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Catalogue of Type Specimens of Beetles (Coleoptera) Deposited in the National Museum, Prague, Czech Republic*

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Catalogue of Type Specimens of Beetles (Coleoptera) Deposited in the National Museum, Prague, Czech Republic* ACTA ENTOMOLOGICA MUSEI NATIONALIS PRAGAE Published 30.vi.2012 Volume 52(1), pp. 297–334 ISSN 0374-1036 Catalogue of type specimens of beetles (Coleoptera) deposited in the National Museum, Prague, Czech Republic* Scarabaeidae: Scarabaeinae: Coprini, Eurysternini, Gymnopleurini and Oniticellini Aleš BEZDĚK1) & Jiří HÁJEK2) 1) Biology Centre ASCR, Institute of Entomology, Branišovská 31, CZ-370 05 České Budějovice, Czech Republic; e-mail: [email protected] 2) Department of Entomology, National Museum, Kunratice 1, CZ-148 00 Praha 4, Czech Republic; e-mail: [email protected] Abstract. Type specimens from the collection of beetles (Coleoptera) deposited in the Department of Entomology, National Museum, Prague, are currently being catalogued. In this part of the catalogue dealing with the Scarabaeoidea, we pre- sent precise information on the types of 77 taxa of the tribe Coprini, four taxa of Eurysternini, 10 taxa of Gymnopleurini and 20 taxa of Oniticellini. Key words. Catalogue, type specimens, National Museum, Scarabaeidae, Scara- baeinae, Coprini, Eurysternini, Gymnopleurni, Oniticellini Introduction This work represents the fi fth part of the catalogue of type specimens of beetles (Coleoptera) deposited in the collection of the National Museum, Prague, Czech Republic (NMP; NMPC when referring to the collection); the fi rst part was published by BEZDĚK & HÁJEK (2009). This part deals with the tribes Coprini, Eurysternini, Gymnopleurini and Oniticellini belonging to the subfamily Scarabaeinae. Along with the paper, a database of all the types including photos for most types and a copy of their original description is available upon request. As in the previous parts, we begin by presenting brief information about the most important collections mentioned in the catalogue: The collection of František Antonín Nickerl (1813–1871), Otakar Nickerl sen. (1838–1920) and Otakar Nickerl jun. (1873–1904) represents one of the oldest and largest collections in NMP. It is important as it contains a number of types from the 19th century, Nicklerls’ active * Catalogue of type specimens in NMPC, part 5 298 BEZDĚK & HÁJEK: Catalogue of types in NMPC: Scarabaeoidea (part V) communication, exchange and purchase of material brought, e.g., the types of Edgar von Harold (1830–1886). The Collection of Vladimír Balthasar (1897–1978) represents the most important specialised collection of the scarabaeoid families in NMP. It contains ca. 40,000 specimens including types of more than 500 species. Although, most of the species were described by V. Balthasar himself, the collection also contains types of some Balthasar’s other contemporaries: Hermann Luederwaldt (1865–1934), Zdeněk Tesař (1907–1985), Eric G. Matthews (born 1932), and Gonzalo Halffter (born 1932). Most recently, François Génier (born 1960) revised a large part of Balthasar’s taxa and described additional new species, of which numerous types are now deposited in Balthasar’s collection as well. Types from the collection of David Král (born 1959) comprise the complete type material (70 holotypes and 445 paratypes) of the collection of this contemporary Czech specialist on Scarabaeoidea. It includes, beside the species he described himself, specimens exchanged with other specialists, e.g., Kimio Masumoto, Jiří Zídek, and Svatopluk Pokorný. For additional information, see also BEZDĚK & HÁJEK (2009, 2010a,b, 2011). Material and methods Recent concepts of higher classifi cation in the Scarabaeoidea are largely inconsistent. The classifi cation used in this catalogue follows SCHOLTZ & GREBENNIKOV (2005). Tribal classifi ca- tion of the subfamily Scarabaeinae is adopted from HANSKI & CAMBEFORT (1991) and partially modifi ed according to MONTREUIL (1998) and VAZ-DE-MELLO (2008): Ateuchini, Canthoni- ni, Coprini, Eucraniini, Eurysternini, Gymnopleurini, Oniticellini, Onitini, Onthophagini, Phanaeini, Scarabaeini and Sisyphini. Because the subfamily Scarabaeinae is species rich, it is catalogued in three parts and particular tribes will be listed in alphabetical order. Within each subfamily/tribe, the genera and species are arranged alphabetically. Each entry includes: – the name of the taxon in original combination. – the name of the taxon in original combination and spelling, with the author and year of description. Pagination, fi gures and plates are also given. – the name-bearing type, number of specimens (including their sex if known) and exact label data. Our remarks are found in square brackets: [p] – preceding data are printed, [hw] – preceding data are handwritten. Separate labels are indicated by a double slash ‘//’ and lines within each label are separated by a slash ‘/’. – the current taxonomic status. – the type condition is mentioned for considerably damaged specimens. – any taxonomic problems and incosistencies are mentioned under Remarks. Full reference to each publication can be found in References. The types of infrasubspecifi c entities, i.e., names not regulated by the International Code of Zoological Nomenclature (ICZN 1999: Art. 1.3.4), are listed separately and briefl y. Acta Entomologica Musei Nationalis Pragae, 52(1), 2012 299 Catalogue Subfamily Scarabaeinae: Coprini Canthidium angulicolle Balthasar, 1939 Canthidium angulicolle Balthasar, 1939a: 111. The holotype is deposited in NMPC (ex coll. V. Balthasar): HOLOTYPE (): Mato grosso / Zobrys [p] // Heller determ. [p] / Choeridium / laterali Har. [hw] // Typ. [p, red label] // angulicolle m. [hw, green label, black margin]’. Current status. Canthidium (Canthidium) angulicolle Balthasar, 1939, see MARTÍNEZ & HALFFTER (1986). Canthidium atomarium Balthasar, 1939 Canthidium atomarium Balthasar, 1939a: 112. One syntype is deposited in NMPC (ex coll. V. Balthasar): SYNTYPE (unsexed specimen): ‘CAYENNE / Kourou Fl. / Le Moult [p] // Typus [p, red label, black margin] // C. atomarium / n. sp. [hw] / Dr. V. Balthasar det. [p] // atomarium m. [hw, green label, black margin]’. Current status. Canthidium (Eucanthidium) atomarium Balthasar, 1939, see MARTÍNEZ & HALFFTER (1986). Canthidium atramentarium Balthasar, 1939 Canthidium atramentarium Balthasar, 1939a: 113. The holotype is deposited in NMPC (ex coll. V. Balthasar): HOLOTYPE (): ‘Peru / W. Schnuse [p] / 28. 12. 03 [hw, green label] // 1910 / 8 [p] // Boucomont det. 1934 [p] / Canthidium sp. ? [hw] // Typus [p, red label, black margin] // C. atramentarium / n. sp. [hw] / Dr. V. Balthasar det. [p] // atramentarium m. [hw, green label, black margin]’. Current status. Canthidium (Eucanthidium) atramentarium Balthasar, 1939, see MARTÍNEZ & HALFFTER (1986). Canthidium basipunctatum Balthasar, 1939 Canthidium basipunctatum Balthasar, 1939a: 114. One syntype is deposited in NMPC (ex coll. V. Balthasar): SYNTYPE (): ‘Peruvia [hw] // Coll. B. / Schwarzer [p] // [blank red label] // Canthidium / basipunctatum [hw] / Dr. V. Balthasar det. [p] // basipunctatum [hw, green label, black margin]’. Current status. Canthidium (Canthidium) basipunctatum Balthasar, 1939, see MARTÍNEZ & HALFFTER (1986). Canthidium caesareum Balthasar, 1939 Canthidium caesareum Balthasar, 1939a: 115. The holotype is deposited in NMPC (ex coll. V. Balthasar): HOLOTYPE (): ‘Rio de Janeiro [p, green label] // Coll. C. Felsche / Kauf 20. 1918 [p, green label] // Typus [p, red label, black margin] // Canth. / caesareum / n. sp. [hw] / Dr. V. Balthasar det. [p] // caesareum m. [hw, green label, black margin]’. 300 BEZDĚK & HÁJEK: Catalogue of types in NMPC: Scarabaeoidea (part V) Current status. Canthidium (Canthidium) caesareum Balthasar, 1939, see MARTÍNEZ & HALFFTER (1986). Canthidium cavifrons Balthasar, 1939 Canthidium cavifrons Balthasar, 1939a: 116. One syntype is deposited in NMPC (ex coll. V. Balthasar): SYNTYPE (unsexed specimen): ‘Espirito Santo [p] // Coll. C. Felsche / Kauf 20. 1918 [p, green label] // Typus [p, red label, black margin] // Canthidium / cavifrons / n. sp. m [hw] / Dr. V. Balthasar det. [p] // cavifrons m. [hw, green label, black margin]’. Current status. Valid species. Canthidium clypeale Harold, 1867 Canthidium clypeale Harold, 1867: 43. One syntype is deposited in NMPC (ex coll. Nickerl): SYNTYPE (unsexed specimen): ‘Harold typ. [hw, green label] // TYPUS [p, red label, black margin] // Canthidium clypeale Harold, Bras. typ. [hw] // Canthidium clypeale Har. Bras. [hw, green label, black margin]’. Current status. Canthidium (Eucanthidium) clypeale Harold, 1867, see MARTÍNEZ & HALFF- TER (1986). Canthidium coerulescens Balthasar, 1939 Canthidium coerulescens Balthasar, 1939a: 117. Two syntypes are deposited in NMPC (ex coll. V. Balthasar): SYNTYPE (unsexed specimen): ‘Santa Inéz / (Ecuad.) / R. Haensch S. [p] // Typus [p, red label, black margin] // Can- thidium / coerulescens n. sp. [hw] / Dr. V. Balthasar det [p] // coerulescens m. [hw, green label, black margin]’. SYNTYPE (unsexed specimen): ‘O. ECUADOR / Jivaria / 17. 12. 05 F. Ohs. S. // Typus [p, red label, black mar- gin]’. Current status. Canthidium (Canthidium) coerulescens Balthasar, 1939, see MARTÍNEZ & HALFFTER (1986). Canthidium convexifrons Balthasar, 1939 Canthidium convexifrons Balthasar, 1939a: 119. One syntype is deposited in NMPC (ex coll. V. Balthasar): SYNTYPE (): ‘Pischindé / Colombia / (Rosenberg) [p] // Coll. C. Felsche / Kauf 20. 1918 [p, green label] // Typus [p, red label, black margin] // Canthidium / convexifrons / n. sp. typ [hw] / Dr. V. Balthasar det [p] // Mus. Nat. Pragae [p] / 26233 [hw] / Inv. [p, orange label] // convexifrons m. [hw, green label, black margin]’. Current status. Canthidium (Canthidium) convexifrons
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