Remarks on Inape Razowski, 1988 from Ecuador with Description of 21 New Species (Lepidoptera: Tortricidae: Euliini)

Nachr. entomol. Ver. Apollo, N. F. 27 (3): 115–130 (2006) 115

Remarks on Inape Razowski, 1988 from Ecuador with description of 21 new species (Lepidoptera: Tortricidae: Euliini)

Józef Razowski and Volker Pelz Prof. Dr. Józef Razowski, Institute of Systematics and Evolution of Animals PAS, Slawkowska 17, PL-31 016 Kraków, Poland; [email protected] Dipl.-Biol. Volker Pelz, Bonnenweg 3, D-53809 Ruppichteroth, Germany; [email protected]

Abstract. The Neotropical genus Inape Razowski, 1988 is species and reassessed the characteristics of this genus. characterized, a list of the known species is provided, 21 new The number of the species of Inape increased thus to 13. species and 2 new subspecies are described from Ecuador: They also characterized Tylopeza Razowski, 1995 as the Inape extraria sp. n. (HT ♂), Inape uncina sp. n. (HT ♂), Inape luteina sp. n. (HT ♂), Inape elegans sp. n. (HT ♂), possible sister-genus of Inape. Inape incarnata sp. n. (HT ♂), Inape epiphanes sp. n. (HT In extensive Euliini material recently collected in Ecua ♂), Inape pompata sp. n. (HT ♂), Inape laterosclera sp. n. dor, mainly on a collecting trip of the second author and (HT ♂), Inape asymmetra sp. n. (HT ♂), Inape commoda sp. n. (HT ♂), Inape homora sp. n. (HT ♀), Inape porpax his wife together with Siska and Dr. Cees Gielis, many sp. n. (HT ♀), Inape cateres sp. n. (HT ♂), Inape eparmuncus new taxa of Inape were found and the number of Inape sp. n. (HT ♂), Inape homeotypa sp. n. (HT ♂), Inape homo species increased now to 34. This new material enabled loga sp. n. (HT ♂), Inape polysparta sp. n. (HT ♀), Inape us to re-examine some generic characters. celypha sp. n. (HT ♀), Inape pseudocelypha sp. n. (HT ♀), Inape geoda sp. n. (HT ♀), Inapes ororia sp. n. (HT ♂), Inape Externally the species of this genus show a distinct sororia coryssa ssp. n. (HT ♂), Inape sororia lojana ssp. n. variation in colouration and markings. A plesiomorphic (HT ♂). The holotypes of the new species are presently in pattern characterizes with more or less preserved trans the collection of Volker Pelz, Ruppichteroth, Germany, and verse markings occasionally combined with an additio eventually will be depoited in the Senckenberg-Museum, nal radial element extending along the distal part of the Frankfurt am Main, Germany. costal edge of the median cell and reaching the apex of Keywords: Andes, cloud forest, high elevation. the wing. The median fascia may be followed by a strong Anmerkungen zur Gattung Inape Razowski, 1988 suffusion reaching the median part of the termen. In aus Ecuador mit Beschreibung von 21 neuen Arten some species in the basal area of the forewing a blotch (Lepidoptera: Tortricidae: Euliini) with an oblique distal edge develops, whose inner area is very often pale. This pattern shows up also in other Zusammenfassung: Die in der Neotropis verbreitete Gattung Inape Razowski, 1988 wird charakterisiert, und die bekannten Neotropical genera of Euliini. The dorsal part of this Arten werden aufgelistet. Aus Ecuador werden 21 Arten und blotch is occasionally reduced to a postbasal fascia which 2 Unterarten neu beschrieben. In der Liste der neuen Taxa is either complete or preserved only in the dorsal area of im englischen Abstract wird in Klammern das Geschlecht des the wing. In some species postmedian and subterminal Holotypus angegeben (siehe Abstract). Die Holotypen der markings are preserved or a subtornal elongate mark neuen Taxa befinden sich zur Zeit in der Sammlung Volker appears. In further species the particular elements may Pelz, Ruppichteroth, Deutschland, und werden letztlich an das Forschungsinstitut und Naturmuseum Senckenberg, fuse or disappear. A sexual dimorphism expressed in the Frankfurt am Main, Deutschland, gelangen. forewing pattern is found in Inape sororia sp. n. in which the ♂♂ have a complete basal blotch while the ♀♀ show Observaciones sobre el género Inape Razowski, 1988 well developed postmedian elements. de Ecuador con la descripción de 21 nuevas especies (Lepidoptera: Tortricidae: Euliini) In the ♂ genitalia a new character found is the presence of a dorsobasal spine-like process of the sacculus; in Resumen: Se dan las características del género Neotropical Inape Razowski, 1988 y se da una lista de las especies cono some species the sacculus is angulated distally, often cidas. Se describen del Ecuador 21 nuevas especies y dos accompanied by a row of spines; similar spines may be subespecies. En la lista de las taxas nuevas en el resumen present in species with simple, not angulate sacculi. inglés se da el sexo del holotipo en paréntesis. Los holotipos Usually the uncus is slender but in a few species (e.g. de las especies nuevas están en la colección Volker Pelz, Rup I. penai Razowski, 1988) is strong or short, very broad pichteroth, Alemania, determinados ultimamente para el postbasally or submedially. Only exceptionally the uncus Senckenberg-Museum, Frankfurt am Main, Alemania. is expanding terminally (e.g. I. extraria sp. n.). The num ber of cornuti is species-specific and is variable in some Introduction species as well, e.g. I. sororia sp. n. and I. pompata sp. n. Inape Razowski, 1988 was described to comprise two Neo ♀ genitalia. The sterigma is broad, with a more or less tropical species. Then Brown (1989) included further two distinct anteostial sclerite. The ductus bursae is broad, species in this genus and Powell et al. (1995) transferred membranous or provided with various sclerites, trans Eulia biremis Meyrick, 1926 to Inape. Razowski (1997, verse or longitudinal placed either in its proximal part 1999) described three additional species from Peru and or in the collicular portion. There are two long sclerites, Ecuador in Inape. Recently Brown & Razowski (2003) one (called here the lateral) with small marginal spines, reviewed the preceding studies, described further five and the other one long, slender, armed with strong spi

© Entomologischer Verein Apollo e. V., Frankfurt am Main 116 nes occurring in innumerous species (named the spiny Acknowledgments sclerite; cf. illustrations). Exceptionally there occurs a We are grateful to Prof. Giovanni Onoré, Pontificia Uni rounded sclerotic plate with strong marginal spines. versidad Católica del Ecuador, Quito, for his efforts and The systematic position of the genus remains unclear. assistance in obtaining permits for collecting. The most closely related genus is Tylopeza Razowski, Further we thank Dr. Wolfgang A. Nässig, Frankfurt am 1995. The only autapomorphy of Inape is still the struc Main, for editing the manuscript and providing helpful ture of the transtilla armed with a pair of dorsosubme comments. dian lobes distinctly separated from one another by means of a concave, well sclerotized median portion. The second author expresses his sincere gratitude to Siska and Dr. Cees Gielis, Lexmond, and Margarita Pelz, Systematic arrangement of species: We could not find Ruppichteroth, for their companionship on the collecting any constant correlation between the character states of trip in autumn 2002 to Ecuador. particular genital or external structures. It is, however, possible to distinguish some groups of species. Thus Systematic part essentially we follow the arrangement by Brown & Razow ski (2003). List of species of Inape (With the country of the type locality provided. New taxa in bold Distribution: All hitherto known species of Inape are dis types.) tributed at high elevations (2000–4000 m a.s.l.) of the Andes from Colombia to Bolivia with only two species Inape Razowski (1988, Acta zool. Cracov. 31: 394). already recorded from Ecuador. The new species from (Type species: Inape penai Razowski, 1988) Ecuador were all found from 1700 m to 4000 m in cloud I. extraria sp. n. — Ecuador forest habitats and confirm therefore a presumable restric I. auxoplaca (Meyrick, 1926) — Colombia tion of the genus to higher elevations of the Andes. I. uncina sp. n. — Ecuador To date nothing is known about their biology and their I. luteina sp. n. — Ecuador immature stages. I. biremis (Meyrick, 1926) — Colombia The holotypes of the new species are at present in the I. papallactana Razowski, 1999 — Ecuador collection of Volker Pelz, Ruppichteroth, Germany, and eventually will be deposited in the Senckenberg-Museum, I. bicornis Razowski, 1999 — Ecuador Frankfurt am Main, Germany. I. elegans sp. n. — Ecuador This publication is the seventh on Ecuadoran Euliini I. commoda sp. n. — Ecuador after Razowski & Pelz (2003) (see complete list of papers I. incarnata sp. n. — Ecuador in the references). I. epiphanes sp. n. — Ecuador Note. Numbers included in descriptions of the labial pal I. pompata sp. n. — Ecuador pus refer to the proportion of their total length to the I. laterosclera sp. n. — Ecuador horizontal diameter of the compound eye. Abbreviations and terms: I. asymmetra sp. n. — Ecuador > road from > to. I. homora sp. n. — Ecuador CVPR Collection Volker Pelz, Ruppichteroth, Germany. I. porpax sp. n. — Ecuador GU Genitalia slide no. I. semuncus Razowski, 1997 — Peru HT Holotype(s). ISEZ Institute of Systematics and Evolution of Animals PAS, I. cateres sp. n. — Ecuador Kraków, Poland. I. eparmuncus sp. n. — Ecuador N, E, S, W compass points. I. penai Razowski, 1988 — Bolivia PAS Polish Academy of Sciences. I. homeotypa sp. n. — Ecuador Phallus formerly aedeagus (aedoeagus); compare Kristensen (2003: 103). I. homologa sp. n. — Ecuador PN National Park. I. circumsetae Brown & Razowski, 2003 — Colombia Prov. Province. I. centrota Brown & Razowski, 2003 — Colombia PT Paratype(s). I. polysparta PUCE Museo de Zoología, Centro de Biodiversidad y Ambi sp. n. — Ecuador ente, Pontificia Universidad Católica del Ecuador, I. reductana Brown & Razowski, 2003 — Peru Quito, Ecuador. I. celypha sp. n. — Ecuador SMFL Lepidoptera collection of Forschungsinstitut und Natur museum Senckenberg, Frankfurt am Main, Germany. I. pseudocelypha sp. n. — Ecuador sta collecting locality. I. sinuata Brown & Razowski, 2003 — Bolivia, new for Stt station. Ecuador

I. geoda sp. n. — Ecuador Diagnosis: The systematic position of I. elegans sp. n. I. clarkeana Brown & Razowski, 2003 — Colombia is somewhat unclear; externally it resembles I. uncina sp. n. but in ♂ genitalia the sacculus of valva is hooked I. sororia sp. n. — Ecuador terminally and the cornuti are straight. I. sororia corryssa ssp. n. — Ecuador I. sororia lojana ssp. n. — Ecuador Description: ♂ (Fig. 18). Wing span 25.5 mm. Head I. xerophanes (Meyrick, 1909) — Peru pale brownish, labial palpus 2.5; thorax rather brown. Forewing fairly broad, termen weakly oblique. Ground I. iantha (Meyrick, 1926) — Colombia colour creamy brownish with weak ferruginous hue and Descriptions of new taxa some darker portions; strigulation and some spots brown. Markings brown consisting of large basal blotch and costal Inape extraria sp. n. half of median fascia extending at the end towards mid- (Figs. 7, 37, 38) termen. Cilia concolorous with ground colour, brown Holotype ♂: “Ecuador: Pichincha-Prov., 1 km W Papallacta- Pass, 3990 m, 0°19'55" S, 78°12'50" W, 22. xii. 2004, leg. between apex and mid-termen. Hindwing whitish, pale Volker Pelz”; GU-2430-V.P. (CVPR, eventually SMFL). ochreous in apex area; strigulation brownish grey; cilia Etymology: The name refers to the systematic position of whitish tinged ochreous towards apex. the species; Latin: extra — outside, -arius — an adjective suf ♂ genitalia (Figs. 43, 44). Uncus as in above mentioned fix. It is defined here as a noun in apposition. species; valva upcurved; sacculus strong terminating in a Diagnosis: External characters of I. extraria sp. n. as in distinct claw-shaped process; processes of transtilla long; numerous other species, e.g. in I. elegans sp. n. or I. incar end of phallus protruding; cornuti two strong spines. nata sp. n., but easily distinguished in connected basal, median and subapical elements of markings; ♂ genitalia ♀ unknown. similar to those of I. papallactana but differing from all Inape incarnata sp. n. known species of Inape in possession of a median lobe of the transtilla situated between the submedian processes (Figs. 14, 45, 46) typical of the genus. Holotype ♂: “Ecuador: Pichincha-Prov., 1 km W Papallacta- Pass, 3990 m, 0°19'55" S, 78°12'50" W, 22. xii. 2004, leg. Description: ♂ (Fig. 7). Wing span 25.0 mm. Head ferru Volker Pelz”; GU-2429-V.P. (CVPR, eventually SMFL). ginous, thorax brownish, brown proximally; labial palpus Etymology: The name refers to the colouration of the fore 3.0, concolorous with head. Forewing weakly expanding wing; Latin: incarnata — flesh coloured. It is defined as a terminally, termen oblique. Ground colour ferruginous; noun in apposition. strigulae and markings brown. Basal blotch indistinct, Diagnosis: Inape incarnata sp. n. is very similar to I. partly fusing with postbasal fascia; this last connecting elegans sp. n. but distinguished by a creamy coloured with median fascia by means of a curved fascia, the latter submedian area of the forewing and a short costal part connected with subapical blotch. Cilia concolorous with of the median fascia; in ♂ genitalia it differs mainly in ground colour. Hindwing whitish cream, tinged ochreous the much longer phallus and a higher number of cornuti at apex; strigulation diffuse, grey; cilia cream. arranged in two groups. ♂ genitalia (Figs. 37, 38). Uncus strongly expanding ter Description: ♂ (Fig. 14). Wing span 26.0 mm. Head minally with weak apical concavity; valva slender; saccu creamy ferruginous, thorax similar with browner marks; lus simple, slender, convex near middle; dorsal part of labial palpus 2.5, cream with ferruginous scales laterally. transtilla with broad, rounded median lobe situated bet Forewing rather slender, costa weakly convex, termen ween short submedian processes; phallus slender, long; moderately oblique. Ground colour creamy suffused one strongly curved at base circular cornutus in vesica. with pale ferruginous along dorsum and in posterior half ♀ not known. of wing; dots and strigulae brown or blackish. Markings blackish consisting of large basal blotch, slender costal Remarks. The systematic position of this species is doubt part of median fascia rather connecting the medio- ful; some characters as broad uncus, simple valva and subdorsal marking and subapical marking. Cilia creamy sacculus are seemingly of a plesiomorphic importance. tinged ferruginous. Hindwing whitish, cream in distal That was a reason to place it at the beginning of the third; cilia cream white. infrageneric system. The structure of the transtilla is, however, specialised. ♂ genitalia (Figs. 45, 46) as in I. elegans sp. n. but valva and sacculus more slender, phallus larger, slenderly Inape elegans sp. n. extending ventro-terminally; cornuti arranged in two (Figs. 18, 43, 44) groups, one group consisting of 6, the other group of 3 Holotype ♂: “Ecuador: Napo-Prov., 5 km W Papallacta, fairly large spines. Laguna Papallacta, 3430 m, 0°22'27" S, 78°9'50"W, 28. x. 2002, sta 40, leg. Gielis & Pelz”; GU-1552-V.P. (CVPR, even ♀ not known. tually SMFL). Etymology: The name refers to the beautiful appearance of Inape uncina sp. n. this large new species; Latin: elegans — elegant. It is defined (Figs. 1, 39, 40) here as a noun in apposition. Holotype ♂: “Ecuador: Zamora-Chinchipe-Prov. 22 km E

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Colour plate 1: Figs. 1–18: Adults of Inape. Fig. 1: I. uncina sp. n., holotype ♂. Fig. 2: I. luteina sp. n., holotype ♂. Fig. 3: I. epiphanes sp. n., holotype ♂. Fig. 4: I. pompata sp. n., holotype ♂. Figs. 5, 6: I. laterosclera sp. n., Fig. 5: holotype ♂, Fig. 6: paratype ♀ (GU-2183-V.P.). Fig. 7: I. extraria sp. n., holotype ♂. Fig. 8: I. asymmetra sp. n., holotype ♂. Fig. 9: I. sinuata Brown & Razowski, 2003, ♀ (GU-2191-V.P.). Fig. 10: I. cateres sp. n., holotype ♂. Fig. 11: I. eparmuncus sp. n., holotype ♂. Fig. 12: I. homeotypa sp. n., holotype ♂. Fig. 13: I. homologa sp. n., holotype ♂. Fig. 14: I. incarnata sp. n., holotype ♂. Fig. 15: I. commoda sp. n., holotype ♂. Figs. 16, 17: I. geoda sp. n., Fig. 16: holotype ♀, Fig. 17: ♀ (GU-1536-V.P.). Fig. 18: I. elegans sp. n., holotype ♂.

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34 35 36

Colour plate 2: Figs. 19–36: Adults of Inape. Fig. 19: I. sororia sp. n., holotype ♂. Fig. 20: I. sororia corryssa ssp. n. holotype ♂. Fig. 21: I. sororia lojana ssp. n., holotype ♂. Fig. 22: I. sororia sp. n., paratype ♂ (GU-2409-V.P.). Fig. 23: I. sororia corryssa ssp. n., paratype ♂ (GU-1631-V.P.). Fig. 24: I. sororia lojana ssp. n., paratype ♂ (GU-2197-V.P.). Fig. 25: I. sororia sp. n., paratype ♀ (GU-1634-V.P.). Fig. 26: I. sororia corryssa ssp. n., paratype ♀ (GU-1703- V.P.). Fig. 27: I. sororia lojana ssp. n., paratype ♀ (GU-1709-V.P.). Fig. 28: I. sororia corryssa ssp. n., paratype ♂ (GU-2194-V.P.). Fig. 29: I. pseudocelypha sp. n., holotype, ♀. Fig. 30: I. celypha sp. n., holotype, ♀. Figs. 31–33: I. homora sp. n., Fig. 31: holotype ♀, Fig. 32: paratype ♀ (GU-1653-V.P.), Fig. 33: paratype ♀ (GU-2266-V.P.). Figs. 34, 35: I. porpax sp. n., Fig. 34: holotype ♀, Fig. 35: paratype ♀ (GU-1596-V.P.). Fig. 36: I. polysparta sp. n., holotype ♀.

Loja, PN Podocarpus, San Francisco Ranger Stt, 2200 m, termen obliquely straight. Ground colour ochreous yel 3°59'15" S, 79°5'37" W, 9. x. 2002, sta 22, leg. Gielis & low suffused pale ferruginous yellow, sparsely sprinkled Pelz”; GU-2185-V.P. (CVPR, eventually SMFL). brownish; venation in major part yellowish. Markings Paratypes (in total 2 ♂♂): 2 ♂♂ (GU-2211-V.P.), same data pale ferruginous partially suffused with brownish consis as HT (CVPR). ting of traces of median and postmedian fasciae repre Etymology: The name refers to the shape of one of the cor nuti; Latin: uncinus — a hook. It is a noun in apposition. sented by elongate, diffuse marks. Cilia concolorous with ground colour. Hindwing whitish cream; apex tinged Diagnosis: In this species the pattern is generalized; in ♂ ochreous; cilia white cream. genitalia the uncus slender; the sacculus similar to that in Ecuadoran I. bicornis Razowski, 1999 and I. semuncus ♂ genitalia (Figs. 41, 42). Uncus as in I. uncina sp. n., Razowski, 1997 from Peru, but I. uncina sp. n. is distinct socius somewhat smaller, processes of transtilla broader; by the broad phallus and the heavily curved cornutus. valva shorter than in mentioned species, sacculus slender See also diagnosis of I. incarnata sp. n. with subterminal thorn at dorsum; one long, very slender Description: ♂ (Fig. 1). Wing span in holotype 18.5 mm, cornutus and two minute ones in vesica. in paratypes 18.0–18.5 mm. Head and thorax rusty ♀ not known. brown with brown marks; labial palpus 2.0, rusty brown. Forewing costa gently convex, termen weakly oblique, Inape epiphanes sp. n. but beneath middle. Ground colour pale ferruginous (Figs. 3, 49, 50) tinged with grey in apical area from apex to mid-ter Holotype ♂: “Ecuador: Zamora-Chinchipe-Prov. 22 km E men, strigulated with brown; creamy spot in middle of Loja, PN Podocarpus, San Francisco Ranger Stt, 2200 m, 3°59'15" S, 79°5'37" W, 9. x. 2002, sta 22, leg. Gielis & proximal edge of median fascia. Markings brown: Post Pelz”; GU-2180-V.P. (CVPR, eventually SMFL). basal fascia dark brown, tapering costally, ill-defined at Etymology: The name refers to the easily distinguishing base; median fascia broad in costal half, atrophying in colouration of the new species; Latin: epiphanes — easy to dorsal part, dark within median cell; submedian fascia distinguish. It is defined as a noun in apposition. from dorsum to this last; subterminal fascia oblique; Diagnosis: In I. epiphanes sp. n. the ground colour of the spots in posterior parts of costa much stronger than dor forewing is ochreous cream, markings dark brown, similar sal spots. Cilia concolorous with ground colour, mixed to I. uncina sp. n., but in ♂ genitalia I. epiphanes sp. n. is brown in costal half of wing. Hindwing creamy brownish; distinct by the broad phallus, the delicate, straight cornu strigulated brownish; cilia dirty cream. tus and a much shorter dorsal processes of the transtilla. Variation. In the paratypes the ground colour is darker From I. bicornis Razowski, 1999, also from Ecuador, the ferruginous except for the posterior area of the wing new species differs mainly in short processes of transtilla especially from middle of median fascia to apex where and a broad phallus. darker brown; markings darker; median fascia extending Description: ♂ (Fig. 3). Wing span 15.0 mm. Head och towards tornus. reous cream, labial palpus 2.8, ochreous cream, thorax ♂ genitalia (Figs. 39, 40). Uncus tapering terminally; darker with brownish bases of tegulae. Forewing more valva rather slender, somewhat up-curved; sacculus slender than in I. uncina sp. n., with more oblique ter short, rounded terminally; dorsal process of transtilla men. Ground colour ochreous cream; strigulation brow almost as long as sacculus; phallus broad with short ter nish; costal part of base of wing more brown. Markings minal portion; cornuti one short weakly bent and one dark brown: Postbasal fascia slender, atrophied dorsally; large strongly curved spines. median fascia reaching beyond middle of wing followed ♀ not known. by costal spots of which the first one is large; subterminal fascia reduced to a few spots. Cilia brownish cream, Inape luteina sp. n. cream ochreous in tornal half of wing. Hindwing brow (Figs. 2, 41, 42) nish cream; cilia creamer. Holotype ♂: “Ecuador: Loja-Prov., 10 km SE Loja, PN Podocarpus, Cajanuma Ranger Stt, 2850 m, 4°6'58" S, ♂ genitalia (Figs. 49, 50). Uncus slender; sacculus broad to 79°10'19" W, 7. x. 2002, sta 20, leg. Gielis & Pelz”; GU-1713- middle then slender; dorsal processes of transtilla broad, V.P. (CVPR, eventually SMFL). Etymology: The name refers to the colouration of the fore Figs. 37–54: Inape, ♂ genitalia. Figs. 37, 38: I. extraria sp. n., holotype, wing; Latin: lutea — yellow, -ina — a suffix defining dimini Fig. 37: genitalia (ventral view), Fig. 38: phallus. Figs. 39, 40: I. uncina tion. It is defined as a noun in apposition. sp. n., holotype, Fig. 39: genitalia (ventral view), Fig. 40: phallus. Figs. Diagnosis: Inape luteina sp. n. is similar to I. uncina sp. n. 41, 42: I. luteina sp. n., holotype, Fig. 41: genitalia (ventral view), Fig. 42: phallus. Figs. 43, 44: I. elegans sp. n., holotype, Fig. 43: genitalia (ventral as the shape of sacculus of valva shows, but distinct by its view), Fig. 44: phallus. Figs. 45, 46: I. incarnata sp. n., holotype, Fig. slender terminal part (reminiscent of that in I. epiphanes 45: genitalia (ventral view), Fig. 46: phallus. Figs. 47, 48: I. asymmetra sp. n.) and the presence of a small dorsal prominence. sp. n., holotype, Fig. 47: genitalia (ventral view), Fig. 48: phallus. Figs. 49, 50: I. epiphanes sp. n., holotype, Fig. 49: genitalia (ventral view), Fig. Description: ♂ (Fig. 2). Wing span 21.0 mm. Head pale 50: phallus. Figs. 51, 52: I. laterosclera sp. n., holotype, Fig. 51: genitalia (ventral view), Fig. 52: phallus. Figs. 53, 54: I. pompata sp. n., paratype ochreous cream, labial palpus 2.5, ochreous cream; tho (GU-2220-V.P.), Fig. 53: genitalia (ventral view), Fig. 54: phallus. — Scale rax ochreous. Forewing costa gently convex; apex sharp; bar for each figure 1.0 mm.

37 39 41

38 40 42

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49 51 53

50 52 54

© Entomologischer Verein Apollo e. V., Frankfurt am Main 122 short; phallus broad; single slender cornutus in vesica. Diagnosis: Inape pompata sp. n. is externally and in ♂ genitalia very similar to I. laterosclera sp. n.; ♂♂ of I. pom ♀ not known. pata sp. n. are larger, more robust, with wing span 17.0– Inape laterosclera sp. n. 18.0 mm (14.0–15.0 mm in I. laterosclera sp. n.), and the (Figs. 5, 6, 51, 52, 73) ground colour of the forewing is more rusty. ♂ genitalia Holotype ♂: “Ecuador: Napo-Prov., 12 km SSE Cosanga, as in I. laterosclera sp. n., but the valva is longer with a 2120 m, 0°37'26" S, 77°48'51" W, 24. x. 2002, sta 36, leg. lenght/width-relation of 3.5 (in I. laterosclera sp. n. 3.1). Gielis & Pelz”; GU-2186-V.P. (CVPR, eventually SMFL). Description: ♂ (Fig. 4). Wing span 17.0–18.0 mm. Head Paratypes (in total 1 ♂, 3 ♀♀): 1 ♂ (GU-2181-V.P.), 3 ♀♀ (GU- 2183-V.P., GU-2184-V.P.): same data as HT (CVPR). grey-brown, labial palpus 2.2, grey-brown, laterally dar Etymology: The specific epithet refers to the lateral position ker with ferruginous admixture; thorax brownish-grey, of the sclerites of the corpus bursae. It is defined as a noun base of tegula brown. Forewing as in I. laterosclera sp. n. in apposition. but ground colour grey-brown with strong rust admixture Diagnosis: Ground colour of forewing brownish; in ♂ especially in outer third where ground colour completely genitalia uncus and phallus of I. laterosclera sp. n. similar rust. Markings as in I. laterosclera sp. n.; cilia brownish to those in I. epiphanes sp. n.; sacculus reminding that grey more cream towards tornus. Hindwing brownish in I. clarkeana Brown & Razowski, 2002 and I. papal cream with weak brownish strigulation; cilia cream. lactana Razowski, 1999 but without terminal process. ♀: unknown. Inape laterosclera sp. n. is very closely related to I. pom ♂ genitalia (Figs. 53, 54). As in I. laterosclera sp. n., but pata sp. n. The differentiating characters are given in the valva longer and only very slightly broadening beyond diagnosis of the latter species. sacculus; phallus longer than in I. laterosclera sp. n. with Description: ♂ (Fig. 5). Wing span in holotype 14.0 mm, one cornutus. In one paratype (GU-1724-V.P.) there is an in ♂ paratye 15.0 mm. Head light grey-brown, labial additional smaller cornutus. palpus 2.2, light grey-brown, laterally with sparse ferrugi Remark: In ♂ genitalia this species is very close to I. nous scales; thorax brownish-grey, base of tegula brown. laterosclera sp. n. The small but constant differences in Forewing as in I. epiphanes sp. n. but with slightly less ♂ genitalia and in habitus as well as their occurrence in oblique termen at least in ♂. Ground colour light grey- different regions (northeastern respectivly southeastern brown with slight yellowish admixture and brown stri part of Ecuador) and at different elevations seem to gulation. Markings brown: Postbasal fascia incomplete support their status as different species. In Euliini also or consisting of two spots; median fascia rather slen other pairs of sibling species in these regions are known der, more or less long, usually not extending beneath Punctapinella cerithiphora median cell; subtornal marking not connecting the for (e.g. Razowski & Pelz, 2004 P. lojana mer; other spots weak, slightly variable; subterminal mar and Razowski & Pelz, 2004). king absent. Cilia brownish-grey to brown, more cream Inape homora sp. n. towards tornus. Hindwing brownish cream with weak (Figs. 31–33, 75) brownish strigulation; cilia cream. Holotype ♀: “Ecuador: Loja-Prov, 10 km SE Loja, PN Podocar ♀ (Fig. 6). Larger than ♂ (wing span 19.0–19.5 mm), with pus, Cajanuma Ranger Stt, 2850 m, 4°6'58" S, 79°10'19" W, forewing ground colour brownish-grey strigulated brown; 7. x. 2002, sta 20, leg. Gielis & Pelz”; GU-2257-V.P. (CVPR, eventually SMFL). subterminal marking ill-defined or absent. Paratypes (in total 3 ♀♀): 1 ♀ (GU-1653-V.P.), same data as ♂ genitalia (Figs. 51, 52). Uncus and processes of trans HT; 2 ♀♀ (GU-2266-V.P, GU-2267-V.P.), same locality, but tilla slender; valva slightly broadening beyond sacculus; 8. x. 2002, sta 21 (CVPR). this last armed with postbasal triangular transverse pro Etymology: The name refers to the similarity and a posible cess; phallus short; cornutus ca 3 times shorter than phal close relation to I. laterosclera sp. n. and its allies; Greek: lus. homora — neighbouring. It is defined as a noun in appo sition. ♀ genitalia (Fig. 73). Proximal part of sterigma well scle Diagnosis: Externally I. homora sp. n. is similar to I. late rotized, rather short; sclerites of colliculum asymmetrical, rosclera sp. n. but in the ♀ genitalia sclerite of the collicu one of them with numerous chiefly marginal spines.

Inape pompata sp. n. Figs. 55–72: Inape, ♂ genitalia. Figs. 55, 56: I. cateres sp. n., holotype, Fig. (Figs. 4, 53, 54) 55: genitalia (ventral view), Fig. 56: phallus. Figs. 57, 58: I. eparmuncus sp. n., holotype, Fig. 57: genitalia (ventral view), Fig. 58: phallus. Figs. Holotype ♂: “Ecuador: Loja-Prov., 10 km SE Loja, PN 59, 60: I. homeotypa sp. n., holotype, Fig. 59: genitalia (ventral view), Podocarpus, Cajanuma Ranger Stt, 2850 m, 4°6'58" S, Fig. 60: phallus. Figs. 61, 62: I. homologa sp. n., holotype, Fig. 61: 79°10'19" W, 8. x. 2002, sta 21, leg. Gielis & Pelz”; GS 2220- genitalia (ventral view), Fig. 62: phallus. Figs. 63, 64: I. commoda sp. n., V.P. (CVPR, eventually SMFL). holotype, Fig. 63: genitalia (ventral view), Fig. 64: phallus. Figs. 65, 66: I. Paratypes (in total 2 ♂♂): same locality as HT, but 7. x. 2002, sororia sp. n., holotype, Fig. 65: genitalia (ventral view), Fig. 66: phallus. sta 20 (GU-2182-V.P., GU-1724-V.P.) (CVPR). Figs. 67, 68: I. sororia sp. n., paratype (GU-2410-V.P.), Fig. 67: genitalia (ventral view), Fig. 68: phallus. Figs. 69, 70: I. sororia corryssa ssp. n., Etymology: The specific epithet refers to the nice appear holotype, Fig. 69: genitalia (ventral view), Fig. 70: phallus. Figs. 71, 72: I. ance of this species. Latin: pompatus — splendid. It is here sororia lojana ssp. n., paratype (GU-2197-V.P.), Fig. 71: genitalia (ventral with defined as a noun in apposition. view), Fig. 72: phallus. — Scale bar for each figure 1.0 mm.

55 57 59

56 58 60

61 63 65

62 64 66

67 69 71

68 70 72

73 74 75

76 77 78

79 80 81

Figs. 73–81: Inape, ♀ genitalia (ventral view). Fig. 73: I. laterosclera sp. n., paratype (GU-2184-V.P.). Fig. 74: I. porpax sp. n., holotype. Fig. 75: I. homora sp. n., holotype. Fig. 76: I. sinuata Brown & Razowski, 2003 (GU-2191-V.P.). Fig. 77: I. polysparta sp. n., holotype. Fig. 78: I. pseudocelypha sp. n., holotype. Fig. 79: I. celypha sp. n., holotype. Figs. 80, 81: I. geoda sp. n., Fig. 80: paratype (GU-1536-V.P.), Fig. 81: holotype. — Scale bar for each figure 1.0 mm.

82 83 84

85 86 87

Figs. 82–87: Inape sororia, ♀ genitalia (ventral view). Figs. 82–84: I. sororia sororia ssp. n., Fig. 82: paratype (GU-1634-V.P.), Fig. 83: paratype (GU- 1553-V.P.), Fig. 84: paratype (GU-1554-V.P.). Fig. 85: I. sororia lojana ssp. n., paratype (GU-1709-V.P.). Figs. 86, 87: I. sororia corryssa ssp. n., Fig. 86: paratype (GU-1703-V.P.), Fig. 87: paratype (GU-2209-V.P.). — Scale bar for each figure 1.0 mm. lum weak; spiny sclerite short; lateral sclerite very large, interfascia somewhat suffused); base of wing ochreous minutely spined along one margin. creamy. Hindwing distinctly strigulated brownish grey. Description: ♂ unknown. ♀ genitalia (Fig. 75). Sterigma rather short, moderately ♀ (Figs. 31–33). Wing span 25.0 mm in HT, 25.0–25.5 mm sclerotized; colliculum broad with weak sclerites; ductus in PT. Head yellow-brown, labial palpus 2.5 yellow- bursae very short; corpus bursae with short spiny sclerite brown; thorax similarly coloured, base of tegula brown. and large lateral sclerite. Forewing costa uniformly convex, termen hardly con cave beneath apex. Ground colour yellowish brown suf Inape porpax sp. n. (Figs. 34, 35, 74) fused and dotted with brown, paler in distal third than anteriorly to median fascia. Markings brown with paler Holotype ♀: “Ecuador: Napo-Prov., 15 km SE Cosanga, Coco drilo, 1850 m, 0°38'56" S, 77°47'34" W, 30. ix. 2002, sta 12, ranges and darker dots: Postbasal fascia ill-defined, base leg. Gielis & Pelz”; GU-2251-V.P. (CVPR, eventually SMFL). ochreous; median fascia diffuse in dorsal area of wing; Paratypes (2 ♀♀): Ecuador: Napo-Prov., 12 km SSE Cosanga, postmedian fascia indistinct, darker at costa, dotted with 2120 m, 0°37'26" S, 77°48'51" W, 24. x. 2002, sta 36, leg. brown. Cilia concolorous with ground colour. Hindwing Gielis & Pelz (GU-1596-V.P., GU- 1597-V.P.) (CVPR). whitish cream, darker on periphery, with indistinct Etymology: The name refers to the shape of the sclerite in strigulation. In paratypes ground colour of forewing the corpus bursae; Latin: porpax — loop. It is a noun in appo darker, more brown (in one paratype even postmedian sition.

Diagnosis: Externally I. porpax sp. n. is similar to I. cely distinct by small spiny sclerite of corpus bursae. pha sp. n., whilst the ♀ genitalia are similar to those of I. homora sp. n. However, I. porpax sp. n. is easily distingui Inape pseudocelypha sp. n. shed by the very long lateral sclerite and spiny sclerite of (Figs. 29, 78) ductus bursae. Holotype ♀: “Ecuador: Pichincha-Prov., 7 km NW Mindo, Sachatamia, 1700 m, 0°1'35" S, 78°45'34" W, 8.–11. xii. Description: ♂ not known. 2004, leg. Volker Pelz”; GU-2547-V.P. (CVPR, eventually ♀ (Figs. 34, 35). Wing span in holotype 26.0 mm, in SMFL). paratypes 24.0 mm. Head pale ferruginous; labial palpus Etymology: The name refers to the similarity of this species to I. celypha sp. n., Greek: pseudos — false. It is a noun in 2.5, pale ferruginous; thorax browner, with darker base apposition. of tegula. Costa of forewing distinctly convex, termen Diagnosis: Inape pseudocelypha sp. n. is externally simi to middle hardly oblique. Ground colour ochreous-fer lar to I. celypha sp. n. and I. porpax sp. n., but with less ruginous with brownish suffusions; brown dots present; markings ill-defined in form of slender median fascia convex forewing costa and in ♀ genitalia distinct by smal marked dark brown at tornus and traces of fasciae in sub ler sclerites of the bursa copulatrix; the ♀ genitalia of terminal part of wing; base of wing creamy suffused grey I. pseudocelypha sp. n. are close to I. homora sp. n. but proximally, edged dark brown distally. Cilia concolorous with weaker sclerites of corpus bursae and broad, well with ground colour, browner from apex to mid-termen. sclerotized sterigma. Hindwing brownish grey; strigulation much darker. Cilia Description: ♂ unknown. ochreous creamy. Variation: Basal part of wing whitish ♀ (Fig. 29). Wing span 20.0 mm; Head and thorax pale tinged grey, indistinctly edged brownish or without distal cinnamon brownish; labial palpus 2.5, pale cinnamon edging; markings and maculation more or less weak. brownish. Costa gently curved outwards, termen fairly ♀ genitalia (Fig. 74) as in I. homora sp. n. but with dis long. Ground colour pale cinnamon dotted and partially tinctly sclerotized proximal part of sterigma and elongate suffused with ferruginous; some other dots and strigulae sclerite of ductus bursae extending in a curved fascia to brown. Basal area occupied by a creamy blotch edged proximal part of corpus; spiny sclerite longer than in the with brown distally; other markings ferruginous brow mentioned species, with very strong spinulation. nish, rather indistinctly edged, sparsely dotted with brown, tinged with blackish brown at tornus. Cilia con Inape celypha sp. n. colorous with ground colour with brownish median line. (Figs. 30, 79) Hindwing brownish cream with diffuse, more grey strigu Holotype ♀: “Ecuador: Napo-Prov., 10 km SSE Cosanga, lation; cilia paler than wing. 2180 m, 0°37'13" S, 77°49'29" W, 23. x. 2002, sta 35, leg. Gielis & Pelz”; GU-1535-V.P. (CVPR, eventually SMFL). ♀ genitalia (Fig. 78). Sterigma rather short, broad with Etymology: The name refers to the colouration of the fore distinct arc-shaped sclerite ventrally; sclerites of ductus wings which when in rest remind a hask, Latin: celypha bursae small; spiny sclerite of bursa copulatrix weak. – hask. It is a noun in apposition. Diagnosis: Inape celypha sp. n. is externally similar to Inape cateres sp. n. I. homora sp. n. but distinguished by the darker greyish (Figs. 10, 55, 56) hindwings; in ♀ genitalia the sclerite of the ductus bursae Holotype ♂: “Ecuador: Azuay-Prov., PN Cajas, Laguna Lla of I. celypha sp. n. is stronger than in I. xerophanes from viuco, 3225 m 2°50'38" S, 79°8'35" W, 5. x. 2002, sta 17, leg. Gielis & Pelz”; GU-1679-V.P. (CVPR, eventually SMFL). which it also differs in having a very small spiny sclerite Paratype (1 ♂): same data as HT (CVPR). of corpus bursae. Etymology: The species name refers to the structure of the Description: ♂ unknown. sacculus; Latin: cateres — armed. It is defined as a noun in apposition. ♀ (Fig. 30). Wing span 25.0 mm; head and thorax pale ferruginous; labial palpus 2.5, pale ferruginous. Apex of Diagnosis: This species externally is similar to I. geoda forewing short, rather rounded, termen slightly oblique, sp. n., I. homologa sp. n. and I. incarnata sp. n. From the straight to middle. Ground colour cinnamon suffused two former species it differs by a narrower forewing and brownish to before middle, hardly mixed ochre in distal the lighter greyish cream hindwings. I. incarnata sp. n. third dotted dark brown. Markings brownish: Postbasal is easily distuinguished from I. cateres sp. n. by the dark fascia brown, base of wing brownish creamy; median fas basal blotch of the forewing; the ♂ genitalia are similar cia slender, connected with subtornal marking; subapical to I. eparmuncus sp. n., but differing in the slender uncus blotch weak, diffuse. Cilia concolorous with posterior not broadening postbasally. part of wing suffused brown in costal half of termen. Description: ♂ (Fig. 10). Wing span 21.5 mm; head and Hindwing greyish, more creamy on periphery; strigu thorax ochreous cream; labial palpus 2.0, grey. Forewing lation dark grey; cilia greyish. somewhat narrower than in I. homologa sp. n. Ground ♀ genitalia (Fig. 79). Sterigma rather short; sclerite of colour light grey, mixed with darker grey scales, light ductus bursae large, similar to that in I. xerophanes but cream along posterior half of costal part of median fascia situated more anteriorly; Inape celypha sp. n. is, however, and last radial veins. Markings dark grey brown with dark

© Entomologischer Verein Apollo e. V., Frankfurt am Main 127 brown spots consisting of costal half of median fascia termen not oblique beneath apex. Ground colour whitish and subterminal triangle. Cilia concolorous with ground grey with white diffuse spots especially in distal half of colour, darker beneath apex. Hindwing light cream; wing and pale ochreous brownish suffusions chiefly in weakly strigulated brown-grey; cilia light cream. basal half; strigulation brownish grey and grey; base of ♂ genitalia (Figs. 55, 56). Uncus slender, tapering ter wing and partially termen suffused brownish ochre or minally; valva beyond sacculus rather slender; sacculus grey, respectively. Markings: Costal spots and costal half convex postbasally armed with distinct dorsopostbasal of median fascia dark brown, similar spot near middle of thorn, with several spines in distal part; dorsal processes wing, all on pale rust suffusions; dorsal spots brownish, of transtilla slender, curved; phallus as long as costa of smaller than costal spots. Cilia brownish, brown between valva; single slender cornutus in vesica. apex and mid-termen, greyer in tornal area. Hindwing greyish tinged brownish in apex area; strigulation weak. ♀ unknown. Cilia grey-white. Inape commoda sp. n. ♂ genitalia (Figs. 57, 58). Uncus strong, broadest postba (Figs. 15, 63, 64) sally, tapering terminally; valva as in I. cateres sp. n.; sac Holotype ♂: “Ecuador: Napo-Prov., 10 km SSE Cosanga, culus less convex, dorsobasal thorn more slender, more 2180 m, 0°37'13" S, 77°49'29" W, 23. x. 2002, sta 35, leg. proximal, spines of posterior part weaker; dorsal pro Gielis & Pelz”; GU-2389-V.P. (CVPR, eventually SMFL). cesses of transtilla slender, strongly curved; phallus and Etymology: The name refers to the similarity of this species cornutus slightly shorter than in I. cateres sp. n. to I. epiphanes sp. n.; Latin: commodus — the same. It is defined here as a noun in apposition. ♀ unknown. Diagnosis: Externally this species resembles I. epiphanes Inape homeotypa sp. n. sp. n. from which it differs in the strongly reduced fore (Figs. 12, 59, 60) wing markings; in ♂ genitalia the sacculus of I. commoda Holotype ♂: “Ecuador: Loja-Prov, 10 km SE Loja, PN Podocar sp. n. is much shorter and broader than in I. epiphanes pus, Cajanuma Ranger Stt, 2850 m, 4°6'58" S, 79°10'19" W, sp. n. and the phallus is somewhat longer. 7. x. 2002, sta 20, leg. Gielis & Pelz”; GU-1715-V.P. (CVPR, eventually SMFL). Description: ♂ (Fig. 15). Wing span 15.0 mm. Head and Paratype (1 ♂): same data as HT (CVPR). thorax ochreous, the latter brown proximally; labial pal Etymology: The specific name refers to a similarity to other pus 2.0. Forewing costa convex, termen fairly oblique. species of Inape in the spinulation of the end of the saccu Ground colour pale ochreous cream with orangeous suf lus; Greek: homos — similar, typus — contents. It is a noun fusions and black-brown dots; markings concolorous, ill- in apposition. defined, consisting of traces of median fascia best visible Diagnosis: The ♂ genitalia of Inape homeotypa sp. n. are in middle of wing. Cilia worn. Hindwing cream; strigula comparable to those of I. eparmuncus sp. n., but easily dis tion indistinct; cilia cream. tinguished by much broader uncus and distinctly angu ♂ genitalia (Figs. 63, 64). Uncus rather short, fairly late sacculus without dorsobasal thorn. broad; valva slender; sacculus simple, short, rather broad; Description: ♂ (Fig. 12). Wing span 22.0 mm. Forewing dorsal processes of transtilla long; phallus extending rather slender with termen slightly oblique. Head brow ventroterminally; cornuti eight unequally large spines. nish, thorax similar in colour, concolorous anteriorly. ♀ not known. Ground colour creamy brown, partially suffused and sprinkled with brown, strigulation sparse, concolorous; Inape eparmuncus sp. n. base of wing suffused brown; costal spots concolorous. (Figs. 11, 57, 58) Markings rudimentary, diffuse, brownish, represented Holotype ♂: “Ecuador: Zamora-Chinchipe-Prov. 22 km E mainly by costal portion of median fascia. Cilia worn, Loja, PN Podocarpus, San Francisco Ranger Stt, 2200 m, concolorous with ground colour. Hindwing whitish tin 3°59'15" S, 79°5'37" W, 9. x. 2002, sta 22, leg. Gielis & Pelz”; GU-1747-V.P. (CVPR, eventually SMFL). ged creamy in apex area; cilia white. Etymology: The specific epithet refers to the shape of the ♂ genitalia (Figs. 59, 60). Uncus very broad postbasally, uncus; Latin: eparma – swelling. It is a noun in apposition. then gradually tapering terminad; socius fairly broad; Diagnosis: Externally I. eparmuncus sp. n. is rather simi valva beyond sacculus slender; sacculus angulate with lar to I. cateres sp. n. and I. geoda sp. n., differing from strong spines in distal half; processes of transtilla slender; the former species by the much darker grey hindwings. phallus in dorsal aspect broad postmedially; one longer I. geoda sp. n. differs from I. eparmuncus sp. n. by the and four much shorter cornuti present. white fascia from beyond middle to apex of forewing. ♂ ♀ unknown. genitalia of I. eparmuncus sp. n. similar to I. cateres sp. n., but the uncus of I. eparmuncus sp. n. is distinctly broad Inape homologa sp. n. ening postbasally. (Figs. 13, 61, 62) Holotype ♂: “Ecuador: Loja-Prov, 10 km SE Loja, PN Podocar Description: ♂ (Fig. 11). Wing span 21.0 mm. Head brow pus, Cajanuma Ranger Stt, 2850 m, 4°6'58" S, 79°10'19" W, nish cream, thorax darker brown, labial palpus 2.0, dark 7. x. 2002, sta 20, leg. Gielis & Pelz”; GU-1708-V.P. (CVPR, brown with some rusty scales. Forewing fairly broad, eventually SMFL).

Etymology: The name concerns the similarity with I. homeo 22 km E Loja, PN Podocarpus, San Francisco Ranger Stt, typa sp. n.; Greek: homos – similar; logos — science. It is a 2200 m, 3°59'15" S, 79°5'37" W, 9. x. 2002, sta 22, leg. Gie noun in apposition. lis & Pelz; GU-2191-V.P. (CVPR). Diagnosis: This species is very similar to I. homeotypa This species was known to date only from the ♀ HT and sp. n., with similar ♂ genitalia, however, easily distin 2 ♂♂ PTs, all from Bolivia. As the abdomina are lost in guished by the shape of the uncus which in I. homologa both paratypes, the ♂ genitalia remain unknown. The sp. n. is shorter and broader postbasally, the more slender structure of the ♀ genitalia allows to suppose that I. phallus, and the lower number (two only) of cornuti. sinuata is closely related to I. laterosclera sp. n. Description: ♂ (Fig. 13). Wing span 22.0 mm. Head and Inape asymmetra sp. n. thorax ochreous rusty; labial palpus 2.0, base of tegula (Figs. 8, 37, 38) brown. Forewing somewhat broader than in I. homeo Holotype ♂: “Ecuador: Azuay-Prov. 22 km SO Gualaceo > typa sp. n. Ground colour brownish, brownish creamy Plan de Milagro, Cordillera Zapote Naida, 3300 m, 6. x. 2000, along posterior half of costal part of median fascia and leg. Volker Pelz”; GU-1033-V.P. (CVPR, eventually SMFL). last radial veins; base of wing brownish; spots along Paratypes: 2 ♂♂, same data as HT (GU-1027-V.P.); (CVPR). costa and termen similarly coloured. Markings brown- Etymology: The name refers to the asymmetry of the saccu grey with dark brown spots consisting of costal half of lus. Greek: a (prefix) — not; sym — common; metros — mea median fascia and subterminal triangle. Cilia concolo sure. It is defined as a noun in apposition. rous with ground colour, browner beneath apex. Hind Diagnosis: Inape asymmetra sp. n. is comparable to I. wing brownish creamy; strigulation more brown-grey; pompata sp. n. and its allies but quite different externally cilia dirty creamy. by a transverse strigulation of the forewing forming more or less complete lines; the ♂ genitalia of the new species ♂ genitalia (Figs. 61, 62). Uncus short, strongly expanding differ from I. pompata sp. n. in a larger uncus, the saccu beyond base, then subtriangular; socius rather slender; lus of the valva with two thorns, and in the asymmetry sacculus broad, with distinct caudal angle marked with of the sacculi. series of spines; phallus rather slender; two moderatly sized cornuti in vesica. Description: ♂ (Fig. 8). Wing span 25.5 mm. Head and thorax brown, labial palpus 2.5, tinged rust. Forewing ♀ unknown. broad, with costa distinctly convex. Ground colour creamy Inape polysparta sp. n. ferruginous, more brownish in distal half of wing, den (Figs. 36, 77) sely strigulated with brown; some strigulae forming more Holotype ♀: “Ecuador: Napo-Prov., 10 km SSE Cosanga, of less complete transverse lines. Cilia brownish with 2180 m, 0°37'13" S, 77°49'29" W, 23. x. 2002, sta 35, leg. slight ferruginous hue. Hindwing dirty cream with slight Gielis & Pelz”; GU-2256-V.P. (CVPR, eventually SMFL). ochreous brownish admixture in distal area; strigulation Etymology: The species name refers to the maculation of pale, greyish; cilia concolorous with ground colour. the forewing; Greek: poly — numerous, sparta – scattered. It is defined as a noun in apposition ♂ genitalia (Figs. 37, 38). Uncus long; valva fairly broad; Diagnosis: Inape polysparta sp. n. is very close to I. sacculus with a larger submedian and a smaller postme homora sp. n. but easily distinguished in ♀ genitalia by dian thorn, asymmetric, the posterior one much larger in broad sterigma and small, posterior sclerites of corpus the right sacculus, minute in the left one; a group of lon bursae. ger spines in the proximal part of the disc; phallus rather short; cornuti arranged in two groups: the anterior group Description: ♂ unknown. represented by a single strong spine, the posterior group ♀ (Fig. 36). Wing span 27.0 mm. Head pale yellowish consisting of 12 much shorter, capitate spines. brown; thorax more brown especially the tegula; labial ♀ not known. palpus 2.3. Forewing similar to that in I. homora sp. n.; ground colour dirty yellow with slight admixture of Inape geoda sp. n. brownish grey, densely dotted dark brown. Markings ill- (Figs. 16, 17, 80, 81) defined, dark brown, consisting of a trace of basal blotch, Holotype ♀: “Ecuador: Napo-Prov., 10 km SSE Cosanga, median fascia represented by costal and median blotches 2180 m, 0°37'13" S, 77°49'29" W, 23. x. 2002, sta 35, leg. Gielis & Pelz”; GU-2168-V.P. (CVPR, eventually SMFL). followed by subapical spot, a part of rudiments of the Further material examined (no PT): 1 ♀, Ecuador: Napo- postmedian fascia. Cilia (rubbed) probably concolorous Prov., 15 km SE Cosanga, Cocodrilo, 1850 m, 0°38'56" S, with ground colour. Hindwing creamy, tinged ochre at 77°47'34" W, 27. x. 2002, sta 39, leg. Gielis & Pelz, GU-1536- apex; strigulation brownish grey; cilia creamy. V.P. (CVPR). ♀ genitalia (Fig. 77). Proximal part of sterigma extending Etymology: The name refers to the general coloration of the forewing; Latin: geodes — sallow. It is defined as a noun in laterally beyond middle; sclerites of ductus bursae weak, apposition. extending towards corpus bursae. Diagnosis: Externally this new species is similar to I. Inape sinuata Brown & Razowski, 2003 homologa sp. n. and I. incarnata sp. n.; in ♀ genitalia I. (Figs. 9, 76) geoda sp. n. is near to I. laterosclera sp. n., but differs in Material examined: 1 ♀: Ecuador: Zamora-Chinchipe-Prov. much broader lateral sclerite of corpus bursae.

Description: ♂ unknown. Description: Dimorphic species. ♀ (Fig. 16). Wing span in HT 28.0 mm. Head pale rust, ♂ (Fig. 19, 22). Wing span 19.6 mm (18.5–20.5 mm; n = labial palpus 2.2, grey-brown; thorax ochreous tinged 6), in holotype 20.5 mm. Head and thorax brownish grey-brown proximally, base of tegula browner. Costa of grey, labial palpus 2.5 in ♂, brownish grey with some red forewing weakly convex; termen slightly oblique, bent. brown scales laterally, base of tegula browner. Costa of Ground color grey-brown suffused whitish in tornal forewing weakly convex, termen slightly oblique, rather fourth of wing and less so along dorsum basad and at straight. Ground colour pale brownish creamy with costa subapically; submedian interfascia tinged ochre; ochreous admixture; strigulation weak, brownish. Mar strigulation grey-brown, spots along wing edges darker; kings sepia-brown: Basal blotch atrophying proximally; a fascia from beyond middle of wing to apex whitish, median fascia distinct proximally, diffuse posteriorly, mixed creamy proximally. Markings dark brown consis not reaching dorsum; brownish lines or strigulae in cos ting of costal half of median fascia and subapical blotch; tal part of suffusion; subtornal marking ill-defined. Cilia subterminal marking paler, rather weak. Cilia brownish, pale brownish, more ochreous in tornal portion. Hind paler towards tornus. Hindwing creamy grey; strigulation wing whitish cream, mixed brownish at apex, with ill- weak. Cilia rather concolorous with wing. defined strigulation. ♀ genitalia (Figs. 80, 81). Sterigma small with rounded ♀ (Fig. 25). Wing span 23.6 mm (21.0–24.5 mm; n = 5). proximal corners; sclerite of ductus bursae extending Labial palpus longer than in ♂ 2.8. Termen of forewing towards proximal portion of corpus bursae; spined somewhat sinuate beneath apex; ground colour suffused sclerite large. brownish or tinged pale ferruginous; dorsal parts of mar Variation. The second ♀ at hand (Fig. 17) is smaller (wing kings present; subterminal fascia more or less distinct. span 25.0 mm) than HT with forewing ground colour Cilia brownish. Hindwing white creamy darker towards slightly mixed with ochreous. This specimen is therefore apex, strigulation rather grey. not included as paratype, but illustrated for reasons of ♂ genitalia (Figs. 65–68). Similar to I. clarkeana but sac comparison. culus with terminal process and a large postbasal lobe; The following specimens were collected at five localities phallus stout; cornuti 1 rather strong and 2–3 much smal in three different mountain chains of the Ecuadoran ler spines. Andes. There are slight but constant differences mainly ♀ genitalia (Figs. 82–84). Sterigma short with broad ante in genitalia between these three population-groups, ostial portion; sclerite of colliculum atrophying; median which may represent allopatric species; but at present sclerite of ductus bursae short, slender proximally; spiny it seems more appropriate to treat them as conspecific sclerite long, accompanied by rounded sclerite situated until more material also from further regions is at hand. in middle of corpus bursae crowned with long spines. Because of the mentioned constant differences we are Inape sororia is found to have three subspecies; besides treating them as subspecies. The sexes were associated the nominotypical one the following two: by their common occurence at all places and their over all similarity. Therefore their combination remains Inape sororia corryssa ssp. n. equivocally until more about the biology of the taxa is (Figs. 20, 23, 26, 28, 69, 70, 86, 87) known. A slight sexual dimorphism is present (to date a Holotype ♂: “Ecuador: Azuay-Prov., PN Cajas, Laguna Lla phenomenon only rarely encountered in Euliini). viuco, 3225 m, 2°50'38" S, 79°8'35" W, 5. x. 2002, sta 17, leg. Gielis & Pelz”; GU-1630-V.P. (CVPR, eventually SMFL). Inape sororia sp. n. Paratypes (in total 6 ♂♂, 4 ♀♀): 3 ♂♂ (GU-1631-V.P., GU- (19, 22, 25, 65–68, 82–84) 2195-V.P.), 3 ♀♀ (GU-1703-V.P., GU-2240-V.P.): same dates as Holotype ♂: “Ecuador: Napo-Prov., 5 km W Papallacta, HT; 3 ♂♂ (GU-2196-V.P.), 1 ♀ (GU-2209-V.P.): same locality Laguna Papallacta, 3430 m, 0°22'27" S, 78°9'50" W, 28. x. but 12. x. 2002, sta 26, leg. Gielis & Pelz; 2 ♂♂ (GU-2232- 2002, sta 40, leg. Gielis & Pelz”; GU-1633-V.P. (CVPR, V.P., GU-2194-V.P.): Ecuador: Azuay-Prov., 25 km S Cuenca, eventually SMFL). Pto. de Tinajilla, 3320 m, 3°9'46" S, 79°1'30" W, 6. x. 2002, sta 18, leg. Gielis & Pelz (CVPR). Paratypes (in total 5 ♂♂, 5 ♀♀): 3 ♀♀ (GU-1634-V.P., GU- 1553-V.P., GU-1554-V.P.), same data as HT (CVPR); 1 ♀ same Etymology: The epithet refers to the heavily spined sclerites data as HT (ISEZ); 1 ♀ same data as HT (PUCE); 3 ♂♂ (GU- of bursa copulatrix; Latin: corryssa — armed. It is defined as 2409-V.P., GU-2410-V.P., GU-2411-V.P.): Ecuador: Napo-Prov., a noun in apposition. 1 km W Papallacta, 3350 m 0°22'25" S, 78°2'10" W, 21.– Diagnosis: Inape sororia corryssa ssp. n. is very close to the 23. xii. 2004, leg. Volker Pelz (CVPR); 1 ♂ same data (ISEZ); nominotypical subspecies but with creamy submedian 1 ♂ same data (PUCE). interfascia at least in costal area of wing and very distinct Etymology: The specific name refers to the similarity to basal blotch in ♂. In ♂ genitalia I. s. corryssa ssp. n. differs some of its congeners both in facies and genitalia., e.g. to I. clarkeana; Latin: sororia — sisterly. It is defined as a noun mainly in the higher number (3 strong, rather large in apposition. and 2-4 small ones) of cornuti, whose positions in the Diagnosis: ♂ genitalia of I. sororia sp. n. resemble those everted vesica are also different; ♀♀ of this subspecies in I. clarkeana but differ in anterior position of the lobe of characterize with large sclerite of ductus bursae. sacculus and the absence of a very long, curved cornutus. Description: ♂ (Figs. 20, 26, 28). Wing span 19.4 mm

(18.0–21.5 mm; n = 7). Head, labial palpus, thorax and ♂ genitalia (Figs. 71, 72). As in the preceding subspecies shape of wings as in the nominotypical subspecies. but sacculus with stronger, downcurved terminal pro Ground colour of forewing pale brownish creamy, cess and larger basal process. Phallus with 3–4 cornuti in slightly suffused with brownish dorsally, with rust pos vesica (2 longer inequally sized and 1–2 small ones). teriorly; strigulation fine, brown; brown suffusion from ♀ genitalia (Fig. 85). Similar to I. s. corryssa ssp. n. but median fascia to terminal area of wing. Markings dark sclerite of ductus bursae smaller and lateral sclerite some brown consisting of postbasal fascia or basal blotch, cos what broader. tal half of median fascia and postmedian marks at costa. Cilia brownish, more creamy at apex, occasionally tinged References with rusty colour. Hindwing white cream, darker apic ally, with weak grey strigulation; cilia whitish. Brown, J. W. (1989): Generic reassignments for Neotropical tor tricid moths (Tortricidae). — Journal of the Lepidopterists’ ♀ (Fig. 23). Wing span 21.0 mm (20.0–22.0 mm; n = 4). Society 43: 313–322. Ground colour of ♀ forewing pale brownish; markings ———, & Razowski, J. (2003): Review of Inape Razowski (Lepido brown, rather weak; basal blotch ill-defined, median fas ptera: Tortricidae: Euliini), with descriptions of five new spe cia reaching tornus, postmedian fascia weak or in form cies. — Acta zoologica cracoviensia 46: 197–208. of a line (anterior edge) followed by a weak suffusion; Kristensen, N. P. (2003): 4. Skeleton and muscles: adults. — Pp. hindwing darker than in ♂. 39–131 in: Kristensen, N. P. (ed.), Lepidoptera, moths and butterflies. Vol. 2: morphology, physiology, and deveop ♂ genitalia (Figs. 69, 70) as in I. s. sororia ssp. n., but ment. — Part 36 in: Fischer, M. (serial ed.), Handbook of ventral edge of uncus distinctly convex and 3 longer, Zoology, vol. IV, Arthropoda: Insecta. — xii + 564 pp.; Berlin, inequally sized, and 2–4 small cornuti present. Sacculus New York (W. de Gruyter). with stronger, more bent terminal process and a larger Powell, J. A., Razowski, J., & Brown, J. W. (1995): Tortricinae. postbasal lobe. — Pp. 138–151 in: Heppner, J. B. (ed.), Atlas of Neotropical Lepidoptera 3. Checklist: Part 2, Hyblaeoidea — Pyraloidea ♀ genitalia (Figs. 86, 87). Sclerite of ductus bursae broadly — Tortricoidea. — Gainesville (Association for Tropical Lepi extending into a slender lateral sclerite; crowned sclerite doptera). distinct; spiny sclerite somewhat variable. Razowski, J. (1997): Euliini (Lepidoptera: Tortricidae) of Peru with description of new taxa and list of the New World genera. Inape sororia lojana ssp. n. — Acta zoologica cracoviensia 40 (1): 79–105. (Figs. 21, 24, 27, 71, 72, 85) ——— (1999): Tortricidae (Lepidoptera) from Ecuador. — Acta zoo Holotype ♂: “Ecuador: Loja-Prov., 10 km SE Loja, PN logica cracoviensia 42: 321–342. Podocarpus, Cajanuma Ranger Stt, 2850 m, 4°6'58" S, 79°10'19" W, 8. x. 2002, sta 21, leg. Gielis & Pelz”; GU-1632- ———, & Pelz, V. (2003a): Tortricidae collected in Ecuador in the V.P. (CVPR, eventually SMFL). years 1996–1999: Euliini (Lepidoptera). — Nachrichten des Entomologischen Vereins Apollo, Frankfurt am Main, N.F. Paratypes (in total 3 ♂♂, 1 ♀): same data as HT, but with 24 (4): 189–207. GU-2198-V.P., GU-2197-V.P., GU-2210-V.P (♂♂) and GU- 1709-V.P. (♀) (CVPR). ———, & ——— (2003b): Descriptions of Netechmodes gen. n. and two Etymology: The subspecific epithet refers to the occurence species of Netechma Razowski, 1991 from Ecuador (Lepi of this taxon in the Loja Province. It is herewith defined as a doptera Tortricidae). — Bollettino di Zoologia agraria e di noun in apposition. Bachicoltura, Ser. II, 35 (1): 17–21. Diagnosis: Inape sororia lojana ssp. n. is very close to the ———, & ——— (2004a): Remarks on Punctapinella Brown, 1991 (Lepidoptera: Tortricidae: Euliini), with descriptions of five two preceding taxa but in the ♂ hindwing light cream new species from Ecuador. — Entomologische Zeitschrift, with only very weak strigulation and not darker towards Stuttgart, 114 (5): 233–236. apex. ♂ genitalia differ mainly in the number and size of ———, & ——— (2004b): Remarks on Seticosta Razowski, with descrip cornuti which is lower with 2 longer, rather large and 1–2 tion of seven new species from Ecuador (Lepidoptera: Tor small ones than in I. s. corryssa ssp. n., but higher than tricidae). — SHILAP Revista de Lepidopterologia, Madrid, in the nominotypical subspecies; in I. s. lojana ssp. n. the 32 (127): 183–194. sacculus is characterized by a larger, more bent terminal ———, & ——— (2005a): New species of Gorytvesica Razowski, 1997 process and a stronger basal process. and Transtillaspis Razowski, 1987 (Lepidoptera: Tortrici dae: Euliini) from Ecuador. — Acta zoologica cracoviensia, Description: ♂ (Figs. 21, 24). Wing span 20.4 mm (19.5– Krakow, 48B (1/2): 57–94. 21.5 mm; n = 4). Head, labial palpus, thorax and shape of ———, & ——— (2005b): Exoletuncus Razowski, 1988 (Lepidoptera: wings as in the nominotypical subspecies. Ground colour Tortricidae: Euliini) from Ecuador, with description of seven of forewing pale brownish cream. Markings as in the pre new species. — SHILAP Revista de Lepidopterologia, Madrid, ceeding subspecies. Hindwing light cream with only very 33: 327–339. weak darker strigulation which is typical for the other ———, & ——— (2005c): Lobogenesis Razowski, 1990 (Lepidoptera: subspecies especially towards apex. Tortricidae: Euliini) from Ecuador, with description of ten new species. — Polski Pismo Entomologiczne, Bydgoszcz, 74: ♀ (Fig. 27). Wing span 25.5 mm (n = 1). Similar to Inape 439–453. sororia corryssa ssp. n. but hindwing with weaker strigu lation. Received: 12. vii. 2005