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September 2001
A checklist of the Limnichidae and the Lutrochidae (Coleoptera) of the world
P. J. Spangler Smithsonian Institution, Washington, DC
C. L. Staines Smithsonian Institution, Washington, DC
P. M. Spangler Smithsonian Institution, Washington, DC
S. L. Staines Smithsonian Institution, Washington, DC
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Spangler, P. J.; Staines, C. L.; Spangler, P. M.; and Staines, S. L., "A checklist of the Limnichidae and the Lutrochidae (Coleoptera) of the world" (2001). Insecta Mundi. 197. https://digitalcommons.unl.edu/insectamundi/197
This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI. Vol. 15. No. 3. Se~tember.2001 151
A checklist of the Limnichidae and the Lutrochidae (Coleoptera) of the world
P. J. Spangler, C. L. Staines, P.M. Spangler, and S.L. Staines Department of Entomology National Museum of Natural History Smithsonian Institution Washington, DC 20560
Abstract. A checklist of the world species of Limnichidae (35 genera, 345 species) and Lutrochidae (1 genus, 11 species) is presented. The author, year of publication and page number, synonyms, distribution by country, and a terminal bibliography are given for each genus and species. Biological information is also reviewed.
Introduction scribed the species and adult behavior of Lutrochus arizonicus Brown and Murvosh. Hilsenhoff and The Limnichidae Erichson (1847) and Lutro- Schmude (1992) reported that adults of L. laticeps chidae Kasap and Crowson (1975) are small semi- Casey (1893) were found in small to large, warm, aquatic beetles; adults of both families most fre- calcareous streams; that adults oviposit on sub- quently live in riparian habitats. Both families merged wood or on travertine rocks; and larvae were previously assigned to the superfamily Dry- were aquatic and were collected on the travertine opoidea (Crowson, 1978); however, they are now rocks. placed in the Byrrhoidea (Lawrence and Newton, Previous world catalogs that included the lim- 1995). From 1975 through 1993, D. P. Wooldridge nichids and lutrochids are by Zaitzev (1910) and published numerous limnichid descriptions and von Dalla Torre (19 11) who recorded 11 genera and revisions but much systematic work remains to be 75 species. Blackwelder (1944) published a check- done for the Limnichidae, especially in the tropical list of the Coleoptera of Mexico, Central America, regions of the Eastern Hemisphere. the West Indies, and South America in which he Not much has been published dealing with the reported 11 genera and 44 species of Limnichidae biology of the families. Paulus (1970) described the and 8 species of Lutrochidae in the nominate genus. larva of the European species Limnichus sericeus Wooldridge (1986b) published a catalog of the Lim- Duftschmid (1825) and the larva and pupa of Pelo- nichidae (including Lutrochus) of North America in chares versicolor Walt1 (1838). Paulus also reported which he included 6 genera and 29 species of that the larvae of both genera are terrestrial and Limnichidae and the single genus, Lutrochus with live in the soil under the "Chlorophyceae-Bryophy- 3 species. Some recent revisonary studies on the ta layers," and reportedly feed on those plants. Limnichidae are those by Wooldridge (1976,1977a, Adults may be found near the waterline on 1977b, l978,1979,1980,1981b, 1982,1983,1987b). branches and grasses hanging over stream banks Additional studies that have added to our knowl- or in riffles, on driftwood, in logjams, and on rocks edge of the Limnichidae are by Darlington (1936), in streams; some are found in splash zones beside fauna of the West Indies; Wu (1937), China; SatG cascades and waterfalls and on dam breasts. Spec- (1965), Formosa; Sat6 (1966, 1976, 1997), Japan; imens of at least five limnichid genera, Pseudeu- Deleve (1968, 1974), Herando and Ribera (2000a), cinetus Heller (1921), Martinius Spilman (1959), Africa; Del&ve(1971,1973), Asia; Wooldridge (1975), Hyphalus Britton (1971, 1973, 1977), Throscinus Western Hemisphere; Putz (1991b), Germany; LeConte (1874) (fide Wooldridge, 1981), andMexico Mascagni (1995), Italy; Putz (1998a, 1998b), China Spilman (1972) are known to live on coral reefs and neighboring countries; Ribera and Hernando (SatG, 1997; Hernando and Ribera, 2000b), coastal (1999), and Hernando and Ribera (2000b), Oriental lagoons, mangrove swamps, or edges of mud or Region. rocky flats along coastal areas. This checklist is an effort to bring the limnichid Brown (1965) discussed characteristic micro- and lutrochid names and pertinent taxonomic liter- habitats of various dryopoids including Lutrochus ature into a single publication and was compiled by Erichson (1847). Brown and Murvosh (1970) de- examining the literature for descriptions, synonyms, 152 Volume 15, No. 3, September, 2001, INSECTA MUNDI and distributional data plus examination of the biroi Pic, 1956:75 collection of the Smithsonian Institution. New Guinea bomansi (DelAve), 1968:245 A checklist of the Limnichidae Tanganyika of the world borneensis Pic, 1923b:7 Borneo ACONTOSCELES Champion, 1924a:27 boucardi Pic, 1923b:7 hydroporoides Champion, 1924a:29 Sumatra Taiwan [Formosa], India brevior (Pic), 1927a:65 tagalog Spilman, 1959:116 Sumatra Philippines convexus (Pic), 1923b:5 yorioi SatG, 1966:60 Philippines Japan (Ryukyus) cribrosus Champion, 1923:253 India, Sri Lanka AFROLIMNICHUS DelBve, 1968:259 curtus Pic, 1935b:471 brevicornis DelAve, 1968:260 Sierra Leone Congo densepunctatus (Pic), 1923b:6 oblongus DelAve, 1974:40 Ivory Coast South Africa (Cape Province) diversus (Pic), 1923b:5 Ivory Coast BABALIMNICHUS SatG, 1994:174 ellypticus Motschulsky, 1858:51 masamii Sat& 1994:175 India Japan (Ryukyus) euryaspis Champion, 1923:250 taiwanus SatG, 1994:174 India [Nilgiri Hills] Taiwan ferax Wooldridge, 1993b:359 Philippines, Borneo BOTHRIOPHORUS Mulsant and Rey, 1852:19 flavicornis Champion, 1923:251 atomus Mulsant and Rey, 1852:20 India (syn., ventetus Ferrari, 1864:108) formosanus Pic, 1935a:2 Europe Taiwan [Formosa] fuscus Motschulsky, 1858:52 BYRRHINUS Motschulsky, 1858:50 India syn., BYRRHININUS Pic, 1922:6 goldbachi Wooldridge, 1987b:310 syn., CYPHONICHUS Sharp, 1902a:679 Ecuador, Paraguay syn., NOTIOCYPHON Blackburn, 1896:272 gracilicornis Pic, 1922:5 syn., PELOCHEROPS Pic, 1923a:3 Brazil, Paraguay, Venezuela amoenus Wooldridge, 1987a:310 grandis (Pic), 1956:74 Belize, British Guiana, Colombia, Ecuador New Guinea andamanus (Pic), 1935a:2 grossepunctatus (DelBve), 1968:249 Andaman Islands Congo angustatus Motschulsky, 1858:51 hargreavesi (Pic), 1935b:470 India, Thailand Sierra Leone antipodium Fauvel, 1903:348 hirsutulus Champion, 1923:250 New Caledonia India [Kumaon] apicalis Pic, 1922:5 hirsutus Pic, 1922:5 Katanga French Guiana, Trinidad atricornis (Pic), 1935a:3 hulstaerti (Pic), 1952:9 Congo, Sierra Leone Kinshasa [Congo] atrolucidus (Pic), 1923a:3 improcerus Wooldridge, 1987b:310 New Guinea Ecuador binhanus (Pic), 1923b:5 inlineatus Pic. 1923c:15 Vietnam India INSECTA MUNDI, Vol. 15, No. 3, September, 2001 153 irregularis Pic, 1922:5 plenus Wooldridge, 1987b:305 Philippines Boliva, Ecuador jacobsoni Pic, 1927a:65 pubiventris Lea:1920:277 Sumatra Australia latus Motschulsky, 185850 punctaticeps Pic, 1927b:49 Bangladesh, Burma, India, Sri Lanka Sumatra leleupi (Delitve), 1968:257 punctatus (Pic), 1922:5 Kivu [Zaire] Philippines leonensis (Pic), 1923b:6 punctulatus (DelGve), 1968:243 Congo, Sierra Leone Upemba [Zaire] maculatus Wooldridge, 1987b:307 reitteri (Pic), 1923b:6 Ecuador Ghana magnus Wooldridge, 1987b:307 rhomboideus Champion, 1923:253 Ecuador, Peri5, India marginatus Champion, 1923:249 rufoapicalis (Pic), 1955:132 India [Kumaon], Pakistan Ruanda, Kiva [Zaire] maroniensis Pic, 1923b:7 rufus Pic, 1935a:Z Brazil, Colombia, French Guiana, Trinidad- Thailand Tobago, Venezuela schoutedeni (Pic), 1952:9 mimicus Wooldridge, 1987b:308 Congo Colombia, Panam6 semirufus Pic, 1923b:8 misellus (Sharp), 1902a:681 Trinidad-Tobago, Venezuela Colombia, Costa Rica, MQxico, Nicaragua, seydeli (Delhve), 1968:252 Panam6 Elisabethville [Zaire] monilicornis Champion, 1923:251 sikkimensis Champion, 1923:249 India [Kumaon] India [Sikkim] neoguineensis (Pic), 1956:75 subglobosus (Delitve), 1968:254 New Guinea Elisabethville [Zaire] niger Pic, 1922:5 subglobosus upembanus (Delitve), 1968:256 Vietnam Upemba [Zaire] nitidicollis Pic, 1956:75 subregularis Pic, 1923c:15 New Guinea Vietnam nitidulus Motschulsky, 1858:51 subtestaceous Pic, 1923b:8 India, Thailand Philippines noctivagus Lea 1920:277 tarawakanus DelBve, 1973:23 Australia Borneo [Sabah], Philippines (Tawi Tawi) oculatus Motschulsky, 1858:52 testaceicornis (Pic), 1923b:5 India Philippines olibroides (Sharp), 1902a:679 testaceipes Pic, 1922:5 (syn., celatus Sharp, 1902a:680) subregularis Pic, 1923b:7 Lapsus Calami Belize, El Salvador, Guatemala, MQxico Sumatra orientalis (Sharp), 1902b:63 tschoffeni (Delitve), 1968:235 Taiwan [Formosa], Malaysia (Borneo) Boma [Congo] padangus Pic, 1927b:49 vatovai (Pic), 1953:113 Sumatra Ethiopia panamensis (Sharp), 1902a:679 vestitus (Sharp), 1902a:681 (syn., mollis Sharp, 1902a:681) Guatemala, MBxico (syn., rufipennis Pic, 1923b:7) vicinus (Delitve), 1968:248 Belize, Colombia, Costa Rica, Guateniala, Congo Panam5 wittei (Delitve), 1968:253 peguensis Pic, 1923c:15 Upemba [Zaire] India 154 Volume 15, No. 3, September, 2001, INSECTA MUNDI
CACCOTHRYPTUS Sharp, 1902b:63 nepalensis Putz, 1998b:360 syn., MACROBYRRHINUS Pic, 1922:4 Nepal (Myagdi] compactus Sharp, 190213363 schillhammeri Putz, 1998b:362 Borneo, Vietnam China [Sichuan] laoseizsis Pic, 1928:2 schuelkei Putz, 1998b:363 Laos China [Sichuan] maculosus Pic, 1923a:3 sichuanensis Putz, 1998b:365 Tonkin China multiseriatus Champion, 1923:224 subelongatus Pic, 1923b:4 India [Bengal] India [Sikkim] ripicola Champion, 1923:223 India [W. Almora] CHIBIDORONUS Sat6, 196659 rouyeri (Pic), 1922:4 aureus Sat6, 1966:60 Sumatra Japan (Ryukyus) testudo Champion, 1923:222 India m.Almora], Nepal CORRINEA Wooldridge, 1980:69 admirandus (Sharp) 1902a:677 CEPHALOBYRRHINUS Pic, 1923b:6 MQxico brevis Wooldridge, 1986a:512 audax Wooldridge, 1980:75 Ecuador MQxico curticornis Pic, 1923b:6 aurata Wooldridge, 1980:71 Costa Rica, Mexico, Nicaragua, Panama, Brazil, Ecuador Venezuela delicata Wooldridge, 1980:75 impressopunctatus Wooldridge, 1977b:32 Colombia, Costa Rica, MQxico,Panam6 Bolivia edita Wooldridge, 1993a:36 lineatus Wooldridge, 1986a:513 Venezuela Costa Rica, Panama incondita Wooldridge, 1980:73 robustus Wooldridge, 1986a:512 Bolivia, Colombia, Ecuador Ecuador, Peru lecta Wooldridge, 1980:74 sedatus Wooldridge, 1986a:513 Bolivia Ecuador parva Wooldridge, 1980:74 Bolivia, Ecuador, Perii CEPHALOBYRRHUS Pic, 1923b:4 placida Wooldridge, 1980:76 bertiae Putz, 1998b:344 Trinidad China mnnan] salebrosa Wooldridge, 1980:75 brevipalpus Putz, 1998b:346 Guatemala China yjrunnan] spicula Wooldridge, 1980:73 emeishanensis Putz, 1998b:347 Ecuador China [Sichuan] splendida Wooldridge, 1980:71 gibbicollis Champion, 1925a:175 Ecuador, Perii India [Kumaon] vianai (Pic), 1939:249 jaechi Putz, 1998b:351 Argentina China [Sichuan] japonicus Champion, 1925a: 176 CYCLOLIMNICHUS DelBve, 1968:270 Japan dentoni Hernando and Ribera 2000a:216 jiangxielzsis Putz, 1998b:354 Cameroon China [Jiangxi] jaechi Hernando and Ribera 2000a:215 jinggangshanensis Putz, 1998b:355 Kenya China [Jiangxi] ovalis Hernando and Ribera 2000a:215 latus Pic, 1923b:4 Cameroon Taiwan [Formosa] presignis DelBve, 1968:272 longipalpis Putz 1998:359 Congo China [Jiangxi] INSECTA MUNDI, Vol. 15, No. 3, September, 2001 155
ERSACHUS Erichson 1847:497 Mkxico, USA (AZ, CA, CO, NM, OR) erichsolzianus Sharp, 1902a:683, pl. 20 nitidulus (LeConte), 1854:117 Guatemala, Mkxico, Panamii Argentina, Brazil, Costa Rica, Ecuador, lugubris Sharp, 1902a:684 MQxico, Nicaragua, Panamii, USA (AL, FL, Guatemala GA, IN, ME, MO, NC, NH, PA, SC, VA) mexicanus Sharp, 1902a:684 obscurus (LeConte), 1854:116 MQxico USA (DC, IL, IN, KS, MD, MS, NC, NY, TN, variegatus Erichson, 1847:497 TX, VA) Colombia optatus Sharp, 1902a:675 Guatemala, MQxico,USA (AL, GA, LA, TX) EULIMNICHUS Casey, 1889:146 pellucidus Wooldridge, 1979:5 acutus Wooldridge, 1979:7 Venezuela Costa Rica, Colombia, El Salvador, Guatemala, perpolitus Casey, 1889:151 Honduras, MQxico, Panamii USA (AZ, CA, NM, TX) analis (LeConte), 1879:515 plebius Sharp, 1902a:675 (syn., arzalis rugiceps Casey, 1912:47) (syn., Limnichoderus convexus Pic, 1922:4) (syn., analis coloradeasis Casey, 1912:47) (syn., Limnichoderus breyeri Pic, 1930:43) MQxico, Panamii, USA (AR, AZ, CA, CO, MO, (syn., Limnichoderusperuvianus Pic, 1954:178) OR, TX, UT, WA) Argentina, Brazil, Colombia, Dominican ater (LeConte), 1854:117 Republic, Ecuador, El Salvador, Guatemala, (syn., guatemalicus Sharp, 1902a:676) Haiti, Honduras, Mkxico, Puerto Rico, Perti, (syn., sculpticeps Casey, 1912:50) Trinidad, Venezuela (syn., Limnichoderus oblongus Pic, 1922:4) rugulosus Wooldridge, 1979:6 Argentina, Belize, Bolivia, Brazil, British USA (IL, MI, WI) Guiana, Colombia, Costa Rica, Cuba, rusticus Wooldridge, 1979:7 Ecuador, Guatemala, Honduras, Jamaica, MQxico Mkxico, PanarnA, Perti, USA (FL, GA, IN, sharpi Wooldridge, 1979:5 LA, MA, MS, NC, SC, TX), Venezuela Mexico californicus (LeConte), 1879:515 sordidus Sharp, 1902a:676 USA (CA, OR) Guatemala coheni Wooldridge, 1979:4 spangleri Wooldridge, 19799 Bolivia, Ecuador El Salvador, Guatemala, Honduras, MQxico confertus Sharp, 1902a:676 subtitus Wooldridge, 1979:8 MQxico, Guatemala USA (AL, FL) corrineae Wooldridge, 1979:3 sublaevis Sharp, 1902a:674 Argentina, Brazil, Venezuela Guatemala, Panamii episternus Sharp, 1902a:673 visendus Wooldridge, 1979:1 Costa Rica, Guatemala, Nicaragua, MQxico MQxico evanescens Casey, 1912:48 USA (CA) EUTHRYPTUS Sharp, 1902a:682 expeditus Wooldridge, 19841357 orbicularis Sharp, 1902a:682 Guatemala, Honduras Guatemala, Mkxico, Panamii impostus Wooldridge, 1978:172 trigonalis Sharp, 1902a:682 (syn. elongatus Pic, 1923b:7; preoccupied) Guatemala Brazil irnprocerus Wooldridge, 1979:4 HYPHALUS Britton, 1971:88 USA OVLA) crowsoni Hernando and Ribera 2000b:240 incultus Wooldridge, 1979:6 Seychelles [Aldabra] Bolivia, Ecuador insularis Britton, 1971:90 langleyae Wooldridge, 1979:3 Australia Ecuador, Per6 kuscheli Britton, 1977:82 montanus (LeConte), 1879:514 New Zealand (syn., laeticulus Sharp, 1902a:674) 156 Volume 15, No. 3, September, 2001, INSECTA MUNDI prolixus Britton, 1977:85 angustus Wooldridge, 1981b:180 New Zealand Colombia, Ecuador, Panam6, Venezuela taekoae Satb, 1997:llO barberi Wooldridge, 1981b:174 Japan (Ryukyus) Haiti, Puerto Rico, Trinidad-Tobago ultimus Britton, 1977:85 conspectus Wooldridge, 1981b:181 New Zealand Ecuador, Perii wisei Britton, 1973:121 cupidus Wooldridge, 1981b:178 New Zealand Brazil curtulus Sharp, 1902a:678 JAECHOBYRRHINUS Putz, 1991a:132 (syn., minutus Pic, 1922:4) amanosius Piitz, 1991a: 132 Costa Rica, Panam6 Turkey durus Wooldridge, 198113: 182 Brazil LICHMINUS Casey, 1889:147 excelsus Wooldridge, 1993a:35 tenuicornis Casey, 1889:155 Venezuela USA (CA) fragosus Wooldridge, 198113: 178 Ecuador LIMNICHITES Casey, 1889:147 itnprioris Blackwelder, 1944:274 browni Wooldridge, 1977a:187 (syn., oblongus BrQthes,1922:297; preoccupied) El Salvador, MBxico Argentina confertus (Sharp), 1902a:676 i~zsularisWooldridge, 1981b:188 Costa Rica, Guatemala, Honduras, Jamaica, Dominican Republic, Haiti, Puerto Rico MBxico, Nicaragua lutrochinus (LeConte), 1879:515 foraminosus Casey, 1912:43 (syn., indiscretus Sharp, 1902a:678) MBxico, USA (AZ, CA, NM, OR, TX, UT) (syn., lanosus Casey, 1912:55) huronicus Casey, 1912:43 Belize, Costa Rica, El Salvador, Guatemala, Canada, USA (IL, MA, MI) Honduras, MQxico, Panam6, USA (AZ, CA, imparatus Wooldridge, 1977a: 181 IL, OK, TX) MQxico magnus Wooldridge, 198113: 179 nebulosus (LeConte), 1879:515 Costa Rica, Guatemala, Mkxico (syn., densissimus Casey, 1912:41) modicus Wooldridge, 1981b:176 MQxico,USA (AR, AZ, CA, OK, OR, TX) Costa Rica, Ecuador, El Salvador, Guatemala, olivaceus (LeConte), 1854:116 Honduras, Panam5 (syn., virginicus Casey, 1912:42) moratus Wooldridge, 198113: 185 USA (AL, AR, CT, DC, IA, IL, IN, MA, MD, MI, Colombia, Costa Rica, Guatemala, MBxico, MO, NC, OH, OK, PA, TX, VA, WV) Panam6 perforatus (Casey), 1889:155 naviculatus (Casey), 1889:156 MBxico, USA (CA) Costa Rica, El Salvador, Guatemala, Honduras, porrectus Wooldridge, 1977a:182 Mexico, Nicaragua, Panam6, Puerto Rico, MBxico USA (AZ, CA, NM, NV,OK, TX) punctatus (LeConte), 1854:116 ovatus (LeConte), 1854:117 (syn., austinianus Casey, 1912:42) (syn., pulvereus Casey, 191254) Canada, USA (AL. DC, GA, IA, IL, MA, MD, MQxico,USA (AL, FL, GA, LA, NC, SC, TX) ME, MI, MN, MS, NH, NJ, NY, OH, OK, PA, placidus Wooldridge, 1987a:330 RI, TX, UT, VA, WI) Costa Rica rudis Wooldridge, 1977a:183 plenus Wooldridge, 198113: 185 Cuba, Jamaica MBxico simplex Wooldridge, 1977a:188 punctiventris (Casey), 1889:159 Costa Rica, MBxico USA (FL) seclusus Wooldridge, 1987a:330 LIMNICHODERUS Casey, 1889:147 Ecuador amoenus Wooldridge, 198113: 179 seriatus (Casey), 1889:158 Ecuador (syn., floridanus Casey, 1912:55) INSECTA MUNDI, Vol. 15, No. 3, September, 2001 157
USA (AL, FL, GA, LA, MS, NC, SC, VA) inornatus Weise, 1877:301 similis Wooldridge, 1981b:182 (syn., punctipennis Baudi, 1870:64) Perii Cyperus vicinus Pic, 1922:4 javanicus Pic, 1922:3 Brazil, Colombia, Ecuador, PanamB, Venezuela Java latiusculus Reitter, 1901:72 LIMNICHOMORPHUS Pic, 1922: 4 Turkestan curtulus Pic, 1922:4 lederi Weise, 1877:302 Vietnam (Tonkin) Caucasus ohbayashii Sat6, 1966:58 lewisi Nakane, 1963:43 Japan (Ryukyus) (syn., paululus Lewis, 1879:12) China, Japan, Russia LIMNICHUS Latreille, 1829:510 longitarsis Champion, 1923:275 aeneipennis Pic, 1935a:l India [Sarda] Vietnam marmoratus Fairmaire, 1898:224 africanus Pic, 1923a:3 Madagascar Ghana micraspis Champion, 1923:276 angustulus Weise, 1877:300 India [Sarda] Sardinia monticola DelBve, 1968:220 astruci Pic, 1922:3 Kenya India murinus Baudi, 1870:63 ater Lea 1920:279 Cyperus Australia nigripes Broun, 1893:1449 aurosericus Jacquelin du Val, 1857 (1859):268 New Zealand (syn., leprieuri Perris, 1864:282) obliteratus Champion, 1923:275 Algeria, France, Italy, Spain India [Kumaon] australis Erichson, 1842:153 orientalis Motschulsky, 1861:112 Australia, Tasmania Sri Lanka brevior Fairmaire, 1898:224 palawanus DelBve, 1973: 18 Madagascar Philippines (Palawan) coomani Pic, 1935a:l picinus Broun, 1881:671 Vietnam New Zealand cuneatus Champion, 1923:275 pruinosus Fleischer, 1909:244 India [Kumaon] Samarkand curtus Pic, 1922:3 pumilio Obenberger, 1916:29 Java Corfu decorus Broun, 1880:247 punctatus Broun, 1880:247 New Zealand New Zealand demoulini DelBve, 1968:217 punctipennis Kraatz, 1858:148 Congo Greece, Italy ellipticus Lea 1920:280 pygmaeus Sturm, 1807:114 Australia (syn., sericeus Stephens, 1832:411) falsus DelBve, 1971:371 (syn., auricomus Reitter, 1877 (1878):180) India (syn., minutus Rey, 1889:66) fragilicornis Wollaston, 1868:80 (syn., variegatus Guillebeau, 1886:2) Cape Verde Islands (syn., pygmaeus var. Reitter, 1887:104) fulvopubens Pic, 1922:3 Turkestan, middle Europe, Caucasus China ruficolor Pic, 1935a:2 globosus Champion, 1923:276 N. Borneo India [Kumaon] rufipes Pic, 1923b:6 incanus Kiesenwetter, 1851:584 Ivory Coast Mediterranean area, southern Europe, rufulopubens Fairmaire, 1897:368 Caucasus, Turkestan Madagascar 158 Volume 15, No. 3, September, 2001, INSECTA MUNDI rufus Pic, 1923b:7 PARATHROSCINUS Wooldridge, 1984c:121 Philippines (Luzon) corrinae Wooldridge, 1990:254 saegeri DelBve, 1968218 Singapore Congo magnus Wooldridge, 1990:254 sericeus Duftschmid, 1825:24 Singapore Europe, Asia murphyi Wooldridge, 1990:251 subchalybeus Reitter, 1881:85 Singapore Caucasus oculatus (Motschulsky), 1858:52 subelongatus Pic, 1922:3 Burma, India Sumbawa orientalis Wooldridge, 1984c:122 subnotatus Champion, 1923:273 Philippines, Vietnam India [Kumaon] sulcatulus Pic, 1922:3 PELOCHARES Mulsant and Rey, 1869:161 Guadeloupe acumiizatus Champion 1923:271 tonkineus Fairmaire, 1888:337 India [Sikkim] Tonkin, Vietnam acutipennis Pic, 1935a:2 trachyformis Pic, 1922:3 Malaysia India [Kumaon] alluaudi DelBve, 1968:226 turkestanicus Pic, 1905:97 Africa Orientale Turkestan basilewskyi (Pic), 1955:131 unistriatus Champion, 1923:274 Ruanda India [Kumaon] biroi (Pic), 1956:75 vagegluttatus Fairmaire, 1898:223 New Guinea Madagascar congoensis DelBve, 1968:221 waelbroecki Delkve, 1968:215 Congo Kinshasha densatus Pic, 1922:3 Java MANDERSIA Sharp, 1902b:62 diversus (Pic), 1922:3 scymnoides Sharp, 1902b:62 Philippines (Los Banos) Burma, China [Shan states], India emarginatus Mulsant and Rey, 1869:161 Europe MARTINIUS Spilman, 1959:118 gibbipennis Champion, 1923:269 ripisaltator Spilman, 1966:124 Europe, Burma, India [Bengal], Sri Lanka Cuba minutus DelBve, 1968:224 tellepontis Spilman, 1959:119 Kivu [Zaire] Panamh muriizus Baudi, 1870:63 temporalis Wooldridge, 1988:314 Cyperus, Caucasus Ecuador oblongulus Champion, 1925b:263 India (Kumaon) MEXICO Spilman, 1972:113 orientalis (Motschulsky), 1861:112 litoralis Spilman, 1972: 114 (syn., horizi DelBve, 1971:368) Mhxico Sri Lanka philippinensis DelBve, 1973:21 PARALIMNICHUS DelBve, 1973:24 Philippines (Tawi Tawi) castaneus (Lea), 1920:279 rugiventrus Champion, 1923:270 Australia (syn., ceylanicus DelBve 1971:370) panamensis Wooldridge, 1983:232 India [Kumaon], Sri Lanka Panamh ryukyuensis Sat6, 1966:57 punctatus (Pic), 1923b:5 China, Japan (Ryukyus) (syn., punctatus var. obscurior Pic, 1923b:5) sulciger Champion, 1923:270 (syn., rufescens Pic, 1923b:5) Sri Lanka Borneo, Guadalcanal, Philippines [Banguey] trifidus Champion, 1923:272 Burma INSECTA MUNDI, Vol. 15, No. 3, September, 2001 159
versicolor Waltl, 1838:273 PLATYPELOCHARES Champion,1923:271 (syn., emarginatus Mulsant and Rey, 1869: 161) syn. WOOLDRIDGEUS Spangler, 1999:181 (var., opacus Reitter, 1881:84) latimargo Champion, 1923:272 Europe, New Caledonia India [Kumaon] vitalisi Pic, 1935a: 1 perforatus (Spangler), 1999:182 Cambodia Malaysia periculosissimus Ribera and Hernando, 1999:107 PHALACRICHUS Sharp, 1902a:678 Loas, Thailand, Vietnam syn., LIMNICHALL4 Casey, 1912:52 petrus Ribera and Hernando, 1999:108 atomarius Sharp, 1902a:679 Malaysia (syn., colonica Casey, 191252) trifidus Champion, 1923:272 Belize, Bolivia, Colombia, Costa Rica, Ecuador, Burma El Salvador, Guatemala, Honduras, MQxico, Nicaragua, Panamh, PerG, Surinam, PSEUDEUCINETUS Heller, 1921:155 Trinidad, Venezuela syn., THAUMASTODUS Champion, 1924a:25 diligens Wooldridge, 1982:389 PSEUDOEUCHINETUS Sate, 1994:173 Bolivia, Brazil, Colombia, Venezuela (misspelling) durus Wooldridge, 1982:387 novabritannica Deliive, 1973:18 Bolivia, Brazil, Ecuador, Paraguay, PerG, Bismarck (New Britain) Venezuela spilmani Spangler, 1995:399 golbachi Wooldridge, 1982:388 Indonesia Paraguay ueizoi Spangler, 1995:396 latus Wooldridge, 1982:386 Malaysia Brazil, Colombia, Ecuador, PerG, Venezuela zygops Heller, 1921:156 punctatus Wooldridge, 1982:389 (syn., Thaumastodus fusiformis Champion, Bolivia, Brazil, Colombia, Panamh, Venezuela 1924a:27) rudis Wooldridge, 1982:385 India, Malaysia, New Ireland, Philippines, Surinam, Venezuela Solomon Islands simplex Wooldridge, 1982:387 Bolivia, Brazil, Venezuela RESACHUS Deliive, 1968:263 spangleri Wooldridge, 1982:386 lineatifrons Deliive, 1968:266 Ecuador, Venezuela Congo sublimus Wooldridge, 1993a:36 striatellus Deliive, 1968:263 Venezuela Congo vianai (Pic), 1938:77 Argentina, Brazil, British Guiana, Trinidad SIMPLOCARINA Pic, 1922:4 syn., CYRTOLIMNICHUS Deliive, 1968:267 PHYSEMUS LeConte, 1854:117 curticollis Pic, 1922:4 syn., DITAPHRUS Casey, 1886:250 (syn., puizctulatus Deliive, 1968:267) excavatus Wooldridge, 1976:182 Congo El Salvador, Guatemala, Honduras latifrons Wooldridge, 1976:180 THROSCINUS LeConte, 1874:51 Panamh aethiops Darlington, 1936:75 levis Wooldridge, 1984a:538 Cayman Islands, Cuba, Dominican Republic, Brazil, Trinidad, Venezuela Haiti, Jamaica, Puerto Rico minutus LeConte, 1854:117 crotchi LeConte, 1874:52 (syn., Ditaphrus scymnoides Casey, 1886:250) Mexico, USA (CA) MQxico, USA (AZ, CA, TX) politus Casey, 1889:162 mirus Wooldridge, 1984a:538 USA (TX) Ecuador punctatus Wooldridge, 1981a:220 punctatus Wooldridge, 1976:180 Colombia Colombia schwarzi Schaeffer, 1904:204 Jamaica, USA (TX) 160 Volume 15, No. 3, September, 2001, INSECTA MUNDI simplex Wooldridge, 1981a:220 busdam additis. Berliner Entomologische Colombia Zeitschrift 14: 49-90. spangleri Wooldridge, 1981a:218 Blackburn, T. 1896. Coleoptera (exclusive of the Ecuador Carabidae). pp. 254-308 in B. Spencer (ed.). Report of the work of the Horn Scientific Expe- TRICHOLIMN2rCHUS Hernando and Ribera, dition to Central Australia. Part 11. Zoology. 2001:153 Dalau and Company. London. maior Hernando and Ribera, 2001:159 Blackwelder, R.E. 1944. Checklist of the Co- Malaysia leopterous Insects of Mexico, Central America, minor Hernando and Ribera, 2001:160 the West Indies, and South America. United Malaysia States National Museum Bulletin 185, Part 2: sabahensis Hernando and Ribera, 2001:160 189-341. Malaysia Brbthes, J. 1922. Descripciitn de Varios Coleopteros de Buenos Aires Annales de la Sociedad de Checklist of the Lutrochidae of the World Ciencias de Argentina 94: 263-305. Britton, E.B. 1971. A new intertidal beetle (Co- L UTROCHUS Erichson, 1847:509 leoptera: Limnichidae) from the Great Barrier BYRROMORPHUS Sharp, 1882:125 Reef. Journal of Entomology (B)40(2): 89-91. acuminatus Grouvelle, 1889:163 Britton, E.B. 1973. Hyphalus wisei sp. n., a new Venezuela intertidal beetle from New Zealand (Coleoptera: arizonicus Brown and Murvosh, 1970:103 1 Limnichidae). Records of the Auckland Insti- USA (AZ) tute and Museum 10: 119-122, 6 figures. geniculatus Chevrolat, 1864:406 Britton, E.B. 1977. Three new intertidal species of Cuba, Haiti Limnichidae (Coleoptera) from New Zealand. germari Grouvelle, 1889:163 Records of the Auckland Institute and Museum Brazil 14: 81-85, 9 figures. gigas Hinton, 1939:23 Broun, T. 1880. Manual of the New Zealand Co- Per& leoptera. Part 1. pp. 1-651. Government Print- laticeps Casey, 1893:580 er. Wellington. USA (AL, AR, DC, IL, IN, MD, MI, MO, OH, Broun, T. 1881. Manual of the New Zealand Co- MS, OK, PA, TN, WI, WV) leoptera. Part 2. pp. 653-744. James Hughes. luteus LeConte, 1852:42 Wellington. USA (NM, OK, TX) Broun, T. 1893. Manual of the New Zealand Co- misellus Grouvelle, 1896:17 leoptera. Part 7. pp. 1395-1504. Government Brazil Printing Office. Wellington. montanus Grouvelle, 1896:16 Brown, H. P. 1965. Microhabitats of some Neotro- Bolivia pica1 riffle beetles (Coleoptera: Dryopoidea). pilula Erichson, 1847:510 American Zoologist 5(2): 151. Brazil Brown, H.P., and C. M. Murvosh. 1970. Lutro- vestitus Sharp, 1882:126 chus arizonicus New Species, with notes on Guatemala ecology and behavior (Coleoptera, Dryopoidea, Limnichidae). Annals of the Entomological So- Acknowledgments ciety of America 63(4): 1030-1035. Casey, T.L. 1886. Descriptive notices of North D. P. Wooldridge, Pennsylvania State Univer- American Coleoptera I. Bulletin of the Califor- sity, Ogontz, and W. E. Steiner, Smithsonian Insti- nia Academy of Sciences 2(6): 157-264. tution, reviewed earlier drafts of this manuscript. Casey, T.L. 1889. 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Pic, M. 1905. Esphces et variktks nouvelles de Pic, M. 1956. Nouveaux Colkopthres de diverses ColkoptBres palkarctiques. L'Echange 21: 97- familles. Annales Historico-Naturales Musei 100. Nationalis Hungarici 7: 71-92. Pic, M. 1922. Nouveautks Diverses. Mklanges Ex- Putz, A. 1991a. Jaechobyrrhi~zus- eine neue Gat- otico-Entomologiques 36: 1-32. tung der Familie Limnichidae aus der Turkei Pic, M. 1923a. Nouveautks Diverses. Mklanges (Col., Limnichidae). Entomologische Nachrich- Exotico-Entomologiques 39: 1-32. ten und Berichte 35(2): 132-134. Pic, M. 192313. Nouveautks Diverses. Mklanges Putz, A. 1991b. Beitrage zur Insektenfauna der Exotico-Entomologiques 40: 1-32. DDR: Coleoptera- Limnichidae. Beitrage Ent. Pic, M. 1923c. Colkopthres exotiques en partie Berlin 41: 375-381. nouveaux. L'Echange 39: 15-16. Piitz, A. 1998a. I. Check list and bibliography of Pic, M. 1927a. Fauna Sumatrensis. Malacoderma- the Limnichide of China and neighbouring coun- ta (Col.). Supplementa Entomologica, Berlin tries (Coleoptera). pp. 337-339 in M. A. Jach 15: 62-65. and L. Ji (eds.). Water beetles of China. Vol. 2. Pic, M. 192713. Fauna Sumatrensis. Malacoderma- Wien: Zoologisch-Botanische Gesellschaft in ta (Col.). Supplementa Entomologica, Berlin ~sterreichand Wiener Coleopterologenverein. 16: 45-49. Putz, A. 1998b. 11. Taxonomic revision of the genus Pic, M. 1928. Nouveautks Diverses. Mklanges Ex- Cephalobyrrhus Pic (Coleoptera). pp. 341-371 otico-Entomologiques 52: 1-32. in M. A. Jach and L. Ji (eds.). Water beetles of Pic, M. 1930. Nouveaux ColkoptBres de diverses China. Vol. 2. Wien: Zoologisch-Botanische familles. Revista de la Sociedad Entomol6gica Gesellschaft in ~sterreichand Wiener Co- de Argentina 3: 43-46. leopterologenverein. Pic, M. 1935a. Nouveautks Diverses. Mklanges Rey, C. 1889. Remarques en passant. L'Echange 5: Exotico-Entomologiques 66: 1-36. 66-67. Pic, M. 193513. Nouveaux ColiloptBres exotiques. Reitter, E. 1877[1878]. in 0. Schneider and H. Annals and Magazine of Natural History (10)6: Leder. Beitrage zur Kenntniss der kaukasis- 470-473. chen Kaferfauna. Verhandlungen des Natur- Pic, M. 1938. Nouveaux ColiloptBres de la Rilpub- forschenden Vereines in Briinn 16: 1-258. lique Argentine. Revista de la Sociedad En- Reitter, E. 1881. Bestimmungs-Tabellen der eu- tom6logica Argentina 10: 77-79. ropaischen Coleopteren. IV. Enthaltend die Pic, M. 1939. Nouveaux ColiloptBres de la Rkpub- Familien: Cistelidae, Georyssidae und Thoric- lique Argentine. Revista de la Sociedad En- tidae. Verhandlungen der kaiserlich-konigli- tom6logica Argentina 10: 249-251. chen zoologisch-botanischen Gesellschaft in Pic, M. 1952. ColkoptBres nouveaux du Congo Wien 31: 67-96. Belge. Annales du Muske Congo belge, Ter- Reitter, E. 19Ol.Weitere Beitrage zue Coleopter- vuren, in-8O, Sciences Zoologiques 12: 9 en-Fauna des russichen Reiches. Deutsche Pic, M. 1953. ColkoptBres nouveaux de YAfrique Entomologische Zeitschrift 1901: 65-84. orientale. Atti Museo Civico di Storia Naturale Ribera, I. and C. Hernando. 1999. Taxonomic di Trieste 19(2): 110-114. revision of Platypelochares Champion (Co- Pic, M. 1954. Ptinidae, Anobiidae, Dermestidae, leoptera: Limnichidae). Koleopterologisch Byrrhidae (Col.). Beitrag zur Fauna Perus, Rundschau 69: 103-110. Jena 4: 176-178. Sat6, M. 1965. The limnichid-beetles of Formosa. Pic, M. 1955. Contributions ~3l'ktude de la fauna Special Bulletin of the Lepidopterists Society of entomologique du Ruanda-Urundi (Mission P. Japan 1965: 121-125. Basilewsky 1953). XI1 Coleoptera Rhipi- Sat6, M. 1966. The Limnichid-Beetles of Japan. phoridae, Oedemeridae, Scaptiidae, Pedilidae, Transactions of the Shikoku Entomological Aderidae, Anthicidae, Dermestidae, Byrrhidae, Society 9(2): 55-62. Anobiidae, Ptinidae, Cleridae, Lymexylonidae Sat& M. 1976. Family Limnichidae. Check-list of et Helodidae. Annales du Muske Congo Belge, the Coleoptera of Japan. 7: 1-2. Tervuren, in-8O, Sciences Zoologiques 63: 121- Sat6, M. 1994. Note on the genus Pseudeucinetus 135. [sic] Heller and its new relative (Coleoptera, Limnichidae). Special Bulletin of the Essa En- tomological Society, Niigata 2: 173-177. 164 Volume 15, No. 3, September, 2001, INSECTA MUNDI
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