog). DISTRIBUTION: Indonesia. Bulletin AMNH OSORIINAE: RHABDOPSIDIUS 1227
Rhabdopsidius [1 species; Ethiopian Region] Rhabdopsidius Fagel, 1955: 117 (species included: vexator). TYPE SPECIES: Osorius vexator Tottenham, fixed by original designation and monotypy. vexator Tottenham, 1953b: 485 (Osorius; Type locality: Kilimanjaro: Western side, 6370 ft). — Fagel, 1955: 118 (Rhabdopsidius; notes). — Fagel, 1960a: 23 (Rhabdopsidius; Tanganyika). DISTRIBUTION: Tanzania.
Rhabdopsis [3 species; Ethiopian Region] Rhabdopsis Fagel, 1955: 111 (species included: africana, leleupi). TYPE SPECIES: Rhabdopsis africana Fagel, fixed by original designation. — Fagel, 1969: 70 (characters; notes; key to species). — Drugmand, 1998a: 223 (list of types in l’Institut royal des Sciences naturelles de Belgique). africana Fagel, 1955: 113 (Rhabdopsis; Type locality: Congo Belge: Kivu: territoire d’Uvira: Haute Sange, 2900 m). — Fagel, 1969: 70 (Rhabdopsis; characters). DISTRIBUTION: Congo. kaboboensis Fagel, 1969: 71 (Rhabdopsis; Type locality: Nord-Katanga: territoire d’Albertville, Kalemie, mont Kabobo, Haute Kiymbi, 18450 m). DISTRIBUTION: Congo. leleupi Fagel, 1955: 115 (Rhabdopsis; Type locality: Congo Belge: Kivu: territoire de Lubero, 2200 m). — Fagel, 1969: 70 (Rhabdopsis; characters). DISTRIBUTION: Congo.
Saegerius [6 species; Ethiopian Region] Saegerius Fagel, 1959: 18 (species included: congoensis, microps, subterraneus, kivuensis, garambanus). TYPE SPECIES: Saegerius garambanus Fagel, fixed by original desig- nation. — Naomi, 1986: 34 (characters). — Naomi, 1987c: 14 (characters; species of Japan). — Newton, 1990: 92 (characters in key; one undescribed Australian species). — Drugmand, 1998a: 223 (list of types in l’Institut royal des Sciences naturelles de Belgique). congoensis Fagel, 1955: 24 (Mimogonus; Type locality: Congo Belge: Basoko). — Fagel, 1959: 19 (Saegerius; characters). DISTRIBUTION: Congo. gabonicus Coiffait, 1967: 275 (Saegerius; Type locality: Bélinga). DISTRIBUTION: Gabon. garambanus Fagel, 1959: 19 (Saegerius; Type locality: Congo Belge: Parc National de la Garamba). DISTRIBUTION: Congo. japonicus Naomi, 1987, see: Mimogonellus. kivuensis Fagel, 1959: 22 (Saegerius; Type locality: Congo Belge: Kibu: territoire de Shabunda, Kiamiseke). — Fagel, 1961: 274 (Saegerius; characters; notes; Ghana). DISTRIBUTION: Congo, Ghana. microps Fagel, 1959: 21 (Saegerius; Type locality: Congo Belge: Kibali-Ituri: territoire d’Epulu, Mambasa-Mungbere, 900 m). DISTRIBUTION: Congo. 1228 OSORIINAE: SAEGERIUS Bulletin AMNH subterraneus Fagel, 1955: 25 (Mimogonus; Type locality: Congo Belge: Bas-Congo: territoire des Cataractes, Kavuya). — Fagel, 1959: 19 (Saegerius; characters). DISTRIBUTION: Congo. yasutoshii Naomi, 1987, see: Mimogonellus.
Tavakilianidia [1 species; Neotropical Region] Tavakilianidia Orousset, 1985a: 60 (species included: anophtalma). TYPE SPECIES: Tavakilia- nidia anophtalma Orousset, fixed by original designation and monotypy. anophtalma Orousset, 1985a: 61 (Tavakilianidia; Type locality: Guyane française: Cacao, alt. 200 m). DISTRIBUTION: French Guiana. Tetrosorius [1 species; Madagascan Region] Tetrosorius Dajoz, 1980: 123 (species included: anosyensis). TYPE SPECIES: Tetrosorius anosyensis Dajoz, fixed by original designation. anosyensis Dajoz, 1980: 123 (Tetrosorius; Type locality: Madagascar Est, station 2087, chaînes Anosyennes, massif de l’Andohahelo, massif Sud, 1880 m). DISTRIBUTION: Madagascar. Thoracogonus [2 species; Madagascan Region] Thoracogonus Coiffait, 1979: 23 (species included: peyrierasi, caecus). TYPE SPECIES: Tho- racogonus peyrierasi Coiffait, fixed by original designation. caecus Coiffait, 1979: 25 (Thoracogonus; Type locality: archipel des Séchelles, Mahé, station des Mornes Seychellois, 900 m). DISTRIBUTION: Seychelles. peyrierasi Coiffait, 1979: 24 (Thoracogonus; Type locality: archipel des Séchelles, Mahé, station des Mornes Seychellois, 900 m). DISTRIBUTION: Seychelles. Thoracoprius [8 species; Oriental Region] Thoracoprius Bernhauer, 1914c: 90 (species included: vulneratus). TYPE SPECIES: Thora- coprius vulneratus Bernhauer, fixed by monotypy. — Notman, 1925: 2 (characters in key). — Scheerpeltz, 1933: 1130 (world catalog supplement). — Blackwelder, 1952: 388 (type species: vulneratus). — Hammond, 1984: 199 (checklist of species of Borneo). denticollis Cameron, 1936a: 34 (Thoracoprius, cited as Thoracoproprius; Type locality: Java: G. Tangkoeban Prahoe, Preanger). DISTRIBUTION: Indonesia. heterocerus Cameron, 1929b: 446 (Thoracoprius, cited as Thoracoproprius; Type locality: Selangor: The Gap). — Scheerpeltz, 1933: 1130 (Thoracoprius; catalog). DISTRIBUTION: Malaysia. laetus Cameron, 1950d: 6 (Thoracoprius; Type locality: Selangor: Bukit Kutu). DISTRIBUTION: Malaysia. longipennis Cameron, 1941a: 204 (Thoracoprius; Type locality: W. Sarawak: Mt. Matang). DISTRIBUTION: Malaysia. Bulletin AMNH OSORIINAE: THORACOPRIUS 1229 major Cameron, 1950d: 5 (Thoracoprius; Type locality: Kuala Lumpur). DISTRIBUTION: Malaysia. malayanus Cameron, 1950d: 5 (Thoracoprius; Type locality: Pahang: Cameron Highlands, 3500 feet). DISTRIBUTION: Malaysia. niger Cameron, 1941a: 204 (Thoracoprius; Type locality: W. Sarawak: Mt. Matang). DISTRIBUTION: Malaysia. vulneratus Bernhauer, 1914c: 91 (Thoracoprius; Type locality: Sarawak). — Scheerpeltz, 1933: 1130 (Thoracoprius; catalog). DISTRIBUTION: Malaysia.
Tumboecus [1 species; Ethiopian Region] Tumboecus Fagel, 1959: 62 (species included: nitidus). TYPE SPECIES: Tumboecus nitidus Fagel, fixed by original designation and monotypy. nitidus Fagel, 1959: 63 (Tumboecus; Type locality: Congo Belge: Tshuapa: territoire de Bikoro, Mabali, lac Tumba). DISTRIBUTION: Congo.
Typhlholotrochus [1 species; Ethiopian Region] Typhlholotrochus Fagel, 1955: 74 (species included: curtipennis, anophthalmus, vagepunctus, helmsi). TYPE SPECIES: Typhlholotrochus curtipennis Fagel, fixed by original desig- nation. — Drugmand, 1998a: 224 (list of types in l’Institut royal des Sciences naturelles de Belgique). curtipennis Fagel, 1955: 75 (Typhlholotrochus; Type locality: Congo Belge: Kivu: territoire de Mwenga, Sud-Ouest Itombwe, Luiko, 2050 m). DISTRIBUTION: Congo.
Typhlobledius [1 species; Australian Region] Typhlobledius Lea, 1906: 200 (species included: cylindricus). TYPE SPECIES: Typhlobledius cylindricus Lea, fixed by monotypy. — Bernhauer and Schubert, 1911: 137 (world catalog; 1 species). — Blackwelder, 1952: 400 (type species: cylindricus). — Herman, 1970: 398 (notes). — Newton, 1983 (characters; transferred to Osoriinae from Oxytelinae). — Newton, 1990: 92 (characters in key; one described and three undescribed Australian species). cylindricus Lea, 1906: 200 (Typhlobledius; Type locality: Hobart, Tasm.). — Bernhauer and Schubert, 1911: 137 (Typhlobledius; catalog). — Newton, 1990: 92 (Typhlobledius). DISTRIBUTION: Australia.
Typhloiulopsis [3 species; Palaearctic Region] Typhloiulopsis Scheerpeltz, 1931: 370 (species included: rudiana, leucadiae). TYPE SPECIES: Typhloiulopsis rudiana Scheerpeltz, fixed by original designation. — Scheerpeltz, 1933: 1128 (world catalog supplement). — Blackwelder, 1952: 401 (type species: rudiana). — Coiffait, 1955b: 263 (characters in key). — Coiffait, 1959b: 216 (characters). — Orousset, 1985c: 181 (characters). 1230 OSORIINAE: TYPHLOIULOPSIS Bulletin AMNH
Numidotyphlus Coiffait, 1959b: 215 (species included: normandi). TYPE SPECIES: Numido- typhlus normandi Coiffait, fixed by original designation and monotypy. — Orousset, 1985c: 181 (synonym of Typhloiulopsis). leucadiae Scheerpeltz, 1931: 375 (Typhloiulopsis; Type locality: Kaligoni auf Levkas). — Scheerpeltz, 1933: 1128 (Typhloiulopsis; catalog). DISTRIBUTION: Greece. normandi Coiffait, 1959b: 216 (Numidotyphlus, cited as Numidopsis; Type locality: Tunisie: environs du Kef). — Coiffait, 1961: 82 (Numidotyphlus; correction of erroneous spelling of original generic combi- nation). DISTRIBUTION: Tunisia. rudiana Scheerpeltz, 1931: 374 (Typhloiulopsis; Type locality: Kephallinia im Gebiete des Bergmassivs des Rudi). — Scheerpeltz, 1933: 1128 (Typhloiulopsis; catalog). DISTRIBUTION: Greece.
Typhlosorius [3 species; Palaearctic Region] Typhlosorius Coiffait, 1958b: 111 (subgenus of Cylindropsis; species included: torressalai, lucidus). TYPE SPECIES: Cylindropsis (Typhlosorius) torressalai Coiffait, fixed by original designation. — Coiffait, 1973b: 223 (valid genus; characters). ibizensis Coiffait, 1973b: 223 (Typhlosorius; Type locality: Iles Baléares, Ibiza, San Anto- nion). — Pons and Palmer, 1996: 165 (Typhlosorius; Balearic endemic; notes; habitat; Eivissa). DISTRIBUTION: Balearic Islands. lucidus Coiffait, 1958b: 114 (Cylindropsis; subgenus Typhlosorius; Type locality: Espagne: Province de Casteillon de la Plana). — Coiffait, 1973b: 223 (Typhlosorius). DISTRIBUTION: Spain. torressalai Coiffait, 1958b: 112 (Cylindropsis; subgenus Typhlosorius; Type locality: Espagne: Pego). — Coiffait, 1973b: 223 (Typhlosorius). DISTRIBUTION: Spain.
Witelsus [1 species; Ethiopian Region] Witelsus Fagel, 1969: 65 (species included: spiniventris). TYPE SPECIES: Witelsus spiniventris Fagel, fixed by original designation and monotypy. spiniventris Fagel, 1969: 66 (Witelsus; Type locality: Cape Province: Humansdorp district, Wit Els Bos). DISTRIBUTION: South Africa.
Tribe Thoracophorini Thoracophorini Reitter, 1909. — Newton, 1990: 86 (characters). — Newton and Thayer, 1992: 59 (tribe of Osoriinae; list of subtribes). — Newton, Thayer, Ashe, and Chandler, 2000: 377 (characters; notes). Thoracophorinae Reitter, 1909: 199 (genera included: Thoracophorus). TYPE GENUS: Thoracophorus (see Newton and Thayer, 1992: 59). — Tóth, 1982: 13 (characters). — Portevin, 1929: 459. Bulletin AMNH OSORIINAE: THORACOPHORINI 1231
Thoracophori Reitter, 1909. — Bernhauer and Schubert, 1910: 24 (world catalog; Espeson, Bothrys, Diplopsis, Rhopalopherus, Thoracophorus, Calocerus, Aneucamptus). — Scheerpeltz, 1933: 1019 (world catalog supplement). — Blackwelder, 1942: 79 (characters). — Blackwelder, 1944: 108 (checklist of species from Mexico, Central and South America, and the West Indies). — Scheerpeltz, 1968a: 2 (catalog of Austrian species).
Subtribe Clavilispinina Clavilispinina Newton and Thayer, 1992: 59 (replacement name for Paralispini Blackwelder; subtribe of Thoracophorini). TYPE GENUS: Clavilispinus. — Newton, Thayer, Ashe, and Chandler, 2000: 377 (characters; notes). Paralispini Blackwelder, 1942: 77, 79 (genera included: Ischiopsaurus, Lispinodes, Neolispinus, Paralispinus). TYPE GENUS: Paralispinus (see Newton and Thayer, 1992: 59). — Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies). — Newton and Thayer, 1992: 59 (based on preoccupied type genus; synonym of Clavilispinina). Paralispinina Blackwelder, 1942. — Arnett, 1963: 263 (characters; list of North American genera).
Allotrochus [2 species; Ethiopian and Madagascan Regions] Allotrochus Fagel, 1955: 78 (species included: curticollis, myllaenoides). TYPE SPECIES: Holotrochus curticollis Fauvel, fixed by original designation. — Coiffait, 1979: 62 (characters; species of Madagascar). — Newton, 1990: 92 (characters in key; one undescribed Australian species). curticollis Fauvel, 1898a: 115 (Holotrochus; Type locality: La Digue, Seychelles; [Note: See lectotype designation by Coiffait, 1979]. Also cited from Madagascar: Diego Suarez; Tamatave). — H. Kolbe, 1910: 20 (Holotrochus; Seychelles). — Bernhauer and Schubert, 1911: 142 (Holotrochus; catalog). — Eichelbaum, 1913: 122 (Holotrochus; Tanzania). — Fagel, 1955: 79 (Allotrochus; Seychelles; Madagascar). — Fagel, 1959: 53 (Allotrochus; Belgian Congo). — Coiffait, 1979: 64 (Allotrochus; lectotype designation: La Digue, Seychelles; characters; Madagascar). DISTRIBUTION: Seychelles, Madagascar, Tanzania, Congo. sparsus Cameron, 1928: 21 (Holotrochus; Type locality: Belgian Congo: Urundi: Kama). — Scheerpeltz, 1933: 1131 (Holotrochus; catalog). — Fagel, 1959: 32, 53 (Allotrochus; synonym of curticollis). ferrugineus Cameron, 1938, see: myllaenoides Cameron, 1950. myllaenoides Cameron, 1950f: 117 (Holotrochus; Type locality: Angola: Dundo: Chiumbe River). — Fagel, 1955: 80 (Allotrochus; notes; Belgian Congo). — Fagel, 1959: 55 (Allotrochus; notes). DISTRIBUTION: Angola, Congo. ferrugineus Cameron, 1938: 5 (Holosus; [preoccupied]; Type locality: Belgian Congo: Eala). — Fagel, 1959: 32, 55 (Allotrochus; synonym of myllaenoides). sparsus Cameron, 1928, see: curticollis Fauvel, 1898. 1232 OSORIINAE: CLAVILISPINUS Bulletin AMNH
Clavilispinus [29 species; Neotropical, Nearctic, Palaearctic, Oriental, Ethiopian, Madagascan, Australian, and Oceanic Regions] Clavilispinus Bernhauer, 1926d: 255 (subgenus of Paralispinus; species included: siargaoanus). TYPE SPECIES: Paralispinus (Clavilispinus) siargaoanus Bernhauer, fixed by monotypy. — Scheerpeltz, 1933: 1009 (world catalog supplement; subgenus of Paralispinus). — Blackwelder, 1952: 102 (type species: siargaoanus). — Moore and Legner, 1974b: 78 (characters; notes). — Moore and Legner, 1975: 166 (catalog; North America north of Mexico). — Moore and Legner, 1979: 144 (characters; notes). — Hammond, 1984: 196 (checklist of species of Borneo). — Frank, 1986a: 365 (checklist of species of Florida). — Irmler, 1991: 85 (key to Neotropical species). — Campbell and Davies, 1991: 92 (checklist of species; Canada). — Newton, 1990: 92 (characters in key; one described Australian species). — Naomi, 1996: 51 (note on use of name in Japan). — Downie and Arnett, 1996: 427 (characters; key to species of Northeastern North America). — Lundgren, 1998: 46 (list of species from Florida; records for other States listed for each species). — Drugmand, 1998a: 207 (list of types in l’Institut royal des Sciences naturelles de Belgique). — Newton, Thayer, Ashe, and Chandler, 2000: 377 (6 Nearctic species; notes; characters in key). Ancaeus Fauvel, 1864b: 56 [= 1865: 60] (species included: megacephalus; preoccupied by Agassiz, 1846, and Adams, 1861). TYPE SPECIES: Ancaeus megacephalus Fauvel, fixed by monotypy. — Gemminger and Harold, 1868: 678 (catalog). — LeConte, 1877: 249 (characters; notes). — Duvivier, 1883: 205 (catalog). — Sharp, 1887: 716 (notes; Mexican and Central American species). — Bernhauer and Schubert, 1910: 19 (world catalog; 11 species). — Leng, 1920: 93 (catalog of North American species). — Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog). — Cameron, 1930: 42 (synonym of Paralispinus). — Scheerpeltz, 1933: 1009 (synonym of Paralispinus). — Blackwelder, 1943: 156 (synonym of Paralispinus; type species magacephalus). — Blackwelder, 1952: 51 (synonym of Neolispinodes; type species: Ancaeus). — Drugmand, 1998a: 206 (list of types in l’Institut royal des Sciences naturelles de Belgique). Paralispinus Bernhauer, 1921b: 67 (preoccupied by Eichelbaum, 1913; replacement name for Ancaeus Fauvel). TYPE SPECIES: Ancaeus megacephalus Fauvel, fixed by objec- tive synonymy with Ancaeus. — Cameron, 1925: 6 (catalog of Indian species). — Cameron, 1930: 42 (characters). — Scheerpeltz, 1933: 1009 (world catalog supplement). — Blackwelder, 1942: 85 (notes). — Blackwelder, 1943: 156 (characters; type species: megacephalus; key to species of West Indies). — Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 291 (synonym of Neolispinodes; type species). — Hatch, 1957: 244 (characters; key to species of Pacific Northwest). — Shibata, 1973c: 21 (checklist of species of Taiwan). — Shibata, 1976: 81 (checklist of species of Japan). Neolispinodes Bernhauer, 1937b: 579 (replacement name for Paralispinus Bernhauer). TYPE SPECIES: Ancaeus megacephalus Fauvel, fixed by objective synonymy with Ancaeus. Bulletin AMNH OSORIINAE: CLAVILISPINUS 1233
— Blackwelder, 1952: 259 (subgenus of Clavilispinus; type species). — Scheerpeltz, 1956a: 506 (key to African species). aruensis Fauvel, 1878, see: exiguus Erichson, 1840. bisulcatus Coiffait, 1978b: 137 (Paralispinus; Type locality: Phuntsholing, 2400 m). DISTRIBUTION: Bhutan. californicus LeConte, 1863a: 59 (Lispinus; Type locality: California, San Jose). — LeConte, 1877: 250 (Ancaeus; characters; California). — Fauvel, 1878d: 178 [= 1878e: 14] (Lispinus; characters; California). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Hatch, 1957: 245 (Paralispinus; characters; Oregon). — Legner and Moore, 1977: 174 (Clavilispinus; found under bark). DISTRIBUTION: USA. canaliculatus Scheerpeltz, 1956a: 482, 508, 512 (Neolispinodes; Type locality: Belgisch- Kongo: Yangambi). DISTRIBUTION: Congo. circumductus Fauvel, 1902: 26 (Ancaeus; Type locality: Nouvelle-Guinee: baie de l’Astrolabe, Stephansort; Nouvelle Bretagne: île Deslacs). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Cameron, 1928d: 96 (Paralispinus; Sumatra). DISTRIBUTION: New Guinea, New Britain, Indonesia. clavicornis Scheerpeltz, 1956a: 507 (Neolispinodes; Type locality: Franz. Kongo: Umg. Brazzaville). DISTRIBUTION: Congo Republic. clavipes Sharp, 1876: 391 (Holotrochus; Type locality: Amazons). — Sharp, 1887: 718 (Ancaeus; Guatemala). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Irmler, 1991: 88 (Clavilispinus; characters; Brazil). DISTRIBUTION: Brazil, Guatemala. collarti Cameron, 1933: 35 (Paralispinus; Type locality: Belgian Congo: Mayumbe: Batasalia). — Scheerpeltz, 1956a: 481, 507 (Neolispinodes; characters; French Congo; Belgian Congo). DISTRIBUTION: Congo, French Equatorial Africa. cordicollis Fauvel, 1904e: 296 (Ancaeus; Type locality: Madagascar: Fort Dauphin). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). DISTRIBUTION: Madagascar. crassicornis Sharp, 1887: 716 (Ancaeus; Type locality: Guatemala, Cubilguitz 1000 feet). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Irmler, 1991: 88 (Clavilispinus; characters; Guatemala). DISTRIBUTION: Guatemala. crepusculus Blackwelder, 1943, see: megacephalus Fauvel, 1865. exiguus Erichson, 1840: 830 (Lispinus; Type locality: Puerto Rico). — Fauvel, 1864b: 55 [= 1865: 59] (Lispinus; characters; Puerto Rico; Mexico). — Solsky, 1872: 314 (Lispinus; Cayenne). — Fauvel, 1878d: 180 [= 1878e: 16] (Lispinus; characters; Alabama; Florida; Louisiana; Mexico; Puerto Rico). — Fleutiaux and Sallé, 1889: 382 (Ancaeus; Guadeloupe Island). — Fauvel, 1903d: 255 (Ancaeus; New Caledonia). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Blatchley, 1910: 484 (Lispinus; characters; Indiana). — Eichelbaum, 1913: 116 (Ancaeus; Tanzania). — Bernhauer, 1920: 3 (Ancaeus; Australia). — Cameron, 1921a: 363, 399 (Ancaeus; characters; Singapore). 1234 OSORIINAE: CLAVILISPINUS Bulletin AMNH
— Bernhauer, 1922b: 165 (Ancaeus; Seychelles). — Cameron, 1928d: 96 (Paralispinus; Sumatra). — Cameron, 1930: 42 (Paralispinus; characters; Assam). — Scheerpeltz, 1933: 1009 (Paralispinus; catalog). — Cameron, 1933f: 73 (Paralispinus; Marquesas Islands). — Blackwelder, 1942: 90 (Pseudolispinodes; subgenus Nacaeus; checklist). — Blackwelder, 1943: 158 (Paralispinus; characters; Cuba; Jamaica; Dominican Republic; Puerto Rico; St. Croix; Guadeloupe; St. Lucia; Grenada). — Cameron, 1949b: 457 (Paralispinus; Formosa). — Blackwelder, 1952: 256 (Nacaeus; subgenus Nacaeus; [Note: Blackwelder 1952: 256 renamed his misidentification of Pseudolispinodes as Nacaeus and listed in Nacaeus the subgenera he described in Pseudolispinodes in 1942; the species he listed in those subgenera therefore should be transferred to Nacaeus]). — Scheerpeltz, 1956a: 481, 506 (Neolispinodes; characters; Belgian Congo). — Miskimen and Bond, 1970: 82 (Paralispinus; St. Croix). — Scheerpeltz, 1971c: 162 (Neolispinodes; Cameroon). — Watanabe and Shibata, 1972: 59 (Paralispinus; Yaku-shima Island). — Shibata, 1974: 33 (Paralispinus; Japan). — Scheerpeltz, 1975a: 242 (Neolispinodes; collecting notes; Brazil). — Legner and Moore, 1977: 174 (Neolispinodes; found under bark). — Newton, 1990: 92 (Clavilispinus). — Irmler, 1991: 86 (Clavilispinus; characters; Colombia; Bolivia; Brazil; Guadeloupe; Guatemala; Cuba; Jamaica; Dominican Republic; Puerto Rico; St. Croix; St. Lucia; Grenada). — Nishida, 1994: 71 (Clavilispinus; Hawaii). — Naomi, 1996: 52 (Clavilispinus; Japan). — Downie and Arnett, 1996: 427 (Clavilispinus; characters; USA). — Nishida, 1997: 60 (Clavilispinus; Hawaii). DISTRIBUTION: USA, West Indies, Mexico, Guatemala, Colombia, French Guiana, Brazil, Bolivia, Marquesas Islands, Hawaii, Taiwan, Japan, Vanuatu, New Caledonia, Australia, New Guinea, Indonesia, Singapore, Myanmar, India, Sri Lanka, Mauritius, Seychelles, Zanzibar, Ethiopia, Tanzania, Congo, Cameroon, Gabon. laevigatus Kraatz, 1859: 188 (Lispinus; Type locality: Ceylan). — Fauvel, 1898a: 114 (Ancaeus; Seychelles). — Fauvel, 1901: 70 (Ancaeus; Colombia; Alabama; Florida; Mexico; Guatemala; Guadeloupe; Grenada; Amazonian region; Brazil; Hawaii; Ethopia; Zanzibar; Gabon; Ceylon; Burma; Suma- tra; Seychelles; Mauritius Islands; Moluccas; New Guinea; New Hebrides; New Caledonia). — Fauvel, 1903d: 255 (Ancaeus; synonym of exiguus). — Sharp, 1908: 539 (Ancaeus; Hawaii). — Bernhauer and Schubert, 1910: 19 (Ancaeus; synonym of exiguus). — H. Kolbe, 1910: 21 (Ancaeus; Seychelles). — Bernhauer, 1922b: 165 (Ancaeus; synonym of exiguus). — Cameron, 1930: 42 (Paralispinus; synonym of exiguus). — Blackwelder, 1943: 158 (Paralispinus; synonym of exiguus). — Irmler, 1991: 86 (Clavilispinus; synonym of exiguus). rufus Fauvel, 1864b: 56 [= 1865: 60] (Lispinus; [Note: Nomen nudum; this name was cited in a paragraph within Fauvel’s account of exiguus; although characters were presented, no indication or suggestion was made that a new taxon was intended, reference was being made to a specimen in a collection that Fauvel considered to be conspecific with exiguus. If Fauvel considered rufus to be conspecific with exiguus then the name was first published as a junior synonym and is an unavailable name (Article 11.6)]). rufescens LeConte, 1863a: 59 (Lispinus; Type locality: Alabama, Mobile). — LeConte, 1877: 249 (Ancaeus; characters; Florida; Alabama; Louisiana). — Fauvel, 1878d: 180 [= 1878e: 16] (Lispinus; synonym of exiguus). — Sharp, 1887: 718 (Ancaeus; notes; Mexico; Guatemala). Bulletin AMNH OSORIINAE: CLAVILISPINUS 1235
— Fauvel, 1903d: 255 (Ancaeus; synonym of exiguus). — Bernhauer and Schubert, 1910: 19 (Ancaeus; synonym of exiguus). — Cameron, 1930: 42 (Paralispinus; synonym of exiguus). — Blackwelder, 1943: 158 (Paralispinus; synonym of exiguus). — Irmler, 1991: 86 (Clavilispinus; synonym of exiguus). fauveli Sharp, 1876: 392 (Holotrochus; Type locality: Amazons). — Fauvel, 1901: 70 (Ancaeus; synonym of laevigatus). — Fauvel, 1903d: 255 (Ancaeus; synonym of exiguus). — Bernhauer and Schubert, 1910: 19 (Ancaeus; synonym of exiguus). — Cameron, 1930: 42 (Paralispinus; synonym of exiguus). — Blackwelder, 1943: 158 (Paralispinus; synonym of exiguus). — Irmler, 1991: 86 (Clavilispinus; synonym of exiguus). aruensis Fauvel, 1878f: 200 (Ancaeus; Type locality: Iles Arou, Wokan). — Fauvel, 1879: 75 (Ancaeus; notes; Gilolo; Dorei). — Fauvel, 1889b: 243 (Ancaeus; notes; New Caledonia). — Fauvel, 1901: 70 (Ancaeus; synonym of laevigatus). — Fauvel, 1903d: 255 (Ancaeus; synonym of exiguus). — Bernhauer and Schubert, 1910: 19 (Ancaeus; synonym of exiguus). — Cameron, 1930: 42 (Paralispinus; synonym of exiguus). — Blackwelder, 1943: 158 (Paralispinus; synonym of exiguus). — Irmler, 1991: 86 (Clavilispinus; synonym of exiguus). pallescens Blackburn, 1885: 126 (Lispinodes; Type locality: near Honolulu). — Blackburn and Sharp, 1885: 224 (Lispinodes; habitat; Hawaii). — Fauvel, 1901: 70 (Ancaeus; synonym of laevigatus). — Fauvel, 1903d: 256 (Ancaeus; synonym of exiguus). — Sharp, 1908: 539 (Ancaeus; synonym of laevigatus). — Bernhauer and Schubert, 1910: 19 (Ancaeus; synonym of exiguus). — Cameron, 1930: 42 (Paralispinus; synonym of exiguus). — Blackwelder, 1943: 158 (Paralispinus; synonym of exiguus). — Legner and Moore, 1977: 174 (Ancaeus; found under bark). — Irmler, 1991: 86 (Clavilispinus; synonym of exiguus). piceus Fauvel, 1902: 27 (Ancaeus; variety of exiguus; Type locality: Nouvelle Guinee: baie Huon, Sattelberg; baie de l’Astrolabe, Stephansort, Erima; Doreï). — Bernhauer and Schubert, 1910: 19 (Ancaeus; synonym of exiguus). obscuripennis Bernhauer, 1926d: 255 (Paralispinus; variety of exiguus; Type locality: Not cited). — Scheerpeltz, 1933: 1009 (Paralispinus; variety of exiguus). — Blackwelder, 1943: 158 (Paralispinus; synonym of exiguus). — Irmler, 1991: 86 (Clavilispinus; synonym of exiguus). fauveli Sharp, 1876, see: exiguus Erichson, 1840. foersteri Bernhauer, 1903, see: Parosorius. globiceps Cameron, 1927, see: nitidissimus Bernhauer, 1905. gracilis Scheerpeltz, 1956a: 481, 507, 508 (Neolispinodes; Type locality: Belgisch-Kongo: Yangambi. Also cited from French Congo). DISTRIBUTION: French Equatorial Africa, Congo. guadeloupensis Irmler, 1991: 89 (Clavilispinus; Type locality: Guadeloupe. Also cited from Brazil). DISTRIBUTION: West Indies, Brazil. incisus Fauvel, 1902: 26 (Ancaeus; Type locality: nord de la Nouvelle-Guinee). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Cameron, 1931: 356 (Paralispinus; New Guinea). DISTRIBUTION: New Guinea. 1236 OSORIINAE: CLAVILISPINUS Bulletin AMNH jeani Herman, 2001: 39 (Clavilispinus; replacement name for piceus Jarrige). DISTRIBUTION: Mascarene Islands. piceus Jarrige, 1957: 110 (Paralispinus; [preoccupied]; Type locality: Saint-Philippe, forêt du Brûlé de Mare-Longue). — Herman, 2001: 39 (Clavilispinus; synonym of jeani). laevigatus Kraatz, 1859, see: exiguus Erichson, 1840. leai Cameron, 1927c: 259 (Paralispinus; Type locality: Fiji: Taveuni). — Scheerpeltz, 1933: 1009 (Paralispinus; catalog). — Coiffait, 1976: 220 (Paralispinus; Tahiti). DISTRIBUTION: Fiji, Society Islands. mariae Hatch, 1957, see: Stictocranius. megacephalus Fauvel, 1864b: 57 [= 1865: 61] (Ancaeus; Type locality: Teapa, État de Tabasco). — Sharp, 1887: 717 (Ancaeus; notes; Mexico). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Irmler, 1991: 89 (Clavilispinus; characters; Paraguay; Venezuela; Brazil; Peru; Bolivia; Mexico; Guadeloupe; Trinidad). DISTRIBUTION: Mexico, West Indies, Venezuela, Peru, Bolivia, Brazil, Paraguay. crepusculus Blackwelder, 1943: 160 (Paralispinus; Type locality: St. Lucia: Duboulay Estate in St. Remy district, 3 miles southeast of Soufriere. Also cited from Cuba, Jamaica, Puerto Rico, and Trinidad). — Irmler, 1991: 89 (Clavilispinus; synonym of megacephalus). minutus Sharp, 1887: 717 (Ancaeus; Type locality: Guatemala, Panzos). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Marcuzzi, 1977: 4 (Neolispinodes; Venezuela). — Irmler, 1991: 88 (Clavilispinus; characters; Guatemala). DISTRIBUTION: Guatemala, Venezuela. nitidus Scheerpeltz, 1956a: 481, 507, 509 (Neolispinodes; Type locality: Luki. Also cited from French Congo and Belgian Congo). DISTRIBUTION: French Equatorial Africa, Congo. nitidissimus Bernhauer, 1905, see: Nacaeus. obscuripennis Bernhauer, 1926, see: exiguus Erichson, 1840. pallescens Blackburn, 1885, see: exiguus Erichson, 1840. paraguensis Irmler, 1991: 89 (Clavilispinus; Type locality: Paraguay). DISTRIBUTION: Paraguay. piceus Fauvel, 1902, see: exiguus Erichson, 1840. piceus Jarrige, 1957, see: jeani Herman, 2001. politus Sharp, 1887: 718 (Ancaeus; Type locality: Guatemala, El Tumbador). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Blackwelder, 1943: 157 (Paralispinus; characters; Guadeloupe; St. Lucia; St. Vincent). — Legner and Moore, 1977: 174 (Neolispinodes; found under bark). — Irmler, 1991: 86 (Clavilispinus; characters; Guadeloupe; Guatemala; Panama; Costa Rica; Mexico; Brazil; Peru; Cuba; St. Lucia; St. Vincent; Jamaica). DISTRIBUTION: West Indies, Mexico, Guatemala, Costa Rica, Peru, Brazil. rulomus Blackwelder, 1943: 160 (Paralispinus; Type locality: Jamaica: 12.5 miles from Kingston on road from Gordon Town to Newcastle, parish of St. Andrew). — Legner and Moore, 1977: 174 (Neolispinodes; found under bark). — Irmler, 1991: 86 (Clavilispinus; synonym of politus). porrectus Sharp, 1887: 717 (Ancaeus; Type locality: Guatemala, Panzos, and San Juan in Vera Paz). Bulletin AMNH OSORIINAE: CLAVILISPINUS 1237
— Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Irmler, 1991: 88 (Clavilispinus; characters; Guatemala; Paraguay). DISTRIBUTION: Guatemala, Paraguay. prolixus LeConte, 1877: 250 (Ancaeus; Type locality: New York). — Fauvel, 1878d: 178 [= 1878e: 14] (Lispinus; characters; New York). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Downie and Arnett, 1996: 427 (Clavilispinus; characters; USA). DISTRIBUTION: USA. punctatus Blackwelder, 1943, see: Nacaeus. pygmaeus Cameron, 1925b: 232 (Paralispinus; Type locality: Selangor, Federated Malay States; The Gap). — Scheerpeltz, 1933: 1009 (Paralispinus; catalog). DISTRIBUTION: Malaysia. rufescens LeConte, 1863, see: exiguus Erichson, 1840. rufescens Hatch, 1957: 245 (Paralispinus; [preoccupied]; Type locality: British Columbia: Creston). — Herman, 2001: 39 (Clavilispinus; junior primary homonym, but species will be moved to another genus). DISTRIBUTION: Canada. rufus Fauvel, 1864 [Note: Nomen nudum], see: exiguus Erichson, 1840. rulomus Blackwelder, 1943, see: politus Sharp, 1887. schedli Scheerpeltz, 1956a: 482, 508, 511 (Neolispinodes; Type locality: Belgisch-Kongo: Yangambi). DISTRIBUTION: Congo. siargaoanus Bernhauer, 1926d: 255 (Paralispinus; subgenus Clavilispinus; Type locality: Siargao, Dapa). — Scheerpeltz, 1933: 1009 (Paralispinus; catalog). DISTRIBUTION: Philippines. singularis Cameron, 1918: 58 (Ancaeus; Type locality: Singapore: Bukit Panjang and Bukit Timah). — Cameron, 1921a: 363 399 (Ancaeus; characters; Singapore). — Scheerpeltz, 1933: 1009 (Paralispinus; catalog). DISTRIBUTION: Singapore. sulcicollis Coiffait, 1976: 231 (Paralispinus; Type locality: Tahiti: vallée Papenoo). — Coiffait, 1977a: 171 (Paralispinus; Tahiti). DISTRIBUTION: Society Islands. vinsoni Jarrige, 1957: 111 (Paralispinus; Type locality: Réunion: Saint-Philippe, forêt du Brûlé de Mare-Longue). DISTRIBUTION: Mascarene Islands. Eulibia [1 species; Palaearctic Region] Eulibia Cameron, 1945: 66 (species included: albizziae). TYPE SPECIES: Eulibia albizziae Cameron, fixed by original designation and monotypy. — Blackwelder, 1952: 156 (type species: albizziae). — Herman, 1970: 425 (moved to Osoriinae). — Newton, 1990: 86 (discussion of tribal placement). albizziae Cameron, 1945: 67 (Eulibia; Type locality: United Provinces: Roorkee, Mohammadur). DISTRIBUTION: India. 1238 OSORIINAE: HETEROTROCHINUS Bulletin AMNH
Heterotrochinus [2 species; Madagascan and Oriental Regions] Heterotrochinus Coiffait, 1983b: 345 (replacement name for Heterotrochus Coiffait). TYPE SPECIES: Holotrochus lineatocollis Cameron, fixed by objective synonymy with Heterotrochus for which a type had been fixed. — Newton, 1990: 86 (discussion of tribal placement). Heterotrochus Coiffait, 1979: 64 (species included: lineatocollis; preoccupied by Haime and Milne-Edwards, 1857). TYPE SPECIES: Holotrochus lineatocollis Cameron, fixed by original designation and monotypy. lineatocollis Cameron, 1936d: 202 (Holotrochus; Type locality: Mauritius: Stanley). — Coiffait, 1978c: 141 (Holotrochidius). — Coiffait, 1979: 66 (Heterotrochus; characters; Mauritius). DISTRIBUTION: Mascarene Islands. osellai Coiffait, 1981f: 346 (Heterotrochus; Type locality: South Andaman: Cinque Island). DISTRIBUTION: Andaman Islands. Holotrochidius [1 species; Palaearctic Region] Holotrochidius Coiffait, 1978b: 141 (species included: holosiformis, lineatocollis). TYPE SPECIES: Holotrochidius holosiformis Coiffait, fixed by original designation. — Newton, 1990: 86 (discussion of tribal placement). holosiformis Coiffait, 1978b: 141 (Holotrochidius; Type locality: Samchi, 300 m). DISTRIBUTION: Bhutan. Hospitalibia [1 species; Oriental Region] Hospitalibia Kistner and Newton, 1999: 239 (species included: thailandica). TYPE SPECIES: Hospitalibia thailandica Kistner and Newton, fixed by original designation and monotypy. thailandica Kistner and Newton, 1999: 245 (Hospitalibia; Type locality: Thailand: Khao Yai National Park, Wung Jumpee, off road near km 36). DISTRIBUTION: Thailand. Ischiopsaurus [3 species; Madagascan and Oriental Regions] Ischiopsaurus Bernhauer, 1929i: 337 (species included: colossus, boettcheri, cordicollis). TYPE SPECIES: Lispinus colossus Bernhauer, fixed by original designation. — Scheerpeltz, 1933: 1012 (world catalog supplement). — Blackwelder, 1942: 88 (type species: boettcheri). — Blackwelder, 1952: 200 (type species: colossus). — Drugmand, 1998a: 212 (cited as Ichiopsaurus; list of types in l’Institut royal des Sciences naturelles de Belgique). boettcheri Bernhauer, 1929i: 338 (Ischiopsaurus; Type locality: Mindanao, Mumungan). — Scheerpeltz, 1933: 1012 (Ischiopsaurus; catalog). DISTRIBUTION: Philippines. colossus Bernhauer, 1902: 172 (Lispinus; Type locality: Port Daufin, Madagaskar). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Bernhauer, 1929i: 338 (Ischiopsaurus). — Scheerpeltz, 1933: 1012 (Ischiopsaurus; catalog). — Paulian, 1941: 141 (Lispinus; cited as colosseus; larval characters; Madagascar). — Scheerpeltz, 1961g: 234 (Ischiopsaurus; Madagascar). DISTRIBUTION: Madagascar. Bulletin AMNH OSORIINAE: ISCHIOPSAURUS 1239 cordicollis Bernhauer, 1929i: 338 (Ischiopsaurus; Type locality: Singapore). — Scheerpeltz, 1933: 1012 (Ischiopsaurus; catalog). DISTRIBUTION: Singapore. Myrmelibia [1 species; Australian Region] Myrmelibia Newton, 1990: 82 (species included: kingi; notes on generic placement). TYPE SPECIES: Glyptoma kingi Lea, fixed by original designation and monotypy. kingi Lea, 1910a: 142 (Glyptoma; Type locality: Sea Lake, Vic. N.S.W.; [Note: See lectotype designation by Newton, 1990]. Also cited from N. S. Wales: Liverpool; Victoria: Birchip). — Scheerpeltz, 1933: 1021 (Calocerus; catalog). — Newton, 1990: 87 (Myrmelibia; lectotype designation: Sea Lake, Vic. N.S.W.; characters; natural history notes; New South Wales; Queensland; Victoria). DISTRIBUTION: Australia. Neolispinus [1 species; Oriental Region] Neolispinus Cameron, 1929b: 439 (species included: crucifer). TYPE SPECIES: Neolispinus crucifer Cameron, fixed by original designation and monotypy. — Scheerpeltz, 1933: 1018 (world catalog supplement). — Blackwelder, 1952: 259 (type species: crucifer). crucifer Cameron, 1929b: 439 (Neolispinus; Type locality: Selangor: The Gap). — Scheerpeltz, 1933: 1018 (Neolispinus; catalog). DISTRIBUTION: Malaysia. Subtribe Glyptomina Glyptomina Newton and Thayer, 1992: 59 (replacement name for Caloceri; subtribe of Thoracophorini). TYPE GENUS: Glyptoma. Caloceri Blackwelder, 1942: 78 (genera included: none included with original description but a year later Blackwelder [1943: 120] included one genus, Glyptoma). TYPE GENUS: Calocerus (see Newton and Thayer, 1992: 59). — Newton and Thayer, 1992: 59 (based on preoccupied type genus; synonym of Glyptomina). — Newton, Thayer, Ashe, and Chandler, 2000: 377 (characters; notes). Espeson [18 species; Neotropical, Nearctic, Ethiopian, Madagascan, Oriental, and Australian Regions] Espeson Schaufuss, 1882b: 168 (species included: moratus). TYPE SPECIES: Espeson moratus Schaufuss, fixed by monotypy. — Schaufuss, 1882: 45 (characters). — Fauvel, 1902: 35 (notes). — Bernhauer and Schubert, 1910: 24 (world catalog; 8 species). — Bernhauer, 1926d: 261 (key to species of Philippines). — Scheerpeltz, 1933: 1019 (world catalog supplement). — Blackwelder, 1943: 144 (characters; type species: moratus; key to species of West Indies). — Blackwelder, 1944: 108 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 152 (type species: moratus). — Scheerpeltz, 1970: 116 (key to species). — Newton, 1990: 93 (characters in key; one undescribed Australian species). — Lundgren, 1998: 46 (unnamed, undescribed species listed from Florida). — Drugmand, 1998a: 209 (list of types in l’Institut royal des Sciences naturelles de Belgique). — Newton, Thayer, Ashe, and Chandler, 2000: 378 (2 undescribed Nearctic species; notes; char- acters in key). 1240 OSORIINAE: ESPESON Bulletin AMNH africanus Cameron, 1935a: 371 (Espeson; Type locality: Stanleyville). — Scheerpeltz, 1970: 126 (Espeson; characters; Zaire). DISTRIBUTION: Congo. angustiformis Cameron, 1950, see: Parespeson. angustissimus Bernhauer, 1926, see: Parespeson. confusus Cameron, 1946: 10 (Espeson; Type locality: Philippines: Mindanao: Momungan). — Scheerpeltz, 1970: 124 (Espeson; characters; Philippines). DISTRIBUTION: Philippines. crassulus Fauvel, 1902, see: Pseudespeson. cruzensis Coiffait, 1981g: 307 (Espeson; Type locality: Ile Galapagos: Ile Santa Cruz, bois au-dessus de Santa Rosa). DISTRIBUTION: Galapagos Islands. curtus Bernhauer, 1926, see: Pseudespeson. dolichurus Lecoq, 1995: 419 (Espeson; Type locality: Cameroun: Edea). DISTRIBUTION: Cameroon. euplectoides Fauvel, 1902: 37 (Espeson; Type locality: Guadeloupe). — Bernhauer and Schubert, 1910: 24 (Espeson; catalog). — Blackwelder, 1943: 145 (Espeson; characters; Guadeloupe; Dominica; Trinidad). — Scheerpeltz, 1970: 118 (Espeson; characters; Guadeloupe; Dominica; Martinique; St. Vincent; Trinidad). DISTRIBUTION: West Indies. franzi Coiffait, 1981g: 307 (Espeson; Type locality: Ile Galapagos: Ile Isabela, environs de Santa Tomas). DISTRIBUTION: Galapagos Islands. gigantulus Bernhauer, 1915, see: Pseudespeson. grossepunctatus Cameron, 1950f: 111 (Espeson; Type locality: Angola: Dundo). — Scheerpeltz, 1970: 126 (Espeson; characters; Angola; Southwest Africa). DISTRIBUTION: Angola, Namibia. indomalayensis Bernhauer, 1926, see: Pseudespeson. lenkoi Scheerpeltz, 1970: 119 (Espeson; Type locality: Südost-Brasilien, Barueri, São Paulo). DISTRIBUTION: Brazil. lombokianus Cameron, 1945, see: Parespeson. luzonicus Bernhauer, 1926, see: Pseudespeson. mexicanus Fauvel, 1902: 37 (Espeson; Type locality: Mexique). — Bernhauer and Schubert, 1910: 24 (Espeson; catalog). — Scheerpeltz, 1970: 117 (Espeson; characters; Mexico; Guatemala). DISTRIBUTION: Mexico, Guatemala. microphthalmus Fauvel, 1902: 36 (Espeson; Type locality: Guyane française). — Bernhauer and Schubert, 1910: 24 (Espeson; catalog). — Scheerpeltz, 1970: 117 (Espeson; characters; French Guiana; Guyana; Venezuela). DISTRIBUTION: French Guiana, Guyana, Venezuela. minor Coiffait, 1981g: 308 (Espeson; Type locality: Ile Galapagos: Ile Isabela, environs de Santa Tomas). DISTRIBUTION: Galapagos Islands. moratus Schauffus, 1882b: 168 (Espeson; Type locality: St. Thomas). — Schaufuss, 1882: 45 (Espeson; Saint Thomas). — Bernhauer and Schubert, 1910: 24 (Espeson; catalog). — Blackwelder, 1943: 146 (Espeson; characters; Cuba; Jamaica; Puerto Rico; St. Thomas; Montserrat; St. Lucia; St. Vincent; Grenada; Trinidad). — Scheerpeltz, 1970: 119 (Espeson; characters; Cuba; Jamaica; Haiti; Puerto Rico; St. Thomas; Monserrat; Guadeloupe; Martinique; Santa Lucia; St. Vincent; Grenada; Trinidad). DISTRIBUTION: West Indies. Bulletin AMNH OSORIINAE: ESPESON 1241 nepalicus Coiffait, 1981, see: Pseudespeson. nevermanni Bernhauer, 1942: 4 (Espeson; Type locality: Kostarika: Reventazon, Ebene Limon, Hamburgfarm). — Scheerpeltz, 1970: 120 (Espeson; characters; Costa Rica; Nicaragua). DISTRIBUTION: Costa Rica, Nicaragua. nitens Fauvel, 1902, see: Pseudespeson. novaeguineae Fauvel, 1902: 37 (Espeson; Type locality: Nouvelle Guinee allemande: Lemien; baie Huon Sattelberg). — Bernhauer and Schubert, 1910: 24 (Espeson; catalog). — Scheerpeltz, 1970: 121 (Espeson; characters; New Guinea). DISTRIBUTION: New Guinea. peyrierasi Lecoq, 1995: 418 (Espeson; Type locality: Seychelles: Mahé, 700 m). DISTRIBUTION: Seychelles. philippinus Bernhauer, 1926 see: Pseudespeson. punctatus Cameron, 1929b: 441 (Espeson; Type locality: Selangor: Bukit Kutu). — Scheerpeltz, 1933: 1020 (Espeson; catalog). — Scheerpeltz, 1970: 124 (Espeson; characters; Malaysia). DISTRIBUTION: Malaysia. scotti Bernhauer, 1922, see: Pseudespeson. subtilis Bernhauer, 1910b: 352 (Espeson; Type locality: Mexiko: Vera-Cruz, Cordoba). — Scheerpeltz, 1933: 1020 (Espeson; catalog). — Scheerpeltz, 1970: 120 (Espeson; characters; Guatemala; El Salvador; Colombia; Venezuela; Brazil). DISTRIBUTION: Mexico, El Salvador, Colombia, Venezuela, Brazil. titschacki Bernhauer, 1941c[= 1951]: 278[= 268] (Espeson; Type locality: Süd-Peru: Sivia, 520 m). — Scheerpeltz, 1970: 121 (Espeson; characters; Peru; Bolivia). DISTRIBUTION: Peru, Bolivia.
Geomitopsis [18 species; Neotropical, Palaearctic, and Ethiopian Regions] Geomitopsis Scheerpeltz, 1931: 388 (species included: zariquieyi, beieri). TYPE SPECIES: Cylindropsis zariquieyi Dodero, by original designation. — Scheerpeltz, 1933: 1128 (world catalog supplement). — Normand, 1935: 363 (list of species of Tunisia). — Blackwelder, 1952: 169 (type species: zariquieyi). — Fagel, 1955: 81 (characters; type species zariquieyi). — Coiffait, 1955b: 263 (characters in key). — Coiffait and Saiz, 1968: 414 (list of Chilean species). — Fagel, 1969: 55 (notes; key to African species). — Saiz, 1973a: 21 (key to species of Chile). — Orousset, 1985a: 63 (characters; notes). — Drugmand, 1998a: 209 (list of types in l’Institut royal des Sciences naturelles de Belgique). Libanotyphlus Coiffait, 1954a: 155 (species included: dubertreti). TYPE SPECIES: Libano- typhlus dubertreti Coiffait, fixed by original designation and monotypy. — Coiffait, 1955b: 263 (characters; type species). — Coiffait, 1959b: 218 (synonym of Geomitopsis). — Orousset, 1985a: 63 (synonym of Geomitopsis). — Drugmand, 1998a: 212 (list of types in l’Institut royal des Sciences naturelles de Belgique). Pseudogeomitopsis Orousset, 1985a: 63 (subgenus of Geomitopsis; species included: remil- leti). TYPE SPECIES: Geomitopsis (Pseudogeomitopsis) remilleti Orousset, fixed by original designation and monotypy. 1242 OSORIINAE: GEOMITOPSIS Bulletin AMNH anatolica Coiffait, 1979b: 316 (Geomitopsis; Type locality: Turquie méridionale: 10 km nord de Mersin). DISTRIBUTION: Turkey. angolensis Fagel, 1969: 58 (Geomitopsis; Type locality: Angola: Alto Chicapa, forêt aux sources du Cuilo). DISTRIBUTION: Angola. beieri Scheerpeltz, 1931: 392 (Geomitopsis; Type locality: Kaligoni auf Levkas). — Scheerpeltz, 1933: 1128 (Geomitopsis; catalog). DISTRIBUTION: Greece. boiteli Normand, 1932: 181 (Geomitopsis; Type locality: Bizerte, Tunisie). — Scheerpeltz, 1933: 1129 (Geomitopsis; catalog). DISTRIBUTION: Tunisia. campanae Saiz, 1973a: 19 (Geomitopsis; Type locality: La Campana, Valparaiso, 750 m, Chili). DISTRIBUTION: Chile. celisi Fagel, 1959: 57 (Geomitopsis; Type locality: Congo Belge: Kivu: territoire de Lubero, Est mont Kibatsiro). — Fagel, 1969: 55 (Geomitopsis; characters). DISTRIBUTION: Congo. chilensis Coiffait and Saiz, 1965b: 577 (Geomitopsis; Type locality: Chili: Curacavi). — Saiz, 1973a: 21 (Geomitopsis; characters). DISTRIBUTION: Chile. dubertreti Coiffait, 1954a: 156 (Libanotyphlus; Type locality: Liban: Beyrouth et environs). DISTRIBUTION: Lebanon. franzi Coiffait, 1978c: 45 (Geomitopsis; Type locality: Barranco de Añavigo près de Arafo, Ténérife, Iles Canaries). — Hernández, Outerelo, and Gamarra, 1994: 192 (Geomitopsis; Canary Islands). DISTRIBUTION: Canary Islands. gabonensis Coiffait, 1967: 277 (Geomitopsis; Type locality: Bélinga). DISTRIBUTION: Gabon. grosclaudei Normand, 1936: 163 (Geomitopsis; Type locality: Tunisie: Radès). DISTRIBUTION: Tunisia. itombwensis Fagel, 1969: 55 (Geomitopsis; Type locality: Kivu: Itombwe, Mulenge, Nyalengwe, 2300 m). DISTRIBUTION: Congo. leleupi Fagel, 1955: 83 (Geomitopsis; Type locality: Congo Belge: Kivu: territoire de Kabare, Nyakasiba, 1900 m). — Fagel, 1959: 56 (Geomitopsis; characters). — Fagel, 1969: 55 (Geomitopsis; characters). DISTRIBUTION: Congo. lusitanica Coiffait, 1965d: 285 (Geomitopsis; Type locality: Portugal, environs de Portimao). DISTRIBUTION: Portugal. mussardi Coiffait, 1965d: 286 (Geomitopsis; Type locality: Maroc, Ouirgane). DISTRIBUTION: Morocco. remilleti Orousset, 1985a: 64 (Geomitopsis; subgenus Pseudogeomitopsis; Type locality: Guyane française: Mont Grand-Matoury, près de Cayenne, alt. 200 m). DISTRIBUTION: French Guiana. tshuapaensis Fagel, 1969: 56 (Geomitopsis; Type locality: Tshuapa: territoire d’Ikela, ruisseau Befali). DISTRIBUTION: Congo. Bulletin AMNH OSORIINAE: GEOMITOPSIS 1243 zariquieyi Dodero, 1918: 104 (Cylindropsis; Type locality: Bonanova et à S. Feliu de Llobregat). — Scheerpeltz, 1931: 392 (Geomitopsis; type species of Geomitopsis; notes). — Scheerpeltz, 1933: 1129 (Geomitopsis; catalog). — Español, 1945: 91 (Geomitopsis; notes; Spain). — Coiffait, 1959b: 218 (Geomitopsis; characters; notes). — Coiffait, 1973b: 223 (Geomitopsis; Spain). DISTRIBUTION: Spain.
Glyptoma [21 species; Neotropical Region] Glyptoma Erichson, 1839b: 32 (without included species, for first included species see Erich- son, 1840; [Note: Glyptoma was first cited in a key (1839) by Erichson who, in 1840, cited it with Thoraxophorus Motschulsky (a lapsus that was changed to Thoracopho- rus by Motschulsky in 1840) as a junior synonym]). TYPE SPECIES: Glyptoma crassi- cornis Erichson, fixed by subsequent designation by Duponchel, 1841: 57. — Erichson, 1840: 908 (first included species: crassicornis, corticinum, costale, denticolle, exile, sculptile; characters). — Redtenbacher, 1849: 757 (characters). — Lacordaire, 1854: 148 (characters; notes; list of species). — Fairmaire and Laboulbène, 1856: 657 (characters). — Kraatz, 1857a: 49 (characters). — Motschulsky, 1860a: 66 (Glyptoma was proposed as a replacement name for Thoraxophorus; discussion). — LeConte, 1861a: 72 (characters). — Fauvel, 1864b: 58 [= 1865: 62] (characters; key to species). — Fauvel, 1871a: 40 [= 1872: 14] (synonym of Thoracophorus). — Fauvel, 1875a: I [= 1875b: 203] (synonym of Thoracophorus). — LeConte, 1877: 249 (characters). — Sharp, 1887: 723 (notes; Mexican and Central American species). — Bernhauer and Schubert, 1910: 25 (synonym of Thoracophorus). — Scheerpeltz, 1933: 1020 (synonym of Thoracophorus). — Blackwelder, 1943: 141 (characters; type species: crassicorne; key to species of West Indies). — Blackwelder, 1944: 108 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 171 (type species). Calocerus Fauvel, 1891: 88 (species included: cicatricosus, crassicornis; preoccupied by LeConte, 1853). TYPE SPECIES: Glyptoma crassicornis Erichson, fixed by subsequent designation by R. Lucas, 1920: 159. — Bernhauer and Schubert, 1910: 26 (world catalog; 13 species). — Bruch, 1928a: 422 (catalog; Argentina). — Scheerpeltz, 1933: 1021 (world catalog supplement). — Blackwelder, 1942: 88 (type species: cicatricosus). — Blackwelder, 1943: 141 (synonym of Glyptoma; type species: cicatricosus). — Blackwelder, 1952: 90 (synonym of Glyptoma; type species: crassicornis). — Scheerpeltz, 1970a (cited as valid genus; [Note: Calocerus is preoccupied so cannot be the valid name of this genus]; key to species). — Drugmand, 1998a: 207 (list of types in l’Institut royal des Sciences naturelles de Belgique). blackburni Sharp, 1880, see: Thoracophorus. boliviana Scheerpeltz, 1970a: 141 (Calocerus; Type locality: Bolivien: Region Chaparé). DISTRIBUTION: Bolivia. bonvouloiri Fauvel, 1867, see: Thoracophorus. brevicristatum Horn, 1871, see: Thoracophorus. 1244 OSORIINAE: GLYPTOMA Bulletin AMNH brevipennis Sharp, 1880, see: Thoracophorus. cicatricosa Motschulsky, 1857a: 494 (Glyptoma; Type locality: Colombie). — Fauvel, 1891: 88 (Calocerus; Venezuela). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 131 (Calocerus; characters; Colombia; Ecuador; Venezuela). DISTRIBUTION: Ecuador, Colombia, Venezuela. cognata Sharp, 1887: 724 (Glyptoma; Type locality: Guatemala, El Reposo). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 138 (Calocerus; characters; Guatemala; Costa Rica). DISTRIBUTION: Guatemala, Costa Rica. costalis Erichson, 1840, see: Thoracophorus. crassicornis Erichson, 1840: 908 (Glyptoma; Type locality: Novae Valenciae, Columbia). — Fauvel, 1864b: 59 [= 1865: 62] (Glyptoma; characters; Colombia). — Fauvel, 1891: 88 (Calocerus; notes; Venezuela). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 134 (Calocerus; characters; Colombia; Ecuador; Venezuela). DISTRIBUTION: Ecuador, Colombia, Venezuela. cribrella Sharp, 1887: 724 (Glyptoma; Type locality: Guatemala, El Tumbador, Zapote, Capetillo; Panama, Volcan de Chiriqui). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 137 (Calocerus; characters; Panama; Guatemala; El Salvador). DISTRIBUTION: El Salvador, Guatemala, Panama. denieri Bernhauer, 1939h: 232 (Calocerus; Type locality: Argentinien: Salta: S. Lorenzo). — Scheerpeltz, 1970a: 140 (Calocerus; characters; Argentina). DISTRIBUTION: Argentina. denticollis Erichson, 1840, see: Thoracophorus. exile Erichson, 1840, see: Thoracophorus. guadelupensis Wendeler, 1930: 182 (Calocerus; Type locality: Trois-Rivieres, Guadelupe). — Scheerpeltz, 1933: 1021 (Calocerus; catalog). — Blackwelder, 1943: 143 (Glyptoma; characters; Guadeloupe). — Scheerpeltz, 1970a: 136 (Calocerus; characters; Guadeloupe). DISTRIBUTION: West Indies. hirticornis Sharp, 1887: 724 (Glyptoma; Type locality: Guatemala, Capetillo). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 136 (Calocerus; characters; Guatemala; Costa Rica). DISTRIBUTION: Guatemala, Costa Rica. inconspicuus Bernhauer, 1929, see: Thoracophorus. kingi Lea, 1910, see: Myrmelibia. klimschi Bernhauer, 1908d: 285 (Calocerus; Type locality: Brasilien: S. Catharina). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 139 (Calocerus; characters; Brazil). DISTRIBUTION: Brazil. laeviceps Sharp, 1887: 723 (Glyptoma; Type locality: Panama, Volcan de Chiriqui 2000 to 4000 feet). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 132 (Calocerus; characters; Panama; El Salvador). DISTRIBUTION: Panama, El Salvador. longipes Sharp, 1887: 723 (Glyptoma; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 133 (Calocerus; characters; Guatemala; Costa Rica). DISTRIBUTION: Guatemala, Costa Rica. Bulletin AMNH OSORIINAE: GLYPTOMA 1245 myrmecophilus Lea, 1910, see: Thoracophorus. nitens Bernhauer, 1908d: 284 (Calocerus; Type locality: Brasilien: S. Paulo). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 138 (Calocerus; characters; Brazil). DISTRIBUTION: Brazil. nitida Bernhauer, 1908d: 284 (Calocerus; Type locality: Paraguay). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 139 (Calocerus; characters; Paraguay). DISTRIBUTION: Paraguay. obscuricollis Bernhauer, 1908d: 284 (Calocerus; Type locality: Brasilien). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 134 (Calocerus; characters; Brazil). DISTRIBUTION: Brazil. opaca Sharp, 1876: 418 (Thoracophorus, cited as Thoraxophorus; Type locality: Ega). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 133 (Calocerus; characters; Brazil). DISTRIBUTION: Brazil. punctatoplicata Solsky, 1870a: 267 (Glyptoma; Type locality: Cayenne). — Solsky, 1872: 313 (Thoracophorus; Cayenne). — Fauvel, 1901: 69 (Calocerus; Colombia; Cayenne; Bolivia; Para). — Bernhauer and Schubert, 1910: 26 (Calocerus; catalog). — Scheerpeltz, 1970a: 137 (Calocerus; characters; French Guiana; Guyana; Venezuela; Brazil). DISTRIBUTION: French Guiana, Guyana, Venezuela, Colombia, Brazil, Bolivia. reichenspergeri Scheerpeltz, 1970a: 135 (Calocerus; Type locality: Südost-Brasilien: Paraná, São Paulo, St. Catharina). DISTRIBUTION: Brazil. ruficolle Fauvel, 1864, see: Thoracophorus. sanctaecatharinae Scheerpeltz, 1970a: 135 (Calocerus; Type locality: Südost-Brasilien: St. Catharina). DISTRIBUTION: Brazil. sculptile Erichson, 1840, see: Thoracophorus. sculptum Blackburn, 1902, see: Thoracophorus. simplex Wendeler, 1955: 191 (Calocerus; Type locality: Brasilien: Nova Teutonia). — Scheerpeltz, 1970a: 140 (Calocerus; characters; Brazil). DISTRIBUTION: Brazil. sordidus Blackburn, 1902, see: Thoracophorus. trinidadensis Blackwelder, 1943: 142 (Glyptoma; Type locality: Trinidad: River Estate near Diego Martin, 8 miles northwest of Port of Spain). — Scheerpeltz, 1970a: 141 (Calocerus; characters; Trinidad). — Legner and Moore, 1977: 174 (Glyptoma; found under bark). DISTRIBUTION: West Indies. zischkai Scheerpeltz, 1970a: 132 (Calocerus; Type locality: Bolivien: Region Chaparé). DISTRIBUTION: Bolivia.
Lispinodes [17 species; Nearctic, Neotropical, Ethiopian, Oriental, and Oceanic Regions] Lispinodes Sharp, 1880: 54 (species included: explicandus). TYPE SPECIES: Lispinodes explicandus Sharp, fixed by monotypy. — Duvivier, 1883: 204 (catalog). — Bernhauer and Schubert, 1910: 19 (world catalog; 17 species). — Cameron, 1925: 6 (catalog of Indian species). — Scheerpeltz, 1933: 1010 (world catalog supplement). 1246 OSORIINAE: LISPINODES Bulletin AMNH
— Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 233 (type species: explicandus). — Scheerpeltz, 1956a: 514 (key to African species). — Hammond, 1984: 196 (checklist of species of Borneo). — Drugmand, 1998a: 212 (list of types in l’Institut royal des Sciences naturelles de Belgique). — Newton, Thayer, Ashe, and Chandler, 2000: 378 (1 undescribed Nearctic species; notes; char- acters in key). africanus Bernhauer, 1927: 49 (Lispinodes; Type locality: Pondoland: Port St. John; Zululand: Eshowe; Natal: Kloof, 1500 ft.). — Scheerpeltz, 1933: 1010 (Lispinodes; catalog). — Scheerpeltz, 1956a: 515 (Lispinodes; characters; South Africa). DISTRIBUTION: South Africa. americanus Bernhauer, 1909c: 226 (Lispinodes; Type locality: Brasilien; Argentinien: Buenos Aires). — Bernhauer and Schubert, 1910: 19 (Lispinodes; catalog). DISTRIBUTION: Brazil, Argentina. bistriatus Fauvel, 1895, see: Lispinus. bisulcatus Fauvel, 1902: 32 (Lispinodes; Type locality: Sumatra, region de Deli). — Bernhauer and Schubert, 1910: 19 (Lispinodes; catalog). — Cameron, 1928d: 96 (Lispinodes; Sumatra). DISTRIBUTION: Indonesia. burgeoni Bernhauer, 1932a: 73 (Lispinodes; Type locality: Haut-Uele: Moto). — Scheerpeltz, 1933: 1010 (Lispinodes; catalog). — Scheerpeltz, 1956a: 482, 514 (Lispinodes; characters; French Congo; Belgian Congo). DISTRIBUTION: Congo, French Equatorial Africa. crassus Sharp, 1908: 542 (Lispinodes; Type locality: Maui, Haleakala, 5000 ft.). — Bernhauer and Schubert, 1910: 19 (Lispinodes; catalog). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. duplicatus Fauvel, 1878f: 197 (Thoracophorus; Type locality: Iles Arou, Wokan; Borneo; Sarawak; Bírmanie). — Bernhauer and Schubert, 1910: 19 (Lispinodes; catalog). DISTRIBUTION: Myanmar, Borneo, Indonesia. explicandus Sharp, 1880: 53 (Lispinodes; Type locality: higher of mountains of Oahu). — Blackburn and Sharp, 1885: 224 (Lispinodes; habitat; Hawaii). — Sharp, 1908: 540 (Lispinodes; characters; Hawaii). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Legner and Moore, 1977: 174 (Lispinodes; found under bark). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. germanus Sharp, 1908: 542 (Lispinodes; Type locality: Hawaii, Hilo, 2000 ft.). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. hawaiiensis Sharp, 1908: 541 (Lispinodes; Type locality: Hawaii, Hilo, 2000 ft.). — Bernhauer and Schubert, 1910: 20 (Lispinodes; cited as hawaiensis; catalog). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. Bulletin AMNH OSORIINAE: LISPINODES 1247 heteroderes Fauvel, 1902, see: Tetrapleurus. kauaiensis Sharp, 1908: 541 (Lispinodes; Type locality: Kauai, Koholuamano). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. mauiensis Sharp, 1908: 541 (Lispinodes; Type locality: Maui, Haleakala, 5000 ft.). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. molokaiensis Sharp, 1908: 540 (Lispinodes; Type locality: Molokai). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. obscurus Sharp, 1908: 541 (Lispinodes; Type locality: Molokai, mts. 4500 ft.). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. oxytelinus Sharp, 1908: 540 (Lispinodes; Type locality: Lanai). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. pallescens Blackburn, 1885, see: Clavilispinus. pallidus Sharp, 1908: 540 (Lispinodes; Type locality: Lanai, 2000 ft.). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Nishida, 1994: 72 (Lispinodes; Hawaii). — Nishida, 1997: 61 (Lispinodes; Hawaii). DISTRIBUTION: Hawaii. quadratus Blackburn, 1885, see: Lispinus. rhodesianus Bernhauer, 1934h: 484 (Lispinodes; Type locality: N. W. Rhodesia: Shigariatombwes. Also cited from Belgian Congo). — Scheerpeltz, 1956a: 514 (Lispinodes; characters; Uganda; Kenya; Tanganyika; Belgian Congo). DISTRIBUTION: Uganda, Kenya, Tanzania, Congo, Zambia. rugosus Fauvel, 1902, see: Tetrapleurus. seriatus Fauvel, 1902, see: Tetrapleurus. tenuipunctus Bernhauer, 1932a: 73 (Lispinodes; Type locality: Bas-Uele: Koteli). — Scheerpeltz, 1933: 1010 (Lispinodes; catalog). — Scheerpeltz, 1956a: 515 (Lispinodes; characters; French Congo; Belgian Congo). DISTRIBUTION: French Equatorial Africa, Congo. Parespeson [3 species; Oriental Region] Parespeson Bernhauer, 1926d: 261 (subgenus of Espeson; species included: angustissimus). TYPE SPECIES: Espeson (Parespeson) angustissimus Bernhauer, fixed by monotypy. — Blackwelder, 1943: 144 (subgenus of Espeson; type species angustissimus). — Blackwelder, 1952: 294 (type species: angustissimus). — Scheerpeltz, 1970: 126, 127 (valid genus; key to species). 1248 OSORIINAE: PARESPESON Bulletin AMNH angustiformis Cameron, 1950d: 3 (Espeson; subgenus Parespeson; Type locality: Selangor: The Gap). — Scheerpeltz, 1970: 128 (Parespeson; characters; Malaysia). DISTRIBUTION: Malaysia. angustissimus Bernhauer, 1926d: 261 (Espeson; subgenus Parespeson; Type locality: Luzon, Provinz Ilocos Norte, Bangui). — Scheerpeltz, 1933: 1020 (Espeson; subgenus Parespeson; catalog). — Scheerpeltz, 1970: 128 (Parespeson; characters; Philippines). DISTRIBUTION: Philippines. lombokianus Cameron, 1945a: 141 (Espeson; subgenus Parespeson; Type locality: Lombok: Sapit, altitude 2000 m). — Scheerpeltz, 1970: 127 (Parespeson; characters; Lombok). DISTRIBUTION: Indonesia.
Pseudespeson [12 species; Neotropical, Palaearctic, Ethiopian, Madagascan, Oriental] Pseudespeson Lecoq, 1994: 299 (species included: rossii, crassulus, crenicollis, gigantulus, indomalayicus, luzonicus, nepalicus, nitens, philippinus, philippinus curtus, and scotti). TYPE SPECIES: Pseudespeson rossii Lecoq, fixed by original designation. — Drugmand, 1998a: 223 (list of types in l’Institut royal des Sciences naturelles de Belgique). crassulus Fauvel, 1902: 35 (Espeson; Type locality: Antilles: Guadeloupe; Saint Vincent). — Bernhauer and Schubert, 1910: 24 (Espeson; catalog). — Blackwelder, 1943: 144 (Espeson; characters; Montserrat; Guadeloupe; St. Lucia; St. Vincent). — Scheerpeltz, 1970: 116 (Espeson; characters; Montserrat; Guadeloupe; Martinique; Santa Lucia; St. Vincent; Grenada). — Lecoq, 1994: 304 (Pseudespeson). DISTRIBUTION: West Indies. crenicollis Fauvel, 1878f: 196 (Thoracophorus; Type locality: Iles Key). — Fauvel, 1879: 75 (Thoracophorus; Gilolo). — Bernhauer and Schubert, 1910: 24 (Espeson; catalog). — Scheerpeltz, 1970: 121 (Espeson; characters; Key Island; Halmahera Island [Gilolo]). — Lecoq, 1994: 304 (Pseudespeson). DISTRIBUTION: Indonesia. curtus Bernhauer, 1926, see: philippinus Bernhauer, 1926. gigantulus Bernhauer, 1915d: 97 (Espeson; Type locality: Deutsch-Ostafrika: West-Usambara). — Scheerpeltz, 1933: 1019 (Espeson; catalog). — Scheerpeltz, 1970: 125 (Espeson; characters; Tanzania). — Lecoq, 1994: 303, 304 (Pseudespeson; characters). DISTRIBUTION: Tanzania. gomyi Lecoq, 1995: 416 (Pseudespeson; Type locality: Moihani, Mohéli, archipel des Comores). DISTRIBUTION: Comoros. indomalayensis Bernhauer, 1926d: 262 (Espeson; Type locality: Polillo). — Scheerpeltz, 1933: 1019 (Espeson; catalog). — Scheerpeltz, 1970: 122 (Espeson; characters; Philippines). — Lecoq, 1994: 304 (Pseudespeson; cited as indomalayicus). DISTRIBUTION: Philippines. luzonicus Bernhauer, 1926d: 262 (Espeson; Type locality: Luzon, Provinz Nueva Vizcaya, Imugan). — Scheerpeltz, 1933: 1019 (Espeson; catalog). — Scheerpeltz, 1970: 123 (Espeson; characters; Philippines). Bulletin AMNH OSORIINAE: PSEUDESPESON 1249
— Lecoq, 1994: 304 (Pseudespeson). DISTRIBUTION: Philippines. minusculus Coiffait, 1982, see: nepalicus Coiffait, 1981. nepalicus Coiffait, 1981d: 334 (Espeson; Type locality: Royal Chitwan, National Park, Theral; [Note: The preceding locality is as cited on page 325 and is a correction of the cita- tion on page 334, “Royal Shitwan, National par , Theral”]). — Lecoq, 1994: 304 (Pseudespeson). DISTRIBUTION: Nepal. minusculus Coiffait, 1982: 135 (Bothrys; Type locality: Nepal: Siwalik, 660 m). — Lecoq, 1994: 304 (Pseudespeson; synonym of nepalicus). nitens Fauvel, 1902: 36 (Espeson; Type locality: Bresil; Guadeloupe). — Bernhauer and Schubert, 1910: 24 (Espeson; catalog). — Blackwelder, 1943: 147 (Espeson; characters; Guadeloupe). — Scheerpeltz, 1970: 117 (Espeson; characters; Guadeloupe; Martinique; Grenada; Trinidad; Venezuela; Brazil). — Lecoq, 1994: 304 (Pseudespeson). — Lecoq, 1994: 304 (Pseudespeson). DISTRIBUTION: Brazil, Venezuela, West Indies. orousseti Lecoq, 1995: 417 (Pseudespeson; Type locality: Philippines, Luzon, Sagada). DISTRIBUTION: Philippines. philippinus Bernhauer, 1926d: 262 (Espeson; Type locality: Luzon, Provinz Laguna, Los Baños. Siargao, Dapa. Masbate, Aroroy. Dinagat. Mindanao, Provinz Lanao, Mumungan). — Scheerpeltz, 1933: 1019 (Espeson; catalog). — Scheerpeltz, 1970: 123 (Espeson; characters; Philippines). — Lecoq, 1994: 304 (Pseudespeson). DISTRIBUTION: Philippines. curtus Bernhauer, 1926d: 262 (Espeson; variety of philippinus; Type locality: Dapa). — Scheerpeltz, 1933: 1019 (Espeson; synonym of philippinus). — Scheerpeltz, 1970: 124 (Espeson; valid species; characters; Philippines). — Lecoq, 1994: 304 (Pseudespeson; subspecies of philippinus). rossii Lecoq, 1994: 302 (Pseudespeson; Type locality: Sierra Leone: Western Area, Number Two River Forest. Also cited from Ivory Coast and Guinea). DISTRIBUTION: Guinea, Sierra Leone, Ivory Coast. scotti Bernhauer, 1922b: 166 (Espeson; Type locality: Seychellen: Silhouette: Umgebung von Pot-à-eau, circa 1500 ft.). — Scheerpeltz, 1933: 1020 (Espeson; catalog). — Scheerpeltz, 1970: 125 (Espeson; characters; Seychelles). — Lecoq, 1994: 304 (Pseudespeson). DISTRIBUTION: Seychelles.
Synaenictus [1 species; Ethiopian Region] Synaenictus Patrizi, 1947: 228 (species included: foveicauda). TYPE SPECIES: Synaenictus foveicauda Patrizi, fixed by monotypy. — Blackwelder, 1952: 367 (type species: foveicauda). — Seevers, 1965: 341 (characters; may be Pselaphidae not Staphylinidae; myrmecophilus species). — Burakowski and Newton, 1992: 38 (uncertain placement in Osoriinae). foveicauda Patrizi, 1947: 229 (Synaenictus; Type locality: Elmenteita, Kenya Colony). DISTRIBUTION: Kenya. 1250 OSORIINAE: LISPININA Bulletin AMNH
Subtribe Lispinina Lispinina Bernhauer and Schubert, 1910. — Arnett, 1963: 262 (characters; list of North American genera). — Newton and Thayer, 1992: 59 (subtribe of Thoracophorini). — Newton, Thayer, Ashe, and Chandler, 2000: 377 (characters; notes). Lispini Bernhauer and Schubert, 1910: 19 (genera included: Ancaeus, Lispinodes, Holosus, Lispinus). TYPE GENUS: Lispinus (see Newton and Thayer, 1992: 59). — Scheerpeltz, 1933: 1009 (world catalog supplement). — Blackwelder, 1942: 77, 78 (characters; key to genera and subgenera). — Blackwelder, 1944: 107 (checklist of species from Mexico, Central and South America, and the West Indies). Lispininae Bernhauer and Schubert, 1910. — Moore, 1964a: 87 (characters; notes). — Moore and Legner, 1974b: 77 (characters; key to genera of America north of Mexico). — Moore and Legner, 1974g: 555 (key to genera of America north of Mexico). — Moore and Legner, 1975: 166 (catalog; North America north of Mexico). — Moore and Legner, 1979: 143 (characters; tabular key to genera of North America north of Mexico). — Downie and Arnett, 1996: 426 (characters; key to genera and species of Northeastern North America). Lispinini Bernhauer and Schubert, 1910. — Blackwelder, 1942: 76, 78 (characters; key to subtribes). — Blackwelder, 1944: 107 (checklist of species from Mexico, Central and South America, and the West Indies). — Hatch, 1957: 243 (characters; key to species of Pacific Northwest). — Arnett, 1963: 242, 262 (key to North American genera; characters; list of North American genera). Lispinus [282 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions] Lispinus Erichson, 1839b: 31 (without included species, for first included species see Erich- son, 1840). TYPE SPECIES: Lispinus attenuatus Erichson, fixed by subsequent designation by Duponchel, 1841: 57. — Erichson, 1840: 828 (characters; first included species: attenuatus, laticollis, striola, angustatus, linearis, tenellus, exiguus). — Lacordaire, 1854: 126 (characters; notes; list of species). — LeConte, 1861a: 72 (characters). — LeConte, 1863: 27 (list of North American species). — Fauvel, 1864b: 40 [= 1865: 44] (characters; list of and key to species). — Gemminger and Harold, 1868: 677 (catalog). — Sharp, 1876: 411 (notes). — LeConte, 1877: 250 (key to species). — Fauvel, 1878d: 176 [= 1878e: 12] (key to North American species). — Fauvel, 1878f: 33 (characters; key to New Guinea species). — Duvivier, 1883: 204 (catalog). — Sharp, 1887: 781 (notes; Mexican and Central American species). — Fauvel, 1897d: 240 (catalog of species of Madeira). — Bernhauer and Schubert, 1910: 21 (world catalog; 94 species). — Blatchley, 1910: 482 (characters). — Leng, 1920: 93 (catalog of North American species). — Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog). — Winkler, 1925: 323 (catalog for Palaearctic region). — Cameron, 1925: 7, 103 (catalog of Indian species). — Cameron, 1930: 51, 452 (characters; key to species of British India). Bulletin AMNH OSORIINAE: LISPINUS 1251
— Scheerpeltz, 1933: 1012 (world catalog supplement). — Bernhauer, 1934h: 483 (key to species of South Africa). — Paulian, 1941: 139 (larval characters). — Blackwelder, 1942: 83 (characters; notes). — Blackwelder, 1943: 129 (characters; type species: attenuatus; key to species of West Indies). — Blackwelder, 1944: 107 (checklist of species from Mexico, Central and South America, and the West Indies). — Shibata, 1973c: 22 (checklist of species of Taiwan). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Moore and Legner, 1975: 168 (catalog; North America north of Mexico). — Shibata, 1976: 82 (checklist of species of Japan). — Moore and Legner, 1979: 146 (characters; notes). — Hammond, 1984: 196 (checklist of species of Borneo). — Newton, 1990: 92 (characters in key; three described Australian species). — Irmler, 1994: 54 (key to Neotropical species). — Naomi, 1996b: 762 (characters; species of Japan). — Drugmand, 1998a: 212 (list of types in l’Institut royal des Sciences naturelles de Belgique). — Newton, Thayer, Ashe, and Chandler, 2000: 377 (2 Nearctic species; notes; characters in key). Paralispinus Eichelbaum, 1913: 117 (subgenus of Lispinus; species included: amaniensis). TYPE SPECIES: Lispinus (Paralispinus) amaniensis Eichelbaum, fixed by monotypy. — Blackwelder, 1952: 290 (subgenus of Lispinus; type species: amaniensis). Pseudolispinodes Bernhauer, 1926d: 258 (species included: longipennis, madurensis, lati- usculus, sinuatus). TYPE SPECIES: Holosus madurensis Bernhauer, 1914, fixed by subsequent designation by Blackwelder, 1942: 88. — Cameron, 1930: 65 (characters). — Scheerpeltz, 1933: 1010 (world catalog supplement). — Blackwelder, 1943: 120 (characters; [Note: Species included in Pseudolispinodes by Black- welder are now in Nacaeus]; type species: madurensis; key to species of West Indies). — Blackwelder, 1952: 327 (subgenus of Lispinus; type species: madurensis). — Abdullah and Qadri, 1970: 114 (key to species of West Pakistan). — Drugmand, 1998a: 223 (list of types in l’Institut royal des Sciences naturelles de Belgique). Spinilus Blackwelder, 1942: 83 (subgenus of Lispinus; species included: bistriatus, sinuatus). TYPE SPECIES: Lispinodes bistriatus Fauvel, fixed by original designation. — Blackwelder, 1952: 355 (synonym of Lispinus; type species). Aulacolispinus Scheerpeltz, 1957c: 223, 239, 240 (subgenus of Lispinus; species included: unistriatus, lineipennis, cheesmani, sutteri, sulcipennis, macropterus, sulcatus, validi- cornis, sulcatulus, germanus, striola). TYPE SPECIES: Lispinus unistriatus Fauvel, fixed by original designation. Stictolispinus Scheerpeltz, 1957c: 241 (subgenus of Lispinus; species included: quadripunc- tulus). TYPE SPECIES: Lispinus quadripunctulus Fauvel, fixed by original designation. abdominalis Bernhauer, 1929i: 355 (Lispinus; Type locality: Luzon, Haight’s Place). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Philippines. acicularis Fauvel, 1904b: 80 (Lispinus; Type locality: Java). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Indonesia. aciculatus Bernhauer, 1927: 47 (Lispinus; Type locality: Gazaland: Chirinda). — Bernhauer, 1934h: 483 (Lispinus; characters in key). DISTRIBUTION: South Africa. acupunctus Eppelsheim, 1895a: 137 (Lispinus; Type locality: Gabon). — Fauvel, 1903e: 235 (Lispinus; Cameroon; Togo; Gabon; Usambara). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). 1252 OSORIINAE: LISPINUS Bulletin AMNH
— Paulian, 1942: 347 (Lispinus; larval characters). — Fagel, 1963a: 339 (Lispinus; Guinea; Ivory Coast). DISTRIBUTION: Guinea, Ivory Coast, Togo, Cameroon, Gabon, Tanzania. acutepunctatus Bernhauer, 1929, see: bernhauerianus Scheerpeltz, 1933. acutepunctatus Bernhauer, 1929i: 344 (Lispinus; Type locality: Luzon: Haight’s Place, Balbalan). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Philippines. aeneiceps Cameron, 1925b: 236, 240 (Lispinus; Type locality: Penang). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). — Cameron, 1936: 4 (Lispinus; Society Islands). — Cameron, 1936a: 29 (Lispinus; Java). — Coiffait, 1976: 220 (Lispinus; Tahiti). DISTRIBUTION: Malaysia, Indonesia, Society Islands. aequalis Fauvel, 1878f: 205 (Lispinus; Type locality: Nouvelle-Guinée, Ansus, Ramoi; Iles Key). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Bernhauer, 1922b: 166 (Lispinus; Seychelles). — Cameron, 1931: 356 (Lispinus; New Guinea). — Cameron, 1937a: 87 (Lispinus; New Guinea). DISTRIBUTION: New Guinea, Indonesia, Seychelles. aequicollis Bernhauer, 1929i: 350 (Lispinus; Type locality: Surigao, Dapa). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Philippines. aequipunctatus LeConte, 1868: 50 (Lispinus; Type locality: Cañada de los Diegos, New Mexico). — LeConte, 1877: 250 (Lispinus; characters). — Fauvel, 1878d: 177 [= 1878e: 13] (Lispinus; characters; New Mexico). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). DISTRIBUTION: USA. aethiops Eppelsheim, 1895, see: Nacaeus. affinis Cameron, 1943a: 336 (Lispinus; Type locality: New Guinea: Moroko). DISTRIBUTION: New Guinea. africanus Bernhauer, 1929, see: Nacaeus. agnatus Fauvel, 1904e: 297 (Lispinus; Type locality: Tananarive; forêt de Moramanga). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). DISTRIBUTION: Madagascar. alutaceipennis Scheerpeltz, 1933: 1012 (Lispinus; replacement name for alutipennis Bernhauer). DISTRIBUTION: Philippines. alutipennis Bernhauer, 1929i: 354 (Lispinus; [preoccupied]; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1012 (Lispinus; synonym of alutaceipennis). alutaceipennis Bernhauer, 1937, see: kenyanus Herman, 2001. alutaceus Fauvel, 1878f: 205 (Lispinus; Type locality: Nouvelle-Guinée, Hatam). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Bernhauer, 1926f: 312 (Lispinus; Buru). DISTRIBUTION: New Guinea, Indonesia. alutipennis Bernhauer, 1921b: 66 (Lispinus; Type locality: Bolivien: Yuracaris). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Bolivia. Bulletin AMNH OSORIINAE: LISPINUS 1253 alutipennis Bernhauer, 1929, see: alutaceipennis Scheerpeltz, 1933. amaniensis Eichelbaum, 1913: 117 (Lispinus; subgenus Paralispinus; Type locality: Amani). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Tanzania. amherstanus Bernhauer, 1926: 20 (Lispinus; Type locality: Unter-Burma: Kawkareik, Amherst Distr.). — Cameron, 1930: 64 (Lispinus; characters; Burma). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Myanmar. andrewesi Cameron, 1930: 56 (Lispinus; Type locality: Nilgiri Hills). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: India. angolanus Cameron, 1950, see: dundoensis Cameron, 1950. anguinus Fauvel, 1864b: 50 [= 1865: 54] (Lispinus; Type locality: Mexique: Teapa; Santo Domingo). — Sharp, 1887: 720 (Lispinus; Mexico). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). — Blackwelder, 1943: 133 (Lispinus; characters; Jamaica; Haiti; Dominican Republic). — Scheerpeltz, 1975a: 242 (Lispinus; collecting notes; Brazil). — Irmler, 1994: 62 (Lispinus; characters; Santo Domingo). DISTRIBUTION: Mexico, West Indies, Brazil. angustatus Erichson, 1840: 829 (Lispinus; Type locality: Madagascar). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). DISTRIBUTION: Madagascar. angustus Bernhauer, 1929i: 342 (Lispinus; [Note: Bernhauer wrote that this species is perhaps identical with macropterus]; Type locality: Luzon, Los Baños, Mount Banahao, Balbalan, Mount Isarog. Mindanao, Mumungan, Surigao, Port Banga. Palawan, Binaluan. Dinagat). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Philippines. antennatus Bernhauer, 1929i: 342 (Lispinus; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Philippines. apicalis Sharp, 1876, see: attenuatus Erichson, 1840. aremicus Blackwelder, 1943, see: linearis Erichson, 1840. argentinus Bernhauer, 1927e: 229 (Lispinus; Type locality: Argentinien: Prov. S. Fe, Fives Lille). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). — Irmler, 1994: 60 (Lispinus; characters; Argentina; Paraguay; Bolivia). DISTRIBUTION: Argentina, Paraguay, Bolivia. armifrons Cameron, 1925b: 236, 242 (Lispinus; Type locality: Selangor, The Gap). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: Malaysia. aper Sharp, 1889, see: Neolosus. assamensis Cameron, 1930: 452 (Lispinus; Type locality: Assam: Naga Hills, alt. 6000 feet). — Scheerpeltz, 1933: 1012 (Lispinus; catalog). DISTRIBUTION: India. attenuatus Erichson, 1840: 828 (Lispinus; Type locality: Puerto Rico). — Fauvel, 1864b: 44 [= 1865: 48] (Lispinus; characters; Puerto Rico). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). 1254 OSORIINAE: LISPINUS Bulletin AMNH
— Blackwelder, 1943: 134 (Lispinus; characters; Puerto Rico; Guadeloupe; Trinidad). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Irmler, 1994: 58 (Lispinus; characters; Puerto Rico; Brazil; Bolivia; Peru). DISTRIBUTION: West Indies, Brazil, Peru, Bolivia. apicalis Sharp, 1876: 413 (Lispinus; Type locality: Para). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Irmler, 1994: 58 (Lispinus; synonym of attenuatus). auratus Irmler, 1994: 64 (Lispinus; Type locality: Vara Blanca, Costa Rica). DISTRIBUTION: Costa Rica. bakeri Bernhauer, 1914c: 80 (Lispinus; Type locality: Philippinen, Los Banos [sic]). — Bernhauer, 1929i: 344 (Lispinus; characters in key; Philippines). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. balbalanensis Bernhauer, 1929i: 343 (Lispinus; Type locality: Luzon: Balbalan). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. banosanus Bernhauer, 1929i: 342 (Lispinus; Type locality: Luzon, Los Baños, Mount Isarog, Mount Maquiling. Dinagat). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. basipennis Bernhauer, 1929i: 351 (Lispinus; Type locality: Luzon, Mount Maquiling, Los Baños). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. beesoni Cameron, 1924, see: Nacaeus. bernhaueri Scheerpeltz, 1933: 1013 (Lispinus; replacement name for praenobilis Bernhauer). DISTRIBUTION: Philippines. praenobilis Bernhauer, 1929i: 351 (Lispinus; [preoccupied]; Type locality: Luzon, Mount Polis, Butac, Haight’s Place, Laguna). — Scheerpeltz, 1933: 1013 (Lispinus; synonym of bernhaueri). bernhauerianus Scheerpeltz, 1933: 1013 (Lispinus; replacement name for acutepunctatus Bernhauer). DISTRIBUTION: Philippines. acutepunctatus Bernhauer, 1929i: 352 (Lispinus; [preoccupied]; Type locality: Luzon, Benguet, Haight’s Place). — Scheerpeltz, 1933: 1013 (Lispinus; synonym of bernhauerianus). pennatus Bernhauer, 1929i: 352 (Lispinus; variety of acutepunctatus, cited as acutipuncta- tus; Type locality: Luzon, Trinidad, Balbalan, Mount Polis, Mount Banahao, Imugan). bicolor Sharp, 1887: 721 (Lispinus; Type locality: British Honduras, Belize, R. Hondo; Guatemala, El Reposo, Cubilguitz). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). DISTRIBUTION: Guatemala, Belize. binaluanensis Bernhauer, 1929, see: Nacaeus. binotatus Fauvel, 1904b: 81 (Lispinus; Type locality: Nouvelle-Guinee: baie de l’Astrolabe: Stephansort; Ile Deslacs). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). DISTRIBUTION: New Guinea. birmanus Fauvel, 1895, see: Nacaeus. bistriatus Fauvel, 1895: 185 (Lispinodes; Type locality: Bírmanie, Bhamo; Annam, Hue; Sikkim, Darjeeling). — Fauvel, 1904a: 44 (Lispinodes; Nilghiris; Sikkim; Bírmanie; Annam; Tonkin). — Bernhauer and Schubert, 1910: 19 (Lispinodes; catalog). Bulletin AMNH OSORIINAE: LISPINUS 1255
— Cameron, 1928d: 96 (Pseudolispinodes; Sumatra). — Cameron, 1930: 66 (Pseudolispinodes; characters; Burma; Sikkim; India; Bihar; Sumatra; Indochina; Philippines). — Scheerpeltz, 1933: 1010 (Pseudolispinodes; catalog). — Blackwelder, 1942: 80, 90 (Lispinus; transferred from Pseudolispinodes; checklist). — Abdullah and Qadri, 1970: 114 (Pseudolispinodes; characters; West Pakistan). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Coiffait, 1981f: 332 (Pseudolispinus; Andaman Islands). DISTRIBUTION: Pakistan, India, Andaman Islands, Myanmar, Vietnam, Indonesia, Philippines. boleticola Bernhauer, 1929i: 355 (Lispinus; Type locality: Philippinen, St. Michael). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. bolivianus Bernhauer, 1929i: 346 (Lispinus; replacement name for opacipennis Bernhauer). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). — Irmler, 1994: 60 (Lispinus; characters; Bolivia; Colombia; Surinam; Paraguay; Panama). DISTRIBUTION: Bolivia, Panama, Colombia, Surinam, Bolivia, Paraguay. opacipennis Bernhauer, 1921b: 66 (Lispinus; [preoccupied]; Type locality: Bolivien: Yuracaris). — Scheerpeltz, 1933: 1013 (Lispinus; synonym of bolivianus). — Irmler, 1994: 60 (Lispinus; synonym of bolivianus). borneensis Cameron, 1930b: 160 (Lispinus; Type locality: North Borneo; Bettotan near Sandakan). DISTRIBUTION: Borneo. boxi Blackwelder, 1943: 137 (Lispinus; Type locality: St. Lucia, upper Roseau Valley, 1 mile south of bridge over Millette River just above its junction with the Roseau River, elevation 750 feet). — Legner and Moore, 1977: 174 (Lispinus; found under bark). DISTRIBUTION: West Indies. brasilianus Irmler, 1994: 65 (Lispinus; Type locality: Teresopolis, S. Catharina, Brasilien). DISTRIBUTION: Brazil. breviceps Bernhauer, 1905: 9 (Lispinus; Type locality: Mexiko). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). DISTRIBUTION: Mexico. brevicollis Fauvel, 1864b: 52 [= 1865: 56] (Lispinus; Type locality: Mexique: Teapa). — Solsky, 1872: 313 (Lispinus; Cayenne). — Sharp, 1887: 722 (Lispinus; Mexico; Guatemala; Nicaragua). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). — Drugmand, 1998a: 223 (Tannea; type in l’Institut royal des Sciences naturelles de Belgique). DISTRIBUTION: Mexico, Guatemala, Nicaragua. brevicornis Kraatz, 1859: 187 (Lispinus; Type locality: India orientalis). — Bernhauer, 1902b: 45 (Lispinus; Ceylon). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Cameron, 1930: 57 (Lispinus; characters; India). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: India, Sri Lanka, “East Indies”. brevipennis Bernhauer, 1929i: 356 (Lispinus; Type locality: Mindoro, San Theodoro). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. brevisulcatus Bernhauer, 1929i: 340 (Lispinus; Type locality: Palawan: Binaluan). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. 1256 OSORIINAE: LISPINUS Bulletin AMNH bruchi Bernhauer, 1921d: 170 (Lispinus; Type locality: Argentinien: Prov. Catamarca). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Argentina. brunneorufus Bernhauer, 1929, see: opaculus Bernhauer, 1929. bryanti Cameron, 1925b: 237, 243 (Lispinus; Type locality: Penang). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Malaysia. burgeoni Cameron, 1928: 12 (Lispinus; Type locality: Belgian Congo: Arebi, Bondo Mabe). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Congo. burunganus Scheerpeltz, 1933: 1013 (Lispinus; replacement name for vulneratus Bernhauer). — Scheerpeltz, 1956a: 482 (Lispinus; French Congo; Belgian Congo). DISTRIBUTION: Congo. vulneratus Bernhauer, 1932a: 74 (Lispinus; [preoccupied]; Type locality: Kivu: Burunga). — Scheerpeltz, 1933: 1013 (Lispinus; synonym of burunganus). butacensis Bernhauer, 1929i: 346 (Lispinus; Type locality: Luzon, Butac). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. caledonicus Fauvel, 1878g: 481 (Lispinus; Type locality: Nouvelle-Calédonie). — Fauvel, 1889b: 244 (Lispinus; New Caledonia). — Fauvel, 1903d: 256 (Lispinus; New Caledonia; île des Pins). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). DISTRIBUTION: New Caledonia, Isle of Pines. californicus LeConte, 1863, see: Clavilispinus. cameroni Scheerpeltz, 1933: 1013 (Lispinus; replacement name for subcoriaceus Cameron). DISTRIBUTION: Borneo. subcoriaceus Cameron, 1928g: 433 (Lispinus; [preoccupied]; Type locality: Borneo: Tutau River). — Cameron, 1928f: 400 (Lispinus; Borneo). — Scheerpeltz, 1933: 1013 (Lispinus; synonym of cameroni). canalis Irmler, 1994: 70 (Lispinus; Type locality: Panama: Canal Zone: Barro Colorado). DISTRIBUTION: Panama. canarii Cameron, 1945: 64 (Lispinus, cited as Lispinis; Type locality: N. Andamans). DISTRIBUTION: Andaman Islands. capensis Bernhauer, 1927: 47 (Lispinus; Type locality: Kapprovinz: Queenstown, 3500 Fuss). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). — Bernhauer, 1934h: 483 (Lispinus; characters in key). DISTRIBUTION: South Africa. carinicollis Bernhauer, 1929i: 346 (Lispinus; Type locality: Mindanao, Kolambugan). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. castaneus Fauvel, 1878, see: Nacaeus. catena Sharp, 1876: 412 (Lispinus; Type locality: Ega). — Sharp, 1887: 719 (Lispinus; notes; Nicaragua). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Blackwelder, 1943: 132 (Lispinus; characters; Trinidad). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Irmler, 1994: 58 (Lispinus; characters; Brazil; Surinam; Bolivia; Ecuador; Colombia; Panama). DISTRIBUTION: Trinidad, Surinam, Brazil, Bolivia, Ecuador, Colombia, Panama, Nicaragua. Bulletin AMNH OSORIINAE: LISPINUS 1257 cavipennis Fauvel, 1898c: 177 (Lispinus; Type locality: Suberbieville). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). DISTRIBUTION: Madagascar. cheesmani Cameron, 1937a: 87 (Lispinus; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft.). — Cameron, 1943a: 337 (Lispinus; characters in key). — Scheerpeltz, 1957c: 223, 240 (Lispinus; subgenus Aulacolispinus; Sumba). DISTRIBUTION: New Guinea. claviger Cameron, 1913, see: Nacaeus. coarcticollis Kraatz, 1859, see: Nacaeus. cognatus Sharp, 1876, see: linearis Erichson, 1840. collaris Bernhauer, 1929i: 354 (Lispinus; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). DISTRIBUTION: Philippines. complanus Bernhauer, 1929i: 354 (Lispinus; subspecies of collaris; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1013 (Lispinus; subspecies of collaris). collarti Cameron, 1933: 35 (Lispinus; Type locality: Belgian Congo: Stanleyville). DISTRIBUTION: Congo. colossus Bernhauer, 1902, see: Ischiopsaurus. complanus Bernhauer, 1929, see: collaris Bernhauer, 1929. congoensis Bernhauer, 1929, see: leonensis Bernhauer, 1929. conradsi Bernhauer, 1937b: 578 (Lispinus; Type locality: Ukerewe Insel: Muruntuguru). DISTRIBUTION: Tanzania. cooki Blackwelder, 1942, see: Nacaeus. cooperi Bernhauer, 1931d: 566 (Lispinus; Type locality: Abyssinia: Jem-Jem Forest, 8,000 feet or above). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Ethiopia. cordilliensis Irmler, 1994: 68 (Lispinus; Type locality: Kolumbien: Cali). DISTRIBUTION: Colombia. cordobensis Bernhauer, 1929h: 187 (Lispinus; Type locality: Mexiko: Cordoba, La Capilla). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). — Irmler, 1994: 59 (Lispinus; characters; Mexico; Guadeloupe; Costa Rica; Guatemala; Cuba; St. Vincent; Grenada; St. Domingo). DISTRIBUTION: Mexico, Guatemala, Costa Rica, West Indies. coriaceus Fauvel, 1895: 185 (Lispinus; Type locality: Bírmanie, Carin Asciuii Cheba, 1200–1300 m; Carin Asciuii Ghécu, 1400–1500 m; Bhamo; Tenasserim, montagnes entre Meekalan et Kyeat, 1000–1400 m; Pulo Penang; Java, monts Tengger, 1100 m, Tcibodas). — Fauvel, 1905e: 78 (Lispinus; Java; Malacca; Pulo Penang; Tenasserim; Burma; Tonkin). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Bernhauer, 1929i: 356 (Lispinus; characters in key; Philippines; Burma; Indonesia). — Cameron, 1930: 62 (Lispinus; characters; Burma; Java; Borneo). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). — Coiffait, 1978b: 112 (Lispinus; Bhutan). — Coiffait, 1982: 41 (Lispinus; Nepal). DISTRIBUTION: Nepal, Bhutan, Myanmar, Vietnam, Borneo, Indonesia, Philippines. coripennis Irmler, 1994: 69 (Lispinus; Type locality: Rio de Janeiro). DISTRIBUTION: Brazil. coryndoni Cameron, 1942b: 322 (Lispinus; Type locality: Chyulu Hills, altitude 5600 feet). DISTRIBUTION: Kenya. 1258 OSORIINAE: LISPINUS Bulletin AMNH costaricensis Irmler, 1994: 64 (Lispinus; Type locality: Costa Rica: Carpintera, 1600 m. Also cited from Panama, Guatemala, Mexico, Honduras, Bolivia, Ecuador, Brazil, Surinam, and French Guiana). DISTRIBUTION: Mexico, Honduras, Guatemala, Costa Rica, Panama, Honduras, Surinam, French Guiana, Brazil, Ecuador, Bolivia. crassicornis Schubert, 1911: 1 (Lispinus; Type locality: Antananarivo, Madagascar). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Madagascar. crenicollis Irmler, 1994: 69 (Lispinus; Type locality: Panama: Chiriqui, E. Escopeta). DISTRIBUTION: Panama. cubensis Irmler, 1994: 63 (Lispinus; Type locality: Cuba: Sierra Rampe). DISTRIBUTION: Cuba. curticollis Bernhauer, 1904, see: tristis Bernhauer, 1929. curticollis Fauvel, 1878f: 204 (Lispinus; Type locality: Iles Key). — Fauvel, 1879: 76 (Lispinus; Gilolo; Pulo Pinang). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Cameron, 1927c: 260 (Lispinus; New Caledonia). — Cameron, 1931: 356 (Lispinus; New Guinea). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Vietnam, Indonesia, New Guinea, New Caledonia. curtipennis Bernhauer, 1929, see: nitidipennis Bernhauer, 1914. curtipennis Bernhauer, 1929, see: Nacaeus. danforthi Blackwelder, 1943, see: Nacaeus. dapanus Bernhauer, 1929i: 339 (Lispinus; Type locality: Siargao, Dapa). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Philippines. dejectus Sharp, 1887: 721 (Lispinus; Type locality: Guatemala, Purula, San Geronimo, Zapote). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Scheerpeltz, 1953: 601 (Lispinus; Guatemala). DISTRIBUTION: Guatemala. depressus Sharp, 1876: 417 (Lispinus; Type locality: Ega). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). DISTRIBUTION: Brazil. difficilis Cameron, 1948a: 224 (Lispinus; Type locality: Ivory Coast: Réserve forestière du Banco). DISTRIBUTION: Ivory Coast. dipterocarpi Cameron, 1930: 452 (Lispinus; Type locality: Middle Andaman). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Andaman Islands. distinctus Cameron, 1933b: 339 (Lispinus; Type locality: British North Borneo: Lumu Lumu, 5500 feet). DISTRIBUTION: Borneo. distinguendus Eichelbaum, 1913: 116 (Lispinus; Type locality: Amani, Berg Bomole). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Tanzania. dollmani Bernhauer, 1934h: 483 (Lispinus; Type locality: N.W. Rhodesia: Mwengwa, 27 40 E., 13 S.; Shigariatombwes). DISTRIBUTION: Zambia. Bulletin AMNH OSORIINAE: LISPINUS 1259 drescheri Cameron, 1936a: 29 (Lispinus; Type locality: Karang Pengalen, Preanger, alt. 750 m). DISTRIBUTION: Indonesia. dubius Bernhauer, 1929, see: medius Bernhauer, 1929. dundoensis Cameron, 1950f: 111 (Lispinus; Type locality: Angola: Dundo: Forêt de la Luachimo). DISTRIBUTION: Angola. angolanus Cameron, 1950f: 111 (Lispinus; variety of dundoensis; Type locality: Angola: Dundo: Forêt de la Luachimo). elongatus Bernhauer, 1904: 12 (Lispinus; Type locality: Sumatra, Si-Rambé). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Cameron, 1928d: 96 (Lispinus; Sumatra). DISTRIBUTION: Indonesia. elongatus Irmler, 1994: 63 (Lispinus; [preoccupied]; Type locality: St. Thomas). — Herman, 2001: 40 (Lispinus; name will be replaced by Irmler). DISTRIBUTION: West Indies. enganoensis Bernhauer, 1929, see: Nacaeus. excellens Bernhauer, 1916b: 18 (Lispinus; Type locality: Java: Telega-Patengan). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Indonesia. exiguus Erichson, 1840, see: Clavilispinus. externepunctus Fauvel, 1903e: 235 (Lispinus; Type locality: Cameroun; Gabon: Libreville). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Cameron, 1952e: 828 (Lispinus; Togo). DISTRIBUTION: Togo, Cameroon, Gabon. fauveli Sharp, 1887, see: Nacaeus. femoralis Blackwelder, 1942, see: Nacaeus. ferrugineus Bernhauer, 1915d: 96 (Lispinus; Type locality: Deutsch-Ostafrika: West-Usambara). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Tanzania. fibulatus Bernhauer, 1932a: 75 (Lispinus; Type locality: Bas-Uele: Koteli). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). — Cameron, 1952e: 828 (Lispinus; Dahomey). DISTRIBUTION: Congo, Benin. flavipennis Fauvel, 1864, see: Nacaeus. formosae Bernhauer, 1922c: 222 (Lispinus; variety of longipennis; Type locality: Suisharyo, Taihorin). — Scheerpeltz, 1933: 1015 (Lispinus; variety of longipennis). — Cameron, 1949b: 458 (Lispinus; cited as valid species; Formosa). DISTRIBUTION: Taiwan. formosanus Cameron, 1949b: 458 (Lispinus; Type locality: Formosa: Taihorin). — Sawada, 1965a: 147 (Lispinus; Formosa). DISTRIBUTION: Taiwan. foveatus Fauvel, 1878, see: Nacaeus. foveiventris Bernhauer, 1929i: 348 (Lispinus; Type locality: Palawan, Binaluan. Mindanao, Port Banga). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Philippines. 1260 OSORIINAE: LISPINUS Bulletin AMNH foveolus Blackwelder, 1943, see: Nacaeus. franckei Wendeler, 1955, see: Holotrochus. fraternus Cameron, 1929b: 438 (Lispinus; Type locality: Selangor: The Gap. Java: Buitenzorg). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Malaysia, Indonesia. fulgens Bernhauer, 1941c: 277 [= 1951: 267] (Lispinus; Type locality: Süd-Peru: Sivia, 520 m). DISTRIBUTION: Peru. fulgidus Cameron, 1940c: 210 (Lispinus; Type locality: Ghum District: Rongdong Valley). DISTRIBUTION: India. fulvescens Wendeler, 1955: 190 (Lispinus; Type locality: Brazilien: Nova Teutonia). DISTRIBUTION: Brazil. fulvus Motschulsky, 1857, see: Nacaeus. funebris Bernhauer, 1921b: 66 (Lispinus; Type locality: Bolivien: Yuracaris, Rio Beni, La Paz-Reyes). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Bolivia. fungosus Herman, 2001: 40 (Lispinus; replacement name for puncticollis Bernhauer, 1929b: 84). DISTRIBUTION: Congo, Cameroon. puncticollis Bernhauer, 1929b: 84 (Lispinus; [preoccupied]; Type locality: Kunungu; Luebo; Kasa Ngombe). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). — Scheerpeltz, 1956a: 482 (Lispinus; French Congo; Belgian Congo). — Scheerpeltz, 1971c: 162 (Lispinus; Cameroon). — Herman, 2001: 40 (Lispinus; synonym of fungosus). galapagosus Coiffait, 1981g: 308 (Pseudolispinodes; subgenus Pseudolispinodes; Type locality: Ile Galapagos: San Cristobal, Station Ch. Darwin). DISTRIBUTION: Galapagos Islands. germanus Cameron, 1937a: 87 (Lispinus; Type locality: New Guinea: Kokoda, alt. 1200 ft.). — Cameron, 1943a: 337 (Lispinus; characters in key). — Scheerpeltz, 1957c: 240 (Lispinus; subgenus Aulacolispinus). DISTRIBUTION: New Guinea. ghesquierei Cameron, 1939: 1 (Lispinus; Type locality: Belgian Congo: Mokanga, Lacs Mokoto). DISTRIBUTION: Congo. gigantulus Bernhauer, 1932a: 74 (Lispinus; Type locality: Haut-Uele: Moku Moto; Moto. Bas-uele: Koteli). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). — Bernhauer, 1942c: 348 (Lispinus; Cameroon). DISTRIBUTION: Congo, Cameroon. glabripennis Bernhauer, 1937: 289 (Lispinus; Type locality: Kenya: Ngong, K.C.). DISTRIBUTION: Kenya. globiceps Cameron, 1928, see: Nacaeus. granadensis Fauvel, 1864, see: insularis Fauvel, 1863. guadeloupae Blackwelder, 1943, see: Nacaeus. guamensis Bernhauer, 1942, see: Nacaeus. hewitti Bernhauer, 1914c: 83 (Lispinus; Type locality: Sarawak; Sumatra: Palembang). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). DISTRIBUTION: Malaysia, Indonesia. Bulletin AMNH OSORIINAE: LISPINUS 1261 hintzi Bernhauer, 1915k: 287 (Lispinus; Type locality: Kamerun: Musake). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). — Bernhauer, 1942c: 347 (Lispinus; Cameroon). — Fagel, 1972: 97 (Lispinus; notes; Sainte Helene). DISTRIBUTION: Cameroon, St. Helena. holosinus Fauvel, 1908: 95 (Lispinus; Type locality: Amani). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Eichelbaum, 1913: 116 (Lispinus; Tanzania). DISTRIBUTION: Tanzania. impar Cameron, 1913, see: Nacaeus. imperialis Bernhauer, 1929b: 82 (Lispinus; Type locality: Haut Uele: Watsa; Moto). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). — Scheerpeltz, 1956a: 482 (Lispinus; French Congo; Belgian Congo). — Levasseur, 1968a: 50 (Lispinus; Central African Republic). DISTRIBUTION: Congo, Central African Republic. impressicollis Motschulsky, 1857, see: Nacaeus. impressithorax Fairmaire, 1849: 290 (Lispinus; Type locality: Taïti). — Fauvel, 1877: 186 (Lispinus; characters; Tahiti). — Fauvel, 1878g: 481 (Lispinus; characters). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Bernhauer, 1922a: 149 (Lispinus; Samoa). — Coiffait, 1976: 220 (Lispinus; Tahiti). DISTRIBUTION: Society Islands, Samoa. imuganensis Bernhauer, 1929i: 347 (Lispinus; Type locality: Luzon, Imugan, Los Baños). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Philippines. insignicollis Fauvel, 1904d: 293 (Lispinus; Type locality: Usambara: Derema, 850 m). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Eichelbaum, 1913: 116 (Lispinus; Tanzania). DISTRIBUTION: Tanzania. insularis Fauvel, 1863: 442 (Lispinus; Type locality: Santo Domingo; Cuba). — Fauvel, 1864b: 49 [= 1865: 53] (Lispinus; characters; Dominican Republic; Cuba; Mexico). — Solsky, 1868a: 144 (Lispinus; Mexico). — Sharp, 1887: 720 (Lispinus; variety of granadensis; characters; Mexico; Guatemala; Antilles). — Fleutiaux and Sallé, 1889: 382 (Lispinus; Guadeloupe Island). — Fauvel, 1901: 70 (Lispinus; Colombia; Mexico; Nicaragua; Guatemala; Cuba; Santo Domingo; Guadeloupe; Saint Vincent; Grenada; Paraguay). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Blackwelder, 1943: 136 (Lispinus; characters; Puerto Rico; Montserrat; Guadeloupe; Dominica; St. Vincent; Grenada). — Irmler, 1994: 63 (Lispinus; characters; Panama; Mexico; Costa Rica; Cuba; Dominica). DISTRIBUTION: West Indies, Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Peru Paraguay. granadensis Fauvel, 1864b: 48 [= 1865: 52] (Lispinus; Type locality: Grenade, Nicaragua). — Solsky, 1872: 313 (Lispinus; Cayenne; Peru). — Sharp, 1887: 719 (Lispinus; notes Mexico; Guatemala; Nicaragua). — Fauvel, 1901: 70 (Lispinus; synonym of insularis). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Paulian, 1941: 142 (Lispinus; [Note: Cited as grandiceps Fauvel; there is not such species name by Fauvel but grandiceps may be an error for granadensis Fauvel which is from the gen- eral region cited by Paulian]; larval characters; Panama, cited as Barro Colorado, Costa Rica). — Blackwelder, 1943: 136 (Lispinus; synonym of insularis). — Irmler, 1994: 63 (Lispinus; synonym of insularis). 1262 OSORIINAE: LISPINUS Bulletin AMNH insularum Bernhauer, 1929i: 345 (Lispinus; Type locality: Luzon, Paete, Zambales, Mount Banahao, Bayombong, Manila. Mindoro, San Theodoro). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Philippines. intermedius Cameron, 1925b: 236, 239 (Lispinus; Type locality: Java, Mt. Tengger; Perak). — Cameron, 1930: 59 (Lispinus; characters; Burma; Java; Sumatra; Perak). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Myanmar, Malaysia, Indonesia. iridescens Bernhauer, 1942, see: Nacaeus. irregularis Blackwelder, 1943, see: Nacaeus. isolatus Cameron, 1949b: 459 (Lispinus; Type locality: Formosa: Kuraru). DISTRIBUTION: Taiwan. jacobsoni Cameron, 1928d: 96 (Lispinus; Type locality: Sumatra: Gunung Singgalang). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Indonesia. jambar Naomi, 1996a: 763 (Lispinus; Type locality: Oku, Kunigami, Yanbaru, Okinawa Is., Nansei Isls.). DISTRIBUTION: Japan. japonicus Cameron, 1930, see: Nacaeus. jocator Tottenham, 1953b: 482 (Lispinus; Type locality: Kilimanjaro, south side, 7500 ft). DISTRIBUTION: Tanzania. jyeri Bernhauer, 1914, see: Nacaeus. kamerunensis Bernhauer, 1929b: 83 (Lispinus; Type locality: Kamerun: Lolodorf). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). — Bernhauer, 1942c: 348 (Lispinus; Cameroon). DISTRIBUTION: Cameroon. kempi Bernhauer, 1926, see: quadrinotatus Fauvel, 1904. kenyanus Herman, 2001: 40 (Lispinus; replacement name for alutaceipennis Bernhauer). DISTRIBUTION: Kenya. alutaceipennis Bernhauer, 1937: 289 (Lispinus; [preoccupied]; Type locality: Kenya: Chania). — Herman, 2001: 40 (Lispinus; synonym of kenyanus). kiwuensis Scheerpeltz, 1933: 1015 (Lispinus; [Note: Cameron (1930d: 405) published a description that lacked generic and specific names but included a type locality and comparison to two species of Lispinus. The name “kiwuensis” was supplied by Scheerpeltz (1933: 1015) and attributed to Cameron, 1930. No evidence is presented that Cameron originated the name. The name was made available by Scheerpeltz therefore he is the author of the name and the date of publication is 1933 (Articles 10.1, 13.1.2, and 50.1)]; Type locality: Kivu: Burunga). DISTRIBUTION: Congo. kokodanus Cameron, 1937a: 88 (Lispinus; Type locality: New Guinea: Kokoda, alt. 1300 ft.). DISTRIBUTION: New Guinea. laetus Sharp, 1876: 417 (Lispinus; Type locality: St. Paulo). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). DISTRIBUTION: Brazil. laevicauda LeConte, 1866, see: Holotrochus. laevicollis Bernhauer, 1929i: 350 (Lispinus; Type locality: Siargao, Dapa). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Philippines. laevigatus Kraatz, 1859, see: Naeceus. laevigatus Bernhauer, 1906, see: laeviusculus Bernhauer, 1910. Bulletin AMNH OSORIINAE: LISPINUS 1263 laevior Fauvel, 1878f: 203 (Lispinus; Type locality: Nouvelle-Guinée, Hatam). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Bernhauer, 1926f: 312 (Lispinus; Buru). DISTRIBUTION: New Guinea, Indonesia. laevipennis Kraatz, 1859: 185 (Lispinus; Type locality: Ceylan). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Cameron, 1930: 54 (Lispinus; characters; Ceylon). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Sri Lanka. laeviusculus Bernhauer, 1910: 22 (Lispinus; replacement name for laevigatus Bernhauer). — Irmler, 1994: 61 (Lispinus; characters; Paraguay; Peru). DISTRIBUTION: Paraguay, Peru. laevigatus Bernhauer, 1906b: 193 (Lispinus; [preoccupied]; Type locality: Paraguay). — Irmler, 1994: 61 (Lispinus; synonym of laeviusculus). lagunae Bernhauer, 1929i: 351 (Lispinus; Type locality: Luzon, Laguna). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Philippines. laticollis Erichson, 1840: 828 (Lispinus; Type locality: Cuba). — Guérin-Méneville, 1857: 45 (Lispinus; characters). — Chevrolat and Fauvel, 1863: 442 (Lispinus; characters; Cuba, Puerto Rico). — Fauvel, 1864b: 45 [= 1865: 49] (Lispinus; characters; Cuba). — Sharp, 1887: 718 (Lispinus; Panama). — Fauvel, 1901: 70 (Lispinus; Colombia; Panama; Cuba). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Blackwelder, 1943: 131 (Lispinus; characters; Puerto Rico; Cuba). — Irmler, 1994: 58 (Lispinus; characters; Cuba; Panama; Costa Rica; Brazil). DISTRIBUTION: West Indies, Brazil, Panama, Costa Rica. latiusculus Bernhauer, 1926d: 260 (Pseudolispinodes; Type locality: Luzon, Provinz Nueva Vizcaya, Imugan). — Scheerpeltz, 1933: 1010 (Pseudolispinodes; catalog). DISTRIBUTION: Philippines. leai Cameron, 1943, see: Nacaeus. leonensis Bernhauer, 1929b: 84 (Lispinus; Type locality: Sierra Leone; Senegal). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Senegal; Sierra Leone; Congo. congoensis Bernhauer, 1929b: 84 (Lispinus; subspecies of leonensis; Type locality: Kongogebiet: Elisabethville). — Scheerpeltz, 1933: 1015 (Lispinus; synonym of leonensis). liepolti Bernhauer, 1937b: 578 (Lispinus; Type locality: Ukerewe Insel: Muruntuguru). — Scheerpeltz, 1956a: 483 (Lispinus; French Congo; Belgian Congo; Uganda; Kenya; Tanganyika). DISTRIBUTION: Congo, Uganda, Kenya, Tanzania. linearis Erichson, 1840: 829 (Lispinus; Type locality: Puerto Cabello, Columbia). — Fauvel, 1878d: 177 [= 1878e: 13] (Lispinus; characters; Arizona; Texas; Mexico; Colombia). — Fauvel, 1891: 89 (Lispinus; notes; Venezuela). — Fall and Cockerell, 1907: 168 (Lispinus; New Mexico). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Blatchley, 1910: 484 (Lispinus; Ohio). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). — Irmler, 1994: 59 (Lispinus; characters; Guadeloupe; Trinidad; Mexico; Guatemala; Panama; British Guiana; Surinam; Venezuela; Ecuador; Colombia; Brazil; Bolivia; Paraguay). DISTRIBUTION: Paraguay, Bolivia, Brazil, Ecuador, Colombia, Venezuela, Guyana, Surinam, West Indies, Panama, Guatemala, Mexico, USA. 1264 OSORIINAE: LISPINUS Bulletin AMNH
punctatus Sharp, 1876: 414 (Lispinus; Type locality: Ega). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Irmler, 1994: 59 (Lispinus; synonym of linearis). modestus Sharp, 1876: 415 (Lispinus; Type locality: Ega). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Irmler, 1994: 59 (Lispinus; synonym of linearis). cognatus Sharp, 1876: 415 (Lispinus; Type locality: Ega). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Irmler, 1994: 59 (Lispinus; synonym of linearis). aremicus Blackwelder, 1943: 131 (Lispinus; Type locality: Trinidad: Arima-Paria road). — Irmler, 1994: 59 (Lispinus; synonym of linearis). lineatopunctatus Bernhauer, 1929i: 340 (Lispinus; Type locality: Mindanao, Dapitan. Luzon: Haight’s Place; Trinidad; Mount Polis; Laguna). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Philippines. montanus Bernhauer, 1929i: 340 (Lispinus; variety of lineatopunctatus; Type locality: Luzon: Mount Polis, Laguna). — Scheerpeltz, 1933: 1015 (Lispinus; variety of lineatopunctatus). lineipennis Fauvel, 1878f: 202 (Lispinus; Type locality: Nouvelle-Guinée, Hatam; Java, Tcibodas). — Fauvel, 1879: 76 (Lispinus; Mysol). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Bernhauer, 1920: 3 (Lispinus; Australia). — Cameron, 1943a: 337 (Lispinus; characters in key). — Scheerpeltz, 1957c: 223, 240 (Lispinus; subgenus Aulacolispinus; Sumba). — Newton, 1990: 92 (Lispinus). DISTRIBUTION: Indonesia, New Guinea, Australia. lispinoides Blackwelder, 1942, see: Nacaeus. listenbarthi Irmler, 1994: 66 (Lispinus; [Note: Cited erroneously as listanbarthi on page 56]; Type locality: Brasilien: Rio de Janeiro). DISTRIBUTION: Brazil. longesulcatus Bernhauer, 1929i: 342 (Lispinus; Type locality: Luzon, Mount Banahao). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Philippines. longicollis Bernhauer, 1929i: 339 (Lispinus; Type locality: Borneo, Sandakan). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Borneo. longicornis Sharp, 1887: 722 (Lispinus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Bierig, 1931: 427 (Lispinus; characters; Panama). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Panama. longipennis Bernhauer, 1915i: 251 (Lispinus; Type locality: Java: Telaga Patengan). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Indonesia. longipennis Bernhauer, 1926, see: luzonensis Herman, 2001. longulus Sharp, 1874, see: Nacaeus. longus Cameron, 1933b: 338 (Lispinus; Type locality: British North Borneo: Lumu Lumu, 5500 feet). DISTRIBUTION: Borneo. Bulletin AMNH OSORIINAE: LISPINUS 1265 lucens Bernhauer, 1906c: 322 (Lispinus; Type locality: Brasilia: S. Catharina). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). — Irmler, 1994: 62 (Lispinus; characters; Brazil). DISTRIBUTION: Brazil. lunaris Irmler, 1994: 70 (Lispinus; Type locality: Surinam: Marowijne Dist. Anapaike village). DISTRIBUTION: Surinam. luzonensis Herman, 2001: 40 (Lispinus; replacement name for longipennis Bernhauer, 1926). DISTRIBUTION: Philippines. longipennis Bernhauer, 1926d: 259 (Pseudolispinodes; [preoccupied]; Type locality: Luzon, Provinz Mountain, Mount Polis). — Scheerpeltz, 1933: 1010 (Pseudolispinodes; catalog). — Blackwelder, 1942: 80 (Lispinus). — Herman, 2001: 40 (Lispinus; synonym of luzonensis). luzonicus Bernhauer, 1929, see: Nacaeus. macerrimus Bernhauer, 1942c: 348 (Lispinus; Type locality: Mont Cameroun: 1800–2000 m., versant Sud Est.). DISTRIBUTION: Cameroon. macropterus Fauvel, 1904b: 78 (Lispinus; Type locality: Celebes sud: Bua-Kraeng, 1700 m). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Scheerpeltz, 1957c: 240 (Lispinus; subgenus Aulacolispinus). DISTRIBUTION: Sri Lanka. madurensis Bernhauer, 1915, see: madurensis Bernhauer, 1914. madurensis Bernhauer, 1914c: 83 (Holosus; Type locality: Chambaganoor, Madura, Ost- indien). — Bernhauer, 1926d: 260 (Pseudolispinodes; characters; Philippines). — Cameron, 1928d: 96 (Pseudolispinodes; synonym of bistriatus). — Cameron, 1930: 66 (Pseudolispinodes; synonym of bistriatus). — Scheerpeltz, 1933: 1010 (Pseudolispinodes; synonym of bistriatus). — Blackwelder, 1942: 80, 81 (Pseudolispinodes; discussion of status of name). DISTRIBUTION: India, Borneo, Philippines. madurensis Bernhauer, 1915i: 251 (Lispinus; [preoccupied]; Type locality: Chambaganoor, Madura, Südindien). — Cameron, 1930: 59 (Lispinus; characters; India). — Cameron, 1930b: 160 (Lispinus; British North Borneo). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). — Blackwelder, 1942: 80, 81 (Pseudolispinodes; synonym of madurensis). magnificus Bernhauer, 1929i: 340 (Lispinus; Type locality: Luzon: Balbalasang). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Philippines. malaisei Scheerpeltz, 1965: 123 (Lispinus; subgenus Lispinus; Type locality: British Sikkim, 200 m, Valley at Tista Bridge). DISTRIBUTION: India. malayanus Cameron, 1925b: 236, 239 (Lispinus; Type locality: Selangor, The Gap, Bukit Kutu [Federated Malay States]). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). DISTRIBUTION: Malaysia. manilensis Bernhauer, 1929i: 350 (Lispinus; Type locality: Luzon, Manila). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. medius Bernhauer, 1929i: 341 (Lispinus; Type locality: Luzon: Bangui). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. 1266 OSORIINAE: LISPINUS Bulletin AMNH
dubius Bernhauer, 1929i: 341 (Lispinus; subspecies of medius; Type locality: Luzon, Bangui). — Scheerpeltz, 1933: 1016 (Lispinus; variety of medius). punctus Bernhauer, 1929i: 341 (Lispinus; subspecies of medius; Type locality: Luzon: Benguet, Baguio). — Scheerpeltz, 1933: 1016 (Lispinus; variety of medius). methneri Bernhauer, 1937b: 577 (Lispinus; Type locality: Deutsch-Ostafrika: Unt. Rufiyi; Usaromo, Puguberg; Narobi b., Tango). DISTRIBUTION: Tanzania, Kenya. microcephalus Fauvel, 1866e: 319 (Lipinus; Type locality: La Réunion: Bourbon). — Fauvel, 1867b: 70 [= 1868a: 74] (Lispinus; characters; Réunion Islands). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Jarrige, 1957: 112 (Lispinus; Réunion). DISTRIBUTION: Mascarene Islands. mindanaoanus Bernhauer, 1929i: 347 (Lispinus; Type locality: Mindanao, Mumungan). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. minor Bernhauer, 1929b: 84 (Lispinus; Type locality: Ituri: La Moto-Madyn; Mayumbe: Makala Ntete). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). — Scheerpeltz, 1956a: 483 (Lispinus; French Congo; Belgian Congo). DISTRIBUTION: Congo. minox Irmler, 1994: 69 (Lispinus; Type locality: Costa Rica: Tapanté). DISTRIBUTION: Costa Rica. minutus Cameron, 1918: 60 (Lispinus; Type locality: Singapore: Mandai). — Cameron, 1921a: 363, 399 (Lispinus; characters; Singapore). — Cameron, 1930b: 160 (Lispinus; British North Borneo). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Singapore, Borneo. mjobergi Cameron, 1928g: 432 (Lispinus; Type locality: Borneo: Mt. Dulit). — Scheerpeltz, 1933: 1016 (Lispinus; cited as mjöbergi; catalog). — Hammond, 1984: 197 (Lispinus; Borneo). — Herman, 2001: 12 (Lispinus; spelling should be mjobergi). DISTRIBUTION: Borneo. modestus Sharp, 1876, see: linearis Erichson, 1840. montanellus Bernhauer, 1929i: 349 (Lispinus; Type locality: Luzon, Mount Polis). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. montanus Bernhauer, 1929, see: lineatopunctatus Bernhauer, 1929. montanus Bernhauer, 1931, see: monticola Scheerpeltz, 1933. monticola Scheerpeltz, 1933: 1016 (Lispinus; replacement name for montanus Bernhauer). DISTRIBUTION: Ethiopia. montanus Bernhauer, 1931d: 565 (Lispinus; [preoccupied]; Type locality: Abyssinia: Jem-Jem Forest, 8,000–9,000 feet). — Scheerpeltz, 1933: 1016 (Lispinus; synonym of monticola). morugae Blackwelder, 1943, see: Nacaeus. natalensis Bernhauer, 1927: 48 (Lispinus; Type locality: Natal: Estcourt). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). — Bernhauer, 1934h: 483 (Lispinus; characters in key). DISTRIBUTION: South Africa. neglectus Cameron, 1950d: 2 (Lispinus; Type locality: Singapore). DISTRIBUTION: Singapore. Bulletin AMNH OSORIINAE: LISPINUS 1267 neotropicus Irmler, 1994: 66 (Lispinus; Type locality: Brasilien: São Paulo). DISTRIBUTION: Brazil. nevermanni Bernhauer, 1942: 3 (Lispinus; Type locality: Kostarika: Vulkan Irazu, Westabhang, 1800–2000 m). DISTRIBUTION: Costa Rica. newtoni Irmler, 1994: 66 (Lispinus; Type locality: Panama: Barro Colorado). DISTRIBUTION: Panama. nigerrimus Bernhauer, 1929i: 346 (Lispinus; Type locality: Masbate, Aroroy. Luzon, Los Baños. Leyte). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. nigrifrons Fauvel, 1863, see: Nacaeus. nitens Cameron, 1925b: 236, 238 (Lispinus; Type locality: Selangor, The Gap; Pahang, Raub, Federated Malay States). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Malaysia. nitidicollis Bernhauer, 1929b: 83 (Lispinus; Type locality: Haut-Uele: Watsa). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Congo. nitidipennis Bernhauer, 1914c: 81 (Lispinus; Type locality: Philippinen: Los Banos [sic]). — Bernhauer, 1929i: 343 (Lispinus; characters in key; Philippines). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). — Blackwelder, 1942: 89, 90 (cited part in Lispinus subgenus Lispinus and part in Pseudolispi- nodes subgenus Pseudolispinodes; [Note: Since it is unclear which “part” of nitidipennis is attributed to which genus I leave the species in Lispinus where it was described]; checklist). DISTRIBUTION: Philippines. curtipennis Bernhauer, 1929i: 343 (Lispinus; variety of nitidipennis; [preoccupied]; Type locality: Luzon: Mount Maquiling). — Scheerpeltz, 1933: 1016 (Lispinus; variety of nitidipennis). tenuipunctus Bernhauer, 1929i: 343 (Lispinus; Type locality: Siargao, Dapa). — Scheerpeltz, 1933: 1016 (Lispinus; variety of nitidipennis). nitidiventris Bernhauer, 1908c: 101 (Lispinus; Type locality: Puguberg bei Usaromo). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). DISTRIBUTION: Tanzania. nitidus Fauvel, 1878f: 203 (Lispinus; Type locality: Nouvelle-Guinée, Hatam). DISTRIBUTION: New Guinea. novoteutonius Wendeler, 1955: 191 (Lispinus; Type locality: Brasilien: Nova Teutonia). DISTRIBUTION: Brazil. obscurellus Fauvel, 1904e: 298 (Lispinus; Type locality: Baie d’Antongil; Tamatave; Fort Dauphin). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Bernhauer, 1922b: 166 (Lispinus; Seychelles). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Madagascar, Seychelles. obscurus LeConte, 1863a: 59 (Lispinus; Type locality: valley of Colorado and Gila River, Arizona). — LeConte, 1877: 250 (Lispinus; characters). — Fauvel, 1878d: 177 [= 1878e: 13] (Lispinus; synonym of linearis). — Fauvel, 1891: 89 (Lispinus; valid species). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). DISTRIBUTION: USA. 1268 OSORIINAE: LISPINUS Bulletin AMNH obsoletipennis Bernhauer, 1929i: 348 (Lispinus; Type locality: Luzon, Atimonan, Los Baños, Butac, Manila). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. obsoletus Bernhauer, 1921b: 67 (Lispinus; Type locality: Bolivien: Yuracaris; Paraguay). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Bolivia, Paraguay. opacinus Bernhauer, 1929i: 346 (Lispinus; Type locality: Mindanao, Kolambugan, Surigao. Basilan). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. opacipennis Bernhauer, 1915d: 96 (Lispinus; Type locality: Deutsch-Ostafrika: West Usambara). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Tanzania. opacipennis Bernhauer, 1921, see: bolivianus Bernhauer, 1929. opaculus Bernhauer, 1929i: 352 (Lispinus; Type locality: Luzon, Mount Banahao, Los Baños, Laguna, Imugan. Pollilo. Mindanao, San Theodoro. Mindanao, Surigao, Port Banga). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. brunneorufus Bernhauer, 1929i: 352 (Lispinus; form of opaculus; Type locality: Not cited). — Scheerpeltz, 1933: 1016 (Lispinus; variety of opaculus). opacus Fauvel, 1864b: 51 [= 1865: 55] (Lispinus; Type locality: Colombie). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). DISTRIBUTION: Colombia. osorioides Wendeler, 1955, see: Holotrochus. papuanus Cameron, 1937a: 88 (Lispinus; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft.). DISTRIBUTION: New Guinea. paradoxus Bernhauer, 1934h: 484 (Lispinus; Type locality: Pondoland). DISTRIBUTION: South Africa. paradoxus Cameron, 1945, see: prodigiosus Herman, 2001. parallelicollis Bernhauer, 1929i: 347 (Lispinus; Type locality: Masbate, Aroroy). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. parallelopipedus Bernhauer, 1929i: 345 (Lispinus; Type locality: Mollukken, Ternate). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Indonesia. parallelus Bernhauer, 1929i: 339 (Lispinus; Type locality: Borneo: Sandakan). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Borneo. paratardus Irmler, 1994: 67 (Lispinus; Type locality: Costa Rica: Esparta). DISTRIBUTION: Costa Rica. parcus Bernhauer, 1929i: 344 (Lispinus; Type locality: Mindanao, Port Banga). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. parvipennis Fauvel, 1866e: 318 (Lispinus; Type locality: La Réunion: Bourbon; Mayotte, Madagascar). — Fauvel, 1867b: 69 [= 1868a: 73] (Lispinus; characters; Réunion Island; Madagascar). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). DISTRIBUTION: Madagascar, Mascarene Islands. pauloensis Irmler, 1994: 71 (Lispinus; Type locality: São Paulo). DISTRIBUTION: Brazil. Bulletin AMNH OSORIINAE: LISPINUS 1269 pennatus Bernhauer, 1929, see: bernhauerianus Scheerpeltz, 1933. perniger Bernhauer, 1929i: 345 (Lispinus; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. picipennis Bernhauer, 1929i: 344 (Lispinus; Type locality: Luzon: Limay, Los Baños). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. planaticollis Bernhauer, 1929i: 348 (Lispinus; Type locality: Mindanao, Port Banga). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. planellus Sharp, 1887, see: Nacaeus. planicollis Bernhauer, 1929i: 342 (Lispinus; Type locality: Mindanao, Mumungan). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. planus Sharp, 1876: 416 (Lispinus; Type locality: Ega). — Fauvel, 1864b: 47 [= 1865: 51] (Lispinus; [Note: Misidentification: Bernhauer and Schubert, 1910: 23 cited linearis Fauvel as a synonym of planus; Fauvel did not describe the species as new, he attributed it to Erichson]; characters; Colombia; Mexico). — Sharp, 1887: 720 (Lispinus; notes; Mexico). — Fauvel, 1891: 89 (Lispinus; synonym of linearis). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Scheerpeltz, 1953: 601 (Lispinus; Guatemala). DISTRIBUTION: Brazil, Colombia, Guatemala, Mexico. politulus Fauvel, 1898a: 114 (Lispinus; Type locality: La Digue). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — H. Kolbe, 1910: 21 (Lispinus; Seychelles). — Bernhauer, 1922b: 165 (Lispinus; Seychelles). DISTRIBUTION: Seychelles. pondoensis Bernhauer, 1934h: 483 (Lispinus; Type locality: Pondoland: Port St. Johns; Zululand: Eshowe; Natal: Kloof, 1500 feet). — Herman, 2001: 13 (Lispinus; spelling corrected to pondoensis). DISTRIBUTION: South Africa. praenobilis Bernhauer, 1926c: 159 (Lispinus; Type locality: Fiji-Inseln). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Fiji. praenobilis Bernhauer, 1929, see: bernhaueri Scheerpeltz, 1933. prodigiosus Herman, 2001: 40 (Lispinus; replacement name for paradoxus Cameron). DISTRIBUTION: Indonesia. paradoxus Cameron, 1945: 141 (Pseudolispinodes; [preoccupied]; Type locality: Sumatra: Fort de Kock). — Herman, 2001: 40 (Lispinus; synonym of prodigiosus). propinquus Cameron, 1936d: 201 (Lispinus; Type locality: Mauritius: Les Mares). — Frank, 1982: 7 (Lispinus; list of parasites). DISTRIBUTION: Mascarene Islands. pseudolucens Irmler, 1994: 68 (Lispinus; Type locality: Costa Rica: Turrialba, cited as Turialba). DISTRIBUTION: Costa Rica. pubiventris Bernhauer, 1904: 10 (Lispinus; Type locality: Africa occidentalis, Gaboon, Libreville). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Eichelbaum, 1913: 116 (Lispinus; Tanzania). DISTRIBUTION: Gabon, Tanzania. 1270 OSORIINAE: LISPINUS Bulletin AMNH pulcher Bernhauer, 1942: 2 (Lispinus; Type locality: Kostarika: Reventazon, Ebene Limon, Hamburgfarm). DISTRIBUTION: Costa Rica. pumilio Bernhauer, 1929i: 349 (Lispinus; Type locality: Basilan). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. punctatellus Bernhauer, 1929i: 345 (Lispinus; Type locality: Luzon, Los Baños. Mindanao, Port Banga). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Philippines. punctatus Sharp, 1876, see: linearis Erichson, 1840. puncticeps Cameron, 1925b: 236, 241 (Lispinus; Type locality: Selangor, The Gap; Pahang, Bentong [Federated Malay States]). — Scheerpeltz, 1933: 1016 (Lispinus; catalog). DISTRIBUTION: Malaysia. puncticollis Bernhauer, 1926d: 260 (Pseudolispinodes; variety of sinuatus; Type locality: Not cited). — Scheerpeltz, 1933: 1010 (Pseudolispinodes; variety of sinuatus). — Cameron, 1949b: 457 (Pseudolispinodes; cited as valid species; China). DISTRIBUTION: China. puncticollis Bernhauer, 1929, see: punctiger Scheerpeltz, 1933. puncticollis Bernhauer, 1929, see: fungosus Herman, 2001. punctiger Scheerpeltz, 1933: 1017 (Lispinus; replacement name for puncticollis Bernhauer, 1929). DISTRIBUTION: Philippines. puncticollis Bernhauer, 1929i: 346 (Lispinus; [preoccupied]; Type locality: Luzon, Limay). punctipennis Bernhauer, 1926, see: sinuatus Bernhauer, 1904. punctipennis Cameron, 1925b: 236, 239 (Lispinus; Type locality: Borneo, Sarawak, Mt. Matang). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Borneo. punctithorax Bernhauer, 1929i: 355 (Lispinus; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Philippines. punctus Bernhauer, 1929, see: medius Bernhauer, 1929. quadratipennis Bernhauer, 1929i: 356 (Lispinus; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Philippines. quadratus Blackburn, 1885: 125 (Lispinodes; Type locality: mountains near Honolulu). — Blackburn and Sharp, 1885: 224 (Lispinodes; habitat; Hawaii). — Fauvel, 1902b: 47 (Lispinus; synonym of impressicollis). — Fauvel, 1903: 150 (Lispinus; synonym of impressicollis). — Fauvel, 1904a: 44 (Lispinus; synonym of impressicollis). — Sharp, 1908: 539 (Lispinus; synonym of impressicollis). — Bernhauer and Schubert, 1910: 22 (Lispinus; synonym of impressicollis). — Cameron, 1930: 60 (Lispinus; synonym of impressicollis). — Scheerpeltz, 1957c: 223 (Lispinus; synonym of impressicollis). — Moore, 1975c: 105 (Lispinus; valid species; Hawaii). — Nishida, 1994: 72 (Lispinus; Hawaii). — Nishida, 1997: 61 (Lispinus; Hawaii). DISTRIBUTION: Hawaii. Bulletin AMNH OSORIINAE: LISPINUS 1271 quadricollis Cameron, 1924b: 160 (Lispinus; Type locality: Almora, Kanari China, 4000 feet above sea). — Cameron, 1930: 55 (Lispinus; characters; India; Assam; Burma). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). — Abdullah and Qadri, 1970: 114 (Lispinus; characters; West Pakistan). DISTRIBUTION: Pakistan, India, Myanmar. quadrillum Fauvel, 1879: 76 (Lispinus; Type locality: Nouvelle Guinée, Dorei). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). DISTRIBUTION: New Guinea. quadrinotatus Fauvel, 1904b: 80 (Lispinus; Type locality: Perak; Java; Celebes sud: Bua-Kraeng, 1700 m). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Bernhauer, 1922c: 222 (Lispinus; Formosa). — Cameron, 1928d: 96 (Lispinus; Sumatra). — Cameron, 1930: 54 (Lispinus; characters; India; Perak; Java; Celebes). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). — Biswas and Biswas, 1985: 141 (Lispinus; India). DISTRIBUTION: India, Indonesia, Taiwan. kempi Bernhauer, 1926: 20 (Lispinus; Type locality: Nordost-Indien: Rotung, 1400 ft.). — Cameron, 1930: 54 (Lispinus; synonym of quadrinotatus). — Scheerpeltz, 1933: 1017 (Lispinus; synonym of quadrinotatus). quadripunctulus Fauvel, 1864b: 45 [= 1865: 49] (Lispinus; Type locality: Caracas, Colombie). — Solsky, 1872: 313 (Lispinus; Cayenne). — Sharp, 1887: 718 (Lispinus; Mexico; British Honduras; Guatemala; Nicaragua; Panama; Colombia; Venezuela). — Fauvel, 1891: 89 (Lispinus; Venezuela). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Blackwelder, 1952: 373 (Nacaeus; subgenus Tannea). — Scheerpeltz, 1957c: 241 (Lispinus; subgenus Stictolispinus; type species of subgenus). — Scheerpeltz, 1960c: 66 (Lispinus; subgenus Stictolispinus; Bolivia). — Irmler, 1994: 61 (Lispinus; characters; Venezuela; Colombia; Ecuador; Bolivia; Brazil; Panama; Costa Rica; Mexico; Guatemala). DISTRIBUTION: Brazil, Bolivia, Ecuador, Colombia, Venezuela, French Guiana, Panama, Costa Rica, Guatemala, Belize, Mexico. terminalis Sharp, 1876: 413 (Lispinus; Type locality: Para). — Bernhauer and Schubert, 1910: 24 (Lispinus; catalog). — Irmler, 1994: 61 (Lispinus; synonym of quadripunctulus). reversus Blackwelder, 1943, see: Nacaeus. rivularis Lea, 1906, see: Nacaeus. robusticollis Bernhauer, 1929, see: Nacaeus. robusticornis Cameron, 1925b: 236, 241 (Lispinus; Type locality: Selangor [Federated Malay States], The Gap, Bukit Kutu; Penang). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Malaysia. rotundicolis Bernhauer, 1929i: 349 (Lispinus; Type locality: Luzon, Los Baños). — Scheerpeltz, 1933: 1017 (Lispinus; cited as rotundicollis; catalog). DISTRIBUTION: Philippines. rubidus Cameron, 1925, see: Nacaeus. rufescens LeConte, 1863, see: Clavilispinus. 1272 OSORIINAE: LISPINUS Bulletin AMNH rufescens Cameron, 1950f: 117 (Holotrochus; [Note: Preoccupied as a secondary homonym]; Type locality: Angola: Dundo). — Fagel, 1959: 32 (Lispinus). DISTRIBUTION: Angola. rufosplendens Bernhauer, 1915g: 181 (Lispinus; Type locality: Neu-Lauenburg). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: New Guinea. rufotestaceus Cameron, 1940c: 211 (Lispinus; Type locality: Ghum District: Tiger Hill, Mangpu). DISTRIBUTION: India. rufulus Fauvel, 1905d: 194 (Lispinus; Type locality: Gabon: Libreville). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). DISTRIBUTION: Gabon. rufus Cameron, 1925b: 236, 237 (Lispinus; Type locality: Penang). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Malaysia. rugipennis Fauvel, 1904b: 80 (Lispinus; Type locality: Java: Tcibodas). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Cameron, 1936a: 29 (Lispinus; Java). DISTRIBUTION: Indonesia. rugosipennis Bernhauer, 1929i: 355 (Lispinus; Type locality: Luzon, Laguna). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Philippines. rugulipennis Bernhauer, 1934h: 483 (Lispinus; Type locality: Zululand: Eshowe). DISTRIBUTION: South Africa. rufus Fauvel, 1864 [Note: Nomen nudum], see: Clavilispinus. santoensis Bernhauer, 1934a: 18 (Lispinus; Type locality: Neue Hebriden: Santo). — Cameron, 1934a: 20 (Lispinus; New Hebrides). DISTRIBUTION: New Hebrides. sarawakensis Cameron, 1925b: 236, 242 (Lispinus; Type locality: Borneo, Mt. Matang; Pengaron, Martapoera). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Borneo. schedli Scheerpeltz, 1956a: 482, 515 (Lispinus; Type locality: Belgisch-Kongo: Hembe Bitale). DISTRIBUTION: Congo. schoutedeni Bernhauer, 1929b: 83 (Lispinus; Type locality: Kasai: Makumbi; Kai Bumba). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Congo. schusteri Irmler, 1994: 68 (Lispinus; Type locality: Brasilien: Petropolis, Rio de Janeiro). DISTRIBUTION: Brazil. sculpturatus Sharp, 1887: 721 (Lispinus; Type locality: Guatemala, near the city, San Geronimo). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). DISTRIBUTION: Guatemala. sculptus Kraatz, 1859, see: Tetrapleurus. secretus Cameron, 1925b: 236, 238 (Lispinus; Type locality: Selangor [Federated Malay States], The Gap). — Cameron, 1928d: 96 (Lispinus; Sumatra). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Malaysia, Indonesia. Bulletin AMNH OSORIINAE: LISPINUS 1273 selangorensis Cameron, 1925, see: Nacaeus. selangorensis Cameron, 1929, see: Nacaeus. semirufus Cameron, 1928: 12 (Lispinus; Type locality: Belgian Congo: Kivu, Burunga). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Indonesia. sericeiventris Bernhauer, 1914, see: Nacaeus. sericeus Coiffait, 1981, see: Nacaeus. setosus Cameron, 1918: 59 (Lispinus; Type locality: Singapore: Woodland). — Cameron, 1921a: 363, 399 (Lispinus; characters; Singapore). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Singapore. setosus Wendeler, 1955, see: Holotrochus. sharpi Cameron, 1920, see: Nacaeus. sidneensis Fauvel, 1878, see: Nacaeus. similis Cameron, 1925b: 236, 240 (Lispinus; Type locality: Penang). — Cameron, 1928d: 96 (Lispinus; Sumatra). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Malaysia. simplex Sharp, 1876: 417 (Lispinus; Type locality: Ega). — Fauvel, 1901: 70 (Lispinus; Colombia; Guyana; Amazonian region). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). DISTRIBUTION: Brazil, Guyana, Colombia. singaporensis Bernhauer, 1929i: 344 (Lispinus; Type locality: Singapore). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Singapore. singularis Gerstaecker, 1867, see: Neolosus. sinuatocollis Bernhauer, 1942: 2 (Lispinus; Type locality: Kostarika: Vara Blanca). DISTRIBUTION: Costa Rica. sinuatus Bernhauer, 1904a: 217 (Holosus; Type locality: Mentawei, Si Oban). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Bernhauer, 1926d: 260 (Pseudolispinodes; characters; Philippines). — Cameron, 1928d: 96 (Pseudolispinodes; Sumatra). — Scheerpeltz, 1933: 1010 (Pseudolispinodes; catalog). — Blackwelder, 1942: 80, 90 (Lispinus; transferred from Pseudolispinodes; checklist). DISTRIBUTION: Indonesia, Philippines. punctipennis Bernhauer, 1926d: 260 (Pseudolispinodes; variety of sinuatus; [preoccupied]; Type locality: Not cited). — Scheerpeltz, 1933: 1010 (Pseudolispinodes; variety of sinuatus). sobrinus Fauvel, 1864b: 43 [= 1865: 47] (Lispinus; Type locality: Caracas). — Fauvel, 1891: 89 (Lispinus; Venezuela). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Marcuzzi, 1977: 5 (Lispinus; Venezuela). — Irmler, 1994: 60 (Lispinus; [Note: Irmler synonymized sobrinus with Lispinus analis Erichson but in a letter to me (V-8-95) stated that analis was a manuscript name; the valid name of the species is sobrinus]; characters; Colombia; Bolivia; Venezuela; Brazil). DISTRIBUTION: Brazil, Bolivia, Colombia, Venezuela. socialis Irmler, 1994: 65 (Lispinus; Type locality: Chilaria, Alto del Parrizal. Also cited from Colombia). DISTRIBUTION: Colombia. sparsepunctatus Cameron, 1923, see: Nacaeus. 1274 OSORIINAE: LISPINUS Bulletin AMNH specialis Cameron, 1947c: 114 (Lispinus; Type locality: Mauritius: Mt. Cocotte). DISTRIBUTION: Mascarene Islands. spectabilis Bernhauer, 1929i: 344 (Lispinus; Type locality: Luzon, Mount Polis). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Philippines. specularis Bernhauer, 1904, see: Nacaeus. spegazzinii Bernhauer, 1933e: 326 (Lispinus; Type locality: Argentinien: La Plata). DISTRIBUTION: Argentina. splendens Bernhauer, 1915d: 95 (Lispinus; Type locality: Deutsch-Ostafrika: West Usambara). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). DISTRIBUTION: Tanzania. splendidicollis Bernhauer, 1929i: 341 (Lispinus; Type locality: Luzon: Laguna, Los Baños). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Philippines. splendidipennis Bernhauer, 1929i: 341 (Lispinus; Type locality: Luzon: Balbalan). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Philippines. splendidus Bernhauer, 1932a: 74 (Lispinus; Type locality: Haut-Uele: Moto). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Congo. strictus Bernhauer, 1909c: 225 (Lispinus; Type locality: Brasilien: Rio Janeiro, Parana: Bella Vista). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). — Irmler, 1994: 62 (Lispinus; characters; Brazil; Argentina). DISTRIBUTION: Brazil, Argentina. strigiventris Kraatz, 1859: 186 (Lispinus; Type locality: India orientalis). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Cameron, 1930: 55 (Lispinus; characters; East Indies). — Nakane, 1963a: 81 (Lispinus; characters; habitus photograph; Japan). DISTRIBUTION: “East Indies”, Japan. striola Erichson, 1840: 829 (Lispinus; Type locality: Columbia; Americae meridionalis insulae). — Guérin-Méneville, 1857: 46 (Lispinus; characters; Colombia; Cuba; île de l’Amérique méridionale). — Chevrolat and Fauvel, 1863: 442 (Lispinus; characters; Cuba, Brazil, Colombia, Mexico). — Fauvel, 1864b: 46 [= 1865: 50] (Lispinus; characters; Mexico; Cuba; Antilles; Colombia; Brazil). — Solsky, 1872: 313 (Lispinus; Cayenne). — Sharp, 1876: 412 (Lispinus; Brazil). — Sharp, 1887: 719 (Lispinus; Mexico; British Honduras; Guatemala; Nicaragua). — Fauvel, 1891: 89 (Lispinus; Venezuela). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Blackwelder, 1943: 135 (Lispinus; characters; Cuba; Trinidad). — Scheerpeltz, 1957c: 240 (Lispinus; subgenus Aulacolispinus). — Scheerpeltz, 1960c: 66 (Lispinus; subgenus Aulacolispinus; Bolivia; Brazil). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Irmler, 1994: 57 (Lispinus; characters; Colombia; Mexico; Puerto Rico; Guatemala; Costa Rica; Panama; Trinidad; Peru; Ecuador; Surinam; French Guiana; Brazil; Paraguay; Bolivia). DISTRIBUTION: Paraguay, Brazil, Bolivia, Peru, Ecuador, Colombia, Venezuela, Surinam, French Guinana, West Indies, Panama, Costa Rica, Nicaragua, Guatemala, Belize, Mexico. Bulletin AMNH OSORIINAE: LISPINUS 1275 subcoriaceus Cameron, 1928, see: cameroni Scheerpeltz, 1933. subcoriaceus Cameron, 1928d: 97 (Lispinus; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Indonesia. sublaevicollis Bernhauer, 1929, see: Nacaeus. sublucens Fauvel, 1895: 184 (Lispinus; Type locality: Tenasserim, Kawkareet; Ile Bengkalis; Java, Tcibodas; Pengalengan, 1300 m). — Fauvel, 1904a: 44 (Lispinus; Nilghiris; N. Kanara; Bírmanie; Tenasserim; Cambodia; île Bengkalis; Java). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Cameron, 1930: 60 (Lispinus; characters; Burma). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). — Coiffait, 1981f: 331 (Lispinus; Andaman Islands). DISTRIBUTION: Indonesia, Cambodia, Myanmar, Andaman Islands, India. subnitens Bernhauer, 1929i: 352 (Lispinus; Type locality: Luzon, Butac, Limay, Laguna, Manila, Los Baños). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Philippines. subnitidus Bernhauer, 1929i: 352 (Lispinus; Type locality: Sumatra: Si Rambe). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Indonesia. subopacus Kraatz, 1859, see: Nacaeus. subpunctipennis Cameron, 1936c: 182 (Lispinus; Type locality: Java: Toeloengagoeng). DISTRIBUTION: Indonesia. subtilipennis Bernhauer, 1929, see: Nacaeus. sulcatulus Cameron, 1925b: 236, 237 (Lispinus; Type locality: New Guinea, Humboldt River). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). — Cameron, 1943a: 337 (Lispinus; characters in key). — Scheerpeltz, 1957c: 240 (Lispinus; subgenus Aulacolispinus). DISTRIBUTION: New Guinea. sulcatus Fauvel, 1904b: 79 (Lispinus; Type locality: Nouvelle-Guinee: Baie Huon; Sattelberg; Friedrich-Wilhelmhafen; Ile Deslacs). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Scheerpeltz, 1957c: 240 (Lispinus; subgenus Aulacolispinus). DISTRIBUTION: New Guinea. sulcicollis Coiffait, 1981g: 308 (Pseudolispinodes; subgenus Pseudolispinodes; Type locality: Ile Galapagos: Ile Santa Cruz, bois au-dessus de Santa Rosa). DISTRIBUTION: Galapagos Islands. sulcipennis Blackburn, 1902: 27 (Lispinus; Type locality: N. Queensland). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Bernhauer, 1920: 3 (Lispinus; Australia). — Cameron, 1927c: 260 (Lispinus; Fiji). — Scheerpeltz, 1957c: 240 (Lispinus; subgenus Aulacolispinus). — Newton, 1990: 92 (Lispinus). DISTRIBUTION: Australia. sutteri Scheerpeltz, 1957c: 223, 239 (Lispinus; subgenus Aulacolispinus; Type locality: W. Sumba: Waimangura). DISTRIBUTION: Indonesia. tardinus Irmler, 1994: 67 (Lispinus; Type locality: Costa Rica: Guapiles). DISTRIBUTION: Costa Rica. 1276 OSORIINAE: LISPINUS Bulletin AMNH tardus Sharp, 1887: 719 (Lispinus; Type locality: Panama, Volcan de Chiriqui 2500 to 4000 feet). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Irmler, 1994: 60 (Lispinus; characters; Mexico; West Indies). DISTRIBUTION: Mexico, Panama, West Indies. tenellus Erichson, 1840, see: Nacaeus. tenuicornis Kraatz, 1859: 187 (Lispinus; Type locality: Ceylan). — Bernhauer and Schubert, 1910: 24 (Lispinus; catalog). — Cameron, 1921a: 363, 399 (Lispinus; characters; Singapore). — Cameron, 1930: 57 (Lispinus; characters; Ceylon). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Sri Lanka, Singapore. tenuipunctus Bernhauer, 1929, see: nitidipennis Bernhauer, 1914. tenuis LeConte, 1863, see: Nacaeus. terminalis Sharp, 1876, see: quadripunctulus Fauvel, 1864. testaceus Kraatz, 1859: 188 (Lispinus; Type locality: Ceylan). — Bernhauer and Schubert, 1910: 24 (Lispinus; catalog). — Cameron, 1930: 64 (Lispinus; characters; Ceylon). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Sri Lanka. teutoniae Irmler, 1994: 71 (Lispinus; Type locality: Nova Teutonia). DISTRIBUTION: Brazil. tikoensis Bernhauer, 1939i: 252 (Lispinus; Type locality: Kamerun: Tiko). DISTRIBUTION: Cameroon. tonkinensis Cameron, 1939a: 22 (Lispinus; Type locality: Tonkin: Hoa Binh). DISTRIBUTION: Vietnam. trichidus Irmler, 1994, see: Neolosus. tripunctatus Bernhauer, 1914c: 82 (Lispinus; Type locality: Sumatra: Si-Rambe). — Cameron, 1928d: 96 (Lispinus; Sumatra). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Indonesia. tristis Bernhauer, 1929i: 356 (Lispinus; replacement name for curticollis Bernhauer). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Vietnam. curticollis Bernhauer, 1904: 12 (Lispinus; [preoccupied]; Type locality: Tonkin, Montes Mauson, 2000–3000 ). — Scheerpeltz, 1933: 1018 (Lispinus; synonym of tristis). unipunctatus Cameron, 1930b: 160 (Lispinus; Type locality: Sarawak; mt. Matang. Also cited from North Borneo). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Borneo. unistriatus Fauvel, 1878f: 202 (Lispinus; Type locality: Nouvelle-Guinée, Ramoi, Hatam). — Fauvel, 1879: 76 (Lispinus; Amberbaki; Mysol). — Bernhauer and Schubert, 1910: 24 (Lispinus; catalog). — Bernhauer, 1926f: 312 (Lispinus; Buru). — Cameron, 1931: 356 (Lispinus; New Guinea). — Cameron, 1937a: 87 (Lispinus; New Guinea). — Cameron, 1943a: 337 (Lispinus; characters in key). — Scheerpeltz, 1957c: 223 (Lispinus; subgenus Aulacolispinus; Flores). DISTRIBUTION: New Guinea, Indonesia. Bulletin AMNH OSORIINAE: LISPINUS 1277 usambarae Fauvel, 1904d: 293 (Lispinus; Type locality: Usambara: Derema, 850 m). — Fauvel, 1907: 11 (Lispinus; Kilimandjaro; Usambara). — Bernhauer and Schubert, 1910: 24 (Lispinus; catalog). — Eichelbaum, 1913: 116 (Lispinus; Tanzania). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). — Scheerpeltz, 1974c: 5 (Lispinus; Sudan). DISTRIBUTION: Tanzania, Sudan. usambaricus Bernhauer, 1915d: 96 (Lispinus; Type locality: Deutsch-Ostafrika: West Usambara). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Tanzania. validicornis Fauvel, 1904b: 79 (Lispinus; Type locality: Queensland: Cooktown). — Bernhauer and Schubert, 1910: 24 (Lispinus; catalog). — Scheerpeltz, 1957c: 240 (Lispinus; subgenus Aulacolispinus). — Newton, 1990: 92 (Lispinus). DISTRIBUTION: Australia. varians Fauvel, 1904e: 296 (Lispinus; Type locality: Tananarive; forêt de Moramanga; Diego Suarez). — Bernhauer and Schubert, 1910: 24 (Lispinus; catalog). DISTRIBUTION: Madagascar. veitchi Bernhauer, 1926c: 159 (Lispinus; Type locality: Fiji: Lantoca). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Fiji. villiersi Cameron, 1948b: 231 (Holotrochus; Type locality: Haute-Guinée: Mont Nimba N.E., alt. 500–700 m). — Fagel, 1959: 32 (Lispinus). DISTRIBUTION: French Guinea. vinsoni Cameron, 1936d: 201 (Lispinus; Type locality: Mauritius: Forest Side). DISTRIBUTION: Mascarene Islands. vulneratus Bernhauer, 1929i: 351 (Lispinus; Type locality: Luzon, Balbalan). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Philippines. vulneratus Bernhauer, 1932, see: burunganus Scheerpeltz, 1933. wagneri Cameron, 1931: 356 (Lispinus; Type locality: Finsch Haven, New Guinea). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: New Guinea. zambalesanus Bernhauer, 1929i: 347 (Lispinus; Type locality: Luzon, Zambales). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). DISTRIBUTION: Philippines.
Nacaeus [41 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions] Nacaeus Blackwelder, 1942: 82 (subgenus of Pseudolispinodes; species included: planellus, nitidissimus, exiguus). TYPE SPECIES: Lispinus planellus Sharp, fixed by original designation. — Blackwelder, 1942: 79, 89 (characters; misidentified as Pseudolispinodes according to Black- welder, 1952: 256, 373; [Note: All of species in the list (p. 89) are now in Nacaeus]). — Blackwelder, 1944: 107 (misidentified as Pseudolispinodes; checklist of species from Mexico, Central and South America, and the West Indies; [Note: All of species in the list are now in Nacaeus]). — Blackwelder, 1952: 256 (type species: planellus). 1278 OSORIINAE: NACAEUS Bulletin AMNH
— Moore and Legner, 1974b: 81 (characters; notes). — Moore and Legner, 1975: 169 (catalog; North America north of Mexico). — Moore and Legner, 1979: 147 (characters; notes). — Frank, 1986a: 365 (checklist of species of Florida). — Newton, 1990: 92 (characters in key; six described Australian species). — Naomi, 1997: 128 (characters; revision of and key to species of Japan). — Downie and Arnett, 1996: 427 (characters). — Lundgren, 1998: 46 (list of species from Florida; records for other States listed for each species). — Drugmand, 1998a: 216 (list of types in l’Institut royal des Sciences naturelles de Belgique). — Newton, Thayer, Ashe, and Chandler, 2000: 377 (2 Nearctic species; notes; characters in key). Liberiella Blackwelder, 1942: 81 (subgenus of Pseudolispinodes; species included: aethiops, cooki). TYPE SPECIES: Pseudolispinodes (Liberiella) cooki Blackwelder, fixed by original designation. — Blackwelder, 1952: 220 (subgenus of Nacaeus; type species). Liberiana Blackwelder, 1942: 82 (subgenus of Pseudolispinodes; species included: femoralis). TYPE SPECIES: Pseudolispinodes (Liberiana) femoralis Blackwelder, fixed by original designation and monotypy. — Blackwelder, 1952: 220 (subgenus of Nacaeus; type species). Rumeba Blackwelder, 1942: 82 (subgenus of Pseudolispinodes; species included: lispinoides). TYPE SPECIES: Pseudolispinodes (Rumeba) lispinoides Blackwelder, fixed by original designation and monotypy. — Blackwelder, 1952: 342 (subgenus of Nacaeus; type species). Tannea Blackwelder, 1952: 373 (subgenus of Nacaeus; proposed for misidentification of Pseudolispinodes by Blackwelder, 1942; species included: tenellus, no other species were cited by name). TYPE SPECIES: Lispinus tenellus Erichson, fixed by original designation and monotypy. aethiops Eppelsheim, 1895a: 138 (Lispinus; Type locality: Gabon). — Fauvel, 1903e: 236 (Lispinus; Cameroon; Togo; Gabon; San Thome; Ivory Coast; Sierra Leone; Ethiopia). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Paulian, 1941: 140 (Lispinus; larval characters). — Bernhauer, 1942c: 347 (Lispinus; Cameroon). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Liberiella; checklist). — Cameron, 1952e: 827 (Lispinus; Togo). — Scheerpeltz, 1956a: 482 (Lispinus; Liberia; Nigeria; Cameroon; French Congo; Belgian Congo; Uganda; Kenya; Tanganyika). — Fagel, 1963a: 338 (Lispinus; Guinea; Ivory Coast). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Ethiopia, Uganda, Kenya, Tanzania, Congo, “Frence Congo”, Gabon, São Tomé and Príncipe, Cameroon, Nigeria, Togo, Liberia, Sierra Leone. africanus Bernhauer, 1929, see: impressicollis Motschulsky, 1857. beesoni Cameron, 1924b: 161 (Lispinus; Type locality: Dehra Dun; Siwaliks). — Cameron, 1930: 61 (Lispinus; characters; India). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Coiffait, 1978b: 112 (Lispinus; Bhutan). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: India, Bhutan. binaluanensis Bernhauer, 1929, see: impressicollis Motschulsky, 1857. birmanus Fauvel, 1895: 184 (Lispinus; Type locality: Bírmanie, Carin Cheba, 1200–1300 m; Carin Asciuii Ghécu, 1400–1500 m; Tenasserim, mont Mooleyit, 1800–1900 m). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). Bulletin AMNH OSORIINAE: NACAEUS 1279
— Cameron, 1930: 58 (Lispinus; characters; Assam, Burma). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Myanmar, India. cameroniellus Naomi, 1997: 141 (Nacaeus; replacement name for selangorensis Cameron, 1929). DISTRIBUTION: Malaysia. selangorensis Cameron, 1929b: 438 (Pseudolispinodes; [preoccupied]; Type locality: Selangor: Bukit Kutu). — Scheerpeltz, 1933: 1010 (Pseudolispinodes; catalog). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Naomi, 1997: 141 (Nacaeus; synonym of cameroniellus). castaneus Fauvel, 1878f: 206 (Lispinus; Type locality: Nouvelle-Guinée, Ansus; Salvatti; Iles Arou, Wokan; Iles Key). — Fauvel, 1879: 76 (Lispinus; Batcian; Sula). — Fauvel, 1898a: 114 (Lispinus; Seychelles). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — H. Kolbe, 1910: 21 (Lispinus; Seychelles). — Bernhauer, 1926c: 160 (Lispinus; characters; Engano). — Cameron, 1927a: 22 (Lispinus; Samoa). — Cameron, 1927c: 260 (Lispinus; Fiji). — Cameron, 1931: 356 (Lispinus; New Guinea). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). — Cameron, 1934a: 20 (Lispinus; New Hebrides). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Frank, 1982: 7 (Nacaeus; list of parasites). DISTRIBUTION: Samoa, Fiji, Vanuatu, New Guinea, Indonesia, Seychelles. claviger Cameron, 1913, see: impar Camerom, 1913. coarcticollis Kraatz, 1859: 186 (Lispinus; Type locality: India orientalis). — Bernhauer and Schubert, 1910: 21 (Lispinus; catalog). — Cameron, 1921a: 363, 399 (Lispinus; characters; Singapore). — Cameron, 1930: 58 (Lispinus; characters; East Indies; Singapore; Malay Peninsula; Borneo). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: “East Indies”, Malaysia, Singapore, Borneo. cooki Blackwelder, 1942: 86 (Pseudolispinodes; subgenus Liberiella; Type locality: Liberia: Mount Coffee). — Blackwelder, 1952: 220 (Nacaeus; subgenus Liberiella). DISTRIBUTION: Liberia. curtipennis Bernhauer, 1929i: 356 (Lispinus; Type locality: Borneo: Sandakan). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Borneo. danforthi Blackwelder, 1943: 125 (Pseudolispinodes; Type locality: Puerto Rico, College Farm 1 mile east of Mayaqüez. Also cited from Guadeloupe). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: West Indies. enganoensis Bernhauer, 1929, see: impressicollis Motschulsky, 1857. fauveli Sharp, 1887, see: tenellus Erichson, 1840. 1280 OSORIINAE: NACAEUS Bulletin AMNH femoralis Blackwelder, 1942: 86 (Pseudolispinodes; subgenus Liberiana; Type locality: Liberia: Mount Coffee). — Blackwelder, 1952: 220 (Nacaeus; subgenus Liberiana). DISTRIBUTION: Liberia. filiformis Walker, 1858, see: impressicollis Motschulsky, 1857. flavipennis Fauvel, 1864, see: tenellus Erichson, 1840. foveatus Fauvel, 1878f: 204 (Lispinus; Type locality: Nouvelle-Guinée, Ansus, Ramoi; Iles Arou, Wokan; Iles Key). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Bernhauer, 1920: 3 (Lispinus; Australia). — Bernhauer, 1926f: 312 (Lispinus; Buru). — Cameron, 1931: 356 (Lispinus; New Guinea). — Cameron, 1937a: 87 (Lispinus; New Guinea). — Scheerpeltz, 1957c: 223 (Lispinus; subgenus Lispinus; Sumba). — Newton, 1990: 92 (Nacaeus). DISTRIBUTION: Indonesia, New Guinea, Australia. foveolus Blackwelder, 1943: 127 (Pseudolispinodes; Type locality: Montserrat). — Legner and Moore, 1977: 174 (Lispinus; found under bark). DISTRIBUTION: West Indies. fulvus Motschulsky, 1857a: 495 (Lispinus; Type locality: Calcutta). — Kraatz, 1859: 187 (Lispinus; India orienali). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Bernhauer, 1929i: 354 (Lispinus; characters in key; Philippines). — Cameron, 1930: 63 (Lispinus; characters; East Indies; Singapore; Philippines). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Philippines, Singapore, India. globiceps Cameron, 1928, see: nitidissimus Bernhauer, 1905. guadeloupae Blackwelder, 1943: 129 (Pseudolispinodes; Type locality: Guadeloupe: Gourbeyre). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: West Indies. guamensis Bernhauer, 1942, see: impressicollis Motschulsky, 1857. impar Cameron, 1913: 322 (Lispinus; Type locality: St. Vincent). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Blackwelder, 1943: 122 (Pseudolispinodes; characters; Cuba; Jamaica; Haiti; Dominican Republic; Montserrat; St. Lucia; St. Vincent; Trinidad). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: West Indies. claviger Cameron, 1913: 321 (Lispinus; Type locality: St. Vincent). — Scheerpeltz, 1933: 1013 (Lispinus; catalog). — Blackwelder, 1943: 122 (Pseudolispinodes; synonym of impar). impressicollis Motschulsky, 1857a: 495 (Lispinus; Type locality: Indes orientales). — Kraatz, 1859: 186 (Lispinus; characters; Ceylon; India orientalis). — Fauvel, 1897d: 240 (Lispinus; Madeira). — Fauvel, 1902b: 47 (Lispinus; Madeira). — Fauvel, 1903: 150 (Lispinus; Ceylon; Japon; Hawaii). — Fauvel, 1904a: 44 (Lispinus; Anamalai). — Bernhauer, 1907: 372 (Lispinus; characters; Japan). — Sharp, 1908: 539 (Lispinus; Hawaii). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). Bulletin AMNH OSORIINAE: NACAEUS 1281
— Bernhauer, 1915g: 180 (Lispinus; Japan, Madagascar, Comoros, Seychelles, German East Africa, Madeira, Hawaii, Indomalayan region). — Cameron, 1921a: 363, 399 (Lispinus; characters; Singapore). — Bernhauer, 1922a: 149 (Lispinus; Indochina). — Bernhauer, 1922b: 166 (Lispinus; Seychelles). — Bernhauer, 1922c: 222 (Lispinus; Formosa). — Cameron, 1927a: 22 (Lispinus; Samoa). — Cameron, 1927c: 260 (Lispinus; Fiji). — Cameron, 1928d: 96 (Lispinus; Sumatra). — Cameron, 1928f: 400 (Lispinus; Borneo). — Bernhauer, 1928: 5 (Lispinus; Sumatra). — Bernhauer, 1929i: 352 (Lispinus; characters in key; Philippines). — Cameron, 1930: 60 (Lispinus; characters; Ceylon; India; Japan; Southwest Africa; Madeira). — Scheerpeltz, 1933: 1014 (Lispinus; catalog). — Scheerpeltz, 1935a: 635 (Lispinus; Australia). — Cameron, 1936a: 29 (Lispinus; Java). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Méquignon, 1942a: 16 (Lispinus; Azores). — Jarrige, 1957: 112 (Lispinus; Réunion). — Scheerpeltz, 1957c: 223 (Lispinus; subgenus Lispinus; Sumba). — Nakane and Sawada, 1960: A122 (Lispinus; Yakushima Island). — Watanabe, 1961: 348 (Lispinus; Izu Islands). — Nakane, 1963a: 82 (Lispinus; characters; habitus photograph Japan). — Smetana, 1963b: 30 (Lispinus; Madeira). — Smetana, 1966: 323 (Lispinus; notes; Czechoslovakia). — Israelson, 1969: 149 (Lispinus; notes; Canary Islands). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Frank, 1982: 7 (Nacaeus; list of parasites). — Yuh, Paik, Kwon, and Lee, 1985: 253 (Lispinus; Korea). — Borges, 1990: Table II (Lispinus; Azores). — Newton, 1990: 92 (Nacaeus). — Nishida, 1994: 72 (Nacaeus; Hawaii). — Hernández, Outerelo, and Gamarra, 1994: 187 (Lispinus; checklist of species; Canary Islands). — Naomi, 1997: 132 (Nacaeus; characters). — Nishida, 1997: 62 (Nacaeus; Hawaii). — Rogé and Dauphin, 1999: 155 (Lispinus; notes; France). DISTRIBUTION: Czechoslovakia, France, Azores, Madeira, Canary Islands, Namibia, Tanzania, Madagascar, Mascarene Islands, Comoros, Seychelles, India, Sri Lanka, Singapore, Indonesia, Borneo, Philippines, Australia, Fiji, Samoa, Mariana Islands, Taiwan, Japan, Korea, Hawaii. filiformis Walker, 1858: 205 (Omalium; Type locality: Ceylon). — Fauvel, 1897d: 240 (Lispinus; synonym of impressicollis). — Fauvel, 1902b: 47 (Lispinus; synonym of impressicollis). — Fauvel, 1903: 150 (Lispinus; synonym of impressicollis). — Fauvel, 1904a: 44 (Lispinus; synonym of impressicollis). — Bernhauer and Schubert, 1910: 22 (Lispinus; synonym of impressicollis). — Cameron, 1930: 60 (Lispinus; synonym of impressicollis). — Scheerpeltz, 1957c: 223 (Lispinus; synonym of impressicollis). africanus Bernhauer, 1929i: 353 (Lispinus; subspecies of impressicollis; Type locality: Deutsch Ostafrika, Usaromo). — Scheerpeltz, 1933: 1014 (Lispinus; variety of impressicollis). binaluanensis Bernhauer, 1929i: 353 (Lispinus; subspecies of impressicollis; Type locality: Palawan: Binaluan). — Bernhauer, 1928: 5 (Lispinus; [Note: Unavailable name here, listed without description]; Sumatra; Philippinus). — Scheerpeltz, 1933: 1014 (Lispinus; variety of impressicollis). 1282 OSORIINAE: NACAEUS Bulletin AMNH
enganoensis Bernhauer, 1929i: 353 (Lispinus; subspecies of impressicollis; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1015 (Lispinus; variety of impressicollis). sublaevicollis Bernhauer, 1929i: 353 (Lispinus; subspecies of impressicollis; Type locality: Mindanao, Mumungan). — Scheerpeltz, 1933: 1015 (Lispinus; variety of impressicollis). subtilipennis Bernhauer, 1929i: 353 (Lispinus; subspecies of impressicollis; Type locality: Luzon, Los Baños, Mount Maquiling. Leyte, Burauen. Basilan, Dinagat. Mindanao, Kolambugan. Siargao, Dapa. Comoren. Madagaskar). — Scheerpeltz, 1933: 1015 (Lispinus; cited as tilipennis; variety of impressicollis). guamensis Bernhauer, 1942a: 41 (Lispinus; subspecies of impressicollis; Type locality: Guam: Agat; Yigo). iridescens Bernhauer, 1942a: 41 (Lispinus; subspecies of impressicollis; Type locality: Guam: Yigo). iridescens Bernhauer, 1942, see: impressicollis Motschulsky, 1857. iriomotensis Naomi, 1997: 137 (Nacaeus; Type locality: Japan: Okinawa Pref.: Kampira fall, Iriomote Is.). DISTRIBUTION: Japan. irregularis Blackwelder, 1943: 124 (Pseudolispinodes; Type locality: Jamaica: vicinity of Trinityville, parish of St. Thomas. Also cited from Cuba and Haiti). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Assing, 1998e: 140 (Nacaeus; Madeira). DISTRIBUTION: West Indies, Madeira. japonicus Cameron, 1930c: 181 (Lispinus; Type locality: Settsu: Katsnuogi). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). — Naomi, 1997: 132 (Nacaeus; characters; Japan). DISTRIBUTION: Japan. jyeri Bernhauer, 1914c: 82 (Lispinus; Type locality: Himalaja: Dehra Dun). — Cameron, 1930: 63 (Lispinus; cited as iyeri; characters; India). — Scheerpeltz, 1933: 1015 (Lispinus; cited as iyeri; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Abdullah and Qadri, 1970: 115 (Lispinus; cited as iyeri; characters; West Pakistan). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: India, Pakistan. leai Cameron, 1943a: 337 (Lispinus; Type locality: Australia: Cairns district). — Newton, 1990: 92 (Nacaeus). DISTRIBUTION: Australia. lispinoides Blackwelder, 1942: 87 (Pseudolispinodes; subgenus Rumeba; Type locality: Liberia: Mount Coffee). — Blackwelder, 1952: 343 (Nacaeus; subgenus Rumeba). DISTRIBUTION: Liberia. longulus Sharp, 1874: 101 (Lispinus; Type locality: Japan; [Note: See lectotype designation by Naomi, 1997]). — Fauvel, 1902b: 47 (Lispinus; synonym of impressicollis). — Bernhauer, 1907: 372 (Lispinus; synonym of impressicollis). — Bernhauer and Schubert, 1910: 22 (Lispinus; synonym of impressicollis). — Bernhauer, 1929i: 353 (Lispinus; subspecies of impressicollis; characters in key; Japan; Formosa). — Cameron, 1930: 60 (Lispinus; synonym of impressicollis). — Scheerpeltz, 1933: 1014 (Lispinus; variety of impressicollis). — Cameron, 1949b: 458 (Lispinus; cited as valid species; Japan). — Watanabe, 1961: 348 (Lispinus; synonym of impressicollis). Bulletin AMNH OSORIINAE: NACAEUS 1283
— Watanabe and Shibata, 1972: 59 (Lispinus; subspecies of impressicollis; Yaku-shima Island). — Naomi, 1997: 129 (Nacaeus; lectotype designation; valid species; notes; characters; Japan). — Watanabe, 1998: 313 (Nacaeus; Japan). — Cho and Ahn, 1999: 178 (Nacaeus; characters; Korea). DISTRIBUTION: Korea, Japan, Taiwan. luzonicus Bernhauer, 1929i: 348 (Lispinus; Type locality: Luzon, Mount Banahao, Los Baños). — Scheerpeltz, 1933: 1015 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Philippines. morugae Blackwelder, 1943: 121 (Pseudolispinodes; Type locality: Trinidad, Herrara Estate near Basse Terre on road from Princes Town to Moruga, Victoria County, ward of Moruga). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: West Indies. newtoniellus Naomi, 1997: 135 (Nacaeus; Type locality: Japan: Okinawa Pref.: Mt. Nagodake, Okinawa Is.). DISTRIBUTION: Japan. nigrifrons Fauvel, 1863: 443 (Lispinus; Type locality: Cuba). — Fauvel, 1864b: 55 [= 1865: 59] (Lispinus; characters; Cuba). — Solsky, 1872: 314 (Lispinus; Cayenne). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Blackwelder, 1943: 128 (Pseudolispinodes; characters; Cuba; Haiti; Montserrat; Guadeloupe; St. Lucia). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: West Indies, French Guiana. sparsepunctatus Cameron, 1923: 391 (Lispinus; Type locality: Haiti). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). — Blackwelder, 1943: 128 (Pseudolispinodes; synonym of nigrifrons). nitidissimus Bernhauer, 1905: 10 (Ancaeus; Type locality: Engano). — Bernhauer and Schubert, 1910: 19 (Ancaeus; catalog). — Cameron, 1928d: 96 (Paralispinus; Sumatra). — Cameron, 1930b: 160 (Paralispinus; British North Borneo). — Scheerpeltz, 1933: 1009 (Paralispinus; catalog). — Blackwelder, 1942: 90 (Pseudolispinodes; subgenus Nacaeus; checklist). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Borneo, Singapore, Indonesia. globiceps Cameron, 1928a: 35 (Lispinus; Type locality: Sumatra; Singapore; Borneo). — Cameron, 1928d: 96 (Paralispinus; synonym of nitidissimus). — Cameron, 1930b: 160 (Paralispinus; synonym of nitidissimus). — Scheerpeltz, 1933: 1009 (Paralispinus; synonym of nitidissimus). — Scheerpeltz, 1933: 1014 (Lispinus; [Note: Scheerpeltz cited globiceps Cameron twice, once (p. 1009) as a synonym of Paralispinus nitidissimus and once (p. 1014) as a valid species in Lispinus; I followed Cameron’s synonymy with nitidissimus]; catalog). planellus Sharp, 1887: 722 (Lispinus; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Bierig, 1931: 427 (Lispinus; characters; Panama). — Scheerpeltz, 1933: 1009 (Paralispinus; catalog). 1284 OSORIINAE: NACAEUS Bulletin AMNH
— Blackwelder, 1942: 90 (Pseudolispinodes; subgenus Nacaeus; checklist; type species of Nacaeus). — Blackwelder, 1952: 256 (Nacaeus; subgenus Nacaeus). DISTRIBUTION: Panama. punctatus Blackwelder, 1943: 161 (Paralispinus; Type locality: Puerto Rico, El Yunque, elevation about 3000 feet). — Irmler, 1991: 85 (Tannea). DISTRIBUTION: West Indies. reversus Blackwelder, 1943: 122 (Pseudolispinodes; Type locality: Hispaniola, Constanza, Dominican Republic, elevation 3000–4000 feet). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: West Indies. rivularis Lea, 1906: 214 (Lispinus; Type locality: Tweed and Richmond Rivers, N.S.W.). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Newton, 1990: 92 (Nacaeus). DISTRIBUTION: Australia. robusticollis Bernhauer, 1929i: 353 (Lispinus; subspecies of impressicollis; Type locality: Luzon, Los Baños, Mount Banahao, Mount Maquiling, Lamao, Mount Isarog, Zambales, Limay. Masbate, Aroroy. Palawan, Binaluan. Mindanao, Mumungan. Borneo. Seychellen). — Scheerpeltz, 1933: 1015 (Lispinus; variety of impressicollis). — Cameron, 1949b: 458 (Lispinus; cited as valid species; Formosa). DISTRIBUTION: Seychelles, Borneo, Philippines, Taiwan. rubidus Cameron, 1925b: 236, 238 (Lispinus; Type locality: Selangor, Federated Malay States; The Gap). — Bernhauer, 1929i: 354 (Lispinus; characters in key; Philippines). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Malaysia, Philippines. selangorensis Cameron, 1925b: 237, 243 (Lispinus; Type locality: Selangor, The Gap). — Naomi, 1997: 140 (Nacaeus). DISTRIBUTION: Malaysia. selangorensis Cameron, 1929, see: cameroniellus Naomi, 1997. sericeiventris Bernhauer, 1914c: 81 (Lispinus; Type locality: Sumatra: Si-Rambe). — Cameron, 1928d: 96 (Lispinus; Sumatra). — Bernhauer, 1928: 5 (Lispinus; Sumatra). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Indonesia. sericeus Coiffait, 1981g: 309 (Pseudolispinodes; subgenus Nacaeus; Type locality: Ile Galapagos: Ile Isabela, environs de Santa Tomas). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Galapagos Islands. sharpi Cameron, 1920, see: specularis Bernhauer, 1904. sidneensis Fauvel, 1878g: 481 (Lispinus; Type locality: Nouvelle-Galles du Sud, Sydney). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Lea, 1910a: 139 (Lispinus; with Termes lactis; Australia). — Bernhauer, 1920: 3 (Lispinus; Australia). — Newton, 1990: 92 (Nacaeus). DISTRIBUTION: Australia. sparsepunctatus Cameron, 1923, see: nigrifrons Fauvel, 1863. Bulletin AMNH OSORIINAE: NACAEUS 1285 specularis Bernhauer, 1904: 11 (Lispinus; Type locality: Ins. Nias). — Fauvel, 1905e: 78 (Lispinus; Java; Sumatra; Nias Island; Singapore). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Bernhauer, 1922b: 165 (Lispinus; Seychelles). — Cameron, 1927c: 260 (Lispinus; Fiji). — Cameron, 1928d: 96 (Lispinus; Sumatra). — Scheerpeltz, 1933: 1017 (Lispinus; catalog). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Scheerpeltz, 1978: 186 (Lispinus; Timor). — Herman, 2001: 13 (Nacaeus). DISTRIBUTION: Seychelles, Singapore, Indonesia, Philippines, Fiji. sharpi Cameron, 1920b: 278 (Lispinus; Type locality: Mandai). — Cameron, 1921a: 363, 399 (Lispinus; characters; Singapore). — Cameron, 1927c: 260 (Lispinus; synonym of specularis). — Cameron, 1928d: 96 (Lispinus; synonym of specularis). — Scheerpeltz, 1933: 1017 (Lispinus; synonym of specularis). — Scheerpeltz, 1978: 186 (Lispinus; synonym of specularis). sublaevicollis Bernhauer, 1929, see: impressicollis Motschulsky, 1857. subopacus Kraatz, 1859: 187 (Lispinus; Type locality: Ceylan). — Fauvel, 1878f: 206 (Lispinus; characters; New Guinea; Aru Islands; Sri Lanka; Salvatti). — Fauvel, 1879: 76 (Lispinus; Moluccas; Celebes). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Bernhauer, 1920: 3 (Lispinus; Australia). — Cameron, 1927c: 260 (Lispinus; Fiji). — Cameron, 1928d: 96 (Lispinus; Sumatra). — Bernhauer, 1929i: 356 (Lispinus; characters in key; Philippines). — Cameron, 1930: 62 (Lispinus; characters; Ceylon; New Guinea; Sumatra). — Scheerpeltz, 1933: 1018 (Lispinus; catalog). — Cameron, 1933f: 74 (Lispinus; Marquesas Islands). — Cameron, 1934a: 20 (Lispinus; New Hebrides). — Cameron, 1936: 4 (Lispinus; Society Islands). — Cameron, 1937a: 87 (Lispinus; New Guinea). — Scheerpeltz, 1954: 110 (Lispinus; Ceylon). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Coiffait, 1976: 220 (Lispinus; Tahiti; Marquesas). — Newton, 1990: 92 (Nacaeus). DISTRIBUTION: Sri Lanka, Indonesia, Philippines, New Guinea, Australia, Vanuatu, Society Islands, Marquesas Islands. subtilipennis Bernhauer, 1929, see: impressicollis Motschulsky, 1857. tenellus Erichson, 1840: 830 (Lispinus; Type locality: Columbia; Americae meridionalis ins. St. Johannis and Puerto Rico). — Fauvel, 1864b: 53 [= 1865: 57] (Lispinus; characters; Mexico; Colombia; Cuba). — Fauvel, 1878d: 179 [= 1878e: 15] (Lispinus; characters; South Carolina; Georgia; Florida; Texas; Mexico; Cuba; Venezuela; Colombia). — Fleutiaux and Sallé, 1889: 382 (Lispinus; Guadeloupe Island). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Scheerpeltz, 1933: 1018 (Lispinus; catalog; Argentina). — Blackwelder, 1942: 89 (Pseudolispinodes; subgenus Pseudolispinodes; checklist). — Blackwelder, 1943: 126 (Pseudolispinodes; characters; Jamaica; Cuba; Haiti; Puerto Rico; St. John; Guadeloupe; St. Vincent; Grenada). — Blackwelder, 1952: 373 (Nacaeus; subgenus Tannea). — Legner and Moore, 1977: 174 (Lispinus; found under bark). — Frank, 1982: 8 (Nacaeus; list of parasites). — Downie and Arnett, 1996: 427 (Nacaeus; cited as tennellus; characters; USA). DISTRIBUTION: Argentina, Colombia, Venezuela, West Indies, Mexico, USA. 1286 OSORIINAE: NACAEUS Bulletin AMNH
tenuis LeConte, 1863a: 60 (Lispinus; Type locality: Southern States). — LeConte, 1877: 250 (Lispinus; characters). — Fauvel, 1878d: 179 [= 1878e: 15] (Lispinus; synonym of tenellus). — Bernhauer and Schubert, 1910: 24 (Lispinus; synonym of tenellus). — Notman, 1920b: 693 (Lispinus; probably valid species; Florida). — Scheerpeltz, 1933: 1018 (Lispinus; synonym of tenellus). — Blackwelder, 1943: 126 (Pseudolispinodes; synonym of tenellus). flavipennis Fauvel, 1864b: 54 [= 1865: 58] (Lispinus; Type locality: Caracas; Teapa, Mexico). — Fauvel, 1878d: 179 [= 1878e: 15] (Lispinus; synonym of tenellus). — Sharp, 1887: 721 (Lispinus; notes; Mexico). — Bernhauer and Schubert, 1910: 24 (Lispinus; synonym of tenellus). fauveli Sharp, 1887: 720 (Lispinus; [preoccupied]; proposed for misidentification of tenel- lus by Chevrolat and Fauvel; Type locality: Mexico, Teapa, Cordova; British Hon- duras, R. Hondo; Guatemala, near the city, Zapote, El Tumbador; Colombia; Cuba). — Chevrolat and Fauvel, 1863: 443 (Lispinus; [Note: Misidentification: Bernhauer and Schubert, 1910: 22 cited tenellus Chevrolat and Fauvel as a synonym of fauveli; Chevrolat and Fauvel did not describe the species as new, they attributed it to Erichson]; characters; Cuba, Mexico, Colombia). — Bernhauer and Schubert, 1910: 22 (Lispinus; catalog). — Blackwelder, 1943: 126 (Pseudolispinodes; synonym of tenellus). tenuis LeConte, 1863, see: tenellus Erichson, 1840.
Neolosus [65 species; Neotropical, Palaearctic, Ethiopian, and Oriental Regions] Neolosus Blackwelder, 1942: 85 (subgenus of Holosus; species included: tachyporiformis, insularis). TYPE SPECIES: Holosus tachyporiformis Motschulsky, fixed by original designation. — Blackwelder, 1952: 260 (elevated to replace the preoccupied Holosus; type species: tachypori- formis). — Hammond, 1984: 197 (checklist of species of Borneo). — Naomi, 1996b: 766 (characters; species of Japan). — Irmler, 1999: 267 (revision of and key to species of South America; characters). Holosus Motschulsky, 1857a: 496 (species included: tachyporiformis, tachiniformis, conuri- formis, mycetoporiformis, olisthaeriformis; preoccupied by Steven, 1829). TYPE SPECIES: Holosus tachiniformis Motschulsky, fixed by subsequent designation by Blackwelder, 1942: 88. — Gemminger and Harold, 1868: 676 (catalog). — Fauvel, 1878f: 198 (characters). — Duvivier, 1883: 203 (catalog). — Bernhauer and Schubert, 1910: 20 (world catalog; 22 species). — Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog). — Bernhauer, 1926d: 256 (key to species of Philippines). — Cameron, 1930: 43 (characters; key to species of British India). — Scheerpeltz, 1933: 1010 (world catalog supplement). — Blackwelder, 1942: 84, 88 (characters; notes; type species). — Blackwelder, 1952: 187 (type species). — Fagel, 1960d: 263 (notes; key to African species). — Shibata, 1973c: 22 (checklist of species of Taiwan). — Drugmand, 1998a: 210 (list of types in l’Institut royal des Sciences naturelles de Belgique). Relinda Blackwelder, 1942: 85 (subgenus of Holosus; species included: mycetoporiformis, olisthaeriformis). TYPE SPECIES: Holosus mycetoporiformis Motschulsky, fixed by original designation. Bulletin AMNH OSORIINAE: NEOLOSUS 1287
Osholus Blackwelder, 1952: 278 (replacement name for the preoccupied Holosus sensu stricto; subgenus of Neolosus). TYPE SPECIES: Holosus tachiniformis Motschulsky, fixed by objective synonymy with Holosus. aberrans Fauvel, 1904b: 78 (Holosus; Type locality: Ceylan). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1930: 51 (Holosus; characters; Ceylon). — Scheerpeltz, 1933: 1010 (Holosus; catalog). DISTRIBUTION: Sri Lanka. ambigenus Fauvel, 1895: 183 (Holosus; Type locality: Bírmanie, Carin Asciuii Cheba, 1200–1300 m; Carin Cheba, 900–1100 m). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1930: 50 (Holosus; characters; Burma). — Scheerpeltz, 1933: 1010 (Holosus; catalog). DISTRIBUTION: Myanmar. aper Sharp, 1889: 466 (Lispinus; Type locality: Kiga.; [Note: See lectotype designation by Naomi, 1996b]. Originally cited from Miyanoshita, Nagasaki, Oyayama, Kashiwagi). — Bernhauer and Schubert, 1910: 21 (Lispinus; cited as asper; catalog). — Nakane, 1963a: 81 (Lispinus; characters; habitus photograph; Japan). — Yuh, Paik, Kwon, and Lee, 1985: 225 (Lispinus; Korea). — Frank, 1982: 7 (Lispinus; cited as asper; list of parasites). — Naomi, 1996b: 767 (Neolosus; lectotype designation: Kiga.; characters; Japan). — Cho and Ahn, 1999: 176, 178 (Neolosus; characters in key; Korea). DISTRIBUTION: Japan, Korea. basicollis Fauvel, 1903b: 166 (Holosus; Type locality: Bussira, Congo belge). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Fagel, 1960d: 272 (Holosus; characters; Cameroon; Gabon; Congo). DISTRIBUTION: Congo, Cameroon, Gabon. distincticollis Bernhauer, 1932a: 73 (Holosus; Type locality: Jaut-Uele: Moku Moto). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Fagel, 1960d: 272 (Holosus; synonym of basicollis). bifoveolatus Bernhauer, 1926d: 258 (Holosus; Type locality: Masbate, Aroroy. Siargao, Dapa. Leyte. Luzon, Provinz Laguna, Los Baños). — Scheerpeltz, 1933: 1010 (Holosus; catalog). DISTRIBUTION: Philippines. boettcheri Bernhauer, 1926d: 257 (Holosus; Type locality: Mindoro, Subaan). — Scheerpeltz, 1933: 1010 (Holosus; catalog). DISTRIBUTION: Philippines. brevipennis Fauvel, 1904b: 77 (Holosus; Type locality: Hong Kong; Hue; Tonkin; Inde; Sumatra). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1928d: 96 (Holosus; Sumatra). — Cameron, 1930: 50 (Holosus; characters; India; Sumatra; Hong Kong; Tonkin). — Scheerpeltz, 1933: 1010 (Holosus; catalog). — Paulian, 1941: 138 (Holosus; larval characters; India). — Abdullah and Qadri, 1970: 114 (Holosus; characters; West Pakistan). DISTRIBUTION: Hong Kong, Vietnam, Indonesia, India, Pakistan. transversus Cameron, 1925b: 232, 234 (Holosus; Type locality: Siwaliks, Dehra Dun). — Cameron, 1928d: 96 (Holosus; synonym of brevipennis). — Cameron, 1930: 50 (Holosus; synonym of brevipennis). — Scheerpeltz, 1933: 1010 (Holosus; synonym of brevipennis). bryanti Cameron, 1925b: 232, 234 (Holosus; Type locality: Borneo, Mt. Matang). — Cameron, 1928d: 96 (Holosus; Sumatra). — Scheerpeltz, 1933: 1010 (Holosus; catalog). DISTRIBUTION: Borneo, Indonesia. 1288 OSORIINAE: NEOLOSUS Bulletin AMNH congoensis Cameron, 1926a: 281 (Holosus; Type locality: Kimuela, Belgian Congo). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1952e: 827 (Holosus; Togo). — Fagel, 1960d: 267 (Holosus; characters; equatorial region of Africa). — Fagel, 1963a: 338 (Holosus; Guinea; Ivory Coast). — Scheerpeltz, 1974c: 5 (Holosus; Sudan). DISTRIBUTION: Sudan, Congo, Togo, Ivory Coast, Guinea. conuriformis Motschulsky, 1857a: 499 (Holosus; Type locality: Indes orientales). — Kraatz, 1859: 185 (Holosus). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1930: 45 (Holosus; characters; East Indies). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1934d: 77 (Holosus; notes on type). DISTRIBUTION: “East Indies”. cordicollis Bernhauer, 1926d: 256 (Holosus; Type locality: Borneo, Sandakan). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Borneo. cribripennis Bernhauer, 1914c: 84 (Holosus; Type locality: Hongkong). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Hong Kong. densus Bernhauer, 1922c: 221 (Holosus; Type locality: Taihorin, Kosempo). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1949b: 457 (Holosus; Formosa). DISTRIBUTION: Taiwan. distincticollis Bernhauer, 1932, see: basicollis Fauvel, 1903. distinguendus Cameron, 1925b: 233 (Holosus; Type locality: Tonkin, Hoabinh; N. India: W. Almora). — Cameron, 1930: 45 (Holosus; characters; Assam). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1934d: 77 (Holosus; synonym of conuriformis). — Coiffait, 1981f: 332 (Holosus; cited as valid species; Andaman Islands). DISTRIBUTION: India, Vietnam. drescheri Cameron, 1936a: 29 (Holosus; Type locality: Java: Batoerraden, G. Slamat). DISTRIBUTION: Indonesia. elegans Bernhauer, 1904: 15 (Holosus; Type locality: Sumatra, Si-Rambé). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1928d: 96 (Holosus; Sumatra). DISTRIBUTION: Indonesia. ecuadoriensis Irmler, 1999: 270 (Neolosus; Type locality: Ecuador: Pichinde, old Quinto- Sto. Domingo road, W. Chiriboza. Also cited from Colombia). DISTRIBUTION: Ecuador, Colombia. ferrugineus Bernhauer, 1926d: 258 (Holosus; Type locality: Siargao, Dapa). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Philippines. ferrugineus Cameron, 1938, see: Allotrochus. formosae Cameron, 1949b: 457 (Holosus; Type locality: Formosa: Musha). DISTRIBUTION: Taiwan. fossulatus Motschulsky, 1857a: 495 (Holotrochus; Type locality: Indes orientales). — Kraatz, 1859: 185 (Holosus). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1928d: 96 (Holosus; Sumatra). — Cameron, 1930: 48 (Holosus; characters; East Indies). Bulletin AMNH OSORIINAE: NEOLOSUS 1289
— Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1934d: 78 (Holosus; notes on type). DISTRIBUTION: “East Indies”, Sumatra. foveolatus Motschulsky, 1857a: 496 (Holotrochus; Type locality: Indes orientales). — Kraatz, 1859: 185 (Holosus). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Bernhauer, 1926d: 257 (Holosus; characters; Philippines). — Cameron, 1930: 48 (Holosus; characters; East Indies; Singapore; Borneo). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1934d: 78 (Holosus; notes on type). DISTRIBUTION: “East Indies”, Philippines, Borneo, Singapore. insularis Fauvel, 1904b: 77 (Holosus; Type locality: Java oriental: Monts Tengger, 1300 m; Borneo: Telang; Sumatra: Deli). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1928d: 96 (Holosus; Sumatra). — Bernhauer, 1928: 5 (Holosus; Sumatra). — Blackwelder, 1942: 90 (Holosus; subgenus Neolosus; checklist). DISTRIBUTION: Borneo, Indonesia. javanus Cameron, 1936a: 29 (Holosus; Type locality: Java: G. Patoeha, alt. 5000 ft. Preanger). DISTRIBUTION: Indonesia. longipennis Cameron, 1925b: 232, 235 (Holosus; Type locality: Nilgiri Hills; Anamalai Hills). — Cameron, 1930: 49 (Holosus; characters; India). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Biswas, 1986: 121 (Holosus; India). DISTRIBUTION: India. lopchuensis Cameron, 1945: 64 (Holosus; Type locality: Darjeeling: Lopchu, altitude 5000 feet). DISTRIBUTION: India. louwerensi Cameron, 1938b: 146 (Holosus; Type locality: E. Java: Res. Kediri, Slawe). DISTRIBUTION: Indonesia. luzonicus Bernhauer, 1926d: 258 (Holosus; Type locality: Luzon, Provinz Nueva Vizcaya, Imugan: Provinz Mountain, Balbalan). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Philippines. macropterus Bernhauer, 1937, see: singularis Gerstaeker, 1867. madurensis Bernhauer, 1914, see: Lispinus. malaisei Scheerpeltz, 1965: 121 (Holosus; Type locality: N.E. Burma: Kambaiti, 7000 ft). DISTRIBUTION: Myanmar. mycetoporiformis Motschulsky, 1857a: 500 (Holosus; Type locality: Indes orientales). — Kraatz, 1859: 185 (Holosus). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1930: 46 (Holosus; characters; Burma). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1934d: 78 (Holosus; notes on type and problems with identification). — Cameron, 1940: 85 (Holosus; characters). — Blackwelder, 1942: 90 (Holosus; subgenus Relinda; checklist). DISTRIBUTION: “East Indies”, Myanmar. nanus Cameron, 1940, see: Holotrochus. navicularis Cameron, 1936a: 28 (Holosus; Type locality: Java: G. Boerangrang; G. Goentoer; G. Patoeha; G. Tangkoeban Prahoe, alt. 4000–5000 ft; Batoerranden, G. Slamat; Sukabumi). DISTRIBUTION: Indonesia. 1290 OSORIINAE: NEOLOSUS Bulletin AMNH obscurus Irmler, 1999: 273 (Neolosus; Type locality: Peru: Pebas). DISTRIBUTION: Peru. olisthaeriformis Motschulsky, 1857a: 500 (Holosus; Type locality: Indes orientales). — Kraatz, 1859: 185 (Holosus). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1930: 47 (Holosus; characters; East Indies). — Scheerpeltz, 1933: 1011 (Holosus; catalog; Philippines). — Cameron, 1934d: 78 (Holosus; notes on type). — Cameron, 1940: 85 (Holosus; characters). — Blackwelder, 1942: 90 (Holosus; subgenus Relinda; checklist). DISTRIBUTION: “East Indies”, Philippines. palawanensis Bernhauer, 1926d: 257 (Holosus; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Philippines. pallidus Bernhauer, 1926d: 258 (Holosus; Type locality: Mindanao, Provinz Surigao, Surigao). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Philippines. papuanus Cameron, 1937a: 87 (Holosus; Type locality: New Guinea: Kokoda, alt. 1200 ft.). DISTRIBUTION: New Guinea. paradosus Bernhauer, 1926d: 256 (Holosus; Type locality: Mindanao, Provinz Surigao, Surigao). — Scheerpeltz, 1933: 1011 (Holosus; cited as paradoxus; catalog). DISTRIBUTION: Philippines. parallelus Cameron, 1950d: 2 (Holosus; Type locality: Kuala Lumpur). DISTRIBUTION: Malaysia. parcestriatus Fauvel, 1904b: 76 (Holosus; Type locality: Java occid.: Pengalengan, 1300 m). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Bernhauer, 1926d: 257 (Holosus; characters; Philippines, Java). — Cameron, 1928f: 400 (Holosus; Borneo). — Bernhauer, 1928: 5 (Holosus; Sumatra). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1936a: 28 (Holosus; Java). DISTRIBUTION: Borneo, Indonesia, Philippines. pennifer Fauvel, 1904b: 76 (Holosus; Type locality: Annam: Phuc-Son). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). DISTRIBUTION: Vietnam. philippinus Bernhauer, 1919: 360 (Holosus; Type locality: Palawan: P. Princesa). — Bernhauer, 1926d: 257 (Holosus; characters; Philippines). — Scheerpeltz, 1933: 1011 (Holosus; [Note: Scheerpeltz, 1933: 1326, also cited this name in Holisus]; catalog). DISTRIBUTION: Philippines. piceolus Bernhauer, 1926d: 258 (Holosus; Type locality: Luzon, Provinz Laguna, Los Baños). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Philippines. plicatus Bernhauer, 1904: 14 (Holosus; Type locality: Sumatra, Siboga). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1921a: 363, 399 (Holosus; characters; Singapore). — Bernhauer, 1926d: 257 (Holosus; characters; Philippines). — Cameron, 1928d: 96 (Holosus; Sumatra). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Singapore, Indonesia, Philippines. Bulletin AMNH OSORIINAE: NEOLOSUS 1291 politulus Fauvel, 1878f: 199 (Holosus; Type locality: Nouvelle-Guinée, Hatam). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). DISTRIBUTION: New Guinea. puncticollis Bernhauer, 1929b: 82 (Holosus; subspecies of tenuicornis; Type locality: Congo da Lemba; Uele: Van Kerkhovenville). — Scheerpeltz, 1933: 1012 (Holosus; variety of tenuicornis). — Bernhauer, 1942c: 347 (Holosus; cited as valid species; Ivory Coast). — Cameron, 1950g: 3 (Holosus; cited as valid species; Belgian Congo). DISTRIBUTION: Congo, Ivory Coast. quadricollis Cameron, 1925: 233, 235 (Holosus; Type locality: Selangor, Federated Malay States; The Gap). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1936a: 28 (Holosus; Java). DISTRIBUTION: Malaysia, Indonesia. reticulatus Fagel, 1960d: 268 (Holosus; Type locality: Congo: Bas-Uele: Koteli). DISTRIBUTION: Congo. rufotestaceus Bernhauer, 1926d: 258 (Holosus; Type locality: Luzon, Provinz Camarines Sur, Mount Isarog. Mindoro, Subaan). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Philippines. rugipennis Cameron, 1945: 64 (Holosus; Type locality: Bengal: Kalimpong, Samsingh, altitude 1800 feet). DISTRIBUTION: India. schoutedeni Bernhauer, 1929b: 82 (Holosus; Type locality: Mayumbe: Mekaia Ntete; Kasai: Makumbi; Ngombe; Haut Uele: Watsa; Kamalembi, Luebo). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Bernhauer, 1942c: 347 (Holosus; Cameroon). — Fagel, 1960d: 270 (Holosus; characters; equatorial Africa). — Levasseur, 1968a: 50 (Holosus; Central African Republic). DISTRIBUTION: Congo, Cameroon, Central African Republic. singularis Gerstaecker, 1867: 30 (Lispinus; Type locality: Ugono). — Fauvel, 1903e: 236 (Holosus; Cameroon; Togo; Congo Gabon; Sierra Leone; Senegal; Ethiopia; Usambara; Natal; Malaysia). — Fauvel, 1907: 11 (Holosus; Kilimandjaro). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Eichelbaum, 1913: 116 (Holosus; Tanzania). — Bernhauer, 1940: 130 (Holosus; Uganda). — Bernhauer, 1942c: 347 (Holosus; Cameroon). — Scheerpeltz, 1956a: 482 (Holosus; Belgian Congo). — Fagel, 1960d: 264 (Holosus; characters; tropical Africa). — Fagel, 1963a: 338 (Holosus; Ivory Coast). DISTRIBUTION: Senegal, Sierra Leone, Ivory Coast, Togo, Cameroon, Gabon, Congo, South Africa, Tanzania, Uganda, Ethiopia, Malaysia. macropterus Bernhauer, 1937b: 579 (Holosus; subspecies of singularis; Type locality: Ost-Usambara). — Fagel, 1960d: 264 (Holosus; variety of singularis). sinuatus Bernhauer, 1904, see: Lispinus. socius Bernhauer, 1926d: 256 (Holosus; Type locality: Mindanao, Provinz Lanao, Mumungan). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Philippines. 1292 OSORIINAE: NEOLOSUS Bulletin AMNH sparsipennis Cameron, 1949b: 457 (Holosus; Type locality: Formosa: Karenko). DISTRIBUTION: Taiwan. strigosus Cameron, 1926a: 282 (Holosus; Type locality: Tshela, Belgian Congo). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1952e: 827 (Holosus; Togo). — Fagel, 1960d: 267 (Holosus; characters; tropical Africa). — Levasseur, 1968a: 50 (Holosus; Central African Republic). — Scheerpeltz, 1971c: 162 (Holosus; Cameroon). DISTRIBUTION: Central African Republic, Congo, Cameroon, Togo. substriatus Fauvel, 1878f: 199 (Holosus; Type locality: Iles Arou, Wokan). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). DISTRIBUTION: Indonesia. sumatrensis Bernhauer, 1904: 15 (Holosus; Type locality: Sumatra). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1928d: 96 (Holosus; Sumatra). DISTRIBUTION: Indonesia. tachiniformis Motschulsky, 1857a: 498 (Holosus; Type locality: Indes orientales). — Kraatz, 1859: 185 (Holosus). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1928d: 96 (Holosus; Sumatra). — Cameron, 1930: 44 (Holosus; characters; East Indies). — Scheerpeltz, 1933: 1011 (Holosus; catalog; Malaysia). — Cameron, 1934d: 78 (Holosus; notes on type). DISTRIBUTION: “East Indies”, Indonesia, Malaysia. tachinoides Cameron, 1928g: 431 (Holosus; Type locality: Borneo: Mt. Murud; Tutau River; Kalabit Country). — Scheerpeltz, 1933: 1011 (Holosus; catalog). DISTRIBUTION: Borneo. tachyporiformis Motschulsky, 1857a: 498 (Holosus; Type locality: Indes orientales). — Kraatz, 1859: 185 (Holosus). — Fauvel, 1905e: 78 (Holosus; Java; Lombock; Sumatra; Borneo; Malacca; Burma). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Cameron, 1921a: 363, 399 (Holosus; characters; Singapore). — Cameron, 1928d: 96 (Holosus; Sumatra). — Cameron, 1930: 46 (Holosus; characters; East Indies). — Scheerpeltz, 1933: 1011 (Holosus; catalog). — Cameron, 1934d: 78 (Holosus; notes on type). — Cameron, 1936a: 28 (Holotrochus; Java). — Blackwelder, 1942: 90 (Holosus; subgenus Neolosus; checklist). — Coiffait, 1978b: 112 (Holosus; Bhutan). DISTRIBUTION: “East Indies”, Borneo, Indonesia, Singapore, Malaysia, Myanmar, Bhutan. tachyporoides Cameron, 1925b: 232, 233 (Holosus; Type locality: Selangor, Federated Malay States; The Gap). — Scheerpeltz, 1933: 1012 (Holosus; catalog). DISTRIBUTION: Malaysia. tenuicornis Fauvel, 1903e: 236 (Holosus; Type locality: Cameroun; Gabon: Libreville). — Bernhauer and Schubert, 1910: 20 (Holosus; catalog). — Scheerpeltz, 1933: 1012 (Holosus; catalog). — Scheerpeltz, 1956a: 482 (Holosus; Cameroon; French Congo; Belgian Congo). — Fagel, 1960d: 270 (Holosus; characters; central Africa). — Herman, 2001: 22 (Neolosus; synonymic notes). DISTRIBUTION: Congo, Cameroon, Gabon, French Equatorial Africa. Bulletin AMNH OSORIINAE: NEOLOSUS 1293
tenuicornis Bernhauer, 1904: 13 (Holosus; [preoccupied; Type locality: Africa occidentalis, Gaboon, Libreville). — Herman, 2001: 22 (Neolosus; synonym of tenuicornis Fauvel). tenuicornis Bernhauer, 1904, see: tenuicornis Fauvel, 1903. tirapensis Biswas and Biswas, 1985: 140 (Holosus; Type locality: India: Arunachal Pradesh, Namdapha Wild Life Sanctuary, Tirap dist.). DISTRIBUTION: India. transversus Cameron, 1925, see: brevipennis Fauvel, 1904. trichidus Irmler, 1994: 71 (Lispinus; Type locality: Bolivien: Yuracaris). — Irmler, 1999: 269 (Neolosus; characters; Bolivia). DISTRIBUTION: Bolivia. wittmeri Coiffait, 1978b: 136 (Holosus; Type locality: Phuntsholing-Thimphu, 1680 m). DISTRIBUTION: Bhutan. zischkai Scheerpeltz, 1951a: 13 (Holosus; Type locality: Chapare, 400 m). — Irmler, 1999: 271 (Neolosus; characters; Bolivia). DISTRIBUTION: Bolivia. Subtribe Thoracophorina Thoracophorina Reitter, 1909. — Arnett, 1963: 262 (characters; list of North American genera). — Burakowski and Newton, 1992: 38 (characters of adults, pupae, and larvae). — Newton and Thayer, 1992: 59 (subtribe of Thoracophorini). — Newton, Thayer, Ashe, and Chandler, 2000: 377 (characters; notes). Aneucamptus [2 species; Neotropical Region] Aneucamptus Sharp, 1887: 725 (species included: excisicollis). TYPE SPECIES: Thoracophorus excisicollis Motschulsky, fixed by monotypy. — Bernhauer and Schubert, 1910: 27 (world catalog; 1 species). — Blackwelder, 1944: 108 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 52 (type species: excisicollis). — Irmler, 1985: 41 (characters; key to neotropical species). dubitans Blackwelder, 1943, see: crassus Sharp, 1876. crassus Sharp, 1876: 419 (Thoracophorus, cited as Thoraxophorus; Type locality: St. Paulo). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Irmler, 1985: 45 (Aneucamptus; characters; Mexico; Panama; Trinidad; Brazil; Bolivia; Peru). DISTRIBUTION: Brazil, Bolivia, Peru, Panama, Mexico, Trinidad. dubitans Blackwelder, 1943: 153 (Thoracophorus; Type locality: Trinidad: cave in Aripo Valley, elevation 2600 feet). — Irmler, 1985: 45 (Aneucamptus; synonym of crassus). excisicollis Motschulsky, 1860a: 68 (Thoracophorus; Type locality: Panama). — Sharp, 1887: 725 (Aneucamptus; Panama). — Bernhauer and Schubert, 1910: 27 (Aneucamptus; catalog). — Irmler, 1985: 44 (Aneucamptus; characters; Panama). DISTRIBUTION: Panama. Dirocephalus [1 species; Neotropical Region] Dirocephalus Silvestri, 1938: 251 (species included: myrmecophilus). TYPE SPECIES: Diro- cephalus myrmecophilus Silvestri, fixed by original designation and monotypy. — Blackwelder, 1952: 129 (type species: myrmecophilus). — Herman, 1970: 425 (moved to Osoriinae). 1294 OSORIINAE: DIROCEPHALUS Bulletin AMNH myrmecophilus Silvestri, 1938: 254 (Dirocephalus; Type locality: Sapucai, Paraguay). DISTRIBUTION: Paraguay.
Euctenopsia [1 species; Neotropical Region] Euctenopsia Bruch, 1942: 137 (species included: ogloblini). TYPE SPECIES: Euctenopsia ogloblini Bruch, by monotypy. — Blackwelder, 1952: 155 (type species: ogloblini). — Herman, 1970: 425 (moved to Osoriinae). ogloblini Bruch, 1942: 137 (Euctenopsia; Type locality: Loreto, Misiones). DISTRIBUTION: Argentina.
Fauva [4 species; Neotropical Region] Fauva Blackwelder, 1952: 165 (replacement name for Diplopsis Fauvel). TYPE SPECIES: Diplopsis alternans Fauvel, fixed by objective synonymy with Diplopsis. Diplopsis Fauvel, 1902: 33 (species included: multicostata, alternans, laeviuscula; pre- occupied by Rafinesque, 1815). TYPE SPECIES: Diplopsis alternans Fauvel, fixed by subsequent designation by Blackwelder, 1952: 129. — Bernhauer and Schubert, 1910: 24 (world catalog; 4 species). — Blackwelder, 1942: 88 (type species: multicostata). — Blackwelder, 1944: 108 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 129 (type species). — Drugmand, 1998a: 207 (list of types in l’Institut royal des Sciences naturelles de Belgique). alternans Fauvel, 1902: 34 (Diplopsis; Type locality: Bahia). — Bernhauer and Schubert, 1910: 24 (Diplopsis; catalog; Brazil). DISTRIBUTION: Brazil. laevipennis Bernhauer, 1906c: 323 (Diplopsis; Type locality: Paraguay). — Bernhauer and Schubert, 1910: 24 (Diplopsis; catalog). DISTRIBUTION: Paraguay. laeviuscula Fauvel, 1902: 35 (Diplopsis; Type locality: Bahia). — Bernhauer and Schubert, 1910: 25 (Diplopsis; catalog; Brazil). DISTRIBUTION: Brazil. multicostata Fauvel, 1902: 34 (Diplopsis; Type locality: Bahia). — Bernhauer and Schubert, 1910: 25 (Diplopsis; catalog; Bahia). DISTRIBUTION: Brazil.
Mesotrochus [1 species; Neotropical Region] Mesotrochus Wasmann, 1890: 317 (species included: paradoxus). TYPE SPECIES: Mesotrochus paradoxus Wasmann, fixed by monotypy. — Bernhauer and Schubert, 1911: 146 (world catalog; 1 species). — Notman, 1925: 3 (characters in key). — Blackwelder, 1944: 112 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 239 (type species: paradoxus). paradoxus Wasmann, 1890: 317 (Mesotrochus; Type locality: Blumenau). — Bernhauer and Schubert, 1911: 146 (Mesotrochus; catalog). DISTRIBUTION: Brazil. Bulletin AMNH OSORIINAE: PARDIROCEPHALUS 1295
Pardirocephalus [2 species; Neotropical Region] Pardirocephalus Bruch, 1942: 130 (species included: cordobensis, loretoensis). TYPE SPECIES: Pardirocephalus cordobensis Bruch, by original designation. — Herman, 1970: 425 (moved to Osoriinae). cordobensis Bruch, 1942: 130 (Pardirocephalus; Type locality: La Granja, sierra de Alta Gracia, Cordoba, with Pheidole obtusopilosa). DISTRIBUTION: Argentina. loretoensis Bruch, 1942: 132 (Pardirocephalus; Type locality: Loreto, Misiones, with Pheidole). DISTRIBUTION: Argentina. Pselaphomimus [1 species; Neotropical Region] Pselaphomimus Bruch, 1942: 134 (species included: amphiphilus). TYPE SPECIES: Pselphomimus amphiphilus Bruch, 1942, by monotypy. — Herman, 1970: 425 (moved to Osoriinae). amphiphilus Bruch, 1942: 136 (Pselaphomimus; Type locality: La Pampa; Rosas, Buenos Aires). DISTRIBUTION: Argentina. Rhopalopherus [2 species; Neotropical Region] Rhopalopherus Bernhauer, 1909c: 227 (species included: gestroi). TYPE SPECIES: Rhopalo- pherus gestroi Bernhauer, fixed by monotypy. — Bernhauer and Schubert, 1910: 25 (world catalog; 1 species). — Scheerpeltz, 1933: 1020 (world catalog supplement). — Blackwelder, 1944: 108 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 340 (type species: gestroi). gestroi Bernhauer, 1909c: 228 (Rhopalopherus; Type locality: Argentinien: Buenos Aires). — Bernhauer and Schubert, 1910: 25 (Rhopalopherus; catalog). — Frank, 1982: 8 (Rhopalopherus; list of parasites). DISTRIBUTION: Argentina. magnipennis Bernhauer, 1925: 33 (Rhopalopherus; Type locality: Argentinien: S. Fé, Fives Lilles). — Scheerpeltz, 1933: 1020 (Rhopalopherus; catalog). DISTRIBUTION: Argentina.
Teiros [1 species; Oriental Region] Teiros Eichelbaum, 1909: 126 (replacement name for Teras Bernhauer). TYPE SPECIES: Teras mirabilis Bernhauer, fixed by objective synonymy with Teras. — Bernhauer and Schubert, 1911: 146 (world catalog; 1 species). — Notman, 1925: 2 (characters in key). — Scheerpeltz, 1933: 1137 (world catalog supplement). — Blackwelder, 1952: 375 (type species). Teras Bernhauer, 1905: 15 (species included: mirabile; preoccupied by Treitschke, 1829, and Hartig, 1840). TYPE SPECIES: Teras mirabile Bernhauer, fixed by monotypy. — Bernhauer and Schubert, 1911: 146 (synonym of Teiros). — Blackwelder, 1952: 376 (type species: mirabile). 1296 OSORIINAE: TEIROS Bulletin AMNH mirabilis Bernhauer, 1905: 16 (Teras; Type locality: Engano). — Bernhauer and Schubert, 1911: 146 (Teiros; catalog). — Cameron, 1928d: 110 (Teiros; Sumatra). — Scheerpeltz, 1933: 1137 (Teiros; catalog). DISTRIBUTION: Indonesia. Tetrapleurus [22 species; Palaearctic, Oriental, and Oceanic Regions] Tetrapleurus Bernhauer, 1914c: 84 (species included: indicus). TYPE SPECIES: Tetrapleurus indicus Bernhauer, fixed by monotypy. — Bernhauer, 1916b: 19 (characters). — Cameron, 1925: 8, 103 (catalog of Indian species). — Bernhauer, 1926d: 262 (key to species of Philippines). — Cameron, 1930: 68 (characters; key to British Indian species). — Scheerpeltz, 1933: 1019 (world catalog supplement). — Blackwelder, 1952: 383 (type species: indicus). — Scheerpeltz, 1970b (key to species). — Shibata, 1973c: 24 (checklist of species of Taiwan). — Hammond, 1984: 199 (checklist of species of Borneo). — Drugmand, 1998a: 223 (list of types in l’Institut royal des Sciences naturelles de Belgique). atriceps Cameron, 1928d: 97 (Tetrapleurus; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 148 (Tetrapleurus; characters; Sumatra). DISTRIBUTION: Indonesia. bakeri Bernhauer, 1926d: 262 (Tetrapleurus; Type locality: Luzon, Provinz Laguna, Mount Maquiling; Los Baños; Mount Banahao: Provinz Mountain, Balbalan. Mindanao, Provinz Surigao, Surigao). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 154 (Tetrapleurus; characters; Philippines). DISTRIBUTION: Philippines. banghaasi Bernhauer, 1926d: 263 (Tetrapleurus; Type locality: Mindanao, Provinz Lanao, Mumungan). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 150 (Tetrapleurus; characters; Philippines). DISTRIBUTION: Philippines. banosanus Bernhauer, 1926d: 263 (Tetrapleurus; Type locality: Luzon, Provinz Laguna, Los Baños). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 146 (Tetrapleurus; characters; Philippines). DISTRIBUTION: Philippines. crenicollis Cameron, 1932a: 124 (Tetrapleurus; Type locality: Pahang: Fraser’s Hill, alt. 4000 feet). — Scheerpeltz, 1970b: 151 (Tetrapleurus; characters; Malaysia). DISTRIBUTION: Malaysia. denticollis Cameron, 1940c: 211 (Tetrapleurus; Type locality: Dehra Dun: Asan River). — Scheerpeltz, 1970b: 145 (Tetrapleurus; characters; India). DISTRIBUTION: India. formosae Bernhauer, 1916b: 20 (Tetrapleurus; Type locality: Formosa: Kosempo). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 153 (Tetrapleurus; characters; Formosa). DISTRIBUTION: Taiwan. fuscipennis Cameron, 1928f: 401 (Tetrapleurus; Type locality: Mt. Penrissen, 4000 feet). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 152 (Tetrapleurus; characters; Borneo). DISTRIBUTION: Borneo. Bulletin AMNH OSORIINAE: TETRAPLEURUS 1297 heteroderes Fauvel, 1902: 30 (Lispinodes; Type locality: Java occidental: Pengalengan, 1300 m). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Bernhauer, 1926d: 263 (Tetrapleurus). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Cameron, 1936a: 30 (Tetrapleurus; Java). — Scheerpeltz, 1970b: 150 (Tetrapleurus; characters; Java). DISTRIBUTION: Indonesia. himalayicus Cameron, 1924b: 161 (Tetrapleurus; Type locality: Mussorie District, Kaligad). — Cameron, 1930: 71 (Tetrapleurus; characters; India). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Abdullah and Qadri, 1970: 114 (Tetrapleurus; characters; West Pakistan). — Scheerpeltz, 1970b: 149 (Tetrapleurus; characters; India). — Legner and Moore, 1977: 174 (Tetrapleurus; found under bark). — Coiffait, 1982: 41 (Tetrapleurus; Nepal). DISTRIBUTION: Pakistan, India, Nepal. indicus Bernhauer, 1914c: 85 (Tetrapleurus; Type locality: Chambaganoor, Madura, südliches Ostindien). — Cameron, 1930: 69 (Tetrapleurus; characters; India). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 152 (Tetrapleurus; characters; India). DISTRIBUTION: India. malayanus Cameron, 1929b: 440 (Tetrapleurus; Type locality: Selangor: The Gap). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 149 (Tetrapleurus; characters; Malaysia). DISTRIBUTION: Malaysia. niasensis Scheerpeltz, 1970b: 155 (Tetrapleurus; Type locality: Insel Nias). DISTRIBUTION: Indonesia. opacus Bernhauer, 1926d: 263 (Tetrapleurus; Type locality: Luzon, Provinz Laguna, Los Baños). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 146 (Tetrapleurus; characters; Philippines). DISTRIBUTION: Philippines. parallelus Bernhauer, 1917e: 41 (Tetrapleurus; Type locality: Formosa: Takao). — Bernhauer, 1922c: 222 (Tetrapleurus; Formosa). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 154 (Tetrapleurus; characters; Formosa). — Naomi, 1996: 52 (Tetrapleurus; cited as paralellus; Taiwan Japan). DISTRIBUTION: Taiwan. rugosus Fauvel, 1902: 31 (Lispinodes; Type locality: Sumatra: Monts Singalang; Java: Tcibodas). — Fauvel, 1904a: 44 (Lispinodes; Nilghiris; Sumatra; Java). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Bernhauer, 1926d: 263 (Tetrapleurus). — Cameron, 1928d: 97 (Tetrapleurus; Sumatra). — Bernhauer, 1928: 5 (Tetrapleurus; Sumatra). — Cameron, 1930: 72 (Tetrapleurus; characters; India; Sumatra; Java). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 145 (Tetrapleurus; characters; Sumatra; Java; Sunda Islands). DISTRIBUTION: India, Indonesia. sauteri Bernhauer, 1916b: 19 (Tetrapleurus; Type locality: Kosempo). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 153 (Tetrapleurus; characters; Formosa). DISTRIBUTION: Taiwan. 1298 OSORIINAE: TETRAPLEURUS Bulletin AMNH sculptus Kraatz, 1859: 188 (Lispinus; Type locality: Ceylan). — Bernhauer and Schubert, 1910: 23 (Lispinus; catalog). — Cameron, 1930: 70 (Tetrapleurus; characters; Ceylon). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 144 (Tetrapleurus; characters; Ceylon). DISTRIBUTION: Sri Lanka. semiopacus Lea, 1927: 273 (Tetrapleurus; Type locality: Fiji: Taveuni). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 155 (Tetrapleurus; characters; Taveuni Island). DISTRIBUTION: Fiji. seriatus Fauvel, 1902: 31 (Lispinodes; Type locality: Bírmanie; Sumatra; Borneo: Sarawak). — Bernhauer and Schubert, 1910: 20 (Lispinodes; catalog). — Cameron, 1928d: 97 (Tetrapleurus; Sumatra). — Cameron, 1930: 69 (Tetrapleurus; characters; Burma; Sumatra; Borneo). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 147 (Tetrapleurus; characters; Sumatra; Borneo). DISTRIBUTION: Myanmar, Indonesia, Borneo. splendidus Bernhauer, 1926d: 263 (Tetrapleurus; Type locality: Palawan, Binaluan). — Scheerpeltz, 1933: 1019 (Tetrapleurus; catalog). — Scheerpeltz, 1970b: 148 (Tetrapleurus; characters; Philippines). DISTRIBUTION: Philippines. subnitidus Motschulsky, 1860a: 66 (Thoracophorus; Type locality: Indes orientales). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Cameron, 1930: 73 (Thoracophorus; characters; East Indies). — Scheerpeltz, 1933: 1021 (Thoracophorus; catalog). — Cameron, 1934d: 78 (transfer from Thoracophorus to Tetrapleurus). — Cameron, 1940: 86 (Tetrapleurus; transferred from Thoracophorus; characters; Ceylon). — Scheerpeltz, 1970b: 147 (Tetrapleurus; characters; India). DISTRIBUTION: “East Indies”, India, Sri Lanka. Thoracophorus [45 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions] Thoracophorus Motschulsky, 1837: 98 (species included: corticinus; cited as Thoraxophorus and later justifiably emended). TYPE SPECIES: Thoracophorus corticinus Motschul- sky, fixed by monotypy. — Erichson, 1840: 908 (cited as Thoraxophorus; synonym of Glyptoma) — Motschulsky, 1840: 197 (emendation of Thoraxophorus which was an error of transcription to Thoracophorus). — Schaum, 1852: 31 (cited as Glyptoma; list of species; Europe). — Lacordaire, 1854: 148 (cited as Thoraxophorus; synonym of Glyptoma). — Jacquelin du Val, 1857: 62 (characters; cited as Glyptoma). — Redtenbacher, 1857: 258 (cited as Glyptoma; characters). — Kraatz, 1857d: 1047 (cited as Thoraxophorus; synonym of Glyptoma). — Motschulsky, 1860a: 66 (Thoracophorus is correct name; discussion). — Fauvel, 1871a: 40 [= 1872: 14] (characters). — Redtenbacher, 1874: 284 (characters; cited as Glyptoma). — Fauvel, 1875a: I [= 1875b: 203] (catalog). — Sharp, 1876: 418 (cited as Thoraxophorus; notes). — Fauvel, 1878d: 175 [= 1878e: 11] (characters). — Mulsant and Rey, 1878: 834 (characters; cited as Glyptoma). — Duvivier, 1883: 204 (catalog; cited as Thoraxophorus). — Sharp, 1887: 725 (notes; Mexican and Central American species). — Heyden, Reitter, and Weise, 1891: 122 (list of species of Europe and Caucasus). — Ganglbauer, 1895: 687 (characters). Bulletin AMNH OSORIINAE: THORACOPHORUS 1299
— Heyden, Reitter, and Weise, 1906: 125 (list of species of Europe and Caucasus). — Bernhauer and Schubert, 1910: 25 (world catalog; 27 species). — Petri, 1912: 50 (list of species of Siebenbürgen). — Leng, 1920: 93 (catalog of North American species). — Winkler, 1925: 323 (catalog for Palaearctic region). — Cameron, 1925: 8 (catalog of Indian species). — Bernhauer, 1926d: 260 (key to species of Philippines). — Porta, 1926: 6 (characters; key to species of Italy). — Bruch, 1928a: 422 (catalog; Argentina). — Cameron, 1930: 72 (characters). — Roubal, 1930: 299 (catalog; Slovakia). — Scheerpeltz, 1933: 1020 (world catalog supplement). — Blackwelder, 1943: 148 (characters; type species: corticinus; key to species of West Indies). — Blackwelder, 1944: 108 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 388 (type species). — Székessy, 1963: 7 (characters). — Coiffait and Saiz, 1968: 460 (characters; Chilean species). — Scheerpeltz, 1968a: 3 (catalog of Austrian species). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Moore and Legner, 1974b: 83 (characters; notes). — Moore and Legner, 1975: 171 (catalog; North America north of Mexico). — Shibata, 1976: 84 (checklist of species of Japan). — Burakowski, Mroczkowski, and Stefan´ska, 1979: 7 (catalog; cited as Thoraxophorus; Poland). — Moore and Legner, 1979: 149 (characters; notes). — M. Dvorˇák, 1979: 109 (some species collected in Slovakia). — Tóth, 1982: 13 (characters; habitus; species of Hungary). — Irmler, 1985: 41 (characters; key to Neotropical species). — Frank, 1986a: 365 (checklist of species of Florida). — Newton, 1990: 92 (characters in key; three described and two undescribed Australian species). — Burakowski and Newton, 1992 (systematic position; characters of immatures stages; discus- sion). — Newton and Thayer, 1992: 18 (correct spelling: Thoracophorus; discussion of original and emended spelling of name). — Bohácˇ, 1993: 40 (list of species of Czech Republic and Slovakia). — Zanetti, 1995: 4 (list of species of Italy). — Downie and Arnett, 1996: 427 (characters). — Lundgren, 1998: 46 (list of species from Florida; records for other States listed for each species). — Drugmand, 1998a: 223 (list of types in l’Institut royal des Sciences naturelles de Belgique). — Newton, Thayer, Ashe, and Chandler, 2000: 377 (377 Nearctic species; notes; characters in key). Thoraxophorus Motschulsky, 1837: 98 [Note: Nomen nudum, an incorrect original spelling (Article 19.1). Stilbogastrus Bernhauer, 1908d: 286 (subgenus of Thoracophorus; species included: nitidus). TYPE SPECIES: Thoracophorus (Stilbogastrus) nitidus Bernhauer, fixed by original designation and monotypy. — Bernhauer and Schubert, 1910: 26 (subgenus of Thoracophorus; catalog; 1 species). — Blackwelder, 1943: 148 (subgenus of Thoracophorus; type species nitidus). — Blackwelder, 1952: 361 (subgenus of Thoracophorus; type species: nitidus). Leipophorus Bernhauer, 1926d: 261 (subgenus of Thoracophorus; species included: minutissimus, alluaudi). TYPE SPECIES: Thoracophorus (Leipophorus) minutissimus Bernhauer, fixed by subsequent designation by Blackwelder, 1942: 88. — Blackwelder, 1943: 148 (subgenus of Thoracophorus; type species minutissimus). — Blackwelder, 1952: 213 (subgenus of Thoracophorus; type species). 1300 OSORIINAE: THORACOPHORUS Bulletin AMNH aequalis Sharp, 1887: 727 (Thoracophorus; Type locality: Guatemala, San Geronimo, Balheu). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Irmler, 1985: 53 (Thoracophorus; characters; Mexico; Guatemala; Panama; Venezuela; Ecuador). DISTRIBUTION: Mexico, Guatemala, Panama, Venezuela, Ecuador. alluaudi Fauvel, 1898a: 115 (Thoracophorus; Type locality: La Digue). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — H. Kolbe, 1910: 21 (Thoracophorus; Seychelles). — Bernhauer, 1922b: 166 (Thoracophorus; Seychelles). — Bernhauer, 1926d: 261 (Thoracophorus; subgenus Leipophorus). — Scheerpeltz, 1933: 1021 (Thoracophorus; subgenus Leipophorus; catalog). DISTRIBUTION: Seychelles. argentinus Bernhauer, 1921d: 170 (Thoracophorus; Type locality: Argentinien: Misiones). — Scheerpeltz, 1933: 1020 (Thoracophorus; catalog). — Irmler, 1985: 57 (Thoracophorus; characters; Paraguay; Brazil; Argentina). DISTRIBUTION: Argentina, Paraguay, Brazil. conicicollis Wendeler, 1955: 191 (Thoracophorus; Type locality: Brasilien: Nova Teutonia). — Irmler, 1985: 57 (Thoracophorus; synonym of argentinus). blackburni Sharp, 1880: 53 (Glyptoma; Type locality: Not cited but presumably Hawaii). — Blackburn and Sharp, 1885: 224 (Thoracophorus; habitat; Hawaii). — Sharp, 1908: 538 (Thoracophorus; characters; Hawaii). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Moore, 1975c: 106 (Thoracophorus; Hawaii). — Nishida, 1994: 74 (Thoracophorus; Hawaii). — Nishida, 1997: 63 (Thoracophorus; Hawaii). DISTRIBUTION: Hawaii. bonvouloiri Fauvel, 1867a: 54 [= 1868: 59] (Glyptoma; Type locality: Santiago). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Coiffait and Saiz, 1968: 461 (Thoracophorus; characters; Chile). — Irmler, 1985: 57 (Thoracophorus; characters; Chile). DISTRIBUTION: Chile. brasilianus Irmler, 1985: 45 (Thoracophorus; Type locality: Brasilien: S. Catharina, Nova Teutonia. Also cited from Bolivia). DISTRIBUTION: Bolivia, Brazil. brevicristatus Horn, 1871a: 332 (Glyptoma; Type locality: Camp Grant, Arizona). — Fauvel, 1878d: 176 [= 1878e: 12] (Thoracophorus; characters; Arizona; Mexico). — Sharp, 1887: 726 (Thoracophorus). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Scheerpeltz, 1933: 1020 (Thoracophorus; catalog). — Blackwelder, 1943: 150 (Thoracophorus; characters; Cuba; Puerto Rico; St. Thomas; St. Croix; Guadeloupe). — Legner and Moore, 1977: 174 (Thoracophorus; found under bark). — Bohácˇ, 1978 (Thoracophorus; larval and pupal characters; discussion of taxonomic position; Cuba). — Irmler, 1985: 48 (Thoracophorus; characters; Mexico; Costa Rica; Cuba; Martinique; St. Thomas; Guadeloupe). DISTRIBUTION: USA, Mexico, Costa Rica, West Indies, Mascarene Islands, Philippines. deletus Fauvel, 1902: 32 (Thoracophorus; variety of brevicristatus; Type locality: La Réunion: Salazie; Philippines: Bato). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; synonym of brevicristatus). — Bernhauer, 1926d: 260 (Thoracophorus; variety of brevicristatus; characters; Philippines). — Scheerpeltz, 1933: 1020 (Thoracophorus; variety of brevicristatus). Bulletin AMNH OSORIINAE: THORACOPHORUS 1301 brevipennis Fauvel, 1889, see: curtipennis Fauvel, 1903. brevipennis Sharp, 1880: 53 (Glyptoma; Type locality: mountains of Oahu). — Blackburn and Sharp, 1885: 224 (Thoracophorus; habitat; Hawaii). — Sharp, 1908: 539 (Thoracophorus; Hawaii). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Nishida, 1994: 74 (Thoracophorus; Hawaii). — Nishida, 1997: 63 (Thoracophorus; Hawaii). DISTRIBUTION: Hawaii. bruchi Bernhauer, 1933e: 326 (Thoracophorus; Type locality: Argentinien: Misiones, Loreto). — Irmler, 1985: 51 (Thoracophorus; characters; Brazil; Ecuador). DISTRIBUTION: Argentina, Brazil, Ecuador. camponoti Irmler, 1985: 50 (Thoracophorus; Type locality: Brasilien: São Paulo). DISTRIBUTION: Brazil. certatus Sharp, 1889: 465 (Thoracophorus; Type locality: Nishimura, Oyayama, Kashiwagi, Nara). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Nakane and Sawada, 1960: A122 (Thoracophorus; Yakushima Island). — Watanabe, 1961: 348 (Thoracophorus; Izu Islands). — Nakane, 1963a: 81 (Thoracophorus; characters; habitus photograph; Japan). — Watanabe and Shibata, 1972: 60 (Thoracophorus; Yaku-shima Island). — Shibata, 1993a: 317 (Thoracophorus; Taiwan). — Naomi, 1996: 53 (Thoracophorus; Japan). DISTRIBUTION: Japan, Taiwan. conicicollis Wendeler, 1955, see: argentinus Bernhauer, 1921. corticinus Motschulsky, 1837: 98 (Thoracophorus; Type locality: Fontainebleau). — Erichson, 1840: 909 (Glyptoma; characters; Italy; France). — Redtenbacher, 1849: 757 (Glyptoma; characters; Austria). — Fairmaire and Laboulbène, 1856: 658 (Glyptoma; characters; France). — Redtenbacher, 1857: 259 (Glyptoma; characters; Austria). — Kraatz, 1857d: 1048 (Glyptoma; characters; Germany). — Schaum, 1859: 33 (Glyptoma; catalog; Europe). — Jacquelin du Val, 1868: 78 (Glyptoma; catalog; Europe). — Fauvel, 1871a: 41 [= 1872: 15] (Thoracophorus; characters; France; Germany; Austria; Italy). — Redtenbacher, 1874: 285 (Glyptoma; characters; Austria). — Seidlitz, 1875: 231 (Glyptoma; characters; Baltic region). — Mulsant and Rey, 1878: 836 (Glyptoma; characters; France). — Seidlitz, 1891: 336 (Glyptoma; characters; Europe). — Seidlitz, 1891a: 358 (Glyptoma; characters; Europe). — Ganglbauer, 1895: 688 (Thoracophorus; characters; middle Europe). — Sainte-Claire Deville, 1907: 12 (Thoracophorus; characters; France). — Reitter, 1909: 200 (Thoracophorus; characters; Germany). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Portevin, 1929: 459 (Thoracophorus; characters; France). — Scheerpeltz, 1933: 1020 (Thoracophorus; catalog). — Horion, 1963: 14 (Thoracophorus; Greece; Bosnia Hercegovina; Yugoslavia; Dalmatia; Italy; Romania; Slovakia; Hungary; Czech Republic; Poland; Austria; Germany; France; Switzerland; Italy). — Lohse, 1964: 19 (Thoracophorus; characters; central Europe). — Legner and Moore, 1977: 174 (Thoracophorus; found under bark). — Tóth, 1982: 14 (Thoracophorus; characters; Hungary). — Lucht, 1987: 84 (Thoracophorus; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux). — Burakowski and Newton, 1992 (Thoracophorus; egg, larval, pupal and adult characters; bionomics; discussion). — Franc, 1993: 307 (Thoracophorus; synoecious myrmecophile; Slovakia). 1302 OSORIINAE: THORACOPHORUS Bulletin AMNH
— Zanetti, 1995: 4 (Thoracophorus; Italy). — Bohácˇ, 1993: 40 (Thoracophorus; Czech Republic; Slovakia). — Segers, 1986: 17 (Thoracophorus; checklist of species; Belgium). DISTRIBUTION: Europe. costalis Erichson, 1840: 909 (Glyptoma; Type locality: America septentrionalis). — LeConte, 1863: 27 (Glyptoma; North America). — Fauvel, 1878d: 175 [= 1878e: 11] (Thoracophorus; characters; Michigan; Pennsylvania; Massachusetts; New Jersey; California; South Carolina; Florida; Louisiana; Texas). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Blatchley, 1910: 484 (Glyptoma; characters; Indiana). — Notman, 1920b: 693 (Thoracophorus; Florida). — Irmler, 1985: 49 (Thoracophorus; characters; Mexico). — Campbell and Davies, 1991: 92 (Thoracophorus; checklist of species; Canada). — Downie and Arnett, 1996: 427 (Thoracophorus; characters; USA). DISTRIBUTION: USA, Mexico. crassus Sharp, 1876, see: Aneucamptus. crenicollis Fauvel, 1878, see: Pseudespeson. curtipennis Fauvel, 1903d: 256 (Thoracophorus; replacement name for brevipennis Fauvel). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). DISTRIBUTION: New Caledonia. brevipennis Fauvel, 1889b: 256 (Thoracophorus; [preoccupied]; Type locality: Yahoué). — Fauvel, 1903d: 256 (Thoracophorus; synonym of curtipennis). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; synonym of curtipennis). deletus Fauvel, 1902, see: brevicristatus Horn, 1871. denticollis Erichson, 1840: 910 (Glyptoma; Type locality: Puerto Rico). — Fauvel, 1864b: 60 [= 1865: 64] (Glyptoma; characters; Puerto Rico). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Blackwelder, 1943: 154 (Thoracophorus; characters; Puerto Rico). — Irmler, 1985: 46 (Thoracophorus; characters; Puerto Rico; Peru). DISTRIBUTION: Peru, West Indies. discretus Sharp, 1887: 727 (Thoracophorus; Type locality: Guatemala, El Tumbador; Panama, Bugaba). — Bernhauer and Schubert, 1910: 25 (Thoracophorus; subgenus Thoracophorus; catalog). — Irmler, 1985: 56 (Thoracophorus; characters; Guatemala; Panama). — Marcuzzi, 1977: 5 (Thoracophorus; Venezuela). DISTRIBUTION: Guatemala, Panama, Venezuela. dubitans Blackwelder, 1943, see: Aneucamptus. duplicatus Fauvel, 1878, see: Lispinodes. elongatus Cameron, 1938c: 173 (Thoracophorus; Type locality: Mauritius: Gorges, Riv. Noire). — Jarrige, 1957: 112 (Thoracophorus; Réunion). DISTRIBUTION: Mascarene Islands. excisicollis Motschulsky, 1860, see: Aneucamptus. exilis Erichson, 1840: 910 (Glyptoma; Type locality: Americae merid. ins. St. Thoma). — Fauvel, 1864b: 61 [= 1865: 66] (Glyptoma; characters; St. Thomas). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Blackwelder, 1943: 153 (Thoracophorus; characters; St. Thomas). — Irmler, 1985: 51 (Thoracophorus; characters; Guadeloupe; St. Lucia; Dominica). DISTRIBUTION: West Indies. filum Sharp, 1887: 728 (Thoracophorus; Type locality: Guatemala, El Tumbador, El Reposo, Zapote). Bulletin AMNH OSORIINAE: THORACOPHORUS 1303
— Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Irmler, 1985: 54 (Thoracophorus; characters; Mexico; Panama; Guatemala; Costa Rica; Brazil). DISTRIBUTION: Mexico, Guatemala, Costa Rica, Brazil. fletcheri Wendeler, 1927: 5 (Thoracophorus; Type locality: Minnesota: Lake Minnetonka [Note: Erroneously cited by Wendeler, 1927 as New York; see Fletcher, 1930]). — Fletcher, 1930: 190 (Thoracophorus; type locality is: Lake Minnetonka, Hennepin County, Minnesota). — Scheerpeltz, 1933: 1020 (Thoracophorus; catalog). DISTRIBUTION: USA. galapagosus Coiffait, 1981g: 306 (Thoracophorus; Type locality: Ile Galapagos: Ile Santa Cruz, 1200 m). DISTRIBUTION: Galapagos Islands. guadalupensis Cameron, 1913: 323 (Thoracophorus; Type locality: Grenada; Haiti). — Scheerpeltz, 1933: 1021 (Thoracophorus; catalog). — Blackwelder, 1943: 151 (Thoracophorus; characters; Cuba; Jamaica; Haiti; Puerto Rico; Dominica; St. Lucia; St. Vincent; Grenada; Trinidad). — Legner and Moore, 1977: 174 (Thoracophorus; found under bark). — Irmler, 1985: 54 (Thoracophorus; cited as gouadalupensis; characters; Mexico; Guadeloupe; Grenada; St. Thomas; Dominica; St. Vincent; Cuba; Panama; British Honduras; Costa Rica; Peru; Brasil). DISTRIBUTION: West Indies, Belize, Costa Rica, Panama, Peru, Brazil. heyeri Wasmann, 1902: 101 (Thoracophorus; Type locality: São Leopoldo, Rio Grande do Sul). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Seevers, 1957: 318 (Thoracophorus; doubtfully termitophilous). DISTRIBUTION: Brazil. inconspicuus Bernhauer, 1929b: 441 (Glyptoma; Type locality: Selangor: The Gap). — Scheerpeltz, 1933: 1021 (Thoracophorus; catalog). DISTRIBUTION: Malaysia. javanus Cameron, 1940a: 103 (Thoracophorus; Type locality: E. Java: Djember). DISTRIBUTION: Indonesia. longicollis Motschulsky, 1860a: 68 (Thoracophorus; Type locality: Nouvelle-Orléans). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). DISTRIBUTION: USA. longipennis Bernhauer, 1926d: 260 (Thoracophorus; Type locality: Luzon, Provinz Nueva Vizcaya, Imugan. Catanduanes, Virac). — Scheerpeltz, 1933: 1021 (Thoracophorus; catalog). DISTRIBUTION: Philippines. minutissimus Bernhauer, 1926d: 261 (Thoracophorus; subgenus Leipophorus; Type locality: Luzon, Provinz Ilocos Norte, Bangui). — Scheerpeltz, 1933: 1021 (Thoracophorus; subgenus Leipophorus; catalog). DISTRIBUTION: Philippines. minutus Cameron, 1950f: 112 (Thoracophorus; Type locality: Angola: Dundo). DISTRIBUTION: Angola. montanus Cameron, 1945: 65 (Thoracophorus; Type locality: Darjeeling: Ghum district: Mangpu). DISTRIBUTION: India. myrmecophilus Lea, 1910a: 141 (Glyptoma; Type locality: Victoria: Forrest; [Note: See lectotype designation by Newton, 1990]). — Scheerpeltz, 1933: 1021 (Calocerus; catalog). — Newton, 1990: 90, 93 (Thoracophorus; lectotype designation; collecting notes; Australia). DISTRIBUTION: Australia. 1304 OSORIINAE: THORACOPHORUS Bulletin AMNH
quadricostatus Bernhauer, 1920: 4 (Thoracophorus; Type locality: Logan Village; Atherton; Bellenden Ker.; Mt. Tambourine). — Scheerpeltz, 1933: 1021 (Thoracophorus; catalog). — Newton, 1990: 90 (Thoracophorus; synonym of myrmecophilus). nitidus Bernhauer, 1908d: 285 (Thoracophorus; subgenus Stilobogastrus; Type locality: Brasilien: S. Catharina). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Stilobogastrus; catalog). — Irmler, 1985: 46 (Thoracophorus; characters; Brazil). DISTRIBUTION: Brazil. obscurior Bernhauer, 1926 [Note: Nomen nudum], see: philippinus Bernhauer, 1926. opacus Sharp, 1876, see: Glyptoma. pallidus Sharp, 1887: 728 (Thoracophorus; Type locality: Guatemala, Cerro Zunil). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). DISTRIBUTION: Guatemala. peruvianus Bernhauer, 1941c: 278 [= 1951: 268] (Thoracophorus; Type locality: Süd-Peru: Sivia, 520 m). DISTRIBUTION: Peru. philippinus Bernhauer, 1926d: 261 (Thoracophorus; Type locality: Luzon, Provinz Laguna, Los Baños; Provinz Nueva Vizcaya, Imugan. Mindoro, San Teodoro. Siargao, Dapa). — Scheerpeltz, 1933: 1021 (Thoracophorus; catalog). DISTRIBUTION: Philippines. obscurior Bernhauer, 1926d: 261 (Thoracophorus; aberration of philippinus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; locality cited: Butao, Mount Banahao, Luzon). — Scheerpeltz, 1933: 1021 (Thoracophorus; synonym of philippinus). proximus Irmler, 1985: 51 (Thoracophorus; Type locality: Mexico: Veracruz, Canyon, Rio Metlac nr. Fortin, 3100 feet. Also cited from Panama). DISTRIBUTION: Mexico. pusillus Sharp, 1887: 727 (Thoracophorus; Type locality: Guatemala, Teleman). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Irmler, 1985: 56 (Thoracophorus; characters; Guatemala). DISTRIBUTION: Guatemala. quadricostatus Bernhauer, 1920, see: myrmecophilus Lea, 1910. rectangulus Blackwelder, 1943, see: rufulus Cameron, 1929. ruficollis Fauvel, 1864, see: sculptilis Erichson, 1840. rufulus Cameron, 1929c: 599 (Thoracophorus; Type locality: British Guiana: Georgetown, Botanic Gardens). — Scheerpeltz, 1933: 1021 (Thoracophorus; catalog). — Irmler, 1985: 49 (Thoracophorus; characters; Grenada; Trinidad; British Guiana; Brazil). DISTRIBUTION: Brazil, Guyana, West Indies. rectangulus Blackwelder, 1943: 150 (Thoracophorus; Type locality: Trinidad, 10 miles north of Arima, elevation 1800 feet). — Irmler, 1985: 49 (Thoracophorus; synonym of rufulus). sahlbergi Irmler, 1985: 50 (Thoracophorus; Type locality: Brasilien: São Paulo). DISTRIBUTION: Brazil. sallaei Sharp, 1887: 725 (Thoracophorus; Type locality: Mexico, Cordova; Guatemala, El Tumbador, Cerro Zunil, Capetillo, Tamahu, Balheu, Cubilguitz). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Irmler, 1985: 46 (Thoracophorus; characters; Mexico; Panama; Costa Rica; Peru). DISTRIBUTION: Mexico, Guatemala, Costa Rica, Panama, Peru. Bulletin AMNH OSORIINAE: THORACOPHORUS 1305 sculptilis Erichson, 1840: 910 (Glyptoma; Type locality: vallis Arguensis Columbiae). — Fauvel, 1864b: 62 [= 1865: 66] (Glyptoma; characters; Colombia). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Irmler, 1985: 56 (Thoracophorus; characters; Mexico; Panama; St. Vincent; Grenada; Guadeloupe; St. Thomas; Colombia). DISTRIBUTION: Colombia, Panama, Mexico, West Indies. ruficollis Fauvel, 1864b: 61 [= 1865: 65] (Glyptoma; Type locality: Mexique: Teapa). — Sharp, 1887: 727 (Thoracophorus; Mexico; Guatemala). — Fleutiaux and Sallé, 1889: 382 (Thoracophorus; Guadeloupe Island). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Irmler, 1985: 56 (Thoracophorus; synonym or sculptilis). sculptus Blackburn, 1902: 26 (Glyptoma; Type locality: Victoria). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Newton, 1990: 90, 93 (Thoracophorus). DISTRIBUTION: Australia. simplex Wendeler, 1930: 181 (Thoracophorus; Type locality: Trois-Rivieres, Guadelupe). — Scheerpeltz, 1933: 1021 (Thoracophorus; catalog). — Blackwelder, 1943: 149 (Thoracophorus; characters; Puerto Rico; Montserrat; Guadeloupe; Dominca; St. Lucia; Grenada). — Legner and Moore, 1977: 174 (Thoracophorus; found under bark). — Irmler, 1985: 53 (Thoracophorus; characters; Guadeloupe). DISTRIBUTION: West Indies. sordidus Blackburn, 1902: 26 (Glyptoma; Type locality: N. Queensland). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Bernhauer, 1920: 4 (Thoracophorus; Australia). — Newton, 1990: 90, 93 (Thoracophorus). DISTRIBUTION: Australia. subnitidus Motschulsky, 1860, see: Tetrapleurus. truquii Sharp, 1887: 726 (Thoracophorus; Type locality: Mexico). — Bernhauer and Schubert, 1910: 26 (Thoracophorus; subgenus Thoracophorus; catalog). — Irmler, 1985: 49 (Thoracophorus; characters; Mexico). DISTRIBUTION: Mexico.
OSORIINAE: TRIBE INCERTAE SEDIS
Sinolispinodes† [1 species; Palaearctic Region] Sinolispinodes† Zhang, 1989: 108 (species included: torosus). TYPE SPECIES: Sinolispinodes torosus Zhang, fixed by original designation and monotypy. torosus† Zhang, 1989: 108 (Sinolispinodes; Miocene fossil; Type locality: China: Shanwang, Shandong). DISTRIBUTION: China.
OXYTELINAE [48 genera; 2008 species; All Regions except the Antarctic] Oxytelinae Fleming, 1821. — Eichelbaum, 1909: 108 (list of genera with some references; generic synonyms; number of genera; number of species in each genus; general distribution of genera). — Reitter, 1909: 163 (characters; including subtribes: Oxytelini, Trogophloeini, Coprophilini, Acrognathini, Deleasterini). — Bernhauer and Schubert, 1910: 3 (world catalog; Piestini, Micropeplini, Pseudopsini, Phloeocharini, Proteinini, Neophonini, Omaliini, Dimerini). — Blatchley, 1910: 461 (characters; included Oxytelini, Osoriini, Omaliini, Proteini; key to Indiana genera). — Cameron, 1921a (characters; key to Singapore species and genera). — Porta, 1926: 5 (characters; used in broad sense; key to genera and species of Italy). — Portevin, 1929: 408 (characters; key to French species). — Böving and Craighead, 1930: 28 (larval characters). — Cameron, 1930: 23 (characters; [sensu lato]; key to tribes). — Joy, 1932: 137 (key to British genera). — Scheerpeltz, 1933: 992 (world catalog supplement). — Bernet Kempers, 1941: 34, 36 (larval characters). — Blackwelder, 1943: 57 (key to West Indian genera). — Tottenham, 1954: 37 (characters; key to British genera and species). — Crowson, 1955: 34 (adult and larval characters in key). — Hatch, 1957: 47, 84 (characters; key to genera of Pacific Northwest). — Palm, 1961a: 4 (characters; key to genera and species of Sweden). — Arnett, 1963: 237 (key to North American genera). — Arnett, 1963: 254 (characters; list of North American genera). — Steel, 1963: 53 (list of species of Malham Tarn; collecting notes; Britain). — Székessy, 1963: 6 (key to genera of Hungary). — Lohse, 1964: 70 (characters; key to central European genera and species). — Moore, 1964a: 87 (characters; notes). — Seevers, 1965: 339 (myrmecophilus species; notes on classification; list of genera of Anisopsis complex). — Kasule, 1966: 265, 268 (larval characters; notes). — Pototskaia, 1967: 17 (larval characters). — Coiffait and Saiz, 1968: 414 (key to Chilean genera). — Kasule, 1968: 118 (larval characters; larval key to some genera). — Scheerpeltz, 1968a: 20 (catalog of Austrian species). — Steel and Woodroffe, 1969: 135 (list of species collected from Isle of Rhum; collecting notes). — Herman, 1970 (characters; revision of generic classification; key to and redescription of genera; list of species included in each genus; anatomical notes; phylogeny). — Britton, 1970: 546 (characters of adults; Australia; notes). — Moore and Legner, 1974g: 553 (key to genera of America north of Mexico). — Moore and Legner, 1974h: 474 (catalog of the taxonomy, biology, and ecology of develop- mental stages of species of America north of Mexico). — Moore and Legner, 1975: 222 (catalog; North America north of Mexico). — Pope, 1977: 24 (list of British species). — Topp, 1978: 306, 309 (larval characters; key to genera). — Burakowski, Mroczkowski, and Stefan´ska, 1979: 80 (catalog; Poland). — Moore and Legner, 1979: 213 (characters; tabular key to genera of North America north of Mexico). — Muona, 1979: 19 (list of Scandinavian species). — Spahr, 1981: 93 (references to specimens from amber and copal). — Tóth, 1981: 95 (list of species of Hortobágy National Park, Hungary). — Watt and McColl, 1982: 18, 19 (characters; notes on New Zealand taxa).
1307 1308 OXYTELINAE Bulletin AMNH
— Newton, 1982a: 14, 16 (larval characters; phylogenetic analysis). — Pajni and Kohli, 1982: 9 (comparative morphology of aedeagus). — Dettner and Schwinger, 1982 (defensive secretions; chemistry; discussion of function). — E. Matthews, 1982: 8, 28 (notes; key to genera of Australia). — Outerelo and Gamarra, 1985: 93 (key to genera of Iberian Peninsula). — Newton, 1985: 203, 204 (notes on austral distribution). — Dettner and Grümmer, 1986 (defensive secretions; quasisynergism; repellency; penetration; evolution). — Dettner, 1987: 30, 31, 32 (defensive glands). — Lohse and Lucht, 1989: 129 (notes). — Frank, 1991: 345 (larval characters). — Dettner, 1991a (effectiveness of quinone-defensive systems). — Frania, 1992 (studies on ecdysis and metamorphosis). — Newton and Thayer, 1992: 13, 59 (summary of current classification; discussion). — Welch, 1993 (ovariole structure and development). — Dettner, 1993a (summary and discussion of defensive secretions and exocrine glands and their bearing on phylogeny). — Lawrence and Newton, 1995: 828 (discussion). — Terlutter, 1995: 5 (list of species collected in Westphalia). — Sparacio, 1995: 144 (characters; species of Sicily). — Klimaszewski, Newton, and Thayer, 1996: 150 (review of subfamily for New Zealand; notes; list of New Zealand genera; number of New Zealand species). — M. Hansen, 1996: 36, 98 (listed in Oxyteline group of subfamilies; list of Danish species). — Downie and Arnett, 1996: 440 (characters; key to species of Northeastern North America). — Ádám, 1996: 239 (list of species collected in Bükk National Park, Hungary). — Czarniawski and Staniec, 1997 (structure of opening of abdominal defensive reservoirs). — M. Hansen, 1997: 125, 177, 179 (autapomorphic features; key adult and larval characters; notes). — Anderson, Nash, and O’Connor, 1997: 16 (annotated checklist; Ireland). — Schülke, 1998i: 125 (supplement to key to genera in Lohse, 1964). — Klimaszewski, 2000: 21, 56 (characters; notes on classification, natural history, distribution, and key to tribes in Canada and Alaska). — Newton, Thayer, Ashe, and Chandler, 2000: 321, 378 (key to Nearctic tribes and genera; characters; notes). Oxytelidae Fleming, 1821: 49 (genera included: Oxytelus, Siagonium, Omalium, Piestus, cited as Piestes, Proteinus, Lesteva). TYPE GENUS: Oxytelus (see Newton and Thayer, 1992: 59). — Gistel, 1856: 389 (list of taxa; cited as Oxeotelidae). — Jeannel and Jarrige, 1949: 311 (characters). — Jarrige, 1952a: 136 (characters). — Coiffait and Saiz, 1968: 401 (Chilean species). — Outerelo and Gamarra, 1985: 74 (key to genera of Iberian Peninsula). Oxytelides Mannerheim, 1830: 9 (genera included: Bledius, Platystethus, Oxytelus, Trogophloeus). TYPE GENUS: Oxytelus (see Newton and Thayer, 1992: 59).
Tribe Coprophilini Coprophilini Heer, 1839. — Reitter, 1909: 164 (Coprophilus, Siagonium). — Bernhauer and Schubert, 1911: 87 (world catalog; Syntomium, Coprostygnus, Deleaster, Zalobius, Asemobius, Coprophilus, Sartallus, Oncophorus, Acrognathus). — Scheerpeltz, 1933: 1074 (world catalog supplement). — Crowson, 1955: 37 (genera). — Hatch, 1957: 85 (characters). — Moore, 1964: 57 (key to Nearctic genera). — Herman, 1970: 362 (characters; type genus). Bulletin AMNH OXYTELINAE: COPROPHILINI 1309
— Muona, 1979: 19 (list of Scandinavian species). — Outerelo and Gamarra, 1985: 101 (key to genera of Iberian Peninsula). — Newton and Thayer, 1992: 59 (tribe of Oxytelinae). — Klimaszewski, Newton, and Thayer, 1996: 150 (list of New Zealand genera; number of New Zealand species). — M. Hansen, 1996: 36, 98 (may not be monophyletic; not clearly defined; list of Danish species). — Newton, Thayer, Ashe, and Chandler, 2000: 379 (characters; notes). Coprophilina Heer, 1839: 198 (genera included: Deleaster, Coprophilus, Syntomium). TYPE GENUS: Coprophilus (see Newton and Thayer, 1992: 59). — Arnett, 1963: 255 (characters; list of North American genera). — Newton and Thayer, 1992: 59 (synonym of Coprophilini). Coprophilini Erichson, 1839b: 30 (genera included: Acrognathus, Coprophilus, Deleaster, Micralymma, Syntomium). TYPE GENUS: Coprophilus (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Coprophilini). Coprophilida Heer, 1839a: 13 (genera included: Deleaster, Coprophilus, Syntomium). TYPE GENUS: Coprophilus (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Coprophilini). Homalotriquitos Solier, 1849: 321 (genera included: Homalotrichus). TYPE GENUS: Homalotrichus (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Coprophilini). Homalotrichites: — Lacordaire, 1854: 153 (genera included: Homalotrichus). TYPE GENUS: Homalotrichus (see Newton and Thayer, 1992: 60, name not latinized so may be unavailable). — Newton and Thayer, 1992: 60 (synonym of Coprophilini). Toxoderi Bernhauer and Schubert, 1911: 91 (genera included: Toxoderus, Crymus). TYPE GENUS: Toxoderus (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Coprophilini; Toxoderi is preoccupied).
Coprophilus [30 species; Nearctic and Palaearctic Regions] Coprophilus Latreille, 1829: 439 (species included: rugosum). TYPE SPECIES: Staphylinus striatulus Fabricius, fixed by action of the Commission in Opinion 1722 (ICZN, 1993a). [Note: The following is of historical interest only. Latreille, 1829: 439 included one species “Omalium rugosum, Gravenhorst” in Coprophilus. The rugosum included in Omalium by Gravenhorst (1802: 115) was attributed to Olivier who cited Fabricius as an earlier use of the name. The author, therefore, of rugosus is Fabricius not Olivier as cited by Blackwelder, 1952: 106 (Bernhauer and Schubert, 1911: 90 cited rugosus Olivier in Coprophilus as a synonym of striatulus and, on page 119, listed rugosus Fabricius in Oxytelus). The name, rugosus Fabricius, is now in Anoty- lus which means that, had there been no ruling by the Commission (Opinion 1722) the name Coprophilus would have had to be moved to Anotylus where it would have been the senior name. The species in Coprophilus would have had to take the next available name, Zonyptilus]. — Latreille, 1832: 89 (characters; notes). — Griffith and Pidgeon, 1832: 296 (characters; [Note: Described as “. . . the body is . . . flat- tened, . . .” so is probably not Coprophilus). — Stephens, 1834: 345 (characters; [Note: Stephens described this genus as depressed; the species in Coprophilus are cylindrical or subcylindrical]). — Westwood, 1838a: 18 (characters; type species). — Erichson, 1839a: 609 (characters). — Heer, 1839: 198 (characters). 1310 OXYTELINAE: COPROPHILUS Bulletin AMNH
— Stephens, 1839: 422 (characters). — Erichson, 1840: 815 (characters). — Laporte, 1840: 189 (characters). — Redtenbacher, 1849: 742 (characters). — Schaum, 1852: 29 (list of species; Europe). — Lacordaire, 1854: 120 (characters; notes; list of species). — Fairmaire and Laboulbène, 1856: 620 (characters). — Jacquelin du Val, 1857: 59 (characters). — Redtenbacher, 1857: 237 (characters; Austrian species). — Kraatz, 1857d: 893 (characters). — Thomson, 1858: 39 (characters). — G. Waterhouse, 1858: 30 (catalog of British species). — Thomson, 1859: 45 (characters; type species: striatulus). — Schaum, 1859: 31 (catalog; European species). — Thomson, 1861: 143 (characters). — Wencker and Silbermann, 1866: 35 (list of species; collecting notes; France). — Gemminger and Harold, 1868: 656 (catalog). — Jacquelin du Val, 1868: 78 (catalog; European species). — Sharp, 1871d: 14 (list of British species). — Fauvel, 1871a: 155 [= 1872: 129] (characters). — Redtenbacher, 1874: 261 (characters). — Fauvel, 1875a:VIII [= 1875b: 210] (catalog). — Mulsant and Rey, 1878: 682 (characters). — Heyden, 1880: 81 (list of species of Siberian region). — Fowler, 1888: 394 (notes). — Heyden, Reitter, and Weise, 1891: 117 (list of species of Europe and Caucasus). — Heyden, 1893: 48 (list of species of Siberian region). — Ganglbauer, 1895: 676 (characters; key to middle European species). — Fauvel, 1897a: 226 (notes; key to Palaearctic species). — Heyden, Reitter, and Weise, 1906: 138 (list of species of Europe and Caucasus). — Bernhauer, 1908e: 327 (discussion; key to species; catalog). — Reitter, 1909: 176 (characters). — Bernhauer and Schubert, 1911: 89 (world catalog; 14 species). — Petri, 1912: 56 (list of species of Siebenbürgen). — Johansen, 1914: 570 (characters). — Winkler, 1925: 338 (catalog for Palaearctic region). — Cameron, 1925: 12 (catalog of Indian species). — Porta, 1926: 35 (characters; species of Italy). — Bruch, 1928a: 423 (catalog; Argentina). — Portevin, 1929: 426 (key to French species). — Cameron, 1930: 171 (characters; key to species of British India). — Roubal, 1930: 321 (catalog; Slovakia). — Joy, 1932: 139 (key to British species). — Scheerpeltz, 1933: 1075 (world catalog supplement). — Kloet and Hincks, 1945: 162 (list of British species). — Tottenham, 1949: 359 (synonym of Elonium; type species: striatulus). — Blackwelder, 1952: 106 (type species: rugosus). — Tottenham, 1954: 38 (cited as Elonium; characters; British species). — Hatch, 1957: 85, 89 (characters; key to species of Pacific Northwest). — Watanabe and Shibata, 1961a (cited as Elonium; revision and key to species of Japan). — Palm, 1961a: 8 (characters; species of Sweden). — Horion, 1963: 161 (list of central European species). — Székessy, 1963: 27 (characters). — Lohse, 1964: 72 (characters; key to central European species). — Moore, 1964: 57 (cited as Elonium; characters). — Pototskaia, 1967: 24, 25 (cited as Elonium; larval characters; key to some genera). Bulletin AMNH OXYTELINAE: COPROPHILUS 1311
— Kasule, 1968: 120 (larval characters). — Scheerpeltz, 1968a: 20 (catalog of Austrian species). — Herman, 1970: 365 (characters; distribution; notes; discussion of nomenclature; list of species). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Moore and Legner, 1975: 242 (catalog; North America north of Mexico). — Scheerpeltz, 1976: 16 (key to species of Himalayan region). — Shibata, 1976: 131 (checklist of species of Japan). — Pope, 1977: 24 (list of British species). — Topp, 1978: 309 (larval characters). — Burakowski, Mroczkowski, and Stefan´ska, 1979: 81 (catalog; cited as Elonium; Poland). — Moore and Legner, 1979: 221 (characters; notes). — Muona, 1979: 19 (list of Scandinavian species; cited as Elonium with Coprophilus as synonym). — Uhlig and Vogler, 1981: 88 (list of some species collected in Germany; habitat and collecting notes). — Ádám, 1985: 250 (cited as Elonium; list of species from southeast Hungary). — Tóth, 1988: 197 (characters; key to subgenera). — Dettner, 1987: 31 (defensive glands). — Tóth, 1991: 96 (key to some species). — Campbell and Davies, 1991: 93 (checklist of species; Canada). — Silfverberg, 1992: 23 (cited as Elonium; list of species of Fennoscandia, Denmark, and the Baltic States). — Thayer, 1992a: 25 (requested to set asida all previous fixations of type species and to designate Staphylinus striatulus Fabricius as type species and to place name on Official List of Generic Names in Zoology). — Tóth, 1993: 376 (key to species with 5 segmented antennal club). — ICZN, 1993a (type species designation). — Bohácˇ, 1993: 42 (cited as Elonium; in list of species of Czech Republic and Slovakia). — Ciceroni and Zanetti, 1995: 10 (list of species of Italy). — M. Hansen, 1996: 98 (list of Danish species). — Downie and Arnett, 1996: 457 (characters; species of Northeastern North America). — Schülke, 1998i: 125 (characters). — Newton, Thayer, Ashe, and Chandler, 2000: 379 (3 Nearctic species; notes; characters in key). Zonyptilus Motschulsky, 1845: 39 (species included: pennifer; [Note: Agassiz (1846: 392) emended this name to Zonoptilus; the change is an unjustified emendation. Zonoptilus was used by Motschulsky in 1857 and again on a plate published in 1858 in the Bul- letin de la Société Impériale des Naturalistes de Moscou. Blackwelder, 1952: 409 stated that Zonyptilus was nomen nudum and that it (as Zonoptilus) was made avail- able by Solsky, 1867: 85. Motschulsky (1845), in a combined characterization, stated that Zonyptilus had the facies of Bledius, the elytral striae of Platystethus and stated the color and approximate size so Article 12.2.6 would apply. Blackwelder also claimed that Motschulsky’s (1857a: 502) second citation (emended to Zonoptilus) did not validate the species. However, Motschulsky in 1857 presented an even more com- plete characterization of the genus and species. Zonyptilus was made available in 1845 but, if not at that time, then certainly in 1857]). TYPE SPECIES: Zonyptilus pennifer Motschulsky, fixed by monotypy. — Motschulsky, 1857a: 502 (cited as Zonoptilus; characters). — Hochhuth, 1862: 98 (notes). — Solsky, 1867: 85 (characters; notes; [Note: Blackwelder, 1952: 409 cited this reference as the first to validate Zonoptilus; see note attached to the generic name]). — Gemminger and Harold, 1868: 646 (catalog). — Fauvel, 1869: 492 (synonym of Coprophilus). — Fauvel, 1871a: 155 [= 1872: 129] (synonym of Coprophilus). — Redtenbacher, 1874: 261 (characters). — Heyden, 1896: 36 (list of species of Siberian region). — Bernhauer and Schubert, 1911: 89 (subgenus of Coprophilus). 1312 OXYTELINAE: COPROPHILUS Bulletin AMNH
— Scheerpeltz, 1933: 1075 (subgenus of Coprophilus). — Blackwelder, 1952: 409 (synonym of Elonium; type species: pennifer). — Horion, 1963: 161 (subgenus of Coprophilus; list of central European species). — Herman, 1970: 366 (synonym of Coprophilus). — Tóth, 1988: 197 (subgenus of Coprophilus; characters; key to species groups). — Smetana, 1998: 157 (comments on spelling and validity of name). Zonoptilus Agassiz, 1847: 392 (emendation of Zonyptilus [Note: Unjustified emendation]). TYPE SPECIES: Zonyptilus pennifer Motschulsky, fixed by objective synonymy with Zonyptilus. adachii Watanabe and Shibata, 1961a: 43 (Elonium; Type locality: Japan: Aomori Pref.: Mt. Iwaki). — Herman, 1970: 367 (Coprophilus). DISTRIBUTION: Japan. alticola Fauvel, 1904b: 93 (Coprophilus; Type locality: Himalaya: Simla). — Bernhauer and Schubert, 1911: 89 (Coprophilus; catalog). — Cameron, 1930: 172 (Coprophilus; characters; India). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Herman, 1970: 367 (Coprophilus). — Abdullah and Qadri, 1970: 120 (Coprophilus; characters; West Pakistan). — Scheerpeltz, 1976: 16 (Coprophilus; characters). DISTRIBUTION: Pakistan, India. araxis Reitter, 1894, see: pennifer Motschulsky, 1845. bimaculatus Luze, 1904: 79 (Coprophilus; Type locality: Trkst., Jagnob, Margef, Fl. Dschid- schigrut, See Kulikulan; [Note: See lectotype designation by Tóth, 1991]. Also cited from Iskan-der-kul). — Bernhauer, 1908e: 330 (Coprophilus; synonym of longicornis). — Bernhauer and Schubert, 1911: 89 (Coprophilus; synonym of longicornis). — Tóth, 1991: 91 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; lectotype designation: Trkst., Jagnob, Margef, Fl. Dschidschigrut, See Kulikulan; valid species). — Tóth, 1993: 376 (Coprophilus; characters). DISTRIBUTION: Tajikistan, Kazakhstan. burphuensis Champion, 1925: 106 (Coprophilus; Type locality: Burphu, Gori Valley, E. Kumaon, alt. 11,500 ft.). — Cameron, 1930: 172 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; India). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Herman, 1970: 367 (Coprophilus). — Abdullah and Qadri, 1970: 120 (Coprophilus; characters; West Pakistan). — Scheerpeltz, 1976: 17 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters). — Coiffait, 1984: 118 (Coprophilus; Pakistan). DISTRIBUTION: Pakistan, India. castoris Campbell, 1979a: 223 (Coprophilus; Type locality: Quebec: Parc Gatineau, Lac Fortune. Also cited from New Brunswick and New York). — Downie and Arnett, 1996: 457 (Coprophilus; characters; Canada; USA). DISTRIBUTION: Canada, USA. chinensis Zheng, 1984: 462 (Coprophilus; subgenus Coprophilus; Type locality: Emei, Sichuan, China). — J. Li, 1993: 22 (Coprophilus; China). DISTRIBUTION: China. difformis Smetana, 1998: 154 (Coprophilus; subgenus Zonyptilus; Type locality: Taiwan: Taichung Hsien, Hsuehshan, Hseuhshan Main Peak, 3650 m). DISTRIBUTION: Taiwan. dimidiatipennis Fauvel, 1895, see: pennifer Motschulsky, 1845. Bulletin AMNH OXYTELINAE: COPROPHILUS 1313 drugmandi Tóth, 1991: 95 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Type locality: Cernavoda, Dobrudja). — Tóth, 1993: 375 (Coprophilus; characters). DISTRIBUTION: Romania. eppelsheimi Fauvel, 1897, see: pennifer Motschulsky, 1845. fauveli Tóth, 1992: 373 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Type locality: “Bakou” [Azerbaidzhan]). DISTRIBUTION: Azerbaijan. formosanus Shibata, 1993: 313 (Coprophilus; Type locality: Sungchuankang, about 2,400 m alt., Nantou Hsien, Taiwan). DISTRIBUTION: Taiwan. fulvomaculatus Adachi, 1935, see: impressus Sharp, 1889. impressus Sharp, 1889: 415 (Coprophilus; Type locality: Fukushima; Sapporo). — Bernhauer and Schubert, 1911: 89 (Coprophilus; catalog). — Nakane, 1963a: 83 (Elonium; characters; habitus photograph; Japan). — Watanabe and Shibata, 1961a: 43, 45 (Elonium; characters; notes; Japan). — Herman, 1970: 367 (Coprophilus). DISTRIBUTION: Japan. fulvomaculatus Adachi, 1935: 125 (Coprophilus; Type locality: Shinnosawa Valley of Chichibu, Musashi Province, Japan). — Watanabe and Shibata, 1961a: 45 (Elonium; synonym of impressus). kashmiricus Cameron, 1941: 145 (Coprophilus; Type locality: Kashmir: Gulmarg). — Herman, 1970: 368 (Coprophilus). DISTRIBUTION: India. laesus Germar, 1832, see: striatulus Fabricius, 1793. lateralis Fauvel, 1875, see: pennifer Motschulsky, 1845. longicollis Cameron, 1941: 145 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Type locality: Darjeeling district: Ghum). — Herman, 1970: 368 (Coprophilus). DISTRIBUTION: India. longicornis Bernhauer, 1903b: 188 (Coprophilus; Type locality: Turkestan). — Bernhauer, 1908e: 330 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; catalog). — Bernhauer and Schubert, 1911: 89 (Coprophilus; catalog). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Herman, 1970: 368 (Coprophilus). — Tóth, 1991: 92 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; Turkestan). — Tóth, 1993: 376 (Coprophilus; characters). — Kashcheev, 1999: 150 (Coprophilus; characters). DISTRIBUTION: “Turkestan”. major Coiffait, 1982: 157 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Type locality: Inde: Cachemire, Tangmarg, Pir Panjal Gebirge, 2600 m). DISTRIBUTION: India. marginalis Reitter, 1894, see: pennifer Motschulsky, 1845. nepalensis Scheerpeltz, 1976: 14 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Type locality: Nepal: Bassislager Yaral bei Pangpoche, 3900 m). DISTRIBUTION: Nepal. nepalicus Coiffait, 1984: 146 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Type locality: Nepal: Bumra Chhurchi Lagna, 3350 m). DISTRIBUTION: Nepal. obscurus Reitter, 1894, see: pennifer Motschulsky, 1845. 1314 OXYTELINAE: COPROPHILUS Bulletin AMNH obscurus Luze, 1904 [Note: Nomen nudum], see: pentatoma Fauvel, 1897. obsoletus Reitter, 1894, see: pennifer Motschulsky, 1845. parvipennis Scheerpeltz, 1972, see: Homalotrichus. pennifer Motschulsky, 1845: 39 (Zonyptilus; [Note: Bernhauer and Schubert, 1911: 39 cited Motschulsky, 1845 as the original decription of pennifer but Blackwelder, 1952: 409 stated that Motschulsky’s name was unavailable and that it was first validated by Sol- sky, 1867: 86; Blackwelder’s decision was incorrect; Motschulsky stated the size and color of pennifer and mentioned other features]; Type locality: Tiflis, Georgia; [Note: See lectotype designation by Tóth, 1988]). — Motschulsky, 1857a: 503 (Zonyptilus, cited as Zonoptilus; characters; Georgie: Tiflis). — Hochhuth, 1862: 99 (Coprophilus; characters; Caucausus). — Solsky, 1867: 86 (Zonyptilus, cited as Zonoptilus; characters; Crimea). — Redtenbacher, 1874: 261 (Zonyptilus, cited as Zonoptilus; characters). — Solsky, 1874: 206 (Zonyptilus, cited as Zonoptilus; characters; Turkestan). — Reitter, 1894: 178 (Zonyptilus, cited as Zonoptilus; characters; Russia; Caucasus). — Reitter, 1894a: 292 (Zonyptilus, cited as Zonoptilus; characters; Caucasus; Armenia; Bagdad; Amur). — Reitter, 1894: 177 (Zonyptilus, cited as Zonoptilus; [Note: Misidentification: Bernhauer and Schubert, 1911: 89 cited lateralis Reitter as a synonym of pennifer; Reitter did not describe the species as new, he attributed it to Fauvel]; characters; Mongolia). — Fauvel, 1897a: 227, 228 (Coprophilus; characters; Hungary; Austria; Astrakan; Turkestan; Georgia; Armenia; Mesopotamie; Amur region). — Bernhauer, 1908e: 329, 330 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; catalog). — Bernhauer and Schubert, 1911: 89 (Coprophilus; catalog). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Scheerpeltz, 1961h: 249 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Afghanistan). — Scheerpeltz, 1963: 5 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Afghanistan). — Lohse, 1964: 72 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; central characters). — Pototskaia, 1965: 877 (Coprophilus; larval description). — Pototskaia, 1967: 25 (Elonium; larval characters). — Herman, 1970: 368 (Coprophilus). — Lucht, 1987: 91 (Coprophilus). — Tóth, 1988: 201 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; lectotype designation: Tiflis, Georgia; characters). — Bohácˇ, 1993: 42 (Elonium; subgenus Zonyptilus; Slovakia; Czech Republic). DISTRIBUTION: Ukraine, Russia, Georgia, Armenia, Azerbaijan, Iraq, Iran, Afghanistan, Turkmenistan, Uzbekistan, Mongolia, China. sellula Kraatz, 1857d: 894, note (Coprophilus; Type locality: Mesopotamia). — Solsky, 1867: 87 (Zonyptilus, cited as Zonoptilus; characters; Mesopotamia). — Eppelsheim, 1892: 346 (Zonyptilus, cited as Zonoptilus; Turkestan). — Reitter, 1894: 178 (Zonyptilus, cited as Zonoptilus; variety of pennifer; characters; Mesopotamia; Transcaspian region). — Reitter, 1894a: 292 (Zonyptilus, cited as Zonoptilus; characters; Mesopotamia; Transcaspian region; Mongolia; east Siberia). — Bernhauer, 1908e: 330 (Coprophilus; synonym of pennifer). — Bernhauer and Schubert, 1911: 89 (Coprophilus; variety of pennifer). — Scheerpeltz, 1933: 1076 (Coprophilus; variety of pennifer). lateralis Fauvel, 1875a:VIII [= 1875b: 210] (Coprophilus; proposed for pennifer as used by Solsky, 1867: 86; Type locality: Crimée; Boukharie, Sibérie orientale). — Fauvel, 1897a: 227, 228 (Coprophilus; characters; Crimea; Margelan; Samarkand; Persia). — Bernhauer, 1908e: 330 (Coprophilus; variety of pennifer). — Bernhauer and Schubert, 1911: 89 (Coprophilus; variety of pennifer). — Scheerpeltz, 1933: 1076 (Coprophilus; variety of pennifer). — Scheerpeltz, 1960b: 67 (Coprophilus; variety of pennifer; Afghanistan). Bulletin AMNH OXYTELINAE: COPROPHILUS 1315
striatipennis Eppelsheim, 1894: 291 (Zonyptilus, cited as Zonoptilus; Type locality: Altai Gebirge). — Fauvel, 1897a: 229 (Coprophilus; type locality is Monts Alai not Monts Altai). — Bernhauer, 1908e: 330 (Coprophilus; synonym of pennifer). — Bernhauer and Schubert, 1911: 89 (Coprophilus; synonym of pennifer). — Scheerpeltz, 1933: 1076 (Coprophilus; monstrosity of pennifer). obscurus Reitter, 1894: 177 (Zonyptilus, cited as Zonoptilus; variety of pennifer; Type locality: Araxes). — Reitter, 1894a: 292 (Zonyptilus, cited as Zonoptilus; characters; Araxes). — Bernhauer, 1908e: 330 (Coprophilus; variety of pennifer). — Bernhauer and Schubert, 1911: 89 (Coprophilus; variety of pennifer). — Scheerpeltz, 1933: 1076 (Coprophilus; variety of pennifer). araxis Reitter, 1894: 178 (Zonyptilus, cited as Zonoptilus; variety of pennifer; Type locality: Araxes). — Reitter, 1894a: 292 (Zonyptilus, cited as Zonoptilus; characters; Arax valley). — Bernhauer, 1908e: 330 (Coprophilus; synonym of pennifer). — Bernhauer and Schubert, 1911: 89 (Coprophilus; variety of pennifer). — Scheerpeltz, 1933: 1076 (Coprophilus; variety of pennifer). obsoletus Reitter, 1894a: 292 (Zonyptilus, cited as Zonoptilus; variety of sellula; Type locality: Araxesthal). — Bernhauer, 1908e: 330 (Coprophilus; synonym of pennifer). — Bernhauer and Schubert, 1911: 89 (Coprophilus; synonym of pennifer). marginalis Reitter, 1894a: 292 (Zonyptilus, cited as Zonoptilus; variety of sellula; Type locality: Nördliche Mongolei; Ostsibirien). — Bernhauer, 1908e: 330 (Coprophilus; variety of pennifer). — Bernhauer and Schubert, 1911: 89 (Coprophilus; variety of pennifer). — Scheerpeltz, 1933: 1076 (Coprophilus; variety of pennifer). eppelsheimi Fauvel, 1897a: 227, 228 (Coprophilus; Type locality: Mongolie, Amdo; Sibérie or., Vladivostok; Chine, Pekin, Tien-Tsin, Shangai). — Bernhauer, 1908e: 330 (Coprophilus; synonym of pennifer). — Bernhauer and Schubert, 1911: 89 (Coprophilus; synonym of pennifer). dimidiatipennis Fauvel, 1900d: 223 (Coprophilus; Type locality: Transcaspienne: Aschabad). — Bernhauer, 1908e: 330 (Coprophilus; variety of pennifer). — Bernhauer and Schubert, 1911: 89 (Coprophilus; variety of pennifer). — Scheerpeltz, 1933: 1076 (Coprophilus; variety of pennifer). pentatoma Fauvel, 1897a: 227, 228 (Coprophilus; Type locality: Mons Alai Turkestan; [Note: See lectotype designation by Tóth, 1991]. Also cited from Baku; Dobrudja). — Bernhauer and Schubert, 1911: 89 (Coprophilus; catalog). — Bernhauer, 1908e: 329, 330 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; catalog). — Scheerpeltz, 1933: 1076 (Coprophilus; synonym of longicornis). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Gridelli, 1934: 72 (Coprophilus; Karakoram region). — Herman, 1970: 368 (Coprophilus). — Tóth, 1991: 94 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; lectotype designation: Mons Alai Turkestan; Turkestan). — Tóth, 1993: 375 (Coprophilus; characters). — Kashcheev, 1999: 149 (Coprophilus; characters). DISTRIBUTION: Azerbaijan, Kyrgyzstan. obscurus Luze, 1904: 79 (Coprophilus; aberration of bimaculatus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]). — Bernhauer, 1908e: 330 (Coprophilus; synonym of pentatoma). — Bernhauer and Schubert, 1911: 89 (Coprophilus; synonym of pentatoma). 1316 OXYTELINAE: COPROPHILUS Bulletin AMNH piceus Solsky, 1867: 88 (Zonyptilus, cited as Zonoptilus; Type locality: Sarepta). — Reitter, 1894: 177 (Zonyptilus, cited as Zonoptilus; characters; Russia; Austria; Hungary). — Reitter, 1894a: 292 (Zonyptilus, cited as Zonoptilus; characters; Russia; Austria; Hungary). — Ganglbauer, 1895: 677 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; Austria; Hungary; southern Russia). — Bernhauer, 1908e: 329, 330 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; catalog; Austria; Germany; Hungary; southern Russia). — Bernhauer and Schubert, 1911: 89 (Coprophilus; catalog). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Horion, 1963: 161 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Russia; Bosnia Hercegovina; Hungary; Slovakia; Czech Republic; Austria; Germany). — Lohse, 1964: 72 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; central Europe). — Smetana, 1964c: 164 (Coprophilus; Czechoslovakia). — Herman, 1970: 368 (Coprophilus). — Ádám, 1987: 138 (Zonyptilus; collecting notes; Hungary). — Lucht, 1987: 91 (Coprophilus; checklist; Germany; Czechoslovakia; Austria). — Tóth, 1988: 199, 202 (Coprophilus; characters; Carpathian Basin). — Bohácˇ, 1993: 42 (Elonium; subgenus Zonyptilus; Slovakia). — Assing, Frisch, Kahlen, et al., 1998: 127 (Coprophilus; Germany). DISTRIBUTION: Europe, Russia. reitteri Luze, 1904a: 69 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Type locality: Aulie-Ata). — Bernhauer, 1908e: 330 (Coprophilus; synonym of schubertii). — Bernhauer and Schubert, 1911: 90 (Coprophilus; synonym of schubertii). — Tóth, 1988: 200 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; valid species; char- acters). DISTRIBUTION: Kazakhstan. rufipennis Reitter, 1894: 178 (Zonyptilus, cited as Zonoptilus; Type locality: Margelan). — Reitter, 1894a: 292 (Zonyptilus, cited as Zonoptilus; characters; Margelan; Alai; Chodshent; Persien). — Bernhauer, 1908e: 329, 330 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; catalog; Margelan; Turcmenistan). — Bernhauer and Schubert, 1911: 90 (Coprophilus; catalog). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Herman, 1970: 368 (Coprophilus). DISTRIBUTION: Uzbekistan, Turkmenistan, Iran. rufitarsis Kashcheev, 1999: 149 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Type locality: Kazakhstan, zapovednik Aksu-Dzhabagly). DISTRIBUTION: Kazakhstan. schubertii Motschulsky, 1860b: 120 (Zonyptilus, cited as Zonoptilus; Type locality: Ostro- gorst, G Voron [sko?, esh?]; [Note: See lectotype designation by Tóth, 1988]). — Fauvel, 1875a:VIII [= 1875b: 210] (Coprophilus; synonym of piceus). — Fauvel, 1897a: 226, 227, 228 (Coprophilus; characters; Russia). — Bernhauer, 1908e: 329, 330 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; catalog; Turkestan; southern Russia). — Bernhauer and Schubert, 1911: 90 (Coprophilus; catalog). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Herman, 1970: 368 (Coprophilus). — Tóth, 1988: 198 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; lectotype designation: Ostrogorst, G Voron [sko?, esh?]; characters; notes). DISTRIBUTION: Russia, “Turkestan”. sellula Kraatz, 1857, see: pennifer Motschulsky, 1845. sexualis Leech, 1939: 259 (Coprophilus; Type locality: Vancouver, B. C. [British Columbia]). — Hatch, 1957: 90 (Coprophilus; characters; British Columbia; Washington; Oregon). Bulletin AMNH OXYTELINAE: COPROPHILUS 1317
— Herman, 1970: 368 (Coprophilus). — Campbell, 1979a: 228 (Coprophilus; characters). DISTRIBUTION: Canada, USA. sibiricus Bernhauer, 1915j: 263 (Coprophilus; Type locality: Ost-Sibirien, Schipka-Gora). — Bernhauer, 1923d: 180 (Coprophilus; characters; Siberia). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Herman, 1970: 368 (Coprophilus). — J. Li, 1993: 22 (Coprophils; China). DISTRIBUTION: Russia, China. simplex Sharp, 1889: 415 (Coprophilus; Type locality: Wada-toge). — Bernhauer and Schubert, 1911: 90 (Coprophilus; catalog). — Watanabe and Shibata, 1961a: 43 (Elonium; characters; notes; Japan). — Herman, 1970: 368 (Coprophilus). DISTRIBUTION: Japan. solskyi Bernhauer, 1908e: 330 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; named for Reitter’s 1894 misidentification of lateralis; Type locality: Dobrutja). — Reitter, 1894a: 291 (Zonyptilus, cited as Zonoptilus; [Note: Misidentification: Bernhauer and Schubert, 1911: 90 cited lateralis Reitter as a synonym of solskyi; Reitter did not describe the species as new, he attributed it to Fauvel]; characters; Dobrutscha). — Bernhauer and Schubert, 1911: 90 (Coprophilus; catalog). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Herman, 1970: 368 (Coprophilus). — Tóth, 1991: 93 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters). — Tóth, 1993: 376 (Coprophilus; characters). — Kashcheev, 1999: 150 (Coprophilus; characters). DISTRIBUTION: Romania. striatipennis Eppelsheim, 1894, see: pennifer Motschulsky, 1845. striatulus Fabricius, 1793: 525 (Staphylinus; Type locality: Germania). — Olivier, 1795:(42): 30 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1911: 90 cited rugosus Olivier as a synonym of striatulus Fabricius; Olivier did not describe the species as new, he attributed it to Fabricius]; characters; England, France). — Panzer, 1795: 354 (Staphylinus; characters; Germany). — Fabricius, 1801: 596 (Staphylinus; characters; Germany). — Turton, 1802: 512 (Staphylinus; characters; Germany). — Curtis, 1829: 29 (Elonium; catalog; Britain). — Stephens, 1834: 345 (Coprophilus; characters; England). — Westwood, 1838a: 18 (Coprophilus; type species). — Erichson, 1839a: 609 (Coprophilus; characters; Germany). — Heer, 1839: 199 (Coprophilus; characters; Switzerland). — Stephens, 1839: 423 (Coprophilus; characters; England). — Erichson, 1840: 816 (Coprophilus; characters; Europe). — Redtenbacher, 1849: 742 (Coprophilus; characters; Austria). — Küster, 1854: 21 (Coprophilus; characters; Europe). — Hardy, 1851: 53 (Coprophilus; notes; England). — Fairmaire and Laboulbène, 1856: 620 (Coprophilus; characters; France). — Redtenbacher, 1857: 237 (Coprophilus; characters; Austria). — Kraatz, 1857d: 894 (Coprophilus; characters; Germany). — Thomson, 1859: 45 (type species of Coprophilus). — Thomson, 1861: 143 (Coprophilus; characters; Scandinavia). — Fauvel, 1871a: 156 [= 1872: 130] (Coprophilus; characters; France; Scandinavia; Britain; Germany; Austria; Italy). — Redtenbacher, 1874: 261 (Coprophilus; characters; Austria). — LeConte, 1875: 170 (Coprophilus; Canada). — Seidlitz, 1875: 245 (Coprophilus; characters; Baltic region). — Fauvel, 1878d: 257 [= 1878e: 93] (Coprophilus; characters; Canada). 1318 OXYTELINAE: COPROPHILUS Bulletin AMNH
— Mulsant and Rey, 1878: 685 (Coprophilus; characters; France). — Fowler, 1888: 394 (Coprophilus; characters; Britain). — Ragusa, 1892: 168 (Coprophilus; Sicily). — Seidlitz, 1891: 352 (Coprophilus; characters; Europe). — Seidlitz, 1891a: 376 (Coprophilus; characters; Europe). — Ganglbauer, 1895: 677 (Coprophilus; characters; Europe). — Fauvel, 1897a: 227, 228 (Coprophilus; characters; Italy; Moldavia; Canada). — Sainte-Claire Deville, 1907: 57 (Coprophilus; characters; France). — Bernhauer, 1908e: 329, 330 (Coprophilus; subgenus Coprophilus; characters; catalog). — Reitter, 1909: 176 (Coprophilus; characters; Germany). — Bernhauer and Schubert, 1911: 90 (Coprophilus; catalog). — Johansen, 1914: 570 (Coprophilus; characters; Denmark). — Eichelbaum, 1914: 331 (Coprophilus; abdominal segmentation; apical segements). — Portevin, 1929: 426 (Coprophilus; cited as striatuius; characters; France). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Leech, 1939: 261 (Coprophilus; characters in key). — Paulian, 1941: 157 (Coprophilus; larval characters). — Palm, 1961a: 8 (Coprophilus; characters; Sweden; Denmark; Norway; Finland). — Horion, 1963: 161 (Coprophilus; Denmark; Sweden; Norway; Britain; Holland; Belgium; France; Switzerland; Italy; Slovenia; Croatia; Bosnia Hercegovina; Romania; Hungary; Czechoslovakia; Poland; Russia; Germany; Austria). — Lohse, 1964: 72 (Coprophilus; subgenus Coprophilus; characters; central Europe). — Pototskaia, 1965: 879 (Coprophilus; larval characters). — Pototskaia, 1967: 26 (Elonium; larval characters). — Szujecki, 1968a: 713 (Coprophilus; Poland). — Herman, 1970: 368 (Coprophilus). — Osella and Zanetti, 1975: 88 (Coprophilus; collected from nest of Talpa europaea; notes; Italy). — Pope, 1977: 24 (Coprophilus; Britain). — Campbell, 1979a: 228 (Coprophilus; characters). — Dettner, 1984 (Coprophilus; chemistry of defensive secretions; structure of glands; discus- sions). — Segers, 1986: 23 (Coprophilus; checklist of species; Belgium). — Ádám, 1987: 138 (Coprophilus; collecting notes; Hungary). — Lucht, 1987: 91 (Coprophilus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzer- land; France; Benelux; Denmark; Sweden). — Tóth, 1988: 202 (Coprophilus; Carpathian Basin). — Nowosad, 1990: 53 (Elonium; collected from mole nest; notes; Poland). — Thayer, 1992a: 25 (requested to place name on Official list of Specific Names in Zoology). — ICZN, 1993a (fixed as type species of Coprophilus; name placed on Official List of Specific Names in Zoology). — Bohácˇ, 1993: 42 (Elonium; subgenus Elonium; Slovakia; Czech Republic). — Hoebeke, 1995 (Coprophilus; characters; habitat and collecting notes; presence confirmed in North America; Ontario; Quebec; New York). — Terlutter, 1995: 6 (Coprophilus; Germany). — Ciceroni and Zanetti, 1995: 10 (Coprophilus; Italy). — Owen, 1999a: 15 (Coprophilus; subterranean capture; Britain). — Owen, 2000: 251 (Coprophilus; pitfall trap collections from underground at roots of trees; Britain). DISTRIBUTION: Europe, Canada, USA. laesus Germar, 1832: 2 (Oxytelus; Type locality: Harcyniae). — Erichson, 1839a: 609 (Coprophilus; synonym of striatulus). — Heer, 1839: 199 (Coprophilus; synonym of striatulus). — Erichson, 1840: 816 (Coprophilus; synonym of striatulus). — Kraatz, 1857d: 894 (Coprophilus; synonym of striatulus). — Mulsant and Rey, 1878: 685 (Coprophilus; synonym of striatulus). Bulletin AMNH OXYTELINAE: COPROPHILUS 1319
— Ganglbauer, 1895: 677 (Coprophilus; synonym of striatulus). — Bernhauer, 1908e: 330 (Coprophilus; synonym of striatulus). — Bernhauer and Schubert, 1911: 90 (Coprophilus; synonym of striatulus). subplagiatus Cameron, 1928c: 558 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; Type locality: Tibet: Jelap La, 13,000 feet). — Cameron, 1930: 173 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters; Tibet). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Herman, 1970: 368 (Coprophilus). — Scheerpeltz, 1976: 17 (Coprophilus; subgenus Zonyptilus, cited as Zonoptilus; characters). DISTRIBUTION: China. vandykei Hatch, 1957, see: Vicelva. weiseri Bernhauer, 1927, see: Homalotrichus. Coprostygnus [4 species; Australian Region] Coprostygnus Sharp, 1886a: 380 (species included: sculptipennis). TYPE SPECIES: Coprostygnus sculptipennis Sharp, fixed by monotypy. — Broun, 1893: 1027 (characters; [Note: Cited as Coprostygnus nov. gen. but followed by a citation to Sharp’s description]). — Bernhauer and Schubert, 1911: 88 (world catalog; 2 species). — Scheerpeltz, 1933: 1075 (world catalog supplement). — Blackwelder, 1952: 106 (type species: sculptipennis). — Herman, 1970: 365 (characters; distribution; notes; list of species). — Klimaszewski, Newton, and Thayer, 1996: 150 (endemic to New Zealand). curvipes Broun, 1921: 484 (Coprostygnus; Type locality: west of Mount Algidus). — Scheerpeltz, 1933: 1075 (Coprostygnus; catalog). — Herman, 1970: 365 (Coprostygnus). DISTRIBUTION: New Zealand. optandus Broun, 1893: 1409 (Coprostygnus; Type locality: Capleston, Westland). — Fauvel, 1900c: 181 (Coprostygnus; New Zealand). — Bernhauer and Schubert, 1911: 88 (Coprostygnus; catalog). — Herman, 1970: 365 (Coprostygnus). DISTRIBUTION: New Zealand. picipennis Broun, 1921: 485 (Coprostygnus; Type locality: Glenhope). — Scheerpeltz, 1933: 1075 (Coprostygnus; catalog). — Herman, 1970: 365 (Coprostygnus). DISTRIBUTION: New Zealand. sculptipennis Sharp, 1886a: 380 (Coprostygnus; Type locality: New Zealand: Greymouth). — Broun, 1893: 1027 (Coprostygnus; [Note: Cited as “C. sculptipennis, n.s.” but followed by a citation to Sharp’s description]; characters; New Zealand). — Fauvel, 1900c: 181 (Coprostygnus; New Zealand). — Bernhauer and Schubert, 1911: 88 (Coprostygnus; catalog). — Herman, 1970: 365 (Coprostygnus). DISTRIBUTION: New Zealand. Homalotrichus [7 species; Neotropical and Australian Regions] Homalotrichus Solier, 1849: 321 (species included: striatus, impressicollis, obscurus, fuscus, luteipes). TYPE SPECIES: Homalotrichus striatus Solier, fixed by subsequent desig- nation by Tottenham, 1939d: 229. — Lacordaire, 1854: 153 (characters; notes; list of species). — Fairmaire and Germain, 1861: 452 (characters). — Fauvel, 1867a: 39 [= 1868: 44] (cited as synonym of Coprophilus; notes; characters; key to Chilean species). 1320 OXYTELINAE: HOMALOTRICHUS Bulletin AMNH
— Gemminger and Harold, 1868: 656 (synonym of Coprophilus). — Bernhauer and Schubert, 1911: 89 (synonym of Coprophilus). — Tottenham, 1939d: 229 (type species designation: striatus). — Blackwelder, 1944: 102 (cited as Coprophilus; checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 190 (type species). — Coiffait and Saiz, 1968: 415 (characters; key to Chilean species). — Herman, 1970: 368 (characters; distribution; notes; list of species). Sharpia Fauvel, 1878g: 488 (species included; banksi; preoccupied by Tournier, 1874, and Bocage, 1878). TYPE SPECIES: Sharpia banksi Fauvel, fixed by monotypy. — Duvivier, 1883: 193 (catalog). — Blackwelder, 1952: 351 (type species). — Herman, 1970: 368 (synonym of Homalotrichus). Toxoderus Fauvel, 1900c: 189 (replacement name for Sharpia Fauvel). TYPE SPECIES: Sharpia banksi Fauvel, fixed by objective synonymy with Sharpia. — Bernhauer and Schubert, 1911: 91 (world catalog; 1 species). — Blackwelder, 1952: 391 (type species). — Herman, 1970: 368 (synonym of Homalotrichus). banksi Fauvel, 1878g: 488 (Sharpia; Type locality: Australie méridionale, Victoria). — Bernhauer and Schubert, 1911: 91 (Toxoderus; catalog). — Herman, 1970: 369 (Homalotrichus). DISTRIBUTION: Australia. fuscus Solier, 1849, see: Thinodromus. impressicollis Solier, 1849: 323 (Homalotrichus; Type locality: Chile: San Carlos). — Kraatz, 1859a: 6 (Homalotrichus; notes). — Fairmaire and Germain, 1861: 452 (Homalotrichus; characters; Chile). — Fauvel, 1864a: 129 (Homalotrichus; notes; Chile). — Fauvel, 1867a: 41 [= 1868: 46] (Coprophilus; characters; Chile). — Bernhauer and Schubert, 1911: 89 (Coprophilus; catalog). — Coiffait and Saiz, 1968: 419 (Homalotrichus; characters; Chile). — Saiz, 1969a: 10 (Homalotrichus; notes). — Herman, 1970: 369 (Homalotrichus). — Scheerpeltz, 1972: 9 (Coprophilus; Argentina). DISTRIBUTION: Chile, Argentina. luteipes Solier, 1849, see: Thinodromus. nahuelbutensis Coiffait and Saiz, 1968: 418 (Homalotrichus; Type locality: Chili: Nahuelbuta, Malleco). DISTRIBUTION: Chile. obscurus Solier, 1849, see: Carpelimus. parvipennis Scheerpeltz, 1972: 64 (Coprophilus; subgenus Coprophilus; Type locality: Argentinien: Rio Negro, El Bolson, 220 m). — Herman, 2001: 14 (Homalotrichus). DISTRIBUTION: Argentina. striatus Solier, 1849: 323 (Homalotrichus; Type locality: Chile: provincia de Valdivia). — Kraatz, 1859a: 6 (Homalotrichus; characters). — Fairmaire and Germain, 1861: 452 (Homalotrichus; characters; Chile). — Fauvel, 1864a: 129 (Homalotrichus; notes; Chile). — Fauvel, 1867a: 42 [= 1868: 47] (Coprophilus; characters; Chile). — Bernhauer and Schubert, 1911: 90 (Coprophilus; catalog). — Coiffait and Saiz, 1968: 417 (Homalotrichus; characters; Chile). — Saiz, 1969a: 10 (Homalotrichus; notes). — Herman, 1970: 369 (Homalotrichus). — Scheerpeltz, 1972: 9 (Coprophilus; Argentina). DISTRIBUTION: Chile, Argentina. Bulletin AMNH OXYTELINAE: HOMALOTRICHUS 1321 substriatus Kraatz, 1859a: 16 (Homalotrichus; Type locality: Chile). — Fairmaire and Germain, 1861: 452 (Homalotrichus; characters; Chile). — Fauvel, 1867a: 42 [= 1868: 47] (Coprophilus; characters; Chile). — Bernhauer and Schubert, 1911: 90 (Coprophilus; catalog). — Coiffait and Saiz, 1968: 419 (Homalotrichus; characters; Chile). — Herman, 1970: 369 (Homalotrichus). — Scheerpeltz, 1972: 9 (Coprophilus; Argentina). — Saiz, Solervicens, and Ojeda, 1989: 52 (Homalotrichus; collecting notes; habitus; Chile). DISTRIBUTION: Chile, Argentina. weiseri Bernhauer, 1927e: 229 (Coprophilus; Type locality: Argentinien: Prov. Catamarca, Peñas Azules). — Scheerpeltz, 1933: 1076 (Coprophilus; catalog). — Herman, 1970: 369 (Homalotrichus). DISTRIBUTION: Argentina.
Tribe Deleasterini Deleasterini Reitter, 1909: 164 (genera included: Syntomium, Deleaster). TYPE GENUS: Deleaster (see Newton and Thayer, 1992: 60). — Portevin, 1929: 428 (characters; key to French genera). — Scheerpeltz, 1933: 1074 (synonym of Coprophili). — Crowson, 1955: 37 (genera). — Hatch, 1957: 84 (characters). — Newton and Thayer, 1992: 60 (synonym of Coprophilini). — M. Hansen, 1996: 37, 98 (separate from Coprophilini; list of Danish species). — Newton, Thayer, Ashe, and Chandler, 2000: 321, 378 (key to Nearctic genera; characters; notes). Pholidiens Mulsant and Rey, 1877: 65 (genera included: Deinopsis, Pholidus, Micropeplus). TYPE GENUS: Pholidus (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Coprophilini). Pholidini Acloque, 1896: 145 (genera included: Pholidus). TYPE GENUS: Pholidus (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Coprophilini). Euphaniae Reitter, 1909: 16 (genera included: Euphanias, Pholidus; [Note: Cited on page 199 as Pholidinae]). TYPE GENUS: Euphanias (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Coprophilini). Syntomiinae Böving and Craighead, 1931: 28 (genera included: Syntomium; larval char- acters). TYPE GENUS: Syntomium (see Newton and Thayer, 1992: 60). — Bernet Kempers, 1941: 34, 36 (larval characters). — Crowson, 1955: 37 (genera). — Newton and Thayer, 1992: 60 (synonym of Coprophilini). Syntomiini: — M. Hansen, 1996: 37, 98 (separate from Coprophilini; list of Danish species). Deleasterini Hatch, 1957: 84, 86 (genera included: Syntomium, Deleaster, Zalobius, Asemo- bius). TYPE GENUS: Deleaster (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Coprophilini).
Deleaster [11 species (1 doubtful); Nearctic, Palaearctic, and Ethiopian Regions] Deleaster Erichson, 1839a: 610 (species included: dichrous). TYPE SPECIES: Anthophagus dichroa Gravenhorst, fixed by monotypy. — Heer, 1839: 198 (characters). — Stephens, 1839: 422 (characters). — Erichson, 1840: 818 (characters). 1322 OXYTELINAE: DELEASTER Bulletin AMNH
— Redtenbacher, 1849: 743 (characters). — Schaum, 1852: 29 (list of species; Europe). — Lacordaire, 1854: 122 (characters; notes; list of species). — Fairmaire and Laboulbène, 1856: 621 (characters). — Gistel, 1856: 389 (list of species). — Jacquelin du Val, 1857: 60 (characters). — Redtenbacher, 1857: 238 (characters; Austrian species). — Kraatz, 1857d: 901 (characters). — G. Waterhouse, 1858: 30 (catalog of British species). — Schaum, 1859: 31 (catalog; European species). — Gredler, 1863: 119 (list of species from Tirol). — Wencker and Silbermann, 1866: 35 (list of species; collecting notes; France). — Gemminger and Harold, 1868: 656 (catalog). — Jacquelin du Val, 1868: 78 (catalog; European species). — Sharp, 1871d: 14 (list of British species). — Fauvel, 1871a: 151 [= 1872: 125] (characters). — Redtenbacher, 1874: 262 (characters). — Fauvel, 1875a:VIII [= 1875b: 210] (catalog). — Mulsant and Rey, 1878: 818 (characters). — Duvivier, 1883: 193 (catalog). — Fowler, 1888: 396 (notes). — Heyden, Reitter, and Weise, 1891: 117 (list of species of Europe and Caucasus). — Ganglbauer, 1895: 677 (characters). — Heyden, Reitter, and Weise, 1906: 137 (list of species of Europe and Caucasus). — Reitter, 1909: 177 (characters). — Bernhauer and Schubert, 1911: 88 (world catalog; 5 species). — Petri, 1912: 56 (list of species of Siebenbürgen). — Winkler, 1925: 338 (catalog for Palaearctic region). — Porta, 1926: 35 (characters; species of Italy). — Roubal, 1930: 321 (catalog; Slovakia). — Joy, 1932: 144 (key to British species). — Scheerpeltz, 1933: 1075 (world catalog supplement). — Jeannel and Jarrige, 1949: 324 (cave species; type species: dichrous). — Kloet and Hincks, 1945: 162 (list of British species). — Tottenham, 1949: 359, 408 (type species: dichrous Gravenhorst; list of British species). — Blackwelder, 1952: 118 (type species: dichroa). — Tottenham, 1954: 38 (characters; British species). — Hatch, 1957: 84, 87 (characters; species of Pacific Northwest). — Kocher, 1958: 95 (checklist of species; Morocco). — Palm, 1961a: 7 (characters; species of Sweden). — Horion, 1963: 159 (list of central European species). — Székessy, 1963: 27 (characters). — Lohse, 1964: 72 (characters; key to central European species). — Moore, 1964: 57 (characters). — Scheerpeltz, 1968a: 20 (catalog of Austrian species). — Herman, 1970: 363 (characters; distribution; notes; list of species). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Moore and Legner, 1975: 243 (catalog; North America north of Mexico). — Shibata, 1976: 130 (checklist of species of Japan). — Pope, 1977: 24 (list of British species). — Topp, 1978: 310 (larval characters). — Burakowski, Mroczkowski, and Stefan´ska, 1979: 80 (catalog; Poland). — Moore and Legner, 1979: 222 (characters; notes). — Muona, 1979: 19 (list of Scandinavian species). — Shibata, 1986a: 118 (supplemental list of species from Taiwan). — Dettner, 1987: 31, 32 (defensive glands). — M. Hansen, Kristensen, Mahler, Pedersen, 1991: 106 (list of Danish species). Bulletin AMNH OXYTELINAE: DELEASTER 1323
— Campbell and Davies, 1991: 92 (checklist of species; Canada). — Silfverberg, 1992: 23 (list of species of Fennoscandia, Denmark, and the Baltic States). — Ciceroni and Zanetti, 1995: 10 (list of species of Italy). — M. Hansen, 1996: 98 (list of Danish species). — Downie and Arnett, 1996: 441 (characters; species of Northeastern North America). — Schülke, 1998i: 125 (characters). — Newton, Thayer, Ashe, and Chandler, 2000: 378 (3 Nearctic species; notes; characters in key). adustus Küster, 1846, see: dichrous Gravenhorst, 1802. bactrianus Semenow, 1900: 684 (Deleaster; Type locality: Bucharia orient.: prov. Hissar: cursus superior fluvii Karatag). — Bernhauer and Schubert, 1911: 88 (Deleaster; catalog). — Scheerpeltz, 1963: 5 (Deleaster; Afghanistan). — Herman, 1970: 364 (Deleaster). — Coiffait, 1979d: 554 (Deleaster; Afghanistan). — Kashcheev, 1999: 149 (Deleaster; characters). DISTRIBUTION: Uzbekistan, Afghanistan. bergi Kashcheev, 1999: 148 (Deleaster; Type locality: okr. g. Almaty). DISTRIBUTION: Kazakhstan. brassicae Scopoli, 1763, see: dichrous Gravenhorst, 1802. concolor LeConte, 1866: 375 (Deleaster; Type locality: San Francisco, California). — Fauvel, 1878d: 256 [= 1878e: 92] (Deleaster; characters; California). — Fall and Cockerell, 1907: 168 (Deleaster; New Mexico). — Bernhauer and Schubert, 1911: 88 (Deleaster; catalog). — Hatch, 1957: 87 (Deleaster; characters; British Columbia; Washington; Oregon). — Herman, 1970: 364 (Deleaster). DISTRIBUTION: Canada, USA. dichrous Gravenhorst, 1802: 188 (Anthophagus; Type locality: Dresdae). — Latreille, 1804: 369 (Lesteva; characters). — Gravenhorst, 1806: 221 (Anthophagus; cited as dichrus). — Mannerheim, 1830: 55 (Lesteva; Germany). — Mannerheim, 1831: 469 (Lesteva; Germany). — Ménétriés, 1832: 147 (Lesteva; Caucasus-Persia region). — Stephens, 1834: 359 (Lesteva; characters; England; Scotland). — Runde, 1835: 23 (Lesteva; characters; Germany). — Lacordaire, 1835: 479 (Lesteva; characters; France). — Guérin-Méneville, 1829: pl. 9, fig. 10 (Lesteva; illustration). — Erichson, 1839a: 611 (Deleaster; characters; Germany). — Heer, 1839: 198 (Deleaster; characters; Switzerland). — Stephens, 1839: 431 (Deleaster; characters; England). — Erichson, 1840: 819 (Deleaster; characters; Germany; France; England). — Dufour, 1843: 37 (Anthophagus; characters; France). — Kiesenwetter, 1844a: 376 (Deleaster; notes; Germany). — Guérin-Méneville, 1844a: 32 (Lesteva; Europe). — Küster, 1846: 47 (Deleaster; characters; Germany; Italy; Switzerland; France; England). — Hochhuth, 1849: 200 (Deleaster; characters). — Redtenbacher, 1849: 743 (Deleaster; characters; Austria). — Kiesenwetter, 1851a: 433 (Deleaster; France). — Fairmaire and Laboulbène, 1856: 621 (Deleaster; characters; France). — Redtenbacher, 1857: 238 (Deleaster; characters; Austria). — Kraatz, 1857d: 902 (Deleaster; characters; Germany). — Hochhuth, 1862: 101 (Deleaster; notes; Russia). — Fauvel, 1865a: 314 [= 1865b: 64] (Deleaster; habitat; France). — Hochhuth, 1872: 168 (Deleaster; notes; Ukraine). — Fauvel, 1871a: 152 [= 1872: 126] (Deleaster; characters; France; Europe; Caucasus; Armenia; Iran). 1324 OXYTELINAE: DELEASTER Bulletin AMNH
— Redtenbacher, 1874: 263 (Deleaster; characters; Austria). — Seidlitz, 1875: 243 (Deleaster; characters; Baltic region). — Mulsant and Rey, 1878: 820 (Deleaster; characters; France). — Fauvel, 1886: 16 [= 1886a: 8] (Deleaster; Morocco). — Fowler, 1888: 396 (Deleaster; characters; Britain). — Seidlitz, 1891: 349 (Deleaster; characters; Europe). — Seidlitz, 1891a: 373 (Deleaster; characters; Europe). — Ganglbauer, 1895: 678 (Deleaster; characters; middle Europe; eastern Mediterranean region). — Fauvel, 1897d: 249 (Deleaster; Morocco). — Fauvel, 1902b: 57 (Deleaster; Algeria; Tunisia). — Sainte-Claire Deville, 1906: 72 (Deleaster; Corsica). — Sainte-Claire Deville, 1907: 56 (Deleaster; characters; France). — Reitter, 1909: 178 (Deleaster; characters; Germany). — Bernhauer and Schubert, 1911: 88 (Deleaster; catalog). — Portevin, 1929: 428 (Deleaster; characters; France). — Bernhauer, 1931e: 9 (Deleaster; Morocco). — Scheerpeltz, 1933: 1075 (Deleaster; catalog). — Olsson, 1934: 67 (Deleaster; Sweden). — Chagnon, 1934 (Deleaster; characters; Canada). — Jules, 1935 (Deleaster; characters; Canada). — C. Koch, 1941: 54 (Deleaster; Morocco). — MacNay, 1946: 213 (Deleaster; Ontario). — Tottenham, 1949: 408 (Deleaster; checklist; Britain). — Jeannel and Jarrige, 1949: 324 (Deleaster; notes; cave species; Spain; Serbia; Romania). — Palm, 1953: 9 (Deleaster; ecological notes; Sweden). — Kocher, 1958: 95 (Deleaster; Morocco). — Palm, 1961a: 7 (Deleaster; characters; Sweden; Denmark; Finland). — Horion, 1963: 159 (Deleaster; Morocco; Iran; Britain; Holland; Belgium; France; Switzerland; Spain; Corsica; Sardinia; Sicily; Italy; Croatia; Bosnia Hercegovina; Albania; Macedonia; Greece; Romania; Hungary; Czechoslovakia; Poland; Russia; Denmark; Sweden; Finland; Germany; Austria). — Lohse, 1964: 72 (Deleaster; characters; central Europe). — Smetana, 1967e: 307 (Deleaster; Turkey). — Szujecki, 1968a: 712 (Deleaster; Poland). — Scheerpeltz, 1968a: 20 (Deleaster; catalog; Austria). — Bolov, 1969: 513 (Deleaster; collecting notes; Russia). — Bolov, 1969a: 328 (Deleaster; collecting notes; Russia). — Herman, 1970: 365 (Deleaster). — Pope, 1977: 24 (Deleaster; Britain). — Bordoni, 1978a: 55 (Deleaster; cave collection; Turkey). — Muona, 1979: 19 (Deleaster; Finland; Russia; Sweden; Denmark). — Burakowski, Mroczkowski, and Stefan´ska, 1979: 80 (Deleaster; catalog; Poland). — Frank, 1982: 13 (Deleaster; list of parasites). — Dettner, Schwinger and Wunderle, 1985 (Deleaster; defensive gland morphology, secretions, function). — Segers, 1986: 23 (Deleaster; checklist of species; Belgium). — Lucht, 1987: 91 (Deleaster; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden). — M. Hansen, Kristensen, Mahler, and Pedersen, 1991: 106 (Deleaster; Denmark). — Silfverberg, 1992: 23 (Deleaster; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania). — Gusarov, 1992a: 776 [= 1993a: 63] (Deleaster; synonymic notes). — Bohácˇ, 1993: 42 (Deleaster; Czech Republic; Slovakia). — Dearborn and Donahue, 1994: 14 (Deleaster; Maine). — Terlutter, 1995: 5 (Deleaster; Germany). — Gerardi and Zanetti, 1995: 143 (Deleaster; collecting notes; Italy). Bulletin AMNH OXYTELINAE: DELEASTER 1325
— Sparacio, 1995: 144 (Deleaster; characters; summary of distribution; notes; Sicily). — Ciceroni and Zanetti, 1995: 10 (Deleaster; Italy). — Downie and Arnett, 1996: 441 (Deleaster; characters; Canada). — Franzen, 1996: 206 (Deleaster; collecting notes; Germany). — M. Hansen, 1996: 98 (Deleaster; checklist; Denmark). — Sterrenburg and Schülke, 1997: 16 (Deleaster; Netherlands). — Czarniawski and Staniec, 1997: 38 (Deleaster; structure of opening of abdominal defensive reservoirs). — Wenzel, 1997: 345 (Deleaster; collecting notes; Germany). — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 45 (Deleaster; Latvia). — Anderson, Nash, and O’Connor, 1997: 16 (Deleaster; Ireland). — Hansen, Jørum, Palm, and Pedersen, 1997: 125 (Deleaster; checklist; Denmark). — Outerelo, Gamarra, and Salgado, 1998: 124 (Deleaster; collected from cave; Spain). — Kashcheev, 1999: 149 (Deleaster; characters). — Eyre, Luff, and Lott, 2000: 33 (Deleaster; Britain). — Herman, 2001: 23 (Deleaster; nomen protectum). DISTRIBUTION: Morocco, Algeria, Tunisia, Europe, Russia, Armenia, Turkey, Iran, USA, Canada. brassicae Scopoli, 1763: 102 (Staphylinus; [Note: This name is older than dichrous. Bern- hauer and Schubert (1911: 88) and other authors cited Schrank, 1798 as the author of brassicae but Schrank attributed the species to Scopoli]; Type locality: Carniola). — Goeze, 1777: 725 (Staphylinus; characters). — Villers, 1789: 424 (Staphylinus; characters). — Gmelin, 1790: 2032 (Staphylinus; page misnumbered as 1032; characters). — Schrank, 1798: 644 (Staphylinus; characters; notes). — Turton, 1802: 516 (Staphylinus; characters; Carniola). — Erichson, 1840: 819 (Deleaster; synonym of dichrous). — Redtenbacher, 1849: 743 (Deleaster; synonym of dichrous). — Kraatz, 1857d: 902 (Deleaster; synonym of dichrous). — Ganglbauer, 1895: 678 (Deleaster; synonym of dichrous). — Bernhauer and Schubert, 1911: 88 (Deleaster; synonym of dichrous). — Herman, 2001: 23 (Deleaster; nomen oblitum). leachii Curtis, 1830: pl. 303 (Lesteva; Type locality: England: Cumberland, near Holme Gate, banks of river Eden). — Stephens, 1834: 359 (Lesteva; synonym of dichrous Gravenhorst). — Erichson, 1839a: 611 (Deleaster; synonym of dichrous). — Stephens, 1839: 431 (Deleaster; synonym of dichroa). — Erichson, 1840: 819 (Deleaster; synonym of dichrous). — Kraatz, 1857d: 902 (Deleaster; synonym of dichrous). — Fauvel, 1871a: 152 [= 1872: 126] (Deleaster; synonym of dichrous). — Mulsant and Rey, 1878: 821 (Deleaster; synonym of dichrous). — Fowler, 1888: 397 (Deleaster; variety of dichrous; characters; Britain). — Ganglbauer, 1895: 678 (Deleaster; variety of dichrous). — Bernhauer and Schubert, 1911: 88 (Deleaster; variety of dichrous). — Scheerpeltz, 1933: 1075 (Deleaster; aberration of dichrous). — Smetana, 1967e: 307 (Deleaster; aberration of dichrous; Turkey). adustus Küster, 1846: 48 (Deleaster; Type locality: Rothenthurmpass in Siebenbürgen). — Redtenbacher, 1849: 743 (Syntomium; characters). — Kraatz, 1857d: 902 (Deleaster; synonym of dichrous). — Fauvel, 1869: 493 (Deleaster; synonym of dichrous). — Fauvel, 1871a: 152 [= 1872: 126] (Deleaster; synonym of dichrous). — Ganglbauer, 1895: 678 (Deleaster; synonym of dichrous). — Bernhauer and Schubert, 1911: 88 (Deleaster; synonym of dichrous). — Scheerpeltz, 1933: 1075 (Deleaster; synonym of dichrous). 1326 OXYTELINAE: DELEASTER Bulletin AMNH
erichsoni Hochhuth, 1851: 57 (Deleaster; Type locality: Gebirge Armeniens; [Note: See lectotype designation by Gusarov, 1992a]). — Fauvel, 1871a: 152 [= 1872: 126] (Deleaster; synonym of dichrous). — Ganglbauer, 1895: 678 (Deleaster; synonym of dichrous). — Bernhauer and Schubert, 1911: 88 (Deleaster; variety of dichrous). — Scheerpeltz, 1933: 1075 (Deleaster; variety of dichrous). — Gusarov, 1992a: 776 [= 1993a: 63] (Deleaster; synonym of dichrous; lectotype designation). testaceicolor Roubal, 1941: 10 (Deleaster; aberration of dichrous; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Ross. subcarp.: Rachov-Mencul; Slov.: Trencin). dryophilus Gistel, 1857: 11 (Deleaster; [Note: Nomen dubium]; Type locality: Styriae). DISTRIBUTION: Austria. erichsoni Hochhuth, 1851, see: dichrous Gravenhorst, 1802. grandiceps† Wickham, 1912: 12 (Deleaster; Miocene fossil; Type locality: Florissant, Colorado). DISTRIBUTION: USA. leachii Curtis, 1830, see: dichrous Gravenhorst, 1802. pectinatus Fauvel, 1882b: 129 (Deleaster; Type locality: Abyssinie). — Bernhauer and Schubert, 1911: 88 (Deleaster; catalog). — Herman, 1970: 365 (Deleaster). DISTRIBUTION: Ethiopia. pekinensis Fairmaire, 1893a: ccxii (Deleaster; Type locality: Peking). — Bernhauer and Schubert, 1911: 88 (Deleaster; catalog). — Herman, 1970: 365 (Deleaster). — J. Li, 1993: 23 (Deleaster; China). DISTRIBUTION: China. taiwanensis Hayashi, 1984: 91 (Deleaster; Type locality: Mt. Lishan, alt. about 2500 m, Nantou Hsien, Taiwan). DISTRIBUTION: Taiwan. testaceicolor Roubal, 1941 [Note: Nomen nudum], see: dichrous Gravenhorst, 1802. trimaculatus Fall, 1910: 115 (Deleaster; Type locality: Cañon City, Colorado). — Scheerpeltz, 1933: 1075 (Deleaster; catalog). — Herman, 1970: 365 (Deleaster). — Moore, 1975d: 174 (Deleaster; Arizona; California). DISTRIBUTION: USA. yokoyamai Adachi, 1935: 126 (Deleaster; Type locality: Shimajima Valley, Shinano Province). — Nakane, 1963a: 83 (Deleaster; characters; habitus photograph; Japan). — Herman, 1970: 365 (Deleaster). — Frank, 1982: 13 (Deleaster; list of parasites). DISTRIBUTION: Japan. Euphanias [5 species; Neotropical and Palaearctic Regions] Euphanias Fairmaire and Laboulbène, 1856: 657 (species included: insignicornis). TYPE SPECIES: Euphanias insignicornis Fairmaire and Laboulbène, fixed by monotypy. — Kraatz, 1857a: 51 (characters). — Redtenbacher, 1857: 991 (characters). — Jacquelin du Val, 1857: 65 (characters). — Kraatz, 1857d: 806 (characters; notes). — Schaum, 1859: 31 (catalog; European species). — Jacquelin du Val, 1868: 79 (catalog; European species). — Gemminger and Harold, 1868: 655 (catalog). Bulletin AMNH OXYTELINAE: EUPHANIAS 1327
— Fauvel, 1869: 493 (synonym of Pholidus). — Fauvel, 1871a: 158 [= 1872: 132] (synonym of Pholidus). — Redtenbacher, 1874: 260 (characters). — Fauvel, 1875a:VIII [= 1875b: 210] (synonym of Pholidus). — Ganglbauer, 1895: 689 (synonym of Pholidus). — Bernhauer and Schubert, 1910: 8 (world catalog; Piestini). — Winkler, 1925: 323 (catalog for Palaearctic region). — Scheerpeltz, 1933: 994 (world catalog supplement). — Blackwelder, 1944: 101 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 157 (type species: insignicornis). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Blinshtein, 1978: 80 (key to species of Mediterranean region). — Newton, 1982a: 6, 8 (characters; transferred to Oxytelinae). — Ciceroni and Zanetti, 1995: 10 (list of species of Italy). Pholidus Mulsant and Rey, 1856a: 7 [= 1856b: 7] (species included: insignis; preoccupied by Rafinesque, 1815, and Gray, 1840). TYPE SPECIES: Pholidus insignis Mulsant and Rey, fixed by monotypy. — Redtenbacher, 1857: 991 (synonym of Euphanias). — Schaum, 1859: 31 (synonym of Euphanias). — Gemminger and Harold, 1868: 655 (synonym of Euphanias). — Fauvel, 1871a: 158 [= 1872: 132] (characters). — Fauvel, 1875a:VIII [= 1875b: 210] (catalog). — Mulsant and Rey, 1880: 408 (characters). — Heyden, Reitter, and Weise, 1891: 117 (list of species of Europe and Caucasus). — Ganglbauer, 1895: 689 (characters). — Heyden, Reitter, and Weise, 1906: 125 (list of species of Europe and Caucasus). — Bernhauer and Schubert, 1910: 8 (synonym of Euphanias). — Porta, 1926: 6 (characters; species of Italy). — Scheerpeltz, 1933: 994 (synonym of Euphanias). — Blackwelder, 1952: 306 (type species: insignis). boiteli Normand, 1936: 163 (Euphanias; Type locality: Tunisie: Bords du lac de Bizerte). — Blinshtein, 1978: 80 (Euphanias; characters). DISTRIBUTION: Tunisia. ganglbaueri Bernhauer, 1908d: 286 (Pholidus; Type locality: Brasilien: Blumenau). — Bernhauer and Schubert, 1910: 8 (Euphanias; catalog). DISTRIBUTION: Brazil. insignicornis Fairmaire and Laboulbène, 1856, see: insignis Mulsant and Rey, 1856. insignis Mulsant and Rey, 1856a: 9 [= 1856b: 9] (Pholidus; Type locality: Hyères; [Note: See lectotype designation by Gusarov, 1997]). — Fauvel, 1871a: 158 [= 1872: 132] (Pholidus; characters; France). — Mulsant and Rey, 1880: 409 (Pholidus; characters; France). — Ganglbauer, 1895: 690 (Pholidus; characters; France). — Fauvel, 1897d: 250 (Pholidus; Algeria). — Fauvel, 1902b: 58 (Pholidus; Algeria; Tunisia; Egypt). — Bernhauer and Schubert, 1910: 8 (Euphanias; catalog). — Portevin, 1929: 458 (Pholidus; characters; France). — Scheerpeltz, 1933: 994 (Euphanias; catalog). — C. Koch, 1934: 33 (Euphanias; characters; Egypt). — C. Koch, 1936b: 126 (Euphanias; Egypt). — Blinshtein, 1978: 80 (Euphanias; characters). — Haghebaert, 1989a: 277 (Euphanias; notes; Camargue). — Sparacio, 1995: 144 (Euphanias; characters; notes; Sicily). — Ciceroni and Zanetti, 1995: 10 (Euphanias; Italy). 1328 OXYTELINAE: EUPHANIAS Bulletin AMNH
— Adorno, Ciceroni, and Zanetti, 1996: 51 (Euphanias; notes; Sicily). — Gusarov, 1997: 278 (Euphanias; lectotype designation; Saudi Arabia). DISTRIBUTION: Algeria, Tunisia, Egypt, Saudi Arabia, Syria, France. insignicornis Fairmaire and Laboulbène, 1856: 657 (Euphanias; Type locality: Hyères). — Redtenbacher, 1857: 991 (Euphanias; characters; Austria). — Fauvel, 1869: 493 (Euphanias; synonym of insignis). — Fauvel, 1871a: 158 [= 1872: 132] (Pholidus; synonym of insignis). — Redtenbacher, 1874: 260 (Euphanias; characters). — Mulsant and Rey, 1880: 410 (Pholidus; synonym of insignis). — Ganglbauer, 1895: 690 (Pholidus; synonym of insignis). — Bernhauer and Schubert, 1910: 8 (Euphanias; synonym of insignis). — Scheerpeltz, 1933: 994 (Euphanias; synonym of insignis). insularis C. Koch, 1934: 33 (Euphanias; subspecies of insignis; Type locality: Sardinien: Cagliari). — Ciceroni and Zanetti, 1995: 10 (Euphanias; subspecies of insignis; Italy). insularis C. Koch, 1934, see: insignis Mulsant and Rey, 1856. pliginskii Bernhauer, 1912: 170 (Euphanias; Type locality: Sebastopol in der Krim). — Scheerpeltz, 1933: 995 (Euphanias; catalog). — Blinshtein, 1978: 80 (Euphanias; characters). DISTRIBUTION: Ukraine. pusanovi Blinshtein, 1976: 78 (Euphanias; Type locality: Cheromorskii zapovednik, Potievskii uchastok). DISTRIBUTION: Russia. Mitosynum [1 species; Nearctic Region] Mitosynum Campbell, 1982a: 687 (species included: vockerothi; habitus). TYPE SPECIES: Mitosynum vockerothi Campbell, fixed by original designation and monotypy. — Downie and Arnett, 1996: 441 (characters; species of Northeastern North America). — Newton, Thayer, Ashe, and Chandler, 2000: 379 (1 Nearctic species; notes; characters in key). vockerothi Campbell, 1982a: 690 (Mitosynum; Type locality: New Brunswick: Kouchi- bouguac N. P.). — Campbell and Davies, 1991: 93 (Mitosynum; checklist of species; Canada). — Downie and Arnett, 1996: 441 (Mitosynum; characters; USA). DISTRIBUTION: Canada. Oxypius [1 species; Australian Region] Oxypius Newton, 1982a: 2 (species included: peckorum; systematic position). TYPE SPECIES: Oxypius peckorum Newton, fixed by original designation. peckorum Newton, 1982a: 10 (Oxypius; Type locality: Australia: Western Australia: Walpole N.P., Tingle Tree; larval description also included). DISTRIBUTION: Australia. Syntomium [8 species; Nearctic and Palaearctic Regions] Syntomium Curtis, 1828: pl. 228 (species included: nigroaeneum). TYPE SPECIES: Syntomium nigroaeneum Curtis, fixed by original designation and monotypy. — Stephens, 1829a: 24 (catalog; Britain). — Stephens, 1834: 329 (characters; habitat). — Westwood, 1838a: 18 (characters; type species). — Heer, 1839: 199 (characters). — Stephens, 1839: 422 (characters). Bulletin AMNH OXYTELINAE: SYNTOMIUM 1329
— Erichson, 1840: 821 (characters). — Redtenbacher, 1849: 743 (characters). — Schaum, 1852: 29 (list of species; Europe). — Lacordaire, 1854: 123 (characters; notes; list of species). — Fairmaire and Laboulbène, 1856: 622 (characters). — Jacquelin du Val, 1857: 60 (characters). — Redtenbacher, 1857: 239 (characters; Austrian species). — Kraatz, 1857d: 890 (characters). — Thomson, 1858: 39 (characters). — G. Waterhouse, 1858: 30 (catalog of British species). — Thomson, 1859: 44 (characters; type species: aeneum). — Schaum, 1859: 31 (catalog; European species). — LeConte, 1861a: 69 (characters). — Thomson, 1861: 140 (characters). — LeConte, 1863: 26 (list of North American species). — Wencker and Silbermann, 1866: 35 (list of species; collecting notes; France). — Gemminger and Harold, 1868: 655 (catalog). — Jacquelin du Val, 1868: 78 (catalog; European species). — Fauvel, 1871a: 156 [= 1872: 130] (characters). — Sharp, 1871d: 14 (list of British species). — Redtenbacher, 1874: 261 (characters). — Fauvel, 1875a:VIII [= 1875b: 210] (catalog). — Mulsant and Rey, 1878: 823 (characters). — Fowler, 1888: 393 (notes; larval characters). — Heyden, Reitter, and Weise, 1891: 117 (list of species of Europe and Caucasus). — Ganglbauer, 1895: 679 (adult and larval characters). — Heyden, Reitter, and Weise, 1906: 137 (list of species of Europe and Caucasus). — Reitter, 1909: 178 (characters). — Bernhauer and Schubert, 1911: 84 (world catalog; 2 species). — Petri, 1912: 56 (list of species of Siebenbürgen). — Johansen, 1914: 580 (characters). — Leng, 1920: 95 (catalog of North American species). — Winkler, 1925: 337 (catalog for Palaearctic region). — Porta, 1926: 35 (characters; species of Italy). — Portevin, 1929: 394 (larval illustration). — Roubal, 1930: 321 (catalog; Slovakia). — Joy, 1932: 140 (key to British species). — Scheerpeltz, 1933: 1074 (world catalog supplement). — Kloet and Hincks, 1945: 162 (list of British species). — Tottenham, 1949: 359, 408 (type species: aeneum P. W. J. Müller = nigroaeneum Curtis; list of British species). — Blackwelder, 1952: 367 (type species: nigroaeneum). — Hatch, 1957: 84, 86 (characters; key to species of Pacific Northwest). — Watanabe and Shibata, 1960: 103 (list of species of world). — Palm, 1961a: 6 (characters; species of Sweden). — Horion, 1963: 157 (list of central European species). — Székessy, 1963: 27 (characters). — Lohse, 1964: 71 (characters; key to central European species). — Moore, 1964: 57 (characters). — Pototskaia, 1967: 24 (larval characters). — Kasule, 1968: 118 (larval characters). — Scheerpeltz, 1968a: 20 (catalog of Austrian species). — Herman, 1970: 372 (characters; distribution; notes; list of species). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Moore and Legner, 1975: 252 (catalog; North America north of Mexico). — Shibata, 1976: 133 (checklist of species of Japan). 1330 OXYTELINAE: SYNTOMIUM Bulletin AMNH
— Pope, 1977: 24 (list of British species). — Topp, 1978: 309 (larval characters). — Burakowski, Mroczkowski, and Stefan´ska, 1979: 80 (catalog; Poland). — Moore and Legner, 1979: 232 (characters; notes). — Muona, 1979: 19 (list of Scandinavian species). — Uhlig, Vogel, and Sieber, 1980: 241 (some species collected in Germany). — Uhlig and Vogler, 1981: 88 (list of some species collected in Germany; habitat and collecting notes). — Muona and Viramo, 1986: 16 (list of species of northeastern Finland). — Dettner, 1987: 31 (defensive glands). — Campbell and Davies, 1991: 93 (checklist of species; Alaska and Canada). — Silfverberg, 1992: 23 (list of species of Fennoscandia, Denmark, and the Baltic States). — Siitonen, 1993: 230 (list of Finnish species). — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species). — Ciceroni and Zanetti, 1995: 10 (list of species of Italy). — M. Hansen, 1996: 98 (list of Danish species). — M. Hansen, Mahler, Palm, and Pedersen, 1996: 241 (additions to list of Danish species). — Downie and Arnett, 1996: 441 (characters; species of Northeastern North America). — Schülke, 1998i: 125 (characters). — Newton, Thayer, Ashe, and Chandler, 2000: 379 (4 Nearctic species; notes; characters in key). Eunonia Casey, 1904: 313 (species included: keeniana). TYPE SPECIES: Eunonia keeniana Casey, fixed by original designation and monotypy. — Bernhauer and Schubert, 1910: 50 (world catalog; 1 species). — Leng, 1920: 94 (catalog of North American species). — Blackwelder, 1952: 157 (type species: keeniana). — Hatch, 1957: 51, 59 (characters; species of Pacific Northwest). — Moore, 1966: 56 (synonym of Syntomium). — Herman, 1970: 372 (synonym of Syntomium). aeneum P. W. J. Müller, 1821: 216 (Omalium; Type locality: Odenbach). — Gyllenhal, 1827: 466 (Omalium; characters; Sweden). — Heer, 1839: 199 (Syntomium; characters; Switzerland). — Erichson, 1840: 821 (Syntomium; characters; Germany; Sweden; England). — Redtenbacher, 1849: 743 (Syntomium; characters; Austria). — Küster, 1854: 23 (Syntomium; characters; Europe). — Hardy, 1851: 53 (Syntomium; notes; England). — Fairmaire and Laboulbène, 1856: 622 (Syntomium; characters; France). — Redtenbacher, 1857: 239 (Syntomium; characters; Austria). — Kraatz, 1857d: 892 (Syntomium; characters; Germany). — Thomson, 1859: 44 (type species of Syntomium). — Thomson, 1861: 141 (Syntomium; characters; Scandinavia). — Fauvel, 1871a: 157 [= 1872: 131] (Syntomium; characters; France; Lapland; Scandinavia; Britain; Germany; Switzerland; Austria; Italy). — Schiødte, 1873: 559 (Syntomium; larval characters). — Fauvel, 1874: 325 [= 1874b: 31] (Syntomium; Russia). — Redtenbacher, 1874: 260 (Synotomium; characters; Austria). — Seidlitz, 1875: 245 (Syntomium; characters; Baltic region). — J. Sahlberg, 1876: 182 (Syntomium; Finland). — Mulsant and Rey, 1878: 825 (Syntomium; characters; France). — Fowler, 1888: 393 (Syntomium; characters; Britain). — Seidlitz, 1891: 349 (Syntomium; characters; Europe). — Seidlitz, 1891a: 373 (Syntomium; characters; Europe). — Ganglbauer, 1895: 681 (Syntomium; characters; north and middle Europe). — Sainte-Claire Deville, 1907: 55 (Syntomium; characters; France). — Reitter, 1909: 178 (Syntomium; characters; Germany). Bulletin AMNH OXYTELINAE: SYNTOMIUM 1331
— Bernhauer and Schubert, 1911: 87 (Syntomium; catalog). — Johansen, 1914: 581 (Syntomium; characters; Denmark). — Portevin, 1929: 429 (Syntomium; characters; France). — Paulian, 1941: 156 (Syntomium; larval characters; France). — Jarrige, 1946: 96 (Syntomium; cited as aencum; natural history notes). — Palm, 1961a: 6 (Syntomium; characters; Sweden; Denmark; Norway; Finland). — Horion, 1963: 157 (Syntomium; Russia; Finland; Sweden; Norway; Denmark; Britain; Holland; Belgium; France; Switzerland; Italy; Croatia; Bosnia; Romania; Czechoslovakia; Poland; Germany; Austria). — Lohse, 1964: 71 (Syntomium; characters; central Europe). — Pototskaia, 1967: 24 (Syntomium; larval characters). — Herman, 1970: 373 (Syntomium). — Pope, 1977: 24 (Syntomium; Britain). — Segers, 1986: 23 (Syntomium; checklist of species; Belgium). — Lucht, 1987: 91 (Syntomium; checklist; Germany; Poland; Czechoslovakia; Austria; Switzer- land; France; Benelux; Denmark; Sweden). — Bohácˇ, 1993: 42 (Syntomium; Slovakia; Czech Republic). — Terlutter, 1995: 5 (Syntomium; Germany). — Ciceroni and Zanetti, 1995: 10 (Syntomium; Italy). DISTRIBUTION: Europe, Russia. nigroaeneum Curtis, 1828: pl. 228 (Syntomium; Type locality: England, Middlesex, Southgate). — Curtis, 1829: 28 (Bolitobius; catalog; Britain). — Stephens, 1834: 330 (Syntomium; characters; England). — Westwood, 1838a: 18 (Syntomium; type species). — Stephens, 1839: 424 (Syntomium; characters; England). — Erichson, 1840: 821 (Syntomium; synonym of aeneum). — Redtenbacher, 1849: 743 (Syntomium; synonym of aeneum). — Hardy, 1851: 53 (Syntomium; synonym of aeneum). — G. Waterhouse, 1858: 30 (Syntomium; synonym of aeneum). — Kraatz, 1857d: 892 (Syntomium; synonym of aeneum). — Mulsant and Rey, 1878: 825 (Syntomium; synonym of aeneum). — Ganglbauer, 1895: 681 (Syntomium; synonym of aeneum). — Bernhauer and Schubert, 1911: 86 (Syntomium; synonym of aeneum). confragosum Mäklin, 1852: 319 (Syntomium; Type locality: insula Sitkha). — Fauvel, 1878d: 258 [= 1878e: 94] (Syntomium; characters; Alaska; Lake Superior). — Bernhauer and Schubert, 1911: 87 (Syntomium; catalog). — Hatch, 1957: 86 (Syntomium; characters; British Columbia). — Herman, 1970: 373 (Syntomium). — Downie and Arnett, 1996: 441 (Syntomium; characters; Canada; USA). DISTRIBUTION: Canada, USA. grahami Hatch, 1957: 86 (Syntomium; Type locality: British Columbia: Stanley). — Herman, 1970: 373 (Syntomium). DISTRIBUTION: Canada. japonicum Watanabe and Shibata, 1960: 103 (Syntomium; Type locality: Itsukaichi, Nishitama, Tokyo, Japan). — Herman, 1970: 373 (Syntomium). DISTRIBUTION: Japan. keenianum Casey, 1904: 314 (Eunonia; Type locality: British Columbia, Metlakatla). — Bernhauer and Schubert, 1910: 50 (Eunonia; catalog). — Hatch, 1957: 69 (Eunonia; characters; British Columbia; Washington; Oregon). — Moore, 1966: 56 (Syntomium). — Herman, 1970: 373 (Syntomium). DISTRIBUTION: Canada, USA. 1332 OXYTELINAE: SYNTOMIUM Bulletin AMNH longicorne Peyerimhoff, 1913: 253 (Syntomium; Type locality: Azerou Tidjer, vers 1300 m). — Scheerpeltz, 1933: 1074 (Syntomium; catalog). — Herman, 1970: 373 (Syntomium). DISTRIBUTION: Algeria. malkini Hatch, 1957: 86 (Syntomium; Type locality: Oregon: Coos Co.: Charleston). — Herman, 1970: 373 (Syntomium). DISTRIBUTION: USA. marusiki Ryabukhin, 1992a: 147 (Syntomium; Type locality: Maradenskaia obl., 35 km severnee Maradana poc. Snezhnaia Dolina). — Ryabukhin, 1993: 162 (Syntomium; translation of Ryabukhin, 1992a; Magadan Province). — Ryabukhin, 1999: 35 (Syntomium; catalog; habitat; distribution; northeastern Russia). DISTRIBUTION: Russia. nigroaeneum Curtis, 1828, see: aeneum P. W. J. Müller, 1821.
Tribe Oxytelini Oxytelini: — Thomson, 1858: 39 (characters; key to genera). — Eichelbaum, 1909: 117 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera). — Reitter, 1909: 163 (Bledius, Platystethus, Haploderus, Oxytelus). — Blatchley, 1910: 462 (characters; included Megalops, Platystethus, Oxytelus, Trogophloeus, Apocellus; key to Indiana genera). — Bernhauer and Schubert, 1911: 87 (world catalog; Coprophili, Actochari, Toxoderi, Oxyteli, Thinobii). — Johansen, 1914: 530 (characters). — Porta, 1926: 34 (characters; species of Italy). — Cameron, 1930: 169 (characters; key to genera of British India). — Scheerpeltz, 1933: 1073 (world catalog supplement). — Scheerpeltz, 1940: 22 (key to Palaearctic genera). — Crowson, 1955: 37 (genera). — Arnett, 1963: 255 (characters; list of North American genera). — Pototskaia, 1967: 17, 21 (larval characters; key to some genera). — Tikhomirova, 1968: 146 (fossil species). — Herman, 1970: 398 (characters; notes). — Hammond, 1975: 145 (notes; characters; key to species of Ceylon). — Muona, 1979: 19 (list of Scandinavian species). — Outerelo and Gamarra, 1985: 100 (key to genera of Iberian Peninsula). — Newton and Thayer, 1992: 60 (tribe of Oxytelinae). — Klimaszewski, Newton, and Thayer, 1996: 150 (list of New Zealand genera; number of New Zealand species). — M. Hansen, 1996: 37, 98 (not clearly defined; list of Danish species). — Newton, Thayer, Ashe, and Chandler, 2000: 322, 380 (key to Nearctic genera; characters; notes). Oxytelida: — Heer, 1839a: 14 (cited as family). Oxytelitae: — Jeannel and Jarrige, 1949: 323 (notes). Oxytelina: — Jakobson, 1908: 446, 466 (characters; key to genera; list of species of Russia and western Europe). — Johansen, 1914: 533 (characters; key to Danish genera). — Arnett, 1963: 255 (characters; list of North American genera). Bulletin AMNH OXYTELINAE: OXYTELINI 1333
Oxyteli: — Bernhauer and Schubert, 1911: 91 (world catalog; Planeustomus, Ancyrophorus, Trogophloeus, Xerophygus, Blediotrogus, Trogactus, Parosus, Apocellus, Oxytelopsis, Delopsis, Oxytelodes, Hoplitodes, Aploderus, Anisopsis, Oxytelus, Platystethus, Bledius, Typhlobledius). — Scheerpeltz, 1933: 1078 (world catalog supplement). Apocellaria Lynch, 1884: 373 (genera included: Apocellus). TYPE GENUS: Apocellus (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Oxytelini). Ecitoclimacini Borgmeier, 1934: 452 (tribe of Aleocharinae; genera included: Ecitoclimax). TYPE GENUS: Ecitoclimax (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Oxytelini). Apocellini Hatch, 1957: 85 (genera included: Apocellus). TYPE GENUS: Apocellus (see Newton and Thayer, 1992: 60). — Newton and Thayer, 1992: 60 (synonym of Oxytelini).
Anisopsidius [1 species; Ethiopian Region] Anisopsidius Fagel, 1960: 8 (species included: quadricollis). TYPE SPECIES: Anisopsis quadri- collis Bernhauer, fixed by original designation and monotypy. — Herman, 1970: 407 (characters; distribution; notes; list of species). quadricollis Bernhauer, 1932a: 76 (Anisopsis; Type locality: Haut-Uele: Moto). — Scheerpeltz, 1933: 1093 (Anisopsis; catalog). — Fagel, 1957a: 35 (Anisopsidius; Ruanda). — Fagel, 1960: 10 (Anisopsidius; characters; Belgian Congo; Ruanda). — Herman, 1970: 408 (Anisopsidius). DISTRIBUTION: Congo, Rwanda.
Anisopsis [1 species; Ethiopian Region] Anisopsis Fauvel, 1904b: 108 (species included: flexuosa, carinata). TYPE SPECIES: Anisopsis flexuosa Fauvel, fixed by subsequent designation by R. Lucas, 1920: 98. — Bernhauer and Schubert, 1911: 109 (world catalog; 2 species). — Scheerpeltz, 1933: 1093 (world catalog supplement). — Blackwelder, 1952: 53 (type species). — Fagel, 1960: 4 (characters). — Herman, 1970: 406 (characters; distribution; notes; list of species). carinata Fauvel, 1904, see: Paroxytelopsis. eichelbaumi Bernhauer, 1927, see: Paroxytelopsis. flexuosa Fauvel, 1904b: 108 (Anisopsis; Type locality: Zambese). — Bernhauer and Schubert, 1911: 109 (Anisopsis; catalog). — Scheerpeltz, 1933: 1093 (Anisopsis; catalog). — Fagel, 1960: 6 (Anisopsis; characters; Zanzibar; Mashonaland). — Herman, 1970: 407 (Anisopsis). DISTRIBUTION: Zimbabwe, “Zambezi”, Zanzibar, Congo, Cameroon. rugosa Fagel, 1960: 7 (Anisopsis; subspecies of flexuosa; Type locality: Congo Belge: Bassin Lukuga. Also cited from Cameroon). guineensis Bernhauer, 1932, see: Paroxytelopsis. longicornis Cameron, 1928, see: Paroxytelopsis. minima Bernhauer, 1927, see: Paroxytelopsis. quadricollis Bernhauer, 1932, see: Anisopsidius. 1334 OXYTELINAE: ANISOPSIS Bulletin AMNH rugosa Fagel, 1960, see: flexuosa Fauvel, 1904. rugulipennis Bernhauer, 1934, see: Paroxytelopsis. tuberculata Cameron, 1938, see: Paroxytelopsis. uelensis Bernhauer, 1927, see: Paroxytelopsis.
Anotylus [356 species (1 doubtful); Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions] Anotylus Thomson, 1859: 44 (species included: sculpturatus). TYPE SPECIES: Oxytelus sculp- turatus Gravenhorst, fixed by monotypy and original designation. — Thomson, 1861: 130 (characters). — Gemminger and Harold, 1868: 648 (synonym of Oxytelus). — Fauvel, 1871a: 189 [= 1872: 163] (synonym of Oxytelus). — Fauvel, 1875a: X [= 1875b: 212] (synonym of Oxytelus). — Seidlitz, 1891: 354 (key to some European species). — Seidlitz, 1891a: 377 (key to some European species). — Heyden, Reitter, and Weise, 1891: 114 (subgenus of Oxytelus; list of species of Europe and Caucasus). — Heyden, 1893: 47 (subgenus of Oxytelus; list of species of Siberian region). — Heyden, 1896: 35 (subgenus of Oxytelus; list of species of Siberian region). — Heyden, Reitter, and Weise, 1906: 141 (subgenus of Oxytelus; list of species of Europe and Caucasus). — Bernhauer and Schubert, 1911: 109 (subgenus of Oxytelus). — Cameron, 1930: 212, 237 (subgenus of Oxytelus; characters in key; key to species of British India). — C. Koch, 1937a: 235 (key to Mediterranean and European species). — Bernhauer, 1939b: 70 (key to species of tetracarinatus group). — Blackwelder, 1943: 91 (subgenus of Oxytelus; type species nitidulus). — Steel, 1948: 188 (subgenus of Oxytelus; characters in key; type species: sculpturatus). — Tottenham, 1949: 363, 410 (type species: sculpturatus Gravenhorst; subgenus of Oxytelus; list of British species). — Blackwelder, 1952: 55 (type species: sculpturatus). — Tottenham, 1954: 46 (subgenus of Oxytelus; characters; key to British species). — Fagel, 1956b: 273 (valid genus; characters; type species: sculpturatus). — Hatch, 1957: 86, 95 (subgenus of Oxytelus; characters; key to species of Pacific Northwest). — Fagel, 1957a: 71 (Ruanda). — Coiffait and Saiz, 1968: 420 (Chilean species). — Herman, 1970: 414 (characters; distribution; notes; list of species). — Shibata, 1973c: 31 (subgenus of Oxytelus; checklist of species of Taiwan). — Tikhomirova, 1973a: 145 (subgenus of Oxytelus; checklist of species of USSR). — Bordoni, 1973j: 660 (list of species from islands near Sicily). — Moore and Legner, 1975: 222 (catalog; North America north of Mexico). — Hammond, 1975: 147, 156 (notes; characters; key to species of Ceylon). — Hammond, 1976 (review of genus; characters; notes on some species; taxonomic history; morphological variation; immature stages; biology; zoogeography; list of some adventitious species; list of fossil species). — Hammond, 1976a (revision and characters of crassicornis species group; key to and phylogeny of species of species group). — Shibata, 1976: 159 (checklist of species of Japan). — Pope, 1977: 25 (list of British species). — Burakowski, Mroczkowski, and Stefan´ska, 1979: 104 (catalog; subgenus of Oxytelus; Poland). — Moore and Legner, 1979: 215 (characters; notes). — Muona, 1979: 19 (list of Scandinavian species). — M. Dvorˇák, 1979: 110 (subgenus of Oxytelus; some species collected in Slovakia). Bulletin AMNH OXYTELINAE: ANOTYLUS 1335
— Uhlig and Vogler, 1981: 91 (list of some species collected in Germany; habitat and collecting notes). — Hammond, 1984: 200 (checklist of species of Borneo). — Ádám, 1985: 250 (list of species from southeast Hungary). — Yuh, Paik, Kwon, and Lee, 1985: 228 (checklist of species of Korea). — Frank, 1986a: 366 (checklist of species of Florida). — Muona and Viramo, 1986: 16 (list of species of northeastern Finland). — Segers, 1986: 26 (checklist of species; Belgium). — Ádám, 1987: 139 (list of six species with collecting notes; Hungary). — Schülke and Uhlig, 1988: 5 (characters). — Lohse and Lucht, 1989: 133 (valid genus). — M. Hansen, Kristensen, Mahler, Pedersen, 1991: 106 (list of Danish species). — Angelini, 1991: 190 (list of species from high elevations of Calabria, Italy). — Campbell and Davies, 1991: 95 (checklist of species; Canada). — Silfverberg, 1992: 23 (list of species of Fennoscandia, Denmark, and the Baltic States). — Navarrete-Heredia and Márquez-Luna, 1993 (cave collections; Mexico). — Welch, 1993: 227 (ovariole structure and development). — Bohácˇ, 1993: 43 (list of species of Czech Republic and Slovakia). — Hernández, Outerelo, and Gamarra, 1994: 186 (checklist of species; Canary Islands). — Ciceroni and Zanetti, 1995: 12 (list of species of Italy). — Terlutter, 1995: 17 (list of some species of Germany). — M. Hansen, 1996: 99 (list of Danish species). — M. Hansen, Mahler, Palm, and Pedersen, 1996: 241 (additions to list of Danish species). — Downie and Arnett, 1996: 447 (characters; key to species of Northeastern North America). — Czarniawski and Staniec, 1997: 37 (structure of opening of abdominal defensive reservoirs). — Lundgren, 1998: 47 (list of species from Florida; records for other States listed for each species). — M. Hansen, Palm, Pedersen, and Runge, 1998: 71 (list of species collected in 1997; Denmark). — Schülke, 1998i: 126 (characters). — Newton, Thayer, Ashe, and Chandler, 2000: 380 (18 Nearctic species; notes; characters in key). — Naomi, Kuranishi, Saito, and Maruyama, 2000: 107 (collected from Kuril Islands and Kam- chatka). — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 58 (necrophilous species in Mexico). Styloxys Gozis, 1886: 15 (species included: rugosus). TYPE SPECIES: Staphylinus rugosus Fabricius, fixed by monotypy. — Blackwelder, 1943: 91 (synonym of Oxytelus). — Steel, 1948: 188 (subgenus of Oxytelus; characters in key; type species: rugosus). — Tottenham, 1949: 363, 410 (cited as Styloxis; type species: rugosus Fabricius; subgenus of Oxytelus; list of British species). — Blackwelder, 1952: 365 (type species: rugosus). — Fagel, 1956b: 271 (valid genus; characters; type species: rugosus). — Lohse, 1964: 83 (subgenus of Oxytelus; cited as Styloxis; characters; key to central European species). — Herman, 1970: 414 (synonym of Anotylus). — Hammond, 1976: 145 (synonym of Anotylus). — Ádám, 1987: 139 (list of three species with collecting notes; Hungary). Oxytelodes Bernhauer, 1908d: 290 (species included: holdhausi). TYPE SPECIES: Oxytelodes holdhausi Bernhauer, fixed by monotypy. — Bernhauer and Schubert, 1911: 108 (world catalog; 1 species). — Blackwelder, 1944: 104 (checklist of species from Mexico, Central and South America, and the West Indies). — Blackwelder, 1952: 282 (type species: holdhausi). — Herman, 1970: 414 (synonym of Anotylus). — Hammond, 1976: 145 (synonym of Anotylus). 1336 OXYTELINAE: ANOTYLUS Bulletin AMNH
Emopotylus Bernhauer, 1910b: 359 (subgenus of Oxytelus; species included: cuernavacanus). TYPE SPECIES: Oxytelus (Emopotylus) cuernavacanus Bernhauer, fixed by monotypy. — Bernhauer and Schubert, 1911: 109 (subgenus of Oxytelus). — Blackwelder, 1943: 91 (subgenus of Oxytelus; type species cuernavacanus). — Steel, 1948: 188 (subgenus of Oxytelus; characters in key; type species: cuernavacanus). — Blackwelder, 1952: 147 (type species: cuernavacanus). — Herman, 1970: 414 (synonym of Anotylus). — Hammond, 1976: 145 (synonym of Anotylus). Boettcherinus Bernhauer, 1936b: 82 (species included: planaticollis, cornutus, militaris, bispinosus, bidentatus). TYPE SPECIES: Oxytelus (Boettcherinus) planaticollis Bernhauer, fixed by subsequent designation by Blackwelder, 1952: 79. — Steel, 1948: 188 (subgenus of Oxytelus; characters in key; type species: planaticollis). — Blackwelder, 1952: 79 (type species: planaticollis). — Herman, 1970: 414 (synonym of Anotylus). — Hammond, 1976: 145 (synonym of Anotylus). Oncoparia Bernhauer, 1936d: 214 (species included: parasita). TYPE SPECIES: Oncoparia parasita Bernhauer, fixed by monotypy. — Blackwelder, 1952: 275 (type species: parasita; notes concerning possible homonymy). — Fagel, 1965a: 338 (characters). — Herman, 1970: 421 (characters; distribution; notes; list of species). — Hammond, 1976: 145 (synonym of Anotylus). Paracaccoporus Steel, 1948: 188 (subgenus of Oxytelus; species included: ocularis). TYPE SPECIES: Oxytelus ocularis Fauvel, fixed by original designation and monotypy. — Herman, 1970: 414 (synonym of Anotylus). — Hammond, 1976: 145 (synonym of Anotylus). Oxytelosus Cameron, 1950c: 92 (species included: abnornalis “and no doubt” mirus). TYPE SPECIES: Oxytelus abnormalis Cameron, fixed by virtual monotypy. — Fagel, 1956b: 274 (valid genus; characters; type species: abnormalis). — Fagel, 1957a: 71 (Ruanda). — Herman, 1970: 414 (synonym of Anotylus; notes). — Hammond, 1976: 145 (synonym of Anotylus). Microxytelus Fagel, 1956b: 272 (species included: nitidifrons). TYPE SPECIES: Oxytelus nitidifrons Wollaston, fixed by original designation and monotypy. — Herman, 1970: 414 (synonym of Anotylus). — Hammond, 1976: 145 (synonym of Anotylus). Oxytelops Fagel, 1956b: 273 (species included: tetracarinatus). TYPE SPECIES: Staphylinus tetracarinatus Block, fixed by original designation and monotypy. — Fagel, 1957a: 71 (Ruanda). — Fagel, 1956b: 273 (Oxytelops). — Herman, 1970: 414 (synonym of Anotylus). — Bordoni, 1973j: 662 (list of species from islands near Sicily). — Hammond, 1976: 145 (synonym of Anotylus). — Burakowski, Mroczkowski, and Stefan´ska, 1979: 111 (catalog; Poland). Pseudodelopsis Fagel, 1957: 3 (species included: scotti, tolaensis, lungwensis). TYPE SPECIES: Pseudodelopsis scotti Fagel, fixed by original designation. — Herman, 1970: 414 (synonym of Anotylus). — Hammond, 1976: 145 (synonym of Anotylus). Anotylops Fagel, 1957: 8 (species included: seydeli). TYPE SPECIES: Anotylops seydeli Fagel, fixed by original designation and monotypy. — Herman, 1970: 423 (notes; list of species). — Hammond, 1976: 145 (synonym of Anotylus). Bulletin AMNH OXYTELINAE: ANOTYLUS 1337
Metoxytelus Coiffait and Saiz, 1968: 422 (species included: sulcicollis). TYPE SPECIES: Oxytelus sulcicollis Gemminger and Harold, fixed by original designation and mono- typy. — Hammond, 1976: 145 (synonym of Anotylus). Neoplatystethus Abdullah and Qadri, 1970: 127 (species included: hameedi, meccii). TYPE SPECIES: Neoplatystethus hameedi Abdullah and Qadri, fixed by original designation. — Hammond, 1976: 146 (synonym of Anotylus). Pseudoplatystethus Abdullah and Qadri, 1970: 129 (subgenus of Neoplatystethus; species included: meccii). TYPE SPECIES: Neoplatystethus (Pseudoplatystethus) meccii Abdullah and Qadri, fixed by original designation. — Hammond, 1976: 146 (synonym of Anotylus). Neopyctocraerus Abdullah and Qadri, 1970: 126 (species included: shafqati). TYPE SPECIES: Neopyctocraerus shafqati Abdullah and Qadri, fixed by original designation and monotypy. — Hammond, 1976: 146 (synonym of Anotylus). Pseudopyctocraerus Abdullah and Qadri, 1970: 125 (subgenus of Platystethus; species included: mahmoodi). TYPE SPECIES: Platystethus (Pseudopyctocraerus) mahmoodi Abdullah and Qadri, fixed by original designation. — Hammond, 1976: 146 (synonym of Anotylus). abnormalis Cameron, 1938e: 202 (Oxytelus; subgenus Anotylus; Type locality: Kenya Colony: Grotte Campbell, prairies alpines du mont Kenya, versant nord-est, alt. 3480 m). — Paulian, 1938: 205 (Oxytelus; larval description; Kenya). — Jeannel and Jarrige, 1949: 331 (Oxytelus; subgenus Anotylus; characters; cave species; Kenya). — Cameron, 1950c: 92 (Oxytelosus). — Fagel, 1956b: 274 (Oxytelosus; type species). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Kenya. aciculatus Bernhauer, 1936b: 84 (Oxytelus; Type locality: Mindanao: Surigao). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Philippines. advena Sharp, 1880, see: nitidifrons Wollaston, 1871. aegyptiacus C. Koch, 1934, see: plagiatus Rosenhauer, 1856. aeneotinctus Cameron, 1930: 215 (Oxytelus; Type locality: Dehra Dun). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Abdullah and Qadri, 1970: 120 (Oxytelus; characters; West Pakistan). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: India. aequicollis Bernhauer, 1936, see: testaceus Motschulsky, 1857. aethiops Bernhauer, 1931d: 567 (Oxytelus; subgenus Anotylus; Type locality: Abyssinia: Mt. Chillalo, ca. 9,000 feet). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Ethiopia. affinis Czwalina, 1870: 420 (Oxytelus; Type locality: Königsberg). — Fauvel, 1872: 176 [= 1873: 18] (Oxytelus; characters; Germany). — Redtenbacher, 1874: 255 (Oxytelus; characters). — Seidlitz, 1875: 248 (Oxytelus; characters; Baltic region). — J. Sahlberg, 1876: 177 (Anotylus; characters; Finland). — Eppelsheim, 1887b: 432 (Oxytelus; synonym of hamatus). — Ganglbauer, 1895: 644 (Oxytelus; synonym of hamatus). — Bernhauer and Schubert, 1911: 113 (Oxytelus; synonym of hamatus). 1338 OXYTELINAE: ANOTYLUS Bulletin AMNH
— Scheerpeltz, 1933: 1097 (Oxytelus; synonym of hamatus). — Scheerpeltz, 1962: 577 (Oxytelus; subgenus Anotylus; cited as valid species; characters; Europe; Mediterranean region). — Horion, 1963: 240 (Oxytelus; characters). — Lohse, 1964: 86 (Oxytelus; subgenus Anotylus; characters; central Europe). — Tóth, 1987: 227 (Anotylus; Carpathian Basin). — Ciceroni and Zanetti, 1995: 12 (Anotylus; Italy). — Kashcheev, 1999a: 57 (Oxytelus; subgenus Anotylus; cited as afinis; Kazakhstan). DISTRIBUTION: Europe. africanus Luze, 1904: 83 (Oxytelus; replacement name for flavipennis Eppelsheim). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — C. Koch, 1932: 652 (Oxytelus; subgenus Anotylus; characters in key). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Libya, Tunisia. flavipennis Eppelsheim, 1889b: 313 (Oxytelus; [preoccupied]; Type locality: Tunis, Monastir; Tripolis). — Fauvel, 1902b: 65 (Oxytelus; Tunisia; Tripolitania). — Luze, 1904: 83 (Oxytelus; synonym of africanus). — Bernhauer and Schubert, 1911: 110 (Oxytelus; synonym of africanus). alienus Cameron, 1943a: 339 (Oxytelus; subgenus Anotylus; Type locality: Australia: Port Darwin). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Australia. aliiceps Cameron, 1950a: 181 (Oxytelus; subgenus Anotylus; Type locality: Kenya Colony: Londiani, 7000 feet). — Herman, 1970: 417 (Anotylus). — Hammond, Morgan, and Morgan, 1979: 217 (Anotylus; gibbulus species group). DISTRIBUTION: Kenya, South Africa. transversefoveolatus Scheerpeltz, 1974: 69 (Oxytelus; subgenus Anotylus; Type locality: Cape Prov.: Groot River, 12 miles NE Plettenbergbaai). — Hammond, 1976: 161 (Anotylus; synonym of aliiceps). alpicola Casey, 1894: 394 (Oxytelus; Type locality: Colorado). — Fall and Cockerell, 1907: 168 (Oxytelus; New Mexico). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: USA. alternans Cameron, 1930: 253 (Oxytelus; subgenus Anotylus; Type locality: Mussoorie. Simla Hills: Theog, 6000 to 8000 feet). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Abdullah and Qadri, 1970: 121 (Oxytelus; characters; West Pakistan). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: India, Pakistan. americanus Mannerheim, 1830, see: insignitus Gravenhorst, 1806. amicus Bernhauer, 1923a: 122 (Oxytelus; Type locality: Korea: Pingan; Mandschurei: Chiau-tou). — Bernhauer, 1922c: 223 (Oxytelus; [Note: Unavailable name here, without description]; Formosa). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). — Yuh, Paik, Kwon, and Lee, 1985: 228 (Anotylus; Korea). — J. Li, 1993: 21 (Anotylus; China). DISTRIBUTION: Taiwan, Korea, China. Bulletin AMNH OXYTELINAE: ANOTYLUS 1339 andrewesi Cameron, 1930: 244 (Oxytelus; subgenus Anotylus; Type locality: Nilgiri Hills). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Scheerpeltz, 1962: 569 (Oxytelus; subgenus Anotylus; characters; India). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: India. angustatus Stephens, 1834, see: nitidulus Gravenhorst, 1802. antennarius Bernhauer, 1907: 377 (Oxytelus; Type locality: Yamanaka). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Japan. anticus Fauvel, 1904e: 304 (Oxytelus; Type locality: Diego-Suarez). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Madagascar. antipodum Bernhauer, 1908: 15 (Oxytelus; Type locality: Dongarra; Serpentine). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Australia. apicalis Fauvel, 1878g: 494 (Oxytelus; Type locality: Australie occidentale, Swan River). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Australia. arecae Cameron, 1929b: 443 (Oxytelus; subgenus Anotylus; Type locality: Penang). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Malaysia. armatifrons Sharp, 1887: 692 (Oxytelus; Type locality: Panama, Bugaba). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Panama. armatus Cameron, 1950, see: lippensi Bernhauer, 1943. armifrons Cameron, 1940b: 182 (Oxytelus; subgenus Emopotylus; Type locality: Hong Kong). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: China. asper Mulsant and Rey, 1878, see: nitidulus Gravenhorst, 1802. asperiventris Fauvel, 1903d: 260 (Oxytelus; Type locality: Nouméa; Tonghoué). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: New Caledonia. athenensis R. Dvorˇák, 1954d: 41 (Oxytelus; subgenus Anotylus; Type locality: Athènes). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Greece. atriceps Fauvel, 1904b: 98 (Oxytelus; Type locality: Celebes: Macassar; Patunuang; Samanga; Bua-Kraeng, 1700 m). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Steel, 1948: 186 (Oxytelus; characters). — Scheerpeltz, 1957c: 226 (Oxytelus; subgenus Anotylus; Sumba). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Indonesia. 1340 OXYTELINAE: ANOTYLUS Bulletin AMNH bacillus Bernhauer, 1915d: 101 (Oxytelus; Type locality: Deutsch-Ostafrika: Unjanjembe). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Tanzania. bakeri Bernhauer, 1915a: 21 (Oxytelus; subgenus Anotylus; Type locality: Philippinen: Luzon: M. Makiling; [Note: See lectotype designation by Hammond, 1976a]). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog). — Bernhauer, 1936b: 82 (Oxytelus; Philippines). — Herman, 1970: 417 (Anotylus). — Hammond, 1976a: 29 (Anotylus; lectotype designation; characters; Philippines). DISTRIBUTION: Philippines. bakeri Cameron, 1941, see: delopsoideus Herman, 1970. balbalanensis Bernhauer, 1936b: 83 (Oxytelus; Type locality: Luzon: Balbalan). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Philippines. barbiellinii Bernhauer, 1908d: 292 (Oxytelus; Type locality: Brasilien: S. Paulo). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Scheerpeltz, 1933: 1094 (Oxytelus; catalog; Argentina). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Argentina, Brazil. basalis Melsheimer, 1844, see: rugosus Fabricius, 1775. belgicus R. Dvorˇák, 1954, see: sculpturatus Gravenhorst, 1806. bernhaueri Ganglbauer, 1898: 400 (Oxytelus; Type locality: Bisamberg bei Wien). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Scheerpeltz, 1962: 577 (Oxytelus; subgenus Anotylus; characters; Europe). — Horion, 1963: 239 (Oxytelus; subgenus Oxytelops; Austria; Hungary; Czech Republic). — Lohse, 1964: 87 (Oxytelus; subgenus Anotylus; characters; central Europe). — Herman, 1970: 417 (Anotylus). — Tóth, 1987: 227 (Anotylus; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Poland; Czechoslovakia; Austria). — Bohácˇ, 1993: 43 (Anotylus; Czech Republic). — Assing, Frisch, Kahlen, et al., 1998: 128 (Anotylus; Germany). — Kashcheev, 1999a: 57 (Oxytelus; subgenus Anotylus; Kazakhstan). DISTRIBUTION: Europe. minarzi Bernhauer, 1936c: 185 (Oxytelus; subgenus Anotylus; Type locality: Hopfengarten in der unmittelbaren Umgebung von Horn). — Scheerpeltz, 1962: 577 (Oxytelus; synonym of bernhaueri). — Herman, 1970: 419 (Anotylus). — Hammond, 1976: 161 (Anotylus; synonym of bernhaueri). christianae Bernhauer, 1939b: 73 (Oxytelus; subgenus Anotylus; Type locality: Gars, Niederdonau). — Scheerpeltz, 1962: 577 (Oxytelus; synonym of bernhaueri). — Herman, 1970: 417 (Anotylus). — Hammond, 1976: 161 (Anotylus; synonym of bernhaueri). besucheti Hammond, 1975: 166 (Anotylus; Type locality: Ceylon: Central Prov.: Hasalaka, near Weragamtota, ca. 250 m). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Sri Lanka. bidentatus Bernhauer, 1936b: 87 (Oxytelus; Type locality: Luzon: Los Baños und Mount Banahao. Mindanao: Momungan und Port Banga). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Philippines. Bulletin AMNH OXYTELINAE: ANOTYLUS 1341 bigemmatus Fauvel, 1905e: 80 (Oxytelus; Type locality: Buitenzorg, Jardin botanique). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Cameron, 1928d: 101, 103 (Oxytelus; subgenus Anotylus; characters in key; Sumatra). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Indonesia. birmanus Cameron, 1930: 240 (Oxytelus; subgenus Anotylus; Type locality: Burma: Ruby Mines. Assam: Manipur). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Scheerpeltz, 1965: 101 (Oxytelus; subgenus Anotylus; Burma). — Abdullah and Qadri, 1970: 124 (Oxytelus; characters; East Pakistan). — Herman, 1970: 417 (Anotylus; [Note: Later workers cited this species in Oxytelus; I have continued to include it in Anotylus because that assignment was based on examination of the type]). — Scheerpeltz, 1976a: 80 (Oxytelus; Nepal). — Coiffait, 1983d: 162 (Oxytelus; Nepal). DISTRIBUTION: Nepal, India, Bangladesh, Myanmar. biroi Steel, 1956a: 64 (Oxytelus; subgenus Anotylus; Type locality: New Guinea: Astrolabe Bay: Friedrich-Wilhelmshafen). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: New Guinea. bispinosus Bernhauer, 1936b: 86 (Oxytelus; Type locality: Mindoro: San Teodoro). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Philippines. bisulcatus Petri, 1891, see: politus Erichson, 1840. boehmi Bernhauer, 1910, see: latiusculus Kraatz, 1859. bogorensis Herman, 2001: 41 (Anotylus; replacement name for longicornis Fauvel). DISTRIBUTION: India, Indonesia. longicornis Fauvel, 1905e: 80 (Oxytelus; [preoccupied]; Type locality: Buitenzorg). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Scheerpeltz, 1933: 1092 (Delopsis; catalog; India). — Herman, 1970: 418 (Anotylus). — Herman, 2001: 41 (Anotylus; synonym of bogorensis). bohemani Bernhauer and Schubert, 1911, see: caffer Erichson, 1840. borealis Motschulsky, 1860, see: nitidulus Gravenhorst, 1802. borneensis Cameron, 1933b: 340 (Delopsis; Type locality: British North Borneo: Kenokok, 3300 feet). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Borneo. borneensis Cameron, 1933: 341, see: tardus Herman, 1970. breviceps Casey, 1894: 385 (Oxytelus; Type locality: New York: Catskills Mts.). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). — Downie and Arnett, 1996: 448 (Anotylus; characters; USA). DISTRIBUTION: USA. brevipennis Fauvel, 1872b: 42 (Oxytelus; Type locality: Mastaganem; Tanger). — Fauvel, 1878: 95 [= 1878a: 15] (Oxytelus; Morocco; Algeria). — Fauvel, 1886: 22 [= 1886a: 14] (Oxytelus; Morocco; Algeria). — Fauvel, 1902b: 65 (Oxytelus; Morocco; Algeria). — Sainte-Claire Deville, 1906: 77 (Oxytelus; Corsica; Spain). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). 1342 OXYTELINAE: ANOTYLUS Bulletin AMNH
— Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Algeria, Morocco, Spain, Corsica. brincki Scheerpeltz, 1974: 67 (Oxytelus; subgenus Anotylus; Type locality: Natal: Royal Natal National Park, The Cascades, 5200 ft.). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: South Africa. bruchi Bernhauer, 1939h: 233 (Oxytelus; subgenus Anotylus; Type locality: Argentinien: Misiones: Loreto, Estacion Experimental). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Argentina. brunneipennis W. J. MacLeay, 1873: 150 (Oxytelus; Type locality: Gayndah). — Fauvel, 1877: 199 (Oxytelus; characters; Australia). — Lea, 1906: 211 (Oxytelus; characters; Australia). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). — Kuschel, 1990: 26 (Anotylus; introduced, Australian species; winged; habitat; New Zealand). DISTRIBUTION: Australia, New Zealand. crookesi Cameron, 1950: 23 (Oxytelus; subgenus Anotylus; Type locality: New Zealand: Auckland: Mt. Roskill). — Herman, 1970: 418 (Anotylus). — Hammond, 1976: 161 (Anotylus; synonym of brunneipennis). brunneus Bernhauer, 1932a: 81 (Oxytelus; Type locality: Watsa; Elisabethville). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Cameron, 1950g: 18 (Oxytelus; subgenus Anotylus; Belgian Congo). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Congo. bryanti Cameron, 1941a: 203 (Oxytelus; subgenus Anotylus; Type locality: W. Sarawak: Mt. Matang). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Malaysia. bubalus Fauvel, 1904b: 97 (Oxytelus; Type locality: Java: Batavia; Malang; Djatinagara; Palabuan; Bali). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Bernhauer, 1926f: 313 (Oxytelus; Buru). — Steel, 1948: 186 (Oxytelus; characters). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Indonesia. bururianus Cameron, 1956: 178 (Oxytelus; Type locality: Urundi: Bururi, 1800–2000 m). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Burundi. cadaverinus Cameron, 1942: 106 (Oxytelus; subgenus Emopotylus; Type locality: Darjeeling district: Ghum, Lepchajagat). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: India. caffer Erichson, 1840: 790 (Oxytelus; Type locality: Promont. Bonae Spei). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). — Scheerpeltz, 1974: 65 (Oxytelus; South Africa; Basutoland). DISTRIBUTION: South Africa, Lesotho, Tristan da Cunha. Bulletin AMNH OXYTELINAE: ANOTYLUS 1343
picipennis Boheman, 1848: 295 (Oxytelus; [preoccupied]; Type locality: Caffraria meridionalis). — Hammond, 1976: 161 (Anotylus; synonym of caffer). bohemani Bernhauer and Schubert, 1911: 110 (Oxytelus; replacement for picipennis Boheman). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 161 (Anotylus; synonym of caffer). christopherseni Brinck, 1948: 28 (Oxytelus; subgenus Anotylus; Type locality: Tristan da Cunha, Big Beach). — Herman, 1970: 417 (Anotylus). — Hammond, 1976: 162 (Anotylus; synonym of caffer). rudebecki Scheerpeltz, 1974: 66 (Oxytelus; subgenus Anotylus; Type locality: Cape Prov.: Buffelagsrivier, 4 miles E. Swellendam). — Hammond, 1976: 162 (Anotylus; synonym of caffer). cameroni Scheerpeltz, 1933: 1095 (Oxytelus; replacement name for occultus Cameron). — Scheerpeltz, 1962: 575 (Oxytelus; subgenus Anotylus; characters; Himalayan region). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: India, Pakistan. occultus Cameron, 1930: 245 (Oxytelus; subgenus Anotylus; [preoccupied]; Type locality: Dehra Dun). — Scheerpeltz, 1962: 575 (Oxytelus; synonym of cameroni). — Abdullah and Qadri, 1970: 121 (Oxytelus; characters; West Pakistan). cameroni Bernhauer, 1936, see: sceptus Herman, 1970. carbonellus Solsky, 1870, see: rugulosus Say, 1831. carinatus Panzer, 1798, see: rugosus Fabricius, 1775. carinipennis Cameron, 1951a: 48 (Oxytelus; subgenus Anotylus; Type locality: Angola: Serra do Moco, Luimbale, alt. 2300 m). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Angola. cavicola Bernhauer, 1926: 22 (Oxytelus; Type locality: Umgebung von Moulmein, Birma). — Cameron, 1930: 257 (Oxytelus; subgenus Anotylus; characters; Burma). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Myanmar. celebensis Fauvel, 1886, see: testaceus Motschulsky, 1857. cephalotes Eppelsheim, 1895c: 66 (Oxytelus; Type locality: Simla im Himalaya). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Cameron, 1930: 239 (Oxytelus; subgenus Anotylus; characters; India). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). — Abdullah and Qadri, 1970: 121 (Oxytelus; characters; West Pakistan). — Scheerpeltz, 1976: 20 (Oxytelus; Nepal). — Scheerpeltz, 1976a: 79 (Oxytelus; Nepal). — Hammond, 1976: 162 (Anotylus; synonyms). — Coiffait, 1984: 118 (Anotylus; Nepal). DISTRIBUTION: Nepal, India, Pakistan. tibialis Schubert, 1908: 624 (Oxytelus; [preoccupied]; Type locality: Kulu, Himalaya). — Bernhauer and Schubert, 1911: 121 (Oxytelus; catalog). — Cameron, 1930: 242 (Oxytelus; subgenus Anotylus; characters; Himalayan region). — Scheerpeltz, 1933: 1103 (Oxytelus; synonym of schuberti). 1344 OXYTELINAE: ANOTYLUS Bulletin AMNH
— Scheerpeltz, 1962: 568 (Oxytelus; synonym of schuberti). — Hammond, 1976: 162 (Anotylus; synonym of cephalotes). — Coiffait, 1984: 118 (Anotylus; cited as valid species; India; Pakistan). schuberti Scheerpeltz, 1933: 1103 (Oxytelus; replacement name for tibialis Schubert). — Scheerpeltz, 1962: 568 (Oxytelus; subgenus Anotylus; characters; Himalayan region). — Herman, 1970: 411 (Oxytelus). — Scheerpeltz, 1976: 20 (Oxytelus; Nepal). — Hammond, 1976: 162 (Anotylus; synonym of cephalotes). championi Fauvel, 1905a: 131 (Oxytelus; Type locality: Guatemala). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Guatemala. chapini Blackwelder, 1943, see: vinsoni Cameron, 1936. chilensis Coiffait and Saiz, 1968, see: complanatus Erichson, 1839. chinkiangensis Bernhauer, 1938: 23 (Oxytelus; subgenus Anotylus; Type locality: Nordwestl. China: Chinkiang). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: China. christianae Bernhauer, 1939, see: bernhaueri Ganglbauer, 1898. christopherseni Brinck, 1948, see: caffer Erichson, 1840. clavatus A. Strand, 1946: 171 (Oxytelus; Type locality: Norway: AK 11: Brønnøya). — Palm, 1961a: 35 (Oxytelus; characters; Norway). — Palm, 1963a: 234 (Oxytelus; subgenus Anotylus; characters; Sweden). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Norway, Sweden. clavicornis Fauvel, 1878f: 219 (Oxytelus; Type locality: Nouvelle-Guinée, Andai). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: New Guinea. clypeonitens Pandellé, 1867a: 171 (Oxytelus; Type locality: Saint-Germain en Laye). — Fauvel, 1872: 174 [= 1873: 16] (Oxytelus; characters; France; Britain; Italy; Sardinia). — Redtenbacher, 1874: 255 (Oxytelus; characters). — Fauvel, 1876a: 57 [= 1876: 242] (Oxytelus; Germany; Spain). — Mulsant and Rey, 1878: 534 (Oxytelus; subgenus Anotylus; characters; France). — Fauvel, 1886: 23 [= 1886a: 15] (Oxytelus; Morocco). — Fowler, 1888: 382 (Oxytelus; characters; Britain). — Ganglbauer, 1895: 643 (Oxytelus; subgenus Anotylus; characters; middle Europe; Italy; Sardinia). — Reitter, 1909: 172 (Oxytelus; subgenus Anotylus; characters; Germany). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Portevin, 1929: 418 (Oxytelus; characters; France). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Scheerpeltz, 1956c: 1094 (Oxytelus; subgenus Anotylus; Cyprus). — Scheerpeltz, 1961e: 135 (Oxytelus; subgenus Anotylus; Rhodes). — Scheerpeltz, 1962: 571 (Oxytelus; subgenus Anotylus; characters; Europe; Mediterranean region; Turkey; Caucasus). — Horion, 1963: 234 (Oxytelus; subgenus Oxytelops; Syria; Turkey; Portugal; Spain; Italy; Dalmatia; Corfu; Bosnia Hercegovina; Montenegro; Serbia; Macedonia; Albania; Greece; Romania; Hungary; Czech Republic; Slovakia; Poland; Russia; France; Switzerland; Belgium; Holland; Britain; Austria; Germany). — Lohse, 1964: 86 (Oxytelus; subgenus Anotylus; characters; central Europe). — Smetana, 1967e: 318 (Oxytelus; subgenus Anotylus; Turkey). — Szujecki, 1968a: 715 (Oxytelus; Poland). Bulletin AMNH OXYTELINAE: ANOTYLUS 1345
— Herman, 1970: 417 (Anotylus). — Coiffait, 1970e: 64 (Oxytelus; Cyprus). — Pope, 1977: 25 (Anotylus; Britain). — Coiffait, 1978d: 166 (Oxytelus; Turkey). — Tóth, 1987: 227 (Anotylus; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark). — Schülke and Uhlig, 1988: 5 (Anotylus; notes; Germany). — Bohácˇ, 1993: 43 (Anotylus; Czech Republic; Slovakia). — Ciceroni and Zanetti, 1995: 12 (Anotylus; Italy). — Dauphin, Laguerre, Tamisier, and Teissier, 1997: 20 (Oxytelus; France). — Assing and Wunderle, 2001: 36 (Anotylus; Cyprus). DISTRIBUTION: Europe, Turkey, Caucasus. cognatus Sharp, 1874: 94 (Oxytelus; Type locality: Nagasaki; Hiogo). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Scheerpeltz, 1929a: 115 (Oxytelus; subgenus Anotylus; notes; China). — Cameron, 1949b: 461 (Oxytelus; Japan). — Nakane, 1963a: 84 (Oxytelus; characters; habitus photograph; Japan). — Watanabe and Shibata, 1965: 317 (Oxytelus; subgenus Anotylus; Japan). — Herman, 1970: 417 (Anotylus). — Yuh, Paik, Kwon, and Lee, 1985: 228 (Anotylus; Korea). — J. Li, 1993: 21 (Anotylus; China). — Watanabe, 1996a: 9 (Anotylus; cave collection; Japan). DISTRIBUTION: Japan, Korea, China. comorensis Lecoq, 1996: 132 (Anotylus; Type locality: l’archipel des Comores: Anjouan, Boungoueni). DISTRIBUTION: Comoros. complanatus Erichson, 1839a: 595 (Oxytelus; Type locality: Mark Brandenburg). — Gyllenhal, 1810: 457 (Oxytelus; [Note: Misidentification: Bernhauer and Schubert, 1911: 111 cited depressus Gyllenhal as a synonym of complanatus Erichson; Gyllenhal did not describe the species as new, he attributed it to Gravenhorst who attributed it to Olivier who attributed it to Paykull]; characters; Sweden). — Heer, 1839: 206 (Oxytelus; characters; Switzerland). — Erichson, 1840: 795 (Oxytelus; characters; Europe). — Redtenbacher, 1849: 738 (Oxytelus; characters; Austria). — Kiesenwetter, 1851a: 427 (Oxytelus; France; Spain). — Hardy, 1851: 52 (Oxytelus; synonym of pallipes). — Wollaston, 1854: 608 (Oxytelus; characters; notes; Madeira Islands). — Rosenhauer, 1856: 82 (Oxytelus; Cadiz). — Fairmaire and Laboulbène, 1856: 612 (Oxytelus; characters; France). — Wollaston, 1857: 200 (Oxytelus; habitat; Madeira Islands). — Redtenbacher, 1857: 233 (Oxytelus; characters; Austria). — Kraatz, 1857d: 858 (Oxytelus; characters; Germany). — Kraatz, 1858b: 127 (Oxytelus; Greece). — Thomson, 1861: 132 (Anotylus; characters; Scandinavia). — Hochhuth, 1862: 92 (Oxytelus; Russia). — Wollaston, 1864: 597 (Oxytelus; notes; Canary Islands). — Fauvel, 1865a: 311 [= 1865b: 61] (Oxytelus; habitat; France). — Wollaston, 1865: 517 (Oxytelus; notes; Madeira Islands; Canary Islands). — Crotch, 1867a: 384 (Oxytelus; Azores). — Pandellé, 1867a: 172 (Oxytelus; characters; France). — Fauvel, 1872: 172 [= 1873: 14] (Oxytelus; characters; France; Europe; Madeira; North Africa; Cyprus; Caramanie; Iran). — Redtenbacher, 1874: 254 (Oxytelus; characters; Austria). — Seidlitz, 1875: 247 (Oxytelus; characters; Baltic region). — J. Sahlberg, 1876: 177 (Anotylus; Finland). 1346 OXYTELINAE: ANOTYLUS Bulletin AMNH
— Fauvel, 1878: 95 [= 1878a: 15] (Oxytelus; Barbary). — Mulsant and Rey, 1878: 527 (Oxytelus; subgenus Anotylus; characters; larval characters; France). — Fauvel, 1886: 22 [= 1886a: 14] (Oxytelus; North Africa). — Fowler, 1888: 381 (Oxytelus; characters; Britain). — Ganglbauer, 1895: 642 (Oxytelus; subgenus Anotylus; characters; north and middle Europe; Mediterranean region). — Fauvel, 1900c: 185 (Oxytelus; New Zealand). — Fauvel, 1902b: 66 (Oxytelus; Barbary; Azores; Madeira; Canary Islands). — Poppius, 1905: 11 (Oxytelus; Shetland Islands). — Sainte-Claire Deville, 1906: 77 (Oxytelus; Corsica). — Reitter, 1909: 171 (Oxytelus; subgenus Anotylus; characters; Germany). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Johansen, 1914: 563 (Oxytelus; subgenus Anotylus; characters; Denmark). — Portevin, 1929: 418 (Oxytelus; characters; France). — Gridelli, 1930: 64 (Oxytelus; Libya). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog; Argentina). — Bernhauer, 1940c: 2 (Oxytelus; Azores; Madeira). — Méquignon, 1942a: 17 (Oxytelus; subgenus Anotylus; Azores). — Coiffait, 1954: 162 (Oxytelus; Canary Islands). — Scheerpeltz, 1956c: 1093 (Oxytelus; subgenus Anotylus; Cyprus). — Scheerpeltz, 1958a: 8 (Oxytelus; subgenus Anotylus; Turkey). — Scheerpeltz, 1960d: 133 (Oxytelus; subgenus Anotylus; Portugal). — Scheerpeltz, 1961d: 119 (Oxytelus; subgenus Anotylus; Italy). — Scheerpeltz, 1961e: 135 (Oxytelus; subgenus Anotylus; Rhodes). — Palm, 1961a: 34 (Oxytelus; characters; Sweden; Denmark; Norway). — Scheerpeltz, 1962: 568 (Oxytelus; subgenus Anotylus; characters; Europe; Mediterranean region; Atlantic Islands; Siberia; Turkey; Caucasus; Turkestan; Afghanistan; Iran). — Smetana, 1963b: 32 (Oxytelus; subgenus Anotylus; Madeira). — Horion, 1963: 232 (Oxytelus; subgenus Anotylus; Madeira; Azores; Canaries; Turkey; Mesopotamia; Turkestan; Iran; Afghanistan; France to Russia; Britain; Norway; Sweden). — Lohse, 1964: 86 (Oxytelus; subgenus Anotylus; characters; central Europe). — Scheerpeltz, 1964: 27 (Oxytelus; subgenus Anotylus; Mongolia). — Scheerpeltz, 1965b: 394 (Oxytelus; subgenus Anotylus; Sicily). — Szujecki, 1968a: 715 (Oxytelus; Poland). — Herman, 1970: 417 (Anotylus). — Smetana, 1970: 54 (Oxytelus; distributional summary; Azores). — Bacchus and Hammond, 1972: 62 (Anotylus; lion dung; Britain). — Hammond, 1976: 174 (Anotylus; notes; adventive species). — Pope, 1977: 25 (Anotylus; Britain). — Frank, 1982: 9 (Anotylus; list of parasites). — Tóth, 1987: 227 (Anotylus; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden). — Nowosad, 1990: 60 (Oxytelus; collected from mole nest; notes; Poland). — Borges, 1990: Table II (Anotylus; Azores). — Kuschel, 1990: 26 (Anotylus; introduced, Palaearctic species; winged; habitat; New Zealand). — Welch, 1993: 227 (Anotylus; ovariole number and ovary structure). — Bohácˇ, 1993: 43 (Anotylus; Czech Republic; Slovakia). — Ciceroni and Zanetti, 1995: 12 (Anotylus; Italy). — Assing and Wunderle, 1999: 437 (Anotylus; Canary Islands). — Kashcheev, 1999a: 57 (Oxytelus; subgenus Anotylus; Kazakhstan). — Assing and Wunderle, 2001: 36 (Anotylus; Cyprus). DISTRIBUTION: Shetland Islands, Europe, Azores, Madeira Islands, Canary Islands, Libya, Turkey, Caucasus, Russia, Iran, Afghanistan, “Turkestan”, Mongolia, New Zealand, Argentina, Chile. Bulletin AMNH OXYTELINAE: ANOTYLUS 1347
pallipes Stephens, 1834: 319 (Oxytelus; [preoccupied]; Type locality: New Forest; near Bristol; Suffolk; Norfolk; Devonshire). — Stephens, 1839: 420 (Oxytelus; characters; England). — Hardy, 1851: 52 (Oxytelus; notes; England). — G. Waterhouse, 1858: 30 (Oxytelus; synonym of complanatus). — Ganglbauer, 1895: 642 (Oxytelus; synonym of complanatus). — Bernhauer and Schubert, 1911: 111 (Oxytelus; synonym of complanatus). — Scheerpeltz, 1962: 568 (Oxytelus; synonym of complanatus). ixellensis R. Dvorˇák, 1954d: 41 (Oxytelus; subgenus Anotylus; Type locality: Belgique: Ixelles). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 162 (Anotylus; synonym of complanatus). chilensis Coiffait and Saiz, 1968: 420 (Anotylus; Type locality: Chili: Santiago). — Hammond, 1976: 162 (Anotylus; synonym of complanatus). consanguineus Cameron, 1934: 18 (Delopsis; Type locality: Nilgiri Hills: Pykara, altitude, 7000 feet). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: India. contiguus Cameron, 1942: 107 (Oxytelus; subgenus Anotylus; Type locality: Kashmir: Gulmarg). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: India. corcyranus Coiffait, 1968b: 95 (Anotylus; subgenus Anotylus; Type locality: Corfou: Route de Kerkyra à Paléokastritsa, à 15 km de Kerkyra). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Corfu. coriaceus Fauvel, 1889b: 250 (Oxytelus; Type locality: Tonghoué). — Fauvel, 1903d: 260 (Oxytelus; New Caledonia). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: New Caledonia. cornutus Bernhauer, 1936b: 86 (Oxytelus; [preoccupied]; Type locality: Mindoro: San Teodoro. Luzon: Los Baños). — Herman, 1970: 417 (Anotylus). — Herman, 2001: 41 (Anotylus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5). DISTRIBUTION: Philippines. coussementi R. Dvorˇák, 1954 [Note: Nomen nudum], see: sculpturatus Gravenhorst, 1806. crassicornis Sharp, 1874: 92 (Oxytelus; Type locality: Japan). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Bernhauer, 1922c: 223 (Oxytelus; Formosa). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Cameron, 1949b: 461 (Oxytelus; Japan). — Watanabe, 1961: 349 (Oxytelus; Izu Islands). — Nakane, 1963a: 84 (Oxytelus; characters; habitus photograph; Japan). — Herman, 1970: 417 (Anotylus). — Hammond, 1976a: 26 (Anotylus; characters; Japan; China; Taiwan). — Yuh, Paik, Kwon, and Lee, 1985: 228 (Anotylus; Korea). DISTRIBUTION: Korea, Japan, Taiwan. crebratus Schubert, 1906: 377 (Oxytelus; subgenus Anotylus; Type locality: Kwai, Usambara). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Tanzania. 1348 OXYTELINAE: ANOTYLUS Bulletin AMNH crenaticollis Fauvel, 1908, see: dilutipennis Fauvel, 1904. crenulatus Broun, 1880, see: rugosus Fabricius, 1775. crenulicollis Bernhauer, 1934, see: stanleyi Cameron, 1933. cribriceps Fauvel, 1878g: 496 (Oxytelus; Type locality: Australie méridionale, Melbourne). — Bernhauer, 1908: 15 (Oxytelus; Australia). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Herman, 1970: 417 (Anotylus). DISTRIBUTION: Australia. cribrum Fauvel, 1905a: 114 (Oxytelus; Type locality: Sikkim: Darjeeling; Bengale: Kurseong). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Cameron, 1930: 214 (Oxytelus; characters; India; Burma; Bengal). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Scheerpeltz, 1965: 100 (Oxytelus; subgenus Oxytelus; Burma). — Abdullah and Qadri, 1970: 120, 124 (Oxytelus; characters; East and West Pakistan). — Herman, 1970: 418 (Anotylus; [Note: This assignment was based on examination of syntypes]). — Scheerpeltz, 1976: 20 (Oxytelus; Nepal). — Coiffait, 1983: 276 (Oxytelus; Kashmir). — Coiffait, 1984: 118 (Oxytelus; Pakistan). — Coiffait, 1984e: 374 (Oxytelus; Nepal). DISTRIBUTION: Pakistan, India, Nepal, Bangladesh, Myanmar. crookesi Cameron, 1950, see: brunneipennis W. J. MacLeay, 1873. cuernavacanus Bernhauer, 1910b: 359 (Oxytelus; subgenus Emopotylus; Type locality: Mexiko: Cuernavaca). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Mexico. currus Herman, 1970: 418 (Anotylus; replacement name for monticola Cameron, 1942 [July]). DISTRIBUTION: India. monticola Cameron, 1942 (July): 107 (Oxytelus; subgenus Anotylus; [preoccupied]; Type locality: Darjeeling district: Ghum, Tiger Hill, altitude 8500–10,000 feet). — Herman, 1970: 418 (Anotylus; synonym of currus). curticornis Cameron, 1941c: 492 (Delopsis; Type locality: Philippines: Luzon: Ballalan). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Philippines. curtipennis Lecoq, 1996: 134 (Anotylus; Type locality: l’archipel des Comores: Mohéli, Miringoni). DISTRIBUTION: Comoros. curtus Bernhauer, 1904, see: wattsensis Blackburn, 1902. darwini Cameron, 1943a: 339 (Oxytelus; subgenus Anotylus; Type locality: Australia: Port Darwin). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Australia. darwinianus Cameron, 1943a: 340 (Oxytelus; subgenus Anotylus; Type locality: N. Australia: Port Darwin). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Australia. deceptor Cameron, 1933b: 341 (Oxytelus; subgenus Anotylus; Type locality: British North Borneo: Lumu Lumu, 5500 feet). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Borneo. Bulletin AMNH OXYTELINAE: ANOTYLUS 1349 decipiens Cameron, 1941a: 203 (Oxytelus; subgenus Anotylus; Type locality: W. Sarawak: Mt. Matang). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Malaysia. delicatus Cameron, 1943a: 340 (Oxytelus; subgenus Anotylus; Type locality: Australia: Port Darwin; Timor: Dilli). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Australia, Indonesia. delopsoideus Herman, 1970: 418 (Anotylus; replacement name for bakeri Cameron). DISTRIBUTION: Philippines. bakeri Cameron, 1941c: 491 (Delopsis; [preoccupied]; Type locality: Philippines: Luzon: Mt. Makiling. Panaon). — Herman, 1970: 418 (Anotylus; synonym of delopsoideus). densus Casey, 1894: 393 (Oxytelus; Type locality: Maryland; Texas). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — Downie and Arnett, 1996: 448 (Anotylus; characters; USA). DISTRIBUTION: USA. denticollis Wendeler, 1955: 193 (Oxytelodes ?; Type locality: Brasilien: Nova Teutonia). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Brazil. dentifer Fauvel, 1878f: 218 (Oxytelus; Type locality: Nouvelle-Guinée, Andai). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: New Guinea. dentifrons Fauvel, 1904b: 107 (Oxytelus; Type locality: Amazone: Pebas. Antilles: Saint-Vincent). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Blackwelder, 1943: 95 (Oxytelus; characters; St. Vincent). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Brazil, West Indies. dilutipennis Fauvel, 1904b: 101 (Oxytelus; Type locality: Libreville, Gabon; [Note: See lectotype designation by Hammond, 1976a]. Also cited from Congo: Kinchassa. Kilimandjaro). — Fauvel, 1907: 16 (Oxytelus; Kilimandjaro; Ethiopia; Sierra Leone; Gabon; Congo; Portuguese Guinea). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Cameron, 1950g: 8 (Oxytelus; Belgian Congo). — Herman, 1970: 418 (Anotylus). — Hammond, 1976a: 30 (Anotylus; lectotype designation: Libreville, Gabon; characters; Gabon; Tanzania; Zaire; Ghana; Nigeria; Cameroon; Rwanda; Ethiopia; Uganda; Angola; Rhodesia; South Africa). DISTRIBUTION: Guinea-Bissau, Ghana, Nigeria, Cameroon, Gabon, Congo, Uganda Ethiopia, Rwanda, Tanzania, Zimbabwe South Africa, Angola. crenaticollis Fauvel, 1908: 96 (Oxytelus; Type locality: Amani, Tanzania; [Note: See lecto- type designation by Hammond, 1976a]. Also cited from Abyssinia). — Bernhauer and Schubert, 1911: 111 (Oxytelus; catalog). — Eichelbaum, 1913: 118 (Oxytelus; Tanzania). — Scheerpeltz, 1933: 1095 (Oxytelus; catalog). — Bernhauer, 1942c: 350 (Oxytelus; Cameroon). — Cameron, 1950g: 9 (Oxytelus; subgenus Tanycraerus; Belgian Congo). — Herman, 1970: 417 (Anotylus). — Hammond, 1976a: 30 (Anotylus; lectotype designation: Amani, Tanzania; synonym of diluti- pennis). 1350 OXYTELINAE: ANOTYLUS Bulletin AMNH
subplagiatus Cameron, 1928: 16 (Oxytelus; Type locality: Zaire: Kivu, Burunga; [Note: See lectotype designation by Hammond, 1976a]). — Scheerpeltz, 1933: 1104 (Oxytelus; catalog). — Herman, 1970: 421 (Anotylus). — Hammond, 1976a: 30 (Anotylus; lectotype designation: Zaire: Kivu, Burunga; synonym of dilutipennis). discipennis Fauvel, 1877: 197 (Oxytelus; Type locality: Australie, Nouvelle Galles du Sud; Tasmanie). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Australia. dispar Lea, 1906: 209 (Oxytelus; Type locality: Hobart, Tasm.). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Australia. disparatus Cameron, 1928d: 103, 104 (Oxytelus; subgenus Anotylus; Type locality: Sumatra: Fort de Kock; also in the Malay Peninsula). — Cameron, 1929b: 444 (Oxytelus; characters in key). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — Hammond, 1975: 158, 161 (Anotylus; notes; characters; Ceylon; India; Nepal; Malaya; Sumatra; Philippines). DISTRIBUTION: Pakistan, China, Nepal, India, Sri Lanka, Malaysia, Indonesia, Philippines. intermedius Cameron, 1930: 249 (Oxytelus; subgenus Anotylus; Type locality: Dehra Dun. Mussoorie). — Scheerpeltz, 1933: 1098 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — Abdullah and Qadri, 1970: 121 (Oxytelus; characters; West Pakistan). — Hammond, 1975: 161 (Anotylus; synonym of disparatus). pauper Cameron, 1930: 249 (Oxytelus; subgenus Anotylus; Type locality: Mussoorie: Arni Gad. Dehra Dun. Ceylon: Colombo; Belligam). — Scheerpeltz, 1933: 1101 (Oxytelus; catalog). — Scheerpeltz, 1962: 573 (Oxytelus; subgenus Anotylus; characters; China; Himalayan region; India; Ceylon). — Herman, 1970: 419 (Anotylus). — Abdullah and Qadri, 1970: 121 (Oxytelus; cited twice, once as paupen; characters; West Pakistan). — Hammond, 1975: 161 (Anotylus; synonym of disparatus). philippinus Cameron, 1946: 10 (Oxytelus; subgenus Anotylus; Type locality: Philippines: Palawan: Binaluan; Luzon: Mt. Makiling). — Herman, 1970: 419 (Anotylus). — Hammond, 1975: 161 (Anotylus; synonym of disparatus). distincticollis Cameron, 1919a: 226 (Oxytelus; subgenus Anotylus; Type locality: Ceylon: Bogawantalawa). — Cameron, 1930: 258 (Oxytelus; subgenus Anotylus; characters; Ceylon). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — Hammond, 1975: 168 (Anotylus; notes; characters; Ceylon). DISTRIBUTION: Sri Lanka. drescheri Cameron, 1936a: 32 (Oxytelus; subgenus Oxytelus; Type locality: Java: G. Tangkoeban Prahoe, Preanger, alt. 4000–5000 ft). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Indonesia. Bulletin AMNH OXYTELINAE: ANOTYLUS 1351 ebonus Blackwelder, 1944: 105 (Oxytelus; replacement name for nitens Fauvel). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Argentina, Uruguay. nitens Fauvel, 1877: xxiv (Oxytelus; [preoccupied]; Type locality: La Plata; Montevideo, Pampas). — Lynch, 1884: 366 (Oxytelus; characters; Argentina). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Scheerpeltz, 1933: 1100 (Oxytelus; catalog). — Blackwelder, 1944: 105 (Oxytelus; synonym of ebonus). elegans Meinert, 1890, see: rugifrons Hochhuth, 1849. elephantis Bernhauer, 1936h: 322 (Oxytelus; subgenus Tanycraerus; Type locality: Kenya Colony: Katamayu R.). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Kenya. emeritus Sharp, 1887: 691 (Oxytelus; Type locality: Guatemala, San Joaquin, San Geronimo). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Guatemala. eppelsheimii Bethe, 1867, see: rugifrons Hochhuth, 1849. exaratus Sharp, 1887: 693 (Oxytelus; Type locality: Guatemala, Balheu in Vera Paz 3000 feet). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Guatemala. exasperatus Kraatz, 1859: 175 (Oxytelus; Type locality: Ceylan). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Cameron, 1921a: 366, 400 (Oxytelus; characters; Singapore). — Cameron, 1930: 261 (Oxytelus; subgenus Anotylus; characters; Ceylon). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Cameron, 1934d: 78 (Oxytelus; synonym of parasitus). — Herman, 1970: 418 (Anotylus). — Hammond, 1975: 158, 162 (Anotylus; notes; characters; Ceylon; Madagascar). DISTRIBUTION: Singapore, Sri Lanka, Madagascar. parasitus Motschulsky, 1860a: 69 (Oxytelus; Type locality: montagnes de Noura-Ellia, Ceylan). — Bernhauer and Schubert, 1911: 117 (Oxytelus; catalog). — Cameron, 1930: 263 (Oxytelus; characters; Ceylon). — Scheerpeltz, 1933: 1101 (Oxytelus; catalog). — Cameron, 1934d: 78 (Oxytelus; notes on type). — Herman, 1970: 410 (Oxytelus). — Hammond, 1975: 162 (Anotylus; synonym of exasperatus). excavatus Motschulsky, 1857, see: inustus Gravenhorst, 1806. excellens Luze, 1904, see: syriacus Eppelsheim, 1881. exiguus Erichson, 1840: 798 (Oxytelus; Type locality: America septentrionalis). — LeConte, 1877: 234, 238 (Oxytelus; characters; middle and southern States). — Casey, 1894: 397 (Oxytelus; characters; New York to Florida; Lake Superior and Texas). — Blatchley, 1910: 469 (Oxytelus; characters; Indiana). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). — Downie and Arnett, 1996: 447 (Oxytelus; characters; USA). DISTRIBUTION: USA. 1352 OXYTELINAE: ANOTYLUS Bulletin AMNH
pygmaeus Melsheimer, 1844: 41 (Oxytelus; Type locality: Pennsylvania). — Casey, 1894: 397 (Oxytelus; synonym of exiguus). — Bernhauer and Schubert, 1911: 112 (Oxytelus; synonym of exiguus). parvulus Melsheimer, 1844: 41 (Oxytelus; Type locality: Pennsylvania). — Casey, 1894: 397 (Oxytelus; synonym of exiguus). — Bernhauer and Schubert, 1911: 112 (Oxytelus; synonym of exiguus). extensicornis Fauvel, 1905e: 79 (Oxytelus; Type locality: Buitenzorg, Jardin botanique). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Cameron, 1928d: 100 (Oxytelus; subgenus Anotylus; Sumatra). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Indonesia. externus Sharp, 1887: 694 (Oxytelus; Type locality: Panama, Bugaba). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Panama. fageli Herman, 1970: 418 (Anotylus; replacement name for scotti Fagel). DISTRIBUTION: Ethiopia. scotti Fagel, 1957: 11 (Anotylus; [preoccupied]; Type locality: Ethiopia: Simien, Arghine, 11,500 ft or higher). fageli Jarrige, 1970, see: jarrigei Hammond, 1976. fairmairei Pandellé, 1867: 172 (Oxytelus; Type locality: H.-Pyrénées). — Kraatz, 1870a: 424 (Oxytelus; characters; Pyrenees). — Fauvel, 1872: 175 [= 1873: 17] (Oxytelus; characters; France; Britain; Germany; Austria). — Mulsant and Rey, 1878: 539 (Oxytelus; subgenus Anotylus; characters; France). — Fowler, 1888: 383 (Oxytelus; characters; Britain). — Ganglbauer, 1895: 644 (Oxytelus; subgenus Anotylus; characters; middle Europe). — Reitter, 1909: 172 (Oxytelus; subgenus Anotylus; characters; Germany). — Bernhauer and Schubert, 1911: 112 (Oxytelus; catalog). — Johansen, 1914: 565 (Oxytelus; subgenus Anotylus; characters; Denmark). — Portevin, 1929: 419 (Oxytelus; characters; France). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Palm, 1961a: 37 (Oxytelus; characters; Sweden; Denmark; Norway; Finland). — Scheerpeltz, 1962: 574 (Oxytelus; subgenus Anotylus; characters; Europe). — Horion, 1963: 236 (Oxytelus; subgenus Oxytelops; Britain; Norway; Sweden; Finland; Russia; Denmark; Holland; Belgium; France; Italy; Slovenia; Slovakia; Czech Republic; Russia; Germany; Austria). — Lohse, 1964: 87 (Oxytelus; subgenus Anotylus; characters; central Europe). — Smetana, 1968: 235 (Oxytelus; subgenus Anotylus; Mongolia). — Herman, 1970: 418 (Anotylus). — Smetana 1975c: 167 (Anotylus; Mongolia). — Pope, 1977: 25 (Anotylus; Britain). — Tóth, 1987: 228, 229, 230 (Anotylus; characters; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden). — Kashcheev, 1989: 287 (Oxytelus; subgenus Anotylus; Mongolia). — Bohácˇ, 1993: 43 (Anotylus; Czech Republic; Slovakia). — Ciceroni and Zanetti, 1995: 12 (Anotylus; Italy). DISTRIBUTION: Europe, Mongolia. transversalis Czwalina, 1870: 419 (Oxytelus; Type locality: Krain; Königsberg). — Fauvel, 1872: 175 [= 1873: 17] (Oxytelus; synonym of fairmairei). — Redtenbacher, 1874: 255 (Oxytelus; characters). — Seidlitz, 1875: 248 (Oxytelus; characters; Baltic region). — Mulsant and Rey, 1878: 539 (Oxytelus; synonym of fairmairei). Bulletin AMNH OXYTELINAE: ANOTYLUS 1353
— Ganglbauer, 1895: 644 (Oxytelus; synonym of fairmairei). — Bernhauer and Schubert, 1911: 112 (Oxytelus; synonym of fairmairei). — Scheerpeltz, 1933: 1096 (Oxytelus; synonym of fairmairei). — Scheerpeltz, 1962: 574 (Oxytelus; synonym of fairmairei). falsus Cameron, 1942: 106 (Oxytelus; subgenus Emopotylus; Type locality: Darjeeling district: Ghum). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: India. flavior Blackburn, 1902, see: impressifrons W. J. MacLeay, 1873. flavipennis Kraatz, 1859: 172 (Oxytelus; Type locality: India orientalis; [Note: See lectotype designation by Hammond, 1976a]). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Cameron, 1930: 215 (Oxytelus; characters; East Indies). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). — Hammond, 1976a: 27 (Anotylus; lectotype designation; characters; East Indies). DISTRIBUTION: “East Indies”. flavipennis Eppelsheim, 1889, see: africanus Luze, 1904. flavipes Stephens, 1834, see: nitidulus Gravenhorst, 1802. flavipes Lacordaire, 1835, see: sculpturatus Gravenhorst, 1806. flavipes Fauvel, 1878, see: serus Herman, 1970. foetidus Cameron, 1930, see: micans Kraatz, 1859. forsteri Scheerpeltz, 1960c: 72 (Oxytelus; subgenus Anotylus; Type locality: Bolivien: Titicacasee, Huatajata, 4000 m). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Bolivia. fortesculpturatus Scheerpeltz, 1965: 160 (Oxytelus; subgenus Anotylus; Type locality: N.E. Burma, Kambaiti, 7000 ft.). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Myanmar. fortipennis Bernhauer, 1936b: 83 (Oxytelus; Type locality: Luzon: Laguna). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Philippines. foveicollis Scheerpeltz, 1965: 159 (Oxytelus; subgenus Anotylus; Type locality: N.E. Burma, Kambaiti, 7000 ft.). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Myanmar. fragilis Sharp, 1887: 691 (Oxytelus; Type locality: Guatemala, El Tumbador, San Joaquin, San Juan in Vera Paz; Panama, Volcan de Chiriqui). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Guatemala, Panama, Colombia. minimus Erichson, 1840: 798 (Oxytelus; [preoccupied]; Type locality: Columbia). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 162 (Anotylus; synonym of fragilis). frater Cameron, 1930: 262 (Oxytelus; subgenus Anotylus; Type locality: Dehra Dun. Mussoorie, Chakrata and Simla districts, alt. 2300 to 8000 feet). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). 1354 OXYTELINAE: ANOTYLUS Bulletin AMNH
— Abdullah and Qadri, 1970: 121 (Oxytelus; characters; West Pakistan). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: India, Pakistan. fraterculus Cameron, 1928d: 104 (Oxytelus; subgenus Anotylus; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Indonesia. fraternus Cameron, 1928d: 102 (Oxytelus; subgenus Anotylus; Type locality: Sumatra: Fort de Kock; also in the Malay Peninsula). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — Shibata, 1993a: 318 (Anotylus; Taiwan). DISTRIBUTION: Malaysia, Indonesia, Taiwan. frugicola Cameron, 1918: 66 (Oxytelus; subgenus Anotylus; Type locality: Singapore: Mandai). — Cameron, 1921a: 367, 400 (Oxytelus; characters; Singapore). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Singapore. funebris Bernhauer, 1907: 376 (Oxytelus; Type locality: Yamanaka). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — J. Li, 1993: 21 (Anotylus; China). DISTRIBUTION: Japan, China. fusciventris Cameron, 1945a: 144 (Oxytelus; subgenus Anotylus; Type locality: Sambawa: B. Aroe Hassa, altitude 2000–2500 feet. Buitenzorg). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Indonesia. ganglbaueri Bernhauer, 1907, see: latiusculus Kraatz, 1859. garambanus Hammond, 1976a: 43 (Anotylus; Type locality: Zaire: Garamba National Park). DISTRIBUTION: Congo. gardneri Cameron, 1930: 456 (Oxytelus; subgenus Anotylus; Type locality: Darjeeling; Rangirum, alt. 6000 feet). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog). — Abdullah and Qadri, 1970: 124 (Oxytelus; characters; East Pakistan). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: India, Bangladesh. gedyei Bernhauer, 1936h: 323 (Oxytelus; subgenus Tanycraerus; Type locality: Kenya Colony: Katamayu R.). — Cameron, 1950g: 16 (Oxytelus; subgenus Anotylus; Belgian Congo). — Herman, 1970: 409 (Oxytelus). — Hammond, 1976: 158 (Anotylus). — Hammond, Morgan, and Morgan, 1979: 217 (Anotylus; gibbulus species group). DISTRIBUTION: Kenya, Congo. gibbulus Eppelsheim, 1878: 122 (Anotylus; Type locality: Hochgebirge östlich von Kasbek, 5000–10,000 ). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Fagel, 1956b: 275 (Oxytelosus). — Scheerpeltz, 1962: 567 (Oxytelus; subgenus Anotylus; characters; Caucasus). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 158 (Anotylus). — Hammond, Morgan, and Morgan, 1979: 215 (Anotylus; characters, composition, distribution, Bulletin AMNH OXYTELINAE: ANOTYLUS 1355
habits, relationships of gibbulus species group; subfossils of species in England and Ontario; currently found in Armenia, Georgia, eastern Siberia; discussion). DISTRIBUTION: Armenia, Georgia, Russia, Britain†, Canada†. glaber Cameron, 1945a: 144 (Oxytelus; subgenus Anotylus; Type locality: Sumatra: Fort de Kock). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Indonesia. glareosus Wollaston, 1854: 610 (Oxytelus; Type locality: Madeira: garden at the Levada). — Wollaston, 1857: 201 (Oxytelus; habitat; Madeira Islands). — Wollaston, 1864: 598 (Oxytelus; characters; notes; Canary Islands). — Wollaston, 1865: 517 (Oxytelus; habitat; Madeira Islands; Canary Islands). — Fauvel, 1902b: 66 (Oxytelus; Madeira). — Fauvel, 1905e: 80 (Oxytelus; Java; Réunion Island; Mauritius; Antilles; Canaries; Madeira). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Bernhauer, 1928: 7 (Oxytelus; Sumatra). — Blackwelder, 1943: 100 (Oxytelus; characters; Jamaica; Dominica; St. Vincent; Grenada). — Smetana, 1963b: 31 (Delopsis; notes; Madeira). — Herman, 1970: 418 (Anotylus). — Hammond, 1975: 158, 163 (Anotylus; notes; characters; Ceylon; India; Malaya; Ghana; Sierra Leone; Canaries; Madeira; Tahiti; Jamaica; Cuba; Haiti; Grenada; Panama). — Hammond, 1976: 174 (Anotylus; notes; adventive species). — Coiffait, 1976: 221 (Oxytelus; Tahiti). — Shibata, 1993a: 318 (Anotylus; Taiwan). DISTRIBUTION: Madeira, Canary Islands, Sierra Leone, Ghana, Mascarene Islands, Pakistan, Sri Lanka, India, Bangladesh, Malaysia, Indonesia, Taiwan, Society Islands, West Indies, Panama. indicus Bernhauer, 1926: 23 (Oxytelus; Type locality: Kumaon, Haldwani Distr.; Sarda, Bengalen). — Cameron, 1930: 210 (Delopsis; characters; India). — Scheerpeltz, 1933: 1092 (Delopsis; catalog). — Abdullah and Qadri, 1970: 122, 124 (Delopsis; characters; East and West Pakistan). — Herman, 1970: 418 (Anotylus). — Hammond, 1975: 163 (Anotylus; synonym of glareosus). gracilicornis Cameron, 1928d: 102 (Oxytelus; subgenus Anotylus; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Indonesia. grafi Reitter, 1909, see: rugosus Fabricius, 1775. granadillae Cameron, 1918: 66 (Oxytelus; subgenus Anotylus; Type locality: Singapore: Keppel Harbour). — Cameron, 1921a: 367, 400 (Oxytelus; characters; Singapore). — Cameron, 1928f: 402 (Oxytelus; Borneo). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Singapore, Borneo. grandiceps Bernhauer, 1931c: 125 (Oxytelus; Type locality: Hongkong). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog). — Scheerpeltz, 1962: 576 (Oxytelus; subgenus Anotylus; characters; China). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: China. gratus Cameron, 1929b: 443 (Oxytelus; subgenus Anotylus; Type locality: Selangor: The Gap). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Malaysia. 1356 OXYTELINAE: ANOTYLUS Bulletin AMNH gregarius Sharp, 1889: 414 (Oxytelus; Type locality: Nikko). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Cameron, 1949b: 461 (Oxytelus; Formosa). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Japan. gridellii Bernhauer, 1927a: 79 (Oxytelus; Type locality: Eritrea: Saganeiti; Adi Caie). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Eritrea. gughensis Fagel, 1957: 14 (Anotylus; Type locality: Abyssinia: Gamo Prov., Bonghé valley, Gughé highland, cited as Cughé highlands, c. 9500–10,000 ft.). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Ethiopia. hamatus Fairmaire and Laboulbène, 1856: 612 (Oxytelus; Type locality: Paris). — Redtenbacher, 1857: 990 (Oxytelus; characters; Austria). — Kraatz, 1857d: 863 (Oxytelus; characters; Germany). — Pandellé, 1867a: 172 (Oxytelus; characters; France). — Czwalina, 1870: 422 (Oxytelus; characters). — Fauvel, 1872: 176 [= 1873: 18] (Oxytelus; characters; France; Germany; Austria; Italy). — Fauvel, 1873b: 124 [= 1873c: 17] (Oxytelus; Russia). — Redtenbacher, 1874: 254 (Oxytelus; characters; Austria). — Mulsant and Rey, 1878: 544 (Oxytelus; subgenus Anotylus; characters; France). — Ganglbauer, 1895: 644 (Oxytelus; subgenus Anotylus; characters; middle Europe; Italy). — Reitter, 1909: 172 (Oxytelus; subgenus Anotylus; characters; Germany). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Portevin, 1929: 419 (Oxytelus; characters; France). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog). — Palm, 1961a: 35 (Oxytelus; characters; Sweden; Norway; Finland). — Scheerpeltz, 1962: 576 (Oxytelus; subgenus Anotylus; characters; Europe; Mediterranean region). — Horion, 1963: 240 (Oxytelus; subgenus Oxytelops; characters; Norway; Sweden; Finland; Russia; France; Belgium; Switzerland; Italy; Romania; Slovakia; Czech Republic; Bosnia Hercegovina; Bulgaria; Germany; Austria). — Lohse, 1964: 87 (Oxytelus; subgenus Anotylus; characters; central Europe). — Herman, 1970: 418 (Anotylus). — Pope, 1977: 25 (Anotylus; Britain). — Tóth, 1987: 228 (Anotylus; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Sweden). — Kashcheev, 1989: 287 (Oxytelus; subgenus Anotylus; Mongolia). — Bohácˇ, 1993: 43 (Anotylus; Czech Republic; Slovakia). — Hodge and Jones, 1995: 33 (Anotylus; characters; Britain). — Ciceroni and Zanetti, 1995: 12 (Anotylus; Italy). — Owen, 1997: 303 (Anotylus; collecting notes; Britain). — Kashcheev, 1999a: 57 (Oxytelus; subgenus Anotylus; Kazakhstan). DISTRIBUTION: Europe, Russia, Mongolia. hameedi Abdullah and Qadri, 1970: 129 (Neoplatystethus; subgenus Neoplatystethus; Type locality: W. Pakistan: Karachi). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Pakistan. hamuliger Fauvel, 1905a: 127 (Oxytelus; Type locality: Guyane sud: Riviere Lunier; bassin du Carsevenne). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: French Guiana. Bulletin AMNH OXYTELINAE: ANOTYLUS 1357 havelkai R. Dvorˇák, 1954 [Note: Nomen nudum], see: sculpturatus Gravenhorst, 1806. helcopterus Bernhauer, 1934: 3 (Oxytelus; subgenus Anotylus; Type locality: Mittel-Szechuan: Hwa-Yin-Shan, 1600 m). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: China. henryi Fernando, 1960a: 127 (Deinopsis; Type locality: Devanagala, Ceylon). — Hammond, 1975b: 172 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Sri Lanka. herculis Bernhauer, 1936g: 237 (Oxytelus; Type locality: Herkulesbad in Südungarn). — Herman, 1970: 418 (Anotylus). — Tóth, 1987: 228, 229 (Anotylus; characters; Romania). DISTRIBUTION: Romania. heterocerus Fauvel, 1904b: 104 (Oxytelus; Type locality: Gabon: Libreville). — Fauvel, 1907: 17 (Oxytelus; Kilimandjaro). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Eichelbaum, 1913: 121 (Oxytelus; Deutsch-Ostafrika). — Cameron, 1950g: 15 (Oxytelus; subgenus Anotylus; Belgian Congo). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Gabon, Congo, Tanzania. heterophthalmus Fauvel, 1904b: 101 (Oxytelus; Type locality: Australie: Nouvelle Galles du Sud). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Australia. hirtulus Eppelsheim, 1895c: 68 (Oxytelus; Type locality: Simla im Himalaya). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Cameron, 1930: 213 (Oxytelus; characters; India; Burma). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog). — Abdullah and Qadri, 1970: 120 (Oxytelus; characters; West Pakistan). — Herman, 1970: 418 (Anotylus). — Coiffait, 1981d: 325 (Oxytelus; Nepal). — Coiffait, 1983d: 162 (Oxytelus; Nepal). — Coiffait, 1984: 118 (Oxytelus; Nepal). — Coiffait, 1984e: 374 (Oxytelus; Nepal). DISTRIBUTION: Pakistan, India, Nepal, Myanmar. holdhausi Bernhauer, 1908d: 290 (Oxytelodes; Type locality: Brasilien: S. Catharina). — Bernhauer and Schubert, 1911: 108 (Oxytelodes; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Brazil. hosodai Ito, 1991: 87 (Anotylus; Type locality: Japan: Yamanashi Pref.: Hauou Lodge, alt. 2400 m, Mt. Houou). DISTRIBUTION: Japan. hostilis Bernhauer, 1936b: 83 (Oxytelus; Type locality: Mindoro: Subaan. Luzon: Mount Maquiling; Mount Isarog; Imugan; Balbalan; Bayombong; Los Baños; Limay; Pagsanjan. Mindanao: Surigao. Biliran). — Herman, 1970: 418 (Anotylus; cited as hostilus). — Frank, 1982: 9 (Anotylus; list of parasites). DISTRIBUTION: Philippines. humilis Gistel, 1857 [Note: Nomen dubium], see: styricola E. Strand [Note: Nomen dubium], 1917. hybridus Eppelsheim, 1878: 121 (Oxytelus; Type locality: Michailowo am Fusse des Suramgebirges; Hochebene von Gomereti 5000 ). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). 1358 OXYTELINAE: ANOTYLUS Bulletin AMNH
— C. Koch, 1932: 653 (Oxytelus; subgenus Anotylus; characters in key). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog). — Pototskaia, 1967: 29 (Oxytelus; larval characters). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Georgia, Turkey. turcicus Bernhauer, 1927b: 90 (Oxytelus; subspecies hybridus; Type locality: Belgraderwald bei Konstantinopel). — Scheerpeltz, 1933: 1097 (Oxytelus; synonym of hybridus). impennis Fauvel, 1877: 198 (Oxytelus; Type locality: Australie, Victoria). — Fauvel, 1878g: 493 (Oxytelus; characters; Australia). — Bernhauer, 1908: 15 (Oxytelus; Australia). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Australia. impressifrons W. J. MacLeay, 1873: 150 (Oxytelus; Type locality: Gayndah). — Fauvel, 1877: 200 (Oxytelus; characters; Australia). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Australia. flavior Blackburn, 1902: 24 (Oxytelus; Type locality: Victoria, Dividing Range and Australian Alps). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — Hammond, 1976: 162 (Anotylus; synonym of impressifrons). inaequalis Fauvel, 1905a: 127 (Oxytelus; Type locality: Republique Argentine: province de Buenos Aires). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — Scheerpeltz, 1975a: 242 (Oxytelus; collecting notes; Brazil). DISTRIBUTION: Argentina, Brazil. incilis Sharp, 1887: 693 (Oxytelus; Type locality: Guatemala, Coatepeque 13000 feet; Panama, San Miquel in the Pearl Islands). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Guatemala, Panama. indicus Bernhauer, 1926, see: glareosus Wollaston, 1854. inornatus Cameron, 1928d: 104, 105 (Oxytelus; subgenus Anotylus; Type locality: Malay Peninsula: Selangor). — Scheerpeltz, 1933: 1097 (Oxytelus; catalog; Sumatra). — Herman, 1970: 418 (Anotylus). — Herman, 2001: 22 (Anotylus; synonymy). DISTRIBUTION: Malaysia, Indonesia. inornatus Cameron, 1929b: 445 (Oxytelus; [preoccupied]; Type locality: Selangor: Bukit Kutu). — Herman, 2001: 22 (Anotylus; synonym of inornatus). inornatus Cameron, 1929, see: inornatus Cameron, 1928. insecatus Gravenhorst, 1806: 189 (Oxytelus; variety of carinatus; Type locality: Not cited). — Gyllenhal, 1810: 453 (Oxytelus; [Note: Misidentification: Bernhauer and Schubert, 1911: 114 cited carinatus variety b Gyllenhal as a synonym of insecatus Gravenhorst; there is no avail- able name]; characters; Sweden). — Mannerheim, 1830: 48 (Oxytelus; [Note: Misidentification: Scheerpeltz, 1933: 1098 cited sculpuratus Mannerheim (= sculpturatus) as a synonym of insecatus Gravenhorst; Mannerheim did not describe the species as new, he attributed it to Gravenhorst]; Finland). Bulletin AMNH OXYTELINAE: ANOTYLUS 1359
— Mannerheim, 1831: 462 (Oxytelus; [Note: Misidentification: Scheerpeltz, 1933: 1098 cited sculpuratus Mannerheim (= sculpturatus) as a synonym of insecatus Gravenhorst; Mannerheim did not describe the species as new, he attributed it to Gravenhorst]; Finland). — Erichson, 1839a: 589 (Oxytelus; characters; Germany). — Heer, 1839: 204 (Oxytelus; characters; Switzerland). — Erichson, 1840: 787 (Oxytelus; characters; Europe). — Redtenbacher, 1849: 736 (Oxytelus; characters; Austria). — Fairmaire and Laboulbène, 1856: 609 (Oxytelus; characters; France). — Redtenbacher, 1857: 232 (Oxytelus; characters; Austria). — Kraatz, 1857d: 852 (Oxytelus; characters; Germany). — Thomson, 1861: 127 (Oxytelus; characters; Scandinavia). — Hochhuth, 1862: 92 (Oxytelus; Russia). — Fauvel, 1871a: 192 [= 1872: 166] (Oxytelus; characters; France; Europe). — Redtenbacher, 1874: 253 (Oxytelus; characters; Austria). — Seidlitz, 1875: 247 (Oxytelus; characters; Baltic region). — J. Sahlberg, 1876: 175 (Oxytelus; Finland). — Mulsant and Rey, 1878: 493 (Oxytelus; characters; France). — Fowler, 1888: 378 (Oxytelus; characters; Britain). — Ganglbauer, 1895: 637 (Oxytelus; characters; Europe). — Montandon, 1908: 78 (Oxytelus; Romania). — Reitter, 1909: 170 (Oxytelus; subgenus Oxytelus; characters; Germany). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Johansen, 1914: 556 (Oxytelus; subgenus Oxytelus; characters; Denmark). — Portevin, 1929: 417 (Oxytelus; characters; France). — C. Koch, 1932: 656 (Oxytelus; characters in key). — Scheerpeltz, 1933: 1098 (Oxytelus; catalog). — Palm, 1961a: 30 (Oxytelus; characters; Sweden; Denmark). — Horion, 1963: 223 (Oxytelus; subgenus Styloxys; Britain; Norway; Sweden; Finland; Russia; Denmark; Ukraine; Poland; Czechoslovakia; Hungary; Romania; Bosnia; Italy; Frnace; Switzerland; Belgium; Holland; Germany; Austria). — Lohse, 1964: 83 (Oxytelus; subgenus Styloxys, cited as Styloxis; characters; central Europe). — Szujecki, 1968a: 714 (Oxytelus; Poland). — Herman, 1970: 418 (Anotylus). — Pope, 1977: 25 (Anotylus; Britain). — Frank, 1982: 9 (Anotylus; list of parasites). — Campbell and Tomlin, 1984: 310 (Anotylus; characters; natural history notes; Ontario). — Tóth, 1987: 228 (Anotylus; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden). — Schülke and Uhlig, 1988: 5 (Anotylus; Germany). — Bohácˇ, 1993: 43 (Oxytelus; subgenus Styloxys, cited as Styloxis; Czech Republic; Slovakia). — Ciceroni and Zanetti, 1995: 12 (Anotylus; Italy). — Downie and Arnett, 1996: 448 (Anotylus; cited as insectatus; characters; Canada). — Czarniawski and Staniec, 1997: 40 (Anotylus; structure of opening of abdominal defensive reservoirs). — Assing, 2001: 75 (Anotylus; collected from xerothermous site; Germany). DISTRIBUTION: Europe, Russia, Canada. nitidus Stephens, 1832: 318 (Oxytelus; Type locality: Suffolk). — Stephens, 1839: 419 (Oxytelus; characters; England). — G. Waterhouse, 1858: 30 (Oxytelus; synonym of insecatus). — Bernhauer and Schubert, 1911: 114 (Oxytelus; synonym of insecatus). insignitus Gravenhorst, 1806: 188 (Oxytelus; variety of carinatus; Type locality: Not cited). — Erichson, 1840: 793 (Oxytelus; characters; America septentrionalis; St. Thomas; Colombia; Brazil). — Guérin-Méneville, 1857: 42 (Oxytelus; characters; Cuba; St. Thomas; Colombia; Brazil; America septentrionale). 1360 OXYTELINAE: ANOTYLUS Bulletin AMNH
— Wollaston, 1857: 199 (Oxytelus; characters; notes; habitat; Madeira Islands). — Chevrolat and Fauvel, 1863: 438 (Oxytelus; characters; Cuba, Saint Thomas, Colombia, Brazil). — Wollaston, 1865: 516 (Oxytelus; habitat; Madeira Islands). — Solsky, 1868a: 144 (Oxytelus; Mexico). — LeConte, 1877: 234, 236 (Oxytelus; characters; New York to Florida and Kansas). — Sharp, 1887: 689 (Oxytelus; Mexico; Guatemala; Nicaragua; Panama; Colombia; Antilles). — Fleutiaux and Sallé, 1889: 381 (Oxytelus; Guadeloupe Island). — Fauvel, 1891: 90 (Oxytelus; notes; Venezuela). — Casey, 1894: 384 (Oxytelus; characters; Atlantic States from New York to Texas; Mexico). — Fauvel, 1902b: 65 (Oxytelus; Madeira). — Blatchley, 1910: 468 (Oxytelus; characters; Indiana). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Notman, 1920b: 6965 (Oxytelus; Florida). — Scheerpeltz, 1933: 1098 (Oxytelus; catalog). — Blackwelder, 1943: 92 (Oxytelus; characters; Cuba; Jamaica; Haiti; Dominican Republic; Puerto Rico; St. Thomas; St. Croix; Antigua; Montserrat; Guadeloupe; Dominica; St. Lucia; St. Vincent; Grenadines; Grenada; Tobago; Trinidad). — Smetana, 1963b: 32 (Oxytelus; subgenus Anotylus; Madeira). — Herman, 1970: 418 (Anotylus). — Hammond, 1976: 174 (Anotylus; notes; adventive species; Brazil; Costa Rica; Mexico; USA; Jamaica; Madeira; Mauritius; Réunion; Tahiti). — Coiffait, 1976: 221 (Oxytelus; Tahiti). — Frank and Thomas, 1981: 403l, 404 (Anotylus; Florida; Costa Rica; Honduras). — Frank, 1982: 9 (Anotylus; list of parasites). — Downie and Arnett, 1996: 448 (Anotylus; characters; USA). — Bain, 1998: 39, 42 (Anotylus; 17th century remains; Massachusetts). DISTRIBUTION: USA, Mexico, Honduras, Guatemala, Nicaragua, Costa Rica, Panama, West Indies, Colombia, Venezuela, Brazil, Madeira Islands, Mascarene Islands, Society Islands. americanus Mannerheim, 1830: 48 (Oxytelus; Type locality: America borealis). — Mannerheim, 1831: 462 (Oxytelus; America borealis). — Erichson, 1840: 793 (Oxytelus; synonym of insignitus). — Guérin-Méneville, 1857: 43 (Oxytelus; synonym of insignitus). — Chevrolat and Fauvel, 1863: 438 (Oxytelus; synonym of insignitus). — Fleutiaux and Sallé, 1889: 381 (Oxytelus; synonym of insignitus). — Casey, 1894: 384 (Oxytelus; synonym of insignitus). — Bernhauer and Schubert, 1911: 114 (Oxytelus; synonym of insignitus). — Blackwelder, 1943: 93 (synonym of insignitus). pumilio Boheman, 1858: 34 (Oxytelus; Type locality: Insula Taiti). — Fauvel, 1891: 91 (Oxytelus; synonym of insignitus). — Bernhauer and Schubert, 1911: 114 (Oxytelus; synonym of insignitus). — Blackwelder, 1943: 93 (synonym of insignitus). intermedius Cameron, 1930, see: disparatus Cameron, 1928. intricatus Erichson, 1840: 794 (Oxytelus; Type locality: Austria; Smyrnae). — Kolenati, 1846: 26 (Oxytelus; Caucasus; Transcaucasus). — Hochhuth, 1849: 190 (Oxytelus; characters). — Redtenbacher, 1849: 738 (Oxytelus; characters; Austria). — Fairmaire and Laboulbène, 1856: 611 (Oxytelus; characters; France). — Redtenbacher, 1857: 233 (Oxytelus; characters; Austria). — Kraatz, 1857d: 859 (Oxytelus; characters; Germany). — Fauvel, 1872: 172 [= 1873: 14] (Oxytelus; characters; France; Spain; Italy; Austria; Turkey; Syria; Iran). — Redtenbacher, 1874: 254 (Oxytelus; characters; Austria). — Fauvel, 1876a: 57 [= 1876: 242] (Oxytelus; Morocco). Bulletin AMNH OXYTELINAE: ANOTYLUS 1361
— Fauvel, 1878: 95 [= 1878a: 95] (Oxytelus; Morocco; Algeria). — Mulsant and Rey, 1878: 524 (Oxytelus; subgenus Anotylus; characters; France). — Fauvel, 1886: 22 [= 1886a: 14] (Oxytelus; Morocco; Tunisia; Algeria). — Ganglbauer, 1895: 642 (Oxytelus; subgenus Anotylus; characters; middle Europe; Mediter- ranean region). — Fauvel, 1902b: 66 (Oxytelus; Barbary). — Montandon, 1908: 78 (Oxytelus; Romania). — Reitter, 1909: 171 (Oxytelus; subgenus Anotylus; characters; Germany). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Portevin, 1929: 418 (Oxytelus; characters; France). — Scheerpeltz, 1933: 1098 (Oxytelus; catalog). — C. Koch, 1941: 55 (intricatus; Morocco). — Scheerpeltz, 1962: 567 (Oxytelus; subgenus Anotylus; characters; Europe; Mediterranean region; Turkey; Caucasus; Iran; Afghanistan; Transcaspian region; Turkestan). — Scheerpeltz, 1963: 6 (Oxytelus; subgenus Anotylus; Afghanistan). — Horion, 1963: 231 (Oxytelus; subgenus Anotylus; Turkey; Caucasus; Iran; Afghanistan; Trans- caspian region; Spain; France; Croatia; Bosnia; Ukraine; Romania; Hungary; Slovakia; Austria; Germany). — Lohse, 1964: 86 (Oxytelus; subgenus Anotylus; characters; central Europe). — Raianu, 1964: 105 (Oxytelus; Romania). — Scheerpeltz, 1965b: 393 (Oxytelus; subgenus Anotylus; Sicily). — Smetana, 1967e: 318 (Oxytelus; subgenus Anotylus; Turkey). — Herman, 1970: 418 (Anotylus). — Tóth, 1987: 228 (Anotylus; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Poland; Czechoslovakia; Austria; France). — Bohácˇ, 1993: 43 (Anotylus; Slovakia). — Ciceroni and Zanetti, 1995: 12 (Anotylus; Italy). DISTRIBUTION: Morocco, Algeria, Tunisia, Europe, Caucasus, Turkey, Syria, Iran, Afghanistan, “Turkestan”. scaber Rosenhauer, 1856: 82 (Oxytelus; Type locality: Granada). — Fauvel, 1873b: 124 [= 1873c: 17] (Oxytelus; synonym of intricatus). — Ganglbauer, 1895: 642 (Oxytelus; synonym of intricatus). — Bernhauer and Schubert, 1911: 114 (Oxytelus; synonym of intricatus). — Scheerpeltz, 1962: 567 (Oxytelus; synonym of intricatus). inustus Gravenhorst, 1806: 188 (Oxytelus; variety of carinatus; Type locality: Not cited). — Erichson, 1839a: 593 (Oxytelus; characters; Germany). — Heer, 1839: 205 (Oxytelus; characters; Switzerland). — Erichson, 1840: 791 (Oxytelus; characters; Europe). — P. Lucas, 1846: 127 (Oxytelus; notes; Algeria). — Redtenbacher, 1849: 737 (Oxytelus; characters; Austria). — Kiesenwetter, 1851a: 427 (Oxytelus; France). — Rosenhauer, 1856: 82 (Oxytelus; Cadiz). — Fairmaire and Laboulbène, 1856: 611 (Oxytelus; characters; France). — Redtenbacher, 1857: 232 (Oxytelus; characters; Austria). — Kraatz, 1857d: 856 (Oxytelus; characters; Germany). — Kraatz, 1858b: 127 (Oxytelus; Crete; Greece). — Peyron, 1858: 433 (Oxytelus; Turkey). — Hochhuth, 1862: 92 (Oxytelus; Russia). — Fauvel, 1872: 170 [= 1873: 12] (Oxytelus; characters; France; Europe; North Africa; Syria; Cyprus; Caramania; Iran). — Redtenbacher, 1874: 253 (Oxytelus; characters; Austria). — Seidlitz, 1875: 247 (Oxytelus; characters; Baltic region). — Fauvel, 1878: 94 [= 1878a: 14] (Oxytelus; Barbary). — Mulsant and Rey, 1878: 511 (Oxytelus; subgenus Anotylus; characters; France). — Fauvel, 1886: 22 [= 1886a: 14] (Oxytelus; North Africa). 1362 OXYTELINAE: ANOTYLUS Bulletin AMNH
— Fowler, 1888: 380 (Oxytelus; characters; Britain). — Ganglbauer, 1895: 640 (Oxytelus; subgenus Anotylus; characters; middle Europe; Mediter- ranean region). — Fauvel, 1902b: 65 (Oxytelus; Barbary). — Sainte-Claire Deville, 1906: 77 (Oxytelus; Corsica). — Reitter, 1909: 171 (Oxytelus; subgenus Anotylus; characters; Germany). — Bernhauer and Schubert, 1911: 114 (Oxytelus; catalog). — Johansen, 1914: 561 (Oxytelus; subgenus Anotylus; characters; Denmark). — Portevin, 1929: 418 (Oxytelus; characters; France). — C. Koch, 1932: 653 (Oxytelus; subgenus Anotylus; characters in key). — Scheerpeltz, 1933: 1098 (Oxytelus; catalog). — West, 1937: 43 (Oxytelus; Faroe Islands). — Paulian, 1941: 167 (Oxytelus; larval characters). — Scheerpeltz, 1956c: 1093 (Oxytelus; subgenus Anotylus; Greece; Cyprus). — Scheerpeltz, 1958a: 8 (Oxytelus; subgenus Anotylus; Turkey). — Scheerpeltz, 1961d: 119 (Oxytelus; subgenus Anotylus; Italy). — Scheerpeltz, 1961e: 135 (Oxytelus; subgenus Anotylus; Rhodes). — Scheerpeltz, 1961h: 250 (Oxytelus; subgenus Anotylus; Afghanistan). — Palm, 1961a: 33 (Oxytelus; characters; Sweden; Denmark). — Horion, 1963: 229 (Oxytelus; subgenus Anotylus; Turkey; Iran; Afghanistan; India; Transbaikal; eastern Russia; Portugal; Spain; Balearics; Italy; Dalmatia; Greece; Crete; Cyprus; France; Switzerland; Belgium; Holland; Britain; Denmark; Sweden; Poland; Czechoslovakia; Germany; Austria; Hungary; Romania; Bosnia Hercegovina; Serbia; Turkey; Russia; Caucasus). — Lohse, 1964: 85 (Oxytelus; subgenus Anotylus; characters; central Europe). — Scheerpeltz, 1965b: 393 (Oxytelus; subgenus Anotylus; Sicily). — Scheerpeltz, 1965c: 497 (Oxytelus; Sicily). — Smetana, 1967e: 318 (Oxytelus; subgenus Anotylus; Turkey). — Pototskaia, 1967: 28 (Oxytelus; larval characters). — Herman, 1970: 418 (Anotylus). — Coiffait, 1970e: 64 (Oxytelus; Taygete). — Jarrige, 1971: 499 (Anotylus; Iran). — Coiffait, 1973a: 270 (Oxytelus; Morocco). — Osella and Zanetti, 1975: 90 (Anotylus; collected from nest of Talpa europaea; notes; Italy). — Pope, 1977: 25 (Anotylus; Britain). — Bordoni, 1982a: 139 (Anotylus; cave collection; Sardinia). — Frank, 1982: 9 (Anotylus; list of parasites). — Tóth, 1987: 228 (Anotylus; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden). — Schülke and Uhlig, 1988: 5 (Anotylus; Germany). — Bohácˇ, 1993: 43 (Anotylus; Czech Republic; Slovakia). — Sparacio, 1995: 145 (Anotylus; characters; notes; Sicily). — Ciceroni and Zanetti, 1995: 12 (Anotylus; Italy). — Assing and Wunderle, 2001: 36 (Anotylus; Cyprus). — Assing, 2001: 75 (Anotylus; collected from xerothermous site; Germany). DISTRIBUTION: Faroe Islands, Europe, Algeria, Syria, Turkey, Iran, Afghanistan, Russia. maxillosus Sperk, 1835: 154 (Bledius; Type locality: Odessa). — Fauvel, 1876a: 57 [= 1876: 242] (Oxytelus; synonym of inustus). — Fowler, 1888: 380 (Oxytelus; synonym of inustus). — Ganglbauer, 1895: 640 (Oxytelus; synonym of inustus). — Bernhauer and Schubert, 1911: 115 (Oxytelus; synonym of inustus). excavatus Motschulsky, 1857a: 503 (Oxytelus; Type locality: Algérie). — Fauvel, 1878: 94 [= 1878a: 14] (Oxytelus; Algeria). — Fauvel, 1886: 22 [= 1886a: 14] (Oxytelus; Algeria). Bulletin AMNH OXYTELINAE: ANOTYLUS 1363
— Fauvel, 1902b: 67 (Oxytelus; Algeria). — Bernhauer and Schubert, 1911: 124 (Platystethus; synonym of oxytelinus). — Hammond, 1976: 162 (Anotylus; synonym of inustus). ixellensis R. Dvorˇák, 1954, see: complanatus Erichson, 1839. jacobsoni Cameron, 1928d: 106 (Oxytelus; subgenus Anotylus; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1098 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Indonesia. jamaicensis Blackwelder, 1943: 102 (Oxytelus; Type locality: Jamaica: main range of Blue Mountains, elevation 5000 to 7000 feet). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: West Indies. japonicus Cameron, 1930c: 183 (Delopsis; Type locality: Kobe: Gomo. Ueno). — Scheerpeltz, 1933: 1092 (Delopsis; catalog). — Herman, 1970: 418 (Anotylus). — Hasegawa and Kanie, 1992: 20 (Anotylus; seashore collection; Japan). — Watanabe, 1998: 313 (Anotylus; Japan). DISTRIBUTION: Japan. jarrigei Hammond, 1976: 158 (Anotylus; replacement name for fageli Jarrige). DISTRIBUTION: Madagascar. fageli Jarrige, 1970: 60 (Delopsis; [preoccupied]; Type locality: Madagascar: Mt. Tsaratanana, Andampy, 750 m). — Hammond, 1976: 158 (Anotylus; synonym of jarrigei). kashmiricus Cameron, 1942: 107 (Oxytelus; subgenus Anotylus; Type locality: Kashmir: Gulmarg). — Herman, 1970: 418 (Anotylus). — Coiffait, 1981e: 350 (Anotylus; Kashmir). DISTRIBUTION: India. kinangopensis Herman, 2001: 41 (Anotylus; replacement name for parasitus Bernhauer). DISTRIBUTION: Kenya. parasitus Bernhauer, 1936d: 213 (Oncoparia; [preoccupied]; Type locality: Kenya: Berg Kinangop, Ostseite, 3200 m). — Herman, 1970: 423 (Oncoparia). — Hammond, 1976: 159 (Anotylus). — Herman, 2001: 41 (Anotylus; synonym of kinangopensis). kokodanus Cameron, 1939b: 140 (Oxytelus; subgenus Anotylus; Type locality: New Guinea: Kokoda, altitude 1200 feet). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: New Guinea. kraatzi Bernhauer and Schubert, 1911: 115 (Oxytelus; replacement name for pulcher Kraatz). — Scheerpeltz, 1933: 1098 (Oxytelus; catalog). — Cameron, 1936a: 31 (Oxytelus; subgenus Anotylus; Java). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: “East Indies”, Myanmar, Malaysia, Indonesia. pulcher Kraatz, 1859: 173 (Oxytelus; [preoccupied]; Type locality: India orientalis). — Fauvel, 1905e: 79 (Oxytelus; Java; Sumbawa; Sumatra; Burma). kraatzi Cameron, 1921a: 367, 400 (Oxytelus; [preoccupied]; replacement name for pulcher Kraatz). kraatzi Cameron, 1925: 16 (Oxytelus; [preoccupied]; replacement name for pulcher Kraatz). 1364 OXYTELINAE: ANOTYLUS Bulletin AMNH
kraatzi Cameron, 1930: 251 (Oxytelus; subgenus Anotylus; [preoccupied]; replacement name for pulcher Kraatz. Also cited from Burma, Malaya Peninsula, Sumatra, and Java). — Watanabe and Shibata, 1976: 329 (Anotylus; Malaysia). kraatzi Cameron, 1921, see: kraatzi Bernhauer and Schubert, 1911. kraatzi Cameron, 1925, see: kraatzi Bernhauer and Schubert, 1911. kraatzi Cameron, 1930, see: kraatzi Bernhauer and Schubert, 1911. kraussi Steel, 1954: 184 (Oxytelus; subgenus Anotylus; Type locality: New Caledonia: Bois du Sud). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: New Caledonia. laetus Cameron, 1930: 252 (Oxytelus; subgenus Anotylus; Type locality: Dehra Dun. Siwaliks: Malhan Range. Almora: Dindihar, alt. 6500 feet). — Scheerpeltz, 1933: 1098 (Oxytelus; catalog). — Abdullah and Qadri, 1970: 121 (Oxytelus; characters; West Pakistan). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: India, Pakistan. laevissimus Fauvel, 1905a: 128 (Oxytelus; Type locality: Ega). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Brazil. lateralis Lea, 1906: 207 (Oxytelus; Type locality: Sydney and Tamworth, N.S.W.). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Australia. laticeps Fagel, 1957: 13 (Anotylus; Type locality: Ethiopia: Simien, E. of Mindagabsa, over 10,000 ft.). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Ethiopia. laticornis Sharp, 1874: 95 (Oxytelus; Type locality: Nagasaki). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Scheerpeltz, 1962: 573 (Oxytelus; subgenus Anotylus; characters; Japan; China). — Herman, 1970: 418 (Anotylus). — Ito, 1992: 63 (Anotylus; Yakushima Island). DISTRIBUTION: Japan. latifrons Sharp, 1887: 693 (Oxytelus; Type locality: Panama, Bugaba). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Panama. latiusculus Kraatz, 1859: 176 (Oxytelus; Type locality: Ceylan). — Fauvel, 1902b: 67 (Oxytelus; Egypt; Syria; China; Sumatra; Madagascar; Gabon; South Africa; Australia). — Fauvel, 1904a: 46 (Oxytelus; Belgaum; Ceylon; China; Annam; Burma; Sumatra; Australia; Madagascar; Syria; Egypt; Ethiopia; Gabon; Natal; Transvaal; Cape). — Fauvel, 1907: 17 (Oxytelus; Nairobi; Mexico). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Cameron, 1921a: 366, 400 (Oxytelus; characters; Singapore). — Bernhauer, 1922c: 223 (Oxytelus; Formosa). — Cameron, 1928d: 101, 103 (Oxytelus; subgenus Anotylus; characters in key; Sumatra). — Cameron, 1929b: 444 (Oxytelus; characters in key). — Scheerpeltz, 1929b: 212 (Oxytelus; notes; Palestine). Bulletin AMNH OXYTELINAE: ANOTYLUS 1365
— Cameron, 1930: 250 (Oxytelus; subgenus Anotylus; characters; India; Ceylon; Malayan Archipelago; Formosa; Australia; Aden; Egypt; Syria). — Scheerpeltz, 1933: 1099 (Oxytelus; catalog). — Bernhauer, 1936b: 84 (Oxytelus; Philippines). — Hafez, 1939a (Oxytelus; larval characters). — Bernhauer, 1942c: 350 (Oxytelus; Cameroon). — Hafez, 1942 (Oxytelus; life history). — Cameron, 1950g: 18 (Oxytelus; subgenus Anotylus; Belgian Congo). — Watanabe, 1961: 349 (Oxytelus; Izu Islands). — Scheerpeltz, 1962: 569 (Oxytelus; subgenus Anotylus; characters). — Herman, 1970: 418 (Anotylus). — Abdullah and Qadri, 1970: 121, 124 (Oxytelus; characters; East and West Pakistan). — Hammond, 1975: 158, 159 (Anotylus; notes; characters; Ceylon; Japan; China; Hong Kong; Taiwan; Australia; New Guinea; Philippines; Vietnam; Malaya; Singapore; Cocos-Keeling Islands; India; Nepal; Madagascar; Somaliland; Kenya; Rhodesia; Botswana; South Africa; South-West Africa; Angola; Congo; Tanzania; Cameroon; Nigeria; Ivory Coast; Egypt; West Aden; Israel; Cyprus). — Hammond, 1976: 175 (Anotylus; notes; adventive species). — Scheerpeltz, 1978: 195 (Oxytelus; subgenus Anotylus; Timor). — Ito, 1992: 63 (Anotylus; Yakushima Island). — J. Li, 1993: 21 (Anotylus; China). — Kashcheev, 1999a: 57 (Oxytelus; subgenus Anotylus; Kazakhstan). DISTRIBUTION: Africa, southwest and southern Asia, Indonesia, Philippines, New Guinea, Australia, Taiwan, Japan, China. pusillus Boheman, 1848: 296 (Oxytelus; [preoccupied]; Type locality: Caffraria orientalis). — Bernhauer and Schubert, 1911: 118 (Oxytelus; catalog). — Eichelbaum, 1913: 121 (Oxytelus; characters; Tanzania). — Scheerpeltz, 1933: 1102 (Oxytelus; catalog). — Herman, 1970: 410 (Oxytelus). — Hammond, 1975: 159 (Anotylus; synonym of latiusculus). sulcifrons Fauvel, 1875a: XI [= 1875b: 213] (Oxytelus; Type locality: Syrie, El-Ghor). — Fauvel, 1902b: 67 (Oxytelus; synonym of latiusculus). — Fauvel, 1904a: 46 (Oxytelus; synonym of latiusculus). — Bernhauer and Schubert, 1911: 115 (Oxytelus; synonym of latiusculus). — Cameron, 1930: 251 (Oxytelus; synonym of latiusculus). — Scheerpeltz, 1962: 570 (Oxytelus; subgenus Anotylus; subspecies of latiusculus; characters; Syria; Lebanon; Palestine). — Hammond, 1975: 159 (Anotylus; synonym of latiusculus). m-elevatus Lea, 1906: 206 (Oxytelus; Type locality: Richmond River, N.S.W.). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Herman, 1970: 410 (Oxytelus; cited as melevatus). — Hammond, 1975: 159 (Anotylus; synonym of latiusculus). ganglbaueri Bernhauer, 1907: 375 (Oxytelus; Type locality: Kanagawa). — Bernhauer and Schubert, 1911: 113 (Oxytelus; catalog). — Scheerpeltz, 1933: 1099 (Oxytelus; synonym of latiusculus). — Scheerpeltz, 1962: 570 (Oxytelus; subgenus Anotylus; subspecies of latiusculus; characters; Japan). — Nakane, 1963a: 84 (Oxytelus; cited as valid species; characters; habitus photograph; Japan). — Watanabe and Shibata, 1965: 317 (Oxytelus; subgenus Anotylus; cited as valid species; Japan). — Hammond, 1975: 159 (Anotylus; synonym of latiusculus). boehmi Bernhauer, 1910b: 256 (Oxytelus; Type locality: Kairo). — Bernhauer and Schubert, 1911: 110 (Oxytelus; catalog). — Scheerpeltz, 1933: 1099 (Oxytelus; synonym of latiusculus). — C. Koch, 1934: 47 (Oxytelus; subgenus Anotylus; subspecies of latiusculus; characters; Egypt). 1366 OXYTELINAE: ANOTYLUS Bulletin AMNH
— Scheerpeltz, 1962: 570 (Oxytelus; subgenus Anotylus; subspecies of latiusculus; characters; Egypt). — Scheerpeltz, 1963b: 426 (Oxytelus; subgenus Anotylus; variety of latiusculus; Egypt). — Hammond, 1975: 159 (Anotylus; synonym of latiusculus). niticeps Bernhauer, 1935: 5 (Oxytelus; subspecies of latiusculus; Type locality: Tempelberg, 900 m, Ginyün b. Bébé, C. China). — Scheerpeltz, 1962: 569 (Oxytelus; subgenus Anotylus; subspecies of latiusculus; characters; China). — Hammond, 1975: 159 (Anotylus; subspecies of latiusculus). leleupi Fagel, 1965a: 341 (Oncoparia; Type locality: Kilimanjaro, Bismarck Hügel, 3100 m). — Herman, 1970: 423 (Oncoparia). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Tanzania. lewisius Sharp, 1874: 95 (Oxytelus; Type locality: Nagasaki). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Bernhauer, 1922c: 223 (Oxytelus; Formosa). — Scheerpeltz, 1933: 1099 (Oxytelus; catalog). — Watanabe, 1961: 349 (Oxytelus; Izu Islands). — Herman, 1970: 418 (Anotylus). — Watanabe and Shibata, 1972: 61 (Oxytelus; Yaku-shima Island). — Hasegawa and Kanie, 1992: 20 (Anotylus; seashore collection; Japan). — J. Li, 1993: 21, 55 (Anotylus; characters; China). — Watanabe, 1996a: 9 (Anotylus; cave collection; Japan). — Watanabe, 1998: 313 (Anotylus; Japan). DISTRIBUTION: Japan, Taiwan, China. similis Cameron, 1930c: 184 (Oxytelus; subgenus Anotylus; Type locality: Kobe: Harada). — Scheerpeltz, 1933: 1104 (Oxytelus; catalog). — Herman, 1970: 419 (Anotylus). — Hammond, 1976: 162 (Anotylus; synonym of lewisius). liliputanus Bernhauer, 1936b: 85 (Oxytelus; Type locality: Luzon: Los Baños). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Philippines. lippensi Bernhauer, 1943b: 277 (Oxytelus; subgenus Tanycraerus; Type locality: Ruanda: L. Karago). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Rwanda, Congo. armatus Cameron, 1950g: 8 (Oxytelus; [preoccupied]; Type locality: Belgian Congo: Myakibumba, près Kikere, alt. 2226 m). — Herman, 1970: 417 (Anotylus). — Hammond, 1976: 162 (Anotylus; synonym of lippensi). lobatus Fauvel, 1904e: 303 (Oxytelus; Type locality: Suberbieville). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Madagascar. loebli Hammond, 1975: 168 (Anotylus; Type locality: Ceylon: Prov. of Uva: Uva Hills, Tollacombe Bungalow, Namunkula, 4000 ft.). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Sri Lanka. longicornis Fauvel, 1905, see: bogorensis Herman, 2001. lucidus Sharp, 1887: 689 (Oxytelus; Type locality: Guatemala, San Juan in Vera Paz; Panama, Volcan de Chiriqui). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). Bulletin AMNH OXYTELINAE: ANOTYLUS 1367
— Scheerpeltz, 1933: 1099 (Oxytelus; catalog; Argentina). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Argentina, Panama, Guatemala. lungwensis Fagel, 1957: 7 (Pseudodelopsis; Type locality: Congo Belge: Kivu: territoire d’Uvira, lac Lungwe, 2700 m). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Congo. luridipennis Luze, 1904: 84 (Oxytelus; Type locality: Seravschan: Serbent; Samarkand: Dargom). — Bernhauer and Schubert, 1911: 117 (Oxytelus; variety of nitidulus). — Scheerpeltz, 1960b: 68 (Oxytelus; subgenus of Anotylus; variety of nitidulus, cited as mitidulus; Afghanistan). — Scheerpeltz, 1963: 6 (Oxytelus; subgenus Anotylus; subspecies of nitidulus; Afghanistan). — Jarrige, 1971: 500 (Anotylus; Iran). — Kashcheev, 1989: 287 (Oxytelus; subgenus Anotylus; cited as valid species; Mongolia). DISTRIBUTION: Mongolia, Uzbekistan, Afghanistan, Iran. luzonicus Bernhauer, 1936b: 85 (Oxytelus; Type locality: Luzon: Mount Maquiling. Mindoro: San Teodoro). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Philippines. madecassus Fauvel, 1904b: 96 (Oxytelopsis; Type locality: Madagascar: Tananarive; forêt de Moramanga). — Bernhauer and Schubert, 1911: 108 (Oxytelopsis; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Madagascar. magdalenae Fauvel, 1905a: 132 (Oxytelus; Type locality: Colombie: Rio Magdalena. Equateur: Pichincha). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Colombia, Ecuador. magniceps Wendeler, 1955: 194 (Oxytelus; subgenus Anotylus; Type locality: Brasilien: Nova Teutonia). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Brazil. mahmoodi Abdullah and Qadri, 1970: 126 (Platystethus; subgenus Pseudopyctocraerus; Type locality: W. Pakistan: Karachi). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Pakistan. malayanus Cameron, 1930a: 154 (Oxytelus; subgenus Anotylus; Type locality: Singapore. Selangor; Kuala Lumpur). — Scheerpeltz, 1933: 1099 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Malaysia, Singapore. ruficeps Cameron, 1930a: 154 (Oxytelus; subgenus Anotylus; variety of malayanus; Type locality: Singapore. Selangor; Kuala Lumpur). — Scheerpeltz, 1933: 1099 (Oxytelus; variety of malayanus). manchuricus Bernhauer, 1939b: 75 (Oxytelus; subgenus Anotylus; Type locality: Mand- schurei, Chandaochedzi; Süd-Mandschurei, Chi-Kuan-Shan). — Scheerpeltz, 1962: 576 (Oxytelus; subgenus Anotylus; characters; China). — Herman, 1970: 418 (Anotylus). — J. Li, 1993: 21 (Anotylus; China). DISTRIBUTION: China. 1368 OXYTELINAE: ANOTYLUS Bulletin AMNH marginatus Weise, 1877: 96 (Oxytelus; Type locality: Jedo). — Sharp, 1889: 415 (Oxytelus; notes; Japan). — Bernhauer and Schubert, 1911: 115 (Oxytelus; catalog). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 158 (Anotylus). — Watanabe, 1996a: 9 (Anotylus; cave collection; Japan). DISTRIBUTION: Japan. sharpianus Cameron, 1930c: 184 (Oxytelus; subgenus Anotylus; Type locality: Kagoshima). — Scheerpeltz, 1933: 1104 (Oxytelus; catalog). — Herman, 1970: 419 (Anotylus). — Hammond, 1976: 162 (Anotylus; synonym of marginatus). maritimus Thomson, 1861: 131 (Anotylus; Type locality: Sandhammar i Skåne). — Hardy, 1851: 51 (Oxytelus; cited as flavipes Stephens; [Note: According to Peter Hammond (personal communication), Hardy’s use of flavipes is a misidentification of maritimus (or perrisii); other authors have cited the name as fulvipes Stephens; there was not such name proposed by Stephens in Oxytelus]; notes; England). — Fauvel, 1869: 492 (Oxytelus; synonym of perrisii). — Fauvel, 1872: 169 [= 1873: 11] (Oxytelus; synonym of perrisii). — Fauvel, 1872: 169 [= 1873: 11] (Oxytelus; cited as fulvipes Hardy and Bold; [Note: There was no such name cited in Hardy and Bold; the citation of fulvipes was probably a misspelling of flavipes which was a misidentification of perrisii]; synonym of perrisii). — Seidlitz, 1875: 247 (Oxytelus; characters; Baltic region). — Mulsant and Rey, 1878: 521 (Oxytelus; synonym of perrisii). — Fowler, 1888: 381 (Oxytelus; characters; Britain). — Ganglbauer, 1895: 640 (Oxytelus; synonym of perrisii). — Ganglbauer, 1895: 640 (Oxytelus; cited as fulvipes Hardy and Bold; [Note: There was no such name cited in Hardy and Bold; the citation of fulvipes was probably a misspelling of flavipes which was a misidentification of perrisii]; synonym of perrisii). — Bernhauer and Schubert, 1911: 117 (Oxytelus; synonym of perrisii). — Bernhauer and Schubert, 1911: 117 (Oxytelus; cited as fulvipes Hardy and Bold; [Note: There was no such name cited in Hardy and Bold; the citation of fulvipes was probably a misspelling of flavipes which was a misidentification of perrisii]; synonym of perrisii). — Pope, 1977: 25 (Anotylus; cited as valid species; Britain). — Lohse and Lucht, 1989: 133 (Anotylus; note). — Welch, 1993: 227 (Anotylus; cited as valid species; ovariole number and ovary structure). — Owen, 1997a: 148 (Anotylus; cited as valid species; collecting sites; Ireland). perrisii Fauvel, 1862b: xxxiv (Oxytelus; Type locality: normannis littoribus). — Fauvel, 1862: 42 (Oxytelus; cited as perrisii; cited as new species; characters; France). — Fauvel, 1872: 169 [= 1873: 11] (Oxytelus; cited as perrisi; characters; France; Scandinavia; Britain; Germany). — Mulsant and Rey, 1878: 521 (Oxytelus; subgenus Anotylus; characters; France). — Fowler, 1888: 381 (Oxytelus; synonym of maritimus). — Ganglbauer, 1895: 640 (Oxytelus; cited as perrisi; subgenus Anotylus; characters; coast of North and West Seas; Atlantic coast of Britain and France). — Fauvel, 1902b: 65 (Oxytelus; cited as perrisi; Tunisia). — Reitter, 1909: 171 (Oxytelus; subgenus Anotylus; cited as perrisi; characters; Germany). — Bernhauer and Schubert, 1911: 117 (Oxytelus; cited as perrisi; catalog). — Johansen, 1914: 561 (Oxytelus; subgenus Anotylus; characters; Denmark). — Portevin, 1929: 417 (Oxytelus; cited as perrisi; characters; France). — C. Koch, 1932: 652 (Oxytelus; subgenus Anotylus; cited as perrisi; characters in key). — Scheerpeltz, 1933: 1101 (Oxytelus; cited as perrisi; catalog). — Palm, 1953: 11 (Oxytelus; notes; Sweden). — Larsson and Gígja, 1959: 74 (Oxytelus; cited as perrisi; biological notes; Iceland). — Palm, 1961a: 33 (Oxytelus; characters; Sweden; Denmark; Norway). — Horion, 1963: 228 (Oxytelus; subgenus Anotylus; cited as perrisi; Sweden; Denmark; Germany; Norway; Britain; Holland; Belgium; France; Sicily; Tunisia). Bulletin AMNH OXYTELINAE: ANOTYLUS 1369
— Lohse, 1964: 85 (Oxytelus; subgenus Anotylus; cited as perrisi; characters; central Europe). — Herman, 1970: 419 (Anotylus; cited as perrisi). — Pope, 1977: 25 (Anotylus; cited as perrisi; cited date of publication for species as 1862; synonym of maritimus). — Lucht, 1987: 93 (Oxytelus; checklist; Germany; Benelux; Denmark; Sweden). — M. Hansen, 1996: 99 (Anotylus; cited as perrisi; synonym of maritimus). oceanus Fauvel, 1862: 292 (Oxytelus; Type locality: Olimae). — Fauvel, 1872: 169 [= 1873: 11] (Oxytelus; synonym of perrisii). — Mulsant and Rey, 1878: 521 (Oxytelus; synonym of perrisii). — Fowler, 1888: 381 (Oxytelus; synonym of maritimus). — Ganglbauer, 1895: 640 (Oxytelus; synonym of perrisii). — Bernhauer and Schubert, 1911: 117 (Oxytelus; variety of perrisii). marmoratus Cameron, 1950a: 181 (Oxytelus; subgenus Anotylus; Type locality: Kenya Colony: Molo, 9000 feet). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Kenya. mashonensis Bernhauer, 1934h: 491 (Oxytelus; subgenus Anotylus; Type locality: Mashona- land: Salisbury). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Zimbabwe. masuriensis Cameron, 1930, see: subsculpturatus Cameron, 1928. maxillosus Sperk, 1835, see: inustus Gravenhorst, 1806. meccii Abdullah and Qadri, 1970: 129 (Neoplatystethus; subgenus Pseudoplatystethus; Type locality: W. Pakistan: Karachi). — Hammond, 1976: 158 (Anotylus). DISTRIBUTION: Pakistan. medius Cameron, 1941c: 491 (Delopsis; Type locality: Philippines: Luzon: Balbalan). — Herman, 1970: 418 (Anotylus). DISTRIBUTION: Philippines. megacephalus Fauvel, 1904b: 99 (Oxytelus; Type locality: Sikkim: Darjeeling; Bengale occid.: Kurseong). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Cameron, 1930: 241 (Oxytelus; subgenus Anotylus; characters; Sikkim; Bengal; India). — Scheerpeltz, 1933: 1099 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — Abdullah and Qadri, 1970: 121, 124 (Oxytelus; characters; East and West Pakistan). — Hammond, Morgan, and Morgan, 1979: 216 (Anotylus; gibbulus species group). — Coiffait, 1984: 118 (Anotylus; Sikkim). DISTRIBUTION: Pakistan, India, Bangladesh. melas Fauvel, 1877: 196 (Oxytelus; Type locality: Tasmanie). — Fauvel, 1878g: 492 (Oxytelus; characters). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Australia. m-elevatus Lea, 1906, see: latiusculus Kraatz, 1859. mendus Herman, 1970: 419 (Anotylus; replacement name for opacus Kraatz). — Lohse and Lucht, 1989: 133 (Anotylus). DISTRIBUTION: Russia, Romania, Hungary, Slovakia, Austria. opacus Kraatz, 1857d: 851 (Oxytelus; [preoccupied]; Type locality: Wien). — Redtenbacher, 1857: 990 (Oxytelus; characters; Austria). — Redtenbacher, 1874: 252 (Oxytelus; characters; Austria). — Mulsant and Rey, 1878: 492 (Oxytelus; characters; Austria). — Ganglbauer, 1895: 637 (Oxytelus; characters; Austria; Hungary; southern Russia). 1370 OXYTELINAE: ANOTYLUS Bulletin AMNH
— Reitter, 1909: 170 (Oxytelus; subgenus Oxytelus; characters; Germany). — Bernhauer and Schubert, 1911: 117 (Oxytelus; catalog). — C. Koch, 1932: 656 (Oxytelus; characters in key). — Scheerpeltz, 1933: 1101 (Oxytelus; catalog). — Horion, 1963: 221 (Oxytelus; subgenus Styloxys; Russia; Romania; Hungary; Slovakia; Austria). — Lohse, 1964: 84 (Oxytelus; subgenus Styloxys, cited as Styloxis; characters; central Europe). — Herman, 1970: 419 (Anotylus; synonym of mendus). — Tóth, 1987: 228 (Anotylus; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Czechoslovakia; Austria). — Bohácˇ, 1993: 43 (Oxytelus; subgenus Styloxys, cited as Styloxis; cited as valid name; Slovakia). methneri Bernhauer, 1937b: 582 (Delopsis; Type locality: N. Uluguru, 1400–1900 m). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Tanzania. micans Kraatz, 1859: 175 (Oxytelus; Type locality: Ceylan). — Fauvel, 1904a: 46 (Oxytelus; [Note: See Oxytelus puncticeps for other species reported by Fauvel but that are misidentifications according to Hammond, 1975: 151]; Nilghiris; Ceylon; Sikkim). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Cameron, 1930: 228 (Oxytelus; subgenus Tanycraerus; characters; India). — Scheerpeltz, 1933: 1099 (Oxytelus; catalog). — Paulian, 1941: 168 (Oxytelus; larval characters; India). — Herman, 1970: 410 (Oxytelus). — Abdullah and Qadri, 1970: 122, 124 (Oxytelus; characters; East and West Pakistan). — Hammond, 1975: 157, 164 (Anotylus; notes; characters; Ceylon; India). — Hammond, 1976: 158 (Anotylus). — Coiffait, 1984: 118 (Tanycraerus, cited as Tanycerus; Pakistan; India). DISTRIBUTION: Pakistan, India, Bangladesh, Sri Lanka. foetidus Cameron, 1930: 230 (Oxytelus; subgenus Tanycraerus; Type locality: India; Bihar; Ceylon). — Scheerpeltz, 1933: 1096 (Oxytelus; catalog). — Herman, 1970: 418 (Anotylus). — Abdullah and Qadri, 1970: 122 (Oxytelus; cited as faetidus; characters; West Pakistan). — Hammond, 1975: 164 (Anotylus; synonym of micans). micropterus Lea, 1906: 203 (Oxytelus; Type locality: Hobart, Tasm.). — Lea, 1910a: 139 (Oxytelus; with Iridomyrmex gracilis; Australia). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Australia. micros Scheerpeltz, 1974: 70 (Oxytelus; subgenus Anotylus; Type locality: Natal: Royal Natal National Park, The Cascades, 5200 ft.). — Hammond, 1976: 159 (Anotylus). DISTRIBUTION: South Africa. miles Cameron, 1928d: 101 (Oxytelus; subgenus Anotylus; Type locality: Sumatra: Fort de Kock). — Scheerpeltz, 1933: 1100 (Oxytelus; catalog). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Indonesia. militaris Bernhauer, 1936b: 86 (Oxytelus; Type locality: Mindanao: Surigao). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Philippines. mimulus Sharp, 1874: 94 (Oxytelus; Type locality: Nagasaki). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Cameron, 1949b: 461 (Oxytelus; Japan). Bulletin AMNH OXYTELINAE: ANOTYLUS 1371
— Nakane, 1963a: 84 (Oxytelus; characters; habitus photograph; Japan). — Herman, 1970: 419 (Anotylus). — Hammond, 1976: 175 (Anotylus; notes; adventive species; China; Siberia; Chile; Illinois). — Yuh, Paik, Kwon, and Lee, 1985: 228 (Anotylus; cited as minulus; Korea). — Watanabe, 1993d (Anotylus; Sarushima Island). — J. Li, 1993: 21 (Anotylus; cited as minulus; China). — Hammond, 1998: 275 (Anotylus; New Jersey). DISTRIBUTION: Japan, Korea, China, Russia, Chile, USA. minarzi Bernhauer, 1936, see: bernhaueri Ganglbauer, 1898. minimus Erichson, 1840, see: fragilis Sharp, 1887. minor Cameron, 1928, see: uncifer Fauvel, 1905. minutus Cameron, 1929b: 444, 445 (Oxytelus; Type locality: Pulo Condore. Penang). — Scheerpeltz, 1933: 1100 (Oxytelus; catalog). — Bernhauer, 1936b: 85 (Oxytelus; Philippines). — Herman, 1970: 419 (Anotylus). — Hammond, 1975: 158, 160 (Anotylus; notes; characters; Ceylon; Nepal; Malaya; Vietnam). DISTRIBUTION: Sri Lanka, Nepal, Malaysia, Vietnam, Philippines. trivialis Cameron, 1940b: 183 (Oxytelus; subgenus Anotylus; Type locality: Annam: Saigon). — Herman, 1970: 421 (Anotylus). — Hammond, 1975: 160 (Anotylus; synonym of minutus). minutus Cameron, 1946, see: spinicornis Hammond, 1975. miriceps Fauvel, 1908: 96 (Oxytelus; Type locality: Amani). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Eichelbaum, 1913: 119 (Oxytelus; Tanzania). — Bernhauer, 1931d: 568 (Oxytelus; Abyssinia). — Cameron, 1950g: 16 (Oxytelus; subgenus Anotylus; Belgian Congo). — Herman, 1970: 419 (Anotylus). — Hammond, Morgan, and Morgan, 1979: 217 (Anotylus; gibbulus species group). DISTRIBUTION: Congo, Tanzania, Ethiopia. mirus Bernhauer, 1915d: 101 (Oxytelus; Type locality: Deutsch-Ostafrika: Kilimandscharo). — Scheerpeltz, 1933: 1100 (Oxytelus; catalog). — Cameron, 1950c: 92 (Oxytelosus?). — Cameron, 1950g: 16 (Oxytelus; subgenus Anotylus; Belgian Congo). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Tanzania, Congo. misellus Cameron, 1950g: 16 (Oxytelus; subgenus Anotylus; Type locality: Belgian Congo: volc. Nyamuragira, Mayumbu, alt. 2100 m). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Congo. mixtus Bernhauer, 1936b: 83 (Oxytelus; Type locality: Luzon: Bangui). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 159 (Anotylus). DISTRIBUTION: Philippines. modestus Bernhauer, 1936b: 84 (Oxytelus; Type locality: Luzon: Los Baños und Bayombong). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Philippines. monodon Fauvel, 1905a: 130 (Oxytelus; Type locality: Bolivie: territoire des Yuracaris; Colombie: Paramba, 1150 m). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Bolivia, Colombia. 1372 OXYTELINAE: ANOTYLUS Bulletin AMNH monticola Cameron, 1942 (July), see: currus Herman, 1970. monticola Cameron, 1942b(May): 323 (Oxytelus; subgenus Anotylus; Type locality: Chyulu Hills, altitude 5600–6600 feet). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Kenya. montivagus Heer, 1841, see: sculpturatus Gravenhorst, 1806. morbosus Cameron, 1942, see: subsculpturatus Cameron, 1928. mortuorum Bernhauer, 1934h: 491 (Oxytelus; subgenus Anotylus; Type locality: Belgian Congo: 18 miles S. W. of Elizabethville). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Congo. munitus Casey, 1894: 385 (Oxytelus; Type locality: Pennsylvania). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Herman, 1970: 419 (Anotylus). — Downie and Arnett, 1996: 448 (Anotylus; characters; USA). DISTRIBUTION: USA. mutator Lohse, 1963: 174 (Oxytelus; subgenus Anotylus; Type locality: Garlstorfer Forst, Kreis Harburg, Niedersachsen). — Lohse, 1964: 85 (Oxytelus; subgenus Anotylus; characters; central Europe). — Hammond, 1968 (Oxytelus; characters; Britain). — Herman, 1970: 410 (Oxytelus). — Korge, 1971a: 6 (Oxytelus; Turkey). — Tóth, 1974: 222, 223 (Oxytelus; characters; Hungary). — Hammond, 1976: 159 (Anotylus). — Pope, 1977: 25 (Anotylus; Britain). — Tóth, 1987: 228 (Anotylus; Carpathian Basin). — Lucht, 1987: 93 (Oxytelus; checklist; Germany; Poland; Austria; Denmark). — Lohse and Lucht, 1989: 133 (Anotylus; note). — Bohácˇ, 1993: 43 (Anotylus; Czech Republic; Slovakia). — Hodge and Jones, 1995: 33 (Anotylus; characters; Britain). — Ciceroni and Zanetti, 1995: 12 (Anotylus; Italy). — Vorst and Huijbregts, 1998 (Anotylus; characters; collecting notes; Netherlands). — Holzer, 1998: 37 (Anotylus; notes; Austria). — Baubet, Tronquet and Callot, 1999 (Anotylus; notes; France). DISTRIBUTION: Britain, France, Netherlands, Denmark, Germany, Poland, Czech Republic, Slovakia, Hungary, Austria, Italy, Turkey. myops Fauvel, 1877: 199 (Oxytelus; Type locality: Australie, Somerset). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 159 (Anotylus). DISTRIBUTION: Australia. myrmecophilus Cameron, 1914: 526 (Oxytelus; subgenus Anotylus; Type locality: S. India, Nilgiri Hills). — Cameron, 1930: 246 (Oxytelus; subgenus Anotylus; characters; India). — Scheerpeltz, 1933: 1100 (Oxytelus; catalog). — Scheerpeltz, 1962: 570 (Oxytelus; subgenus Anotylus; characters; China; India). — Abdullah and Qadri, 1970: 121 (Oxytelus; characters; West Pakistan). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Pakistan, India, China. nanus Erichson, 1840: 797 (Oxytelus; Type locality: America septentrionalis). — LeConte, 1877: 234, 238 (Oxytelus; characters; Carolina). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Notman, 1920b: 695 (Oxytelus; Florida). Bulletin AMNH OXYTELINAE: ANOTYLUS 1373
— Herman, 1970: 419 (Anotylus). — Downie and Arnett, 1996: 449 (Anotylus; characters; USA). DISTRIBUTION: USA. neotomae Hatch, 1957: 96 (Oxytelus; subgenus Anotylus; Type locality: Oregon: Corvallis). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: USA. nevesi Scheerpeltz, 1951, see: plagiatus Rosenhauer, 1856. niger LeConte, 1877: 235 (Oxytelus; Type locality: San Francisco, California; British Columbia). — Casey, 1894: 392 (Oxytelus; characters; California; British Columbia). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Hatch, 1957: 97 (Oxytelus; characters; British Columbia). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: USA, Canada. nigripennis Bernhauer, 1936b: 82 (Oxytelus; Type locality: Mindanao: Surigao und Momungan; Luzon: Los Baños). — Herman, 1970: 419 (Anotylus). DISTRIBUTION: Philippines. niobe Fernando, 1959: 95 (Oligota; Type locality: Ceylon: Dambulla). — S. Williams, 1978: 168, 169 (Anotylus; notes). DISTRIBUTION: Sri Lanka. nitens Fauvel, 1877, see: ebonus Blackwelder, 1944. nitescens Bernhauer, 1942: 5 (Oxytelus; subgenus Tanycraerus Type locality: Kostarika: Vara Blanca, 1700 m). — Herman, 1970: 410 (Oxytelus). — Hammond, 1976: 159 (Anotylus). DISTRIBUTION: Costa Rica. nitidifrons Wollaston, 1871a: 411 (Oxytelus; Type locality: St. Helena). — Wollaston, 1877: 36 (Oxytelus; characters; St. Helena). — Fauvel, 1898a: 115 (Oxytelus; Seychelles). — Fauvel, 1902b: 65 (Oxytelus; Madeira). — Fauvel, 1904a: 45 (Oxytelus; Nilghiris; Burma; Philippines; Java; Seychelles; Mauritia; Réunion; Zanzibar; Gabon; Saint Helena; Madeira; Hawaii). — H. Kolbe, 1910: 20 (Oxytelus; Seychelles). — Bernhauer and Schubert, 1911: 116 (Oxytelus; catalog). — Cameron, 1921a: 367, 400 (Oxytelus; characters; Singapore). — Cameron, 1930: 256 (Oxytelus; subgenus Anotylus; characters; India; Burma). — Scheerpeltz, 1933: 1100 (Oxytelus; catalog). — Bernhauer, 1936b: 82 (Oxytelus; Philippines). — Méquignon, 1942a: 17 (Oxytelus; subgenus Anotylus; Azores). — Fagel, 1956b: 273 (Microxytelus). — Jarrige, 1963a: 120 (Microxytelus; India). — Smetana, 1963b: 32 (Oxytelus; subgenus Anotylus; Madeira). — Herman, 1970: 419 (Anotylus). — Abdullah and Qadri, 1970: 121 (Oxytelus; characters; West Pakistan). — Smetana, 1970: 54 (Oxytelus; distributional summary; Azores). — Fagel, 1972: 99 (Microxytelus; notes; Sainte Helene). — Hammond, 1975: 158, 164 (Anotylus; notes; characters; Ceylon; China; Japan; India; Laos; Java; Vietnam; Mauritius; Madagascar; Madeira; St. Helena; Hawaii; Palau Islands). — Hammond, 1976: 176 (Anotylus; notes; adventive species). — Moore, 1978: 489 (Anotylus; Hawaii). — Borges, 1990: Table II (Anotylus; Azores). — Nishida, 1994: 70 (Anotylus; Hawaii). 1374 OXYTELINAE: ANOTYLUS Bulletin AMNH