DOCSLIB.ORG

The Ammonoids from the Grès Du Kahla Supérieur of Timimoun

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
The Ammonoids from the Grès Du Kahla Supérieur of Timimoun Fossil Record 13 (1) 2010, 13–34 / DOI 10.1002/mmng.200900009 The ammonoids from the Grs du Kahla suprieur of Timimoun (Middle-early Late Tournaisian; Gourara, Algeria) Dieter Korn*,1, Volker Ebbighausen2 and Jrgen Bockwinkel3 1 Museum fr Naturkunde Berlin, Invalidenstraße 43, 10115 Berlin, Germany. E-mail: [email protected] 2 Engstenberger Hhe 12, 51519 Odenthal, Germany. E-mail: [email protected] 3 Dechant-Feinstraße 22, 51375 Leverkusen, Germany. E-mail: [email protected] Abstract Received 18 June 2009 The ammonoids from two very prolific horizons in the upper Kahla Sandstone (Tour- Accepted 7 August 2009 naisian, Early Carboniferous) of Timimoun (Gourara, Algeria) are described monogra- Published 19 February 2010 phically. Nine new ammonoid species are introduced: Imitoceras altilobatum n. sp., Triimitoceras amplisellatum n. sp., Kazakhstania inequalis n. sp., Acrocanites imperfec- Key Words tus n. sp., Xinjiangites scalaris n. sp., and Becanites canalifer n. sp. from the North African Kazakhstania-Acrocanites Assemblage as well as Acrocanites recurvus n. sp., Ammonoidea Becanites singularis n. sp., and Becanites inflateralis n. sp. from the Pericyclus-Progo- Mississippian niatites Assemblage. The first of these is one of the most diverse ammonoid faunas Carboniferous known from this time interval. North Africa taxonomy Introduction Prodromites, Muensteroceras, and Protocanites. Its similarity to the fauna of the Chouteau Limestone led Miller & Collinson (1951) to assume that the Middle and early Late Tournaisian ammonoids belong two are time equivalents. to the poorest known representatives in the Palaeozoic Kentucky – Early, middle, and late Osagean ammonoid history of the clade. In some regions with otherwise assemblages (Work & Mason 2003, 2004, 2005) rich Early Carboniferous ammonoid records, this time show a succession similar to the occurrences in Al- interval is represented by black shales and cherts, geria. The Cave Run Lake fauna (Nancy Member of which are extremely poor in macrofossils. Therefore, the Borden Formation; early Osagean) with Kazakh- ammonoid faunas from this interval are reported from stania, Furnishoceras, and Muensteroceras (Work & only a few regions. Mason 2005) probably represents a time equivalent Missouri – The Chouteau Limestone and Northview of the lower assemblage from the Grs de Kahla Shale yielded ammonoid faunas (Smith 1903; Miller suprieur. & Collinson 1951), which probably belong in the Belgium – The assemblages first described by de Ko- Middle Tournaisian or earliest Late Tournaisian. ninck (1844, 1880) and later revised by Delpine From these two formations, the genera Imitoceras, (1940) from the Calcaire de Vaulx and the Calcaire Masonoceras, Gattendorfia (?), Prodromites, Xin- de Calonne contain the genera Acrocanites, Xinjian- jiangites, Goniocyclus (restricted to the Chouteau gites, and Becanites (Calcaire de Vaulx) as well as Limestone), and Protocanites are known. Imitoceras, Pericyclus, Xinjiangites, Temertassetia, Indiana – The Rockford Limestone was very produc- Jerania, Bouhamedites, and Progoniatites (Calcaire tive for ammonoid faunas (Hall 1860; Miller 1891; de Calonne); they have probably an earliest Late Gutschick & Treckman 1957); they most likely Tournaisian age. come from the earliest Late Tournaisian. The fauna Thuringia – Phosphatic nodules in the Rußschiefer near is composed of the genera Imitoceras, Gattendorfia, Zadelsdorf yielded ammonoids of the genera Gatten- * Corresponding author # 2010 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim 14 Korn,D.etal.:AmmonoidsfromtheGre` s du Kahla supe´rieur dorfia, Pericyclus, Acrocanites, Xinjiangites, Steno- and Becanites (Sheng 1984; Liang & Wang 1988, cyclus, Progoniatites, and Becanites (?) (Schinde- 1991; Ruan 1995). Parts of this fauna are, in its wolf 1922, 1926, 1939). This appears to be a mixed genus composition, very similar to the assemblage fauna, of which at least some elements have a Late from the Grs de Kahla suprieur. Tournaisian age. New South Wales – Xinjiangites and Protocanites oc- Montagne Noire – The Horizon Lydiennes (Ly- cur in the Namoi Formation together with a priono- diennes Formation) yielded a small ammonoid fauna ceratid (Delpine 1941; Campbell et al. 1983). The with Globimitoceras, Gattendorfia, Goniocyclus, Ni- fauna may be a time equivalent of the Grs de Kah- grocyclus, and Eocanites (Bhm 1935; Korn & Feist la suprieur. 2007); this fauna suggest that a Middle Tournaisian age is represented. North African ammonoid faunas from the Middle and Kazakhstan – Librovitch (1940) described species of early Late Tournaisian have been described from var- the genera Acutimitoceras, Paragattendorfia (?), ious places: Gattendorfia, Karagandoceras, Kazakhstania, Xin- –– Area of Taouz in the Tafilalt (Anti-Atlas) – Faunas jiangites, Asiacyclus, and Eocanites from the Kassi- from the lower part of the Oued Znagui Formation ni Beds and the genera Asiacyclus, Xinjiangites, and yielded the genera Acutimitoceras (?), Gattendorfia, Terektytes from the Terekty Beds. The Kassini Beds Jdaidites, Goniocyclus, and Protocanites (Korn et al. in particular yielded an assemblage similar to the 2002, 2007); they can be attributed to the Middle Grs de Kahla suprieur. Tournaisian. North Xinjiang – The lower Heishantou (Dongguloba- –– Aguelmous, Ma’der region (Anti-Atlas) – time sitao) Formation contains a rich ammonoid fauna equivalent faunas contain Globimitoceras, Goniocy- with the genera Gattendorfia, Kazakhstania, Xin- clus, and Protocanites (Ebbighausen & Bockwinkel, jiangites, Pericyclus, Zhifangoceras, Protocanites, 2007). 500 km Figure 1. The outcrop of Palaeozoic rocks in northern Africa and Iberia and the geographic position (marked by an asterisk) of the fossil localities in the Grs de Kahla suprieur near Timimoun. museum-fossilrecord.wiley-vch.de # 2010 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim Fossil Record 13 (1) 2010, 13–34 15 –– Oued Smide north of the Ma’der – another small fauna with only Gattendorfia and Acrocanites was described by Korn et al. (2007). –– Area of Taouz in the Tafilalt (Anti-Atlas) – The fauna from the middle part of the Oued Znagui For- mation is composed of Irinoceras, Triimitoceras, Muensteroceras, Mouydiria, Ouaoufilalites, Bouha- medites, Progoniatites, Pericyclus, Helicocyclus, Je- rania, and Becanites (Korn et al. 2003a). This fauna has an early Late Tournaisian age and can be inter- preted as time equivalent with the younger of the here described faunas (i.e. samples TIM-A1 and TIM-C1). –– Area of Timimoun (Gourara, Algeria) – The fossili- ferous section was discovered by Meyendorff (1939), who mentioned Early Tournaisian ammo- noids (Ebbighausen et al. 2004). Some ammonoids from the Grs de Kahla suprieur were figured by Conrad (1984). These represent the genera Acroca- nites and Becanites. New findings from this locality will be described in this article. The problematicum Duodecimedusina stella has already been described by Korn et al. (2008). –– Oued Temertasset and Hassi Habadra in the Mouy- dir (Algeria) – the Argiles de Teguentour are al- ready well known for their ammonoid faunas (Ter- mier & Termier 1950; Follot 1953; Conrad & Pareyn 1968; Conrad 1984). In these articles, only species of the genera Imitoceras, Pericyclus, Muen- steroceras, Rotopericyclus, and Progoniatites were figured. A detailed monographic description of the fauna will be published in a parallel article (Korn et al. 2010). Geological setting The ammonoid faunas described here come from two localities 28 and 15 km west-southwest of Timimoun (Gourara, western Algeria), at the south-western margin of the Grand Erg Occidental (Figs 1, 2). Carboniferous sedimentary rocks are exposed between the sand dunes of the Grand Erg, extending over a distance of approxi- mately 40 km in a SW–NE erosion window that ranges in width between a few hundreds of metres and 10 km. The rocks are gently inclined towards the north-east. Their succession (Conrad 1984) consists mainly of shales with several intercalated sandstone formations; carbonates are very rare and restricted to distinct hori- zons. The Carboniferous succession of Timimoun begins with the Grs de Kahla, a more than 300 m thick succession of shales with prominent sandstone interca- lations, of which the lower portion at least partly has Figure 2. Stratigraphic section of the Early Carboniferous (Mis- a Late Devonian age (Conrad 1984). The Grs de sissippian) rock succession near Timimoun with the position of the assemblages from the Grs de Kahla suprieur (After Ru- Kahla suprieur has a thickness of about 80 m and zhencev & Bogoslovskaya 1971; Riley 1990; Korn et al. 2004, contains two major shaly portions, of which the lower 2007). For a description of the faunas from the Ammonellip- one yielded a fauna of the Gattendorfia-Kahlacanites sites-Merocanites Assemblage and Bollandoceras assemblages, Assemblage (Ebbighausen et al. 2004; Korn et al. see Ebbighausen et al. (2010) and Bockwinkel et al. (2010). # 2010 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim museum-fossilrecord.wiley-vch.de 16 Korn,D.etal.:AmmonoidsfromtheGre` s du Kahla supe´rieur Figure 3. Stratigraphic scheme of the Early Carboniferous (Mississippian) chronostratigraphy and ammonoid zonation, with corre- lation of the North African ammonoid assemblages (after Korn et al. 2004, 2007). Highlighted the position of the assemblages from the Grs de Kahla suprieur. 2007). In the upper of the two shaly intervals, four A comparable fauna with Kazakhstania, but with the horizons with ammonoids were
Load more

Recommended publications
  • Suture with Pointed Flank Lobe and ... -.: Palaeontologia Polonica
    210 JERZY DZIK Genus Posttornoceras Wedekind, 1910 Type species: Posttornoceras balvei Wedekind, 1910 from the mid Famennian Platyclymenia annulata Zone of the Rhenisch Slate Mountains. Diagnosis. — Suture with pointed flank lobe and angular or pointed dorsolateral lobe. Remarks. — Becker (1993b) proposed Exotornoceras for the most primitive members of the lineage with a relatively shallow dorsolateral lobe. The difference seems too minor and continuity too apparent to make this taxonomical subdivision practical. Becker (2002) suggested that this lineage was rooted in Gundolficeras, which is supported by the data from the Holy Cross Mountains. The suture of Posttornoceras is similar to that of Sporadoceras, but these end−members of unrelated lin− eages differ rather significantly in the geometry of the septum (Becker 1993b; Korn 1999). In Posttornoceras the parts of the whorl in contact with the preceding whorl are much less extensive, the dorsolateral saddle is much shorter and of a somewhat angular appearance. This is obviously a reflection of the difference in the whorl expansion rate between the tornoceratids and cheiloceratids. Posttornoceras superstes (Wedekind, 1908) (Figs 154A and 159) Type horizon and locality: Early Famennian at Nehden−Schurbusch, Rhenish Slate Mountains (Becker 1993b). Diagnosis. — Suture with pointed tip of the flank lobe and roundedly angulate dorsolateral lobe. Remarks.—Gephyroceras niedzwiedzkii of Dybczyński (1913) from Sieklucki’s brickpit was repre− sented by a specimen (probably lost) significantly larger than those described by Becker (1993b). The differ− ence in proportions of suture seem to result from this ontogenetic difference, that is mostly from increase of the whorl compression with growth. Distribution. — Reworked at Sieklucki’s brickpit in Kielce.
    [Show full text]
  • ABHANDLUNGEN DER GEOLOGISCHEN BUNDESANSTALT Abh
    ©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at ABHANDLUNGEN DER GEOLOGISCHEN BUNDESANSTALT Abh. Geol. B.-A. ISSN 0016–7800 ISBN 3-85316-14-X Band 57 S. 167–180 Wien, Februar 2002 Cephalopods – Present and Past Editors: H. Summesberger, K. Histon & A. Daurer Morphometric Analyses and Taxonomy of Oxyconic Goniatites (Paratornoceratinae n. subfam.) from the Early Famennian of the Tafilalt (Anti-Atlas, Morocco) VOLKER EBBIGHAUSEN, RALPH THOMAS BECKER & JÜRGEN BOCKWINKEL*) 13 Text-Figures and 1 Table A contribution to IGCP 421 Morocco Upper Devonian Ammonoids Morphometry Taxonomy Contents Zusammenfassung ...................................................................................................... 167 Abstract ................................................................................................................. 167 1. Introduction ............................................................................................................. 169 2. Locality and Methods .................................................................................................... 169 3. Morphometric Analyses of Dar Kaoua Paratornoceratinae ................................................................. 170 3.1. Size Distribution .................................................................................................... 170 3.2. Shell Parameters ................................................................................................... 172 4. Systematics ............................................................................................................
    [Show full text]
  • ICS Subcommission Annual Report 2015
    INTERNATIONAL UNION OF GEOLOGICAL SCIENCES INTERNATIONAL COMMISSION ON STRATIGRAPHY Chair Prof. Stanley FINNEY, Department of Geological Sciences, California State University at Long Beach, Long Beach, CA 90840, USA TEL: 1-562-985-8637 (office); FAX: 1-562-985-8638; E-mail: [email protected] Vice Chair Prof. Shanchi PENG, Nanjing Institute of Geology & Palaeontology, Chinese Academy of Sciences, 39 East Beijing St., Nanjing 210008, China TEL and FAX: 86-25-8328 2135; E-mail: [email protected] Secretary General Prof. Paul R. BOWN, Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, UK TEL: 44-0-20-7504-2431 office; FAX 44-0-20-7388-7614; E-mail: [email protected] December 2015 Compiled ICS Subcommission Annual Reports for 2015 SUBCOMMISSION ON QUATERNARY STRATIGRAPHY ANNUAL REPORT 2015 Chair Prof. Martin J. HEAD, Department of Earth Sciences, Brock University, St. Catharines, Ontario L2S 3A1, Canada TEL: 1-905-688-5550 ext 5216 (office); FAX: 1-905-641-8186; E-mail: [email protected] Vice Chair Prof. Brad PILLANS, Research School of Earth Sciences, The Australian National University, Canberra, Act, 0200, Australia Tel: +61-2-6125 9644 (Office); +61-2-6249 1507 (Lab); E-mail: [email protected] Secretary Dr. Jan ZALASIEWICZ, Department of Geology, University of Leicester, Leicester, LE1 7RH, UK Tel: +44 (0)116 252 3928; Fax: +44 (0)116 252 3918; E-mail: [email protected] & [email protected] 1. TITLE OF CONSTITUENT BODY Subcommission on Quaternary Stratigraphy (SQS) 2. OVERALL OBJECTIVES, AND FIT WITHIN IUGS SCIENCE POLICY 1.
    [Show full text]
  • Alpinites and Other Posttornoceratidae (Goniatitida, Famennian)
    Mitt. Mus. Nat.kd. Berl., Geowiss. Reihe 5 (2002) 51-73 10.11.2002 Alpinites and other Posttornoceratidae (Goniatitida, Famennian) R. Thomas Becker' With 6 figures, 1 table and 3 plates Summary The rediscovery of the supposedly lost type allows a revision of Alpinites Bogoslovskiy, 1971, the most advanced genus of the Posttornoceratidae. The type-species, Alp. kayseri Schindewolf, 1923, is so far only known from the Carnic Alps. Alp. schultzei n. sp. from the eastern Anti-Atlas of Morocco is closely related to AZp. kajruktensis n. sp. (= Alp. kayseri in Bogoslovskiy 1971) from Kazakhstan. A second new and more common species of southern Morocco, Alp. zigzag n. sp., is also known from the Holy Cross Mountains (Poland). The taxonomy and phylogeny of other Posttornoceratidae are discussed. The holotype of Exotornoceras nehdense (Lange, 1929) was recovered and is re-illustrated; it is conspecific with Exot. superstes (Wedekind, 1908). The genus and species is also here first recorded from Morocco. Post. weyeri Korn, 1999 is a subjective synonym of Post. posthurnurn (Wedekind, 1918) in which strongly biconvex growth lines, as typical for the family, are observed for the first time. Goniatites lenticularis Richter, 1848 is a nomen dubium within Discoclyrnenia, Clyrnenia polytrichus in Richter (1848) is a Falcitornoceras. It seems possible to distinguish an extreme thin and trochoid Disco. haueri (Miinster, 1840) from the tegoid Disco. cucullata (v. Buch, 1839). Various taxa are excluded from the Posttornoceratidae. Posttornoceras sapiens Korn, 1999 forms the type-species of Maidero- ceras n. gen.. Discoclyrnenia n. sp. of Miiller (1956) is assigned to Maid. rnuelleri n.
    [Show full text]
  • Paleontological Contributions
    THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS May 15, 1970 Paper 47 SIGNIFICANCE OF SUTURES IN PHYLOGENY OF AMMONOIDEA JURGEN KULLMANN AND JOST WIEDMANN Universinit Tubingen, Germany ABSTRACT Because of their complex structure ammonoid sutures offer best possibilities for the recognition of homologies. Sutures comprise a set of individual elements, which may be changed during the course of ontogeny and phylogeny as a result of heterotopy, hetero- morphy, and heterochrony. By means of a morphogenetic symbol terminology, sutural formulas may be established which show the composition of adult sutures as well as their ontogenetic development. WEDEKIND ' S terminology system is preferred because it is the oldest and morphogenetically the most consequent, whereas RUZHENTSEV ' S system seems to be inadequate because of its usage of different symbols for homologous elements. WEDEKIND ' S system includes only five symbols: E (for external lobe), L (for lateral lobe), I (for internal lobe), A (for adventitious lobe), U (for umbilical lobe). Investigations on ontogenetic development show that all taxonomic groups of the entire superorder Ammonoidea can be compared one with another by means of their sutural development, expressed by their sutural formulas. Most of the higher and many of the lower taxa can be solely characterized and arranged in phylogenetic relationship by use of their sutural formulas. INTRODUCTION Today very few ammonoid workers doubt the (e.g., conch shape, sculpture, growth lines) rep- importance of sutures as indication of ammonoid resent less complicated structures; therefore, phylogeny. The considerable advances in our numerous homeomorphs restrict the usefulness of knowledge of ammonoid evolution during recent these features for phylogenetic investigations.
    [Show full text]
  • Sepkoski, J.J. 1992. Compendium of Fossil Marine Animal Families
    MILWAUKEE PUBLIC MUSEUM Contributions . In BIOLOGY and GEOLOGY Number 83 March 1,1992 A Compendium of Fossil Marine Animal Families 2nd edition J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions . In BIOLOGY and GEOLOGY Number 83 March 1,1992 A Compendium of Fossil Marine Animal Families 2nd edition J. John Sepkoski, Jr. Department of the Geophysical Sciences University of Chicago Chicago, Illinois 60637 Milwaukee Public Museum Contributions in Biology and Geology Rodney Watkins, Editor (Reviewer for this paper was P.M. Sheehan) This publication is priced at $25.00 and may be obtained by writing to the Museum Gift Shop, Milwaukee Public Museum, 800 West Wells Street, Milwaukee, WI 53233. Orders must also include $3.00 for shipping and handling ($4.00 for foreign destinations) and must be accompanied by money order or check drawn on U.S. bank. Money orders or checks should be made payable to the Milwaukee Public Museum. Wisconsin residents please add 5% sales tax. In addition, a diskette in ASCII format (DOS) containing the data in this publication is priced at $25.00. Diskettes should be ordered from the Geology Section, Milwaukee Public Museum, 800 West Wells Street, Milwaukee, WI 53233. Specify 3Y. inch or 5Y. inch diskette size when ordering. Checks or money orders for diskettes should be made payable to "GeologySection, Milwaukee Public Museum," and fees for shipping and handling included as stated above. Profits support the research effort of the GeologySection. ISBN 0-89326-168-8 ©1992Milwaukee Public Museum Sponsored by Milwaukee County Contents Abstract ....... 1 Introduction.. ... 2 Stratigraphic codes. 8 The Compendium 14 Actinopoda.
    [Show full text]
  • Abstracts and Program. – 9Th International Symposium Cephalopods ‒ Present and Past in Combination with the 5Th
    See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/265856753 Abstracts and program. – 9th International Symposium Cephalopods ‒ Present and Past in combination with the 5th... Conference Paper · September 2014 CITATIONS READS 0 319 2 authors: Christian Klug Dirk Fuchs University of Zurich 79 PUBLICATIONS 833 CITATIONS 186 PUBLICATIONS 2,148 CITATIONS SEE PROFILE SEE PROFILE Some of the authors of this publication are also working on these related projects: Exceptionally preserved fossil coleoids View project Paleontological and Ecological Changes during the Devonian and Carboniferous in the Anti-Atlas of Morocco View project All content following this page was uploaded by Christian Klug on 22 September 2014. The user has requested enhancement of the downloaded file. in combination with the 5th International Symposium Coleoid Cephalopods through Time Abstracts and program Edited by Christian Klug (Zürich) & Dirk Fuchs (Sapporo) Paläontologisches Institut und Museum, Universität Zürich Cephalopods ‒ Present and Past 9 & Coleoids through Time 5 Zürich 2014 ____________________________________________________________________________ 2 Cephalopods ‒ Present and Past 9 & Coleoids through Time 5 Zürich 2014 ____________________________________________________________________________ 9th International Symposium Cephalopods ‒ Present and Past in combination with the 5th International Symposium Coleoid Cephalopods through Time Edited by Christian Klug (Zürich) & Dirk Fuchs (Sapporo) Paläontologisches Institut und Museum Universität Zürich, September 2014 3 Cephalopods ‒ Present and Past 9 & Coleoids through Time 5 Zürich 2014 ____________________________________________________________________________ Scientific Committee Prof. Dr. Hugo Bucher (Zürich, Switzerland) Dr. Larisa Doguzhaeva (Moscow, Russia) Dr. Dirk Fuchs (Hokkaido University, Japan) Dr. Christian Klug (Zürich, Switzerland) Dr. Dieter Korn (Berlin, Germany) Dr. Neil Landman (New York, USA) Prof. Pascal Neige (Dijon, France) Dr.
    [Show full text]
  • High Resolution Stratigraphy of the Devonian-Carboniferous Transitional Beds in the Rhenish Mountains
    Mitt. Mus. Nat.kd. Berl., Geowiss. Reihe 6 (2003) 79-124 10. 11.2003 High resolution stratigraphy of the Devonian-Carboniferous transitional beds in the Rhenish Mountains Dieter Korn' & Dieter Weyer2 With 26 figures, 1 table, and 4 plates Summary The Devonian-Carboniferous Boundary sections at Hasselbachtal, Oese, Apricke, and Ober-Rodinghausen, all located at the northern margin of the Rhenish Mountains, were measured in detail. A semi-quantitative evaluation of the carbonate content and resulting carbonate curves permitted a highly exact correlation of these sections. This result is supported by data on ammonoid records and volcanoclastic horizons. Carbonate fluctuations within the Hangenberg Limestone are regarded as showing a 100000 years cyclicity. A new ammonoid genus Hasselbachia n. gen. and the species Paprothites ruzhencevi n. sp. are described. Among the rugose corals, a new species Hillaxon hassel n. sp., is erected. Key words: Ammonoidea, Rugosa, Devonian-Carboniferous Boundary, Rhenish Mountains, biostratigraphy, lithostratigraphy. correlation. Zusammenfassung Die DevodKarbon-Grenzprofile Hasselbachtal, Oese, Apricke und Ober-Rodinghausen, alle auf der Nordflanke des Rem- scheid-Altenaer Sattels gelegen, wurden detailliert aufgenommen. Die halbquantitative Ermittlung des Karbonatgehaltes und daraus resultierenden Karbonatkurven eignen sich fur eine sehr genaue Korrelation dieser Profile, die durch Funde von Ammonoideen sowie durch vulkanoklastische Horizonte unterstutzt wird. Wechsel irn Karbonatgehalt werden als 100000Jahres- Zyklizitat gedeutet. Die neue Ammonoideen-Gattung Hasselbachia n. gen. sowie die Art Paprothites ruzhencevi n. sp. werden beschrieben. Unter den rugosen Korallen wird die neue Art Hillaxon hassel n. sp. errrichtet. Schliisselworter: Ammonoidea, Rugosa, DevodKarbon-Grenze, Rheinisches Schiefergebirge. Biostratigraphie, Lithostrati- graphie, Korrelation. Introduction 1984, Higgs et al. 1993), ostracods (e.g., Bless 1983, Bless & Groos-Uffenorde 1984, G.
    [Show full text]
  • Early Carboniferous; Morocco)
    Fossil Record 9(1) (2006), 87–129 / DOI 10.1002/mmng.200600003 A new ammonoid fauna from the Gattendorfia-Eocanites Genozone of the Anti-Atlas (Early Carboniferous; Morocco) Ju¨ rgen Bockwinkel*,1 & Volker Ebbighausen**,2 1 Dechant-Feinstraße 22, D-51375 Leverkusen, Germany 2 Engstenberger Ho¨ he 12, D-51519 Odenthal, Germany Received August 4, 2005, accepted September 19, 2005 Published online zzz With 40 figures Key words: ammonoids, biostratigraphy, Early Carboniferous, Anti-Atlas, Morocco. Abstract From the eastern part of the Tafilalt (Anti-Atlas, Morocco), rich assemblages of ammonoids of the Gattendorfia-Eocanites Genozone are described. The detailed investigation of an outcrop near Mfis in the south-eastern Tafilalt yielded a diverse fauna with 24 species, of which 13 are new. The genus Weyerella n. gen. is newly erected with Weyerella protecta n. sp. as type species. Furthermore, the following new species are described: Paragattendorfia aboussalamae n. sp., Acutimitoceras hollardi n. sp., Acutimitoceras posterum n. sp., Acutimitoceras mfisense n. sp., Acutimitoceras occidentale n. sp., Imitoceras oxydentale n. sp., Kazakhstania nitida n. sp., Zadelsdorfia debouaaensis n. sp., Weyerella minor n. sp., Eocanites simplex n. sp., Eocanites rtbeckeri n. sp., and Eocanites dkorni n. sp. The state of preservation of the ammonoids as limonitic steinkern specimens per- mits a detailed study of sutures and conch ontogeny. Schlu¨ sselwo¨ rter: Ammonoidea, Biostratigraphie, Unter-Karbon, Anti-Atlas, Marokko. Zusammenfassung Aus dem o¨ stlichen Tafilalt (Anti-Atlas, Marokko) wird eine Ammonoideen-reiche Abfolge aus der Gattendorfia-Eocanites Genus-Zone beschrieben. Ein im Detail untersuchtes Profil bei Mfis hat eine individuenreiche und diverse pelagische Fauna mit 24 Arten geliefert, von denen 13 neu sind.
    [Show full text]
  • Palaeobiogeographic Relationships and Diversity of Upper Devonian Ammonoids from Western Australia
    Records of the Western Australian Museum Supplement No. 58: 385-401 (2000). Palaeobiogeographic relationships and diversity of Upper Devonian ammonoids from Western Australia R. Thomas Becker Museum fur Naturkunde, Invalidenstr. 43, D-10115 Berlin, Germany e-mail: [email protected] Abstract - Upper Devonian ammonoids from the Canning Basin of Western Australia represent one of the most diverse faunas globally known. It consists of cosmopolitan (pantropical), endemic and "spot" taxa (with disjunct distribution in few widely separated basin). Endemism is low at the generic but very significant (ca. 50%) at the species level. Linked with regional facies change and eustatic influences, there were alternating episodes with low­ diverse, relatively highly endemic or with species-rich and rather cosmopolitan faunas. Faunal similarities both in the Frasnian and Famennian were closest with Germany, slightly less with North Africa, SW England, the Ardennes, and the Montagne Noire. Frasnian faunal links with the Timan and eastern North America were severed after the Frasnian-Famennian boundary whilst relationships with the Urals and Poland became closer. Faunal similarities were clearly more dependent on regional facies developments of plates than on their spatial distance. The regional diversity curve reflects both global extinctions and radiations as well as effects of Canning Basin structural evolution. Three major extinctions, the Bugle Gap, Lower Kellwasser and Upper Kellwasser Events occurred in the Frasnian. Not a single ammonoid
    [Show full text]
  • Package 'Paleotree'
    Package ‘paleotree’ December 12, 2019 Type Package Version 3.3.25 Title Paleontological and Phylogenetic Analyses of Evolution Date 2019-12-12 Author David W. Bapst, Peter J. Wagner Depends R (>= 3.0.0), ape (>= 4.1) Imports phangorn (>= 2.0.0), stats, phytools (>= 0.6-00), jsonlite, graphics, grDevices, methods, png, RCurl, utils Suggests spelling, testthat Language en-US ByteCompile TRUE Encoding UTF-8 Maintainer David W. Bapst <[email protected]> BugReports https://github.com/dwbapst/paleotree/issues Description Provides tools for transforming, a posteriori time-scaling, and modifying phylogenies containing extinct (i.e. fossil) lineages. In particular, most users are interested in the functions timePaleoPhy, bin_timePaleoPhy, cal3TimePaleoPhy and bin_cal3TimePaleoPhy, which date cladograms of fossil taxa using stratigraphic data. This package also contains a large number of likelihood functions for estimating sampling and diversification rates from different types of data available from the fossil record (e.g. range data, occurrence data, etc). paleotree users can also simulate diversification and sampling in the fossil record using the function simFossilRecord, which is a detailed simulator for branching birth-death-sampling processes composed of discrete taxonomic units arranged in ancestor-descendant relationships. Users can use simFossilRecord to simulate diversification in incompletely sampled fossil records, under various models of morphological differentiation (i.e. the various patterns by which morphotaxa originate from one another), and with time-dependent, longevity-dependent and/or diversity-dependent rates of diversification, extinction and sampling. Additional functions allow users to translate simulated ancestor-descendant data from simFossilRecord into standard time-scaled phylogenies or unscaled cladograms that reflect the relationships among taxon units.
    [Show full text]
  • Armatites Kaufmanni N. Sp., the First Late Devonian Goniatite with Ventral Spines
    See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/260719399 Armatites kaufmanni n. sp., the first Late Devonian goniatite with ventral spines Article in Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen · March 2014 DOI: 10.1127/0077-7749/2014/0393 CITATIONS READS 2 119 1 author: Dieter Korn Museum für Naturkunde - Leibniz I… 297 PUBLICATIONS 2,712 CITATIONS SEE PROFILE All in-text references underlined in blue are linked to publications on ResearchGate, Available from: Dieter Korn letting you access and read them immediately. Retrieved on: 02 September 2016 N. Jb. Geol. Paläont. Abh. 271/3 (2014), 349–352 Article E Stuttgart, March 2014 Armatites kaufmanni n. sp., the first Late Devonian goniatite with ventral spines Dieter Korn With 2 figures Abstract: Armatites kaufmanni n. sp. is described from an early Famennian (Late Devonian) oc- currence of Braunau (Ense near Bad Wildungen) in the Rhenish Mountains Germany). It is the first goniatitic ammonoid with ventral shell extensions in the form of two rows of prominent, laterally flattened spines. Key words: Ammonoidea, Famennian, Rhenish Mountains, shell sculpture. 1. Introduction shell extensions. They occur often in species, in which a flattened or concave venter is separated from the flanks The development of shell extensions in the form of by an angular margin. The oldest of these is the late longitudinal skids, ventrolateral or ventral parabolic Early Devonian genus Gyroceratites v. Meyer, 1831, ribs and spines is a repeated process in the evolution of but they become more important in the Famennian. the Devonian and earliest Carboniferous ammonoids.
    [Show full text]