National Park Service U.S. Department of the Interior

Natural Resource Program Center

Annotated Checklist of Vascular Flora Hovenweep National Monument

Natural Resource Technical Report NPS/NCPN/NRTR—2009/207 ON THE COVER Hackberry (Celtis reticulata) from Hackberry Unit, Hovenweep National Monument, Colorado. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Hovenweep National Monument

Natural Resource Technical Report NPS/NCPN/NRTR—2009/207

Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741

Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

May 2009

U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientifi c community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer-reviewed to ensure that the information is scientifi cally credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner.

The Natural Resource Technical Report series is used to disseminate the peer-reviewed results of scientifi c studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service’s mission. The reports provide contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. Current examples of such reports include the results of research that addresses natural resource management issues; natural resource inventory and monitoring activities; resource assessment reports; scientifi c literature reviews; and peer- reviewed proceedings of technical workshops, conferences, or symposia.

Views, statements, fi ndings, conclusions, recommendations, and data in this report are solely those of the author(s) and do not necessarily refl ect views and policies of the U.S. Department of the Interior, National Park Service. Mention of trade names or commercial products does not constitute endorsement or recommendation for use by the National Park Service.

Printed copies of reports in this series may be produced in limited quantity, and are only available as long as the supply lasts. You may send a request to:

Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

This report is also available electronically from the Northern Colorado Plateau Network website, http://science.nature.nps.gov/im/units/ncpn/index.cfm, or at http://www.nature.nps.gov/publications/NRPM/index.cfm.

Please cite this publication as: Fertig, W. 2009. Annotated checklist of vascular fl ora: Hovenweep National Monument. Natural Resource Technical Report NPS/NCPN/NRTR—2009/207. National Park Service, Fort Collins, Colorado.

NPS D-63, May 2009 ii Annotated Checklist of Vascular Flora: Hovenweep National Monument Contents

Figures ...... v Tables ...... v Abstract ...... vii Acknowledgements ...... ix

1 Introduction ...... 1 1.1 Background and justifi cation ...... 1 1.2 Objectives ...... 1 1.3 Study area ...... 1 1.3.1 History and setting ...... 1 1.3.2 Geology ...... 2 1.3.3 Vegetation ...... 2 1.3.4 Previous botanical studies ...... 2

2 Methods ...... 7

3 Results ...... 9 3.1 Summary of the fl ora of Hovenweep National Monument ...... 9 3.2 Life form and biogeographic diversity ...... 11 3.3 Plants of special concern ...... 11 3.4 Non-native species and noxious weeds ...... 11 4 Discussion ...... 15

5 Literature Cited ...... 17

Appendices ...... 21 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family and scientifi c name with taxonomic and biogeographic notes...... 23 Appendix B. Potential (unconfi rmed) taxa of Hovenweep National Monument...... 57 Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument...... 78 Appendix D. Confi rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes...... 94

Contents iii

Figures

Figure 1.3.1. Location of Hovenweep National Monument, San Juan County, Utah, and Montezuma County, Colorado...... 3 Figure 1.3.4. Increase in the number of plant taxa confi rmed as present in Hovenweep National Monument by decade, 1972–2006...... 5

Tables

Table 1.3.4. Chronology of plant-collecting efforts in Hovenweep National Monument...... 4 Table 3.1-1. Statistical summary of the fl ora of Hovenweep National Monument...... 9 Table 3.1-2. Statistical summary of the fl ora of Hovenweep National Monument, by state...... 10 Table 3.3. Plant species of conservation concern from Hovenweep National Monument...... 13 Table 3.3.1. Noxious weed species of Hovenweep National Monument...... 14

Contents v

Abstract

In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Hovenweep National Monument. This entailed a thorough review of existing literature, re-examination of specimens from the Hovenweep National Monument herbarium and other state and national museum collections, and additional fi eld work to corroborate unverifi ed species reports or locate new species. Based on this study, 340 vascu- lar plant taxa have been confi rmed or reliably reported as occuring in the monument (273 are known from validated herbarium specimens and 67 from unvouchered literature reports). Another 123 spe- cies formerly considered part of the monument fl ora have now been shown to be falsely reported. The Utah portion of Hovenweep National Monument contains 218 species (6% of the state fl ora) and the Colorado segments contain 268 species (8.5% of the entire Colorado fl ora). At least 16 of these species are tracked as species of concern by state natural heritage programs, but none are federally listed as threatened, endangered, or sensitive. Introduced plants make up 16.7% of the monument fl ora (57 species), of which 12 species are considered noxious weeds by the states of Utah and Colorado. Fur- ther surveys in Hovenweep National Monument are likely to fi nd additional new species (about 270 are known from similar habitats outside the monument boundary), especially among late-fl owering, weedy, or wetland taxa.

Contents vii

Acknowledgements

I would like to thank the following individuals for their assistance with this project: Dr. Angela Even- den, botanist and former leader of the NCPN program in Moab, was an active proponent of this proj- ect and helped secure funding for fi eld work and herbarium studies in 2006. Vicki Webster, curator of museum collections for the National Park Service Southeast Utah Group (SEUG), provided access to all mounted and unmounted herbarium specimens from Hovenweep National Monument (HOVE), as well as workspace within the museum vaults. Charlie Schelz, biologist with SEUG, provided research permits for work in 2005–2006 and shared his unmounted plant collections. Libby Nance, former database specialist with the NCPN, assisted with development and quality control of the monument’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories on the monument. Drs. Stanley Welsh and Duane At- wood provided access to herbarium collections at Brigham Young University (BRY). Dr. Michael Piep provided access to collections at the Utah State University (Intermountain) herbarium. Dr. Ron Hart- man and Ernie Nelson provided access and specimen loans from the Rocky Mountain Herbarium at the University of Wyoming. Thanks to the technical reviewers (Dr. Angela Evenden, Dr. Dusty Perkins, and Libby Nance) for their comments and suggestions. To any others who I may have inadvertently forgotten, please accept my apologies (and thanks).

Research was conducted under permit # HOVE-2006-SCI-0002 for study number HOVE-00010. Funding was provided by the National Park Service through the University of Wyoming.

Contents ix

et al. 2002). This eff ort has entailed reviewing ex- 1 Introduction isting species checklists, corroborating the iden- tifi cation of specimens in park museum and her- In 2006, the National Park Service (NPS) estab- barium collections, locating additional specimen lished a cooperative agreement with the University records from other museums, mining relevant of Wyoming, through the Rocky Mountains Co- literature and datasets, and conducting fi eld sur- operative Ecosystem Studies Unit, to produce a se- veys to locate new taxa or confi rm unsubstantiated ries of technical reports summarizing the vascular reports.† All of these data have been synthesized plant fl ora of each of the 16 national park units of in NPSpecies, the park service’s standardized the Northern Colorado Plateau Network (NCPN). database for documenting park status (present, These reports are based on existing information probably present, unconfi rmed, falsely reported), in the park service’s NPSpecies database system abundance, residency, nativity, and rarity. With the and augmented by further data mining, reviews of production of a technical series of reports on the state and regional herbaria, and new fi eld surveys fl ora of each park, the NPSpecies data will now be to resolve the status of many reported and uncon- available to a broader audience. fi rmed species. The objective of these reports is to summarize the current state of knowledge of the fl ora of each park for the benefi t of NPS biologists 1.2 Objectives and interpretive staff , outside researchers, and the The objectives of this project were to: general public. The following report addresses the vascular fl ora of Hovenweep National Monu- 1. Document the current status (present, ment, in southeast Utah and southwest Colorado. reported, potential, or falsely reported) of Other NCPN park units are addressed in separate all vascular plant species that have been reports. cited as occurring in Hovenweep National Monument based on a review of herbarium collections and past literature. 1.1 Background and justifi cation One of the primary missions of the NPS is “to con- 2. Develop an annotated checklist of the fl ora serve unimpaired the natural and cultural resourc- of Hovenweep National Monument with es and values of the national park system for the supplemental information on taxonomic enjoyment of this and future generations” (NPS synonyms, distribution, within-park abun- 2000). Due to their high degree of protection and dance, fl owering period, growth form, nativ- distribution across a wide variety of habitat types, ity, habitat, and sources for each record. the nation’s national park units make a signifi - cant contribution to the protection of biological 1.3 Study area diversity (Noss and Cooperrider 1994). Unfortu- nately, the full contribution of the NPS system is 1.3.1 History and setting not known, as many park units have traditionally In 1854, Mormon settlers exploring southeastern lacked comprehensive biological inventories or Utah found the unusual stone towers and other ar- monitoring programs to identify the composition cheological remains of the Hovenweep area. These and condition of their biota. Knowing the status structures were apparently constructed by small of park fl ora and fauna is critical for eff ective park communities of Puebloan farmers between 1235 management and for systematically identifying and and 1240 A.D. The towers were likely occupied fi lling gaps in the nation’s protected area network for no more than 70 years and then abandoned (Margules and Pressey 2000). as the Puebloans migrated to southern Arizona and New Mexico, following a prolonged drought. To rectify this shortcoming, Congress passed the Pioneer western photographer William Henry National Parks Omnibus Management Act in 1998 Jackson visited the area in 1874, and popularized to provide guidance and funding for an inventory the name Hovenweep (a Ute word meaning “de- and monitoring program across the entire Nation- serted valley”) for the area. Archeologists from the al Park System. Since 2000, the NCPN Inventory Smithsonian Institution surveyed the Hovenweep and Monitoring Program has been developing and area in 1918, and recommended that the structures verifying species lists of vertebrates and vascular be protected by the federal government. President plants for each park unit in the network (Evenden

†Taxa refer to all named species, subspecies, and varieties.

Chapter 1: Introduction 1 Warren G. Harding established Hovenweep Na- 1.3.3 Vegetation tional Monument by Presidential Proclamation in Rocky mesa rims and slopes in the Square Tower, March 1923 (Evenden et al. 2002). Holly, Hackberry, and Cutthroat Castle units con- Hovenweep National Monument (NM) currently tain woodlands dominated by Utah juniper (Junipe- consists of six separate units totaling 318 hectares rus osteosperma)/two-needle pinyon (Pinus edulis) (785 acres) in southeastern San Juan County, Utah, and cliff rose (Purshia mexicana var. stansburyana), and adjacent southwestern Montezuma County, with an understory of basin big sagebrush (Artemi- Colorado (Figure 1.3.1). The Cajon, Square Tower, sia tridentata var. tridentata), muttongrass (Poa Holly, Hackberry (or Horseshoe), and Cutthroat fendleriana), broom snakeweed (Gutierrezia saro- Castle units are located on Cajon Mesa along both thrae), and slenderleaf rabbitbrush (Chrysotham- sides of the state line, about 30 miles north of nus viscidifl orus var. stenophyllus). Open areas on Four Corners. The Goodman Point unit is located mesa tops consist mostly of basin big sagebrush about 20 km to the east in the Montezuma Valley, or Wyoming big sagebrush (A. tridentata var. wyo- northwest of Cortez, Colorado. The four Colora- mingensis) and broom snakeweed with occasional do units (Holly, Hackberry, Cutthroat Castle, and Utah juniper. Snakeweed is especially abundant Goodman Point) are embedded within or border in areas of high sheep grazing during the fi rst half the U.S. Bureau of Land Management’s Canyon of of the twentieth century (Evenden et al. 2002). At the Ancients National Monument, established by Square Tower, shadscale (Atriplex confertifolia) and President William Clinton in 2000. cheatgrass (Bromus tectorum) co-dominate with sagebrush and rabbitbrush on dry lower slopes. Within Hovenweep NM, elevation ranges from Seeps below the canyon rim contain stands of Fre- 1,585 meters (5,200 feet) at the Cajon Unit to 2,060 mont cottonwood (Populus fremontii) and netleaf meters (6,760 feet) at Goodman Point (Evenden et hackberry (Celtis reticulata). al. 2002). The area averages 12 inches (305 mm) of annual precipitation. Mean annual temperature is The Cajon unit is the driest area of Hovenweep 52° F, but ranges from lows of –10° F in winter to and consists mostly of black sagebrush (Artemisia over 100° F in summer (Evenden et al. 2002). nova)-broom snakeweed-shadscale-galleta (Hilar- ia jamesii) vegetation. Colorado’s Goodman Point 1.3.2 Geology unit consists of extenstive Utah juniper/two-nee- dle pinyon pine woods interspersed by basin big The scattered units of Hovenweep NM are locat- sagebrush and meadows of smooth brome (Bro- ed on a relatively fl at, sedimentary tableland that mus inermis), squirreltail (Elymus elymoides), and is irregularly dissected by small canyons draining western wheatgrass (Elymus smithii). A Fremont into the San Juan River basin. The canyon bottoms cottonwood-golden currant (Ribes aureum) forest and lower slopes consist of light gray-to-maroon borders a small riparian corridor. mudstones and shales of the late-Jurassic Morri- son Formation. These sediments were deposited 1.3.4 Previous botanical studies on a broad fl oodplain by a meandering river drain- ing higher mountains to the west and south more The Four Corners region received much attention than 140 million years ago (Chronic and Chronic from early botanical explorers, but the oldest con- 2004). The Morrison layers are topped by 1–2 fi rmed fl oristic study of the Hovenweep area dates ledges of massive gray-to-brownish sandstone of to Richart (1941), an NPS employee who cata- the Cretaceous-age Dakota Formation (deposited logued the fl ora and fauna of several southwestern 92–96 million years ago). The Dakota Sandstone is parks. The earliest collections in the Hovenweep derived from beach or strand line deposits along National Monument herbarium (HOVE) were the shore of an inland sea. Seeps and springs are made in 1972, by Dale Schmidt and colleagues often associated with the relatively porous Dakota conducting botanical surveys in association with Formation (Chronic and Chronic 2004). Younger archeological research (Table 1.3.4, Figure 1.3.4). Cretaceous shales and sandstones of the Mancos Over the next three years, Dale and Marilyn Shale and Mesa Verde group eroded off the Ca- Schmidt, “Redfi eld,” and William Litzinger docu- jon Plateau during regional uplift in the Tertiary. mented 151 vascular plant taxa for Hovenweep Today, the mesa tops are often covered by Qua- NM (Litzinger and Winter 1976). Another 31 spe- ternary, wind-blown silts, which formed the fer- cies reported for Hovenweep during this time pe- tile farmlands used by the Puebloan culture in the riod (Schelz and Moran 2003) were based on Litz- thirteenth and fourteenth centuries. inger collections made at other archeological sites outside of the monument.

2 Annotated Checklist of Vascular Flora: Hovenweep National Monument Hovenweep National Monument Utah

491

Colorado

Utah

Cutthroat Castle Unit

Goodman Hackberry Unit Point To Hwy 191 Unit ! Holly Unit

Square Tower Unit Visitor

Center !

To Cortez, CO Cajon Unit

Aneth

To Four Corners !

Established March 2, 1923 Legend: 1,585–2,060 m elevation Park boundary (NPS) (5,200–6,760 ft elevation) Intermittent streams 318 ha (785 ac) Perennial streams Canyons of the Ancients (BLM) Rivers Bureau of Land Management (BLM) 0241 Kilometers Paved roads Navajo & Ute Mtn. Reservations (IR) 0241 Unpaved roads Private Miles Unimproved/4WD roads State

MAP CREATED BY ANETH WIGHT, NCPN (DERIVED FROM EVENDEN ET AL. 2002)

Figure 1.3.1. Location of Hovenweep National Monument, San Juan County, Utah, and Montezuma County, Colorado.

Chapter 1: Introduction 3 Over the next 25 years, no fl oristic research oc- Since 2003, vegetation mappers have added three curred at Hovenweep NM except for a period of new species reports for the Hovenweep NM fl ora brief collecting and rare plant surveys by Mari- (Von Loh et al. 2008). Kim Andersen added one lyn Colyer, of Mesa Verde National Park, during new species during a noxious weed inventory of 1992–1994. From 2001 to 2003, Charles Schelz of the monument (Dewey and Andersen 2005). Re- the NPS Southeast Utah Group conducted a com- becca Harms collected one new species for the prehensive literature and herbarium review and fl ora during the NCPN springs and seeps survey fl oristic inventory to fi ll gaps in the monument’s (Springer et al. 2006). In 2006, I added 13 addi- species list. Over this period, Schelz added more tional species during surveys of each of the monu- than 300 specimens to the HOVE herbarium and ment’s six subunits. vouchered 104 new plant species (Schelz 2002; 2003). Schelz and Moran (2003) compiled a check- list of the monument fl ora recognizing another 62 species as reported (but not vouchered) (Appen- dix A).

Table 1.3.4. Chronology of plant-collecting efforts in Hovenweep National Monument. # of new Year Collector(s) Comments taxa 1941 R. S. Richart ? Roland Richart (1941) compiled the earliest checklist of the fl ora of Hovenweep and Yucca House national monuments. 1972 D. Schmidt & 16 Dale Schmidt made the earliest collections for the HOVE herbarium. Redfi eld 1973 D. Schmidt, M. 59 Dale and Marilyn Schmidt and “Redfi eld” (fi rst name unknown) added 59 new Schmidt, & Redfi eld taxa to the monument species list. 1974– W. J. Litzinger & M. 76 William Litzinger added 70 new species to the monument fl ora during a three- 1976 Schmidt year project, collecting plant species associated with archeological sites in Hovenweep NM and surrounding areas of Cajon Mesa (now within the Canyon of the Ancients National Monument). Marilyn Schmidt contributed 6 additional species in 1974. 1992– M. Colyer & Oppelt 3 Marilyn Colyer, biologist from Mesa Verde National Park, documented three 1994 new species during rare plant surveys of the monument. 2001– C. Schelz 91 Charles Schelz, of the NPS Southeast Utah Group, initiated a multi-year fl oristic 2002 survey of the monument and added 91 new taxa to the Hovenweep fl ora. 2003 C. Schelz, K. 14 (+5 Kim Andersen collected one new species and reported one additional taxon Andersen, & reported) for the Hovenweep NM fl ora during a survey of noxious weeds (Dewey and NCPN vegetation Andersen 2005). Three new species were reported (but not vouchered) by Von mapping crew Loh et al. (2008). Schelz (2002, 2003) concluded his fl oristic survey with 14 new species records. 2005 R. Harms 1 One new species confi rmed during seeps and springs inventory of Springer et al. (2006). 2006 W. Fertig 13 Walter Fertig collected vouchers for 13 new species in 2006. The earliest collection date is not known for an additional 62 unvouchered reports cited by Schelz and Moran (2003).

4 Annotated Checklist of Vascular Flora: Hovenweep National Monument 300

250

200

150

100 Number of vascular taxa

50

0 1972 1976 1994 2003 2006

Year

Figure 1.3.4. Increase in the number of plant taxa confi rmed as present in Hovenweep National Monument by decade, 1972–2006.

Chapter 1: Introduction 5

context of Utah and Colorado), population size 2 Methods in the monument (derived from Von Loh et al. 2008, and fi eld observations), fl owering period, In July 2005, I reviewed all mounted specimens distribution across general vegetation types, na- in the Hovenweep NM herbarium (housed in tivity, data source (collector and collection num- the NPS Southeast Utah Group offi ce in Moab, ber from a sample specimen or literature cita- Utah) to determine whether each was correctly tion), year of earliest collection (from herbarium identifi ed to species, subspecies, or variety fol- records), and relevant synonymy and taxonomic lowing the nomenclature of Welsh et al. (2003). problems (based on review of Barkworth et As necessary, I made corrections and annotations al. 2003, 2007; Barneby 1989; Cronquist 1994; to each specimen and to the voucher records in Cronquist et al. 1972, 1977, 1984, 1997; Dorn the monument’s Automated National Cataloging 2001; Flora of North America Editorial Commit- System (ANCS+) and NPSpecies database. Based tee 1993, 1997, 2000, 2002a, 2002b, 2003, 2005, on this herbarium work and a review of monu- 2006a, 2006b, 2006c; Holmgren et al. 2005; and ment references (Dewey and Andersen 2005; Weber and Wittmann 2001). The annotations to Schelz and Moran 2003; Springer et al. 2006), I these lists and label data from the HOVE her- developed four lists: barium review were then incorporated into the NPSpecies database for the monument. 1. A revised checklist of species that were pres- ent in the park (confi rmed by an authenti- Work in 2006–2007 focused on confi rmation of cated voucher; included in Appendices A reported and potential species for the monument and D); fl ora through fi eld surveys and review of collec- tions from state and regional herbaria. I made one 2. A list of species that were reported in the lit- collecting trip to Hovenweep NM, in June 2006, erature but not corroborated with a voucher to target missing species, emphasizing under- (equivalent to “probably present” in NPSpe- sampled wetland and weedy taxa. Herbarium cies terminology; included in Appendices A studies were undertaken at Brigham Young Uni- and D); versity (BRY), Utah State University (USU), and 3. A list of potential species (“unconfi rmed” the University of Wyoming (RM) in March, June, in NPSpecies) known from the vicinity of and October 2006, and February and March Hovenweep National Monument using an 2007. I also documented monument specimens unpublished list generated by Dr. N. Duane from the Virtual Herbarium website of the New Atwood of Brigham Young University and York Botanical Garden (http://sciweb.nybg.org/ Albee et al. (1988) (included in Appendix B); science2/hcol/intf/index.asp). Finally, I found and additional new species reports by examining unmounted collections by NPS Southeast Utah 4. A list of species that are now known to be Group and NCPN researchers and by reviewing misidentifi ed or highly questionable (Ap- plot data from recent vegetation mapping and pendix C). wetland inventories (Von Loh et al. 2008; Spring- Each list was annotated with supplemental infor- er et al. 2006). mation on life form, geographic range (within the

Chapter 2: Methods 7

the University of Wyoming, and the New York 3 Results Botanical Garden Virtual Herbarium but found no additional species reports for Hovenweep 3.1 Summary of the fl ora of NM. Field surveys in 2006 netted 13 new spe- cies and a review of collections from the NCPN Hovenweep National Monument seeps and springs inventory project (Springer et Of the 606 vascular plant vouchers in the HOVE al. 2006) added one additional taxon. Overall, the herbarium that I examined in 2005 (excluding documented fl ora of Hovenweep National Mon- 142 specimens collected from outside the monu- ument has increased by 5.1% since 2005, bringing ment), 141 were misidentifi ed (23.2%), 63 were the total number of taxa to 273 (Tables 3.1-1, 3.1- revised to a more current name (10.4%), 70 had 2, Appendices A, D). variety or subspecies names added (11.6%), and 332 were found to be correctly determined Currently, 67 additional taxa have been reported (54.8%). Based solely on these collections, the for Hovenweep NM but have yet to be corrobo- confi rmed fl ora of Hovenweep National Monu- rated with a voucher specimen (Table 3.1-1, Ap- ment stood at 259 taxa (Fertig 2006). From 2005 pendix A). Of these, 63 were reported by Schelz to 2007, I reviewed herbarium collections at and Moran (2003) and the remainder by Dewey Brigham Young University, Utah State University, and Andersen (2005) and Von Loh et al. (2008).

Table 3.1-1. Statistical summary of the fl ora of Hovenweep National Monument. Number Number Category confi rmed additionally Total in park reported for park Taxonomic diversity Total taxa 273 67 340 (including varieties and subspecies) Full species 265 64 329 (excluding varieties and subspecies) Families 52 8 60 Life form diversity Tree taxa 4 0 4 Shrub taxa 51 8 59 Perennial forb taxa 121 28 149 Annual forb taxa 55 20 75 Perennial graminoid taxa 33 8 41 Annual graminoid taxa 9 3 12 Fern taxa 0 0 0 Biogeographic diversity Introduced taxa 45 12 57 Native taxa Locally endemic taxa 1 0 1 Regionally endemic taxa 24 6 30 Disjunct taxa 0 0 0 Peripheral taxa 19 5 24 Sparse taxa 4 1 5 Widespread taxa 180 43 223 Total native taxa 228 55 283 The number of taxa and families is based on taxonomic concepts of Welsh et al. (2003). See Appendices for an explanation of the various categories.

Chapter 3: Results 9 Table 3.1-2. Statistical summary of the fl ora of Hovenweep National Monument, by state. Present in park Reported for park Total Category UT CO UT CO NA* UT CO Taxonomic diversity Total taxa 189 236 29 32 22 218 268 (including varieties and subspecies) Full species 184 230 26 31 22 210 261 (excluding varieties and subspecies) Families 47 56** 3 1 5 50 57** Life form diversity Tree taxa 4 4 0 0 0 4 4 Shrub taxa 41 43 7 1 2 48 44 Perennial forb taxa 67 112 15 16 6 82 128 Annual forb taxa 45 42 6 10 9 51 52 Perennial graminoid taxa 23 31 1 4 4 24 35 Annual graminoid taxa 9 4 0 2 1 9 6 Fern taxa 0 0 0 0 0 0 0 Biogeographic diversity Introduced taxa 28 38 2 7 4 30 45 Native taxa Locally endemic taxa 0 1 0 0 0 0 1 Regionally endemic 15 15 5 4 0 20 19 taxa Disjunct taxa 0 0 0 0 0 0 0 Peripheral taxa 2 17 0 1 3 2 18 Sparse taxa 0 4 0 0 1 0 4 Widespread taxa 144 161 22 20 14 166 181 Total native taxa 161 198 27 25 18 188 223 The number of taxa and families is based on taxonomic concepts of Welsh et al. (2003) for Utah species and Weber and Wittmann (2001) for Colorado taxa. See Appendices for an explanation of the various categories.

*Not Available. Reported for Hovenweep by Schelz & Moran (2003) without a state or subunit affi liation (these are indicated by a ? in the state distribution column in Appendix A)

**The number of families recognized in Colorado is higher than that in Utah, or for the total area, because of the rec- ognition of several segregate families within Caryophyllaceae, Liliaceae, and Ranunculaceae by Weber and Wittmann (2001). The number of families for Utah and the total area follow the criteria of Welsh et al. (2003).

Taken together, the number of confi rmed and 2006). At least 12 of these species were found to reported vascular plant taxa in Hovenweep Na- occur in the monument based on fi eld collections tional Monument stands at 340 taxa (Table 3.1-1, or relocated herbarium specimens in 2006. An- Appendix A). other 40 species were eliminated from the poten- tial list based on their low probability of occurring Entering the 2006 fi eld season, I recognized in the monument. Most of these were species of 322 taxa as potentially occurring in Hovenweep montane coniferous forest or wetland habitats in NM, based on the proximity of the monument the Abajo Range or the San Juan River drainage. to known populations (Albee et al. 1988; Fertig The current number of potential species that I am

10 Annotated Checklist of Vascular Flora: Hovenweep National Monument recognizing for Hovenweep National Monument Hovenweep NM contains 283 native plant spe- is 270 (Appendix B). cies. Of these, 188 occur in the Utah units of the monument and 223 in the Colorado portions. Of the original pool of species documented or More than half (65.6%) of the monument’s spe- reported for Hovenweep NM, I found that 113 cies (223 species in all) are widespread across taxa were falsely reported based on misidentifi ed Utah, Colorado, or western North America (Ta- voucher specimens in the HOVE herbarium. I bles 3.1-1, 3.1-2). Thompson’s indigo-bush (Pso- consider an additional 10 species cited (but not rothamnus thompsoniae var. thompsoniae) is the vouchered) by Schelz and Moran (2003) and the only local endemic currently known from Hov- NCPN vegetation mapping team (Von Loh et al. enweep NM. Thirty additional species (8.8% of 2008) to be questionable based on the absence the total fl ora) are regional endemics restricted to of suitable habitat or the distance of Hovenweep the Colorado Plateau. Twenty-nine species (8.5%) NM to the known or expected range of each spe- are either sparsely distributed or at the very edge cies (Appendix C). of their contiguous range in Utah or Colorado (Tables 3.1-1, 3.1-2). The confi rmed and reported fl ora of the Utah portion of Hovenweep NM consists of 218 taxa. These species represent 6.1% of the 3,594 native 3.3 Plants of special concern and naturalized taxa documented within the state Hovenweep NM has no plants listed as threat- of Utah by Welsh et al. (2003) and 33.1% of the ened or endangered under the U.S. Endangered state’s 151 plant families (Table 3.1-2). Within San Species Act (Table 3.3) and none are currently Juan County, the monument fl ora captures 15.9% considered offi cial NPS species of concern of the 1,370 taxa reported by Welsh et al. (2003). (Schelz and Moran 2007). One species, Thomp- At the ecoregional scale, the Utah units of Hov- son’s indigo-bush, was formerly a candidate (3C) enweep NM contain 12.9% of the 1,694 species for potential listing, but currently has no formal reported for the Utah portion of the Colorado protective status. This species was not previously Plateau (Shultz et al. 2006). known from Colorado, but was discovered by Schelz in the Cutthroat Unit in 2001 (Schelz and Species richness is higher within the Colorado Moran 2003). It is not currently tracked by the units of Hovenweep NM. In all, 268 taxa are con- Colorado Natural Heritage Program. Two other fi rmed or reported from the state, representing Colorado species are considered sensitive by the 8.5% of the 3,145 species in the fl ora of Colorado U.S. Bureau of Land Management in Colorado (Weber and Wittmann 2001) and 35.6% of the (Cryptantha [Oreocarya] osterhoutii and Swertia 160 recognized plant families (Table 3.1-2). The utahensis [Frasera paniculata]), and a third (Calo- monument contains 25.9% of the 1,033 plant taxa chortus fl exuosus) is listed as sensitive by the U.S. reported for Montezuma County (Kartesz 2003). Forest Service (Table 3.3). In all, eight Hoven- The Colorado segment of the monument con- weep NM species are tracked as species of con- tains 14.1% of the 1,898 species in the state’s por- cern by the Colorado Natural Heritage Program tion of the Colorado Plateau ecoregion (Kartesz (2007). The Utah Division of Wildlife Resources 2003). Conservation Data Center (UDWR 1998) moni- tors seven additional taxa from the Utah portion 3.2 Life form and biogeographic of the monument (Table 3.3). diversity Although ecologically dominant, tree and shrub 3.4 Non-native species and noxious species make up only 18.5% of the confi rmed weeds and reported plant taxa of Hovenweep NM (Ta- Introduced plant species (not historically native ble 3.1-1). Perennial forbs are the most species- to Utah, Colorado, or North America) make up rich life form in the monument’s fl ora, with 149 16.8% of the confi rmed and reported fl ora of known and reported taxa (43.8% of the total fl o- Hovenweep NM (Table 3.1-1). This fi gure is high- ra). Annual forbs and grass-like plants comprise er than the statewide average of 13.5% non-native about 26% of the fl ora. Like trees and shrubs, pe- species in Utah (Fertig 2007; Welsh et al. 2003) rennial graminoids are ecologically important but and 15.6% for Colorado (Rejmanek and Randall relatively species-poor, representing only 12% of 1994). Of the monument’s 57 introduced taxa, 12 the total fl ora. No ferns and fern-allies have been are listed as noxious by the states of Utah (USDA- documented in the monument (Table 3.1-1). NRCS 2009) or Colorado (CDA 2009) (Table 3.4).

Chapter 3: Results 11 Plants on the Colorado state noxious weed list fall the goal for species on list B and voluntary con- into three categories with diff ering management trol, education, and research is the objective for objectives and requirements (Colorado Noxious species on list C. Currently, seven species from Weed Act of 1996 Title 35 Article 5.5). Weeds on Hovenweep NM are on List B and fi ve are on List list A are designated for full eradication wherever C; none are on List A (Table 3.4). they occur, while containment of infestations is

12 Annotated Checklist of Vascular Flora: Hovenweep National Monument Table 3.3. Plant species of conservation concern from Hovenweep National Monument. TNC State heritage Family Species Common name global Legal status status rank Asteraceae Haplopappus spinulosus spiny goldenweed G5T3? UTCDC: Watch, S2 (Compositae) var. paradoxus Boraginaceae Cryptantha osterhoutii Osterhout’s cryptanth G2G3 CO BLM: Sensitive CNHP: SOC, S2 (Oreocarya osterhoutii) Boraginaceae Cryptantha pterocarya var. wing-nut cryptanth G5T5 UTCDC: cycloptera Peripheral, S2 Cactaceae Opuntia phaeacantha var. large pricklypear G5T5 UTCDC: major* Peripheral, S2S3 Cucurbitaceae Cucurbita foetidissima* buffalo gourd G5 UTCDC: Peripheral, S3? Fabaceae Astragalus coltonii var. Moab milkvetch G4T3? CNHP: SOC, S2 (Leguminosae) moabensis Fabaceae Psorothamnus Thompson’s indigo- G3?T3? USFWS: 3C UTCDC: Watch, (Leguminosae) thompsoniae var. bush S3? thompsoniae Gentianaceae Swertia utahensis Utah swertia G4 CO BLM: Sensitive CNHP: SOC, S1 (Frasera paniculata) Liliaceae Calochortus fl exuosus sinuous mariposa G4 USFS Region 2: CNHP: SOC, S2 (Calochortaceae) Sensitive Linaceae Linum aristatum broom-fl ax G5 CNHP: SOC, SH (Mesynium aristatum)* Polygonaceae Eriogonum palmerianum Palmer’s wild G4 CNHP: SOC, S1 buckwheat Portulacaceae Portulaca halimoides dwarf purslane G5 CNHP: SOC, S1 Scrophulariaceae Penstemon breviculus shortest penstemon G3 UTCDC: Watch, S1? Scrophulariaceae Penstemon lentus var. Abajo penstemon G4T3T4 UTCDC: Watch, lentus S2? Scrophulariaceae Penstemon utahensis* Utah penstemon G4 CNHP: SOC, S2 Solanaceae Chamaesaracha coronopus false nightshade G5 UTCDC: Peripheral, S1 Zygophyllaceae Kallstroemia parvifl ora* orange kallstroemia G5 UTCDC: Peripheral, S1

This list is derived from the Colorado Natural Heritage Program (2007) and Utah Division of Wildlife Resources (1998).

Species with an * are reported from the monument; all others are confi rmed as present with a voucher specimen.

Codes: TNC (The Nature Conservancy) global rank assesses abundance and conservation priority on a scale of 1–5 (1 being extremely vulnerable and 5 being secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank, Q = taxonomic questions, U = unknown, and NR = not ranked. Under legal status, USFWS = U.S. Fish and Wildlife Service. C2 = Category 2 candidate (a former category used for taxa that might warrant being proposed for threatened or endangered status following additional research). 3C = Category 3 candidates (species dropped from consideration for listing). USFS = U.S. Forest Service. CO BLM = U.S. Bureau of Land Management, Colorado Offi ce. Under the column State heritage status, a comparable scoring system to that used in the “TNC global rank” column is used to measure state abundance (S) in Colorado and Utah. UTCDC = Utah Conservation Data Center (Utah Division of Wildlife Resources); these listings include conservation categories adopted by the state natural heritage program to prioritize endemic and rare plant taxa (UDWR 1998). Those categories are: Historical (presumed extinct or only known historically through their range), Rare (plants with rangewide viability concerns), Watch (regional endemics without rangewide viability concerns), Peripheral (rare or uncommon in Utah, but more common rangewide), Infrequent (plants occur infrequently over western U.S.), Taxonomic Problems (validity of species, subspecies, or variety has been questioned), Additional Data (species appear rare, but more information needed before being assigned to a different category). CNHP = Colorado Natural Heritage Program; in these listings, SOC denotes plants on the state “Species of Concern” list and Watch indicates species on the watch list that may ultimately warrant listing as species of concern (Colorado Natural Heritage Program 2007)

Chapter 3: Results 13 Table 3.4. Noxious weed species of Hovenweep National Monument. State weed Family Species Common name Park status list Asteraceae Carduus nutans musk thistle Present in CO Colo List B, UT Noxious Asteraceae Centaurea repens Russian knapweed Present in CO Colo List B, UT (Acroptilon repens) Noxious Asteraceae Arctium minus burdock Present in CO Colo List C Asteraceae Cirsium arvense Canada thistle Present in CO Colo List B, UT (Breea arvense) Noxious Chenopodiaceae Halogeton glomeratus halogeton Reported in UT, Colo List C CO Convolvulaceae Convolvulus arvensis fi eld bindweed Present in CO Colo List C, UT Noxious Geraniaceae Erodium cicutarium stork’s-bill Present in UT, Colo List B Observed in CO Poaceae Aegilops cylindrica jointed goatgrass Reported in CO Colo List B (Cylindropyrum cylindricum) Poaceae Bromus tectorum cheatgrass Present in UT, Colo List C (Anisantha tectorum) Observed in CO Scrophulariaceae Linaria dalmatica Dalmatian toadfl ax Present in CO Colo List B (L. genistifolia ssp. dalmatica) Scrophulariaceae Verbascum thapsus woolly mullein Present in CO Colo List C Tamaricaceae Tamarix chinensis fi ve-stamen tamarisk Present in UT, Colo List B (T. ramosissima) Observed in CO List derived from the Colorado Department of Agriculture (2009) and U.S. Department of Agriculture Natural Resources Conservation Service (2009).

14 Annotated Checklist of Vascular Flora: Hovenweep National Monument other 270 plant taxa may occur within the monu- 4 Discussion ment (Appendix B). Should all of these potential species be found, the documented fl ora of the Studies of the fl ora of Hovenweep National monument would increase by nearly 44%. Many Monument date to Richart’s 1941 checklist. Field of the “missing” potential or reported taxa con- work was most intensive in the early 1970s, in as- sist of cryptic species that fl ower late in the grow- sociation with archeological studies, and in the ing season, or occur in undersampled wetland or fi rst few years of the twenty-fi rst century, follow- weedy habitats. Confi rmation of reported species ing initiation of the NCPN inventory and moni- should remain a high priority, as some of these toring program. Presently, the monument fl ora taxa may ultimately prove to be erroneous. Addi- consists of 340 vascular plant species confi rmed tional studies would also be useful to better refi ne by voucher specimens (273) or reported in the the abundance and distribution of species of high literature (67) (Appendix A). Based on available management interest (especially rare plants and habitats and proximity to other populations, an- invasive non-natives).

Chapter 4: Discussion 15

Cronquist, A., A. H. Holmgren, N. H. Hol- 5 Literature Cited mgren, J. L. Reveal, and P. K. Holmgren, eds. 1977. Volume 6, The monocotyledons. Albee, B. J., L. M. Shultz, and S. Goodrich. 1988. Intermountain fl ora: Vascular plants of the Atlas of the vascular plants of Utah. +Occa- Intermountain West, U.S.A. Bronx, N.Y.: sional Publ. 7. Salt Lake City: Utah Museum New York Botanical Garden. 584 pp. of Natural History. 670 pp. ——. 1984. Volume 4, Subclass Asteridae (except Barkworth, M. E., K. M. Capels, S. Long, and M. Asteraceae). Intermountain fl ora: Vascular B. Piep, eds. 2003. Magnoliophyta: Com- plants of the Intermountain West, U.S.A. melinidae (in part): Poaceae, part 2. Flora of Bronx, N.Y.: New York Botanical Garden. North America north of Mexico, Volume 25. 573 pp. New York: Oxford University Press. 783 pp. Cronquist, A., N. H. Holmgren, and P. K. Hol- Barkworth, M. E., K. M. Capels, S. Long, L. K. mgren, eds. 1997. Volume 3, Part A, Subclass Anderton, and M. B. Piep, eds. 2007. Mag- Rosidae (except Fabales). Intermountain noliophyta: Commelinidae (in part): Poace- fl ora: Vascular plants of the Intermountain ae, part 1. Flora of North America north West, U.S.A. Bronx, N.Y.: New York Botani- of Mexico, Volume 24. New York: Oxford cal Garden. 446 pp. University Press. 911 pp. Dewey, S., and K. Andersen. 2005. An inventory Barneby, R. C. 1989. Volume 3, Part B. Fabales. of invasive non-native plants in Hovenweep In Cronquist, A., A. H. Holmgren, N. H. National Monument (2003): Final report. Holmgren, J. L. Reveal, and P. K. Holmgren, Utah State University, Logan Ut. Weed Sci- eds. Intermountain fl ora: Vascular plants of ence Research Project Report # SD0518a. the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 279 pp. Dorn, R. D. 2001. Vascular plants of Wyoming. Third edition. Cheyenne, Wy.: Mountain Chronic, H., and L. Chronic. 2004. Pages of West Publishing. 412 pp. stone: Geology of the Grand Canyon and plateau country national parks and monu- Evenden, A., M. Miller, M. Beer, E. Nance, S. ments. Second edition. Seattle, Wa.: The Daw, A. Wight, M. Estenson, and L. Cudlip. Mountaineers. 173 pp. 2002. Northern Colorado Plateau Vital Signs Network and Prototype Cluster plan Colorado Department of Agriculture (CDA). for natural resources monitoring: Phase 2009. Colorado noxious weed list. http:// I report, October 1, 2002 [two volumes]. wsprod.colostate.edu/cwis79/Noxious_ National Park Service, Northern Colorado weeds/noxious_weeds.cfm. Plateau Network, Moab, Ut. 138 pp + app. Colorado Natural Heritage Program. 2007. Fertig, W. 2006. Herbarium and fi eld studies of Tracked vascular plant species (8/16/2007 vascular plant fl ora of HOVE for National version). www.cnhp.colostate.edu/tracking/ Park Service Inventory and Monitoring pro- vascular.html. gram. Invesigator’s Annual Report, National Cronquist, A. 1994. Volume 5, Asterales. In Park Service. 2 pp. Cronquist, A., A. H. Holmgren, N. H. Fertig, W. 2007. Introduced and naturalized Holmgren, J. L. Reveal, and P. K. Holmgren, plants of Utah. Sego Lily 30(5):7–11. eds., Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: Flora of North America Editorial Committee. New York Botanical Garden. 496 pp. 1993. Pteridophytes and gymnosperms. Flora of North America north of Mexico, Cronquist, A., A. H. Holmgren, N. H. Holmgren, Volume 2. New York: Oxford University and J. L. Reveal, eds. 1972. Volume 1, Geo- Press. 475 pp. logical and botanical history of the region: Its plant geography and a glossary. The ——. 1997. Magnoliophyta: Magnoliidae and vascular cryptogams and the gymnosperms. Hamamelidae. Flora of North America Intermountain fl ora: Vascular plants of the north of Mexico, Volume 3. New York: Ox- Intermountain West, U.S.A. Bronx, N.Y.: ford University Press. 590 pp. New York Botanical Garden. 270 pp.

Chapter 5: Literature Cited 17 ——. 2000. Magnoliophyta: Alismatidae, Are- National Park Service. 2000. Management poli- cidae, Commelinidae (in part), and Zin- cies. Washington, D.C.: U.S. Department of giberidae. Flora of North America north Interior, National Park Service. Publication of Mexico, Volume 22. New York: Oxford #NPS D1416. 137 pp. University Press. 352 pp. Noss, R. F., and A. Y. Cooperrider. 1994. Saving ——. 2002a. Magnoliophyta: Commelinidae (in nature’s legacy: Protecting and restoring part): Cyperaceae. Flora of North America biodiversity. Washington, D.C.: Island Press. north of Mexico, Volume 23. New York: 416 pp. Oxford University Press. 608 pp. Rejmanek, M., and J.M. Randall. 1994. Invasive ——. 2002b. Magnoliophyta: Liliidae: Liliales and alien plants in California: 1993 summary Orchidales. Flora of North America north and comparison with other areas in North of Mexico, Volume 26. New York: Oxford America. Madrono 41(3):161–177. University Press. 723 pp. Richart, R. S. 1941. Fauna and fl ora of Hoven- ——. 2003. Magnoliophyta: Caryophyllidae, part weep and Yucca House National Monu- 1. Flora of North America north of Mexico. ment. Southwest National Monuments Volume 4. New York: Oxford University Supplement for June 1941. Press. 559 pp. Schelz, C. 2002. General fl oristic survey of Hov- ——. 2005. Magnoliophyta: Caryophyllidae, part enweep National Monument: Annual report, 2. Flora of North America north of Mexico. 2002. National Park Service Southeast Utah Volume 5. New York: Oxford University Group, Moab, Ut. 77 pp. Press. 656 pp. Schelz, C. 2003. General fl oristic survey of Hov- ——. 2006a. Magnoliophyta: Asteridae, part 6: enweep National Monument: Annual report, Asteraceae, part 1. Flora of North America 2003. National Park Service Southeast Utah north of Mexico. Volume 19. New York: Group, Moab, Ut. 77 pp. Oxford University Press. 579 pp. Schelz, C., and M. Moran. 2003. Plants, Hoven- ——. 2006b. Magnoliophyta: Asteridae, part 7: weep National Monument. National Park Asteraceae, part 2. Flora of North America Service, Southeast Utah Group, Moab, Ut. north of Mexico. Volume 20. New York: 8 pp. Oxford University Press. 666 pp. ——. 2007. Southeast Utah Group threatened, ——. 2006c. Magnoliophyta: Asteridae, part 8: endangered and species of concern. Nation- Asteraceae, part 3. Flora of North America al Park Service, Moab, Ut. 3 pp. north of Mexico. Volume 21. New York: Shultz, L. M., R. D. Ramsey, and W. Lindquist. Oxford University Press. 616 pp. 2006. Revised atlas of Utah plants. College Holmgren, N. H., P. K. Holmgren, and A. Cron- of Natural Resources, Utah State University, quist. 2005. Volume 2, Part B, Subclass Dille- Logan, Ut. http://earth.gis.usu.edu/plants. niidae. Intermountain fl ora: Vascular plants Springer, A. E., L. E. Stevens, and R. Harms. of the Intermountain West, U.S.A. Bronx, 2006. Inventory and classifi cation of selected N.Y.: New York Botanical Garden. 488 pp. National Park Service springs on the Colo- Kartesz, J. T. 2003. Synthesis of the North Ameri- rado Plateau. Northern Arizona University, can fl ora, Version 2.0. Flagstaff , Az. NPS Cooperative Agreement #CA 1200-99-009; Task # NAU-118. Litzinger, W. J., and J. C. Winter. 1976. Annotated checklist of the vascular plants of Cajon U.S. Department of Agriculture Natural Re- Mesa, Colorado–Utah. Hovenweep 1975: sources Conservation Service (USDA- Archeological Report No. 2. Archeological NRCS). 2009. Utah state-listed noxious Report. San Jose State University, San Jose, weeds. http://plants.usda.gov/java/noxious?r Ca. ptType=State&statefi ps=49. Margules, C. R., and R. L. Pressey. 2000. Sys- Utah Division of Wildlife Resources (UDWR). tematic conservation planning. Nature 1998. Inventory of sensitive species and 405:243–253. ecosystems in Utah. Endemic and rare plants of Utah: An overview of their distribution

18 Annotated Checklist of Vascular Flora: Hovenweep National Monument and status. Report prepared for the Utah Weber, W. A., and R. C. Wittmann. 2001. Colo- Reclamation Mitigation and Conservation rado fl ora: Western Slope. Third edition. Commission and U.S. Department of the Boulder, Co.: University Press of Colorado. Interior. 566 pp + app. 488 pp. Von Loh, J., G. Wakefi eld, A. Wight, A. Evenden, Welsh, S. L., N. D. Atwood, S. Goodrich, and L. and J. Coles. 2008. Vegetation classifi cation C. Higgins. 2003. A Utah fl ora. Third edition, and mapping project report, Hovenweep revised. Provo, Ut: Brigham Young Univer- National Monument. Natural Resource sity. 912 pp. Technical Report NPS/NCPN/NRTR— 2008/092. National Park Service, Fort Col- lins, Co.

Chapter 5: Literature Cited 19

Appendices Synonyms/Taxonomic notes (Appendices A, B, and C) Pertinent synonyms are included for species with alterna- tive scientifi c names in other recent state and regional fl or- The following appendices contain annotated checklists of as (Barkworth et al. 2003, 2007; Barneby 1989; Cronquist the vascular fl ora of Hovenweep National Monument. 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn 2001; Appendix A lists all species that have been confi rmed for Flora of North America Editorial Committee 1993, 1997, the fl ora with an authenticated voucher specimen (includ- 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; Hol- ing extant and historical records) or that have been reli- mgren et al. 2005). Only species not otherwise listed in the ably reported for the monument in the literature (Dewey Utah or Colorado species name columns are included. No and Andersen 2005; Schelz and Moran 2003; Springer et eff ort has been made to identify additional synonyms from al. 2006; Von Loh et al. 2008). Vouchers were examined monographs and fl oras preceding the early 1970s. from the HOVE herbarium (housed in the National Park Common name (Appendices A, B, C, and D) Service’s Southeast Utah Group offi ce in Moab, Utah), and collections from Brigham Young University (BRY), Common names are derived primarily from Welsh et al. Utah State University (USU), the University of Wyoming (2003) and the Intermountain Flora (Barneby 1989; (RM), and the digital museum database of the New York Cronquist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Botanical Garden (NY). Species entries in Appendix A are Holmgren et al. 2005). organized alphabetically by family and genus. Life form (Appendices A, B, C, and D) Appendix B includes species that may potentially occur Life form is based on the stature and growth habit of typi- within Hovenweep NM based on the proximity of the cal members of a species at reproductive maturity. Trees monument to other populations or the presence of suit- are woody perennials with a single stem or trunk and are able habitat. typically over 3.5 m tall. Shrubs are woody perennials with one to many trunks arising from near the base of the Appendix C lists species that have been previously report- plant and are usually less than 3.5 m tall. Perennial Forbs ed for Hovenweep NM but are now considered errone- (PerF) are broad-leaved dicots or monocots with non- ous because they are based on misidentifi ed specimens or woody stems (at least above ground level) that live for mul- questionable literature records. tiple years. Annual Forbs (AnnF) are broad-leaved dicots or monocots with non-woody stems that complete their Appendix D contains the same confi rmed, historical, and life cycle (mature, fl ower, and die) in one year. Perennial reported taxa as in Appendix A, but organized by life form Graminoids (PerG) are linear-leaved, grass-like monocots rather than by family in order to better meet the needs of that live for multiple years. Annual Graminoids (AnnG) NPS interpretive specialists. are linear-leaved, grass-like monocots that complete their life cycle (mature, fl ower, and die) in one year. Ferns and The codes and references for each fi eld in the appropriate fern-allies (Fern) are non-fl owering vascular plants that appendices are summarized below. reproduce by spores. Family (Appendices A, B, C, and D) Range UT/CO (Appendices A, B, and C) Family concepts and nomenclature follow A Utah Flora, Range represents the distribution of a species within Utah third edition (Welsh et al. 2003). Alternative family names or Colorado and each state’s contribution to the plant’s and treatments from other regional fl oras (Dorn 2001; overall global range. Usually these values are the same for Flora of North America Editorial Committee 1993, 2002b; both states, but in some cases a species might be wide- Holmgren et al. 2005; Weber and Wittman 2001) are listed spread in one state, but considered sparse, peripheral, or in parentheses. introduced in the other. Disjunct (Disj) taxa have their Species name (Appendices A, B, C, and D) Utah or Colorado distribution separated from the main, contiguous portion of their range by a gap of more than Because Hovenweep NM straddles Utah and Colorado, 800 km (ca 500 miles). Introduced (Intro) taxa are those two sets of species names are provided using the scientifi c that are not native to Utah, Colorado, or North America names and taxonomic concepts of the most recent fl ora of but have become naturalized (breeding on their own each state (Weber and Wittmann 2001; Welsh et al. 2003). without human assistance). Local Endemics (LocEn) are Scientifi c names are listed in alphabetical rather than phy- taxa whose entire global range is restricted to an area of logenetic order and according to the Utah names of Welsh less than 16,500 km2 (ca 6,370 mi2, or 1° of latitude × 2° et al. (2003). of longitude). Peripheral (Periph) taxa have a widespread global distribution but occur at the margin of their contig-

Chapter 6: Appendices 21 uous range in Utah or Colorado and occupy less than 5% CO (Appendices A, C) of the state’s area (usually only within a few miles of the This column indicates whether a given species has been state border). Regional Endemics (RegEn) are taxa with a documented in the Colorado portion of Hovenweep NM 2 global range of 16,500–250,000 km (an area about the size with a voucher (X) or by an observation (O). Species that of the state of Wyoming). Sparse taxa occur widely across are reported for the monument without the state indicated Utah, Colorado, or North America but their range within are marked by a ?. a state is small and patchy, with populations restricted to specialized or uncommon habitats. Widespread (Wide) Source (Appendix A) taxa have global ranges exceeding 250,000 km2 and occur Sources provide the evidence for the assigned park sta- over at least 10% of each state. tus. In the case of Present and Historical taxa, sources are Park status (Appendices A, C, and D) herbarium records (with collector, collection number, and repository) that were corroborated during this study. For Park status identifi es whether a species is Present (Pres), reported taxa, the source is a published or unpublished Historical (Hist), or Reported (Rep) for Hovenweep NM. reference citing the species within the monument. Speci- Species are considered Present if a confi rmed voucher or men repositories include the Hovenweep NM herbarium documented observation has been made within monu- (HOVE), Brigham Young University (BRY), Utah State ment boundaries since 1970. Historical species have been University (USU), New York Botanical Garden (NY), and confi rmed with a voucher from the monument but have the University of Wyoming’s Rocky Mountain Herbarium not been relocated since before 1970. Reported species (RM). are listed for the monument in the literature (Dewey and Andersen 2005; Schelz and Moran 2003; Springer et al. Year doc. (Appendix A) 2006; Von Loh et al. 2008), but have not been corrobo- Year documented indicates the fi rst year in which a spe- rated with a voucher specimen. Species that may poten- cies was confi rmed or reported to occur in Hovenweep tially occur in the monument are listed in Appendix B. Re- NM. This year does not necessarily correspond with the jected taxa (listed in Appendix C) have either been falsely date of the voucher specimen cited under Source. “NA” reported (FalsRep) in the literature based on misidenti- indicates that the fi rst year is not known (used mostly for fi ed herbarium specimens or are considered Questionable “reported” taxa). (Ques?) because monument reports are from well outside their known or expected range. Comments (Appendices A, B, C, and D) Pop. size (Appendix A) This column includes additional comments on the status of a particular species in the monument, the place of ori- Population size refers to the abundance and frequency gin for introduced species, legal status, and other data. with which a species occurs within Hovenweep NM. Abundance data are derived from Dewey and Andersen Nativity (Appendix D) (2005), Von Loh et al. (2008), and personal observations. Nativity indicates whether a species is native to Utah and Abundant (Abun) taxa have large populations, have a Colorado or Introduced (Intro) based on Fertig (2007). broad ecological amplitude (often being dominant in one- to-several vegetation types), or occur extensively across Flower time (Appendix D) the monument. Common (Com) taxa have large local populations but may be restricted to a single vegetation Flower time refers to the months in which a species is most type or do not occur across the entire monument. Un- likely to be in fl ower. NA indicates a species that is a non- common (Unc) taxa have low-to-medium-sized popula- fl owering fern or gymnosperm. tions, typically have a narrow ecological amplitude (or at Habitat (Appendix D) least contribute relatively little to total vegetative cover), or have a small range across the monument. Rare taxa usu- The distribution of species across generalized habitats in ally have low population numbers and are restricted to a Hovenweep NM is indicated by an “x” in the columns single location within the monument. A “?” indicates that PinJun through Wet and is derived from the Hovenweep population size is unknown. NM vegetation classifi cation (Von Loh et al. 2008; Welsh et al. 2003). PinJun = Pinyon-Juniper woodlands/rimrock UT (Appendices A, C) (including pockets of sandy soils and sagebrush). PonP- This column indicates whether a given species has been DF = Ponderosa Pine-Douglas Fir woodlands. Wet = Wet- documented in the Utah portion of Hovenweep NM with lands (including hanging gardens). Additional comments a voucher (X) or by an observation (O). Species that are are included for species that are restricted to a particular reported for the monument without the state indicated are soil/geologic type or are found mostly in disturbed sites. marked by a ?.

22 Annotated Checklist of Vascular Flora: Hovenweep National Monument Central America is Helper, is Helper, Carbon Co., UT (Trelease s.n. MO) Comments record for Colorado? Central America c name with 2001 1972 1973 2001 2002 1975 Native to NA locality Type Year Year doc. 1975 State NA NA NA Native to 1975 (HOVE) 20020912.17 (HOVE) 20030611.02 (HOVE) 20010501 T (HOVE) 20020403.02 (HOVE) 20010907b (HOVE) Moran (2003) Source (HOVE) Moran (2003) Moran (2003) Moran (2003) (HOVE) Pop. size UT CO Pres X OPres Unc X Schelz 2103 X Com Schelz Pres X XPres Unc Schelz Pres X Rare X Schelz Rare Schelz Pres X X Unc Schelz Rep O ? Schelz and Pres X X Unc Schelz 2124 Park stat. Rep O ? Schelz and RepRep O ? ? ? Schelz and ? Schelz and Pres X X Unc Litzinger 631 Wide Wide RegEn Periph Wide Intro Wide RegEn Range UT/CO Wide Wide Intro Wide PerF Wide/ PerF Wide/ Shrub RegEn/ AnnF Intro/ Shrub Wide/ Shrub RegEn/ Life form Shrub Wide/ AnnF Wide/ AnnF Intro/ Fendler’s Fendler’s barberry squawbush Shrub Wide/ squawbush Shrub Wide/ spider milkweed pallid milkweed redroot pigweed Harriman’s Harriman’s yucca narrowleaved yucca Common name western poison ivy prostrate pigweed blood amaranth datil yucca Shrub Wide/

var. var. var. angustissima R. trilobata simplicifolia R. trilobata trilobata Synonyms/ notes Taxonomic Rhus radicans, T. Rhus radicans, T. radicans A. graecizans baccata Y. angustissima Y. var. simplicifolia exus trilobata trilobata asperula Rhus aromatica ssp. Toxicodendron Toxicodendron rydbergii Asclepias cryptoceras Rhus aromatica ssp. forma Asclepias asperula ssp. Amaranthus retrofl Amaranthus blitoides Yucca harrimaniae Yucca Yucca Yucca angustissima Amaranthus cruentus

Species name var. exus harrimaniae trilobata simplicifolia asperula Rhus aromatica var. Toxicodendron rydbergii Asclepias cryptoceras cryptoceras Berberis fendleri Berberis fendleri Amaranthus blitoides Yucca baccataYucca baccata Yucca baccata Y. Utah Yucca angustissima Colorado Rhus aromatica var. Asclepias asperula var. Amaranthus retrofl Amaranthus cruentus Yucca harrimaniae Yucca var. vascular plant taxa of Hovenweep National Monument, organized by family and scientifi rmed, historical, and reported Anacardiaceae Anacardiaceae Asclepiadaceae Berberidaceae Anacardiaceae Amaranthaceae Agavaceae Agavaceae Family Asclepiadaceae Amaranthaceae Amaranthaceae Agavaceae Appendix A. Confi taxonomic and biogeographic notes.

Chapter 6: Appendices 23

eld Co., eld Comments is Monument Park [Colorado National Monument], near Grand Junction, Mesa Co., CO (Osterhout 6138 RM). CO BLM: Sensitive is Paria Canyon, Garfi UT (Jones 5297p POM) of var. of var. echinoides 1975 1973 NA Year Year doc. 2001 2001 locality Type NA 1975 1973 1972 locality Type c name with taxonomic and (HOVE) (HOVE) Moran (2003) Source (HOVE) 20011005 J (HOVE) Moran (2003) (HOVE) 20020503.15 (HOVE) Oppelt 1047 (HOVE) Pop. size UT CO PresPres X X Unc XRep Schelz 2085 Unc ? Litzinger 608 ? ? Schelz and Park stat. PresPres X X Rare X Schelz 2133 Unc Schelz Rep O OPres ? X X Schelz and Unc Schelz 1962 Pres X Unc Schelz Pres X Unc Colyer & Wide Wide Wide Range UT/CO Wide RegEn Wide Wide Periph RegEn AnnF Wide/ AnnF Wide/ Life form AnnF Wide/ PerF RegEn/ PerF Wide/ PerF Wide/ AnnF Periph/ PerF RegEn/ thick-sepaled cryptanth Oregon-grape Shrub Wide/ sand dune cryptanth Common name slender cryptanth Osterhout’s cryptanth plateau yellow cryptanth yellow-eye cryptanth wing-nut cryptanth plateau cryptanth

& Synonyms/ notes Taxonomic Includes vars. fulvocanescens echinoides

var. ava avoculata elachantha Cryptantha crassisepala Cryptantha fendleri Cryptantha gracilis Oreocarya osterhoutii Oreocarya fl Oreocarya fulvocanescens Cryptantha pterocarya

Species name avafl Oreocarya var. var. avoculata Cryptantha crassisepala elachantha Cryptantha fendleri Cryptantha fl UtahMahonia repens Mahonia repens Berberis repens Colorado Cryptantha pterocarya Cryptantha gracilis Cryptantha osterhoutii Cryptantha fl Cryptantha fulvocanescens cycloptera rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Boraginaceae Boraginaceae Boraginaceae Berberidaceae Family Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Appendix A. Confi biogeographic notes, cont.

24 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments Year Year doc. 1975 NA 2002 NA 1973 1974 2001 2003 2006 2006 c name with taxonomic and Source (HOVE) Moran (2003) 20020912.21 (HOVE) Moran (2003) (HOVE) (HOVE) 20011005 G (HOVE) (HOVE) (HOVE) (HOVE) Pop. size UT CO Park stat. Pres ORep X O Unc Litzinger 656 ? Schelz and PresRep X Unc O Schelz ? Schelz and Pres X XPres Com Schmidt 59 Pres X UncPres X Schmidt 65 Rare Schelz X Rare Schelz 2109 Pres X O Unc Fertig 22613 Pres X Unc Fertig 22618 Range UT/CO Wide NA Wide NA Wide Wide Wide RegEn Wide Wide Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ PerF RegEn/ PerF Wide/ Common name common pricklypear Utah pricklypear brittle pricklypear large pricklypear cupseed stickseed showy stoneseed western stoneseed spindle-rooted bluebell claretcup PerF Wide/ Arizona pincushion .

var. Vars. Vars. var. (2003) - (2003) (2003). (2003) Vars. Vars. erinacea Flora of North Flora of North Synonyms/ notes Taxonomic Var. not Var. recognized in Flora of North America considered a few- spined variant of O. polyacantha not Vars. recognized in Flora of North America L. texana, redowskii cupulata Includes var. Includes var. inermis. not recognized in America var. Escobaria vivipara arizonica. not recognized in America brachyartha Lappula marginata Echinocereus triglochidiatus Opuntia erinacea O. polyacantha var. Lithospermum incisum Lithospermum ruderale Mertensia oblongifolia Coryphantha vivipara

var. var. Species name var.

var. utahensis erinacea UtahLappula occidentalis cupulata Colorado Echinocereus triglochidiatus melanacanthus Opuntia erinacea var. Opuntia fragilis Opuntia fragilis Opuntia phaeacantha major Lithospermum incisum Lithospermum ruderale Mertensia fusiformis Coryphantha vivipara arizonica Opuntia erinacea var. rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Boraginaceae Family Cactaceae Cactaceae Cactaceae Cactaceae Boraginaceae Boraginaceae Boraginaceae Cactaceae Cactaceae Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 25 is near Emery, Emery Co., UT (Jones 5443 RM) Comments 2002 1975 1975 locality Type Year Year doc. 1973 1974 1972 1972 2006 1975 NA 2003 eld 77 eld c name with taxonomic and (HOVE) (HOVE) (HOVE) Source 20010503.14 (HOVE) 1049/1050 (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) Moran (2003) al. (i2008) & Schelz & Moran (2003) Pop. size UT CO Pres XPres X Unc Schelz 2115 Rare Litzinger 678 Pres X Rare Litzinger 679 Park stat. Pres X X UncPres Schelz X XPres Unc X Colyer Pres X X Com Redfi X Com Litzinger 752 Pres X Com Fertig 22614 Pres O X Com Litzinger 656 Rep O ? Schelz and Rep O O Unc Loh et Von NA RegEn RegEn Range UT/CO Wide Wide Wide Wide Wide Wide RegEn Wide Shrub RegEn/ Life form PerF Wide/ Shrub Wide/ PerF Periph/ PerF Wide/ Shrub RegEn/ PerF Wide/ y AnnF Wide/ ower shhook basin saltbush Shrub Wide/ stalked orach AnnF RegEn/ Castle Valley Castle Valley saltbush Common name Eastwood’s Eastwood’s sandwort sleepy catchfl four-wing four-wing saltbush shadscale Shrub Wide/ berry pricklypear plains pricklypear Whipple’s cholla small-fl fi cactus

var. var. var.

S. var.

& & (2003) (2003) S. orus. spina A. tridentata robusta caput-medusae Synonyms/ notes Taxonomic Eremogone eastwoodiae eastwoodiae, A. eastwoodiae occidentalis Vars. not Vars. recognized in Flora of North America trichophora Flora of North America includes vars. rufi S. parvifl Includes contortus terrae-canyonae

var. Atriplex saccaria A. saccaria Eremogone fendleri Atriplex canescens A. canescens Atriplex gardneri A. cuneata Opuntia phaeacantha Atriplex confertifolia Opuntia polyacantha whipplei Sclerocactus whipplei intermedius

var. var.

Species name var. utahensis saccaria eastwoodiae canescens cuneata Atriplex gardneri var. Atriplex saccaria var. UtahOpuntia phaeacantha phaeacantha Colorado Arenaria fendleri var. Silene antirrhina Silene antirrhina Atriplex canescens var. Atriplex confertifolia Atriplex gardneri var. Opuntia polyacantha polyacantha Opuntia whipplei Cylindropuntia Sclerocactus whipplei roseus rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Chenopodiaceae Chenopodiaceae Cactaceae Family Caryophyllaceae (Alsinaceae) Caryophyllaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Cactaceae Cactaceae Cactaceae Appendix A. Confi biogeographic notes, cont.

26 Annotated Checklist of Vascular Flora: Hovenweep National Monument Eurasia Eurasia Comments noxious weed (List C). Native to Eurasia Asia 2001 Native to 2001 Native to Year Year doc. 1973 NA CO state 1975 2001 2001 1974 1975 1975 Native to NA 2003 c name with taxonomic and 20010824 VA 20010824 VA (HOVE) 20010824 G (HOVE) Source (HOVE) Moran (2003) (HOVE) (HOVE) 20020912.14 (HOVE) (HOVE) (HOVE) (HOVE) Moran (2003) (HOVE) Pop. size UT CO Pres X Unc Schelz Pres X Unc Schelz Park stat. Pres XRep X ? Unc Litzinger 508 ? ? Schelz and Pres X OPres Rare Litzinger 754 X Unc Schelz 2118 Pres X Unc Schelz Pres X O Com Schmidt 81 Pres O X Rare Litzinger 650 Pres X O Unc Litzinger 800 Rep O ? Schelz and Pres X Unc Schelz 2122 Intro Intro Range UT/CO Wide Intro Wide Wide Wide Wide Wide Intro Wide Wide AnnF Intro/ Life form AnnF Wide/ AnnF Wide/ AnnF Wide/ AnnF Sparse/ AnnF Wide/ lambsquarter AnnF Intro/ summer- cypress Common name halogeton AnnF Intro/ oakleaf goosefoot spiny hopsage Shrub Wide/ narrow-leaf goosefoot pitseed goosefoot greasewood Shrub Wide/ winterfat Shrub Wide/ Russian thistle AnnF Intro/ desert goosefoot silvery goosefoot

var.

var. var. (2003). . Vars. not . Vars. Synonyms/ notes Taxonomic C. salinum C. berlandieri zschackei Ceratoides lanata, Eurotia lanata recognized in Flora of North America S. pestifer C. pratericola C. incanum incanum, C. incanum occidentale Chenopodium album Chenopodium glaucum Chenopodium berlandieri Krascheninnikovia lanata Halogeton glomeratus Chenopodium leptophyllum Chenopodium desiccatum Sarcobatus vermiculatus Chenopodium incanum

Species name

album lanata var. var. var. var. var. Chenopodium album Chenopodium album UtahBassia scoparia Bassia sieversiana Kochia scoparia Colorado salinum berlandieri Krascheninnikovia lanata Halogeton glomeratus Chenopodium leptophyllum Grayia spinosa Atriplex grayi A. spinosa Chenopodium desiccatum Sarcobatus vermiculatus Salsola tragus Salsola australis S. kali, iberica, Chenopodium fremontii incanum Chenopodium glaucum rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Chenopodiaceae Chenopodiaceae Chenopodiaceae Family Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae (Sarcobataceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 27 Eurasia Comments noxious weed (List C). Native to Eurasia 1975 1974 Native to Year Year doc. 1975 NA 2002 2002 CO state NA 2001 2003 1975 NA 1975 c name with taxonomic and (HOVE) (HOVE) Source (HOVE) Moran (2003) 20020912.18 (HOVE) 20020912.25 (HOVE) Moran (2003) 20010911 A (HOVE) al. (2008) (HOVE) Moran (2003) (HOVE) Pop. size UT CO Pres X Unc Litzinger 755 Pres X Rare Litzinger s.n. Park stat. Pres XRep X ? Unc Litzinger 799 ?Pres ? O X Schelz and Unc Schelz Pres X Unc Schelz Rep ? ? ? Schelz and Pres X Unc Schelz Rep O Unc Loh et Von Pres X X Com Litzinger 811 Rep ? ? ? Schelz and Pres X Rare Litzinger 810 Wide Intro Range UT/CO Wide Wide Wide Intro Wide Wide Wide Wide Wide NA AnnF Wide / Life form Shrub Wide/ Shrub Wide/ Shrub Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Torrey’s Torrey’s seepweed spinach AnnF Intro / Common name fringed sagebrush Bigelow’s Bigelow’s sagebrush sand sagebrush burdock PerF Intro/ common yarrow western spiderwort Louisiana wormwood common pussytoes bur ragweed AnnF Wide/ Louisiana wormwood var. var. vars. & (2006) Flora of North fruticosa Synonyms/ notes Taxonomic occidentalis; not recognized in America Includes vars. occidentalis scopulorum Franseria acanthicarpa

ssp. ssp. lifolius Suaeda moquinii S. nigra, Tradescantia Tradescantia occidentalis Artemisia bigelovii fi Ambrosia acanthicarpa Achillea lanulosa A. millefolium Artemisia ludoviciana albula Antennaria parvifolia Artemisia ludoviciana mexicana

Species name var. var. var. lifolia Oligosporus torreyana Suaeda torreyana var. Tradescantia occidentalis UtahSpinacia oleracea Spinacia oleracea Colorado Artemisia bigelovii Artemisia fi Artemisia frigida Artemisia frigida Artemisia ludoviciana Arctium minus Arctium minus Ambrosia acanthicarpa lanulosa Achillea millefolium Artemisia ludoviciana albula Antennaria parvifolia mexicana rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Chenopodiaceae Commelinaceae Chenopodiaceae Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi biogeographic notes, cont.

28 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments noxious weed. CO state noxious weed (List B). Native to Eurasia Year Year doc. 2002 1975 2002 1975 1972 1975 2001 1973 2003 Utah state eld 5 eld c name with taxonomic and Source 20020607.16 (HOVE) 20020924.3 (HOVE) 20020607.06 (HOVE) 20010403 H (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) Pop. size UT CO Park stat. PresPres X Unc X Schelz X Unc Schelz Pres O X Unc Schelz Pres X OPres Unc X Schelz OPres Com X Redfi X Com Litzinger 829 Pres X X Unc Schelz 2132 Pres XPres Unc Schmidt 33 X Unc Fertig 22622 Range UT/CO Wide Wide Wide Wide Wide Wide Wide Wide Intro Life form Shrub Wide/ Shrub Wide/ Shrub Wide/ Shrub Wide/ PerF Wide/ Shrub Wide/ Shrub Wide/ Common name California brickellbush basin big sagebrush Wyoming big sagebrush black sagebrush budsage Shrub Wide/ arrowleaf balsamroot rough brickellbush Mohave brickellbush musk thistle PerF Intro/

var. (2006) vars. not A. arbuscula Synonyms/ notes Taxonomic nova; recognized in Flora of North America

ssp. ssp. var. macrolepis Seriphidium novum Brickellia californica tridentatum wyomingense Picrothamnus desertorum Balsamorhiza sagittata Brickellia microphylla Seriphidium tridentatum Seriphidium tridentatum scabra var. Brickellia oblongifolia

var. Species name var. var. var. nova UtahArtemisia nova var. Colorado Brickellia californica Brickellia microphylla Artemisia spinescens Artemisia tridentata Artemisia tridentata Balsamorhiza sagittata scabra tridentata wyomingensis Brickellia oblongifolia Carduus nutans Carduus nutans linifolia rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 29 C. orus noxious weed. CO state noxious weed (List B). Native to Eurasia Comments specimens tend towards viscidifl 2006 1972 1973 2001 UT state Year Year doc. 2002 2001 HOVE 1973 1975 1972 eld 25 eld c name with taxonomic and (HOVE) 20020607.2 (HOVE) (HOVE) 20010824 C (HOVE) Source 2002912.31 (HOVE) 20010830 B (HOVE) (HOVE) 20010824 S (HOVE) (HOVE) Pop. size UT CO Pres X Rare Fertig 22617 Pres X X Unc Schelz Pres X Unc Litzinger 550 Pres X Unc Schelz Park stat. Pres O XPres Unc X Schelz Rare Schelz Pres X X Com Schmidt 41 Pres X X Unc Schelz Pres X X Unc Redfi Wide Wide Wide Intro Range UT/CO Periph Wide Wide Wide Wide PerF Sparse/ PerF Wide/ AnnF Wide/ PerF Intro/ Life form Shrub Wide/ Shrub Wide/ Shrub Wide/ Shrub Wide/ hairy golden- aster dwarf rabbitbrush hispid golden- aster stevia dusty- maiden Russian knapweed Common name Greene’s Greene’s rabbitbrush rose-heath PerF Wide/ graystem rabbitbrush glabrate rabbitbrush

var.

var. var. var. var. includes hispida hololeucus, C. villosa hispida, H. villosa var. Synonyms/ notes Taxonomic Aster arenosus C. nauseosus var. Ericameria nauseosa hololeuca Ericameria nauseosa graveolens, E. nauseosa glabrata; C. nauseosus oreophilus

var. var. villosa minor var. var. Heterotheca villosa Chrysothamnus depressus Heterotheca villosa Chaenactis stevioides Leucelene ericoides Chrysothamnus nauseosus Chrysothamnus greenei gnaphalodes Chrysothamnus nauseosus graveolens

Species name var. var. villosa minor Chrysothamnus depressus Chrysopsis villosa var. Chrysothamnus nauseosus Chaenactis stevioides Chaetopappa ericoides UtahCentaurea repens Acroptilon repens Colorado gnaphalodes Chrysothamnus greenei Chrysopsis villosa var. Chrysothamnus nauseosus graveolens rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi biogeographic notes, cont.

30 Annotated Checklist of Vascular Flora: Hovenweep National Monument eld Co., eld Comments is Bromide Pass, Henry Mtns, Garfi UT (Jones 5695bh US) nox-ious weed. CO state noxious weed (List B). Native to Eurasia Year Year doc. 2002 locality Type 2001 Utah state NA NA NA 2003 NA 2002 1973 c name with taxonomic and Source 20020912.29 (HOVE) 20020912.41 (HOVE) Moran (2003) Moran (2003) Moran (2003) (HOVE) Moran (2003) 20020912.16 (HOVE) (HOVE) Pop. size UT CO Park stat. Pres X Unc Schelz Pres X Unc Schelz Rep O O ?Rep Schelz and Rep O ? O O Schelz and ?Pres Schelz and X Unc Schelz 2087 Rep O O ? Schelz and Pres X Rare Schelz Pres X Com Litzinger 327 Range UT/CO RegEn Intro Wide Wide Wide Wide RegEn Wide Wide Life form PerF RegEn/ PerF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ Shrub RegEn/ Shrub Wide/ Shrub Wide/ Common name Caineville thistle Canada thistle PerF Intro/ New Mexico thistle wavy-leaved thistle Canadian horseweed gray hawksbeard rush rabbitbrush whitestem rabbitbrush slenderleaf rabbitbrush var. var.

by

& var. var. (2006) (2006) orus albicaulis, Synonyms/ notes Taxonomic bipinnatum integrifolium Vars. not Vars. recognized in Flora of North America Vars. not Vars. recognized in Flora of North America Ericameria nauseosa juncea C. nauseosus var. Ericameria nauseosa speciosa Included in var. Included in var. viscidifl some authors Includes vars. horridum

var. var. var. orus Cirsium calcareum C. arizonicum Conyza canadensis Cirsium neomexicanum Chrysothamnus nauseosus Cirsium undulatum intermedia junceus Chrysothamnus nauseosus speciosus stenophyllus Chrysothamnus viscidifl

Species name var. var. var. var. orus

calcareum glabrata UtahChrysothamnus nauseosus Colorado Cirsium neomexicanum var. neomexicanum Cirsium calcareum var. junceus Cirsium undulatum undulatum Conyza canadensis var. Crepis intermedia Psilochenia Chrysothamnus nauseosus speciosus Chrysothamnus viscidifl stenophyllus Cirsium arvense Breea arvensis rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 31 is Kanab, Kane Co., UT (Thompson s.n. GH) Comments 2001 1973 2001 1975 NA locality Type 2003 1972 Year Year doc. 1975 2002 1973 1975 c name with taxonomic and 20011005 K (HOVE) (HOVE) (HOVE) (HOVE) Moran (2003) (HOVE) 20010824 T (HOVE) Source (HOVE) 20020503.19 (HOVE) 20010430 K (HOVE) (HOVE) Pop. size UT CO PresPres X Unc XPres Schelz O Unc Schmidt 15 Pres X Unc Schelz 295 XRep Unc O Litzinger 594 O ? Schelz and PresPres X X Unc X Schelz 2119 Abun Schelz Park stat. Pres X Unc Litzinger 324 Pres O X Unc Schelz Pres X Unc Schelz Pres X X Unc Litzinger 823 Wide Wide Wide Wide RegEn Wide Wide Range UT/CO Wide Wide RegEn Wide PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Shrub Wide/ Life form AnnF Wide/ PerF Wide/ PerF RegEn/ AnnF Wide/ ower ower ower eabane western hawksbeard pretty daisy AnnFspreading Wide/ daisy Navajo fl Utah daisy PerF RegEn/ Hopi blanketfl broom snakeweed Common name prairie sunfl thrift goldenweed spiny goldenweed common sunfl

var.

var. (2006) concinnus; bellidiastrum E. concinnus var. includes var. subglaber E. bellidiastrum var. Synonyms/ notes Taxonomic petiolaris armerioides S. armerioides Xanthisma spinulosum paradoxum Vars. not Vars. recognized in Flora of North America Includes var.

var. da da Erigeron concinnus Psilochenia occidentalis Erigeron bellidiastrum Erigeron divergens Erigeron utahensis Gutierrezia sarothrae Stenotus armerioides Gaillardia pinnatifi Machaeranthera pinnatifi paradoxa Helianthus annuus Helianthus petiolaris

Species name var.

var. var. var. var. ssp. var. da concinnus occidentalis Gaillardia pinnatifi Erigeron bellidiastrum Erigeron divergens divergens Erigeron pumilus var. Erigeron utahensis utahensis Gutierrezia sarothrae Haplopappus armerioides armerioides UtahCrepis occidentalis var. Colorado Haplopappus spinulosus paradoxus Helianthus annuus lenticularis Helianthus petiolaris fallax rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi biogeographic notes, cont.

32 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments Europe NA 2001 1975 2006 Year Year doc. NA 1973 2002 1973 Native to NA 1972 2002 c name with taxonomic and Moran (2003) 20010824 U (HOVE) (HOVE) (HOVE) Source Moran (2003) (HOVE) (HOVE) (HOVE) Moran (2003) (HOVE) (HOVE) Pop. size UT CO Rep ? ?Pres ? Schelz and X Unc Schelz Pres X X Unc Litzinger 743 Pres O X Unc Fertig 22623 Park stat. Rep O ?Pres X Schelz and Pres Unc Litzinger 633 X Unc Schelz 2077 Pres X X Unc Litzinger 802 Rep O ? Schelz and Pres X X UncPres Schelz 2130 X Unc Schelz 2078 Wide Wide Wide Wide Range UT/CO Wide Wide Wide Intro Wide Wide Sparse PerF Wide/ PerF Wide/ PerF Wide/ Life form PerF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ showy rush- pink hoary aster PerF Wide/ Douglas’ groundsel common hyalineherb Common name gumweed aster sowthistle desert- dandelion rock- goldenrod prickly lettuce AnnF Intro/ marsh-elder AnnF Wide/ Canyonlands hymenoxys Colorado rubber-plant

var. var. var. var. var. lifolius orus, canescens ivesiana, paucifl lifolius lifolius lifolius accidus accidus fl S. longilobus, S. fl H. fi megacephalus, H. fi tomentosus Synonyms/ notes Taxonomic Aster canescens, Dieteria canescens var. Haplopappus nuttallii, Xanthisma grindelioides grindelioides petradoria var. Tetraneuris ivesiana H. fi lugens, H. fi var.

var.

accidus var. ora douglasii lifolius oribunda Machaeranthera canescens Machaeranthera grindelioides Lygodesmia Lygodesmia grandifl Petradoria pumila Solidago var. Senecio fl Tetraneuris acaulis Tetraneuris H. argentea Hymenopappus fi cinereus Malacothrix sonchoides xanthifolia fl Picradenia richardsonii

var. Species name var.

var.

var. var. ora ora var. pumila longilobus lifolius oribunda Machaeranthera canescens canescens Machaeranthera grindelioides grindelioides Hymenoxys acaulis ivesiana UtahHymenopappus fi cinereus Colorado Lygodesmia Lygodesmia grandifl grandifl Malacothrix sonchoides Petradoria pumila var. Lactuca serriola Lactuca serriola Senecio douglasii var. Hymenoxys richardsonii fl Iva xanthifolia Cyclachaena rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 33 noxious weed. CO state noxious weed (List C). Native to Eurasia Comments Europe Eurasia 2002 1973 Utah state NA 1973 Year Year doc. 2001 1973 1973 Native to 2003 1973 2003 Native to c name with taxonomic and (HOVE) (HOVE) Moran (2003) 20010430 A (HOVE) Source (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) 20030611.03 (HOVE) 20010501 B Pop. size UT CO PresPres X Unc Schelz s.n. X Unc Schelz 2096 Rep ? ? ? Schelz and Pres X X Unc Schelz Park stat. PresPres X X Unc O Litzinger 595 Unc Schmidt 37 Pres O X Unc Schelz 2101 Pres X RarePres Schmidt s.n. O X Unc Schelz Pres X Unc Schelz Wide Intro Wide Wide Range UT/CO Wide Intro Wide Wide Intro Wide PerF Wide/ PerF Intro/ PerF Wide/ PerF Wide/ Life form PerF Wide/ Shrub Wide/ PerF Wide/ PerF Intro/ Shrub Wide/ eld showy goldeneye fi bindweed lance-leaved stonecrop basin groundsel Common name hoary townsendia yellow salsify PerF Intro/ spineless horsebrush common dandelion slender wirelettuce shortspine horsebrush

&

var. var. (2006) Vars. not Vars. Includes ssp. lanceolatum subalpinum Synonyms/ notes Taxonomic minor. minor. recognized in Flora of North America S. minor spinosa T. spinosa T.

major var. var. ora ora cinale Amerosdeum lanceolatum Convolvulus arvensis multifl Heliomeris multifl Stephanomeria tenuifolia Packera multilobata Townsendia Townsendia incana Tragopogon dubius Tetradymia Tetradymia canescens Tetradymia spinosa Taraxacum Taraxacum offi

Species name var. var. ora ora cinale Sedum lanceolatum Convolvulus arvensis multifl Stephanomeria tenuifolia tenuifolia UtahSenecio multilobatus Colorado Viguiera multifl Townsendia Townsendia incana Tragopogon dubius Tetradymia Tetradymia canescens Tetradymia spinosa Taraxacum Taraxacum offi rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Crassulaceae Convolvulaceae Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi biogeographic notes, cont.

34 Annotated Checklist of Vascular Flora: Hovenweep National Monument Eurasia is Left Fork Sawtooth Canyon, House Range, Millard Co., UT (Holmgren & Holmgren 11292 NY) Comments Europe Asia Asia is Westwater, is Westwater, Grand Co., UT (Jones s.n. US) 2001 2001 Native to NA 1992 locality Type Year Year doc. 1973 1975 Native to 2001 Native to 2001 Native to 1975 locality Type c name with taxonomic and 20010403 J (HOVE) 20010430 G (HOVE) Moran (2003) (HOVE) Source (HOVE) (HOVE) 20010403 M (HOVE) 20011005 R (HOVE) (HOVE) Pop. size UT CO Pres X Rare Schelz Pres X X Unc Schelz Rep O O ? Schelz and Pres X Rare Colyer s.n. Park stat. Pres X XPres Com X Litzinger 482 O Unc Litzinger 606 Pres X Unc Schelz Pres X Unc Schelz Pres X X Unc Litzinger 480 Wide Intro Wide RegEn Range UT/CO Sparse Intro Intro Intro Wide PerF Wide/ AnnF Intro/ PerF Wide/ Shrub RegEn/ Life form AnnF Wide/ AnnF Intro/ PerF Wide/ ixweed AnnF Intro/ ax perennial rockcress desert European madwort Holboell’s Holboell’s rockcress small-leaved greasebush Common name western tansy- mustard fl blue mustard AnnF Intro/ little-pod false fl rockcress orum micranthum, A. parvifl var. A. simplex Name may be misapplied (Holmgren et al. 2005) Synonyms/ notes Taxonomic

var. ssp. orum pallens Boechera holboellii pinetorum Boechera pulchra var. Alyssum parvifl Boechera perennans Forsellesia meionandra Descurainia pinnata halictorum Descurainia sophia Camelina microcarpa

var.

Species name

var. var. micranthum pinetorum perennans Arabis holboellii var. Arabis pulchra pallens Alyssum minus var. Arabis perennans var. UtahGlossopetalon spinescens microphyllum Colorado Descurainia pinnata osmiarum Descurainia sophia Chorispora tenella Chorispora tenella Camelina microcarpa rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Crossosomataceae (Celastraceae) Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 35 is St. George, Washington Co., UT (Jones 1636 NY) Comments Europe Africa Europe 1975 2001 1974 locality Type 1975 1973 2001 2001 Year Year doc. 1973 2001 1973 Native to NA Native to 1975 Native to c name with taxonomic and (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) 20011005 I (HOVE) 20010403 G (HOVE) Source (HOVE) 20011005 F (HOVE) (HOVE) Moran (2003) 20010430 N (HOVE) Pop. size UT CO Pres X OPres Unc X Litzinger 485 X Unc Schelz 2141 Pres X X Com Litzinger 325 Pres X O Unc Litzinger 481 Pres XPres X Unc Schmidt 69 X Unc Schelz Pres X Rare Schelz Park stat. Pres X O Unc Schmidt 73 PresPres X Unc X Schelz O Unc Litzinger 484 Rep O O ? Schelz and Pres X Unc Schelz Wide Periph Wide Wide Wide RegEn Periph Range UT/CO Wide Wide Intro Intro Intro AnnF Wide/ AnnF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ PerF RegEn/ Life form PerF Wide/ AnnF Intro/ AnnF Intro/ AnnF Intro/ ax-leaved prairie pepperwort hairy-pod pepperwort Jones’ pepperwort wedgeleaf draba sharp-leaf twinpod Colorado Plateau bladderpod spectacle-pod AnnF Wide/ Common name prince’s-plume PerF Wide/ fl plainsmustard tumble mustard African mustard clasping pepperwort Welsh et al. Welsh (2003) include var. macrocarpum Synonyms/ notes Taxonomic Sisymbrium linifolium

var. orum pinnata Lepidium lasiocarpum Lepidium densifl Lepidium montanum jonesii Draba cuneifolia rectipes Schoenocrambe linifolia Physaria acutifolia australis P. wislizenii var. Sisymbrium altissimum Malcolmia africana Lepidium perfoliatum

var. var. Species name var. orum orum cuneifolia acutifolia Lepidium lasiocarpum lasiocarpum Lepidium montanum jonesii Lepidium densifl densifl Physaria rectipes Lesquerella Schoenocrambe linifolia Draba cuneifolia var. UtahDithyrea wislizenii Dimorphocarpa Colorado Stanleya pinnata Stanleya pinnata Physaria acutifolia var. Sisymbrium altissimum Malcolmia africana Lepidium perfoliatum rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix A. Confi biogeographic notes, cont.

36 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments 1973 Year Year doc. 2001 2002 NA NA 1973 1972 2006 2001 2002 NA NA 1972 c name with taxonomic and 20010403 D (HOVE) Source 20010824 K (HOVE) (HOVE) Moran (2003) (2003) (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) Moran (2003) Moran (2003) (HOVE) Moran Pop. size UT CO Pres X O Unc Schelz Park stat. PresPres X Rare Schelz X Rare Schelz 2134 Rep O ? Schelz and Pres X X Unc Litzinger 484 Pres X X Com Schmidt 89 Pres X Unc Fertig 22619 Pres XPres Rare Schelz 2092 X O Unc Schelz 2062 Rep O ORep ? ? Schelz and ?Pres ? XRep X Schelz and ? Abun Schelz 2116 ? ? Schelz and Periph Range UT/CO Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide AnnF Wide/ Life form PerG Wide/ PerF Wide/ Shrub Wide/ Shrub Wide/ PerF Wide/ ower ddle- eld dodder eld AnnF Wide/ long-beak fi mustard Common name blackcreeper sedge western sedge PerG Wide/ elk sedge PerG Wide/ heart-leaf jewelfl green ephedra Ross’ sedge PerG Wide/ Torrey’s Torrey’s ephedra alkali bulrush PerG Wide/ alpine pennycress gourdbuffalo PerF Wide/ Utah juniper Shrub Wide/ fi

var. var. var. exa boottii, paludosus, Synonyms/ notes Taxonomic S. cordatus cordatus var. C. brevipes, pityophila Scirpus maritimus var. Schoenoplectus maritimus montana

ssp. Streptanthus cordatus Streptanthella longirostris Ephedra viridis maritimus paludosus Noccaea montana N. montana Ephedra torreyana Cucurbita foetidissima Sabina osteosperma Grammica campestris

Species name var. viridis Streptanthus cordatus UtahStreptanthella longirostris Colorado Carex praegracilis Carex praegracilis Carex rossii Carex rossii C. defl Carex occidentalis Carex occidentalis Carex geyeri Carex geyeri Thlaspi montanum Scirpus maritimus Bolboschoenus montanum Ephedra torreyana Ephedra viridis var. Cucurbita foetidissima Juniperus osteosperma Cuscuta pentagona rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Family Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cruciferae (Brassicaceae) Cyperaceae Ephedraceae Ephedraceae Cucurbitaceae Cupressaceae Cuscutaceae (Convolvulaceae) Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 37 tuckeri Comments is Buckskin Mts, Kaibab Plateau “on the so. edge of Utah”, Kane Co., UT (Jones s.n. POM). CO BLM Sensitive is Butler San Wash, Juan Co., UT (Eastwood 141 NY) locality of Q. havardii var. is Low Pass E of Kens Lake, San Juan Co., UT (Welsh & Higgins 23630 BRY) Year Year doc. 1973 locality Type 1975 2002 locality Type 1973 1975 1973 Type NA c name with taxonomic and Source (HOVE) (HOVE) 20020912.6 (HOVE) (HOVE) (HOVE) (HOVE) Moran (2003) Pop. size UT CO Park stat. Pres X Unc Schmidt 56 Pres X X Unc Schelz 2170 Pres X Rare Schelz Pres X O Unc Litzinger 326 Pres XPres O X Unc Litzinger 830 Unc Schmidt 97 Rep ? ? ? Schelz and Range UT/CO RegEn Wide RegEn Wide Wide RegEn Wide Life form PerF Wide/ Shrub RegEn/ PerF Wide/ AnnF Wide/ golden Common name Utah swertia PerF RegEn/ corydalis Eastwood’s Eastwood’s oak Fendler’s Fendler’s spurge oakGambel’s Treeshinnery oak Wide/ Shrub RegEn/ ridge-seeded spurge Q. x Synonyms/ notes Taxonomic welshii Euphorbia fendleri Q. undulata Hybrid between Q. gambelii Euphorbia glyptosperma

ssp. x tuckeri Corydalis curvisiliqua occidentalis Chamaesyce fendleri eastwoodiae Quercus Chamaesyce glyptosperma var. Quercus gambelii Species name x occidentalis gambelii UtahChamaesyce fendleri Colorado Swertia utahensis Frasera paniculata Corydalis aurea var. Quercus eastwoodiae Chamaesyce glyptosperma Quercus welshii Quercus havardii Quercus gambelii var. rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Euphorbiaceae Family Gentianaceae Fumariaceae Fagaceae Euphorbiaceae Fagaceae Fagaceae Appendix A. Confi biogeographic notes, cont.

38 Annotated Checklist of Vascular Flora: Hovenweep National Monument noxious weed (List C). Native to Eurasia noxious weed (List B). Native to Europe Comments Eurasia Eurasia. Eurasia Eurasia Eurasia. Eurasia and Africa NA CO state 1973 CO state Year Year doc. 1975 Native to 1975 Native to 2002 Native to 2001 1975 Native to 2006 Native to 1975 2002 Native to c name with taxonomic and Moran (2003) (HOVE) Source (HOVE) (HOVE) 20020607.01 (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) Pop. size UT CO Rep O ? Schelz and Pres X O Unc Schelz 2071 Park stat. Pres X Unc Litzinger 575 Pres X X Unc Schelz 2064 Pres X Unc Schelz Pres O X Unc Schelz 2117 Pres X Rare Litzinger 824 Pres O X Unc Fertig 22616 Pres X O Unc Litzinger 322 Pres X Unc Schelz 1959 Intro Intro Range UT/CO Intro Intro Intro Wide Intro Intro Wide Intro AnnG Intro/ Life form AnnG Intro/ PerG Intro/ PerG Intro/ PerG Wide/ jointed goatgrass stork’s-bill AnnF Intro/ Common name red brome AnnG Intro/ Japanese chess smooth brome blue grama PerG Wide/ oats AnnG Intro/ crested wheatgrass purple three- awn redtop PerG Intro/

var. var. ssp. B. Synonyms/ notes Taxonomic includes commutatus A. cristatum A. sativa cristatum, A. cristatum pectinatum, A. pectinatum, A. sibiricum, A. mongollicum A. glauca, purpurea fendleriana, A. fendleriana A. alba, gigantea

var. longiseta Agropyron desertorum Cylindropyrum cylindricum Erodium cicutarium Bromopsis inermis Avena fatua Avena sativa gracile Agrostis stolonifera var.

var. Species name inermis Agropyron cristatum Aegilops cylindrica UtahErodium cicutarium Colorado Bromus rubens Anisantha rubens Bromus japonicus Bromus japonicus B. arvensis; sativa Bouteloua gracilis Chondrosum Bromus inermis var. Agrostis stolonifera Aristida purpurea Aristida purpurea fatua Avena rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Geraniaceae Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 39 to South America noxious weed (List C). Native to Eurasia Comments is top of Salina Pass, Sevier Co., UT (Jones 5447 POM) Eurasia Eurasia. Eurasia NA Native 1973 CO state Year Year doc. 1973 locality Type 2001 Native to 2003 NA Native to 1973 NA Native to c name with taxonomic and Moran (2003) (HOVE) Source (HOVE) 20010824 N (HOVE) (HOVE) Moran (2003) (HOVE) Moran (2003) Pop. size UT CO Rep O ? Schelz and Pres X O Abun Litzinger 305 Park stat. Pres X X Unc Schelz 2137 Pres O X Unc Schelz 2112 Rep ? ? ? Schelz and Pres X O Unc Litzinger 310 Rep O ? Schelz and Pres X Unc Schelz Intro Intro Range UT/CO Wide Wide Intro Wide Intro Intro Life form PerG Wide/ AnnG Intro/ PerG Intro/ PerG Intro/ Chilean chess AnnG Intro/ cheatgrass AnnG Intro/ Common name Salina wildrye PerG Wide/ western wheatgrass barnyard- grass squirreltail PerG Wide/ tall wheatgrass intermediate wheatgrass

var. & includes smithii elymoides, ponticus, hispidus, Synonyms/ notes Taxonomic ambiguus Welsh et al. Welsh (2003) include E. microstachya (E. muricata Agropyron smithii; vars. palmeri microstachya) var. Sitanion hystrix E. elongatus var. Agropyron elongatum E. hispidus var. Agropyron intermedium, Elytrigia intermedia

ssp. Echinochloa crus- galli tectorum smithii ponticum intermedium barbulatum Species name Echinochloa crus- galli Bromus trinii Bromus trinii B. berteroanus UtahBromus tectorum Anisantha Colorado Elymus smithii Pascopyrum Elymus salinus Leymus salina E. salina, L. Elymus elongatus Thinopyrum Elymus elymoides Elymus elymoides E. elymoides Elymus hispidus Thinopyrum rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix A. Confi biogeographic notes, cont.

40 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments Eurasia Europe. 1974 2006 Year Year doc. 2002 2001 2001 1975 NA Native to 1975 Native to 2001 NA NA c name with taxonomic and (HOVE) (HOVE) Source 20020607.03 (HOVE) (HOVE) 20010430 F (HOVE) (HOVE) Moran (2003) 20010430 B (HOVE) (HOVE) Moran (2003) Moran (2003) Pop. size UT CO Pres X X Com Litzinger 312 Pres X Unc Fertig 22627 Park stat. Pres O X Unc Schelz Pres X X Unc Schelz 1958 Pres XPres Rare X Schelz ORep Unc ? Litzinger 477 ? ? Schelz and Pres X Unc Schelz Pres O XRep Unc ? Schelz 2111 ?Rep ? O O Schelz and ? Schelz and Wide Wide Range UT/CO Wide Wide Wide Intro Wide Intro Wide Wide Wide PerG Wide/ Life form PerG Wide/ AnnG Wide/ PerG Intro/ galleta PerG Wide/ fowl mannagrass Common name foxtail barley PerG Wide/ slender wheatgrass hairy tridens PerG Wide/ six-weeks fescue meadow fescue rabbit barley AnnG Intro/ foxtail muhly PerG Wide/ sandhill muhly PerG Wide/ junegrass PerG Wide/

& ssp. G. elata ora latiglumis, trachycaulus, Synonyms/ notes Taxonomic Includes E. trachycaulus var. E. alaskanus ssp. A. caninum, subsecundum, A. trachycaulum, A. latiglume Schedonorus pratensis leporinum, H. leporinum Includes vars. glauca, hirtella, octofl K. cristata, pyramidata, K. nitida ora Elymus trachycaulus Critesion glaucum H. murinum Erioneuron pilosum Koeleria macrantha Muhlenbergia andina Muhlenbergia pungens oraoctofl Vulpia Species name Hilaria jamesii Hilaria jamesii Pleuraphis jamesii UtahElymus trachycaulus Colorado Hordeum murinum Hordeum jubatum Critesion jubatum Glyceria striata Glyceria striata Erioneuron pilosum Festuca octofl Festuca pratensis Festuca pratensis Lolium pratense, Koeleria macrantha Muhlenbergia andina Muhlenbergia pungens rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Gramineae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 41 Comments Eurasia and Africa Europe Eurasia Eurasia and Africa 2001 NA Year Year doc. 1975 1973 Native to 1975 2001 Native to 2001 Native to NA 2003 2001 Native to c name with taxonomic and 20010830 D (HOVE) Moran (2003) Source (HOVE) 20010830 J (HOVE) (HOVE) (HOVE) (HOVE) Moran (2003) (HOVE) 20011005 S (HOVE) Pop. size UT CO Pres XRep Rare ? Schelz ? ? Schelz and Park stat. Pres X X Unc Schelz 1950 Pres X Unc Schelz Pres X X Unc Schelz 2110 Pres X X Com Schelz 1966 Pres X Unc Schelz 2102 RepPres O ? XPres X Schelz and X Unc X Schelz 2139 Unc Schelz Wide Wide Range UT/CO Wide Intro Intro Wide Intro Periph Intro Wide AnnG Wide/ PerG Wide/ Life form PerG Wide/ AnnG Intro/ PerG Intro/ PerG Intro/ AnnG Wide/ PerG Intro/ false buffalograss littleseed ricegrass Common name Sandberg’s Sandberg’s bluegrass rabbitfoot- grass Kentucky bluegrass muttongrass PerG Wide/ Canada bluegrass Bigelow’s Bigelow’s bluegrass bulbous bluegrass common reed PerG Wide/ P. & P. P. secunda P. and all longiligula; P. cusickii P. Synonyms/ notes Taxonomic Includes gracillima, P. juncifolia, P. sandbergii, canbyi, P. P. scabrella, P. nevadensis, P. ampla vars. of P. fendleriana P. ssp. includes reports of epilis P. communis P. Piptatherum micranthum Phragmites australis Polypogon monspeliensis Species name Oryzopsis micrantha Phragmites australis UtahMunroa squarrosa Monroa squarrosa Colorado Poa secunda Poa secunda Polypogon monspeliensis Poa pratensis Poa pratensis agassizensis P. Poa fendleriana Poa fendleriana longiligula, P. Poa compressa Poa compressa Poa bigeloviiPoa bulbosa Poa bigelovii Poa bulbosa rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix A. Confi biogeographic notes, cont.

42 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments Europe 1975 NA 2001 Year Year doc. 2006 NA Native to 2001 2002 1973 2001 1974 2003 1975 1975 c name with taxonomic and (HOVE) Moran (2003) 20010824 E (HOVE) Source (HOVE) Moran (2003) 20010501 J (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) Pop. size UT CO Pres X XRep Unc ? Litzinger 682 ?Pres ? O X Schelz and Unc Schelz Park stat. Pres X Rare Fertig 22620 Rep O ? Schelz and Pres X X Unc Schelz Pres X X Unc Schmidt 65 Pres X X Unc Schelz 2090 Pres X Rare Litzinger 336 Pres O X Unc Schelz 2063 Pres X O Unc Litzinger 576 Pres X O Unc Litzinger 302 Pres X X Unc Schelz 2099 Sparse Wide Wide Range UT/CO Wide Intro Wide Wide Wide Wide Wide Wide Wide Periph PerG Wide/ Life form PerF Wide/ AnnF Wide/ PerG Wide/ PerG Wide/ PerG Wide/ ower alkali sacaton PerG Wide/ spike dropseed sand dropseed PerG Wide/ Common name smallfl dragonhead German iris PerF Intro/ Ives’ phacelia AnnF Wide/ corrugate phacelia Torrey’s rushTorrey’s PerG Wide/ longstyle rush PerG Wide/ Baltic rush PerG Wide/ needle-and- thread Indian ricegrass desert needlegrass

var. var. ora montanus; Synonyms/ notes Taxonomic Var. not Var. recognized in Cronquist et al. (1984) Moldavica parvifl balticus, J. balticus var. includes var. mexicanus J. arcticus H. comata comata Oryzopsis hymenoides Jarava speciosa orum ater Phacelia crenulata Sporobolus contractus Sporobolus cryptandrus Sporobolus airoides ssp. Hesperostipa comata Dracocephalum parvifl hymenoides speciosum

var. Species name

var. orum corrugata Sporobolus contractus Sporobolus cryptandrus Stipa comata UtahSporobolus airoides airoides Colorado Juncus arcticus Juncus arcticus Iris germanica Iris germanica Phacelia crenulata var. Phacelia ivesiana Phacelia ivesiana comata Dracocephalum parvifl Juncus torreyi Juncus torreyi Juncus longistylus Juncus longistylus Stipa hymenoides Achnatherum Stipa speciosa Achnatherum rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Juncaceae Iridaceae Hydrophyllaceae Hydrophyllaceae Labiatae (Lamiaceae) Juncaceae Juncaceae Gramineae (Poaceae) Gramineae (Poaceae) Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 43 is near CisCo., Grand Co., UT (Jones s.n. POM) is near Monticello, San Juan Co., UT (Eastwood 9 POM) Europe Eurasia Comments is Kanab, Kane Co., UT (Thompson s.n. GH) NA locality Type 2001 locality Type 2001 1994 2005 Native to 1975 1973 Native to Year Year doc. NA 1973 locality Type 1973 c name with taxonomic and Moran (2003) 20010811 A (HOVE) 20011005 W (HOVE) Oppelt 1046 (HOVE) s.n. (HOVE) (HOVE) (HOVE) Source Moran (2003) (HOVE) 20010430 P (HOVE) Pop. size UT CO Rep O O ? Schelz and Pres X X Rare Schelz Pres O XPres Unc Schelz X Rare Colyer & Pres X Unc Harms, R. Pres XPres O X Unc X Litzinger 757 Unc Litzinger 337 Park stat. Rep O ? Schelz and Pres X X Unc Schmidt 74 Pres X X Com Schelz RegEn RegEn Periph Periph Intro Wide Intro Range UT/CO Periph Wide RegEn PerF RegEn/ PerF RegEn/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Intro/ Life form PerF Wide/ PerF Wide/ AnnF RegEn/ owered rimrock milkvetch Moab milkvetch painted milkvetch crescent milkvetch Torrey’s milkvetch catnip PerF Intro/ mock pennyroyal common horehound Common name woolly milkvetch small- fl milkvetch Synonyms/ notes Taxonomic

var. var. var. moabensis Astragalus desperatus Astragalus coltonii var. ceramicus Astragalus amphioxys Astragalus calycosus Astragalus ceramicus Marrubium vulgare Hedeoma drummondii Astragalus mollissimus Astragalus nuttallianus thompsoniae micranthiformis

Species name var. var. var. var. var. var. var. moabensis Astragalus mollissimus Astragalus desperatus desperatus Astragalus amphioxys Astragalus calycosus calycosus Astragalus ceramicus ceramicus amphioxys Nepeta cataria Nepeta cataria Marrubium vulgare UtahHedeoma drummondii Colorado thompsoniae Astragalus coltonii var. Astragalus nuttallianus micranthiformis rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Family Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix A. Confi biogeographic notes, cont.

44 Annotated Checklist of Vascular Flora: Hovenweep National Monument L.

Europe Comments for Colorado & (Weber Wittmann 2001 consider previous reports erroneous) Europe Europe reports of argenteus from Von Loh et al. (2008) 2001 Native to NA Year Year doc. 1975 State record 2002 Native to 1973 Native to 2001 2002 Includes 2001 2006 2001 c name with taxonomic and 20010824 B (HOVE) Moran (2003) Source (HOVE) 20020607.07 (HOVE) (HOVE) (HOVE) 20020912.28 (HOVE) 20011005 U (HOVE) (HOVE) 20010503.21 (HOVE) Pop. size UT CO Pres X Unc Schelz Rep O O ? Schelz and Park stat. Pres X Rare Litzinger 653 Pres O X Unc Schelz Pres X O Unc Schmidt s.n. Pres X Rare Schelz 2135 Pres X Unc Schelz Pres O XPres Unc Schelz X Unc Fertig 22621 Pres X Rare Schelz Intro Wide Range UT/CO Periph Intro ntro RegEn Wide Wide RegEn Wide Life form PerF Intro/ AnnF Intro/ PerF Wide/ PerF RegEn/ PerF Wide/ alfalfa PerF Intro/ dwarf lupine AnnF Wide/ Common name rush peteria PerF NA/ yellow sweet- clover white sweet- clover sand lupine PerF RegEn/ Utah lupine PerF Wide/ Rocky Mountain milkvetch Fort Wingate milkvetch northern sweetvetch by

for

ssp. var. included var. cinalis A. adanus M. offi in sativa caudatus, L. argenteus Synonyms/ notes Taxonomic some authors L. caudatus utahensis Includes reports of UT Includes var. Includes var. pabulare utahensis cinale Lupinus pusillus Astragalus scopulorum Peteria scoparia Melilotus offi Lupinus ammophilus Hedysarum boreale Lupinus caudatus var. Astragalus wingatanus

Species name var.

var. pusillus utahensis cinalis Melilotus alba Melilotus albus M. albus; Medicago sativa Medicago sativa M. sativa UtahAstragalus scopulorum Colorado Melilotus offi Lupinus pusillus var. Lupinus polyphyllus ammophilus boreale Astragalus wingatanus Hedysarum boreale Lupinus caudatus var. rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Family Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 45 ower Comments of Utah is near Kanab, Kane Co., UT (Thompson s.n. GH). USFS R2 Sensitive. Eurasia is near Kanab, Kane Co., UT (Thompson s.n.) Europe in N AZ, near Kanab, or along the Colorado River in SE UT or AZ (Thompson s.n. US). USFWS: 3C Year Year doc. NA 1976 2006 State fl 1973 locality Type 2002 Native to 1975 locality Type 2003 Native to 2001 locality Type c name with taxonomic and Source Moran (2003) (HOVE) (HOVE) (HOVE) 20020607.04 20010403 A (HOVE) al. (2008) & Schelz & Moran (2003) 20010503 A (HOVE) Pop. size UT CO Park stat. Rep ? ?Pres ? X X Schelz and Unc Litzinger 510 Pres X Unc Fertig 22625 Pres X X Unc Schmidt 82 Pres X Rare Schelz Pres X O Unc Schelz Rep O Unc Loh et Von Pres X Rare Schelz Range UT/CO Wide Wide Wide Periph Intro Wide Intro LocEn Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Shrub LocEn/ Common name false Solomon’s- seal foothills death camas sego-lily PerF Wide/ sinuous mariposa asparagus PerF Intro/ pink funnel- lily red clover PerF Intro/ Thompson’s Thompson’s indigo-bush amplexicaule Synonyms/ notes Taxonomic Brodiaea paysonii M. racemosum var.

var. orum cinalis exuosus Psorothamnus thompsoniae Maianthemum amplexicaule Toxicoscordion paniculatum Calochortus nuttallii Calochortus fl Asparagus offi Androstephium brevifl thompsoniae

var. Species name orum cinalis exuosus UtahPsorothamnus thompsoniae thompsoniae Colorado Smilacina racemosa Zigadenus paniculatus nuttallii Calochortus Calochortus fl Asparagus offi Androstephium brevifl Trifolium pratenseTrifolium pratense Trifolium rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Leguminosae (Fabaceae) Family Liliaceae (Convallariaceae or Ruscaceae) Liliaceae (Melanthiaceae) Liliaceae (Calochortaceae) Liliaceae (Calochortaceae) Liliaceae (Asparagaceae) Liliaceae (Alliaceae, Themidaceae) Leguminosae (Fabaceae) Appendix A. Confi biogeographic notes, cont.

46 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments Eurasia material is var. cronquistii. locality Type is 75 miles W of Blanding and 10 miles E of Hite, San Juan Co., UT (Cronquist 9030 NY) NA 1974 Year Year doc. 1973 1974 1973 1973 NA 1976 NA Native to 1974 HOVE c name with taxonomic and Moran (2003) 20020425.01 (HOVE) Source 20010403 Q (HOVE) (HOVE) (HOVE) (HOVE) Moran (2003) (HOVE) Moran (2003) (HOVE) Pop. size UT CO Rep ? ?Pres ? X X Schelz and Unc Schelz Park stat. Pres X XPres Unc X Schelz Pres X RarePres O Litzinger 370 X Unc Schmidt 92 Rep Unc ? Litzinger 573 ? ? Schelz and Pres X Unc Litzinger 894 Rep O ? Schelz and Pres X Rare Litzinger 332 Periph Wide Range UT/CO Wide Periph Wide Wide Periph Wide Intro RegEn Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ AnnF Intro/ PerF RegEn/ ax AnnF Wide/ ax PerF Wide/ broom-fl Common name mallow slenderleaf globemallow small-leaf globemallow fragrant sand- verbena trailing four- o’clock blue fl white-stem blazingstar scarlet globe- common mallow Colorado stickleaf

var. var.

var. elata & var. harrisii cronquistii, Synonyms/ notes Taxonomic incarnata L. lewisii lewisii M. obscura, A. gracilis elliptica, A. nana var. Includes ssp. coccinea M. marginata var. M. cronquistii, N. cronquistii

ssp. Adenolinum lewisii Acrolasia albicaulis aristatum Sphaeralcea leptophylla Sphaeralcea parvifolia dissecta Nuttallia marginata Sphaeralcea coccinea Species name lewisii Linum perenne ssp. Mentzelia albicaulis UtahLinum aristatum Mesynium Colorado Sphaeralcea leptophylla Sphaeralcea parvifolia Abronia fragrans Abronia elliptica A. fragrans Allionia incarnata Allionia incarnata A. incarnata Mentzelia marginata Sphaeralcea coccinea Malva neglecta Malva neglecta rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Linaceae Loasaceae Linaceae Family Malvaceae Malvaceae Nyctaginaceae Nyctaginaceae Loasaceae Malvaceae Malvaceae Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 47 material is var. glandulosa is Labyrinth Canyon, San Juan Co., UT (Newberry s.n. NY) Comments 1973 HOVE 1973 locality Type NA Year Year doc. NA 2001 2003 NA NA NA 1973 1974 c name with taxonomic and (HOVE) (HOVE) Moran (2003) Source Moran (2003) (HOVE) (HOVE) Moran (2003) Moran (2003) Moran (2003) (HOVE) (HOVE) Pop. size UT CO Pres X O Unc Schmidt 98 Pres X O Unc Schmidt 100 Rep O ? Schelz and Rep O ? Schelz and Park stat. Pres X XPres Unc O Schelz 2079 XRep Unc O Schelz 2073 Unc Schelz and Rep O ? Schelz and Rep O ? Schelz and Pres X X UncPres Litzinger 350 O X Abun Litzinger 375 Wide Wide Wide Wide Range UT/CO Wide Periph Wide Wide Wide Wide Wide PerF Wide/ PerF Wide/ Life form PerF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ Tree Wide/ showy four- o’clock singleleaf ash Shrub Wide/ desert olive Shrub Wide/ narrow-leaf four-o’clock Common name long-tube evening- primrose pale evening- primrose northern willow-herb white-stem evening- primrose spotted coral- root two-needle pinyon Cooper’s Cooper’s broomrape

var. & from O. (1993), ora, ora ciliatum, Includes vars. multifl pubescens, glandulosa F. neomexicana F. O. decumbens, M. glabra Synonyms/ notes Taxonomic Includes var. Includes var. runcinata E. ciliatum var. E. watsonii, adenocaulon maculata C. maculata UT Flora O. cooperi Includes multifl

ssp. ora pallida Forestiera pubescens Oxybaphus linearis Mirabilis multifl Oenothera pallida ssp. Epilobium ciliatum Oenothera cespitosa marginata Corallorhiza maculata Oenothera albicaulis Orobanche ludoviciana

Species name var. var. ora pallida linearis Forestiera pubescens Epilobium ciliatum Fraxinus anomala Fraxinus anomala UtahMirabilis linearis var. Colorado Mirabilis multifl Oenothera caespitosa marginata Oenothera pallida var. Corallorhiza maculata Oenothera albicaulis Orobanche ludoviciana Pinus edulis Pinus edulis cooperi rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Oleaceae Onagraceae Oleaceae Nyctaginaceae Nyctaginaceae Family Onagraceae Onagraceae Orchidaceae Onagraceae Orobanchaceae Pinaceae Appendix A. Confi biogeographic notes, cont.

48 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments Year Year doc. NA 1973 1973 2001 1973 1994 1973 NA 1973 2001 c name with taxonomic and Source Moran (2003) (HOVE) (HOVE) 20010501 M (HOVE) (HOVE) 1049/1050 (HOVE) 20011005 Q (HOVE) Moran (2003) (HOVE) 20011005 A (HOVE) Pop. size UT CO Park stat. Rep O OPres Unc X Schelz and X Unc Schelz 2061 Pres X X Unc Litzinger 530 Pres O X Unc Schelz Pres X O Unc Schmidt 2 Pres O XPres Unc X Colyer X Unc Schelz Rep O O ? Schelz and Pres X O Unc Litzinger 557 Pres X Rare Schelz Range UT/CO Wide Wide Wide Wide Wide Periph Periph Periph Wide Sparse Life form PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ Common name long-leaf phlox slender phlox AnnF Wide/ dwarf gilia AnnF Wide/ spreading gilia woolly plantain spreading eriastrum shy gilia AnnF Wide/ common gilia AnnF Wide/ prickly-phlox Shrub Wide/ ball-head gilia PerF Wide/

var. ssp.

& & & var. Synonyms/ notes Taxonomic Includes vars. longifolia stansburyi humilis Includes vars. patagonica, gnaphaloides, spinulosa, breviscapa Includes vars. inconspica sinuata, G. ophthalmoides, G. clokeyi, transmontana leptomeria Leptodactylon pungens pungens, Linanthus pungens Gilia congesta congesta Gilia polycladon

var. Plantago patagonica Gilia leptomeria Aliciella Microsteris gracilis Phlox gracilis Leptodactylon pungens congesta Eriastrum diffusum Ipomopsis congesta Ipomopsis polycladon

Species name var. leptomeria humilior UtahPlantago patagonica Colorado Phlox longifolia Phlox longifolia Leptodactylon pungens Ipomopsis pumila Ipomopsis pumila Gilia pumila Eriastrum diffusum Gilia inconspicua Gilia sinuata Gilia leptomeria var. Ipomopsis congesta congesta Microsteris gracilis var. Ipomopsis polycladon rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Plantaginaceae Family Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 49 rmed Comments Eurasia is Aquarius Plateau, Co., Wayne UT (Ward 722 GH) to be confi for HOVE material Eurasia. Year Year doc. 1975 1973 1975 Native to NA NA 1975 2006 locality Type 2001 2002 needs Var. 1973 1973 2001 Native to c name with taxonomic and Source 20020916.5 (HOVE) (HOVE) (HOVE) Moran (2003) Moran (2003) (HOVE) (HOVE) 20010907 C (HOVE) 20020607.09 20010430 M (HOVE) (HOVE) 20010824 F (HOVE) Pop. size UT CO Park stat. Pres X O Com Schelz Pres X O Unc Schmidt 18 Rep ORep ? ? ?Pres Schelz and ? X Schelz and Rare Litzinger 684 Pres X Rare Fertig 22626 PresPres X Unc Schelz X Unc Schelz Pres X XPres Unc X Schelz X Unc Schmidt 22 Pres X X Unc Litzinger 479 Pres O X Unc Schelz Range UT/CO Wide Wide Wide Periph Wide Wide Wide Periph Wide Periph Intro Intro Life form PerF Wide/ Shrub Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ AnnF Intro/ Common name slender wild buckwheat canaigre dock PerF Wide/ crisp-leaf wild buckwheat spreading wild buckwheat desert trumpet Baker’s wild Baker’s buckwheat redroot wild buckwheat arid wild buckwheat cushion wild buckwheat wild Palmer’s buckwheat curly dock PerF Intro/ yard knotweed

var. var. var. E. ssp. & foliosum Synonyms/ notes Taxonomic Includes var. Includes var. velutinum lancifolium E. microthecum var. E. arcuatum arcuatum E. ovalifolium purpureum P. aviculare P. aviculare

var. var. var. var. atum Eriogonum corymbosum corymbosum simpsonii Eriogonum ovalifolium Rumex hymenosepalus Eriogonum divaricatum Eriogonum infl Eriogonum jamesii ovalifolium Eriogonum microthecum Polygonum arenastrum Eriogonum umbellatum subaridum Eriogonum racemosum Eriogonum palmerianum

var. var. var.

Species name var. var. var. var. atum atum avescens UtahEriogonum corymbosum corymbosum Colorado Eriogonum ovalifolium Rumex hymenosepalus Eriogonum divaricatum Eriogonum infl infl Eriogonum jamesii fl Eriogonum microthecum simpsonii Polygonum aviculare Eriogonum umbellatum subaridum racemosum Eriogonum palmerianum Eriogonum racemosum Rumex crispus Rumex crispus rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Polygonaceae Family Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Appendix A. Confi biogeographic notes, cont.

50 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments E North America locality is Toquerville, Washington Co., UT (Jones 1716 GH) Eurasia Year Year doc. 1975 2003 2001 2001 2001 NA Native to 2001 1975 2001 1975 Type 1975 Native to c name with taxonomic and Source (HOVE) (HOVE) (HOVE) 20010824 R (HOVE) 20010501 W (HOVE) Moran (2003) 20010501 S (HOVE) (HOVE) 20011005 N (HOVE) (HOVE) 20010403 K (HOVE) Pop. size UT CO Park stat. Pres X O Unc Litzinger 614 PresPres X X Rare Schelz 2169 Pres Unc Schelz s.n. X Unc Schelz PresRep X Rare Schelz O ? Schelz and Pres O X Unc Schelz PresPres X Unc Litzinger 599 X Unc Schelz Pres O X Com Litzinger 490 Pres X O Unc Schelz Range UT/CO Wide Wide Wide Periph Intro Wide Wide Wide Wide Intro Wide Life form AnnF Wide/ AnnF Wide/ PerF Wide/ AnnF Periph/ AnnF Intro/ PerF Wide/ Shrub Wide/ Shrub Wide/ ower Common name pygmy-fl rock-jasmine western rock- jasmine white virgin’s- bower common purslane dwarf purslane Nuttall’s Nuttall’s larkspur alder-leaf alder-leaf mountain mahogany squaw-apple Shrub Wide/ Utah serviceberry bur buttercup AnnF Intro/ pale larkspur PerF Wide/ &

subilifera Synonyms/ notes Taxonomic P. parvula P. D. nelsonii puberulenta, subumbellata, var. Includes ssp. C. testiculata D. scaposum

var. Portulaca halimoides Delphinium nuttallianum Androsace occidentalis Androsace septentrionalis Clematis ligusticifolia Ceratocephala orthoceras Delphinium andersonii Cercocarpus montanus Peraphyllum ramosissimum Amelanchier utahensis scaposum

Species name var. UtahPortulaca halimoides Colorado Delphinium nuttallianum Androsace occidentalis Androsace septentrionalis Clematis ligusticifolia Delphinium andersonii Portulaca oleracea Portulaca oleracea Ranunculus testiculatus scaposum Cercocarpus montanus Peraphyllum ramosissimum Amelanchier utahensis rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Portulacaceae Family Ranunculaceae (Helleboraceae) Primulaceae Primulaceae Ranunculaceae Ranunculaceae (Helleboraceae) Portulacaceae Ranunculaceae Rosaceae Rosaceae Rosaceae Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 51 Comments 2003 1975 1973 1972 1975 1972 Year Year doc. 1975 1972 1972 NA 1974 1974 2001 2001 eld 30 eld eld 62 eld c name with taxonomic and (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) Source (HOVE) (HOVE) (HOVE) Moran (2003) 20011005.22 (HOVE) (HOVE) (HOVE) 20010824 V (HOVE) Pop. size UT CO PresPres X Unc X Schelz 2083 X Com Schelz 2084 Pres XPres O X UncPres O Litzinger 531 Com Redfi X Unc Litzinger 600 Pres X X Unc Litzinger 335 Park stat. Pres O X Unc Litzinger 372 Pres X OPres Unc X Litzinger 307 Rep X ? Unc Redfi ? ? Schelz and Pres X X Unc Schelz Pres XPres Rare Schmidt 44 X Unc Schelz 2168 Pres O X Unc Schelz Wide Wide Wide Wide Wide Wide Range UT/CO Wide Wide Wide Wide Wide RegEn Wide Wide Tree Wide/ Shrub Wide/ Shrub Wide/ Life form AnnF Wide/ PerF Wide/ Shrub Wide/ Shrub Wide/ PerF Wide/ PerF RegEn/ AnnF Wide/ PerF Wide/ ax Fremont cottonwood antelope bitterbrush roseWoods’ Shrub Wide/ western chokecherry cliff-rose Shrub Wide/ coyote willow Shrub Wide/ Common name Wright’s Wright’s bird’s-beak bastard toadfl trumpet gooseberry golden currant desert paintbrush Eastwood’s Eastwood’s paintbrush blue-eyed Mary Wyoming paintbrush

var. ssp. var. var. var. dubia, C. P. deltoides P. fremontii ultramontana, R. manca Cowania mexicana exigua Synonyms/ notes Taxonomic C. pallida aureum C. angustifolia var. applegatei martinii C. linariaefolia ora wislizenii melanocarpa Populus deltoides ssp. Purshia stansburiana Salix exigua S. exigua Padus virginiana ssp. wrightii Comandra umbellata Castilleja chromosa Castilleja linariifolia Collinsia parvifl Cordylanthus Castilleja scabrida

Species name var. var. ora fremontii stansburyana melanocarpa scabrida Populus fremontii var. Salix exigua stenophylla Purshia mexicana var. Purshia tridentata Purshia tridentata Rosa woodsii Rosa woodsii R. woodsii Comandra UtahPrunus virginiana var. Colorado umbellata pallida Ribes aureum Ribes aureum Ribes leptanthum Ribes leptanthum R. aureum Castilleja chromosa Castilleja linariifolia Collinsia parvifl Cordylanthus wrightii Castilleja scabrida var. rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Salicaceae Salicaceae Rosaceae Rosaceae Rosaceae Santalaceae Rosaceae Family Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Appendix A. Confi biogeographic notes, cont.

52 Annotated Checklist of Vascular Flora: Hovenweep National Monument is Mancos, Montezuma Co., CO Earle, (Baker, s.n. & Tracy RM) is St. George, Washington Co., UT (Jones 5110h US) noxious weed (List B). Native to Europe Comments is between Hatch’s & Wash Monticello, San Juan Co., UT (Eastwood s.n. CAS) is 2.5 miles N of Kanab, Kane Co., UT (Maguire, Maguire, & Piranian 12279 UTC) 1974 2001 locality Type 1973 2001 locality Type 2003 CO state Year Year doc. 2001 locality Type NA locality Type c name with taxonomic and (HOVE) 20010824 J (HOVE) (HOVE) 20010501 Q (HOVE) HOVE V-1 HOVE V-1 (HOVE) Source 20010503.04 (HOVE) Moran (2003) Pop. size UT CO Pres XPres X Unc Litzinger 301 X Unc Schelz Pres XPres Rare Schmidt 38 X Unc Schelz Pres X Rare Andersen Park stat. Pres X O Unc Schelz Rep O ? Schelz and RegEn RegEn RegEn Periph Intro Range UT/CO Periph RegEn PerF RegEn/ PerF NA/ PerF RegEn/ PerF Wide/ PerF Intro/ Life form PerF Wide/ PerF RegEn/ ax Colorado penstemon Abajo penstemon shortest penstemon undulate penstemon Dalmatian toadfl Common name Utah penstemon Palmer’s Palmer’s penstemon P. coloradoensis P. Synonyms/ notes Taxonomic

ssp. var. var. var. dalmatica lentus coloradoensis Penstemon palmeri Penstemon linarioides Penstemon lentus var. Penstemon breviculus Penstemon eatonii undosus ssp. Penstemon utahensis eglandulosus Species name

var. var. var. lentus Penstemon palmeri eglandulosus Penstemon lentus var. Penstemon breviculus Penstemon eatonii undosus UtahLinaria dalmaticagenistifolia Linaria Colorado Penstemon utahensis rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Family Scrophulariaceae Appendix A. Confi biogeographic notes, cont.

Chapter 6: Appendices 53 noxious weed (List C). Native to Eurasia Comments noxious weed (List B). Native to Eurasia C North America, may be introduced in UT. 1974 1975 1973 1975 CO state Year Year doc. 2002 1972 1973 CO state 1973 1973 2001 2003 Native to eld 54 eld c name with taxonomic and (HOVE) (HOVE) (HOVE) (HOVE) Source (HOVE) (HOVE) (HOVE) (HOVE) (HOVE) 20010824 P (HOVE) Andersen (2005) Pop. size UT CO Pres X X Com Litzinger 651 Pres XPres X X Unc Schelz 2121 Unc Schmidt 50 Pres X Unc Litzinger 744 Park stat. PresPres X X Unc X Schelz 2166 Unc Redfi Pres X O Unc Litzinger 309 Pres X OPres Unc X Schmidt 53 X Unc Schelz 2106 Pres X Unc Schelz Rep O ? Dewey & Wide Wide Wide Intro Range UT/CO Wide Wide Intro Wide Wide Sparse Wide PerF Periph/ AnnF Wide/ PerF Intro/ Life form PerG Wide/ Tree Wide/ Shrub Intro/ PerF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ ve-stamen Angel’s Angel’s trumpet pale wolfberry Shrub Wide/ false nightshade woolly mullein Common name broad-leaved cattail netleaf hackberry tamarisk Fendler’s Fendler’s cymopterus onion spring- parsley Fendler’s Fendler’s ground-cherry cut-leaf nightshade fi

var. var. Synonyms/ notes Taxonomic reticulata C. glomeratus P. fendleri, P. fendleri, P. P. hederaefolia fendleri T. pentandra T. orum var. var. Chamaesaracha coronopus Verbascum Verbascum thapsus Cymopterus fendleri Physalis hederifolia cordifolia Cymopterus bulbosus ramosissima

Species name orumtrifl Solanum var.

var. Physalis hederifolia Chamaesaracha coronopus Datura wrightii Datura wrightii pallidumLycium D. meteloides pallidum Lycium Utah Verbascum thapsus Colorado Typha latifoliaTypha latifolia Typha Celtis reticulata Celtis reticulataCymopterus acaulis C. laevigata fendleri fendleri Cymopterus bulbosus Solanum trifl Tamarix chinensisTamarix Tamarix rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Solanaceae Solanaceae Solanaceae Solanaceae Scrophulariaceae Family Typhaceae Ulmaceae (Celtidaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Solanaceae Tamaricaceae Appendix A. Confi biogeographic notes, cont.

54 Annotated Checklist of Vascular Flora: Hovenweep National Monument P. and Pinus parasitic on Juniperus osteosperma monophylla Colorado from Colorado according & to Weber Wittmann (2001) Comments parasitic on edulis NA 2006 Typically 2001 1975 New to 2003 Not known Year Year doc. 2001 Typically 1975 NA c name with taxonomic and Moran (2003) (HOVE) (HOVE) (HOVE) Moran (2003) Source (HOVE) (HOVE) Moran (2003) Pop. size UT CO Rep ? ? ? Schelz and Pres X Unc Fertig 22624 Pres X X Com Schelz 2167 Pres X X Rare Litzinger 476 Rep O ? Unc Schelz and Park stat. Pres X Unc Schelz 2164 Pres X XRep Unc O Litzinger 583 O ? Schelz and Wide Wide RegEn Periph NA Range UT/CO Wide Wide Wide AnnF Periph/ PerF Wide/ PerF RegEn/ PerF Wide/ PerF RegEn/ Life form PerF Wide/ PerF Wide/ PerF Wide/ at-seed orange kallstroemia juniper mistletoe Colorado Plateau spring-parsley basin white- cup spring- parsley sticky spring- parsley Common name fl lomatium prostrate vervain pinyon dwarf- mistletoe Synonyms/ notes Taxonomic L. simplex

ssp. ora ed (i.e., could be UT or CO). Kallstroemia parvifl Phoradendron juniperinum Cymopterus purpureus Lomatium triternatum platycarpum Cymopterus purpurascens Cymopterus newberryi Arceuthobium divaricatum

Species name ssp. var. ora Kallstroemia parvifl Phoradendron juniperinum Cymopterus purpureus purpureus Lomatium triternatum platycarpum Cymopterus purpurascens UtahCymopterus newberryi Colorado Verbena bracteataVerbena bracteata Verbena Arceuthobium divaricatum rmed, historical, and reported vascular plant taxa of Hovenweep National Monument, organized by family scientifi Zygophyllaceae Viscaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Family Verbenaceae Viscaceae Appendix A. Confi biogeographic notes, cont. X = documented with a voucher. O = documented by an observation. for the monument without state specifi ? = reported

Chapter 6: Appendices 55 Comments America CO BLM Sensitive USFWS 3C Type locality is Elk Type Mts, San Juan Co., UT (Rydberg & Garrett 9320 NY) Wide NA Sparse Periph RegEn NA NA Periph Wide Range (UT/CO) Wide NA Wide Wide RegEn Life form Common name Bailey’s yuccaBailey’s sea purslane Shrubtumble pigweed Periph/ woolly tidestromia PerF AnnF Wide/ AnnF Intro/IntroJones’ amsonia Native to tropical Periph/ PerF RegEn/ box elder Tree Wide/ Cutler’s milkweedCutler’s PerF RegEn/ bigseed milkweed PerF RegEn/ dwarf milkweedbroadleaf milkweed PerF PerF Periph/ Wide/ James’ cryptanth PerF Wide/ Fremont’s mahoniaFremont’s Shrub Wide/ climbing milkweed PerF Periph/ orange milkweed PerF Wide/ San Juan cryptanth PerF RegEn/

ssp. var. by some A. Synonyms/ notes Taxonomic A. negundo interius Included in involucrata auhors C. jamesii Funastrum cynanchoides heterophyllum interius terminalis Cladothrix lanuginosa lanuginosa T. ssp. Asclepias tuberosa ssp. Oreocarya pustulosa C. pustulosa Oreocarya suffruticosa

Species name

var. var. rmed) taxa of Hovenweep National Monument. interior pustulosa jamesii Utah Colorado Yucca baileyi Yucca Sesuvium verrucosum Sesuvium verrucosum Amaranthus albus Amaranthus albus Tidestromia lanuginosa lanuginosa Amsonia jonesii Amsonia jonesii Asclepias cutleri Asclepias tuberosa var. Asclepias involucrata Asclepias involucrata Asclepias latifoliaAsclepias Asclepias latifolia macrosperma Cryptantha cinerea var. Mahonia fremontii Mahonia fremontiiCryptantha cinerea var. Berberis fremontii Sarcostemma cynanchoides hartwegii Acer negundo Negundo aceroides Agavaceae Aizoaceae Amaranthaceae Amaranthaceae Apocynaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Boraginaceae Berberidaceae Boraginaceae Asclepiadaceae Family Aceraceae Appendix B. Potential (unconfi

56 Annotated Checklist of Vascular Flora: Hovenweep National Monument Type locality is S Type of Moab, Grand Co., UT (Rydberg & Garrett 9110 US) Comments USFWS: 3C Wide Wide Periph RegEn Periph NA NA Range (UT/CO) RegEn Wide Wide Wide Periph PerF Wide/NA PerF Wide/ Life form PerF Periph/ ower snowberry ower Shrub Wide/ ve-horn smotherweed AnnF Intro/Intro Native to Eurasia long-fl orachRydberg’s AnnF RegEn/ rufous-spine pricklypear hedgehog Simpson’s cactus blue elderberry Shrub Wide/ Common name bindweed heliotrope AnnFmatted tiquilia Wide/ PerF RegEn/ slenderspine pricklypear greenmollyfi PerF Wide/ Four Corners orachthinleaf orach AnnF RegEn/ AnnF Wide/ wing-nut cryptanth AnnFrecurved cryptanth Wide/ AnnF Wide/

Flora O.

in var.

caerulea, var. (2003) ssp. polyacantha polyacantha of North America (2003) Included with Opuntia polyacantha var. Flora of North America S. cerulea Includes var. simpsonii Synonyms/ notes Taxonomic Included with polyacantha triangularis

ssp. orus minor Symphoricarpos longifl Atriplex pachypoda californica var. Cryptantha pterocarya Euploca convolvulacea

var. Species name

var. spina rmed) taxa of Hovenweep National Monument, cont. orus trichophora rufi Utah Colorado Sambucus caerulea Sambucus coeruleaSymphoricarpos longifl S. nigra Atriplex argentea rydbergii Opuntia polyacantha var. Tiquilia latiorTiquilia Opuntia polyacantha var. Coldenia hispidissima Pediocactus simpsonii Pediocactus simpsonii Bassia americanaBassia hyssopifolia Kochia americana Bassia hyssopifolia Atriplex pleiantha Proatriplex pleiantha Atriplex prostrata Atriplex patula A. patula Cryptantha recurvata Cryptantha recurvata Heliotropium convolvulaceum Cryptantha pterocarya pterocarya Caprifoliaceae (Adoxaceae) Caprifoliaceae Chenopodiaceae Cactaceae Chenopodiaceae Boraginaceae (Ehrehtiaceae) Cactaceae Cactaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Boraginaceae Boraginaceae Appendix B. Potential (unconfi Family Boraginaceae

Chapter 6: Appendices 57 Comments Type locality is San Type San Juan Juan River, Co., UT (Brandegee s.n. GH) Wide Wide Wide Wide Range (UT/CO) Sparse Wide Wide Wide Wide Wide Periph RegEn Life form c aster c PerF Wide/ Common name Rio Grande seepwillow Shrub Wide/ blueleaf aster PerF Wide/ Pacifi leafy aster AnnF Wide/ winged pigweed poverty-weedNuttall’s AnnF AnnFbarbwire Russian thistle Wide/ Wide/ AnnF Intro/Intro Native to Eurasia sand wormwood PerF Wide/ American bugseed AnnF Wide/ common ragweed AnnF Wide/ seepweedWright’s longleaf brickellbush Shrub Periph/ Shrub Wide/ Brandegee’s siltbushBrandegee’s Shrub RegEn/

var.

var. var.

& var. ssp. ssp. ssp. B. emoryi cus, A. cus, ca glauca, Eurybia Synonyms/ notes Taxonomic Includes var. var. glauca adscendens, A. adscendens, Symphyotrichum ascendens Includes var. Includes var. americanum frondosum campestris rydbergii pacifi campestris pacifi longifolia A. artemisiifolia elatior cus Oligosporus Corispermum americanum Cycloloma atriplicifolium Monolepis nuttalliana Atriplex brandegeei Oligosporus pacifi Ambrosia artemisiifolia Species name rmed) taxa of Hovenweep National Monument, cont. brandegeei scouleriana Utah Colorado Baccharis salicina Baccharis salicina Aster glaucodes Eucephalus glaucus Herrickia glauca Aster frondosus Brachyactis frondosa Symphyotrichum Cycloloma atriplicifolia Monolepis nuttalliana Salsola paulseniiZuckia brandegeei var. Salsola paulsenii Artemisia campestris var. Aster ascendens Virgulaster ascendens A. chilensis Brickellia longifolia Brickellia longifolia B. longifolia Corispermum americanum Baccharis wrightii Baccharis wrightii Ambrosia artemisiifolia Compositae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi Family Chenopodiaceae (Asteraceae) Compositae (Asteraceae)

58 Annotated Checklist of Vascular Flora: Hovenweep National Monument is Paria, nidulum Kane Co., UT (Jones 5290a POM). Comments weed (List B). Native to Europe weed. CO state noxious weed (List B). Native to Europe Wide Wide Wide Wide Range (UT/CO) NA Life form Arizona thistle PerFpretty thistle Wide/NA locality of var. Type PerF Wide/ green rabbitbrush Shrub Wide/ greenish rabbitbrush Shrub Wide/ Common name spear-leaf rabbitbrushspear-leaf Shrub Wide/ Rothrock’s thistleRothrock’s PerF RegEn/ bull thistle PerF Intro/Intro CO state noxious spotted knapweed PerF Intro/Intro Utah state noxious

Flora

var. var. var.

var. in & Not var. ssp. (2006) consimilis, Flora of North rothrockii. Includes vars. arizonicum nidulum not Variety recognized in Intermountain Flora (1994), included in C. arizonicum bipinnatum of North America 2006) Ericameria nauseosa var. E. nauseosa oreophila, C. nauseosus oreophilus Synonyms/ notes Taxonomic recognized for Utah in America Lorandersonia linifolia micranthos, C. biebersteinii

ssp. ssp. orus orus pulchellum Cirsium calcareum var. viscidifl Chrysothamnus viscidifl Chrysothamnus nauseosus consimilis Chrysothamnus linifolius

Species name

var. var. rmed) taxa of Hovenweep National Monument, cont. orus orus pulchellum Cirsium rothrockii C. arizonicum Cirsium arizonicum Cirsium calcareum var. viscidifl Utah Colorado Chrysothamnus viscidifl Chrysothamnus nauseosus consimilis Cirsium vulgare Cirsium vulgare Centaurea maculosa Acosta maculosa C. stoebe Chrysothamnus linifolius Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi Family Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 59 Comments Type locality Type is Montezuma Canyon, E of Monticello, San Juan Co., UT (Rydberg & Garrett 9692 NY) Type locality is Type Willow Creek, San Juan Co., UT (Eastwood 48 CAS) Range (UT/CO) RegEn RegEn RegEn Wide Wide Wide Periph RegEn Life form Common name leafybase gumweed PerF RegEn/ Brandegee’s sandplantBrandegee’s AnnF RegEn/ slenderleaf daisy PerF RegEn/ bush encelia Shrub Wide/NA western goldenrodcrustweed PerF Wide/ slender cudweedlowland cudweed AnnF AnnF Wide/ AnnF Wide/NA Wide/ Rydberg’s gumweedRydberg’s PerF Periph/ erect gumweed PerF RegEn/ UT

Vars. Vars. (2006)

ssp. Includes

Included Flora of G. var. in pre- in var. (2006) (2006) . wetherillii. Flora of North G. squarrosa Flora of North Synonyms/ notes Taxonomic brandegeei. var. not recognized in America G. arizonica E. resinifera, frutescens frutescens 2003 editions of Flora E. sparsifolius aphanactis. in in America Included in hirsutula North America sparsifolius stenophylla Gnaphalium uliginosum Erigeron utahensis var. var.

var. Species name rmed) taxa of Hovenweep National Monument, cont. sparsifolius Utah Colorado Dicoria brandegeei Dicoria brandegeei D. canescens Encelia frutescens resinosa Erigeron utahensis var. Euthamia occidentalis Euthamia occidentalis Solidago occidentalis Glyptopleura marginata Gnaphalium exilifolium Gnaphalium palustre Gnaphalium palustre Grindelia aphanactis Grindelia aphanactis G. nuda Grindelia fastigiata Grindelia fastigiata Grindelia laciniata Grindelia arizonica Appendix B. Potential (unconfi Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

60 Annotated Checklist of Vascular Flora: Hovenweep National Monument Type locality is Type Willow Creek, San Juan Co., UT (Eastwood 45 CAS) Comments Type locality is 6.4 Type km S of Gateway, Mesa Co., CO (Tomb 648 TEX) Type locality is Type Cedar Canyon, Iron Co., UT (Garrett 6051 US) Type locality Type is desert S of Hanksville, Wayne Co., UT (Stanton 328 US). RegEn Periph RegEn RegEn Range (UT/CO) Wide NA Periph NA Wide Sparse AnnF Wide/NA Life form ower AnnF RegEn/ liform desert- liform aristate aster PerFfi dandelion RegEn/ copperweed Shrub Wide/ Arizona rush-pink PerF RegEn/ Common name western rush-pink PerF NA/ blue lettuce PerF Wide/ spindly goldenbush Shrub RegEn/ sand sunfl Rusby’s goldenbushRusby’s Shrub Wide/ poverty-weed PerF Wide/ Arizona hymenoxys PerF Wide/

Flora var.

var.

L. by some var. var. var. var. H. vars. not vacans, Dieteria canescens aristata glabrata Synonyms/ notes Taxonomic M. linearis, rigida, M. canescens Included in dianthopsis authors L. pulchella, oblongifolia, Mulgedium pulchellum Hesperodoria scopulorum, Chrysothamnus scopulorum Includes deserticola rusbyi axillaris; recognized in of North America (2006). arizonica

ssp.

var. Lygodesmia arizonica Lygodesmia Machaeranthera canescens aristata Lygodesmia Lygodesmia doloresensis pulchella Lactuca tatarica Tetraneuris acaulis Tetraneuris acaulis T.

var.

Species name

var. var. var. ora rmed) taxa of Hovenweep National Monument, cont. arizonica Utah Colorado Oxytenia acerosa Oxytenia acerosa Iva acerosa Malacothrix glabrata M. californica arizonica Machaeranthera canescens aristata Lygodesmia Lygodesmia grandifl Haplopappus scopulorum hirtellus Helianthus anomalus Haplopappus rusbyi Isocoma drummondii H. drummondii, I. Hymenoxys acaulis var. Lactuca tatarica pulchella Iva axillaris Iva axillaris I. axillaris Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 61 Comments Type locality is Type Cottonwood Wash, 1 1/2 miles N of San Juan Co., Bluff, UT (Maguire 13509 GH) Type locality is Type “Southern Utah” (Parry 109 GH) Range (UT/CO) Periph Periph Sparse Wide Wide Wide Wide Wide Wide Life form PerF Wide/ ower PerF Periph/ Common name woolly paperfl prenanthella AnnF Wide/ scale glandweedannual townsendia PerF Wide/NA AnnF Wide/ brown-plume wire- lettuce Hopi-tea PerF Periph/ scapose greenthread PerF Wide/ dogweed PerF Wide/NA broom groundsel PerFMissouri goldenrod Wide/ PerF Wide/ three-nerve goldenrod PerF Wide/ var. var.

Vars. Vars. var. ora, (2006). (2006) (2006). ora belenidium Flora of North Flora of North Flora of North Synonyms/ notes Taxonomic var. Vars. not recognized Vars. in America Vars. not recognized Vars. in America S. missouriensis missouriensis. not recognized in America Dyssodia pentachaeta S. velutina sparsifl ora Thelesperma megapotamicum Thelesperma subnudum Senecio spartioides Solidago missouriensis Stephanomeria paucifl

Species name

var. var. rmed) taxa of Hovenweep National Monument, cont. ora spartioides Utah Colorado Psilostrophe tagetina Psilostrophe tagetina Prenanthella exigua Prenanthella exigua exigua Lygodesmia Townsendia annuaTownsendia annua Townsendia Senecio spartioides var. Thelesperma megapotamicum Thelesperma subnudum subnudum Thymophylla acerosaThymophylla pentachaeta belenidium Dyssodia acerosa Solidago missouriensis Solidago velutina Solidago velutinaStephanomeria paucifl S. sparsifl Compositae (Asteraceae) Appendix B. Potential (unconfi Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

62 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments Type locality is Type near Epsom Creek [between Bluff & Mexican Hat], San Juan Co., UT (Eastwood 36 CAS) Type locality is E Type bank of Green River W of Moab (Cronquist 8994 NY) Range (UT/CO) Periph NA Wide Wide Wide Wide NA Wide Wide Life form PerF Wide/ Common name western tansy-mustard AnnF Wide/ resinbush Shrub RegEn/ Selby’s rockcressSelby’s Indian mustard PerFblack mustard Wide/ AnnF mustardhare’s-ear Intro/Intro Native to Asia AnnFCalifornia tansy- AnnFmustard Intro/Intro Intro/Intro Native to Europe Native to Europe Holboell’s rockcressHolboell’s PerF Wide/ golden crownbeard AnnF Wide/ hurtleaf woodyaster PerF RegEn/ western tansy-mustard AnnF Wide/ Fendler’s rockcressFendler’s PerF Wide/

.

var. D. var. by some var var. var. var. Synonyms/ notes Taxonomic Included in pinnata osmiarum authors stylosa retrofracta, B. holboellii secunda Machaeranthera tortifolia imberbis spatifolia, A. fendleri. not recognized Vars. in Holmgren et al. (2005)

ssp. intermedia lipes Descurainia californica Descurainia pinnata ssp. Ximenesia encelioides Boechera fendleri B. fendleri Boechera retrofracta A. holboellii Descurainia incisa fi

var. var. Species name rmed) taxa of Hovenweep National Monument, cont. lipes intermedia exauriculata fi imberbis Utah Colorado Vanclevea stylosaVanclevea Chrysothamnus Verbesina encelioides Verbesina var. Arabis selbyiBrassica juncea Boechera selbyi Brassica nigra Brassica juncea Conringia orientalis Brassica nigra Conringia orientalis Descurainia californica Descurainia pinnata Sinapis nigra var. Xylorhiza tortifolia var. Descurainia pinnata var. Arabis holboellii Arabis holboellii fendleri secunda Appendix B. Potential (unconfi Family Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

Chapter 6: Appendices 63 Comments valley of the Virgin near St. George, Co., UT Washington (Parry 19 NY) Wide Range (UT/CO) Wide Sparse Periph Periph Wide Wide RegEn Wide Periph Wide Life form PerF Wide/ three-square bulrush PerG Wide/ Common name Parish’s spikerushParish’s PerG Wide/NA common spikerush PerG Wide/ ovalpurse AnnF Wide/ prairie pepperwortSt. George pepperwort AnnF AnnF Wide/ Wide/NA locality is Type bladderpodFendler’s PerFsilver bladderpod Periph/ smoothleaf prince’s- PerFplume golden thelypody Wide/ golden sedge PerF RegEn/ PerG Wide/ dwarf draba AnnF Wide/ western tansy-mustard AnnF Wide/ E. &

var. var. E. polyphyllus, polyphyllus Synonyms/ notes Taxonomic Scirpus pungens var. Schoenoplectus pungens longispicatus, Schoenoplectus pungens uniglumis Includes erythropoda Hornungia procumbens

var. orum nelsonii ramosum Schoenoplectus pungens Lepidium densifl var. ludoviciana integrifolia macrostachya Hymenolobus procumbens Descurainia pinnata ssp.

var. orum Species name rmed) taxa of Hovenweep National Monument, cont. ramosum nelsonii georginum Utah Colorado Eleocharis parishii Scirpus pungens longispicatus Lepidium densifl var. Descurainia pinnata var. Lepidium lasiocarpum var. Physaria fendleriPhysaria ludoviciana Lesquerella fendleri Lesquerella Stanleya integrifolia Stanleya pinnata Thelypodiopsis aurea Thelypodiopsis aurea Carex aureaEleocharis palustris Eleocharis Carex aurea C. garberi Draba reptansHutchinsia procumbens Draba reptans Cyperaceae Cyperaceae Cruciferae (Brassicaceae) Appendix B. Potential (unconfi Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cyperaceae Cyperaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

64 Annotated Checklist of Vascular Flora: Hovenweep National Monument (2003) Flora of North Type locality is Type about 18 km N of Juab Co., Lynndyl, UT (Cronquist & Thorne 11839 NY) locality is Type near St. George, Co., UT Washington (Palmer s.n. US) Comments weed (List B). Native to Europe Not included for UT in America Wide RegEn NA Periph Periph Wide NA Wide Range (UT/CO) Wide Wide Periph RegEn PerF Wide/ Life form eld horsetail eld Fern Wide/ Utah dove-weed AnnF RegEn/ Great Basin centaury PerFwhite-margined swertia Wide/ thyme-leaved spurge AnnF Wide/ centauryBuckley’s AnnF Periph/ Common name Hall’s bluestemHall’s PerG Wide/ tiny spurge AnnF Wide/NA Nelson’s scouring-rushNelson’s Ferngreenleaf manzanita Sparse/ Shrub Wide/ big bluestem PerG Wide/ Russian-olive Tree Intro/Intro CO state noxious Arizona three-awn PerG Periph/ Cutler’s ephedraCutler’s fi Shrub RegEn/ E. E. and Euphorbia serpyllifolia C. arizonicum Synonyms/ notes Taxonomic Euphorbia micromeria Hybrid between laevigatum variegatum platyphylla Croton texensis Centaurium exaltatum Frasera albomarginata Chamaesyce serpyllifolia Andropogon hallii ssp. Andropogon gerardii Elaeagnus angustifolia Ephedra cutleri

var. Species name var. nelsonii x rmed) taxa of Hovenweep National Monument, cont. chrysocomus gerardii Utah Colorado Croton texensis Centaurium exaltatum Swertia albomarginata utahensis Centaurium calycosum Andropogon gerardii var. Andropogon gerardii var. Chamaesyce serpyllifolia Arctostaphylos patula Arctostaphylos patula Chamaesyce micromeria Equisetum Elaeagnus angustifolia Aristida arizonica Aristida arizonica Ephedra viridis viscida Equisetum arvense Equisetum arvense Euphorbiaceae Gentianaceae Gentianaceae Gentianaceae Gramineae (Poaceae) Gramineae (Poaceae) Euphorbiaceae Ericaceae Euphorbiaceae Equisetaceae Appendix B. Potential (unconfi Family Elaeagnaceae Gramineae (Poaceae) Ephedraceae Equisetaceae

Chapter 6: Appendices 65 Specimens from HOVE herbarium are from just outside the park. Comments Type locality Type is Springdale, Co., UT Washington (Jones 6075 POM) Periph Periph Wide Periph Wide Wide Range (UT/CO) Periph Periph Wide Wide NA Life form uffgrass PerG Wide/ New Mexican muhly PerG Wide/ fl little barley AnnG Wide/ annual wheatgrass AnnG Intro/Intro Native to Asia sixweeks muhly AnnG Sparse/ tufted lovegrass AnnG Wide/ thickspike wheatgrass PerG Wide/ Common name black gramaJones’ reedgrass PerG PerG Wide/ Wide/ sixweeks grama AnnG Wide/ desert saltgrass PerG Wide/ big sandreed PerG Periph/ M.

thsii var. var. barbata stricta

griffi var. var. albicans, A. rothrockii Includes reports of arsenei Agropyron triticeum dasystachyum; includes var. riparium, A. albicans var. albicans Synonyms/ notes Taxonomic Includes vars. & lanceolatus, A. dasystachyum ora Dasyochloa pulchella Muhlenbergia depauperata Muhlenbergia paucifl Eremopyrum triticeum barbatum eriopodum Calamagrostis scopulorum Species name rmed) taxa of Hovenweep National Monument, cont. ora Hordeum pusillum Critesion pusillum Erioneuron pulchellum Muhlenbergia depauperata Muhlenbergia paucifl Utah Colorado Eragrostis pectinacea Eragrostis pectinacea Eremopyrum triticeum Bouteloua eriopoda Chondrosum Calamagrostis scopulorum Elymus lanceolatus Elymus lanceolatus E. lanceolatus Bouteloua barbata Chondrosum Calamovilfa gigantea Distichlis spicata Distichlis stricta D. spicata Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix B. Potential (unconfi Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

66 Annotated Checklist of Vascular Flora: Hovenweep National Monument Type locality is near Type Milford, Beaver Co., UT (Jones 2106) HOVE herbarium are from just outside the park. Native to Eurasia and Africa Comments Periph Periph Periph Wide Wide Wide Sparse Wide Wide Range (UT/CO) Life form giant dropseedMormon needlegrass PerG PerG Wide/ Wide/ mesa dropseed PerG Wide/ prairie wedgegrass PerG Wide/ green bristlegrassIndiangrass AnnG Intro/Intro Native to Eurasia PerG Wide/ little bluestem PerG Wide/ water polypogon PerG Intro/Intro Specimens from weeping alkaligrass PerG Intro/NA Native to Eurasia ditch polypogon PerG Intro/Intro Common name witchgrassvine mesquite AnnG PerG Wide/ Wide/ bundle panicgrass PerG Wide/

x

var. ssp. var. var. S. obtusata obtusata Andropogon scoparius, S. scoparium neomexicanum semiverticillata Hybrid between monspeliensis P. Agrostis stolonifera Synonyms/ notes Taxonomic P. occidentale, P. tennessense, P. huachucae, P. Dichanthelium acuminatum fasciculatum, D. lanuginosum fasciculatum

var. exuosus avenaceum Sphenopholis obtusata Schizachyrium scoparium Polypogon viridis Agrostis occidentale Polypogon interruptus acuminatum Species name exuosusfl Sporobolus rmed) taxa of Hovenweep National Monument, cont. Sporobolus giganteus Sporobolus giganteus Stipa arida Stipa arida Achnatherum aridum Sporobolus fl Setaria viridisSorghastrum nutans Sorghastrum Sphenopholis Setaria viridis obtusata Puccinellia distans Schizachyrium scoparium Utah Colorado Polypogon semiverticillatus Panicum capillare Panicum capillare Panicum obtusum Panicum obtusum Polypogon interruptus Panicum acuminatum Dichanthelium Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix B. Potential (unconfi Family Gramineae (Poaceae)

Chapter 6: Appendices 67 Type locality is Type W of Comb Reef, San Juan Co., UT (Cronquist 9123 NY). UT BLM: Sensitive, USFWS: former C2 candidate Comments USFWS 3C NA Wide Wide LocEn Wide Wide Wide Range (UT/CO) Wide Periph RegEn Periph Life form PerG Wide/ ddleleaf AnnF Wide/ eld mint eld PerF Wide/ plains beebalmcamel-thorn AnnF milkvetchCronquist’s Periph/ PerF PerF LocEn/ Intro/Intro Native to Asia dwarf pennyroyalfi PerF Periph/ mountain rush PerG Wide/ Tracy’s rushTracy’s PerG Wide/ Common name toad rush AnnG Wide/ New Mexico feathergrass leafy fi namaHowell’s AnnF RegEn/ Torrey’s phaceliaTorrey’s AnnF Periph/

orma , J.

. var. & J. by var J. tracyi, canadensis Includes ssp. californicum Includes xiphoides Included in ensifolius montanus some authors saximontanus f brunnescens Synonyms/ notes Taxonomic ambiguus Includes vars. bufonius, occidentalis, halophilus, Mentha arvensis M. arvensis Juncus saximontanus Juncus ensifolius neomexicana Nama densum

var. var. var. Species name var. rmed) taxa of Hovenweep National Monument, cont. orum Mentha arvensis glabrata Monarda pectinata Monarda pectinata Alhagi maurorum Alhagi maurorum Astragalus cronquistii Astragalus cronquistii Hedeoma nanum montanus Juncus ensifolius Utah Colorado brunnescens Juncus ensifolius Nama densum Nama retrorsum Nama retrorsum parvifl Stipa neomexicana Hesperostipa Phacelia integrifolia Phacelia integrifolia Juncus bufonius Juncus bufonius Labiatae (Lamiaceae) Labiatae (Lamiaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Juncaceae Juncaceae Hydrophyllaceae Hydrophyllaceae Appendix B. Potential (unconfi Family Gramineae (Poaceae) Hydrophyllaceae Juncaceae

68 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments Type locality is Type ‘southern Utah’, Kane Co?, UT (Bishop s.n. GH) Type locality Type is Montezuma Canyon, San Juan Co., UT (Eastwood s.n. POM) Type locality is Type ‘southern Utah’, (Parry 32 ISC) USFWS: 3C Type locality is Type Grand Westwater, Co., UT (Jones s.n. POM) locality is Type Emery Green River, Co., UT (Jones s.n. POM) NA Wide RegEn RegEn Wide NA RegEn RegEn RegEn Range (UT/CO) RegEn NA Life form Common name Bishop’s milkvetchBishop’s PerF RegEn/ Hopi milkvetchstraggling milkvetch PerF PerF RegEn/ RegEn/ yellow milkvetch PerF Wide/ great rushy milkvetch PerF Wide/ Moenkopi milkvetch PerFNaturita milkvetch RegEn/ PerF RegEn/ Patterson’s milkvetchPatterson’s PerFlongstipe milkvetch RegEn/ PerF RegEn/ Eastwood’s milkvetchEastwood’s PerF RegEn/ Preuss’ milkvetch PerF RegEn/ gravel milkvetch AnnF Wide/NA

var.

var. by some avus Synonyms/ notes Taxonomic episcopus candicans Included in var. Included in var. preussii authors

var. var. avus fl A. Astragalus fl Astragalus lentiginosus palans Astragalus lonchocarpus Astragalus naturitensis Astragalus praelongus lonchopus Astragalus eastwoodiae

var.

Species name

var. var. avus rmed) taxa of Hovenweep National Monument, cont. latus avus Utah Colorado Astragalus fl Astragalus fucatus Astragalus fucatus Astragalus lentiginosus palans fl Astragalus lonchocarpus Astragalus moencoppensis Astragalus naturitensis Astragalus pattersonii Astragalus pattersonii Astragalus praelongus lonchopus Astragalus eastwoodiae Astragalus preussii var. Astragalus episcopus A. episcopus Astragalus sabulonum Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix B. Potential (unconfi Family Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

Chapter 6: Appendices 69 Comments Type locality is Type near St. George, Co., UT Washington (Parry 41 NY) Type locality Type is Springdale, Co., UT Washington (Jones 5259 POM) locality is 10 Type miles E of Zion Kane Co., UT NP, (Hitchcock 19013 isotype NY) Type locality is Type “Kanab” (but probably from farther east (Thompson s.n. GH) Type locality is Type Professor Valley Ranch, Grand Co., & Taylor UT (Welsh 14632 BRY) Periph Wide Wide Range (UT/CO) NA RegEn NA Periph Wide RegEn Wide Life form Common name blackbrush lupine AnnF Wide/ Nuttall’s licoriceNuttall’s PerF Wide/ white prairie-clover PerF Wide/ Zion sweetpea PerF RegEn/ Zion milkvetch PerF RegEn/ Great Basin lupine AnnF Wide/NA Kanab prairie-clover PerF RegEn/ woolly dalea PerF Periph/ silverleaf lupine PerF Wide/ Moab lupine PerF RegEn/ shortstem lupine AnnF Wide/

cutleri by some var. var. included in var. pusillus avescens Synonyms/ notes Taxonomic L. rubens, var. authors lepidota Petalostemon candidum, P. occidentale D. fl L. caudatus, argenteus argophyllus, L. caudatus

var.

ssp. var. var. var. zionis moabensis Lathyrus brachycalyx var. Lupinus pusillus rubens oligophylla Dalea lanata terminalis Lupinus argenteus var. Lupinus caudatus argophyllus

var. var. var.

var. var. Species name var. avescens rmed) taxa of Hovenweep National Monument, cont. zionis moabensis avescens Utah Colorado Lathyrus brachycalyx var. Glycyrrhiza lepidota Glycyrrhiza lepidota G. lepidota Astragalus zionis zionis Lupinus pusillus intermontanus Lupinus pusillus rubens Dalea fl fl Lupinus argenteus var. Dalea lanata terminalis Dalea oligophylla Dalea candida Lupinus brevicaulis Lupinus brevicaulis Lupinus caudatus argophyllus Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix B. Potential (unconfi Family Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

70 Annotated Checklist of Vascular Flora: Hovenweep National Monument Comments Type locality is near Type Wilson Mesa, Grand Co., UT (Rydberg & Garrett 8367 NY) America locality is near Type Emery Green River, Type locality is Ep- Type som Creek, San Juan Co., UT (Eastwood 21 CAS) Co., UT (Jones s.n. POM) Wide Periph NA Wide Wide Range (UT/CO) RegEn NA Wide Wide Wide RegEn Life form PerF RegEn/ PerF Wide/ Shrub RegEn/ ax AnnF Sparse/ owered Common name slenderleaf scurfpea PerF RegEn/ black locust Tree Intro/Intro Native to E North sand lily PerF RegEn/ Nevada onion PerF Wide/ silvery sophoraAmerican vetch PerFSan Juan onion PerF Wide/NA Wide/ PerF Wide/ rush scurfpea PerF RegEn/ large-fl breadroot starry false Solomon’s- seal small yellow fl narrow-leaf dunebroom Lambert’s locoweedLambert’s PerF Wide/

A.

var. lambertii var. Synonyms/ notes Taxonomic Vars. not recognized Vars. in Barneby (1989) Welsh et al. Welsh (2003) includes atrorubens cristatum Psoralea megalantha, Pediomelum megalanthum megalanthum Includes var. Includes var.

var. lifolia Vicia americana americana Psoralidium lanceolatum stellatum Pediomelum megalanthum Oxytropis lambertii

var. Species name var. lifoliafi Parryella rmed) taxa of Hovenweep National Monument, cont. bigelovii Utah Colorado Robinia pseudoacacia Robinia pseudoacacia Eremocrinum albomarginatum Sophora stenophylla Vicia americana americana Allium macropetalum Allium macropetalum Allium nevadense Allium nevadense Psoralidium junceum Psoralea juncea Psoralidium lanceolatum stenophyllum Oxytropis lambertii var. Smilacina stellata Maianthemum Linum australe Mesynium australe Parryella fi Pediomelum megalanthum Leguminosae (Fabaceae) Liliaceae (Anthericaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix B. Potential (unconfi Family Leguminosae (Fabaceae) Liliaceae (Convallariaceae or Ruscaceae) Linaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae)

Chapter 6: Appendices 71 Type locality is Type Paradox Valley, Montrose Co., CO Comments Specimen at HOVE from outside the locality park. Type Kane is Paria River, Co., UT (Cronquist 10202 MO) Not recognized for et al. UT by Welsh (2003) Type locality is Type Willow Creek, San Juan Co., UT (Eastwood s.n.) Type locality is Type Willow Creek, San Juan Co., UT (Eastwood 31 CAS) Wide Periph Wide Wide RegEn Wide Range (UT/CO) Periph NA RegEn Wide Wide NA Wide Life form PerF Wide/ PerF Wide/ PerF Wide/ ax PerF RegEn/ owered gaura owered AnnF Wide/ ower sandpuffs ower AnnF Wide/ Walker’s camissoniaWalker’s AnnF Wide/ lavender-leaf sundropslavender-leaf PerF Wide/ scarlet gaura PerF Wide/ small-fl Common name Paria evening-primrose PerF RegEn/ hairy-throat evening- primrose small-fl Utah yellow-fl trailing four-o’clock AnnF Periph/ gooseberry-leaf globemallow sand-verbenaWooton’s AnnF RegEn/ western evening- primrose wing-seed stickleaf PerF Wide/

var.

ssp. lavandulifolius C. hartwegii Synonyms/ notes Taxonomic Abronia micrantha, T. Abronia micrantha, T. pedunculatus S. grossulariifolia grossulariifolia wootonii T.

ssp. navajoensis wootonii Oenothera caespitosa ssp. Calylophus lavandulifolius Oenothera coronopifolia Tripterocalyx Tripterocalyx micranthus hirsutissima Nuttallia pterosperma Sphaeralcea grossulariifolia carneus Tripterocalyx var. Species name oramollis Gaura rmed) taxa of Hovenweep National Monument, cont. navajoensis wootonii Camissonia walkeri Camissonia walkeri Gaura coccineaGaura parvifl Gaura coccinea Utah Colorado Oenothera caespitosa var. Calylophus lavandulifolius Oenothera coronopifolia Linum subteres Allionia choisyi Tripterocalyx micranthus Tripterocalyx carneus Tripterocalyx var. Oenothera elata Oenothera elata Mentzelia pterosperma Sphaeralcea grossulariifolia Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Appendix B. Potential (unconfi Family Linaceae Nyctaginaceae Nyctaginaceae Nyctaginaceae Onagraceae Loasaceae Malvaceae

72 Annotated Checklist of Vascular Flora: Hovenweep National Monument Type locality is 37.9 Type miles SW of Green Emery Co., UT River, (Ownbey & Ownbey 2146 ) Type locality is Type “plains of the San Juan River in S Colorado or adjacent Utah” (Brandegee s.n.) locality Type is Springdale, Co., UT Washington (Jones 5247 NY) Type locality is Type “sand banks of the Green River”, UT (Cruetzfeldt s.n. NY) Comments Type locality is Type Armstrong & White canyons [Natural Bridges NM], San Juan Co., UT (Rydberg & Garrett 9410 NY) USFS R2 Sensitive Wide NA RegEn RegEn NA RegEn Wide Range (UT/CO) RegEn Sparse PerF RegEn/ Life form PerF RegEn/ San Rafael prickly- poppy slender bog orchid PerF Wide/ San Juan gilia PerF RegEn/ shrubby gilia PerF RegEn/ carmine gilia PerF giliaGunnison’s RegEn/ AnnF RegEn/ Common name northern bog orchid PerF Wide/ Bridges evening- primrose giant helleborine PerF Wide/

var. Flora . Vars. not . Vars. Platanthera ora recognized in of North America (2002) Gilia congesta frutescens sparsifl Synonyms/ notes Taxonomic Includes aquilonis, P. huronensis frutescens Ipomopsis congesta ssp. Limnorchis ensifolia Platanthera Limnorchis hyperborea ora Species name ora rmed) taxa of Hovenweep National Monument, cont. arenicola frutescens sparsifl Argemone corymbosa var. Gilia haydenii Gilia haydenii Aliciella haydenii Gilia subnudaIpomopsis congesta var. Ipomopsis gunnisonii Ipomopsis gunnisonii Gilia gunnisonii Aliciella subnuda Utah Colorado Habenaria sparsifl var. Oenothera longissima Oenothera longissima Epipactis gigantea Epipactis gigantea Habenaria hyperborea Papaveraceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Orchidaceae Appendix B. Potential (unconfi Family Onagraceae Orchidaceae Orchidaceae

Chapter 6: Appendices 73 Comments Type locality is San Type San Juan Juan River, Co., UT (Brandegee 1172 GH) Willow Creek, San Juan Co., UT (Eastwood 10 CAS) Type locality is Type Emery Green River, Co., UT (Jones s.n. POM) Type locality is Type near Butler Wash, San Juan Co., UT (Eastwood 134 CAS) NA Wide Wide Wide Range (UT/CO) Wide RegEn Wide RegEn RegEn Wide RegEn NA RegEn Life form PerF Wide/ AnnF Wide/ Shrub RegEn/ AnnF Wide/ AnnF RegEn/ Shrub RegEn/ Shrub RegEn/ y wild y owered gilia owered PerF Wide/ Common name winged wild buckwheat thorny milkwort Shrubthorny milkwort Periph/ Shrubcushion milkwort Wide/NA locality is Type PerF Wide/ carpet phlox PerF Wide/ rimrock wild buckwheat nodding wild buckwheat San Rafael gilia PerF RegEn/ bent wild buckwheat AnnF wild Hooker’s buckwheat RegEn/ long-fl Grand Valley desert Grand Valley trumpet butterfl buckwheat San Juan wild buckwheat

var. alatum glabrata, ora var. viscidula (2005) & Flora of North Synonyms/ notes Taxonomic Vars. not recognized Vars. in America Includes ssp. hoodii, acanthoclada P. intermontana P. subspinosa

oralongifl Gilia

exum var. var. Eriogonum cernuum Eriogonum corymbosum orbiculatum Phlox hoodii Eriogonum fusiforme Eriogonum leptocladon ramosissimum

oralongifl Ipomopsis

Species name atum var. var. var. var. var. var. exumfl Stenogonum rmed) taxa of Hovenweep National Monument, cont. cernuum fusiforme Utah Colorado Eriogonum alatum Pterogonum alatumEriogonum cernuum E. alatum var. Polygala acanthoclada acanthoclada Polygala acanthoclada intricata Polygala subspinosa Polygala subspinosa subspinosa P. Eriogonum corymbosum orbiculatum Ipomopsis roseata Ipomopsis roseataPhlox hoodii canescens Gilia roseata Eriogonum fl Eriogonum hookeri Eriogonum hookeri Ipomopsis longifl Eriogonum infl var. Eriogonum leptocladon papiliunculi Eriogonum leptocladon ramosissimum Polygonaceae Polygonaceae Polygalaceae Polygalaceae Polygalaceae Polygonaceae Polemoniaceae Polemoniaceae Polygonaceae Polygonaceae Appendix B. Potential (unconfi Family Polemoniaceae Polygonaceae Polygonaceae Polygonaceae

74 Annotated Checklist of Vascular Flora: Hovenweep National Monument is near Type locality is Bluff, locality is Bluff, Type San Juan Co., UT s.n. CAS) (Wetherill Comments Type locality of var. locality of var. Type longilobum Price, Carbon Co., UT (Jones 5590j POM) Type locality is Type along the San Juan San Juan Co., River, UT (Eastwood 124 CAS) Wide RegEn Wide Wide Wide RegEn Wide Wide Range (UT/CO) Wide Wide Wide Shrub Wide/ Life form PerF Wide/ AnnF RegEn/ AnnF Wide/ alcove columbine PerF RegEn/ marsh buttercup PerF Wide/ birchleaf buckthorn Shrub Wide/NA dwarf mountain mahogany Common name Shockley’s wild Shockley’s buckwheat sulfur wild buckwheat PerF wild Wetherill’s Wide/ buckwheat sausageleaf talinum PerFtiny mousetail Wide/NA AnnF Sparse/ slender lip-fern Fern Wide/ Apache plumebush oceanspray Shrub Shrub Wide/ Wide/ smooth wild buckwheat

Flora var.

var. & Vars. not Vars. R. cymbalaria cymbalaria. recognized in of North America (1997) intricatus Synonyms/ notes Taxonomic Includes vars. longilobum shockleyi brevifolius

var. ssp. longilobum Eriogonum umbellatum umbellatum Halerpestes cymbalaria saximontana Stenogonum salsuginosum var.

Species name var. rmed) taxa of Hovenweep National Monument, cont. Utah Colorado Aquilegia micrantha Aquilegia micrantha Rhamnus betulifoliaCercocarpus intricatus Cercocarpus intricatus C. ledifolius Fallugia paradoxa Fallugia paradoxa Frangula betulifolia Eriogonum umbellatum umbellatum Eriogonum wetherillii Eriogonum wetherillii Ranunculus cymbalaria Myosurus minimus Myosurus minimus Cheilanthes feei brevifoliumTalinum Cheilanthes feei Phemeranthus Holodiscus dumosus Holodiscus discolor Eriogonum shockleyi Eriogonum shockleyi Eriogonum salsuginosum Ranunculaceae (Helleboraceae) Rhamnaceae Rosaceae Rosaceae Polygonaceae Polygonaceae Ranunculaceae Ranunculaceae Polypodiaceae (Adiantaceae) Portulacaceae Rosaceae Polygonaceae Appendix B. Potential (unconfi Family Polygonaceae

Chapter 6: Appendices 75 eld Co., UT Comments Type locality is Type Aquarius Plateau, Garfi 462 GH) (Ward Wide Wide Range (UT/CO) RegEn NA Wide Wide RegEn Periph Periph Wide Wide Wide RegEn Wide Wide Life form Tree Wide/ ower AnnF Wide/ ower PerF Wide/ Common name cleavers AnnF Wide/ rock spiraea PerF Wide/ Colorado bedstrawred bluets PerF RegEn/ PerF Periph/ yellow willow Shrub Wide/ beardlip penstemon PerFdusty penstemon Wide/ PerF RegEn/ beaked penstemon PerF Wide/ annual paintbrushyellow monkeyfl AnnF Wide/ Goodding’s black Goodding’s willow fendlerbush Shrub Wide/ reddish monkeyfl veined penstemon PerF RegEn/ bush penstemon PerF Wide/NA dwarf lousewort PerF Wide/

minor watsonii vallicola var. ssp. var. var. Synonyms/ notes Taxonomic Spiraea caespitosa P. bridgesii P. rupicola Leiostemon ambiguus

var. orus torreyi Galium spurium Galium coloradoense Salix lutea S. rigida Petrophyton caespitosum Penstemon barbatus var. Penstemon comarrhenus Penstemon rostrifl Penstemon angustifolius venosus Pedicularis centranthera

var.

var. Species name orum var. rmed) taxa of Hovenweep National Monument, cont. orus coloradoense barbatus laevissimus Utah Colorado Galium aparine echinospermon Galium multifl var. Houstonia rubra Salix eriocephala watsonii Salix gooddingii Salix gooddingii S. nigra Petrophyton caespitosum Penstemon comarrhenus Penstemon rostrifl Fendlera rupicolaCastilleja exilis Fendlera falcataMimulus guttatus rupicola F. Castilleja minor Mimulus guttatus C. minor Mimulus rubellus Mimulus rubellus Penstemon barbatus var. Penstemon angustifolius venosus Pedicularis centranthera Penstemon ambiguus var. Rubiaceae Rubiaceae Rubiaceae Salicaceae Salicaceae Appendix B. Potential (unconfi Family Rosaceae Scrophulariaceae Scrophulariaceae Saxifragaceae (Hydrangeaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae

76 Annotated Checklist of Vascular Flora: Hovenweep National Monument . Duane Atwood Comments Type locality Type is Mancos, Montezuma Co., Earle, & CO (Baker, 76 NY) Tracy “Southern Utah” (Parry 85) Native to C North America Range (UT/CO) RegEn Periph Wide Wide Wide Periph Wide Sparse could be considered as potentially occurring in could be considered Life form PerF RegEn/ PerF Wide/ ah, and Montezuma County, Colorado), but have ah, and Montezuma County, Common name Natural Bridges penstemon water speedwell PerF Intro/Intro Native to Europe lomatiumParry’s PerFthicket-creeper Wide/NA locality is Type Shrub Wide/ rock spring-parsley PerF Wide/ blunt-leaf spike-moss Fern wolfberryAnderson’s long-leaf ground- Wide/ Shrubcherry Wide/NA James’ potatobuffalobur PerF Periph/ AnnF Intro/ southern cattail PerG Wide/ mutica var. Synonyms/ notes Taxonomic petraea (Albee et al. 1988), and records of species from just outside the monument in Hovenweep National of species from (Albee et al. 1988), and records sonorae Penstemon strictiformis Veronica anagallis- Veronica aquatica Parthenocissus vitacea var. Aletes petraeus C. petraeus, Pteryxia

Species name Atlas of the Vascular Flora of Utah Atlas of the Vascular var. rmed for the monument. Species on this list are derived from a checklist of potential species for NCPN parks prepared by Dr. N by Dr. a checklist of potential species for NCPN parks prepared derived from rmed for the monument. Species on this list are rmed) taxa of Hovenweep National Monument, cont. Utah Colorado Penstemon strictiformis Veronica anagallis- Veronica aquatica Selaginella mutica Selaginella mutica S. mutica Lomatium parryi Parthenocissus vitacea Lycium andersonii Lycium Physalis longifolia Physalis virginiana Solanum jamesiiSolanum rostratum Solanum jamesii Solanum rostratum Typha domingensisTypha domingensis Typha Cymopterus terebinthinus petraeus Appendix B. Potential (unconfi Family Scrophulariaceae Scrophulariaceae Selaginellaceae Umbelliferae (Apiaceae) Vitaceae Solanaceae Solanaceae Solanaceae Solanaceae Typhaceae Umbelliferae (Apiaceae) the monument, but are not included here. the monument, but are The vascular plant species listed in this table are known from the vicinity of Hovenweep National Monument (San Juan County, Ut the vicinity of Hovenweep National Monument (San Juan County, known from The vascular plant species listed in this table are University in 2001, the of Brigham Young Appendix C (especially those collected by William just outside Hovenweep) Monument herbarium. Some species from Litzinger from not previously been reported or confi been reported not previously

Chapter 6: Appendices 77 . C. . . . . ed ed ed ed ed ed avoculata avoculata A. blitoides Bassia scoparia Amsonia jonesii C. fl C. fl C. osterhoutii . Type locality is . Type ed specimen of avoculata based on a misidentifi specimen of based on a misidentifi specimen of Type locality is Palisades, Type Mesa Co., CO (Crandall s.n. RM) based on a misidentifi specimen of based on a misidentifi specimen of USFWS 3C (2003) based on a specimen from HOVE herbarium (Litzinger 739) from outside the park. Moran (2003) based on a misidentifi (2003) based on a specimen from HOVE herbarium (Litzinger 737) collected outside the park based on a misidentifi specimen of fl Grand Co., Courthouse Wash, UT (Eastwood s.n. CAS) based on a misidentifi specimen of UT CO Comments FalsRep O Report from HOVE herbarium FalsRep O Report from HOVE herbarium Park stat. FalsRep O Report from HOVE herbarium FalsRep O Report from HOVE herbarium FalsRep O Reported by Schelz & Moran FalsRep O Report from Schelz & FalsRep O Reported by Schelz & Moran FalsRep O Report from HOVE herbarium FalsRep O Report from HOVE herbarium Range NA NA Wide RegEn Wide RegEn Wide RegEn RegEn Life form AnnF Periph/ AnnF Wide/ PerF Wide/ PerF RegEn/ PerF Wide/ PerF RegEn/ PerF Wide/ PerF RegEn/ PerF RegEn/ ower Common name California amaranth Torrey’s Torrey’s amaranthus common dogbane Paradox cryptanth showy milkweed slender cryptanth whorled milkweed long-fl cryptanth Baker’s Baker’s cryptanth

& Synonyms/ notes Taxonomic angustifolium Includes vars. cannabinum, glaberrimum, A. galioides ora Apocynum cannabinum Oreocarya paradoxa Asclepias speciosa Cryptantha tenuis Asclepias subverticillata Oreocarya bakeri Oreocarya longifl Species name ora Amaranthus californicus Amaranthus torreyi Apocynum cannabinum Cryptantha paradoxa Utah Colorado Asclepias speciosa Cryptantha tenuis Asclepias subverticillata Cryptantha bakeri Cryptantha longifl Family Amaranthaceae Amaranthaceae Apocynaceae Boraginaceae Asclepiadaceae Boraginaceae Asclepiadaceae Boraginaceae Boraginaceae Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument. Appendix C. Rejected (falsely reported

78 Annotated Checklist of Vascular Flora: Hovenweep National Monument L. C. ed var. var. var. var. . ed ed cuneata A. gardneri A. argentea L. ruderale berlandieri ed specimen of ed specimen of and var. and var. . Native to Eurasia var. var. based on a misidentifi specimen of rydbergii Moran (2003) based on a misidentifi based on a misidentifi specimen of (2003) based on HOVE specimen (Litzinger 773) collected outside the park. (2003) based on HOVE specimen (Litzinger 821) collected outside the park. utahensis Moran (2003) based on a misidentifi ruderale al. (2008), but HOVE is well outside of the known range of this taxon. Report probably based on var. eastwoodiae. (2003), but HOVE is well outside the known range of this species. Native to Eurasia (2003) based on misidentifi specimens of album UT CO Comments FalsRep O Report from HOVE herbarium FalsRep O O Reported by Schelz & FalsRep O Report from HOVE herbarium FalsRep ? ? Reported by Schelz & Moran FalsRep ? ? Reported by Schelz & Moran Park stat. FalsRep O Reported by Schelz & Ques? O Loh et Reported by Von Ques? O Reported by Schelz & Moran FalsRep O Reported by Schelz & Moran Wide Range Intro Wide Wide Wide Wide Wide Intro Periph Life form PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ PerF Periph/ Shrub Periph/ Common name southwestern stoneseed yellow beeplant Rocky Mountain beeplant wedge orach AnnF Wide/ Fremont’s Fremont’s goosefoot gromwell AnnF Intro/ Fendler’s Fendler’s sandwort New Mexico saltbush garden orach AnnF Intro/ Synonyms/ notes Taxonomic C. fremontii orum Chenopodium fremontii Buglossoides arvensis Lithospermum multifl Eremogone fendleri Atriplex hortensis

Species name var. orum fendleri Chenopodium fremontii fremontii Lithospermum arvense Cleome lutea Cleome lutea Cleome serrulata Cleome serrulata Arenaria fendleri var. Atriplex truncata Atriplex truncata Utah Colorado Lithospermum multifl Atriplex hortensis Atriplex obovata Atriplex obovata Chenopodiaceae Family Boraginaceae Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) Caryophyllaceae (Alsinaceae) Chenopodiaceae Boraginaceae Chenopodiaceae Chenopodiaceae Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

Chapter 6: Appendices 79 C. A. A. ed ed .

. albula. albula. A. ludoviciana A. ludoviciana var. var. var. . . ed specimen of ed specimen of ed specimen of ed specimens of ed specimens of . albula albula (2003) based on a specimen in HOVE herbarium (Litzinger 890) from outside the park. (2003), but HOVE is well beyond the known range of this species. Native to C North America. Moran (2003) based on a misidentifi Gnaphalium exilifolium Moran (2003) based on a misidentifi herbarium based on a misidentifi ludoviciana ludoviciana (2003) based on misidentifi specimens of var. herbarium based on misidentifi Chaetopappa ericoides (2003) based on misidentifi specimens of var. stevioides herbarium based on misidentifi UT CO Comments FalsRep O Reported by Schelz & Moran Ques? ? ? Reported by Schelz & Moran FalsRep O Reported by Schelz & FalsRep O Reported by Schelz & FalsRep ? ? Reports from HOVE Park stat. FalsRep O O Reported by Schelz & Moran FalsRep O Reports from HOVE FalsRep O Reported by Schelz & Moran FalsRep O Reports from HOVE Range Wide Wide Wide Wide Periph Wide Wide Wide Wide Life form PerF Wide/ PerF Sparse/ PerF Wide/ PerF Wide/ PerF Wide/ Common name cutleaf agoseris Louisiana wormwood giant ragweed AnnF Intro/ low pussytoes PerF Wide/ tarragon PerF Wide/ Louisiana wormwood Louisiana wormwood alkali aster PerF Wide/ hoary dusty- maiden

ssp.

& var. ora Flora of Synonyms/ notes Taxonomic A. ludoviciana candicans A. parvifl Vars. not Vars. recognized in Cronquist (1994) and North America (2006) montana Includes var. Includes var. douglasii

. da ssp. ssp. ssp. orus laciniata Agoseris glauca var. Antennaria dimorpha Oligosporus dracunculus glaucus Artemisia ludoviciana incompta Artemisia ludoviciana ssp latiloba Artemisia ludoviciana ludoviciana Chaenactis douglasii paucifl

da trifi Ambrosia var. var. var. var. orus Almutaster Species name laciniata Agoseris glauca var. Ambrosia trifi Antennaria dimorpha Artemisia dracunculus glauca Utah Colorado Artemisia ludoviciana incompta Artemisia ludoviciana latiloba Artemisia ludoviciana ludoviciana Chaenactis douglasii Aster paucifl Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

80 Annotated Checklist of Vascular Flora: Hovenweep National Monument

. C. var.

var. var. ed ed H. villosa minor. minor. gnaphalodes . . Type locality is . Type C. nauseosus C. nauseosus var. ) based on a var. ed specimen of ed specimen ed foliosa C. villosa gnaphalodes Type locality is W of Leeds Type along road to Silver Reef Campgrounds, Washington Co., UT (Semple & Heard 7859 NY) Moran (2003) based on a misidentifi var. var. misidentifi of Moran (2003, as nauseosus (2003) based on a specimen from HOVE herbarium (Litzinger 373) from outside the park. (2003) based on a specimen from HOVE herbarium (Litzinger 766) from outside the park. CO state noxious weed (List C). Native to Eurasia gnaphalodes Canaan Ranch, Kane Co., UT (Jones 6066c POM) based on a misidentifi specimen of based on a misidentifi specimen of UT CO Comments FalsRep O Reported by Schelz & Park stat. FalsRep O Reported by Schelz & FalsRep O Reported by Schelz & Moran FalsRep O Reported by Schelz & Moran FalsRep O Report from HOVE herbarium FalsRep O Report from HOVE herbarium Range NA RegEn Wide Intro NA RegEn Life form Shrub RegEn/ PerF RegEn/ Shrub Wide/ Shrub Periph/ Shrub RegEn/ Common name sand rabbitbrush Zion goldenaster lanceleaf rabbitbrush pretty rabbitbrush chicory PerF Intro/ dune rabbitbrush

var. var. var. Ericameria nauseosa arenaria Synonyms/ notes Taxonomic H. villosa foliosa, H. zionensis Ericameria pulchella, Lorandersonia baileyi, C. baileyi Ericameria nauseosa turbinata

var. var. orus Chrysopsis zionensis Chrysothamnus viscidifl lanceolatus Cichorium intybus Chrysothamnus nauseosus turbinatus

var. Species name var. var. var. orus Chrysopsis zionensis Chrysothamnus nauseosus arenarius Chrysothamnus viscidifl lanceolatus Utah Colorado Cichorium intybus Chrysothamnus nauseosus turbinatus Chrysothamnus pulchellus baileyi Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

Chapter 6: Appendices 81 . G. S. . . ed ed collected oribunda fl ed specimen ed tenuifolia longilobus ed specimens of Hymenoxys Stephanomeria var. var. ed specimen of ed specimen of var. var. var. Aster ascendens (2003, without variety) based on a misidentifi of outside the park. (2003), but HOVE is well beyond the known range of locality is this species. Type Co., Springdale, Washington UT (Jones 5249a US). (2003), but HOVE is outside the known range of this locality is CisCo., taxon. Type Grand Co., UT (Jones s.n. US) Moran (2003) based on a misidentifi Moran (2003) based on a misidentifi dougalsii laciniata (2003) and Von Loh et al. (2003) and Von (2008), but all records in HOVE herbarium are based on misidentifi G. sarothrae based on a misidentifi specimen of richardsonii based on a misidentifi specimen of tenuifolia UT CO Comments FalsRep O Reported by Schelz & Moran Ques? ? ? Reported by Schelz & Moran Ques? ? ? Reported by Schelz & Moran Park stat. FalsRep O Reported by Schelz & FalsRep O Reported by Schelz & FalsRep O O Reported by Schelz & Moran FalsRep O Report from HOVE herbarium FalsRep O Report from HOVE herbarium Range NA Wide RegEn Wide Wide Periph RegEn Wide Life form PerF RegEn/ PerF Wide/ PerF Wide/ Shrub Wide/ PerF NA/ PerF Periph/ Common name Dixie rock- daisy aspen daisy PerF Wide/ curly-cup gumweed desert bahia PerF RegEn/ many-headed groundsel thread snakeweed Colorado hyalineherb rush skeleton- plant

Flora of (2006) desertorum Synonyms/ notes Taxonomic Vars. not recog- Vars. nized in North America (2006) Bahia nudicaulis var. Vars. not Vars. recognized in Flora of North America

var. var. var. lifolius Erigeron speciosus Grindelia squarrosa serrulata Platyschkuhria integrifolia desertorum Senecio multicapitatus Gutierrezia microcephala parvulus Hymenopappus fi Lygodesmia Lygodesmia juncea

var. var. Species name var. var. Erigeron speciosus Perityle tenella macranthus Grindelia squarrosa serrulata Platyschkuhria integrifolia desertorum Senecio spartioides multicapitatus Utah Colorado Gutierrezia microcephala Lygodesmia Lygodesmia juncea Family Compositae (Asteraceae) (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

82 Annotated Checklist of Vascular Flora: Hovenweep National Monument C. ed

. ed . Native to Native to T. incana T. S. tenuifolia collected outside ed specimen of ed specimen of ed specimen of var. ed specimen of var. tenuifolia. Moran (2003) based on a misidentifi Moran (2003) based on a misidentifi Lactuca serriola Europe pentagona the park. microphyllum. herbarium based on a misidentifi (2003) based on a specimen from HOVE herbarium (Litzinger 352) from outside the park. based on a misidentifi specimen of var. southwestern US, but not Utah or Colorado. (2003) based on misidentifi specimens of (2003) based on specimen from HOVE herbarium (Litzinger 698) from outside the park. (2003) based on specimen from HOVE herbarium (Litzinger 820) from outside the park. UT CO Comments FalsRep O Reported by Schelz & FalsRep O O Reported by Schelz & Park stat. FalsRep O Report from HOVE FalsRep O Reported by Schelz & Moran FalsRep O Reported by Schelz & Moran FalsRep O O Reported by Schelz & Moran FalsRep O Reported by Schelz & Moran Wide Range Intro NA Wide NA Periph Wide Life form AnnF Intro/ Shrub RegEn/ AnnF Wide/ PerF NA/NA FalsRep O Report from HOVE herbarium PerF Wide/ PerF Wide/ AnnF Wide/ AnnF NA/ atglobe Common name spiny-leaf sow- thistle spiny greasebush white-plume wire-lettuce Wright’s Wright’s wirelettuce showy townsendia rough mule’s- ears rough cocklebur fl dodder

ssp. (2006). scabra orifera Synonyms/ notes Taxonomic Forsellesia meionandra S. exigua exigua T. fl ssp. X. pensylvanicum, X. saccharatum; vars. not recognized in Flora of North America Cuscuta umbellata Stephanomeria exigua Xanthium strumarium Grammica umbellata

var. var. Species name orifer Sonchus asper Sonchus asper Stephanomeria exigua Glossopetalon spinescens meionandrum Utah Colorado Stephanomeria wrightii Townsendia Townsendia fl Wyethia scabra Wyethia scabra Scabrethia scabra Xanthium strumarium canadense Family Compositae (Asteraceae) Compositae (Asteraceae) Crossosomataceae (Celastraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Convolvulaceae (Cuscutaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

Chapter 6: Appendices 83 L. L. . Native . jonesii. jonesii from outside var. var. ed specimens of ed specimens of ed specimens of ed specimens of . gracilipes (2003), but HOVE is well beyond the known range of this species (2003), but HOVE is well beyond the known range of this species. Native to Europe (2003) based on a specimen in HOVE herbarium (Litzinger 582) from outside the park. herbarium were based on misidentifi Schoenocrambe linifolia (2003), but HOVE is outside the known range of this taxon. herbarium were based on misidentifi var. montanum to Asia herbarium were based on misidentifi the park herbarium were based on misidentifi montanum UT CO Comments Ques? O Reported by Schelz & Moran Ques? O O Reported by Schelz & Moran FalsRep O Reported by Schelz & Moran FalsRep O Previous reports from HOVE Park stat. Ques? ? ? Reported by Schelz & Moran FalsRep O Previous reports from HOVE FalsRep O Previous reports from HOVE FalsRep O Previous reports from HOVE Range NA Intro Wide Wide Wide NA Intro Sparse Life form PerF RegEn/ PerF Wide/ PerF Wide/ PerF Wide/ AnnF Intro/ AnnF Periph/ ower PerF Wide/ ower eld Common name twinpod blunt-leaf yellowcress cousin thelypody pale madwort AnnF Intro/ sand dune wallfl shy wallfl fi pepperwort poor-man’s poor-man’s pepperwort Newberry’s Newberry’s

var. var. var. Included capitatum Synonyms/ notes Taxonomic T. rhomboideum T. E. asperum asperum, E. capitatum purshii. in var. in Holmgren et al. (2005) P. newberryi P. racemosa Rorippa curvipes Alyssum alyssoides Erysimum asperum Erysimum inconspicuum Neolepia campestris Lepidium virginicum

var. Species name var. ne curvipes affi Rorippa curvipes var. Alyssum alyssoides purshii Erysimum asperum Erysimum inconspicuum Lepidium campestre Utah Colorado Thelypodium integrifolium var. Lepidium virginicum medium Physaria newberryi Cruciferae (Brassicaceae) Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

84 Annotated Checklist of Vascular Flora: Hovenweep National Monument . . ed . . . and orum Type Type ed ed ed ed ed nelsonii . densifl x Q. welshii var. S. maritimus C. praegracilis S. utahensis Lepidium E. ed specimen of orum based on a misidentifi specimen of Atriplex confertifolia. locality is upper valley of the Virgin, Kane Co., UT (Siler s.n. ISC) Moran (2003) based on a misidentifi based on a misidentifi specimen of based on misidentifi specimen of (2003) based on a collection from HOVE herbarium (Litzinger 712) from outside the park. based on a misidentifi specimen of Native to Europe (2003) based on misidentifi specimens of Mahonia repens (2003) based on a specimen from HOVE herbarium (Litzinger 724) collected outside the park based on a misidentifi specimen of densifl UT CO Comments FalsRep O Report from HOVE herbarium FalsRep O Report from Schelz & FalsRep O Report from HOVE herbarium FalsRep O Report from HOVE herbarium FalsRep O Report from Schelz & Moran FalsRep O Report from HOVE herbarium Park stat. FalsRep O O Report from Schelz & Moran Ques? O Reported by Schelz & Moran FalsRep O Report from HOVE herbarium Wide NA Wide Wide Range Wide Wide Wide RegEn Intro Shrub RegEn/ Fern Wide/ Life form PerF Wide/ Shrub Wide/ PerF RegEn/ AnnF Intro/ eld roundleaf buffaloberry pale bulrush PerG Sparse/ smooth scouring-rush Common name Liddon sedge PerG Wide/ shorthorn spurge turbinella live- oak felwort PerF Wide/ Moab thelypody fi pennycress

var. var. pallidus E. kansanum E. robusta, brachycera robusta Synonyms/ notes Taxonomic Hippochaete laevigata Tithymalus Tithymalus brachyceras Quercus turbinella Thelypodium integrifolium Species name gracilipes Shepherdia rotundifolia Equisetum laevigatum Scirpus pallidus Scirpus pallidus S. atrovirens Euphorbia brachycera Quercus turbinella Thelypodium integrifolium var. Carex petasata Carex petasata Swertia perennis Swertia perennis Utah Colorado Thlaspi arvense Thlaspi arvense Elaeagnaceae Equisetaceae Cyperaceae Euphorbiaceae Fagaceae Family Cruciferae (Brassicaceae) Cyperaceae Gentianaceae Cruciferae (Brassicaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

Chapter 6: Appendices 85 . B. . Utah ed ed ed ed Bromus inermis E. smithii B. tectorum E. smithii. ed specimen of . Native to Eurasia Native to Europe based on a misidentifi specimen of (2003) based on a specimen from HOVE herbarium (Litzinger 732) from outside the park. Moran (2003) based on a misidentifi based on a misidentifi specimen of state noxious weed. CO noxious weed (List B). Native to Eurasia based on a misidentifi specimen of japonicus (2003) based on a specimen from HOVE herbarium (Litzinger 735) from outside the park. based on a misidentifi specimen of (2003) based on a specimen from HOVE herbarium (Litzinger 668) from outside the park. UT CO Comments FalsRep O Report from HOVE herbarium FalsRep O Reported by Schelz & Moran FalsRep O Report from Schelz & FalsRep O Report from HOVE herbarium Park stat. FalsRep O Report from HOVE herbarium FalsRep O Reported by Schelz & Moran FalsRep O Report from HOVE herbarium FalsRep O Reported by Schelz & Moran Range Intro Wide Intro Intro Wide Wide Wide Wide Life form PerG Wide/ PerG Wide/ eld sandbur eld AnnG Wide/ Common name tall fescue PerG Intro/ sideoats grama PerG Wide/ nodding brome rye chess AnnG Intro/ quackgrass PerG Intro/ fi Canada wildrye blue wildrye PerG Wide/

var. & var. orus Synonyms/ notes Taxonomic Lolium arundinaceum, Schedonorus phoenix, S. arundinaceus Includes vars. curtipendula caespitosa Bromus porteri, B. frondosus, Bromopsis porteri Cenchrus paucifl E. canadensis canadensis glaucus Festuca arundinacea Bouteloua curtipendula Bromopsis porteri Cenchrus longispinus Elymus canadensis Species name arundinacea Bouteloua curtipendula Bromus anomalus Festuca Utah Colorado Bromus secalinus Bromus secalinus Cenchrus longispinus Elymus canadensis Elymus repens Elytrigia repens Agropyron repens Elymus glaucus Elymus glaucus E. glaucus Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

86 Annotated Checklist of Vascular Flora: Hovenweep National Monument . P. ed ed . Native . Native to P. fendleriana P. S. viridis ed specimen of ed specimen of . Native to Eurasia based on a misidentifi specimens of herbarium based on a misidentifi (2003) based on a specimen from HOVE herbarium (Litzinger 713) from outside the park. Native to Eurasia based on a misidentifi specimen of to Eurasia (2003) based on a specimen from HOVE herbarium (Litzinger 720) from outside the park. Native to Eurasia pratensis Moran (2003) based on a misidentifi Elymus elongatus Eurasia and Africa (2003) based on a specimen from HOVE herbarium (Litzinger 736) from outside the park. (2003) based on a specimen from HOVE herbarium (Litzinger 811) from outside the park. UT CO Comments FalsRep OFalsRep Report from HOVE herbarium O Reports from HOVE FalsRep O Reported by Schelz & Moran FalsRep O Report from HOVE herbarium FalsRep O Reported by Schelz & Moran FalsRep O Report from Schelz & FalsRep O Reported by Schelz & Moran FalsRep O Reported by Schelz & Moran Park stat. Intro Wide Intro Intro Intro Range Intro Wide Wide PerG Intro/ AnnG Intro/ Life form PerG Intro/ roughstalked bluegrass yellow bristlegrass fowl bluegrass PerG Wide/ cultivated rye AnnG Intro/ Common name perennial ryegrass timothy PerG Intro/ scratchgrass PerG Wide/ alpine timothy PerG Wide/ pumila, ssp. Pennisetum glaucum Synonyms/ notes Taxonomic Muhlenbergia asperifolia commutatum Species name Poa trivialis Poa trivialis Setaria glauca Setaria glauca S. pumila Poa palustris Poa palustris Secale cereale Secale cereale Lolium perenne Lolium perenne Muhlenbergia asperifolia Phleum alpinum Phleum Phleum pratense Phleum pratense Utah Colorado Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

Chapter 6: Appendices 87

A. A. var. . ed ed ed ed . . Native to Polypogon J. longistylis. A. calycosus J. arcticus avus fl ed specimen of ed specimen of var. avus (2003) based on a specimen from HOVE herbarium (Litzinger 775) from outside the park. Native to Eurasia based on a misidentifi specimen of semiverticillatus eastern and southwestern North America, but outside of Utah or Colorado. based on a misidentifi specimen of based on a misidentifi specimen of (2003) based on a specimen from HOVE herbarium (Litzinger 697) from outside the park. calycosus. based on a misidentifi specimen of Moran (2003) based on a misidentifi Moran (2003) based on a misidentifi fl wingatanus UT CO Comments FalsRep O Reported by Schelz & Moran FalsRep O Report from HOVE herbarium FalsRep O Report from HOVE herbarium FalsRep O Reported by Schelz & Moran Park stat. FalsRep O Report from HOVE herbarium FalsRep O Report from Schelz & FalsRep O Report from Schelz & Range Sparse Wide Wide Periph Periph Wide Wide Life form AnnG NA/NA FalsRep O Report from HOVE herbarium PerF Wide/ PerF Wide/ PerF Periph/ Common name sheathed dropseed spike dropseed PerG Wide/ thread rush PerG Periph/ poverty rush PerG Wide/ purple sage Shrub Wide/ silver-leaved silver-leaved milkvetch Hayden’s Hayden’s milkvetch pliant milkvetch

& dudleyi, Synonyms/ notes Taxonomic Includes vars. congestus, tenuis. J. dudleyi var. var. liformis exuosus Sporobolus contractus Poliomintha incana Astragalus argophyllus martinii Astragalus haydenianus Astragalus fl

Species name var. var. liformisfi Juncus orus exuosus exuosus Sporobolus contractus Sporobolus vaginifl Juncus fi Poliomintha incana Astragalus argophyllus Astragalus bisulcatus haydenianus Astragalus fl fl Utah Colorado Juncus tenuis Juncus tenuis Family Gramineae (Poaceae) Gramineae (Poaceae) Juncaceae Labiatae (Lamiaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Juncaceae Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

88 Annotated Checklist of Vascular Flora: Hovenweep National Monument . L. ed

A. . ed ed ed ed Not and .

A. nuttallianus utahensis Peteria scoparia A. amphioxys Hedysarum boreale. A. mollissimus . Native to SW US ed specimen of var. var. var. thompsoniae amphioxys. micranthiformis Z. paniculatus specimen of (2003) based on a specimen from HOVE herbarium (Litzinger 934) from outside the park. (2003) based on misidentifi specimens of var. sabulonum but not to Utah or Colorado. based on a misidentifi specimen of var. (2008), but HOVE is beyond the known range of this taxon. Report probably based on based on a misidentifi specimen of var. native to Utah or Colorado. Moran (2003) based on a misidentifi based on a misidentifi specimen of boreale caudatus based on a misidentifi UT CO Comments FalsRep ? ? Reported by Schelz & Moran FalsRep O Report from HOVE herbarium Park stat. Ques? O Loh et al. Reported by Von FalsRep ? ? Report from HOVE herbarium FalsRep O Report from Schelz & FalsRep O Report from HOVE herbarium Wide Range NA Wide RegEn Wide Wide Life form PerF Wide/ PerF RegEn/ PerF NA/NA FalsRep ? ? Report from HOVE herbarium Tree Periph/ Common name Gila milkvetch PerF NA/NA FalsRep O Report from Schelz & Moran taper-tip oniontaper-tip PerF Wide/ mountain death camas Newberry’s Newberry’s milkvetch Utah milkvetch PerF Wide/ silvery lupine PerF Wide/ ashen milkvetch New Mexico locust Synonyms/ notes Taxonomic Includes var. Includes var. escalantinus Allium acuminatum Anticlea elegans Astragalus newberryi Lupinus argenteus Robinia neomexicana

Species name var. var. Allium acuminatum Zigadenus elegans Astragalus gilensis Astragalus newberryi newberryi Utah Colorado argenteus Lupinus argenteus Astragalus utahensis Astragalus tephrodes Robinia neomexicana Liliaceae (Alliaceae) Liliaceae (Melanthiaceae) Family Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

Chapter 6: Appendices 89 M. G. O. . . ed ed ed cooperi M. marginata S. leptophylla O. albicaulis . var. var. . ed specimen of ed specimen of ed specimen of Moran (2003) based on a misidentifi based on a misidentifi specimen of based on a misidentifi specimen of Moran (2003) based on a misidentifi marginata (2003) based on a specimen from HOVE herbarium (Litzinger 704) from outside the park. (2003) based on a specimen from HOVE herbarium (Litzinger 561) from outside the park. Native to Eurasia. inconspicua (2003) based on a specimen from HOVE herbarium (Litzinger 586) from outside the park. based on a misidentifi specimen of Moran (2003) based on a misidentifi ludoviciana UT CO Comments FalsRep O Report from Schelz & FalsRep OFalsRep Report from HOVE herbarium O Report from HOVE herbarium FalsRep O Report from Schelz & FalsRep O Reported by Schelz & Moran Park stat. FalsRep O Reported by Schelz & Moran FalsRep O Reported by Schelz & Moran FalsRep O Report from HOVE herbarium FalsRep O O Report from Schelz & Wide NA NA NA Range Wide Intro Wide RegEn Wide PerF RegEn/ PerF Periph/ Life form AnnF Wide/ PerF Intro/ PerF RegEn/ PerF Wide/ Wyoming stickleaf narrowleaf globemallow Common name desert stickleaf PerF Wide/ branching groundsmoke English plantain broadlobe gilia AnnF Wide/ pale evening- primrose scarlet gilia PerF Wide/ clustered broomrape

var. ora ora aggregata M. multifl M. pumila multifl Synonyms/ notes Taxonomic Aliciella hutchinsifolia Gilia aggregata var.

ssp.

ssp. ora Nuttallia multifl Gayophytum ramosissimum Plantago lanceolata Oenothera pallida trichocalyx Ipomopsis aggregata aggregata Aphyllon fasciculatum

var.

Species name . var var. ora pumila Sphaeralcea angustifolia Gayophytum ramosissimum Mentzelia multifl Mentzelia pumila trichocalyx Oenothera pallida Gilia hutchinsifolia Ipomopsis aggregata aggregata Utah Colorado Orobanche fasciculata Plantago lanceolata Malvaceae Onagraceae Family Loasaceae Loasaceae Onagraceae Polemoniaceae Polemoniaceae Orobanchaceae Plantaginaceae Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

90 Annotated Checklist of Vascular Flora: Hovenweep National Monument

E. ed

is a var. var. Type Type ed simpsonii. P. lentus P. caudatus P. angustifolius P. var. var. P. fremontii P. and Var. ed specimen of var. ed specimen of var. specimens ed ed specimen of coloradoensis. venosus lentus. locality is Osmer, Kane Co., locality is Osmer, UT (Siler s.n. GH) Moran (2003) based on a misidentifi undosus. of ssp. Moran (2003) based on misidentifi var. var. USFWS 3C Moan (2003) based on a misidentifi var. var. (2003) based on misidentifi specimens of microthecum (2003) based on a specimen from HOVE herbarium (Litzinger 729) from outside the park. (2003) based on a specimen from HOVE herbarium (Litzinger 569) from outside the park. Great Plains taxon. based on a misidentifi specimen of wislizenii UT CO Comments FalsRep O Report by Schelz & FalsRep O Report from Schelz & Park stat. FalsRep O Report from Schelz & FalsRep O O Report from Schelz & Moran FalsRep O Reported by Schelz & Moran FalsRep O Reported by Schelz & Moran FalsRep O Report from HOVE herbarium Range Periph NA RegEn Wide Wide Periph Wide Life form PerF Wide/ PerF RegEn/ Shrub RegEn/ PerF NA/ AnnF Wide/ recracker Common name fi Siler’s Siler’s penstemon Comb wild Wash buckwheat Sawatch knotweed blackbrush Shrub Wide/ balsam poplar Tree Wide/ Eaton’s .

& ssp & sawatchense Synonyms/ notes Taxonomic P. sawatchense P. var. Includes vars. californica balsamifera trichocarpa, P. trichocarpa

var.

var. Eriogonum clavellatum Polygonum douglasii Penstemon angustifolius caudatus Coleogyne ramosissima Populus balsamifera Penstemon eatonii eatonii

var. Species name var. var. var. Penstemon linarioides sileri Eriogonum clavellatum Utah Colorado Polygonum douglasii johnstonii Coleogyne ramosissima Populus balsamifera Penstemon eatonii eatonii Scrophulariaceae Family Polygonaceae Polygonaceae Rosaceae Salicaceae Scrophulariaceae Scrophulariaceae Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

Chapter 6: Appendices 91

C. ed ed . . C. ed ed ed and

purpureus purpureus Cymopterus L. triternatum Cymopterus Chamaesaracha Chamaesaracha fendleri var. . Native to E North . Native to C North var. var. ed specimen of . Native to E North var. var. platycarpum. purpureus (2003) based on misidentifi specimens of acaulis based on a misidentifi specimen of purpureus (2003) based on a specimen from HOVE herbarium (Litzinger 657) from outside the park. Moran (2003) based on a HOVE herbarium specimen (Litzinger 706) from outside the park. based on a misidentifi specimen of coronopus America based on a misidentifi specimen of coronopus America. Moran (2003) based on a misidentifi reticulata America. ssp. (2003) based on misidentifi specimens of UT CO Comments Park stat. FalsRep O O Report from Schelz & Moran FalsRep O Report from HOVE herbarium FalsRep O Reported by Schelz & Moran FalsRep O Reported by Schelz & FalsRep O Report from HOVE herbarium FalsRep O Report from HOVE herbarium FalsRep O Report from Schelz & FalsRep O Report from Schelz & Moran Range RegEn Wide Wide Wide Intro Intro Wide Wide Life form PerF RegEn/ PerF Wide/ PerF Wide/ PerF Intro/ PerF Intro/ Tree Intro/ PerF Wide/ Common name juniper lomatium milfoil lomatium Rocky Mountain penstemon coyote tobacco AnnF Wide/ Carolina nightshade silverleaf nightshade common hackberry giant lomatium

var. dum Synonyms/ notes Taxonomic L. dissectum multifi Lomatium juniperinum Lomatium grayi Penstemon strictus Nicotiana attenuata Solanum carolinense Solanum elaeagnifolium Celtis occidentalis Lomatium dissectum

Species name var. grayi Lomatium juniperinum var. Penstemon strictus Utah Colorado Nicotiana attenuata Solanum carolinense Solanum elaeagnifolium Celtis occidentalis Lomatium dissectum eatonii Lomatium grayi Umbelliferae (Apiaceae) Family Scrophulariaceae Solanaceae Solanaceae Solanaceae Ulmaceae (Celtidaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont.

92 Annotated Checklist of Vascular Flora: Hovenweep National Monument (2003) based on a HOVE herb- arium specimen (Litzinger 774) from outside the park. CO state noxious weed (List C). Native to Eurasia UT CO Comments FalsRep O Reported by Schelz & Moran Park stat. ed literature (Von Loh et al. 2008; Schelz & Moran 2003) or from Loh et al. 2008; Schelz & Moran 2003) or from (Von ed literature Range Intro ed (i.e., could be UT or CO). Life form ed or collected outside the monument (False Reports FalsRep) because Hovenweep NM Common name puncture vine AnnF Intro/ Synonyms/ notes Taxonomic Tribulus Tribulus terrestris Species name Tribulus Tribulus terrestris Utah Colorado Family Zygophyllaceae Appendix C. Rejected (falsely reported or questionable) taxa of Hovenweep National Monument, cont. The vascular plant species listed in this table have been reported for Hovenweep National Monument from published and unpublish for Hovenweep National Monument from The vascular plant species listed in this table have been reported misidentifi vouchers were because corroborating now believed to be erroneous herbarium specimens, but are is well outside the known or expected range of species (Questionable Ques?). for the monument without state specifi X = documented with a voucher; O by an observation; ? reported

Chapter 6: Appendices 93 gravelly soils x Wet Comments Jun Pin- Sage Flower time Aug Park status Native Pres Mar–Jun x Native Pres May–Sep x Saline clays Fendler’s barberryFendler’s Oregon-grape Native chollaWhipple’s Pres saltbushfour-wing Native Native May–Jun Native Presshadscale Rep Pres Apr–Jun May–Jul May–Sep x x x x Native x x Pres x x May–Sep x x Sandy soils x Clayey to squawbushsquawbush Nativewestern poison ivy Pres Native Native May–Jul Rep Pres x May–Jun May–Jul x x x Fremont cottonwood netleaf hackberry Native Presnarrowleaved yucca Nativedatil yucca Apr–May Pres yuccaHarriman’s x May–Jul Native Native Rep x Pres Apr–May x x Apr–Jun x x x Sandy soils x Castle Valley saltbush Common name Nativity basin saltbushspiny hopsage Nativewinterfat Pres Nativegreasewood Jun–Sep Pres Native Apr–Jul Native Pres x Pres x May–Jul May– x x x two-needle pinyon Native Pres NA x x Gambel’s oakGambel’s Native Pres Apr–Jun x x x

ssp. ssp. forma trilobata Atriplex canescens simplicifolia Rhus aromatica trilobata Toxicodendron rydbergii wislizenii harrimaniae Yucca Rhus aromatica ssp. Atriplex gardneri Krascheninnikovia lanata Populus deltoides Quercus gambelii

var. var. var. var. Species name var. var. var. var. var. lanata var. Mahonia repensOpuntia whippleiAtriplex canescens canescens Mahonia repens Cylindropuntia whipplei Atriplex confertifolia Atriplex confertifolia Rhus aromatica trilobata Toxicodendron rydbergii Berberis fendleri Berberis fendleri Yucca angustissima Yucca baccataYucca angustissima Yucca harrimaniae Yucca harrimaniae Rhus aromatica baccata Yucca simplicifolia Atriplex gardneri cuneata Celtis reticulata Celtis reticulata Atriplex gardneri Grayia spinosaKrascheninnikovia lanata Sarcobatus vermiculatus Sarcobatus vermiculatus Atriplex grayi utahensis Populus fremontii fremontii Quercus gambelii Pinus edulis Pinus edulis gambelii Utah Colorado taxa of Hovenweep National Monument, organized by life form with ecological notes. rmed, historical, and reported Berberidaceae Cactaceae Chenopodiaceae Chenopodiaceae Anacardiaceae Anacardiaceae Berberidaceae Shrubs Agavaceae Agavaceae Agavaceae Anacardiaceae Chenopodiaceae Ulmaceae (Celtidaceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae (Sarcobataceae) Salicaceae Family Trees Fagaceae Pinaceae Appendix D. Confi

94 Annotated Checklist of Vascular Flora: Hovenweep National Monument gravelly saline soils Wet Comments Jun Pin- Sage Flower time Park status Native Pres Jul–Sep x x Native Pres Aug–Sep x Rock crevices Native Pres Aug–Oct x x Greene’s rabbitbrushGreene’s Native Presgraystem Jul–Sep x x Wyoming big sagebrush Mohave brickellbush Native Presdwarf rabbitbrush Native Jun–Aug Pres x Jul–Sep x x x California brickellbush rough brickellbush Native Pres Aug–Oct x x Sandstone Common name Nativity rush rabbitbrush Native Rep Aug–Oct x x rabbitbrush glabrate rabbitbrush Native Pres Aug–Oct x x basin big sagebrush Native Pres Jul–Sep x x Bigelow’s sagebrushBigelow’s Nativesand sagebrush Pres Aug–Octfringed sagebrush Native x Rep Nativeblack sagebrush Pres Aug–Oct xbudsage Native Jul–Sep x Pres x x Sep–Oct Rocky areas Native x x Pres x Apr–Jun x x Clayey to

junceus lifolius ssp. ssp. var. var. var. scabra Chrysothamnus nauseosus gnaphalodes wyomingense Brickellia oblongifolia Chrysothamnus depressus Brickellia microphylla var. Chrysothamnus nauseosus Chrysothamnus nauseosus graveolens tridentatum Seriphidium tridentatum Seriphidium novum desertorum Seriphidium tridentatum

var. var.

Species name var.

junceus lifoliafi Oligosporus var. var. var. linifolia scabra Chrysothamnus nauseosus gnaphalodes Chrysothamnus depressus Chrysothamnus greenei Chrysothamnus greenei Brickellia californica Brickellia californica Brickellia oblongifolia var. Brickellia microphylla var. Chrysothamnus nauseosus graveolens Chrysothamnus nauseosus Artemisia tridentata wyomingensis Artemisia fi Artemisia frigidaArtemisia nova nova Artemisia frigida Artemisia spinescens Picrothamnus Artemisia bigelovii Artemisia bigelovii Utah Colorado Artemisia tridentata tridentata rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi Family Shrubs, cont. Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 95 Wet Comments x Jun Pin- Sage Flower time Aug Park status Native Rep Jul–OctNative Pres x May–Jun x x x Native Pres Aug–Oct x x Native Pres Jul–Sep x x Native Pres Apr–Jun x x Native Pres Mar–Jun x x Native Pres May– Common name Nativity crisp-leaf wild buckwheat Utah serviceberry mountain alder-leaf Nativemahogany Pres Apr–Jun x x x squaw-apple Native Pres May–Jun x x whitestem rabbitbrush singleleaf ashprickly-phlox Native Pres Native Pres Apr–May May–Jul x x x slenderleaf rabbitbrush broom snakeweed Nativespineless horsebrush Pres Nativeshortspine Pres Jul–Octhorsebrush Jun–Sep x x x small-leaved greasebush Utah juniper ephedraTorrey’s green ephedra Native Nativeshinnery oak Pres Pres Native NA Pres NA oakEastwood’s Native NA Native Pres x Pres Apr–Jun x Apr–Jun x x x x x x Sandy soils Thompson’s indigo- Thompson’s bush desert olive Native Rep Apr–May x var. var.

speciosus var. var. var. eastwoodiae x orus corymbosum Eriogonum corymbosum var. Peraphyllum ramosissimum Chrysothamnus nauseosus Chrysothamnus viscidifl stenophyllus Forsellesia meionandra Ephedra viridis tuckeri Psorothamnus thompsoniae thompsoniae

Species name

var.

speciosus var. var. var. var. eastwoodiae Quercus x orus corymbosum Cercocarpus montanus Cercocarpus montanus Amelanchier utahensis Amelanchier utahensis Peraphyllum ramosissimum Leptodactylon pungens Leptodactylon pungens Eriogonum corymbosum var. Chrysothamnus nauseosus Chrysothamnus viscidifl stenophyllus Fraxinus anomala Fraxinus anomala Gutierrezia sarothrae Gutierrezia sarothrae canescensTetradymia canescens Tetradymia spinosaTetradymia spinosa Tetradymia Glossopetalon spinescens microphyllum Juniperus osteosperma Sabina osteosperma Ephedra torreyanaEphedra viridis viridis Ephedra torreyana Quercus welshiiQuercus Psorothamnus Quercus havardii thompsoniae thompsoniae Forestiera pubescens Forestiera pubescens Utah Colorado rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Rosaceae Rosaceae Rosaceae Polemoniaceae Polygonaceae Family Shrubs, cont. Compositae (Asteraceae) Oleaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Crossosomataceae (Celastraceae) Cupressaceae Ephedraceae Ephedraceae Fagaceae Fagaceae Leguminosae (Fabaceae) Oleaceae Appendix D. Confi

96 Annotated Checklist of Vascular Flora: Hovenweep National Monument Wet Comments Jun Pin- xx Sage May Flower time Park status Native Rep Apr–Jun x x Native Pres Apr–JunNative x Pres May–Jun x Native Pres x May–Jun x x ve-stamen tamarisk ve-stamen Intro Pres Apr–Sep x antelope bitterbrush Native roseWoods’ Prescoyote willow Apr–Julgolden currant Native Native x Prestrumpet gooseberry Pres Native Native May–Jul x Apr–Aug Prespale wolfberry Rep Mar–Jul Apr–Jun Native x Pres Mar– x x x x x western chokecherry Native Prescliff-rose May–Jul x Native x Pres May–Jun x x Common name Nativity Arizona pincushion Native Pres Apr–Aug x x pallid milkweed Native Pres Apr–Jun x x fi spider milkweed Native Pres May–Jul x x plateau yellow cryptanth yellow-eye cryptanth Osterhout’s cryptanth showy stoneseedwestern stoneseed Nativespindle-rooted Nativebluebell Pres Pres Apr–Jul Apr–Jun x x x x plateau cryptanth Native Pres May–Jun x x Mancos Shale

ssp. ssp. ava avoculata Salix exigua melanocarpa Purshia stansburiana Coryphantha vivipara Padus virginiana Asclepias cryptoceras Asclepias asperula asperula Oreocarya fulvocanescens

var. Species name var. var.

avafl Oreocarya avoculatafl Oreocarya var. arizonica cryptoceras Rosa woodsiiSalix exigua stenophylla Ribes aureum Rosa woodsii Ribes leptanthum Ribes aureum pallidumLycium Ribes leptanthum pallidum Lycium Purshia mexicana stansburyana Purshia tridentata Purshia tridentata Coryphantha vivipara var. Prunus virginiana melanocarpa Cryptantha fl Tamarix chinensisTamarix Asclepias asperula ramosissima Tamarix asperula Asclepias cryptoceras var. Utah Colorado Cryptantha fl Lithospermum ruderale Lithospermum ruderale Mertensia fusiformis Mertensia oblongifolia Cryptantha fulvocanescens Cryptantha osterhoutii Oreocarya osterhoutii Lithospermum incisum Lithospermum incisum rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Rosaceae Salicaceae Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Solanaceae Rosaceae Rosaceae Appendix D. Confi Family Shrubs, cont. Rosaceae Boraginaceae Tamaricaceae Forbs Perennial Asclepiadaceae Asclepiadaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Cactaceae

Chapter 6: Appendices 97 areas areas Wet Comments Jun Pin- xx Sage Flower time Aug Park status Native PresNative Jul–Oct Pres Jul–Oct x x x x Native Pres Apr–Jul x x Native Rep Apr–May x x Native Pres May–Sep x x ower shhook cactus shhook Common name Nativity Louisiana wormwood Louisiana wormwood arrowleaf balsamroot musk thistle Intro Pres Jun–Sep x x Disturbed small-fl fi Eastwood’s sandwort large pricklypear Nativeberry pricklypear Rep Nativeplains prickly-pear Apr–Jul Pres Native Pres Apr–Jul May–Jun x xwestern spiderwort Native x x Rep May– Utah pricklypear Nativebrittle prickly-pear Rep Native Pres May–Jun Jun–Jul x x x x claretcupcommon pricklypear Native Native Pres Pres May–Jun Apr–Jun x x x common yarrowcommon pussytoes Native Native Presburdock Rep May–Sep May–Jul x x x Intro x Pres x Jul–Sep x Disturbed

var. var. mexicana albula intermedius Artemisia ludoviciana ssp. macrolepis Eremogone fendleri Opuntia phaeacantha Opuntia polyacantha Sclerocactus whipplei var. Tradescantia occidentalis brachyartha Echinocereus triglochidiatus Opuntia erinacea Achillea lanulosa Artemisia ludoviciana ssp.

var.

Species name var. var. var. var. mexicana phaeacantha polyacantha roseus major albula Artemisia ludoviciana var. Balsamorhiza sagittata Balsamorhiza sagittata Carduus nutans Carduus nutans Arenaria fendleri eastwoodiae Opuntia phaeacantha var. Opuntia polyacantha var. Sclerocactus whipplei var. Tradescantia occidentalis Opuntia fragilisOpuntia phaeacantha var. Opuntia fragilis Echinocereus triglochidiatus melanacanthus Opuntia erinacea erinacea Opuntia erinacea utahensis Utah Colorado Achillea millefolium lanulosa Antennaria parvifolia Antennaria parvifolia Arctium minus Arctium minus Artemisia ludoviciana var. rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Caryophyllaceae (Alsinaceae) Cactaceae Cactaceae Cactaceae Commelinaceae Cactaceae Cactaceae Family Perennial Forbs, cont. Cactaceae Cactaceae Cactaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi

98 Annotated Checklist of Vascular Flora: Hovenweep National Monument areas areas Wet Comments Jun Pin- xx Sage Flower time Aug Park status ower Native Pres Apr–Aug x eabane Native Pres May– Common name Nativity Hopi blanketfl spiny goldenweed Native Pres Apr–Sep x x Morrison F Russian knapweed Intro Pres Jun–Sep x Disturbed thrift goldenweed Native Pres May–Jun x x rose-heathhispid golden-aster Native Nativehairy golden-aster Pres Pres Native May–SepCanada thistle Pres May–Sep x x May–Sep Intro x Pres x Jul–Sep x x Disturbed gray hawksbeard Nativewestern hawksbeard Native Pres Presspreading daisy May–Jul May–JulNavajo fl Native x x Pres x May–Sep x x x Caineville thistleNew Mexico thistle Native Native Preswavy-leaved thistle Rep Jun–Sep Native May–Jun Rep x Jun–Sep x x x x Utah daisy Native Rep Apr–Jun x

var. var. da

var. da paradoxa Stenotus armerioides Machaeranthera pinnatifi Heterotheca villosa minor Heterotheca villosa villosa Psilochenia occidentalis Erigeron divergens Erigeron concinnus Cirsium calcareum Cirsium neomexicanum Cirsium undulatum Erigeron utahensis

var. var. var. var. var. var. dapinnatifi Gaillardia Species name var. var. var. var.

var. paradoxus neomexicanum Centaurea repens Acroptilon repens Utah Colorado Haplopappus armerioides armerioides Haplopappus spinulosus var. Chaetopappa ericoides Leucelene ericoides Chrysopsis villosa Chrysopsis villosa Cirsium arvense Breea arvensis minor villosa Crepis occidentalis Erigeron divergens divergens Erigeron pumilus concinnus occidentalis Cirsium neomexicanum var. Cirsium undulatum Crepis intermedia Psilochenia intermedia calcareum undulatum Cirsium calcareum Erigeron utahensis utahensis Gaillardia pinnatifi rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Appendix D. Confi Family Perennial Forbs, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 99 areas areas areas Wet Comments Jun Pin- xx Sage Flower time Aug Park status Native Pres May–Oct x x Native PresNative Apr–Jun Pres May– x Native Rep Jun–Aug x x eld bindweed eld Intro Pres Jun–Aug Disturbed Common name Nativity common hyalineherb canyonlands hymenoxys Colorado rubber- plant common dandelion Intro Pres Apr–Dec x x x Disturbed Douglas’ groundsel Nativebasin groundsel Pres May–Octslender wirelettuce Native Native x Pres Pres Apr–Aug x Jun–Sep x x x x showy rush-pinkhoary aster Native Repgumweed aster May–Jun Native Native xrock-goldenrod Pres Rep May–Sep x Native May–Jul x Pres x Jul–Sep x x x x hoary townsendia Nativeyellow salsify Pres Apr–Jul Intro x Pres May–Jul x x x Disturbed showy goldeneye Nativefi Pres Jun–Sep x x x lance-leaved stonecrop Holboell’s rockcressHolboell’s Native Rep May–Jul x x x

var. var. ora var. ora lifolius cinale ora accidus oribunda cinereus fl multifl Hymenopappus fi var. acaulis Tetraneuris Picradenia richardsonii var. douglasii Stephanomeria tenuifolia Lygodesmia grandifl Lygodesmia Machaeranthera canescens Machaeranthera grindelioides Petradoria pumila Senecio fl major Heliomeris multifl var. lanceolatum Boechera holboellii pinetorum

var. ora var. var. Species name var.

lifolius var. ora cinaleoffi Taraxacum canescens tenuifolia var. ora var. var. ora oribunda cinereus fl grandifl Hymenopappus fi var. Hymenoxys acaulis offi Taraxacum ivesiana Hymenoxys richardsonii var. Senecio multilobatus Packera multilobata Stephanomeria tenuifolia Utah Colorado Lygodesmia grandifl Lygodesmia var. Machaeranthera canescens Machaeranthera grindelioides grindelioides Petradoria pumila pumila Senecio douglasii longilobus incanaTownsendia dubiusTragopogon incana Townsendia dubius Tragopogon Viguiera multifl Convolvulus arvensis Convolvulus arvensis multifl Sedum lanceolatum Amerosdeum Arabis holboellii pinetorum rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Family Perennial Forbs, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Convolvulaceae Crassulaceae Cruciferae (Brassicaceae) Appendix D. Confi

100 Annotated Checklist of Vascular Flora: Hovenweep National Monument seleniferous soils areas areas Wet Comments Jun Pin- xx x x x Disturbed Sage Flower time Aug Aug Park status Native Pres Jun–Aug x x Native PresNative Apr–Aug Pres Apr–Jul x x x Native Pres Apr–Jul x x ower ower ax-leaved Common name Nativity smallfl dragonhead mock pennyroyal Native Pres May– Jones’ pepperwort Nativesharp-leaf twinpod Pres NativeColorado Plateau Apr–Sep Presbladderpod xfl May–Julplainsmustard x x x perennial rockcress Nativedesert rockcress Pres Mar–Jul Native Pres x Mar–Jun x x x prince’s-plume Native Pres Apr–Sep x x Often on common horehound Intro Pres May–Oct Disturbed heart-leaf jewelfl gourdbuffalo Native Rep May– alpine pennycress Native Rep Apr–Aug x catnip Intro Pres Jun–Sep x x x Fendler’s spurgeFendler’s golden corydalis NativeUtah swertia Native PresGerman iris Pres May–Sep Apr–Sep Native x Pres x Intro x Jun–Jul Rep x Apr–May x Sandy soils Cultivated

var. var. orum occidentalis jonesii Dracocephalum parvifl Physaria acutifolia Boechera perennans Boechera pulchra pallens Lepidium montanum var. pinnata Noccaea montana Corydalis curvisiliqua ssp.

var. var.

Species name var. var. var. orum jonesii Hedeoma drummondii Hedeoma drummondii Physaria acutifolia acutifolia Physaria rectipes Lesquerella rectipes Arabis perennans Arabis pulchra pallens Lepidium montanum var. Schoenocrambe linifolia Schoenocrambe linifolia Stanleya pinnata Stanleya pinnata perennans Utah Colorado Marrubium vulgare Marrubium vulgare Streptanthus cordatus Streptanthus cordatus Thlaspi montanum montanum Cucurbita foetidissima Cucurbita foetidissima Nepeta cataria Nepeta cataria Chamaesyce fendleriCorydalis aurea Chamaesyce fendleri occidentalis Swertia utahensisIris germanicaDracocephalum Frasera paniculata parvifl Iris germanica rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Labiatae (Lamiaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix D. Confi Family Perennial Forbs, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Labiatae (Lamiaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cucurbitaceae Labiatae (Lamiaceae) Euphorbiaceae Fumariaceae Gentianaceae Iridaceae Labiatae (Lamiaceae)

Chapter 6: Appendices 101 areas areas areas areas Wet Comments x Jun Pin- xx xx Sage Flower time Aug Aug Aug Park status Native Pres May– Native Pres Apr–Jul x x Common name Nativity woolly milkvetch NativeRocky Mountain milkvetch Pres Feb–Jun x x rimrock milkvetch Native Rep May–Jul x x painted milkvetch NativeMoab milkvetch Rep Native Apr–Jun Pres x Apr–Jun x x x yellow sweet-clover Intro Pres May–Oct x Disturbed crescent milkvetch Native milkvetchTorrey’s Pres Native Mar–Jun Pres Apr–Jul x x x Fort Wingate milkvetch rush peteriared clover Intro Pres Intro Rep May–Oct x x Disturbed northern sweetvetch Native Pres May– Utah lupine Native Pres May– asparagus Intro Pres Jun–Jul x Disturbed pink funnel-lily Native Pres Apr–May x x sand lupinealfalfa Native Pres Jun–Aug Intro x Pres x May–Sep Sandy soils x Disturbed

var. var. cinalis cinale orum thompsoniae ceramicus Astragalus desperatus Astragalus mollissimus var. Astragalus coltonii moabensis Astragalus amphioxys Astragalus calycosus Astragalus ceramicus var. Peteria scoparia Hedysarum boreale Lupinus caudatus utahensis Androstephium brevifl Lupinus ammophilus

var. var. var. var. var. Species name var. cinalisoffi Asparagus cinalisoffi Melilotus orum thompsoniae desperatus amphioxys ceramicus Astragalus scopulorum Astragalus scopulorum Astragalus mollissimus var. Astragalus coltonii moabensis Astragalus desperatus var. Astragalus amphioxys var. Astragalus calycosus calycosus Astragalus ceramicus var. Melilotus offi Utah Colorado Astragalus wingatanus Astragalus wingatanus pratenseTrifolium pratense Trifolium Hedysarum boreale boreale Lupinus caudatus utahensis Asparagus offi Androstephium brevifl Lupinus polyphyllus ammophilus Medicago sativa Medicago sativa rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Family Perennial Forbs, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Asparagaceae) Liliaceae (Alliaceae, Themidaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix D. Confi

102 Annotated Checklist of Vascular Flora: Hovenweep National Monument Wet Comments Jun Pin- xxx Sage Flower time Aug Park status Native Pres Mar–Jun x x Native PresNative May–Sep PresNative x Mar–Oct Pres x x Apr–SepNative Rep x x May– x Native Rep Jun–Aug x Native Pres Apr–Jun x x Native Rep May–Jul x x ax Native Pres Apr–Aug x x Common name Nativity spotted coral-root broomrapeCooper’s Native Native Rep Pres Jun–Jul May–Jul x x x sego-lilyfalse Solomon’s-seal Native Rep Nativefoothills death camas Jun–Jul Pres Jun–JulColorado stickleaf xscarlet globe-mallow Native x Native Pres Pres x slenderleaf globemallow Apr–Oct Apr–Octsmall-leaf globemallow x xfragrant sand- verbena x x trailing four-o’clock Nativenarrow-leaf four- o’clock Rep Apr–Sep x sinuous mariposa Native Pres Apr–Junblue fl x northern willow- herb showy four-o’clock Native Preslong-tube evening- primrose Apr–Jul x pale evening- primrose

ssp. ora exuosus dissecta . marginata Orobanche ludoviciana amplexicaule paniculatum ssp. Oxybaphus linearis Adenolinum lewisii Oenothera cespitosa ssp Oenothera pallida pallida

var. var. Species name var. ssp. ora multifl Mirabilis exuosusfl Calochortus cooperi marginata Orobanche ludoviciana var. Smilacina racemosa Maianthemum Zigadenus paniculatus Toxicoscordion Sphaeralcea coccinea Sphaeralcea coccinea Sphaeralcea leptophylla Sphaeralcea leptophylla Mirabilis linearis linearis Calochortus fl Calochortus nuttallii Calochortus nuttallii Linum perenne lewisii Mentzelia marginata Nuttallia marginata Sphaeralcea parvifolia Sphaeralcea parvifolia Abronia fragransAllionia incarnata Abronia elliptica Allionia incarnata Utah Colorado Epilobium ciliatumOenothera caespitosa Epilobium ciliatum var. Mirabilis multifl Oenothera pallida pallida Corallorhiza maculata Corallorhiza maculata rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Orobanchaceae Liliaceae (Convallariaceae or Ruscaceae) Liliaceae (Melanthiaceae) Malvaceae Malvaceae Nyctaginaceae Appendix D. Confi Family Perennial Forbs, cont. Liliaceae (Calochortaceae) Liliaceae (Calochortaceae) Linaceae Loasaceae Malvaceae Nyctaginaceae Nyctaginaceae Onagraceae Onagraceae Nyctaginaceae Onagraceae Orchidaceae

Chapter 6: Appendices 103 areas areas Wet Comments Jun Pin- Sage Flower time Park status Native Pres May–Jul x x Slickrock Native Pres Jun–Oct x x Native Pres Jun–Oct x x Native Pres Apr–Aug x x Native Pres Jun–Oct x x Native Pres Jul–Aug x x ax Intro Pres Jul–Aug x x Disturbed ax Native Pres Apr–Aug x x Dalmatian toadfl Common name Nativity bastard toadfl Eastwood’s Eastwood’s paintbrush shortest penstemon Native Pres May–Jun x x long-leaf phloxdesert trumpet Native wild Baker’s Native Presbuckwheat Pres Apr–Jul Mar–Oct x x x Sandy soils ball-head gilia Native Pres Jun–Augslender wild xbuckwheat x cushion wild buckwheat arid wild buckwheat Native Prescurly dock Jun–Oct x Intro x Pres May–Sep x Disturbed redroot wild buckwheat desert paintbrushWyoming Nativepaintbrush Pres Apr–Jun x x pale larkspur larkspurNuttall’s Native Native Pres Pres Apr–Jun Apr–Jul x x x canaigre dockwhite virgin’s-bower Native Native Pres Pres Jul–Aug Apr–Jun x x

var. ssp.

atum var. ovalifolium subaridum scaposum dalmatica Castilleja scabrida Eriogonum infl Eriogonum jamesii Ipomopsis congesta congesta Eriogonum microthecum simpsonii var. Eriogonum racemosum Eriogonum umbellatum var. Comandra umbellata Delphinium nuttallianum Delphinium andersonii var.

var. var. var. var. var. Species name

atum var. racemosum subaridum pallida scaposum atum avescens scabrida Linaria dalmatica Linaria genistifolia Penstemon breviculus Penstemon breviculus Eriogonum infl Eriogonum jamesii fl infl Ipomopsis congesta congesta Phlox longifolia Phlox longifolia Eriogonum microthecum simpsonii Eriogonum ovalifolium Eriogonum ovalifolium Rumex crispus Rumex crispus Eriogonum racemosum var. Eriogonum umbellatum var. Utah Colorado Castilleja linariifolia Castilleja linariifolia Castilleja chromosa Castilleja chromosa Castilleja scabrida Delphinium nuttallianum Comandra umbellata var. Rumex hymenosepalus Rumex hymenosepalus Clematis ligusticifoliaDelphinium andersonii Clematis ligusticifolia var. rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Scrophulariaceae Scrophulariaceae Polygonaceae Polygonaceae Family Perennial Forbs, cont. Polemoniaceae Polemoniaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Ranunculaceae (Helleboraceae) Santalaceae Polygonaceae Ranunculaceae Ranunculaceae (Helleboraceae) Appendix D. Confi

104 Annotated Checklist of Vascular Flora: Hovenweep National Monument areas areas areas Wet Comments Jun Pin- xx xx Sage Flower time Aug May Park status Native Pres May–Jul x Native Pres Aug–Sep x Native Pres May–Oct x x Native Pres Apr–May x x Native Pres Mar– Native Pres Apr–Jun x x at-seed lomatium at-seed Native Pres May–Jul x x Common name Nativity prostrate vervain Native Rep May–Sep x x Disturbed undulate penstemon Native PresAbajo penstemon May–Jul NativeColorado penstemon Pres x May–Jun x x x fl Palmer’s penstemonPalmer’s Native Rep May– pinyon dwarf- mistletoe juniper mistletoe Native Pres Jul–Sep x Utah penstemonwoolly mullein Native Pres Intro Apr–Jun Pres x Jun–Aug x x x Disturbed false nightshade Native Pres Apr–Jul x Disturbed Fendler’s ground- Fendler’s cherry Fendler’s Fendler’s cymopterus onion spring-parsley Nativesticky spring-parsley Pres Native Apr–Maybasin white-cup Repspring-parsley x Apr–May x x x Colorado Plateau spring-parsley

var. var. var. var. var. var. coloradoensis platycarpum Penstemon eatonii undosus Penstemon lentus lentus Penstemon linarioides ssp. Lomatium triternatum ssp. Penstemon palmeri eglandulosus Arceuthobium divaricatum Phoradendron juniperinum Chamaesaracha coronopus Physalis hederifolia cordifolia Cymopterus fendleri Cymopterus purpurascens Cymopterus purpureus

var. var. var. var. var. var. Species name platycarpum purpureus Penstemon eatonii undosus Penstemon lentus lentus Lomatium triternatum ssp. bracteataVerbena bracteata Verbena Penstemon palmeri eglandulosus Utah Colorado Arceuthobium divaricatum Phoradendron juniperinum Penstemon utahensis thapsusVerbascum Penstemon utahensis thapsus Verbascum Chamaesaracha coronopus Physalis hederifolia fendleri Cymopterus acaulis fendleri Cymopterus bulbosus Cymopterus bulbosus Cymopterus newberryi Cymopterus newberryi Cymopterus purpurascens Cymopterus purpureus var. rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Appendix D. Confi Family Perennial Forbs, cont. Scrophulariaceae Umbelliferae (Apiaceae) Scrophulariaceae Scrophulariaceae Verbenaceae Scrophulariaceae Viscaceae Viscaceae Scrophulariaceae Scrophulariaceae Solanaceae Solanaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae)

Chapter 6: Appendices 105 areas areas areas areas Morrison F areas areas areas Wet Comments x Jun Pin- Sage Flower time Aug Park status Native Pres Jul–Sep x Native Pres Apr–Jun x x y Native Pres May– Common name Nativity goosefoot halogeton Intro Rep Jul–Aug x Disturbed prostrate pigweed Native Rep Jun–Sep Disturbed blood amaranthredroot pigweed Intro Intro Rep Pres Jun–Sep Jun–Sep Cultivated Disturbed wing-nut cryptanth Nativecupseed stickseed Pres Native Apr–Jul Pres x Apr–Jun x x x Disturbed thick-sepaled cryptanth sand dune cryptanth Nativeslender cryptanth Rep Native May–Jul Pres x Apr–Jul x x x sleepy catchfl stalked orach Native Pres Aug–Sep x Mancos & summer-cypress Intro Pres Aug–Oct x x x Disturbed lambsquarter Intro Pres Jun–Sep Disturbed pitseed goosefoot Native Pres Jul–Sep x x Disturbed silvery goosefoot Nativeoakleaf goosefoot Pres Nativenarrow-leaf Pres Jun–Aug Aug–Sep x x desert goosefoot Native Rep Jun–Sep x x x exus elachantha Cryptantha pterocarya Lappula marginata Cryptantha crassisepala var. Atriplex saccaria Chenopodium album Chenopodium berlandieri Chenopodium incanum Chenopodium glaucum Chenopodium leptophyllum Chenopodium desiccatum

var.

exusretrofl Amaranthus Species name var. cycloptera elachantha album berlandieri salinum incanum Halogeton glomeratus Halogeton glomeratus Amaranthus blitoides Amaranthus blitoides Utah Colorado Amaranthus cruentusAmaranthus retrofl Amaranthus cruentus Lappula occidentalis cupulata Cryptantha gracilisCryptantha pterocarya var. Cryptantha gracilis Cryptantha fendleri Cryptantha fendleri Cryptantha crassisepala var. Silene antirrhina Silene antirrhina Atriplex saccaria saccaria Bassia scoparia Bassia sieversiana Chenopodium album var. Chenopodium album var. Chenopodium glaucum var. Chenopodium leptophyllum Chenopodium desiccatum Chenopodium fremontii var. rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Chenopodiaceae Family Annual Forbs Amaranthaceae Amaranthaceae Amaranthaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Caryophyllaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Appendix D. Confi

106 Annotated Checklist of Vascular Flora: Hovenweep National Monument areas areas areas areas areas areas areas areas Wet Comments x Jun Pin- Sage Flower time Aug Park status Native Pres Mar–Aug x x Native Rep Jul–Sep x Native Pres Apr–Jun x x ax Intro Pres Apr–Jul Disturbed ower Native Pres Jun–Sep x Disturbed ower Native Pres Jun–Sep x x x ixweed Intro Pres Mar–Aug x x Disturbed Common name Nativity western tansy- mustard fl Russian thistle Intro Pres Jun–Sep Disturbed spectacle-pod Native Pres Apr–Sep x x spinach seepweedTorrey’s Nativebur ragweed Pres Intro Jun–Sep Native Pres Rep May–Sep Jul–Oct x x x x Cultivated Disturbed stevia dusty-maiden NativeCanadian Preshorseweed Apr–Julpretty daisy x Native x Pres May– common sunfl prairie sunfl marsh-elder Native Rep Aug–Oct x Disturbed prickly lettuce Intro Pres Jul–Sep x x Disturbed sowthistle desert- dandelion European madwort Introlittle-pod false fl Pres Apr–Jul x x blue mustard Intro Pres Apr–Jun Disturbed

ssp. orum halictorum wislizenii Suaeda moquinii Conyza canadensis Helianthus annuus Helianthus petiolaris Alyssum parvifl Descurainia pinnata

var. var. var. var. Species name var. var. fallax Descurainia sophia Descurainia sophia Salsola tragus Salsola australis Utah Colorado Dithyrea wislizenii Dimorphocarpa Spinacia oleraceaSuaeda torreyana torreyana Spinacia oleracea Ambrosia acanthicarpa Ambrosia acanthicarpa Chaenactis stevioides Chaenactis stevioides Conyza canadensis glabrata Erigeron bellidiastrum Erigeron bellidiastrum Helianthus annuus lenticularis Helianthus petiolaris ssp. Iva xanthifolia Cyclachaena xanthifolia Lactuca serriola Lactuca serriola Malacothrix sonchoides Malacothrix sonchoides Camelina microcarpa Camelina microcarpa Alyssum minus micranthum Chorispora tenella Chorispora tenella Descurainia pinnata osmiarum rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Cruciferae (Brassicaceae) Appendix D. Confi Family Annual Forbs, cont. Chenopodiaceae Cruciferae (Brassicaceae) Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

Chapter 6: Appendices 107 areas areas areas areas Disturbed areas areas areas Wet Comments Jun Pin- Sage Flower time Aug Park status Native Pres Mar–Jun x x Native Pres Apr–Aug x x Native Pres Mar–Jun x x Native Pres Apr–Jun x x ddle- ax Native Rep Apr–Jun x x owered eld dodder eld Native Rep Jul–Sep x x Common name Nativity white sweet-clover Intro Pres May–Oct x x Disturbed wedgeleaf draba Nativeprairie pepperwort Pres Nativehairy-pod Pres Feb–Julpepperwort May–Jul x x x x clasping pepperwort Intro Pres May–Jul x x Disturbed broom-fl African mustard Intro Rep Apr–Jul Disturbed white-stem blazingstar common mallow Intro Rep Apr–Oct Disturbed tumble mustard Intro Pres May– long-beak fi mustard fi ridge-seeded spurge Native Rep Jun–Aug x x Disturbed stork’s-bill Intro Pres Feb–Oct x x Disturbed small-fl milkvetch dwarf lupine Native Rep Apr–Jul x x Sandy soils corrugate phacelia NativeIves’ phacelia Pres May–Jun Native x Pres Apr–Jun x x x orum micranthiformis Draba cuneifolia Lepidium densifl Lepidium lasiocarpum Streptanthella longirostris Chamaesyce glyptosperma Astragalus nuttallianus var. Lupinus pusillus Phacelia crenulata

var. Species name var. var. orum orum densifl lasiocarpum micranthiformis Draba cuneifolia cuneifolia Lepidium densifl var. Lepidium lasiocarpum var. Melilotus alba Melilotus albus Lepidium perfoliatum Lepidium perfoliatum Utah Colorado Linum aristatum Mesynium aristatum Malcolmia africana Malcolmia africana Mentzelia albicaulisMalva neglecta Acrolasia albicaulis Malva neglecta Sisymbrium altissimum Sisymbrium altissimum Streptanthella longirostris Chamaesyce glyptosperma Cuscuta pentagona Grammica campestris Erodium cicutarium Erodium cicutarium Lupinus pusillus pusillus Phacelia crenulata corrugata Phacelia ivesianaAstragalus nuttallianus var. Phacelia ivesiana rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Family Annual Forbs, cont. Cruciferae (Brassicaceae) Leguminosae (Fabaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Linaceae Cruciferae (Brassicaceae) Loasaceae Malvaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Euphorbiaceae Cuscutaceae (Convolvulaceae) Geraniaceae Leguminosae (Fabaceae) Hydrophyllaceae Hydrophyllaceae Leguminosae (Fabaceae) Appendix D. Confi

108 Annotated Checklist of Vascular Flora: Hovenweep National Monument areas areas areas areas areas Wet Comments Jun Pin- xx Sage Flower time Aug Park status Native Pres Mar–Jun x Native Rep May–Oct x Native Pres Mar–Oct x x Native Pres Apr–Jun x Native Pres May– ower rock- ower Common name Nativity orange kallstroemia Native Rep Jul–Sep Disturbed white-stem evening- primrose woolly plantain spreading eriastrum Native Nativeshy gilia Prescommon gilia Pres May–Jun Mar–Augspreading gilia x Nativedwarf gilia Native Repslender phlox Native Pres x x Pres May–Junspreading wild Apr–Jul Native Nativebuckwheat x Apr–Jun Pres Rep x x x Apr–Jun Mar–Jun x x x x x x Palmer’s wild Palmer’s buckwheat yard knotweed Intro Pres May–Oct Disturbed dwarf purslanecommon purslane Native Intro Pres Rep May–Sep Jun–Sep x Disturbed western rock- jasmine pygmy-fl jasmine bur buttercup Intro Pres Mar–Jun x x Disturbed cut-leaf nightshade Native Rep May–Sep x x Disturbed Wright’s bird’s-beakWright’s Native trumpetangel’s Pres Native Jul–Sep Pres x Apr–Oct x x x Sandy soils blue-eyed Mary Native Pres Apr–Jun x x ora ora orum Gilia leptomeria Microsteris gracilis Eriogonum palmerianum Androsace septentrionalis orthoceras

var. oraparvifl Kallstroemia Species name var. oraparvifl Collinsia orumtrifl Solanum Eriastrum diffusumGilia inconspicua Gilia leptomeria Eriastrum diffusum leptomeria Gilia sinuata Ipomopsis polycladonIpomopsis pumila Ipomopsis polycladon Microsteris gracilis Ipomopsis pumila Eriogonum divaricatum Eriogonum divaricatum humilior Oenothera albicaulis Oenothera albicaulis Plantago patagonica Plantago patagonica Eriogonum palmerianum Polygonum aviculare Polygonum arenastrum Utah Colorado Portulaca halimoidesPortulaca oleracea Portulaca halimoides Portulaca oleracea Androsace occidentalis Androsace occidentalis Androsace septentrionalis Ranunculus testiculatus Ceratocephala Datura wrightiiSolanum trifl Datura wrightii Cordylanthus wrightii Cordylanthus wrightii Collinsia parvifl Kallstroemia parvifl rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polygonaceae Appendix D. Confi Family Annual Forbs, cont. Onagraceae Plantaginaceae Polygonaceae Polygonaceae Portulacaceae Portulacaceae Primulaceae Primulaceae Ranunculaceae Solanaceae Solanaceae Scrophulariaceae Scrophulariaceae Zygophyllaceae

Chapter 6: Appendices 109 areas areas areas areas x Wet Comments Jun Pin- xx xxx Sage Flower time Aug Aug Aug Park status Intro Pres Jun–Aug x x Disturbed Common name Nativity elk sedgewestern sedgeblackcreeper sedge Native Native Native Pres Pres Rep May– Jun–Aug Apr–Jul x x Ross’ sedge Native Pres May– alkali bulrushcrested wheatgrass Native Intro Pres Pres Jun–Sep Jun–Aug x x x Disturbed redtopblue grama Introsmooth brome Pres Native Jun–Sep Intro Pres Pres Jul–Sep Jun–Aug x x x x x Disturbed purple three-awn Native Pres Mar–Sep x x tall wheatgrass Intro Rep Jun–Aug x Disturbed squirreltail Native Pres May– intermediate wheatgrass Salina wildryewestern wheatgrass Native Nativeslender wheatgrass Pres Pres Native Jun–Jul Jun–Aughairy tridens Pres x x Jun–Aug Native x x x Pres May–Jul x x x

var.

ssp. ssp. maritimus paludosus Bromopsis inermis longiseta barbulatum intermedium

Species name var. Carex occidentalisCarex praegracilis Carex occidentalis Carex praegracilis Carex geyeri Carex geyeri Utah Colorado Carex rossii Carex rossii Scirpus maritimus Bolboschoenus Agropyron cristatum Agropyron desertorum Agrostis stoloniferaAristida purpurea Agrostis stolonifera Aristida purpurea Bromus inermis inermis Bouteloua gracilis Chondrosum gracile Elymus elongatus Thinopyrum ponticum Elymus elymoides Elymus elymoides Elymus hispidus Elymus salinus Thinopyrum Elymus smithii Leymus salina Elymus trachycaulus Elymus trachycaulus Pascopyrum smithii Erioneuron pilosum Erioneuron pilosum rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Cyperaceae Cyperaceae Family Graminoids Perennial Cyperaceae Cyperaceae Cyperaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Confi

110 Annotated Checklist of Vascular Flora: Hovenweep National Monument areas areas areas areas areas areas Wet Comments Jun Pin- xx xx x x x Disturbed Sage Flower time Aug Aug Aug Park status Common name Nativity meadow fescue Intro Rep Jun–Aug x Disturbed fowl mannagrass Nativegalleta Pres Jul–Aug Native Pres May– x foxtail barley Native Pres Jun–Aug x x x Disturbed junegrassfoxtail muhlysandhill muhly Native Native Preslittleseed ricegrass Rep Native Jun–Aug Native Repcommon reed Jul–Sep Rep x Jul–Sepbulbous bluegrass Jun–Aug Native x Intro x x Rep Pres Jul–Sep x x Apr–Jul x x x x x Disturbed Canada bluegrass Intromuttongrass Pres Jun–Aug Native x Pres May– x x Kentucky bluegrass Intro Pres May– Sandberg’s bluegrassSandberg’s Native Presalkali sacaton Apr–Sep Native x Pres x Jun–Aug x x x x Disturbed spike dropseedsand dropseed Native Rep Native Jun–Sep Pres Jun–Aug x x x Disturbed micranthum Sporobolus airoides

var. Species name Festuca pratensis Festuca pratensis Utah Colorado Glyceria striataHilaria jamesii Glyceria striata Hilaria jamesii Hordeum jubatum Critesion jubatum Koeleria macranthaMuhlenbergia andina Koeleria macrantha Muhlenbergia andina Muhlenbergia pungens Muhlenbergia pungens Oryzopsis micrantha Piptatherum Phragmites australisPoa bulbosa Phragmites australis Poa bulbosa Poa compressaPoa fendleriana Poa compressa Poa fendleriana Poa pratensis Poa pratensis Poa secundaSporobolus airoides airoides Poa secunda Sporobolus contractus Sporobolus contractus Sporobolus cryptandrus Sporobolus cryptandrus rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Appendix D. Confi Family Perennial Graminoids, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

Chapter 6: Appendices 111 areas areas areas areas areas areas areas areas Wet Comments Jun Pin- xx Sage Flower time May Park status Common name Nativity Baltic rushlongstyle rush rushTorrey’s broad-leaved cattail Native Native Native Pres Pres Native Presjointed goatgrass Pres Jun–Aug May–Jul Jun–Aug Intro Jun–Aug Rep May–Jun x x x x Disturbed needle-and-thread NativeIndian ricegrass Presdesert needlegrass May–Jul Native Native Pres x Pres May–Jul Apr–Jun x x x x x oats Intro Pres Jun–Aug x Disturbed Japanese chessred brome Intro Pres Jun–Aug Intro x Pres Mar–Jun x x Disturbed cheatgrass Intro Pres May–Jul x x Disturbed Chilean chess Intro Rep Apr–May x Disturbed barnyard-grass Intro Rep Aug–Sep x Disturbed six-weeks fescuerabbit barley Native Pres Apr–Jun Intro x Pres May–Jul x x x Disturbed false buffalograss Native Pres Jun–Sep x x Disturbed Bigelow’s bluegrassBigelow’s Native Pres Mar– rabbitfoot-grass Intro Pres Jun–Aug x ater sativa ssp. var. ora cylindricum Hesperostipa comata hymenoides speciosum Polypogon monspeliensis Species name

sativa fatua Avena var. oraoctofl Vulpia var. Juncus longistylusJuncus torreyi latifoliaTypha Juncus longistylus Aegilops cylindrica Juncus torreyi latifolia Typha Cylindropyrum Stipa comata comata Stipa hymenoidesStipa speciosa Achnatherum Juncus arcticus Achnatherum Juncus arcticus Utah Colorado Avena fatua Avena Bromus japonicusBromus rubens Bromus japonicus Anisantha rubens Bromus tectorum Anisantha tectorum Bromus trinii Bromus trinii Echinochloa crus-galli Echinochloa crus-galli Festuca octofl Hordeum murinum Critesion glaucum Munroa squarrosa Monroa squarrosa Poa bigelovii Poa bigelovii Polypogon monspeliensis rmed, historical, and reported taxa of Hovenweep National Monument, organized by life form with ecological notes, cont. Juncaceae Juncaceae Typhaceae Annual Graminoids Gramineae (Poaceae) Family Perennial Graminoids, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Juncaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Confi

112 Annotated Checklist of Vascular Flora: Hovenweep National Monument The Department of the Interior protects and manages the nation’s natural resources and cultural heritage; provides scientifi c and other information about those resources; and honors its special responsibilities to American Indians, Alaska Natives, and affi liated Island Communities.

NPS D-63, May 2009 National Park Service U.S. Department of the Interior

Natural Resource Program Center

Natural Resource Program Center 1201 Oak Ridge Drive, Suite 150 Fort Collins, Colorado 80525 www.nature.nps.gov

EXPERIENCE YOUR AMERICA™