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uoslry' C d ,{g BIIerlsnY urelsod\ Jo dladpugrd (eeecelng dgurey eq1uo salou JIIuouoxEI Only species: R. euphemiae(F. Muell.) P. G. Wilson comb. nov. Nemat.olepiseuphemiae F. Muell., Fragm. 3:149 t.25 (Apr. 1863).- Pheb.rliumeuphemiae (F. Muell.J C. A. Gardn., Enum. pl. Austial. Occid. 7_0(1931).-Type: Near Cape Arid, Maxuell (iso. K, photo seen). P. baxte.riqenth., Fl. Austial. 1:345 (30 May 1863).-M. baxtiri (Benth.) lngler in.Engler et Prantl, Nat. Pflanzenfam. 3/4:t4S \1896). T-ype': W. Au'tralia. sourhcoasr. Ba.\/.,r, tholo.:K. photo seen.;. Distribution: Found near the south coast of Western from the Eyre Range east to Mt. Ragged. _ _This species was placed in the genus Nematolepis by Mueller because it has lrgulatestamens and becausehe thought (ilcorectly) that the petalswere united. The seed, anther, and other floral characters iik lt with- ph"b,:ttium secl, Gonioclados-which section includes p. rude and p. anceps,It is also closely related to_tlre _genus Chorilaenq. AII these taxa have the hilum surrounded by i ligament-like border, a character not found elsewherein the tribe Boronieae. Rhqdinoth.tmunusls unique in the family Rutaceaein having sub-orbicular tetra- colporate pollen grains, whereas in all other genera they are ellipsoidal and ldcolporate.

DIPLOLAENA R. Br. Diplolaena ferruginea P. G. Wilson sp. nov. Folir,lareelliplicaca2_\lcm.-uprale\ia.BldndUlo-o-punctala.glabravel , glabrescenlia. suOIu\ pallrOa ,\elultna. _( apt(uld ca. Z cm lara: bracLeaele ugtneo_lOmentosde,e\ternae angusre-rflcngllrare\.LJ, tl mm tongae..tn-lernae ljneari_acuminatdel pelala brcclej, p.lulo ofe\ror

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'9182 'Hrxgd: olo" 696I xr's'Llrnour Luo{ Itu 0I r?J.ta.^Iu reo,erg *rrJ';fiylr't;ffi 'Nnlqe[8 'Lun8uolLuur €c 'Iunl"clns JollpulpnlrSool 'runlP,{"i3'Lunu3€tu ] 'slq!tllols "ru8rts snl{ls :runsolrd es.ledsrunuE^O 'req€15'sueuurnloo rotl^arq snaslc slul le 'o€3uol -lEd,spue ruruZ.I Ec'oelnce'eele^ooBre{lue:esoanJle^'exeUulealcie'u8uolLuru8I 'ea'Elenuoll€ €urleladelueLuBJu :e8uolrutu g.Z Bc '€le^elc 'essErc"ulFdes "luoLu€JU:€solrd 'ErqEI3 'BlEIn.Ide rs€q 'eqdrourrp Eurruels elplrr^-o^r?gaprlted lolllealdeqns e8uol tutu 8 :er'e1u,roqo 'elecuqllrr eleled eln-leqnd'el€lnclcle lellle3ld€qns tSuol LuLu '!eeplollep 'rtEloeltrerqlq'I8uol 'llellJqrnl_!]sn8u? t-Z elsnSuelo.\ ele^o eledes tutu € ea Il{aalped:sor€ll)ie 'rr.rellosserol{ e8uol urul elreeull lloIIoJ :r8uol url'u €c SI-l'esntqo'eteeun.qo-"ueeull'elElotloj '€]€uutcl le^ t srprqcrqr rtncrrre:elluars.rq€I3 Ie^ elqe[3 Z-€ e!lo{ llnraqnd IIntueU '^ou'ds uoslrlA O d clB^€lJ Bluorog relSuA ( qlueg) aB.rpuBreleH pes 'ursYlNoxos

'rlleJuoq '3uo1 f,sotuJuro]AIJSuap puP urc 7 ec 'pro^oqo 01 cr1drl1a,{lptorq ere sa^?el erll nlotldato.tcltu nt ul.roJed,(1 eq1 rt1 'urrll 1o 'iuoruels 3uo1 ,{re,t puu se.teel l€U sBq }r :ur8pelunt{ pus 8uu?punt{ uee,rl3q punoJ sr rlcrq,\\ 'qltloB tlPuotuun)p'r?A loJ sI Jsrllouv e^oq? peqlJcsep ,t:leu€^ eql .roJ sr eserll euo 'souo8elEc culcedse{ul el"elc 01 elqvcqcerd Jo 'stuJoJ .reedd8lr soop eseqlJo lcurtsrp lsolu arll ,{luo JoJ lnq lueJ3up snoleunu strlqrqxesarceds eql ?eJ? eplt\ slql ra^O re^Q{ uoslqclnl tr oql se rluou J"J s€ ls€oc qlnos eqt tuo{ sr op7dacotctw g setcedsetll Jo uollngu]slp eql filamenta incurva, pilosula, sepalinistereliuscLrlis, ca. 2 mm longis, globulo magno verrucoso terminatis, petalinis brevioribus,teretiusculis, ca. 1.2 mm longis; antherae + aequales,ca. 0.8 mm longae, breviter apiculatae. Discus puberulus, albus, in n'ussas4 pulviniformes petalinas divisus. Ovar:um humile, densepilosulum; sessilequadrilobum, ca. 6 mm latum. album. TypeaP. G. Wikott'7108,28 km W of Ravensthorpeand 12 knr N of Ravensthorpe-Ongerup Road,l4.vii. 1968(holo.:PERTH, iso.:K, MEL). Undershrub ca. 0.3 rrr high; branchlets minutely puberulous in two opposite lines. Leaves subsessile,temate, glabrousi leafletssubterele to sub- clavate,ca. 5 mm long. Flowers axillary, solitary, pendulous. turb- inate, ca. 2 mm long, conspicuouslybibracteolate. ovate, ca. 3 mm iong, glabrescent.Petals broadly elliptic, ca. 8 mm long, glabrescent,greenish- cream 10 reddish-brown.Stamens all fertile; filaments incurved, pilosulose, the antisepaloussub-terete, ca.2 mm long, with a large, abaxial muricare swelling at the apex, the antipetalousshorter, sub-terete,ca. I.2 mm long, with no apical swelling; anthers cordate, ca.0.8 mm long, shortly white- apiculate, Disc puberulous to glabrous, pale green, divided into four anti- petalous cushion-shapedmasses. Ovary denselypubesceDt, equal to disc in height; stigma sessile,hemispherical ca. 0 6 mm wide, with four rounded lobes,papiJlose, pale green. Distributio : Western Australia, soutlt-westregion, from Gnowangerup east to the West River. Cnowangerrrp,C. A. Gat(lner2091]'10 mi N of Borden,Mr. J.!nes,10.rx.1960: l5 mi W of Ongerup.K Newbq 352|.3mi N of Ongerup,K. Newbet2443,2144:55 km N of Brener Bay,P. G. Wilson5:102;4 mi E of WestRlvet, F. Lulllitz 5399. Boronia octandrq has externally an appearalce similar to B,crassifolia. It differs fron that speciesin having almost terete leaflets, large sepaline stamens which have a massiveapical swelling,a divided disc, and a small stigma. It also has tl'reanlhers consistentlysimilar and ferlile (a character approached by some forns of B.uassi/olia). A divided disc is found in the two heterandrous speciesB. tetrundra and B. purdieana, but in these the disc masscsare opposite to the sepals whereas in B. octandra they are alternate to them.

Boronia sect.Cyanothamnus (Lind].) F. Muell. Boronia coerulescensF. Muell. subsp.spicata P. G. Wilson subsp.nov. Ramulierecti, derse pilosuli. Folia erectavel leviterdivaricata, anguste-oblonga usque anguste-cuneatavel elliptica,integra, 5 l0 mm longa,obtusa, pilosula, supra concavi. Floies in racemospiciformi folioso aggregati. Pedicelli ca. 2 nlm longi, pilosuli,bracteolis parvis basalibus ,ornati. Sepala anguste-ovala, ca. 4 mm longa,proninenter glanduioso-punctata, pilosula.Ovarium pilosulum. Tfpe: J. W. Gteen1500,2 mi S of Wubin,erect shrub 50 cm high,petals white ringed blue on dorsalside, yellow sand, 22.viii. 1957 (holo.:PERTH). Branches erect, densely pilosulose.Leaves erect or slightly spreading, narrow-oblong to narrow-cuneateor elliptic, entire, 5-10 mm long, obtuse, pilosulose, upper surface concave. Flowers in dense leafy, spike-like . Pedicelsca. 2 mm long, pilosulose,with a pair of small basalbracteoles. Sepals narrow ovate, ca.4 mm long, prominently glandular punctate, pilosulose. Ovary pilosulose. Distribution: WesterrrAustralia, betweenWubin and Muntadgin. 6mi Sof Wubin,T. l.4Aplin 565: Bet\\eenMollerin and Koorda, W/. E. Blackall3452; MuntadgiD,E. T. Bailet 727.

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Sepalsovate- acuminate, ca 3 mrn long, very sparsely stellate-pilose, red. Petals valvate, ovate, ca_7_mm long (enlarging slightly in fruit), acute, loosely stellate-pilose outside, glabrous within except near apex, pale pink, midrib prominent. Stamens 2 5-3 mm long (the antisepalous slightly longer than the antipetalous); fila- ments subterete, somewhat swollen and veffucose at apex, sparsely stellate- hirsute; anthers cordate, ca. I mm long, apex rounded witfi an acuminate and recurved apiculum ca 1_length of loculi; pollen orange. Disc entire, pale red, glabrous. Ovary hemispherical, glabrous; style shofi, terete, ca. 0.2 mm long, glabrous; stigma capitate. Seed reniform,3.5 mm lonq; ourcr resra membranous,dull; placentalendocarp persistent,thick: raphe-insigniflcant; chalaza near base on adaxial margin. Distribution: Western Australia, between Lake King and the Johnston Lakes. HattersHilI (40kln NE of Lake King townshj.p),Oct. 1929,H. Steednan32 (in fruit). This is similar to B. erici;folia, a speciesfound much farther north between Moora and the Wongan Hills, B, revoluta differs most noticeably in having muci longer hairson stem and petals.a periojatetnol sessile.;Iiaf, and larger flowers. Since the collectionsseen have both been made on iron- stone hills it would appear probable that B. reyoluta is restdcted to this type of habitat. B. eticiJblia grows on yellow sand or loam.

Boronia sect. Boronia Boronia oxyantha Turcz. var. brevicalyx (Benth.) P. G. Wilson comb. nov. B. lanuginosaEndl. var. brevicalyx Benth., Fl. Austral. l:317 0863).- 7.1'per"Phillips River, Herb. Mueller", (iso.:MEL). The type of Boronia lanuginosa was thought by Endlicher to have come from King George Sound on the south-west coast of Western Australia. Ben- tham, who had rot seen the type specimen, assumed that the short orieinal descdption applied to a Boronia which had been collected by Robert Brown in the same area. Reference to the type of B. lanuginosa (Ferd. Bauer, Nalur- historisches Museum, Wien) shows that it belongs in fact to a speciesfound along the nodh coast of Australia and which receivedthe I aler nam€of B. artemi- misifolia F. -Muell. The correct name for the speciesBentham had in mind, which came from the south coast, is -8. strictq Barll. Bentham's vartetv "brevi- calyx" is not, however, closely related to this species nor to B. laiuginosa, but to ,8. ox)a thq. Although conspecific with the latter it differs sufficiently to warrant varietal recognition; in var. oxyqntha the branches are puberulous and the sepalsca. 5 mm long and acuminate, while in var. brevicalyx the btanches are hirtellous and the sepalsare acute and ca. 2 mm long. The variability within tar. bre calyx suggeststhat it may grade into the typical variety but for this to be confirmed further collecting is necessaryin the Ravensthorpe-Fitzgerald R. areawhere both are Iound.

Bororia capitata Benth. subsp. clayata P. G. Wilson subsp. nov. Folia carnosasaepe sub-clavata, basinr versus sparse ciliata aliter Blabra.Bracteolae naviculiformes,latae. Sepala manifeste ciliall. aliter glabra.apice incrassata; petala glabra. Twe: l'/.5 mi W of LakeGrace, .!. ,S.George 348,3.ix. 1959 Oolo.:PERTH, iso.:K). Leaves fleshy, often sub-clavate, sparsely ciliate towards base, otherwise glabrous. Bracteolesnaviculiform. broad. Sepals prominently ciliate other- wise glabrous, apex thickened. Petals glabrous. Distributiotl : Western Australia, south-west. Corrigin-Nyabing district. 1 mi W of Corfigin,,(. Ne".',bq2610; K]ulin, A. Ashby43: Tarin Rock, C. A. Gardner1745; 2 mi N of Nyabing,K. Newbey439. 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[ - S urteuurd 1:,r elqorlojir; er1o1 ilir:aqnd as:eds lyireanf nluti ur rureli ',rou ds uos1rL1 n ,1 r1u6.4,ruluorog 'sledes .soqcu"Jq luocsoqld reln8uer-r1,{l,troleu pue prte s3^Eel luecseqnd Brir.teq uI o,{\l l€qlo eqt luo{ sreJlp lr :,{lle8tlrd J?eu €eJ€ Il€urs " uro4 u,rou) ,{luo sr sirJedsqnslEJrd{l JqJ uotlnqulsrp Jo s"al" e:}€Jedese,ltq prrr ecuereed -dp ur lJurlsrp ars pJpl^rp sr DtolldtD.g qcrq,r olur sercedsqnsoeJrll erTJ ' .O .y.Suopurf 665Vn[ov 'JaArl{ ,"rT€]lsnv leJeAJ€Jr'{prle uollessng ueo,rleg lso,r-qlnos vtelse{\ :uo!fiqulslT 'snorqslS sJ"led .elelncrdu ,{lelnuru .o}"rTrce,peJ 'snoJq€I8 'J?ln8uEul .slelsnlc .es"q fl,\\oll"u sledas leurtujol ll?rrrs ur sJe,rolC J€elJo oprsJeqlre uo.rJ luollncap seurl pasrBlur eso|d ,:apualsseqcuerg '(): ,dull],{S osr'HJXEd:oloq) €S6l x €'dnculer{ )c€lg.TlSt,to;y.O V :adtJ 'prqclS plElad: ptpt Jtde al.tutru.rrqnr .etErltJe.elqelS'EUelnBueuL rt\r)ffueplpdos:rtpBrrSSp un^rpd LunlnldRcrr s.roll i.rqe.siorrle.eidrlr::sl;ds.n.r5\Lulscq 'uAr-'JuoJ .tsolrd p.rdn\'erlJlJl-clns'etJtJeJEptlol snq4uelncop siaurl ur ,selltel8 llnLu€l{ '^ou'dsqns .q1ueg uoslr/ry9 d slIIrErE'dsqns e1el1ilucego.rog alibusaggregali..Sepala imbricata, late ovata\el senli-orbicularia,ca. 1 mm longa,losea, apiceoblusa, indistincte subterminaliter apiculata. Stamina erecta; filamenta cla--vata, alii qrantum vertucosa;antherae cofdatae acute apiculatae, ca. 1 nl'n longae.Discus planus, simplex.Ovarium hemisphericum; stylus breviter c)lindricus, ca. 1 mm longus;stigma subl capitatum,4-sulcatum. Type:Abovt 20 mi S of Mt. Raggedon road to IsraeliteBay, growing on limestonerubble, plant 50 cm high,petals pink, 3.x.1970,P. G. Wilson10083 (holo.:PERTH, iso.:K). Small ericoid shrub ca. 50 cm high. Branchlets faintly glandular-verrucose, minutely puberulous in 1wo opposite sunken grooves, otherwise glabrous. Leavesternately trifoliolate or pinlately 5-foliolate, ca. 1-5 cm long, coriaceous, glandular-verrucose(when dry), glabrous; petiole semiterete,2-4 mm long; leaflets narrowly obcuneate, ca. 12 x | 5 mm, apex obtuse to rounded. Floweis completely glabrous, in small terminal clusten not exceedingthe leaves.Pedicel ca. 3.mm long, glabrescent, medially minutely bibracteoiate. Sepals imbricate, broadly ovate to semi-orbicular, ca. I mm long, thick. Perals imbricate, broadly ovate, ca. 5 mm long, pink; apex obtuse,indistinctly subterminally apiculate. Stamens erect, the antisepalous slightly exceedingthe stigma; filaments clavate, somewhatverrucose; anthers cordate, acutely apiculate,ca. 1 mm long. Disc flat, simple. Ovary hemispherical;style shortly cylindrical, ca. I mm long; stigma sub-capitate, 4-grooved, Fruit not seel. This specieshas only been recorded from the type locality in the south- east of Western Australia. It is found on thin limestone soil both in sand-heath and in mallee vegetation. BetweenMt. Raggedand IsraeliteBay, malle€on limestone,28)<.1967, J. S. Beard53Ol (KINCSPARK. PERTH)

Boronia crenulata Sm. var. gracilis (Benth.) P. G. Wilson comb. nov B. viminea var. graciln Benth., Fl. Austral. 1:325 (1863).,T1tpe: Drwnmond 1848colln. no. 92 (holo.:K, photo seen). This taxon is convarietal with B. vimineq Lindl., B. tenuifoliq Bartl., a\d B- machardiqnq F. Muell. lt differs from the typical variety moit conspicuously in the floral charactem; i\ yat- crenulata the pedicels are short, often thick, and tlre sepals ciliate, whereas in vat. gracilis the pedicels are slender and the sepalseciliate.

Boronia sect. Imbricatae Engl. Engler, Nat. Pflanzenfam. 3/4:136 (1896). Lectotlpe: B. cymosa ErLdl. This section consists of three series which may be distilguished by the following key:- Sepals persistent. Staminal-filament apex not swollen; funicle expanding into an aril in centre of adaxial surface of seed ser. Imbricatae Staminal-filament apex swollen; funicle slender, no aril present ser. Ovatae Sepalsfalling in fruit; funicle slender ser. Pedunculatae

SeriesImbricatae Only species:B. cymosaEndl.

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'IpurI DJqns g sr sJr.rJssnlt ur sarlads JJrllo ^Fro erJJ 'qloolus 'snouerqLuelll ?1s3]r3lrlo :lEcndllls 'll?rus 'tueul unlrq :snocnpEc'urql d.ttcopue l?tueJ€ld :.I€lnctuel fllEplosdrllo pees -€lg uor od ol pexuB 'el€lncrd"-e:lq,r sJeqtu" :xedu el"cunru Jo l"crd"-qns'a}e.re} uo o,rs E qlJ./d el"rTrc + s}uoru?lg I?uluels luelsrs:od sledeg are clearly distincl.ln Philoth?ca the petals are soft. the stamens are all fertile with versatile anthers, and the carpels have a termiiral sterile portion which is produced in fruit as a short beak. In D nmon,lita the petalsire glumaceous, only the antipetalous stamens bear anthers (and these are dorsifixed), and the ovary and fruit are rounded at the apex. The following four speciesare klown:- 1, Drummondita ericoidesHarvey, l.c. Philothecq ericoirles(Harv-) F. Muell., Fragm. 7:21 (1869).-Zype: ,,Near the summit of the White Peak, a detachedhill near Moresbls Range,, (iso.:K, photo seen). 2. Drummondita miniata (C. A. Gardn.) P. G. Wilson comb. nov. Philotheca miniatq C. A. Gardn., J. Roy. Soc. W. Aust. 14:79 (1928). Type: near Cue, C. A. Gardner Jul. 1927; east of Cue, W. D. Campbell June i902 (syn.:PERTH), (lecto.:July 192'7colln. PERTH). 3. Drummondita hassellii (F. Muell.) P. G. Wilson con.rb.nov. Philothecahqssellii F. Muell., South. Sci. Rec.3:3 (1883). Iype..,,North- eastof Janamonjup'.A. y. Har-rel/(holo.:MEL). This speciesis very variable in flower and size but one form apDears to warrant varietal _recognition. The two infraspecific taxa may be reiognised by the following key : Leaves semi-tereteto obovoid, 3 12 mm long, smooth var. hassellii Leaves slender-terete,ca, 17 mm long, rugulose var. longifolia Var. longifolia P. G. Wilson var. nov. _. Folialeviter divaricata, aDguste-teretia, ca. 17_(20)x I mm,rugulosa (statu sicco), nitida. Floresusque 25 n'lmlongi; petalafilamentaque rubra. Tf!:r_g. .A. Gadner,15.\i. 1929,Peak Charles, in granitecrevices, 1300 feet alt. (hoto.: PERTH.iso.:K). _ Leaves somewhat spreading, slender-terete,ca. 17 (20) x 1 mm, rugulose (when dry), glossy. Flowers to 25 mm long, petals and filaments red. Distribution: Western Australia. Known only from the type locality ca. 40 km W of Dowak, which is between Norseman and Esperance. 4. Drummondita calida (F. Muell.) P. G. Wilson comb. nov. "In Philotheca calida F. M:nell., Fragm. 7 :2I (1869).-Type: montibus ad flumen Gilbert's River", R. Daintree (holo.:MEL). Distribution : Northern Queensland.

UROCARPUS Drumm. ex Harv. UrocarpusDrumm. ex Harv., J. Bot. Kew Misc.7:54 (Feb. or Mar. 1855).- Type: U. phebalioidesDrumm. ex Harv. Asterclasia s:cl. Urocqrpus (Harv.) Benth., Fl. Austral. 1:362 (1863). AsterolqsiqF. Muell,, Trans. Phil. Soc. Vict. 1:10 (Sept.-Nov. 1855).- Lectotlpe (Baillon, 1872): A. trymalioides F. Muell. Pleur,andropsisBaill., Adansonia 10:305 (1872). Type: P.phebalioides (F. Muell.) Baill. ly'ote. F. Mueller, (1855) in the same paper described two speciesin the genus Asterolasia, A. trymalioides and. A. phebalioides. The latter was r.emoved by Baillon uho placedit in his monotypic genusPleurandropsrs: thus the formei speciesbecame the lectolypeof Asrerolqsia. The correct genedc name for the Western Australian speciesis (Jrocarpus, th-is being th€ garliest of the three generic synonyms cited above. The spe;ie; of New South Wales and Victoria could be consideredsectionally or geneiically LIz

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