Baseline Survey of the Terrestrial Invertebrate Fauna of Barrow Island

RECORDS OF THE WESTERN AUSTRALIAN MUSEUM 83 013–112 (2013) SUPPLEMENT

Baseline survey of the terrestrial invertebrate fauna of Barrow Island

Jonathan D. Majer1, Shae K. Callan1, Karl Edwards1,2, Nihara R. Gunawardene1 and Christopher K. Taylor1

1 Curtin Institute for Biodiversity and Climate, Curtin University, PO Box U1987, Perth, Western Australia 6845, Australia. Email: [email protected]

2 Chevron Australia Pty Ltd, 256 St Georges Terrace, Perth, Western Australia 6000, Australia.

ABSTRACT – Barrow Island is Western Australia’s second largest offshore island and its flora and fauna have been able to evolve without major human disturbances. Chevron Australia Pty Ltd and its Joint Venture Participants made an application to construct a plant to liquefy natural gas on the island in 2001. One of the conditions under which approval was granted was the implementation of a rigorous biosecurity effort to ensure that no non-indigenous species (NIS) are introduced or allowed to establish on the island. To fulfil this condition it was first necessary to characterise what was already present on the island. A series of surveys have been performed using a purpose-designed sampling protocol in order to provide baseline data on the existing terrestrial invertebrates on Barrow Island. A total of 1,873 morphospecies were sampled but subsequent surveys and taxonomic developments have increased the count to 2,397. This compares with an estimated species richness of 2,481 terrestrial invertebrate species on the island. Composition of the fauna varied considerably between the wet and dry seasons and between years, even when samples were taken during the same month. Composition also varied with distance from the coast, which may be associated with soil type and vegetation association. Twenty five non-indigenous species and seven putative non-indigenous species have been found, all of which are believed to have been present prior to commencement of the Gorgon Gas Development project.

KEYWORDS: species inventory, biodiversity survey, non-indigenous species, offshore island.

INTRODUCTION causing undesirable environmental consequences (Government of Western Australia 2007). In other Barrow Island is Western Australia’s second words, if a non-indigenous species was to arrive on largest offshore island. Separated from the island, there should be at least an 80% certainty the mainland for 8,000 years, uninhabited by that it would be detected. indigenous people and un-colonised by Europeans, Barrow Island’s flora and fauna have been able Although this was a straightforward undertaking to evolve without major human disturbances. for plants and vertebrate animals because these Discovery of large gas deposits in the north-west components of the biota are well known on Barrow shelf prompted Chevron Australia Pty Ltd and its Island (Chevron Texaco Pty Ltd. 2005; Moro and Joint Venture Participants (see acknowledgements) Macauley 2010a, b and c), lack of data made this a to make an application to construct a plant to problematic task for the invertebrates. There had liquefy natural gas on the island in 2001. A number been a few taxon-specific surveys performed in the of conditions accompanied final approval of the 1970s through to the 1990s (Solem 1997—land snails; Gorgon Project. Included amongst these was the Perry 1972—termites; Smithers and Butler 1985— need to implement a rigorous biosecurity effort to dragonflies; Smithers 1984a—psocopterans; Smithers ensure that no non-indigenous species (NIS) are and Butler 1983—butterflies; Smithers 1984 b, 1988— introduced or allowed to establish on the island. neuropterans) but most of these were not exhaustive, Furthermore, the conditions required the design having been carried out during short, single visits and implementation of a detection program with a to the island. Recognising this deficiency, Chevron detection power of 0.8, or an appropriate alternative contracted entomologists at Curtin University power, for detecting NIS at an early enough stage to carry out a baseline survey of the terrestrial that eradication could be attempted without invertebrate fauna of Barrow Island (BWI). 14 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR

FIGURE 1 Monthly total rainfall and mean monthly minimum and maximum temperatures for the period covering the Pilot Study (green arrows), the Gas Plant survey (red arrows) and the Non-Indigenous Species surveys (black arrows).

Prior to commencing the survey, it was necessary baseline species list of indigenous and non- to design a sampling protocol that was cost- indigenous species inhabiting Barrow Island which effective, while still being adequate to sample the was subsequently used in conjunction with the species present. A transect-based approach formed potential NIS threats to the island to develop the the basis of the protocol, and built upon a design surveillance program, described in the papers by that had been used for many years in rehabilitated Whittle et al. (2013 a, b). mine sites for sampling invertebrates from all strata; ranging from the litter layer up to the trees PILOT STUDY (shrubs in the case of Barrow Island) (Allen 1989). The pilot study was performed twice, once Once the sampling protocol had been agreed upon, during the week beginning 22 April 2005 and once two separate, but complementary surveys were during the week of 1 May 2005, immediately after performed. The Gas Plant (GP) survey targeted a significant rain event (Figure 1). Seven native the area where the gas processing plant was to vegetation sites and three disturbed sites (airport, be built and was designed to sample the species old rubbish dump and accommodation camp) were most likely to be affected by the development. sampled. Sampling was based upon a 90 m transect This positioning of the transects was designed to and included pitfall trapping, herb and shrub enable us to elucidate the major determinants of vacuum sampling (10 swathes at 90o to the transect), the composition of the invertebrate fauna, including shrub beating (10 small trees or shrubs close to soil type, vegetation association, distance from the each pitfall trap), litter sampling using Winkler coast, and season. Paired transects were set up both sacks (Agosti et al. 2000) and hand collecting for close to and distant from the proposed gas plant in one hour during both daytime and night. Pitfall order to provide a basis for ecological monitoring trap configurations included small (43 mm internal of potential edge effects. The second survey was diameter) versus large (110 mm) traps, 10 versus 20 designed to detect whether any non-indigenous small diameter traps, and 5 versus 10 days of small species (NIS) already exist on the island and diameter trapping. targeted areas of current or previous disturbance – The resulting material was sorted to areas where NIS are most likely to be found. morphospecies and individual groups were sent The results of the two surveys have been to the relevant taxonomists for determination, comprehensively reported on by Callan et al. (2011). resulting in a list of approximately 550 The results are summarised here and findings have morphospecies. Then, for selected groups, the been updated in the light of further taxonomic number of species per sampling technique was information being provided. This formed the summed, the estimated total number of species TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 15 was calculated from species accumulation curves, Finally, invertebrates were hand collected from a the efficacy of each method was compared, and the range of microhabitats in a 100 m radius around complementarity between the different methods each transect for 1 person-hour during the day and was assessed. From this, a standardised sampling 2 person-hours during the night. Light traps were protocol was agreed upon, and is summarised in also run at a third of the GP and NIS sites, selected Figure 2. to maximise their spread across the island. The finalized protocol was once again based on a 90 m transect, with 10 small-diameter pitfall traps GAS PLANT (GP) SURVEY with ethylene glycol preservative, run for 5 day periods following a 12 hour digging-in period. Leaf Twelve sites were sampled during this survey, litter was taken from close to each pitfall trap and representing coastal dunes, floodplain vegetation, coarse-sieved to remove large litter fragments. The and a range of limestone sites ranging from flats resulting material was placed in Winkler sacks and to ridges. Wherever possible, sites of comparable invertebrates were extracted over a 4-day period. vegetation were in duplicate pairs, with one site Three vegetation vacuum swathes were taken in 30 proximal to the future gas plant and the other some x 15 m areas parallel with the sampling transect. distance away (Figure 3a). This was designed to To complement this, 10 small trees or shrubs were characterise the fauna close to the areas that will sampled by beating with a stout stick over a net. be disturbed and to enable monitoring of any spill-

FIGURE 2 Standardised sampling protocol used for the GP and NIS surveys. Further details are explained in the text; design of the protocol was modiﬁed in some of the NIS sites to allow for irregular shape of the areas and the existence of structures of interest, such as buildings. 16 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR

,19(57(%5$7(678'<6,7(6 ,19(57(%5$7(678'<29(59,(:

m 7HUPLQDO7DQNV m

%DUURZ,V %DUURZ,V

0HWUHV .LORPHWUHV &RRUGLQDWH6\VWHP*'$0*$=RQH &RRUGLQDWH6\VWHP*'$0*$=RQH

5HI *25B$B5HY'DWH6HS&$,&,.4 5HI *25B%B5HY 'DWH6HS&$,&,.4

*3 *3 *3

*3 1RUWK:KLWH V +''6LWH %HDFK 7 3 /1*3ODQW 2QVKRUH 3LSHOLQH *3

:$3(7/DQGLQJ *3

*3; $GPLQLVWUDWLRQ

*3 8WLOLWLHV *3 *3 7HUPLQDO7DQNV

:DWHU'LVSRVDO6LWH /1* 7RZQ 3ODQW 3RLQW

$GPLQLVWUDWLRQ 8WLOLWLHV

:$2LO%DVH &RQVWUXFWLRQ 9LOODJH :$2LO&DPS

&& $LUSRUW

&& /(*(1' &RQVWUXFWLRQ ,QYHUWHEUDWH6XUYH\7UDQVHFW 9LOODJH (QG3RLQW 6WDUW3RLQW /(*(1' )HHG*DV3LSHOLQH5RXWH )HHG*DV3LSHOLQH5RXWH ,QIUDVWUXFWXUH ,QIUDVWUXFWXUH

FIGURE 3 Location of the (A) undisturbed (GP) and (B) disturbed (NIS) survey sites where invertebrates were sampled using the standardised sampling protocol. Note some of the infrastructure positions have changed since the time when these surveys were undertaken. over impacts on the environment by comparing by species matrices for each taxonomic group are proximal and distant sites within each vegetation provided in the electronic supplements to Callan type, the identity of which can be obtained by et al. (2011). Vouchers of most of the species are consulting Appendix C1 of the EIS/ERMP for the housed in the Entomology Laboratory at Curtin Gorgon Development on Barrow Island (Chevron University and WA Department of Agriculture Texaco Pty Ltd. 2005). Sampling was performed and Food. They will be deposited in the Western using the protocol shown in Figure 2 and was Australian Museum upon cessation of the program. carried out twice, once in the fortnight beginning A non-metric multi-dimensional scaling (NMDS) 15 March 2006 at the end of the wet season and ordination, using Bray-Curtis matrices of non- again in the period beginning 25 September 2006 standardised data for the most frequently occurring at the end of the dry season. The pattern of rain species in the GP survey (350 taxa), plus the around these times is shown in Figure 1 and recent environmental variables sampling time, distance falls prior to the first sampling period had resulted from coast, soil type and vegetation type is shown in a flush of green vegetation, which was not so in Figure 4a. The wet season and dry season evident during the second period. samples are separated into two distinct clusters, A total of 1,123 morphospecies were collected whose species composition is significantly different during the two periods, comprising four Mollusca (ANOSIM global R = 0.979, P = 0.001). Within species, 165 Arachnida species, nine Malacostraca each cluster, the relative positioning of sites along species, 11 Chilopoda species, two Diplopoda axis 2 was reasonably similar (correlation of axis species, 34 Collembola species and 898 Insecta 2 scores in the two seasons r = 0.52; df =10, P = species. The full list of species encountered during 0.05) and was associated with distance from coast the GP survey is given in Appendix 1 and the sites (ANOSIM global R = 0.166, P = 0.030) and weakly TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 17

FIGURE 4 NMDS ordination of the GP sites (top) and the NIS sites (bottom), both of which were sampled twice. The left-hand cluster of the GP ordination delimits the March 2006 wet season and the right hand cluster delimits the September 2006 dry season samples; the left- and right-hand clusters of the NIS ordination delimit the May 2006 and May 2007 samples, respectively. 18 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR

TABLE 1 List of conﬁrmed (Status C) or putative (Status ?) non-indigenous species found during the pilot, GP, NIS and subsequent surveillance surveys carried out between 2005 and 2012. Those highlighted in bold were collected in the subsequent surveillance survey.

Order/Family Family/Subfamily Non-Indigenous Species of Invertebrate Status

Araneae Pholcidae Artema atlanta (Giant daddy long legs spider) C

Araneae Pholcidae Crossopriza lyoni (Daddy long-legs spider) C

Araneae Scytodidae Dictis striatipes (Spitting spider) C

Araneae Oecobidae Oecobius navus (Urban wall spider) C

Araneae Theridiidae Latrodectus hasseltii (Red-backed spider) ?

Blattodea Blattidae Periplaneta americana (American cockroach) C

Blattodea Blatellidae Supella longipalpa (Brown banded cockroach) C

Coleoptera Anobiidae Lasioderma serricorne C

Coleoptera Cleridae Necrobia ruﬁpes (Red-legged ham beetle) C

Coleoptera Dermestidae Attagenus sp. ?

Coleoptera Dermestidae Dermestes haemorrhoidalis C

Coleoptera Dermestidae Dermestes ater C

Coleoptera Languriidae Leucohimatium arundinaceum C

Coleoptera Tenebrionidae Tribolium confusum (Confused ﬂour beetle) ?

Collembola Hypogastruridae Hypogastrura sp. cf. vernalis ?

Collembola Isotomidae Isotoma sp. cf. viridis ?

Diptera Psychodidae Psychoda alternata C

Formicidae Cerapachyiinae Cerapachys longitarsus (Cannibal ant) C

Formicidae Formicinae Paratrechina longicornis (Black crazy ant) C

Formicidae Myrmicinae Cardiocondyla nuda (Sneaking ant) ?

Hemiptera Anthocoridae Montandoniola sp. ?

Hemiptera Aphididae Aphis craccivora (Black legume aphid) C

Hemiptera Aphididae Aphis gossypii (Cotton aphid) C

Hemiptera Miridae Nesidiocoris tenuis (Tomato mirid ) C

Isopoda Porcellionidae Porcellionides pruinosus (Delicate slater) C

Lepidoptera Pyralidae Plodia interpunctella (Indian Meal Moth) C

Lepidoptera Pyralidae Plutella xylostella (Diamondback moth) C

Psocoptera Liposcelidae Liposcelis bostrychophila C

Psocoptera Liposcelidae Liposcelis entomophila C

Psocoptera Psyllipsocidae Dorypteryx domestica C

Thysanoptera Thripidae Frankliniella schultzei (Tomato thrips) C

Thysanura Ctenolepismatidae Ctenolepisma longicaudata (Grey silverfish) C TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 19 with soil type (global R = 0.159, P = 0.057) but not were found during this survey and also during the significantly so with vegetation association (global various components of the surveillance program R = 0.078, P = 0.174). Although these variables are described by Whittle et al. (2013a) and implemented inevitably correlated, this indicates the importance by us. There are 25 confirmed non-indigenous of distance from coast and the progressively species and seven putative species, whose status is diminishing role of soil type and vegetation as uncertain. All of these species are believed to have determinants of faunal composition. been present prior to commencement of the Gorgon Gas Development project. The status of each NON-INDIGENOUS SPECIES SURVEY group was confirmed by consulting the literature, pest databases and by feedback from the relevant Twenty five disturbed sites were sampled during specialists for each taxonomic group. The putative this component of the survey. Sites were either species may well be cosmopolitan species that actively used (e.g., accommodation camp, barge have been spread by natural means, such as aerial landing, airport, warehouse, etc.), disused (e.g., plankton in the case of the Collembola species, or former workshop and administration areas) or may occur in any Australian environment once restored (e.g., former waste disposal areas or sites it is disturbed, as may be the case with the ant of old accommodation camps). All sites were chosen Cardiocondyla nuda. Others may be synanthropic because if any invertebrates had been previously species whose status as a native to Australia introduced, they are more likely to be found in remains a matter of debate, as is the case with the these areas of previous or current human activity. red-backed spider, Latrodectus haselltii (Griffiths et The NIS sites were more widely distributed across al. 2005). Of the confirmed non-indigenous species, the island (Figure 3b) than the GP survey sites several are well-known stored product insects, such but the same sampling protocol (Figure 2) was as most of the beetles and moths, and a number adopted, although sometimes modified to suit site are associated with buildings and other structures, conditions. Sampling was performed in the week which is the case for all of the spiders. beginning 6 May 2006 and was repeated in the week beginning 1 May 2007. These periods were originally chosen as being the optimal period for RICHNESS OF THE TERRESTRIAL sampling, as it follows on from the summer rain INVERTEBRATE FAUNA period. However, although this was the case during The combined results of the Pilot, GP and NIS 2006, hardly any rain fell during the entire period surveys give a total of 1,873 morphospecies of of 2007 (Figure 1). terrestrial invertebrates. If the sites and periods for A total of 1,468 morphospecies were sampled in the Pilot, GP and NIS sampling programs are taken the 25 sites, represented by eight Mollusca species, together and analysed for species accumulation 240 Arachnida species, 12 Malacostraca species, by the EstimateS computer package (Colwell 13 Chilopoda species, three Diplopoda species, 2000; Colwell and Coddington 1994), the Chao2 31 Collembola species and 1,161 Insecta species. estimator predicts a total terrestrial invertebrate The full list of species encountered during the richness of 2,481 species, indicating that the survey NIS survey is given in Appendix 1 and the sites is not complete. A similar analysis of those orders by species matrices for each taxonomic group are that were sufficiently diverse for the procedure provided in the electronic supplements to Callan et to be reliably performed enables a consideration al. (2011). to be made about the groups that were under- The NMDS ordination of the 13 NIS sites that sampled (Table 2). In this case it was only possible were sampled in both 2006 and 2007 and using to perform the analysis on the GP and NIS data the 316 most frequently occurring taxa are shown and only on taxa that were diverse enough to in Figure 4b. The two sampling periods separate analyse; the numbers therefore do not exactly out into two clearly demarcated clusters which match those from the global analysis. The analysis are significantly different (global R = 0.687, P = indicates that most taxa had not been completely 0.001), indicating the high degree of variation in sampled by GP and NIS surveys. Notable examples fauna between years, probably due to differences include Acarina (taxonomic verification had not in the pattern and amount of rainfall. Although been completed at the time), Mantodea (possible environmental variables were included in the method bias), Neuroptera (reason unknown), analysis, no influences on the fauna of disturbance Heteroptera (unavailability of taxonomists), type or time since rehabilitation could be detected. Coleoptera (high diversity), Diptera (high diversity) This may well reflect the large variation in the type and Hymenoptera (samples initially only identified of disturbance and also the fact that rehabilitated from one sampling period). However, subsequent areas were extremely heterogeneous in their sampling has increased the species count, with composition and stage of rehabilitation. values for most taxa rising to, or even exceeding, Table 1 shows the non-indigenous species that the estimated total of species (Table 2). 20 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Percentage of estimated species sampled to date he Chao2 estimator, plus plus he Chao2 estimator, n. Species and morphospecies sampled to date Percentage of estimated species sampled by GP and NIS survey Chao2 estimate of total species 15 40 37.5 19 47.5 Species and morpho-species sampled in baseline survey Thysanura 9Odonata 4Phasmatodea 6Orthoptera 91Mantodea 10Blattodea 23Isoptera 14Psocoptera 17 12Thysanoptera 39 4Homoptera 6 94Heteroptera 106 27Neuroptera 16 147 75.0Coleoptera 278 23Diptera 169 16 100.0Lepidoptera 179 18 100.0 85.8Hymenoptera 269 43 62.5 100.0 142 87.5 125 94.4 170 381 90.1 11 256 8 240 66.2 7 498 99 21.6 86.5 73.0 10 66.0 23 74.5 18 91.7 54.0 25 200.0 46 116.7 93.4 108 62.5 28 100.0 156 388 112.5 201 138.9 187 106.9 373 76.1 22.4 91.8 101.8 78.5 77.9 74.9 Collembola 42 48 87.5 73 152.1 Isopoda 14 14 100.0 16 114.3 ScorpionidaPseudoscorpionida 13Acarina 53 4Araneae 202 16 112 214 4 81.3 47.3 94.4 100.0 13 141 212 8 81.3 125.9 99.1 200.0 Pulmonata 9 15 60.0 9 60.0 the number of species sampled to date. The percentagethe baseline survey to date are also show The of predicted species sampled by the number of species sampled to date. t and as predicted by groups during the GP and NIS baseline survey Numbers of morphospecies sampled from the most species-rich Insecta Entognatha Malacostraca Myriapodous groups ARTHROPODA Arachnida MOLLUSCA Gastropoda Higher Taxon Lower Taxon TABLE 2 TABLE TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 21

TABLE 3 Dichotomous key for new species additions to the baseline (Based on specimens that are mature and of the appropriate sex for identiﬁcation to species).

1. Specimen aligns with species in the Curtin Specimen is not identifiable to species BWI Reference Collection ...... 2 (possibly new species) ...... 12

Specimen does not align with species in 8. Any available information is sought on Curtin BWI Reference Collection ...... 3 species from taxonomist and from published 2. Residence status of species on BWI already sources: known – Species is a known non-native species for If only one or two specimens of species in Australia: Curtin BWI Reference Collection: Chevron informed immediately. specimens bottled or pinned and added Species is a known native species for to collection Australia ...... 9 Numerous specimens available in Curtin BWI 9. Species known from northern Western Reference Collection: Australia ...... 10 specimens kept together with bulk Species not known from northern Western specimens in vial and stored in permanent wet collection. Australia ...... 11 10. Species is vagile, disperses easily, and likely 3. Residence status of species on BWI unknown: to be detected by collection method and Taxonomist for Order or Family available and sampling regime: willing to take specimens ...... 4 most likely naturally dispersed to BWI No taxonomist available ...... 5 from mainland. Added to baseline, 4. Taxonomist identifies specimen(s) as included in annual report. previously collected on BWI ...... 6 Species is sedentary and/or difficult to detect: Taxonomist identifies specimen(s) as not most likely native to BWI but not previously collected on BWI ...... 7 collected previously. Added to baseline, 5. Specimen identifiable to species by in-house included in annual report. parataxonomist ...... 9 11. Species is vagile, disperses easily, and likely Specimen not identifiable to species (no keys to be detected by collection method and available): sampling regime:

specimen designated morphospecies most likely human assisted. Chevron number and stored for future informed immediately. confirmation of species if taxonomist Species is sedentary and/or difﬁcult to detect- or taxonomic keys become available. Provisionally added to baseline, included most likely native to BWI but not in annual report. collected previously. Added to baseline, 6. Specimen is returned to Curtin and included in annual report. incorporated into collection ...... 2 12. Genus/Family known from Australia: Specimen is not returned to Curtin: species likely to be native to BWI but taxonomist queried for identification not collected previously, designated notes/photographs if Curtin BWI morphospecies number. Added to Reference Collection does not contain a baseline, included in annual report. representative specimen Genus/Family not known from Australia 7. Specimen is identiﬁable to species ...... 8 ...... 11 22 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR

SURVEILLANCE collected in previous trapping efforts (NIS Survey) that focussed on areas external to the buildings. Based on five exemplar invasive invertebrates species, Barrow Island was classed into three The surveillance programme is now in its zones of invasion susceptibility. This was based on third year of implementation, with a field survey analyses of potential entry points, establishment undertaken at the six sites each summer and winter and spread of these exemplars onto the island. season (March and September). This was based on the significant seasonality observed in the GP Risk Zones were identified where the likelihood survey and to ensure that natural variability in of entry and suitability of the environment for invertebrate populations is adequately sampled. establishment were medium to high probability. To date, the surveillance has increased the record These zones fell predominantly within six sites on of indigenous invertebrate species to 2,397. Each the island; these were the Accommodation Camp, additional species collected was scrutinised for Airport, Barge Landing, Horizontal Directional its potential to be a NIS and expert advice was Drilling site, Materials Off-loading Facility and sought to assess whether its presence on the island finally the Gorgon Gas Development Footprint. had been assisted by Gorgon’s activities. At the The surveillance methods were chosen from the time of writing no specimen has been identified suite of methods used in the baseline surveys in as a potential introduction, and new species have accordance with suggestions made by a Quarantine either been placed as ones that were not detected Expert Panel. These analyses have been reported in in the baseline surveys or as vagrant species from Whittle et al. (2009; 2013b) and Barrett et al. (2010) the mainland that are easily transported by strong and are summarised in Whittle et al. (2013a). winds to BWI. Within the first year of construction (2009– 2010), after ministerial approval was granted, a DISCUSSION preliminary survey was conducted to test the efficacy of the surveillance program (Seismic A number of developments have occurred since Survey 2009–2010) and to reinforce the knowledge the 2005–7 surveys, notably further determinations of established NIS invertebrates on the island (Built from taxonomists, a dedicated survey of Collembola and the implementation of the surveillance Environment Survey 2010; Collembola Survey 2010). program. Figure 5 shows the accumulation This yielded new additions to the baseline species of species through the progression of these list, as well as hitherto undetected non-indigenous developments, reaching a current total of 2,397 as species (Greenslade 2010). Previously undetected of mid-2013. The slowing of accumulation of species NIS were identified as synanthropic species (Table is commensurate with the asymptote calculated by 1) that inhabited the built environment (e.g., some the Chao2 estimator, suggesting that this may be of the stored-products insects) and hence were not close to the actual value for the island. One of the issues that the Gorgon Project faces is the determination of whether a newly discovered species on the island is a native species hitherto undocumented for the island or a species that has been introduced as a result of Gorgon related activities. Terrestrial invertebrates are comprised of a variety of life forms and life histories, making them difficult to completely assess during surveys. No matter how many samples or collection methods are employed, a variety of issues may impede the collection of any one species at any one time. Some have cryptic life histories and hence are not readily captured, while others display large variances in seasonal populations and hence may not be captured in one given year or season. Also, Barrow Island is an offshore island less than 60 km from the coast and is in an area that experiences annual cyclonic events. Larger, strong fliers can be assisted by strong winds, or smaller, planktonic FIGURE 5 Increment in number of morphospecies forms can drift across the ocean in large enough through the various surveys and populations to colonise an island. All these factors taxonomic developments on Barrow make the determination on the native status of a Island that have taken place between newly encountered species quite difficult. We have 2005 and mid-2012. TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 23 developed a dichotomous key of questions to aid (2000). Biodiversity of the Southern Carnarvon Basin. determining the native status of a species (Table 3). Records of the Western Australian Museum, Supplement 61: 595 pp. In terms of duration, range of methods used, and taxonomic breadth, this is probably the most Callan, S.K., Majer, J.D., Edwards, K. and Moro, D. (2011). Documenting the terrestrial invertebrate fauna of comprehensive terrestrial invertebrate survey Barrow Island, Western Australia. Australian Journal of ever undertaken in Australia. Whether or not Entomology 50: 323–343. the island’s species richness is high or low in Colwell, R.K. (2000). User’s guide to Estimates 6. URL comparison with mainland sites is difficult to http://viceroyeeb.uconn.edu/estimates. say. Although surveys exist for the Pilbara and Colwell, R.K. and Coddington, J.A. (1994). Estimating the Southern Carnarvon Basin (George et al. 2011; terrestrial biodiversity through extrapolation. Burbidge et al. 2000), the sampling methods and Philosophical Transactions of the Royal Society of London taxonomic coverage are not the same. However, Series B: Biological Sciences, 345: 101–118. our survey does provide important baseline Chevron Texaco Pty Ltd. (2005). Gorgon development on data for the terrestrial invertebrate fauna of an Barrow Island technical report: Flora and vegetation. arid Australian island and a Carnarvon Basin Report No. R03201. Chevron Texaco Pty Ltd.: Perth. ecosystem. This information can be used for George, A.S., McKenzie, N.L. and Doughty, P. (eds.) environmental monitoring and as a background (2011). A biodiversity survey of the Pilbara Region of for future surveillance programs. Yet the fact that Western Australia, 2002–2007. Records of the Western the differences in faunal composition between the Australian Museum, Supplement 78 (part 1): 311 pp. same months in different years can be as great as Government of Western Australia (2007). Ministerial the difference between seasons within a given year Environmental Statement No. 748, vol. 2009. http:// needs to be taken into account when considering www.gorgon.com.au/03-man_environment/pdf/ MIS%20748%20Complete.pdf. the temporal aspects of the program. Greenslade, P. (2010). Report of survey of Collembola on Barrow Island, June 6th to 10th 2010. Report to Chevron ACKNOWLEDGEMENTS Australia Pty Ltd: Perth. The Gorgon Project is operated by an Australian Griffiths, J.W., Peterson, A.M. and Vink, C.J. (2005). subsidiary of Chevron, Chevron Australia Pty Ltd, Molecular insights into the biogeography and species status of New Zealand’s endemic Latrodectus spider and is a joint venture of Chevron Australia Pty Ltd species L. katipo and L. atritus (Araneae: Theridiidae). (approximately 47 per cent), ExxonMobil (25 per Journal of Arachnology 33: 776–784. cent) and Shell (25 per cent), Osaka Gas (1.25 per Moro, D. and Macauley, I. (2010a). A guide to the reptiles cent), Tokyo Gas (one per cent) and Chubu Electric and amphibians of Barrow Island. Chevron Australia Pty Power (0.417 per cent). The authors are grateful to Ltd: Perth. Chevron Australia Pty Ltd for on-island support Moro, D. and Macauley, I. (2010b). A guide to the birds of with logistics, and the Gorgon Project Partners Barrow Island. Chevron Australia Pty Ltd: Perth. for financial support to undertake the project. Moro, D. and Macauley, I. (2010c). A guide to the mammals Karen Edward, Rebecca Severtson, Tim Lardner, of Barrow Island. Chevron Australia Pty Ltd: Perth. Dr Anita Lyons, Morgan Lythe, Natalie Randall, Perry, D.H. (1972). Some notes on the termites (Isoptera) Brad Scanlon, Adele Scarfone, Dusty Severtson and of Barrow Island and a check list of species. Western Callum Smithyman all assisted the project, either in Australian Naturalist 12: 52–55. the field and/or in the laboratory. Smithers, C.N. (1984a). The Psocoptera of Barrow and Boodie Islands, Western Australia. Entomologica REFERENCES Scandanavica 15: 21–226. Smithers, C.N. (1984b).The Neuroptera of Barrow and Agosti, D., Majer, J., Alonso, E. and Schultz, T. (eds.) nearby islands off the west coast of Western Australia. (2000). Ants: Standard methods for measuring and Australian Entomological Magazine 11: 61–68. monitoring biodiversity. Biological Diversity Handbook Smithers, C.N. (1988). Four additional antlion records Series. Smithsonian Institution Press: Washington from Barrow Island, Western Australia (Neuroptera: D.C. Myrmeleontidae). Australian Entomological Magazine Allen, N.T. (1989). A methodology for collecting 15: 2. standardized biological data for planning and Smithers, C.N. and Butler, W.H. (1983). The butterflies monitoring reclamation and rehabilitation programs. (Lepidoptera: Hesperioidea and Papilionoidea) of In: Majer, J.D. (ed.) Animals in primary succession: The Barrow and nearby islands, Western Australia. The role of fauna in reclaimed land: 179–205. Cambridge Western Australian Naturalist 15: 141–145. University Press: Cambridge. Smithers, C.N. and Butler, W.H. (1985). Dragonflies Barrett, S., Whittle, P., Mengersen, K. and Stoklosa, R. and damselflies (Odonata) from Barrow and nearby (2010). Biosecurity threats: the design of surveillance islands off the coast of Western Australia. Australian systems, based on power and risk. Environmental and Entomological Magazine 12: 9–12. Ecological Statistics 17: 503–519. Solem, A. (1997). Camaenid land snails from western and Burbidge, A.H., Harvey, M.S. and McKenzie, N.L. (eds.) central Australia (Mollusca: Pulmonata: Camaenidae). 24 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR

Records of the Western Australian Museum, Supplement indigenous species of invertebrates on Barrow Island. 50: 1461–1906. Records of the Western Australian Museum, Supplement Whittle, P.J., Barrett, S., Jarrad F., Murray J., Mengersen 83: 113–130. K., Hardie D., Nietrzeba A., Stoklosa R., Parkes J. and Whittle, P.J., Jarrad, F.C., Barrett, S., Martin, P., Majer, J.D., Majer J. (2009). Design of detection surveillance program Stoklosa, R. and Mengersen, K. (2013b). A method for for non-indigenous species of terrestrial invertebrates, designing complex biosecurity surveillance systems: plants and vertebrates on Barrow Island, Report to Detecting non-indigenous species of invertebrates on Chevron Australia Pty Ltd: Bruce, ACT. Cooperative Barrow Island. Diversity and Distributions DOI10.111/ Research Centre for National Plant Biosecurity. ddi.12056. Whittle, P.J., Jarrad, F.C. and Mengersen, K. (2013a). Design of the quarantine surveillance for non- MANUSCRIPT RECEIVED 10 DECEMBER 2012; ACCEPTED 20 MARCH 2013. TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 25 from the Other Collection veys out such as the Collembola out such veys carried 10001000 10001000 00001000 01101000 11111100 11000000 11000000 and during the subsequent surveillance program. Surveillance includes sur 010111010 110001110 010000100 111011010 111011110 101111110 010001000 000000001 000000001 000000001 veys beltianus interioris contrarius cf. cf. cf. sp. 1 0 sp. 1 0 sp. 2 1 sp. 3 1 sp. 4 0 sp. 5 1 Quistrachia barrowensis Rhagada convicta Stenopylis coarctata Gastrocopta mussoni Pupoides Pupoides Eremopeas Truncatella Isometroides “barrow” Lychas Lychas Lychas Lychas Lychas Aops “ops” Urodacus butleri Survey and the Built Environment Survey. Survey and the Built Environment Seismic Study (S1 and S2) and from the commencement of construction (C1–C4). Other collections include additional sur (S1 and S2) from the commencement of construction Seismic Study Taxonomic list of species sampled during the GP and NIS sur Taxonomic FamilyGASTROPODA Camaenidae Genus Species GP NIS S1 S2 C1 C2 C3 C4 Camaenidae cf. Hydrobiidae Unknown sp. 1 0 Helicodiscidae Pupillidae Pupillidae Pupillidae Subulinidae Truncatellidae ARACHNIDA: Scorpionida Buthidae Buthidae Buthidae Buthidae Buthidae Buthidae Urodacidae Urodacidae APPENDIX 1 26 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collections Seismic 1 Seismic 2 C1 C2 00000 00000100 00100000 10100000 11000000 11101000 01101111 11100010 10000000 00010001 11100101 00100000 10100001 00000001 01000000 00100000 00000001 00000001 10000001 11000000 00000000 01000000 00000001 00100000 01100001 11000000 sp. BI.155 sp. BI.111 Bimichaelia cf. Speleorchestes Amblyomma limbatum Amblyomma triguttatum Haemaphysalis ratti SuborderARACHNIDA: Acari Astigmatina Family cf. Glycyphagidae Subfamily Genus Unknown Species sp. BI.514 GP 2006 NIS 2006 NIS 2007 Astigmatina Unknown sp. BI.510 Astigmatina Unknown sp. BI.511 0 0 1 Astigmatina Unknown sp. BI.512 Endeostigmata Alycidae Endeostigmata Alycidae Unknown sp. BI.132 Endeostigmata Micropsammidae Unknown sp. BI.513 Endeostigmata Nanorchestidae Ixodida Argasidae Unknown sp. BI.426 Ixodida Ixodidae Ixodida Ixodidae Ixodida Ixodidae Mesostigmata Ascidae UnknownAsca sp. BI.407 nr Mesostigmata Ascidae Unknown sp. BI.433 Mesostigmata cf. Laelapidae Unknown sp. BI.411 Mesostigmata cf. Laelapidae Unknown sp. BI.429 Mesostigmata cf. Laelapidae Unknown sp. BI.430 Mesostigmata cf. Laelapidae Unknown sp. BI.432 Mesostigmata cf. Macrochelidae Unknown sp. BI.409 Mesostigmata cf. Phytoseiidae Unknown sp. BI.415 Mesostigmata Dinychidae Unknown sp. 1 Mesostigmata Dinychidae Unknown sp. 2 Mesostigmata Laelapidae Unknown sp. 1 Mesostigmata Laelapidae Unknown sp. BI.412 Mesostigmata Laelapidae Unknown sp. BI.413 Mesostigmata Laelapidae Unknown sp. BI.414 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 27 Other Collections Seismic 1 Seismic 2 C1 C2 00000 00100000 01100000 00000001 00000001 10000000 01100000 10000000 00000001 11000001 00001000 00100010 11100111 01100110 10100000 00100000 00100000 11101101 11100000 00100100 11101110 11000000 11100000 01000000 00000000 11101111 11100101 incrustata trirostrata sp. 1 sp. 2 sp. BI.212 sp. BI.242 sp. BI.222 sp. BI.241 sp. BI.213 sp. BI.223 sp. BI.239 sp. BI.227 sp. BI.207 cf. cf. Amblyseius Amblyseius Scapheremaeus Scapheremaeus Eremaeozetes Galumna Incabates Peloribates Peloribates Africoribates Neoliodes Ceroribatula Ceroribatula Ceroribatula monozygata SuborderMesostigmata Family Oplitidae Subfamily Genus Unknown Species sp. 1 GP 2006 NIS 2006 NIS 2007 Mesostigmata Oplitidae Unknown sp. 2 Mesostigmata Pachylaelapidae Unknown sp. BI.406 1 0 0 Mesostigmata Phytoseiidae Mesostigmata Phytoseiidae Mesostigmata Uropodidae Unknown sp. 1 Mesostigmata Uropodidae Unknown sp. 2 Mesostigmata Uropodoidea Unknown sp. 1 Mesostigmata Uropodoidea Unknown sp. BI.431 Mesostigmata Unknown sp. BI.410 Mesostigmata Unknown sp. BI.427 Mesostigmata Unknown sp. BI.428 Oribatida Cymbaeremaeidae Oribatida Cymbaeremaeidae Oribatida Eremaeozetidae Oribatida Euphthiracaridae Unknown sp. BI.247 Oribatida Galumnidae Oribatida Haplozetidae Oribatida Haplozetidae Oribatida Haplozetidae Oribatida Mycobatidae Oribatida Neoliodidae Oribatida Neoliodidae Unknown sp. BI.219 Oribatida Neoliodidae Unknown sp. BI.249 Oribatida Oribatulidae Oribatida Oribatulidae Oribatida Oribatulidae 28 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collections Seismic 1 Seismic 2 C1 C2 00100000 01100111 00000000 10000011 01000100 11100111 11100111 11100111 00100111 11100000 10000000 00000001 11100100 00001000 00100000 10001000 01100000 01001000 00000000 11101101 00000100 11100011 11101110 11101000 00001011 11101011 00100000 colossus sp. BI.237 sp. BI.201 sp. BI.225 sp. BI.246 sp. BI.228 sp. BI.229 sp. BI.187 sp. BI.230 sp. BI.191 sp. BI.126 sp. 1 sp. 2 sp. 2 nr group Reticuloppia reticulata Zygoribatula Bipassalozetes Walzia cf. cf. Fovoribatula brevisetosa Lucoppia Oribatula runcinata Zygoribatula cycloporosa Zygoribatula magna Zygoribatula Zygoribatula cf. Passalozetes Paralycus Pedrocortisella calmorum Licnodamaeus Adamystis cf. Chausseria warrengense Erythracarus ciliatus Pedidromus Pedidromus Tarsotomus abruptus Tarsotomus anubis Tarsotomus SuborderOribatida Family Oribatulidae Subfamily Genus Species GP 2006 NIS 2006 NIS 2007 Oribatida Oribatulidae Oribatida Oribatulidae Oribatida Oribatulidae Oribatida Oribatulidae Oribatida Oribatulidae Oribatida Oribatulidae Oribatida Oribatulidae Oribatida Oribatulidae Oribatida Passalozetidae Oribatida Passalozetidae Oribatida Pediculochelidae Oribatida Pedrocortisellidae Oribatida Pheroliodidae Unknown sp. 1 Oribatida Phthiracaridae Unknown sp. BI.245 Oribatida Plateremaeidae Oribatida Scheloribatidae Unknown sp. BI.238 Oribatida Zetomotrichidae Unknown sp. BI.211 Prostigmata Adamystidae Prostigmata Anystidae Anystinae Prostigmata Anystidae Erythracarinae Prostigmata Anystidae Erythracarinae Prostigmata Anystidae Erythracarinae Prostigmata Anystidae Erythracarinae Prostigmata Anystidae Erythracarinae Prostigmata Anystidae Erythracarinae Prostigmata Anystidae Erythracarinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 29 Other Collections Seismic 1 Seismic 2 C1 C2 11100100 00100111 11101101 11101100 00000000 00001111 00000010 11100000 11101011 00000001 00001000 11100000 01000010 11101101 00000101 11100000 11100111 11000000 10100000 11100101 10000010 11101000 11000000 11101110 00000001 11000101 11100101 thori cronini sp. BI.130 sp. nr sp. nr sp. 1 sp. 1 sp. BI.120 sp. BI.153 sp. BI.107 sp. 1 sp. BI.123 sp. BI.105 sp. 2 sp. BI.116 sp. BI.127 sp. BI.106 sp. BI.113 sp. BI.118 sp. BI.182 sp. BI.109 sp. BI.114 Bdellodes Biscirus Spinibdella Microcaeculus Neocaeculus Coptocheles Molothrognathus crucis Hemicheyletia bakeri Cunaxa Microsmaris Pollux Wartookia Abrolophus Caeculisoma Caeculisoma Charletonia Unknown sp. BI.162 Erythrites Erythrites Erythrites Erythrites Erythroides Leptus Callidostomatinae Callidostomatinae Callidostomatinae Callidostomatinae Callidostomatinae Erythraeidae Erythraeidae Erythraeidae Erythraeidae SuborderProstigmata Family Bdellidae Subfamily Odontoscirinae Genus Species GP 2006 NIS 2006 NIS 2007 Prostigmata Bdellidae Spinibdellinae Prostigmata Bdellidae Spinibdellinae Prostigmata Caeculidae Prostigmata Caeculidae Prostigmata Calligonellidae Prostigmata Calligonellidae Prostigmata Camerobiidae Unknown sp. 2 Prostigmata Camerobiidae Unknown sp. BI.134 Prostigmata Cheyletidae Prostigmata Cryptognathidae Unknown sp. 1 Prostigmata Cunaxidae Prostigmata Cunaxidae Unknown sp. 2 Prostigmata Erythraeidae Balaustiinae Prostigmata Erythraeidae Balaustiinae Prostigmata Erythraeidae Balaustiinae Prostigmata Prostigmata Prostigmata Prostigmata Prostigmata Erythraeidae Prostigmata Erythraeidae Erythraeinae Prostigmata Erythraeidae Erythraeinae Prostigmata Erythraeidae Erythraeinae Prostigmata Erythraeidae Erythraeinae Prostigmata Erythraeidae Erythraeinae Prostigmata Erythraeidae Leptinae 30 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collections Seismic 1 Seismic 2 C1 C2 11000010 00000000 00100000 00100010 00000001 00100000 10000000 10000000 00100000 00000010 00100010 10000111 00000101 00000001 10100010 00000001 00000000 00000001 00000001 11001111 00000010 01001111 00000001 00000001 10001001 11000000 00000000 sp. unknown sp. BI.172 sp. BI.173 sp. 1 sp. 1 sp. 2 sp. BI.112 sp. BI.178 sp. 1 sp. BI.185 sp. BI.152 sp. BI.184 sp. BI.183 Erythrites Pronematulus Phanolophus Andrevella cf. Myrmicotrombium Rainbowia Eupodes Eupodes cf. Anoplocheylus Spaherotarsus cf. Smaris Eryngiopus Eustigmaeus Austrateneriffia hirstii SuborderProstigmata Family Erythraeidae Leptinae Subfamily Unknown Genus sp. BI.122 Species GP 2006 NIS 2006 NIS 2007 Prostigmata Erythraeidae Prostigmata Erythraeidae Prostigmata Erythraeidae Prostigmata Erythraeidae Prostigmata Erythraeidae Unknown sp. BI.176 Prostigmata Eupodidae Prostigmata Eupodidae Prostigmata Eupodidae Unknown sp. BI.174 Prostigmata Iolinidae Pronematinae Prostigmata Pseudocheylidae Prostigmata Raphignathidae Unknown sp. BI.141 Prostigmata Smarididae Hirstiosomatinae Prostigmata Smarididae Phanolophinae Prostigmata Smarididae Smaridinae Prostigmata Stigmaeidae Prostigmata Stigmaeidae Prostigmata Stigmaeidae Unknown sp. BI.196 Prostigmata Tarsonemidae Unknown sp. BI.192 Prostigmata Teneriffiidae Prostigmata Tenuipalpidae Unknown sp. BI.197 Prostigmata Tetranychidae Unknown sp. BI.169 Prostigmata Tetranychidae Unknown sp. BI.195 Prostigmata Tetranychidae Unknown sp. BI.198 Prostigmata Trombidiidae Unknown sp. BI.143 Prostigmata Trombidiidae Unknown sp. BI.151 Prostigmata Trombidiidae Unknown sp. BI.194 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 31 Other Collection Other Collections Seismic 1 Seismic 2 C1 C2 00000 00000001 01000000 00100100 01100000 00100100 10000000 10100000 00021010 11001000 10001000 10001000 0 00001000 0 01000000 0 01010100 1 11101010 1 10101110 1 10000000 1 11000100 1 11101100 1 10011110 1 00111100 0 10001000 sp. n. sp. 1 0 sp. 1 0 sp. VWF 1023 0 sp. VWF 1022 1 sp. 1 1 Tuckerella Carepalxis Araneus eburniventris Araneus exsertus Araneus lutulentus Araneus rhombocephalus Argiope protensa Argiope trifasciata Austracantha minax Backobourkia collina cf. Dolophones Larinia montagui Larinia Lariniaphora ragnhildae Neoscona Neoscona theisi Psittacinus SuborderProstigmata Family Tuckerellidae Subfamily GenusInfraorder Species Family GP 2006 Genus NIS 2006 NIS 2007 Species GP NIS S1 S2 C1 C2 C3 C4 Prostigmata Tydeidae Unknown sp. 2 ProstigmataARACHNIDA: Araneae Araneomorphae Araneidae Unknown sp. BI.156 1 0 0 Prostigmata Unknown sp. BI.163 Prostigmata Unknown sp. BI.170 Araneomorphae Araneidae Prostigmata Unknown sp. BI.177 Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Genus 1 sp. 1 1 Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae Araneomorphae Araneidae 32 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 0001000 00020000 10000010 00101110 10001010 10000100 10001110 01000000 10000000 10000110 11000110 00000100 00000001 00010000 00001000 10000010 10000000 10000000 00001110 10000000 00000000 00000000 1 10000000 1 10001110 1 10010000 0 10010000 1 10000000 sp. 1 0 sp. 1 0 sp. 2 1 sp. 1 1 sp. 1 0 sp. 1 0 sp. 2 0 sp. 1 0 sp. 1 0 Zygiella Clubiona Clubiona Orthobula Supunna funerea Supunna picta Deinopis Forsterina Forsterina Wandella waldockae Meedo houstoni Oreo capensis Eilica Encoptarthria Infraorder Family Genus Species GP NIS S1 S2 C1 C2 C3 C4 Araneomorphae Araneidae Araneomorphae Clubionidae Araneomorphae Clubionidae Araneomorphae Corinnidae Genus 1 sp. 1 0 1 Araneomorphae Corinnidae Genus 2 sp. 1 0 Araneomorphae Corinnidae Araneomorphae Corinnidae Araneomorphae Corinnidae Araneomorphae Deinopidae Araneomorphae Desidae Araneomorphae Desidae Araneomorphae Dictynidae Genus 1 sp. 1 1 Araneomorphae Dictynidae Genus 2 sp. 1 0 Araneomorphae Dictynidae Genus 2 sp. 2 0 Araneomorphae Filistatidae Araneomorphae Gallieniellidae Araneomorphae Gallieniellidae Araneomorphae Gnaphosidae Araneomorphae Gnaphosidae Araneomorphae Gnaphosidae Genus 1 sp. 1 1 Araneomorphae Gnaphosidae Genus 1 sp. 2 0 Araneomorphae Gnaphosidae Genus 1 sp. 3 0 Araneomorphae Gnaphosidae Genus 1 sp. 4 0 Araneomorphae Gnaphosidae Genus 1 sp. 5 1 Araneomorphae Gnaphosidae Genus 1 sp. 6 0 Araneomorphae Gnaphosidae Genus 1 sp. 7 1 Araneomorphae Gnaphosidae Genus 1 sp. 8 1 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 33 Other Collection 0010000 10001000 10000010 00000010 10010000 10001000 10000000 10000000 10000000 00000000 00010000 00000000 00100000 00001000 00000010 10100000 00000100 00000100 10000000 11101000 0 11020000 0 10000000 1 10000000 0 01001100 1 10001000 0 00020001 0 00001010 sp. 1 0 sp. 1 0 sp. 1 1 sp. 1 0 sp. VWF 166 0 Hemicloea Asadipus phaleratus Asadipus yundamindra Bigenditia zuytdorp Lamponega Lamponella Lamponina scutata Erigone prominens Laetesia Hoggicosa snelli Hogna Knoelle clara Infraorder Family Genus Species GP NIS S1 S2 C1 C2 C3 C4 Araneomorphae Gnaphosidae Genus 1 sp. 9 0 Araneomorphae Gnaphosidae Genus 2 sp. 1 1 Araneomorphae Gnaphosidae Genus 3 sp. 1 0 Araneomorphae Gnaphosidae Genus 3 sp. 2 0 1 Araneomorphae Gnaphosidae Genus 4 sp. 1 1 Araneomorphae Gnaphosidae Genus 4 sp. 2 1 Araneomorphae Gnaphosidae Genus 4 sp. 3 0 Araneomorphae Gnaphosidae Genus 5 sp. 1 0 Araneomorphae Gnaphosidae Genus 6 sp. 1 0 Araneomorphae Gnaphosidae Genus 6 sp. 2 1 Araneomorphae Gnaphosidae Genus 6 sp. 3 0 Araneomorphae Gnaphosidae Genus 7 sp. 1 1 Araneomorphae Gnaphosidae Araneomorphae Lamponidae Araneomorphae Lamponidae Araneomorphae Lamponidae Araneomorphae Lamponidae Araneomorphae Lamponidae Araneomorphae Lamponidae Araneomorphae Linyphiidae Araneomorphae Linyphiidae Genus 1 sp. 1 0 Araneomorphae Linyphiidae Genus 1 sp. 2 0 Araneomorphae Linyphiidae Genus 3 sp. 1 0 Araneomorphae Linyphiidae Araneomorphae Lycosidae Araneomorphae Lycosidae Araneomorphae Lycosidae 34 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 10001000 10021110 10011000 10001010 10011000 01001000 00000000 10000000 10111000 10000000 00000001 10000000 10000000 10000000 10000000 10000000 11011010 00011000 00000001 00000000 10000000 11001110 11000110 1 10000100 1 10101110 0 10000000 1 10100000 sp. VWF 161 1 sp. A 1 sp. 1 1 sp. 1 1 sp. 3 0 cf. sp. 2 0 sp. 1 1 sp. 3 1 sp. 4 0 sp. 5 0 sp. 1 1 sp. 2 1 sp. 1 0 sp. 2 1 sp. 3 1 sp. 1 0 sp. 2 1 Venator Cheiracanthium Nephila edulis Oecobius navus Grymeus Myrmopopaea Myrmopopaea Opopaea Opopaea Opopaea Opopaea Opopaea Orchestina Orchestina Pelicinus saaristoi Xestaspis inconspiqua Xestaspis Xestaspis Xestaspis Oxyopes Oxyopes Infraorder Family Genus Species GP NIS S1 S2 C1 C2 C3 C4 Araneomorphae Lycosidae Araneomorphae Miturgidae Araneomorphae Miturgidae Genus 1 sp. 1 1 Araneomorphae Miturgidae Genus 2 sp. 1 1 Araneomorphae Miturgidae Genus 3 sp. 1 1 Araneomorphae Miturgidae Genus 3 sp. 2 1 Araneomorphae Nephilidae Araneomorphae Oecobiidae Araneomorphae Oonopidae Genus 1 sp. 1 1 Araneomorphae Oonopidae Genus 2 sp. 1 1 Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oonopidae Araneomorphae Oxyopidae Araneomorphae Oxyopidae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 35 Other Collection 1001110 10011000 11041110 11011110 11031000 00001100 10170100 11031100 00001000 10010000 00001010 11001100 00001000 01010000 10000000 00000100 11011110 0 10000000 0 11001000 1 10011110 1 00001100 1 10031000 0 10000110 1 10000100 1 00011110 0 10011010 0 00010100 sp. 3 0 sp. 4 0 sp. 5 0 sp. 6 0 sp. 7 1 sp. 1 0 sp. 1 0 sp. 2 0 sp. 1 1 sp. 1 1 sp. 2 0 1 Oxyopes Oxyopes Oxyopes Oxyopes Oxyopes Tibellus Artema atlanta Crossopriza lyoni Trichocyclus nigropunctatus Cryptoerithus occultus Cryptoerithus Cryptoerithus Nomindra leeuweni Wydundra barrow Wydundra kennedy Prodidomus woodleigh Clynotis albobarbatus Cytaea Grayenulla australensis Grayenulla Grayenulla Infraorder Family Genus Species GP NIS S1 S2 C1 C2 C3 C4 Araneomorphae Oxyopidae Araneomorphae Oxyopidae Araneomorphae Oxyopidae Araneomorphae Oxyopidae Araneomorphae Oxyopidae Araneomorphae Philodromidae Araneomorphae Pholcidae Araneomorphae Pholcidae Araneomorphae Pholcidae Genus 1 sp. 1 0 Araneomorphae Pholcidae Araneomorphae Prodidomidae Araneomorphae Prodidomidae Araneomorphae Prodidomidae Araneomorphae Prodidomidae Araneomorphae Prodidomidae Araneomorphae Prodidomidae Araneomorphae Prodidomidae Araneomorphae Prodidomidae Genus 1 sp. 1 0 Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Genus 1 sp. 1 0 Araneomorphae Salticidae Genus 2 sp. 1 0 Araneomorphae Salticidae Genus 3 sp. 1 0 Araneomorphae Salticidae Genus 4 sp. 1 0 Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae 36 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 1111110 00000001 01000100 11041100 10001000 10001000 11010100 10000000 00011000 10001000 10011100 10001000 11100000 01010000 00001000 10101000 10001010 11100100 10001010 00011100 0 00000001 1 10001010 1 11011110 0 11100000 0 11170100 0 00010000 0 01010100 sp. 1 0 sp. 2 1 sp. 1 0 sp. 2 1 1 sp. 4 1 sp. 5 0 sp. 6 1 sp. 1 0 sp. 1 0 sp. 2 0 sp. 3 1 sp. 1 0 sp. 1 0 sp. 1 1 sp. 1 1 sp. 1 1 sp. 2 1 sp. 3 1 sp. 4 1 Holoplatys desertina Holoplatys Holoplatys Lycidas Lycidas Lycidas Lycidas Lycidas Myrmarachne Neon Neon Neon Opisthoncus Paraplatoides Pellenes bitaeniata Simaetha tenuior Zebraplatys Zenodorus orbiculatus Dictis striatipes Anyphops Heteropoda hermitis Irileka iridescens Neosparassus Neosparassus Neosparassus Neosparassus Infraorder Family Genus Species GP NIS S1 S2 C1 C2 C3 C4 Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Salticidae Araneomorphae Scytodidae Araneomorphae Segestriidae Genus 1 sp. 1 0 Araneomorphae Selenopidae Araneomorphae Sparassidae Araneomorphae Sparassidae Araneomorphae Sparassidae Araneomorphae Sparassidae Araneomorphae Sparassidae Araneomorphae Sparassidae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 37 Other Collection 0000100 10010000 10001000 11151110 10041000 10020000 10030000 11031100 10001110 10101110 00001110 00000100 11011000 00001000 00000100 10000100 11000110 00000000 10000000 10001110 10000110 11011110 11000000 00001000 1 10000000 0 00001000 0 10000010 sp. 1 0 sp. 1 0 sp. 1 1 0 sp. 1 1 sp. 2 0 sp. 5 0 sp. 6 0 sp. 1 0 sp. 1 1 sp. 1 1 sp. 1 1 sp. 2 0 sp. 1 0 sp. 2 0 Pediana horni Pediana tenuis Phonognatha Tetragnatha Argyrodes Emertonella Emertonella Euryopis Euryopis Latrodectus hasselti Latrodectus Rhomphaea Steatoda Yagunimena Yagunimena Diaea Diaea Infraorder Family Genus Species GP NIS S1 S2 C1 C2 C3 C4 Araneomorphae Sparassidae Araneomorphae Sparassidae Araneomorphae Tetragnathidae Araneomorphae Tetragnathidae Araneomorphae Theridiidae Araneomorphae Theridiidae Araneomorphae Theridiidae Araneomorphae Theridiidae Araneomorphae Theridiidae Araneomorphae Theridiidae Genus 1 sp. 1 0 Araneomorphae Theridiidae Genus 1 sp. 2 0 Araneomorphae Theridiidae Genus 1 sp. 3 0 Araneomorphae Theridiidae Genus 1 sp. 4 1 Araneomorphae Theridiidae Genus 1 sp. 6 0 Araneomorphae Theridiidae Genus 2 sp. 1 0 Araneomorphae Theridiidae Genus 2 sp. 2 1 Araneomorphae Theridiidae Genus 3 sp. 1 1 Araneomorphae Theridiidae Genus 4 sp. 1 0 Araneomorphae Theridiidae Araneomorphae Theridiidae Araneomorphae Theridiidae Araneomorphae Theridiidae Araneomorphae Theridiidae Araneomorphae Theridiidae Araneomorphae Thomisidae Araneomorphae Thomisidae Araneomorphae Thomisidae Genus 1 sp. 1 1 38 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 0001000 00000010 10000000 10001010 00010000 10001100 11111010 11030000 10000000 00001000 10121000 10031110 11111110 11101010 10011110 00021000 10021000 10041000 00040000 10000000 10001010 10000000 00000001 1 11021110 sp. 1 0 sp. 1 0 sp. 2 0 sp. 3 0 sp. 1 0 sp. 2 0 sp. 1 0 sp. 1 0 sp. 2 1 sp. 3 0 sp. 4 0 sp. 1 1 sp. 2 0 sp. 1 0 sp. 2 0 p000100000 sp.0 p000000001 sp.0 p000000001 sp.0 sp. 1 0 sp. 3 0 cf. sp. 1 1 1 Sidymella Stephanopis Stephanopis Stephanopis Tharpyna Tharpyna Australutica Habronestes Habronestes Habronestes Habronestes Minasteron minusculum Neostorena Neostorena Argoctenus Argoctenus Missulena Idiommata Aurecocrypta Conothele Conothele Aname Barychelidae Ctenizidae Infraorder Family Genus Species GP NIS S1 S2 C1 C2 C3 C4 Araneomorphae Thomisidae Genus 1 sp. 2 0 Araneomorphae Thomisidae Araneomorphae Thomisidae Araneomorphae Thomisidae Araneomorphae Thomisidae Araneomorphae Thomisidae Araneomorphae Thomisidae Araneomorphae Zodariidae Araneomorphae Zodariidae Genus 1 sp. 1 1 Araneomorphae Zodariidae Genus 2 sp. 1 0 Araneomorphae Zodariidae Araneomorphae Zodariidae Araneomorphae Zodariidae Araneomorphae Zodariidae Araneomorphae Zodariidae Araneomorphae Zodariidae Araneomorphae Zodariidae Araneomorphae Zoridae Araneomorphae Zoridae Araneomorphae Zoridae Genus 1 sp. 1 0 Araneomorphae Zoridae Genus 2 sp. 1 0 Mygalomorphae Actinopodidae Mygalomorphae Barychelidae Mygalomorphae Mygalomorphae Mygalomorphae Ctenizidae Mygalomorphae Nemesiidae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 39 Other Collection Other Collection 1011110 1100110 0011010 0001010 0000110 1001010 01000000 00000000 00001000 00000001 1 00001010 0 1 1000010 0 1 0000110 1 1 1001110 0 1 0010000 0 1 0011000 1 1 0000000 sp. 2 1 sp. 3 1 sp. 4 1 sp. 1 0 pallidulus .s.1 0 0000010 sp. n. .s.0 1 0000000 sp. n. pseudopyrgoniscus sp. 1 1 1 sp. 2 1 1 sp. nov. 32sp. nov. 1 1 sp. nov. 53sp. nov. 0 0 .s.0 0 0000001 sp. n. .s.1 0 0000000 sp. n. cf. Aname mainae Aname Aname Aname Yilgarnia Acanthodillo Barrowdillo Barrowdillo Buddelundia hirsuta Buddelundia humphreysi Buddelundia Buddelundia Buddelundia Buddelundia Spherillo Hanoniscus Laevophiloscia yalgoonensis Porcellionides pruinosus Alloniscus Infraorder Family GenusFamily Species GP Genus NIS S1 S2 Species C1 C2 GP C3 NIS C4 S1 S2 C1 C2 C3 C4 Mygalomorphae Nemesiidae MALACOSTRACA: Isopoda Armadillidae Mygalomorphae Nemesiidae Armadillidae Mygalomorphae Nemesiidae Armadillidae Armadillidae Mygalomorphae Nemesiidae Armadillidae Mygalomorphae Nemesiidae Armadillidae Armadillidae Armadillidae Armadillidae Armadillidae Gen. n. sp. 1 n. sp. 1 1 Armadillidae Gen. n. sp. 2 n. sp. 0 1 Armadillidae Oniscidae Philosciidae Porcellionidae Scyphacidae 40 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR 00000 10000000 10000000 01000000 01000000 11000000 11000000 10000000 10000010 11000000 01000000 11111101 11000020 01000021 11100000 11100010 11000001 11000011 11000011 sp. 1 sp. 1 sp. unknown Tuoba laticeps Cryptops EthmostigmusArthrorhabdusCormocephalus curtipes Cormocephalus paucispinus Scolopendra strigosus Scolopendra turneri Pilbarascutigera laeta Thereuopoda morsitans incola Thereuopoda UnixenusPhryssonotus mjobergi novaehollandiae OrderCHILOPODA Geophilida Family Subfamily Genus Species Genus 1 GP sp. 1 NIS S1 S2 C1 C2 C3 C4 Geophilida Genus 2 sp. 1 Geophilida Genus 3 sp. 1 Geophilida Genus 4 sp. 1 Geophilida Lithobiida Genus 1 sp. 1 0 1 0 ScolopendridaScolopendrida Cryptopidae Scolopendrida ScolopendridaeScolopendrida ScolopendridaeScolopendrida Otostigminae ScolopendridaeScolopendrida Scolopendrinae ScolopendridaeScolopendrida Scolopendrinae ScolopendridaeScutigerida Scolopendrinae ScolopendridaeScutigerida Scolopendrinae Scutigerida Scutigeridae Scolopendrinae Scutigeridae DIPLOPODA Scutigeridae PolydesmidaPolyxenidaPolyxenida Paradoxosomatidae Polyxenidae Synxenidae Genus 1 sp. 1 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 41 Other Collection 0000001 0000001 0000001 0000001 0000001 0000001 0000001 0000001 0000001 0000001 0000001 0000000 0000000 0000000 0000000 0000000 0 0 0000001 0 0 0000001 0 0 0000001 0 0 0000001 1 0 0000000 1 0 0000000 1 0 0000000 1 0 0000000 prominentia vernalis kingi sp. VN48 0 0 sp. 2 0 0 sp. 3 0 0 sp. nr sp. VN69 0 0 sp. 3 0 0 sp. cf. sp. 1 0 0 sp. VN61 0 0 sp. 4 0 0 sp. 2 0 0 .s.0 0 0000001 sp. n. sp. 1 0 0 sp. 2 0 0 .s.0 0 0000001 sp. n. sp. VN40 1 0 sp. VN71 0 1 sp. 2 0 1 sp. 2 0 1 sp. cf. sp. VN39 1 0 Kenyura Collophorura Isotoma viridis Rastriopes Paraxenylla Folsomides loftyensis Folsomides Proisotoma brisbanensis Psammisotoma Sminthurides Brachystomella Acherontiella Paraxenylla Axelsonia Folsomides Stenognathellus cf. Oudemansia Oudemansia Caufrenyllodes cf. Corynephoria Brachystomella Calx Hypogastrura Xenylla yucatana Archisotoma cf. Pseudanurida glauerti Family Genus Species GP NIS S1 S2 C1 C2 C3 C4 ENTOGNATHA: Collembola ENTOGNATHA: Bourletiellidae Hypogastruridae Isotomidae Isotomidae Isotomidae Isotomidae Sminthurididae Brachystomellidae Hypogastruridae Hypogastruridae Isotomidae Isotomidae Katiannidae Neanuridae Neanuridae Neanuridae Odontellidae Arrhopalitidae Bourletiellidae Brachystomellidae Entomobryidae Hypogastruridae Hypogastruridae Isotomidae Isotomidae Neanuridae 42 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 0000000 0000000 0000010 0000010 0000000 0000010 0001000 0010000 0000000 0000010 0000010 0100010 0000010 0000010 0000010 0100100 0001010 1100000 1 1 0000000 1 1 0000010 1 1 0000010 0 1 0000110 1 1 0000010 1 1 0000010 0 0 1001100 1 0 1100010 gp VN28 gp albidorsalis reticulata vulgaris absoloni thermophila sp. VN37 1 0 sp. VN33 1 0 .s.0 0 0000100 sp. n. sp. white 0 0 sp. 1 0 0 sp. 2 1 1 sp. VN35 0 0 sp. 1 1 0 sp. VN25 0 1 sp. VN2 1 1 sp. 2 1 1 sp. 3 0 1 sp. 1 0 0 sp. VN15 1 1 sp. 1 sp. cf. sp. cf. sp. VN17 1 1 sp. sp. 1 1 1 sp. cf. sp. VN47 1 1 sp. 1 1 1 sp. VN42 1 1 Mesaphorura Tullbergia Acanthocyrtus Lepidosira Folsomides parvulus Folsomides Folsomides Sphaeridia Calx Sminthurinus Pseudachorutes Prorastriopes Prorastriopes Brachystomella dianae Lepidocyrtus Sminthurinus Corynephoria Corynephoria Corynephoria Rastriopes Hemisotoma Stenognathellus Corynephoria Corynephoria Prorastriopes Corynephoria Pygicornides torridus Family Genus Species GP NIS S1 S2 C1 C2 C3 C4 Tullbergiidae Tullbergiidae Entomobryidae Entomobryidae Isotomidae Isotomidae Isotomidae Sminthurididae Entomobryidae Katiannidae Neanuridae Bourletiellidae Bourletiellidae Brachystomellidae Entomobryidae Katiannidae Bourletiellidae Bourletiellidae Bourletiellidae Bourletiellidae Isotomidae Katiannidae Bourletiellidae Bourletiellidae Bourletiellidae Bourletiellidae Sminthurididae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 43 Other Collection Other Collection 00000 0111010 0110010 0111110 1101010 1111110 111011000 140011010 100000010 000011000 000011100 001011000 110000000 111040100 100000000 111001110 1 1 1001010 1 1 0011010 1 1 0011010 1 1 1101110 1 1 1111110 1 1 1111110 grp hartmeyeri sp. nov. 1 sp. nov. sp. nov. 2 sp. nov. sp. nov. 3 sp. nov. sp. nov. 4 sp. nov. cf. sp. 8 VN19 queenslandica vulgaris mitchelli coeruleopicta sp. 3 0 0 sp. cf. sp. VN36 1 1 sp. cf. sp. 1 0 0 sp. VN10 1 1 sp. cf. sp. cf. sp. VN6 1 1 Acrotelsella devriesiana Acrotelsella Acrotelsella Acrotelsella Acrotelsella Ctenolepisma longicaudata Anisolepisma Heterolepisma Acanthocyrtus Seira Temeritas Corynephoria Acanthocyrtus barrowensis Acanthocyrtus Drepanosira Lepidobrya Drepanura cinquelineata Drepanura Metacoelura Family Genus SpeciesFamily GP NIS Subfamily S1 S2 Genus C1 C2 Species C3 GP C4 NIS S1 S2 C1 C2 C3 C4 Entomobryidae Entomobryidae Sminthuridae INSECTA: Zygentoma INSECTA: Lepismatidae Ctenolepismatinae Bourletiellidae Entomobryidae Entomobryidae Lepismatidae Ctenolepismatinae Entomobryidae Entomobryidae Entomobryidae Lepismatidae Ctenolepismatinae Entomobryidae Paronellidae Lepismatidae Ctenolepismatinae Lepismatidae Ctenolepismatinae Lepismatidae Ctenolepismatinae Lepismatidae Ctenolepismatinae 1 Gen. nov. 1 sp. nov. 1 1 0 0 Lepismatidae Ctenolepismatinae 1 Gen. nov. 2 sp. nov. Lepismatidae Ctenolepismatinae 2 Gen. nov. 1 sp. nov. Lepismatidae Heterolepismatinae Lepismatidae Heterolepismatinae 44 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 0 110000000 000001000 000001000 000011000 010000000 110001000 000001000 110000000 110001100 011001010 111001000 010000000 110001000 111111010 000000001 000000001 000000001 aussa caurus cf. cf. n. sp.110000000 n. n. sp.110000000 n. sp. 5 Hemianax papuensis Austroagrion pindrina Ischnura aurora Ischnura heterosticta Lestes concinnus Diplacodes bipunctata Orthetrum caledonicum Pantala flavescens Candovia pallida Denhama Hyrtacus Hyrtacus Marcenia Vasilissa walkeri Unknown Aposthonia spinulosa Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 INSECTA: Odonata INSECTA: Aeshnidae Coenagrionidae Coenagrionidae Coenagrionidae Lestidae Libellulidae Libellulidae Libellulidae INSECTA: Phasmida INSECTA: Diapheromeridae Necrosciinae Phasmatidae Lonchodinae Phasmatidae Lonchodinae Phasmatidae Lonchodinae Phasmatidae Lonchodinae Phasmatidae Phasmatinae Phasmatidae INSECTA: Embioptera INSECTA: Oligotomidae INSECTA: Orthoptera INSECTA: Acrididae Catantopinae Catantopin sp. 1 Acrididae Catantopinae Catantopin sp. 10 Acrididae Catantopinae Catantopin sp. 2 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 45 Other Collection 00001 000000001 000000001 000000001 000000001 000000001 010000000 000000010 000000010 000000100 000000010 000000010 000000100 100000000 010000000 010000000 010000000 100000000 100000000 100000000 100000000 010000000 100000000 100000000 010000000 100000000 010000000 grp australis modesta sp. 11 sp. 1 0 0 0 0 sp. 1 sp. 06-1 sp. sp. 07-1 cf. sp. 06-1 sp. 06-3 sp. ANIC 4 sp. 06-1 sp. 06-1 . 47 sp. 06-1 Macrotona Goniaeoidea Perbellia picta Oecanthus rufescens Torbia Goniaea furcifera Goniaeoidea Stropis nigrovitellina Lepidogryllus Trigonidium cf. ANIC Gen. Nov cf. cf. Eumecistes gratiosus Goniaea vocans Macrotona Macrotona Macrotona Micreola Pespulia Urnisiella Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Acrididae Catantopinae Catantopin sp. 3 Acrididae Catantopinae Catantopin sp. 4 Acrididae Catantopinae Catantopin sp. 5 Acrididae Catantopinae Acrididae Catantopinae Gryllidae Oecanthinae Tettigoniidae Phaneropterinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Eumastacidae Morabinae Unknown sp. 06-2 Gryllidae Gryllinae Gryllidae Trigonidiinae Acrididae Catantopinae ANIC Gen. No. 95 sp. 1 Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Eumastacidae Morabinae Genus 06-4 sp. Gryllacrididae Gryllacridinae Genus 06-2 sp. Gryllacrididae Gryllacridinae Unknown sp. 06-2 46 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 010000000 100000000 100000000 010000000 010000000 010000000 010000000 010000000 010000000 010000000 100000000 100000000 010000000 010000000 010000000 010000000 100001000 100000010 010001000 100000100 010001000 101000000 100000010 010001000 010001000 010000010 100000010 grp sp. 1 grp sp. 2 magnifica sp. 07-1 sp. 07-1 sp. 07-1 sp. 06-1 sp. z sp. z1 sp. 06-1 sp. a sp. cf. ANIC 23 sp. cf. sp. nr sp. 07-1 sp. 06-1 Apterogryllus palpatus Aritella Eurygryllodes gorimuis Eurygryllodes gorimuis Teleogryllus commodus Yarrita Marinna Oecanthus Conocephalus Conocephalus Conocephalus upoluensis Coryphistes Macrazelota Macrotona roseipennis Macrotona Austracris guttulosa Hadrogryllacris Myara Eurygryllodes Oecanthus angustus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Gryllacrididae Gryllacridinae Unknown sp. 06-3 Gryllidae Eneopterinae Genus 06-1 sp. Gryllidae Eneopterinae Genus 06-3 sp. Gryllidae Gryllinae Gryllidae Gryllinae Gryllidae Gryllinae Gryllidae Gryllinae Gryllidae Gryllinae Gryllidae Gryllinae Gryllidae Mogoplistinae Gryllidae Oecanthinae Gryllidae Oecanthinae Unknown sp. 07-1 Tettigoniidae Conocephalinae 9 ANIC Gen. nov. sp. 06-1 Tettigoniidae Conocephalinae Tettigoniidae Conocephalinae Tettigoniidae Conocephalinae Acrididae Catantopinae Acrididae Catantopinae 28 Gen. nov. sp. 07-1 Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Cyrtacanthacridinae Gryllacrididae Gryllacridinae Genus 06-1 sp. Gryllacrididae Gryllacridinae Gryllidae Eneopterinae Gryllidae Gryllinae Gryllidae Oecanthinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 47 Other Collection 100001000 010001000 100001000 110000000 010000000 110000000 110000000 110000000 010000000 110000000 110000000 000001110 000001110 011000010 010001010 010001010 010001010 110001000 110001000 110001000 110000010 010001000 110000010 010000100 110000000 110000000 110000000 grp meekappa parvula sp. 07-1 sp. nov. sp. 1 ANIC sp. 5 nr sp. 07-1 sp. 1 sp. 06-1 sp. 06-1 sp. 07-1 sp. nov. sp. nov. sp. ANIC 9 sp. 06-1 Rufocephalus Conocephalus Conocephalus Throscodectes Acrida conica Desertaria Pardillana limbata Valanga irregularis cf. Ornebius Aritella Kalyra goparinga Aiolopus thlassinus Stropis Birubia Oecanthus adyeri Trigonidium Ablectia rufescens Goniaeoidea Peakesia Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Tettigoniidae Conocephalinae Tettigoniidae Conocephalinae Tettigoniidae Tettigoniinae Acrididae Acridinae Acrididae Catantopinae Acrididae Catantopinae Gen. 95 Acrididae Catantopinae Acrididae Catantopinae Unknown sp. 06-A Acrididae Cyrtacanthacridinae Eumastacidae Morabinae Genus 06-1 sp. Gryllidae Gryllinae Eumastacidae Morabinae Morabin sp. 1 Gryllidae Mogoplistinae Acrididae Catantopinae 28 Gen. nov. sp. 1 Gryllacrididae Gryllacridinae Unknown sp. 06-4 Gryllidae Gryllinae Gryllidae Mogoplistinae Acrididae Acridinae Acrididae Catantopinae Eumastacidae Morabinae Genus 06-2 sp. Gryllacrididae Gryllacridinae Unknown sp. Gryllidae Gryllinae Gryllidae Oecanthinae Gryllidae Trigonidiinae Acrididae Catantopinae Acrididae Catantopinae Acrididae Catantopinae 48 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 110000000 110000000 111000010 010001010 010001010 110001010 110001010 111000000 011001110 110011010 110001010 110001010 111001000 110001010 110000110 110011110 111001110 100000000 100101110 110000010 010000000 010000000 110001000 110000000 010000000 kimberleyensis minor sp. 06-1 sp. ANIC 3 sp. 06-1 sp. 06-1 sp. 06-1 sp. 06-1 sp. 06-1 cf. cf. sp. 07-1 sp. 1 sp. 07-1 sp. 07-2 sp. 07-1 sp. 07-1 sp. 1 Bolbe Archimantis Archimantis Pseudomantis Sphodropoda Salmonites Monistria pustulifera Pycnostictus seriatus Happarana Eurepella mjobergi Conocephalus Antipodectes brevicaudus Goniaeoidea rufotestacea Turana kiwani Coryphistes glabriceps Craspedogryllacris Endacusta Caedicia Polichne Macrotona curvicostalis Paraoxypilus Phthersigena cf. Archimantis cf. cf. cf. cf. Pseudomantis Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Gryllidae Eneopterinae Pyrgomorphidae Pyrgomorphinae Acrididae Acridinae Acrididae Catantopinae Gryllidae Eneopterinae Tettigoniidae Conocephalinae Tettigoniidae Tettigoniinae Acrididae Catantopinae Gryllidae Eneopterinae Acrididae Catantopinae Gryllacrididae Gryllacridinae Gryllidae Mogoplistinae Unknown sp. 06-1 Gryllidae Phalangopsinae Tettigoniidae Phaneropterinae Tettigoniidae Phaneropterinae Acrididae Catantopinae INSECTA: Mantodea INSECTA: Amorphoscelidae Paraoxypilinae Amorphoscelidae Paraoxypilinae Mantidae Iridopteryginae Mantidae Mantinae Mantidae Mantinae Mantidae Mantinae Mantidae Mantinae Mantidae Mantinae Mantidae Mantinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 49 Other Collection 110100010 010011110 010001010 110000100 110010100 010000100 001000001 110001100 110111010 111011110 110101010 110111010 010000010 110001000 111100001 110001000 100000000 010010000 111001010 110011100 111001010 110011000 110001110 110101110 110001000 ministralis tibialis sp. VN3 sp. 07-1 sp. nr sp. VN13 nr. nr. undesc. sp. 1 undesc. sp. 2 undesc. sp. 3 undesc. sp. 4 Ellipsidion UnknownUnknownUnknownUnknown sp. 1 Unknown sp. 2 sp. 3 sp. 4 sp. 5 Supella longipalpa Rhodomantis Orthodera Calolampra aspera Periplaneta americana Anamesia uniformis Eppertia Platyzosteria Platyzosteria sublobata Platyzosteria tricaudata Platyzosteria Platyzosteria Platyzosteria Platyzosteria Pseudophyllodromiinae Pseudophyllodromiinae Pseudophyllodromiinae Pseudophyllodromiinae Pseudophyllodromiinae Pseudophyllodromiinae Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Ectobiidae Ectobiidae Ectobiidae Ectobiidae Ectobiidae Ectobiidae Blattelllinae Mantidae Mantinae Ectobiidae Unknown sp. 6 Mantidae Orthoderinae Ectobiidae Unknown sp. 7 INSECTA: Blattodea INSECTA: Blaberidae Diplopterinae Ectobiidae Unknown sp. a Blaberidae Diplopterinae undesc. gen. sp. VN2 Blattidae Blattinae Blattellidae Blattidae Polyzosteriinae Blattidae Polyzosteriinae Blattidae Polyzosteriinae Blattidae Polyzosteriinae Blattidae Polyzosteriinae Blattidae Polyzosteriinae Blattidae Polyzosteriinae Blattidae Polyzosteriinae Blattidae Polyzosteriinae Polyphagidae Unknown sp. 1 50 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 011011110 110011000 010100010 000000110 110100000 010010000 000110000 000000001 011011100 000100010 010110100 110000000 010000000 011111010 111110010 100001010 111111110 110000010 111111000 010000000 010000000 111001100 110000000 rctuosus hartmeyeri magnus peracutus reticulatus vicinus erosus nr a nr cf. nr sp. 1 cf. cf. sp. 1 Species T nr Species B Species N Drepanotermes Heterotermes ferox Heterotermes intermedius Schedorhinotermes Schedorhinotermes Amitermes abruptus Amitermes Amitermes cf. Drepanotermes diversicolor Drepanotermes rubriceps Microcerotermes serratus Nasutitermes Nasutitermes eucalypti Nasutitermes longipennis Nasutitermes triodiae Tumulitermes Tumulitermes hastilis Unknown Caecilius Ectopsocus Ectopsocus Ectopsocus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 INSECTA: Isoptera INSECTA: Rhinotermitidae Heterotermitinae Rhinotermitidae Heterotermitinae Rhinotermitidae Rhinotermitinae Rhinotermitidae Rhinotermitinae Termitidae Amitermitinae Termitidae Amitermitinae Termitidae Amitermitinae Termitidae Amitermitinae Termitidae Amitermitinae Termitidae Amitermitinae Termitidae Amitermitinae Termitidae Nasutitermitinae Termitidae Nasutitermitinae Termitidae Nasutitermitinae Termitidae Nasutitermitinae Termitidae Nasutitermitinae Termitidae Nasutitermitinae Termitidae Nasutitermitinae INSECTA: Psocoptera INSECTA: Amphientomidae Amphientomid 1 Species H Caeciliusidae Ectopsocidae Ectopsocidae Ectopsocidae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 51 Other Collection 00100 00000000 110000000 010000100 010001100 010000000 101110110 000000001 110000110 010000010 000000010 000000001 010100011 110101110 010010010 000010000 011001000 110111010 001000001 000000100 111100000 101000000 111100100 110000000 010000000 Species P Species R Species A Species O Species F Species I sp. 2 1 sp. 1 Lepinotus Lepinotus Ectopsocus Ectopsocus Pteroxanium Liposcelis bostrychophila Liposcelis entomophila Liposcelis Peripsocus fici Cladioneura foliata Barrowia insularis Dorypteryx domestica cf. cf. Andrewarthaia kellyana Desmothrips broomei Cranothrips kartus Cranothrips Merothrips Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Ectopsocidae Ectopsocidae Ectopsocidae Unknown Species Q Unknown Gen. n. Species M Lepidopscocidae Liposcelididae Liposcelid 1 Species D Liposcelididae Liposcelid 2 Species E 0 0 0 0 Liposcelididae Liposcelid 3 Species J Liposcelididae Liposcelid 4 Species K Liposcelididae Liposcelid 5 Species L Liposcelididae Liposcelididae Liposcelididae Peripsocidae Philotarsidae Philotarsid 1 Species G Pseudocaeciliidae Psocidae Psyllipsocidae Trogiidae Trogiidae INSECTA: Thysanoptera INSECTA: Aeolothripidae Aeolothripidae Melanthripidae Melanthripidae Merothripidae 52 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 111111110 010000000 110011110 010000100 100010010 111111110 001000000 001001110 011100000 110001110 000010010 001001110 111000100 110000010 111101110 101111000 111100000 100011000 111101000 111111110 011100110 010011100 111110010 001000000 011101110 001100000 ritchei sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 0 0 1 0 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. n cf. Nesothrips Horistothrips Stomothrips Allothrips stannardi cf. Gastrothrips Neosmerinthothrips Nesothrips Priesneriella citricauda Scotothrips Akainothrips Baenothrips Biconothrips Brakothrips cf. cf. Corroboreethrips Gynaikothrips Haplothrips Heligmothrips Holothrips Jacotia rhodorcha Katothrips Kellyia Klambothrips Ostlingothrips corini Podothrips ardis Podothrips barrowi Podothrips Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Phlaeothripidae Idolothripinae Phlaeothripidae Idolothripinae Phlaeothripidae Idolothripinae Phlaeothripidae Idolothripinae Phlaeothripidae Idolothripinae Phlaeothripidae Idolothripinae Phlaeothripidae Idolothripinae Phlaeothripidae Idolothripinae Unknown sp. 1 Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 53 Other Collection 00000 010000000 000000110 111111110 010000000 110000000 111010110 111111110 011111110 111000100 010101010 001001110 001000110 110100000 000000010 100001100 110000010 110000000 110000000 100000000 110011110 100001010 100000100 010011010 sp. n sp. 1 sp. n. sp. 1 0 1 0 0 sp. 1 sp. 1 sp. n. sp. 1 sp. 2 p 000000110 sp. Dendrothrips Podothrips Rhopalothripoides luteus Senithrips psomus cf. Neohydatothrips Anaphothrips barrowi Caprithrips Frankliniella schultzei Pseudanaphothrips achaetus Rhamphothrips Scirtothrips Thrips parvispinus Thrips safrus Thrips Cicadetta Cicadetta Taurella viridis Illyria major Macrotristria extrema Dysoliarus unicornis Oliarus Ozoliarus bullockorum Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Phlaeothripidae Phlaeothripinae Thripidae Dendrothripinae Thripidae Sericothripinae Thripidae Thripinae Thripidae Thripinae Thripidae Thripinae Thripidae Thripinae Thripidae Thripinae Thripidae Thripinae Thripidae Thripinae Thripidae Thripinae Thripidae Thripinae INSECTA: Hemiptera: Auchenorrhyncha INSECTA: Cicadidae Tibicininae Cicadidae Tibicininae Cicadidae Tibicininae Cicadidae Cicadinae Cicadidae Cicadinae Cicadidae Gen. 1 sp. 1 Caliscelidae Unknown A Caliscelidae Unknown B Cixiidae Cixiinae Cixiidae Cixiinae Cixiidae Cixiinae 54 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 010000100 110001100 010000010 010001110 010100010 111111110 010111110 111111110 111011110 000100000 110011110 110100010 A 110001100 n. sp.100000010 n. p 110001110 sp. n. sp.011000000 n. p 010011100 sp. p 000000010 sp. p 010000000 sp. p 111011110 sp. A 110001010 B 111001110 A 111011110 B 111001110 C 100000110 p 010000000 sp. sp. 1 sp. 2 Massila Sardia pluto rostrata Toya Cleotyche Thanatodictya Falcophantis westcotti Barsac Barsac cf. Siphanta patruelis Galela parva Amyclin Phaconeura Bilbilicallia Bilbilicallia Lipocallia Lipocallia Lipocallia Euricania Alleloplasis Alleloplasis Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Delphacidae Delphacinae Delphacidae Delphacinae Delphacidae Delphacinae Unknown B Delphacidae Delphacinae Unknown C Delphacidae Delphacinae Unknown D Delphacidae Delphacinae Unknown E Delphacidae Delphacinae Unknown F 0 1 0 1 Dictyopharidae Dictyopharinae Dictyopharidae Dictyopharinae Eurybrachidae Unknown A Eurybrachidae Unknown B Flatidae Flatinae Flatidae Flatinae Flatidae Flatinae Flatidae Flatinae Flatidae Flatinae Fulgoridae Aphaeninae Fulgoridae Amyclinae Meenoplidae Nisiinae Nogodinidae Lipocalliinae Nogodinidae Lipocalliinae Nogodinidae Lipocalliinae Nogodinidae Lipocalliinae Nogodinidae Lipocalliinae Ricaniidae Tropiduchidae Bladininae Tropiduchidae Bladininae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 55 Other Collection 010000010 110000010 110011000 100001010 010000000 110101110 110100100 100001010 100000010 100000100 110011110 000000001 110101110 010000000 110000010 110101110 000000001 110001000 110000100 110000000 110101010 110000110 p 010000000 sp. sp. 1A A 011001100 p 010000010 sp. sp. 1A sp. 9 p 010000110 sp. Oligaethus Arawa Exitianus nanus Exitianus plebeius Gunawardenea linnaei Limotettix incertus Linacephalus foveolatus Chiasmus varicolor Horouta Horouta austrina Horouta darwini Horouta perparvus Maiestas knighti Maiestas Maiestas Maiestas Amrasca Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Tropiduchidae Tropiduchinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Gen. n. C Cicadellidae Deltocephalinae Genus L sp. Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Unknown A Cicadellidae Deltocephalinae Unknown F Cicadellidae Deltocephalinae Unknown H Cicadellidae Ulopinae Cicadellidae Ulopinae Unknown B Cicadellidae Deltocephalinae Unknown G Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Unknown D Cicadellidae Deltocephalinae Unknown E Cicadellidae Typhlocybinae 56 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 001000 010000010 010000100 111100100 110000100 111000000 010000010 111011110 110000010 111001110 111011100 111001110 010100010 110000000 011001000 111100010 110001100 110001010 010001000 010000010 sp. nov. sp. 23A sp. 69 sp. 70 n. sp. 1 p 111000000 sp. p 000000010 sp. p 111101110 sp. p 010001000 sp. n. sp.010001010 n. sp. 1A 1 1 1 p 110000000 sp. A 111001100 Austroasca histrionicula Austroasca viridigrisea Anzygina Anzygina Anzygina Empoascanara Empoascanara peregrina Empoascanara Euacanthella impressa Goniagnathus Hecalus australis Batracomorphus adventitiosus Batracomorphus angustatus Batracomorphus Ipoides hackeri Balclutha incisa Balclutha rosea Balclutha saltuella Balclutha Nesoclutha Nesoclutha phryne Nesoclutha Hishimonus Orosius Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Cicadellidae Typhlocybinae Cicadellidae Typhlocybinae Cicadellidae Typhlocybinae Cicadellidae Typhlocybinae Cicadellidae Typhlocybinae Cicadellidae Typhlocybinae Cicadellidae Typhlocybinae Cicadellidae Typhlocybinae Cicadellidae Typhlocybinae ZA sp. Cicadellidae Typhlocybinae ZA sp. 2 Cicadellidae Typhlocybinae ZA sp. 2A Cicadellidae Euacanthellinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Iassinae Cicadellidae Iassinae Cicadellidae Iassinae Cicadellidae Eurymelinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 57 Other Collection 111001110 111011110 110001000 010000110 010000010 100001010 100000000 100001010 100000000 100000000 110000000 000000001 000000001 010001110 110001010 100000000 C 010000000 sp. n. ’large white’ sp. n. ’large sp. n. ’small’ sp. n. ’white face’ p 111011010 sp. A 111111110 B 111111110 C 110001110 p 010000010 sp. p 100000000 sp. sp. n sp. n n. sp. 28 n. sp.011010010 n. p 110011110 sp. G 110001010 p 110000100 sp.

Diemoides Orosius Orosius canberrensis Orosius orientalis Orosius Orosius Orosius Mapochiella Paradorydium Paradorydium Paradorydium ? Mayawa capitata Mayawa Mayawa Soractellus Thamnophryne Stirellus fatigandus Stirellus Kahavalu Austroagallia torrida Maiestas Newmanniana mullensis Newmanniana Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Eupelicinae Cicadellidae Eupelicinae Cicadellidae Eupelicinae Cicadellidae Eupelicinae Cicadellidae Eupelicinae Unknown D Cicadellidae Eupelicinae Unknown E Cicadellidae Eupelicinae Unknown G Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Deltocephalinae Cicadellidae Ulopinae Cicadellidae Agalliinae Cicadellidae Deltocephalinae BIsD sp. 01 Cicadellidae Deltocephalinae Cicadellidae Tartessinae Cicadellidae Tartessinae 58 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 111000000 111111110 000100100 110000000 110000000 110000000 110000010 110000000 000000001 100000000 000001100 000000001 100000000 010000000 110011010 000000010 110000000 110000000 000000001 100000000 110000000 100000000 110000000 110000000 p 110011100 sp. Pascoepus viridiceps Protartessus spinosus Zaletta webbi Rigula Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Cicadellidae Idiocerinae Cicadellidae Tartessinae Cicadellidae Macropsinae Unknown A Cicadellidae Deltocephalinae Unknown A Cicadellidae Tartessinae Unknown A Cicadellidae Typhlocybinae Unknown A Cicadellidae Tartessinae Unknown B 0 1 0 0 Cicadellidae Deltocephalinae Unknown B Cicadellidae Typhlocybinae Unknown B Cicadellidae Tartessinae Unknown C Cicadellidae Deltocephalinae Unknown C Cicadellidae Typhlocybinae Unknown C Cicadellidae Tartessinae Unknown D Cicadellidae Typhlocybinae Unknown D Cicadellidae Typhlocybinae Unknown E Cicadellidae Deltocephalinae Unknown E Cicadellidae Deltocephalinae Unknown F Cicadellidae Typhlocybinae Unknown G Cicadellidae Deltocephalinae Unknown H Cicadellidae Deltocephalinae Unknown I Cicadellidae Typhlocybinae Unknown J Cicadellidae Typhlocybinae Unknown K Cicadellidae Typhlocybinae Unknown L Cicadellidae Typhlocybinae Unknown M Cicadellidae Idiocerinae Membracidae Centrotinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 59 Other Collection 110000000 110001000 110000000 010011110 010111110 100000100 001000110 111110110 010010000 111111110 111111110 111111110 011000000 110001010 000011110 010000010 010000000 010000000 110011010 110000000 100000000 110001000 scutellaris p 110010100 sp. D 100000000 sp. 1 sp. 2 sp. 3 RIPT001 sp. nr ORIU001 MONT001 META001 AGRI001 Metacanthus Aphis craccivora Aphis gossypii Aphis Acizzia Acizzia Acizzia Acizzia Riptortus Melanacanthus Orius Montandoniola nr Agriopocoris Mictis profana Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 INSECTA: Hemiptera: Sternorrhyncha INSECTA: Aleyrodidae Unknown A Aleyrodidae Unknown B Aleyrodidae Unknown C Aleyrodidae Unknown sp. 1 Aleyrodidae Unknown sp. 2 Aphididae Aphididae Aphididae Coccoidea Unknown sp. Pseudococcidae Unknown sp. Psyllidae Psyllidae Psyllidae Psyllidae Triozidae Unknown sp. INSECTA: Hemiptera: Heteroptera INSECTA: Alydidae Alydinae Alydidae Anthocoridae Anthocorinae Anthocoridae Anthocoridae Unknown ANTH001 Berytidae Coreidae Coreinae Coreidae Coreinae Gen. nov. COLP001 Coreidae Coreinae 60 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00001 010000000 110111110 110001110 100000000 100000000 010001110 010000000 110000000 010000000 010010010 010001110 000000001 010001010 100000000 010001110 110001110 010001100 010010000 010001010 111001100 000010000 111111010 110101110 100000010 010000000 010000000 GRAL001 BLAE001 SYST001 SYST002 GEOC001 GERM001 DINO001 DICY001 SING001 SING002 FULV002 FULV001 PERI001 TRIL001 CREO001 FRON001 OPER001 nr Singhalesia Graviclava Dinomachus Fulvius Creontiades Fronsetta Opera nr Blaena Systelloderes Systelloderes Geocoris Germalus Limnogonus fossarum gilguy nr Gen. nov. Gen. nov. Nesidiocoris tenuis Singhalesia Singhalesia Fulvius nr Peritropis Trilaccus nr nr nr Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Coreidae Pseudophloeinae Cydnidae Cydnidae Unknown CYDN001 Cydnidae Unknown CYDN002 Cydnidae Unknown CYDN003 Cydnidae Unknown CYDN005 Cydnidae Unknown sp. Q075-1 0 0 0 0 Dipsocoridae Unknown DIPS001 Enicocephalidae Enicocephalinae Enicocephalidae Geocoridae Geocorinae Geocoridae Geocorinae Gerridae Gerrinae Heterogastridae Miridae Bryocorinae Miridae Bryocorinae Miridae Bryocorinae Miridae Bryocorinae Miridae Cylapinae Miridae Cylapinae Miridae Cylapinae Miridae Cylapinae Unknown FULV003 Miridae Deraeocorinae Miridae Mirinae Miridae Mirinae Unknown MIRI001 Miridae Orthotylinae Miridae Orthotylinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 61 Other Collection 00000 100000000 110001110 100000000 010000000 011001100 100000000 010000000 110001110 010001110 010000000 100000000 010100000 110010110 010111110 100000010 111101110 111001110 100000000 111111110 110001110 000001110 100001000 010001000 000001000 100100000 110011110 schellenbergii ornata RANK001 SOPH001 MACR001 HALL001 SEJA001 CAMP001 NABI001 OXYC001 EOCA001 cf. MENE001 KALK001 cf. BACH001 BRIZO001 Macrotylus Sejanus Eocanthocona Menestheus Kalkadoona Bacheusia Brizocoris Rankiniella Sophocoris Coridromius pilbarensis nr Hallodapus nr Sejanus albisignatus Campylomma Nabis Nysius vinitor Oxycarenus Stenophyella macreta nr Oechalia nr nr Ippatha nr nr Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Miridae Orthotylinae Miridae Orthotylinae Miridae Orthotylinae Unknown AUST001 Miridae Orthotylinae Miridae Orthotylinae Miridae Orthotylinae Unknown ORTH001 Miridae Orthotylinae Unknown ORTH002 1 0 0 0 Miridae Orthotylinae Gen. nov. ZANC002 Miridae Orthotylinae Unknown ZANC001 Miridae Orthotylinae Unknown ZANC003 Miridae Orthotylinae Unknown ZANC004 Miridae Phylinae Miridae Phylinae Miridae Phylinae Miridae Phylinae Miridae Phylinae Unknown PHYL001 Nabidae Nabinae Orsillidae Oxycarenidae Pachygronthidae Pentatomidae Asopinae Pentatomidae Asopinae Pentatomidae Pentatominae Pentatomidae Pentatominae Pentatomidae Pentatominae Pentatomidae Pentatominae Pentatomidae Pentatominae 62 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 01000 110000110 110000000 110000010 000001110 100001010 011011100 000011010 010101000 010000000 010000000 000100000 000001000 010000000 000001000 101000000 101000000 100000010 100000000 100010100 000001010 110100100 100000000 110000100 110000010 010000010 100000000 desertus ONC001 PIEZ001 ROEB001 sp. nr DYSD001 PLOI001 0 1 1 0 sp. 1 sp. 1 PRIST001 sp. 1 sp. 1 sp. 1 LEPT001 sp. 1 PLIN001 PLIN002 PLIN003 PARA001 Oncocoris Piezodorus Roebournea Ploiaria Pristhesancus Paradrymus nr nr nr Oncocoris Cephaloplatus pertyi Dysdercus nr Ploiaria Pseudobargylia Coranus aridellus Coranus granosus nr Ectomocoris Peirates Oncocephalus Leptocoris Liorhyssus Plinthisus Plinthisus Plinthisus nr Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Pentatomidae Pentatominae Pentatomidae Pentatominae Pentatomidae Pentatominae Pentatomidae Pentatominae Pentatomidae Phyllocephalinae Pyrrhocoridae Reduviidae Emesinae Reduviidae Emesinae Reduviidae Emesinae Reduviidae Emesinae Unknown EMES001 Reduviidae Emesinae Unknown EMES002 Reduviidae Harpactocorinae Reduviidae Harpactocorinae Reduviidae Harpactocorinae Reduviidae Peiratinae Reduviidae Peiratinae Reduviidae Reduviinae Unknown FLIG001 Reduviidae Stenopodainae Reduviidae Unknown brown’ sp. ’large Rhopalidae Serinethinae Rhopalidae Rhyparochromidae Plinthisinae Rhyparochromidae Plinthisinae Rhyparochromidae Plinthisinae Rhyparochromidae Rhyparochrominae Rhyparochromidae Rhyparochrominae Unknown COLE001 Rhyparochromidae Rhyparochrominae Unknown LETH001 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 63 Other Collection 00000 10110010 100000000 100000000 100000100 110000110 110000000 010000000 100000000 010000000 010000000 010000000 100001010 000010010 010000000 010000000 010000000 010000000 100000000 010010110 100011000 100010000 100000000 010100010 100010100 trilobata STIZ001 UDEO001 MORB001 DICR001 cf. COPI001 INOM001 LASI001 TING001 TING002 TING003 sp. 1 sp. 1 0 complex complex complex Stizocephalus Udeocoris Copium Inoma Lasiacantha Dieuches notatus nr nr Morbora Dicranocephalus Diplocysta nr nr nr Tingis Tingis Tingis Pseudencyoposis bicornis Suhpalacsa barrowensis Suphalomitus difformis Spermophorella Stenobiella Calochrysa extranea Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Rhyparochromidae Rhyparochrominae Unknown LETH002 Rhyparochromidae Rhyparochrominae Unknown LETH003 Rhyparochromidae Rhyparochrominae Unknown MYOD001 Rhyparochromidae Rhyparochrominae Rhyparochromidae Rhyparochrominae Rhyparochromidae Rhyparochrominae Unknown RHYP001 Rhyparochromidae Rhyparochrominae Unknown sp. ’long head’ 1 0 0 0 Rhyparochromidae Rhyparochrominae Unknown dark’ sp. ’small round Rhyparochromidae Rhyparochromidae Unknown sp. Scutelleridae Odontotarsinae Stenocephalidae Tingidae Tinginae Tingidae Tinginae Tingidae Tinginae Tingidae Tinginae Tingidae Tinginae Tingidae Tinginae Tingidae Tinginae INSECTA: Neuroptera INSECTA: Ascalaphidae Ascalaphinae Ascalaphidae Ascalaphinae Ascalaphidae Ascalaphinae Berothidae Berothidae Chrysopidae Chrysopinae 64 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 10001000 110010010 100000000 110001110 011101110 011101110 011111110 110001000 010000000 110011010 010000000 010000000 010010000 010000000 010000000 010000010 110100000 010001010 010000000 010000000 100000000 000010000 000010010 001021000 010000000 tillyardi congestus exilis fuscus kimminsi rieki peterseni conspurcata sp. 1 sp. 1 sp. 1 sp. nov. cf. sp. nov. cf. cf. cf. cf. sp. n. cf. cf. cf. sp. 1 Mallada Cryptoscenea Bandidus Plesiochrysa ramburi cf. Mallada signatus cf. Coniopteryx Heterocornis nigricornis Mestressa subfasciata Glenoleon Bandidus Bandidus Bandidus Bandidus cf. Eophanes distinctus Froggattisca Myrmeleon acer Myrmeleon croceicollis Protoplectron Protoplectron peterseni Stilbopteryx napoleo Heoclisis Xantholeon Lasioderma serricorne Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Chrysopidae Chrysopinae Chrysopidae Chrysopidae Coniopterygidae Aleuropteryginae Coniopterygidae Coniopteryginae Coniopterygidae Myrmeleontidae Acanthaclisinae Myrmeleontidae Dendroleontinae Myrmeleontidae Distoleontinae Myrmeleontidae Distoleontinae Myrmeleontidae Distoleontinae Myrmeleontidae Distoleontinae Myrmeleontidae Distoleontinae Myrmeleontidae Distoleontinae Myrmeleontidae Myrmeleontinae Myrmeleontidae Myrmeleontinae Myrmeleontidae Myrmeleontinae Myrmeleontidae Myrmeleontinae Myrmeleontidae Myrmeleontinae Myrmeleontidae Stilbopteryginae Myrmeleontidae Myrmeleontidae INSECTA: Coleoptera INSECTA: Anobiidae Anobiidae Unknown sp. 1 0 Anobiidae Unknown sp. 2 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 65 Other Collection 00000 10000000 010000000 001000000 110001010 010000000 000000001 010000000 110000000 010000000 010000010 110010110 100001010 000100010 000401000 001000010 010001110 112211110 010000010 010000000 010000010 010000000 000001000 000001000 110000000 100000000 110000000 sp. 1 sp. 1 sp. 2 sp. 3 sp. 4 sp. 5 1 1 0 0 sp. 6 sp. 7 sp. 8 sp. 1 Egestrina Formicomus Formicomus Formicomus Formicomus Formicomus Formicomus Formicomus Formicomus Mecynotarsus Bostrichopsis jesuita Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Anobiidae Unknown sp. 3 Anthicidae Anthicidae Anthicidae Anthicidae Anthicidae Anthicidae Anthicidae Anthicidae Anthicidae Anthicidae Anthicidae Unknown sp. 1 Anthicidae Unknown sp. 2 Anthicidae Unknown sp. 3 Anthicidae Unknown sp. 5 Apionidae Unknown sp. 1 Apionidae Unknown sp. 2 Apionidae Unknown sp. 3 Apionidae Unknown sp. 4 Apionidae Unknown sp. 5 Apionidae Unknown sp. 6 Apionidae Unknown sp. 7 Apionidae Unknown sp. 8 Biphyllidae Unknown sp. 1 1 Biphyllidae Unknown sp. 2 Biphyllidae Unknown sp. 3 Bostrichidae 66 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 10000000 000000001 000000001 000000001 000000001 000000001 112001000 000000001 000011000 000010000 000001000 000001000 000001000 011000000 100000000 100000000 100000000 100001000 110000000 110000000 100001000 010010010 000000010 010000000 010000000 010000000 sp. 1 0 0 1 0 sp. 1 sp. 1 sp. 1 sp. 1 sp. 2 sp. 3 sp. 4 sp. 5 sp. 6 sp. 1 Ankleineella sulcicollis Unknown Merimna atrata Neocuris discoflava Synechocera Unknown Adelotopus Calosoma schayeri Clivina Clivina Clivina Clivina Clivina Clivina Sphallomorpha Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Bostrichidae Unknown sp. 1 Bostrichidae Unknown sp. 2 Bostrichidae Unknown sp. 3 Bostrichidae Unknown sp. 4 Bostrichidae Unknown sp. 5 Brentidae Brentidae Buprestidae Buprestidae Buprestidae Buprestidae Buprestidae Unknown sp. 2 Buprestidae Unknown sp. 3 Carabidae Carabidae Carabidae Carabidae Carabidae Carabidae Carabidae Carabidae Carabidae Carabidae Unknown sp. 1 Carabidae Unknown sp. 10 0 Carabidae Unknown sp. 11 Carabidae Unknown sp. 12 Carabidae Unknown sp. 13 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 67 Other Collection 00001 10000000 110000000 110001000 110001000 010001000 010001000 100000000 100011110 010001000 010001000 100001000 110400000 010000000 010000000 010000000 010011010 110000000 110001010 010000000 010000000 010000000 010000000 010000000 000000001 000001100 000000001 Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Carabidae Unknown sp. 14 Carabidae Unknown sp. 15 Carabidae Unknown sp. 16 Carabidae Unknown sp. 17 Carabidae Unknown sp. 18 Carabidae Unknown sp. 19 Carabidae Unknown sp. 2 0 0 0 0 Carabidae Unknown sp. 20 Carabidae Unknown sp. 21 Carabidae Unknown sp. 22 Carabidae Unknown sp. 23 Carabidae Unknown sp. 24 Carabidae Unknown sp. 25 Carabidae Unknown sp. 26 Carabidae Unknown sp. 27 Carabidae Unknown sp. 28 Carabidae Unknown sp. 29 Carabidae Unknown sp. 3 Carabidae Unknown sp. 30 Carabidae Unknown sp. 31 Carabidae Unknown sp. 32 Carabidae Unknown sp. 33 Carabidae Unknown sp. 34 Carabidae Unknown sp. 35 0 Carabidae Unknown sp. 36 Carabidae Unknown sp. 37 Carabidae Unknown sp. 38 68 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 00000000 100001010 000011000 000001000 000001000 000000001 000010000 000000001 110001000 100000010 110000010 110001100 000000001 000000001 110101010 010000000 000001000 000001000 000001000 110011010 000021010 000000001 100010000 100000010 010000010 010000000 laminosus cf. Uracanthus maculatus Piesarthrius Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Carabidae Unknown sp. 4 Carabidae Unknown sp. 40 Carabidae Unknown sp. 41 Carabidae Unknown sp. 42 Carabidae Unknown sp. 43 Carabidae Unknown sp. 44 Carabidae Unknown sp. 45 0 0 1 0 Carabidae Unknown sp. 5 Carabidae Unknown sp. 6 Carabidae Unknown sp. 7 Carabidae Unknown sp. 8 Carabidae Unknown sp. 9 Cerambycidae Cerambycinae Cerambycidae Cerambycidae Unknown sp. 1 Cerambycidae Unknown sp. 10 Cerambycidae Unknown sp. 11 Cerambycidae Unknown sp. 12 Cerambycidae Unknown sp. 17 Cerambycidae Unknown sp. 2 Cerambycidae Unknown sp. 3 Cerambycidae Unknown sp. 4 Cerambycidae Unknown sp. 5 Cerambycidae Unknown sp. 6 1 Cerambycidae Unknown sp. 7 Cerambycidae Unknown sp. 8 Cerambycidae Unknown sp. 9 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 69 Other Collection 11000 00200000 010000000 010000000 006001000 002000000 110301010 110001110 010101110 110011010 112120100 000000001 010001000 011001000 010101000 110001010 013000010 110010010 000000001 100001010 010000000 100101010 100000000 100000010 010001000 000001000 110201110 Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Chrysomelidae Bruchinae Unknown sp. 1 Chrysomelidae Bruchinae Unknown sp. 2 Chrysomelidae Bruchinae Unknown sp. 3 Chrysomelidae Bruchinae Unknown sp. 4 Chrysomelidae Unknown sp. 1 Chrysomelidae Unknown sp. 10 Chrysomelidae Unknown sp. 11 1 1 0 0 Chrysomelidae Unknown sp. 12 Chrysomelidae Unknown sp. 13 Chrysomelidae Unknown sp. 14 Chrysomelidae Unknown sp. 15 Chrysomelidae Unknown sp. 16 Chrysomelidae Unknown sp. 17 Chrysomelidae Unknown sp. 18 Chrysomelidae Unknown sp. 19 Chrysomelidae Unknown sp. 2 Chrysomelidae Unknown sp. 20 Chrysomelidae Unknown sp. 21 Chrysomelidae Unknown sp. 22 Chrysomelidae Unknown sp. 23 Chrysomelidae Unknown sp. 24 Chrysomelidae Unknown sp. 25 Chrysomelidae Unknown sp. 26 Chrysomelidae Unknown sp. 27 1 Chrysomelidae Unknown sp. 28 Chrysomelidae Unknown sp. 29 Chrysomelidae Unknown sp. 3 70 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 01110 11000010 000001000 000011010 110000110 000001000 000000001 110001010 000000010 110100000 110121010 010000000 010111000 000000001 000001000 000001110 113711010 000001010 017000000 001000000 010200000 010000000 110001110 000001000 015111010 111301010 111001110 sp. 1 sp. 1 Myriochile semicincta Allelidea Necrobia rufipes Opilo Coccinella transversalis Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Chrysomelidae Unknown sp. 30 Chrysomelidae Unknown sp. 4 Chrysomelidae Unknown sp. 5 Chrysomelidae Unknown sp. 6 Chrysomelidae Unknown sp. 7 Chrysomelidae Unknown sp. 8 Chrysomelidae Unknown sp. 9 1 1 1 1 Cicindelidae Cleridae Tarsosteninae Unknown sp. 2 Cleridae Cleridae Cleridae Cleridae Unknown sp. 1 Cleridae Unknown sp. 2 Coccinellidae Coccinellidae gen. n. sp. 1 Coccinellidae gen. n. 2 sp. 1 Coccinellidae Unknown sp. 1 Coccinellidae Unknown sp. 10 Coccinellidae Unknown sp. 2 Coccinellidae Unknown sp. 3 Coccinellidae Unknown sp. 4 Coccinellidae Unknown sp. 5 Coccinellidae Unknown sp. 6 0 Coccinellidae Unknown sp. 7 Coccinellidae Unknown sp. 8 Coccinellidae Unknown sp. 9 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 71 Other Collection 00000 10101110 010000010 112030110 112700110 015531110 017300000 000021110 010501100 011031110 010001100 000011000 000000001 000000001 101200000 110001110 110001010 110201110 110001110 110001000 110011110 010000000 010100000 112421110 112021010 100020000 100000000 tragocephalus sp. 1 sp. 2 sp. 1 c.f. sp. 1 sp. 1 sp. 1 sp. 2 sp. 3 sp. 4 sp. 5 sp. 6 sp. 7 sp. 8 Alloparmulus Alloparmulus Hypothenemus Acantholophus Gonipterus Lixus Myllocerus Myllocerus Myllocerus Myllocerus Myllocerus Myllocerus Myllocerus Myllocerus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Colydiidae Unknown sp. 1 Corylophidae Corylophidae Curculionidae Scolytinae Curculionidae Scolytinae Unknown sp. 1 Curculionidae Scolytinae Unknown sp. 2 Curculionidae Scolytinae Unknown sp. 3 0 0 1 1 Curculionidae Scolytinae Unknown sp. 4 Curculionidae Scolytinae Unknown sp. 5 Curculionidae Scolytinae Unknown sp. 6 Curculionidae Scolytinae Unknown sp. 7 Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Unknown sp. 1 Curculionidae Unknown sp. 10 1 Curculionidae Unknown sp. 11 Curculionidae Unknown sp. 12 Curculionidae Unknown sp. 13 72 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 10000 01001000 100001000 101000000 100000000 010000000 010010010 000000001 110011000 110000010 110000110 010000000 010001110 110001010 110000010 011000000 000800100 110101000 011031110 010000000 100211000 000210000 000001000 001000000 110000000 000000001 sp. 1 p 000000001 sp. sp. 1 sp. 2 sp. 1 sp. 2 sp. 3 sp. 4 0 sp. 1 Anthrenocerus Attagenus Dermestes ater Dermestes haemorrhoidalis Orphinus Orphinus Trogoderma Trogoderma Trogoderma Trogoderma Aphanocephalus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Curculionidae Unknown sp. 14 Curculionidae Unknown sp. 16 Curculionidae Unknown sp. 17 Curculionidae Unknown sp. 18 Curculionidae Unknown sp. 19 Curculionidae Unknown sp. 2 Curculionidae Unknown sp. 20 0 1 0 2 Curculionidae Unknown sp. 3 Curculionidae Unknown sp. 4 Curculionidae Unknown sp. 5 Curculionidae Unknown sp. 6 Curculionidae Unknown sp. 7 Curculionidae Unknown sp. 8 Curculionidae Unknown sp. 9 Dermestidae Dermestidae Dermestidae Dermestidae Dermestidae Dermestidae Dermestidae Dermestidae Dermestidae Dermestidae Dermestidae Unknown sp. 1 Discolomidae Discolomidae Unknown sp. 1 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 73 Other Collection 01000 00000010 110000000 010001110 110000000 010000000 100100000 110000000 100010000 000000001 000000001 002000001 000000001 010001000 110000000 110010000 000001010 000001010 000000001 110000100 100000000 110011100 100000000 100000000 010000000 000000001 010000000 sp. 1 sp. 1 sp. 2 sp. 1 Allodessus Cybister tripunctatus Eretes australis Limbodessus Limbodessus Holoparamecus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Dytiscidae Dytiscidae Dytiscidae Dytiscidae Dytiscidae Elateridae Unknown sp. 1 Elateridae Unknown sp. 10 1 0 0 0 Elateridae Unknown sp. 11 Elateridae Unknown sp. 12 Elateridae Unknown sp. 13 Elateridae Unknown sp. 14 Elateridae Unknown sp. 15 Elateridae Unknown sp. 2 Elateridae Unknown sp. 3 Elateridae Unknown sp. 4 Elateridae Unknown sp. 5 Elateridae Unknown sp. 6 Elateridae Unknown sp. 7 Elateridae Unknown sp. 8 Elateridae Unknown sp. 9 Endomychidae Geotrupidae Unknown sp. 2 Geotrupidae Unknown sp. 3 Geotrupidae Unknown sp. 4 1 Geotrupidae Unknown sp. 5 Geotrupidae Unknown sp. 6 Gyrinidae Unknown sp. 1 74 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 00031100 110000000 110000000 110000000 000001000 110001000 110000000 112011100 110101000 010000110 010000100 010000000 010000110 110000100 1002410100 010101110 110100110 112051010 100100100 012000000 111000000 110010000 000100000 000011000 001000000 001000000 sp. 1 sp. 2 sp. 1 sp. 2 sp. 1 0 0 1 0 sp. 1 sp. 1 sp. 1 sp. 2 sp. 3 sp. 4 sp. 5 sp. 4 1 Halacritus lividus Liparochrus Liparochrus Enochrus Enochrus Cryptolestes Leucohimatium arundinaceum Lithostygnus Carphurus Simoderus Simoderus Simoderus Simoderus Simoderus Unknown Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Heteroceridae Unknown sp. 1 Histeridae Hybosoridae Hybosoridae Hydrophilidae Hydrophilidae Laemophloeidae Languridae Lathridiidae Lathridiidae Unknown sp. 1 Lathridiidae Unknown sp. 2 Lathridiidae Unknown sp. 3 Lathridiidae Unknown sp. 4 Lathridiidae Unknown sp. 5 Melandryidae Unknown sp. 1 Melyridae Melyridae Melyridae Melyridae Melyridae Melyridae Melyridae Unknown sp. 1 Melyridae Unknown sp. 3 Melyridae Melyridae Unknown sp. 5 Melyridae Unknown sp. 7 Melyridae Unknown sp. 8 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 75 Other Collection 31110 10000000 100000000 100001010 102000010 010000000 010000000 010000000 000001000 000100000 000000100 110021100 110000000 110000000 000000010 000001010 000001000 000001000 110000000 000000001 100000000 101000000 100100010 110000000 110000000 100400000 111000010 sp. 1 sp. 2 sp. 3 sp. 4 sp. 5 sp. 6 sp. 1 sp. 2 sp. 3 sp. 1 sp. 1 sp. 1 sp. 1 Glipostena Glipostena Glipostena Glipostena Glipostena Glipostena Litochrus Litochrus Litochrus Phycosecis Pselaphaulax Tmesiphorus Diplocotes Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Mordellidae Mordellidae Mordellidae Mordellidae Mordellidae Mordellidae Nitidulidae gen. n. sp. 1 0 0 5 4 Nitidulidae Unknown sp. 3 Nitidulidae Unknown sp. 4 Oedemeridae Unknown sp. 1 Phalacridae Phalacridae Phalacridae Phalacridae Unknown sp. 1 Phalacridae Unknown sp. 2 Phalacridae Unknown sp. 3 Phalacridae Unknown sp. 4 Phycosecidae Phycosecidae Unknown sp. 1 Pselaphidae Pselaphidae Pselaphidae Unknown sp. 1 Ptiliidae Unknown sp. 1 Ptiliidae Unknown sp. 2 0 Ptiliidae Unknown sp. 3 Ptiliidae Unknown sp. 4 Ptinidae 76 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 081110 110010010 110000000 111000110 110000000 000001000 110000000 000000001 000100000 000001000 010000000 000011000 110000000 000000001 010000000 010000000 000010000 000001000 100010010 100000000 000000001 100000000 110000010 110000010 000001000 000100000 sp. 1 sp. 1 sp. 2 sp. 3 sp. 4 sp. 1 sp. 2 1 1 0 0 sp. 1 sp. 1 sp. 2 sp. 3 sp. 4 sp. n. sp. n. sp. 1 sp. 1 sp. 2 sp. 3 sp. n. Ectrephes Polyplocotes Polyplocotes Polyplocotes Polyplocotes Ptinus Ptinus Diastictus Carneodon Carneodon Carneodon Carneodon Acheilo pedarius Acheilo Allothnonius Gnaphalopoda Heteronyx Heteronyx Heteronyx Neophyllotocus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Ptinidae Ptinidae Ptinidae Ptinidae Ptinidae Ptinidae Ptinidae Ptinidae Unknown sp. 1 0 0 10 Ptinidae Unknown sp. 2 Pythidae Unknown sp. 1 Rhipiphoridae Unknown sp. 1 Rhipiphoridae Unknown sp. 2 Rhipiphoridae Unknown sp. 3 Scarabaeidae Aphodiinae Scarabaeidae Dynastinae Scarabaeidae Dynastinae Scarabaeidae Dynastinae Scarabaeidae Dynastinae Scarabaeidae Dynastinae Unknown sp. 1 Scarabaeidae Melolonthinae Scarabaeidae Melolonthinae Scarabaeidae Melolonthinae Scarabaeidae Melolonthinae Scarabaeidae Melolonthinae Scarabaeidae Melolonthinae Scarabaeidae Melolonthinae Scarabaeidae Melolonthinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 77 Other Collection 00000 00001000 010000010 110000010 110100010 010000000 100001010 000100000 100100000 000000001 111001000 010300000 007000000 110000000 001010100 100300000 110001010 110001010 100000000 010000000 110100110 110000010 010000010 010000000 110000110 100000000 110001000 sp. 1 sp. 1 1 1 0 0 sp. 2 sp. 1 sp. 2 sp. 1 Onthophagus glabratus Onthophagus mjobergi Onthophagus murchinsoni Onthophagus Tesserodon tenebroides Euconnus Euconnus Silvanus Silvanus Scaphysoma Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Scarabaeidae Scarabaeinae Scarabaeidae Scarabaeinae Scarabaeidae Scarabaeinae Scarabaeidae Scarabaeinae Scarabaeidae Scarabaeinae Scraptidae Unknown sp. 1 Scydmaenidae Scydmaenidae Scydmaenidae Unknown sp. 1 Scydmaenidae Unknown sp. 2 Silvanidae Silvanidae Silvanidae Unknown sp. 1 Silvanidae Unknown sp. 2 Staphylinidae Staphylinidae Unknown sp. 1 Staphylinidae Unknown sp. 10 Staphylinidae Unknown sp. 11 Staphylinidae Unknown sp. 12 Staphylinidae Unknown sp. 13 Staphylinidae Unknown sp. 14 Staphylinidae Unknown sp. 15 Staphylinidae Unknown sp. 16 Staphylinidae Unknown sp. 18 0 Staphylinidae Unknown sp. 2 Staphylinidae Unknown sp. 3 Staphylinidae Unknown sp. 4 78 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 10011000 100001010 100000000 010000000 010000000 111000000 100001010 010000000 110001000 110001000 110141110 113000100 010000000 110000010 110000000 010000000 110000010 110000000 010100000 110001000 110011000 110211010 110001000 110000010 000000001 110000000 sp. 1 sp. 1 0 1 0 0 sp. 1 sp. 2 sp. 1 sp. 1 sp. 2 sp. 1 sp. 1 sp. 2 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 1 sp. 2 0 sp. 1 Boreosaragus Brises Celibe Celibe Chromomoea Ectyche nana Gonocephalum Gonocephalum Metistete Mithippia Mithippia Nocar Oectosis cylindrica Pterohelaeus Saragus Sobas Sympetes Thorictosoma Thorictosoma Trachyscelis ciliaris Tribolium confusum Ulomoides Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Staphylinidae Unknown sp. 5 Staphylinidae Unknown sp. 6 Staphylinidae Unknown sp. 7 Staphylinidae Unknown sp. 8 Staphylinidae Unknown sp. 9 Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae Tenebrionidae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 79 Other Collection 00000100 110001000 000010000 010000000 010000000 010000000 000000001 010000000 000010000 110010110 1102116110 110012110 110123110 000007010 110007110 111109010 010004010 110002010 110001010 010011000 110000000 010001000 100002100 sp. 1 sp. 1 sp. 2 sp. 3 p 010010010 sp. sp. 1 Phytoliromyza Unknown Omorgus Omorgus Omorgus Elenchus varleyi Mengenilla cf. Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Tenebrionidae Tenebrionidae Unknown sp. 2 Tenebrionidae Unknown sp. 3 Trogidae Trogidae Trogidae INSECTA: Strepsiptera INSECTA: Elenchidae Mengenillidae INSECTA: Diptera INSECTA: Agromyzidae Phytomyzinae Agromyzidae Unknown barr-01 Agromyzidae Agromyzidae Unknown barr-02 Agromyzidae Agromyzidae Unknown barr-03 Agromyzidae Agromyzidae Unknown barr-04 Agromyzidae Agromyzidae Unknown barr-05 Agromyzidae Asilidae Unknown Asilidae barr-01 Asilidae Unknown Asilidae barr-02 Asilidae Unknown Asilidae barr-03 Asilidae Unknown Asilidae barr-04 Asilidae Unknown Asilidae barr-05 Asilidae Unknown Asilidae barr-06 Asilidae Unknown Asilidae barr-07 Asilidae Unknown Asilidae barr-08 Asilidae Unknown Asilidae barr-09 Asilidae Unknown Asilidae barr-10 1 80 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 00012000 100004110 100001000 100001000 100003010 100010000 100002010 000011000 000010000 110010100 114028110 1130517110 015023000 100052110 100020100 101030100 100000010 100010000 111010000 111000110 1111164110 110021000 100002000 100013010 100000100 110000000 sp. 1 sp. 1 Comptosia Geron Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Asilidae Unknown Asilidae barr-11 Asilidae Unknown Asilidae barr-12 Asilidae Unknown Asilidae barr-13 Asilidae Unknown Asilidae barr-14 Asilidae Unknown Asilidae barr-15 Asilidae Unknown Asilidae barr-16 Asilidae Unknown Asilidae barr-17 1 0 0 0 Bombyliidae Lomatiniae Bombyliidae Bombyliidae Unknown Bombyliidae barr-01 Bombyliidae Unknown Bombyliidae barr-02 Bombyliidae Unknown Bombyliidae barr-03 Bombyliidae Unknown Bombyliidae barr-04 Bombyliidae Unknown Bombyliidae barr-05 Bombyliidae Unknown Bombyliidae barr-06 Bombyliidae Unknown Bombyliidae barr-07 Bombyliidae Unknown Bombyliidae barr-08 Bombyliidae Unknown Bombyliidae barr-09 Calliphoridae Unknown Calliphoridae barr-01 Calliphoridae Unknown Calliphoridae barr-02 Calliphoridae Unknown Calliphoridae barr-03 Calliphoridae Unknown Calliphoridae barr-04 Calliphoridae Unknown Calliphoridae barr-05 Calliphoridae Unknown Calliphoridae barr-06 1 Calliphoridae Unknown Calliphoridae barr-07 Calliphoridae Unknown Calliphoridae barr-08 Calliphoridae Unknown Calliphoridae barr-09 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 81 Other Collection 04010 12007000 100022010 100000000 011001000 000001000 1140150110 114012110 110000110 112015110 110000110 010000010 113023110 010000000 011000110 010000110 010000010 111000110 010032010 011003010 001010010 010003010 110011010 010000100 010015100 010022100 010000010 Canacidae barr-01 Cecidomyiidae barr-01 Cecidomyiidae barr-02 0 1 0 0 Cecidomyiidae barr-03 Cecidomyiidae barr-04 Cecidomyiidae barr-05 Cecidomyiidae barr-06 Cecidomyiidae barr-07 Cecidomyiidae barr-08 Cecidomyiidae barr-09 Cecidomyiidae barr-10 Cecidomyiidae barr-11 Ceratopogonidae barr-01 Ceratopogonidae barr-02 Ceratopogonidae barr-03 Ceratopogonidae barr-04 Ceratopogonidae barr-05 Ceratopogonidae barr-06 Ceratopogonidae barr-07 Ceratopogonidae barr-08 0 Ceratopogonidae barr-09 Ceratopogonidae barr-10 Ceratopogonidae barr-11 Plesiotethina australis Unknown Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Calliphoridae Unknown Calliphoridae barr-10 Calliphoridae Unknown Calliphoridae barr-11 Calliphoridae Unknown Calliphoridae barr-12 Canacidae Canacidae Cecidomyiidae Unknown Cecidomyiidae Unknown Cecidomyiidae Unknown Cecidomyiidae Unknown Cecidomyiidae Unknown Cecidomyiidae Unknown Cecidomyiidae Unknown Cecidomyiidae Unknown Cecidomyiidae Unknown Cecidomyiidae Unknown Cecidomyiidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown Ceratopogonidae Unknown 82 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 32000 10000000 010003110 010008000 110003000 010001000 110005110 010000010 100001000 000001000 000003000 112028010 0100112110 111011110 013016110 010113100 113013110 010000000 110000100 100001010 011004110 000021000 000010010 011102110 100000000 Chamaemyiidae barr-01 Chironomidae barr-01 Chironomidae Chironomidae barr-02 Chironomidae Chironomidae barr-03 Chironomidae Chironomidae barr-04 Chironomidae Chironomidae barr-05 Chironomidae Chironomidae barr-06Chironomidae 0 1 0 0 Chironomidae barr-07 Chironomidae sp. 1 sp. 1 sp. 1 sp. 1 sp. 2 p 010003100 sp. Birobitella Conioscinella Ensiferella Thaumatomyia Thaumatomyia Coelopa allaudi Aedes bancroftianus Aedes Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Chamaemyiidae Unknown Chironomidae Unknown Chironomidae Unknown Chironomidae Unknown Chironomidae Unknown Chironomidae Unknown Chironomidae Unknown Chironomidae Unknown Chloropidae Oscinellinae Chloropidae Oscinellinae Chloropidae Chloropidae Chloropidae Chloropidae Unknown barr-01 Chloropidae 1 1 11 0 5 27 1 1 0 Chloropidae Unknown barr-05 Chloropidae Chloropidae Unknown barr-08 Chloropidae Chloropidae Unknown barr-09 Chloropidae Chloropidae Unknown barr-10 Chloropidae Chloropidae Unknown barr-11 Chloropidae Chloropidae Unknown barr-12 Chloropidae Chloropidae Unknown barr-13 Chloropidae Coelopidae Coelopidae Unknown Coelopidae barr-01 Conopidae Unknown Conopidae barr-01 1 Cryptochetidae Unknown Cryptochetidae barr-01 Culicidae Culicidae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 83 Other Collection 10000 10000000 000001000 110000000 100000000 010000000 010000000 000000001 010002000 110000110 010009010 011000000 010005010 010000000 010003000 111023110 010021100 100000000 110001100 010000010 000010000 010001010 110000010 010001000 000001000 000001000 000002000 sp. 1 sp. 1 0 0 0 0 sp. 1 sp. 1 sp. 1 Aedes vigilax Anopheles amictus Microphor Sympcnus Drapetus Zeros Hydrellia Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Culicidae Culicidae Culicidae Gen. 2 sp. Culicidae Unknown Culicidae barr-01 Culicidae Unknown Culicidae barr-03 Dolichopodidae Microphorinae Dolichopodidae Sympycninae Dolichopodidae Unknown Dolichopodidae barr-01 Dolichopodidae Unknown Dolichopodidae barr-02 Dolichopodidae Unknown Dolichopodidae barr-03 Dolichopodidae Unknown Dolichopodidae barr-04 Dolichopodidae Unknown Dolichopodidae barr-05 Dolichopodidae Unknown Dolichopodidae barr-06 Dolichopodidae Unknown Dolichopodidae barr-07 Dolichopodidae Unknown Dolichopodidae barr-08 Dolichopodidae Unknown Dolichopodidae barr-09 Dolichopodidae Unknown Dolichopodidae barr-10 Drosophilidae Unknown barr-02 Drosophilidae Drosophilidae Unknown barr-03 Drosophilidae Empididae Tachydromiinae Empididae Unknown Empididae barr-01 Empididae Unknown Empididae barr-02 Empididae Unknown Empididae barr-03 Empididae Unknown Empididae barr-04 0 Ephydridae Gymnomyzinae Unknown sp. 1 Ephydridae Ilytheinae Ephydridae 84 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 10000 00000001 000001000 1100213110 0100743110 1100110100 1100216110 010012010 011015110 110002110 011010000 111010110 000001000 010010110 010001110 000001100 002000000 111002010 110028010 010002000 011012100 010043010 000002000 000000110 111022110 010100110 1121216100 vestustissima sp. 1 cf. sp. 1 sp. 1 sp. 1 0 Allotrichia nr Desmometopa varipalpus Musca Musca Dorniphora Megaselia scalaris Megaselia Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Ephydridae Ephydridae Unknown Ephydridae barr-01 Ephydridae Unknown Ephydridae barr-02 Ephydridae Unknown Ephydridae barr-03 Ephydridae Unknown Ephydridae barr-04 Ephydridae Unknown Ephydridae barr-05 Fergusoninidae Unknown barr-01 Fergusoninidae 1 1 2 0 Heleomyzidae Unknown Heleomyzidae barr-01 Keroplatidae Keroplatinae Unknown barr-01 Sciaroidea Lauxaniidae Unknown Lauxaniidae barr-01 Lonchaeidae Unknown Lonchaeidae barr-01 Milichiidae Milichiidae Unknown Milichiidae barr-01 Milichiidae Unknown Milichiidae barr-02 Muscidae Muscidae Muscidae Unknown Muscidae barr-01 Muscidae Unknown Muscidae barr-02 Muscidae Unknown Muscidae barr-03 Muscidae Unknown Muscidae barr-04 Muscidae Unknown Muscidae barr-05 Phoridae Phoridae Phoridae Phoridae Unknown Phoridae barr-01 Phoridae Unknown Phoridae barr-02 Phoridae Unknown Phoridae barr-03 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 85 Other Collection 10001000 101008010 100038100 000010110 111005010 010000100 010000000 112001110 110000100 1105216110 110006000 010009100 010001100 100010000 000012000 000020000 000001000 011034110 110013110 111000110 110000000 010502010 001000100 010000000 111004110 100000000 000011100 spp gp. nyctias cf. sp. 1 sp. 1 sp. 2 sp. 1 Tomasvaryella Tomasvaryella Psychoda alternata Scenopinus albicinctus Riekiella Riekiella Australosepsis niveipennis Tabanus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Phoridae Unknown Phoridae barr-04 Phoridae Unknown Phoridae barr-05 Pipunculidae Pipunculidae Platystomatidae Unknown Platystomatidae barr-01 Platystomatidae Unknown Platystomatidae barr-02 Psychodidae Psychodidae Unknown Psychodidae barr-02 Sarcophagidae Unknown barr-01 Sarcophagidae Sarcophagidae Unknown barr-02 Sarcophagidae Sarcophagidae Unknown barr-03 Sarcophagidae Sarcophagidae Unknown barr-04 Sarcophagidae Scatopsidae Unknown Scatopsidae barr-01 Scenopinidae Scenopininae Scenopinidae Scenopinidae Scenopinidae Unknown Scenopinidae barr-01 Scenopinidae Unknown Scenopinidae barr-02 Sciaridae Unknown Sciaridae barr-01 Sciaridae Unknown Sciaridae barr-02 Sciaridae Unknown Sciaridae barr-03 Sepsidae Sepsinae Sepsidae Unknown Sepsidae barr-01 Sepsidae Unknown Sepsidae barr-02 0 Sphaeroceridae Unknown barr-01 Sphaeroceridae Syrphidae Unknown Syrphidae barr-01 Tabanidae 86 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 03100 110012100 010142110 010002000 010001000 000000010 100003100 102004110 111000110 110001100 011000100 014020000 010000100 010000000 102000100 100000100 010000000 000001000 000200100 000010100 010001000 010001100 010004000 110000000 010000000 sp. 1 sp. A sp. Lep 161 Genus Group Patanothrix nr Taenogera Erioptera pilipes Digama marmorea Anthela Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Tabanidae Unknown barr-01 Tabanidae Tachinidae Unknown barr-01 Tachinidae Tachinidae Unknown barr-02 Tachinidae Tachinidae Unknown barr-03 Tachinidae Tachinidae Unknown barr-04 Tachinidae Tachinidae Unknown barr-05 Tachinidae Tachinidae Unknown barr-06 Tachinidae 1 0 0 0 Tachinidae Unknown barr-07 Tachinidae Tephritidae Unknown barr-01 Tephritidae Tephritidae Unknown barr-02 Tephritidae Tephritidae Unknown barr-03 Tephritidae Tephritidae Unknown barr-04 Tephritidae Tephritidae Unknown barr-05 Tephritidae Tephritidae Unknown barr-06 Tephritidae Tephritidae Unknown barr-07 Tephritidae Tephritidae Unknown barr-08 Tephritidae Tethinidae Unknown barr-01 Tethinidae Therevidae Therevidae Tipulidae Tipulidae Unknown barr-01 Tipulidae Tipulidae Unknown barr-02 Tipulidae Tipulidae Unknown barr-03 Tipulidae INSECTA: Lepidoptera INSECTA: Aganaidae Aganainae Anthelidae Anthelinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 87 Other Collection 00000 010000000 010000000 010000000 111001000 010000000 110000100 010000000 110000000 110000000 010000000 100000000 010000000 010000000 110000000 010000000 110000000 010000000 110000000 010000000 010000000 110000000 010000000 010000000 110000000 110100000 aperta sp. Lep 166 sp. Lep 167 cf. sp. Lep 150 sp. Lep 109 sp. Lep108 sp. Lep 072 sp. Lep 096 p 000000001 sp. sp. Lep 018 sp. Lep 145 Nausinoe Protolechia Scirpophaga Anthela Anthela Aloa marginata Utetheisa pulchelloides Amata Amata Unknown Achyra affinitalis Achyra massalis cf. Deuterarcha xanthomela Diaphania indica Dysallacta negatalis Glycythyma leonina Hellula hydralis Sameodes iolealis Sceliodes cordalis Spoladea recurvalis Unknown Unknown Uresiphita ornithopteralis Scoparia anthraias nr cf. Unknown Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Anthelidae Anthelinae Anthelidae Anthelinae Arctiidae Arctiinae Arctiidae Arctiinae Arctiidae Ctenuchinae Arctiidae Ctenuchinae cf. Tineidae Unknown sp. 2 0 1 0 0 Crambidae Crambinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Pyraustinae Crambidae Scopariinae Crambidae Scopariinae Unknown sp. Lep 147 Crambidae Shoenobiinae Gelechiidae Gelechiidae 88 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00200 110000000 110000000 110000000 110100000 110000000 100000000 010000000 110100000 110200000 010000000 110000000 110000000 010100000 100000000 100000000 110000000 010000000 110000000 110000000 010000000 010000000 110000000 albitacta poecilatis Gelechioidea sp. Lep 058 sp. 1 sp. 2 sp. Lep 014 sp. Lep 002 sp. Lep 035 sp. Lep 008 1 1 0 0 sp. Lep 121 nr nr sp. Lep 141 sp. Lep 151 p 000000001 sp. sp. Lep 178 Gelechioidea sp. Lep 044 Gelechioidea sp. Lep 058 Gelechioidea sp. Lep 087 Gelechioidea sp. Lep 087 Msp. 039 Msp. 041 Msp. 043 Msp. 046 Msp. 047 Macrobathra Macrobathra Cleora Comostola cf. cf. Unknown cf. cf. Chlorocoma Unknown Dichromodes Dichromodes Dichromodes Taxeotis intextata Scopula Acrocercops Unknown Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Gelechioidea Gelechoidea Cosmopteriginae Gelechoidea Cosmopteriginae Gelechoidea Geometridae Ennominae Geometridae Geometrinae Geometridae Geometrinae Geometridae Lacentiinae Geometridae Oenochrominae Geometridae Oenochrominae Geometridae Oenochrominae Geometridae Oenochrominae Geometridae Sterrhinae Gracillariidae Lasiocampidae Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 89 Other Collection 00000 100000000 110000100 100000000 100000000 110000000 010000000 110000000 010000000 010000000 010000000 010000000 010000000 010000000 110000000 010000000 110000000 010000000 010000000 010000000 010000000 010000000 010000000 010000010 100000000 010000000 110000000 Msp. 050 Msp. 053 Msp. 054 Msp. 073 Msp. 076 Msp. 086 Msp. 092 0 1 0 0 Msp. 093 Msp. 106 Msp. 111 Msp. 117 Msp. 118 Msp. 123 Msp. 130 Msp. 140 Msp. 142 Msp. 144 Msp. 148 Msp. 169 Msp. 172 Msp. 177 Msp. 183 Msp. 190 sp. Lep 180 sp. Lep 009 Nacaduba biocellata Theclinesthes miskini Unknown Euproctis Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lycaenidae Polyommatinae Lycaenidae Polyommatinae Lycaenidae Lymantriidae 90 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 010000000 100000000 100000000 110000000 100000000 100000000 110000000 010000000 010000000 010000000 010000000 010000000 110000000 010000000 110000000 110000000 000000001 110100100 110000000 110000000 110000100 100000000 100000000 010000000 110000100 110000000 110000000 marmarinopa sp. Lep 185 sp. Lep 113 sp. Lep 100 sp. Lep 120 sp. Lep 179 nr sp. Lep 005 Unknown Acontia clerana Acontia crocata Eublemma anachoresis Eublemma vestalis Ipanica cornigera Athetis tenuis Eremechroa Proteuxoa interferens Proteuxoa Proteuxoa Unknown Achaea janata Anomis involuta Dasypodia selenophora Diatenes aglossoides Diatenes Donuca memorabilis Eudesmeola lawsoni Eudocima materna Grammodes pulcherrima Mocis alterna Ophiusa parcemacula Pandesma submurina Pantydia capistrata Pantydia Prorocopis melanochorda Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Nepticulidae Noctuidae Acontiinae Noctuidae Acontiinae Noctuidae Acontiinae Noctuidae Acontiinae Noctuidae Agaristinae Noctuidae Amphipyrinae Noctuidae Amphipyrinae Noctuidae Amphipyrinae Noctuidae Amphipyrinae Noctuidae Amphipyrinae Noctuidae Amphipyrinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae Catocalinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 91 Other Collection 10000000 010000000 010000000 010000000 010000000 010000000 010000000 010000000 100000000 010000000 110000000 110000000 110000000 010000000 100000000 010100000 100010000 110000000 100000000 100000000 110000000 010000000 010000000 010000000 010000000 010000000 110000000 sp. Lep 081 sp. Lep 112 sp. Lep 114 sp. Lep 129 sp. Lep 119 sp. Lep 067 sp. Lep 128 sp. Lep 037 sp. Lep 165 sp. Lep 101 sp. Lep 034 sp. Lep 083 sp. Lep 124 sp. Lep 127 sp. Lep 171 0 sp. Lep 174 sp. Lep 182 Catoblemma Unknown Unknown Unknown Unknown Leucania cruegeri Leucania Leucania stenographa Helicoverpa punctigera Heliocheilus atrilinea Heliocheilus cladotus Heliothis punctifera Unknown Unknown Unknown Unknown Agrotis munda Chrysodeixis argentifera Pataeta cf. Unknown Unknown Unknown Unknown Unknown Unknown Armactica columbina Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Noctuidae Catocalinae Noctuidae Catocalinae Noctuidae cf. Hadeninae Noctuidae cf. Hadeninae Noctuidae Hadeninae Noctuidae Hadeninae Noctuidae Hadeninae Noctuidae Heliothinae Noctuidae Heliothinae Noctuidae Heliothinae Noctuidae Heliothinae Noctuidae Heliothinae Noctuidae Heliothinae Noctuidae Hypeninae Noctuidae Hypeninae Noctuidae Noctuinae Noctuidae Plusiinae Noctuidae Stictopterinae Noctuidae Noctuidae Noctuid sp. 1 Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Nolidae Chloephorinae 92 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 010000000 010000000 110000000 100000000 110000000 110000000 010000000 110000000 010000000 010000000 000000001 000000001 110000000 000003100 000000001 010100000 100000000 010000000 100000000 010000000 000000001 010001000 010010000 010000000 110000000 100000000 sp. Lep 059 sp. Lep 110 0 1 0 0 sp. Lep 146 sp. Lep 188 sp. Lep 078 sp. Lep 011 Phycitine Earias chlorodes Earias parallela Unknown Ochrogaster lunifer Danaus chrysippus Junonia villida Philobota Philobota Maroga melanostigma Papilo demoleus Belenois java teutonia Plutella xylostella Lepidoscia Paracharactis cf. Etiella behri Plodia interpunctella Phostria odontostica Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Nolidae Chloephorinae Nolidae Chloephorinae Nolidae Chloephorinae Notodontidae Thaumetopoeinae Nymphalidae Danainae Nymphalidae Nymphalinae Oecophoridae Oecophorinae Oecophoridae Oecophorinae Oecophoridae Oecophorinae Unknown sp. Lep 149 Oecophoridae Oecophorinae Unknown sp. Lep 184 Oecophoridae Oecophorinae Unknown sp. Lep 187 Oecophoridae Xyloryctinae Oecophoridae Oecophorid sp. 2 Papilionidae Pieridae Pierinae Plutellidae Psychidae Psychidae Pterophoridae Unknown sp. Lep 170 Pyralidae Phycitinae Pyralidae Phycitinae Pyralidae Phycitinae Pyralidae Phycitinae Unknown sp. Lep 135 Pyralidae Pyralidae Unknown sp. Lep 091 Pyraloidea Unknown sp. Lep 057 Pyraloidea Unknown sp. Lep 074 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 93 Other Collection 00000 010000000 010000000 010000000 010000000 000000001 110100000 110000000 010000000 010000000 010000000 010000000 010000000 010000000 010000000 000000110 000001110 101000000 000001110 100000000 001010100 000011010 100000000 010000110 group plebeia castor sp. Lep 095 p 100111000 sp. sp. nr sp. 1 cf. sp. 2 Hippotion celerio Agrius convulvuli Hyles lineata livornicoides Edosa Amegilla Braunsapis Leioproctus Ceylalictus perditellus Lasioglossum Lasioglossum ebeneum Lasioglossum erythrurum Lasioglossum Lipotriches flavoviridis Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Pyraloidea Unknown sp. Lep 105 Pyraloidea Unknown sp. Lep 116 Sphingidae Macroglossinae Sphingidae Sphinginae Sphingidae Tineidae Perissomasticinae Tineidae Unknown sp. 1 1 0 0 0 Tineidae Unknown sp. Lep 061 Tineidae Unknown sp. Lep 175 Tineidae Unknown sp. Lep 176 Tortricidae Olethreutinae Unknown sp. Lep 136 Tortricidae Olethreutinae Unknown sp. Lep 137 Tortricidae Tortricinae Unknown sp. Lep 189 Tortricidae Unknown sp. Lep 173 Zyganoidea Unknown sp. Lep 098 INSECTA: Hymenoptera: Bees INSECTA: Apidae Apinae Apidae Xylocopinae Apidae Xylocopinae Unknown Allodapini sp. Colletidae Colletinae Halictidae Halictinae Halictidae Halictinae Halictidae Halictinae Halictidae Halictinae Halictidae Halictinae Halictidae Nomiinae 94 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 11100 000001000 000001000 011111110 100000000 001000100 001001010 001000000 100000110 100000110 101011010 100010000 111001100 101001100 101001110 010011010 010000000 010001110 100001100 010001110 000001000 110010100 100000000 100000110 110010000 sp. 1 sp. 1 sp. 1 sp. 1 Lipotriches Megachile Pleistodontes Dolichurus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Halictidae Nomiinae Megachilidae Megachilinae INSECTA: Hymenoptera: Wasps INSECTA: Agaonidae Agaoninae Ampulicidae Aphelinidae Unknown sp. 1 Aphelinidae Unknown sp. 2 Aphelinidae Unknown sp. 3 0 0 0 0 Aphelinidae Unknown sp. 4 Bethylidae Bethylinae Unknown sp. 11 Bethylidae Bethylinae Unknown sp. 15 Bethylidae Epyrinae Unknown sp. 1 Bethylidae Epyrinae Unknown sp. 10 Bethylidae Epyrinae Unknown sp. 12 Bethylidae Epyrinae Unknown sp. 13 Bethylidae Epyrinae Unknown sp. 14 Bethylidae Epyrinae Unknown sp. 16 Bethylidae Epyrinae Unknown sp. 18 Bethylidae Epyrinae Unknown sp. 19 Bethylidae Epyrinae Unknown sp. 2 Bethylidae Epyrinae Unknown sp. 20 Bethylidae Epyrinae Unknown sp. 22 Bethylidae Epyrinae Unknown sp. 3 Bethylidae Epyrinae Unknown sp. 4 Bethylidae Epyrinae Unknown sp. 5 Bethylidae Epyrinae Unknown sp. 6 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 95 Other Collection 01100 01000110 110001010 100001000 100000100 000010000 000001000 100000000 010000010 010000110 010001110 010000000 110101110 000010000 101000100 100000000 100000100 101111110 101001110 100001110 100001100 100001110 110001100 110001110 100000110 110001100 100000010 sp. 21 sp. 19 sp. 33 0 1 0 0 sp. 36 sp. 40 sp. 51 sp. 52 sp. 8 sp. 54 Apenesia Chelonus Chelonus Chelonus Chelonus Phanerotoma Phanerotoma Phanerotoma Spathius Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Bethylidae Epyrinae Unknown sp. 7 Bethylidae Epyrinae Unknown sp. 8 Bethylidae Epyrinae Unknown sp. 9 Bethylidae Pristocerinae Braconidae Braconinae Unknown sp. 55 Braconidae Chelioninae Braconidae Chelioninae Braconidae Chelioninae Braconidae Chelioninae Braconidae Chelioninae Braconidae Chelioninae Braconidae Chelioninae Braconidae Doryctinae Braconidae Euphorinae Unknown sp. 2 Braconidae Euphorinae Unknown sp. 26 Braconidae Euphorinae Unknown sp. 30 Braconidae Unknown sp. 1 Braconidae Unknown sp. 10 Braconidae Unknown sp. 11 Braconidae Unknown sp. 14 Braconidae Unknown sp. 15 Braconidae Unknown sp. 16 Braconidae Unknown sp. 17 Braconidae Unknown sp. 20 1 Braconidae Unknown sp. 22 Braconidae Unknown sp. 23 Braconidae Unknown sp. 24 96 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 10100 00100100 100000000 111001110 101001010 100001110 010000100 010000100 110000000 010000010 101001110 010000100 010001110 010001110 010101110 010101000 010001100 010001110 010101000 010001000 101000000 010000000 010001000 110001110 110000110 100000010 100000100 Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Braconidae Unknown sp. 25 Braconidae Unknown sp. 27 Braconidae Unknown sp. 28 Braconidae Unknown sp. 3 Braconidae Unknown sp. 32 Braconidae Unknown sp. 34 Braconidae Unknown sp. 35 0 1 0 0 Braconidae Unknown sp. 37 Braconidae Unknown sp. 39 Braconidae Unknown sp. 4 Braconidae Unknown sp. 41 Braconidae Unknown sp. 42 Braconidae Unknown sp. 43 Braconidae Unknown sp. 44 Braconidae Unknown sp. 45 Braconidae Unknown sp. 46 Braconidae Unknown sp. 47 Braconidae Unknown sp. 48 Braconidae Unknown sp. 49 Braconidae Unknown sp. 5 Braconidae Unknown sp. 50 Braconidae Unknown sp. 53 Braconidae Unknown sp. 6 Braconidae Unknown sp. 7 1 Ceraphronidae Unknown sp. 1 Ceraphronidae Unknown sp. 10 Ceraphronidae Unknown sp. 11 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 97 Other Collection 00110 10001000 100001010 100000010 010000000 010000000 010000000 010000100 110000110 110000110 110000100 100000000 100000100 110001110 100001010 000000100 000010000 100000110 000001000 101001110 110000100 100000000 011000000 011000100 000001010 000010100 010000000 sp. 1 sp. 6 sp. 8 Antrocephalus Haltichiella Acanthostethus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Ceraphronidae Unknown sp. 12 Ceraphronidae Unknown sp. 13 Ceraphronidae Unknown sp. 15 Ceraphronidae Unknown sp. 16 Ceraphronidae Unknown sp. 17 Ceraphronidae Unknown sp. 18 Ceraphronidae Unknown sp. 19 0 1 0 0 Ceraphronidae Unknown sp. 2 Ceraphronidae Unknown sp. 3 Ceraphronidae Unknown sp. 4 Ceraphronidae Unknown sp. 5 Ceraphronidae Unknown sp. 6 Ceraphronidae Unknown sp. 7 Ceraphronidae Unknown sp. 8 Chalcididae Chalcididae Chalcididae Unknown sp. 1 Chalcididae Unknown sp. 10 Chalcididae Unknown sp. 2 Chalcididae Unknown sp. 3 Chalcididae Unknown sp. 4 Chalcididae Unknown sp. 5 Chalcididae Unknown sp. 7 Chalcididae Unknown sp. 8 0 Chalcididae Unknown sp. 9 Chrysididae Unknown sp. 1 Crabronidae 98 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00010 00001010 000000110 100011010 100000000 101000000 010111100 000001000 000001000 000000010 000010000 000001000 110001000 000001000 110001100 100001100 100011110 101101100 110001010 000001010 000001000 000001000 000001000 000001000 100000100 110011110 011011110 sp. 23 sp. 2 sp. 4 sp. 1 sp. 10 Arnactophilus Bembecinus Bembix Lyroda Tachysphex Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Crabronidae Crabronidae Crabronidae Crabronidae Crabronidae Crabronidae Unknown sp. 12 Crabronidae Unknown sp. 17 0 0 0 0 Crabronidae Unknown sp. 18 Crabronidae Unknown sp. 19 Crabronidae Unknown sp. 21 Crabronidae Unknown sp. 22 Diapriidae Ambositrinae Unknown sp. 3 Diapriidae Belytinae Unknown sp. 4 Diapriidae Diapriinae Unknown sp. 1 Diapriidae Diapriinae Unknown sp. 2 Dryinidae Gonatopodinae Unknown sp. 3 Dryinidae Gonatopodinae Unknown sp. 6 Dryinidae Unknown sp. 1 Dryinidae Unknown sp. 10 Dryinidae Unknown sp. 11 Dryinidae Unknown sp. 12 Dryinidae Unknown sp. 13 Dryinidae Unknown sp. 14 Dryinidae Unknown sp. 2 1 Dryinidae Unknown sp. 4 Dryinidae Unknown sp. 5 Dryinidae Unknown sp. 7 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 99 Other Collection 11100 00010000 010000000 001001000 000111110 001000100 001001010 001011010 001011100 001010000 001011000 100000010 000010000 000010100 110000110 010001110 000000010 000001000 000001110 110011110 101000110 011111110 110000001 010001110 001111000 001011110 001111110 n. sp. sp. 1 0 Elasmus arumburinga Elasmus auratiscutellum Elasmus bellicaput Elasmus broomensis Elasmus curticornis Elasmus emma ero Elasmus flavipropleurum Elasmus formosus nakomara Elasmus funereus Elasmus neofunereus Elasmus tenebrosus Quadrastichus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Dryinidae Unknown sp. 8 Dryinidae Unknown sp. 9 Encyrtidae Unknown sp. 1 Encyrtidae Unknown sp. 2 Encyrtidae Unknown sp. 4 Encyrtidae Unknown sp. 5 Encyrtidae Unknown sp. 6 0 0 1 0 Encyrtidae Unknown sp. 7 Encyrtidae Unknown sp. 8 Encyrtidae Unknown sp. 9 Eucharitidae Unknown sp. 1 Eucharitidae Unknown sp. 2 Eulophidae Eulophidae Eulophidae Eulophidae Eulophidae Eulophidae Eulophidae Eulophidae Eulophidae Eulophidae Eulophidae Eulophidae Eulophidae Unknown sp. 1 Eulophidae Unknown sp. 10 Eulophidae Unknown sp. 2 100 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00110 10011010 001011010 001011110 001111100 001011110 001011010 000111100 100000000 011010000 010010000 010001110 000001010 111011110 110000010 100010000 101011000 110001110 100000010 101100000 101101110 000001000 111011110 001001110 111001110 101001110 100011010 sp. 11 Sycophila cf. Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Eulophidae Unknown sp. 3 Eulophidae Unknown sp. 4 Eulophidae Unknown sp. 5 Eulophidae Unknown sp. 6 Eulophidae Unknown sp. 7 Eulophidae Unknown sp. 8 Eulophidae Unknown sp. 9 0 0 0 0 Eupelmidae Unknown sp. 1 Eupelmidae Unknown sp. 10 Eupelmidae Unknown sp. 11 Eupelmidae Unknown sp. 12 Eupelmidae Unknown sp. 13 Eupelmidae Unknown sp. 2 Eupelmidae Unknown sp. 3 Eupelmidae Unknown sp. 4 Eupelmidae Unknown sp. 5 Eupelmidae Unknown sp. 6 Eupelmidae Unknown sp. 7 Eupelmidae Unknown sp. 8 Eupelmidae Unknown sp. 9 Eurytomidae Eurytomidae Unknown sp. 1 Eurytomidae Unknown sp. 10 Eurytomidae Unknown sp. 2 1 Eurytomidae Unknown sp. 3 Eurytomidae Unknown sp. 4 Eurytomidae Unknown sp. 5 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 101 Other Collection 01000 00010000 010011100 010001000 010011110 000001110 100001100 110001110 000010000 000001010 000001000 100011100 110011000 010001010 100101100 110000100 101000000 100001110 100100100 110010000 011001110 010001110 110001000 110001110 100000000 100000000 100001000 sp. 1 sp. 2 Acanthinevania Acanthinevania Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Eurytomidae Unknown sp. 6 Eurytomidae Unknown sp. 7 Eurytomidae Unknown sp. 8 Eurytomidae Unknown sp. 9 Evaniidae Evaniidae Evaniidae Unknown sp. 3 0 0 0 0 Evaniidae Unknown sp. 4 Evaniidae Unknown sp. 5 Evaniidae Unknown sp. 6 Figitidae Eucoilinae Unknown sp. 1 Figitidae Eucoilinae Unknown sp. 2 Figitidae Eucoilinae Unknown sp. 3 Ichneumonidae Unknown sp. 1 Ichneumonidae Unknown sp. 2 Ichneumonidae Unknown sp. 3 Ichneumonidae Unknown sp. 4 Ichneumonidae Unknown sp. 5 Ichneumonidae Unknown sp. 6 Ichneumonidae Unknown sp. 7 Megaspilidae Unknown sp. 1 Mutillidae Unknown sp. 1 Mutillidae Unknown sp. 11 Mutillidae Unknown sp. 12 1 Mutillidae Unknown sp. 13 Mutillidae Unknown sp. 14 Mutillidae Unknown sp. 15 102 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 01000010 010001000 010000000 010000000 010001000 110010000 010000000 010000100 010010000 011011000 110011010 000010000 000001000 000001110 000010000 110001000 100001000 110001000 100001010 100001110 111011110 100001000 000000010 001010100 001011100 000000100 Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Mutillidae Unknown sp. 16 Mutillidae Unknown sp. 17 Mutillidae Unknown sp. 18 Mutillidae Unknown sp. 19 Mutillidae Unknown sp. 2 Mutillidae Unknown sp. 20 Mutillidae Unknown sp. 21 0 1 0 0 Mutillidae Unknown sp. 22 Mutillidae Unknown sp. 23 Mutillidae Unknown sp. 24 Mutillidae Unknown sp. 25 Mutillidae Unknown sp. 26 Mutillidae Unknown sp. 27 Mutillidae Unknown sp. 28 Mutillidae Unknown sp. 29 Mutillidae Unknown sp. 3 Mutillidae Unknown sp. 4 Mutillidae Unknown sp. 5 Mutillidae Unknown sp. 6 Mutillidae Unknown sp. 7 Mutillidae Unknown sp. 8 Mutillidae Unknown sp. 9 Mymaridae Unknown sp. 1 Mymaridae Unknown sp. 10 0 Mymaridae Unknown sp. 11 Mymaridae Unknown sp. 2 Mymaridae Unknown sp. 3 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 103 Other Collection 00010 001001000 000001000 100000000 001011110 001000110 001010110 010001000 100001010 100001000 100000010 100000010 100000010 110001100 110001010 100000010 010000010 010000000 010001000 010001010 010000000 101011010 100000010 100000000 110000010 010000000 100000000 sp. 1 sp. 4 sp. 5 sp. 7 sp. 8 Inostemma Inostemma Inostemma Inostemma Inostemma Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Mymaridae Unknown sp. 4 Mymaridae Unknown sp. 5 Mymaridae Unknown sp. 6 Mymaridae Unknown sp. 7 Mymaridae Unknown sp. 8 Mymaridae Unknown sp. 9 Ormyridae Unknown sp. 1 0 0 0 0 Perilampidae Unknown sp. 1 Platygastridae Platygastrinae Platygastridae Platygastrinae Platygastridae Platygastrinae Platygastridae Platygastrinae Platygastridae Platygastrinae Platygastridae Platygastrinae Unknown sp. 10 Platygastridae Platygastrinae Unknown sp. 11 Platygastridae Platygastrinae Unknown sp. 12 Platygastridae Platygastrinae Unknown sp. 14 Platygastridae Platygastrinae Unknown sp. 15 Platygastridae Platygastrinae Unknown sp. 16 Platygastridae Platygastrinae Unknown sp. 17 Platygastridae Platygastrinae Unknown sp. 18 Platygastridae Platygastrinae Unknown sp. 2 Platygastridae Platygastrinae Unknown sp. 3 Platygastridae Platygastrinae Unknown sp. 9 Platygastridae Unknown sp. 13 Platygastridae Unknown sp. 19 Platygastridae Unknown sp. 6 104 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 110101000 110011110 010010100 110001100 110010100 101010000 001001100 001001110 001011100 001001010 001011110 000111010 001001010 100000000 000001000 110000000 110000000 010010000 011001000 010010000 110101110 110011010 110000110 110011110 110001010 110001010 sp. 4 sp. 3 sp. 6 sp. 1 sp. 2 sp. 8 sp. 1 sp. 1 sp. 2 sp. 1 sp. 2 sp. 29 sp. 38 sp. 39 sp. 2 Pachycurjus Mirobaeoides cf. Fabriogenia Agenoideus Ctenostegus Turneromyia Turneromyia Notanisus Sycoscapter Sycoscapter Olixon Olixon Ceratobaeus Ceratobaeus cf. Oxyscelio Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Pompilidae Pepsinae Pompilidae Pepsinae Pompilidae Pompilinae Pompilidae Pompilinae Pompilidae Pompilinae Pompilidae Pompilinae Pompilidae Unknown sp. 5 1 0 1 0 Proctotrupidae Unknown sp. 1 Pteromalidae Ormorcerinae Pteromalidae Sycoryctinae Pteromalidae Sycoryctinae Pteromalidae Unknown sp. 1 Pteromalidae Unknown sp. 4 Pteromalidae Unknown sp. 5 Pteromalidae Unknown sp. 6 Pteromalidae Unknown sp. 7 Rhopalosomatidae Rhopalosomatidae Scelionidae Scelioninae Scelionidae Scelioninae Scelionidae Scelioninae Scelionidae Scelioninae Scelionidae Scelioninae Unknown sp. 1 Scelionidae Scelioninae Unknown sp. 11 Scelionidae Scelioninae Unknown sp. 12 Scelionidae Scelioninae Unknown sp. 14 Scelionidae Scelioninae Unknown sp. 15 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 105 Other Collection 01110 100001000 110000100 101001000 100010000 111001110 010000110 010000100 010011000 010001110 010101010 010001100 011001110 111001100 010000110 110101110 110001110 100000000 110001110 100001100 100100100 100000110 010001110 010001110 010011000 111001110 100111110 Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Scelionidae Scelioninae Unknown sp. 16 Scelionidae Scelioninae Unknown sp. 19 Scelionidae Scelioninae Unknown sp. 21 Scelionidae Scelioninae Unknown sp. 22 Scelionidae Scelioninae Unknown sp. 23 Scelionidae Scelioninae Unknown sp. 26 Scelionidae Scelioninae Unknown sp. 28 0 1 0 1 Scelionidae Scelioninae Unknown sp. 30 Scelionidae Scelioninae Unknown sp. 31 Scelionidae Scelioninae Unknown sp. 32 Scelionidae Scelioninae Unknown sp. 33 Scelionidae Scelioninae Unknown sp. 35 Scelionidae Scelioninae Unknown sp. 37 Scelionidae Scelioninae Unknown sp. 4 Scelionidae Scelioninae Unknown sp. 40 Scelionidae Scelioninae Unknown sp. 6 Scelionidae Scelioninae Unknown sp. 8 Scelionidae Telenominae Unknown sp. 10 Scelionidae Telenominae Unknown sp. 13 Scelionidae Telenominae Unknown sp. 17 Scelionidae Telenominae Unknown sp. 18 Scelionidae Telenominae Unknown sp. 20 Scelionidae Telenominae Unknown sp. 24 Scelionidae Telenominae Unknown sp. 25 Scelionidae Telenominae Unknown sp. 27 Scelionidae Telenominae Unknown sp. 3 Scelionidae Telenominae Unknown sp. 5 106 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 01000 00001000 110000010 100001010 000010000 000000101 111001110 110001000 010001000 010001000 000010000 000001000 100000000 010000000 010001000 010001000 110000000 100000000 110001000 100001110 000001000 000001000 100001000 000001000 010000010 000001000 globosus sp. 1 sp. 1 sp. 2 1 1 0 0 sp. 3 sp. 1 cf. sp. 1 sp. 7 sp. 8 sp. 10 sp. 9 sp. 1 sp. 4 sp. 3 sp. 2 Probethylus Caetrathynnus cf. Radumeris radula Prionyx Prionyx Prionyx Sphex Tanaostigmodes Unknown Caetrathynnus Caetrathynnus cf. Chilothynnus Eirone Eirone Epactiothynnus Gen. n. cf. Gymnothynnus Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Scelionidae Telenominae Unknown sp. 7 Scelionidae Telenominae Unknown sp. 9 Sclerogibbidae Scoliidae Signiphoridae Unknown sp. 1 Sphecidae Sphecidae Sphecidae Sphecidae Tanaostigmatidae Tetracampidae Tiphiidae Thynninae Tiphiidae Thynninae Tiphiidae Thynninae Tiphiidae Thynninae Tiphiidae Thynninae Tiphiidae Thynninae Tiphiidae Thynninae Tiphiidae Thynninae Tiphiidae Thynninae Unknown sp. 11 Tiphiidae Thynninae Unknown sp. 12 Tiphiidae Thynninae Unknown sp. 6 Tiphiidae Unknown sp. 13 Tiphiidae Unknown sp. 14 0 Tiphiidae Unknown sp. 15 Tiphiidae Unknown sp. 16 TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 107 Other Collection 11000 000001000 111100000 011000000 010000100 100001010 011000110 000001000 100000000 001011100 001001000 000001010 001011000 001011000 100000000 100000000 110000000 010110000 010000000 010000000 010000010 010000000 000100000 011100000 p 010000000 sp. sp. JDM 1103 sp. JDM 1170 sp. JDM 942 Aenictus turneri Amblyopone Cerapachys brevis Cerapachys longitarsus Cerapachys ruficornis Cerapachys Cerapachys Cerapachys Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Tiphiidae Unknown sp. 17 Torymidae Megastigminae Unknown sp. 2 Torymidae Megastigminae Unknown sp. 5 Torymidae Megastigminae Unknown sp. 6 Torymidae Monodontomerinae Unknown sp. 1 Torymidae Monodontomerinae Unknown sp. 4 Torymidae Toryminae Unknown sp. 3 0 1 0 0 Trichogrammatidae Unknown sp. 1 Trichogrammatidae Unknown sp. 2 Trichogrammatidae Unknown sp. 3 Trichogrammatidae Unknown sp. 4 Trichogrammatidae Unknown sp. 5 Trichogrammatidae Unknown sp. 6 Trichogrammatidae Unknown sp. 7 Trichogrammatidae Unknown sp. 9 Vespidae Eumeninae Unknown sp. 1 Vespidae Eumeninae Unknown sp. 2 INSECTA: Hymenoptera: Ants INSECTA: Formicidae Aenictinae Formicidae Amblyoponinae Formicidae Cerapachyiinae Formicidae Cerapachyiinae Formicidae Cerapachyiinae Formicidae Cerapachyiinae Formicidae Cerapachyiinae Formicidae Cerapachyiinae 108 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 100000000 011000000 010000000 111101100 111111110 111011110 111111110 010011000 011101100 000000010 111111110 011011110 111111010 111111110 111101110 111111110 010100000 011111110 111111110 111011110 001001000 010000000 111111110 111111110 111011110 111101110 complex sp. micans sp. JDM 433 sp. JDM 527 sp. JDM 78 sp. JDM 981 cf. JDM 1129 Sphinctomyrmex duchaussoyi Arnoldius Doleromyrma rottnestense Iridomyrmex agilis Iridomyrmex anceps Iridomyrmex cephaloinclinus Iridomyrmex chasei Iridomyrmex coeruleus Iridomyrmex discors Iridomyrmex dromus Iridomyrmex exsanguis Iridomyrmex gibbus Iridomyrmex hartmeyeri Iridomyrmex minor Iridomyrmex mjobergi Iridomyrmex sanguineus Iridomyrmex tenuiceps Ochetellus flavipes Ochetellus Tapinoma Tapinoma Rhytidoponera Rhytidoponera crassinoda Rhytidoponera taurus Rhytidoponera tyloxys Camponotus capito Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Formicidae Cerapachyiinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Dolichoderinae Formicidae Ectatomminae Formicidae Ectatomminae Formicidae Ectatomminae Formicidae Ectatomminae Formicidae Formicinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 109 Other Collection 111111110 111111110 110011110 010010000 000100000 111111110 011001110 110010110 111001110 000000010 111011110 111111110 000011110 100000000 111001010 111111110 111011010 111110000 010000000 111111110 111011110 010010000 111010000 111011110 111111110 sp. gp sp. complex sp. sp. JDM 1116 complex sp. evae aeneovirens cf. evae JDM 1158 bruneus JDM 520 nr JDM 545 sp. JDM 1063 sp. JDM 532 sp. JDM 897 minutula JDM 916 Camponotus Camponotus discors Camponotus donnellani Camponotus Camponotus evae zeuxis Camponotus fieldeae Camponotus scratius Camponotus simpsoni Melophorus Melophorus insularis Melophorus ludius Melophorus ludius sulla Melophorus marius Melophorus Melophorus Melophorus Melophorus Melophorus turneri Nylanderia braueri glabrior Opisthopsis haddoni rufoniger Paraparatrechina minutula Paraparatrechina Paratrechina longicornis Polyrhachis ammonoeides Polyrhachis ?melanura Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae 110 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 00000 100010000 111101110 000000100 111010000 111011010 010100000 011111000 111111110 000100000 000000001 110001110 000010110 111101100 100001000 000010000 110000000 111110000 000000001 000111010 010000000 010000000 101011010 110111110 101000000 111111110 gp sp. complex sp. sp. JDM 1009 sp. JDM 1010 sp. JDM 703 0 1 0 0 sp. JDM 807 sp. JDM 808 sp. JDM 1131 queenslandica JDM 1132 puryi JDM 889 sp. JDM 1133 sp. JDM 268 sp. JDM 865 Polyrhachis gravis Polyrhachis inconspicua Polyrhachis senilis Polyrhachis Polyrhachis Polyrhachis seducta Polyrhachis Polyrhachis Polyrhachis Polyrhachis bohemia Stigmacros termitoxena Leptanilla swani Cardiocondyla atalanta Cardiocondyla nuda Carebara Crematogaster laeviceps chasei Crematogaster Meranoplus dimidiatus Meranoplus fenestratus Meranoplus Meranoplus Meranoplus Meranoplus Monomorium antipodum Monomorium arenarium Monomorium disetigerum Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Formicinae Formicidae Leptanillinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae TERRESTRIAL INVERTEBRATE FAUNA OF BARROW ISLAND 111 Other Collection 010010000 001001100 010100010 111011110 110111110 011011000 011011100 011000000 111111110 010110100 110101100 000011000 011000000 110111110 110001010 000000001 110011010 110000000 010000110 111111110 111111110 110001010 001001000 110001000 001001000 111000000 complex sp. sp. JDM 1021 gp sp. n. sp. variabilis tricosa rubriceps JDM 1175 sydneyense JDM 101 nr sp. JDM 1134 sp. JDM 536 sp. JDM 684 sp. JDM 1230 sp. JDM 1142 nr sp. JDM 1128 Monomorium eremophilum Monomorium euryodon Monomorium fieldi Monomorium insolescens Monomorium laeve Monomorium leae Monomorium punctulatum Monomorium Monomorium sydneyense Monomorium Pheidole mjobergi Pheidole Pheidole Pheidole Pheidole Solenopsis belisarius Solenopsis clarki Strumigenys Tetramorium sjostedti Tetramorium spininode Tetramorium striolatum Anochetus rectangularis Anochetus renatae Hypoponera Leptogenys Leptogenys Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Myrmicinae Formicidae Ponerinae Formicidae Ponerinae Formicidae Ponerinae Formicidae Ponerinae Formicidae Ponerinae 112 J.D. MAJER, S. K. CALLAN, K. EDWARDS, N. R. GUNAWARDENE AND C. K. TAYLOR Other Collection 110111110 011100000 010111110 100000000 000010000 010101110 n. sp. sp. JDM 1130 Odontomachus ruficeps Pachycondyla denticulata Pachycondyla lutea Discothyrea Probolomyrmex latalongus Tetraponera punctulata Family Subfamily Genus Species GP NIS S1 S2 C1 C2 C3 C4 Formicidae Ponerinae Formicidae Ponerinae Formicidae Ponerinae Formicidae Proceratiinae Formicidae Proceratiinae Formicidae Pseudomyrmecinae