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KKNOWN COLEOPTERA FROM FRANKLIN Co., MISSOURI Reputedly the largest order of insects, this group includes plant feeders such as leaf beetles, long-horned beetles, and many scarabs; scavengers and coprophages such hide, carrion, and dung beetles; and effective predator among the ground and water beetles. Some beetles are of economic importance as pests of crop and ornamental plants, and stored grain pests. A well-known invasive species is the Japanese beetle Popilia japonica which has spread from the east toward Missouri in recent decades. Most of the species below occur or are likely to at Shaw Nature Reserve. Many on this list are identified to genus level only, a function of both the hugeness of the group, and of much taxonomic work remaining to be done. Records compiled from bugguide.net, especially the exhaustive records of Mr. Jack Foreman, plus those by J. Trager and Jon Rapp. Photos here are all by James C. Trager, taken at SNR. They represent the species whose name is directly above the image. Suborder Archostemata Harpalus pensylvanicus Harpalus spp. Family Cupedidae Lebia analis Tenomerga cinerea Lebia atriventris Lebia solea Suborder Adephaga Lebia viridipennis Lebia viridis Family Carabidae Lebia vittata Acupalpus testaceus Paraclivina bipustulata Agonum sp. Platynus decentis Amara sp. Poecilus chalcites Amphasia sericea Poecilus sp. Anisodactylus caenus Scaphinotus elevatus Anisodactylus harrisii Aspidoglossa subangulata Badister neopulchellus Bembidion sp. Bradycellus festinans Bradycellus tantillus Calosoma scrutator Chlaenius laticollis Chlaenius sp. Chrysochus auratus Cicindela purpurea Stenolophus dissimilis Cicindela repanda Stenolophus lecontei Cicindela sexguttata Stenolophus lineola Cicindelidia rufiventris Stenolophus ochropezus Colliuris pensylvanica Tachys proximus Dicaelus purpuratus Telephanus atricapillus Dicaelus sp. Thalpius dorsalis Galerita sp. Family Dytiscidae Acmaeodora pulchella Acilius sp. Acmaeodora tubulus Family Gyrinidae Dineutus ciliatus Dineutus discolor Family Haliplidae Haliplus triopsis Peltodytes muticus Chrysobothris cf. femorata Suborder Polyphaga Family Callirhipidae Family Aderidae Zenoa picea Aderus sp. Elonus hesperus Family Cantharidae Emelinus melsheimeri Atalantycha bilineata Zonantes hubbardi Atalantycha neglecta Chauliognathus marginatus Family Anthicidae Chauliognathus pensylvanicus Anthicus lutulentus Notoxus murinipennis Omonadus floralis Stricticollis tobias Tomoderus sp. Family Anthribidae Euparius marmoreus Piesocorynus mixtus Polemius repandus Toxonotus cornutus Podabrus flavicollis Podabrus poricollis Family Attelabidae Podabrus rugulosus Eugnamptus angustatus Podabrus tricostatus Merhynchites bicolor Rhagonycha mollis Silis sp. Family Bostrichidae Trypherus sp. Lichenophanes bicornis Tytthonyx erythrocephala Trogoxylon parallelipipedum Xylobiops basilaris Family Cerambycidae Acanthocinus sp. Family Brentidae Anelaphus parallelus Arrhenodes minutus Ataxia crypta Trichapion sayi Batyle suturalis Desmocerus palliates Family Buprestidae Dryobius sexnotatus Agrilaxia flavimana Eburia quadrigeminata Agrilus sp. Ecyrus dasycerus Elytrimitatrix undata Capraita sexmaculata Capraita thyamoides Charidotella sexpunctata Chelymorpha cassidea Chrysomela knabi Enaphalodes atomarius Chrysomela scripta Euderces picipes Colaspis brunnea Eupogonius pauper Colaspis sp. Goes debilis Cryptocephalus leucomelas Graphisurus despectus Cryptocephalus mutabilis Graphisurus fasciatus Cryptocephalus venustus Knullian cincta Cryptocephalus sp. Leptostylus transversus Dectes texanus Necydalis mellita Deloyala guttata Neoclytus acuminatus Derospidea brevicollis Obrium maculatum Diabrotica barberi Odontotaenius disjunctus Diabrotica cristata Oeme rigida Diabrotica undecimpunctata Plinthocoelium suaveolens Disonycha admirabila Prionus imbricornis Disonycha glabrata Psyrassa unicolor Donacia sp. Saperda discoidea Gibbobruchus mimus Saperda tridentata Glyptoscelis sp. Smodicum cucujiforme Kuschelina cf. fimbriata Stenocorus cinnamopterus Labidomera clivicollis Sternidius alpha Luperaltica nigripalpis Sternidius punctatus Neochlamisus sp. Sternidius sp. Odontota scapularis Strangalia sexnotata Pachybrachis sp. Tetraopes tetrophthalmus Parchicola tibialis Typocerus cf. confluens Paria cf. fragariae Xylotrechus colonus Paria sp. Phyllecthris gentilis Family Ceratophytidae Phyllotreta bipustulata Ceratphytum pulsator Rhabdopterus sp. Scolytus sp. Xanthonia sp. Family Chrysomelidae Amblycerus robiniae Aulacothorax copallina Altica chalybea Capraita circumdata Family Cleridae Xyleborus celsus Cymatodera undulata Enoclerus sp. Family Dermestidae Neorthopleura thoracica Anthrenus verbasci Placopterus thoracicus Dermestes lardarius Family Coccinelidae Family Dryopidae Anatis labiculata Helichus lithophilus Coleomegilla maculata Coccinella septempunctata Family Elateridae Cycloneda munda Aeolus mellilus Harmonia axyridis Aeolus sp. Alaus oculatus Family Curculionidae Conoderus bellus Arrhenodes minutus Conderus lividus Chalcodermus sp. Conoderus vespertinus Conotrachelas anaglypticus Glyphonyx sp. Contrachelas buchanani Megapenthes rufilabris Conotrachelas elegans Melanotus sp. Contrachelas fisssunguis Paradonus pectoralis Conotrachelas seniculus Curculio sp. Family Elmidae Stenelmis sp. Family Erotylidae Ischyrus quadripunctatus Languria bicolor Languria trifasciata Cyclorhipidion bodoanum Cyrtepistomus castaneus Eubulus obliquefasciatus Haplorhynchites aeneus Hormops abducens Megalodacne fasciata Lignyodes bischoffi Pseudischyrus extricates Lignyodes fraxini Toramus pulchellus Lignyodes helvolus Listronotus appendiculatus Family Eucnemidae Listronotus humilis Adelothyreus sp. Listronotus sp. Hylochares nigricornis Monarthrum fasciatum Odontocorynus umbellae Family Hydrophilidae Pheloconus cribricollis Enochrus blatchleyi Pheloconus hispidus Enochrus sp. Phyrdenus divergens Hydrochara sp. Rhyssomatus lineaticollis Tropisternus collaris Smicronyx spp. Xanthonia sp. Family Laemophloeidae Laemophloeus biguttatus Family Nitidulidae Cryptarcha ampla Family Lampyridae Epuraea aestiva Lucidota atra Glischrochilus fasciatus Photinus pyralis Lobiopa undulata Photuris sp. Omosita nearctica Pleotomus davisii Pyractomena angulata Family Oedomeridae Oxacis trimaculata Family Latridiidae Corticaria sp. Family Phalacridae Melanopthalma sp. Stilbus sp. Family Leiodidae Family Phengodidae Platypsyllus castoris Phengodes sp. Family Lucanidae Family Ptilodactylidae Lucanus capreolus Ptilodactyla carinata Family Lycidae Family Ptinidae Calopteron reticulatum Ptinus bimaculatus Calopteron terminale Trichodesma gibbosa Plateros sp. Tricorynus sp. Family Melandryidae Family Pyrochroidae Micronotus sericans Dendroides canadensis Neopyrochroa femoralis Family Meloidae Neopyrochroa flabellata Epicauta atrata Pedilus terminalis Epicauta cf. funebris Epicauta pennsylvanica Family Rhipiphoridae Epicauta vittata Macrosiagon limbata Meloe campanicollis Nemognatha nemorensis Nemognatha sp. Family Mordellidae Hoshihananomia octopunctata Mordella insulata Mordella marginata Family Scarabaeidae Necrodes surinamensis Aphodius pseudolividus Nicrophorus orbicollis Aphodius sp. Nicrophorus pustulatus Ataenius imbricatus Ataenius sp. Atinus monilicornis Callistethus marginatus Canthon sp. Cyclocephala lurida Dichotomius carolinus Diplotaxis sp. Dynastes tityus Eucanthus impressus Euphoria sepulcralis Family Staphylinidae Germarotes sp. Arthmius sp. Macrodactylus subspinosus Bledius sp. Odonteus sp. Conotelus sp. Pelidnota punctata Coproporus laevis Phanaeus vindex Gyrophaenina sp. Phyllophaga quercus Homaeotarsus sp. Phyllophaga spp. Lobrathium sp. Mycetoporus sp. Paederus sp. Philonthus rufulus Pinophilus sp. Platydracus sp. Popilia japonica Tomarus gibbosus Rugilus angularis Trichiopeltastes delta Tachyporus sp. Trichiotinus piger Trichiotinus viridans Family Tenebrionidae Family Scirtidae Alobates sp. Contacyphon sp. Bolitotherus cornutus Scirtes tibialis Cynaeus angustus Scirtes sp. Diaperis maculata Lobopoda punctulata Neatus tenebrioides Family Silvanidae Platydema erythrocerum Ahasverus advena Platydema ruficorne Statira sp. Tenebrio molitor Family Scraptiidae Allopoda lutea Family Silphidae Family Tetratomidae Penthe pimelia Family Trogidae Omorgus sp. Family Trogossitidae Temebroides sp. Family Zopheridae Colydium lineola .
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  • Conspicuousness, Phylogenetic Structure, and Origins of Müllerian

    Conspicuousness, Phylogenetic Structure, and Origins of Müllerian

    www.nature.com/scientificreports OPEN Conspicuousness, phylogenetic structure, and origins of Müllerian mimicry in 4000 lycid beetles from all zoogeographic regions Michal Motyka1, Dominik Kusy1, Michal Masek1, Matej Bocek1, Yun Li1, R. Bilkova1, Josef Kapitán2, Takashi Yagi3 & Ladislav Bocak1* Biologists have reported on the chemical defences and the phenetic similarity of net-winged beetles (Coleoptera: Lycidae) and their co-mimics. Nevertheless, our knowledge has remained fragmental, and the evolution of mimetic patterns has not been studied in the phylogenetic context. We illustrate the general appearance of ~ 600 lycid species and ~ 200 co-mimics and their distribution. Further, we assemble the phylogeny using the transcriptomic backbone and ~ 570 species. Using phylogenetic information, we closely scrutinise the relationships among aposematically coloured species, the worldwide diversity, and the distribution of aposematic patterns. The emitted visual signals difer in conspicuousness. The uniform coloured dorsum is ancestral and was followed by the evolution of bicoloured forms. The mottled patterns, i.e. fasciate, striate, punctate, and reticulate, originated later in the course of evolution. The highest number of sympatrically occurring patterns was recovered in New Guinea and the Andean mountain ecosystems (the areas of the highest abundance), and in continental South East Asia (an area of moderate abundance but high in phylogenetic diversity). Consequently, a large number of co-existing aposematic patterns in a single region and/or locality is the rule, in contrast with the theoretical prediction, and predators do not face a simple model-like choice but cope with complex mimetic communities. Lycids display an ancestral aposematic signal even though they sympatrically occur with diferently coloured unproftable relatives.