appointed as an extern for the hospital, his Phycological Trailblazer momentum was interrupted by his being called No. 21 to serve in the military. But his family came to his rescue and persuaded his younger brother to Édouard Bornet take Bornet’s place, an arrangement that was then possible. Bornet was able to connue his studies (originally printed in the Phycological newsleer. 2004. without problems. He came under the wing of Vol. 40 No. 2) Dr. J. H. Léveillé, an authoritave mycologist of the me. Léveillé was careful not to encourage Jean-Bapse Edouard Bornet was born Bornet into a scienfic career without having on Sept. 2, 1828, in Guérigny, , the eldest sufficient means. Bornet was not deterred, and of four sons. His father was a cashier at the in 1851 he had his first publicaon in the Chaussade Foundry. Bornet received his Annales des Sciences Naturalles, a report on the educaon at the college in Nevers. He oen was fungal genus Meliola. The paper was an irritant to his instructors and was threatened accompanied by two plates, which revealed his with expulsion, but the fact arsc talents. Bornet that he was consistently at became acquainted with the top of the class saved the eminent botanists of him from being expelled the me who oen (Flahault, 1924). His gathered at Léveillé’s summers and holidays home. were spent collecng and At a me when Bornet idenfying plants around lacked the resources to his hometown. A senior complete his medical official at the foundry studies, upon the advice of where his father was , Gustav employed noced his Thuret proposed that interest and arranged to Bornet become his loan the young Bornet a assistant, an offer which microscope to encourage Bornet accepted him, and that proved to be (Woelkerling & Lamy, a decisive step in molding 1998). The two went to his future career. Versailles in 1852, where Upon graduang they spent the summer from college with honors in collecng and studying 1846, Bornet was happy to plants, especially the escape from a possible cryptogams. Before the bureaucrac life. His family Édouard Bornet [from Bullen de la Société end of 1852, Bornet and directed him toward Mycologique de France vol. 29, 1913]. Thuret established medicine because of his themselves in Cherbourg, love of nature (and plants). and despite the rigors of He headed for , where his life was eased by the winter season, they iniated their studies on the support of the parents of the young lady marine . They quickly realized that the who was to become his future wife and life-long marine vegetaon and the reproducve state of companion. Ever the eager student, Bornet the algae were very different in the winter divided his me with the faculty of medicine, the months than that of late summer. But Thuret Sorbonne, and the Museum. When he was became ill, brought on both by exhauson and 1 Fig. 1. Cutleria mulfida (Turner) Grev. Pl. IX in Thuret & Fig. 2. Helminthora divaricata (C. Agardh) J. Agardh, Bornet (1878). Pl. XXXII in Thuret & Bornet (1878). the winter condions at Cherbourg. Bornet maneuver over slippery boulders, and he was recognized the need to complete his medical preoccupied with liing the spirits of his paent, training, and so he returned to Paris to take his Thuret. final exams in November of 1854. He presented The constant dampness of the Cherbourg a thesis on the sexual reproducon in both climate eventually led to Thuret’s decision to animals and plants, stressing the plants. He spend the winter at Cannes on the Riviera. He earned his ‘medical qualificaon’ in August 1855 rented a villa and returned the following winter. and soon returned to Cherbourg. Eventually, Thuret purchased a site on the The two men complemented each other Anbes coast where he built a home and where perfectly. Despite having different personalies, he, with Bornet, would carry out their research. a strong friendship and an effecve working The two colleagues had an ideal locaon to relaonship developed between Bornet and pursue their research on marine algae, the home Thuret. Thuret was tall, with a light complexion becoming full of notes, drawings, and (maybe due to his grey hair), spoke slowly, and manuscripts in various stages of compleon. carried himself with a stately demeanor, while Their research encompassed every group of Bornet was short, with a dark complexion, and marine algae, and it was their collaboraon that spoke with much animaon. Thuret came across solved the difficult queson of sexual as slightly English, while Bornet was the reproducon in the red algae. Papenfuss (1955) quintessenal Frenchman (Farlow, 1876). The credited Bornet & Thuret (1866, 1867) as the young doctor was well trained and always ready first to clearly describe sexual reproducon in a to work, an expert photographer, a skilled arst, number of red algae. Thuret employed the arst alert in body and mind. He had the agility to Riocreux, who rendered the illustraons with 2 both great accuracy member, or and beauty. It was corresponding only during the member, of some heat of the summer 27 scienfic that Thuret and sociees around Bornet would move the world northward to spend (Guignard, 1912; me briefly vising De Wildeman, family and friends 1913). in Paris and then on Bornet and his to the Atlanc wife bought a coast before holiday home at returning to Cosne in the center Anbes in the of France, where autumn. they had both a In 1875 Fig. 3. Nemoderma ngitana Schousboe. Pl. I, figs 8-10 in garden and a Thuret suffered a Bornet, 1892. working laboratory. fatal heart aack at The students were the age of 57 also welcomed (Bornet, 1875). He le his library, his collecons, there, as they were in their apartment in Paris. and a number of unfinished manuscripts to Bornet and the students would carry on Bornet, along with a generous endowment to smulang conversaons as they walked along allow Bornet to connue his work the Loire River. The Bornets spent their summers independently. Bornet completed much of the at Cosne and spent the rest of the year in Paris. collaborave work (Bornet & Thuret, 1876, 1880; Toward the end of his long life, Bornet worked to Thuret & Bornet, 1878). Madame Henri Thuret, put his algal herbarium and other affairs into Thuret’s sister-in-law, purchased the villa with order. He donated his collecons, which included the hope that Bornet would stay on and the Thuret herbarium, to the Paris Museum connue the work he had done with Thuret. But (Woelkerling & Lamy, 1998). His health declined Bornet decided to rere to Paris, where he and toward the end of 1910, and he died on 18 his wife occupied the first floor of an old December 1911, at the age of 83. His devoted building. Their apartment became a meeng wife was looked aer by Bornet’s many friends place for younger sciensts. Even though Bornet and survived him by another 10 years (DeToni, had no formal links to a University, he was 1910; Flahault, 1924). regarded as a font of knowledge, to whom many came for scienfic advice. These included Bornet, É. 1875. M. Gustave-Adolphe Thuret. Equisse Maurice Gomont, E. P. Munier-Chalmas, Paul biographique. Ann. Sci. Nat. Bot., sér. 6, 2: Hariot, Charles Flahault, and Anna Vickers 308-360. (Woelkerling & Lamy, 1998). Bornet & Flahault's ____. 1888. Algues du voyage au Golfe de Tadjoura recueillies par M. L. Faurot. Jour. de Bot. 2: 17-20. (1886-1888) monograph of heterocyst- ____. 1892. Les Algues de P.-K.-A. Schousoe, récoltées au containing Nostocaceae has been accepted by Maroc & dans la Méditerranée de 1815 à 1829. the Code to serve as the later starng date of Mémoires de la Société naonale Sciences nomenclature on this algal group. Bornet & naturelles et mathémaques de Cherbourg 28: Flahault (1889) also collaborated on a detailed 165-376, pl. 1-3. examinaon of the algae (e.g., Ostreobium, ______& C. Flahault. 1886-1888. Révision des Gomona, Masgocoleus, et al.) that lived Nostocacées héterocystées connues dans les within calcareous substrates. Bornet had the principal herbiers de France. Ann. Sci. Bot., sér. 7, highest reputaon as a scienst and was a 3: 323-381; 4: 343-373; 5: 51-129; 7: 177-262. 3 [Arcle 13.1(e) of the Internaonal Code of Nomenclature for algae, fungi, and plants (Melbourne Code) treats the four parts of the "Révision" as having been simultaneously published on 1 January 1886.] ____ & ____. 1889. Sur quelques plantes vivant dans le test calcaire des mollusques. Bull. Soc. Bot. France 36: 147-178, pls. VI-XII. ____ & G. Thuret. 1866. Note sur la fécondaon des Floridées. Mém. Soc. Imp. Sci. Nat. Cherbourg 12: 257-262 ____ & ____. 1867. Recherches sur la fécondaon des Floridées. Ann. Sci. Nat.Bot., sér. 5, 7: 137-166, pls 11-13. ____ & ____. 1876. Notes algologiques 1. G. Masson, Paris, 90 pp., 25 pls. ____ & ____. 1880. Notes algologiques 2. G. Masson, Paris, 123 pp., 24 pls. ____ & C. Flahault. 1888. Note sur deux nouveaux genres d’algues perforantes. Jour. de Bot. 2: 161-165. De Toni, G. B. 1912. Edoardo Bornet (1828-1911). Nuova Notarisia 23: 25-42, portrait. De Willdenow, É. 1913. Édouard Bornet (1828- 1911). Bull. Soc. Roy. Belgique 52: 95-110. Flahault, C. 1924. Gustave Thuret – Edouard Bornet. Revue Algologique 1: 4-23, 1 pl. Guignard, L. 1912. Noce sur M. Édouard Bornet. Bull. Soc. Bot. Fr. 59: 257-301. Papenfuss, G. F. 1955. Classificaon of the algae. In: A century of progress in the natural sciences 1853-1953. California Acad. Sci., San Francisco, pp.115-224. Thuret, G., & E. Bornet. 1878. Études phycologiques. Analyses d’algues marines. [v] + iii + 105 pp., 51 pls. G. Masson, Paris. Woelkerling, W. J., & D. Lamy. 1998. Nongeniculate coralline red algae and the Paris Muséum: systemacs and scienfic theory. Muséum naonal d’Historie naturelle, ADAC. viii +767 pp.

I am grateful to Mrs. Lynda Entwisle (Sydney, Australia) for her generous help in translang the Flahault (1924).

Michael J. Wynne University of Michigan

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