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1758 to the End of the Second Millennium. Iii. Oxyteline Group

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1758 to the End of the Second Millennium. Iii. Oxyteline Group CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM. III. OXYTELINE GROUP LEE H. HERMAN BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY NUMBER 265 NEW YORK : 2001 Recent issues of the Bulletin may be purchased from the Museum. Lists of back issues of the Bulletin, Novitates, and Anthropological Papers published during the last five years are available at World Wide Web site http://nimidi.amnh.org. Or address mail orders to: American Museum of Natural History Library, Central Park West at 79th St., New York, N.Y. 10024. TEL: (212) 769- 5545. FAX: (212) 769-5009. E-MAIL: [email protected] CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM. III. OXYTELINE GROUP LEE H. HERMAN Curator, Division of Invertebrate Zoology American Museum of Natural History BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Number 265, pages 1067–1806 Issued July 18, 2001 (Parts I–VII) Copyright © American Museum of Natural History 2001 ISSN 0003-0090 CATALOG OF THE STAPHYLINIDAE, PARTS I–VII PART I, pages 1–650 Introduction Brief History of Taxonomic Studies of the Staphylinidae Research History • Biographical Sketches Overview of Classification Omaliine Group Dasycerinae • Empelinae • Glypholomatinae • Micropeplinae • Microsilphinae • Neophoninae • Omaliinae • Proteininae • Protopselaphinae PART II, pages 651–1066 Tachyporine Group Habrocerinae • Olisthaerinae • Phloeocharinae • Tachyporinae • Trichophyinae PART III, pages 1067–1806 Oxyteline Group Apateticinae • Osoriinae • Oxytelinae • Piestinae • Trigonurinae PART IV, pages 1807–2440 Staphylinine Group Euaesthetinae • Leptotyphlinae • Megalopsidiinae • Oxyporinae • Pseudopsinae • Solieriinae • Steninae PART V, pages 2441–3020 Staphylinine Group, continued Staphylininae Diochini • Maorothiini • Othiini • Platyprosopini • Staphylinini (Amblyopinina • Hyptiomina • Anisolinina • Philonthina) PART VI, pages 3021–3840 Staphylinine Group, continued Staphylininae, continued Staphylinini (Quediina • Staphylinina • Tanygnathinina • Xanthopygina • Incertae Sedis) • Xantholinini Staphylinidae: Subfamily Incertae Sedis Protactinae† PART VII, pages 3841–4218 Bibliography Index iii CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM ABSTRACT This catalog (published in seven parts, all released on the same day) is based on only the published lit- erature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylin- inae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or descrip- tion are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or syn- onyms are listed for each valid name. Annotated subsequent references are presented for all names. Dis- tributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Alesˇ Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies. Address correspondence to: Lee H. Herman, Curator, Division of Invertebrates, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA ([email protected]). Please send comments, corrections, or updates to either the postal or e-mail address. Reprints of new publications would be helpful and appreciated. iv CONTENTS PART III OXYTELINE GROUP Oxyteline Group . 1067 Apateticinae . 1067 Osoriinae . 1071 Eleusinini . 1072 Leptochirini . 1094 Osoriini . 1145 Thoracophorini . 1230 Clavilispinina . 1231 Glyptomina . 1239 Lispinina . 1250 Thoracophorina . 1293 Osoriinae: Tribe Incertae Sedis . 1305 Oxytelinae . 1307 Coprophilini . 1308 Deleasterini . 1321 Oxytelini . 1332 Thinobiini . 1491 Oxytelinae: Tribe Incertae Sedis . 1778 Piestinae . 1781 Trigonurinae . 1803 PARTS I, II, IV–VII Introduction, History, and Biographical Sketches . Part I Omaliine Group . Part I Tachyporine Group . Part II Staphylinine Group . Parts IV–VI Staphylinidae: Subfamily Incertae Sedis . Part VI Protactinae† . Part VI Bibliography . Part VII Index . Part VII v APATETICINAE [2 genera; 24 species; Oriental and Palaearctic Regions] Apateticinae Fauvel, 1895. — Newton and Thayer, 1992: 55 (summary of current classification). — Lawrence and Newton, 1995: 827 (discussion). — M. Hansen, 1996: 33, 36 (relatively basal position within superfamily; listed in Oxyteline group of subfamilies but should probably be omitted from family). Apateticae Fauvel, 1895: 190 (genera included: Apatetica, Trygaeus). TYPE GENUS: Apa- tetica (see Newton and Thayer, 1992: 55). — M. Hansen, 1997: 78, 79, 80, 135, 166, 180 (characters supporting clade; autapomorphic features; description; notes). Apateticini Fauvel, 1895. — Eichelbaum, 1909: 115 (list of genera with some references; generic synonyms; number of genera; number of species in each genus; general distribution of genera). — Newton, 1982a: 14 (larval characters). Apatetica [19 species; Oriental and Palaearctic Regions] Apatetica Westwood, 1848: 86 (species included: lebioides). TYPE SPECIES: Apatetica lebioides Westwood, fixed by monotypy. — Fauvel, 1895: 190 (characters; notes). — Bernhauer and Schubert, 1910: 4 (world catalog; 12 species). — Winkler, 1925: 323 (catalog for Palaearctic region). — Cameron, 1925: 1 (catalog of Indian species). — Cameron, 1930: 26 (characters; key to species of British India). — Scheerpeltz, 1933: 992 (world catalog supplement). — Jansson, 1947: 1 (characters; discussion). — Blackwelder, 1952: 411 (type species: lebioides). — Abdullah and Qadri, 1970: 115 (cited as Apatelica; key to species of East Pakistan). — Tikhomirova, 1973a: 142 (checklist of species of USSR). — Shibata, 1976: 76 (checklist of species of Japan). — Lawrence and Newton, 1995: 827 (Apateticinae). Trygaeus Sharp, 1874a: 420 (species included: princeps). TYPE SPECIES: Trygaeus princeps Sharp, fixed by monotypy. — Duvivier, 1883: 203 (catalog; cited as Trigaeus). — Lewis, 1893a: 248 (synonym of Apatetica). — Fauvel, 1895: 190 (synonym of Apatetica). — Bernhauer and Schubert, 1910: 4 (synonym of Apatetica). — Cameron, 1930: 26 (synonym of Apatetica). — Jansson, 1947: 5 (characters; may be valid genus or synonym of Nodynus). — Blackwelder, 1952: 414 (type species: princeps). Idiocheila Frivaldszky, 1883: 135 (species included: spinipennis). TYPE SPECIES: Idiocheila spinipennis Frivaldszky, fixed by monotypy. — Fauvel, 1895: 190 (synonym of Apatetica). — Bernhauer and Schubert, 1910: 4 (synonym of Apatetica). — Cameron, 1930: 26 (synonym of Apatetica). — Blackwelder, 1952: 412 (type species: spinipennis). birmana Cameron, 1930: 33 (Apatetica; Type locality: Upper Burma: Seinghku Valley, alt. 5000 feet). — Scheerpeltz, 1933: 992 (Apatetica; catalog). L DISTRIBUTION: Myanmar. 1067 1068 APATETICINAE: APATETICA Bulletin AMNH birmanica Jansson, 1947: 7 (Apatetica; Type locality: Kambaiti at 7000 feet). L DISTRIBUTION: Myanmar. brunnipes Ritsema, 1879: 46 (Apatetica; Type locality: Sumatra). — Fauvel, 1895: 192 (Apatetica; characters; Sumatra; Borneo). — Bernhauer and Schubert, 1910: 4 (Apatetica; catalog; Sumatra, Borneo). — Cameron, 1928d: 90 (Apatetica; Sumatra). L DISTRIBUTION: Sumatra, Borneo. caeruleipennis Cameron, 1930: 32 (Apatetica; Type locality: Manipur). — Scheerpeltz, 1933: 992 (Apatetica; catalog). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). L DISTRIBUTION: India, Bangladesh. callosa Jansson, 1947: 10 (Apatetica; Type locality: Kambaiti, 6000 feet). L DISTRIBUTION: Myanmar. indica Cameron, 1930: 29 (Apatetica; Type locality: Bhutan). — Scheerpeltz, 1933: 992 (Apatetica; catalog). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). L DISTRIBUTION: Bhutan, Bangladesh. intermedia Cameron, 1930: 30 (Apatetica; Type locality: Burma: Ruby Mines; Bengal). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). L DISTRIBUTION: Myanmar, India, Bangladesh. javanica Sharp, 1892: 61 (Trygaeus; Type locality: Mount Poentjak, West Java). — Fauvel, 1895: 194 (Apatetica; characters; Bengal; Burma; Java). — Bernhauer and Schubert, 1910: 4 (Apatetica; catalog; Java; East Indies). — Cameron, 1930: 29 (Apatetica; characters; Assam; Bengal). — Scheerpeltz, 1933: 993 (Apatetica; catalog). — Abdullah and Qadri, 1970: 115 (Apatetica; characters; East Pakistan). L DISTRIBUTION: India, Bangladesh, Java, Sumatra. sumatrana Cameron, 1930d: 326 (Apatetica; variety of javanica; Type locality: Gunung Singgalang). — Scheerpeltz, 1933: 993 (Apatetica; variety of javanica). laevicollis Fauvel, 1904b: 86 (Apatetica; Type locality: Tonkin central: Chiem-Hoa). — Bernhauer and Schubert, 1910: 4 (Apatetica; catalog; Tonkin). L DISTRIBUTION:
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