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ivist stlin di leontologi e trtigrfi volume IPH noF P ppF IVQEIWH tuly PHIR e xi qix exh igsi yp ri pewsv egregrshei @gyviyieD ehireqeA pyw ri i vehsxsex @wshhvi sesgA yp wyxi ex qsyqsy I PB I ve e ehe D weiy wyxeqxe 8 exhie sxys eeivedX perury TD PHIRY eptedX tune IQD PHIR uey wordsX possil snsetD rhyphideD new genus nd speE its glol pleontologil mening of its mrine verteE iesD predtorD widdle rissiD pleoenvironmentF rte funsF egrding the ulkshieferzoneD topof weride estrtF e new genus nd speies of fossil rhyphide vimestoneD dted vte vdininD PRH ± PQW w followE redodromeus sngiorgiensis genF nFD nF spF @goleopterD rhyphiE deD iodromeineAD with well developed symmetril mndilesD ing tokr et lF @PHIPAD it ws so fr renowned for shorter thn the hedD suretngulr pronotum nd slender tiieD its vertertesD mostly fishes ut lso few reptiles longer thn femorD is desried from the ulkshieferzone @uppermost @vomrdo PHHIY enesto et lF PHHQY intori IWWHY memer of the weride vimestoneA of the wiss side of wonte n intori 8 vomrdo PHHUAF roweverD lso rthropodsD qiorgioF rhyphide re terrestril goleopter well represented suh s onhostrns @intori IWWHY intori 8 frmE in the wesozoi fossil reord y the extint fmily iodromeineF p to nowD more thn QQ fossil speies elonging to IQ gener of iodroE ill IWWIA nd mysidiens @vrghi et lFD sumittedAD meine hve een desriedF he new speies is prt of olletion of re importnt in the ulkshieferzone fossil ssemE fossil insets tht inludes memers of t lest six different ordersF lgesF fsed on the eology of orresponding extnt txD this inset ssemE huring the fieldwork rried out etween IWWU lge provides new insights on the pleoenvironment of wonte n qiorgio re during the vte vdininF he ssemlge inludes terresE nd PHHQ in the vower ulkshieferzone t the l wr tril tx tht support the presene of emerged lnd nd tx with longE site h ner werideD out IT speimens of insets @oth lsting quti lrvl stge @s leopter nd iphemeropterA tht onE dult nd lrvl formsA hve een olletedF ynly two firm the presene of stle fresh wter sinsF speimens hve een so fr desried s new genus nd speies of iphemeropter @urzeminski 8 vomrdo PHHIAF sn the sme pperD urzeminski 8 vomrdo sntrodution @PHHIA desried lso speimen of xotoupes @goleopE terD ymmtideA from the vower weride vimestoneF wonte n qiorgio is fossil vgersttte known È th eentlyD new speies of hsyleptide @erheogntA entury due to industril exploittion of sine the IW from the pper ulkshieferzone ws desried y ituminous shles s well s for the numerous pleonE fehley 8 tokr @PHIIAF hese findings mke the tologil exvtions tht took ple minly in the lst ulkshieferzone of wonte n qiorgio unique site enturyF sn timeD it hs yielded severl remrkle fossils for widdle rissi insetsF of mrine fishes nd reptiles s well s invertertes nd e herein desrie new speies nd genus of terrestril plnt reminsF vying ross the stlinEwiss iodromeine @goleopterD rhyphideAF rhypE oundry to the outh of the geresio vke @pigF IAD this re is now in the orld reritge vist of xigy for hyde inludes the single extnt sufmily rhypE I hiprtimento di ienze dell err erdito hesio99D i wngiglli QRD sEPHIQQ wilnoD stlyF iEmilX lurFstrddunimiFitD ndreFtintoridunimiFit P hiprtimento di ienze egrrie e emientli E niversitÁ degli tudi di wilnoD i qiovnni gelori P E PHIQQ wilnoD stlyF iEmilX mtteoFmontgndunimiFit B gorresponding uthor IVR trd vFD wontgn wF 8 intori eF pigF I E l wr site h E geogrphi positionF the depositionl sinF elterntivelyD it fell diretly on hine @two gener desriedX rhyphus nd ystoE losomAD for totl of six speies @wddison et lF PHHWA the wter surfe while flyingD floted for some time nd the extint sufmily iodromeine @onomrenko undergoing rpid initil dey in the wrm superfiil IWUUAF iodromeine is well represented in the fossil reE wters efore sinking to the ottomD where uril nd ord from the rissi to the greteous nd it is muh fossiliztion then took pleF woreoverD sine floting more diverse thn rhyphine @onomrenko inset remins re sujeted to predtionD they were IWWIAF et presentD t lest QQ fossil speies elonging proly trnsported only for short distnesF he to IQ gener of iodromeine hve een desried @oE time lg etween deth nd uril determined the exE nomrenko IWUUY orino et lF PHHUY rong IWVQY ti 8 tent of soft tissues deyX when the ody rehed the en PHIIY ng et lF PHIPY onomrenko 8 olkov ottom nd possilyD during initil postEuril ompE PHIQAF he oldest gener re from the erminl ermin tion of the sedimentD the disrtiultion eme pprE @onomrenko 8 olkov PHIQA nd only one other entF sn the speimenD the ody lyout is preserved so genus @ogdodromeus onomrenkoD IWUUA is from the we n infer tht dey ws time limited nd the deE rissi @onomrenko IWUUAF elsoD during our preliE positionl environment ws without ny urrent tht minry survey on this inset olletionD the undesried ould helpin the disperslof the somewht loose ody speimens were tenttively ssigned to six orders prtsF nmely iphemeropterD yrthopterD leopterD imE he degree of onservtion of the other speimens iopterD remipter nd goleopterF fsed on the urE vries gretlyD proly reording strong vritions in rent knowledge out the eology of these ordersD sed the tphonomil historyF sn few instnesD only hedD on oth extnt nd fossil speiesD we provide new inE thorx nd domen outline is preservedD while in other sights on the pleoenvironment of wonte n qiorgio ses lso ppendges reminedF sn lmost oneEhlf of t lest during the vte vdininF the fossils soft tissues preservtion oursD proly due to phosphtiztion proessD sometimes preserving inE ternl strutures in detilF snset tphonomy he speimen under study preserves the generl lyout of the odyF xonethelessD ody segments nd wteril nd methods legsD nd to lesser extentD hed prts nd domen re he speimen hs een olleted in lolity h in l wrD ner displed from their ntomil position in the living weride @gnton iinoD witzerlndA nd is deposited t wuseo gnE nimlF his lyout suggests tht the speimen went tonle di tori xturle di vugno @wgxAF peimen mnipultion through first stge of dey in sueril ndGor superE nd mesures were ompleted with the uxiliry use of the stereo miE fiil quti environmentD during whih soft tissues rosope vei wS nd n oulr mirometerF smges were quired prtilly degrdedF hereforeD either the inset died y digitl mer gnon ios RSH mounted on mhinery in order to nd prtilly deyed on the groundD its originl hiE sn the smple t different fous lyers tht were mounted with erE ttD nd ws then trnsported y high9 energy eventD ene tker IFH @tudent iditionAF hrwings were mde with mer luid tthed to the mirosopeF suh s flood following hevy rins or just windD to e new goleopter from the vdinin of wonte n qiorgio IVS pigF P E eA redodromeus sngiorE giensis genF nD spF nF vength from pex of mndiules to pex of elytr a IHFV mmF fA drwing of the speimenF ystemti pleontology hesription elmost omplete impression in dorsl viewD lkE yrder goleopter vinneusD IUSV ing ntenneF red nd domen in nturl positionD uorder edephg hellenergD IVHT single disrtiulted elytron preservedF vength from pex of mndiules to pex of elytr of IHFV mmY width pmily rhyphide homsonD IVSU t se of elytr of RFT mm @lengthEwidth rtio PFQRAF ufmily iodromeine onomrenkoD IWUU ell three legs on the right side preservedF wid nd qenus redodromeus genF novF hind femor rther lrge nd roustY mximum width of ype speiesX redodromeus sngiorgiensis nF spF y monotypyF right mid femor is HFUW mmF iie thin nd slenderF he preserved trsi @sCssA elongted nd slimY only two itymologyX he nme derived from the vtin ver predo99D fore trsomers distinguishleD first one enlrgedF yn whih mens to preyD onsidering its strong nd shrp mndiles tht we supposed to e used to th preyD nd the suffix ±dromeus99D whih the left side preserved only ssCsss femor nd sss tiiF mens runner99D from its ursory legsF ight fore tiie @length IFWP mmA hrouring simple vous typiusX ite hD l wr ner werideD wendrisioD gnE sulte ntenn lener orgn with enlrged distl re ton iinoD witzerlndF nd n pil spur @HFPR mmAF pive trsi @totl length hignosisX he generl hitus of the new txon @genus or speiesAD simple sulte ntenn lener in the distl prt of the tiieD IFTP mmA on s right leg lerly visileF lender tiieF the metox seprting the thorx nd domenD the metepisternum red igD longer thn widerD length of QFH mm rehing the mesooxl vities prompts its sription to the to the mndiles exluded nd of QFRP mndiles inludedD fmily rhyphideD sufmily iodromeineF he new txon is deE width t the eyes PFUT mmF wndiles symmetril sried s new genus nd speies due to the presene of unique set of nd tringulrD roust nd shrpwith smll tooth in hrtersX well developed symmetril mndilesY retngulr pronoE tumD longitudinl groove sentY short metventriteY nd ig hed with the nterior seondEthird of the inner mrginF otl lrge eyes on the upper surfeF length of IFQV mmD width HFWT mmD lengthEwidth rtio of IFRRF rtil impression of lil plp visile etween redodromeus sngiorgiensis spF nF mndilesF pigF PeD f ronotum wide @QFI mmAD nterior mrgin slightly onve with pointed pies @left sideAD posterior mrgin rolotypeX wgx VRSIF strightF itymologyX he nme derived from the olleting re of edomen prtilly visile with light impressions wonte n qiorgio @witzerlndGstlyAF of segmenttion preserved on the left sideF vength of hignosisX s for the genusD eing the only speiesF IVT trd vFD wontgn wF 8 intori eF domen IFS times the length of mesoE nd metthores se nd on n often noxi ottomD s indited y ommon lminted limestone or mrlyElimestone lyers ominedF nd the lmost generl sene of ioturtion @intori hisussion IWWHY intori 8 enesto IWWHY intori 8 vomrdo