ISSN 1211-8788 Acta Musei Moraviae, Scientiae biologicae (Brno) 89: 25–51, 2004
The water beetles (Coleoptera: Hydradephaga) of Afghanistan, with particular emphasis on the collection of the Moravian Museum Brno, Czech Republic
JIØÍ HÁJEK 1 & HANS FERY 2 1Dept. of Entomology, National Museum, Kunratice 1, CZ-148 00 Praha 4, Czech Republic; e-mail: [email protected] 2Räuschstraße 73, D-13509 Berlin, Germany; e-mail: [email protected]
HÁJEK J. & FERY H. 2004: The water beetles (Coleoptera: Hydradephaga) of Afghanistan, with emphasis on the collection of the Moravian Museum Brno, Czech Republic. Acta Musei Moraviae, Scientiae biologicae (Brno) 89: 25–51. – We present all known records of Hydradephaga (Coleoptera: Gyrinidae, Haliplidae, Dytiscidae) from Afghanistan, including recently studied material. The latter was chiefly collected during five Czech scientific expeditions to Afghanistan in 1960’s and is now deposited in the Moravian Museum Brno (Czech Republic). Fourteen species are recorded from Afghanistan for the first time, and eleven first records from other countries are provided. Altogether four species of the family Gyrinidae, four species of the family Haliplidae, of which one is in doubt, and 52 species of the family Dytiscidae, of which two are in doubt, are now known from Afghanistan. Only four of these taxa are known to be endemic to this country. Members of the families Hygrobiidae and Noteridae have never been recorded. Key words. Coleoptera, Gyrinidae, Haliplidae, Dytiscidae, Afghanistan, distribution, first records
Introduction Although several biological expeditions have ventured to Afghanistan in the past, the country remains one of the least explored areas in the Palaearctic region. To our knowledge, water beetles have been collected during at least sixteen expeditions; eleven of them were led by other than Czech scientists: J. Klapperich (Germany) in 1952–1953 (OCHS 1955, GUÉORGUIEV 1963); H.G. Amsel (Germany) in 1956 (GUIGNOT 1961); H.G. Amsel & G. Ebert (Germany) in 1966 (AMSEL 1967); K. Lindberg (Sweden) in 1957–1958 (GUIGNOT 1960, GUÉORGUIEV 1963), 1959–1960 (GUÉORGUIEV 1963, 1975) and 1962 (GUÉORGUIEV 1975); R. Yosu (Japan) in 1960 (SATÔ 1966); F. Kasy, A. & E. Vartian (Austria) in 1963 and 1965 (GUÉORGUIEV 1967); Y. Arita (Japan) in 1970 (SATÔ 1972); G. Meurgues & G. Ledoux (France) in 1974 (LEGROS 1977). The five expeditions to Afghanistan organised by Czech biologists took place between 1963 and 1967. The rich entomological material from these expeditions is deposited in the collection of the Moravské zemské muzeum Brno (Moravian Museum Brno) and has been previously studied by several specialists. This material also includes water beetles. The aim of the present paper is to summarise all data on water beetles from Afghanistan (Gyrinidae, Haliplidae, Dytiscidae) based on the literature, the material deposited in Brno, and a few additional data coming from other sources.
25 J. HÁJEK & H. FERY
Material and methods All records from Afghanistan found in the literature are presented for each species. Studied material data are ordered as follows: number of specimens, label texts, and depository. Our remarks are given in square brackets. Data on material stemming from four Czech expeditions are followed by abbreviations in square brackets, which are taken from JAKEŠ & POVOLNÝ (1967): “J” stands for “O. Jakeš expedition 1963–64”; “Š” stands for “J. Šimek expedition 1965–66”; “PT” stands for “D. Povolný & F. Tenora expedition 1966”; “P” stands for “D. Povolný, J. Gaisler, Z. Šebek & F. Tenora expedition 1967”. Material from the remaining expedition (“D. Povolný expedition 1965”) is relatively limited and comes only from the environs of Jalalabad; it is not marked by any abbreviation. Recent names of the provinces and localities are unified according to The Times Comprehensive Atlas of the World, 10th edition (London, 2001). Specifications of localities are taken from standard sources such as Andrees Handatlas (Bielefeld, Leipzig, 1890), Gazetteer of Afghanistan (Washington, D.C., 1992), Stiehlers Handatlas (Gotha, 1928/30), The World Atlas (Moscow, 1967), Knauers Grosser Weltatlas (Munich, 1999), and Microsoft Encarta World Atlas 2000. We supplemented these sources with data from DUPREE (1971), ADAMEC (1972, 1985), and SCHÜTZE & KLEINFELD (1997). We adopt the style of The Times Atlas for the presentation of the coordinates of many localities. The Appendix contains names that we assumed would not to be commonly known. These names are marked with an asterisk in the text. Short characteristics of the distribution are cited for the species recorded from Afghanistan or other countries for the first time. We refer to the Catalogue of Palaearctic Coleoptera (MAZZOLDI 2003, NILSSON 2003a, b, VONDEL 2003) for information about a more general distribution of each species.
Abbreviations The following acronyms are used for denoting the depositories: CGW ...... coll. G. Wewalka, Wien, Austria CHF ...... coll. H. Fery, Berlin, Germany, property of the NMW COK ...... coll. O. N. Kabakov, St. Petersburg, Russia JHCP ...... coll. J. Hájek, deposited in the Národní Muzeum, Praha, Czech Republic MRTO ...... Museo Regionale di Scienze Naturali, Torino, Italy (M. Daccordi) MZMB ...... Moravské zemské muzeum, Brno, Czech Republic (V. Kubáò) NHMB ...... Naturhistorisches Museum Basel, Switzerland (M. Brancucci) NMPC ...... Národní muzeum, Praha, Czech Republic (J. Hájek) NMW ...... Naturhistorisches Museum Wien, Austria (M. A. Jäch) SMNK . . . . Staatliches Museum für Naturkunde Karlsruhe, Germany (G. Ebert, A. Riedel) ZSM . . . . . Zoologische Staatssammlung, München, Germany (M. Baehr, M. Kühbandner)
26 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan
List of recorded species
Family Gyrinidae Subfamily Gyrininae
Aulonogyrus (s.str.) concinnus (Klug, 1834)
References. OCHS (1955: 47): “Kandahar-Kuna* 950 m; 2.II., 7.II. und 7.III.1953”, “Tangi-Saitan* bei Kabul 1750 m; 27.V.1952”, and “Kabul 1740 m; 20.VII.1952”. Material studied. Afghanistan: 96 exs., “N. Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 18.9.–21.9.1964, 470 m” [J73, irrigation canal] (MZMB). Armenia: 1 ex., “Armenia, 13–14.7.2003, Ararat prov., 1300–2100 m, Khosrov nature reserv., 39°59’N 44°53’E, V Zieris lgt.”, J. Hájek det. 2003, coll. V. Zieris (Pardubice, Czech Republic). Turkmenistan: 1 ex., “Turkmenia – Chardzhou [= Chärjew], Dargan Ata [ca. 40.50N 61.57E], 2.–4. 6. [19]92, Z. Kleteèka lgt.”, J. Hájek det. 1999 (JHCP). Notes. A South Palaearctic species. It occurs in southern and Central Europe, North Africa, Turkey, Syria, Iraq, Iran, Afghanistan, China and Mongolia (MAZZOLDI 1995). First records from Armenia and Turkmenistan.
Gyrinus (s.str.) distinctus Aubé, 1838
References. OCHS (1955: 46): “Kuschke, ex. Coll. Hauser”; exact locality unknown. OCHS (1955: 46) assumed that the single specimen was collected in northern Afghanistan near the Turkmenistan border.
Subfamily Orectochilinae
Orectochilus (s.str.) afghanus Ochs, 1955
References. OCHS (1955: 49): “Nuristan*, Bashgultal* 1100–1200 m; 10.–17.IV.1953” (type locality), and “Pagman-Gebirge*, ca. 30 km westlich von Kabul aus Höhen von 2400 m, 6.VII.1952, resp., 2300 m, 30.V.1952”.
Orectochilus (s.str.) nuristanicus nuristanicus Ochs, 1955
References. OCHS (1955: 48): “Nuristan*, Kutiau*, 1400 m, 10.V.1953” (type locality).
Family Haliplidae
Haliplus (Haliplinus) fluviatilis Aubé, 1836
References. GUIGNOT (1960: 320): “Sirouas* (entre Moqor* et Qalat*), au bord d’un ruisseau, sous les pierres, 1650 m, 8 avril 1958”, and “Chevaki* (près Kaboul*), steppe, sous les pierres, 1800 m, 4 oct. 1957”. GUÉORGUIEV (1963: 216): “Balkh* (Turkestan afghan), marécage, alt. 385 m, 1 ♂ à 19.V.1959 (K. Lindberg)”.
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 27 J. HÁJEK & H. FERY
Haliplus (Liaphlus) maculatus Motschulsky, 1860
References. VONDEL (1991: 119): “1 ♀, W. Afghanistan, Hari Rud*, Herat*, 20.vii.1949, Danish Central Asian Expedn, St. no. 4, Ex stomach of Lohivanellus indicus, Haliplus fulvus, J. Balfour-Brown det. 1964 (BMNH) [= British Museum of Natural History, London]”.
[? Haliplus (Liaphlus) variegatus Sturm, 1834]
References. GUÉORGUIEV (1963: 216) (as Haliplus variegatus pallidior J. Müller, 1900): “Kandahar* (31.36N 65.41E), alt. 950 m, 1 ♂ et 5 ♀ à 13.II.1953 (J. Klapperich)”. Notes. VONDEL (1991: 133) called the occurrence of this species in Afghanistan into doubt. According to him, the specimens probably belong to another species such as H. abbreviatus Wehncke, 1880 or H. villiersi Legros, 1972.
Peltodytes caesus (Duftschmid, 1805)
References. GUÉORGUIEV (1963: 215): “environs de Kaboul*, alt. 1740 m, 1 exempl. à 16.IX.1952 (J. Klapperich)”. Material studied. Afghanistan: 1 ex., “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Murièaq*, 16.6.1964, 400 m” [J28, at light] (MZMB).
Family Dytiscidae Subfamily Agabinae
Agabus (Gaurodytes) biguttatus (Olivier, 1795)
References. GUIGNOT (1960: 320, 321) (as Agabus (Dichonectes) biguttatus, A. (D.) nitidus (FABRICIUS, 1801), and Agabus (D.) nitidus ab. nigricollis ZOUBKOFF, 1833): “Siah Guerd* (vallée du Ghourband*), rivière et au bords, 1920 m, 28 juill. 1957”, “Haouz-Mahiyan* (75 km à l’ouest de Kaboul*), marécage, 2470 m, 4 avril 1958”, “Chevaki* (près Kaboul*), steppe, sous les pierres, 1800 m, 4 oct. 1957”, “Kotal-Lachva* (entre Qal’éh Chahrak* et Chibarg*, Hézaradjat*), en bas du col, dans torrent et au bords, 2500 m, 22 août 1957”, “Tang- Djebel Baz* ou Tang-Kharzar* (entre Khvadjah Tchicht* et Tang-Azao*, Hzaradjat*), dans torrent et au bord, 1850 m, 24 août 1957”, “Sirouas* (entre Moqor* et Qalat*), au bord d’un ruisseau, sous les pierres, 1650 m, 8 avril 1958”, and “Badgah* (près Hari Roud*, à 30 km à l’ouest de Daoulet Yar*, Hézaradjat*), bassin d’une source, 2350 m, 16 août 1957”. GUÉORGUIEV (1963: 219, 200) (as A. (D.) biguttatus, A. (D.) biguttatus ab. balearicus GUIGNOT, 1953, and A. (D.) nitidus): “Pir-Khil* (environs 55 km NE d’Orozgan*), bassin d’une source, alt. 2350 m, ... à 13.VI.1960 (K. Lindberg)”, “Masdjed-Tchoubi*, Chileh Hamman* (en bas du col de Sabzzak*), bassin d’une source, alt. 2190 m, ... (K. Lindberg)”, “Qal’eh-Darreh Zang* (région de Beltchiragh*), terrain humide au pied de rochers, sous pierres, alt. 1520 m, ... à 26.V.1956 (K. Lindberg)”, and “Mazanah* (vallée du Ghourband*, NO de Kaboul*), fossé, alt. 2080 m, ... à 13.V.1959 (K. Lindberg)”, and “Col de Sabzzak*, Paropamise* à 65 km au NE de Hérat*, mare de torrent, ... 4.VI.1959 (K. Lindberg)”. SATÔ (1966: 255) (as A. (D.) biguttatus): “1 ♀, Kabul, Afghanistan, June 17, 1960”. SATÔ (1972: 49): “1 ex., Band-i-Amir*, July 29, 1970, Y. Arita leg.” (as A. (D.) biguttatus), and “1 ex., Parakh*, Panjshir* (alt.2,700 m), July 6, 1970, Y. Arita leg.” (as A. (D.) nitidus).
28 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan
Agabus (Gaurodytes) bipustulatus (Linnaeus, 1767)
References. GUIGNOT (1960: 321): “Rah-Kol* (à l’ouest de Dival Kol*, Hézaradjat*), étang Dand Karkatak*, 2950 m, 13 août 1957”.
Agabus (Gaurodytes) conspersus (Marsham, 1802)
References. GUIGNOT (1960: 321): “Andkoï*, mare dans une steppe de sable, 325 m, 17 oct. 1957”, and “Kouh-Bibtchéh Baran* (au nord de Farah*), mare de torrent, 900 m, 27 avril 1958”. GUÉORGUIEV (1963: 220): “Col de Sabzzak*, Paropamise* à 65 km au NE de Hérat*, mare de torrent, 2090 m, 1 ♂ à 4.VI.1959 (K. Lindberg)”. SATÔ (1966: 255): “1 ♂, 1 ♀, Borak*, Afghanistan, July 12, 1960”, and “1 ♂, 1 ♀, Ishkashim*, Afghanistan, Sept. 2, 1960”.
Agabus (Gaurodytes) dichrous Sharp, 1878
References. GUÉORGUIEV (1963: 221): “Tchakaran* (entre Baharaknet* et Soufian*, Badakhchan*), bassin d’une source, alt. 1740 m, 1 ♂ à 26.VII.1960 (K. Lindberg)”, and “Zebak* (Badakhchan*), marécage, alt. 2550 m, 1 ♀ à 16.VII.1959 (K. Lindberg)”.
Agabus (Gaurodytes) glazunovi (Zaitzev, 1953)
References. GUÉORGUIEV (1975: 99): “Qal’eh-Darrech Zang*, 1500 m, (26.V.1959), 1 ♂ (K. Lindberg leg.)”, and “Pandjab* (Hezaradjat), Decht Ghoudjour*, 2680 m (4.VII.1962), 1 ♂ (K. Lindberg leg.)”.
Agabus (Gaurodytes) solskii Jakovlev, 1897
References. GUIGNOT (1960: 321) (as Agabus (Gaurodytes) bipustulatus solskyi): “Soltan Khel* (au nord de Ghazni*), sous pierres, près d’un cours d’eau, 2230 m, 11 sept. 1957”. SHAVERDO (2002: 124): “Afghanistan, Behsud*, 2800 m, 10.7.1971, Leg. Kabakov O. N.”
Agabus (Gaurodytes) winkleri (Gschwendtner, 1923)
References. GUÉORGUIEV (1963: 219) (as Agabus (Dichonectes) pamiricus GUIGNOT, 1955) from: Afghanistan: “Bamvardar Aoudak* (Pandjchir*, 140 km NE de Kaboul*), petit lac, alt. 2740 m, 1 ♂ à 3.VII.1960 (K. Lindberg)”, “Ichkachim* (à l’entrée du Vakahn*), sous pierres, alt. 2500 m, 1 ♂ (K. Lindberg)”, “Yelgah*, lac Mohammed Zaher* (à l’est du Koul-Choghnan* = lac Chiva*), alt. 3150 m, 2 ♂ à 10.VIII.1960 (K. Lindberg)”, “Yaoulang*, bord occidental du Pamir de Chivah*, source, alt. 2800 m, 1 ♂ et 1 ♀ à 26.VII.1959 (K. Lindberg)”, and “Pamir de Chivah* (entre Yaoulang* et Doavi*), mares de rivières, alt. 2580 m, 29 ♂ et 5 ♀ (K. Lindberg)”. LEGROS (1977: 16) (as Anagabus [sic!] (Dichonectes) pamiricus): “Khot-Gaï* (Shahidan*), prov. Paktia*, vers 2500 m, 20–VIII–1974, 16 exemplaires (G. Meurgues et G. Ledoux)”.
Material studied. Afghanistan: 1 ♂, “Umg. [= environs] Kabul Afghanistan” (CHF). Notes. The latter male was keyed to Agabus basalis (GEBLER, 1830). A.N. Nilsson (pers. comm.), however, suggested that the specimen be included under A. winkleri. Any
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 29 J. HÁJEK & H. FERY identification should be regarded as provisional since the Agabus guttatus-group species – in particular those from Central Asia – are in urgent need of revision.
Platambus lindbergi Guéorguiev, 1963
References. GUÉORGUIEV (1963: 218): “Pagham* (près de Kaboul*), bord de torrent, alt. 2450 m, 1 ♂ à 15.VII.1960 (K. Lindberg)” (holotype). BRANCUCCI (1988: 202): “Kabul district, 1740 m, 6.VII.1952, J. Klapperich”.
Platambus sogdianus (Jakovlev, 1897)
References. SATÔ (1972: 49) (as Agabus (Anagabus) vatelloides Régimbart, 1899): “9 exs., Parakh*, Panjshir* (alt. 2,700 m), July 6, 1970, Y. Arita leg.” WEWALKA (1975: 156) (as Platambus holzschuhi Wewalka, 1975): “1 ♀, Afghanistan, östl. Bande-amir*, 15.7.1971, leg. C. Holzschuh”. LEGROS (1977: 14) (as Anagabus ledouxi Legros 1977): “Afghanistan: Bamyian*, bords de la rivière Kalu* (Prov. de Bamyian*), au pied de la Ville Rouge* 13–VIII–1974 (G. Meurges et G. Ledoux)”; data repeated in BRANCUCCI (1982: 120), who stated the synonymy of Anagabus ledouxi.
Material studied. Afghanistan: 1 ♂, “SO [= SE]-Afghanistan, Safed Koh*, S-Seite [= southern slope], Kotkai*, 2350 m, 14.–23.6.1966, G. Ebert leg.” (SMNK). 1 ♀, “O [= E]-Afghanistan, Salang-Paß*, N-Seite [= northern slope], (Khinjan*) 2100 m, 5.–11.7.1966, G. Ebert leg.” (SMNK). 1 ♀, “O [= E]-Afghanistan, Kabulschlucht*, 22 km östl. [= east of] Kabul, 1650 m, Amsel u. Ebert leg.”, on reverse “3.7.1966” (SMNK). Kyrgyzstan: 1 ♀, “Kyrgyzstan, Alaj G.K., Majdan [= Maydan in SW Kyrgyzstan, ca. 40.02N 72.04E] (right) gorge, 2270 m, 13.VII.1997, leg. Dolin (K-12/97)” (NMW). Notes. This species occurs in mountains of Central Asia (Uzbekistan, Tajikistan, Afghanistan), Pakistan and Nepal. First record from Kyrgyzstan.
Platynectes (Gueorguievtes) kashmiranus kashmiranus J. Balfour-Browne, 1944
References. GUÉORGUIEV (1972: 45) (as Platynectes kashmiranus): “Nuristan*, Bashgultal*, 1100 m, 11.IV.1953, 1 ♂ (J. Klapperich)”; data repeated in NILSSON (1998: 117). Notes. According to Š•ASTNÝ (2003: 230) this record needs verification.
Subfamily Colymbetinae
Colymbetes fuscus (Linnaeus, 1758)
References. GUIGNOT (1961: 233): “Kaboul*, 1800 m”. GUÉORGUIEV (1967: 474): “Khurd-Kabul*, SO von Kabul, 1900 m, 18.VI.1965, 1 ♂; 26.VI.1965, 1 ♀ et 3.VII.1965, 1 ♀ (Kasy & Vartian)”.
Colymbetes semenowi (Jakovlev, 1896)
References. GUÉORGUIEV (1963: 221): “Kaboul*, fossé dans un jardin public, alt. 1800 m, 1 ♀ à 20.VIII.1959 (K. Lindberg)”.
30 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan
Material studied. Afghanistan: 1 ♂, “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 11.6.–15.6.1964, 470 m” [J26, at light], J. Š•astný det. 2000 (MZMB). 1 ♀, “C Afghanistan, Prov. Kabul, A. Gayoun”, “Kabul, 5.7.1965”, J. Š•astný det. 2000 (MZMB). 1 ♂, “8.VII.1965. Afgh., Khurd-Kabul SO v. [= SE of] Kabul, 1900 m, Kasy & Vartian” (very immature specimen) (NMW). The following female was identified by J. Š•astný, with doubt: 1 ♀, “C Afghanistan, Dasht-I Nawar*, Ghazni* Prov.”, Colymbetes sp., J. Š•astný det. (MZMB). Notes. The male specimen collected by Kasy & Vartian and deposited in NMW appears to belong to a series of specimens of which GUÉORGUIEV (1967) studied one female and identified it as C. fuscus (see that species below). In our opinion, the specimen represents C. semenowi as it has more than 20 rather closely spaced stridulatory apparatus ridges on the hind margin of the second abdominal segment. However, we believe that C. semenowi needs to be revised and may only be a variant of C. fuscus.
Rhantus (s.str.) suturalis (MacLeay, 1825)
References. GUIGNOT (1960: 321) (as Rhantus pulverosus (STEPHENS, 1828)): “Sirouas-Salar* (entre Cheikhabad* et Soltan Khel*, route de Kaboul* à Ghazni*), dans mare et aux bords, 2230 m, 11 sept. 1957”. GUÉORGUIEV (1967: 474): “Khurd-Kabul*, SO von Kabul, 1900 m, 3.VII.1965, 1 ♂ (Kasy & Vartian)”, “40 km SW Kabul*, 2300 m, 22.VII.1965, 1 ♂ et 1 ♀ (Kasy & Vartian)”, and “Paghman*, 30 km NW Kabul, 2500 m, 27.VII.1965, 1 ♀ (Kasy & Vartian)”. Material studied. Afghanistan: 3 exs., “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 25.5.–10.6.1964, 470 m” [J21] (MZMB). 1 ex., same data but 8.6.1964 [J24, at light] (MZMB). 2 exs., same data but 11.6.–15.6.1964 [J26, at light] (MZMB). 1 ex., same data but 1.9.–25.9.1964 [J66, at light] (MZMB). 1 ex., same data but 12.11.1964 [J99] (MZMB). 3 exs., “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Mangan*, 28.6.1964, 640 m” [J35] (MZMB). 1 ♀, “SO [= SE]-Afghanistan, Safed Koh*, S-Seite [= southern slope], Kotkai*, 2350 m, 14.–23.6.1966, G. Ebert leg.” (SMNK).
Subfamily Dytiscinae Tribe Cybistrini
Cybister (s.str.) limbatus (Fabricius, 1775)
References. GUIGNOT (1960: 321): “Ali Cheng* (Laghman*), marécages près d’une source, 1130 m, 30. janv. 1958”.
Cybister (s.str.) tripunctatus lateralis (Fabricius, 1798)
References. GUIGNOT (1954: 201) (as Cybister tripunctatus (OLIVIER, 1795)) from “Afghanistan”, no exact localities. GUIGNOT (1960: 321) (as C. tripunctatus asiaticus SHARP, 1882): “Bokan* (entre Ghourmatch* et Bala Morghab*), mare, 460 m, 21 oct. 1957”. GUÉORGUIEV (1963: 221) (as C. tripunctatus gotschi HOCHHUTH, 1846): “Kaboul*, dans chambre d’habitation, alt. 1800 m, 1 ♂ et 1 ♀ à 22.VIII.1960 (K. Lindberg).” Material studied. Afghanistan: 1 ♂, “N Afghanistan, Prov. Kataghan*, coll. J. Šimek”, “6 km w Kunduz*, 24.5.1966, 400 m” [Š44] (MZMB).
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 31 J. HÁJEK & H. FERY
Cybister (Melanectes) sugillatus Erichson, 1834
Material studied. Afghanistan: 1 ♀, “E Afghanistan, Nengarhar* prov., Shewa*, 4.3.1966, L.D. Haws leg.” (MZMB). Notes. C. sugillatus is a predominantly Oriental species. It occurs in India, Sri Lanka and Indonesia. In the Palaearctic region it reaches Tibet, Nepal, Bhutan and China (WEWALKA 1975). First record from Afghanistan.
Cybister (Scaphinectes) lateralimarginalis torquatus (Steven, 1829)
References. GSCHWENDTNER (1938: 75): “Seistan*, Afghanistan” as C. lateralimarginalis var. hedeni ZAITZEV, 1908. GUIGNOT (1960: 321) (as C. lateralimarginalis (DEGEER, 1774)): “Kaboul*, Djabir Ansar*, étang d’eau saumàtre, 1800 m, 18 mars 1958”. GUIGNOT (1961: 233) (as C. lateralimarginalis): “Kaboul*, 1800 m”. GUÉORGUIEV (1963: 222) (as C. lateralimarginalis): “Kaboul*, fossé dans un jardin public, alt. 1800 m, 1 ♀ à 20.VIII.1959 (K. Lindberg). GUÉORGUIEV (1967: 477) (under the name C. lateralimarginalis): “Kabul, 1800 m, 8.VIII.1963, 1 ♂ et 1 ♀ (Kasy & Vartian)”, “10 km NW von Kabul, 26.VII.1965, 4 ♂♂ et 3 ♀♀ (Kasy & Vartian)”, and “Sumpfgebiet O von Kabul, 20.VII.1963, 1 ♂ et 1 ♀ (Kasy & Vartian)”. SATÔ (1966: 255) (as C. lateralimarginalis): “1 ♂, Kabul, Afghanistan, June 17, 1960”. Material studied. Afghanistan: 2 ♀♀, “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 20.3.–1.4.1964, 470 m” [J7] (MZMB). 1 ♂, “N Afghanistan, Prov. Mazar-i-Sharif*, Tashkurgan*, 3.11.1962, 800 m” (MZMB). 1 ♂, “C Afghanistan, Kabul prov., Kabul, 24.6.1965, Habibullah lgt.” (MZMB). 1 ♀, same data but 7.7.1965, Mahd Munir lgt. (MZMB). 1 ♂, “O [= E]-Afghanistan, Kabulschlucht*, 22 km östl. [= east of] Kabul, 1650 m, 5.–12.6.1966, Amsel u. Ebert leg.” (SMNK). Notes. We provisionally assign the Afghan records of Cybister lateralimarginalis to C. lateralimarginalis torquatus in accordance with the distributional data given in NILSSON (2003a, b). The subspecies was described from China. ZAITZEV (1953, 1972) compared it with the nominotypical form and stated that it has a more elongated body shape that is slightly broadened beyond midlength, more convex elytra, and a rectangular posterior angle of the metafemora. However, we are unable find any differences between Afghan and Central European specimens.
Tribe Dytiscini
Dytiscus persicus Wehncke, 1876
References. ROUGHLEY (1990: 470): “Narin*, Afghanistan”. Notes. Dytiscus persicus is closely related to D. marginalis Linnaeus, 1758 and D. pisanus Laporte, 1835. We studied the holotype and several other specimens of D. persicus from Iran and Turkey and came to the conclusion that the best diagnostic character for the species is the coloration of the abdominal segments. ROUGHLEY (1990: 427; fig. 22C) erroneously illustrated the metacoxal process with a concave inner margin. However, the margin is clearly convex (GSCHWENDTNER 1938: 46, FERY 2002: 29).
32 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan
Dytiscus tianshanicus Gschwendtner, 1923
References. ROUGHLEY (1990: 494) from “Afghanistan”, no additional data provided.
Tribe Eretini
Eretes griseus (Fabricius, 1781)
References. GUIGNOT (1961: 233) [as Eretes sticticus succinctus (Klug, 1834)]: “Polichomri*, 700 m”.
Material studied. Afghanistan: 4 ♂♂, “O [= E] Afghanistan, Prov. Nengrahar*, Povolný & Tenora”, “Tor- Khama*, 23.3.[19]66, 900 m” [PT41] (MZMB).
Notes. The genus Eretes was recently revised by MILLER (2002), who recognised two species in the central part of Asia: E. sticticus occurs in the Old World from the Middle East throughout Africa, reaching Cyprus in the north, and in the New World from Peru to southern USA. E. griseus occurs from Indonesia and northernmost Australia (Darwin) to Japan, the Philippines, Sichuan province of China and Vladivostok, Russia; it ranges further throughout southern Asia to Africa including Madagascar, and reaches southern Europe in the north (MILLER 2002). In Miller’s opinion, the material identified as E. sticticus succintus belongs to E. griseus.
Eretes sticticus (Linneaus, 1767)
Material studied. Afghanistan: 1 ♂, 1 ♀, “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 25.5.–31.5.1964, 470 m” [J20, at light] (MZMB). 1 ♂, same data, but 8.6.1964 [J24, at light] (MZMB). 4 ♂♂, “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Herat*, 24.7.–30.7.1964, 1000 m” [J56, tank, spring] (MZMB). Kazakhstan: 1 ♀, “Kazakhstan, Guriev [= Atyraü, ca. 47.06N 51.54E], 12.9.1987, S. Murzin leg.”, J. Hájek det. 1999 (JHCP). Tajikistan: 1 ♂, “SW Tadzikistan, Shartuz [ca. 180 km SWS Dushanbe, ca. 37.15N 68.08E], 15.6.1989, at light, I. Kováø leg.”, J. Hájek det. 1999 (JHCP). Notes. A cosmopolitan species (see also under E. griseus). However, primary data from Central Asia are missing or in need of revision. First records from Kazakhstan, Tajikistan and Afghanistan.
Tribe Hydaticini
[?Hydaticus (Guignotites) fabricii (MACLEAY, 1825)]
References. GUIGNOT (1954: 201): “Afghanistan”, no additional data provided. GUÉORGUIEV (1967: 474): “Nuristan*, Petschtal*, 10.–13.VII.1965, 1 ♂ (Kasy & Vartian)”. Notes. According to WEWALKA (1979) H. fabricii is replaced in Afghanistan by H. ricinus. We have not been able to study the specimens cited by GUÉORGUIEV (1967) and GUIGNOT (1954). We therefore follow the opinion of WEWALKA (1979), which has been also adopted by NILSSON (2003a, b).
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 33 J. HÁJEK & H. FERY
Hydaticus (Guignotites) grammicus Germar, 1830
Material studied. Afghanistan: 1 ♂, “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab, 6.8.–7.8.1964, 470 m” [J58] (MZMB). Turkey: 6 exs., “Turcia, 16.VI.2001, Edirne vil. [= prov.], Edirne env., flooded meadow, drain”, “Turkey 2001 Expedition, GPS 41°37’N, 26°37’E, 225 m, M. Fikáèek, J. Hájek & J. Straka leg.”, J. Hájek det. 2001 (JHCP). Uzbekistan: 1 ♀, “Uzbekistan, Zeravshan Mts., pr. Agalik [ca. 15 km SWS Samarkand, ca. 39.31N 66.34E], 19.5.1990, 1300 m, pools near the river, J. Strejèek lgt.”, J. Strejèek det. 1992 (JHCP). Notes. A widespread Palaearctic species. It occurs from southern Europe throughout Asia Minor, Iran, and Central Asia to China and Japan. First records from Turkey, Uzbekistan and Afghanistan.
Hydaticus (Guignotites) histrio Clark, 1864
References. GUÉORGUIEV (1967: 474): “Khurd-Kabul*, SO von Kabul, 1900 m, 3.VII.1965, 1 ♂ (Kasy & Vartian)”, and “Nimla*, 40 km SW von Dschelalabad*, 23.VI.1965, 1 ♂ (Kasy & Vartian)”. Material studied. Afghanistan: 11 exs., “O [= E] Afghanistan, Prov. Nengrahar*, Povolný & Tenora”, “Tor- Khama*, 23.3.[19]66, 900 m” [PT41] (MZMB). 1 ex., “O [= E] Afghanistan, Prov. Nengrahar*, [D. Povolný]”, “Nimla*, 40 km SW Jalalabad*, 25.6.1965” (MZMB). 1 ♂, “O [= E]-Afghanistan, Kabulschlucht*, 22 km östl. [= east of] Kabul, 1650 m, 5.–12.6.1966, Amsel u. Ebert leg.” (SMNK).
Hydaticus (Guignotites) ricinus WEWALKA, 1979
References. WEWALKA (1979: 129): “O. Afghanistan, Sarobi*, 1100 m, 10.10.1961, leg. G. Ebert”. See also the records and notes under H. fabricii.
Material studied. Afghanistan: 1 ♀, “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Bokan*, 10.8.–11.8.1964, 1000 m” [J61, stream] (MZMB). This female could not be determined beyond doubt (see notes under H. fabricii). Notes. H. ricinus was described from Southern India and recorded from Afghanistan, Pakistan, Nepal and most regions of continental SE Asia (WEWALKA 1979). Along with H. ricinus we may also expect H. ponticus Sharp, 1882, to occur in Afghanistan; the latter species is known from India, Pakistan, Iran and the Near East.
Prodaticus pictus Sharp, 1882
Material studied. Afghanistan: 1 ♀, “O [= E] Afghanistan, Prov. Nengrahar*, D. Povolný et coll. “, “Tor- Khama*, 3.4.1967, 850 – 900 m” [P87] (MZMB). Syria: 1 ♀, “Syria, Ayyash, 10 km W Deir ez-Zur [= Dayr az Zawr], 35°25’N, 40°01’E, 219 m, 19.V.2001, A. Reiter leg.”, J. Hájek det. 2001 (JHCP). Turkmenistan: 1 ♀, “Turkmenia, Kugitang Mts. [= Khrebet Kugitangtau, mountain range at the border between south-eastern Uzbekistan and eastern Turkmenistan, ca. 37.30–38.00N 66.30–68.50E], Karljuk [= Karlyuk, ca. 37.35N 66.20E], 1.V.[19]91, Tusov lgt.”, J. Š•astný det. 1992 (JHCP). 6 ♂♂, “Kugitang [= Khrebet Kugitangtau], Umg. v. [= environs of] Karljuk [= Karlyuk], 490–520 m, 1.5.[19]89”, “Turkmenien, warmer bach [= warm stream], V.G. Dolin” (NHMB).
34 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan
Uzbekistan: 1 ♀, “Uzbekistan, Babatak – Sev. Kette [Babatak refers most probably to the Babatag Khrebet (mountain range) SW Dushanbe, ca. 37.30N 68E, in the border region between Tajikistan and Uzbekistan], 1400 m, 2.6.1987”, J. Hájek det. 2003 (NHMB). Notes. This species is widespread in the Middle East and is known from the Arabian Peninsula, Iran and Pakistan. ALI (1978) gave also Iraq. First records from Syria, Turkmenistan, Uzbekistan, and Afghanistan.
Subfamily Hydroporinae Tribe Bidessini
Hydroglyphus geminus (Fabricius, 1792)
References. GUIGNOT (1960: 320) (as Guignotus pumilus germinus [sic!]): “Tchehel Tan* (près Kaboul*), étang, 2000 m, 4 oct. 1957”, “Bosbal* (entre Ghourmatch* et Bala Morghab*), ruisseau et aux bords, 600 m, 21 oct. 1957”, “Chahr Safa* (entre Qandahar* et Qalat*), étang, 1240 m, 9 sept. 1957”, and “Guerechk*, étang du jardin public, 865 m, 29 avril 1958”. GUIGNOT (1961: 233) (as Guignotus pusillus ab. geminus): “Straße Gulbahar*-Sarobi*, Artemisia-Steppe, 1600 m”. Material studied. Afghanistan: 1 ex., “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 10.6.1964, 470 m” [J25, at light] (MZMB). 1 ex., same data but 11.6.–15.6.1964 [J26, at light] (MZMB). 1 ex., same data but 29.6.1964 [J36, at light] (MZMB). 2 exs., same data but 30.6.–2.7.1964 [J37, at light] (MZMB). 2 exs., same data but 10.7–13.7.1964 [J44, at light] (MZMB). 1 ex., same data but 10.7.1964 [J46, at light] (MZMB). 1 ex., “N Afghanistan, Prov. Kataghan*, coll. J. Šimek”, “10 km sö [= SE] Kunduz*, 1.4.1966, 450 m” [Š26] (MZMB).
Hydroglyphus signatellus (Klug, 1834) Material studied. Afghanistan: 3 exs., “N Afghanistan, Prov. Herat*, coll. J. Šimek”, “Bala Murghab*, 10.7.–13.7.1964, 470 m” [J44, at light] (MZMB). 4 exs., “N Afghanistan, Prov. Kataghan*, J. Šimek”, “10 km sö [= SE] Kunduz*, 1.4.1966, 450 m” [Š26] (MZMB). Notes. A widely distributed African and Palaearctic species. It occurs from North Africa and southern Europe throughout the Near East to Central Asia; in the east it reaches Tajikistan and Pakistan. First record from Afghanistan.
Tribe Hydroporini
Deronectes abnormicollis Semenov, 1900
References. GUÉORGUIEV (1963: 217): “Tang-Kharzar* (Hézaradjat*), torrent, alt. 1850 m, 1 ♀ à 25.VIII.1957 (K. Lindberg)”, “Gazestan* (entre Talaqan* et Kechm*), étang, alt. 1690 m, 10 ♂ à 12.VII.1959 (K. Lindberg)”, and “Mazahan* (vallée du Ghourband*, NO de Kaboul*), fossé, alt. 2080 m, 1 ♂ et 1 ♀ à 13.V.1959 (K. Lindberg)”; records cited without precise data in WEWALKA (1971: 136). FERY & HOSSEINIE (1998: 230): “Oizir*, S. du col d’Unaï*, Prov. de Maïdan* 14.VIII.75”, “Sefir Kuh*, Kuschke”, “Afghanistan, Mazanah*, 13.5.1959, Dr. K. Lindberg Leg.”, “Afghanistan, Gazestan*, 12.7.1959,
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 35 J. HÁJEK & H. FERY
Dr. K. Lindberg Leg.”, “Afghanistan, Tang-Kharzar*, 24.8.1957, Dr. K. Lindberg Leg.”, “Paghman-Geb.*, 2800 m, 26.VIII., O. Afghan. 1953, J. Klapperich”, and “J. Klapperich, Pagmangebirge*, 2300 m, 25.6.1952, O. Afghanistan”.
Material studied. Afghanistan: 2 ♂♂, 1 ♀, “Afghanistan, Behsud*, 2800m, 10.7.1971, leg. Kabakov O.N.” (COK). 2 ♀♀, “Afghanistan, 40 km NW Ghazni*, 2500 m, 28.5–12.6.1972, leg. Kabakov O.N.” (COK).
Deronectes afghanicus Wewalka, 1971
References. WEWALKA (1971: 139): “Nuristan*, Bashgultal* in 1200 m, 10.4.1953, leg. J. Klapperich” (type locality); data repeated in FERY & HOSSEINIE (1998: 239). Material studied. Afghanistan: 15 exs., “NO. [= NE] Afghan. 1953, J. Klapperich”, “Nuristan* 1200 m, Bashgultal* 10.IV.” (NHMB, MRTO, CGW, CHF). 2 exs., same data but 11.4.1953 (NHMB). 1 ♂, “J. Klapperich, Bashgultal* 1200 m, Nuristan* 10.4.1953, Afghanistan” (CHF).
Deronectes danielssoni Fery & Hosseinie, 1998
References. GUÉORGUIEV (1963: 217) (as Deronectes longipes SHARP, 1882): “Tang-Kharzar* (Hézaradjat*), torrent, alt. 1850 m, 1 ♂ à 25.VIII.1957 (K. Lindberg)”; record repeated in WEWALKA (1971: 140) (without precise data), and in FERY & HOSSEINIE (1998: 242) (description of the specimens under consideration as D. danielssoni).
Deronectes roberti Fery & Hosseinie, 1998
References. GUÉORGUIEV (1963: 217) (as Deronectes longipes): “Orozgan* (environs 175 NNE Quandhar*), bord de rivière, alt. 2100 m, 7 exempl. à 10.VI.1960 (K. Lindberg)”; record repeated in WEWALKA (1971: 140) (without precise data), and in FERY & HOSSEINIE (1998: 242) (description of the specimens under consideration as D. roberti).
Deronectes vestitus (Gebler, 1848)
Material studied. Afghanistan: 1 ♂, 2 ♀♀, “Afghan. Ghor Šarak*, 00m [sic!], 10.VII.1970, Kabakov [leg.]”, “Deronectes [20]01 persicus Peschet, Shaverdo det.” (COK). 1 ♀, “Afghan. Ghor Šarak*, 2800m, 10.VII.1970, Èistäkov [leg.]”, “Deronectes [20]01 persicus Peschet, Shaverdo det.” (COK). Notes. The one male D. vestitus studied coincides well with specimens from other countries, in particular with respect to their aedeagi. On the other hand, the three females have a rather weak, wave-like and twisted structure before the apex of the elytra (for expansion, see FERY & HOSSEINIE 1998: 236); the view that one might indeed believe these to belong to Deronectes persicus Peschet, 1914, which to date is only known from Iran. In our opinion, however, the description of a new taxon does not seem to be justified. D. vestitus has been recorded only from northern Kazakhstan, Russia (Western Siberia), Tajikistan and Uzbekistan. First record from Afghanistan.
36 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan
Deronectes sp.
Material studied. Afghanistan: 1 ♂, 1 ♀, “Afghan. Laghman*, Šamakat* 1500 m, 9.4.1972, Kabakov” (COK). 1 ♀, “Afghanistan, 40 km NW Ghazni*, 2500 m, 28.5–12.6.1972, leg. Kabakov O.N.” (COK). 1 ex., “Afghan. Ghazni*, W Moqur* 2300 m, 26.9.1972, Kabakov” (COK). Notes. The specimens from the three localities cannot be keyed safely to any of the known Deronectes and very probably represent three different and so far undescribed species. A thorough treatment of these deficiencies is due, and will be addressed in a future publication.
Hydroporus discretus Fairmaire & Brisout in Fairmaire, 1859
References. GUIGNOT (1960: 320) (as Hydroporus discretus woerndlei Zimmermann, 1919): “Badgah* (près Hari Roud*, à 30 km à l’ouest de Daoulet Yar*, Hézaradjat*), bassin d’une source, 2350 m, 16 août 1957”. Notes. NILSSON (2003a, b) did not record this species from Afghanistan.
Hydroporus glasunovi Zaitzev, 1905
References. GUÉORGUIEV (1963: 216): “Koul-Chogan* (lac Chiva*), bord du lac, bassin central, alt. 3100 m, 1 ♂ et 4 ♀ à 10.VIII.1960 (K. Lindberg)”.
Nebrioporus (s.str.) airumlus (Kolenati, 1845)
References. GUIGNOT (1960: 320) (as Potamonectes airumlus): “Bend Amir*, Lac Bend-Haïbat*, 3200 m, 2 août 1957”, “pentes herbeuses au bord du lac Bend-Ghambar*, 3190 m, 2 août 1957”, “Tang-Djebel Baz* ou Tang-Kharzar* (entre Khvadjah Tchicht* et Tang-Azao*, Hzaradjat*), dans torrent et au bord, 1850 m, 24 août 1957”, “Doab*, marécage près d’une source, 1450 m, 7 oct. 1957”, “Paghman* (près Kaboul*), sous pierres, bord de ruisseau, 2225 m, 18 mai 1958”, and “Bokan* - Qal’éh Nou*, steppe, sous pierres, 850 m, 24 oct. 1957”. Material studied. Afghanistan: 78 exs., “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 1.5–10.5.1964, 470 m” [J13, at light], M. Toledo det. 2000 (MZMB).
[?Nebrioporus (s.str.) elegans (Panzer, 1794)]
References. SATÔ (1966: 255) (as Potamonectes elegans): “11 ♂♂, Kabul, Afghanistan, June 10 & 17, 1960”, and “1 ♂, Doab-by Shewa*, Afghanistan, Sept. 10, 1960”. Notes. Although we could not study Satô’s material, we assume that he confused the widespread N. airumlus with the strictly European N. elegans.
Nebrioporus (s.str.) indicus Sharp, 1882 Material studied. Afghanistan: 1 ex., “O [= E] Afghanistan, Prov. Nengrahar*, Povolný & Tenora”, “2 km sw Jalalabad*, 18.3.[19]66, 600 m” [PT67, irrigation canals], M. Toledo det. 2000 (MZMB). 1 ex., “Afghanistan, Paktya*, Janny-Chel*, 2100 m, 23.VII.1964”, M. Toledo det. 2000 (MZMB).
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 37 J. HÁJEK & H. FERY
Notes. The species has previously been known only from northern Pakistan (Baltistan), northern India (Kashmir, Himachal Pradesh, Uttar Pradesh) and Nepal (TOLEDO 1998). First record from Afghanistan.
Nebrioporus (s.str.) mascatensis (Régimbart, 1897)
References. GUIGNOT (1960: 320) (as Potamonectes mascatensis): “Kouh-Bibtchéh Baran* (au nord de Farah*), mare de torrent, 900 m, 27 avril 1958”. Material studied. Afghanistan: 9 exs., “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 22.9.1964, 470 m” [J89], R. Mouchamps det. 1968, M. Toledo revid. 2000 (MZMB).
Notes. NILSSON (2003a, b) did not record this species from Afghanistan.
Nebrioporus (s.str.) ressli (Wewalka, 1974) Material studied. Afghanistan: 3 exs., “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 1.5.–10.5.1964, 470 m” [J13, at light] (MZMB). 10 exs., “N Afghanistan, Prov. Maimana*, coll. O. Jakeš”, “Rašíd*, 18.5.1964, 700 m” [J17] (MZMB). 3 exs., “N Afghanistan, Prov. Maimana*, coll. O. Jakeš”, “Èašma- i Yanbulaq*, 19.5.1964, 650–800 m” [J18/1], R. Mouchamps det. 1967 as Potamonectes sulphuricola (ZAITZEV, 1951), all material M. Toledo revid. 2000. (MZMB). Notes. The species has previously been known only from Iran. First record from Afghanistan.
Tribe Hydrovatini
Hydrovatus bonvouloiri Sharp, 1882
Material studied. Afghanistan: 1 ♀, “O [= E] Afghanistan, Prov. Nengrahar*, D. Povolný et coll.”, “Jalalabad, 19.–20.4.1967, 580 m” [P101] (MZMB). 1 ♂, 2 ♀♀, same data, but 21.–22.4.1967 [P103] (MZMB). 3 ♀♀, same data, but 24.4.1967 [P106] (MZMB). Notes. An Oriental species, originally described from “India bor.” and recorded from India, Sri Lanka, China, Japan, continental SE Asia, Sumatra, Java and the Philippines (BISTRÖM 1997). First record from Afghanistan.
Tribe Hygrotini
Herophydrus musicus (Klug, 1834) Material studied. Afghanistan: 1 ex., “N Afghanistan, Prov. Herat*, coll. J. Šimek”, “Bala Murghab*, 10.7.–13.7.1964, 470 m” [J44, at light] (MZMB). Notes. This species is widespread in the southern Palaearctic from North Africa and southern Europe to western and Central Asia and China. It reaches as far as India and Burma (NILSSON 1995). First record from Afghanistan.
38 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan
Hygrotus (Coelambus) confluens (Fabricius, 1787)
References. GUIGNOT (1960: 320): “Bokan* (entre Ghourmatch* et Bala Morghab*), mare, 460 m, 21 oct. 1957”. FERY (2003: 161): “Rašíd*, 18.5.1964, 700 m (17)” and “Herat-Gazergah*, 23.7.1964, 1000 m (54)”.
Material studied. Afghanistan: 1 ♀, “N Afghanistan, Prov. Maimana*, coll. O. Jakeš”, “Rašíd*, 18.5.1964, 700 m” [J17] (MZMB). 1 ♀, “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Herat-Gazergah*, 23.7.1964, 1000 m” [J54] (MZMB).
Hygrotus (Coelambus) enneagrammus (Ahrens, 1833)
References. FERY (2003: 168): “Afghanistan, Kuschke, Coll. Hauser 1896”, “Bala Murghab*, 1.6.–10.6.1964, 470 m (23)”, “Bala Murghab*, 30.6.–2.7.1964, 470 m (37)”, and “Murièaq*, 16.6.1964, 400 m (28)” (MZMB). Material studied. Afghanistan: 1 ex., “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 25.5.–10.6.1964, 470 m” [J21] (MZMB). 1 ♀, same data but 1.6.–10.6.1964 [J23] (MZMB). 1 ♀, same data but 30.6.–2.7.1964 [J37] (MZMB). 1 ♂, “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Murièaq*, 16.6.1964, 400 m” [J28, at light] (MZMB).
Hygrotus (Coelambus) impressopunctatus (Schaller, 1783)
References. FERY (2003: 147): “Afghanistan, Kuschke, Coll. Hauser 1896”, and “Band.i.Amir*, 2700 25.VIII.1975”.
Hygrotus (Coelambus) lernaeus (Schaum, 1857)
Material studied. Afghanistan: 1 ♀, “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 8.6.1964, 470 m” [J24, at light] (MZMB). 1 ♂, same data but 11.–15.6.1964 [J26, at light] (MZMB). 1 ♂, “Kouh-Tcehel Mastan*, 45 km W Herat*, No A 1018. Alt 1300 m, 7.6.1962”, “Afghanistan, K. Lindberg”, “A. 1018 K” (MRTO) Uzbekistan: 1 ♀, “Uzbekistan, Serawshan [Zerawshan Khrebet Mts.], Aman-Kutan [ca. 39.19N 66.56E], 2000 m, 1.06.1996, S. Bajduk lg”, H. Fery det. 2002 (JHCP). Notes. This species is widely distributed in the eastern Mediterranean and reaches eastwards to Uzbekistan, Afghanistan and Iran. The Afghan specimens show slight differences in their genitalia as compared with specimens from the eastern Mediterranean, and may represent a subspecies or even a new species. The status of these specimens must be clarified by further material from Afghanistan and adjacent areas. First records from Uzbekistan and Afghanistan.
Hygrotus (Coelambus) parallellogrammus (Ahrens, 1812)
References. FERY (1992: 121): “Afghanistan”, no additional data available. FERY (2003: 152): “Afghanistan, Kuschke, Coll. Hauser 1896”, and “Ost-Afghanistan, 69 40’L/34 35’B, Khurd-Kabul* 2000 m, SO v. Kabul, 11.6.1971, leg C. Holzschuh”.
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 39 J. HÁJEK & H. FERY
Hyphoporus pacistanus Guignot, 1959
Material studied. Afghanistan: 3 ♂♂, 4 ♀♀, “O [= E] Afghanistan, Prov. Nengrahar*, Povolný & Tenora”, “2 km sw Jalalabad*, 18.3.[19]66, 600 m” [PT67, irrigation canals] (MZMB). 2 ♀♀, “Kabul, Klapp[erich], II.[19]53”, “Hyphoporus sp.” [hw by Guéorguiev] (NMPC).
Notes. GUIGNOT (1959) described this species from Rawalpindi (Punjab prov.) in northern Pakistan. We compared our material to the holotype and two paratypes stored in ZSM. First record from Afghanistan.
Hyphoporus severini Régimbart, 1892
References. VAZIRANI (1977: 47): “Afghanistan, Seistan*”, no additional data given.
Tribe Hyphydrini
Hyphydrus gschwendtneri Guignot, 1942
Material studied. Afghanistan: 1 ♀, “O [= E]. Afghanistan, Prov. Paktia*”, “Khost*, 1200 m, 27.7.1964” (MZMB).
Notes. According to BISTRÖM (1982), this species occurs in Pakistan and northern India. First record from Afghanistan.
Subfamily Laccophilinae
Laccophilus minutus (Linnaeus, 1758)
References. GUIGNOT (1960: 320): “Bokan* (entre Ghourmatch* et Bala Morghab*), mare, 460 m, 21 oct. 1957” (f. typ.), and “Chindand* (entre Hérat* et Farh*), étang, 1110 m, 5 sept. 1957” (ab. mougeli Guignot, 1937).
Notes. NILSSON (2003a, b) did not record this species from Afghanistan.
Laccophilus poecilus Klug, 1834 Material studied. Afghanistan: 1 ex., “N Afghanistan, Prov. Herat*, coll. O. Jakeš”, “Bala Murghab*, 25.5.–10.6.1964, 470 m” [J21] (MZMB). 1 ex., “N Afghanistan, Prov. Kataghan*, coll. J. Šimek”, “Kunduz*, 22.5.1966, 400 m” [Š43] (MZMB). Uzbekistan: 2 ♂♂, “Uzbekistan, Zeravshan Mts., pr. Agalik [ca. 15 km SWS Samarkand, ca. 39.31N 66.34E], 19.5.1990, 1300 m, pools near the river, J. Strejèek lgt.”, J. Hájek det. 2001 (JHCP).
Notes. This species is known from almost the whole Palaearctic Region (BRANCUCCI 1983). First records from Uzbekistan and Afghanistan.
40 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan
Discussion The present paper summarises all data on the Hydradephaga from the territory of Afghanistan that we are aware of. Four species of the family Gyrinidae, four species of the family Haliplidae (including Haliplus variegatus that we include with reservations) and 52 species of the family Dytiscidae (including Nebrioporus elegans and Hydaticus fabricii that we include with reservations) are now known from Afghanistan; 14 of them are recorded as new from the country. There are no records of Hygrobiidae and Noteridae from Afghanistan. The list is still very inadequate and cannot be regarded as final. Continuation of the research on water beetles in Afghanistan is very desirable as most specimens were collected at lights. This method selects for species that are frequent and good fliers. Nevertheless, the basic zoogeographical characteristics of the Afghan water beetle fauna can be outlined. The species can be divided into several groups, with most species representing two main types of distribution: 1. Widely distributed Western Palaearctic species, which generally range throughout most of Europe, the Near East, Central Asia and reach eastwards to the Himalayas (e.g. Hygrotus confluens, Colymbetes fuscus). 2. Central Asian species, distributed mainly in the mountains and high plains of the Russian Central Asia, Uzbekistan, Kyrgyzstan, Kazakhstan, Tajikistan, Afghanistan, Pakistan, Kashmir and Nepal (e.g. Deronectes abnormicollis, Platambus sogdianus). 3. A small group containing Oriental species, which have the western border of their range in Afghanistan (e.g. Hydaticus ricinus). 4. A small group of species occurring in desert habitats in the Arabian peninsula and Iran and reaching Afghanistan, and possibly further regions in Central Asia, in the north-east (e.g. Prodaticus pictus). 5. Finally, a group of widely distributed “Old World” species such as Rhantus suturalis or Cybister tripunctatus. Only four species – all from running waters – appear to be endemic to Afghanistan: Orectochilus nurestanicus, O. afghanus, Deronectes danielssoni, and D. roberti.
Acknowledgements We would like to thank all the curators and other colleagues mentioned above for their help, and in particular Vítìzslav Kubáò (Brno, Czech Republic) for putting the Afghanistan water beetle material at our disposal. We are obliged to Helmut Schütze (Gleichen, Germany) and Robert Danielsson (Lund, Sweden) for their valuable help in the identification of some Afghan localities. We also thank Mario Toledo (Brescia, Italy) and Jaroslav Š•astný (Liberec, Czech Republic) for identifying the material of
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 41 J. HÁJEK & H. FERY
Nebrioporus and Colymbetes, respectively. We are indebted to David Boukal (Èeské Budìjovice, Czech Republic) for correcting the English of an earlier version of our manuscript. The present study was partly supported by a Ministry of Culture of the Czech Republic grant (MK0 CEZ 99F0201).
References
ADAMEC L. W. 1972: Historical and political gazetteer of Afghanistan vol. 1. Badakhshan province and northeastern Afghanistan. Akademische Druck- und Verlagsanstalt, Graz, xiii + 257 pp. + several maps. ADAMEC L. W. 1985: Historical and political gazetteer of Afghanistan vol. 6. Kabul and southeastern Afghanistan. Akademische Druck- und Verlagsanstalt, Graz, xx + 847 pp + several maps. ALI H. A. 1978: A list of some aquatic beetles of Iraq (Coleoptera: Dytiscidae). Bulletin of the Natural History Research Centre 7: 11–14. AMSEL H. G. 1967: Die zweite Deutsche Afghanistan-Expedition 1966 der Landessammlungen für Naturkunde Karlsruhe. Ein Reisebericht. Beiträge zur naturkundlichen Forschung in Südwestdeutschland 26: 3–14. BISTRÖM O. 1997: Taxonomic revision of the Genus Hydrovatus Motschulsky (Coleoptera, Dytiscidae). Entomologica Basiliensia 19(1996): 57–584. BRANCUCCI M. 1982: Les Platambus du sous-gengre Anagabus (Col., Dytiscidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 55: 115–124. BRANCUCCI M. 1983: Révision des espèces est-paléarctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey 31/32: 241–426. BRANCUCCI M. 1988: A revision of the genus Platambus Thomson (Coleoptera, Dytiscidae). Entomologica Basiliensia 12: 165–239. DUPREE N. H. 1971: An historical guide to Afghanistan. Afghan Tourist Organization Publication Number 5. Jagra, Tokyo, ix + 333pp. + 1 map. FERY H. 1992: Coelambus lagari n. sp. und Coelambus sanfilippoi n. sp. aus dem westlichen Mittelmeergebiet (Coleoptera: Dytiscidae). Entomologische Zeitschrift 102: 113–132. FERY H. 2002: Nomenklatorische und taxonomische Notizen zu einigen Dytiscidae (Coleoptera). Entomologische Zeitschrift 112: 25–30. FERY H. 2003: Dytiscidae: V. Taxonomic and distributional notes on Hygrotus Stephens, with emphasis on the Chinese fauna and a key to the Palearctic species (Coleoptera), pp. 133–193. In: JÄCH M. A. & JI L. (eds.): Water beetles of China. Vol. III. Zoologisch-Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien, VI + 572 pp. FERY H. & HOSSEINIE S. 1998: A taxonomic revision of Deronectes Sharp, 1882 (Insecta: Coleoptera: Dytiscidae) (part II). Annalen des Naturhistorischen Museums in Wien 100B: 219–290. GSCHWENDTNER L. 1938: Monographie der paläarktischen Dytiscidae. IX. Dytiscinae. Koleopterologische Rundschau 24: 33–76. GUÉORGUIEV V. B. 1963: Contribution à l’étude des coléoptères hydrocanthares (Haliplidae et Dytiscidae d’Afghanistan). Opuscula Entomologica 28: 215–222. GUÉORGUIEV V. B. 1967: Österreichische entomologische Expeditionen nach Persien und Afghanistan, Beiträge zur Coleopterologie, Teil X: Dytiscidae. Annalen des Naturhistorischen Museums in Wien 70(1966): 473–477. GUÉORGUIEV V. B. 1972: Notes sur les Agabini (Coleoptera, Dytiscidae), II. Révision des genres Platynectes Rég. et Colymbinectes Falk. Izvestija na Zoologitjeskija Institut s Musei Sofia 34: 33–62. GUÉORGUIEV V. B. 1975: Description d’un nouveau Agabus du Népal et notes sur quelques espèces du sous- genre Dichonectes Guignot (Coleoptera, Dytiscidae). Acta Zoologica Bulgarica 1: 97–103. GUIGNOT F. 1954: Quarante-et-unième note sur les hydrocanthares. Revue Française d’Entomologie 21: 195–202. GUIGNOT F. 1959: Dytiscides du Pakistan occidental. Bulletin Mensuel de la Société Linnéenne de Lyon 28: 8–11.
42 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan
GUIGNOT F. 1960: Haliplides et dytiscides de l’Afghanistan (Col.) (Contribution à l’étude de la faune d’Afghanistan. 7). Bulletin de la Société Zoologique de France 84(1959): 319–321. GUIGNOT F. 1961: Ergebnisse der Deutschen Afghanistan-Expedition 1956 der Landessammlungen für Naturkunde Karlsruhe, Dytiscidae (Coleoptera). Beiträge zur naturkundlichen Forschung in Südwestdeutschland 19 (1960): 233. JAKEŠ O. & POVOLNÝ D. 1967: Beiträge zur Kenntnis der Fauna Afghanistans (Sammelergebnisse von O. Jakeš 1963–64, D. Povolný 1965, D. Povolný & Fr. Tenora, 1966, J. Šimek 1965–66). Reisebericht und Charakteristik des Sammelgebietes. Èasopis Moravského Musea, Vìdy Pøírodní 52(Suppl.): 9–34. KLAPPERICH J. 1954: Auf Forschungsreisen in Afghanistan. Entomologische Blätter 50: 107–118. LEGROS C. 1977: Un dytiscide nouveau d’Afghanistan (Col. Hydrocanthares). Bulletin de la Société Entomologique de France 82: 14–16. MAZZOLDI P. 1995: Gyrinidae: Catalogue of Chinese Gyrinidae (Coleoptera), pp. 155–172. In: JÄCH M. A. & JI L. (eds.): Water beetles of China. Vol. I. Zoologisch-Botanische Gesellchaft in Österreich and Wiener Coleopterologenverein, Wien, 410 pp. MAZZOLDI P. 2003: Family Gyrinidae, pp. 26–30. In: LÖBL I. & SMETANA A. (eds.). Catalogue of Palaearctic Coleoptera, 1. Archostemata – Myxophaga – Adephaga. Apollo Books, Sternstrup, 819 pp. MILLER K. B. 2002: Revision of the genus Eretes Laporte, 1833 (Coleoptera: Dytiscidae). Aquatic Insects 24: 247–272. NILSSON A. N. 1995: Noteridae and Dytiscidae: Annotated check list of the Noteridae and Dytiscidae of China (Coleoptera), pp. 35–96. In: JÄCH M. A. & JI L. (eds.): Water beetles of China. Vol. I. Zoologisch- Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien, 410 pp. NILSSON A. N. 1998: Dytiscidae: V. The genus Platynectes Régimbart in China, with a revision of the dissimilis- complex (Coleoptera), pp. 107–121. In: JÄCH M. A. & JI L. (eds.): Water beetles of China. Vol. II. Zoologisch-Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien, 371 pp. NILSSON A. N. 2003a: Catalogue of Palearctic Dytiscidae. Internet version as present on June 16th 2003 at: http://www.bmg.umu.se/biginst/andersn/Cat_main.htm. NILSSON A. N. 2003b: Family Dytiscidae, pp. 35–78. In: LÖBL I. & SMETANA A. (eds.): Catalogue of Palaearctic Coleoptera, 1. Archostemata – Myxophaga – Adephaga. Apollo Books, Sternstrup, 819 pp. OCHS G. 1955: Die Gyrinidae (Coleoptera) der Afghanistan-Expedition (1952 u. 1953) J. Klapperichs. Entomologisk Tidskrift 76: 46–51. ROUGHLEY R. E. 1990: A systematic revision of species of Dytiscus Linnaeus (Coleoptera: Dytiscidae). Part 1. Classification based on adult stage. Quaestiones Entomologicae 26: 383–557. SATÔ M. 1966: Some species of the aquatic coleoptera from Afghanistan and Pakistan. Results of the Kyoto University scientific Expedition to Karakorum Hindukush 1955 8: 255–256. SATÔ M. 1972: Records of aquatic Coleoptera from Afghanistan, India, Laos, E. Pakistan and S. Vietnam. The Entomological Review of Japan 24: 49–53. SCHÜTZE H. & KLEINFELD F. 1997: Carabusformen Sibiriens und Zentral-Asiens. Privately printed, Gleichen, Fürth, 198 pp. SHAVERDO H. V. 2002: New data on distribution of some species of Agabus Leach and Ilybius Erichson (Coleoptera: Dytiscidae). Koleopterologische Rundschau 72:123–124. Š•ASTNÝ J. 2003: Dytiscidae: X. Review of Platynectes subgen. Gueorguievtes Vazirani from Southeast Asia (Coleoptera), pp. 217–259. In: JÄCH M. A. & JI L. (eds.): Water beetles of China. Vol. III. Zoologisch- Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien, VI + 572 pp. TOLEDO M. 1998: Dytiscidae: II. The genus Nebrioporus Régimbart, 1906 in China (Coleoptera), pp. 69–91. In: JÄCH M. A. & JI L. (eds.): Water beetles of China. Vol. II. Zoologisch-Botanische Gesellchaft in Österreich and Wiener Coleopterologenverein, Wien, 371 pp. VAZIRANI T. G. 1977: Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India Miscellaneous Publication Occasional Paper 6(1976): 1–111. VONDEL B. J. VAN 1991: Revision of the Palaearctic species of Haliplus subgenus Liaphlus Guignot (Coleoptera: Haliplidae). Tijdschrift voor Entomologie 134: 75–144. VONDEL B. J. VAN 2003: Family Haliplidae, pp. 30–33. In: LÖBL I. & SMETANA A. (eds.). Catalogue of Palaearctic Coleoptera, 1. Archostemata – Myxophaga – Adephaga. Apollo Books, Sternstrup, 819 pp.
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 43 J. HÁJEK & H. FERY
WEWALKA G. 1971: Revision eines Teiles der Gattung Deronectes Sharp (Col.) mit vier Neueschreibungen. Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen 22(1970): 126–142. WEWALKA G. 1975: Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Dytiscidae, unter Berücksichtigung einiger Arten aus benachbarten Gebieten. Entomologica Basiliensia 1: 151–165. WEWALKA G. 1979: Revision der Artengruppe des Hydaticus (Guignotites) fabricii (MacLeay), (Col., Dytiscidae). Koleopterologische Rundschau 54: 119–139. ZAITZEV F.A. 1953: Fauna SSSR, Nasekomye Žestkokrylye, Tom 4, plavuncovye i vertjaèky. Izdatelstvo Akademii nauk SSSR, Moskva – Leningrad, 376 pp [in Russian]. ZAITZEV F. A. 1972: Fauna of the USSR 58. Coleoptera IV. Families: Amphizoidae, Hygrobiidae, Haliplidae, Dytiscidae, Gyrinidae. Israel Program for Scientific Translations, Jerusalem, 401 pp. [English translation of ZAITZEV (1953)].
44 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan Appendix
Appendix
List of localities Although we cross-referenced several comprehensive works on Afghan localities, we cannot be certain that all data given below are correct. This is due to the following facts: • local names change in time • different Afghan tribes use different local names • borders and names of provinces have been changing • spellings of localities in entomological publications depend on the language The following are explanations for the localities that seem to correspond to the data given on the labels or cited in the literature. Several localities exist with the same name; there are, however, only a few cases and they are given additional explanations. In these cases the different possibilities for one name are denoted with numbers in brackets.
We used the following routine for the identification of localities: (1) Find the same or a similar name in a reasonable position in MS Encarta 2000. If no success – (2) find the same or a similar name in another source, and – (3) use the position from the other source to find the current name in Encarta. In both cases – (4) get the latitude and longitude from Encarta. All localities are arranged alphabetically. Different spellings or names are presented in brackets. Ali Cheng: in the Laghman prov: ca. 34.47N 70.05E. Andkoï (= Andkhvoy): in the Faryab prov., ca. 36.57N 65.07E. Badakhchan (= Badachschan, Badakhshan): a province in NE Afghanistan. Badgah: in the Ghowr prov., ca. 34.31N 65.28E. Baharaknet (= Baharak, Baharestan): in the Badakhshan prov., ca. 37.00N 70.53E. Bala Bokan (= Bala Bowkan): in the Badghis prov., in NW Afghanistan, ca. 35.34N 63.37E. Two nearby localities with the name Bokan (= Bowkan) are situated ca. 10 km north of Bala Bokan (see under Bokan). Bala Morghab (= Bala Murghab, Morghab): in the Badghis prov., in NW Afghanistan, ca. 35.33N 63.20E. Bala Murghab: see Bala Morghab. Balkh: a province in N Afghanistan. Bamvardar Aoudak: (most probably = Awdak): in the Parvan prov., ca. 35.30N 69.51E. Bamyian (= Bamian): (1) a province ca. 150 km W Kabul; (2) a village of same name, ca. 34.49N 67.31E. Bande-amir: see Band-e Amir.
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 45 J. HÁJEK & H. FERY Appendix
Band-e Amir: Several names listed in the “Gazetteer of Afghanistan” (1992: 104), come into consideration, some denoting streams and others “populations”. However, we are sure that the name refers to an area in the Bamian prov. that contains “a series of natural dams across the bed of the Band-i-Amir [a stream]” (ADAMEC 1985: 88). These dams form several lakes (DUPREE 1971: 128 ff.); the co-ordinates are ca. 34.50N 67.12E. Band.i.Amir, Band-i-Amir: see Band-e Amir. Bashgultal (= Bashghul valley, Bashghul): river in the NE part of the Konar prov., near the Pakistani border; ca. 35.40N 71.20E. Behsud: ca. 10 km ENE Jalalabad, ca. 34.26N 70.30E. Beltchiragh (= Belcherag): in the Faryab prov. (= the former Meymaneh or Maimana prov.), ca. 35.49N 65.14E. Bend Amir: see Band-e Amir. Bend-Ghambar (= Band-i-Kambar): a lake in the Band-e Amir area; see under that name. Bend-Haïbat (= Bend-Haïbat, Band-i-Haibat): a lake in the Band-e Amir area (see under this name). Bokan (= Bowkan): (1) a village in the Badghis prov., in NW Afghanistan; (2) two nearby localities of the same name, ca. 35.42N 63.32E; (3) another locality named Bala Bokan (= Bala Bowkan) is situated ca. 10 km S of Bokan (see under Bala Bokan). Borak: We have not been able exactly to identify this locality; there are several possibilities: (1) Kotal Borak (= Kowtal-e Borak), a pass in Farah prov., 32.58N 63.43E; (2) Borak Kaly, in the Paktia prov., 33.08N 68.35E; (3) several localities with the names Barak; (4) Borjak, in the Vardak prov., ca. 34.34N 68.11E. Bosbal (most probably = Bozbai): in the Badghis prov., in NW Afghanistan, ca. 35.27N 63.22E; however, Bozbai is situated ca. 10 km S of Morghab and not “entre Ghourmatch et Bala Morghab” as indicated by GUIGNOT (1960: 320). Èašma-i Yanbulaq (= Cheshme-ye Yanbolaq): in the Faryab prov. (= the former Meymaneh or Maimana prov.), ca. 35.48N 63.56E. Chahr Safa (= Shar-e Safa): in the Zabol prov., ca. 31.49N 66.21E. Cheikhabad (= Sheykhabad, Saydabad): in the Vardak prov., ca. 34.05N 68.45E. Chevaki (most probably = Sevak): in the Vardak prov., ca. 34.16N 68.37E. We have found several other localities with the name “Sevak”, but all of them are further away from Kabul, while GUIGNOT (1960: 320) wrote “près Kaboul*” [= near Kabul]. Chibarg: not found on any map; according to the data of the Lindberg expedition in 1957, it should be Sheberg (= Qeshlag-e Sheberg), in the Ghowr prov., ca. 34.14N 64.05E. Chileh Hamman (most probably = Haman): in the Herat prov., ca. 34.35N 63.08E. Chindand (= Shindand): ca. 100 km N Farah, in Farah province, ca. 33.18N 62.10E. Chiva (= lac Chiva, Chiva lake, Shiva lake, Kowl-e Shiveh, Kul-i-Shiwa, Koul-Chogan): in the Badakhshan prov., ca. 37.23N 71.21E.
46 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan Appendix
Chivah (= Pamir de Chivah): a mountain region, highlands east of the Shiva lake, in Badakhstan prov. Col d’Unaï (= Unai pass, Kotal-i-Unai, Kowtal-e Wonay): in the Vardak prov., ca. 34.27N 68.22E. Dand Karkatak: not found on any map; according to GUIGNOT (1960: 321) a pond near Rah-Kol (see under that name), in the Vardak prov. Daoulet Yar (= Dowlat Yar): in the Ghowr prov., ca. 34.33N 65.46E. Dasht-i Nawar (= Dasht-e Navar): in Ghazni prov., ca. 33.35N 67.48E. Decht Ghoudjour: not found on any map. Dival Kol: four nearby localities may correspond to this name: (1) Dival Qowl (34.20N 67.41E), (2) Dival Qowl (= Diwal Qowl) (34.24N 67.52E), (3) Diwal Qol (34.18N 67.53E), and (4) Gardan Diwal (ca. 34.30N 68.15E), all in the Vardak prov. Djabir Ansar: not found on any map; according to R. Danielsson (pers. comm.): “near Kabul, 1850 m a.s.l., lake Chodar Soualhim”. Doab: Doab Mekh-i-Zarin, 1450 m a.s.l. (R. Danielsson, pers. comm.), in the Bamian prov., ca. 35.14N 67.58E. Doab-by Shewa (= Doabe Shiwa): not found on any map but certainly in the Chiva region. Doavi: we have found three localities which might come into consideration: (1) Doabi (ca. 36.33N 70.20E), (2) Doab (ca. 36.50N 70.31E), and (3) Do Abi Yaftal (ca. 37.16N 70.33E); all in Badakhstan prov. (NE-Afghanistan). Dschelalabad: see Jalalabad. Farah: a province in SW Afghanistan and the capital of this province, ca. 32.22N 62.06E. Farh: see Farah. Gazergah: in the Herat prov., ca. 10 km NE Herat, ca. 34.22N 62.13E. Gazestan: in the Takhar prov., in NE Afghanistan, ca. 36.39N 69.47E. Ghazni (= Ghasni): a province SW of Kabul and the capital of this province, ca. 33.32N 68.25E. Ghor Šarak: see Qal’éh Chahrak. Ghourband [vallée de] (= Ghourband valley, Ghorband valley): ca. 75 km NW Kabul; the stream is named Darya-ye Ghowr Band. Ghourmatch (= Ghowrmach, Ghormaè): in the Badghis prov., in NW Afghanistan, ca. 35.44N 63.47E. Guerechk (= Gerechk, Girishk): in the Kandahar prov., ca. 100 km WNW Kandahar, ca. 31.49N 64.33E. Gulbahar (= Golbahar): in the Kapisa prov., ca. 80 km N Kabul, ca. 35.08N 69.17E. Haouz-Mahiyan (= Hauz-i-Mahya, Sar-i-Chahma): in the Vardak prov., ca. 34.25N 68.31E; according to ADAMEC (1985: 701) “8½ miles east of Unai pass”; according to R. Danielsson (pers. comm.) “65 km W Kabul”. Hari Roud: see Hari Rud. Hari Rud (= Harirud, Hari Roud, Heri Rud): a river in NW Afghanistan (the Herat and Ghowr provinces).
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 47 J. HÁJEK & H. FERY Appendix
Herat: (1) a province in NW Afghanistan, and (2) the capital of this province, ca. 34.20N 62.11E. Herat-Gazergah: see Gazergah. Hézaradjat (= Hazarajat, Hazarehjat, Hazara (“THE TIMES ATLAS”), Hasara (“STIEHLERS HANDATLAS”), Hazaristan): a region W of Kabul, between Herat and Kabul (see ADAMEC 1985: 251 for more details). Hzaradjat: see Hézaradjat. Ichkachim: see Ishkashim. Ishkashim (= Ashkasham, Eshkashim): in the Badakhshan prov., ca. 36.42N 71.33E. Jalalabad: the capital of the Nangarhar prov., ca. 34.25N 70.25E. Janny-Chel (= Janekkhel): in the Paktia prov.; ca. 33.28N 69.07E. Kaboul (= Kabol, Kabul): the capital of Afghanistan. Kabulschlucht (= Kabul Gorge): ca. 25 km E of Kabul. Kalu [rivière] (= Kalu river): in the Vardak (or Bamian) prov., ca. 34.39N 68.03E; the village of Kalu (= Kalow) has the same co-ordinates. Kandahar: a province in SE Afghanistan and the capital of this province, ca. 31.36N 65.41E. Kandahar-Kuna: according to KLAPPERICH (1954: 117), the ruins of the ancient Kandahar, situated 5 km W of today’s Kandahar. Kataghan prov.: the Kondoz province (= Kunduz prov.) in N Afghanistan. Kechem (= Kischm): in the Badakhshan prov. in NE Afghanistan, ca. 36.48N 70.06E. Khinjan (= Khenjan): (1) a village, (2) a river, and (3) a region, in Baghlan prov., ca. 35.36N 68.56E. Khost (= Khowst): in the Paktia prov., 33.20N 69.55E. Khot-Gaï: see Kotkai. Khurd-Kabul (= Khord Kabul, Khvord Kabul [= little Kabul]): ca. 25 km SE of Kabul, in the Kabul prov., ca. 34.23N 69.23E; the collector C. Holzschuh gave the co- ordinates 34.35N 69.40E; we believe that his data are inaccurate. Khvadjah Tchicht (= Khvajeh Chesht, Khwajachisht, Chwadscha-gischt (“Stiehlers Handatlas”), Chesht-e Sharif): (1) a village in the Herat prov., 34.21N 63.44E; (2) there exists also the stream Darreh-ye Khvajeh Chesht, ca. 34.19N 63.43E. Kotal-Lachva: not found on any map, most probably a pass; according to GUIGNOT (1960: 320) situated between Qal’éh Chahrak and Chibarg (see under these names), and thus in the Ghowr prov. Kotkai (= Kotgay): in the Paktia prov., 33.59N 69.50E. Kouh-Bibtchéh Baran (= Kuh-e Bibicheh Baran, Bibcha Baran, Bibchi Baran): a mountain range in Farah prov., ca. 20 km NW of Farah, ca. 32.28N 61.54E. Kouh-Tcehel Mastan: not found on any map, according to the label data “45 km W Herat”, thus certainly in the Herat prov. Koul-Chogan (= lac Chiva, Shiva lake): see under Chiva. Kunduz (= Kondoz): (1) a province in N Afghanistan, and (2) the capital of this province, ca. 36.43N 68.51E.
48 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan Appendix
Kutiau (= Kutya): in the Konar prov., ca. 35.16N 71.29E. Laghman: a province NE of Kabul. Maïdan [prov.]: the Vardak province, W of Kabul. Maimana prov. (= Meymaneh prov.): a province in NW Afghanistan, today known as the Faryab province. Mangan: in the Badghis prov., in NW Afghanistan, ca. 35.27N 63.09E. Masdjed-Tchoubi (= Masjed-e Chobi, Masjed-e Chubi): in the Herat prov., ca. 34.34N 63.05E. Mazahan: a misspelling of “Mazanah”; see under this name. Mazanah: in the Parvan prov., NW of Kabul, ca. 34.59N 68.37E. Mazar-i-Sharif (= Mazar-e Sharif): (1) capital of the Balkh prov.; (2) a province formerly named “Mazar-e Sharif” or “Samangan”. Mohammed Zaher [lac]: this lake was not found on any map; according to GUÉORGUIEV (1963: 219) east of “lac Chiva”, thus probably in Badakhstan prov. Moqor: in the Ghazni prov., ca. 32.48N 67.46E. Moqur: see Moqor. Murièaq (= Murichaq): in the Badghis prov., in NW Afghanistan, ca. 35.48N 63.08E. Narin: we have not been able to precisely locate this place; we assume that it is a region or a village of that name, SW of Baghlan city, in Baghlan prov., ca. 36.04N 69.07N. Nengarhar prov.: the Nangarhar province, E of Kabul. Nengrahar: a misspelling of “Nengarhar”; see under that name. Nimla (= Nemla, Nemleh, Memlah): ca. 40 km SW of Jalalabad, in the Nangarhar prov., ca. 34.18N 70.06E. Nuristan: a region in the Konar and Laghman provinces, ca. 200 km NE of Kabul. Oizir: not found on any map, however, this locality must be located close to the “col d’Unaï” (see under that name). Orozgan: see Oruzgan. Oruzgan: (1) Oruzgan province, ca. 200 km SW of Kabul; (2) a homonymous village in that province, ca. 32.55N 66.38E. Pagham [“near Kabul”]: an incorrect spelling of “Paghman”; see under that name. Pagman: the Paghman mountain range, W of Kabul. Paghman-Geb.: see Pagman. Pagman-Gebirge: see Pagman. Paktia (= Paktiya, Paktya): the Paktia province, ca. 150 km SSE of Kabul. Paktya: see under Paktia. Pandjab (= Panjab, Panjao): in the Bamian prov., ca. 34.23N 67.01E. Pandjchir (= Panjshir valley): (1) a valley in the Parvan prov., NE of Kabul, drained by the river of the same name; (2) the village Tel-e Panjshir, ca. 35.33N 69.51E. Parakh: in the Parvan prov., ca. 35.16N 69.28E. Paropamise: the Paropamisus mountain range, in NW Afghanistan, chiefly in the Herat, the Badghis and Ghowr provinces. Petschtal (= valley of the Peè Dara, Darrah-ye Pich stream): in the Konar prov., ca. 34.52N 71.09E.
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 49 J. HÁJEK & H. FERY Appendix
Pir-Khil: not found on any map; according to GUÉORGUIEV (1963: 219) ca. 55 km NE of Orozgan, thus located at ca. 33.17N 67.06E. Polichomri (= Pol-e Khomri, Pol-i-Chumri): (1) a village in the Baghlan prov., ca. 35.58N 68.43E; (2) a stream south of that village is named Darya-ye Pol-e Khomri, in the Baghlan prov., ca. 35.05N 68.36E. Qalat: in the Zabol prov., in S Afghanistan, ca. 32.06N 66.53E. Qal’éh Chahrak (= Shahrak): in the Ghowr prov., 34.06N 64.18E. Qal’eh-Darrech Zang (= Qal’eh-Darreh Zang, Darreh Zang): a village in the Faryab prov., ca. 35.37N 65.16E, situated in the valley of Dara-i Zang (= Darreh-ye Zang) of the stream Darya-ye Darreh-ye Zang. Qal’eh-Darreh Zang: see under Qal’eh-Darrech Zang. Qal’éh Nou (= Qal’e-ya Now, Qala-i Naw): in the Badghis prov., in NW Afghanistan, ca. 34.59N 63.07E. Qandahar: see Kandahar. Quandhar: see Kandahar. Rah-Kol (= Sar-e Raqowl): in the Vardak prov., 34.20N 67.51E. Rašíd (= Rashid): in the Badghis prov., in NW Afghanistan, ca. 35.47N 63.43E. Sabzzak [col de]: Sabzak pass, a pass in the Kuh-e Band-e Sabzak mountain range, in the Herat or Badghis provinces, in NW Afghanistan, ca. 34.36N 63.06E. Safed Koh (= Safid Kuh, Sefid-Kuh (“Stiehlers Handatlas”)): a mountain range in the Nangarhar and Paktia provinces, ca. 50 km S Jalalabad, ca. 34.00N 70.30E. Salang-Paß (= Salang pass): in the Baghlan prov., ca. 35.18N 69.03E. Šamakat (=Shamakat): in the Laghman prov., ca. 34.43N 70.01E. Sarobi (= Sarowbi): ca. 50 km E Kabul, in the Kabul prov., 34.36N 69.44E. Sefir Kuh: most probably a misspelling of “Sefid Kuh”. According to the known distribution of Deronectes abnormicollis, we assume that the name “Sefir Kuh” means a mountain range that is termed Qasa Murg in “The Times Atlas”, ca. 100–200 km E of Herat, S of Khvadjah Tchicht and S of the river Hari-Rud (see also Ochs’ notes on a “Kuschke” label under Gyrinus distinctus). “Stiehlers Handatlas” still contains “Sefid Kuh” (beside another mountain range “Sefid Kuh” which is identical with “Safed Koh”; see under that name), it is seemingly a name used in the 19th and at the beginning of the 20th century. “Gazetteer of Afghanistan” mentions a mountain range, which seems to be identical with that under consideration: Safedkoh (= Kuh-e Sefed), ca. 34.06N 63.22E. Another mountain range (N of the Hari-Rud river) is named Safed Koh = Selseleh-ye Sefid Kuh, ca. 34.30N 63.30E. Seistan: a region in south-western Afghanistan and eastern Iran. The Afghan part belongs to the Farah and Nimruz provinces and includes a lake called “Daryacheh-ye Sistan”, “Hamun-e Helmand” (the river Helmand feeds this lake), or “Hamun-e Saberi”; moreover, there exists a village of “Sistan” in the Nimruz prov. (ca. 30.14N 61.36E) and another village in Iran (Sistan va Baluchistan prov.) called “Band-e Sistan” or “Band-i-Seistan” (ca. 30.50N 61.45E, directly on the border of Afghanistan). Shahidan: a fort in the Paktia prov., ca. 34.10N 69.40E. Shewa (= Shewah, Shiveh): in the Nangarhar prov., ca. 34.34N 70.34E.
50 Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 Hydradephaga of Afghanistan Appendix
Siah Guerd (= Siah Gerd): in the Parvan prov., ca. 34.59N 68.50E. Sirouas: not found on any map; according to GUIGNOT (1960: 320) this locality is situated between Moqor and Qalat; according to R. Danielsson (pers. comm.): “32.30N 67.30E, 1660 m”, thus the village should be situated in the Zabol prov. Sirouas-Salar: not found on any map; according to GUIGNOT (1960: 321) this locality is situated between Cheikhabad and Soltan Khel (both in the Vardak prov., see under the latter two local names), on the road from Kabul to Ghazni. We found a village called “Salar” which corresponds well to the Guignot description, co-ordinates ca. 33.54N 68.40E. Soltan Khel (= Soltan Kheyl): in the Vardak prov., ca. 33.52N 68.37E; on the road from Kabul to Ghazni. Soufian (= Sufian): in the Badakhshan prov., ca. 36.44N 71.04E. Talaqan (= Taloqan, Taluquan, Talogan, Talachan): the capital of the Takhar prov. in NE Afghanistan, ca. 36.44N 69.32E. Tang-Azao (= Tangi Azao, Robat-e Tangi Azow): in the Ghowr prov., 34.07N 64.12E. Tang-Djebel Baz: see under Tang-Kharzar. Tangi-Saitan (= Tangi Sayedan): ca. 10 km SSW of Kabul, 34.25N 69.07E. Tang-Kharzar (= Kharzar or Tang-Djebel Baz): in the Herat prov., 34.17N 63.58E. Tashkurgan (= Kholm): (1) a village in the Balkh prov. (= former Mazar-i-Sharif prov.), ca. 36.41N 67.41E; (2) a river named Tashkurgan Rud (= Darya-ye Samangan), ca. 36.40N 67.41E. Tchakaran (= Chakaran): in the Badakhshan prov., ca. 36.54N 71.03E. Tchehel Tan: (1) we have found one locality named Cheheltan, in Lowgar prov. (33.56N 68.52E), and (2) another one named Chehek Tan, in the Bamian prov. (34.43N 67.37E); both are situated “près” [= near] Kabul. Tor-Khama (= Tor Kham, Towr Kham): ca. 80 km ESE of Jalalabad, at the border to Pakistan, in the Nangarhar prov., 34.07N 71.05E. Vakahn (= Wakhan, Seseleh-ye Kuh-e Vakhan, Khrebet Vakhanskiy): (1) a mountain range in S Tajikistan and NE Afghanistan (the Badakhshan prov.), ca. 36.40–37.10N 71.30–74.00E; (2) the name is also in use for the corridor to China (DUPREE 1971: 269). Ville Rouge (= Red Village, Shar-i-Zohak): according to DUPREE (1971: 125) ca. 17 km E of Bamian “at the confluence of the Bamyian and Kalu Rivers”; in the Bamian prov., ca. 34.53N 68.01E. Yaoulang: not found on any map; according to GUÉORGUIEV (1963: 219) on the western slope of the “Pamir de Chiva”, thus probably in Badakhstan prov. Yelgah: not found on any map; according to GUÉORGUIEV (1963: 219) east of “lac Chiva”, thus probably in Badakhstan prov. Zebak (= Zibak): in the Badakhshan prov., ca. 36.31N 71.20E.
Acta Mus. Moraviae, Sci. Biol. (Brno), 89, 2004 51