bioRxiv preprint doi: https://doi.org/10.1101/2020.09.13.295170; this version posted September 14, 2020. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY-NC-ND 4.0 International license. 1 Title: Tripartite holobiont system in a vent snail broadens the concept of chemosymbiosis 2 Authors: 3 Yi Yang1, Jin Sun1, Chong Chen2, Yadong Zhou3, Yi Lan1, Cindy Lee Van Dover4, Chunsheng 4 Wang3,5, Jian-Wen Qiu6, Pei-Yuan Qian1* 5 Affiliations: 6 1 Department of Ocean Science, Division of Life Science and Hong Kong Branch of the 7 Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), The Hong 8 Kong University of Science and Technology, Hong Kong, China 9 2 X-STAR, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), 2-15 10 Natsushima-cho, Yokosuka, Kanagawa 237-0061, Japan 11 3 Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry 12 of Natural Resources, Hangzhou, China 13 4 Division of Marine Science and Conservation, Nicholas School of the Environment, Duke 14 University, Beaufort, NC, United States 15 5 School of Oceanography, Shanghai Jiao Tong University, Shanghai, China 16 6 Department of Biology, Hong Kong Baptist University, Hong Kong, China 17 Correspondence to: 18 Pei-Yuan Qian:
[email protected] bioRxiv preprint doi: https://doi.org/10.1101/2020.09.13.295170; this version posted September 14, 2020. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity.