J. Jpn. Bot. 89: 279–330 (2014)

Siebold and Zuccarini’s Type Specimens and Original Materials from Japan, Part 5. Angiosperms. Dicotyledoneae 4

a, b,† c Shinobu Akiyama *, Gerard Thijsse , Hans-Joachim Esser d and Hideaki Ohba

aDepartment of Botany, National Museum of Nature and Science, 4-1-1, Amakubo, Tsukuba, 305-0005 JAPAN; bNationaal Herbarium Nederland, Naturalis Biodiversity Center, P. O. Box 9514, THE NETHERLANDS; †Present address: Department of Botany, Naturalis Biodiversity Center, P. O. Box 9517, THE NETHERLANDS; cBotanische Staatssammlung München, Menzinger Straße 67, D-80638 München, GERMANY; dDepartment of Botany, the University Museum, the University of Tokyo, 7-3-1, Hongo, Tokyo, 113-0033 JAPAN *Corresponding author: [email protected]

(Accepted on May 10, 2014)

This is part 5 of the list of type specimens and original materials of the taxa described by Siebold and Zuccarini from Japan. Part 5 concerns the taxa belonging to the families ranging from Saxifragaceae to Euphorbiaceae according to the 12th Engler system (Melchior 1964). The followings five new combination names are proposed. These are: Amygdalus persica L. f. pendula (Siebold) H. Ohba & S. Akiyama, stat. nov., Amygdalus persica L. f. stellata (Siebold & de Vriese) H. Ohba & S. Akiyama, stat. nov., Amygdalus persica L. f. versicolor (Siebold) H. Ohba & S. Akiyama, comb. et stat. nov., Armeniaca mume (Siebold & Zucc.) K. Koch f. pendula (Siebold) H. Ohba & S. Akiyama, comb. et stat. nov., Cerasus itosakura (Siebold) H. Ohba & S. Akiyama, comb. nov. (Continued from J. Jpn. Bot. 89: 209–229, 2014)

Key words: Japanese flora, lectotypification, Siebold collections, Zuccarini.

Saxifragaceae Jap. 1: 19, t. 6 (1835); in Abh. Math.-Phys. Cl. Cardiandra Siebold & Zucc., Fl. Jap. 1: 119 Königl. Bayer. Akad. Wiss. 4(2): 192 (1845). (1835); in Abh. Math.-Phys. Cl. Königl. Bayer. Present treatment: Deutzia crenata Siebold & Akad. Wiss. 4(2): 192 (1845). Zucc. Type: Cardiandra alternifolia (Siebold) Japanese name: Utsugi. Siebold & Zucc. Lectotype (designated here): Japonia. Bürger Present treatment: Cardiandra Siebold & (L0170287) [Category 5 (Fl. Jap. 1: t. 6); the Zucc. branch on the far right corresponds with the plant illustrated in Table 6 of Flora Japonica] Deutzia Thunb. (Fig. 116). 184) Deutzia crenata Siebold & Zucc., Fl. Original material: Herbar. de Siebold

—279— 280 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 coll. 1829 (L0170284, excl. central flowering a local variety, var. sieboldiana (Maxim.) H. branch) [Category 2]. Legit in Japonia Bürger. Hara (D. sieboldii Koernicke) with different Communicavit v. Siebold, Herbarium Zuccarinii tendencies from var. scabra in leaf shape and (M0120974) [Category 2]. Japonia. [Siebold teeth of filaments. s.n.] (MAK S0889) [Category 2]. A specimen collected by Siebold in 1829, The specimen designated here as the L0170284, consists of two species, D. crenata lectotype of Deutzia crenata Siebold & Zucc. and D. scabra, i.e., a branch with flowers was not annotated by Siebold and/or Zuccarini, mounted in the central part of the sheet but the branch on the far right corresponds surrounded by an elongate branch of D. crenata particularly well with the plant illustrated corresponds well with the drawing in Table 7 in Table 6 of their Flora Japonica under of Flora Japonica under D. scabra, which is D. D. crenata. Both the far right branch of the scabra var. sieboldiana. specimen and the branch in the illustration have One of the original specimens collected by widely to moderately ovate petiolate leaves with Bürger, M0120974, consists of two branches. an acute apex and widely cuneate to truncate or Both branches have narrowly oblong-ovate sometimes shallowly cordate base and filaments leaves (with acute or acuminate apex and with terminal teeth, and are within the range of truncate or widely rounded base), which are D. crenata. sessile or nearly so, except the uppermost ones Siebold and Zuccarini distinguished Deutzia on the flowering branches. The right hand crenata from D. scabra Thunb. by 1) the shape branch has a paniculate inflorescence somewhat of leaves, i.e. short petiolate, widely ovate transitional to a pyramidal inflorescence with with acute apex, rounded or subcordate base, numerous flowers. The filaments have two rather lightly or finely “tenuiter” crenulate margins distinct teeth at the apex. We consider these vs. subsessile or very short petiolate, ovate with plants to be D. crenata, but doubt whether they acuminate apex, rounded or subcordate base, might be Deutzia scabra var. sieboldiana still serrulate margins, 2) stellate hairs on upper remains. surface with 4–6 vs. 3 or 4 arms, and 3) filaments with two terminal teeth vs. without teeth. Except 185) Deutzia gracilis Siebold & Zucc., Fl. for the presence of teeth on the filaments these Jap. 1: 22, t. 8 (1835); in Abh. Math.-Phys. Cl. two species are not clearly distinguished by the Königl. Bayer. Akad. Wiss. 4(2): 192 (1845). features given by them. Present treatment: Deutzia gracilis Siebold Usually D. crenata is distinguished by the & Zucc. petiolate leaves (usually oblong with acute Japanese name: Hime-utsugi. or acuminate apex) and dentate filaments; D. Lectotype (designated here): Japonia. scabra has sessile leaves (usually ovate with Herbar. de Siebold coll. 1829 (L0176009) obtuse apex) on flowering branches and toothless [Category 2] (Fig. 117). filaments. In and around Nagasaki the range of Original material: Legit Bürger in Japonia. variation of the leaves, especially in the shape Communicavit v. Siebold, Herbarium Zuccarinii and length of the petioles, and also teeth on the (M0120975) [Category 2]. Japonia. Herb. filaments, overlaps between the two species. Zuccarinii (M0120976) [Category 2]. Japonia. In D. scabra the leaves are usually ovate or [Siebold s.n.] (MAK S0890) [Category 2]. orbicular-ovate or widely oblong-ovate, although in the plants around Nagasaki the leaves show a Hoteia C. Morren & Decne. tendency toward being oblong to oblong-ovate, 186) Hoteia thunbergii Siebold & Zucc. as illustrated in Table 7 of Flora Japonica. As in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. pointed by Hara (1957), these plants belong to Wiss. 4(2): 191 (1845). October 2014 The Journal of Japanese Botany Vol. 89 No. 5 281

Present treatment: Astilbe thunbergii (Siebold Japonia. Herb. Burger (Communicavit Bürger, & Zucc.) Miq., Ann. Mus. Bot. Lugd.-Bat. 3: 96 Hb. Siebold) (L0103049) [Category 2, 5 (left- (1867). hand branch, Fl. Jap. 1: t. 57 II)]. Japanese name: Aka-shôma. Ohba and Akiyama (2013b) recognized Lectotype (designated here): Legit in Hydrangea buergeri Siebold & Zucc. as Japonia & Communicavit de Siebold anno 1842, a variety of H. acuminata because in the Herbarium Zuccarinii (M0120971) [Category 2] text Siebold wrote: Occurrit varietas foliis (Fig. 118). robustioribus longioribusque ... Original material: In Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii 189) Hydrangea alternifolia Siebold in (M0120970) [Category 2b]. Japonia. [Siebold Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. s.n.] (MAK S0408) [Category 2]. Nat. Cur. 14(2): 692 (1828). Siebold and Zuccarini published Hoteia Present treatment: Cardiandra alternifolia thunbergii with a description based on their (Siebold) Siebold & Zucc., Fl. Jap. 1: t. 65 materials. Although they cited Spiraea (1839); 121, t. 66 (1840, versim.). aruncus of Thunberg’s Flora Japonica (1784, Japanese name: Kusa-ajisai; Kusasimots’ke p. 211), there is no indication that they studied (Siebold 1828), Kusa-Kaku (Siebold). Thunberg’s specimens in UPS. Lectotype (designated here): Japonia. Hb. Itoo Keiske ex Owari (L0420368) [Category 2, Hydrangea L. (see Ohba and Akiyama 2013a, 3] (Fig. 119). 2013b, 2014). Original material: Japonia. Communicavit 187) Hydrangea acuminata Siebold & Dr. Bürger, Hb. v Siebold (L0043362) [Category Zucc., Fl. Jap. 1: 110, t. 56, 57 I (1839). 2]. Legit in Japonia et Communicavit Siebold Present treatment: Hydrangea serrata 1839, Herbarium Zuccarinii (M0120979) (Thunb.) Ser. in DC., Prodr. 4: 15 (1830) (var. [Category 2]. Legit in Japonia & Communicavit serrata). de Siebold (M0120981) [Category 2b]. In Japanese name: Yama-ajisai. Japonia legit Bürger. Communicavit d. Siebold Lectotype (Ohba and Akiyama 2013b): (M0120980) [Category 2b]. Japonia. Herb. Siebold (L0043373). Hydrangea alternifolia was originally Original material: Legit in Japonia et described by Siebold in 1828 and transferred to Communicavit d. Siebold, Herbarium Zuccarinii Cardiandra by Siebold and Zuccarini in 1839. (M0153946) [Category 2]. Legit Dr Siebold The specimen in M annotated “Cardiandra in Japonia. Communicavit Prof. Zuccarini alternifolia S & Z” by Zuccarini, M0120979, is anno 1843, Herbarium Zuccarinii (M0153945) considered to have been given by Siebold and is [Category 2b]. In Japonia legit et Communicavit part of the original material. Siebold, Herb. Zuccarinii (M0153948) [Category 2b]. 190) Hydrangea azisai Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 188) Hydrangea acuminata Siebold & 14(2): 689 (1828). Siebold & Zucc., Fl. Jap. 1: Zucc. var. buergeri Siebold & Zucc., Fl. Jap. 1: 104, t. 51 (1839). 111, t. 57 II (1839), as “Bürgeri.” Present treatment: Hydrangea macrophylla Present treatment: Hydrangea serrata (Thunb.) Ser. in DC., Prodr. 4: 15 (1830) (f. (Thunb.) Ser. in DC., Prodr. 4: 15 (1830) (var. macrophylla). serrata). Japanese name: Ajisai; Azisai (Siebold Japanese name: Yama-ajisai. 1828). Lectotype (Ohba and Akiyama 2013b): Lectotype (Ohba and Akiyama 2014): 282 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

Japonia. Hb. von Siebold (L0043369) [Category Lectotype (Ohba and Akiyama 2014): 2, 5 (lower branch, Fl. Jap. 1: t. 51)]. Japonia. Herb. Siebold (L0043367) [Category Original material: Legit in Japonia et 5, right hand branch (Fl. Jap. 1: t. 92, upper) and Communicavit Siebold anno 1839, Herbarium left hand branch (Fl. Jap. 1: t. 92, lower)]. Zuccarinii (M0153951) [Category 2]. In Japonia legit Bürger. Communicavit d. Siebold, Herb. 194) Hydrangea cordifolia Siebold & Zucc., Zuccarinii (M0153952) [Category 2b]. Japonia. Fl. Jap. 1: 113, t. 59. II (1839). [Siebold s.n.] (MAK S0433) [Category 3]. Present treatment: Hydrangea petiolaris Japonia. [Siebold s.n.] (MAK S0837) [Category Siebold & Zucc. 2]. Japanese name: Tsuru-ajisai. The specimen in M annotated “Hydrangea Lectotype (Ohba and Akiyama 2014): Azisai Sieb. & Zuccar.” by Zuccarini, Japonia. Communicavit Bürger, Hb. von Siebold M0153951, is considered to have been given by (L0043365) [Category 2]. Siebold and is part of the original material. Original material: Legit in Japonia Bürger. Communicavit d. Siebold, Herbarium Zuccarinii 191) Hydrangea azisai Siebold var. foliis (M0153957) [Category 2]. In Japonia legit aureo-variegatis Siebold & Zucc., Fl. Jap. 1: Bürger. Communicavit d. Siebold, Herb. 105 (1839), nom. illeg. [phrase name] Zuccarinii (M0153958) [Category 2b]. In Present treatment: probably Hydrangea Japonia legit Bürger. Communicavit d. Siebold, macrophylla (Thunb) Ser. Herb. Zuccarinii (M0153959) [Category 2b]. In Original material: not found in L, M, or Japonia legit Bürger. Communicavit d. Siebold, MAK. Herb. Zuccarinii (M0153960) [Category 2b]. Japonia. [Siebold s.n.] (MAK S0850) [Category 192) Hydrangea belzonii Siebold & Zucc., 2]. Fl. Jap. 1: 109, t. 55 (1939). Present treatment: Hydrangea serrata 195) Hydrangea hortensia Siebold in Nova (Thunb.) Ser. var. yesoensis (Koidz.) H. Ohba in Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Iwatsuki et al. (eds.), Fl. Jap. 2b: 90 (2001). Cur. 14: 688 (1828), nom. superfl. Japanese name: Ezo-ajisai. Present treatment: probably Hydrangea Lectotype (Ohba and Akiyama 2013b): macrophylla (Thunb.) Ser. (var. macrophylla). Japonia. Hb. von Siebold (L0329008) [Category Original material: not found in L, M, or 2, 5 (upper right-hand branch, Fl. Jap. 1: t. 55)]. MAK. Original material: Hb. Siebold (L0043370) Hydrangea hortensia is regarded as a variant [Category 2]. Legit Bürger in Japonia. name for Hydrangea hortensis J. Sm. Communicavit d. Siebold, Herbarium Zuccarinii (M0153955) [Category 2]. In Japonia legit 196) Hydrangea involucrata Siebold in Bürger. Communicavit d. Siebold, Herb. Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Zuccarinii (M0153954) [Category 2b]. Japonia. Nat. Cur. 14: 691 (1828). Siebold & Zucc., Fl. [Siebold s.n.] (MAK S0851) [Category 2]. Jap. 1: 118, t. 63, 64 (1839). Present treatment: Hydrangea involucrata 193) Hydrangea bracteata Siebold & Zucc., Siebold Fl. Jap. 1: 176, t. 92 (1841). Japanese name: Tama-ajisai; Ginbaisoo and Present treatment: Hydrangea petiolaris Kinbaisoo (Siebold 1828) Siebold & Zucc. Lectotype (Ohba and Akiyama 2014): Japanese name: Tsuru-ajisai. Japonia. Hb. von Siebold (L0043366) [Category October 2014 The Journal of Japanese Botany Vol. 89 No. 5 283

2, 5 (upper branch, flowers with doubled petals, 199) Hydrangea japonica Siebold var. Fl. Jap. 1: t. 64, left hand branch)]. konkaku Siebold in Nova Acta Phys.-Med. Original material: Japonia. Hb. von Siebold, Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 689 in July to Aug. (L0101479) [Category 2, 5 (1828). Siebold & Zucc, Fl. Jap. 1: 106 (1839). (lower branch, flowers with single petals, Fl. Present treatment: Hydrangea serrata Ser. Jap. 1: t. 64, right hand branch)]. Japonia. Hb. var. japonica (Siebold) H. Ohba & S. Akiyama von Siebold (L0103032) [Category 2, 5 (Fl. Jap. in Bull. Natl. Sci. Nat. Sci, B, 39: 179 (2013). 1: t. 64, left hand branch)]. Legit in Japonia et Japanese name: Konkaku (Siebold 1828). Communicavit Siebold anno 1839, Herbarium Lectotype (Ohba and Akiyama 2013b): Zuccarinii (M0153961) [Category 2]. Japonia. Hb. von Siebold (L0103054). The specimen in M annotated “Hydrangea involucrata S. & Z.” by Zuccarini, M0153961, 200) Hydrangea otaksa Siebold & Zucc., Fl. is considered to have been given by Siebold and Jap. 1: 105, t. 52 (1839). is part of the original material. Present treatment: Hydrangea macrophylla (Thunb.) Ser. ‘Otaksa.’ 197) Hydrangea japonica Siebold in Nova Japanese name: Ajisai [Otaksa-ajisai as Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. the horticulture name]; Otaksa (Siebold and Cur. 14(2): 689 (1828). Siebold & Zucc., Fl. Jap. Zuccarini 1839). 1: 106, t. 53 (1839). Lectotype (Ohba and Akiyama 2014): Present treatment: Hydrangea serrata Japonia. Hb. Siebold (L0113146) [Category 2]. (Thunb.) Ser. var. japonica (Siebold) H. Ohba Original material: Japonia. Hb. Siebold, in & S. Akiyama in Bull. Natl. Sci. Nat. Sci, B, 39: July (L0328086) [Category 2]. Legit in Japonia 179 (2013). Bürger. Communicavit d. Siebold, Herbarium Japanese name: Beni-gaku; Kakusoo Zuccarinii (M0153965) [Category 2]. In Japonia (Siebold 1829). legit Bürger. Communicavit d. Siebold, Herb. Lectotype (Ohba and Akiyama 2013a): Zuccarinii (M0153966) [Category 2b]. Japonia. Hb. v Siebold, in July (L0043374) [Category 2]. 201) Hydrangea paniculata Siebold in Nova Original material: Japonia. Hb. Siebold, in Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. June (L0329010) [Category 2]. Cur. 14(2): 691 (1828). Siebold & Zucc., Fl. Jap. The original specimen, L0329010, is also the 1: 115, t. 61 (1839). lectotype of H. japonica var. benkaku Siebold. Present treatment: Hydrangea paniculata Siebold. 198) Hydrangea japonica Siebold var. Japanese name: Nori-utsugi, Nori-no-ki. benkaku Siebold in Nova Acta Phys.-Med. Lectotype (Ohba and Akiyama 2014): Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 689 Japonia. Hb. von Siebold (L0043375) [Category (1828), nom. superfl. Siebold & Zucc., Fl. Jap. 2]. 1: 106 (1839). Original material: Legit Bürger in Japonia. Present treatment: Hydrangea serrata Communicavit d. Siebold, Herbarium Zuccarinii (Thunb.) Ser. var. japonica (Siebold) H. Ohba & (M0153968) [Category 2]. In Japonia legit S. Akiyama. Bürger. Communicavit d. Siebold, Herb. Japanese name: Benkaku (Siebold 1828). Zuccarinii (M0153969) [Category 2b]. Japonia. Lectotype (Ohba and Akiyama 2013b): [Siebold s.n.] (MAK S0846) [Category 2]. Japonia. Hb. Siebold, in June (L0329010) The specimen in M annotated “Hydrangea [Category 2]. paniculata S. & Z.” by Zuccarini, M0153968, is 284 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 considered to have been given by Siebold and is 1: 11, t. 58 (1839). part of the original material. Present treatment: Hydrangea serrata (Thunb.) Ser. (var. serrata). 202) Hydrangea petiolaris Siebold & Zucc., Japanese name: Yama-ajisai; ‘Ama-tsja’ in Fl. Jap. 1: 108, t. 54 (1835). sensu Siebold (1830). Present treatment: Hydrangea petiolaris Type: Type of Hydrangea serrata Ser. Siebold & Zucc. Hydrangea thunbergii Siebold is a Japanese name: Tsuru-ajisai; Jama demari superfluous name for H. serrata (Thunb.) Ser., (Siebold and Zuccarini 1835). but H. thunbergii [sensu Siebold, excl. type] is Lectotype (Ohba and Akiyama 2014): the plant called Amatya (Amats'ja in Siebold, Japonia. Hb. von Siebold, in July (L0043364) 1828) in Japan, i.e., H. serrata var. thunbergii [Category 2, 3]. H. Ohba & S. Akiyama (Ohba and Akiyama 2013b). A specimen, “Japonia. Cultivated in 203) Hydrangea sitsitan Siebold in Nova Hortus Botanicus, Hb. von Siebold, in July,” Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. (L0043372), is an authentic specimen of H. Cur. 14(2): 692 (1828). thunbergii Siebold [sensu Siebold] based on our Present treatment: Hydrangea serrata Ser. category 2 criteria. Another specimen, “Japonia. var. serrata f. prolifera (Regel) Nakai, Siebold [Bürger s.n.],” (L0103047, branch in upper left), Hydrang. Kaisetsu: 14 (1938). is also an authentic specimen of H. thunbergii Japanese name: Shichidanka. Siebold [sensu Siebold] based on our category 2 Lectotype (Ohba and Akiyama 2013b): criteria. Japonia. Hb. von Siebold (L0043368) [Category 2]. Mitellopsis Meisn. Original material: Japonia. Hb. von Siebold 206) Mitellopsis japonica Siebold & Zucc. (L0170300) [Category 2, 3]. Japonia. [Siebold in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. s.n.] (MAK S0819) [Category 3]. Wiss. 4(2): 190 (1845). nom. nud. The lectotype specimen, L0043368, is also Present treatment: Mitella japonica Maxim. the lectotype specimen of H. stellata Siebold & in Bull. Acad. Imp. Sci. St. Pétersb. 11: 431 Zucc. (1867). Japanese name: Ô-charumeru-sô. 204) Hydrangea stellata Siebold & Zucc., Original material: Japonia. In montium locis Fl. Jap. 1: 112, t. 59. I (1839), nom. superfl. for humidis. Siebold s.n. (L0176011) [Category 3]. H. sitsitan Siebold. The Japanese name of Mitellopsis japonica, Present treatment: Hydrangea serrata Ser. Charumeru-sô, was not appeared in the var. serrata f. prolifera (Regel) Nakai, Siebold protologue but appears on the original label of Hydrang. Kaisetsu: 14 (1938). Siebold on sheet L0176011. Although there is no Japanese name: Shichidanka. annotation by Siebold or Zuccarini, we regard Lectotype (Ohba and Akiyama 2013b): this specimen to be original material. Japonia. Hb. von Siebold (L0043368) [Category 2] (Fig. 133). Parnassia L. 207) Parnassia mucronata Siebold & Zucc. 205) Hydrangea thunbergii Siebold in Nova in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Wiss. 4(2): 169 (1845). Cur. 14(2): 690 (1828), nom. superfl. for H. Present treatment: Parnassia palustris L. var. serrata (Thunb.) Ser. Siebold & Zucc., Fl. Jap. multiseta Ledeb., Fl. Ross. 1: 263 (1842). October 2014 The Journal of Japanese Botany Vol. 89 No. 5 285

Japanese name: Umebachi-sô. Mus. Bot. Lugd.-Bat. 3: 99 (1867), nom. superfl. Lectotype (designated here): Legit in Present treatment: Philadelphus satsumi Japonia & Communicavit de Siebold anno 1842, Siebold ex Lindl. & Paxt. Herbarium Zuccarinii (M0120972) (Fig. 120). Japanese name: Baika-utsugi. Original material: Legit Dr Siebold in Lectotype (Ohba et al. 2005): Japonia. Japonia. Communicavit Prof. Zuccarini anno Siebold s.n. (L0329021). 1843, Herbarium Zuccarinii (M0120973) The original material of P. satsumanus [Category 2b]. Siebold, Keiske s.n. (L0170309), is the The lectotype specimen, M0120972, has only specimen with Siebold’s annotation no annotation by Siebold and/or Zuccarini, “Philadelphus satsumanus S” but it was not however Zuccarini retained the name Parnassia cited by Miquel (1867). punctulata S. et Z. on the label. The epithet Philadelphus satsumanus Siebold was “mucronata” apparently derives from the reported by Siebold and Co. (1856) and description “... differt ... foliis omnibus validly published by Miquel in 1867, but it mucronato-acutis ...” or “[cauline leaves] sessili is a superfluous name for P. satsumi Siebold e basi cordata suborbiculari mucronato.” ex Lindl. & Paxt. Miquel cited the name, “P. The epithet punctulata appears to be related satsumi Sieb. in hort.—Paxt. Fl. Gard. II. n. 375 to the “ferrygino-punctatis” feature of the [p. 102–103] icon. xylogr.” in synonymy, and leaves. It might be thought that mucronata and four specimens collected by Siebold (L0329021, punctulata are candidates for the epithet. this is the lectotype of P. satsumanus Siebold Siebold s.n. in Leiden (L0170304) may be ex Miq.), Keiske (L0329023), Mohnike original material. Three plants on the right side (L0170308) and Oldham 258 (L 908, 24- of the sheet accompany Siebold’s annotation 200). These four specimens were used for the “Parnassia palustris an nova species” and description of Philadelphus satsumanus by “Parnassia Japonica, Sieb;” and a plant of Miquel. the left side (no. 134) accompanies a sketch Hu (1955) distinguished P. satsumi Siebold of a gland showing a mucronate (or bifurcate) ex Lindl. & Paxt. and P. satsumanus Siebold apex. The epithet mucronata might possibly be ex Miq. by the pubescence of the leaves, i.e. derived from this characteristic of the gland. uniformly pubescent in P. satsumanus and Siebold s.n. in Leiden (L0170302), with glabrous, except pubescent primary nerves and his annotation “Parnassia palustris. Nov. lusus axils on lower surface, in P. satsumi. Miquel’s Higo.” is also attributed to this species, but it is original material (syntypes) used for the not certain that it is original material. description of P. satsumanus, however, have nearly glabrous leaves except for the veins and Philadelphus L. the axils. 208) Philadelphus satsumanus Siebold, Catalogue: 5 (1856), nom. nud. 209) Philadelphus satsumi Siebold [in Present treatment: Philadelphus satsumi sched., nom. nud.] ex Lindl. & Paxt. in Paxton’s Siebold ex Lindl. & Paxt. Fl. Gard. 2: 102, fig. 188 (1852). Japanese name: Baika-utsugi. Present treatment: Philadelphus satsumi Original material: Japonia. Keiske s.n. Siebold ex Lindl. & Paxt. (L0170309) [Category 2]. Japanese name: Baika-utsugi. Original material: not found in L, M, or 208b) Philadelphus satsumanus Siebold MAK. [Catalogue: 5 (1856), nom. nud.] ex Miq., Ann. The name Philadelphus satsumi Siebold was 286 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 published by Lindley and Paxton in 1852. Hu L0327094. The epithet of the provisional name, (1955) reported Siebold’s specimen in “Roy. Ribes Jabusansasi, is derived from the Japanese Hort. Soc. Gard., Wisley” as being the type. name of the plant. We cannot choose the Leiden specimens as original material because they have Platycrater Siebold & Zucc., Fl. Jap. 1: 62 (1837 no indication of Siebold’s and/or Zuccarini’s or 1838); in Abh. Math.-Phys. Cl. Königl. Bayer. description in 1845. Akad. Wiss. 4(2): 192 (1845). Type: Platycrater arguta Siebold & Zucc. Saxifraga L. Present treatment: Platycrater Siebold & 212) Saxifraga cortusifolia Siebold & Zucc. Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 190 (1845), as “cortusaefolia.” 210) Platycrater arguta Siebold & Zucc., Fl. Present treatment: Saxifraga cortusifolia Jap. 1: 64, t. 27 (1835); in Abh. Math.-Phys. Cl. Siebold & Zucc. Königl. Bayer. Akad. Wiss. 4(2): 192 (1845). Japanese name: Jinji-so. Present treatment: Platycrater arguta Siebold Lectotype (designated here): Legit in & Zucc. Japonia et Communicavit d. Siebold, Herbarium Japanese name: Baika-amacha; Bai kwa ama Zuccarinii (M0120982) [Category 2] (Fig. 123). tsja (Siebold and Zuccarini 1835). Lectotype (designated here): Japonia. Schizophragma Siebold & Zucc., Fl. Jap. 1: 58 Herbar. de Siebold, in 1829 (L0104524) (1837 or 1838); in Abh. Math.-Phys. Cl. Königl. [Category 2] (Fig. 121). Bayer. Akad. Wiss. 4(2): 192 (1845). Original material: Japonia. Siebold s.n. Type: Schizophragma hydrangeoides (L0104521) [Category 2]. Legit in Japonia Siebold & Zucc. & Communicavit de Siebold anno 1842, Present treatment: Schizophragma Siebold & Herbarium Zuccarinii (M0120977) [Category Zucc. 2]. In Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M012978) [Category 213) Schizophragma hydrangeoides Siebold 2b]. & Zucc., Fl. Jap. 1: 60, t. 26 (1837 or 1838), t. 100 I (1839); in Abh. Math.-Phys. Cl. Königl. Ribes L. Bayer. Akad. Wiss. 4(2): 192 (1845). 211) Ribes fasciculatum Siebold & Zucc. Present treatment: Schizophragma in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. hydrangeoides Siebold & Zucc. Wiss. 4(2): 189 (1845). Japanese name: Iwagarami; Tsuru-demari Present treatment: Ribes fasciculatum (Siebold and Zuccarini 1837 or 1838). Siebold & Zucc. Lectotype (designated here): Japonia. Japanese name: Yabu-sannzashi. Herbar. de Siebold coll. 1829 (L0104501) (Fig. Lectotype (designated here): In Japonia 124). legit Bürger. Communicavit d. Siebold, Herb. Original material: Japonia. Hb. Siebold Zuccarinii (M0153974) [Category 2b] (Fig. (L0104500) [Category 2]. Japonia. Nagasaki. 122). Siebold s.n. (L0104499) [Category 3]. Original material: In Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii Rosaceae (M0153975) [Category 2b]. Amygdalus L. There are two specimens annotated “Ribes 214) Amygdalus persica L. var. stellata Jabusansasi” by Siebold in L, L0327091 and Siebold & de Vriese in Ann. Hort. Bot. (Fl. October 2014 The Journal of Japanese Botany Vol. 89 No. 5 287

Jardin) 2: 65, t. [6] (1859). (L0329142, this is also original material of Present treatment: Amygdalus persica L. f. Prunus mume) [Category 2]. Japonia. [Bürger stellata (Siebold & de Vriese) H. Ohba & S. s.n.] (L0328748) [Category 2]. Akiyama, stat. nov. Basionym: Amygdalus Armeniaca mume Siebold (1830) is an persica L. var. stellata Siebold & de Vriese invalid name. Koch (1853) made a new in Ann. Hort. Bot. (Fl. Jardin) 2: 65 (1859) ≡ combination, Armeniaca mume (Siebold & Prunus persica (L.) Batsch. var. β. vulgaris Zucc.) K. Koch based on Prunus mume Siebold (DC.) Maxim. f. stellata (Siebold & de Vriese) & Zucc. Makino in Bot. Mag. (Tokyo) 22: 119 (1908). Japanese name: Genpei-kiku-momo (nov.); 216) Armeniaca mume Siebold var. a. Kiku-momo, Genji-guruma (Makino 1908). floribus simplicibus plenisque niveis Siebold, Makino (1908) reported “Kiku-momo” and Syn. Pl. Oeconom.: 69 (1830), nom. illegit. “Genji-guruma” as Japanese names, but this [phrase name] plant has both rose and white flowers. A new Present treatment: a cultivar of Armeniaca Japanese name, Genpei-kiku-momo [meaning mume (Siebold & Zucc.) K. Koch. plant with narrow petals and with both rose and Original material: not found in L, M, or white flowers], is presented here. MAK. Lectotype (designated here): Illustration t. [6] in Ann. Hort. Bot. (Fl. Jardin) 2: t. [6, 217) Armeniaca mume Siebold var. b. between pages 64 and 65] (1859). floribus simplicibus plenisque roseis Siebold, We designate the illustration in the protologue Syn. Pl. Oeconom.: 69 (1830), nom. illegit. as the lectotype. The plant illustrated in the table [phrase name] has double flowers with both pink and white Present treatment: a cultivar of Armeniaca narrow petals. In L, M and MAK there are some mume (Siebold & Zucc.) K. Koch. specimens of the cultivar of Amygdalus persica, Original material: not found in L, M, or but none of them represents var. stellata Siebold MAK. & de Vries. One of them in L, L0102983, is a specimen annotated “Amygdalus persica 218) Armeniaca mume (Siebold & Zucc.) erecta, Genpei too” by Siebold. “Genpei too” K. Koch var. praecocissima rubiflora Siebold means having both pink and white flowers. That & de Vriese in de Vriese, Tuinb. Fl. 1: 1, t. 1 specimen resembles var. stellata with bicolored (1855), nom. illegit. [phrase name] double flowers, but the petals are wide. In M Present treatment: a cultivar of Armeniaca there is also a specimen, M0154003, named mume (Siebold & Zucc.) K. Koch. Persica vulgaris var. flore pleno albo with Original material: Illustration in de Vriese, double petaled flowers, but appears to be only Tuinb. Fl. 1: 1, t. 1 (1855). white. According to the original description and the illustration, praecocissima rubiflora appears to Armeniaca L. be an early flowering double flowered cultivar 215) Armeniaca mume Siebold, Syn. Pl. with rose colored petals like ‘Kankôbai.’ Oeconom.: 69 (1830), nom. nud. Present treatment: Armeniaca mume (Siebold Aronia Mitch. & Zucc.) K. Koch, Hort. Dendrol.: 140 (1853). 219) Aronia asiatica Siebold & Zucc., Fl. Original material: Japonia. Herbar. de Siebold Jap. 1: 87, t. 42 (1839); in Abh. Math.-Phys. Cl. coll. 1829 [and 1827] [flowers: Mar.1829 and Königl. Bayer. Akad. Wiss. 4(2): 131 (1845). leaves: 亥 六 月 June 1827 by lunar calendar] Present treatment: Amelanchier asiatica 288 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

Endl. in Walp., Rep. 2: 53 (1843). M0153991 collected by Bürger, and which was Japanese name: Zaifuri-boku; Zaifuri, cited by Aldasoro et al. as an isolectotype, as the Zaifuribok (Siebold and Zuccarini 1839). lectotype. Lectotype (designated here): Herb. de Siebold in 1829 (L0176017) [Category 2] (Fig. 222) Crataegus cuneata Siebold & Zucc. 125). in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Original material: Japonia. [April. Jedo]. Wiss. 4(2): 130 (1845). Siebold s.n. (L0170335) [Category 3]. Present treatment: Crataegus cuneata Siebold & Zucc. Ceraseidos Siebold & Zucc. in Abh. Math.- Japanese name: Sanzashi. Phys. Cl. Königl. Bayer. Akad. Wiss. 3(3): 743 Lectotype (designated here): Legit Bürger in (1843); 4(2): 122 (1845). Japonia. Communicavit de Siebold, Herbarium Present treatment: Cerasus Mill. Zuccarinii (M0153734) [Category 2] (Fig. 128). Type: Ceraseidos apetala Siebold & Zucc. Original material: In Japonia legt. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Bürger comcvt. de Siebold, Herb. Zuccarinii Wiss. 3(3): 744, t. V 2 (1843); 4(2): 122 (1845). (M0153733) [Category 2b]. The specimens in L, Siebold s.n. (L0170337 220) Ceraseidos apetala Siebold & Zucc. and L0170338), with accompanying slips with in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. the Japanese name “Crataegus sansasi, Jap. and Wiss. 3(3): 744, t. V 2 (1843); 4(2): 122 (1845). サン ザシ ” by Siebold, are also representative. Present treatment: Cerasus apetala (Siebold The specimen, Siebold (L0170336), with the & Zucc.) Ohle ex H. Ohba in J. Jpn. Bot. 67: Japanese name “Crataegus oosansasi, J[ap.]” by 277 (1992) (var. apetala). Siebold, also has no evidence of being original Japanese name: Miyama-chôji-zakura. material. Lectotype (designated here): Japonia. Siebold s.n. (L0176026) [Category 2] (Fig. 126). Kerria DC. 223) Kerria japonica (L.) DC. var. a Crataegus L. Floribus simplicibus Siebold, Oeconom.: 69 221) Crataegus alnifolia Siebold & Zucc. (1830), nom. illegit. [phrase name] in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Present treatment: Kerria japonica (L.) DC. Wiss. 4(2): 130 (1845). in Trans. Linn. Soc. London 12: 157 (1817) (f. Present treatment: Sorbus alnifolia (Siebold japonica). & Zucc.) K. Koch in Ann. Mus. Bot. Lugd.-Bat. Japanese name: Hitoe-yamabuki. 1: 249 (1864). Original material: Japonia. [Siebold s.n.] Japanese name: Azuki-nashi. (MAK S0261a) [Category 2]. Lectotype (designated here): Japonia. Legit Bürger in Japonia, communicavit de Siebold, 224) Kerria japonica (L.) DC. var. b. Herbarium Zuccarinii (M0153991) [Category 2] Floribus plenis Siebold, Oeconom.: 69 (1830), (Fig. 127). nom. illegit. [phrase name] Original material: Herb. Bürger (L0329395) Present treatment: Kerria japonica (L.) DC. [Category 2]. f. plena C. K. Schneid., Ill. Hand. Laubholzk. 1: Aldasoro et al. (2004) selected Bürger s.n. 501 (1905). (L0245120) as the lectotype, but the specimen Japanese name: Yae-yamabuki. has no annotation or other information to Original material: not found in L, M, or indicate it to be original material. We designate MAK. October 2014 The Journal of Japanese Botany Vol. 89 No. 5 289

225) Kerria japonica (L.) DC. var. c corresponds well with this variety but lacks Floribus simplicibus foliis argentatis Siebold, Siebold and/or Zuccarini’s handwriting. Oeconom.: 69 (1830), nom. illegit. [phrase The illustration of Table 99 II represents name] well the variegated form of Kerria japonica, Present treatment: Kerria japonica (L.) DC. although there is no explanation by Siebold and ‘Variegata.’ Zuccarini. Yamaguchi et al. (2003) reported that Japanese name: Fuiri-yamabuki. the illustration of Table 98 of Flora Japonica Original material: not found in L, M, or was based on this specimen. MAK. 229) Kerria tetrapetala Siebold, Syn. Pl. 226) Kerria japonica (L.) DC. var. α. sejo- Oeconom.: 69 (1830), nom. nud. jama-buki Siebold & Zucc., Fl. Jap. 1: 183 Present treatment: Rhodotypos scandens (1841), as “Sejô jama buki.” (Thunb.) Makino in Bot. Mag. (Tokyo) 27: 126 Present treatment: Kerria japonica (L.) DC. (1913). f. plena C. K. Schneid., Ill. Hand. Laubholzk. 1: Japanese name: Shiro-yamabuki; 501 (1905). Sirojamabuki (Siebold 1830). Japanese name: Yae-yamabuki; Sejô jama Original material: Herb. von Siebold buki (Siebold and Zuccarini 1841). (L0329269) [Category 2]. Japonia. Keiske Lectotype (designated here): Hb. von [Sukerok] s.n. (L0175824) [Category 3]. Siebold (L0328743) [Category 5 (Fl. Jap. 1: t. 98 The specimen, L0329269, is also the III)] (Fig. 129). lectotype of Rhodotypos kerrioides Siebold & Zucc. (see p. 293). 227) Kerria japonica (L.) DC. var. β. hitoje- jamabuki Siebold & Zucc., Fl. Jap. 1: 183 Malus L. (1841), as “Hitoje jamabuki.” 230) Malus floribunda Siebold, Cat. Rais. 5 Present treatment: Kerria japonica (L.) DC. (1856), nom. nud. in Trans. Linn. Soc. London 12: 157 (1817) (f. Present treatment: probably Malus halliana japonica). Koehne, Gatt. Pomac.: 27 (1890). Japanese name: Hitoe-yamabuki; Hitoje Japanese name: probably Kaidô. jamabuki. (Siebold and Zuccarini 1841). Original material: not found in L, M, or Lectotype (designated here): Illustration in MAK. Siebold & Zucc., Fl. Jap. 1: t. 98 I (1841). 231) Malus [Jap.] Ringo Siebold, Cat. Rais. 228) Kerria japonica (L.) DC. var. γ. fuiri- 5 (1856), nom. nud. jamabuki Siebold & Zucc., Fl. Jap. 1: 183 Present treatment: Malus domestica Borkh., (1841), as “Fuiri jamabuki.” Theor. Prakt. Handb. Forstbot. 2: 1272 (1803). Present treatment: Kerria japonica (L.) DC. Japanese name: Ringo. ‘Variagata.’ Original material: Japonia. Siebold Japanese name: Fuiri-yamabuki; Fuiri (L0329359) [Category 3]. jamabuki. (Siebold and Zuccarini 1841). Lectotype (designated here): Illustration in 232) Malus spectabilis Desf. var. [Jap.] Siebold & Zucc., Fl. Jap. 1: t. 99 II (1841). Kaido Siebold, Cat. Rais. 5 (1856), nom. nud. The specimen, Herb. Siebold (L0328745), Present treatment: Malus halliana Koehne, with the note “Kerria japonica DC. Sieb. & Gatt. Pomac.: 27 (1890). Zucc., Fl. jap. t. 98. variet. foliis maculatis,” Japanese name: Kaidô. 290 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

Original material: Japonia. Siebold s.n. 235) Persica stellata Siebold, Cat. Rais.: 36 (L0329350) [Category 3]. Japonia. Siebold s.n. (1863), nom. nud. (L0329358) [Category 3]. Present treatment: Amygdalus persica L. f. stellata (Siebold & de Vriese) H. Ohba & S. 233) Malus toringo Siebold, Cat. Rais.: 5 Akiyama. (1856). Japanese name: Genpei-kiku-momo. Present treatment: Malus toringo (Siebold) Original material: not found in L, M, or Siebold, Cat. Rais.: 5 (1856). De Vriese, Tuinb.- MAK. Fl. 3: 368, t. [17, between pages 368 and 369], excl. fig. fl. (1856). 236) Persica vulgaris Mill. var. versicolor Japanese name: Zumi; Ko-ringo; Kimi-zumi Siebold, Cat. Rais.: 36 (1863). (Siebold and de Vriese 1856). Present treatment: Amygdalus persica L. f. Type: Type of Sorbus toringo Siebold in versicolor (Siebold) H. Ohba & S. Akiyama, Jaarb. Kon. Ned. Maatsch. Aanm. Tuinb. 1848: comb. et stat. nov. Basionym: Persica vulgaris 47 (1848). Mill var. versicolor Siebold, Cat. Rais.: 36 When Siebold published Malus toringo (1863) ≡ Prunus persica (L.) Batsch f. versicolor Siebold (Cat. Rais.: 5, 1856) validly, he adopted (Siebold) Koidz. in Bot. Mag. (Tokyo) 44: 94 the epithet of Sorbus toringo Siebold (1848) (1930). without referring to that name. According to Japanese name: Genpei-momo. ICN Art. 41.4 it is treated as a new combination, Original material: not found in L, M, or Malus toringo (Siebold) Siebold, not as the MAK. name of a new species. In the same year de Vriese wrote Malus toringo Siebold (Tuinb. Prunus L. Fl. 3: 368, t. [17, between pages 368 and 369], 237) Prunus domestica L. var. [a] zumomo 1856) with referring to Sorbus toringo Siebold Siebold, Syn. Pl. Oeconom.: 68 (1830). and Siebold’s Catalogue Raisonné. Present treatment: probably a cultivar of Prunus domestica L. Persica Mill. Japanese name: Sumomo. 234) Persica pendula Siebold, Cat. Rais.: 36 Original material: not found in L, M, or (1863). MAK. Present treatment: Amygdalus persica L. f. pendula (Siebold) H. Ohba & S. Akiyama, stat. 238) Prunus domestica L. var. [b] nov. Basionym: Persica pendula Siebold, Cat. awozumomo Siebold, Syn. Pl. Oeconom.: 68 Rais.: 36 (1863) ≡ Prunus persica (L.) Batsch (1830). var. pendula (Siebold) Koidz. in Bot. Mag. Present treatment: probably a cultivar of (Tokyo) 44: 95 (1930). Prunus domestica L. Japanese name: Beni-shidare-momo. Japanese name: Ao-sumomo. Lectotype (designated here): Japonia. Original material: not found in L, M, or Siebold s.n. (L0102984) [Category 5] (Fig. 130). MAK. The lectotype specimen is not annotated as ‘Persica pendula’ but has Siebold’s annotation 239) Prunus domestica L. var. [c] “Amygdalus persica pendula” and the Japanese sirozumomo Siebold, Syn. Pl. Oeconom.: 68 name “Sdale momo” [it is now written ‘Sudare’ (1830). or ‘Shidare’ and means a pendulous form of Present treatment: probably a cultivar of Amygdalus persica in Japanese]. Prunus domestica L. October 2014 The Journal of Japanese Botany Vol. 89 No. 5 291

Japanese name: Shiro-sumomo. Plantae Oeconomicarum (1830, No. 368[sic]) Original material: not found in L, M, or as the source of the Siebold name. Number 368 MAK. of Siebold’s Synopsis Plantae Oeconomicarum is neither Cerasus nor Prunus but a species of 240) Prunus donarium Siebold, Syn. Pl. Gramineae, “CCXVII. GRAMINEA, Makomo, Oeconom.: 68 (1830), nom. nud. Japon.” Present treatment: Cerasus serrulata (Lindl.) G. Don in Loudon, Hort. Brit.: 480 (1839). 242) Prunus jamasakura Siebold, Syn. Pl. Japanese name: Shiogama(-sakura); Sakura Oeconom.: 68 (1830), nom. nud. (Siebold 1830). Present treatment: Cerasus jamasakura Original material: Japonia. Siebold s.n. (Siebold ex Koidz.) H. Ohba in J. Jap. Bot. 67: (L0834824) [Category 2, 3]. Japonia in templis. 278 (1992). Legit v. S. (MAK S0251, upper branch) Japanese name: Yamazakura. [Category 2]. Original material: Japonia. [Siebold s.n.] (L0834829) [Category 3]. Japonia. K[Keiske] 241) Prunus itosakura Siebold, Syn. Pl. s.n. (L0329119) [Category 2, 3]. Japonia. Legit Oeconom.: 68 (1830). v. S.[Siebold s.n.] (MAK S0247) [Category 2]. Present treatment: Cerasus itosakura Japonia. [Siebold s.n.] (MAK S0249) [Category (Siebold) H. Ohba & S. Akiyama, comb. nov. 2]. Basionym: Prunus itosakura Siebold, Syn. Pl. Prunus jamasakura, native to Japan, was Oeconom.: 68 (1830). validly published by Koidzumi (1911), but he Japanese name: Shidarezakura, Itozakura; did not designate a lectotype. The specimen, Itosakura (Siebold 1830). L0834829, consists of three branches with Lectotype (designated here): Japonia. Nagas. simple petaled flowers. The leaves on a post [aki]. Siebold s.n. (L0102980) [Category 2] (Fig. flowering branch are narrowly oblong-obovate 131). or elliptic with glabrous surfaces and margins Original material: Japonia. Siebold s.n. and single (partly double) serrations and have (L0102977) [Category 2]. Japonia. [Siebold s.n.] interpetiolar glands. These features characterize (L0102981) [Category 2]. Cerasus jamasakura. Ohba mistakenly treated Prunus itosakura Siebold as an illegitimate name (nomen nudum) 243) Prunus macrophylla Siebold & Zucc. and adopted Cerasus spachiana Lavallée ex in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. E. Otto (in Hamburger Garten-Blumenzeitung Wiss. 4(2): 122 (1845), non Poir. (1816). 35: 57, 1879) according to Sargent (1888) Present treatment: Laurocerasus zippeliana for it (Ohba 2001, 2012, Ohba et al. 2007). Miq., Fl. Ned. Ind., Eerste Bijv. (also as Fl. Ind. However, it is recognized as a legitimate name Batav.) 1(1): 367 (1855). with a short diagnosis. Siebold wrote on this Japanese name: Bakuchi-no-ki. species: “Cerasus ramulis pendulis florentibus Lectotype (designated here): Japonia. adornans idolorum sacra.” The “Cerasus ramulis Siebold s.n. (L0329088) [Category 2] (Fig. 132). pendulis” [Cerasus with pendulous branchlets], Original material: Legit Bürger in Japonia. is considered sufficient for a diagnosis. In Communicavit de Siebold, Herbarium Siebold’s time, this was the only flowering Zuccarinii (M0154001) [Category 2]. In Japonia cherry with pendulous branchlets. legt Bürger comct. de Siebold, Herb. Zuccarinii Maximowicz (1883) published Prunus (M0154002) [Category 2b]. Japonia. Siebold pendula Siebold ex Maxim. and cited Synopsis s.n. (L0329091) with Siebold’s annotation 292 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

“Laurocerasus japonica S[iebold]” and 246) Prunus mume Siebold & Zucc. var. Japanese name “ バクチノキ [Bakuchi-no-ki in pendula Siebold in Jaarb. Kon. Ned. Maatsch. Japanese].” Japonia. Siebold s.n. (L0176030) Tuinb. 1848: 47 (1848). with Siebold’s annotation “Laurocerasus Present treatment: Armeniaca mume (Siebold japonica Siebold.” & Zucc.) K. Koch f. pendula (Siebold) H. Ohba Two specimens, Siebold s.n. (L0329091 & S. Akiyama, comb. et stat. nov. Basionym: and L0176030), have only Siebold’s annotation Prunus mume Siebold & Zucc. var. pendula “Laurocerasus japonica S[iebold]” and also the Siebold in Jaarb. Kon. Ned. Maatsch. Tuinb. Japanese name “ バクチノキ [Bakuchi-no-ki in 1848: 47 (1848). Japanese]” on L0329091, and are considered to Japanese name: Shidare-ume. be derived from the same source as lectotype. Lectotype (designated here): Japonia. Siebold s.n. (MAK S0551, left) [Category 3] 244) Prunus mume Siebold & Zucc., Fl. (Fig.134). Jap. 1: 29, t. 11 (1836); in Abh. Math.-Phys. Cl. Original material: Japonia. s.n. (MAK Königl. Bayer. Akad. Wiss. 4(2): 122 (1845). S0551, right) [Category 3]. Japonia. Present treatment: Armeniaca mume (Siebold & Zucc.) K. Koch, Hort. Dendrol.: 140 (1853). 247) Prunus spinulosa Siebold & Zucc. Japanese name: Ume; Mume (Siebold and in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Zuccarini 1836). Wiss. 4(2): 122 (1845). Lectotype (designated here): Japonia. Present treatment: Laurocerasus spinulosa Siebold s.n. (L0329140) [Category 3, 5 (Fl. Jap. (Siebold & Zucc.) C. K. Schneid., Ill. Handb. t. 11 I)] (Fig. 133). Laubh. 1: 649, f. 354 o–p, f. 355 f (1906). Original material: Japonia. Legit v. Japanese name: Rin-boku. S.[Siebold] (L0329135, MAK S0259) [Category Lectotype (designated here): Legit in 2]. Legit Bürger in Japonia Communicavit de Japonia et Communicavit de Siebold, Herbarium Siebold, Herbarium Zuccarinii (M0153997) Zuccarinii (M0154004) [Category 2] (Fig. 135). [Category 2]. In Japonia legt. Bürger comct. Original material: In Japonia legt. v. Siebold, de Siebold, Herb. Zuccarinii (M0153998) Herb. Zuccarinii (M0154005) [Category 2b]. [Category 2b]. In Japonia legt. Bürger comct. de Siebold, Herb. Zuccarinii (M0153999) Pyrus L. [Category 2b]. Legit Dr Siebold in Japonia. 248) Pyrus gracilis Siebold & Zucc. in Abh. Communicavit Prof. Zuccarini, Herbarium Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. Zuccarinii (M0154000) [Category 2b]. Japonia. 4(2): 131 (1845). [Siebold s.n.] (MAK S0373) [Category 2]. Present treatment: Sorbus gracilis (Siebold & Japonia. Planta culta introducta in H. Sieb. Zucc.) K. Koch, Hort. Dendrolog.: 178 (1853). [Siebold s.n.] (MAK S0553) [Category 2]. Japanese name: Nankin-nanakamado. Lectotype (designated here): Legit. Bürger in 245) Prunus mume Siebold & Zucc. var. Japonia. Communicavit de Siebold (M0154368) floribus purpureis semiplenis adjecto ramulo [Category 2] (Fig. 136). foliifero Siebold & Zucc., Fl. Jap. 1: 30, t. 11 III Original material: Japonia. Siebold (1836), nom. invalid. [phrase name] (L0329393) [Category 2]. Present treatment: probably a cultivar of A specimen, Herb. Bürger (L0043355), Armeniaca mume (Siebold & Zucc.) K. Koch. has an accompanying type slip, Pyrus gracilis Original material: not found in L, M, or Siebold & Zucc., in Hara’s hand, but the MAK. specimen is not regarded as original material October 2014 The Journal of Japanese Botany Vol. 89 No. 5 293 because there is no evidence of the original 4(2): 125 (1845). description. Present treatment: Rhodotypos scandens (Thunb.) Makino in Bot. Mag. (Tokyo) 27: 126 Raphiolepis Lindl. (1913). 249) Raphiolepis japonica Siebold & Zucc., Japanese name: Shiro-yamabuki; Siro Fl. Jap. 1: 162, t. 85 (1841); in Abh. Math.-Phys. jamabuki. Cl. Königl. Bayer. Akad. Wiss. 4(2): 130 (1845). Lectotype (designated here): Herb. von Present treatment: Raphiolepis indica (L.) Siebold (L0329269) (Fig. 138). Lindl. var. umbellata (Thunb.) H. Ohashi Original material: Japonia. Siebold s.n. f. umbellata (Thunb.) Hatus. in J. Geobot. (L0329268) [Category 3]. Japonia. Siebold (Kanazawa) 25: 126 (1978). s.n. (L0175824) [Category 3]. Legit in Japonia Japanese name: Sharin-bai; Hama mokkok’ Bürger. Communicavit de Siebold anno 1842, (Siebold and Zuccarini 1841). Herbarium Zuccarinii (M0153995) [Category Lectotype (designated here): Japonia. Hb. 2]. Legit in Japonia & Comunicavit de Siebold von Siebold (L0176032) [Category 2, 3] (Fig. anno 1842, Herbarium Zuccarinii (M0153994) 137). [Category 2]. Original material: Japonia. Hb. von Siebold The lectotype is also the original material of (L0176031) [Category 3]. Legit Bürger in Kerria tetrapetala Siebold, nom. nud. Japonia. Communicavit de Siebold, Herbarium Zuccarinii (M0153736) [Category 2]. In Japonia Rosa L. legt. Bürger comct. v. Siebold, Herb. Zuccarinii 252) Rosa iwara Siebold in Cat. Raisonné (M0153735) [Category 2b]. des Pl. du Japon, 1856: 6 (1856), nom. nud. Present treatment: Rosa ×iwara Siebold 250) Raphiolepis mertensii Siebold & Zucc., ex Regel, Index Sem. Hort. Petrop. 1861: 53 Fl. Jap. 1: 164 (1841); in Abh. Math.-Phys. Cl. (1861). Königl. Bayer. Akad. Wiss. 4(2): 130 (1845). Japanese name: Ko-hamanasu. Present treatment: Raphiolepis indica (L.) Original material: Japonia. Legit Siebold, Lindl. var. umbellata (Thunb.) H. Ohashi Herb. Zuccarinii (M0154345) with Siebold’s f. umbellata (Thunb.) Hatus. in J. Geobot. annotation “Rosa Iwara Sieb.” [Category 2]. (Kanazawa) 25: 126 (1978). Rosa ×iwara is a natural hybrid between Japanese name: Maruba-sharin-bai. Rosa multiflora and R. rugosa, and in Japan was Original material: not found in L, M, or rarely cultivated in gardens as an ornamental MAK. during Siebold’s stay (Ohba and Akiyama 2013a). Rosa iwara Siebold was validly Rhodotypos Siebold & Zucc., Fl. Jap. 1: 185 published by Regel [Index Sem. Hort. Petrop. (1841); in Abh. Math.-Phys. Cl. Königl. Bayer. 1861: 53, 1861]. As Regel did not designate a Akad. Wiss. 4(2): 125 (1845). lectotype, we designate the following specimen Type: Rhodotypos kerrioides Siebold & as the lectotype. Zucc. Lectotype (designated here): Japonia. Present treatment: Rhodotypos Siebold & Legit Siebold, Herb. Zuccarinii (M0154345) Zucc. with Siebold’s annotation “Rosa Iwara Sieb.” and Crépin’s annotation “Rosa rugosa × R. 251) Rhodotypos kerrioides Siebold & multiflora” (Fig. 139) [Category 2]. Zucc., Fl. Jap. 1: 187, t. 99 (1841); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 253) Rosa polyantha Siebold & Zucc. in 294 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Herb. Zuccarinii (M0154013) [Category 2b]. Wiss. 4(2): 128 (1845). Present treatment: Rosa multiflora Thunb. in 256) Rubus tokkura Japon ex Siebold, Syn. Murray, Syst. Veg. ed. 14: 474 (1784). Pl. Oeconom.: 65 (1830), nom. nud. Japanese name: No-ibara. Present treatment: Rubus coreanus Miq., Lectotype (Ohba and Akiyama 2013a): Ann. Mus. Bot. Lugd.-Bat. 3: 34 (1867). Japonia. Legit Bürger, communicavit Siebold, Japanese name: Tokkuri-ichigo; Tokkuri- Herbarium Zuccarinii (M0154318). itsigo (Siebold 1830). Original material: not found in L, M, or Rubus L. MAK. 254) Rubus ribifolius Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Sieversia Willd. Wiss. 4(2): 127 (1845). 257) Sieversia dryadoides Siebold & Zucc. Present treatment: Rubus ×ribifolius Siebold in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. & Zucc. Wiss. 4(2): 125 (1845). Japanese name: Hama-kiichigo. Present treatment: Geum pentapetalum (L.) Lectotype (designated here): Legit in Makino in Bot. Mag. (Tokyo) 24: 32 (1910). Japonia & Communicavit Zuccarini, Herbarium Japanese name: Chinguruma. Zuccarinii (M0120864) [Category 2] (Fig. 140). Lectotype (designated here): Japonia. [Sukerok s.n.] (L0175827) [Category 2] (Fig. 255) Rubus thunbergii Siebold & Zucc. 141). in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Original material: Legit v. Siebold in Wiss. 4(2): 126 (1845). Japonia. Communicavit de Siebold, Herbarium Present treatment: Rubus hirsutus Thunb., Zuccarinii (M 0154382) [Category 2]. Dissert. Bot. Med. Rubo.: 7, 10 (1813). Japanese name: Kusa-ichigo. Sorbus L. Lectotype (designated here): Japan. 258) Sorbus japonica Siebold, Syn. Pl. (Thunberg s.n., UPS-Herb. Thunberg no. Oeconom.: 67 (1830), nom. nud. 12266). Present treatment: Aria japonica Decne. in Siebold and Zuccarini cited Rubus idaeus Nouv. Arch. Mus. Hist. Nat. 10: 166 (1874). [auct. non L.: Thunb., Fl. Jap.: 216, 1784] Japanese name: Urajiro-no-ki; Urasironoki as a synonym of Rubus thunbergii. Seven (Siebold 1830). specimens in Thunberg’s collection (UPS, Herb. Original material: Japonia. Siebold s.n. Thunberg nos. 12261–12267) are identified (L0366764) [Category 2, 3]. as Rubus idaeus. We have selected no. 12266 as the lectotype which appears to be the most 259) Sorbus toringo Siebold in Jaarb. Kon. representative among them. Ned. Maatsch. Aanm. Tuinb. 1848: 47 (1848). Specimens used for the description of Rubus Present treatment: Malus toringo (Siebold) thunbergii by Siebold and Zuccarini are as Siebold, Cat. Rais.: 5 (1856). De Vriese, Tuinb.- follows: Fl. 3: 368, t. [17, between pages 368 and 369] Legit in Japonia & Communicavit de Siebold excl. fig. fl. (1856). anno 1842, Herbarium Zuccarinii (M0154011) Japanese name: Zumi; Ko-ringo, Kimi-zumi [Category 2]. In Japonia legit Bürger comcvt. v. (Siebold & de Vriese 1856) Siebold, Herb. Zuccarinii (M0154012) [Category Lectotype (designated here): Japonia. 2b]. In Japonia legit Bürger comcvt. v. Siebold, Siebold s.n. (L0176022) [Category 2] (Fig. 142). October 2014 The Journal of Japanese Botany Vol. 89 No. 5 295

Siebold effectively published Sorbus toringo 262) Spiraea thunbergii Siebold, Syn. Pl. with the description “Arbuste nain à feuilles Oeconom.: 65 (1830), non Blume (1827). florales elliptiques et raméales trilobées. Couvert Present treatment: Spiraea thunbergii Siebold au premier printemps de fleurs blanches et ex Blume, Bijdr. 17: 1115 (1827). rosées moins grandes que celles de notre Pyrus Japanese name: Yuki-yanagi; Jukiyanagi spectabilis.” (Siebold 1830). Veldkamp selected the specimens L0176022 Lectotype (designated here): Japan. and L0329344 as the neotype of Sorbus toringo Thunberg s.n. (L0144310, van Royen Siebold. L0329344 (Siebold s.n.) has neither Herbarium) (Fig. 144). annotation nor notes by Siebold himself, but This is a new name for the Spiraea reported L0176022 has Siebold’s annotation “Sorbus by Thunberg as S. crenata L. in his Flora tôringo” and is regarded as original material. Japonica (1784, p. 210). Siebold cited Spiraea crenata [misprinted as “erenata”][auct. non Spiraea L. Pallas; Thunb., Fl. Jap.: 210, 1784] as a synonym 260) Spiraea japonica Siebold, Syn. Pl. of Spiraea thunbergii. From Spiraea crenata Oeconom.: 65 (1830), nom. nud. in Thunberg’s collection the lectotype must be Present treatment: Spiraea cantoniensis selected. Lour., Fl. Cochinch. 1: 322 (1790). Thunberg’s collections were deposited Japanese name: Ko-demari; Kogomebana mainly in UPS (Spiraea crenata: nos. 12070– (Siebold 1830). 12072) and partly in the van Royen Herbarium Original material: Japonia. Siebold (now in L). Since Siebold’s annotation “Dolichos (L0176039) [Category 2]. polystachyos Th.” is on a Thunberg specimen of Honda (1938) considered this to be Spiraea Wisteria in the van Royen collection, we believe thunbergii Siebold & Zucc. that Siebold had an opportunity to examine Thunberg’s specimens in the van Royen 261) Spiraea prunifolia Siebold & Zucc., Fl. Herbarium and we have selected a lectotype Jap. 1: 131, t. 70 (1840); in Abh. Math.-Phys. Cl. of Spiraea thunbergii from the van Royen’s Königl. Bayer. Akad. Wiss. 4(2): 125 (1845). collection. Present treatment: Spiraea prunifolia Siebold Spiraea thunbergii Siebold was published & Zucc. validly by Blume in 1827, earlier than by Japanese name: Shijimi-bana; Fage bana Siebold himself in 1830. So Spiraea thunbergii (Siebold and Zuccarini 1840). Siebold (1830) is a later homonym (see below Lectotype (designated here): Japonia. Hb. no. 262b). von Siebold (L0176040) [Category 2] (Fig. Specimens used for the preparation of 143). Siebold’s Spiraea thunbergii are as follows: Original material: Japonia. Siebold s.n. Japonia. Siebold s.n. (L0176042) [Category (L0329293) [Category 2]. Legit in Japonia. 2]. Japonia. Hb. von Siebold (L0176041) Bürger. Communicavit de Siebold anno 1842, [Category 2]. Herbarium Zuccarinii (M0154017) [Category The specimen L0176041 is considered to 2]. be the one used for the description of Spiraea Although the specimen M0154017 was thunbergii Siebold & Zucc., and the basis for transferred to M from Siebold in 1842 (two the illustration in Flora Japonica, Tab. 69. years later than the publication of the name), we L0176042 is the lectotype of Spiraea thunbergii believe that it is original material. Siebold ex Blume. 296 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

262b) Spiraea thunbergii Siebold ex Blume, Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Bijdr. 17: 1115 (1827). Akad. Wiss. 3(3): 740, t. IV 2 (1843); 4(2): 191 Present treatment: Spiraea thunbergii Siebold (1845). ex Blume. Present treatment: Stephanandra incisa Japanese name: Yuki-yanagi. (Thunb.) Siebold & Zucc. ex Zabel in Gart.- Lectotype (designated here): Japonia. Zeitung (Berlin) 4: 510 (1885). Siebold s.n. (L0176042, left) [Category 2] (Fig. Japanese name: Kogome-utsugi. 145). Lectotype (designated here): Japonia. A specimen, L0176041, with Siebold’s Siebold s.n. (L0329272) [Category 2] (Fig. 147). annotation was collected in 1829 and is not Original material: Legit in Japonia & original material. Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0154024) [Category 262c) Spiraea thunbergii Siebold & Zucc., 2]. Fl. Jap. 1: 130, t. 69 (1840), nec Blume (1827) non Siebold (1830); in Abh. Math.-Phys. Cl. Stranvaesia Lindl. Königl. Bayer. Akad. Wiss. 4(2): 124 (1845). 264) Stranvaesia digyna Siebold & Zucc. Present treatment: Spiraea thunbergii Siebold in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. ex Blume, Bijdr. 17: 1115 (1827). Wiss. 4(2): 129 (1845), as “Stranvaisia.” Japanese name: Yuki-yanagi; Juki janagi, Iwa Present treatment: Pourthiaea villosa janagi (Siebold and Zuccarini 1840). (Thunb.) Decne. in Nouv. Arch. Mus. Hist. Nat. Lectotype (designated here): Japonia. 10: 147 (1874) (var. villosa). Siebold s.n. (L0176041, left) [Category 2] (Fig. Japanese name: Ke-kamatsuka. 146). Lectotype (designated here): Legit. Bürger Original material: Japonia. Siebold s.n. comcvt. de Siebold, Herbarium Zuccarinii (L0176042) [Category 2] (Fig. 145). Legit in (M0154027) [Category 2] (Fig. 148). Japonia Bürger. Communicavit de Siebold Original material: Legit. Bürger in Japonia. anno 1842, Herbarium Zuccarinii (M0154020) Communicavit de Siebold, Herbarium Zuccarinii [Category 2]. Legit in Japonia. Bürger. (M0154028) [Category 2b]. In Japonia legt. Communicavit Siebold anno 1839, Herbarium Bürger comvt. de Siebold, Herb. Zuccarinii Zuccarinii (M0154021) [Category 2]. Legit (M0154026) [Category 2b]. in Japonia. Bürger comcvt. d. Siebold, Herb. Zuccarinii (M0154022) [Category 2b]. Legit Leguminosae Dr Siebold in Japonia. Communicavit Prof. Caesalpinia L. Zuccarini, anno 1843 (M0154369) [Category 265) Caesalpinia japonica Siebold & Zucc. 2b]. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 117 (1845). Stephanandra Siebold & Zucc. in Abh. Math.- Present treatment: Caesalpinia decapetala Phys. Cl. Königl. Bayer. Akad. Wiss. 3(3) 739 (Roth) Alston in Trimen, Handb. Fl. Ceylon 6: (1843); 4(2): 191 (1845). 89 (1931). Type: Stephanandra flexuosa Siebold & Japanese name: Jaketsu-ibara. Zucc. (= S. incisa (Thunb.) Siebold & Zucc. ex Lectotype (designated here): Japonia Zabel). in monte Wunzen. Siebold s.n. (L0325745) Present treatment: Stephanandra Siebold & [Category 2] (Fig. 149). Zucc. Original material: Legit in Japonia et Communicavit de Siebold anno 1839, 263) Stephanandra flexuosa Siebold & Herbarium Zuccarinii (M0153878) [Category October 2014 The Journal of Japanese Botany Vol. 89 No. 5 297

2]. (1825). Siebold and Zuccarini described Caesalpinia Japanese name: Natamame (Siebold 1830). japonica based on Siebold’s collection and Original material: not found in L, M, or cited Caesalpinia crista L. in Thunberg’s Flora MAK. Japonica (1784, p. 179). The lectotype specimen L0325745 is a representative specimen upon 269) Dolichos icurvatus Thunb. var. [b] which the description of Caesalpinia japonica Floribus purpureis seminibus roses Siebold, was prepared. Regarding Caesalpinia japonica Syn. Pl. Oeconom.: 55 (1830), nom. illegit. to be a new species described by Siebold and [phrase name] Zuccarini, we select the lectotype from the Present treatment: probably a cultivar of collections with annotation by Siebold and Canavalia gladiata (Jacq.) DC., Prodr. 2: 404 Zuccarini. (1825). Japanese name: Natamame (Siebold 1830). Crotalaria L. Original material: not found in L, M, or 266) Crotalaria eriantha Siebold & Zucc. MAK. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 121 (1845). 270) Dolichos lablab L. var. [a] Floribus Present treatment: Crotalaria sessiliflora L., purpureis seminibus nigris Siebold, Syn. Pl. Sp. Pl. ed. 2: 1004 (1763). Oeconom.: 55 (1830), nom. illegit. [phrase Japanese name: Tanuki-mame. name] Lectotype (designated here): Japonia. Present treatment: probably a cultivar of Siebold (L0423600) [Category 2] (Fig. 150). Lablab purpureus (L.) Sweet, Hort. Brit.: 481 Original material: Legit Dr Siebold in (1823). Japonia. Communicavit Prof. Zuccarini anno Japanese name: Ingenmame (Siebold 1830). 1843, Herbarium Zuccarinii (M0153881) Original material: not found in L, M, or [Category 2]. Legit in Japonia Bürger. MAK. Communicavit de Siebold, Herbarium Zuccarinii (M0153882) [Category 2b]. 271) Dolichos lablab L. var. [b] Floribus albis seminibus albis Siebold, Syn. Pl. Dolichos L. Oeconom.: 55 (1830), nom. illegit. [phrase 267) Dolichos hassjoo Siebold, Syn. Pl. name] Oeconom.: 55 (1830), nom. nud. Present treatment: probably a cultivar of Present treatment: Mucuna pruriens (L.) DC. Lablab purpureus (L.) Sweet, Hort. Brit.: 481 var. utilis (Wall. ex Wight) Burck in Ann. Jard. (1823). Bot. Buitenzorg 11: 187 (1893). Japanese name: Ingenmame (Siebold 1830). Japanese name: Hasshô-mame; Original material: not found in L, M, or Hassjoomame (Siebold 1830). MAK. Original material: Japonia. [Siebold s.n.] (L0104303) [Category 2]. 272) Dolichos umbellatus Thunb. var. [a] Seminibus roseis leguminibusque pendulis 268) Dolichos incurvatus Thunb. var. [a] Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. Floribus albis seminibus albis Siebold, Syn. illegit. [phrase name] Pl. Oeconom.: 55 (1830), nom. illegit. [phrase Present treatment: a cultivar of Vigna name] unguiculata (L.) Walp., Repert. Bot. Syst. 1: 779 Present treatment: probably a cultivar of (1842). Canavalia gladiata (Jacq.) DC., Prodr. 2: 404 Japanese name: Sasage (Siebold 1830). 298 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

Original material: Japonia. Siebold s.n. Glycine Willd. (L0367305) [Category 2]. 276) Glycine soja Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 273) Dolichos umbellatus Thunb. var. [b] 4(2): 119 (1845). Verdcourt in Kew Bull. 24: Seminibus nigris leguminibusque pendulis 256 (1970). Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. Present treatment: Glycine max (L.) Merr. illegit. [phrase name] subsp. soja (Siebold & Zucc.) H. Ohashi in J. Present treatment: a cultivar of Vigna Jap. Bot. 57: 30 (1982). unguiculata (L.) Walp. in Repert. Bot. Syst. 1: Japanese name: Tsuru-mame. 779 (1842). Lectotype (designated here): Japonia. Japanese name: Kurosasage (Siebold 1830). Siebold s.n. (L0103661) [Category 2] (Fig. 152). Original material: Japonia. Siebold s.n. Original material: Legit Bürger in Japonia. (L0367300) [Category 2b]. Communicavit de Siebold, Herbarium Zuccarinii (M0153853) [Category 2]. In Japonia legt. 274) Dolichos umbellatus Thunb. var. [c] Bürger. Comcvt. de Siebold, Herb. Zuccarinii Seminibus nigris leguminibusque erectis (M0153854) [Category 2b]. Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. One of the original materials, M0153853, illegit. [phrase name] has an accompanying label indicating Present treatment: a cultivar of Vigna lectotypification by Tepljacova in 1990, but we unguiculata (L.) Walp. in Repert. Bot. Syst. 1: were unable to find a paper by Tepljacova on 779 (1842). this matter. Original material: not found in L, M, or MAK. Glycyrrhiza L. A specimen, Siebold s.n. (L0367301) with 277) Glycyrrhiza kansoo Siebold, Syn. Pl. Siebold’s annotation “Dolichos kuronagasasake, Oeconom.: 58 (1830), nom. nud. S.” appears to be part of the original material. Present treatment: probably Glycyrrhiza sp. Japanese name: Kanzô. Dumasia DC. Original material: not found in L, M, or 275) Dumasia truncata Siebold & Zucc. MAK. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 119 (1845). Lespedeza Michx. Present treatment: Dumasia truncata Siebold 278) Lespedeza argyraea Siebold & Zucc. & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Japanese name: No-sasage. Wiss. 4(2): 120 (1845). Lectotype (designated here): Legit Bürger in Present treatment: Lespedeza cuneata (Dum. Japonia. Communicavit de Siebold, Herbarium Cours.) G. Don, Gen. Hist. 2: 307 (1832). Zuccarinii (M0153850) [Category 2] (Fig. 151). Japanese name: Medo-hagi. Original material: In Japonia legt v. Siebold, Lectotype (designated here): Legit in Japonia Herb. Zuccarinii (M0153851) [Category 2b]. Bürger. Communicavit de Siebold, Herbarium H. E. Hazelhorst selected Pierot specimen Zuccarinii (M 0153856) [Category 2] (Fig. 153). (L0043333 and L0043334) as holo- or lectotype, Original material: Japonia. Siebold s.n. and also Siebold s.n. (L0043336) and Bürger s.n. (L0103929) [Category 2]. In Japonia legt. (L0043335) as syntypes in 1977. Although these Bürger comcvt. de Siebold, Herb. Zuccarinii specimens are fine and well representative, none (M0153857) [Category 2b]. has indication that they are original material. October 2014 The Journal of Japanese Botany Vol. 89 No. 5 299

Pachyrhizus DC. Original material: Japonia. Siebold 279) Pachyrhizus thunbergianus Siebold s.n. (L0176057) [Category 2]. Siebold s.n. & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. (L0367322, middle right and left) [Category 2]. Akad. Wiss. 4(3): 237 (1846), as “Pachyrrhizus.” Present treatment: Pueraria lobata (Willd.) 282) Phaseolus bundoo Siebold, Syn. Pl. Ohwi in Bull. Tokyo Sci. Mus. no. 18: 16 Oeconom.: 57 (1830), nom. nud. (1947). Present treatment: Vigna radiata (L.) R. Japanese name: Kuzu; Kats, Kudsu, Kudsu Wilczek, Flore du Congo Belge et du Ruanda- Kadsura, Kadsune (Siebold and Zuccarini 1846) Urandi: 6 (1954). Type: Type of Dolichos hirsutus Thunb. in Japanese name: Yae-nari; Bundoo or Trans. Linn. Soc. Lond. 2: 339 (1794). Futanarimame (Siebold 1830). This was a new name for Dolichos hirsutus Original material: Japonia. Siebold s.n. Thunb. under Pachyrhizus. (L0367326) [Category 2]. Specimens used for the preparation of the treatment of Pachyrhizus thunbergianus by 283) Phaseolus vulgaris L. var. [a] Siebold and Zuccarini are as follows: Seminibus albis Siebold, Syn. Pl. Oeconom.: 57 Legit in Japonia. Siebold s.n. (L0327145). (1830). nom. illegit. [phrase name] Japonia. Siebold s.n. (L0327149). Japonia. Present treatment: probably a cultivar of Siebold s.n. (L0327148). Legit Bürger in Phaseolus vulgaris L., Sp. Pl. 723 (1753). Japonia. Communicavit de Siebold, Herbarium Japanese name: Sirotorukusun (Siebold Zuccarinii (M0153873). In Japonia legt. 1830). Bürger comcvt de Siebold, Herb. Zuccarinii Original material: not found in L, M, or (M0153874). In Japonia legt. Bürger comcvt de MAK. Siebold, Herb. Zuccarinii (M0153875). 284) Phaseolus vulgaris L. var. [b] Phaseolus L. Seminibus rubis Siebold, Syn. Pl. Oeconom.: 57 280) Phaseolus atsuki Siebold, Syn. Pl. (1830). nom. illegit. [phrase name] Oeconom.: 57 (1830). nom. nud. Present treatment: probably a cultivar of Present treatment: Vigna angularis (Willd.) Phaseolus vulgaris L., Sp. Pl.: 723 (1753). Ohwi & H. Ohashi in J. Jap. Bot. 44: 29 (1969) Japanese name: Akasasage (Siebold 1830). (var. angularis). Original material: not found in L, M, or Japanese name: Azuki. MAK. Original material: Japan. Siebold s.n. The specimen, Siebold s.n. (L0367309) (L0367315) [Category 2]. Japonia. Siebold s.n. with Siebold’s annotation “Phaseolus aka (L0176058) [Category 2]. Japonia. Siebold s.n. nagasasake,” appears to be part of the original (L0367322, lower left) [Category 3]. material because the Japanese word “aka” means red. 281) Phaseolus atsuki var. ooatsuki Siebold, Syn. Pl. Oeconom.: 57 (1830), nom. nud. 285) Phaseolus vulgaris L. var. [c] Present treatment: a cultivar of Vigna Seminibus atro virentibus Siebold, Syn. Pl. angularis (Willd.) Ohwi & H. Ohashi (var. Oeconom.: 57 (1830). nom. illegit. [phrase angularis). name] Syn.: Phaseolus chrysanthos Savi f. ooatsuki Present treatment: probably a cultivar of (Siebold) Honda, Pl. Siebold.: 555 (1935). Phaseolus vulgaris L., Sp. Pl.: 723 (1753). Japanese name: Ooatsuki (Siebold 1830). Japanese name: Aosasage (Siebold 1830). 300 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

Original material: not found in L, M, or Present treatment: probably a cultivar of MAK. Glycine max (L.) Merr. Japanese name: Sinsjumame (Siebold 1830). 286) Phaseolus vulgaris L. var. [d] Original material: not found in L, M, or Seminibus nigris Siebold, Syn. Pl. Oeconom.: MAK. 57 (1830). nom. illegit. [phrase name] Present treatment: probably a cultivar of 291) Soja japonica Siebold var. [d] Phaseolus vulgaris L., Sp. Pl.: 723 (1753). Seminibus nigris globosis Siebold, Syn. Pl. Japanese name: Kurotorukusun (Siebold Oeconom.: 56 (1830). nom. illegit. [phrase 1830). name] Original material: not found in L, M, or Present treatment: probably a cultivar of MAK. Glycine max (L.) Merr. Japanese name: Kuromame (Siebold 1830). Soja Moench. Original material: not found in L, M, or 287) Soja japonica Siebold, Syn. Pl. MAK. Oeconom.: 56 (1830). nom. nud., as “Sooja.” Present treatment: Glycine max (L.) Merr., 292) Soja japonica Siebold var. [e] Interp. Rumph. Herb. Amboin.: 274 (1917). Seminibus nigris combressis Siebold, Syn. Japanese name: Daizu; Sooju (Siebold 1830), Pl. Oeconom.: 56 (1830). nom. illegit. [phrase Daisu (Siebold 1830). name] Original material: not found in L, M, or Present treatment: probably a cultivar of MAK. Glycine max (L.) Merr. Japanese name: Kurotokorosun (Siebold 288) Soja japonica Siebold var. [a] 1830). Seminibus albis Siebold, Syn. Pl. Oeconom.: 56 Original material: not found in L, M, or (1830). nom. illegit. [phrase name] MAK. Present treatment: probably a cultivar of Glycine max (L.) Merr. 293) Soja nomame Siebold, Syn. Pl. Japanese name: Daisu (Siebold 1830). Oeconom.: 56 (1830), nom. nud. Original material: not found in L, M, or Present treatment: Chamaecrista nomame MAK. (Siebold) H. Ohashi. in J. Jap. Bot. 64: 215 (1989). 289) Soja japonica Siebold var. [b] Japanese name:. Kawara-ketsumei; Seminibus albo bruscis Siebold, Syn. Pl. Nomame, Jawaraketsmai (Siebold 1830). Oeconom.: 56 (1830). nom. illegit. [phrase Original material: Japonia. H. B. [Hortus name] Botanicus in Dezima]. Siebold s.n. (L0325738) Present treatment: probably a cultivar of [Category 2]. Glycine max (L.) Merr. The specimen cited above has an Japanese name: Tobimame (Siebold 1830). accompanying slip with a description of flower, Original material: not found in L, M, or but the specimen has no flowers. MAK. Sophora L. 290) Soja japonica Siebold var. [c] 294) Sophora angustifolia Siebold & Zucc. Seminibus fuscis Siebold, Syn. Pl. Oeconom.: in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. 56 (1830). nom. illegit. [phrase name] Wiss. 4(2): 118 (1845). October 2014 The Journal of Japanese Botany Vol. 89 No. 5 301

Present treatment: Sophora flavescens Sol. in Zuccarini 1839). Aiton, Hort. Kew. 2: 43 (1789). Original material: not found in L, M, or Japanese name: Kurara. MAK. Lectotype (designated here): Japonia. Siebold s.n. (L0421682) [Category 2] (Fig. 154). 298) Wisteria japonica Siebold & Zucc., Fl. Original material: Legit Bürger in Japonia. Jap. 1: 88, t. 43 (1839); in Abh. Math.-Phys. Cl. Communicavit de Siebold, Herbarium Zuccarinii Königl. Bayer. Akad. Wiss. 4(2): 118 (1845). (M0121586) [Category 2]. “Legit Dr Siebold Present treatment: Wisteria japonica Siebold in Japonia. Communicavit Prof. Zuccarinii & Zucc. anno 1845, Herbarium Zuccarinii” and “In Japanese name: Natsu-fuji; Ko-fudsi (Siebold Japonia legt. Bürger comcvt. de Siebold, Herb. and Zuccarini 1839). Zuccarinii” [This specimen has two labels] Lectotype (designated probably by S. T. (M0121585) [Category 2b]. Dunn, 1912): Japonia. Herb. de Siebold (in) 1829 (L0059625) [Category 2] (Fig. 157). Wisteria Nutt. Original material: Legit Bürger in Japonia. 295) Wisteria brachybotrys Siebold & Zucc., Communicavit de Siebold, Herbarium Fl. Jap. 1: 92, t. 45 (1839); in Abh. Math.-Phys. Zuccarinii (M0153889) [Category 2]. In Japonia Cl. Königl. Bayer. Akad. Wiss. 4(2): 118 (1845). legt Bürger comcvt de Siebold, Herb. Zuccarinii Present treatment: Wisteria brachybotrys (M0153890) [Category 2b]. In Japonia legt Siebold & Zucc. Bürger comct de Siebold, Herb. Zuccarinii Japanese name: Yama-fuji; Jamma Fudsi. (M0153891) [Category 2b]. Lectotype (designated by Compton and Lack 2012): Japonia. Herb. de Siebold (L0176059) Geraniaceae [Category 2] (Fig. 155). Geranium L. Original material: Legit in Japonia de 299) Geranium thunbergii Siebold & Zucc. Siebold. Communicavit de Siebold, Herbarium in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Zuccarinii (M0153884) [Category 2]. Wiss. 4(2): 136 (1845). Present treatment: Geranium thunbergii 296) Wisteria chinensis (Sims) DC. var. Siebold & Zucc. beni-fudsi Siebold & Zucc., Fl. Jap. 1: 90 Japanese name: Gen-no-shôko. (1839), as “sinensis.” Lectotype (designated here): Japonia. Present treatment: Wisteria floribunda Nagasaki, Kosido. Sept. Oct. Herb. Thunberg (Willd.) DC., Prodr. 2: 390 (1825). s.n. (L0059752) (Fig. 158). Japanese name: Fuji; Beni-fudsi (Siebold and Original material: Japonia. Thunberg s.n. Zuccarini 1839). (L0059753). Lectotype (designated here): Japonia. Herb. This is a new name for Geranium reported de Siebold in 1829 (L0170538) [Category 5 (Fl. by Thunberg as G. palustris L. (UPS Thunberg Jap. t. 44)] (Fig. 156). collection nos. 15733 and 15734) in his Flora Japonica (1784, p. 268). Siebold & Zuccarinii 297) Wisteria chinensis (Sims) DC. var. siro- (1845) cited Geranium palustris as a synonym fudsi Siebold & Zucc., Fl. Jap. 1: 90 (1839) , as of Geranium thunbergii. So from Geranium “sinensis.” palustris in Thunberg’s collection the lectotype Present treatment: probably a cultivar of must be selected. Wisteria floribunda (Willd.) DC. As mentioned under Spiraea thunbergii Japanese name: Shiro-fudsi (Siebold and (p. 295) we believe that Siebold was able to 302 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 examine Thunberg’s specimens in the van Two specimens, Siebold s.n. (L0015981) and Royen Herbarium. Siebold s.n. (L0015982), collected in Japan in L Thunberg wrote: “pedunculis bifloris bear no annotation by Siebold or Zuccarini. longissimis declinatis, petalis integris, foliis quinquelobis incisis. ... Nagasaki, in Kosido, Cicca L. alibi...” This description well corresponds 301) Cicca flexuosa Siebold & Zucc. in Abh. to Thunberg’s specimens, (L0059752) and Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. (L0059753) in the van Royen collection and 4(2): 143 (1845). Siebold’s specimens (L0176062, L0176063) Present treatment: Phyllanthus flexuosus identifiedG. thunbergii by Siebold himself. (Siebold & Zucc.) Müll. Arg. in DC., Prodr. The following specimens were also used for 15(2): 324 (1866). the preparation of the treatment of Geranium Japanese name: Koban-no-ki. thunbergii by Siebold & Zuccarini: Lectotype (designated here): Legit in Japonia. Siebold s.n. (L0176062). Japonia. Japonia Bürger comcvt. v. Siebold, Herbarium Siebold s.n. (L0176063). Japonia. Herb. Zuccarinii (M0120849) [Category 2] (Fig. 160). Thunberg (L0059753). Original material: Japonia. Siebold s.n. According to ICN Art. 38, Siebold and (L0159844) [Category 2]. Japonia. [Siebold s.n.] Zuccarini’s Geranium thunbergii is regarded as a (MAK S1595) [Category 2]. Japonia. [Siebold new species, although Yeo (1992) did not accept s.n.] (MAK S1596) [Category 2]. Geranium thunbergii Siebold & Zucc. as such. Croton L. 299b) Geranium thunbergii Siebold ex 302) Croton siraki Siebold, Syn. Pl. Lindl. & Paxton in Paxton’s Flow. Gard. 1: 186, Oeconom.: 30 (1830), nom. nud. f. 115 (1851). Present treatment: Neoshirakia japonica Present treatment: Geranium thunbergii (Siebold & Zucc.) Esser in Blumea 43: 129 Siebold & Zucc. (1998). Japanese name: Gen-no-shôko. Japanese name: Shiraki. Lectotype (designated here): Illustration in Original material: Japonia. Siebold s.n. Paxton’s Flow. Gard. 1: 186, f. 115 (1851). (L0022693) [Category 2]. Japonia. Siebold s.n. Though Yeo (1992) regarded Lindl. & (L0022695) [Category 2]. Japonia. [Siebold s.n.] Paxton as the author of Geranium thunbergii (MAK S1592) [Category 2, 3]. Siebold, the name was legitimately published by Siebold and Zuccarini as mentioned above. 302b) Croton siraki Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. Euphorbiaceae 4(2): 144 (1845). nom. nud. Antidesma L. Present treatment: Neoshirakia japonica 300) Antidesma japonicum Siebold & Zucc. (Siebold & Zucc.) Esser. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Japanese name: Shiraki. Wiss. 4(3): 212 (1846). Original material: Legit v. Siebold in Japonia Present treatment: Antidesma japonicum et Communicavit, Herbarium Zuccarinii Siebold & Zucc. (M0120848) [Category 2b]. Japonia. [Siebold Japanese name: Yama-hihatsu. s.n.] (MAK S1592) [Category 2]. These are also Lectotype (designated here): Japonia. the original material for Croton Siraki Siebold [Siebold s.n.] (MAK S1587) [Category 2] (Fig. (see above). 159). October 2014 The Journal of Japanese Botany Vol. 89 No. 5 303

Glochidion J. R. Forst. & G. Forst. Lectotype (designated here): Communicavit 303) Glochidion obovatum Siebold & Zucc. de Siebold anno 1842, Herbarium Zuccarinii in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. (M0088455) (Fig. 163). Wiss. 4(2): 143 (1845). Original material: Japonia. [Siebold s.n.] Present treatment: Glochidion obovatum (MAK S1542) [Category 2]. Siebold & Zucc. Siebold and Zuccarini (1845) compared Japanese name: Kanko-no-ki. Phyllanthus lepidocarpus with “Ph. Niruri Lectotype (designated here): Legit Bürger in Thunb. Fl. Jap. p. 56” with a question mark. In Japonia. Communicavit v. Siebold, Herbarium L and M there are no specimens determined as Zuccarinii (M0120852) [Category 2] (Fig. 161). P. lepidocarpus by Siebold or Zuccarini. The Original material: Japonia. Siebold s.n. lectotype designated here, M0088455, was (L0030948) [Category 2]. In Japonia legt. annotated at first as “Phyllanthus Niruri L.” Bürger comunic. v. Siebold, Herb. Zuccarinii and then “P. echinatus S. & Z.” by Zuccarini. (M0120853) [Category 2b]. Legit Dr Siebold Phyllanthus echinatus Siebold & Zucc. was in Japonia. Communicavit Prof. Zuccarini anno not published, but the epithet echinatus has a 1843 (M0120851) [Category 2b]. Japonia. meaning similar to lepidocarpus. We consider Siebold s.n. (MAK S1534) [Category 2]. that Siebold and Zuccarini described P. Japonia. Siebold s.n. (MAK S1535) [Category lepidocarpus based on this specimen. 2]. 306) Stillingia japonica Siebold & Zucc. Mercurialis L. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. 304) Mercurialis leiocarpa Siebold & Zucc. Wiss. 4(2): 145 (1845). in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Present treatment: Neoshirakia japonica Wiss. 4(2): 145 (1845), as “lejocarpa.” (Siebold & Zucc.) Esser in Blumea 43(1): 129 Present treatment: Mercurialis leiocarpa (1998). Siebold & Zucc. Japanese name: Shiraki. Japanese name: Yama-ai. Lectotype (redesignated here): Legit. v. Lectotype (designated here): Legit. in Siebold in Japonia et Communicavit, Herbarium Japonia & Communicavit de Siebold, Herbarium Zuccarinii (M0120845) [Category 2] (Fig. 164). Zuccarinii (M0120841) [Category 2] (Fig. 162). Original material: In Japonia legt. Bürger Original material: Japonia. Siebold comunic. v. Siebold, Herb. Zuccarinii (L0080015) [Category 2]. In Japonia legt (M0120843) [Category 2b]. In Japonia legt. Bürger comunic. v Siebold, Herb. Zuccarinii Bürger comunic. v. Siebold, Herb. Zuccarinii (M0120842) [Category 2b]. Japonia. [Siebold (M0120844) [Category 2b]. s.n.] (MAK S1538) [Category 2]. Japonia. Peter van Welzen designated a specimen [Siebold s.n.] (MAK S1539) [Category 2]. “Japonia. Siebold s.n. (L0081028)” as the lectotype (Esser 1998) but this is not original Phyllanthus L. material because it lacks an annotation by 305) Phyllanthus lepidocarpus Siebold & Siebold and/or Zuccarini. Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 143 (1845). References Present treatment: Phyllanthus lepidocarpus Akiyama S., Thijsse G., Esser H.-J. and Ohba H. Siebold & Zucc. (see T. Kurosawa in Acta 2014. Siebold and Zuccarini’s type specimens and original materials from Japan, part 4. Angiosperms. Phytotax. Geobot. 52: 18, 2001) Dicotyledoneae 3. J. Jpn. Bot. 89: 209–229. Japanese name: Ko-mikan-sô. Aldasoro J. J., Aedo C., Garmendia F. M., de la Hoz F. P. 304 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

and C. Navarro. 2004. Revision of Sorbus Subgenera and his collaborators from Japan. J. Jpn. Bot. 88: 197– Aria and Torminaria. Syst. Bot. Monogr. 69: 1–148. 211. Compton J. and Lack H. W. 2012. The discovery, naming Ohba H. and Akiyama S. 2013b. A revision of the species and typification of Wisteria floribunda and W. of Hydrangea (Hydrangeaceae) described by Siebold brachybotrys (Fabaceae) with notes on associated and Zuccarini part 1. Bull. Natn. Mus. Nat. Sci., Ser. B, names. Willdenowia 42: 219–240. 39: 173–194. Esser H.-J. 1998. Neoshirakia, a new name for Shirakia Ohba H. and Akiyama S. 2014. A revision of the species of Hurus. (Euphorbiaceae). Blumea 43: 129–130. Hydrangea (Hydrangeaceae) described by Siebold and Hara H. 1957. Critical notes on some specimens of East- Zuccarini part 2. Bull. Natn. Mus. Nat. Sci., Ser. B, 40: Asiatic plants in foreign herbaria (8). J. Jap. Bot. 32: 29–45. 134–140. Ohba H., Akiyama S., and Thijsse G. 2005. Miquel’s new Hara H. 1978. Comments on the East Asiatic plants (5). J. taxa of the vascular plants described from Japan in Jpn. Bot. 53: 232–238. Prolusio florae japonicae and some other works. In: Honda M. 1938. Plantae Sieboldianae. In: Japanisch- Ohba H. and Boufford D. E. (eds.), The Botanical Deutsche Gesellschaft, Tokyo (ed.), Siebold Kenkyu シ Collections: Proceedings of the Symposium 'Siebold ーボルト研究 , pp. 517–596. Iwanami shoten, Tokyo. in 21st Century' Held at the University Museum, the Hu S.-Y. 1955. A monograph of the genus Philadelphus University of Tokyo, in 2003. The University Museum, (continued). J. Arnold Arbor. 36: 325–368. the University of Tokyo, Bulletin no. 41: 31–140. Koidzumi G. 1911. Notes on Japanese Rosaceae. II. Bot. Ohba H., Kawasaki T., and Tanaka H. 2007. Flowering Mag. (Tokyo) 25: 74–76. Cherries of Japan, new ed. 263 pp. Yama-kei Publishers Koidzumi G. 1911. Notes on Japanese Rosaceae. III. Bot. Co. Ltd., Tokyo. Mag. (Tokyo) 25: 183–188. Sargent C. S. 1888. Plant notes. Prunus pendula. Garden Koidzumi G. 1913. Conspectus Rosacearum Japonicarum. and Forest 1: 196. J. Coll. Sci. Imp. Univ. Tokyo 34(2): 1–312. Thunberg C. P. 1784. Flora Japonica. J. G. Mülleriano, Makino T. 1908. Observations on the Flora of Japan. Bot. Leipzig. Mag. (Tokyo) 22: 113–120. Yamaguchi T., Fujii S., Tamura M. N., Nagamasu H., Kato Maximowicz C. P. 1883. Diagnoses plantarum novarum N., Wada H., and Yamamoto A. 2003. The specimens asiaticarum, V. Bull. Acad. Imp. Sci. Saint-Pétersbourg of Japanese plants collected by Ph. F. von Siebold, 29: 51–228. H. Bürger and von Siebold's collaborators, and now Ohba H. 2001. Prunoideae. In: Iwatsuki K,. Boufford D. housed in the National Herbarium of the Netherlands E. and Ohba H. (eds.), Flora of Japan, 2b: 124–144. and the Botanische Staatssammlung München. Calanus Kodansha Scientific Inc., Tokyo. (Bull. Aitsu Mar. Stat. Kumamoto Univ. Jap.) Special Ohba H. (ed.) 2012. Tetsuya Kawasaki’s Illustrated Works No. 5, 191–576. of Japanese Flowering Cherries. 222 pp. Aboc-sha, Yeo P. F. 1992. A revision of Geranium L., in south-west Kamakura. China. Edinb. J. Bot. 49: 123–211. Ohba H. and Akiyama S. 2013a. Rosa collected by Siebold

秋 山 忍 a， G. Thijsseb， H.-J. Esserc， 大 場 秀 章 d： Siebold と Zuccarini が日本から記載した分類群レクト タイプと原資料，第五部．被子植物．双子葉植物綱 四 Siebold と Zuccarini は， 主 に Siebold 自 身 お よ び， ボルト・コレクションから Siebold と Zuccarini が記載 日本人植物学者，Bürger らの採集品からなるシーボル に用いた標本（原資料）を標本上に記された手記ならび ト・コレクションを研究し，日本から多数の新種や新変 に記載文などを参考に特定した．また，タイプが指定さ 種を発表し，日本植物相の分類学的研究に多大な貢献を れていない分類群については，特定した原資料のなかか 行った．しかしながら，それらの新植物の発表に際して ら記載に最もよく合致する標本をレクトタイプとして は大半で標本の引用がなく，またタイプの指定がなされ 選定した． なかった． 本稿はその第五部で，被子植物双子葉植物綱（ユキノ Siebold と Zuccarini が命名した学名の安定化などの シタ科からトウダイグサ科）を収載した．レクトタイプ 目的で，ライデン，ミュンヘン，牧野植物標本館（首都 の選定に関連して，とくに Deutzia crenata（ウツギ）， 大学東京），東京大学総合研究博物館に収蔵されるシー Parnassia mucronata（ウメバチソウ），Philadelphus October 2014 The Journal of Japanese Botany Vol. 89 No. 5 305 satsumanus（バイカウツギ）， Sorbus toringo（ズミ）， ズミの学名について，原 (1978) はすでに小泉が詳 Geranium thunbergii（ゲンノショウコ）， Phyllanthus 述 し て い る と し，Malus toringo (Siebold) Siebold が lepidocarpus（コミカンソウ）では選定について問題点 正 名 で あ る と し た． し か し， 小 泉 (1911, 1913) は などをコメントした． Malus toringo Siebold の 出 典 を Siebold の Catalogue Amygdalus persica f. pendula（ ベ ニ シ ダ レ モ モ ）， Raisonné (1856) と し た が， 原 は de Vriese と し Amygdalus persica f. stellata（ゲンペイキクモモ，新 Catalogue Raisonné に は 触 れ な か っ た．Catalogue 称），Amygdalus persica f. versicolor（ゲンペイモモ）， Raisonné (1856) での Malus toringo の発表は短い記載 Armeniaca mume f. pendula（シダレウメ），Cerasus を伴い新種としての要件を備えているが，同じ 1856 年 itosakura（エドヒガン）の新学名を提唱した． に de Vriese によって Sorbus toringo Siebold にもとづ エドヒガン（シダレザクラ）に Siebold が与えた く Malus toringo (Siebold) Siebold が Tuinbouw-Flora Prunus itosakura は， こ れ ま で、 記 載 を 伴 わ な い 裸 3 巻 368 ページに発表された．国際藻類・菌類・植物命 名とみなされてきたが，解説文中に “Cerasus ramulis 名規約第 41 条 4 項に従い，Malus toringo Siebold の学 pendulis（下垂性の小枝をもつサクラ）” とあり，当時 名は Sorbus toringo Siebold にもとづく新組合せとし， の日本産自生種中で下垂性サクラはエドヒガンしかな その出典を Siebold の Catalogue Raisonné (1856) とす く，この種の特徴を記述したものと理解することがで るのが適切である． きる．なお，Siebold はサクラは属としては Prunus に （a 国立科学博物館植物研究部， 分類しているので，この解説文中の Cerasus は属名と b オランダ・国立標本館 してではなく，サクラ類の意味で用いている．そこで 現所属：オランダ・国家生物多様性解析センター植物部門， Prunus itosakura を有効名と断定し，上記の新学名を提 c ドイツ・ミュンヘン州立植物標本館， 唱した． d 東京大学総合研究博物館） 306 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 307 Siebold & Zucc. (L0176009). Zucc. & Siebold Deutzia gracilis Deutzia ig. 117. Lectotype of of Lectotype 117. ig. F Siebold & Zucc. (L0170287). Zucc. & Siebold Deutzia crenata Deutzia ig. 116. Lectotype of of Lectotype 116. ig. F 306 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 307 Siebold (L0420368). Siebold Hydrangea alternifolia Hydrangea ig. 119. Lectotype of of Lectotype 119. ig. F Siebold & Zucc. (M0120971). Zucc. & Siebold Hoteia thunbergii thunbergii Hoteia ig. 118. Lectotype of of Lectotype 118. ig. F 308 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 309 Siebold & Zucc. (L0104524). Zucc. & Siebold Platycrater arguta Platycrater ig. 121. Lectotype of of Lectotype 121. ig. F Siebold & Zucc. (M0120972). Zucc. & Siebold Parnassia mucronata Parnassia ig. 120. Lectotype of of Lectotype 120. ig. F 308 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 309 Siebold & Zucc. (M0120982). Zucc. & Siebold Saxifraga cortusiifolia Saxifraga ig. 123. Lectotype of of Lectotype 123. ig. F Siebold & Zucc. (M0153974). Zucc. & Siebold Ribes fasciculatum Ribes ig. 122. Lectotype of of Lectotype 122. ig. F 310 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 311 Siebold & Zucc. (L0176017). Zucc. & Siebold Aronia asiatica Aronia ig. 125. Lectotype of of Lectotype 125. ig. F Siebold & Siebold Zucc. Schizophragma hydrangeoides Schizophragma (L0104501). ig. ig. 124. of Lectotype F 310 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 311 Siebold & Zucc. (M0153991). Zucc. & Siebold Crataegus alnifolia Crataegus ig. 127. Lectotype of of Lectotype 127. ig. F Siebold & Zucc. (L0176026). Zucc. & Siebold Ceraseides apetala Ceraseides ig. 126. Lectotype of of Lectotype 126. ig. F 312 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 313 Siebold & & Siebold sejô-jama-buki var. [α.] [α.] var. Kerria japonica Kerria Zucc. (L0328743). Zucc. ig. 129. Lectotype of of Lectotype 129. ig. F Siebold & Zucc. (M0153734). Zucc. & Siebold Crataegus cuneata Crataegus ig. 128. Lectotype of of Lectotype 128. ig. F 312 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 313 Siebold (L0102980). Siebold Prunus itosakura Prunus ig. 131. Lectotype of of Lectotype 131. ig. F Siebold (L0102984). Siebold Persica pendula Persica ig. 130. Lectotype of of Lectotype 130. ig. F 314 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 315 Siebold & Zucc. (L0329140). Zucc. & Siebold Prunus mume Prunus ig. 133. Lectotype of of Lectotype 133. ig. F Siebold & Zucc. (L0329088). Zucc. & Siebold Prunus macrophylla Prunus ig. 132. Lectotype of of Lectotype 132. ig. F 314 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 315 Siebold & Zucc. (M0154004). Zucc. & Siebold Prunus spinulosa Prunus ig. 135. Lectotype of of Lectotype 135. ig. F Siebold Siebold pendula Siebold & Zucc. var. var. Zucc. & Siebold Prunus mume Prunus (MAK S0551, left). S0551, (MAK ig. 134. Lectotype of of Lectotype 134. ig. F 316 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 317 Siebold & Zucc. (L0176032). Zucc. & Siebold Raphiolepis japonica Raphiolepis ig. 137. Lectotype of of Lectotype 137. ig. F Siebold & Zucc. (M0154368). Zucc. & Siebold Pyrus gracilis Pyrus ig. 136. Lectotype of of Lectotype 136. ig. F 316 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 317 Siebold ex Regel (M0154345). Regel ex Siebold Rosa iwara Rosa ig. 139. Lectotype of of Lectotype 139. ig. F Siebold & Zucc. Rhodotypos kerrioides (L0329269). ig. 138. Lectotype of F 318 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 319 Siebold & Zucc. Sieversia dryadoides (L0175827). ig. 141. Lectotype of F Siebold & Zucc. (M0120864). Zucc. & Siebold Rubus ribifolius Rubus ig. 140. Lectotype of of Lectotype 140. ig. F 318 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 319 Siebold & Zucc. (L0176040). Zucc. & Siebold Spiraea prunifolia Spiraea ig. 143. Lectotype of of Lectotype 143. ig. F Siebold & Zucc. (L0176022). Zucc. & Siebold Sorbus toringo Sorbus ig. 142. Lectotype of of Lectotype 142. ig. F 320 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 321 Siebold ex Blume (L0176042, (L0176042, Blume ex Siebold Spiraea thunbergii Spiraea left). ig. 145. Lectotype of of Lectotype 145. ig. F Siebold (L0144310). Siebold Spiraea thunbergii Spiraea ig. 144. Lectotype of of Lectotype 144. ig. F 320 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 321 Siebold & Zucc. Stephanadra flexuosa (L0329272). ig. 147. Lectotype of F Siebold & Zucc. (L0176041, (L0176041, Zucc. & Siebold Spiraea thunbergii Spiraea left). ig. 146. Lectotype of of Lectotype 146. ig. F 322 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 323 Siebold & Zucc. (L0325745). Zucc. & Siebold Caesalpinia japonica Caesalpinia ig. 149. Lectotype of of Lectotype 149. ig. F Siebold & Zucc. (M0154027). Zucc. & Siebold Stranvaesia digyna Stranvaesia ig. 148. Lectotype of of Lectotype 148. ig. F 322 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 323 Siebold & Zucc. (M0153850). Zucc. & Siebold Dumasia truncata Dumasia ig. 151. Lectotype of of Lectotype 151. ig. F Siebold & Zucc. (L0423600). Zucc. & Siebold Crotalaria eriantha Crotalaria ig. 150. Lectotype of of Lectotype 150. ig. F 324 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 325 Siebold & Zucc. (M0153856). Zucc. & Siebold Lespedeza argyraea Lespedeza ig. 153. Lectotype of of Lectotype 153. ig. F Siebold & Zucc. (L0103661). Zucc. & Siebold Glycine soja Glycine ig. 152. Lectotype of of Lectotype 152. ig. F 324 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 325 Siebold & Zucc. Wisteria brachybotrys (L0176059). ig. 155. Lectotype of F Siebold & Zucc. Sophora anguistifolia (L0421682). ig. 154. Lectotype of F 326 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 327 Siebold & Zucc. (L0059625). Zucc. & Siebold Wisteria japonica Wisteria ig. 157. Lectotype of of Lectotype 157. ig. F

beni-fudsi Siebold & Zucc. var. var. Zucc. & Siebold Wisteria chinensis Wisteria Siebold & Zucc. (L0170538). Zucc. & Siebold ig. 156. Lectotype of of Lectotype 156. ig. F 326 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 327 Siebold & Zucc. (MAK S1587). (MAK Zucc. & Siebold Antidesma japonicum Antidesma ig. 159. Lectotype of of Lectotype 159. ig. F Siebold & Zucc. (L0059752). Zucc. & Siebold Geranium thunbergii Geranium ig. 158. Lectotype of of Lectotype 158. ig. F 328 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 329 Siebold & Zucc. (M0120852). Zucc. & Siebold Glochidion obovatum Glochidion ig. 161. Lectotype of of Lectotype 161. ig. F Siebold & Zucc. (M0120849). Zucc. & Siebold Cicca flexuosa Cicca ig. 160. Lectotype of of Lectotype 160. ig. F 328 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月 October 2014 The Journal of Japanese Botany Vol. 89 No. 5 329 Siebold & Zucc. Phyllanthus lepidocarpus (M0088455). ig. 163. Lectotype of F Siebold & Zucc. (M0120841). Zucc. & Siebold Mercurialis leiocarpa Mercurialis ig. 162. Lectotype of of Lectotype 162. ig. F 330 植物研究雑誌 第 89 巻 第 5 号 2014 年 10 月

Fig. 164. Lectotype of Stillingia japonica Siebold & Zucc. (M0120845).