TULANE STUDIES in GEOLOGY and PALEONTOLOGY Volume 30, Number 2 August 27, 1997

TULANE STUDIES IN GEOLOGY AND PALEONTOLOGY Volume 30, Number 2 August 27, 1997

NEW SPECIES OF MOLLUSCA FROM THE ENTRERRIENSE FORMATION (UPPER MIOCENE} OF CHUBUT PROVINCE ARGENTINA AND SPECIES NOT PREVIOUSLY REPORTED FROM THIS FORMATION PART II - GASTROPODA

RODOLFO F.J . BRUNET PUERTO MADRYN, CHUBUT PROVINCE ARGENTINA

CONTENTS .·· Page I. ABSTRACT ...... 61 II. INTRODUCTION ...... 61 III. ACKNOWLEDGMENTS ...... 61 IV. SYSTEMATIC PALEONTOLOGY ...... 61 V. LITERATURE CITED ...... 97

I.ABSTRACT III. ACKNOWLEDGMENTS The second part of this work contains I wish to thank Tulane University for its the descriptions and systematics of gas help in the publication of this second part, tropods obtained from the upper levels of and all the persons who contributed to its the Entrerriense Formation (Late realization. I am particularly grateful to Miocene), outcropping at various localities Dra. Emily H . Vokes, Tulane University, in the Valdes Peninsula, Chubut Province, for her enormous help in preparation of Argentina. Herein are treated 62 species, the manuscript and the moral support of which 37 are new to science, the that she offered to me. I also acknowledge remainder are either previously described the Centro Nacional Patag6nico (CEN or closely related to described species. PAT), which permitted the use of their Forty-two species are new to the fauna, Servicio Fotografico, as well as the techni which consolidates the affinities with the cal help of Sr. Franciso Pertini. Panamanian, Caribbean, and Neozelandic Bioprovinces. IY. SYSTEMATIC PALEONTOLOGY II. INTRODUCTION Class GASTROPODA This paper is a continuation of a previ Order ARCHAEOGASTROPODA ous study examining the molluscan fauna Superfamily PATELLOIDA of the Entrerriense Formation (Late Family ACMAEIDAE Miocene) of the Valdes Peninsula, Chubut Subfamily PECTINODONTINAE Province, Argentina, and with the material Genus PECTINODONTA Dall, 1882 contained herein we increase the number PECTINDONTA sp. of species of Gastropoda recorded in the Plate 1, figure ·1 formation to a total of 128, of which 83 are Description: Shell coniform, small, curved, newly described. With regard to the his low ·and thin; apex blunt, subcentral, only a bit torical considerations and stratigraphy, anterior. Muscle scar in form of a horseshoe the reader is kindly referred to Part I with aperture anterior. Ornamented with fine (Brunet, 1995). We continue supporting concentric growth lines, in some examples dark the relationship of these strata with the concentric bands and fine radial striae, corre upper levels of Parana, Entre Rios sponding to original coloration of shell. Province, Argentina, reported by Bravard Hypotype: MPEF-Pl-212; length 10 mm, (1858), and consider both synchronous and height 4.1 mm, width 8.1 mm. . corresponding to a paleogeographic event Locality: Puerto Piramide, Valdes Perunsula, not yet elucidated. Chubut Province, Argentina.

61 Tulane Studies in Geology and Paleontology Vol. 30 62

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420 s

P. Bajos

LDES P. Hercules Punta Delgada

0 50 100 150

SN MILCIMSTRDS

Text-figure 1. Map of Argentina, showing location of Valdes Peninsula. Inset map with fossil localities: 1) Puerto Piramide; 2) Punta Pardela; 3) Playa Fracasso; 4) Baliza Punta Flecha. Discussion: We assign this example to shorter ribs arising at middle of summit, the genus Pectinodonta, because of the . extending to margin; strong concentric cords low, arcuate shell, with a large blunt apex. crossing over ribs, giving to them a tuberculate Dall (1882) reports Pectinodonta arcuata aspect. Projection of ribs (long and short) pro from the Recent of the West Indies. This is ducing a scalloped lip. With transmitted light the first record of the genus from the sh~ll showing radiating striae chestnut-yellow Entrerriense Formation of South America. in color, corresponding to original coloration of shell. Family PATELLIDAE Hypotype: MPEF-PI-213; length 20.i mm, Subfamily NACELLINAE height 9.5 mm, width 15.1 mm. Genus NACELLA Schumacher, 1817 Locality: Puerto Piramide, Valdes Peninsula, Subgenus PATINIGERA Dall, 1905 Chubut Province, Argentina. NACELLA (PATINIGERA) aff. N.(P.) DEAURATA (Gmelin, 1791) Discussion: The example described pre Plate 1, figure 2 sents conchological characters similar to Description: Shell relatively thin, oval, slight the living species N.(P.) deaurata, the car ly elevated. Apex situated on anterior third. acters differentiating them is the presence Strong radial ribs initially thin at apical level, of short intercalated ribs between the rapidly becoming wider near margin, other longer ones and a strong concentric orna- No.2 Entrerriense Gastropoda 63

mentation. The Recent form lives from rounded, curving gradually toward base; lack Latitude 38° South, to the Magellanic ing umbilicus. region and the Kerguelen Islands. We Hypotypes: MPEF-Pl-215; height 11.5 mm, record this genus and subgenus for the width 11.5 mm (female example). MPEF-Pl-216; first time from the Entrerriense height 7 mm, width 4 mm (male example). Formation of South America. MPEF-Pl-217; height 3.8 mm, width 2.2 mm. Locality: Puerto Piramide, Valdes Peninsula, Superfamily TROCHACEA Chubut Province, Argentina. Family TROCHIDAE Subfamily MONODONTINAE Discussion: From the Entrerriense Genus TEGULA Lesson, 1835 Formation two species have been TEGULA ORBIGNYANA (Pilsbry, 1900) described as Littorina unicostalis Plate 1, figure 3 Borchert, 1901, and Littorina paranensis Description: Shell trocoid, with a high spire; Borchert, 1901, both from Parana five very convex, rounded whorls. Protoconch (Argentina); the first also has been report smooth; sutures very marked; with deep umbili ed from the Camacho Formation of cus. Peristome rounded, thin; columella with Uruguay (Figueiras and Broggi, 1971-73). three teeth, in some examples only two, the The specimen figured here (pl. 1, fig. 4; third disappearing. Aperture lacking internal MPEF-PI-215) possibly represents a denticles. Ornamented with tuberculated spiral female example, in which case the other striae, 12 to 13 in number; base with ten to 11 example (pl. 1, fig. 5; ·MPEF-PI-216) smooth striae, in some examples tuberculated. should be a male, ornamented with fine Hypotype: MPEF-Pl-214; height 7.1 mm, and coarse spiral cords. We have also width 10 mm. observed other examples (pl. 1, fig. 6; Locality: Punta Norte, Valdes Peninsula, MPEF-PI-217) with a very conical form Chubut Province, Argentina. and conspicuous peripheral carina, which resemble those from the area of Discussion: Our examples show no dif Governor's Harbour, Eleuthra, Bahama ferences from specimens .living from Mar Islands, figured by Bequaert (1943, pl. 5, del Plata to Santa Cruz (Argentina). They fig. 5). probably should be placed in the subgenus Agathistoma Olsson and Harbison, 1953. Tegula patagonica (d'Orbigny, 1840), LITTORINA SAXATILIS (Olivi, 1792) which has been cited from the Entrer Plate 1, figure 7 riense Formation, differs by having denti Description: Shell turbinate, variable in cles on the inner margin of the lip. shape, but higher than wide, heavy; three or four whorls increasing rapidly, the last two Superfamily LITTORINOIDEA thirds of total height, whorls convex; suture Family LITTORINIDAE deep; without visible ornamentation. Aperture Subfamily LITTORININAE semicircular, outer lip thin, smooth. Columellar Genus LITTORINA Ferussac, 1822 area wide, heavy, smooth, expanding toward LITTORINA aff. L. ZICZAC (Gmelin, 1791) basal margin and toward body-whorl. A small Plate 1, figures 4-6 umbilicus beside columellar margin (juvenile Description: Shell small, conical, almost as example). Apparently color in this species vari high as wide, moderately heavy; five o~ six able, with yellow, orange, grayish-orange, or rather flat :whorls (specimens very corroded); grayish-maroon in adults, with one or more spi suture well marked and smooth. Body-whorl ral bands, white, gray-black, gray, or dark pur half of total height, very convex, with an obtuse ple. Our specimen with uniform yellowish col peripheral carina. In some apparently gerontic oration, with dark purple band in middle of examples a discontinuous subsutural carina penultimate whorl and another subsutural (Bequaert, 1943). First two whorls smooth, band on body-whorl, coloration considered to be sculpture only visible on base, consisting of fine original color. spiral cords. Aperture pyriform; outer lip thin, Hypotype: MPEF-Pl-218; height 5 mm, width united directly to body-whorl. Columellar area 3mm. moderately long and wide, oblique within, Locality: Puerto Piramide, Valdes Peninsula, smooth, at times rather concave; inner lip Chubut Province, Argentina. 64 Tulane Studies in Geology and Paleontology Vol. 30

this species from the Entrerriense Discussion: Littorina saxatilis occurs in Formation of South America, as well as the boreal and arctic areas. In North the southernmost record of the species America it extends from Cape Barrow, presently known. Southampton Id., and Hudson's Bay, to southern New Jersey. It also is found in LITTORINA PIRAMIDESIA, n. sp. Post-Pliocene deposits that represent Plate 1, figure 8 brackish lagoons near Branford, Description: Shell of small size, turriform, Connecticut (Knight, 1934, in Bequaert, heavy, high-spired, whorls convex; body-whorl 1943, p. 10). This is the first record for almost twice height of spire, very globose with

PLATE 1 Figures Page 1. PECTINODONTA sp...... 61 MPEF-PI-212 (hypotype); length 10 mm, height 4.1 mm, width 8.1 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 2. NACELLA (PATINIGERA) aff. N.(P.) DEAURATA (Gmelin, 1791) ...... 62 MPEF-PI-213 (hypotype); length 20.1 mm, height 9.5 mm, width 15.1 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 3. TEGULA ORBIGNYANA (Pilsbry, 1900) ...... 63 MPEF-PI-214 (hypotype); height 7.1 mm, width 10 mm. Locality: Punta Norte, Valdes Peninsula, Chubut Province, Argentina. 4. LITTORINA aff. L. ZICZAC (Gmelin, 1791) (female) ...... 63 MPEF-PI-215 (hypotype); height 11.5 mm, width 11.5 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 5. LITTORINA aff. L. ZICZAC (Gmelin, 1791) (male) ...... 63 MPEF-PI-216 (hypotype); height 7 mm, width 4 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 6. LITTORINA aff. L. ZICZAC (Gmelin, 1791) ...... 63 MPEF~PI-217 (hypotype); height 3.8 mm, width 2.2 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 7. LITTORINA SAXATILIS (Olivi, 1792) ...... 63 MPEF-PI-218 (hypotype); height 5 mm, width 3 mm. ' Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 8. LITTORINA PIRAMIDESIA, n. sp...... 64 MPEF-PI-219 (holotype); height 7.2 mm, width 4.8 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 9. LITTORINANEBULOSATESSELLATAPhilippi, 1847 ...... 66 MPEF-PI-220 (hypotype); height 3.9 mm, width 2.6 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Provjnce, Argentina. 10. VITRINELLA PARDELENSIS, n. sp...... 66 MPEF-PI-222 (holotype); height 2 mm, diameter 5.1 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 11. CIRCULUS PARDELENSIS, n. sp ...... 66 MPEF-PI-223 (holotype); height 1.3 mm, width 3.1 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 12. TEINOSTOMA (AEPYSTOMA) aff. T.(A.) ANDRIUM Woodring, 1957 ...... 67 MPEF-PI-224 (hypotype); height 2.4 mm, diameter 4 mm. Locality: Baliza Punta Flecha, Puerto Madryn, Chubut Province, Argentina. 13. CREPIDULAANCESTRA, n. sp ...... 74 MPEF-PI-234 (holotype); height 21 mm, width 20.3 mm, diameter 6 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 14. CREPIDULAANCESTRA, n. sp...... 74 MPEF-PI-235 (paratype); height 26 mm, width 19.5 mm, diameter 12 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. No.2 Entrerriense Gastropoda 65

PLATE 1 66 Tulane Studies in Geology and Paleontology Vol. 30

heavy peripheral carina. Aperture semicircular; VITRINELLA PARDELENSIS, n. Sp. outer lip heavy, very curved; columellar area Plate 1, figure 10 consisting of a heavy callus folded back over Description: Shell of small size, porcelaneous, body-whorl, beginning within aperture and compressed, smooth, widely umbilicate, diame forming angle with basal lip, which also is wide. ter of shell 5.1 times diameter of umbilicus. Base convex; umbilicus not covered by columel Three and one-half convex whorls, the last infe. lar callus. Although appearing smooth, under riorly flat. Aperture strongly oblique, basal out microscope (X 40) fine spiral striae present. line subrectangular due to superior extension of Holotype: MPEF-PI-219; height 7.2 mm, peristome, inferior retraction and parallel width 4.8 mm. nature of the columellar and outer margins. Type locality: Puerto Piramide, Valdes Columellar margin folded back over whorl, Peninsula, Chuput Province, Argentina. extending to umbilicus, but not obscuring it. Derivatio nominis: Named for the type locali Ornamented with heavy growth lines. ty. Holotype: MPEF-Pl-222; height 2 = · diam· eter 5.1 mm, diameter of umbilicus 1 mm. Discussion: Littorina piramidesia, n . Type locality:. Punta Pardela, Valdes sp., is characterized and differentiated Peninsula, Chubut Province, Argentina. from other related species by the develop Derivatio nominis: Named for the type locali· ment of the columellar callus, by the very ty. convex whorls, and the constant presence of an umbilicus in juvenile and adult spec Discussion: Vitrinella pardelensis, n. sp., imens. In apparently female examples, is rather similar to V. fioridiana Pilsbry there are microscopic spiral striae, while and McGinty, 1946, which differs in hav in male examples the spire is shorter and ing a proportionally larger umbilicus, an the ornamentation consists of fine spiral aperture that is oblique but not typically cords. "teinostomiform" as in V. pardelensis. The columellar margin does not cover the LITTORINA NEBULOSA TESSELLATA basal whorl and the the aspect of the aper Philippi, 1847 ture laterally is rounded and basally sub Plate 1, figure 9 triangular. Vitrinella helicoidea C.B. Description: Shell very small; spire conical, Adams, 1850, differs in having a parietal low, but pointed; last whorl very globose, with division of the lip into which penetrates out peripheral carina, double or more than the penultimate whorl, heavy periumbili height of spire. Suture very marked, shoulder of cal carina and the upper margin of the whorls obtusely anguled by depth of suture. peristome is curved forward; the diameter Aperture subcircular, outer lip thin. Columellar of the shell is 3.3 times the diameter of the area long, wide and smooth, inner lip thick umbilicus. Vitrinella praecox Pilsbry and ened, almost straight; ending abruptly below, McGinty, , 1946, differs by having a peri separated by a slight depression in the outer umbilical carina, a peristome oblique but lip. Neither umbilicus nor columellar depres not "teinostomiform" and the diameter of sion. the shell is 3. 75 times the diameter of the Hypotype: MPEF-Pl-220; height 3.9 mm, umbilicus. We note for the first time this width 2.6 mm. genus from the Entrerriense Formation of Locality: Puerto Piramide, Valdes Peninsula, South America. Chubut Province, Argentina. Genus CIRCULUS Jeffreys, 1865 Discussion: We assign this example to CIRCULUS PARDELENSIS, n. sp. the variety tessellata because of the angu Plate 1, figure 11 lation of the shoulder and the very globose Description: Shell very small, heavy, spire body-whorl. This is the first record from moderately low, exposed, peripherially rounded. the Entrerriense Formation of South America. Suture weakly marked, with a wide parasutur al groove beginning at protoconch and having Family VITRINELLIDAE six to eight poorly developed fine spiral striae. Genus VITRINELLA C.B. Adams, 1850 Protoconch smooth; penultimate whorl orna mented with five heavy spiral cords and ulti- No.2 Entrerriense Gastropoda 67 mate with 12 heavy cords. Aperture oblique, what elevated, last whorl very large. subquadrate, with extended upper lip, the Protoconch smooth. Peristome typical, with lower and upper almost straight, the outer upper margin projecting forward and lower barely convex. Umbilicus very deep, with strong withdrawn. Umbilicus open, on the basis of an periurobilical growth lines only. immature example, with slight development of Holotype: MPEF-Pl-223; height 1.3 mm, a callus. Sculture of spiral cords crossed by width 3.1 mm. radiating cords, creating a cancellate ornamen Type locality: Punta Pardela, Valdes tation over entire surface. Peninsula, Chubut Province, Argentina. Hypotype: MPEF-PI-224; height 2.4 mm, Derivatio nominis: Named for the type locali diameter 4 )llill. ty. Locality: Baliza Punta Flecha, Puerto Madryn, Chubut Province, Argentina. Discussion: Relatively near living forms are known from the Atlantic and the Discussion: This form presents charac~ Pacific. Circulus trilix (Bush, 1897) ranges ters very similar to the holotype of from Cape Hatteras to Cuba, and is distin Woodring (1957, p. 70, pl. 1~, figs. 9-11), guished by being tricarinate and lacking which also is an immature example. The the parasutural groove. Woodring (1957) dimensions of the holotype are: height 2 cites this species from the Chipola (Early mm, diameter 4. 7 mm. This is the first Miocene), Shoal River (Middle Miocene), record for the genus Teinostoma and the and Gatun (Late Miocene) formations; also subgenus Aepystoma from the Entrerri with tricarinate forms. Circulus cerrosen ense Formation of South America. sis Bartsch, 1907, from Santa Catalina Island, California, Baja California, and Superfamily CERITHIOIDEA the Gulf of California, may be considered Family TURRITELLIDAE the Pacific homologue of C. trilix. Genus TURRITELLA Lamarck, 1799 Cyclostremiscus glyptomphalus Pilsbry TURRITELLA HEIMI, n. sp. and Olsson, 1952, from the Pleistocene of Plate 2, figure 1 Panama, may be distinguished with diffi Description: Shell ttirriculate, of medium culty from C. trilix; therefore, those size. Six and one-half teleoconch whorls, with authors suggested the existence of a "race" two smooth nepionic whorls. Suture deep and or a subspecies. Circulus cosmius Bartsch, shoulder very marked. Whorls flat. First teleo 1907, from Ecuador is rather near C. cer conch whorl with foµr cords, the remaining roensis, but is characterized by a narrow with five, consisting of one strong anterior and curvature of the upper margin of the peri one strong posterior, with three weaker inter stome, directed downward, extending for mediate cords, between each of these a very fine ward, lacking the parasutural groove. secondary cord; behind the posterior and bor Circulus occidentalis Pilsbry and Olsson, dering the shoulder one fine cord and, in 1941, from the Pliocene of Ecuador, has advance of the anterior, another fine cord. the margin of the peristome directed Suture with a secondary cord ~hove and anoth below, lacks the parasutural groove and er below. In adult examples, upper and infrasu resembles C. cosmius. Circulus philippii tural cords disappearing and on last whorls two (Cantraine, 1842) also has a deep umbili or three very fine cords between primaries. cus, is ornamented with a much greater Heavy growth lines causing some cords to number of spiral cords and lacks the para appear tuberculate, especially the primaries. sutural groove. Base with a heavy peripheral cord and four low flat cords, between these one or two fine sec Genus TEINOSTOMA H. Adams and ondary cords. Aperture apparently oval, with a A. Adams, 1854 thin columellar border, columella convex. Subgenus AEPYSTOMA Woodring, 1957 Growth lines typical of T. conspicabilis TEINOSTOMA (AEPYSTOMA) aff. T.(A.) (Miocene of Australia) and of the T. hybrida ANDRIUM Woodring, 1957 group (Cretaceous-Eocene of Europe), according Plate 1, figure 12 to Guillaume (1924). Description: Shell of small size, heavy, con Holotype: MPEF-Pl-225; height 18.1 mm, vex, with three and one-half whorls, spire some- width 5.8 mm, apical angle 16°. 68 Tulane Studies in Geology and Paleontology Vol. 30

Type local.ity: Puerto Piramide, Vaides Turritella iheringia Figueiras and Broggi, Peninsula, Chubut Province, Argentina. 1976, has seven cords per whorl, with the Derivatio nominis: Dedicated to the late geol upper strongest and forming the shoulder; ogist Dr. Arnoldo Heim. the base has five flat cords and the suture is within a deep groove. Turritella indeter-. Discussion: Turritella heimi, n. sp., dif minata Borchet, 1901, has convex whorls, fers from T. americana (Bravard, 1858), the suture within a deep groove, with nine which has only four primary cords per cords and finer secondaries. Turritella whorl and one irregular fine cord on the heimi, n. sp., has five principal cords, the early whorls, while on the last whorl there anterior and the posterior stronger, and may be intercalated up to two intermedi an intermediary secondary cord; the ate cords; all cords with granulations, over suture is within a deep groove, with one the. early teleoconch whorls. Turritella suprasutural cord and one infrasutural iheringiana Figueiras and Broggi, 1976, cord. differs in having each whorl with seven subequal cords; except the upper, which is TuRRITELLA FIGUEIRAS!, n. sp. stronger and peripheral to the shoulder. Plate 2, figure 3 The sutural area lacks cords and the base Description: Shell of medium size, whorls has five flat cords.· Turritella indetermina flat, suture within a deep groove, shoulder con ta Borchert, 1901, differs in having convex spicuous. Ornamented with three very heavy whorls, with nine cords, and with finer cords, barely granulose, and one cord of equal secondary cords. strength in front of anterior suture; in front of thls, three fine secondary cords, situated within TuRRITELLA BROGGII, n. sp. suture; between anterior suture and cord situ Plate 2, figure 2 ated in advance of it, two fine secondary cords; Description: Shell turriform, of medium size, between anterior suture and the middle, one whorls flat, suture inconspicuous, without fine low central cord and four very fine cords on either sutural depression or shoulder. Twelve each side; between the middle and posterior and one-half whorls, two smooth nepionic suture, one fine central cord and three very fine whorls; first teleoconch whorls with three cords on either side; above posterior suture, on smooth, irregular cords, the remaining with shoulder, three or four fine cords, with some of three primary cords, somewhat heavy, equal these heavier than others. Base with one heavy and granulose; on last whorl one anterior cord peripheral cord and four flat cords, with three added, also granulose and corresponding to to six extremely fine secondary cords between periphero-basal cord; between each pair of prin these. Columella straight and aperture sub cipal cords one fine, low secondary cord with quadrate. No growth lines seen on any speci small granules. Base with a heavy peripheral mens. cord and.five flatter cords, with one secondary Holotype: MPEF-PI-227; height 24 mm; cord between each of them. Columella straight width 7.9 mm, apical angle 15°. and aperture subquadrate; no growth lines seen Type locality: Puerto Piramide, Valdes on any specimens. Peninsula, Chubut Province, Argentina. Holotype: MPEF-PI-226; height 20 mm, Derivatio nominis: Dedicated to the late pale width 6 mm, apical angle 19°. ontologist Dr. Alfredo Figueiras. Type locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. Discussion: Turritella figueirasi, n. sp., Derivatio nominis: Dedicated to the malacolo differs from T. americana (Bravard, 1858), gist Sr. Jorge Broggi. which has four granulose cords with two intermediary cords intercalated. Turritella Discussion: Turritella broggii, n. sp., dif indeterminata Borchert, 1901, has strong fers from T. americana (Bravard, 1858), ly·convex whorls, nine. cords on each which has four granulose primary cords whorl, which increase in number on the per whorl and an inconsistant fine cord on body-whorl, with one secondary cord inter the first whorls, while on the last whorl calated between the primaries. We have there may be intercalated up to two inter no information on the aperture of this mediate cords, also with granulations. species: Turritella iheringiana Figueiras No.2 Entrerriense Gastropoda 69 and Broggi, 1976, has seven subequal spi figures of Woodring (1957, pl. 83, figs. 4, 5, ral cords, with the uppermost strongest; 9, 14) there are two or three heavy cords the base with five ·cords, and an apical located on the anterior half of the whorls. angle of 20°. Turritella broggii, n. sp., has Turritella gatunesis rhytodes Woodring, three primary cords, without a suprasu 1957, has the last whorls very compressed tural cord, a distinct number of secondary posteriorly, without a well-defined shoul cords, the base with five primary cords, a der, approximately 16 to 18 cords covering single secondary cord between these; lack the entire whorl, from suture to suture, of ing a sutural groove. Turritella heimi, n. these three or four most pronounced. sp., has five primary cords, with one finer Turritella adela Woodring, 1957, has secondary cord between them, one supra approximately nine primary cords of vari sutural cord and another subsutural cord; ous strength and very fine secondary the base with five primary cords and one intermediate cords. Turritella mataru or two fine intermediary cords; the aper cana Hodson, 1926, has numerous cords ture is oval with a ·convex columella. that run from suture to suture, without a well-defined shoulder and with very deep TvRRITELLA FLECHENSIS, n . sp. sutures. Plate 2, figure 4 Description':' Sh.ell slender, rather thin, TvRRITELLA HODSON!, n . sp. whorls convex, suture shallow, shoulder wide, Plate 2, figure 5 very oblique and barely concave or straight. Description: Shell slender, with sutures bare Whorls with one heavy anterior cord and one ly visible; protoconch broken; first preserved posterior on ·margin of shoulder, somewhat whorls with three cords, the posterior very finer; between these four or five finer cords and heavy, the medial of the strength of the posteri between the middle of all cords, three or four or and the anterior somewhat more salient and much finer tertiary threads; in front of primary appressed to suture, at times almost covering anterior cord, one fine cord situated within it. On last whorls posterior cord becoming flat suture. Shoulder with three or four fine cords. tened, the surface becoming concave, carrying Base with one heavy peripheral cord and six in this concavity one fine cord; anteriorly con flatter cords; between each o( these one finer tinuing with median cord, followed by another secondary cord. Growth lines as in T. conspica concavity and finally the conspicuous anterior bilis (Miocene of Australia) and the T. hybrida cord, projecting as a ledge over shallow suture. group (Cretaceous-Eocene of Europe), following Aperture presumably suboval, with columella Guillaume (1924). very concave and outer lip with a depression Holotype: MPEF-Pl-228; height 19 mm, caused by projection of external groove. widtl\ 6 mm, apical angle 16°. Apparently base with one heavy, almost flat, Type locality: Baliza Punta Flecha, Puerto peripheral cord. Madryn, Chubut Province, Argentina. Holotype: MPEF-PI-229; height 29 mm, Deriuatio nominis: Named for the type locali width 9.8 mm. ty. Type locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. Discussion: Turritella fiechensis, n. sp., Derivatio nominis: Dedicated to the late pale differs from T. indeterminata Borchert, ontologist Dr. Floyd Hodson. 1901, which has three spiral cords 'on the upper part of the whorl, these followed by Discussion: Turritella hodsoni, n. sp., is six even stronger, on the last whorl even rather similar to T. montanitensis Hodson, more numerous; aperture not seen in the 1926, from the Oligo-Miocene of holotype. Turritella gatunensis gatunensis Venezuela, especially the paratype illus Conrad, 1857, has two primary cords, one trated by Hodson (1926, pl. 16, fig. 6), in the middle of the whorl and another on which differs from our holotype by having the anterior third or quarter, with numer a submedial cord on the early teleoconch ous secondary cords, and with other whorls, in the form of a smooth 'V", with extremely fine threads, observable under one anterior cord and another posterior a microscope on the last whorl; the base one, markedly granulose, due to the with numerous fine spiral cords. In the growth lines and fine intermediate cords, 70 Tulane Studies in Geology and Paleontology Vol. 30

the secondaries also granulose. Turritella sutural cord. Turritella montanitenesis montanitensis olcotti Hodson, 1926, has saladilloensis Hodson, 1926, has the space cords with a lesser number of granules, between the primary anterior cord and the but more conspicuous, especially on supra- ·primary posterior cord more concave, the

PLATE2 Figures Page 1. TURRITELLA HEIM!, n. sp...... 67 MPEF-PI-225 (holotype); height 18.1 mm, width 5.8 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 2. TURRITELLA BROGGII, n. sp...... 68 MPEF-PI-226 (holotype); height 20 mm, width 6 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 3. TURRITELLA FIGUEIRAS!, n. sp...... 68 MPEF-PI-227 (holotype); height 24 mm, width 7.9 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 4. TURRITELLA FLECHENSIS, n. sp...... 69 MPEF-PI-228 (holotype); height 19 mm, width 6 mm. Locality: Baliza Punta Flecha, Puerto Madryn, Chubut Province, Argentina. 5. TURRITELLA HODSON!, n. sp...... 69 MPEF-PI-229 (holotype); height 29 mm, width 9.8 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 6. TURRITELLAPIRAMIDESIA, n. sp...... 72 MPEF-PI-230 (holotype); height 16 mm, width 6.1 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 7. TURRITELLAALZOLAl, n, sp...... 72 MPEF-PI-231 (holotype); height 20.1 mm, width 7 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 8. SPIROCOLPUS FERUGLIOI, n. sp...... 73 MPEF-PI-232 (holotype); height 18.6 mm, width 12 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 9. PETALOCONCHUS sp...... 73 MPEF-PI-233 (hypotype); height 6 mm, width 3.8 mm. Locality: Baliza Punta Flecha, Puerto Madryn, Chubut Province, Argentina. 10. HIPPONIX FLECHENSIS, n. sp...... 73 MPEF-PI-236 (holotype); height 1.8 mm, length 4 mm, width 3.9 mm. Locality: Baliza Punta Flecha, Puerto Madryn, Chubut Province, Argentina. 11. CAPULUS PIRAMIDENSIS, n. sp ...... 74 MPEF-PI-235 (holotype); height 3 mm, length 6 mm, width 6 mm. Loc.ality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 12. POLINICES (EUSPIRA) CONSIMILIS (lhering, 1897) ...... 75 MPEF-PI-237 (hypotype); height 7 mm, width 8.5 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 13. POLINICES (EUSPIRA) ORTMANN! (lhering, 1907) ...... 75 MPEF-PI-238 (hypotype); height 9 mm, width 7.2 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 14. ATAXOCERITHIUM PARDELENSIS, n . sp...... 75 MPEF-PI-239 (holotype); height·9.1 mm, width 4 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 15. ATAXOCERITHIUM PIRAMIDENSIS, n. sp...... 76 MPEF-Pl-240 (holotype); height 18.6 mm,° width 7 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 16. BALCIS FLECHENSIS, n. sp...... : ...... 76 MPEF-PI-241 (holotype); height 6 mm, width 2 mm. Locality: Baliza Punta Flecha, Puerto Madryn, Chubut Province, Argentina. No.2 Entrerriense Gastropoda 71

PLATE2 72 Tulane Studies in Geology andPaleontology Vol. 30

primary posterior cord weaker, the second cords; in the suture there are only two primary cord heavier and the two anterior cords, one suprasutural and the other primaries joined. Observing the figures of infrasutural. On the base there is one Hodson, we have some doubt that the peripheral cord and four additional cords, example figured as a paratype of T. mon with one or two secondaries between tanitensis (Hodson, 1926, pl. 16, fig. 6) is them. the same species, we believe that it is referable to another species of Turritella, TURRITELLA ALZOLAI, n. sp. somewhat similar to T. hodsoni n. sp., but Plate 2, figure 7 distinguished from it by the presence of a Description: Shell turriform, of medium size. surface oblique and straight above the Ten and one-half whorls, with two smooth nepi anterior cord, while that of T. hodsoni is onic whorls. Suture deep, shoulder well-marked completely concave. and oblique, with whorls very convex. First teleoconch whorls with three primary cords and TuRRITELLA PIRAMIDESIA, n. sp. one secondary cord between them, the middle Plate 2, figure 6 primary heavier and projecting; later whorls Description: Shell of medium size, heavy and with five primary cords, intercalated between slender; eight and one-half flat whorls, two these one to three secondary cords. At level of smooth nepionic whorls, suture not visible; stitural groove three or four suprasutural cords, shoulder well marked by a granulose posterior the second of these heavier, and one fine cord cord. Early teleoconch whorls with three spiral situated at crest of shoulder of whorl. Base with cords, the central twice the width of other two six doubled, flat cords. Columella convex, with and the posterior granulose, with one fine cord heavy columellar callus covering one-third of occupying the space of the suture; cords base. Aperture subquadrate. increasing in number up to five, one of these Holotype: MPEF-PI-231; height 20.1 mm, becoming heaviest. In more adult whorls, width 7mm. between principal cords finer smooth secondary Type locality: Puerto Piramide, Valdes cords intercalated, up to five in number, one of Peninsula, Chubut Province, Argentina. these always becoming heavier. Principal cords Derivatio nominis: Dedicated to Dr. Rodolfo following, the posterior and the median, becom Mendez Alzola. ing granulose, but anterior one smooth. Base with one peripheral cord and seven low, convex Discussion: Turritella alzolai, n. sp., dif cords with one fine secondary cord between fers from T. gatunensis gatunesis Conrad, each of these. Columella curved and aperture 1857 (in Woodring, 1957, pl. 23, figs . 4, 5, oval. No growth lines visible. 9, 14), which has the whorls more com Holotype: MPEF-PI-230; height 16 mm, pressed posteriorly, causing the shoulder width 6.1 mm. to be less well defined, with only two pri Type locality: Puerto Piramide, Valdes mary cords and numerous secondary Peninsula, Chubut Province, Argentina. cords, the primaries diminishing in Derivatio nominis: Named for the type locali strength on the last whorls; the base is ty. sculptured with fine spiral cords. Turritella gatunensis rhytodes Woodring, Discussion: Turritella piramidesia, n. 1957, has the .whorls more compressed sp., differs from T. broggii, n. sp:, which posteriorly, with deeper sutures; in this has a less well-marked shoulder, three subspecies one or two of the primary cords equaI granulose cords, with a fourth cord tend to increase in size, thus the last intercalated on the last whorl, and only whorl has five primary cords and numer one fine granulose secondary cord, which ous secondary cords on the shoulder of all is intercalated between the primaries; the of the whorls, which always reach the columella is straight and the aperture suture. Turritella flechensis, n. sp., has subquadrate. Turritella heimi, n. sp., has one heavy anterior cord and one heavy four cords on the early teleoconch whorls posterior one, with three or four fine sec and five primary cords on the last whorls, ondary cords and between these three or with one anterior cord and another heavy four very fine tertiary cords. The shoulder posterior one and three finer intermediate has three or four fine cords and the base No. 2 Entrerriense Gastropoda 73 has six flat cords and one fine intermedi Genus PETALOCONCHUS ate cord. The columella is more convex Lamarck, 1799 and lacks a columellar callus. PETALOCONCHUS sp. Plate 2, figure 9 Genus SPIROCOLPUS Finlay, 1927 Description: Shell with only nepionic whorls SPIROCOLPUS FERUGLIOI, n. sp. and a few teleoconch whorls preserved. Proto Plate 2, figure 8 conch with two or three smooth whorls, contin Description: Shell turriform, very heavy, with uing with flat, smooth spiral bands, seven in sutures poorly marked; shoulder conspicuous, number. Interior totally concealed by lime. straight and rather convex owing to great Hypotype: MPEF-Pl-233; height 6 mm, width development of posterior cord. Ornamented 3.8mm. with one heavy posterior cord at periphery of Locality: Baliza Punta Flecha, Puerto shoulder, with conspicuous granulations, this Madryn, Chubut Province, Argentina. followed by two very :fine cords and three cords even finer than posterior one, also granulose, Discussion: Up to this time, we have with smooth interspaces, equal in width to obtained only two examples of this form. cords , except interspace between middle and Because of the conchological characters, anterior cords, which is twice width of cords; including the .protoconch and part of the one fine and apparently smooth suprasutural teleconch consisting of almost flat bands, cord below and poorly defined fine cords at level apparently without ornamentation, we of surface of shoulder completing ornamenta assign these tentatively to the genus tion. Base and columella lacking, as well as Petaloconchus. In this case, this would be early whorls. No growth lines visible. the first record of this genus from the Holotype: MPEF-Pl-232; height 18.6 mm; Entrerriense Formation of South America. width 12 mm. Type locality: Punta Pardela, Valdes Penin Superfamily HIPPONICOIDEA sula, Chubut Province, Argentina. Family HIPPONICIDAE Deriuatio nominis: Dedicated to Dr. Egidio Genus HIPPONIX Defrance, 1819 Feruglio. · HIPPONIX FLECHENSIS, n. sp. Plate 2, figure 10 Discussion: Spirocolpus feruglioi, n. sp., Description: Shell in form of a cap, small, differs from S. ambulacrum (Sowerby, thin; apex opistogyre, small, pointed, turned to 1846) and from S. ambulacrum pyra the left, not extending beyond shell margin. midesia (lhering, 1907), by having one Aperture suborbicular; lip crenulated by projec heavy crenulated peripheral cord at the tion of ribs. Muscle scar in·form of a horseshoe, shoulder and only three principal cords, open anteriorly; with an internal perilabral with one fine suprasutural cord and, at groove, corresponding to position of fixed disk. the level af the surface of the shoulder, Ornamented with 42 heavy radial ribs (2 per fine spiral cords. Spirocolpus breantiana mm), with interspaces equal in width to ribs; (d'Orbigny, 184 7) is more slender, with other shorter ribs intercalated, beginning at only three heavy cords, of which only the middle of height of shell, extending to peris posterior is generally crenulated. These tome; heavy growth lines giving an almost can increase on the last whorls up to six in cellate aspect. number; the shoulder is much less con Holotype: MPEF-PI-236; height 1.8 mm, spicuous and the suture deeper. length 4 mm, width 3.9 mm. Spirocolpus iheringi (Cossmann, 1898) Type locality: Baliza Punta Flecha, Puerto has only three cords, with the posterior Madryn, Chubut Province, Argentina. stronger, but less conspicuous than jn S. Derivatio nominis: Named for the type locali feruglioi. Evidently this species is closely ty. related to the species of Spirocolpus from Patagonia and to the species from the time Discussion: Hipponix fiechensis, n. sp., period of Eocene-Middle Oligocene in New differs from H. oligocostata Brunet, 1995, Zealand (Marwick, 1957). which has 37 radial ribs, without interme diate ribs. The height/width index is 0.54 Family VERMETIDAE and the widtMength index is 0.85, where- 74 Tulane Studies in Geology and Paleontology Vol. 30

as in H. flechensis these indices are: d'Orbigny, 1847, has the shell very round height/width 0.46 and width/length 0.97. ed and the external surface weakly Hipponix costata Erdemann and Morra, rugose, without conspicuous radial ribs. It 1985, from the Patagonian of Santa Cruz differs from Crepidula unguiformis (Argentina) has more numerous ribs Lamarck, 1822, because in spite of its (approximately 50), without intermediate depressed form and its concave septum, it ribs, and its indices are: height/width 0.23 does not have conspicuous ribs nor a longi and width/length 0.89. Hipponix conicus tudinal carina on the septum. We believe (Shumacher, 1817), living in the western that we may consider it a valid ancestor to Pacific (Fremantle, Australia), has heavier C. dilatata var. patagonica. The high form ribs (1 per mm) with very fine interspaces, resembles C. protea d'Orbigny, 1835, but without intermediate ribs and the length that has a concave septum and a smooth of the shell is 2 cm. surface. Crepidula onyx Sowerby, 1824, has the septum slightly concave and is Superfamily CREPIDULOIDEA smooth exteriorly. From Parana (Entre Family CREPIDULIDAE Rios) Crepidula paranensis Borchert, Genus CREPIDULA Lamarck, 1799 1901, has been described from a unique CREPIDULA ANCESTRA, n. sp. example having an oval form, with a blunt Plate 1, figures 13, 14 apex, which does not ex~end beyond the Description: Shell of medium size, subcircu margin; the septum is broken, preventing lar to oval, low to high. Protoconch curved at an a valid comparison . . Closs (1970) has cited angle of 90°. Apex small, pointed or very strong C. paranensis and C. protea from the and blunt, projecting beyond posterior margin. Middle Miocene of Brazil. Consequently, Septum small or very deep, depending on thick this is the first· record of the genus in the ness of shell, with a medial longitudinal carina Entrerriense Formation of Patagonia. divided into two zones; edge of septum convex, in form of an "S," with a deep notch to the left. Family CAPULIDAE Pedal muscle scar semicircular, appressed to Genus CAPULUS Montfort, 1810 right insertion of septum. Ornamented with CAPULUS PIRAMIDENSIS, n. sp. low, radial ribs, apparently without spines, with Plate 2, figure 11 interspaces equal in width to ribs and with Description: Shell in form of a cap, small but marked growth lines. heavy, apex opistogyre, small, very pointed, not Holotype: MPEF-Pl-234; height 21 mm, extending beyond margin of shell, protoconch width 20.3 mm, diameter 6 mm (low form). clearly visible. Aperture circular, labrum crenu Paratype: MPEF-Pl-235; height 26 mm, lated by projection of ribs. Muscle-scar in form width 19.5 mm, diameter 12 mm (high form). of horseshoe, open anteriorly. External orna Type locality: Puerto Piramide, Valdes Pen mentation consisting of numerous radial ribs (5 insula, Chubut Province, Argentina. per mm), where these are crossed by growth Derivatio -nominis: Denominated thus for lines, rugae developed; interspaces more nar having characters of various species of row than ribs and crossed by fine growth lines; Crepidula. some ribs divided. Holotype: MPEF-Pl-235; height 3 mm, length Discussion: Crepidula ancestra, n. sp., 6 mm, width 6 mm. presents characters common to various Type locality: Puerto Piramide, Valdes Penin living species. We consider both forms the sula, Chubut Province, Argentina. same, due to the characteristics of the sep Derivatio nominis: Named for the type locali tum and the presence of the radial ribs. ty. The low form is rather close to C. dilatata Lamarck, 1822, with its convex septum, Discussion: Capulus piramidensis, _n. but that has a smooth surface. It resem sp., differs from C. ladyae Brunet, 1995, bles C. aculeata (Gmelin, 1791), by the which is lower and less broad, with indices surface covered with ribs and the longitu of height/length: 0.42 and width/length: dinal carina of the septum, but that has 0.83; while those of C. piramidensis are: the margin of the septum concave. height/length 0.50 and width/length 1.00. Crepidula dilatata var. patagonica It has five ribs per mm, whereas C. ladyae No. 2 Entrerriense Gastropoda 75

has three per mm and the apex extends POLINICES (EUSPIRA) ORTMANN! past the labral margin. It is not compara lhering, 1907 ble with C. compressus Smith, 1891, living Plate 2, figure 13 in the Argentine littoral (8.5 x 2.5 x 8 Description: Shell small, somewhat globose, mm), C. incurvatus (Gmelin, 1791), living spire relatively high, with 4 1/2 whorls, includ from North Carolina to Brasil (16 x IO ing nepionic. Suture shallow. Body-whorl occu mm), or with C. hungaricus (Linne, 1767), pying 76% of total height. Aperture semilunar, living from the coast of Iceland to the occupying 71 % of height of body-whorl; inner lip Mediterranean Sea (diameter 5 cm), which almost straight, with conspicuous posterior cal are much larger species. We may deduce lus,the exterior narrow. Umblicus deep, without that in the Entrerriense Sea there arose a funicule. Ornamented with fine growth lines. "race" of Capulus of very small size, in Hypotype: MPEF-Pl-238; height 9 mm, width contrast to those living in the Atlantic 7.2mm. today. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. Superfamily NATICOIDEA Family NATICIDAE Discussion: Polinices (Euspira) ortman Subfamily POLINICINAE ni is relatively common in the outcrops of Genus POLINICES Montfort, 1810 the Valdes Peninsula, we have collected at Subgenus EUSPIRAAgassiz, 1838 various times 38 examples in perfect con POLINICES (EUSPIRA) CONSIMILIS dition, coming from Punta Pardela and (lhering, 1897) Puerto Piramide principally. Natica Pla~e 2, figi.ire 12 ovoidea Philippi (Ortmann, 1900, p. 380) Description: Shell medium-sized, low-spired, is a synonym of Polinices ortmanni the last whorl large and globose, representing Ihering, 1907. four-fifths of total height; suture strongly chan neled, .whorls very convex. Aperture occupying Suborder HETEROGLOSSA three-fourths of body-whorl, semilunar in Superfamily CERITHIOPSOIDEA shape. Columella straight, with an inconspicu Family CERITHIOPSIDAE ous callus and without funicule. -Umbilicus Subfamily CERITHIELLINAE deep; peristome narrow. Ornamented only by Genus ATAXOCERITHIUM Tate, 1893 fine growth lines. ATAXOCERITHIUM PARDELENSIS, n. sp. Hypotype: MPEF-PI-237; height 7 mm, width Plate 2, figure 14 8.5=. Description: Shell small, turriculated and Locality: Puerto ·Piramide, Valdes Peninsula, heavy. Spire with 4 112 whorls and three Chubut Province, Argentina. smooth nepionic whorls. Sutures marked with deep postsutural groove, superior margin of Discussion: Polinices (Euspira) consim teleoconch whorls strongly projecting as a ilis is relatively abundant in the outcrops result of postsutural groove. The last whorls. at the Valdes Peninsula (Chubut). It has ornamented with three spiral cords up to penul been recorded from sediments of Oligocene timate and with four on the ultimate, the sec age at Los Manantiales, La Cuevas, and ond much finer than the others, on both whorls Monte Entrance and from the Patagoni the anterior cord more conspicuous; axial cords ense of Golfo San Jorge. Ihering (1897) from 14 to 17 in 'number, not reaching suture cited Natica consimilis from Monte Leon and forming nodules where they cross spirals. (Sta. Cruz) and, from Parana (Entre Rios), Within suture one fine intrasutural cord. Base Borchert (1901) mentioned Natica entr slightly convex, almost flat, without ornamenta erriana, which we consider a. synonym of tion, except for peripheral basal cord. Columella P.(E.) consimilis. This taxon differs from convex, with evidently siphonal canal short and P.(E.) ortmanni Ihering, 1907, mentioned barely curved. Aperture subquadrate. from Yegua Quemada and Los Manatiales Holotype: MPEF-Pl-239; height 9.1 mm, (Monte Leon Formation, Oligocene), by width4mm. having a more globose form, a lower spire, 'J'Ype locality: Punta Pardela, Valdes Penin a larger aperture, and deeper sutures. sula, Chubut Province, Argentina. Derivatio nominis: Named for the type locali- 76 Tulane Studies in Geology and Paleontology Vol. 30

ty. width 7 mm. Type locality: Puerto Piramide, Valdes Penin Discussion: Ataxocerithium pardelensis, sula, Chubut Province, Argentina. n. sp., differs from A. Juliana (lhering, Derivatio nominis: Named for the type locali 1907), which has more convex whorls, ty. without the typical postsutural depression of A. pardelensis. The type specimen of Discussion: Ataxocerithium piramiden Ihering's species has a postsutural cord, sis, n. sp., differs from A. Juliana (Ihering, from 18 to 20 axial cords per whorl and 1907), from the Superpatagoniense of San three spiral cords on all of the whorls. Julian (Sta. Cruz), which has one fine Ataxocerithium trespunta (Ihering, 1907), smooth postsutural cord, from 18 to 20 from the Patagoniense of Caho Tres axial cords per whorl, the whorls very con Puntas, has seven spiral cords and 30 vex B,nd narrow, with deeper sutures and axial cords per whorl, the base is orna the base smooth. Ataxocerithium trespun mented with eight simple cords and it has ta (lhering, 1907), from the middle a straight columella. It differs from A. Patagoniense of Caho Tres Puntas (Sta. pullum (Philippi, 1845), which lacks the Cruz) has seven spiral cords per whorl and postsutural groove, has two to three cords 30 axial cords; the base has eight simple on the base, whorls always with three spi cords; the columella is straight and the ral cords, seeming on the last whorl not aperture subquadrangular. Ataxocerithi tuberculate, but rectangular. um pullum (Philippi, 1845) has the sutures less deep, the whorls flatter, quad ATAXOCERITHIUM PIRAMIDENSIS, n. sp. rangular tubercules on the last whorl and Plate 2, figure 15 only two peripheral basal cords. The genus Description: Shell medium-sized, turriform, Ataxocerithium is noted here for the first with 8 1/2 whorls plus two smooth nepionic time from the Entrerriense Formation of whorls. Sutures deep, with two fine intrasutur Argentina. Figueiras and Broggi (1986) al cords. Ornamented with three. spiral cords on cited Cerithiopsis entrerriensis from the all whorls and from 18 to 24 axial cords cross Camacho Formation of Uruguay, as the ing spirals to form tubercules. Base weakly con unique representative of the genus. vex, with three fine peripheral cords. Columella convex, with siphonal canal short and oblique. Superfamily EULIMOIDEA Aperture oval. Family EULIMIDAE Holotype: MPEF-PI-240; height 18.6 mm, Genus BALCIS Leach, 1847 PLATE 3 Figures Page 1. TROPHON LEANZA!, n. sp...... 78 MPEF-PI-116 (holotype); height 30.6 mm (incomplete), width 20.7 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 2. TROPHON HARRINGTONI, n. sp...... 78 MPEF-Pl-124 (holotype); height 23.9 mm (incomplete), width 19.7 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 3. TROPHON PLICATUS var. SANTACRUZENSIS Ihering, 1897 ...... 79 MPEF-Pl-128 (hypotype); height 38.8 mm (incomplete), width 25 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 4. TROPHON CONTORTUS, n. sp...... 80 MPEF-Pl-024 (holotype); height 32.8 mm (incomplete), width 27.2 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 5. TROPHON RETRORSUM, n. ·sp...... 79 MPEF-Pl-180 (holotype); height 58 mm (incomplete), width 44.2 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 6. "TROPHON" VOKESAE, n. sp...... 81 MPEF-Pl-019 (holotype); height 50 mm (incomplete), width 38.5 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. No. 2 Entrerriense Gastropoda 77

PLATE 3 78 Tula.ne Studies in Geology and Paleontology Vol. 30

BALCIS FLECHENSIS, n. sp. All heav:y intercostal cords prolonged over Plate 2, figure 16 lamellae, their projection over the edges giving Description: Shell very small, slender, 9 112 these a -crenulated aspect. Axially, between whorls; all almost flat, body-whorl rounded, lamellae and · crossing cords, from nine to 15 with strong basal carina; sutures weakly very fine threads. In juvenile examples, axial impressed. Aperture oval, external lip broken; cords more conspicuous, giving a cancellate columella arched. Without external ornamenta appearance. tion. Holotype: MPEF-Pl-116; height 30.6 mm Holotype: MPEF-Pl-241; height 6 mm, width (lacking approximately three whorls), width 2mm. 20. 7 mm. Body-whorl: length 26 mm. Aperture: Type locality: Baliza Punta Flecha, Puerto length 11 mm, width 7.5 mm. Siphonal canal: Madryn, Chubut Province, Argentina. ·length 7 mm, width 1 mm. Derivatio nominis: Named for the type locali Type locality: Punta Pardela, Valdes ty. Peninsula, Chubut Province, Argentina. Derivatio nominis: Dedicated to the late geol Discussion: Balcis fiechensis, n . sp., dif ogist Dr. Armando F. Leanza. fers from B. cetia Woodring, 1970, which has the suture profoundly impressed and Discussion: Trophon leanzai, n. sp., dif very convex whorls. Balcis aulaca fers from T retrorsum, n. sp., which has a Woodring, 1970, has very convex whorls, greater number of lamellae (13) on the the suture is better marked and there is body-whorl, lacks the fine spiral cords no peripheral carina on the body-whorl. alternating with the primaries, and has Balcis lipara Woodring, 1970, has some the spire proportionally lower. The what convex whorls., a heavy columellar siphonal canal is strongly oblique toward callus, no peripheral carina on the body the left and the umbilicus forms the pari whorl and the two last whorls globose. etal margin of the siphonal canal. Trophon Balcis jacululum (Maury, 1917) has deep contortus, n. sp., differs by having a strong er sutures, more convex whorls and lacks constriction at the beginning of the a peripheral carina on the body-whorl. si2honal canal, better development of the This unique example represents the first lamellae and the lamellar angulations, record of the genus Balcis from the En which turn at the shoulder abruptly in a trerriense Formation of Soutli America. horizontal direction. The shoulder ramp is wider and horizontal. Trophon harring Superfamily MURICOIDEA toni, n . sp., differs in having nine to 11 Family MURICIDAE Rafmesque, 1815 lamellae, a strong constriction at the Subfamily TROPHONINAE beginning of the siphonal canal, faint spi Cossmann, 1903 ral ornamentation, and lamellar angula Genus TROPHON Montfort, 1810 tions upon reaching the shoulder. The TROPHON LEANZA!, n. sp. shoulder ramp is broader and horizontal, Plate 3, figure 1 without lamellar projections, and there is Description: Shell medium-sized, relatively a straight siphonal canal. Trophon plica thin. Spire elevated; whorls high and narrow. tus var. santacruzensis Ihering, 1897, is Aperture subtriangular, with concave columella more similar to T. plicatus (Lightfoot, forming an angle of almost 90 degrees with pos 1786) and thus is not comparable. This terior lip; obtuse posterior angulation; outer lip species is based on 44 examples. curved and crenulated. Siphonal canal recurved and long, with umbilicus long and inconspicu ous. Fasciole created by growth lines imbricated TROPHON HARRINGTON!, n. sp. one above the other. Last whorl ·with nine rela Plate 3, figure 2 tively strong axial lamellae forming a slightly Description: Shell small, muriciform, elongat elevated angulation at shoulder level, continu- ed; spire long, composed of six to eight whorls, -. ing up to the suture. Shoulder smooth, at an flattened over suture and nearly vertical on angle of 45 degrees. Spiral ornamentation con lower part, forming an almost acute angle. sisting of 15 heavy cords with finer alternating Each whorl with nine to eleven projecting axial . ones; one heavy cord forming shoulder of whorl. lamellae, thin at margins, almost straight and No. 2 Entrerriense Gastropoda 79

pointed on the angle. Lamellae somewhat vari cose and recurved inward, principally on last Plate 3, figure 3 whorl. Lamellae crossed by fine, regularly . Descr~ption : Shell fusiform, heavy, with rela marked spiral striae, and barely visible axials. tively high spire. Aperture oval with reflected labrum; columella concave with marked callus· Last whorl twice the height of spire. Peristome siphonal canal wide and oblique. Fasciol ~ somewhat rounded, with margin smooth and formed by imbricated and projecting growth folded; siphonal canal narrow and elongated, lines; umbilicus poorly defined. Body-whorl oblique toward the left, with a narrow and long with nine laminar varices, strongly angled and umbilicus. curved almost backward to shoulder level· Holotype: MPEF-Pl-124; height 23.9 mm shoulder not flattened. On early whorls marked (lacking approximately three whorls and the spiral striae crossing over laminae, later whorls protoconch), width 19. 7 mm. Aperture: length almost smooth. 11 mm, width 7 mm. Siphonal canal: length 7 Hypptype: MPEF-PI-128; height 38.8 mm = , width 1 mm. (lacking protoconch), width 25 mm. Aperture: Type locality: Puerto Piramide, Valdes Penin length 15 mm, width 8 mm. Siphonal carial: sula, Chubut Province, Argentina. length 8.5 mm, width 2 mm. Derivato nominis: Dedicated to the late geolo Locality: Punta Pardela, Valdes Peninsula, gist Dr. Horacio Harrington. Chubut Province, Argentina.

Discussion: Trophon harringtoni, n. sp., Discussion: Trophon plicatus var. san differs from T. orbignyi Carcelles, 1946, tacruzensis differs from T. plicatus (Light which has the spire more elevated and the foot, 1786) by having a higher spire, the whorls more narrow; the height/width siphonal canal twisted, more oblique, and index of the holotype and 10 paratypes of wider; with fine spiral striations, very visi T. orbignyi varies between 1.53 and 2.47, ble on the early whorls and inconspicuous while the indices of the seven examples of on the body-whorl. The holotype of T. harringtoni vary between 1.21 and 1.32 Ihering's species has 12 varices on the and apparently the spire does not exceed body whorl in contrast to our specimen four whorls. The latter has more devel with only nine. The shoulder is flattened oped lamellae, a narrower siphonal canal, and the siphonal canal is wider. According oblique to the left. Trophon plicatus var. to Ihering (1907), specimens from Parana, santacruzensis Ihering, 1897, from the as well as those from Punta Ninfas and Early and Middle Patagoniense of Santa from Golfo Nuevo (Chubut Province), are Cruz, Argentina, is much smaller in size, poorly preserved, making it difficult to with well developed lamellae. The shoul assign them to the Entrerriense Forma der ramp is flattened, the siphonal canal tion. Therefore, because of this doubt, we longer and the umbilicus indefinite. assign this well-preserved specimen to Trophon retrorsum, n. sp., has a much this species. larger size, a greater number of varices, and strong spiral ornamentation that pro TROPHON RETRORSUM, n. sp. duces crenulations on the lamellar mar Plate 3, figure 5 gins . The lip is strongly beveled, the Description: Shell subfusiform with a short siphonal canal very oblique and the spire, whorls angulose, last w~orl triple the umbilicus long and deep. Trophon contor height of spire. Aperture oval, with strong pos tus, n. sp., has strong axial lamellae, con tero-external angulation; lip heavy and beveled; spicuous spiral ornamentation that pro strong columellar callus, columell~ con~ve pos duces strong crenulations on the lamellar teriorly, ending in one final lamma; s1phonal margins and a siphonal canal that is canal broad and short; umbilicus long and de~p , longer and curved dorsally. We consider T. turned toward siphonal canal. Ornamentation harringtoni a valid and unique predeces on last whorl of slightly projecting lamellae, sor of T. orbignyi. folded over whorl, 13 in number, c~ossed by inconspicuous spiral cords, altern~tmg heavy and fine, at times doubled, producmg crenula tions along lamellar margins; lamellae TROPHON PLICATUS var. SANTACRUZENSIS an~at Ihering, 1897 ed at shoulder level, but elevated only slightly 80 Tulane Studies in Geology and Paleontology Vol. 30 above it. Subsutural ramp radially wrinkled by curved; spire short. The holotype with only 2 growth lines and weak lamellar projections. 1/2 whorls, these somewhat oblique, wider than Penultimate whorl with only five lamellae, all high, increasing rapidly in diameter, separated situated on final third of whorl, the remainder by a shallow suture. Ornamentation of last with no lamellae, only inconspicuous cords; also whorl consisting of nine thin lamellae, curved on antepenultimate whorl, with latter two toward peristome, at shoulder of wh9rl turning whorls having four simple spiral cords. abruptly in a horizontal direction; lamellae fes. Holotype: MPEF-PI-180; height 58 mm (only toned with five to eight spiral cords passing three whorls), width 44.2 mm. Body-whorl: over them; minor cords intercalated at rate of length 50 mm. Aperture: length 24 mm, width one between two principal cords; from middle of 17 mm. Siphonal canal: length 12 mm, width 5 body-whorl aperturally cords becoming finer, mm. the principal cords disappearing. Shoulder Type locality: Punta Pardela, Valdes Penin ramp of last whorl practically flat, and extend· sula, Chubut Province, Argentina. ing over it lamellae merging with growth lines, Derivatio nominis: Denominated thus for the up to level of suture. Besides lamellae, only twisted shell. marked growth lines constituting axial orna mentation. On penultimate whorl, same type of Discussion: We have observed above the ornamentation, except on shoulder ramp, with differences between T. retrorsum, n. sp., T. three weak spiral cords crossing lamellar pro leanzai, n. sp., and T. harringtoni, n. sp. jections. Columella curved, with convexity at Trophon retrorsum differs from T. contor beginning of siphonal canal, ending in a lamina; tus, n. sp., by being of a larger size, with a fasciole long and narrow, with strong wrinkles. greater number oflamellae (13) and incon Holotype: MPEF-PI-024; height 32 .8 mm spicuous spiral ornamentation, lacking (incomplete), width 27.2 mm. Body-whorl: crenulations on the lamellar margins. The length 30 mm. Aperture: length 13 mm, width 8 aperture is oval, the siphonal canal very mm. S"iphonal canal: length 11 mm. wide and very oblique toward the . left, Type locality: · Punta Pardela, Valdes Penin· with a conspicuous umbilicus that forms sula, Chubut Province, Argentina. the inner body-whorl wall of the siphonal Derivatio nominis: Denominated thus for the canai. The differences presented between twist of the shell. Trophon patagonicus (Sowerby, 1846), from the lower Patagoniense of Santa Discussion: Trophon contortus,· n. sp., Cruz (Early Oligocene), Trophon gever differs from Trophon pelseneri (Smith, sianus (Pallas, 1774), living on the 1915) in having famellae that do not cross Patagonian coast, and Trophon retrorsum, over the shoulder ramp, a wider siphonal n. sp., permits us to consider the new canal and less developed lamellae. It dif· species as an intermediate stage between fers from Pagodula vaginatus (Cristofori Trophon patagonicus and the Recent vari and Jan, 1832), which lacks the spiral eties of Trophon geversianus, which is ornamentation, in being of a much smallE>,. characterized by the presence of more or size, having fewer lamellae, axial orna less well developed lamellae and marked mentation elevated above the shoulder spiral ornamentation or by almost equal ramp, and the siphonal canal longer and development of the lamellae and the spiral straighter. Pagodula coulmanensis (Smith, ornamentation, such as we observe in the 1907) has a lesser number of ribs, these Golfo San Jose, Chubut Province, being elevated over the shoulder ramp, Argentina (Vokes, 1991B). with less developed spiral ornamentation and a longer and straighter siphonal canal. We have already established the TROPHON CONTORTUS, n. sp. differences between this species and the Plate 3, figure 4 other species of Trophon s.s. described Description: Shell small, muriciform; aper herein. ture subquadrate; labrum thin with marked crenulations corresponding to projection of spi Genus TROPHON sensu lato ral cords. Strong prolongation of postero-exter nal labral angle. Siphonal canal long and The typical species of Trophon have a No. 2 Entrerriense Gastropoda 81

medium to large, thin, globose shell, with Derivatio nominis: Dedicated to the paleon prominent axial ornamentation. The tologist Dra. Emily H. Vokes of Tulane genus Xanthochorus Fischer, 1884, is University, New Orleans, Louisiana (U.S.A.). characterized by a generally low spire, with spiral ornamentation almost equal to Discussion: "Trophon" vokesae, n . sp., or more developed than the axial, with differs from Trophon patagonicus (Sow conspicuous tubercules on the intersec erby, 1846) by the greater development of tions. The examples, which are here the lamellae, by the greater number of spi described as "Trophon s.l.", have spiral ral cords, by the lamellar prolongations ornamentation rather similar to directed backward, and by the curved Xanthochorus, but with strong varices in siphonal canal. Trophon geversianus para the manner of the genus Trophon. naensis (Borchert, 1901) has less devel oped axial ornamentation, a smooth lip, "TROPHON" VOKESAE, n. sp. and lacks the angular projections on the Plate 3, figure 6 shoulder, a less developed fasciole and a Description: Shell muriciform, higher than straight siphonal canal. Trophon broggii, wide, aperture subtriangular, siphonostomate; n. sp., differs in the weaker development lip thin with undulations that correspond to of the lamellae, which are fewer in num projections of spiral cords; siphonal canal not ber (eight), weaker development of the pri long and slightly curved toward right. Spire low mary spiral ornamentation and a greater (in holotype, which lacks two whorls plus proto number of secondary cords. "Trophon" conch), with whorls wider than high and slight piramidensis, n. sp., has a ficoid form, a ly oblique, enlarging rapidly in diameter; low spire, axial ornamentation consisting suture somewhat deep aild irregular. External of eight heavy cords, a homogeneous spiral ornam~tation of strong undulating lamellar ornamentation, lacking differentiation processes, ten in number on last whorl. between primary and secondary cords. It Lamellae 4 to 5 mm in height; in middle of lacks an umbilicus and has an almost body-whorl forming an angular prolongation straight siphonal canal. This species is reaching to shoulder of whorl; these curving based on 55 adult examples and 73 juve toward right and extending over subsutural niles. .ramp up to suture, at this level 2.5 to 4 mm in height. In interlamellar spaces six convex prin "TROPHON" BROGGII, n. sp. cipal cords, the first forming shoulder of whorl; Plate 4, figure 1 between these from one to three lower convex Description: Shell of medium size, four cords, running spirally. From last primary cord whorls (lacking the protoconch); aperture oval; up to fasciole, 12 or 13 low convex cords. On labrum broadly projecting toward outer side, subsutural ramp of body-whorl from three to with marked undulations formed by projection five barely convex cords. Axial growth lines of spiral cords; strong prolongation of posterio passing over spiral cords, giving them an undu external angle. Siphonal canal short, barely lating aspect. Subsutural ramp oblique down oblique, recurved upward; spire short, com ward owing to great development of primary posed of oblique whorls, wider than high, cord delineating it. Penultimate whorl with ten increasing rapidly in diameter, separated by axial laminar processes 2 mm in height, with shallow suture. Body-whorl four to five times less marked angulations and four spiral cords; higher than spire. Ornamentation formed by subsutural ramp with growth lines only. fine, slightly elevated axial lamellae, eight in Columella slightly curved, endillg at level of number, strongly crenulated, forming a marked umbilicus with a heavy laminar process, angulation on shoulder of whorl; prolonged per Umbilicus long but shallow, with a well devel pendicularly above subsutural ramp to suture. oped fasciole. Subsutural ramp oblique and forming an angle Holotype: MPEF-Pl-019; height 50 mm of 110 degrees relative to anterior whorl, the (incomplete), width 38.5 mm. Body-whorl: latter ornamented by seven or eight low, flat length 40 mm. Aperture: length 25 mm, width cords and a heavy cord forming margin of sub 15mm. sutural ramp. Body-whorl with five primary Type locality: Punta Pardela, Valdes Penin spiral cords, between these from four to seven sula, Chubut Province, Argentina. finer, flat cords, being more numerous (from 20 82 Tulane Studies in Geology and Paleontology Vol. 30 to 25) from last primary cord to umbilical chink. have low varices, others with the varices Lamellae disappearing on two first whorls, more or less well developed are similar in being replaced by small axial cords, crossed by appearance to "T." vokesae, n. sp., but the from five to eight spiral cords. Columella almost juvenile forms of both are easily distin· st raight, enlarged at beginning of siphonal guishable by the greater development of canal and terminating sharply. Umbilicus long the axial ornamentation in "T." vokesae. and shallow; strong fasciole with marked folds. This species is based upon 39 adult exam Holotype: MPEF-Pl-022; height 66.1 mm ples and 40 juveniles. (incomplete), width 48 mm; height of spire 12 mm. Body-whorl: length 55 mm. Aperture: length 46 mm, width 16 mm. (?)"TROPHON" PIRAMIDENSIS, n. sp. Type locality: Punta Pardela, Valdes Penin Plate 4, figure 2 sula, Chubut Province, Argentina. Description: Shell ·ficoid, broad at level of Derivatio nominis: Dedicated to Sr. Jorge shoulder of body-whorl, reducing notably its Broggi, Uruguayan malacologist and paleontol width toward siphonal canal. Spire short, ogist. whorls low and globose. Body-whorl constitut ing approximately 81% of total height (lacking Discussion: "Trophon" broggii, n. sp., one whorl and the protoconch), globose in poste· differs from Trophon geversianus rior part, with shoulder rounded. Aperture sub· paranaensis (Borchert, 1901), which has a triangular; columella curved and long; posterior proportionally more developed spire, a labium concave, at a 90 degree angle to body· smooth lip; no lamellar angulosity at the whorl; strong posterior angulation; labrum shoulder, fewer spiral cords (three), and a beveled, thin and directed toward outside, with straight siphonal canal. Trophon patagoni small crenulations corresponding to projection cus (Sowerby, 1846) has a great number of of spiral cords; siphonal canal long, narrow and lamellae (12), with undifferentiated and straight. Fasciole formed by imbricated, barely fewer spiral cords (from three to seven). In projecting growth lines, lacking an umbilicus. the latter, the lamellar angulations are External ornamentation consisting of eight directed upward, but in Trophon broggii, heavy, projecting axial cords, forming a small, n. sp., they are extended laterally. inconspicuous tubercule at shoulder level; these "Trophon" piramidensis, n. sp., has a ficoid cords lost on anterior third of body-whorl; spi· form, a low spire, axial ornamentation rally seven or eight cords above shoulder, and consisting of eight heavy cords, homoge laterally up to 32 flat cords, alternating heavy neous spiral ornamentation, lacking dif and weak. ferentiation between primary and sec Holotype: MPEF-Pl-119; height 28. 6 mm ondary cords; it lacks an umbilicus and (lacking one whorl plus the protoconch), width has a straight siphonal canal. Some adults 19 mm. Aperture: length 15 mm, width 9 mm.

PLATE4 Figures Page 1. "TROPHON" BROGGII, n. sp...... 81 MPEF-PI-022 (holotype); height 66.1 mm (incomplete), width 48 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 2. (?)"TROPHON" PIRAMIDENSIS, n. sp...... · .82 MPEF-Pl-119 (holotype); height 28.6 mm (incomplete), width 19 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 3. FUEGOTROPHON FLINT!, n. sp...... · .84 MPEF-Pl-121 (holotype); height 29.6 mm (incomplete), width 17.5 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 4. FUEGOTROPHON TURNER!; n. sp...... · · .84 MPEF-Pl-122 (holotype); height 23.5 mm (incomplete), width 16.4 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 5. XYMENOPSIS FLECHENSIS, n. sp...... · .85 MPEF-Pl-025 (holotype); height 32.5 mm (incomplete), width 20.5 mm. Locality: Baliza Punta Flecha, Puerto Madryn, Chubut Province, Argentina. No. 2 Entrerriense Gastropoda 83

PLATE4 TULANE STUDIES IN GEOLOGY AND PALEONTOLOGY Volume 30, Number 2 August 27, 1997

NEW SPECIES OF MOLLUSCA FROM THE ENTRERRIENSE FORMATION (UPPER MIOCENE} OF CHUBUT PROVINCE ARGENTINA AND SPECIES NOT PREVIOUSLY REPORTED FROM THIS FORMATION PART II - GASTROPODA

RODOLFO F.J . BRUNET PUERTO MADRYN, CHUBUT PROVINCE ARGENTINA

CONTENTS .·· Page I. ABSTRACT ...... 61 II. INTRODUCTION ...... 61 III. ACKNOWLEDGMENTS ...... 61 IV. SYSTEMATIC PALEONTOLOGY ...... 61 V. LITERATURE CITED ...... 97

61 Tulane Studies in Geology and Paleontology Vol. 30 62

,. _) >.

420 s

P. Bajos

LDES P. Hercules Punta Delgada

0 50 100 150

SN MILCIMSTRDS

PLATE 1 66 Tulane Studies in Geology and Paleontology Vol. 30

PLATE2 72 Tulane Studies in Geology andPaleontology Vol. 30

PLATE 3 78 Tula.ne Studies in Geology and Paleontology Vol. 30

PLATE4 84 Tulane Studies in Geology and Paleontology Vol. 30

Siphonal canal: length 6 mmr width 1 mm. ed, without postero-extemal angulation. Type locality: Puerto Piramide, Valdes The lip is not beveled and the siphonal Peninsula, Chubut Province, Argentina. canal is oblique. Fuegotrophon asperum, n. Derivatio nominis: Named thus for the type sp., is fusiform, with the spire more ele locality. vated (50% of the total height), and the whorls wider and high. There is one Discussion: (?)"Trophon" piramidensis, strong cord at the level of the shoulder on n. sp., presents characters atypical of the the last whorl. "muriciforms." It differs from "T." vokesae, n. sp., and "T. " broggii, n. sp., for the rea sons expressed in the discussion of each of FUEGOTROPHON TURNER!, n. sp. these. This species is based upon 26 adult Plate 4, figure 4 examples and 15 juveniles, in some of Description: Shell broad and low; last whorl these there are varices that form a small, also broad and relatively short. Aperture almost acute angulation at the shoulder; in other semicircular; labrum rounded with crenulations examples the axial cords are less elevated, corresponding to projections of spiral cords. but equal in number. Columella concave; siphonal canal short, nar row, and oblique. Fasciole consisting of imbri Genus FUEGOTROPHON Powell, 1951 cated growth lines, umbilicus long but shallow. FUEGOTROPHON FLINT!, n. sp. Body-whorl inflated, with seven heavy, project· Plate 4, figure 3 ing cords; spirally from 16 to 20 cords, crossed Description: Shell fusiform, spire not elevat by numerous fine, corrugated axial cordlets. ed; w;horls rounded. Aperture pyriform;. col Penultimate whorl with only six spiral cords umella concave posteriorly and convex anterior and nine much heavier axial cords. ly, with well developed columellar callus. Holotype: MPEF-PI-122; height 23. 5 mm Labrum recurved with weak posterior angula (lacking two whorls and the protoconch), width tion, crenulated along entire margin by projec 16:4 mm. Aperture: length 9.5 mm, width 6.5 tion of spiral cords; siphonal canal long, oblique mm. Siphonal canal: length 6 mm, width 1 mm. and relatively wide. Fasciole .consisting of TJpe locality: Puerto Piramide, Valdes Penio imbricated growth lines; no umbilicus. sula, Chubut Province, Argentina. Ornamentation of body-whorl consisting of cer Derivatio nominis: Dedicated to the late geol· tain growth lines, more marked than the rest, ogist Dr. Juan C.M. Turner. intercalated as weak axial cords. Spiral cords more developed, four at level of shoulder, one Discussion: Fuegotrophon turneri, n. sp., stronger forming shoulder of whorl, and lateral differs from Fuegotrophon pallidus ly from 16 to 19 spiral cords. Axially innumer (Broderip, 1833) in having a less fusiform able fine, corrugated striae. Penultimate whorl shell, a broader spire, a semicircular aper rounded, without shoulder and with nine spiral ture, a shorter siphonal canal, and by hav cords, plus lamelliform axial cordlets. ing an umbilicus. It differs fro m Holotype: MPEF-PI-121; height 29.6 mm Fuegotrophon flinti, n. sp., by having (lacking two whorls, plus the protoconch), width heavier, more marked axial cords, a short 17.5 mm. Aperture: length 11.5 mm, width 7 er, oblique siphonal canal, and a conspicu mm. Siphonal canal: length 6.5 mm, width 1.5 ous umbilicus. Fuegotrophon asperum , n. mm. sp., is fusiform, . with inconspicuous axial Type locali ty: Puerto Piramide, Valdes ornamentation, but strong spiral orna· Peninsula, Chubut Province, Argentina. mentation, and a strong cord forming the Derivatio nominis: Dedicated to the late geol shoulder of the body-whorl. It lacks an ogist Dr. Richard Foster Flint. umbilicus.

Discussion: Fuegotrophon flinti, n . sp., FUEGOTROPHON ASPERUM, n. sp. differs from Fuegotrophon turneri, n. sp., Plate 5, figure 1 which has a greater development of the Description: Shell fusiform, small, with six axial ornamentation, consisting of true convex whorls, the last almost twice height of cords. The subsutural ramp is not differ spire. Two last whorls with a strong cord form· entiated and the aperture is totally round- ing a shoulder. Aperture oval, with outer hp No. 2 Entrerriense Gastropoda 85

beveled and crenulated by projection of spiral Madryn, Chubut Province, Argentina. cords; siphonal canal narrow, long and. oblique. Derivatio nominis: Named for the type locali External ornamentation consisting of nine spi ty. ral cords, plus two on subsutural ramp; on last whorl four or five poorly defined axial cords. It Discussion: Xymenopsis flechensis, n. is not possible to detail other characters of spec sp., differs from Xymenopsis muriciformis imen due to incrustation of bryozoans and state (King and Broderip, 1832) in having fewer of fossilization of shell. axial cords (eight), weaker development of Holotype: MPEF-PI-023; height 18 mm, the spiral ornamentation, with a preva width 7.9 mm. Body-whorl: length 13 mm. lence of the axial. ornamentation over the Aperture: length 5 mm, width 3 mm. Siphonal spiral. The aperture with the peristome is canal: length 3 mm. much more closed anteriorly, and there is Type locality: Puerto Piramide, Valdes Penin a wide umbilicus. Xymenopsis falklandi sula, Chubut Province, Argentina. cus (Strebel, 1908) has better development Deriuatio nominis: Denominated thus for the of the spiral ornamentation, a greater roughness of the shell. number of axial cords and lacks an umbili cus. Xymenopsis michelii, n. sp., differs in Discussion: Fuegotrophon asperum, n. having lower, .wider, more globose whorls. sp ., is most similar to the species The platform is undifferentiated and the Fuegotrophon pallidus (Broderip, 1833), axial ornamentation inconspicuous. The but differs from it in having a lesser num aperture is more elongated, the lip not ber of spiral cords and in having the aper beveled, the siphonal canal is straight and ture represent 62% of the length of the there is no umbilicus. Xymenopsis aperture-siphonal canal, whereas in the quenseli, n, sp., has ten strong axial cords, first species it is 69%. It differs from F. which end in angulations at the level of flinti, n. sp., and F. turneri, n. sp., for the the shoulder. There is a strong constric reasons previously discussed under each tion that begins at the siphonal canal. The of these species. platform is concave and there is no umbilicus. Genus XYMENOPSIS Powell, 1951 XYMENOPSIS MICHELII, n. sp. XYMENOPSIS FLECHENSIS, n. sp. Plate 5, figure 2 Plate 4, figure 5 Description: Elongated shell with rounded Description: Shell fusiform, whorls rounded whorls, spire high; last whorl representing 81% (only 2 1/2 whorls preserved); aperture oval, of total height. External ornamentation consist labrum festooned by projection of spiral cords, ing of fine axial and spiral cords. Aperture siphonal canal rather long and curved; suture broad, elliptical, with strong columellar callus; shallow and irregular. External ornamentation labrum rounded with crenulations correspond consisting of eight heavy, wide and rounded, ing to projection of spiral ornamentation; single or double ·cords, running axially; dimin siphonal canal short, wide and slightly curved; ishing in thickness aperturally; from 15 to 17 fasciolar area consisting of imbricated growth marked spiral cords, passing over axials; last lines and a long but shallow umbilicus. whorl rounded, with a length more than double Holotype: MPEF-PI-118; height 34.2 mm that of spire, without forming a shoulder, but (lacking approximately one whorl plus the pro axial cords continuing up to suture; penulti toconch), width 18.8 mm. Aperture: length 11 mate whorl with eleven heavy longitudinal mm, width 8 mm. Siphonal canal: length 7 mm, cords, crossed by five finer spiral cords, passing width 1.5 mm. over primaries. Columella slightly concave, end Type locality: Puerto Piramide, Valdes Penin ing at canal as a lamina. Umbilicus long, wide sula, Chubut Province, Argentina. and deep; fasciole well developed, with strong Derivatio nominis: Dedicated to the diver Sr. folds. Ricardo Micheli. Holotype: MPEF-Pl-025; height 32.5 mm (incomplete), width 20.5 mm. Body-whorl: Discussion: Xymenopsis michelii, n . sp., length 28 mm. Aperture: length 14 mm, width 8 differs from X. albidus (Philippi, 1846) by mm. having a less developed columellar callus Type locality: Baliza Punta Flecha, Puerto and unequal axial ornamentation, without 86 Tulane Studies in Geology and Paleontology Vol.30

defined cords. The peristome is not reflect cords on the body-whorl, with less deveJ. ed outward, and the crenulations are opment of the spiral ornamenation. It much less conspicuous. The umbilicus is lacks the shoulder ramp, which is replaced longer and less deep. Xymenopsis fiechen in this. case by a perisutural groove. The sis, n. sp., has more globose whorls and siphonal canal is closed at the beginning more defined axial ornamentation in the by a projection of the lip. form of heavy cords. The aperture is semi circular, the siphonal canal oblique and XYMENOPSIS PENINSULARIS, n. Sp. the umbilicus of greater size. Xymenopsis Plate 5, figure 4 quenseli, n. sp., has the whorls not so glo Description: Shell ficoid, very thin, with short bose, with a greater development of the spire and globose whorls. Body-whorl broad at spiral ornamentation and a well defined beginning, diminishing rapidly in diameter. shoulder. The platform is concave, the Aperture pyriform, columella concave, with a axial ornamentation well defined and con conspicuous columellar callus. Labrum strongly spicuous, with on some whorls small angu crenulated by projection of external spiral la tions at the level of the shoulder. cords; siphonal canal moderately long, closed at Xymenopsis peninsularis, n. sp., has a· beginning by a projection of lip, more open strong constriction at the beginning of the anteriorly. .Fasciole composed .of imbricated but siphonal canal, ornamentation consisting weakly projecting growth lines. Ornamentation of strong rounded cords, a deep perisutur of body-whorl consisting of heavy tuberculate al groove and a semi-closed siphonal cords beginning at level of suture, eight in num· canal. her; disappearing in middle of whorl. Five spi· ral cords on shouHler and from 18 to 20 on lat eral face; between these finer threads interca· XYMENOPSIS QUENSELI, n. sp. lated. At suture a deep groove bordering all Plate 5, figure 3 whorls, crossed only by growth striae. Description: Shell relatively short and broad, Holotype: MPEF-Pl-123; height 20 mm (lack· with low whorls. Aperture almost semicircular, ing two whorls and the protoconch), width 14.7 with concave columella, posterior lip at a 45 mm. Aperture: length 8 mm, width 5 mm. degree angle to body-whorl, with poorly defined Siphonal canal: length 6 mm, width 1 mm. posterior angulation, remainder of lip recurved, Pype locality: Puerto Piramide, Valdes Penin· with crenulations corresponding to projection of sula, Chubut Province, Argentina. spiral cords. Strong constriction at beginning of Derivatio nominis: Dedicated to the Valdes short, straight, oblique siphonal canal. Fasciole Peninsula, site of the type locality. with imbricated growth lines, no umbilicus. Body-whorl ornamented by ten heavy axial Discussion: Xymenopsis peninsularis, n. cords. Strong _spiral ornamentation of 15 to 18 sp., differs from X. fiechensis, n. sp., X. cords, passing over axial cords, alternating michelii, n. sp., and X. quenseli, n. sp., for between heavy and weak, that forming shoul the reasons mentioned in the treatment of der heavier. Shoulder oblique, with weak each of these species. It differs from the growth lines and two or three weak spiral 'stri living representatives of the genus by the ae. presence of a conspicuous perisutural Holotype: MPEF-PI-120; height 24 mm (lack groove. ing two whorls and the protoconch), width 16.4 mm. Aperture: length 10 mm, width 7.5 mm. Siphonal canal: length 5 mm, width 2 mm. Genus PAGO DULA Monterosato, 1884 Pype locality: Puerto Piramide, Valdes Penin PAGODULA CARCELLESI, n. sp. sula, Chubut Province, Argentina. Plate 5, figure 5 Derivatio nominis: Dedicated to the late geol Description: Shell fusiform, strong; spire high ogist Dr. Percy Quensel. and stepped (approximately 20% of total height); apex with three angular whorls and Discussion: We have established above two nepionic whorls. Aperture subpyriform. the differences between Xymenopsis Labrum arcuate, corrugated toward outside, quenseli, n. sp., X. fiechensis, n. sp., and X. with strong posterior angulation. Columellar michelii, n. sp. Xymenopsis peninsularis, callus inconspicuous. Siphonal canal long, nar· n. sp., has more globose whorls, eight axial row and open, straight in juvenile examples, No. 2 Entrerriense Gastropoda 87

but somewhat curved in adults. Fasciole long, PAGODULA .aff. P. CARDUELIS (Watson, 1883) formed by weakly marked imbricate growth Plate 5, figure 6 lines. Ornamentation consisting of strong, wide Description: Shell muriciform, spire high, Jamellae, on body-whorl starting from begin whorls slightly rounded. Last whorl with a ning of siphonal canal, extending separately height of 71 % of the total; spire with six whorls; from body-whorl and upon reaching shoulder aperture ovoid; columella concave. Labrum becoming markedly elevated and twisted back beveled and reflected. Siphonal canal long and ward. On other whorls, lamellae beginning at narrow. Body-whorl ornamented with ten axial suture, running separated from wall of whorl, lamellae, elevated above the shoulder. markedly elevated at level of shoulder and Hypotype: MPEF-PI-125; height 22 mm (only curved backward. Shoulder of whorls showing four whorls preserved), width 16. 7 mm. growth Jines in form of an "S," reaching to Locality: Puerto Piramide, Valdes Peninsula, suture. Well-marked spiral striae on all lamel Chubut Province, Argentina. Jar surfaces and an axial ornamentation con sistirig of fine undulating striae, better marked Discussion: Given the state of the shell, on siphonal canal. which had to be extracted by plastering Holotype : MPEF-PI-129; height 57.3 mm the shell in situ, we assign the species (lacking approximately 1/4 of the siphonal with reservation to Pagodula carduelis canal), width 34.5 mm. Aperture: length 19 mm, (Watson, 1883), living off the eastern coast width 12 mm. Siphonal canal: length not mea of Australia. We have also studied another surable, width 3.5 mm. Height of lamellae two examples, also in a bad state of above body-whorl: 6 mm (average). preservation. Type locality: Playa Fra~asso, Valdes Penin sula, Chubut Province, Argentina. Subfamily THAIDINAE Derivatio nominis: Dedicated to the eminent Genus STRAMONITA Schumacher, 1817 malacologist Alberto Carcelles. STRAMONITA HOMOGENEUS, n. sp. Plate 5, figure 7 Discussion: Pagodula carcellesi, n. sp., Description: Shell heavy, oval-oblong, with differs from Padogula carduelis (Watson, whorls markedly convex, the last three times 1883), living in eastern Australia, which the size of the spire; aperture subtriangular; has a much smaller size (32 mm; BMNH labrum crenulated, crenulations prolonged type collection 87.2.9.571-2), with the toward interior of aperture; siphonal canal spire proportionally higher (29.4% of the short and slightly curved; suture shallow. total height), and with a greater develop Ornamentation of last whorl provided by axial ment of the labral margin and of the pos ribs formed only by growth lines, in places more tero-lateral angulation. It has strong spi notable (in some examples forming small nod ral ornamentation on the early whorls and ules on shoulder of whorl). Spiral ornamenta the columellar callus is more developed. tion consisting of barely projecting, almost flat Pagodula coulmanensis (E.A. Smith, cords, from 35 to 45 in number; among these 1907), living in the Antarctic Zone, has a more conspicuous ones forming shoulder of much smaller size (10.4 mm; BMNH type whorl, with two or three more distributed on collection 1905.9.25.02), with fewer lamel external face. Growth lines running longitudi lae on the body-whorl (six), straight nally, c:r;ossing cords. Penultimate whorl also whorls and the shoulder ramp also with weak axial cords and six spiral cords, straight. The spire is proportionally more forming the shoulder. Suture having small elevated (39% of total height), the lamel oblique folds, forming growth lines upon reach lae elevated above the shoulder and the ing suture. External lip slightly angulate (very siphonal canal proportionally shorter. This angulate in examples possessing tubercules at species is based on seven adult examples, shoulder). Columella concave, with post in varying states of preservation, collected siphonal thickening forming internal margin of exclusively from Playa Fracasso over a umbilical zone. Umbilicus scarcely suggested; period of ten years. This is the first record fasciole strong, beginning from parietal lip and of this genus in the Entrerriense forming external rim of umbilical zone. Formation of South America. Holotype: MPEF-Pl-026; height 45.7 mm, width 29 mm; height of spire 13 mm. Body whorl: length 38 mm. Aperture: length 20 mm, 88 Tulane Studies in Geology and Paleontology Vol.30

width 12 mm. Genus ILYANASSA Stimpson, 1865 Type locality: Punta Pardela, Valdes Penin ILYANASSA sp. sula, Chubut Province, Argentina. Plate 6, figure 1 Deriuatio nominis: Named thus for the incon Description: Th this unique example there is spicuous ornamentation of the shell. preserved only part of the last whorl and a small portion of the penultimate. The last whorl Discussion: Stramonita homogeneus, n. apparently wide and strong, the outer lip bro sp., differs from Stramonita haemastoma ken, but it is possible to see lirae interiorly, floridiana (Conrad, 1837) in having an which end at interior margin of lip. Colwnella umbilicus, a greater development of the very strong, convex, with one heavy fold at the fasciole, and weaker axial ribs. It differs level of the pillar. Columellar callus very wide, from Stramonita haemastoma haemas curving over part of wall of body-whorl. toma (Linne, 1767) by the absence of Siphonal canal short, deep and ob1ique toward tubercules, lacking the palatal sinus and left. Basal fasciole reaching to middle of body by the presence of an umbilicus. This new whorl, witli four heavy plications, very project species is similar to the subspecies of ing and curved. Exteriorly, in spite of wear, Stramonita haemastoma from Rio de appearing to be ornamented. with strong axial Janeiro and to Stramonita haemastoma striae. floridiana from the Florida peninsula. Hypotype: MPEF-Pl-242; measurement Having obtained a considerable number of impractical. specimens permits us to confirm that Locality: Puerto Piramide, Valdes Peninsula, already in the Tertiary there is a great .Chubut Province, Argentina. variability of forms, such as the presence of the umbilicus, although poorly marked, Discussion: Ilyanassa sp., in spite of the indicating a tendancy toward the disap poor state of preservation, is distinguish pearance of the same. In the future the able from I. arata (Say, 1824), which has a reader will be informed on the variability straight siphonal canal, less conspicuous of this species in the Tertiary of fasciole and spiral ornamentation. Patagonia. llyanassa irrorata (Conrad, 1863) does not have the heavy columellar plicae and has Family NASSARIIDAE cancellate ornmentation. Ilyanassa sex·

PLATES Figures Page L FUEGOTROPHON ASPERUM, n. sp...... · · .84 MPEF-Pl-023 (holotype); height 18 mm, width 7.9 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argeqtina. 2. XYMENOPSIS MICHELII, n. sp...... · .85 MPEF-PI-118 (holotype); height 34.2 mm (incomplete), width 18.8 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 3. XYMENOPSIS QUENSELI, n . sp...... 86 MPEF-Pl-120 (holotype); height 24 mm (incomplete), width 16.4 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 4. XYMENOPSIS PENINSULARIS, n. sp...... · · .86 MPEF-Pl-123 (holotype); height 20 mm (incomplete), width 14.7 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 5. PAGODULA CARCELLESI, n. sp...... · · · · · · · · · ·86 MPEF-Pl-129 (holotype); height 57.3 mm (incomplete), width 34.5 mm. Locality: Playa Fracasso, Valdes Peninsula, Chubut Province, Argentina. 6. PAGODULA aff. P. CARDUELIS (Watson, 1883) ...... · · · · · ·87 MPEF-Pl-125 (hypotype); height 22.mm (incomplete), width 16.7 mm (incomplete). Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 87 7. STRAMONITA HOMOGENEUS, n. sp...... · · · · · · · · · · · ·. MPEF-Pl-026 (holotype); height 45.7 mm, width 29 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. No. 2 Entrerriense Gastropoda 89

PLATE 5 90 Tulane Studies in Geology and Paleontology Vol. 30 dentata (Conrad, 1843) has a lip with six Discussion: We attribute this juvenile denticles, a sigmoidal columellar lip, and example to the species A. (A.) beckii axial and spiral ornamentation. Ilyanassa because of its general aspect, its proto wilmingtonensis Gardner, 1948, has a conch, its. very marked shoulder, bordered more elongate form, a shallow straight with tubercules, and its two columellar siphonal canal, slightly oblique toward the plications, typical of the species. This is right with axial and spiral ornamentation. the first record of this species from the Ilyanassa isogramma Dall, 1892, has one Entrerriense Formation of South America. inconspicuous columellar plication and a poorly differentiated anterior fasciole. ADELOMELON (ADELOMELON) FERUSSACII Ilyanassa obsoleta (Say, 1822) has one (Donovan, 1824) heavy columellar plication, with axial and Plate 6, figure 6 spiral ornamentation. llyanassa granifera Description: Shell of medium size, fi ve convex (Conrad, 1868) has cancellate ornamenta whorls, two forming protoconch; last whorl tion. Ilyanassa schizopyga Dall, 1892, is inflated, 5 112 times height of spire, without subumbilicate, with a straight anterior tubercules; spire short and pointed. Lip convex, siphonal canal and cancellate ornamenta with anal canal inconspicuous, but marked. tion. lyanassa johnsoni (Dall, 1892) has Aperture wide, with maximum amplitude in the columellar callus exceedingly devel anterior third, siphonal canal broken. oped and axial and spiral ornamentation. Columella strongly convex posteriorly, then con Ilyanassa scalaspira (Conrad, 1868) dif tinuing almost straight, with two heavy plica fers by its inconspicuous columellar plica tions. Columellar callus very heavy, margined tion and its ornamentation consisting of by a glassy surface; fasciole oblique, with heavy spaced axial cords and fine spiral marked margin and numerous very imbricate striae. Ilyanassa harpuloides (Conrad, plications. Ornamented with heavy axial cords 1844) has the fasciole-poorly developed, caused by a great development of growth lines, shallow siphonal canal and cancellate with fine intermediate cords. ornamentation. This is the first record of Hypotype: MPEF-PI-243; height 63.2 mm, the genus Ilyanizs:sa from the Entrerriense width 37 mm. Formation of South America. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. Family VOLUTIDAE Subfamily ZIDONINAE Discussion: The figured specimen pre Genus ADELOMELON Dall, 1906 sents conchological characters similar to Subgenus ADELOMELON s.s. A. ferussacii , differing only by the pres ADELOMELON (AnELOMELON) BECKI! ence of two· columellar plications, but in (Broderip, 1836) the figure given by Clench and Turner Plate 6, figure 2 (1964, pl. 98)·it is possible to see the pres Description: Shell juvenile, very small, 4 112 ence of only two columellar plications; whorls, With two nepionic whorls; last whorl therefore, this is the first record of this very elongate and narrow, with deep suture, species from the Entrerriense Formation shoulder marked, very wide, edged with con of South America. spicuous tubercules. Lip joined to body-whorl very low, forming an angulation -as. a conse Subgenus PACHYCYMBIOLA quence of projection of tubercules, then describ Ihering, 1907 ing a gentle curve, closing in at top of siphonal AnELOMELON (PACHYCYMBIOLA) aff. A.(P.) canal. Columella with heavy callus and two col BRASILIANA (Lamarck, 1811) umellar plications, the anterior stronger than Plate 6, figure 3 the posterior. Shell practically smooth, with Description: We have only part of the last only fine-growth lines and some inconspicuous, whorl and posterior edge of the lip. Body-whorl widely spaced spiral striae. with conspicuous tubercules, which continue Hypotype: MPEF-PI-259; height 30.3 mm, forward in the form of fine cords. Posterior width 13 mm. labral margin very angulate. Columella convex Locality: Puerto Piramide, Valdes Peninsula, its entire length; heavy columellar callus and Chubut Province, Argentina. two plications, the anterior much heavier than the posterior. No. 2 Entrerriense Gastropoda 91

Hypotype: MPEF-Pl-244; measurement or labral angulation, from three to four impractical. columellar plications, but we believe that Locality: Puerto Piramide, Valdes Peninsula, we can consider our species a good ances Chubut Province, Argentina. tor. This is the first record of the genus Odontocymbiola in the Entrerriense Discussion: We assign this specimen Formation of South America. with some doubt to A. (P.) brasiliana on the basis of the heavy posterior labral angula tion, the wide callus, and the presence of Family OLIVIDAE two unequal columellar plications. This is Subfamily OLIVINAE the first record of the subgenus in the Genus OLIVANCILLARIA d'Orbigny, 1840 Enterrriense Formation of South America. OLIVANCILLARIA PRISCA Ihering, 1907 Plate 6, figure 7 Subfamily ODONTOCYMBIOLINAE Description: Shell of mediilm size, oliviform, Genus ODONTOCYMBIOLA Clench and spire low, conical, apex pointed; suture chan Turner, 1964 neled in final part of last whorl; body-whorl ODONTOCYMBIOLA PARDELENSIS, n. sp. occupying 87% of total height. Aperture subtri Plate 6, figure 4 angular, anteriorly very wide. Columella with Description: Shell of medium size, only last three very heavy oblique plications, fasciole whorl preserved, that relatively narrow, with a very large, reaching almost to middle of length row of tubercules at shoulder. Aperture wide, of aperture. Columellar callus conspicuous, lip attached to body-whorl at a sharp angle, extending to canalicular suture, without cover gradually opening so that aperture is wider in ing spire. Ornamented with undulating growth the anterior third. Siphonal canal short but lines. wide. Columella convex posteriorly, practically Hypotype: MPEF-PI-247; height 15 mm, straight and oblique anteriorly, with two very width6 mm. heavy plications, the anterior more conspicu Locality: Puerto Piramide, Valdes Peninsula, ous . Columellar callus very developed. Chubut Province, Argentina. Ornamented with tubercules prolonged axially by heavy plications. Fasciole oblique, subparal Discussion: We consider that this exam lel to columella, not very high, with heavy ple presents the characters described by curved plications. Ihering (1907) for 0. prisca, from the Holotype: MPEF-PI-245; height incomplete, uppermost Entrerriense of Parana. width59mm. Figueiras and Broggi (1971-73) mention Type locality: Punta Pardela, Valdes Penin the species from the Camacho Formation sula, Chubut Province, Argentina. of Uruguay on the basis of internal and Defiuatio nominis: Named for the type locali external molds with shell remains. ty. Philippi (1893) described, on the basis of a indeterminable mold, Oliva platensis; Discussion: Odontocymbiola pardelen therefore, Ihering decided to denominate sis, n. sp., differs from 0 americana the Parana species as 0 . prisca. (Reeve, 1856) by its outline, which is wider at the shoulder level, appearing less OLIVANCILLARIA FRACASSOENSIS, n. sp. cylindrical, by having only two columellar Plate 6, figure 5 plications, a better developed callus, a Description: Shell of medium size, heavy. wider siphonal canal, and less conspicuous Spire broken; suture canaliculated at least on tubercules. It differs from the varieties of last whorl; aperture very large, rhomboidal, lip 0. americana, such as 0. cleryaria Petit, forming a sharp angle with body-whorl and a 1856, which has three columellar plica small posterior siphonal pseudocanal, greatest tions. Odontocymbiola magellanica diameter of aperture in anterior third, then (Gmelin, 1791) differs by generally lacking notably narrowing and forming short and nar tubercules, having a much wider last row anterior siphonal canal. Columella curved, whorl, and a columella with two to five with two conspicuous plications at beginning of small plications. Odontocymbiola subno anterior third. Columellar callus well devel dosa (Leach) (in Clench and Turner, 1964, oped, not extending to cover spire. Basal fasci p. 155, pls. 95, 96), has one heavy posteri- ole relatively small in proportion; with strong 92 Tulane Studies in Geology and Paleontology Vol. 30 cords that are a continuation of conspicuous with heavy cords as ornamentation. In 0. axial cords ornamenting shell. auricularia the fasciole is smooth, occupy Holotype: MPEF-Pl-248; height incomplete, ing more than 113 of the surface of the width 36mm. shell, and is delimited by a fine groove. Type locality: Playa Fracasso, Valdes Penin Olivancillaria carcellesi Klappenbach, sula, Chubut Province, Argentina. 1965, has only one .basal plication and Derivatio _nominis: Named for the type locali seven small columellar plications, a deep ty. er anal canal, the width the aperture is greatest anteriorly, and the siphonal canal Discussion: Olivancillaria fracassoen is shorter and wider. It differs from 0. sis, n . sp., differs from 0. auricularia teaguei· Klappenbach, 1964, which has a (Lamarck, 1810) by its more· elongated height of only approximately 25 mm, a col form, an aperture posteriorly narrow, with umellar callus slightly developed, a long its maximum diameter in the anterior constricted aperture, ten oblique plica third; its longer and narrower siphonal tions and four longitudinal to the columel canal; its columellar plications located lar axis. Olivancillaria urceus (Roding, more poster iorly, and its small fasciole 1798) (= 0. brasiliana Lamarck, 1810) has

PLATE 6 Figures Page 1. ILYANASSA sp...... 88 MPEF-r: -242 (hypotype); measurement impractical. LocGLlity: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 2. ADELOMELON (ADELOMELON) BECKI! (Broderip, 1836) ...... 90 MPEF-PI-259 (hypotype); height 30.3 mm, width 13 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 3. ADELOMELON (PACHYCYMBIOLA) aff. A.(P.) BRASILIANA (Lamarck, 181-1) ...90 MPEF-PI-244 (hypotype); measurement impractical. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 4. ODONTOCYMBIOLA PARDELENSIS, n. sp...... 91 MPEF-Pl-245 (holotype); height incomplete, width 59 mm. Locality: Punta Pardela, Valdes Peninsula, Chubut Province, Argentina. 5. OLIVANCILLARIA FRACASSOENSIS, n. sp...... 91 MPEF-Pl-248 (holotype); height incomplete, width 36 mm. Locality: Playa Fracasso, Valdes Peninsula, Chubut Province, Argentina. 6. ADELOMELON (ADELOMELON) FERUSSACII (Donovan, 1824) ...... 90 ~EF-PI-243 (hypotype); height 63.2 mm, width 37 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 7. OLIVANCILLARIAPRISCAihering, 1907 ...... 91 MPEF-Pl-247 (hypotype); height 15 mm, width 6 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 8. OLIVELLA aff. 0. PLATA (lhering, 1909) ...... 94 MPEF-Pl-251 (hypotype); height 6 mm, width 3 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 9. OLIVANCILLARIA aff. 0. URETA! Klappenbach, 1965 ...... · · .94 MPEF-Pl-249 (hypotype); height 17.5 mm, width 7.2 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 10. OLIVELLA aff. 0 . PUELCHA (Duclos, 1840) ...... · · · .95 MPEF-PI-250 (hypotype); height 8.5 mm, width 4 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 11. OLIVELLA FRACASSOENSIS, n. sp ...... · · · .9 4 MPEF-PI-252 .(holotype); height 9 mm, width 3.2 mm. Locality: Playa Fracasso, Valdes Peninsula, Chubut Province, Argentina. 12. PRUNUM sp...... · , · · · · · ·95 MPEF-Pl-253 (hypotype); height 22 mm, width 9 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. No.2 Entrerriense Gastropoda 93

9 11

PLATE 6 94 Tulane Studies in Geology and Paleontology Vol. 30

the last whorl conical, with its greatest Locality: Puerto Piramide, Valdes Peninsula, diameter posteriorly, a more reduced cal Chubut Province, Argentina. lus, only one basal plication and small col umellar plications, a straight lip, a fasciole Discussion: We relate this specimen to occupying 1/3 of the surface of the shell, 0. puelcha basically by the presence of a and a short and wide siphonal canal. single columellar plication. Figueiras and Broggi (1971-73) mention this species 0LIVANCILLARIA aff. 0 . URETA! from the Barranca de San Pedro, Colonia Klappenbach, 1965 (Camacho Formation); consequently this Plate 6, figure 9 is the first record from the Entrerriense Formation of Argentina. From the Middle Description: Shell oliviform, spire low, coni Miocene of Brasil, Closs (1970), cited cal, apex pointed. Suture channeled on final Olivella (Olivina) bullula (Reeve, 1850) part of last whorl. Callus weak, but extending and Olivella (Toroliva) goliath Olsson, in an inconspicuous form to sutural channel, 1922. without covering spire. Columella almost straight with five to six oblique plications, sepa OLIVELLA aff. 0. PLATA (Ihering, 1909) rated by a groove of the basal plication. Plate 6, figure 8 Fasciolar band inconspicuous. Aperture occupy Description: Shell small, slender, spire thin, ing four-fifths of total length. Ornamented with whorls ·high, suture deep; last whorl narrow; undulating growth lines. callus slightly marked, extending past aper· Hypotype: MPEF-Pl-249; height 17.5 mm, ture, but not reaching to suture. Lip weakly width 7.2 mm. curved. Columella barely concave, with one Locality: Puerto Piramide, Valdes Peninsula, wide basal plication and a second very marked Chubut Province, Argentina. plication above dorsum of shell, separated from the first by a conspicuous groove. Discussion: We associate this example Hypotype: MPEF-Pl-251; height 6 mm, width with the species 0. uretai, because of the 3mm. presence of five to six fine columellar pli Locality: Puerto Piramide, Valdes Peninsula, cations, differing in their configuration ~ Chubut Province, Argentina. being thinner, and presenting an aperture less wide than the living form, but we con Discussion: This species is living from sider it a valid ancestor to the latter. This the coast of the Province of Buenos Aires is the first record of this species from the to the Golfo Nuevo. It has been recorded Entrerriense Formation of South America. from the Pampeano of Puerto Militar, Previously there has been noted from this Ihering (1907) citing ·it as Oliuancillaria formation 0 . prisca and 0 . urceus (Roding, auricularia plata, and from the 1798), cited as 0 . brasiliana, by Nieves de Interensenadense of Punta Piedras. The Medina (1962) from the well at Chuy fossil examples differ from the living ones (Uruguay), at 124 meters depth. only by the greater posterior expansion of the callus, which reaches tci the suture. Subfamily OLIVELLINAE This is the first report of this species from Genus OLIVELLA Swainson, 1831 the Entrerriense Formation of South OLIVELLA aff. 0. PUELCHA (Duclos, 1840) America. Plate 6, figure 10 Description: Shell very small, conical, elon OLIVELLA FRACASSOENSIS, n. sp. gated, extended, spire sharp; whorls high and Plate 6, figure 11 narrow. Aperture extended posteriorly, expand Description: Shell small, elongated and slen· ing in advance. Sutural channel. deep. der, spire very high and pointed; sutures deep; Columella with only one anterior plication, sep whorls of spire convex. Body-whorl correspond· arated by a groove of the basal plication. Well ing to 87% of total height; aperture enlarging developed callus reaching suture, without cov very little anteriorly, its length representing ering spire. Ornamented with growth lines. 4 7% of total height. Callus reaching sutural Fasciole well marked. channel, without covering spire. Only one col Hypotype: MPEF-Pl-250; height 8 .5 mm, umellar plication, separated from basal plica width4mm. tion by a groove. Columella rather concave due No. 2 Entrerriense Gastropoda 95

to presence of conspicuous indentation begin Formation); while from the marine ning at siphonal canal. Fasciole inconspicuous Miocene of southern Brasil they cited and smooth. Prunum prunum (Gmelin, 1791). This is Holotype: MPEF-PI-252; height 9 mm, width the first record of the genus Prunum in 3.2 mm. Body-whorl: length 7.8 mm. Aperture: the Entrerriense Formation of Argentina. length 4.2 mm. Type locality: Playa Fracasso, Valdes Penin Subclass HETEROBRANCHIA sula, Chubut Province, Argentina. Order ALLOGASTROPODA Derivatio nominis: Named for the type locali Superfamily PYRAMIDELLOIDEA ty. Family PYRAMIDELLIDAE Subfamily TURBONILLINAE Discussion: Olivella fracassoensis, n. sp., Genus TURBONILLA Risso, 1826 differs from 0. puelcha, in having only one TuRBONILLA URUGUAYENSIS Pilsbry, 1897 columellar plication; it is much higher and Plate 7, figure 1 narrower, with less expansion of the aper Description: Shell turriform, slender, strong. ture anteriorly, spire much higher, and Ten almost straight whorls, with conspicuous columellar indentation. It differs from 0. sutures; first whorls smooth, later whorls with plata by having one very wide basal plica fine oblique ribs, on last whorls these becoming tion and a second plication over the dor heavier, tabulate, and vertical. Aperture sub sum of the shell; it has a lower spire and quadrate, with columella practicaly vertical, narrower body-whori. Olivella carolinae simple and rather tilted toward base. Base only Gardner, 1948, has an ovoid-conical form, with growth lines. being of much greater size (16.2 mm in Hypotype: MPEF-PI-254; height 7.7 mm, height), the aperture is wider anteriorly, width 2.1 mm. the spire proportionally lower, the body Locality: Puerto Piramide, Valdes Peninsula, whorl much wider and lacking the col Chubut Province, Argentina. umellar indentation. Discussion: The fossil specimen presents Family MARGINELLIDAE the conchological characters of the living Genus PRUNUM Herrmansen, 1852 species. Figueiras and Broggi (1971-73) PRUNUMsp. list Turbonilla sp. from the Camacho Plate 6, figure 12 Formation of Uruguay, undoubtedly Description: Shell medium-sized, elongate, assignable to the subgenus Pyrgiscus. relatively oviform; spire low, aperture apparent From the marine Miocene of southern ly narrow, outer lip destroyed. Columella Brasil is listed for the genus Pyramidella straight, with four heavy plications; ·columellar Lamarck, 1799, P candida Morch, 1875, callus appearing to reach only to middle of and for the genus Turbonilla Risso, 1826, aperture; siphonal canal destroyed. T. (Pyrgiscus) interrupta Totten, 1835. Hypotype: MPEF-PI-253; height 22 mm, Brown and Pilsbry (1913) proposed as width9mm. forms representing the genus Turbonilla Locality: Puerto Piramide, Valdes Peninsula, from the Gatun Formation (Late Miocene), Chubut Province, Argentina. T. bartschiana and T. gatunensis. This is the first record of the genus Turbonilla in Discussion: The example described is the Entrerriense Formation of Argentina. unique, being found almost destroyed. We tentativaly place it in the genus Prunum Subgenus PYRGISCUS Philippi, 1841 because of the apparently low spire, the TURBONILLA (PYRGISCUS) sp. lip begining below the spire, with a sharp Plate 7, figure 2 angle, and with four columellar plications. Description: Shell very small, thin, with 9 1/2 Castellanos (1967) lists Marginella convex whorls; sutures conspicuous; first prunum (Gmelin, 1791) from the Antilles, whorls smooth, the following whorls with fine Brasil, Uruguay, and Argentina, as far aXial ribs, on last whorls these becoming some south as Gplfo San Matias. Figueiras and what heavier, with addition of heavy spiral Broggi (19!11-1973) cited Prunum pruni cords. Base with strong spiral striations. formis (lhering, 1907) from the Barranca Aperture subquadrate; columella almost de San Pedro, Uruguay (Camacho straight. 96 Tulane Studies in Geology and Paleontology Vol. 30

Hypotype: MPEF-PI-255; height 5 mm, width Quequen (Prov. Bs. Aires), has the spire 1.lmm. more elevated, the aperture beginning Locality: Puerto Piramide, Valdes Peninsula, below the ·shoulder of the body-whorl is Chubut Province, Argentina. wider and lacks the anterior expansion. According to the author there are fine spi Discussion: This unique example is ral striations. This is the first record for almost complete, but in bad condition. It is the Entrerriense Formation of South characterized by the heavy spiral orna America. mentation on the last whorls; therefore, it is assigned with doubt to the subgenus Family RETUSIDAE Pyrgiscus. From the marine Miocene of Genus VOLVULELLA Newton, 1891 Brasil T.(P.) interrupta Totten, 1835, has VOLVULELLA aff. V. PHOINICOIDES been recorded; consequently, this is the (Gardner, 1937) first record from the Entrerriense Plate 7, figure 4 Formation of Argentina. Description: Shell of small size, cylindrical, very thin; height three times width. Apical Subclass OPISTOBRANCHIA spine short, twisted, leaving a depresion Order CEPHALASPIDEA between spine and shell. Outer lip thin, anteri Superfamily PHILINOIDEA? orly expanded forming a flange. Strong columel Family SCAPHANDRIDAE lar callus. Microscopically (X 40) no ornamenta Genus ACTEOCINA Gray, 1847 tion seen. ACTEOCINA PIRAMIDESIA, n. sp. Hypotype: MPEF-Pl-257; height 3 mm, width Plate 7, figure 3 1.1 mm. Description: Shell very small, slender, sub Locality: Puerto Piramide, Valdes Peninsula, cyliiidrical. Spire low, visible only in apertural Chubut Province, Argentina. view, dorsally only the protoconch visible, lack ing in this example. Posterior margin of lip very Discussion: This specimen is the only elevated, reaching same height as spire, lateral e,xample obtained to date. From the con margin par.allel to body-whorl, with aperture chological characters it is near V. phoini very narrow; anteriorly lip opening widely. Base coides, especially in its apical characters of columella broken; therefore, plications and its measurements. It differs by the unknown. Exteriorly smooth under microscope structure of the anterior end, with a con (X40). spicuous columellar callus. This is the Holotype: MPEF-PI-256; height 6 mm, width first record of the genus Volvulella from 3mm. the Entrerriense Formation of South Type locaiity: Puerto Piramide, Valdes America. Peninsula, Chubut ProVince, Argentina. Derivatio nominis: Named for the type locali Subclass PULMONATA ty. Order STYLOMMATOPHORA Suborder ELASMOGNATHA Discussion: Acteocina piramidesia, n . Superfamily SUCCINEOIDEA sp., differs from A. bullata (Kiener, 1834), Family SUCCINEIDAE living in the West Indies, which has the Subfamily SUCCINEINAE lip straight, without anterior expansion, Genus SUCCINEA Draparnaud, 1801 the aperture wider and spiral striations as SUCCINEA MERIDIONALIS d'Orbigny, 1846 ornamentation. Acteocina elachista Wood Plate 7, figure 5 ring, 1970, has a higher spire, the aper Description: Shell diminutive, thin and frag ture beginning below the shoulder of the ile, translucent, imperforate; consisting of three body-whorl is much wider and without convex whorls, the last representing more th~ anterior expansion. Acteocina rusa two-thirds of total height; spire short with lin Gardner, 1937, has a higher spire, the ear sutures, deep and inclined. Aperture elon '' aperture begins below the shoulder of the gate-oval, wider anteriorly, holoperistomate. whorl; it is much wider, lacks an anterior Exteriorly smooth, with only growth lines .. expansfon, and is ornamented with axial Hypotype: MPEF-Pl-258; height 7 niin, width striations. Acteocina candei (d'Orbigny, 6 mm. Body-whorl: length 6 mm. 1842), living from the Antilles to Puerto Locality: Puerto Piramide, Valdes ~eninsula,

\~ No.2 Entrerriense Gastropoda 97

Chubut Province, Argentina. V. LITERATURE CITED Discussion: The presence of a pulinonate BEQUAERT, J .C., 1943, The genus Littorina in gastropod is not new to the Tertiary of the Western Atlantic: Johnsonia, v. 1, no. 7, Patagonia. Ihering (1907) mentioned for p. 1-27, pls. 1-7. the Entrerriense of Parana, as well as for BORCHERT, A., 1901, Die Molluskenfauna und the gray sands of the Rionegrense, the das Alter der Parana-Stufe: Neues Jahrb. genera Strophocheilus, Chilina, Ampul Min., Geol., Pal., v. 14, p. 5-78, pls. 5-10. laria, Diplodon and Corbiculella; also in BRAVARD, A., 1858, Monografia de los terrenos that work are described species of the marinos terciarios de las cercanfas de genus Littorina from waters typically Parana: Reprinted in Anal. Mus. Nae. Buenos brackish. But this is the first mention of Aires, v: 3, p. 45-94 (1883-1891). the genus Succinea from the Entrerriense BROWN, A.P., and H.A. PILSBRY, 1913, Fauna Formation of South America. We have no of the Gatun Formation, Isthmus of Panama doubts with respect to the stratigraphic - II: Acad. Nat. Sci. Philadelphia, Proc., v. 64, position of this example, having been p. 500-519, pls. 22-26, 5 text-figs. obtained in depth and sieved from the sed BRUNET, R.F.J., 1995, New species of Mollusca iments. from the Entrerriense Formation (Upper

2 3 1 5

PLATE 7 Figures Page '· 1. TURBONILLA URUGUAYENSIS Pilsbry, 1897 ...... 95 MPEF-PI-254 (hypotype); height 7.7 mm, width 2.1 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 2. TURBONILLA (PYRGISCUS) sp...... 95 MPEF-PI-255 (hypotype); height 5 mm, width Ll mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 3. ACTEOCINA PIRAMIDESIA, n. sp...... 96 MPEF-PI-256 (holotype); height 6 mm, width 3 mm. 4 Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 1 4. VOLVULELLA aff. V. PHOINICOIDES (Gardner, 1937) ...... 96 MPEF-PI-257 (hypotype); height 3 mm, width 1.1 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 5. SUCCINEA MERIDIONALIS d'Orbigny 1846 ...... 96 MPEF-PI-258 (hypotype); height 7 mm, width 6 mm. Locality: Puerto Piramide, Valdes Peninsula, Chubut Province, Argentina. 98 Tulane Studies in Geology and Paleontology Vol.30

Miocene) of Chubut Province, Argentina, and HODSON, F., 1926, Venezuelan and Caribbean species not previously reported from this for Turritellas: Bulls. Amer. Paleontology, v. 11, mation, Part I, Gastropoda and Scaphopoda: no. 45, p. 1-50, pis. 1-30. Tulane Stud. Geol. Paleont., v. 28, nos. 1-2, p. !HERING, H. von, 1897, Os Molluscos dos ter 1-56, pis. 1-5, 1 text-figure. renos Terciarios de Patagonia: Rev. Mu s. CASTELLANOS; Z., 1967, Catalogo de los Paulista (Sao Paulo), v. 2, p. 217-382, pis. 1·7. moluscos marinos bonaerense: Anal. Com. !HERING, H . von, 1907, Les Mollusques fos Inv. Cient. (La Plata), v. 8, 365 p., 26 pis. siles du Tertiaire et du Cretace Superieur de CLENCH, W.J., and R.D. TURNER, 1964, The !'Argentine: Anal.' Mus. Nae. Buenos Aires, v. Subfamilies Volutinae, Zidoninae, 14 (Ser. 3, v. 7), p. 1-611, pis. 1-18. Odontocymbiolinae, and Calliotectinae in the MARWICK, J ., 1957, New Zealand genera of Western Atlantic: Johnsonia, v. 4, no .. 43, p. Turritellidae, and species of Spirocolpus: 129-180, pis. 80-114. New Zealand Geo!. Surv., Paleont. Bull. 27, p. CLOSS; D., 1970, Estratigrafia da Bacia de 5-55, 5 pis., 5 text-figs. Pelotas, Rio Grande do Sul: Iheringia, Geol., MEDINA, N.P. , 1962, Notas ·sobre moluscos de no. 3, p. 3-76, 11 figs. Edad Entrerriana, procedentes de una per· DALL, W.H., 1882, On certain Limpets and foraci6n en el Chuy (Dto. de Rocha) (Uru· Chitons from the deep waters of the eastern guay): Rev. Mus. Arg. Cienc. Nat. United States: U.S. Natl. Mus., Proc., v. 4, p. "Bernar.dino Rivadavia," Cienc. Zool. , v. 8, no. 400-414. 16, p. 213-212, pl. 1. FIGUEIRAS, A., and J . BROGGI, 1971-1973, ORTMANN, A. , 1900', Synopsis of the collec· Estado actual de nuestros conocimientos tion~ of invertebrate fossils made by the sobre los moluscos f6siles del Uruguay, parte Princeton Expedition to southern Patagonia: III (cont.): Soc. Malac. Uruguay, Comun., v. Amer. Jour. Sci., v. 10, p. 368-381. 3,, no. 21, p. 131-154; no. 23-24, p. 203-240. PHILIPPI, R.A., 1893, Descripci6n de algunos FIGUEIRAS, A, and J. BROGGI, 1976, Nuevas f6siles Terciarios de la Republica Argentina: especies de gastropodos marinos de la Anal. Mus. Nae. Chile, Min. Geol. Paleont., Formaci6n Camacho (Entrerriana) (Mioceno 16 p., 4 pis. Superior de Uruguay): Mus. Nat. Hist. VOKES, E.H., 1991A, Collecting Trophons in Montevideo, Comun. Paleont., v. 1, no. 6, p. Argentina; part I, Tierra de! Fuego: American 135-147, pl. 1. Conchologist, v. 19, no. 1, p. 7-10, 9 te~t-figs., FIGUEIRAS, A., and J . BROGGI, 1986, Nuevas lmap. especies de gastr6podos marinos de la VOKES, E.H., 1991B, Collecting Trophons in Formaci6n Camacho (Mioceno Superior de Argentina; part II, the Valdes Peninsula: Uruguay), parte II: Soc. Malac. Uruguay, American Conchologist, v. 19, no. 2, p. 8-11, 1 Comun., v. 6, no. 50, p. 341-353, pl. 1 pl., 9 text-figs., 1 map. GUILLAUME, L., 1924, Essai sur la classifica WOODRING, W.P., 1957-1982, Geology and tion des Tt:rrritelles, aussi que sur leur evolu paleontology of Canal Zone and adjoining tion et leurs migrations, depuis le debut des parts of Panama: U.S. Geo!. Surv. , Prof. temps tertiaires: Bull. Soc. Geol. France, Paper 306 (in 6 parts), 759 p., 124 pis., 5 text· (Ser.4) v. 24, p. 281-311, pis. 10, 11, 33 text figs., 1 map. figs.