In the Caucasus Region
Multivariate analysis, systematics, and distribution of Galium sect. Orientigalium Ehrend. (Rubiaceae) in the Caucasus region IVAN SCHANZER & FRIEDRICH EHRENDORFER ABSTRACT SCHANZER, I. & F. EHRENDORFER (2002). Multivariate analysis, systematics, and distribu - tion of Galium sect. Orientigalium Ehrend. (Rubiaceae) in the Caucasus region. Candollea 57: 329-357. In English, English abstract. Galium sect. Orientigalium Ehrend. is a species rich and polymorphic clade centred in the western Irano-Oriental region, from where it extends to the Iberian Peninsula in the W, the Leba - non in the S, and the Caucasus region in the N. Preliminary chromosome counts and correlated pollen diameter measurements show that the clade includes several polyploid complexes. One of them, particularly diverse in the Caucasus, is the G. subvelutinum alliance. On the basis of an extensive evaluation of morphological differential characters, eighteen populations of this alliance were studied in the field, together with additional herbarium material resulting in a total of four hundred and three individuals scored for twenty differential characters and eventually arranged in twenty-six OTUs. This matrix was subjected to cluster analysis, multidimensional scaling, and dis - criminant analyses. The resulting tree reveals five major OTU clusters. Their structure, affinities, discontinuities and clinal connections are illustrated by 2-dimensional scatter plots and maps. As a consequence, it is possible to separate two informal species groups within the Caucasian mem - bers of the G. subvelutinum alliance: the ± suffruticose and more xerophilous G. mite group and the ± herbaceous and rather mesophilous G. hyrcanicum group. Both apparently contain diploid and polyploid members and are occasionally linked by hybridization. Within the G.
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