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CLAUDE W. HIBBARD MEMORIAL VOLUMES (1 -5) Index and Bibliography of Late Cenozoic Freshwater Mollusca of.Western North America Dwight W. Taylor The Fossil Catfishes of North America John G. Lundberg Studies on Cenozoic Paleontology and Stratigraphy (Papers Presented at Hibbard Memorial Symposium, May, 1974, in Ann Arbor) Geology and Paleontology of the Early Pleistocene (Late Blancan) White Rock Fauna from North-Central Kansas Ralph E. Eshelman Fishes of the Pliocene Glenns Ferry Formation, Southwest Idaho Geraid R. Smith and Fishes of the Miocene-Pliocene Deer Butte Formation, Southeast Oregon Peter G. Kimmel Museum of Paleontology The University of Michigan Ann Arbor, Michigan 48 104 RSHES OF THE PLIOCENE GLENNS FERRY FQP.MAT!OH. SOUTHWEST IDAHO Gerald R. Smith University of Michigan Museum of Paleontology and Division of Fishes, Museum of Zoology FISHES OF THE MIOCENE-PLIOCENE DEER BUTTE FORMATION, SOUTHEAST OREGON Peter G. Kimmel University of Michigan Museum of Paleontology PAPERS ON PALEONTOLOGY NO. 14 1935 Claude W. Nibbard Memorial Volume 5 Claude W. Hibbard Memorial Volume 5 Papers on Paleontology No. 14 The Museum of Paleontology University of Michigan Ann Arbor, Michigan 48 104 Gerald R. Smith, Director Edited by N.E.Friedland APPRECIATION AND ACKNOWLEDGMENT IS EXPRESSED TO THE 1962 CLASS OF THE UNIVERSITY OF MICHIGAN FOR ESTABLISHING AN INSTITUTE OF SCIENCE AND TECHNOLOGY PUBLISHING FUND. THIS FUND AND A CONTRIBUTION BY DWIGHT W. TAYLOR HAVE AIDED IN THE PUBLICATION OF THIS WORK. December 1.1975 CONTENTS Fishes of the Pliocene Glenns Ferry Formation. Southwest Idaho: GERALD R . SMITH ABSTRACT ......................................... iv Idadon newgenus .............................. 45 INTRODUCTION ..................................... 1 Idadon condoninnus (Cope) .................... 47 SPECIES ACCOUNTS................................. 5 Idadon hibbardi new species .................... 49 FamilySalmonidae ................................. 5 Genus Gila Baird and Guard ...................... 50 Subfamily Salmoninae............................ 5 Gila milleri new species ........................ 50 Genus Rhabdofario Cope ....................... 5 Genus Richardsonius Girard ....................... 50 Rhabdofario lacustris Cope ................... 7 Richardsonius durranti new species .............. 50 Genus Oncorhynchus Suckley .................. 11 Cyprinidae incertae sedis ......................... 51 Oncorhynchus salax new species .............. 11 Family Ictaluridae ................................. 51 Subfamily Coregoninae .......................... 15 Genus Ictalurus Rafiiesque ....................... 51 Genus Prosopium Milner ....................... 15 Ictalurus vespertinus Miller and Smith ............ 51 Prosopium prolixus new species .............. 17 Ictaluridae incertae sedis ......................... 53 Unidentified Salmonines......................... 18 Family Centrarchidae .............................. 53 Family Catostomidae .............................. 19 Genus Archoplites Gill ........................... 53 Genus Chasmistes Jordan ......................... 19 Archoplites taylori Miller and Smith ..............53 Chasmistes spatulifer Miller and Smith ............ 19 FamilyCottidae .................................. 55 Genus Catostomus Lesueur ....................... 21 Kerocottus new genus ........................... 55 Catostomus shoshonensis Cope ..................29 Kerocottus divaricatus (Cope).................. 55 Catostomus cristatus Cope ..................... 3 1 Kerocottus pontifex (Cope) .................... 59 Catostomus owyhee (Miller and Smith) ........... 33 Kerocottus hypoceras (Cope) ................... 59 Catostomus arenatus Miller and Smith ............33 Genus Myoxocephalus (Steller) Tilesius ..............61 Family Cyprinidae ................................. 35 Myoxocephalus idahoensis new species............ 61 Genus Prychocheilus Agassiz ...................... 35 Myoxocephalus antiquus new species ............. 61 Ptychocheilus arcifems (Cope) .................. 35 Genus Cottus Linnaeus .......................... 62 Genus Acrocheilus Agassiz ........................ 37 Cottus bairdi Girard, species group ............... 62 Acrocheilus Iatus (Cope)....................... 37 Cottidae incertae sedis ........................... 62 GenusOrthodon Girard .......................... 39 Orthodon hadrognathus new species .............. 39 DISCUSSION....................................... 62 Genus Mybcheilus Agassiz ........................ 41 Zoogeography .................................... 62 Mylocheilus robustus (Leidy) ................... 41 Ecology ......................................... 65 Mylocheilus inflexus (Cope) .................... 43 Evolution ....................................... 66 Genus Mylopharodon Ayres ....................... 45 ACKNOWLEDGMENTS ............................... 67 Mylopharodon hagermanensis Uyeno .............45 LITERATURE CITED ................................ 68 Fishes of the Miocene-Pliocene Deer Butte Formation. Southeast Oregon: PETER G . KIMMEL ABSTRACT ........................................ 69 Genus Orthodon Guard .............................81 Orthodotz hadrognathus Smith..................... 82 INTRODUCTION .................................... 69 Orthodon onkognathus new species................. 82 Family Salmonidae ................................... 70 Genus Ptychocheilus Agassiz ......................... 82 Subfamily Salmoninae .............................. 7 1 Ptychocheilus arciferus (Cope)..................... 82 Paleolox new genus ............................. 71 Cyprinidae, incertae sedis ........................... 82 Paleolox larsoni new species .................... 71 Family Ictaluridae ................................... 82 Genus Rhabdofario Cope ......................... 73 Genus Ictalums Rafinesque .......................... 82 Rhabdofario carinaturn new species .............. 73 Ictalurus vespertinus Miller and Smith ............... 82 Salmoninae, incertae sedis ........................ 77 Family Centrarchidae ................................. 83 Subfamily Coregoninae ............................. 79 Genus Archoplites Gill............................. 83 Genus Prosopium Milner ......................... 79 Archoplites taylori Miller and Smith ................ 83 Prosopium prolixus Smith ..................... 79 Family Cottidae ..................................... 83 Family Catostomidae ................................. 79 Genus Cottus Linnaeus ............................. 83 Genus Catostomus Lesueur .......................... 79 Cottus calcatus new species ....................... 83 Genus Chasmistes Jordan ........................... 79 Genus Kerocottus Smith ............................ 85 Chasmistes spatulifer Miller and Smith ...............79 Kerocottus divaricatus (Cope) ..................... 85 Genus Myoxocephalus (Steller) Tilesius ................ 85 FamilyCyp~inidae................................... 79 Myoxocephalus sp .............................. 85 Genus Acrocheilus Agassiz .......................... 79 Cottidae, incertae sedis ............................. 85 Acrocheilus lafus (Cope) ......................... 79 Genus Idadon Smith ............................... 81 DISCUSSION....................................... 85 Idadon condonianus (Cope) .......................81 Depositional Environment ........................... 85 Idadon hibbardi Smith ........................... 81 Ecology ......................................... 86 Genus Mylochellus Agassiz .......................... 81 Evolution ....................................... 86 Mylocheilus inflexus (Cope) ....................... 81 ACKNOWLEDGMENTS ............................... 87 Mylocheilus robustus (Leidy) ...................... 81 LITERATURE CITED ................................87 ABSTRACT ?fie fish fauna of the Glenns Ferry Formation comprises about 30 species of freshwater fishes represented by disartic- ulated fossils that occur in lacustrine and fluviatile sediments along the Snake River Plain between Hagerman and the Oregon border in southwest Idaho. The Glenns Ferry is one of several sedimentary and volcanic formations along the western Snake kver Plain that contain the characteristic Lake Idaho fish fauna (see Kimmel's description of the Deer Butte fishes in this volume). Most of the Glenns Ferry Formation is Pliocene in age; a younger section, containing a different vertebrate fauna, is middle Pleistocene. Lake Idaho refers to a series of more or less continuous lacustrine habitats that existed during late Miocene and Pliocene time in the area, probably under structural control of the system of volcanic activity and caldera collapse now centered in the Yellowstone region. The fauna has been regarded as representative of the Blancan North American land mammal age, but associated K-Ar dates of 4.4-6.2 m.y. in the Glenns Ferry Formation near Hagerman indicate that a reevaluation is necessary. The fishes include representatives of six families: Salrnonidae (trouts, whtefish), Catostomidae (suckers), Cyprinidae (minnows), Ictaluridae (catfish), Centrarchidae (sunfish), and Cottidae (sculpins). The salmonids in the fauna include a large extinct trout, Rhabdofario; a landlocked, planktivorous salmon, Oncorhynchus; two or three species of uncertain affinities; and a large whitefish in the genus Prosopium. Catostomids are abundantly represented by at least one species of Chasmistes and about five species of Catostomus. Cyprinids show the greatest