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A New Xinjiangchelyid Turtle from the Middle Jurassic of Xinjiang, China and the Evolution of the Basipterygoid Process in Mesozoic Turtles Rabi Et Al
A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles Rabi et al. Rabi et al. BMC Evolutionary Biology 2013, 13:203 http://www.biomedcentral.com/1471-2148/13/203 Rabi et al. BMC Evolutionary Biology 2013, 13:203 http://www.biomedcentral.com/1471-2148/13/203 RESEARCH ARTICLE Open Access A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles Márton Rabi1,2*, Chang-Fu Zhou3, Oliver Wings4, Sun Ge3 and Walter G Joyce1,5 Abstract Background: Most turtles from the Middle and Late Jurassic of Asia are referred to the newly defined clade Xinjiangchelyidae, a group of mostly shell-based, generalized, small to mid-sized aquatic froms that are widely considered to represent the stem lineage of Cryptodira. Xinjiangchelyids provide us with great insights into the plesiomorphic anatomy of crown-cryptodires, the most diverse group of living turtles, and they are particularly relevant for understanding the origin and early divergence of the primary clades of extant turtles. Results: Exceptionally complete new xinjiangchelyid material from the ?Qigu Formation of the Turpan Basin (Xinjiang Autonomous Province, China) provides new insights into the anatomy of this group and is assigned to Xinjiangchelys wusu n. sp. A phylogenetic analysis places Xinjiangchelys wusu n. sp. in a monophyletic polytomy with other xinjiangchelyids, including Xinjiangchelys junggarensis, X. radiplicatoides, X. levensis and X. latiens. However, the analysis supports the unorthodox, though tentative placement of xinjiangchelyids and sinemydids outside of crown-group Testudines. A particularly interesting new observation is that the skull of this xinjiangchelyid retains such primitive features as a reduced interpterygoid vacuity and basipterygoid processes. -
Tetrapod Track Assemblages from Lower Cretaceous Desert Facies In
Palaeogeography, Palaeoclimatology, Palaeoecology 507 (2018) 1–14 Contents lists available at ScienceDirect Palaeogeography, Palaeoclimatology, Palaeoecology journal homepage: www.elsevier.com/locate/palaeo Tetrapod track assemblages from Lower Cretaceous desert facies in the Ordos Basin, Shaanxi Province, China, and their implications for Mesozoic T paleoecology ⁎ Lida Xinga,b,c, Martin G. Lockleyd, , Yongzhong Tange, Anthony Romiliof, Tao Xue, Xingwen Lie, Yu Tange, Yizhao Lie a State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Beijing 100083, China b School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China c State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China d Dinosaur Trackers Research Group, CB 172, University of Colorado at Denver, PO Box 173364, Denver, CO 80217-3364, USA e Shaanxi Geological Survey, Xi'an 710054, Shaanxi, China f School of Biological Sciences, The University of Queensland, Brisbane, Queensland 4072, Australia ARTICLE INFO ABSTRACT Keywords: Tetrapod ichnofaunas are reported from desert, playa lake facies in the Lower Cretaceous Luohe Formation at Ichnofacies Baodaoshili, Shaanxi Province, China, which represent the first Asian example of an ichnnofauna typical of the Brasilichnium Chelichnus Ichnofacies (Brasilichnium sub-ichnofacies) characteristic of desert habitats. The mammaliomorph Sarmientichnus tracks, assigned to Brasilichnium, -
Investigating Sexual Dimorphism in Ceratopsid Horncores
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2013-01-25 Investigating Sexual Dimorphism in Ceratopsid Horncores Borkovic, Benjamin Borkovic, B. (2013). Investigating Sexual Dimorphism in Ceratopsid Horncores (Unpublished master's thesis). University of Calgary, Calgary, AB. doi:10.11575/PRISM/26635 http://hdl.handle.net/11023/498 master thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Investigating Sexual Dimorphism in Ceratopsid Horncores by Benjamin Borkovic A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE DEPARTMENT OF BIOLOGICAL SCIENCES CALGARY, ALBERTA JANUARY, 2013 © Benjamin Borkovic 2013 Abstract Evidence for sexual dimorphism was investigated in the horncores of two ceratopsid dinosaurs, Triceratops and Centrosaurus apertus. A review of studies of sexual dimorphism in the vertebrate fossil record revealed methods that were selected for use in ceratopsids. Mountain goats, bison, and pronghorn were selected as exemplar taxa for a proof of principle study that tested the selected methods, and informed and guided the investigation of sexual dimorphism in dinosaurs. Skulls of these exemplar taxa were measured in museum collections, and methods of analysing morphological variation were tested for their ability to demonstrate sexual dimorphism in their horns and horncores. -
Paleobios 32:1–42, September 7, 2015 Paleobios
PaleoBios 32:1–42, September 7, 2015 PaleoBios OFFICIAL PUBLICATION OF THE UNIVERSITY OF CALIFORNIA MUSEUM OF PALEONTOLOGY Edwin A. Cadena and James F. Parham (2015). Oldest known marine turtle? A new protostegid from the Lower Cretaceous of Colombia. Cover illustration: Desmatochelys padillai on an early Cretaceous beach. Reconstruction by artist Jorge Blanco, Argentina. Citation: Cadena, E.A. and J.F. Parham. 2015. Oldest known marine turtle? A new protostegid from the Lower Cretaceous of Colombia. PaleoBios 32. ucmp_paleobios_28615. PaleoBios 32:1–42, September 7, 2015 Oldest known marine turtle? A new protostegid from the Lower Cretaceous of Colombia EDWIN A. CADENA1, 2*AND JAMES F. PARHAM3 1 Centro de Investigaciones Paleontológicas, Villa de Leyva, Colombia; [email protected]. 2 Department of Paleoherpetology, Senckenberg Naturmuseum, 60325 Frankfurt am Main, Germany. 3John D. Cooper Archaeological and Paleontological Center, Department of Geological Sciences, California State University, Fullerton, CA 92834, USA; [email protected]. Recent studies suggested that many fossil marine turtles might not be closely related to extant marine turtles (Che- lonioidea). The uncertainty surrounding the origin and phylogenetic position of fossil marine turtles impacts our understanding of turtle evolution and complicates our attempts to develop and justify fossil calibrations for molecular divergence dating. Here we present the description and phylogenetic analysis of a new fossil marine turtle from the Lower Cretaceous (upper Barremian-lower Aptian, >120 Ma) of Colombia that has a minimum age that is >25 million years older than the minimum age of the previously recognized oldest chelonioid. This new fossil taxon,Desmatochelys padillai sp. nov., is represented by a nearly complete skeleton, four additional skulls with articulated lower jaws, and two partial shells. -
The First Occurrence of Eurygnathohippus Van Hoepen, 1930 (Mammalia, Perissodactyla, Equidae) Outside Africa and Its Biogeograph
TO L O N O G E I L C A A P I ' T A A T L E I I A Bollettino della Società Paleontologica Italiana, 58 (2), 2019, 171-179. Modena C N O A S S. P. I. The frst occurrence of Eurygnathohippus Van Hoepen, 1930 (Mammalia, Perissodactyla, Equidae) outside Africa and its biogeographic signifcance Advait Mahesh Jukar, Boyang Sun, Avinash C. Nanda & Raymond L. Bernor A.M. Jukar, Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington DC 20013, USA; [email protected] B. Sun, Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China; University of Chinese Academy of Sciences, Beijing 100039, China; College of Medicine, Department of Anatomy, Laboratory of Evolutionary Biology, Howard University, Washington D.C. 20059, USA; [email protected] A.C. Nanda, Wadia Institute of Himalayan Geology, Dehra Dun 248001, India; [email protected] R.L. Bernor, College of Medicine, Department of Anatomy, Laboratory of Evolutionary Biology, Howard University, Washington D.C. 20059, USA; [email protected] KEY WORDS - South Asia, Pliocene, Biogeography, Dispersal, Siwalik, Hipparionine horses. ABSTRACT - The Pliocene fossil record of hipparionine horses in the Indian Subcontinent is poorly known. Historically, only one species, “Hippotherium” antelopinum Falconer & Cautley, 1849, was described from the Upper Siwaliks. Here, we present the frst evidence of Eurygnathohippus Van Hoepen, 1930, a lineage hitherto only known from Africa, in the Upper Siwaliks during the late Pliocene. Morphologically, the South Asian Eurygnathohippus is most similar to Eurygnathohippus hasumense (Eisenmann, 1983) from Afar, Ethiopia, a species with a similar temporal range. -
Comparative Bone Histology of the Turtle Shell (Carapace and Plastron)
Comparative bone histology of the turtle shell (carapace and plastron): implications for turtle systematics, functional morphology and turtle origins Dissertation zur Erlangung des Doktorgrades (Dr. rer. nat.) der Mathematisch-Naturwissenschaftlichen Fakultät der Rheinischen Friedrich-Wilhelms-Universität zu Bonn Vorgelegt von Dipl. Geol. Torsten Michael Scheyer aus Mannheim-Neckarau Bonn, 2007 Angefertigt mit Genehmigung der Mathematisch-Naturwissenschaftlichen Fakultät der Rheinischen Friedrich-Wilhelms-Universität Bonn 1 Referent: PD Dr. P. Martin Sander 2 Referent: Prof. Dr. Thomas Martin Tag der Promotion: 14. August 2007 Diese Dissertation ist 2007 auf dem Hochschulschriftenserver der ULB Bonn http://hss.ulb.uni-bonn.de/diss_online elektronisch publiziert. Rheinische Friedrich-Wilhelms-Universität Bonn, Januar 2007 Institut für Paläontologie Nussallee 8 53115 Bonn Dipl.-Geol. Torsten M. Scheyer Erklärung Hiermit erkläre ich an Eides statt, dass ich für meine Promotion keine anderen als die angegebenen Hilfsmittel benutzt habe, und dass die inhaltlich und wörtlich aus anderen Werken entnommenen Stellen und Zitate als solche gekennzeichnet sind. Torsten Scheyer Zusammenfassung—Die Knochenhistologie von Schildkrötenpanzern liefert wertvolle Ergebnisse zur Osteoderm- und Panzergenese, zur Rekonstruktion von fossilen Weichgeweben, zu phylogenetischen Hypothesen und zu funktionellen Aspekten des Schildkrötenpanzers, wobei Carapax und das Plastron generell ähnliche Ergebnisse zeigen. Neben intrinsischen, physiologischen Faktoren wird die -
Proganochelys Quenstedti, to Investigate the Early Evolution of the Adductor Chamber and the Sensorial Anatomy in This Taxon
UNIVERSIDADE DE SÃO PAULO FFCLRP - DEPARTAMENTO DE BIOLOGIA PROGRAMA DE PÓS-GRADUAÇÃO EM BIOLOGIA COMPARADA Patterns of morphological evolution in the skull of turtles: contributions from digital paleontology, neuroanatomy and biomechanics Padrões de evolução morfológica no crânio das tartarugas: contribuições da paleontologia digital, neuroanatomia e biomecânica Gabriel de Souza Ferreira RIBEIRÃO PRETO - SP 2019 UNIVERSIDADE DE SÃO PAULO FFCLRP - DEPARTAMENTO DE BIOLOGIA PROGRAMA DE PÓS-GRADUAÇÃO EM BIOLOGIA COMPARADA Patterns of morphological evolution in the skull of turtles: contributions from digital paleontology, neuroanatomy and biomechanics Padrões de evolução morfológica no crânio das tartarugas: contribuições da paleontologia digital, neuroanatomia e biomecânica Gabriel de Souza Ferreira Supervisor: Prof. Dr. Max Cardoso Langer Co-supervisor: Profa. Dra. Madelaine Böhme Tese apresentada à Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto da USP, como parte das exigências para a obtenção do título de Doutor em Ciências, Área: BIOLOGIA COMPARADA. RIBEIRÃO PRETO - SP 2019 Autorizo a reprodução e divulgação total ou parcial deste trabalho, por qualquer meio convencional ou eletrônico, para fins de estudo e pesquisa, desde que citada a fonte FICHA CATALOGRÁFICA Ferreira, Gabriel de Souza Patterns of morphological evolution in the skull of turtles: contributions from digital paleontology, neuroanatomy and biomechanics. 190 p. : il. ; 30cm Tese de doutorado, apresentada ao Departamento de Biologia da Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto/USP – Área de concentração: Biologia Comparada. Orientador: Langer, Max Cardoso. Co-orientadora: Böhme, Madelaine 1. Computed tomography. 2. Digital endocast. 3. Finite-Element Analysis. 4. Testudinata. 5. Skull. Name: Ferreira, Gabriel de Souza Title: Patterns of morphological evolution in the skull of turtles: contributions from digital paleontology, neuroanatomy and biomechanics. -
Phylogenetic Relationships Among Extinct and Extant Turtles: the Position of Pleurodira and the Effects of the Fossils on Rooting Crown-Group Turtles
Contributions to Zoology, 79 (3) 93-106 (2010) Phylogenetic relationships among extinct and extant turtles: the position of Pleurodira and the effects of the fossils on rooting crown-group turtles Juliana Sterli1, 2 1 CONICET - Museo Paleontológico Egidio Feruglio, Av. Fontana 140, 9100 Trelew, Chubut, Argentina 2 E-mail: [email protected] Key words: molecules, morphology, phylogeny, Testudinata, Testudines Abstract Taxonomic nomenclature ........................................................ 94 Taxonomic sampling ................................................................ 94 The origin and evolution of the crown-group of turtles (Crypto- Character sampling ................................................................. 95 dira + Pleurodira) is one of the most interesting topics in turtle Phylogenetic analyses ............................................................. 95 evolution, second perhaps only to the phylogenetic position of Results ............................................................................................... 97 turtles among amniotes. The present contribution focuses on Morphological analysis with extinct taxa .......................... 97 the former problem, exploring the phylogenetic relationships Molecular analyses .................................................................. 97 of extant and extinct turtles based on the most comprehensive Morphological and molecular analysis excluding phylogenetic dataset of morphological and molecular data ana- extinct taxa ................................................................................ -
1 APPENDIX S1 to Evers, Barrett & Benson 2018
APPENDIX S1 to Evers, Barrett & Benson 2018: Anatomy of Rhinochelys pulchriceps (Protostegidae) and marine adaptation during the early evolution of chelonioids Table of Contents ADDITIONAL CT DATA INFORMATION p. 2 ADDITIONAL ANATOMICAL ILLUSTRATIONS p. 3 CHARACTER MODIFICATIONS p. 24 SCORING SOURCES p. 46 CHARACTER OPTIMIZATION p. 51 Methods p. 51 Results p. 52 PCA DATA p. 80 PCA USING MEASUREMENTS OF COLLINS (1970) p. 83 Methods p. 83 Results p. 83 INSTITUTIONAL ABBREVIATIONS p. 85 REFERENCES p. 86 1 ADDITIONAL CT DATA INFORMATION TABLE S1.1. Information about Rhinochelys specimens that were CT scanned for this study. Taxonomy (sensu Voxel size Specimen number Holotype Scanning facility CT Scanner Data availability Reference Collins [1970]) (mm) NHMUK Imaging and Nikon XT H MorphoSource Media CAMSM B55775 R. pulchriceps R. pulchriceps 0.0355 This study Analysis Center 225 ST Group M29973 NHMUK Imaging and Nikon XT H MorphoSource Media NHMUK PV R2226 R. elegans R. elegans 0.0351 This study Analysis Center 225 ST Group M29987 NHMUK Imaging and Nikon XT H MorphoSource Media NHMUK PV OR43980 R. cantabrigiensis R. cantabrigiensis 0.025 This study Analysis Center 225 ST Group M29986 NHMUK Imaging and Nikon XT H MorphoSource Media Evers & Benson CAMSM B55783 - R. cantabrigiensis 0.0204 Analysis Center 225 ST Group M22140 (2018) NHMUK Imaging and Nikon XT H MorphoSource Media CAMSM B55776 - R. elegans 0.0282 This study Analysis Center 225 ST Group M29983 NHMUK Imaging and Nikon XT H MorphoSource Media NHMUK PV OR35197 - R. elegans 0.0171 This study Analysis Center 225 ST Group M29984 2 ADDITIONAL ANATOMICAL ILLUSTRATIONS The following illustrations are provided as additional guides for the description provided in the main text of this paper. -
Norntatesamerican MUSEUM PUBLISHED by the AMERICAN MUSEUM of NATURAL HISTORY CENTRAL PARK WEST at 79TH STREET, NEW YORK, N.Y
NorntatesAMERICAN MUSEUM PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, N.Y. 10024 Number 3048, 13 pp., 6 figures, 1 table July 28, 1992 Dracochelys, a New Cryptodiran Turtle from the Early Cretaceous of China EUGENE S. GAFENEY' AND XIANGKUI YE2 ABSTRACT A turtle skull from the Early Cretaceous Tugulu internal nares, pterygoid forming a cleft-shaped series in the Wuerho district ofXinjiang province, opening for the palatine artery, and flattened skull People's Republic of China, is a new genus of eu- show a close similarity to Hangaiemys, usually cryptodiran turtle. Dracochelys bicuspis is unique placed in the Sinemydidae/Macrobaenidae. How- among cryptodires in possessing a very narrow ever, we place Dracochelys as Eucryptodira incer- maxillary triturating surface with paired cusps on tae sedis, recognizing that the "Macrobaenidae" the labial ridge at the premaxilla-maxilla suture. is presently diagnosed only by characters primitive The very large foramen palatinum posterius, large for Eucryptodira. INTRODUCTION In 1973, Ye figured and briefly mentioned ilar to Hangaiemys, described by Sukhanov an "amphichelydian skull" from the Early and Narmandakh (1974) from the Early Cre- Cretaceous Tugulu series in the Wuerho dis- taceous (Aptian-Albian) of Mongolia (Mon- trict of Xinjiang province, People's Republic golian People's Republic). Hangaiemys is of China. Further preparation and study of known from well-preserved skulls and shells, this specimen shows that it is a new genus of and, on the basis of the shell morphology, cryptodire. The purpose of this paper is to has been placed in the family Macrobaenidae describe and name this new taxon. -
20 New Candidates (For Evaluation in 2020)
20 New candidates (for evaluation in 2020) Bolivia: -Torotoro Brazil: -Caminhos dos Cânions do Sul -Serido China: -Longyan -Xingyi DPR Korea: -Mount Paektu Ecuador: -Napo Sumaco -Tungurahua France: -Armorique -Normandie-Maine Germany: -Ries Iran: -Aras Jordan: -Mujib Mexico : -Huasteca Potosina New Zealand: -Waitaki Whitestone Poland : -Land of Extinct Volcanoes Sweden: -Platåbergen Thailand: -Khorat UK: -Mourne Gullion Strangford Vietnam: -Ly Son-Sa Huynh 3 Extension requests < 10 %: Czech Republic : -Boheminan Paradise Germany: -Thuringia Inselsberg – Drei Gleichen -Vulkaneifel Disclaimer The Secretariat of UNESCO does not represent or endorse the accuracy of reliability of any advice, opinion, statement or other information or documentation provided by the States Parties to the Secretariat of UNESCO. The publication of any such advice, opinion, statement or other information documentation on the website and/or on working documents also does not imply the expression of any opinion whatsoever on the part of the Secretariat of UNESCO concerning the legal status of any country, territory, city or area or of its boundaries. Applicant UNESCO Global Geopark Torotoro, Bolivia Geographical and geological summary Location of the Torotoro Andean Geopark, Aspiring Unesco, in Central Bolivia, South America. Location of the Torotoro Andean Geopark, Aspiring Unesco, in Potosí Department, Bolivia. 1. Physical and human geography Aspiring Torotoro Andean Geopark, located in the Province Charcas, North of Potosí Department, Bolivia, has the same limits of the Municipality of Torotoro. The most used access occurs through the city of Cochabamba, whose distance is 134 km. From Potosí city, the distance is 552 km. The Torotoro's coordinates are 18°08'01"S and 65°45'47"W, and the area is 118,218 km². -
Lihoreau-Et-Al.Merged-PDF-Proo
The Libycosaurus (Hippopotamoidea, Artiodactyla) intercontinental dispersal event at the early late Miocene revealed by new fossil remains from Kasserine area, Tunisia Fabrice Lihoreau, El Mabrouk Essid, Hayet Khayati Ammar, Laurent Marivaux, Wissem Marzougui, Rodolphe Tabuce, Rim Temani, Monique Vianey-Liaud, Gilles Merzeraud To cite this version: Fabrice Lihoreau, El Mabrouk Essid, Hayet Khayati Ammar, Laurent Marivaux, Wissem Marzougui, et al.. The Libycosaurus (Hippopotamoidea, Artiodactyla) intercontinental dispersal event at the early late Miocene revealed by new fossil remains from Kasserine area, Tunisia. Historical Biology, Taylor & Francis, 2021, 33 (2), pp.146-158. 10.1080/08912963.2019.1596088. hal-02082905 HAL Id: hal-02082905 https://hal.umontpellier.fr/hal-02082905 Submitted on 4 Nov 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. Historical Biology For Peer Review Only The Libycosaurus (Hippopotamoidea, Artiodactyla) intercontinental dispersal event at the early Late Miocene revealed by new fossil remains from Kasserine area,