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The Carnivora (Mammalia) from the Middle Miocene Locality of Gračanica (Bugojno Basin, Gornji Vakuf, Bosnia and Herzegovina)
Palaeobiodiversity and Palaeoenvironments https://doi.org/10.1007/s12549-018-0353-0 ORIGINAL PAPER The Carnivora (Mammalia) from the middle Miocene locality of Gračanica (Bugojno Basin, Gornji Vakuf, Bosnia and Herzegovina) Katharina Bastl1,2 & Doris Nagel2 & Michael Morlo3 & Ursula B. Göhlich4 Received: 23 March 2018 /Revised: 4 June 2018 /Accepted: 18 September 2018 # The Author(s) 2018 Abstract The Carnivora (Mammalia) yielded in the coal mine Gračanica in Bosnia and Herzegovina are composed of the caniform families Amphicyonidae (Amphicyon giganteus), Ursidae (Hemicyon goeriachensis, Ursavus brevirhinus) and Mustelidae (indet.) and the feliform family Percrocutidae (Percrocuta miocenica). The site is of middle Miocene age and the biostratigraphical interpretation based on molluscs indicates Langhium, correlating Mammal Zone MN 5. The carnivore faunal assemblage suggests a possible assignement to MN 6 defined by the late occurrence of A. giganteus and the early occurrence of H. goeriachensis and P. miocenica. Despite the scarcity of remains belonging to the order Carnivora, the fossils suggest a diverse fauna including omnivores, mesocarnivores and hypercarnivores of a meat/bone diet as well as Carnivora of small (Mustelidae indet.) to large size (A. giganteus). Faunal similarities can be found with Prebreza (Serbia), Mordoğan, Çandır, Paşalar and Inönü (all Turkey), which are of comparable age. The absence of Felidae is worthy of remark, but could be explained by the general scarcity of carnivoran fossils. Gračanica records the most eastern European occurrence of H. goeriachensis and the first occurrence of A. giganteus outside central Europe except for Namibia (Africa). The Gračanica Carnivora fauna is mostly composed of European elements. Keywords Amphicyon . Hemicyon . -
Oldest Mephitine Cranium and Its Implications for the Origin of Skunks
Oldest mephitine cranium and its implications for the origin of skunks MIECZYSEAW WOLSAN Wolsan, M. 1999. Oldest mephitine cranium and its implications for the origin of skunks. -Acta Palaeontologica Polonica 44,2,223-230. -- Recent molecular studies have recognized the skunks (Mephitinae) to be the sister taxon of a clade comprising the Procyonidae and Mustelidae. These findings are inconsistent with the traditional placement of the skunks among mustelids, which is based on mor- phological evidence from extant taxa. This paper reports on a well-preserved cranium of a young individual, recovered from the middle Miocene deposits (MN 7+8, about 11-12 Ma) of Steinheim am Albuch, Germany. The fossil is the holotype of Palaeo- mephitis steinheimensis Jager, 1839, which is here recognized as a senior subjective syn- onym of Trochotherium cyamoides Fraas, 1870 (consequently, Palaeomephitis Jager, 1839 is a senior subjective synonym of Trochotherium Fraas, 1870). The specimen is identified as the oldest and most primitive mephitine cranium known to date, approach- ing the primitive morphology for the Mephitinae. It exhibits a combination of mephitine (accessory middle-ear chamber, lateral swelling of the squamosal) and mustelid (mus- telid suprameatal fossa) synapomorphies,corroborating the view that skunks are derived from a mustelid ancestor. Its auditory bulla shows a slightly inflated and relatively large caudal entotympanic, which indicates that the uninflated and relatively small caudal entotympanics of adult mephitines, as well as their hypertrophied ectotympanics, are not primitive (as hitherto assumed) but derived, providing a synapomorphy that supports a sister-group relationship between the Mephitinae and Lutrinae. Key words : Palaeomephitis, Trochotherium,Mephitinae, Mustelidae, Carnivora, phy- logeny, taxonomy, morphology. -
University of Florida Thesis Or Dissertation Formatting
UNDERSTANDING CARNIVORAN ECOMORPHOLOGY THROUGH DEEP TIME, WITH A CASE STUDY DURING THE CAT-GAP OF FLORIDA By SHARON ELIZABETH HOLTE A DISSERTATION PRESENTED TO THE GRADUATE SCHOOL OF THE UNIVERSITY OF FLORIDA IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY UNIVERSITY OF FLORIDA 2018 © 2018 Sharon Elizabeth Holte To Dr. Larry, thank you ACKNOWLEDGMENTS I would like to thank my family for encouraging me to pursue my interests. They have always believed in me and never doubted that I would reach my goals. I am eternally grateful to my mentors, Dr. Jim Mead and the late Dr. Larry Agenbroad, who have shaped me as a paleontologist and have provided me to the strength and knowledge to continue to grow as a scientist. I would like to thank my colleagues from the Florida Museum of Natural History who provided insight and open discussion on my research. In particular, I would like to thank Dr. Aldo Rincon for his help in researching procyonids. I am so grateful to Dr. Anne-Claire Fabre; without her understanding of R and knowledge of 3D morphometrics this project would have been an immense struggle. I would also to thank Rachel Short for the late-night work sessions and discussions. I am extremely grateful to my advisor Dr. David Steadman for his comments, feedback, and guidance through my time here at the University of Florida. I also thank my committee, Dr. Bruce MacFadden, Dr. Jon Bloch, Dr. Elizabeth Screaton, for their feedback and encouragement. I am grateful to the geosciences department at East Tennessee State University, the American Museum of Natural History, and the Museum of Comparative Zoology at Harvard for the loans of specimens. -
Hyaenodonts and Carnivorans from the Early Oligocene to Early Miocene of Xianshuihe Formation, Lanzhou Basin, Gansu Province, China
Palaeontologia Electronica http://palaeo-electronica.org HYAENODONTS AND CARNIVORANS FROM THE EARLY OLIGOCENE TO EARLY MIOCENE OF XIANSHUIHE FORMATION, LANZHOU BASIN, GANSU PROVINCE, CHINA Xiaoming Wang, Zhanxiang Qiu, and Banyue Wang ABSTRACT Fieldwork in the 1990s produced a few records of carnivorous mammals (cre- odonts and carnivorans) in the early Oligocene through early Miocene of Lanzhou Basin, Gansu Province, China. Although only five taxa are known so far, most repre- sented by fragmentary material, the new carnivore assemblage is unique and shows little resemblance to faunas elsewhere in eastern Asia. The early Oligocene record is represented by a single premolar of a Hyaenodon, referable to H. pervagus. In the early Miocene, a new species of Hyaenodon, H. weilini sp. nov., represented by sev- eral cheek teeth, is the last record of this genus in east Asia. A horizontal ramus of an amphicyonid, here identified as Ictiocyon cf. I. socialis, is the first record of this taxon in Asia, and a very small musteloid lower carnassial, although difficult to identify to genus, is clearly a new record because nothing of this small size has been known in Asia. The overall faunal composition seems to suggest affinity with northern Eurasia rather than southern Asia. Xiaoming Wang. Department of Vertebrate Paleontology, Natural History Museum of Los Angeles County, 900 Exposition Blvd., Los Angeles, California 90007, United States [email protected] Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, P.O. Box 643, Beijing 100044, China. Zhanxiang Qiu. Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, P.O. Box 643, Beijing 100044, China. -
Sin Título De Diapositiva
SISTEMÀTICA MOLECULAR, FILOGEOGRAFIA I GENÈTICA DE LA CONSERVACIÓ DE MUSTÈLIDS I DE MACACS Josep Marmi i Plana Tesi Doctoral 2004 Departament de Ciències Experimentals i de la Salut UNIVERSITAT POMPEU FABRA Sistemàtica molecular, filogeografia i genètica de la conservació de mustèlids i de macacs Memòria presentada per Josep Mª Marmi Plana per optar al grau de Doctor per la Universitat Pompeu Fabra. Aquest treball ha estat realitzat sota la direcció del Dr. Xavier Domingo i Roura al Departament de Ciències Experimentals i de la Salut de la Universitat Pompeu Fabra. Dr. Xavier Domingo i Roura Josep Mª Marmi i Plana Director de Tesi Barcelona, març de 2004 Dipòsit legal: B.43715-2004 ISBN: 84-688-8523-1 Als meus pares per haver-me ajudat tant durant tots aquests anys Als meus germans A la Queralt A GRAÏMENTS En aquest apartat vull agrair l’ajuda que m’han ofert tot un seguit de persones des que em vaig començar a introduir en el món de la genètica de poblacions i de l’evolució molecular. Sobretot vull agrair als Drs Alfredo Ruíz i Xavier Domingo que em donessin una oportunitat per iniciar-me productivament en aquestes especialitats. Sota la supervisió del primer vaig cursar - juntament amb el col·lega Oscar Ramírez i dins de la llicenciatura de Biologia - l’assignatura de Pràctiques en Empreses i Institucions Públiques amb l’objectiu de realitzar un treball recopilatori d’estudis de diversitat genètica en fauna dels Països Catalans. Amb el segon, després de col·laborar- hi durant una mica més d’un any (entre el setembre de 1998 i el desembre de 1999), vaig iniciar la tesi doctoral que ara presento. -
AMERICAN MUSEUM NOVITATES Published by Thz American MUSEUM of NATURAL HISTORY Number 485 New York City Aug
AMERICAN MUSEUM NOVITATES Published by THz AmERICAN MUSEUM OF NATURAL HISTORY Number 485 New York City Aug. 25, 1931 56.092T EXPLORATIONS, RESEARCHES AND PUBLICATIONS OF PIERRE TEILHARD DE CHARDIN, 1911-1931 WITH MAP AND LEGEND SHOWING CHIEF FOSSIL COLLECTING AREAS OF CHINA, 1885-1931 By HENRY FAIRFIELD OSBORN In recent years the Department of Vertebrate Paleontology of the American Museum has enjoyed the cooperation of Pierre Teilhard de Chardin, especially in the Eocene fauna of France from the Paleocene stage up to the close of the Phosphorites or Lower Oligocene time, also through explorations and researches first reported to the Mus6e National d'Histoire Naturelle of Paris in the year 1920. The Ameri- can Museum Central Asiatic Expedition of the summer of 1930 was fortunate in obtaining the cooperation of Teilhard in the field. On February 10, 1931, he addressed the Osborn Research Club and sum- marized the results of his explorations in China, accompanying his address by the map which is reproduced herewith. The present article is prepared as a brief outline of the principal observations made by Teil- hard in the Eocene of France and Belgium and as a summary of the observations made in China, concluding with an abstract of the address of February 10 above mentioned, a bibliography of his writings now contained in the Osborn Library of the American Museum, and a map showing twenty-three of the chief fossil collecting areas of China pre- pared by Teilhard for the present publication. OLIGOCENE TO MIDDLE EOCENE Under the direction of Marcellin Boule, now the ranking verte- brate palheontologist of France, Teilhard began his studies in 1911 with a synthetic review of the creodonts and true carnivores, of the PHOS- PHORITES DU QUERCY, the famous fissure horizon of Eocene to Oligo- cene age, tracing their phylogeny (1915, p. -
Abstract Volume 7Th Swiss Geoscience Meeting Neuchâtel, 20Th – 21St November 2009
Abstract Volume 7th Swiss Geoscience Meeting Neuchâtel, 20th – 21st November 2009 6. Darwin, Evolution and Palaeontology 198 6. Darwin, Evolution and Palaeontology Lionel Cavin Schweizerische Paläontologische Gesellschaft (SPG/SPS), Kommission der Schweizerischen Paläontologischen Abhandlungen (KSPA), Swiss Commission of Palaeontology Award 6.1 Becker D., Berger J.-P., Hiard F., Menkveld-Gfeller U., Mennecart B. & Scherler L.: Late Aquitanian mammals from the Engehalde locality (Bern, Switzerland) 6.2 Brühwiler T., Bucher H., Goudemand N., Ware D., Hermann E.: Smithian ammonoids (Early Triassic): explosive evolu- tionary radiation following the Permian/Triassic mass extinction 6.3 Buffetaut, E.: Darwinian dinosaurs : missing links or evolutionary failures ? Symposium 6: Darwin, Evolution and Palaeontology 6.4 Girault F. E. & Thierstein H. R.: Diatom assemblage turnover in the NW-Pacific at the 2.73 Ma climate transition 6.5 Girault F. E., Weller A. F. & Thierstein H. R.: Neogene global cooling: diatom size variability in a changing ocean 6.6 Goudemand N., Orchard M., Tafforeau P., Urdy S., Brühwiler T., Bucher H., Brayard A., Galfetti T., Monnet C., Lebrun R. & Zollikofer C.: Paleobiology of Early Triassic conodonts: implications of newly discovered fused clusters imaged by X-ray synchrotron microtomography 6.7 Klug C., Kröger B., Kiessling W., Mullins G.L., Servais T., Frýda J., Korn D. & Turner S.: The Devonian Nekton Revolution 6.8 Klug C., Schulz H. & De Baets K.: Red trilobites with green eyes from the Devonian of Morocco 6.9 Klug C., Schweigert G., Fuchs D. & Dietl G.: First record of a belemnite preserved with beaks, arms and ink sac from the Nusplingen Lithographic Limestone (Kimmeridgian, SW Germany) 6.10 Lavoyer T., Bertrand Y. -
An Arctomorph Carnivoran Skull from the Phosphorites Du Quercy and the Origin of Procyonids
An arctomorph carnivoran skull from the Phosphorites du Quercy and the origin of procyonids MIECZYSEAW WOLSAN and BRIGI'ITE LANGE-BAD& Wolsan, M. & Lange-BadrC, B. 1996. An arctomorph carnivoran skull from the Phosphorites du Quercy and the origin of procyonids. Acta Palaeontologica Poloni- ca 41, 3, 277-298. The size and morphological characteristics of a skull of an arctomorph carnivoran mammal from Mouillac (old collection of the Phosphorites du Quercy, of unknown age) in France closely match those of the holotype of the earliest known procyonid Pseudobassaris riggsi and another skull referred to this species, both from old collections of the Phosphorites du Quercy (Caylus and Mouillac), probably earliest Late Oligocene in age. The skull is more primitive in morphology than those of ~seudobassarisrig&i and every other known procyonid, approaching a hypothetical primitive procyonid morphotype. The only, but methodologically fundamental, departure from this morphotype is the lack of the procyonid supra- meatal fossa, which is the crucial synapomorphy of the family Procyonidae. To explain the phylogenetic and taxonomic status of the arctomorph represented by the skull, three competing hypotheses are put forward. Hypothesis A, which considers the arctomorph as an individual of Pseudobassaris riggsi, assumes that the procyonid suprameatal fossa first appeared in a common ancestor of Pseudo- bassaris and other procyonids but was still of variable occurence within Pseudo- bassaris riggsi. Hypothesis B, which proposes the arctomorph as a member of a new Pseudobassaris species ancestral to Pseudobassaris riggsi, concludes that the procyonid suprameatal fossa arose in Pseudobassaris riggsi and in the Procyonidae independently, excluding Pseudobassaris from the procyonids. -
Comms GSN 23 Complete
A Reference Section for the Otavi Group (Damara Supergroup) in Eastern Kaoko Zone near Ongongo, Namibia P.F. Hoffman1,2,*, S.B. Pruss3, C.L. Blättler4, E.J. Bellefroid5 & B.W. Johnson6 1School of Earth & Ocean Sciences, University of Victoria, Victoria, BC, Canada, V8P 5C2 2Department of Earth & Planetary Sciences, Harvard University, Cambridge, MA 02138, USA 3Department of Geosciences, Smith College, Northampton, MA 01065, USA 4Department of Geophysical Sciences, University of Chicago, Chicago, IL 60637, USA 5Department of Earth & Planetary Sciences, McGill University, Montreal, QC, Canada, H3A 0E8 6Department of Geological & Atmospheric Sciences, Iowa State University, Ames, IA 50011-1027, USA * 1216 Montrose Ave., Victoria, BC, Canada, V8T 2K4. email: <[email protected]> Abstract : A reference section for the Otavi Group (Damara Supergroup) in the East Kaoko Zone near Ongongo is proposed and described. The section is easily accessible, well exposed, suitable for field excursions, and well documented in terms of carbonate lithofacies, depositional sequences and stable- isotope chemostratigraphy. The late Tonian Ombombo Subgroup is 355 m thick above the basal Beesvlakte Formation, which is not included in the section due to poor outcrop and complex structure. The early- middle Cryogenian Abenab Subgroup is 636 m thick and the early Ediacaran Tsumeb Subgroup is 1020 m thick. While the section is complete in terms of formations represented, the Ombombo and lower Abenab subgroups have defined gaps due to intermittent uplift of the northward-sloping Makalani rift shoulder. The upper Abenab and Tsumeb subgroups are relatively thin due to erosion of a broad shallow trough during late Cryogenian glaciation and flexural arching during post-rift thermal subsidence of the carbonate platform. -
Carnivorous Mammals from the Middle Eocene Washakie Formation, Wyoming, USA, and Their Diversity Trajectory in a Post-Warming World
Journal of Paleontology, Volume 95, Memoir 82, 2021, p. 1–115 Copyright © The Author(s), 2021. Published by Cambridge University Press on behalf of The Paleontological Society. All rights reserved. This is an Open Access article, distributed under the terms of the Creative Commons Attribution licence (http://creativecommons.org/licenses/ by/4.0/), which permits unrestricted re-use, distribution, and reproduction in any medium, provided the original work is properly cited. 0022-3360/21/1937-2337 doi: 10.1017/jpa.2020.74 Carnivorous mammals from the middle Eocene Washakie Formation, Wyoming, USA, and their diversity trajectory in a post-warming world Susumu Tomiya,1,2,3 Shawn P. Zack,4 Michelle Spaulding,5 and John J. Flynn2,6 1Center for International Collaboration and Advanced Studies in Primatology, Kyoto University Primate Research Institute, 41-2 Kanrin, Inuyama, Aichi 484-8506, Japan <[email protected]> 2Negaunee Integrative Research and Gantz Family Collections Centers, Field Museum of Natural History, 1400 South Lake Shore Drive, Chicago, Illinois 60605, USA 3University of California Museum of Paleontology, Berkeley, California 94720, USA 4Department of Basic Medical Sciences, University of Arizona College of Medicine, 425 N Fifth Street, Phoenix, Arizona 85004, USA <[email protected]> 5Department of Biological Sciences, Purdue University Northwest, 1401 S US Highway 421, Westville, Indiana 46391, USA <mspauldi@ pnw.edu> 6Richard Gilder Graduate School and Division of Paleontology, American Museum of Natural History, 200 Central Park West, New York, New York 10024, USA <jfl[email protected]> Abstract.—The middle Eocene Washakie Formation of Wyoming, USA, provides a rare window, within a single deposi- tional basin, into the faunal transition that followed the early Eocene warming events. -
Miocene Carnivorans from the Vallès-Penedès Basin (NE Iberian Peninsula)
Departament de Biologia Animal, de Biologia Vegetal i d’Ecologia Unitat d’Antropologia Biològica Miocene carnivorans from the Vallès-Penedès Basin (NE Iberian Peninsula) Josep Maria Robles Giménez Tesi Doctoral 2014 A mi padre y familia. INDEX Index .......................................................................................................................... 7 Preface and Acknowledgments [in Spanish] ....................................................... 13 I.–Introduction and Methodology ........................................................................ 19 Chapter 1. General introduction and aims of this dissertation .......................... 21 1.1. Aims and structure of this work .............................................................. 21 Motivation of this dissertation ................................................................ 21 Type of dissertation and general overview ............................................. 22 1.2. An introduction to the Carnivora ............................................................ 24 What is a carnivoran? ............................................................................. 24 Biology .................................................................................................... 25 Systematics and phylogeny ...................................................................... 28 Evolutionary history ................................................................................ 42 1.3. Carnivoran anatomy ............................................................................... -
Evolution of Hypercarnivory: the Effect of Specialization on Morphological and Taxonomic Diversity
Paleobiology, 30(1), 2004, pp. 108±128 Evolution of hypercarnivory: the effect of specialization on morphological and taxonomic diversity Jill A. Holliday and Scott J. Steppan Abstract.ÐThe effects of specialization on subsequent morphological evolution are poorly under- stood. Specialization has been implicated in both adaptive radiations that result from key inno- vations and evolutionary ``dead ends,'' where specialized characteristics appear to limit subse- quent evolutionary options. Despite much theoretical debate, however, empirical studies remain infrequent. In this paper, we use sister-group comparisons to evaluate the effect of morphological specialization to a particular ecological niche, hypercarnivory, on subsequent taxonomic and mor- phological diversity. Six sets of sister groups are identi®ed in which one clade exhibits hypercar- nivorous characteristics and the sister clade does not. Comparison results are summed across the categories ``hypercarnivore'' and ``sister group.'' We also evaluate whether increasing degrees of specialization are correlated with decreasing phenotypic variation. Results presented here indicate that specialization to hypercarnivory has no effect on taxonomic diversity, but a strong effect on subsequent morphological diversity related to the jaws and dentition, and that increasing special- ization does not correlate with morphological diversity except in the most specialized saber- toothed taxa, which exhibit higher variance than less specialized morphs, possibly due to selection on other characteristics. Jill A. Holliday and Scott J. Steppan. Department of Biological Science, Florida State University, Talla- hassee, Florida 32306-1100. E-mail: [email protected], E-mail: [email protected] Accepted: 21 May 2003 Introduction pattern (e.g., Price and Carr 2000), and others ®nd no effect at all (e.g., Wiegmann et al.