Novitates PUBLISHED by the AMERICAN MUSEUM of NATURAL HISTORY CENTRAL PARK WEST at 79TH STREET, NEW YORK, N.Y

AMERICAN MUSEUM Novitates PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, N.Y. 10024 Number 3062, 9 pp., 8 figures, 5 maps June 10, 1993

A Review of the Cuban Ground Spiders of the Family Gnaphosidae (Araneae, Gnaphosoidea)

GIRALDO ALAYON G.1 AND NORMAN I. PLATNICK2

ABSTRACT

Sixteen species ofGnaphosidae are known from is described for the first time. Sergiolus kastoni Cuba. A new genus, Cubanopyllus, is described for Platnick and Shadab, S. cyaneiventris Simon, Ce- Litopyllus inconspicuus Bryant, the male ofwhich sonia bilineata (Hentz), C. irvingi (Mello-Leitio), is described for the first time. A new species ofthe and Urozelotes rusticus (L. Koch) are newly re- Caribbean gnaphosine genus Microsa, M. cubitas, corded from Cuba. is described. The male of Cesonia cincta (Banks)

INTRODUCTION Only a few arachnologists have reported sequently synonymized with G. sericata (L. spiders ofthe family Gnaphosidae from Cuba. Koch). The most important work on the fau- Banks (1909, 1914) described two species, na to date is Bryant (1940), in which seven Eilicina cincta and Callilepis grisea (both now gnaphosid species were recorded, and three placed in Cesonia), from La Habana and Pi- genera (Caridrassus, Paramyrmecion, and nar del Rio, respectively. Franganillo (1926) Eilicina) were newly established (all three ge- described Gnaphosa simplex, which was sub- neric names have been synonymized in sub-

' Kalbfleisch Research Fellow, Department of Entomology, American Museum of Natural History; Curator of Arachnida, Museo Nacional de Historia Natural, Ciudad de La Habana, Cuba. 2 Chairman and Curator, Department of Entomology, American Museum of Natural History; Adjunct Professor, Department of Biology, City College, City University of New York; Adjunct Professor, Department of Entomology, Cornell University.

Figs. 1, 2. Microsa cubitas, new species, male palp. 1. Ventral view. 2. Retrolateral view. sequent revisions). Platnick (1975) and Plat- seum of Comparative Zoology (MCZ, cour- nick and Shadab (1975, 1980a, 1980b, 1981, tesy of H. W. Levi), the Museum National 1982) revised the Nearctic elements of the d'Histoire Naturelle, Paris (MNHN, courtesy family, including Cuban records from ma- of C. Rollard), the Museo Nacional de His- terial deposited in North American collec- toria Natural, Havana (MNHNC), and the tions. Finally Alayon (1992, in press) de- National Museum ofNatural History, Wash- scribed two new species, one in the genus ington (USNM, courtesy of J. Coddington). Zelotes and the other in Camillina. We thank M. U. Shadab (AMNH) for help In this paper we report on specimens from with illustrations, and C. D. Dondale of the Cuban collections taken, over the last two Centre for Land and Biological Resources decades, in various parts of the island. We Research, Ottawa, and J. A. Murphy of describe one new genus (including the first Hampton, England, for helpful comments on known males of its type species, Litopyllus a draft of the manuscript. inconspicuus Bryant), one new species of the Caribbean gnaphosine genus Microsa, and the SYSTEMATICS male of Cesonia cincta (Banks), and present five new records for Cuba as well as new lo- Gnaphosa sericata (L. Koch) cality data for most ofthe remaining species. Map 1 The format of the descriptions and stan- Pythonissa sericata L. Koch, 1866: 31, pl. 2, figs. dard abbreviations of morphological terms 21, 22 (male holotype from Baltimore, Mary- follow those used in Platnick and Shadab land, in BMNH, examined). (1975). All measurements are in millimeters. Gnaphosa simplex Franganillo, 1926:49 (male ho- Material was obtained from the collections lotype from Santa Clara, Cuba, lost). First syn- ofthe American Museum ofNatural History onymized by Bryant, 1940: 390. (AMNH), the Natural History Museum, Gnaphosa sericata: Platnick and Shadab, 1975: London (BMNH, courtesy ofP. Hillyard), G. 61, figs. 143-149. Alayon (CGA), the Instituto de Ecologia y DISTRIBUTION: Found from New York Sistematica, Academia de Ciencias de Cuba south to Guatemala; Cuban records include (IES, courtesy of L. F. de Armas), the Mu- the following provinces: Cienfuegos (Sole- 1993 ALAYON G. AND PLATNICK: CUBAN GROUND SPIDERS 3 dad), La Habana (San Antonio de los Banios), Pinar del Rio (Valle de San Juan; Playa Gal- afre, San Juan y Martinez), and Villa Clare (Santa Clara). NATURAL HISTORY: In Cuba, found under stones in humid, semiopen areas. Microsa cubitas, new species Map 1. Cuba, showing records ofGnaphosa sericata (closed circles), Microsa cubitas (cross), Cu- Figures 1, 2; Map 1 banopyllus inconspicuus (open circles), and Lito- TYPE: Male holotype from Loma de la Car- pyllus cubanus (triangles). idad, Sierra de Cubitas, Camaguey province, Cuba (June 15, 1984; L. F. de Armas), deposited in MNHNC. and interdistances: AME 0.06, ALE 0.06, ETYMOLOGY: The specific name is a noun PME 0.07, PLE 0.06; AME-AME 0.05, AME- in apposition taken from the type locality. ALE 0.02, PME-PME 0.06, PME-PLE 0.05, DIAGNOSIS: Males resemble those of Mi- ALE-PLE 0.06; MOQ length 0.20, front width crosa chickeringi Platnick and Shadab but lack 0.18, back width 0.20. Leg spination: femora: the distal prolateral lobe found on the palp I pO-1-1; III rO-O-1; IV pO-O-1; patellae III, of that species (figs. 1, 2; cf. Platnick and IV rO-0-0; tibiae: I v2-2-2; II vlr-2-2; III dO- Shadab, 1977, figs. 4-6). 0-0; IV dO-0-0, pO-1-0; metatarsi: I, II v2-0- MALE: Total length 1.88. Carapace 0.88 0; IV pO-2-2, v2-0-1p. Dorsum of abdomen long, 0.77 wide. Femur II 0.62 long. Eye sizes with transverse white band at about half its and interdistances: AME 0.04, ALE 0.05, length. Embolus long, basally thickened (fig. PME 0.04, PLE 0.06; AME-AME 0.03, AME- 3); retrolateral tibial apophysis constricted at ALE 0.02, PME-PME 0.05, PME-PLE 0.05, tip (fig. 4). ALE-PLE 0.06; MOQ length 0.14, front width NEW REcORD: The newly discovered male 0.1 1, back width 0. 12. Leg spination: femur was taken 5 km S ofYaguajay, Sancti Spiritus IV pO-0-0; tibiae: I vlr-lr-0; II vir-ir-ip; III province, Cuba, in May 1978 (L. F. de Ar- dl-0-0, pO-1-0, vlp-2-2; metatarsi: I vO-0-0; mas, CGA). II vO-lr-0; III pO-O-i, vO-0-2, rO-O-1; IV pO- 0-1, vO-0-2, rl-0-1. Palp with retrolateral pa- Cesonia grisea (Banks) tellar apophysis, without retrolateral tibial Map 2 apophysis, without distal prolateral lobe (figs. Callilepis grisea Banks, 1914: 639, fig. 2 (female 1, 2). holotype from Pinar del Rio, Cuba, in AMNH, FEMALE: Unknown. examined). OTHER MATERLAL EXAMINED: None. Cesonia grisea: Platnick and Shadab, 1980b: 384, DISTRIBUTION: Known only from the type figs, 144, 145. locality. DISTRIBUTION: Endemic to Cuba; recorded from Crocodile (formerly Jacksonville), Isle Cesonia cincta (Banks) of Youth (formerly Isle of Pines), Batey del Figures 3, 4; Map 2 Medio, Meneses, Sancti Spiritus province; La Eilica cincta Banks, 1909: 157, pl. 45, fig. 8 (female Punta, 5 km from La Mula, Guama, Santiago holotype from La Habana, La Habana province, de Cuba province. Cuba, in MCZ, examined). NATURAL HISTORY: Specimens have been Cesonia cincta: Platnick and Shadab, 1980b: 380, taken on bromeliads (Tillandsia sp.) in wet figs. 114, 115. forests. DIAGNOSIS: Males, newly described here, resemble those of C. bryantae Platnick and Cesonia bilineata (Hentz) Shadab but can be distinguished by the lon- Map 2 ger, basally thicker embolus (figs. 3, 4). Herpyllus bilineatus Hentz, 1847: 456, pl. 24, fig. MALE: Total length 3.33. Carapace 1.61 5 (male and female syntypes from North Car- long, 1.71 wide. Femur II 0.96 long. Eye sizes olina and Alabama, destroyed). 4 AMERICAN MUSEUM NOVITATES NO. 3062

Figs. 3, 4. Cesonia cincta (Banks), male palp. 3. Ventral view. 4. Retrolateral view.

Cesonia bilineata: Platnick and Shadab, 1980b: Cesonia irvingi: Platnick and Shadab, 1 980b: 382, 342, figs. 1-7. figs. 1 34-1 39. DISTRIBUTION: Eastern North America, DISTRIBUTION: Known only from southern from Manitoba south to Florida and Cuba Florida, the Bahamas, and Cuba: Cayo Can- (Puerto Manati, Las Tunas province). tiles, Canarreos Archipelago, Isle of Youth NATURAL HISTORY: Cuban specimens have (formerly Isle of Pines). been taken in sandy and marshy places on NATURAL HISTORY: In Cuba this species the border of coastal forests. lives in coastal forest, under stones and bark. Cesonia irvingi (Mello-Leitao) Map 2 Cubanopyllus, new genus Herpyllus australis Fox, 1938: 233, pl. 2, fig. 1 TYPE SPECIES: Litopyllus inconspicuus Bry- (female holotype from Key West, Monroe ant. County, Florida, in USNM, examined). ETYMOLOGY: The generic name is a con- Herpyllus irvingi Mello-Leitio, 1944: 4 (new name for Herpyllus australis Fox, believed to be pre- traction ofCuban Litopyllus and is masculine occupied in Herpyllus by Drassus australis in gender. Holmberg). DIAGNOSIS: Cubanopyllus can be recog- nized most easily by genitalic characters: the rounded spermathecae and enlarged median ducts of females, and palpi with the retrolateral tibial apophysis distally swollen, the median apophysis sinuous and folded at the base of the embolus, and the embolus short, thick, and distally acute. Specimens can be distinguished from other North American gnaphosid genera as follows: from Gnaphosa, Map 2. Cuba, showing records ofCesonia cincta Callilepis, and Eilica by lacking a retromar- (closed circles), C. grisea (cross), C. bilineata (open ginal cheliceral keel or lamina; from Dras- circle), and C. irvingi (square). sodes by lacking deeply notched trochanters; 1 993 ALAYON G. AND PLATNICK: CUBAN GROUND SPIDERS 5 from Scotophaeus by the short embolus; from gynum with anterior margin and plate; sper- Drassyllus, Zelotes, Trachyzelotes, Urozel- mathecae rounded. otes, and Camillina by the absence ofa meta- tarsal preening comb; from Scopodes by hav- Cubanopyllus inconspicuus (Bryant), ing the posterior eye row slightly procurved; new combination from Nodocion, Haplodrassus, Orodrassus, Figures 5-8; Map 1 Herpyllus, Scotophaeus, Rachodrassus, Sco- Litopyllus inconspicuus Bryant, 1940: 393, fig. 176 podes, and Litopyllus by the rounded sper- (female holotype from Soledad, Cienfuegos mathecae; and from Sosticus by lacking an province, Cuba, in MCZ, examined). epigynal scape. DESCRIPTION: Total length 2.8-4.1. Cara- DIAGNOSIS: With the characters of the ge- pace oval in dorsal view, truncated anteriorly nus and genitalia as in figures 5-8. and posteriorly, widest between coxa II and MALE: Total length 2.86. Carapace 1.38 III, slightly elevated in area ofthoracic groove, long, 1.00 wide. Femur II 0.91 long. Eye sizes yellowish beige with some filiform setae on and interdistances: AME 0.07, ALE 0.07, margins, setae longer in ocular area. Thoracic PME 0.08, PLE 0.09; AME-AME 0.03, AME- groove longitudinal, straight. From front, an- ALE 0.01, PME-PME 0.03, PME-PLE 0.03, terior eye row strongly procurved. AME cir- ALE-PLE 0.05; MOQ length 0.23, front width cular, dark; other eyes oval, light. AME and 0.17, back width 0.19. Leg spination typical ALE subequal, largest; PME smallest. AME for genus. Palp as in figures 5, 6. separated by their radius, almost touching FEMALE: Described by Bryant (1940). ALE; PME separated by their diameter, from MATERIAL EXAMINED: The holotype, plus PLE by less than their radius; MOQ roughly one male taken at Puerto Manati, Las Tunas square. Clypeal height equal to or slightly province, Dec. 1981 (L. F. de Armas, IES) greater than AME radius. Cheliceral pro- and one female taken at Cuabales de San Fco. margin with two tiny teeth and one denticle, Matanzas, May 5, 1984 (L. F. de Armas, IES). retromargin with one denticle. Mouthparts DISTRIBUTION: Known from Cienfuegos pale yellowish orange; endites slightly con- and Las Tunas provinces (map 1). verging distally, obliquely depressed, with NOTE: The type locality is in Cienfuegos weak scopula; labium subquadrate, anterior province, not in Oriente as indicated by Plat- margin rounded; sternum broad in middle nick and Shadab (1 980a). The eastern locality with stiff setae at margins, rebordered, with with the same name was never visited by past slight extensions between coxae. Leg formula local or foreign collectors (P. Alayo, personal 4123. Typical leg spination pattern (only sur- commun.). In early years, North American faces bearing spines listed): femora: I dl-1- collectors often visited Soledad (a site well 1, pO-0-1; II dl-1-1, pO-1-1; III, IV dl-l-1, known because of the presence of the Atkins p0- 1 -1, rO- 1-1; patella III p0- 1-0, rO- 1-1; tib- Garden and the former laboratories of the iae: I vlr-Ir-1p; II vlr-2-lp; III dl-0-0, p1- MCZ). 1-1, v2-2-2, rO-1-1; IV pl-1-i, v2-2-2, rl-l- NATURAL HISTORY: Specimens have been 1; metatarsi: I, II v2-0-0; III p1-2-2, v2-2-0, collected under stones in coastal areas. rl-1-2; IV pl-2-2, v2-2-0, rl-2-2. Tarsi and metatarsi with sparse scopulae, more devel- Litopyllus cubanus (Bryant) oped in females; tarsi with two dentate claws Map 1 and claw tufts. Trochanters only shallowly Paramyrmecion cubanum Bryant, 1940: 395, figs. notched. Distal leg segments with dorsal tri- 173, 175 (female holotype from Soledad, Cien- chobothria. Metatarsi III with preening brush. fuegos province, Cuba, in MCZ, examined). Abdomen white with yellow anterior scutum Litopyllus cubanus: Platnick and Shadab, 1980a: in males, grayish white in females. Six spin- 21, figs. 36-39. nerets, anterior laterals long, with three large DISTRIBUTION: Known only from Florida, piriform gland spigots. Palp with short, sin- Bimini, and Cuba (city ofLa Habana and the uous embolus, folded median apophysis, and type locality). membranous conductor. Retrolateral tibial NATURAL HISTORY: Specimens have been apophysis long, slightly swollen distally. Epi- taken under stones in semiopen places (bor- 6 AMERICAN MUSEUM NOVITATES NO. 3062

Figs. 5-8. Cubanopyllus inconspicuus (Bryant). 5. Male palp, ventral view. 6. Same, retrolateral view. 7. Epigynum, ventral view. 8. Same, dorsal view. ders of woods), and in the nests of Rattus has been found under stones in coastal sem- norvegicus. ideciduous forest.

Sergiolus minutus (Banks) Sergiolus kastoni Platnick and Shadab Map 3 Map 3 Poecilochroa minuta Banks, 1898: 185 (male ho- Sergiolus kastoni Platnick and Shadab, 1981: 13, lotype from Brazos County, Texas, in MCZ, ex- figs. 24-29 (male holotype and female paratype amined). from Gainesville, Alachua County, Florida, in Sergiolus minutus: Platnick and Shadab, 1981: 20, AMNH, examined). figs. 48-53. DISTRIBUTION: Known only from Florida DISTRIBUTION: Eastern United States, Ja- and Cuba (La Morena, Carretera El Socucho, maica, and Cuba (9 km S Pinar del Rio city, Puerto Padre, Las Tunas province). and Soroa, both in Pinar del Rio province). NATURAL HISTORY: In Cuba this species NATURAL HISTORY: In Cuba this species has been found under stones in dry and open has been found under stones in semihumid areas the forest. along coast. Sergiolus cyaneiventris Simon Eilica bicolor Banks Map 3 Map 4 Sergiolus cyaneiventris Simon, 1893: 311 (female Eilica bicolor Banks, 1896: 60 (male holotype from holotype from Florida, in MNHN, examined). Punta Gorda, Charlotte County, Florida, in - Platnick and Shadab, 1981: 24, figs. 60-65. MCZ, examined). - Platnick, 1975: 10, figs. 1, 20-23. DISTRIBUTION: Eastern United States and Caridrassus wheeleri Bryant, 1940: 392, figs. 166, Cuba (Guanahacabibes, Pinar del Rio prov- 167 (female holotype from Cienaga de Zapata, ince). Matanzas province, Cuba, in MCZ, examined). NATURAL HISTORY: In Cuba this species First synonymized by Platnick, 1975: 10. 1993 ALAYON G. AND PLATNICK: CUBAN GROUND SPIDERS 7

Map 3. Cuba, showing records ofSergiolus min- Map 4. Cuba, showing records ofEilica bicolor utus (closed circles), S. cyaneiventris (cross), and (closed circles), Zelotes holguin (open circle), S. kastoni (open circle). Camillina elegans (triangle), Camillina sp. (square), and Urozelotes rusticus (crosses). DISTRIBUTION: California south to Hon- duras, east to Jamaica and Cuba: El Abra, collected at Canalejas, Cayo Romano, Ca- Sierra de Casas, Isle of Youth (formerly Isle maguey province, Cuba. of Pines). NATURAL HISTORY: In Cuba this species Urozelotes rusticus (L. Koch) has been collected under stones in semide- Map 4 ciduous forest. Prosthesima rustica L. Koch, 1872: 309 (female holotype from "Trient," Trento, Trentino-Alto Zelotes holguin Alayon Adige, Italy, in BMNH, examined). Map 4 Urozelotes rusticus: Platnick and Murphy, 1984: 24, figs. 55-58. Zelotes holguin Alay6n, 1992: 2, figs. IA, B (female holotype from Loma de la Cruz, Holguin, DISTRIBUTION: Cosmopolitan (synan- Holguin province, Cuba, in MNHNC, exam- thropic). In Cuba: San Antonio de los Baiios, ined). and Medicine School (in laboratory) at Ma- DISTRIBUTION: Known only from the type rianao, both in La Habana province. locality. NATURAL HISTORY: In Cuba this species NATURAL HISTORY: The holotype was taken has been taken only in houses and buildings. under a stone in a replanted pine forest. DISCUSSION Camillina elegans (Bryant) The spider family Gnaphosidae is not well Map 4 represented in Cuban collections, despite all Eilicina elegans Bryant, 1940: 391, figs. 165, 169 the collecting done during this century. Prior (female holotype from Maisi, Guantanamo to this contribution only nine species had been province, Cuba, in MCZ, examined). reported, distributed primarily in the western Camillina elegans: Platnick and Shadab, 1982: 4, and southcentral parts of the main island, figs. 1-4. with few records from eastern provinces. DISTRIBUTION: Florida to Cura9ao; Ha- Hence our collecting trips over the last two waii; Marshall Islands; Papua New Guinea; decades concentrated on the eastern portion Angola. In Cuba the species is known only of the Cuban archipelago. Nevertheless, the from the holotype. total number of species is not high. In this NATURAL HISTORY: In other areas, the spe- paper we report seven additional species for cies has been collected in sandy places, hu- a total of 16, six of which appear to be en- man dwellings, various crop fields, on the demics: Cesonia cincta (Banks), Cesonia gri- shores of ponds, in cleared woods, and in sea (Banks), Cubanopyllus inconspicuus (Bry- herbaceous strata at the edge of forests. ant), Microsa cubitas new species, Camillina sp., and Zelotes holguin Alayon. Camillina sp. It is probable that areas such as the Sierra Map 4 de Guaniguanico (Pinar del Rio province), Cienaga de Zapata (Matanzas province), Si- NOTE: A description of this new species is erra de Guamuhaya (Sancti Spiritus prov- forthcoming (Alayon, in press). The type was ince), Sierra Maestra (Santiago de Cuba prov- 8 AMERICAN MUSEUM NOVITATES NO. 3062

groups (Platnick and Shadab, 1980b), two of which are present in Cuba: the elegans group with C. cincta, C. grisea, and C. irvingi, and the bilineata group with C. bilineata. The genus has 30 species, mostly Nearctic in distribution but with some Neotropical and An- tillean elements. In Cuba C. cincta appears Map 5. Cuba, showing some of the principal to be the most widespread species (see map areas of endemism: Sierra de Guaniguanico (1), 2). Cienaga de Zapata (2), Sierra de Guamuhaya (3), Sergiolus is the second best represented ge- Sierra Maestra (4), Cuchillas del Toa (5), and the nus, with three species: S. minutus, S. kas- south coast of Guantanamo province (6). toni, and S. cyaneiventris. The genus is found in temperate North America, with one spe- ince), Cuchillas del Toa (Holguin and cies extending into northern Canada, five into Guantanamo provinces), and the south coast Mexico, one into the West Indies, and one of Guantanamo province (see map 5) will be each endemic to the islands of Guadalupe found to contain additional new species or and Hispaniola. In Cuba S. minutus is the records of this spider family, because they most widespread species (see map 3). are zones ofhigh endemism for other animals The Cuban fauna ofCamillina includes two and plants. species, C. elegans and a new species; the first Only three genera are represented by more has only an isolated record from the eastern than a single species. Cesonia is the most tip of Cuba (see map 4), the second in the speciose, with four species represented. The northeastern part. It is likely that additional genus has been divided into four species species of this genus occur in Cuba.

REFERENCES Alay6n G., G. Hentz, N. M. 1992. Nueva especie de Zelotes (Araneae: 1847. Descriptions and figures ofthe araneids Gnaphosidae) de Cuba. Poeyana 422: ofthe United States. J. Boston Soc. Nat. 1-4. Hist. 5: 443-478. In press. Nueva especie de Camillina (Araneae: Koch, L. Gnaphosidae) de Cuba. Poeyana. 1866. Die Arachniden-Familie der Drassiden. Banks, N. Nurnberg, pts. 1-6, pp. 1-304. 1896. New North American spiders and mites. 1872. Beitrag zur Kenntniss der Arachniden- Trans. Am. Entomol. Soc. 23: 57-77. fauna Tirols. Zeits. Ferdinandeums Ti- 1898. Some new spiders. Can. Entomol. 30: rol Vorarlberg, ser. 3, 17: 239-328. 185-188. Mello-Leitao, C. F. de 1909. Arachnida of Cuba. Estac. Centr. Agr. 1944. Descripcion de Barbiellinia hirsuta g.n., Cuba, 2nd Rep., pt. 2, pp. 150-174. sp.n. y notas de nomenclatura aracnol- 1914. New West Indian spiders. Bull. Am. ogica. Com. Zool. Mus. Hist. Nat. Mon- Mus. Nat. Hist. 33: 639-642. tevideo 1(21): 1-4. Bryant, E. B. Platnick, N. I. 1940. Cuban spiders in the Museum of Com- 1975. A revision of the spider genus Eilica parative Zoology. Bull. Mus. Comp. (Araneae, Gnaphosidae). Am. Mus. Zool. 86: 247-554. Novitates 2578: 19 pp. Fox, I. Platnick, N. I., and J. A. Murphy 1938. Notes on North American spiders ofthe 1984. A revision of the spider genera Trachy- families Gnaphosidae, Anyphaenidae, zelotes and Urozelotes (Araneae, Gna- and Clubionidae. Iowa State College J. phosidae). Am. Mus. Novitates 2792: Sci. 38: 227-243. 30 pp. Franganillo B., R. P. Platnick, N. I., and M. U. Shadab 1926. Aracnidos nuevos o poco conocidos de 1975. A revision ofthe spider genus Gnaphosa la Isla de Cuba. Bol. Soc. Entomol. Es- (Araneae, Gnaphosidae) in America. paiia 9: 42-68. Bull. Am. Mus. Nat. Hist. 155: 1-66. 1993 ALAYON G. AND PLATNICK: CUBAN GROUND SPIDERS 9

1977. A new genus of the spider subfamily 1981. A revision ofthe spider genus Sergiolus Gnaphosinae from the Virgin Islands (Araneae, Gnaphosidae). Am. Mus. (Araneae, Gnaphosidae). J. Arachnol. 3: Novitates 2717: 41 pp. 191-194. 1982. A revision of the American spiders of 1980a. A revision of the North American spi- the genus Camillina (Araneae, Gna- der genera Nodocion, Litopyllus, and phosidae). Ibid., 2748: 38 pp. Synaphosus (Araneae, Gnaphosidae). Simon, E. Am. Mus. Novitates 2691: 26 pp. 1893. Descriptions d'especes et de genres nou- 1980b. A revision of the spider genus Cesonia veaux de l'ordre des Araneae. Ann. Soc. (Araneae, Gnaphosidae). Bull. Am. Mus. Entomol. France 62: 299-330. Nat. Hist. 165: 337-385.

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