FISHERY SURVEY OF THE PASCAGOULA RIVER SYSTEM I I i
COMPLETION REPORT < FISHERY SURVEY OP THE PASCAGOULA RIVER SYSTEM i
PREPARED BY:
• IP Stephen T. Ross, S. David Wilkins and John S. Peyton
HH Department of Biological Sciences Ip University of Southern Mississippi Hattiesburg, MS 39406-5018
TO • iP Mississippi Department of Wildlife Conservation •P.O.Southport BoxCenter,451 Jackson,Ellis at MSHwy. 80 39205-0451 ^ Attn.: Mr. Harry Barkley m 17 May, 1989 i i < i STATE: Mississippi PROJECT NO. F-89
PROJECT TITLE: Fishery Survey of the Pascagoula River System
STUDY TITLE: A survey of recreationally and commercially important
fishes of the Pascagoula, Chickasawhay and Leaf Rivers
PROJECT PERIOD: 1 March, 1988 - 28 February, 1989
ABSTRACT
We surveyed fishermen and sampled fishes on 165 km of the Pascagoula River drainage (Pascagoula, Leaf and Chickasawhay rivers) during April - September, 1988. The drainage supports a
recreational fishery for catfish (Ictalurus and Pvlodictus) and sunfish (Lepomis spp.) , although the number of recreational anglers was likely reduced during the study period by abnormally low water flow. Fishes nximerically dominant in the catch by hook "hnd line fishermen included bluegill and longear sunfish and channel catfish. These same species were dominant by weight in the catch. Trotline fishermen caught primarily channel and blue catfish.
The study area supports a diverse native freshwater fish fauna of at least 83 species. More species would be included if sampling were increased, especially on the tributary streams and backwater areas. • 2
INTRODUCTION • In contrast to many states in which the major recreational HPI fishing effort occurs primarily in natural lakes or reservoirs, Mississippi's recreational fishermen also spend a considerable 9 portion of their time fishing rivers and streams. This is especially true for anglers in the southeim portion of the state • where there is an abundance of natural flowing water (Miranda and HPI Frese 1989). Little information on angler use patterns exists, however, for one of the largest river systems in the state, the [|P Pascagoula River system. The Pascagoula River system includes the Leaf, Chickasawhay, and Escatawpa Rivers, and flows from north to • south in the southeast portion of the state. Hp Mississippi has one of the most diverse freshwater fish faunas in the United States, possessing approximately 200 species of iP ^ native freshwater fishes; at least 124 species occur, in the ^ Pascagoula River system (Ross and Brenneman,. in prep.). While many of these species are small minnows and darters, there are also many which grow to large sizes and which are included in the recreational or commercial fish catches. Distributional data for the larger species, such as redhorses, catfishes, buffalos, and basses are especially limited, as these species are rarely retained
for museum fish collections. OBJECTIVES
Study Objectives: The primary study objectives were to determine angler usage and fish distributions for the lower portions of the Leaf, Chickasawhay and Pascagoula rivers.
Job Objectives: Job_J,. Determine angler usage and catch data for the Leaf,
Chickasawhay and Pascagoula rivers. Job__2. Determine the distribution and relative abundance of fishes in the Leaf, Chickasawhay and Pascagoula rivers, with emphasis on larger species of recreational or
commercial importance. Job_3.. Identify and enumerate all fishes collected in Job 2. Catalog all collected fish specimens for ^einnanent storage in the University of Southern Mississippi Museum
of Ichthyology. Job—4. Determine length—weight regressions for species counted and measured in Job 1, using data from specimens
collected in Job 2.
METHODS AND MATERI2VLS
•lob 1. Angler survey. We followed preliminary sampling guidelines prepared by the Mississippi Department of Wildlife Conservation. The guidelines f i
4 f called for a sampling regime of 72 field days for a two-man field ¥ team, or 144 man days. Actual dates of sampling (Table 1) varied slightly (69 actual field days) from this plan, due primarily to
equipment and scheduling problems.
Each river section within the study area (Figure 1) was
divided into an upstream and a downstream reach, with stream
reaches sampled equally each month. For samples in March, July and
August, sampling was only on weekends. Samples in April, May and
June were stratified between weekends and weekdays. All sample
dates were chosen randomly within each month and stratum. The
sections of the three rivers which were sampled and river distances
were:
IA. Upper Pascagoula River: Merrill downstream to Wilkinsons Ferry (George County); 30.1 km (18.8 mi).
IB. Lower Pascagoula River: Wilkinsons Ferry downstream to the Wade Bridge (Jackson County) ; 40.3 km (25^!2 mi) .
2A. Upper Leaf River: Beaumont downstream to McLain (Perry and Greene Counties); 25.6 km (16.0 mi).
2B. Lower Leaf River: McLain downstream to Merrill (Green and George Counties); 25.6 km (16 mi).
3A. Upper Chickasawhay River: Leakesville downstream to entrance of Crawford Creek (Green County); 20.5 km (12.8 mi) .
3B. Lower Chickasawhay River: Entrance of Crawford Creek downstream to Merrill (Greene and George Counties); 23.0 km (14.6 mi). K P
Table 1 Actual Collection Dates for the Pascagoula Fishery if! Survey- 1988 i Date Water Bodv Stream Reach f April 02 Chickasawhay River Lower 04 Pascagoula River Lower P 09 Pascagoula River Lower I I 10 Pascagoula River Upper 11 Leaf River Upper 12 Chickasawhay River Lower P 19 Pascagoula River Upper 23 Leaf River Upper 24 Chickasawhay River Upper P 26 Leaf River Lower 28 Chickasawhay River Upper
May 05 Leaf River Upper 06 Pascagoula River Upper 07 Leaf River Lower 08 Chickasawhay River Upper 09 Chickasawhay River Lower 10 Pascagoula River Upper P 11 Leaf River Upper 12 Pascagoula River Upper 13 Chickasawhay River Upper 16 Chickasawhay River Upper 17 Pascagoula River Lower 18 Chickasawhay River Lower 20 Pascagoula River Lower 21 Leaf River Upper 22 Pascagoula River Lower 23 Pascagoula River Lower 24 Leaf River Lower 25 Chickasawhay River Upper 26 Leaf River Upper p 27 Leaf River Lower 28 Leaf River Lower 29 Leaf River Upper P 30 Pascagoula River Upper 31 Chickasawhay River Lower
June 02 Leaf River Upper I 04 Pascagoula River Upper 05 Chickasawhay River Lower k 06 Pascagoula River Lower r 08 Leaf River Lower ii 09 Pascagoula River Upper i p 10 Leaf River Upper 13 Chickasawhay River Upper 14 Chickasawhay River Lower 18 Leaf River Upper 19 Chickasawhay River Upper 21 Chickasawhay River Lower 22 Pascagoula River Upper 23 Pascagoula River Lower 24 Leaf River Lower 25 Pascagoula River Lower 29 Chickasawhay River Lower 30 Pascagoula River Upper
July 02 Chickasawhay River Lower 16 Pascagoula River Upper 17 Chickasawhay River Upper 23 Leaf River Upper 30 Pascagoula River Lower 31 Leaf River Lower
August 06 Chickasawhay River Upper 07 Pascagoula River Lower 13 Leaf River 14 Chickasawhay River Lower 21 Pascagoula River Upper 27 Leaf River Upper
September 03 Pascagoula River Lower 04 Leaf River Upper 17 Chickasawhay River Lower 18 Chickasawhay River Upper
I 7
Data were recorded from anglers using a roving survey method as described in Malvestuto (1983) . An example data sheet is shown in Figure 2. Surveys were done using an outboard boat and all
Scimpling trips originated and ended at the same boat landing. The decision on whether to survey anglers on the outgoing or return leg of the sampling trip was decided by a coin toss.
Job 2. Distribution and relative abundance of fishes
The field work for Job 2 was done in conjunction with Job 1.
Because of the uncertain time demands of Job 1, caused by varying numbers of fishermen to interview, the sampling design for Job 2 was qualitative rather than quantitative. Fishes were sampled by hoop nets (4', 1" mesh with two throats) / gill nets (30' x 5' with
3" mesh) and seines (20' bag, 1/8" mesh; 10* straight, 1/8"mesh) . Hoop nets and gill nets were set near the beginning ^of each sampling day and then checked near the end of the day, or the following day if the sampling schedule permitted. Sampling sites were varied during the study so that habitats representative of both main channel, baclcwaters and tributary streams were represen ted. We classified as a backwater any body of water not directly connected to the main channel. Tributaries were generally sampled at their junction with the main cheinnel or no further than 1 km upstream from the main channel. No more than two collections were generally made at the same site and we attempted to intensively seine at least two locations per sample day.
All fishes that were retained for accession into the mH Figure 1. Amap of the Pascagoula River drainage in southeastern Mississippi showing the sampling areas. LEAKESVILLE
BEAUMONT LEAF RIVER
HWY98 N McLAIN CHICKASAWHAY RIVER
MERRILL HWY57 AREA HWY98 SAMPLED PASCAGOULA RIVER LUCEDALE
BENNDALE HWY26
BLACK CREEK
1 3 9 12 HWY57 KILOMETERS WADE 10
£s f
Figure 2. An example of the form used during the fishery survey
of the Pascagoula River drainage.
I ii I s hi ii
i k Southern Streams Creel Survey
Sample Date ; Sample Number
County Township ; Range : Section
Stream Location
Stream Level Water Temperature Air Temp..
Weather Conditions
Main Channel
No. of Anglers Sex: M: F, Slate and county that fishing triporiginated from
Target species Rshing from: bsffik : boat.
Rshing Gear:
# rods : time fished
hoop nets: size number time fished
trot/set lines: # hooks iff lines time fished
Species caught: Length (SL)
Number of times fished annually 13
Because of the effect of sampling effort on species composi
tion, comparisons of fish diversity between•rivers must address the
problem of unequal sample sizes and sampling efficiency. We used
the rarefaction procedure (Hurlbert 1971; Ludwig and Reynolds
(1988) to determine the expected number of species for a given
sampling effort for each of the three rivers. The expected number
of species in a random sample (size n) from a community is the sum
of the probabilities that each species will be included in the
scimple (James and Rathbun 1981) . Rarefaction was calculated using
a BASIC program in Ludwig and Reynolds (1988).
Job 4. Length-Weight Regressions
Based on the larger fishes collected in Job 2, we determined
regressions for length-weight data. We then used these regressions
to determine weight data from the length measurements of^Job 1.
Fishes were weighed using either a Salter 5 kg spring balance, or
appropriately sized Pesola spring balances (20 kg, 2.5 kg, 1 kg,
0.5 kg).
RESULTS AND DISCUSSION
Job 4. Determination of Lenath/Weiaht Relationships
Length and weight data were obtained for 211 specimens representing 14 of the taxa taken by anglers (Table 2) . In addition, we recorded weights and lengths for five species
(Ancmilla rostrata. Dorosoma ceoedianum. Ictiobus bubalus. Muail 14 cephalus and Lepisosteus oculatus) which were not taken in the recreational or commercial fishery (Appendix I).
Initially, following recommendations of Ricker (1975) and Laws and Archie (1981) , we used functional (Model II) regressions to estimate weights of fishes caught by anglers. Model II regressions are appropriate when both the dependent and independent variables have an error component. However, we also calculated weights of the fishes based on a Model I regression and found that this model provided a more accurate representation of fish weights (Table 3) .
Using the field weighed fishes as a control, the Model II regres sion consistently overestimated weights, having an average percent deviation from actual weights of +17.2%. In contrast, the Model
I regression slightly underestimated actual weights with an average percent deviate of -6.1%.
We were unable to capture two species, freshwater drum
(Aplodinotus grunniensl and spotted sunfish (Lepomis punctatus), that were also very uncommon in angler catches, and we only captured one specimen of striped bass (Mprone saxatilis). Because of this we did not determine weight-length relationships for the freshwater drum and striped bass; we used material from all species of sunfish to estimate weights of L. punctatus. In addition, because of very limited weight-length data for several other species (Ictalurus furcatus. Lepomis oulosus. Micropterus punctulatus and M. salmoides) we used regressions based on all material for each genus. 15
Job 1. Angler Survey The fishermen suarvey forms were returned to the Mississippi Department of Wildlife Conservation in January of 1989 for analysis. Consequently, we will present only limited data on fisheimien. Overall, we interviewed 383 anglers in 173 groups. The majority of persons interviewed were using fishing rods (373 persons, 145 groups), while only 36 groups (84 persons) were using trotlines, with 10 of the latter group also using fishing rods. Only two interviews were with hoopnet fishermen. Extremely low river flow during late May and in June likely reduced the number of anglers. During this period navigation by
outboard powered boats was difficult. The primary target species (Figure 3) listed by the 145 groups of rod fishermen were sunfish (40%) and catfish (39%). Bass were listed as a target species by only 8% of the fishing parties. All of the trotline and hoopnet fishermen were fishing for catfish. The catch overall was dominated numerically by channel catfish (31%), bluegill (28%) and longear sunfish (23%) (Table 4) . The catch by rod fishermen was similar, being dominated by bluegill (35%), longear sunfish (29%) and catfish (23%). Hoopnet fishermen caught primarily carpsucker (Carmiodes spp.) , and flathead and blue catfish, while trotline fishermen caught primarily channel and blue CQ-'tfish. Compared to the suggested target species for rod anglers, sunfishes were over-represented while catfish were under-represent ed. 1 f f 16 f f f
Table 2. Weight - length regressions (Model I) for fishes from the Pasca- goula River drainage. Regressions are for natural log transformed data; lengths are measured in standard length (SL); weights are measured in grams.
Soecies Rearession rz N
Caroiodes spp. Log (Wt) = -07.50 + 2.45 Log (Length 0.985 18 Ictalurus ounctatus Log (Wt) = -10.39 + 2.87 Log (Length 0.967 20 Ictalurus spp. Log (Wt) = -11.12 + 3.00 Log (Length 0.976 24 Pvlodictus olivaris Log (Wt) = -11.27 + 3.04 Log (Length 0.984 30 AmbloDlites ariommus Log (Wt) = -09.89 + 2.93 Log (Length 0.951 16 Leoomis macrochirus Log (Wt) = -10.69 + 3.09 Log (Length ^^0.959 32 Leoomis meaalotis Log (Wt) = -10.64 + 3.08 Log (Length 0.978 46 Leoomis microloohus Log (Wt) = -10.58 + 3.06 Log (Length 0.993 13 Leoomis spp. Log (Wt) = -10.66 + 3.08 Log (Length 0.971 79 Pomoxis annularis Log (Wt) = -11.62 + 3.17 Log (Length 0.984 10 Microoterus sop. Log (Wt) = -11.75 + 3.18 Log (Length 0.981 7
• fe • • i 17
Table 3. A comparison of measured and calculated average weights for fishes collected from the Pascagoula River drainage, 1988. Calculated weights are based on a Model I regression.
TAXON Measured Weight Calculated Weight Number X Oiffei Caroiodes 682.2 440.5 18 -35.42 Ictalurus furcatus 982.5 822.6 4 -16.27 [. Dunctatus 401.5 408.1 20 1.66 Pvlodictus olivaris 1095.2 1075.0 30 -1.84 Ambloolites ariommus 110.2 109.9 16 -0.29 Leooriis qulosus 90.0 85.5 1 -5.03 Leoomis macrochirus 36.7 36.3 32 -1.18 Leoocnis meqalotis 32.9 32.4 46 -1.40 Leoomis mfcroloohus 112.6 112.7 13 0.03 Hicrooterus ounctulatus 213.3 241.5 3 13.21 Hicrooterus salmoides 857.5 826.7 4 -3.59 Pomoxis annularis 99.4 98.7 10 -0.71 P. niqromaculatus 165.0 117.3 1 -28.90
Average 0ifference: -6.13 18 Based on weight data (Table 5) , the catch overall was dominated by channel (36.4%), flathead (18.6%) and blue catfish (18.3%). Major species caught by rod fisheraen were channel catfish (41%) and bluegill (21%), while channel (75%) and blue catfish (15%) were gravimetric dominants in the catch by trotline fishermen. The most important species, by weight, for hoopnet fishermen were flathead catfish (48%), blue catfish (38%) and carpsuckers (14%).
Jobs 2 & 3. Relative Abundance of Fishes in the Chickasawhav. Leaf
and Pascaaoula rivers.
Overall we captured 26,627 fishes, of which 24,802 were iden tified to species (Table 6). The remaining 1,826 specimens (primarily post-lairval and early juvenile forms) were identified to the generic or family level. The fishes represented 83 species in 41 genera and 18 families. The samples were dominated numerically by the blacktail shiner (Notroois yenustus) which comprised 66.7% of the catch. Other abundant species included Notroois lonairostris. longnose shiner (4.4%), Hvboanathus nuchalis. Mississippi silvery minnow (3.2%), Gambusia affinis, mosquitofish (3.2%), Leoomis macrochirus. bluegill sunfish (2.9%), Leoomis meaalotis. longear sunfish (1.9%), Ammocrypta beani, naked sand darter (1.7%) , Microotenis sal'moides. largemouth bass (1.2%), Notroois texanus. weed shiner (1.1%) and Hvboosis stpreriana, gravel chub (1.1%). f f 19
Figure 3. The primary fish species sought after by rod fishermen in the Pascagoula River drainage. Data are based on 325 persons
in 145 fishing parties. CO s CRAPPIE o BASS CO E-« W AOTTfflNG C!J CATFISH
SUNFISH
0 5 10 15 20 25 30 35 40 45 50 PERCENT OF ANGLER GROUPS 21
Table 4. Hunbers of Fishes Caught by Fishermen During the Pascagoula Fishery Survey, 1988.
All Fishermen Pod Fishermen HoooNets Trotlines*
H X M % M % M % Catostomidae Caroiodes sp. 12 1.5 0 0.0 12 44.4 0 0.0 Ictaluridae Ictalurus furcatus 46 5.8 24 3.8 5 18.5 17 11.8 Ictalurus ounctatus 243 30.5 143 22.8 0 0.0 100 69.4 Pvlodictis olivaris 20 2.5 5 0.8 9 33.3 6 4.2 Centrarchidae Ambloolites arioninus 4 0.5 4 0.6 0 0.0 0 0.0 Leoomis qulosus 3 0.4 3 0.5 0 0.0 0 0.0 Leoomis macrochirus 219 27.5 217 34.7 0 0.0 2 1.4 Leoomis meoalotis 186 23.4 182 29.1 0 0.0 4 2.8 Leoomis microloohus IS 1.9 6 1.0 0 0.0 9 6.3 Leoomis ounctatus 2 0.3 2 0.3 0 0.0 0 0.0 Hicrooterus ounctulatus 17 2.1 13 2.1 0 0.0 4 2.8 Microoterus salmoides 13 1-6 13 2.1 0 0.0 0 0.0* Pomoxis annularis 10 1.2 8 1.3 0 0.0 2 1.4 Pomoxis nioromaculatus 2 0.3 2 0.3 0 0.0 0 0.0 Sciaenidae Aolodinotus orunniens 2 0.3 2 0.3 0 0.0 0 0.0 Percichthyidae Horone saxatilis 2 0.3 1 0.2 1 3.7 0 0.0
Totals 796 625 27 144
both rods and trotlines used by fishing groups 22
Table 5. Weights of Fishes Caught by Fishermen During the Pascagoula Fishery Survey, 1988.
All Fishermen Rod Fishermen HoooNets Trotlines* Total Ueiqht % Total Ueiqht X Total Ueiqht X Total Ueiqht % Catostomidae Carpiodes so. 8677.T 4.7 0 0.0 8677.1 14.2 0.00 0.0 Ictaluridae Ictalurus furcatus 33649.5 18.3 3004.1 4.2 23182.7 37.8 7462.7 14.9 Ictalurus ounctatus 66922.6 36.4 29461.2 40.7 0.0 0.0 37461.4 74.7 Pvlodictis olivaris 34256.3 18.6 1875.9 2.6 29433.0 48.0 2947.8 5.9 Centrarchidae Ambloolites ariommus 343.8 0.2 343.8 0.5 0.0 0.0 0.0 0.0 Lepomis qulosus 379.8 0.2 379.8 0.5 0.0 0.0 0.0 0.0 Leoomis macrochirus 15502.0 8.4 15422.1 21.3 0.0 0.0 80.0 0.2 Leoocnis meoalotis 93124.4 5.1 9075.3 12.5 0.0 0.0 239.1 0.5 Leoomis microloohus 1786.1 1.0 785.5 1.1 0.0 0.0 1000.6 2.0 Leoomis ounctatus 111.6 0.1 111.6 0.2 0.0 0.0 0.0 0.0 Microoterus ounctulatus 3165.1 1.7 2585.6 3.6 0.0 0.0 579.6 1.2 Microoterus salmoides 8581.6 4.7 8581.6 11.9 0.0 0.0 0.0 0.0 Pomoxis annularis 965.6 0.5 616.0 0.9 0.0 0.0 349.5 0.7 Pomoxis niqromaculatus 142.4 0.1 142.4 0.2 0.0 0.0 0.0 0.0
Total 183798.4 72385.0 61292.8 50120.61
* both rods and trotlines used by fishing groups
We collected 68 species from the Pascagoula River. The
samples were numerically dominated by blacktail shiners (59%) in all of the three habitat types (main channel, backwaters and tributaries (Tcible 7) . Other commonly occurring species in the main channel were Hvbocmathus nuchalis (8.4%), Notroois lonai- rostris (4.8%) and Trinectes maculatus. hogchoker (3.2%).
Centrarchids, especially Pomoxis niaromaculatus. black crappie
(10.3%), largemouth bass (8.4%) and bluegill sunfish (6.4%) were abundant in the backwater areas. Mosquitofish (8.7%) were also 23
Table 6. Fishes collected from the Pascagoula River drainage, including the Leaf, Chickasawhay and Pascagoula rivers, 1988.
Species Number of Fish Percent
Petromyzontidae Ichthvomvzon aaaei 004
Lepisosteidae Leoisosteus oculatus .012 Leoisosteus osseus .004
Anguillidae Anouilla rostrata .016
Clupeidae Alosa chrvsochloris 3 .012 Dorosoma ceoedianum 1 .004 Dorosoma petenense 49 .198
Hiodontidae Hiodon teraisus .004
Cyprinidae Ericvmba buccata 8 .032 Hvbocmathus havi 204 .823 Hvboonathus nuchalis 806 3.250 Hvbopsis aestivalis 5 .02.0 Hvbopsis storeriana 262 1.056 Hvbopsis winchelli 109 0.439 Nocomis leptocephalus 2 .008 Notemiaonus crvsoleucas 12 .048 Notropis atherinoides 180 .726 Notropis chrvsocephalus 38 .153 Notropis lonairostris 1084 4.371 Notropis maculatus 101 .407 Notropis roseipinnis 12 .048 Notropis texanus 283 1.141 Notropis venustus 16555 66.749 Notropis volucellus 41 . 165 Qpsopoeodus emiliae 290 1.169 Pimephales viailax 201 .810
Catostomidae Carpiodes cvprinus 6 .024 Carpiodes velifer 36 .145 Erimvzon tenuis 2 .008 Hvpentelium niaricans 4 .016 Ictiobus bubalus 4 .016 Minvtrema melanops 1 . 004 24
Species Number of Pish Percent
Ictaluridae Ictalurus f-urcatus 4 016 Ictalurus natalis 2 008 Ictalurus punctatus 56 226 ' Noturus ovrinus 1 004 Noturus leptacanthus 28 113 Noturus nocturnus 17 069 Pvlodictis olivaris 32 129
Esocidae Esox americanus 9 .036 Esox nicer 51 .206
Aphredoderidae Aphredoderus savanus 74 .298
Atherinidae Labidesthes sicculus 119 .480
Fundulidae Fundulus blairae 22 .089 Fundulus chrvsotus 8 .032 Fundulus notatus 12 .048 Fundulus notti 16 .065 Fundulus olivaceus 91 .367
Poeciliidae Gambusia affinis 788 3.177
Centrarchidae Ambloplites ariommus 35 .141 Centrarcfaus macropterus 10 .040 Elassoma zonatum 1 .004 Enneacanthus aloriosus 6 .024 Lepoittis cvanellus 10 .040 Lepomis oulosus 17 .069 Lepomis macrochirus 722 2.911 Lepomis marainatus 19 .077 Lepomis meaalotis 465 1.875 Lepomis microlophus 20 .081 Lepomis punctatus 6 .024 Micropterus punctulatus 71 .286 Micropterus salmoides 303 1.222 Pomoxis annularis 36 .145 Pomoxis niaromaculatus 234 .943
Mugilidae Muail cephalus 004 r
I 25 Species Ntmber of Fish Percent t Percichthyidae Morone saxatilis . 004 t Percidae Anunocrvota beani 412 1.661 Ammocrvota vivax 1 .004 Etheostoma chlorosomuni 5 .020 t Etheostoma fusiforme 1 .004 Etheostoma histrio 102 .411 Etheostoma lynceum 4 .016 I Etheostoma oroeliare 24 .097 Etheostoma stiomaeum 8 .032 Etheostoma swaini 53 .214 I Percina lenticula 24 .097 Percina niarofasciata 92 .371 Percina solera 52 .210 Percina shumardi 12 .048 I Percina sp. 1 (Gulf Logperch) 44 .177 Percina sp. 3 (Pearl River Channel Darter)14 .056 I Percina viqil 5 .020 Soleidae Trinectss maculatus 353 1.423 I *** Total *** 24,802 I I I I I I I I f 26
Table 7. Fishes collected from the Pascagoula River system, 1988.
Main Channel Backwaters Tributaries Total Species ILi. % % % N. % [ , Lepisosteidae ||| Lepisosteus osseus 1 1 .01 Anguillidae Ancmilla rostrata f 1 1 .27 2 .03 Clupeidae Alosa chrvsochloris 1 .02 1 .01 Dorosoma cepedianum 1 .06 1 .01 a Dorosoma petenense 28 • .51 1 .06 29 .38 •o• oto Cyprinidae toto Hvboanathus havi 24 .44 1 .27 25 .33 Hvboonathus nuchalis 460 8.36 25 6.83 485 6.31 Hvbopsis aestivalis 2 .04 2 .03 Hvbopsis storeriana 83 1.51 4 .22 2 .55 89 1.16 Hvbopsis winchelli 50 .91 2 .11 3 .82 55 .72 Nocomis leptocephalus 1 .02 1 .27 2 .03 Notropis atherinoides 105 1.91 26 7.10 131 1.71 Notropis chrvsocephalus 5 1.37 5 .07 Notropis lonairostris 265 4.81 3 .17 6 1.64 274 3.57 Notropis maculatus 3 .05 13 .72 16 .21 Notropis roseipinnis 3 .82 3 .04 Notropis texanus 67 1.22 30 1.66 44 12.02 141 1.84 Notropis venustus 3601 65.43 801 44.23 158 43.17 4560 59.37 Notropis volucellus 19 .35 19 .25 Opsopoeodus emiliae 39 .71 55 3.04 1 .27 95 1.24 Pimephales viailax 31 .56 26 1.44 57 .74
Catostomidae Carpiodes cvprinus 1 .02 1 .01 Carpiodes velifer 4 .07 4 .05 Ictiobus bubalus 2 .04 2 .03
Ictaluridae Ictalurus furcatus 1 1 .01 Ictalurus natalis 1 .27 1 .01 Ictalurus punctatus 3 .05 1 .27 4 .05 Noturus leptacanthus 1 .02 1 .27 2 .03 Noturus nocturnus 6 1.64 6 .08 Pvlodictis olivaris 8 .15 8 .10
Esocidae Esox americanus 1 .02 3 .17 4 .05 Esox nicrer 4 .07 17 .94 21 .27
I r
I 27
Main Channel Backwaters Tributaries Total r Species N. % N. % N. % N. Aphredoderidae r Aphredoderus savanus 7 .13 10 .55 1 .27 18 .23
• Atherinidae I Labidesthes sicculus 7 .13 51 2.82 58 .76 Fundulidae Fundulus blairae 2 .11 2 .03 I Fundulus chrvsotus 2 .04 3 .17 5 .07 Fundulus notatus 5 ,28 5 .07 Fundulus notti 1 .06 1 .01 I Fundulus olivaceus 4 .07 37 2.04 41 .53 Poeciliidae I Gambusia affinis 1 .02 157 8.67 158 2.06 Centrarchidae Ambloplites ariommus 5 .09 1 .27 6 .08 I Centrarchus macropterus 8 .44 8 .10 Enneacanthus aloriosus 1 .02 5 .28 6 .08 Lepomis crulosus 4 .22 4 . 05 Lepomis macrochirus 93 1.69 116 6.41 5 1.37 214 2.79 Lepomis marainatus 1 .02 5 .28 6 .08 Lepomis meaalotis 102 1.85 67 3.70 6 1.64 175 2.28 Lepomis microlophus 11 .20 1 .06 12 .16 I Lepomis punctatus 1 .02 1 .01 Micropterus punctulatus 19 .35 19 .25 Micropterus salmoides 41 .74 152 8.39 11 3.01 204 2.66 Pomoxis annularis 6 .11 10 .55 16 .21 Pomoxis niaromaculatus 12 .22 186 10.27 198 2.58
Mugilidae Muail cephalus 1 .02 1 .01
Percidae Ammocrvpta beani 158 2,87 10 .55 1 .27 169 2.20 Etheostoma fusiforme 1 .06 1 .01 Etheostoma histrio 22 .40 35 9.56 57 .74 Etheostoma Ivnceum 1 .02 1 .01 Etheostoma proeliare 10 .55 10 .13 Etheostoma swaini 5 .09 6 1.64 11 .14 Percina lenticula 2 .04 2 .04 Percina niarofasciata 1 .02 1 .27 2 .03 Percina sciera 10 .18 1 .27 11 .14 • Percina shumardi 2 .04 2 .03 28
Main Channel Backwaters Tributaries Total Species Ni N. N. IL_ %
Percina sp. l 3 .05 13 .72 16 .21 Percina sp. 3 5 .09 5 .07 ' Percina viail .27 1 .01
Soleidae Trinectes maculatus 175 3.18 1 .06 12 3.28 188 2.45
*** Totals *** 5,504 1,811 366 7,681
I I I • I I I I 29
coimon in the bac3cwater habitats. In addition to blacktail shiners, the most abundant species in the tributaries were other minnows (especially Notropis texanus (12%), Notropis atherinoides. emerald shiner (7.1%) and the Mississippi silvery minnow (6.8%)), and the harlequin darter, Etheostoma histrio (9.6%). The species-sample cuive (Fig. 4) for the Pascagoula River system indicated that there were sufficient samples to describe the fish fauna of the main channel. However, the data for baclcwater and tributary streams are more limited, with the curves for both habitats not reaching a horizontal asymptote. The lognormal distribution (Fig. 5) indicates that the system was well sampled overall, but also that there are at least 14 additional species that would be obtained by further sampling with similar gear types. The potential presence of additional species can be estimated by ®x't^3^polating the curve to the left of the ordinate and ''counting the number of species present in the rarer abundance groups. We collected 64 species from the Leaf River system (Table 8) .
In collections from the main channel the blacktail shiner was the numerical dominant (67.3%), followed by longnose shiners (5.8%), bluegill (3.4%), Mississippi silvery "minnows (3.3%), Hvbocmathus hayi, cypress minnow (2.2%) and longear sunfish (2.1%). Only one backwater habitat was present in the study section of the Leaf River so that the number of samples for this habitat type is limited. In the backwater area Notemiaonus crvsoleucas. golden shiner (17.5%), mosquitofish (17.5%), bluegill (15.9%) and blacktail shiners (12.7%) were among the most abundant species. 30
Figure 4. Fish species-sample cuirves for the Pascagoula, Leaf and Chickasawhay rivers, 1988.
ii I k
• CO ip/as©A(i®(y)iLA yj O LU CL CO UL o cr LU GO ALL HABITATS MAIN CHANNEL LU -I > BACKWATERS § SIDE STREAMS ID S 3 O
CO LU O LU CL CO LL O QC LU CD ALL HABITATS MAIN CHANNEL LU > • SIDE STREAMS § 3
3 O
©(l=30©KAiAWIHlAV CO LU O LU Q. -/ CO LL O oc LU CD
ril-u riMoi 1 rv 1 o
a MAIN CHANNEL LU > •— BACKWATERS I- i 3 0 SIDE STREAMS 3 1 2 1 1 3 7 O 20 30 40 50 60 CUMULATIVE NUMBER OF SAMPLES f 32
Figure 5. Fish species abundance patterns for the Pascagoula, Leaf and Chickasawhay rivers, 1988, See text for further explanation.
I NUMBER OF SPECIES NUMBER OF SPECIES NUMBER OF SPECIES
O CO o CD ro CXI o A 1—I ' • »
CO - m &
m [AiO
Oiiil CD -
ro
cn 34
Table 8. Fishes collected from the Leaf River system, 1988
Main Channel Backwaters Tributaries Tota 1 Species IL % IL. % IL. % IL. % . Lepisosteidae Lepisosteus oculatns 2 .02 1 .09 3 .03 Clupeidae Alosa chrvsochloris 1 .01 1 .01 Dorosoma petenense 19 .23 19 .20 Hiodontidae Hiodon teraisus 1 .01 1 .01 Cyprinidae Ericvmba buccata 6 .07 6 .06 Hvboanathus havi 177 2.16 177 1.88 Hvboanathus nuchalis 268 3.26 149 12.81 417 4.42 Hvbopsis aestivalis 3 .04 3 .03 Hvbopsis storeriana 140 1.71 38 3.27 178 1.89 Hvbopsis vinchelli 25 .30 3 .26 28 .30 Notemiaonus crvsoleucas! 12 .15 11 17.46 23 .24 Notropis atherinoidgs 33 .40 Notropis chrvsQcephalus 33 .35 ; 33 .40 33 2.84 66 Notropis lonairostris .70 475 5.79 163 14.02 638 Notropis maculatus 6.76 2 .02 2 Til 4 Notropis roseipinnis .04 1 .01 1 .01 Notropis texanus 92 1.12 48 4.13 140 1.48 Notropis Venustus 5525 67.30 8 12.70 420 36.11 5953 63 .09 Notropis volucellus 8 .10 1 .09 9 .10 Opsopoeodus emiliae 145 1.77 5 7.94 72 6.19 222 2.35 Pimephales viailax 57 .69 57 .60 Catostomidae Cairoiodes velifer 1 .01 1 .01 Hvpentelium niaricans 3 .04 2 .17 5 Ictiobus bubalus .05 1 .01 1 .01 Minvtrema melanops 1 .01 1 1.59 . 2 .02 • Ictaluridae Ictalurus furcatus 3 .04 3 .03 Ictalurus punctatus 11 .13 1 .09 12 .13 Noturus leptacanthug 15 .18 15 1.29 30 .32 Pvlodictis olivaris 20 .24 2 .17 22 .23 Esocidae Esox americanns 1 .01 1 .01 Esox niaer 6 E .07 6 .06 35
Main Channel Backwaters Tributaries Total Species N. N. IL. i JL. % Aphredoderidae Aphredoderus savanus .06 5 .05
Atherinidae Labidesthes sicculus 52 63 18 1,55 70 .74
Fundulidae Fundulus blairae 14 .17 5 7.94 19 .20 Fundulus notatus 6 .07 6 .06 Fundulus notti 2 .02 2 .02 Fundulus olivaceus 38 .46 11 .95 49 .52
Poeciliidae Gambusia affinis 162 1.97 11 17.46 7 .60 180 1.91
Centrarchidae Ambloplites arioininus 12 .15 4 .34 16 .17 Elassoitia zonatum 1 .01 1 .01 Lepomis cvanellus 6 .07 6 .52 12 .13 Lepomis oulosus 3 .04 1 1.59 2 .17 6 .06 Lepomis macrochirus 275 3.35 10 15.87 24 2.06 309 3.27 Lepomis meaalotis 169 2.06 23 1.98 192 2.03 Lepomis microlophus 4 .05 4 .04 Lepomis punctatus 3 .04 3 .03 Micropterus punctulatus 32 .39 6 .52 38 .40 Micropterus salmoides 28 .34 2 3.17 7 .60 37 .39 Pomoxis annularis 11 .13 2 3.17 1 .09 14 .15 Pomoxis niaromaculatus 15 .18 7 11.11 4 .34 26 .28
Percidae Ammocrvpta beani 73 .89 18 1.55 91 .96 Ammocrvpta vivax 1 .01 1 .01 Etheostoma chlorosomum 4 .05 3 .26 7 .07 Etheostoma histrio 17 .21 8 .69 25 .26 Etheostoma Ivnceiim 3 .04 1 .09 4 .04 Etheostoma oroeliare 12 .15 11 .95 23 .24 Etheostoma stiomaeum 8 .10 8 .69 16 .17 Etheostoma swaini 27 .33 9 .77 36 .38 Percina lenticula 1 .01 1 .01 Percina niarofasciata 35 .43 29 2.49 64 . 68 Percina sciera 18 .22 5 .43 23 .24 Percina shumardi 1 .01 1 .01 Percina sp. l 16 .19 2 .17 18 .19
Soleidae Trinectes maculatus 69 84 6 .52 75 .79
*** Totals *** 8;210 63 1,163 9,436 k 6- 36 The blacktail shiner was the numerical dominant in the tributary streams (36.1%), followed by longnose shiners (14%), Mississippi silvery minnows (12.8%), Opsopoeodus emiliae. pugnose minnow (6.2%), weed shiners (4.1%) and Hvbopsis storeriana. gravel chub
(3.3%).
The species-sample curve (Fig. 4) for the Leaf River indicates that the main channel was well sampled; however, more species could
be expected from tributaries. Data for backwater areas are not
plotted.
The lognormal distribution (Fig. 5) also indicates that the main channel was well sampled. The species-abundance pattern fits the lognormal curve closely and most of the curve is revealed (e.g. to the right of the y-axis) . Based on an extrapolation of the cuirve to the left of the y—axis, at least seven additional species
^ could be expected from the Leaf River for collections made'*with the
same gear types as the present study. We collected 64 species from the Chickasawhay River system. The blacktail shiner was the most abundant species from the main channel (81%) and tributaries (72%) of the Chickasawhay River
system (Table 9). Mosquitofish (39.2%) were numerical dominants in the backwater areas. Other numerically abundant species in the main channel were longnose shiners (4.1%), naked sand dairters
(2.5%) , longear sunfish (1.4%) , Pimephales viailax. bullhead minnow (1.3%) and hogchokers (1.2%) . In addition to mosquitofish, longear sunfish (15.4%), blacktail shiners (12.8%) and Notropis maculatus.
shiner (8.4%) were abundant in samples from backwater 37
Table 9. Fishes collected from the Chickasawhay River system, 1988.
Main Channel BacJcwaters Tributaries Total Soecies N. 1 IL. % IL. % iL. % Lepisosteidae Leoisosteus oculatus 1 .10 1 .01
Anguillidae Anouilla rostrata 2 .03 2 .02
Clupeidae Alosa chrvsochloris 1 .01 1 .01 Dorosoma oetenense 1 .01 1 .01
Cyprinidae Hvbocmathus havi 2 .03 2 .02 Hvboanathus nuchalis 17 .25 32 3.24 49 .56 Hyboosis storeriana 33 .48 33 .38 Hybopsis winchelli 28 .41 1 .12 29 .33 Notroois atherinoides 4 .06 11 1.11 1 .12 16 .18 Notroois lonairostris 282 4.11 11 1.11 39 4.60 332 3.82 Notroois maculatus 83 8.41 83 .95 Notropis roseioinnis 8 .94 .8 .09 Notroois texanus 8 .12 41 4.84 49 .56 Notroois venustus 5565 81.04 126 12.77 609 71.90 6300 72.41 Notroois volucellus 14 .20 14 .16 I Oosoooeodus emiliae 4 .06 36 3.65 10 1.18 50 .57 Pimeohales viailax 87 1.27 87 1.00 Catostomidae I Caroiodes cvorinus 5 .51 5 .06 Caroiodes velifer 9 .13 22 2.23 31 .36 Erimvzon tenuis 1 .10 1 .12 2 .02 i Hvoentelium niaricans 1 .01 1 .01
Ictaluridae Ictalurus natalis 1 .12 1 .01 I Ictalurus ounctatus 13 1.53 13 .15 Noturus ovrinus 26 .38 1 .12 27 .31 Notums leotacanthus 1 .01 9 1.06 10 .11 Noturus nocturnus 2 .03 9 1.06 11 .13 Pvlodictis olivaris 4 .06 4 .05 k Esocidae Esox americanus 4 .41 4 .05 Esox nicer 24 2.43 24 .28 k Aphredoderidae k Aphredoderus savanus 44 .64 7 .71 51 .59 38
Main Channel Backwaters Tributaries Total Soecies IL. % IL. % N. % N. %
Atherinidae Labidesthes sicculus 5 .07 2 .20 2 .24 9 .10
Fundulidae Fiindulus blairae 1 .01 5 .51 6 .07 Fundulus chrvsotus 3 .30 3 .03 Fundulus notatus 1 .01 1 .01 Fundulus notti 6 .61 7 .83 13 .15 Fundulus olivaceus 5 .07 6 .71 11 .13
Poeciliidae Gambusia affinis 66 .96 387 39.21 15 1.77 468 5.38
Centrarchidae Airibloolites arioramus 10 .15 7 .83 17 .20 Centrarchus macroDterus 2 .20 2 .02 Leoomis cvanellus 4 .06 4 .05 Lepomis oulosus 1 .01 9 .91 10 .11 Leoomis macrochirus 67 .98 152 15.40 14 1.65 233 2.68 Leoomis marainatus 13 1.32 13 .15 Lepomis meaalotis 94 1.37 9 .91 17 2.01 120 1.38 Lepomis microloohus 2 .03 2 .02 Leoomis ounctatus 1 .10 1 .12 2 .02 Microoterus ounctulatus 16 .23 1 .10 3 I .35 20 .23 Microoterus salmoides 44 .64 27 2.74 71 .82 Pomoxis annularis 8 .12 1 .10 9 .10 k Pomoxis niaromaculatus 21 .31 21 .24
Percichthyidae i Morone saxatilis 1 .01 1 .01
Percidae I Ammocrypta beani 168 2.45 168 1.93 Etheostoma chlorosomum 1 .10 1 .01 Etheostoma histrio 25 .36 3 .35 28 .32 Etheostoma oroeliare 1 .01 1 .10 2 .02 I Etheostoma swaini 2 .03 10 1.18 12 .14 Percina lenticula 17 .25 4 .47 21 .24 Percina niorofasciata 31 .45 15 1.77 46 .53 I Percina sciera 23 .33 23 .26 Percina shumardi 9 .13 9 .10 Percina sp. 1 12 .17 12 .14 I Percina sp. 3 9 .13 9 .10 Percina viail 4 .06 4 .05
Soleidae I Trinectes maculatus 85 1.24 4 .41 89 1.02
*** Totals *** 6, I 867 987 847 8, 701
I 39 habitats. Collections from the tributary streams were numerically dominated by weed shiners (4.8%), longnose shiners (4.6%), longear sunfish (2%), Percina niarofasciata. blaclcbanded darter (1.8%), mosquitofish (1.8%), bluegill (1.7%) and channel catfish (1.5%),
in addition to blacktail shiners. The species-sample curve for the main channel of the ChicJc- asawhay River (Figure 4) shows a general increase with sample number across most of the abscissa, indicating that sampling was only marginally sufficient. Backwater and tributary streams both would require more sampling to adequately characterize the fish species composition. The species abundance data showed considerable variation in the modal area of the lognormal cuirve (Fig. 5) . Such variation can greatly affect the shape of the curve and thus both the estimates of sampling sufficiency and the potential number of rare species. Based on the number of fish species taken in our collections (Tables 7—9), the Pascagoula River system (68 fish species) was somewhat more diverse than the Leaf (64) or Chickasawhay (64) systems. However, comparisons of total species numbers can be biased by sampling effort. To control, in part, for sampling effects we used rarefaction curves to compare species richness of the three river systems (Fig. 6) . Using a standard sample size of 700 specimens, the three systems are virtually identical in species richness. Such a high similarity is not unexpected given the close and general similarity in size of the three areas. Species composition is also very similar between the three river 40 systems. An index of similarity, Morisita's Index (Brewer and Zar 1984), gives values of >95% for all comparisons. Overall, the Pascagoula River drainage supports a rich native fish fauna. Our recording of 83 species for the 165 km study area is very close to the 86 species listed by Swift et al. (1986) for the entire Pascagoula drainage. A number of species that Swift et • (1986) listed as being questionable for the Pascagoula drainage Hiodon teraisus. Hvboanathus havi. Cairoiodes cvorinus. C. velifer, Ictiobus bubalus. Ictalurus furcatus. Enneacanthus
crloriosus and Etheostoma chlorosomum) have been verified as occurring in the Pascagoula drainage by this study. In addition, one of the more common minnows that we recorded in our samples, H. nuchalis, was listed by Swift et al. (1986) as not occurring in the Pascagoula drainage. Enneacanthus qloriosus was recorded by « Clemmer et al. (1975) and Miller and Clemmer (1980) as occurring in the Pascagoula River drainage in Bluff Creek near Vancleave. In a more recent comparison, Ross and Brenneman (in prep.) have recorded 124 species from the entire Pascagoula drainage. However, the present study also added two species, Etheostoma
proeliare and Fundulus blairae. which were not included in the"
listing by Ross and Brenneman. 41
Rarefaction curves for fish species from the Pascagoula, and. Chickasawhay river systems. See text for further explanation. (FiSwsor CO yj o LU Q- co UL O cn ULl CQ 2 3 •— All Habitats Z ^— Backwater Q UJ a— Main Channel 1- o— Side Streams o LU CL X LU 100 200 300 400 500 600 700 800
CO LU o LU •l CO IJL o cc LU CQ All Habitats Main Channel Q Side Streams LU ts LU Q-
2 100 200 300 400 500 600 700 800
CO yj o LU CL CO LL O CC LU CQ 2 3 All Habitats Z Backwater Q LU Main Channel H— O Side Streams LU CL X LU 100 200 300 400 500 600 700 800 SAMPLE SIZE 43
CONCLUSIONS
The Pascagoula River drainage suppoirts a small recreational fishery with target species being primarily channel catfish and sunfish. There is presently minimal commercial fishing (e.g. use of hoopnets) . The catch by anglers is dominated numerically by sunfish, followed by channel catfish; in terms of weight catfish rank highest in importance. The degree of fishing activity was likely depressed during the study period because of abnormally low river flow which occurred from mid-May to July. 165 km study section of the Pascagoula drainage possesses ^ diverse native fish fauna of at least 83 species. Additional species would likely be added with further collecting effort, especially by sampling over a longer time period
ACKNOWLEDGEMENTS We thank Harry Barkley and Ron Garavelli (MDWC) for making this study possible, and Ken Rich and David Robinson (MDWC) for providing the outboard motors used in the study. We are grateful to Aileen Minchew, Julie Kelley, Bill Brenneman and Barbara Viskup for helping with field work. 44
LITERATURE CITED Brower, J. E. and J. H. Zar. 1984. Field and laboratory methods for general ecology, second edition. Wm. C. Brown, Dubuque. Clemmer, G. H., R. D. Suttkus and J. S. Ramsey. 1975. A prelimi- nary checklist of endangered and rare fishes of Mississippi,
p. 6-11. In; A Preliminary List of Rare and Threatened Vertebrates in Mississippi. Mississippi Game and Fish
Commission, Jackson. S. H. 1971. The nonconcept of species diversity: a critique and alternative parameters. Ecology 52:577-586. James, F. C. and S. Rathbun. 1981. Rarefaction, relative abund
ance, and diversity of avian communities. The Auk 98:785-
800. Laws, E. A. and J. W. Archie, 1981. Appropriate use of regression analysis in marine biology. Marine Biology 65:13-1§. Ludwig, J, A. and J. F. Reynolds. 1988. Statistical Ecology: A Primer on Methods and Computing. John Wiley and Sons, New
York. Malvestuto, S. P. 1983. Chapter 21, Sampling the recreational fishery, p. 397-419. In: L. A. Nielsen, and D. L".~~Johnson (eds.). Fisheries Techniques. American Fisheries Society,
Bethesda. May, R. M. 1975. Patterns of species abundances and diversity,
P* 81—120. In: M. L. Cody and J. M. Diamond (eds.) , Ecology and Evolution of Communities. Belknap Press, Cambridge. f 46 Appendix I. Lengths (mm SL) and Weights (g) of Fishes From the Chickasawhay, I Leaf and Pascagoula Rivers, 1988. Species Lenath Weiaht
Ambloplites ariommus 135.0 90.0 140.0 100.0 155.0 130.0 • 175.0 210.0 125.0 70.0 140.0 110.0 105.0 50.0 f 155.0 140.0 155.0 150.0 I 105.0 39.0 I 140.0 120.0 120.0 60. 0 185.0 200.0 I 140.0 105.0 155.0 125.0 130.0 65.0
total weight. 1764.0
1 Ancniilla rostrata I 715.0 740.0 645.0 660.0
il total weight. 1400.0 Carpiodes spp. 360.0 1100.0 345.0 930.0 325.0 770.0 200.0 225.0 210.0 250.0 240.0 410.0 325.0 780.0 225.0 340.0 350.0 930.0 335.0 940.0 310.0 680.0 I 340.0 930.0 I 350.0 840.0 355.0 980.0 360.0 880.0 I 325.0 770.0 205.0 255.0 I 210.0 270.0 total wt. 12280.0 I I 47
Species Length Weight
CarDiodes velifer 130.0 60.0
Dorosoma cepedianim 265.0 370.0 f Ictalurus furcatus 220.0 160.0 195.0 110.0 265.0 250.0 f 570.0 3410.0 total weight. 3930.0
Ictalurus punctatus 160.0 60.0 185.0 100.0 430.0 1200.0 240.0 250.0 260 320.0 305 430.0 295 420.0 345 660.0 285.0 380.0 305.0 440.0 295.0 410.0 195.0 125.0 280.0 295.0 315.0 420 305.0 320 340.0 520 345.0 620 205.0 110. 370.0 610.0 305.0 340.0
total weight. 8030.0
Ictiobus bubalus 470.0 2800.0 305.0 810.0 355.0 1240.0 385.0 1850.0
total weight. 6700.0
Lepisosteus oculatus 715.0 1800.0 605.0 1270.0 355.0 200.0
total weight. 3270.0
Lepomis gulosus 135.0 90.0 p 48
Species Length Weight
Lepomis macrochirus 95.0 45.0 125.0 60.0 135.0 65.0 90.0 25.0 65.0 9.0 69.0 11.0 68.0 10.0 130.0 90.0 85.0 20.0 70.0 10.0 90.0 20.0 70.0 10.0 65.0 10.0 115.0. 60.0 90.0 20.0 95.0 30.0 115.0 60.0 140.0 120.0 110.0 50.0 120.0 60.0 135.0 95.0 110.0 45.0 110.0 45.0 100.0 32.0 95.0 28.0 75.0 15.0 85.0 20.0 75.0 15.5 95.0 30.5 70.0 11.5 • 65.0 9.0 125.0 43.5
total weight. 1175.0
Lepomis megalotis 110.0 55.0 115.0 45.0 100.0 32.0 i 57.0 8.0 105.0 40.0 I 120.0 72.0 110.0 50.0 125.0 80.0 80.0 20.0 I 80.0 15.0 115.0 60.0 80.0 20.0 I 80.0 20.0 80.0 15.0 I 80.0 20.0 I 49
Species Lencrbh
(continued) 125.0 70.0 75.0 15.0 125.0 80.0 115.0 50. 0 110.0 45.0 105.0 40.0 125.0 80.0 110.0 40.0 110.0 40.0 80.0 18.5 115.0 55.0 95.0 28.0 80. 0 17.5 50,0 4.0 75.0 14.0 60.0 7.0 100.0 35.0 115.0 47.0 110.0 50.0 105.0 40. 0 80.0 16.0 105.0 38.0 90.0 21.5 60.0 7.0 70.0 11.0 85.0 19.5 80.0 17.5 70.0 10.0 75.0 14.5 65.0 10.5 80.0 17.5
total weight. 1511.0 Lepomis roicrolophus 165.0 140.0 150.0 120.0 140.0 90. 0 140.0 100.0 190.0 235.0 • 150.0. 110. 0 105.0 40.0 85.0 19.0 175.0 180.0 85.0 20.5 145.0 90.0 165.0 150.0 165.0 170.0 total weight. 1464.5 I
/ I 50 Species Length Weight
Micropterus punctulatus 225.0 240.0 250.0 250.0 195.0 150.0
total weight. 640.0
Micropterus salmoides 455.0 2250 320.0 830 190.0 160 210.0 190
total weight. 3430.0
Morone saxatilis 360.0 900.0
Muail cephalus 235.0 250.0
Pomoxis annularis 145 60 160 80 155 75 i 220 240 175 110 205 220.0 135 52.0 i 140 62.0 135 52.0 145 120.0
total weight. 1071.0
Pomoxis niaromaculatus 175.0 165.0
Pvlodictis olivaris 310.0 480.0 310.0 420.0 I 390.0 850.0 520.0 2710.0 620.0 4100.0 I -430 1360.0 415 1130.0 340 590.0 I 395 930.0 395 930.0 355 645.0 395.0 1020.0 I 270.0 290.0 225.0 290.0 330.0 490.0 I 305.0 460,0 290.0 340.0 I I 51
Species Lencrfch Weight
Pvlodictis olivaris (cont.) 515.0 2150 530.0 2310 345.0 640 295.0 390 255.0 230 410.0 1020 215.0 160,0 215.0 160.0 445.0 1300.0 520.0 2250 545.0 2800 515.0 2280 210.0 130
total weight. 32855.0
\