Mesozoic Decapod Diversity with an Emphasis on the Early Cretaceous (Albian) of Spain
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MESOZOIC DECAPOD DIVERSITY WITH AN EMPHASIS ON THE EARLY CRETACEOUS (ALBIAN) OF SPAIN A dissertation submitted to Kent State University in partial fulfillment of the requirements for the degree of Doctor of Philosophy by Adiël A. Klompmaker August, 2012 Dissertation written by Adiël A. Klompmaker M.S., Utrecht University, 2006 Ph.D., Kent State University, 2012 Approved by ________________________________, Advisor Rodney M. Feldmann ________________________________, Committee Member Carrie E. Schweitzer ________________________________, Committee Member Joseph D. Ortiz ________________________________, Committee Member Ferenc A. de Szalay Accepted by _________________________________, Chair, Department of Geology Daniel K. Holm _________________________________, Dean, College of Arts and Sciences Timothy Moerland ii TABLE OF CONTENTS LIST OF FIGURES ...................................................................................................................... xii LIST OF TABLES ........................................................................................................................xvi LIST OF APPENDICES ............................................................................................................xviii ACKNOWLEDGEMENTS ..........................................................................................................xix 1. SUMMARY. ................................................................................................................................1 2. INTRODUCTION .......................................................................................................................3 3. SYSTEMATIC PALEONTOLOGY .........................................................................................13 3.1. Revision of the family Gastrodoridae (Crustacea, Decapoda), with description of the first species from the Cretaceous.......................................................................................13 Abstract.................................................................................................................14 Introduction...........................................................................................................14 Stratigraphy and Locality......................................................................................15 Systematic Paleontology.......................................................................................16 Superfamily Gastrodoroidea Van Bakel et al., 2008 (new status)...........16 Family Gastrodoridae Van Bakel et al., 2008............................23 iii Genus Gastrodorus von Meyer, 1864...........................23 G. cretahispanicus new species........................24 Genus Eogastrodorus Van Bakel et al., 2008...............27 Acknowlegdments................................................................................................28 References.............................................................................................................29 Figures..................................................................................................................38 Tables....................................................................................................................42 3.2. The Cretaceous crab Rathbunopon: revision, a new species and new localities........44 Abstract.................................................................................................................44 Introduction...........................................................................................................45 Locality and Stratigraphy......................................................................................45 Systematic Paleontology.......................................................................................46 Superfamily Homolodromioidea Alcock, 1900.......................................46 Family Prosopidae von Meyer, 1860..........................................46 Genus Rathbunopon Stenzel, 1945................................47 R. obesum (Van Straelen, 1944).......................50 R. schrattenkalkensis n. sp...............................55 Evolution and Paleoenvironment..........................................................................59 Acknowlegdements...............................................................................................60 References.............................................................................................................61 Figures..................................................................................................................67 Table.....................................................................................................................73 3.3. Etyid crabs (Crustacea, Decapoda) from mid-Cretaceous reefal strata of Navarra, northern Spain....................................................................................................................74 iv Abstract.................................................................................................................74 Introduction...........................................................................................................75 Systematic Palaeontology.....................................................................................76 Superfamily Etyoidea Guinot and Tavares, 2001....................................76 Family Etyidae Guinot and Tavares, 2001.................................76 Genus Xanthosia Bell, 1863 sensu lato.........................76 X. koskobiloensis sp. nov..................................79 X. cf. X. koskobiloensis sp. nov........................82 Genus Etyxanthosia Fraaije et al., 2008........................85 E. fossa (Wright and Collins, 1972).................85 Genus Caloxanthus A. Milne-Edwards, 1864...............87 C. paraornatus sp. nov.....................................87 Discussion.............................................................................................................90 Acknowlegdements...............................................................................................94 References.............................................................................................................95 Figures................................................................................................................102 Tables..................................................................................................................108 3.4. New European localities for coral-associated, Cretaceous decapod crustaceans.....110 Abstract...............................................................................................................110 Introduction.........................................................................................................111 Localities and Stratigraphy.................................................................................112 Systematics.........................................................................................................113 Superfamily Callianassoidea Dana, 1852..............................................113 Family ?Upogebiidae indet.......................................................114 v Family Callianassidae indet. 1..................................................114 Family Callianassidae indet. 2..................................................116 Family ?Callianassidae indet. 3................................................117 Discussion...........................................................................................................118 Acknowlegdements.............................................................................................119 References...........................................................................................................120 Figures................................................................................................................126 3.5. Peak diversity of Cretaceous galatheoids (Crustacea, Decapoda) from northern Spain................................................................................................................................130 Abstract...............................................................................................................130 Introduction.........................................................................................................131 Geologic setting and previous research..............................................................131 Systematic Palaeontology...................................................................................132 Superfamily Galatheoidea Samouelle, 1819..........................................133 Family Galatheidae Samouelle, 1819.......................................133 Genus Eomunidopsis Vía Boada, 1981.......................133 E. aldoirarensis sp. nov..................................134 E. navarrensis (Van Straelen, 1940)..............139 E. orobensis (Ruiz de Gaona, 1943)...............144 Genus Paragalathea Patrulius, 1959...........................148 P. multisquamata Vía Boada, 1981................149 P. ruizi Van Straelen, 1940............................153 P. straeleni Ruiz de Gaona, 1943...................156 Genus Hispanigalathea gen. nov................................159 vi H. pseudolaevis sp. nov..................................162 H. tuberosa sp. nov.........................................164 Family Munidopsidae Ortmann, 1898......................................166 Genus