____17. plana -- ____62. (Gomphus) graslinellus -- Pronghorn CHECKLIST OF ____18. -- Blue-ringed Dancer Clubtail ( AND ) ____19. -- Blue-tipped Dancer ____63. Gomphus (Gomphus) militaris -- Sulpher-tipped ____20. Argia translata -- Dusky Dancer Clubtail (125 ) ____21. antennatum -- Rainbow Bluet ____64. Hagenius brevistylus -- Dragonhunter ____22. -- Azure Bluet ____65. Ophiogomphus rupinsulensis -- Rusty Snaketail With Data Sources, Notes, and Historical Records. ____23. Enallagma basidens -- Double-striped Bluet [Probably O. westfalli] Last updated 25 January 2002 ____24. Enallagma carunculatum -- ____66. Ophiogomphus severus -- Pale Snaketail ____25. Enallagma civile -- ____67. Progomphus obscurus -- Common Sanddragon ____26. -- Turquoise Bluet ____68. Stylogomphus albistylus -- Least Clubtail ____27. Enallagma exsulans -- ____69. Stylurus amnicola -- Riverine Clubtail ____28. -- Skimming Bluet ____70. Stylurus intricatus -- Brimstone Clubtail ____29. Enallagma praevarum -- ____71. Stylurus plagiatus -- Russett-tipped Clubtail ____30. Enallagma signatum -- - - SPIKETAILS ____31. Enallagma traviatum westfalli -- - 1 SPECIES ____32. -- Vesper Bluet ____33. barberi -- Desert Forktail ____72. obliqua -- Arrowhead Spiketail ____34. -- Plains Forktail (MACROMIINAE) - - CRUISERS ____35. -- Mexican Forktail - 4 SPECIES ____36. -- Black-fronted Forktail ____37. Ischnura hastata -- ____73. transversa -- Stream Cruiser ____38. -- Western Forktail ____74. Macromia illinoiensis -- River Cruiser ____39. -- Fragile Forktail ____75. Macromia pacifica -- Gilded River Cruiser ____40. Ischnura verticalis -- ____76. Macromia taeniolata -- Royal River Cruiser Above: Photograph of Argia moesta male taken in ____41. Nehalennia gracilis -- Sphagnum Sprite CORDULIIDAE (CORDULIINAE) - - EMERALDS August 1994 near the Big Caney River, Chautauqua ____42. Telebasis salva -- Desert Firetail County, Kansas By Roy J. Beckemeyer. - 9 SPECIES ODONATA (ANISOPTERA) – DRAGONFLIES Prepared by Roy J. Beckemeyer - 83 SPECIES ____77. (Tetragoneuria) costalis -- Stripe-winged Formatted by Jim Mason (30 August 2002) Baskettail PETALURIDAE - - PETALTAILS ____78. Epitheca (Tetragoneuria) cynosura -- Common ODONATA (ZYGOPTERA) – DAMSELFLIES - 1 SPECIES Baskettail ____79. Epitheca (Tetragoneuria) petechialis -- Dot-winged - 42 SPECIES ____43. Tachopteryx thoreyi -- Gray Petaltail Baskettail ____80. Epitheca (Epicordulia) princeps -- Prince Baskettail - - DARNERS -- BROAD-WINGED DAMSELFLIES - ____81. Neurocordulia molesta -- Smoky Shadowdragon - 10 SPECIES 3 SPECIES ____82. Neurocordulia xanthosoma -- Orange ____44. constricta -- Lance-tipped Darner Shadowdragon ____1. Calopteryx maculata -- Ebony Jewelwing ____45. Aeshna interrupta lineata -- Variable Darner ____83. Somatochlora linearis -- Mocha Emerald ____2. americana -- American Rubyspot ____46. Aeshna multicolor -- Blue-eyed Darner ____84. Somatochlora ozarkensis -- Ozark Emerald ____3. Hetaerina titia -- Smoky Rubyspot ____47. Aeshna umbrosa -- ____85. Somatochlora tenebrosa -- Clamp-tipped Emerald -- SPREADWING DAMSELFLIES ____48. junius -- Common Green Darner - - SKIMMMERS - 5 SPECIES ____49. Anax longipes -- Comet Darner - 40 SPECIES ____50. Basiaeshchna janata -- Springtime Darner ____4. grandis -- Great Spreadwing ____51. vinosa -- Fawn Darner ____86. Brechmorhoga mendax -- Pale-faced Clubskimmer ____5. Lestes disjunctus australis -- Common Spreadwing ____52. heros -- Swamp Darner ____87. elisa -- Calico Pennant ____6. Lestes eurinus -- Amber-winged Spreadwing ____53. Nasiaeschna pentacantha -- Cyrano Darner ____88. Celithemis eponina -- Halloween Pennant ____7. Lestes rectangularis -- Slender Spreadwing ____89. Celithemis fasciata -- Banded Pennant ____8. Lestes unguiculatus -- Lyre-tipped Spreadwing - - CLUBTAILS ____90. Celithemis verna -- Double-ringed Pennant - 18 SPECIES -- POND DAMSELS ____91. fugax -- Checkered Setwing - 34 SPECIES ____54. lentulus -- Stillwater Clubtail ____92. Dythemis velox -- Swift Setwing ____55. Arigomphus submedianus -- ____93. simplicicollis -- Eastern Pondhawk ____9. sp. -- Red Damsel ____56. spinosus -- Black-shouldered ____94. -- Great Pondhawk ____10. -- Paiute Dancer Spinyleg ____95. umbrata -- Band-winged Dragonlet ____11. Argia apicalis -- Blue-fronted Dancer ____57. -- Flag-tailed Spinyleg ____96. Ladona deplanata -- ____12. -- Seepage Dancer ____58. designatus -- Eastern Ringtail ____97. Leucorrhinia intacta -- Dot-tailed Whiteface ____13. Argia fumipennis violacea -- ____59. Gomphus () externus -- Plains Clubtail ____98. comanche -- Comanche Skimmer ____14. -- Kiowa Dancer ____60. Gomphus (Gomphurus) ozarkensis -- Ozark ____99. Libellula composita -- Bleached Skimmer ____15. Argia moesta -- Clubtail ____100. Libellula cyanea -- Spangled Skimmer ____16. Argia nahuana -- ____61. Gomphus (Gomphurus) vastus -- Cobra Clubtail ____101. Libellula flavida -- Yellow-sided Skimmer ____102. Libellula incesta -- Slaty Skimmer and Fauna of Kansas for 1977", Tech. Publ. of the 17. Beckemeyer, R.J., 1998, "Some Kansas state and ____103. Libellula luctuosa -- Widow Skimmer State Biol. Surv. of Kansas, 6:1-35. county Odonata records for 1998", Argia: The ____104. Libellula pulchella -- Twelve-spotted Skimmer news journal of the Dragonfly Society of the ____105. Libellula saturata -- Flame Skimmer 7. Cringan, M.S., 1978, "Dragonflies and damselflies Americas, 10(4):26. ____106. Libellula semifasciata -- Painted Skimmer of McKinney Marsh", The Emporia State ____107. Libellula vibrans -- Great Blue Skimmer University Research Studies, XXVII(3):1-28. [One 18. Charlton, R., April, 1998, Personal ____108. Orthemis ferruginea -- Roseate Skimmer of the few studies of the seasonal occurrence of communication. Prof. Charlton has noted the ____109. Pachydiplax longipennis -- Blue Dasher Odonata in Kansas.] existence of three male speciemens of Aeshna ____110. Pantala flavescens -- Wandering Glider interrupta lineata in the Kansas State University ____111. Pantala hymenaea -- Spot-winged Glider 8. Huggins, D.G., 1983, "New Records of Odonata", collection. There are 2 from Riley County and one ____112. Perithemis tenera -- Eastern Amberwing in "New Records of the Fauna and Flora of from Marion County. ____113. Plathemis lydia -- Kansas for 1982", Tech. Publ. of the State Biol. ____114. Plathemis subornata -- Surv. of Kansas, 13:24-25. 19. Beckemeyer, R.J., 2000, "Some county Odonata ____115. Sympetrum ambiguum -- Blue-faced records for Kansas and for 1999 and Meadowhawk 9. Huggins, D. G., and G.L. Harp, 1985, "The Nymph 2000", Argia: The news journal of the Dragonfly ____116. Sympetrum corruptum -- Variegated of Gomphus (Gomphurus) ozarkensis Westfall Society of the Americas, 12(3):27-28. Meadowhawk (Odonata: Gomphidae)", J. Ks. Entomol. Soc., ____117. Sympetrum costiferum -- Saffron-winged 58(4):656-661. Meadowhawk ____118. Sympetrum internum -- Cherry-faced 10. Huggins, D.G., and M.F. Moffett, 1988, "Proposed Meadowhawk Biotic and Habitat Indices for Use in Kansas ____119. Sympetrum obtrusum -- White-faced Streams", Rept. No. 35 of the Kansas Biological Meadowhawk Survey. ____120. Sympetrum occidentale fasciatum -- Western Meadowhawk 11. Beckemeyer, R.J., 1995, "Some county records ____121. Sympetrum rubicundulum -- Ruby Meadowhawk for Kansas and Oklahoma", Argia: The news ____122. Sympetrum vicinum -- Yellow-legged journal of the Dragonfly Society of the Americas, Meadowhawk 7(3):28-29 ____123. Tramea carolina -- Carolina Saddlebags ____124. Tramea lacerata -- Black Saddlebags 12. Beckemeyer, R.J., 1996, "First record of ____125. Tramea onusta -- Red-mantled Saddlebags Brechmorhoga mendax from Kansas", Argia: The news journal of the Dragonfly Society of the Americas, 8(2):29-30 SOURCE DATA: 13. Todd, R., 1996 & 1997, Personal communication. 1. Banks. N., 1894, "On a collection of neuropteroid [Mr. Todd passed on to me voucher specimens of from Kansas", Ent. News, 4:178-180. Celithemis verna, Dythemis velox, and Tramea carolina, all of which are thus newly added to the 2. Tucker, 1907, "Some results of desultatory list. He also verified the occurrence of Anax collections of insects of and Kansas", longipes for Crawford Co. These specimens are Univ. Ks. Sci. Bull. (Odonata), 4:78-79. being placed in the KBS collection.]

3. Kennedy, C.H. 1917, "The Dragonflies of 14. Charlton, R., June 1997, Personal Kansas", Univ. KS Dept. Ent. Bull. 11:129-145 communication. Prof. Charlton has collected .[Kennedy's pioneering effort listed 64 species of Erythemis vesiculosa from Geary Co., and Lestes Odonata for the state.] eurinus from Pottawatomie Co. , both new to Kansas, and has collected Anax longipes from 4. Allison, V.C., 1921, "Some dragonflies of Woodson Co. southeastern Kansas", Trans. Kansas Acad. Sci., 30:45-58. 15. Beckemeyer, R., March, 1997, Notes taken on a group of specimens on loan to the author from the 5. Huggins, D.G., P.M. Liechti, and D.W. Roubik, USNM. Single specimens of Argia immunda from 1976, "Species Accounts for Certain Aquatic Cherokee County and from Macroinvertebrates from Kansas (Odonata, Labette County were found. In addition, Labette Hemiptera, Coleoptera and Sphaeriidae)", in County was added to the occurrence records for "New Records of the Fauna and Flora of Kansas Arigomphus submedianus. for 1975", Tech. Publ. of the State Biol. Surv. of Kansas, 1:13-77. 16. Beckemeyer, R.J., 1998, "Some midwestern Odonata records for 1997", Argia: The news 6. Huggins, D.G., 1978, "Additional Records of journal of the Dragonfly Society of the Americas, Kansas Odonata", in "New Records of the Flora 9(4):6. D.G. Huggins and verified by Roy Beckemeyer. There is no Epicordulia) Can. Entomol., 98:897-902); that is, making NOTES: locality record available for the photo other than it having both Epicordulia and Tetragoneuria subgenera of Epitheca. been taken in the state (somewhere in the southeastern 1. This list is arranged taxonomically by portion of Kansas). No. 77. Garrison, following the advice of K.J. Tennessen, SUBORDER : FAMILY, then alphabetically by synonomized Tetragoneuria williamsoni with T. costalis I : Species within each FAMILY. Entries No. 42. Telebasis salva is added to the list based on a have followed this advice (Tennessen, Personal include: Entry No. Genus species Author who specimen in the Florida State Collection of that Communication, 1995) as well, replacing the records of T. described species (In parentheses if species is was the source for the listing by Westfall and May of this williamsoni with . now assigned to a different genus than when species in Kansas. Mike May and Bill Mauffray kindly described by author), Date described -- Common tracked down the specimen, so the species is hereby added Name. No. 79. While Paulson (1996, Personal Communication) to the list vide Mauffray: A single envelope in the FSCA with and some other authorities consider Epitheca petechialis to the following data: "Telebasis salva Hagen 1 male, 1 be a synonym of Epitheca costalis, I follow the advice of 2. References used to update nomenclature include: female, Kansas, Sumner County, Hunters Mill Pond, 1 Tennessen (1995, Personal Communication), and retain R.W. Garrison (1991, "A Synonomic List of the September 1936, Eldon Kile (Collector), Ex Williamson- species status for E. petechialis pending further New World Odonata", Argia, 3(2):1-30); M.J. Kennedy Collection, Ex M.J. Westfall Collection". investigation. Westfall, Jr., & M.L. May, (1996, "Damselflies of North America", Scientific Publishers, Gainesville, Update: Mark O'Brien of the Univ. of Museum of FL);K.J. Tennessen, (1977, "Rediscovery of No. 95. Erythrodiplax umbrata is added based on a Zoology located 8 additional specimens that had been Epitheca costalis", Ann. Ent. Soc. Am., 70(2):267- specimen from the USNM collected by Gary F. Hevel on 11 collected by Eldon Kile (same location) on Sept. 1, 1936, 273); K.J. Tennessen, (1995, Personal July, 1964 from Oswego in Labette County (teneral male, and 4 that were collected Sept. 11, 1936. All were Communication regarding genus Epitheca); determined by R.W. Garrison, 1986). determined by L.K. Gloyd. Mark has donated a pair of the Dennis Paulson (1996, Personal Communication specimens to us for the Kansas Biological Survey regarding nomenclature). Please note the Update: A female specimen was collected in Sedgwick Co. Collection. following regarding the cited entries: by Roy J. Beckemeyer on 8 June, 1999. No. 9. The Amphiagrion sp. was identified by Huggins Update #2: Betsy Betros of Lenexa, Kansas photographed 3. Common names are from D.R. Paulson & S.W. (1976) as distinct from A. saucium and A. abbreviatum, T. salva (male) in Barber County, Kansas on 5 Sept. 1999. Dunkle, 1996, "Common names of North and therefore to be a new species. It remains to be Photo id by R.J. Beckemeyer. American dragonflies and damselflies, adopted by described and named, and was not listed by Garrison. the Dragonfly Society of the Americas", ARGIA, No. 45. Aeshna interruptaq lineata is added to the list (The late Mrs. L.K. Gloyd studied Amphiagrion for 8(2). Link to Internet Version of List based on three male specimens in the Kansas University many years. Rosser Garrison has told me that she had collection discovered by Ralph Charlton (1998). Labels carry examined over 3000 specimens of the genus and was 4. Entries 43 and 84 (indicated by "*" ) are listed as the following information: Specimen #1: "T.F. Winburn coll / planning on describing the "mid-American" form (AZ to "Species in Need of Conservation" in the State of Manhattan, Ks. / July '29"; Specimen #2: "Coll. / W C Dick / the Midwestern states) as A. mesonum. This name Kansas. Kansas / Marion Co. / Fall 1958"; Specimen #3: "Manh'tn, was in use as a manuscript name, and many workers Ks. / '23 Sept. / Aeshna male / male app = lineata / color during that time period therefore described 5. Entries marked with "^" were listed by Kennedy. female = interna". Amphiagrion from the region as A. sp. Some workers today apparently consider Amphiagrion as a species No. 65. Ophiogomphus rupinsulensis is known only from with gradual clinal variation from east to west. The nymphal material for Kansas. Cook & Daigle (1985) problem remains to be closed. Thanks to Rosser determined that specimens previously identified as this Garrison (Personal Communication, Sept. 1995) for species in and were really a new discussing the issue with me.) In their recent master species, O. westfalli. It is likely that the Kansas specimens work on the Zygoptera of North America, Westfall and are also of this new species but I have chosen to retain this May (1996) state: "There is evidence that populations entry as O. rupinsulensis until verification by examination of from the midwest, southwest into , probably the specimens has been accomplished. repesent a third species, but their status is as yet unclear. Amphiagrion not clearly assignable to either A. No. 74. Huggins listed and Garrison recognized both abbreviatum or A. saucium are known from: AZ, CO, Macromia illinoiensis and M. georgina as valid species. IL, IN, IA, KS, MI, NE, NM, ND, OK, SD, WI." However, T.W. Donnelly and K.J. Tennessen recently No. 14. Argia immunda is added to the list based on a declared M. georgina to be a subspecies of M. illinoiensis, specimen from the USNM that was collected from and the Kansas specimens they had inspected to be either Cherokee County on 27 April, 1963 by Gary F. Hevel intermediate forms (subspecies indeterminate) or of the (Male, determined by M. Davis, 1975) subspecies M. i. georgina (1994, "Macromia illinoiensis and georgina: A Study of Their Variation and Apparent No. 37. I have followed Garrison(1991) and Westfall & Subspecific Relationship (Odonata: Corduliidae)". Bull. of May (1996) by transferring Anomalagrion hastatum into American Odonatology, 2(3):27-61). I have therefore the genus Ischnura as I. hastata (Say), 1839. removed Macromia georgina from the Kansas list.

No. 41. Nehalennia gracilis is added to the list based on a No. 77-80. I follow Tennessen (1995, Personal photograph published in Insects in Kansas (2000, 3rd Communication) who recommends Walker's treatment Edition, S.C. White & G.A. Salsbury, Kansas Department of (1966, "On the generic status of Tetragoneuria and Agriculture, Topeka, KS: p. 52). The photo was identified by "Sympetrum semicinctum Say and its nearest County] HISTORICAL RECORDS: allies (Odonata)", Ent. News, 62: 153-163) 3. Anomalagrion hastatum [= Ischnura hastata (No. 37)] description of Sympetrum occidentale 4. Anax junius (No. 48) 1. Records of C.H. Kennedy (1917) fasciatum (No. 115). 5. Argia apicallis [sic] [= Argia apicalis] (No. 11) 6. Argia putridia [sic] [= Argia moesta (No. 15)] In addition to the 57 species noted by the "^" annotation Kennedy also listed a species which has subsequently 7. Argia violacea [= Argia fumipennis violacea (No. 13)] above, the following 7 species were also identified as been Synonomized: 8. Celethemis [sic] elisa [= Celithemis elisa (No. 86)] occuring in Kansas by Kennedy (1917). However, 9. Celethemis [sic] eponina [=Celithemis eponina (No. Huggins (Huggins et al, 1976) was unable to 1. Macromia australensis - Syn. of M. georgina - 87)] substantiate the occurrence of the first 5 when he Subsp. of M. illinoiensis (No. 72 - See Note 2). 10. Dromogomphus spoliatus (No. 56) reviewed the material in the Kansas University 11. Enallagma aspersum (No. 22) collection. Because there are also no recent records of Finally, the following species listed by Kennedy in 1917 12. Enallagma civile (No. 25) occurrence, we have listed them as Unsubstantiated: are synonyms of species in the current list: 13. Enallagma signatum (No. 30) 14. Epicordulia princeps [= Epitheca (Epicordulia) 1. Lestes forcipatus Rambur 1842. Huggins could 1. Agrion maculatum - Syn. of Calopteryx princeps (No. 79)] not locate Kennedy's material. Given the maculata (No. 1). 15. Ischnura verticalis (No. 40) confusion in old records between Lestes 16. Lestes unguiculatus (No. 8) disjunctus and Lestes forcipatus, these records 2. Hetaerina tricolor - Syn. of Hetaerina titia 17. Libellua auripennis [There are no current collection may all refer to L. disjunctus australis. See (No.3). records or known voucher species. None of the currently Walker, E.M., 1952, The Lestes disjunctus and documented species of Libellula from southeast Kansas forcipatus comples (Odonata: Lestidae), 3. Argia moesta var. putrida - Syn. of Argia could be easily confused with auripennis. The species Transactions of the American Entomological moesta (No. 15). has been recorded in Oklahoma and Missouri, and could Society, 78:59-74 very well occur or have occurred in Kansas. I thus 4. Æschna constricta - Syn. of Aeshna constricta choose to add L. auripennis to the list of (Burmeister 1839). 2. (No. 43). Unsubstantiated species of Odonata for Kansas.] Kennedy's material could not be located by 18. Libellula basalis [= Libellula luctuosa (No. 102)] Huggins, but all material identified as A. 5. Æschna multicolor - Syn. of Aeshna multicolor 19. Libellula cyanea (No. 98) saucium in the KU collection in 1975 was (No. 45). 20. Libellula pulchella (No. 104) determined by him to be Amphiagrion sp. nov. 21. Libellula vibrans (No. 108) (See Note 2). 6. Platycordulia xanthosoma - Syn. of 22. Macromia taeniolata (No. 75) Neurocordulia xanthosoma (No. 81). 23. Mesothemis simplicicollis [= 3. Morse 1895. No Kansas (No. 92)] specimens were found by Huggins in the KU 7. Perithemis domita var. tenera - Syn. of 24. Pachydiplax longipennis (No. 110) collection. Perithemis tenera (No. 111). 25. Pantala hymenaea (No. 112) 26. Pantala flavescens (No. 111) 4. (Walsh 1863). Only material 8. Sympetrum ambigua - Syn, of Sympetrum 27. Perithemis domitia [= Perithemis tenera (No. 113) from and was found in the KU ambiguum (No. 112). 28. Plathemis lydia [=Libellula lydia (No. 103)] collection by Huggins. 29. Sympetrum albifrons [= Sympetrum ambiguum (No. 2. Records of V.C. Allison (1921) 114) Note that for Tramea carolina (Linnaeus 1763). Kennedy 30. Sympetrum corruptum (No. 115) referenced material from Pratt & Clark Counties. Huggins (1975) reported that Allison's Crawford County 31. Sympetrum vicinum (No. 121) Huggins determined that the Clark Co. specimen was T. collection had apparently been lost, and he did not 32. Sympetrum semicinctum [Probably = Sympetrum onusta, but could not locate the Pratt Co. specimen. comment further on Allison's listing of 34 species. occidentale fasciatum (No. 119), as all other Kansas Ragan Todd of Pittsburg, KS (Crawford Co.) collected Allison noted that "'The Book', Howard; Hagen's specimens available that were labelled as semicinctum this species recently, thus justifying adding it back onto 'Neuroptera'; 'The Dragon Flies of Indiana' by E.B. were the latter species.] the list. Williamson, were used in this work". There is no 33. Tramea lacerata (No. 123) indication that he had any of his specimens verified by 34. Tramea onucta (No. 124) In addition, Huggins (et al, 1976) determined the any contemporary authorities on Odonata. I list his 34 following specimens of Kennedy to have been species and comment briefly on them here. Since there Misidentified: are no vouchers, the one species that would constitute and addition to the list is carried as Unsubstantiated. 1. Lestes congener Hagen 1861. Kennedy's Douglas Co. record was determined by 1. Æschna pentacantha [= Nasiaeschna pentacantha of Huggins to be Lestes unguiculatus (No. 8). this list (No. 52)] 2. Æschna verticalis [I strongly suspect this to be 2. Sympetrum semicinctum (Say 1839). Huggins Aeshna constricta. Allison noted that the species was determined that all adult specimens in the KU "Not rare", and A. constricta is present in Crawford collection matched Walker's (1951,