Journal of Animal Diversity Online ISSN 2676-685X Volume 3, Issue 1 (2021)

Review Article http://dx.doi.org/10.52547/JAD.2021.3.1.6

An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Paolo Parenti

Department of Earth and Environmental Sciences, University of Milano-Bicocca, Piazza della Scienza, 1, 20126 Milano, Italy *Corresponding author : [email protected]

Abstract A checklist of the damselfishes of the world, family Pomacentridae, is presented. A total of 798 nominal species belonging to 423 valid species and 29 genera is included. Most of the species are grouped in five genera: Chromis (109 species), Pomacentrus (82), Chrysiptera (40), Stegastes (39), and Amphiprion (29), but 12 genera contain only one or two species. The following main taxonomic decisions are taken: Chrysiptera personata Fowler, 1946 is a new synonym of Chrysiptera rex (Snyder 1909); Dischistodus notophthalmus (Bleeker, 1853) is the valid Received: 23 November 2020 name for the species known as Dischistodus melanotus (Bleeker, 1858). Accepted: 30 December 2020 Chaetodon rotundus Linnaeus, 1758 and Chaetodon rotundatus Lacepède, 1802 Published online: 30 June 2021 are senior synonyms of Abudefduf bengalensis (Bloch, 1787). Perca japonica Bloch, 1792 is a senior synonym of Chromis notata (Temminck and Schlegel, 1843); Pomacentrus niomatus De Vis, 1884 is a senior synonym of Stegastes fasciolatus (Ogilby, 1889); Glyphisodon sculptus Peters, 1855 is a senior synonym of Abudefduf notatus (Day, 1870). All these senior synonyms are herein regarded as nomina oblita. Neopomacentrus flavicauda is proposed as nomen novum Neopomacentrus xanthurus Allen and Randall, 1980. Stegastes adustus (Troschel, 1865) is the valid name for the species known as Stegastes dorsopunicans (Poey, 1868) and Wangia Fowler, 1954 is an available name not invalidated by Wanga Chen 1943.

Key words: Chrysiptera personata, Glyphidodon sculptus, nomen protectum, nomen oblitum, nomen novum

Introduction to forked. The mouth is small with a single row of conical to incisiform teeth, in some genera followed Damselfishes are typical inhabitants of tropical marine by 1–3 rows of conical or flattened teeth, no teeth on reef environments, primarily in the Indo-Pacific, vomer and palatines, and the snout length usually although a few species are found in fresh and brackish shorter than eye diameter. The trunk, fin bases and water (Neopomacentrus aquadulcis, N. taeniurus, head are scaled. The scales are ctenoid on the trunk, Pomacentrus taeniometopon, and Stegastes otophorus) opercle and preopercle, but cycloid elsewhere. The (Kohda, 1988; Allen, 1989; 1991; Jenkins and Allen, lateral line is interrupted. The body color is 2002) and a number of species lives in cooler extremely variable, from brown or black to bright temperate seas (e.g. Australia and New Zealand). blue, yellow or orange, also with a dark ocellus on Damselfishes are small percoid fishes usually not the dorsal fin in several species (Nelson et al., 2016). exceeding 10–15 cm in total length. They are characterized by having an ovate to elongate, The life cycle is characterized by five stages: egg, laterally compressed body, a single dorsal fin with larva, settling larva, juvenile, and adult. Eggs are VIII–XVII (usually XII–XIII) spines and 9–21 soft guarded by the male. Damselfishes display a rays, anal fin with II spines and 9–16 soft rays, remarkable ecological and behavioral diversity, pectoral fins with 14–22 rays, and caudal fin rounded generally occurring amongst living or dead corals,

This article is published with open access on www.jad.lu.ac.ir | © Lorestan University Press 37 Parenti 38 atoll lagoons, the vicinity of dropoffs on outer reefs, (Amphiprioninae, Chrominae, Lepidozyginae, and and other inshore areas usually at depths shallower Pomacentrinae) are considered by Nelson et al. (2016), than 20 m. The majority of species are omnivorous. but about 40 new species have been described in the Damselfishes are frequently territorial, particularly last five years (Fricke et al., 2020). A molecular study algal-feeding species. Some live in association with by Cooper et al. (2009) demonstrated that anemones (Amphiprion) and a few are cleaners. Amphiprioninae are nested within Pomacentrinae and Detailed ecomorphological studies have revealed that that Chrominae are still valid if the genera pomacentrid diversification is not due to a single Acanthochromis Gill, 1853 and Altrichthys Allen, adaptive radiation with an early burst of 1999 are moved to Pomacentrinae. Also, they erected diversification, but that “repeated ecological radiations two new subfamilies Abudefdufinae and Stegastinae have characterized the diversification of damselfishes” (Cooper et al., 2009). However, these two names are (Frédéric et al., 2013). A comprehensive treatment of invalid since they are not in accordance with the rules general biology and ecology of damselfishes has been of the Zoological Code of Nomenclature, as explained recently published (Frédéric and Parmentier, 2016). by Cooper and Santini (2016). The correct name for Pomacentridae have no economic importance except these two new subfamilies are Glyphisodontinae and for the aquarium trade. Twenty-nine genera (Allen, Microspathodontinae, respectively (Table 1). 1999b) with 387 species, arranged into 4 subfamilies

Table 1: Classification of genera and species in the family Pomacentridae.

No. of Subfamily Tribe Genus species Azurina 2 Chrominae Bonaparte, 1831 Chromis 109 Dascyllus 11 Glyphisodontinae Richardson, 1844 Abudefduf 21 Lepidozyginae Allen, 1975 Lepidozygus 1 Hypsypops 1 Microspathodon 4 Microspathodontinae Jordan and Evermann, 1898 Microspathodontini Nexilosus 1 Similiparma 2 Mecaenichthys 1 Parma 10 Microspathodontinae Jordan and Evermann, 1898 Parmini Whitley, 1929 Plectroglyphidodon 10 Stegastes 39 Amphiprion 29 Pomacentrinae Bonaparte, 1831 AmphiprioniniGill, 1859 Premnas 1 Acanthochromis 1 Altrichthys 3 Amblyglyphidodon 11 Amblypomacentrus 3 Cheiloprion 1 Chrysiptera 40 Dischistodus 7 Pomacentrinae Bonaparte, 1831 Pomacentrini Bonaparte, 1831 Hemiglyphidodon 1 Neoglyphidodon 9 Neopomacentrus 16 Pomacentrus 82 Pomachromis 4 Pristotis 2 Teixeirichthys 1

Many new species have been described in the last ten plus morphology and genetic markers, that reveal the years and an updated checklist of damselfishes is existence of species complexes and geographic needed, as testified by recent papers citing outdated variations which might be difficult to solve (e.g. the reference works. For example, Arango et al. (2019) Western Atlantic species of the genus Stegastes stated that “The Pomacentridae (damselfishes and Jenyns, 1840 as shown by de Souza et al., 2017). anemonefishes) is one of the largest families of reef In this study several taxonomic decisions have been fishes, with over 330 described species (Michael, undertaken, and they are summarized in the Results 2008)”. The presence of cryptic species have been section before the checklist of species. recently challenged by Souza et al. (2017) who emphasized inconsistencies between color patterns,

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Material and Methods Glyphisodon Lacepède, 1802: 542. Masculine. Type species Glyphisodon moucharra Lacepède 1802. Type Original names, authorship and dates have been by subsequent designation by Jordan 1917: 64. checked on ECoF (Eschmeyer’s Catalog of Fishes) Glyphidodon Agassiz, 1846: 164. Unjustified (Fricke et al., 2020) and in the original publication. emendation of Glyphisodon Lacepède, 1802. Type material was examined for three previously Euschistodus Gill, 1862: 145. Masculine. Type species unplaced species, namely Chrysiptera personata Euschistodus declivifrons Gill 1862. Type by Fowler, 1946; Glyphisodon sculptus Peters, 1855 and subsequent designation by Jordan and Evermann Glyphidodon fallax Peters, 1855. 1898: 1560. In the present annotated checklist each genus and Nexilaris Jordan and Evermann, 1896: 512. species is arranged alphabetically, bearing in mind Masculine. Type species Euschistodus concolor Gill, the subfamily classification as proposed by Cooper et 1862. Type by original designation. al. (2009) and modified by Cooper and Santini Nexilarius Jordan and Evermann, 1898: 1559. (2016) should be adopted when phylogenetic studies Masculine. Unjustified emendation of Nexilaris Jordan of generic interrelationships are considered (Table 1). and Evermann, 1896. For each species, the type locality, type material, Indoglyphidodon Fowler, 1944b: 25. Masculine. Type synonyms, and distribution are given. Scientific species Indoglyphidodon abbotti Fowler 1944. Type names and authorship were checked by consulting by original designation). the original references in the digital library BHL or in Tropical Indo-Pacific and Atlantic Oceans. 14 species the conventional library as recorded in ECoF (Fricke according to Kuiter (1993); 18 species according to et al., 2020). Allen (1991) and 20 species according to Nelson et al. (2016); 21 species are listed here. The list of institutions where the type material is located is in Appendix. Abudefduf abdominalis (Quoy and Gaimard, 1825) Glyphisodon abdominalis Quoy and Gaimard, 1825: Results 390. (Type locality, Hawaiian Islands, U.S.A). Holotype: MNHN 8327. At present there are 798 nominal species belonging Distribution: Johnston Atoll and Hawaiian Islands. to 423 valid species grouped in 29 genera. The Abudefduf bengalensis (Bloch, 1787) following results, which are not reported in or Chaetodon rotundus Linnaeus, 1758: 277. (Type disagree with the ECoF, are here summarized. locality, Indian Ocean). No types known. Chaetodon rotundus Linnaeus, 1758 and Chaetodon Chaetodon rotundatus Lacepède, 1802: 452. rotundatus Lacepède, 1802 are senior synonyms of Misspelling or new name for Chaetodon rotundus Abudefduf bengalensis (Bloch, 1787); Perca Linnaeus, 1758. japonica Bloch, 1792 and its nomina nova are all Chaetodon bengalensis Bloch, 1787: 110, Pl. 213 senior synonyms of Chromis notata (Temminck and (fig. 2). (Type locality, Bay of Bengal, India). Schlegel, 1843); Pomacentrus niomatus De Vis, Syntypes: ZMB 2722 (2), ?8771 (dry). On page 82 of 1884 is a senior synonym of Stegastes fasciolatus Bloch's Ichthyologie, v. 6. (Ogilby, 1889); Glyphisodon sculptus Peters, 1855 of Labrus macrogaster Lacepède, 1801: 430, 477, Pl. Abudefduf notatus (Day, 1870) and all these senior 19 (fig. 3). (Type locality, Sea of Indies). Syntypes: synonyms are invalidated by reversal of precedence MNHN A-8126 (1, dry), 9442 (1, dry). (ICZN, Art. 29.3) to preserve stability of zoological Glyphidodon affinis Günther, 1862: 41. (Type names. Chrysiptera personata Fowler, 1946 is a new locality, China). Holotype: BMNH 1851.12.27.175. junior synonym of Chrysiptera rex (Snyder, 1909). Glyphisodon palmeri Ogilby, 1913: 87, Pl. 22 (fig. Neopomacentrus fallax (Peters, 1855) is the senior 2). (Type locality, Moreton Bay, Queensland, synonym of N. taeniurus (Bleeker, 1856). The genus Australia). Holotype: QM I.1059. Paratypes: AFAQ name Wangia Fowler, 1954 is available and is a (1) whereabouts unknown. junior synonym of Teixeirichthys Smith, 1953. Distribution: Pakistan to southern New Guinea and Species accounts Rottnest Island (Western Australia) and Port Macquarie (New South Wales, Australia) north to Infraclass Teleostei Müller, 1845 Vietnam, China and southern Japan. Division Acanthopterygii Artedi, 1738 Comments: Chaetodon rotundus Linnaeus, 1758 has Subdivision Percomorphaceae Wiley and Johnson, been described having XXIII dorsal fin spines and 3 2010 dorsal soft rays, but, Cuvier (1830: 462) considered Series Ovalentaria Smith and Near, 2012 this measure a misprinting for the correct number of Family Pomacentridae Bonaparte, 1831 dorsal fin elements as XIII/13. Under his Genus Abudefduf Forsskål, 1775 Glyphisodon bengalensis Cuvier wrote: «Je pense Abudefduf (subgenus of Chaetodon) Forsskål, 1775: même qu’il faut ici chercher le chaetodon rotundus xiii, 59. Masculine. Type species Chaetodon sordidus de Linnaeus […] etr sa description est très conforme Forsskål 1775. Type by subsequent designation by à ce que nous voyons dans nos glyphisodon; […] Or, Jordan and Evermann, 1917: 33.

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 39 Parenti 40 qui a jamais vu un chétodon, et même un poisson Abudefduf declivifrons Gill, 1862 quelconque, avec vingt-trois aguillons et seulement Euschistodus declivifrons Gill, 1862: 146. (Type trois rayons mous?» Other features (a rounded body, locality, Cape San Lucas, Baja California, Mexico). presence of five bands on the body as described and Syntypes: (orig. 7) USNM 9332 (now 10). figured by Bloch (1787) fit with the species A. Distribution: Gulf of California to Acapulco, Mexico. bengalensis. Chaetodon rotundus Linnaeus, 1758 and Abudefduf hoefleri (Steindachner, 1881) Chaetodon rotundatus Lacepède, 1802 has not been Glyphidodon hoefleri Steindachner, 1881a: 159. used as a valid name since 1899, while Chaetodon (Type locality, Gorée, Senegal). Holotype: NMW bengalensis Bloch, 1787 has been used as a valid 76421. Species illustrated and described in more name in over 25 publications by more than 10 detail in Steindachner 1881b: 43 [page 27 of authors in the last 50 years. Therefore, under Article separate], Pl. 5 (fig. 2). 23.9.2 of the Code, Chaetodon bengalensis Bloch, Heliastes bicolor Rochebrune, 1880: 166. (Type 1787 is here regarded as a nomen protectum and locality, Casamance River, Senegal). No types known. Chaetodon rotundus Linnaeus, 1758 and Chaetodon Distribution: Capo Verde Islands, Islands of Gulf of rotundatus Lacepède, 1802 are here regarded as Guinea, and Benin. nomina oblita. To give evidence of this decision the Comments: Type catalog for Heliastes bicolor in following 25 works are produced: Allen (1975), Dor Bauchot et al. 1978: 34 with Sauvage as author. A (1984), Allen and Swainston (1988), Randall et al. synonym of Chromis chromis (Linnaeus 1758) (1990), Allen (1991), Kuiter (1992), Kuiter (1993), according to Monod (1973: 424).Valid as Chromis Francis (1993), Goren and Dor (1994), Mohsin and bicolor (Lloris and Rucabado 1990: 845); Ambak (1996), Allen (1997; Randall et al. (1997), questionable a synonym of Abudefduf hoefleri Myers (1999), Johnson (1999), Kuo and Shao (1999), (Steindachner, 1881) according to Allen (1991: 234). Nakabo (2000), Sadovy and Cornish (2000), Hutchins (2001), Allen (2001), Allen and Adrim Abudefduf lorenzi Hensley and Allen, 1977 (2003), Golani and Bogorodsky (2010), Motomura et Abudefduf lorenzi Hensley and Allen 1977: 109, Figs. al. (2010), Allen and Erdmann (2012), Larson et al. 1–2. (Type locality, Tenibuli Bay, Isabel Island, (2013), Psomadakis et al. (2020). Solomon Islands, w. Pacific, ca. 8°25'S, 159°50'E). Holotype: FMNH 23564. Paratypes: AMS I.12069 (1), Abudefduf caudobimaculatus Okada and Ikeda, 15360-001 (1), 16694-003 (1), 16710-001 (2); BMNH 1939 1858.4.21.307 (2); BPBM 9501 (4), 15998 (1); CSIRO Abudefduf caudobimaculatus Okada and Ikeda, 1939: C282 (1), C452 (1), C1778 (1); FMNH 23563 (1), 169, Fig. 6. (Type locality, Ishigaki Island, Okinawa 40675 (1); RMNH 886 (1), 11818 (1 or 2), 14932–33 Prefecture, Ryukyu Islands, Japan). Holotype: YCM- (1, 1), 27463 (1), 28162 [ex RMNH 11817] (1); P 30435 [ex Okada coll. spec. 2596]. USNM 216250 (1); WAM P.24924-001 (1). Abudefduf sp.: Kato 2011: 136, unnumbered figs. Distribution: Indo-Australian Archipelago: Philippines (Type locality, Ishigaki Island). and Moluccas to the Solomon Islands; Palau in Distribution: Ryukyu Islands, southern Japan and Micronesia. Pari Island, Seribu Islands, Jakarta, Indonesia. Comments: Resurrected from synonymy with A. Abudefduf margariteus (Cuvier, 1830) vaigiensis by Wibowo et al. (2017). Glyphisodon margariteus Cuvier in Cuvier and Valenciennes, 1830: 470. (Type locality, Mauritius). Abudefduf concolor (Gill, 1862) Syntypes: MNHN 0000-9443 (1, dry), 0000-8373 (1). Euschistodus concolor Gill 1862: 145. (Type locality, Glyphidodon rhyncholepis Bleeker, 1869: 230. (Type Panama). Syntypes or possible syntypes: BMNH locality, Réunion, western Mascarenes, southwestern 1862.5.21.22 [ex USNM] (1); USNM 4355 (2, lost), Indian Ocean). Holotype: RMNH 6507. 4357–59 (1, 2, 1), 4361–62 (12, 1), 30986 (1). Glyphidodon maculipinnis Bliss, 1883: 55. (Type Pomacentrus robustus Günther, 1862: 17. (No locality, Mauritius, Mascarenes, southwestern Indian locality). Holotype: BMNH 1855.9.19.273. Ocean). Syntypes: MCZ 6092 (4), USNM 153569 Distribution: Lower Gulf of California to Peru [ex. MCZ 6092] (1). including Galapagos Islands. Distribution: Mauritius and Réunion. Abudefduf conformis Randall and Earle, 1999 Abudefduf natalensis Hensley and Randall, 1983 Abudefduf conformis Randall and Earle, 1999: 335, Abudefduf natalensis Hensley and Randall, 1983: 2, figs. 1–4. (Type locality, Entrance of Taiohai Bay, Figs. 1, 2, 3a. (Type locality, Reef off Sodwana Bay, Nuku Hiva, Marquesas Islands). Holotype: BPBM Kwazulu, South Africa, 12 m). Holotype: BPBM 12476. Paratypes: AMS I.39228.001 (2); BPBM 22734. Paratypes: BMNH 1917.5.1.6 (1), 12482 (2), 25718 (25), 25756 (2); CAS 204162 (2); 1980.11.5.1 (1); BPBM 22732–33 (1, 1); CAS 04 MNHN 1998-1480 (4); NSMT-P 55946 (2); USNM (1); MNHN 1980-1498 (1); RUSI 514 [ex 76-16] 352406 (2). (27), 9121 (1), 9914 (1); USNM 223079 (1); WAM Distribution: Marquesas Islands. P.26529-001 (2).

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Distribution: South Africa to Madagascar and Western Chaetodon marginatus Bloch, 1787: 98, Pl. 207. Mascarenes. (Type locality, Antilles). No types known. Chaetodon mauritii Bloch, 1787: 109, Pl. 213 (Fig. Abudefduf nigrimargo Wibowo, Koeda, Muto and 1). (Type locality, Brazil). No types known. Motomura, 2018 Chaetodon sargoides Lacepède, 1802: 453, , Pl. 10 Abudefduf nigrimargo Wibowo, Koeda, Muto and (Fig. 3). (Type locality, Martinique Island, West Motomura, 2018: 472, Figs. 1–5, 6a–b. (Type Indies). No types known. locality, Off Jia-le-shui, Kenting, Taiwan, 22°00'N, Glyphisodon moucharra Lacepède, 1802: 542, 543. (Type 120°52'E; purchased at Hengchun fish market). locality, Brazil, Arabia, East Indies). No types known. Holotype: NMMB-P26624. Paratypes: KAUM-I Sparus fasciatus Gronow in Gray, 1854: 60. (Type 39257, 109043 (1, 1); NMMB P26625, P27395, locality, Indian Ocean). Syntypes: BMNH P27396 (1, 1, 1). 1853.11.12.105 [Gronovius coll.] (1, skin). Distribution: Taiwan North to Southern Japan. Apogon quinquevittatus Blyth, 1858: 272. (Type Abudefduf notatus (Day, 1870) locality, Port Blair, Andaman Islands). No types known. Glyphisodon sculptus Peters, 1855a: 456. (Type Abudefduf ascensionis Fowler, 1919: 217, Fig. 4. locality, Mozambique). Syntypes: ZMB 2738 (2). (Type locality, Ascension Island). Holotype: USNM Glyphidodon notatus Day, 1870: 521. (Type locality, 42315. Paratypes: USNM 42314 (2), 42316 (4), Andaman Islands). Syntypes: (7) ZSI F1433 (1), 273309 (1). A.424 (6). Distribution: Rhode Island to Uruguay, Capo Verde Abudefduf clarki Snyder, 1911: 534. (Type locality, Islands and tropical eastern Atlantic to Angola. Pool at Tanegashima, Osumi Islands, Japan). Abudefduf septemfasciatus (Cuvier, 1830) Holotype: USNM 68234. Paratypes: CAS-SU 21423 Glyphisodon septemfasciatus Cuvier in Cuvier and (2); USNM 74534 (1), 74588 (2). Valenciennes, 1830: 463. (Type locality, Mauritius). Indoglyphidodon abbotti Fowler, 1944b: 26, Fig. Syntypes: MNHN 0000-8372 (2). (Type locality, Tropical Indian Ocean). Holotype: ? Abudefduf paee Curtiss, 1939: 115. (Type locality, USNM 44537. Lagoon near Tautira, Tahiti, Society Islands). No Chrysiptera paucifasciata Fowler, 1946: 152, Figs. types known. 17–18. (Type locality, Aguni Shima, Ryukyu Islands, Distribution: Western Indian Ocean to Tuamotu Japan). Holotype: ANSP 72014. Paratypes: ANSP Archipelago and Line Islands South to Eastern 72013 (1), 72015–16 (1, 1). Australia North to Japan. Distribution: South Africa, Seychelles and Christmas Island to New Guinea North to Japan. Abudefduf sexfasciatus (Lacepède, 1801) Comments: Glyphisodon sculptus Peters, 1855 is ? Chaetodon abudafur var. hanni Forsskål, 1775: 65, here treated as senior synonym of Abudefduf notatus xiii. (No locality stated, presumed Red Sea). No (Day, 1870). Glyphisodon sculptus has not been used types known. as a valid name since 1899, while the latter has been Labrus sexfasciatus Lacepède 1801: 430, 477, Pl. 19 (fig. used as a valid name in over 25 publications by more 2). (Type locality, Indo-West Pacific). Syntypes: MNHN than 10 authors in the last 50 years. Therefore, under A.121 (2, poor condition), ?MNHN 5136 (1) Malabar. Article 23.9.2 of the Code, Abudefduf notatus (Day, Glyphisodon melanurus Cuvier, 1829: 180. (no 1870) is here regarded as a nomen protectum whereas locality) Available from footnote as, "Gl. melanurus, Glyphisodon sculptus Peters, 1855 is here regarded N., ou labre six bandes, Lacép. III, xix, 2." as nomen oblitum. To give evidence that reversal of Abudefduf coelestinus Cuvier (ex Solander) in Cuvier precedence is applicable the following list of 25 and Valenciennes 1830: 464, Pl. 135. (Type locality, publications is cited: Allen (1975), Akimichi and Indian Ocean). Syntypes: MNHN 0000-5136 (1) Sauchomal (1982), Chang et al. (1983), Allen and Malabar, A-0121 (2, poor condition). Steene (1988), Allen (1991), Myers (1991), Randall Distribution: Red Sea and western Indian Ocean, and Anderson (1993), Allen and Smith-Vaniz (1994), Persian Gulf, Socotra, Seychelles, Comoros, Chen et al. (1995), Heemstra (1995), Randall (1995), Madagascar and Mascarenes eastward to Marshall Donaldson (1996), Randall et al. (1997), Ganaden Islands, Marquesas and Gambier islands; Australia and Lavapie-Gonzales (1999), Randall and Lim North to Japan. (2000), Allen and Adrim (2003), Choi et al. (2003), Comments: Glyphisodon coelestinus Cuvier is Manilo and Bogorodsky (2003), Cooper et al. (2009), considered valid by Winterbottom et al. (1989) in Motomura et al. (2010), Allen and Erdmann (2012), their list of fishes of Chagos Archipelago. However, Fricke et al. (2013), Koeda et al. (2016), Kwun et al. G. coelestinus is a redescription and a new name of (2017), Zajonz et al. (2019). Labrus sexfasciatus Lacepède as mentioned by Cuvier (1830: 464). Chaetodon abudafur var. hanni Abudefduf saxatilis (Linnaeus, 1758) Forsskål, 1775 is placed into synonymy with Chaetodon saxatilis Linnaeus, 1758: 276. (Type Abudefduf bengalensis (Bloch 1787) by the ECoF locality, India and other localities). Syntypes: BMNH following Dor (1984: 179). However A. bengalensis 1853.11.12.105 [Gronovius coll.] (1, skin), NHRM is not known from the Red Sea. Forsskål’s LP 60 (1), ZMUU Linn. coll. 49 (1).

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 41 Parenti 42 description is very concise and includes the presence 0000-8321 (1), 0000-8326 (1) Sulawesi, 0000-8497 of five large bands on body. (1) Red Sea, A.8127 (1). Glyphisodon waigiensis Cuvier in Cuvier and Abudefduf sordidus (Forsskål, 1775) Valenciennes, 1830: 457. (Type locality, Waigeo, Chaetodon sordidus Forsskål 1775: 62, xiii. (Type Indonesia; Rawak Island, Bismarck Archipelago). No locality, Jidda, Saudi Arabia, Red Sea). Holotype: types known. ZMUC P56264 (dry skin, poor condition). Holotype Chaetodon tyrwhitti Bennett, 1830: unnumbered figured in Klausewitz and Nielsen 1965: Pl. 34. pages, Pl. 25. (Type locality, South coast of Sri ? Glyphidodon adenensis Günther in Playfair and Lanka). No types known. Günther, 1866: 83, Pl. 11 (Fig. 1). (Type locality, Glyphisodon quadrifasciatus Bleeker, 1847: 17. Aden, Yemen, northwestern Indian Ocean). (Type locality, Jakarta, Java, Indonesia). Syntypes: Holotype: BMNH 1867.3.9.317 (right half skin). ?BMNH 1862.2.28.28 (1). Glyphidodon leucopleura Day, 1877: 385, Pl. 83 Abudefduf quinquilineatus von Bonde, 1934: 452, Pl. (Fig. 4). (Type locality, Andaman Islands, east of 23 (Fig. 6). (Type locality, Zanzibar fish market, Bay of Bengal). Syntypes (2) and/or Day specimens: Tanzania, western Indian Ocean). Holotype: ? AMS I.95 (1) Bay of Bengal, NMW 60982 (1), ZSI Whereabouts unknown. 1434 (1, figured specimen). Distribution: Red Sea and western Indian Ocean to Abudefduf tridentatus Clark, 1938: 181. (Type locality, Tuamotu Archipelago and Line Islands south to Taiohae Bay, Nukuhiva Island, Marquesas Islands). Australia north to Japan. Mediterranean Sea (Red Sea Holotype: CAS 5533. Paratypes: CAS 5534 (1). immigrant); Hawaiian Islands introduction. Distribution: Red Sea and western Indian Ocean to Comments: Often erroneously placed in synonymy Pitcairn Group and Hawaiian Islands; Australia north with A. saxatilis (i.e. Lloris and Rucabado, 1990: to Japan. 843; Goren and Galil 1998: 57) which is a complex Abudefduf sparoides (Quoy and Gaimard, 1825) of three species, one of which is A. vaigiensis Glyphisodon sparoides Quoy and Gaimard, 1825: (Kailola, 1987; Myers, 1991). 394. (Type locality, Mauritius). Syntypes: MNHN Abudefduf whitleyi Allen and Robertson, 1974 0000-8332 (1). Abudefduf whitleyi Allen and Robertson 1974: 154, Pl. Pomacentrus zanzibarensis Von Bonde, 19(34: 452, 1. (Type locality, Pixie Reef, Great Barrier Reef, off Pl. 23 (Fig. 5). (Type locality, Shearwater Patches, Cairns, Queensland, Australia, 16°32'S, 145°51'E, 3 north of Zanzibar, Tanzania, western Indian Ocean, m). Holotype: AMS I.16634-001. Paratypes: AMS depth 17 fathoms). Holotype: Whereabouts unknown. IA.894 (1), IA.1772 (1), IB.3112 (1), I.15638-007 (7), Distribution: Southwestern Indian Ocean including I.15642-001 (10), I.15681-004 (9), I.15683-003 (67); Madagascar, Aldabra, Mauritius and Réunion. BPBM 13605-06 (3, 1); USNM 207939-40 (1, 2). Abudefduf taurus (Müller and Troschel, 1848) Distribution: Great Barrier Reef, Coral Sea and New Glyphisodon taurus Müller and Troschel in Caledonia. Schomburgk, 1848: 674. (Type locality, Barbados, Genus Acanthochromis Gill, 1863 West Indies). Possible types: ZMB 2716 (1). Acanthochromis Gill, 1863: 214. Masculine. Type Euschistodus analogus Gill, 1863: 220. (Type locality, species Dascyllus polyacanthus Bleeker, 1855. Type Aspinwall, Panama). Holotype: USNM 6419. by original designation. Distribution: Southern Florida and Caribbean Sea; Heptadecanthus Alleyne and Macleay, 1877: 343. Cape Verde Islands and off Senegal to Angola. Masculine. Type species Heptadecanthus longicaudis Abudefduf troschelii (Gill, 1862) Alleyne and Macleay. 1877. Type by monotypy. Glyphidodon troschelii Gill 1862: 150. (Type Indo-Australian Archipelago; one species. locality, Cape San Lucas, Baja California, Mexico). Acanthochromis polyacanthus (Bleeker, 1855) Syntypes: (about 100) BMNH 1862.5.21.18-20; Dascyllus polyacanthus Bleeker 1855c: 503. (Type FMNH 221 (1); MCZ 35978 [ex USNM 8173] (2); locality, Batjan, Moluccas Islands, Indonesia). USNM 8173 (50+), 8180 (7), 9347 (8). Holotype: ?BMNH 1862.2.28.33. Bleeker specimens: Distribution: Baja California to Peru including AMS B.8246 (1) Malay. Galapagos Islands and other offshore islands. Heptadecanthus longicaudis Alleyne and Macleay, Abudefduf vaigiensis (Quoy and Gaimard, 1825) 1877: 343, Pl. 15 (Fig. 3). (Type locality, Cape Glyphisodon vaigiensis Quoy and Gaimard, 1825: 391. Grenville, Queensland, Australia). Syntypes: AMS (Type locality, Waigeo, Indonesia). Lectotype: MNHN I.16356-001 [ex MAMU F820] (3). 0000-5317. Paralectotypes: MNHN 0000-5317 (4). Heptadecanthus brevipinnis De Vis, 1885: 872. Lectotype designated by Wibowo et al. (2017). (Type locality, Queensland Coast, Australia). Glyphisodon rahti Cuvier in Cuvier and Holotype: QM I.947. Valenciennes 1830: 456. (Type locality, New Heptadecanthus maculosus De Vis, 1885: 873. (Type Guinea; Red Sea; Java, Indonesia; Sulawesi, locality, Cardwell, Queensland, Australia. Type Indonesia). Syntypes: MNHN 0000-5315 (1) Java, material whereabouts unknown.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Abudefduf jordani Seale, 1906: 53, Fig. 2. (Type Amblyglyphidodon batunaorum Allen, 1995 locality, Shortland Island, Solomon Islands, Pacific). Amblyglyphidodon batunai Allen, 1995: 86, Fig. 1. Holotype: BPBM 1162. Paratypes: (6) ANSP [ex (Type locality, Lagoon next to shore, south side of BPBM] 87028 [ex BPBM] (1), 91529 [ex BPBM] Bunaken Island, Manado Bay, n. Sulawesi, ca. (1), 93275 (1); BPBM 1115-19 (now 0), 1302 (now 1°42'N, 124°57'E, 5-6 m). Holotype: NCIP 6132. 0); USNM 109381 (1). Paratypes: NCIP 6133 (1); USNM 332339 (6); WAM Chromis desmostigma Fowler and Bean, 1928: 51. P.25248-004 (1), P.30152-001 (3), P.30812-002 (1). (Type locality, Tataan Island, Tawi Tawi Islands). Distribution: Malaysia and Indonesia East to Palau, Holotype: USNM 89954. Paratypes: USNM 93520 Yap and New Ireland (Papua New Guinea), South to (3), 98241 (2). Northwestern Australia and Solomon Islands. Distribution: Indonesia, Philippines, to Solomon Islands, Amblyglyphidodon curacao (Bloch, 1787) Vanuatu and northeastern Australia and Melanesia. Chaetodon curacao Bloch, 1787: 106, Pl. 212 (fig. Comments: Homalogristes guntheri Alleyne and 1). (Type locality, Southern America, Curacao Island Macleay, 1876 listed by Allen (1991: 235) as synonym [probably Indian Ocean]). Holotype: ZMB 8772 (dry, of Acanthochromis polyacanthus is actually a junior lost). On page 79 of Bloch's Ichthyologie, vol. 6. synonym of the grouper Epinephelus coioides Glyphisodon trifasciatus Bleeker, 1847a: 19. (Type (Hamilton, 1822). locality, Jakarta, Java, Indonesia). Syntypes: RMNH Genus Altrichthys Allen, 1999 6483 (2 of 56). Appeared first as name only in Altrichthys Allen, 1999a: 23. Masculine. Type Bleeker 1845: 523; also described as new in Bleeker species Chromis azurelineatus Fowler and Bean, (1847b: 156). 1928. Type by original designation. Glyphisodon smaragdinus Brevoort, 1856: 264 [12], Philippines. Three species. Pl. 6 (fig. 3). (Type locality, Ryukyu Islands, Japan). No types known. Altrichthys alelia Bernardi, Longo and Quiros, 2017 Amblyglyphidodon nudirostrum Fowler, 1944a: 178, Fig. Altrichthys alelia Bernardi, Longo and Quiros, 2017: 32. (Type locality, Vanuatu). Holotype: ANSP 71387. 48, Figs. 1–3, 4 (left). (Type locality, San José, Distribution: Gulf of Thailand to Ryukyu Islands; Busuanga Island, Philippines, 12.1911°N, 120.1022°E, Melanesia to Samoa, Marshall Islands, and Kiribati. depth 3 meters). Holotype: PNM 15195. Paratypes: CAS 241438, 3 adult specimens; CAS 241439, 8 Amblyglyphidodon flavilatus Allen and Randall, 1980 juvenile individuals. Amblyglyphidodon flavilatus Allen and Randall, Distribution: Busuanga Island, Philippines. 1980: (6) 14, Figs. 4–5. (Type locality, Taba, 3 km south of H. Steinitz Marine Laboratory, Gulf of Altrichthys azurelineatus (Fowler and Bean, 1928) Aqaba, Rea Sea, 14 m). Holotype: BPBM 13398. Chromis azurelineatus Fowler and Bean, 1928: 57, Pl. 5. Paratypes: BMNH 1976.9.20.8-9 (2); BPBM 14671 (Type locality, Port Uson, west of Pinas Island, (1), 18121 (1), 19892 (1), 20347 (1), 20474 (2); HUJ Philippines). Holotype: USNM 89957. Paratypes: ANSP 8372 (1); MNHN 1976-119 (2), 1977-454 (2); 80317 (3); USNM 96398-99 (10, 5), 96424–25 (6, 1). USNM 216478 (13); WAM P.25619-001 (2). Distribution: Philippines. Distribution: Red Sea and Gulf of Aden. Altrichthys curatus Allen, 1999 Amblyglyphidodon flavopurpureus Allen, Erdmann Altrichthys curatus Allen, 1999a: 24, Figs. 1–3. and Drew, 2012 (Type locality, Cuyo Island, Palawan Province, Amblyglyphidodon flavopurpureus Allen, Erdmann Philippines). Holotype: UPMSI 1998-000002-FI. and Drew in Allen and Erdmann, 2012: 1126, Figs. Paratypes: UPMSI 1998-000003-FI (10); USNM 1–5. (Type locality, Numamuran Strait, 2°27.178'S, 348456 (94); WAM P.31400-001 (6), 31406-001 (3), 134°31.832'E, Cenderawasih Bay, West Papua 31407-001 (8). Province, Indonesia, depth 8–15 meters). Holotype: Distribution: Northern Palawan Province, Philippines. MZB 20578. Paratypes: MCZ 20587 (6). Genus Amblyglyphidodon Bleeker, 1877 Distribution: Cenderawasih Bay, West Papua, Amblyglyphidodon (subgenus of Glyphidodon) Bleeker, Indonesia. 1877c: 92, 94. Masculine. Type species Glyphisodon Amblyglyphidodon indicus Allen and Randall, 2002 aureus Cuvier 1830. Type by original designation. Also Amblyglyphidodon indicus Allen and Randall, 2002: appeared in Bleeker 1877d: 40. 140, Figs.1–3. (Type locality, Villingili Island, North Tropical Indo-West Pacific; 11 species. Malé Atoll, Maldive Islands). Holotype: WAM Amblyglyphidodon aureus (Cuvier, 1830) P.26514-008. Paratypes: AMS I.41006-001 (1); Glyphisodon aureus Cuvier (ex Kuhl and Van Hasselt) BPBM 13877 (2), 18901 (1), 18924 (7), 32970 (2); in Cuvier and Valenciennes, 1830: 472. (Type locality, CAS 214469 (1); NCIP 6182-4 (3); NSMT-P 61776 Java, Indonesia). Holotype: RMNH D2407. (1); ROM 46338 (3), 71277 (17), 71278 (15), 72634 Distribution: Eastern Indian Ocean to Fiji north to (10); USNM 366837 (1); WAM P.25523-008 (1), Marshall Islands and Ryukyu Islands. P.25527-001 (2), P.26514-006 (4).

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 43 Parenti 44

Distribution: Red Sea and Indian Ocean from East Africa Müller, 1840. Type by subsequent designation by and Madagascar east to Andaman Sea and Sumatra. Jordan and Evermann (1898: 1569). Indo-Australian Archipelago. Three species. Amblyglyphidodon leucogaster (Bleeker, 1847) Glyphisodon leucogaster Bleeker, 1847a: 26. (Type Amblypomacentrus breviceps (Schlegel and Müller, locality, Jakarta, Java, Indonesia). Holotype: RMNH 1840) 6849 (1 of 7). Bleeker specimens: AMS B.8238 Glyphisodon breviceps Schlegel and Müller, 1840: 23. [from Day] (1); NMV 46138 (1), BMNH (Type locality, Indian Ocean, SW. coast of Sumatra, 1862.2.28.92. Appeared first as name only in Bleeker Indonesia). Syntypes: RMNH 900 (2) SW. Sumatra. (1845: 523). Also in Bleeker (1847b: 156). Pristotis trifasciatus Bleeker, 1848: 637. (Type Distribution: Red Sea and western Indian Ocean to locality, Bima, Sumbawa Island, Lesser Sunda Islands, Melanesia, Micronesia, and Samoa; Great Barrier Indonesia). Syntypes: RMNH 6542 (some of 7). Reef north to Ryukyu Islands. Pomacentrus nematopterus Bleeker, 1852a: 285. (Type locality, Ambon Island, Molucca Islands, Amblyglyphidodon melanopterus Allen and Randall, Indonesia). Holotype: 6542 (1 of many). 2002 Pomacentrus beauforti Fowler and Bean, 1928: 115, Amblyglyphidodon melanopterus Allen and Randall, Pl. 13. (Type locality, Catbalogan, Samar Island, 2002: 146, Figs. 5–7. (Type locality, East side of Philippines). Holotype: USNM 89962. Paratypes: Monuate Island, Tongatapu, Tonga). Holotype: (109) ANSP 80291 (1), 80318 (4); BPBM 10558 [ex BPBM 37967. Paratypes: AMS I.41006-001 (1); ANSP 80318 and USNM 97911] (1); USNM 97208- BPBM 37926 (1); CAS 214470 (1); MNHN 2001- 12 (1, 4, 4, 98, 3). 2673 (1); NSMT-P 61777 (1); ROM 72658 (1); Distribution: Philippines, New Guinea, New Britain, USNM 337888 (23), 337939 (2). Solomon Islands, and Northeastern Queensland. Distribution: Tonga. Amblypomacentrus clarus Allen and Adrim, 2000 Amblyglyphidodon orbicularis (Hombron and Amblypomacentrus clarus Allen and Adrim, 2000: Jacquinot, 1853) 51, Fig. 1. (Type locality, Banggai Harbor, Banggai Glyphisodon orbicularis Hombron and Jacquinot in Island, Banggai Islands, Sulawesi, Indonesia, Jacquinot and Guichenot, 1853: 52, Pl. 5 (Fig. 6). 1°34.84'S, 123°20.01'E, depth 18 meters). Holotype: (No locality; probably Fiji or Samoa). Holotype: NCIP 6186. Paratypes: WAM P.31497-002 (3). MNHN 0000-8535 [or 8335] (poor condition). Distribution: Cambodia, Indonesia and Philippines; Distribution: Samoa, Fiji, Tonga and New Caledonia. probably more widespread. Amblyglyphidodon silolona Allen, Erdmann and Amblypomacentrus vietnamicus Prokofiev, 2004 Drew, 2012 Amblypomacentrus vietnamicus Prokofiev, 2004: Amblyglyphidodon silolona Allen, Erdmann and 121, Figs. 1–2. (Type locality, Eastward of N'achang Drew in Allen and Erdmann, 2012: 1130. (Type City, Che Island, off Vietnam, South China Sea). locality, North Button Island, 12°18.560'N, Holotype: ZIN 53059. Paratypes: ZIN 53060 (1). 93°04.070'E, Andaman Islands, depth 6 meters). Distribution: South China Sea. Holotype: WAM P.33253-006. Paratypes: WAM P.33230092 (7). Genus Amphiprion Bloch and Schneider, 1801 Distribution: Northeastern Indian Ocean: Andaman Amphiprion Bloch and Schneider, 1801: 200. Islands. Masculine. Type species Lutjanus ephippium Bloch, 1790. Type by subsequent designation. Type designated Amblyglyphidodon ternatensis (Bleeker, 1853) by Griffith (1834: 160). Glyphisodon ternatensis Bleeker, 1853c: 137. (Type Prochilus Bleeker (ex Klein), 1864: 185. Masculine. locality, Ternate, Moluccas, Indonesia). Syntypes: Type species Lutjanus ephippium Bloch, 1790. Type by RMNH. subsequent designation. Type designated by Jordan Glyphisodon schlegelii Bleeker, 1853c: 138. (Type (1917: 39) validates Prochilus Klein, 1778, not locality, Ternate, Molucca Islands, Indonesia). available. Objectively invalid; preoccupied by Prochilus Holotype: RMNH 6490 (1 of 3). Bleeker specimens: Illiger, 1811 in Mammalia, Prochilus Cuvier, 1816 in BMNH 1862.2.28.27 (1). fishes, and Prochilus Brullé, 1835 in Orthoptera. Glyphidodon nigrifrons MacLeay, 1883: 271. (Type Actinicola (subgenus of Amphiprion) Fowler, 1904: locality, Southeastern coast of New Guinea). Holotype: 533. Masculine. Type species Lutjanus percula AMS I.9195 (1). Lacepède, 1802. Type by original designation. Abudefduf philippinus Fowler, 1918: 54, Fig. 21. Phalerebus Whitley, 1929: 216. Masculine. Type (Type locality, Philippines). Holotype: ANSP 47531. species Amphiprion akallopisos Bleeker, 1853. Type Paratypes: ANSP 47532 (1). by original designation. Distribution: Indo-Australian Archipelago. Paramphiprion Wang, 1941: 89. Masculine. Type Genus Amblypomacentrus Bleeker, 1877 species Paramphiprion hainanensis Wang, 1941. Amblypomacentrus Bleeker, 1877: 68. Masculine. Type by original designation. Type species Glyphisodon breviceps Schlegel and Twenty-nine species (Allen, 1991; Allen et al., 2008).

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Comments: based on type species Prochilus Bleeker, Amphiprion chrysogaster Cuvier, 1830 1864 is a synonym of Amphiprion Bloch and Amphiprion chrysogaster Cuvier in Cuvier and Schneider, 1801 and not of Premnas Cuvier, 1816 as Valenciennes, 1830: 400. (Type locality, Mauritius; recorded in the Catalog of Fishes (Fricke et al., 2020). Réunion Island). Syntypes: MNHN 8506 (1) Mauritius, 8507 (1) Réunion. Amphiprion akallopisos Bleeker, 1853 Amphiprion fusciventris Bennett, 1832: 165. (Type Amphiprion akallopisos Bleeker, 1853d: 281. (Type locality, Mauritius, Mascarenes, southwestern Indian locality, Priaman, Indonesia). Holotype: RMNH Ocean). Holotype: BMNH 1856.2.15.10. 6446 (1 of 9). Amphiprion mauritiensis Schultz, 1953: 196, Pl. 9 Distribution: Western Indian Ocean to Java Sea (not (Fig. I). (Type locality, Mauritius, Mascarenes, recorded from Maldives and Sri Lanka). Southwestern Indian Ocean). Holotype: MCZ 6093. Amphiprion akindynos Allen, 1972 Paratypes: MCZ 5800 (now 1), 5801 (2), 5802 (2), Amphiprion akindynos Allen, 1972: 153, Figs. 76–80. 46313 [ex MCZ 6093] (?2); USNM 153679 [ex MCZ (Type locality, South reef face about 1 mi. offshore, 5800] (1), 153680 [ex MCZ 6093] (2). One Tree Island, Queensland, Australia, 0–15 ft). Distribution: Madagascar; Réunion, Mauritius and Holotype: USNM 204287. Paratypes: AMS IA.2626 Rodrigues (Mascarenes). (1), IA.4063–64 (1, 1), IA.6802 (3), IB.1402 (1), Amphiprion chrysopterus Cuvier, 1830 IB.2367–68 (1, 1), IB.3875–78 (1, 1, 1, 1), IB.5105 Amphiprion chrysopterus Cuvier in Cuvier and (1), IB.5108 (1), IB.6054 (1), IB.6059 (1); ANSP Valenciennes, 1830: 401. (No locality). Holotype: 93228 (1); BPBM 11100 (1); USNM 204288-93 (9). MNHN (not located). Distribution: Great Barrier Reef, New Caledonia and Distribution: Coral Sea to Marshalls East to Society Loyalty Islands. and Tuamotus. Amphiprion allardi Klausewitz, 1970 Amphiprion clarkii (Bennett, 1830) Amphiprion allardi Klausewitz, 1970: 182, Figs. 1–6. Anthias clarkii Bennett, 1830: unnumbered page, Pl. (Type locality, Bamburi near Mombasa, East African 29. (Type locality, South coast of Sri Lanka). No coast). Holotype: USNM 204239. Paratypes: BPBM types known. Spelled clarckii by Valenciennes in 9421–24 (2, 13, 2, 9); BMNH 1861.5.2.10 (1), Cuvier and Valenciennes (1833: 504). Lectotype 1902.1.16.90 (1), 1913.4.7.94-95 (2), 1938.12.6.12 designated by Fricke (1999: 369) as the specimen (1), 1960.11.15.66-67 (2), 1968.5.30.121 (1); MCZH figured in Bennett's Pl. 29. 4489 (3); SMF 10290–96 (3, 5, 41 or 42, 1, 2, 1, 30), Amphiprion xanthurus Cuvier in Cuvier and 10297-98 (2 skeletons); 10311 (38 or 39), 10327 (2 Valenciennes, 1830: 402. (Type locality, Mauritius, or 3), 10328 (4); USNM 204240–44 (1, 14, 20, 1, 1). Mascarenes, Southwestern Indian Ocean). Syntypes: Distribution: Western Indian Ocean from Kenya to Durban. MNHN 0000-8106 (1), 0000-8108 (1). Amphiprion barberi Allen, Drew and Kaufman, Sparus milii Bory de Saint-Vincent, 1830: 130. (Type 2008 locality, Shark Bay, Western Australia). No types Amphiprion barberi Allen, Drew and Kaufman, known. Figured in v. 17, 1831: Pl. 113 (Fig. 2). 2008: 107, Figs. 1–4. (Type locality, Namena Island, Amphiprion melanostolus Richardson, 1842: 390. approximately 17°07'S, 179°04'E, Lomaiviti Group, (Type locality, Depuch Island, Western Australia). Fiji, depth 5–12 meters). Holotype: WAM P.32261- No types known. 001. Paratypes: BPBM 10808 (4); USNM 204274 Amphiprion japonicus Temminck and Schlegel, 1842: (5), 204275, 238890 (20), 334855 (2), 335157 (6), 66. (Type locality, Japan). Holotype: RMNH 873. 338150 (2); WAM P-32332-002 (6). Amphiprion chrysargyrus Richardson, 1846: 254. Distribution: Fiji, Tonga and Samoa. (Type locality, China). Holotype (?): BMNH 2004.11.5.5. Amphiprion bicinctus Rüppell, 1830 Amphiprion boholensis Cartier, 1874: 96. (Type locality, Amphiprion bicinctus Rüppell, 1830: 139, Pl. 35 Bohol, Philippines). Holotype: Whereabouts unknown. (Fig. 1). (Type locality, El Tûr, Sinai coast, Egypt, Amphiprion papuensis MacLeay, 1883: 271. (Type Gulf of Suez or Massawa, Eritrea, Red Sea). locality, D'Entrecasteaux Group, New Guinea). Lectotype: SMF 1460. Paralectotypes: ?BMNH Holotype: AMS I.9240. 1845.10.29.89, ?1860.11.9.78-79; SMF 2716 (1), Amphiprion snyderi Ishikawa, 1904: 11, Pl. 5. (Type 4934 (1), 17518 (1, skeleton). Lectotype established locality, Ogasawara-jima, Bonin Islands, Japan). by Dor, 1984: 180. Holotype: Whereabouts unknown. Distribution: Red Sea and Chagos Archipelago. Distribution: North Indian Ocean to Melanesia and Amphiprion chagosensis Allen, 1972 Micronesia (except Marshalls), North to Southern Japan. Amphiprion chagosensis Allen, 1972: 161, Figs. 81– Comments: as reported by Heemstra et al. 2004, 82. (Type locality, Diego Garcia Atoll, Chagos Fricke (1999) was incorrect in listing Amphiprion Archipelago, Indian Ocean, 2–8 ft). Holotype: USNM xanthurus Cuvier, 1830 in synonymy with A. allardi 204301. Paratypes: USNM 204302–04 (2, 1, 2). as this latter species is not known from the Distribution: Chagos Archipelago. Mascarenes.

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 45 Parenti 46

Amphiprion ephippium (Bloch, 1790) Distribution: Lord Howe and Norfolk Islands. Lutjanus ephippium Bloch, 1790: 121, Pl. 250 (Fig. Amphiprion melanopus Bleeker, 1852 2). (Type locality, Tranquebar, India). Syntypes: Amphiprion melanopus Bleeker, 1852: 561. (Type ZMB 2766 (1). On page 98 of Bloch's Ichthyologie locality, Ambon Island, Moluccas Islands, Indonesia). (Bloch, 1797). Holotype: RMNH 6444 (probably the largest of 5). Amphiprion calliops Schultz, 1966: 72, unnumb. Fig. Amphiprion monofasciatus Thiollière in Montrouzier, (Type locality, Indian Ocean but exact locality 1857: 476 [198]. (Type locality, Woodlark Island, unknown). Holotype: USNM 197554. Paratypes: South Pacific). No types known. USNM 82781 (1), 231708 [ex 236481-F1] (6). Prochilus macrostomus Bleeker, 1877a: 136. (Type Distribution: Andaman and Nicobar Islands to locality, Ambon Island, Molucca Islands, Indonesia). Moluccas. Holotype: Whereabouts unknown. Figured in Bleeker Amphiprion frenatus Brevoort, 1856 (1877b: Pl. 401, Fig. 5). Amphiprion frenatus Brevoort, 1856: 263 [11], Pl. 6 Amphiprion arion De Vis, 1884: 450. (Type locality, (fig. 4). (Type locality, Naha, Okinawa Island, South Sea). Holotype: QM I.2688. Ryukyu Islands, Japan). Neotype: USNM 71702. Amphiprion verweyi Whitley, 1933: 85. (Type Based on a drawing. Neotype listed in USNM locality, Northwest Islet, Capricorn Group, database, but location of designation not found. Queensland, Australia). Holotype: AMS IA.5063. Prochilus polylepis Bleeker, 1877a: 135. (Type Paratypes: AMS IA.5064–65 (1, 5). locality, Ambon Island, Molucca Islands, Indonesia; Distribution: Bali to Queensland and New Caledonia, southeastern New Guinea). Syntypes: (3). Figured in Micronesia, Fiji, Samoa, Tonga, Society Islands. Bleeker (1877b: Pl. 401, fig. 6). Amphiprion nigripes Regan, 1908 Distribution: Singapore, Malaysia, Brunei, Thailand, Amphiprion nigripes Regan, 1908: 230, Pl. 24 (Fig. Vietnam, western Indonesia, Philippines, China, 2). (Type locality, Maldives, Indian Ocean). Taiwan and Southern Japan. Holotype: BMNH 1908.3.23.101. Amphiprion fuscocaudatus Allen, 1972 Distribution: Maldives and Sri Lanka. Amphiprion fuscocaudatus Allen, 1972: 134, Figs. Amphiprion ocellaris Cuvier, 1830 64–65. (Type locality, Beacon Island, off Port Amphiprion ocellaris Cuvier in Cuvier and Victoria, Seychelles Islands, 40–50 ft). Holotype: Valenciennes, 1830: 399. (Type locality, Sumatra, ANSP 109813. Paratypes: ANSP 109814 (1), BPBM Indonesia). Holotype: BMNH. 10841 [not 10491] (1). Amphiprion melanurus Cuvier in Cuvier and Distribution: Seychelles and Aldabra. Valenciennes, 1830: 400. (Type locality, Sumatra, Amphiprion latezonatus Waite, 1900 Indonesia). Holotype: BMNH 2011.5.17.4. Amphiprion latezonatus Waite, 1900: 201, Pl. 34. Amphiprion bicolor Castelnau, 1873: 92. (Type (Type locality, Lord Howe Island). Holotype: AMS locality, Darwin, Northern Territory, Australia). I.4349. Holotype: MNHN A-0705. Distribution: Eastern Australia, New Caledonia, Lord Distribution: Andaman Islands to Ryukyu Islands. Howe and Norfolk Islands. Amphiprion omanensis Allen and Mee, 1991 Amphiprion latifasciatus Allen, 1972 Amphiprion omanensis Allen and Mee in Allen, Amphiprion latifasciatus Allen, 1972: 138, Figs. 66– 1991: 47, 220, Fig. bottom left. (Type locality, Ras 67. (Type locality, Ambariobe Bay, Nosy Be, Kanasa, Barr Al Hikman, Oman, Arabian Sea, 5 m). Madagascar). Holotype: USNM 204299. Paratypes: Holotype: CAS 73147. Paratypes: BPBM 33955–56 USNM 204300 (7). (1, 1); CAS 73148 (2), 73164 (1); WAM P.30108- Distribution: Madagascar. 001 (1), P.30109-001 (2), P.30110-001 (1). Distribution: Arabian Sea coast of Oman and Socotra. Amphiprion leucokranos Allen, 1973 Amphiprion leucokranos Allen, 1973b: 319, Figs. 1, Amphiprion pacificus Allen, Drew and Fenner, 2010 2a, 3. (Type locality, outer reef front of the n. end of Amphiprion pacificus Allen, Drew and Fenner, 2010: Kranket Island, Madang, New Guinea, 5°11'24"S, 131, Figs. 1–2. (Type locality, Wallis Island, Ile Uvea, 145°50'54"E, 7 m). Holotype: USNM 207358. West side of outer reef, North-west of Point Paratypes: AMS I.16483-001 (2), I.16484-001 (1); Vaha'a'Utu, 13°16'50"S, 176°15'55"W, depth 6–12 USNM 207359 (2). meters). Holotype: USNM 373925. Paratypes: USNM Distribution: Northeastern Melanesia: West Papua 393941 (1), 393942 [as 39342] (1), 393561 (1). and Papua New Guinea to Solomon Islands. Distribution: Papua New Guinea, Fiji, Tonga, Samoa, Wallis Island. Amphiprion mccullochi Whitley, 1929 Amphiprion mccullochi Whitley, 1929: 213. (Type Amphiprion percula (Lacepède, 1802) locality, Lord Howe Island). Holotype: AMS Perca tysonis Walbaum, 1792: 351. (Type locality, IA.1962. Paratypes: AMS I.5392 (2, now 1), I.5729– South seas). No types known. Appeared without a 30 (1, 1), IA.1963–67 (1, 1, 1, 1, 1). species name in Tyson (1772: 247) in the genus Perca.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Lutjanus percula Lacepède, 1802: 194, 239. (Type Amphiprion trifasciatus Cuvier in Cuvier and locality, Port Praslin, New Britain Island, Bismarck Valenciennes, 1830: 395. (Type locality, Molucca Archipelago). Holotype: Whereabouts unknown. Islands). Holotype: MNHN. Based on Commerson manuscript. Amphiprion intermedius Schlegel and Müller, 1840: Amphiprion tunicatus Cuvier in Cuvier and 18. (Type locality, Molucca Islands, Indonesia). No Valenciennes, 1830: 399, Pl. 132 (Fig. 2). (Type types known. locality, Vanikoro Island, Santa Cruz Islands, Coracinus vittatus Gronow in Gray, 1854: 57. (Type southwestern Pacific). Syntypes: MNHN 0000-7992 locality, Indian Ocean and Ambon Island, Molucca (2). Islands, Indonesia). Syntypes: BMNH 1853.11.12.33. Distribution: New Guinea East to Solomon Islands Amphiprion bifasciatus annamensis Chevey, 1932: and Vanuatu, South to Queensland (Australia) and 99, Pl. 39. (Type locality, South coast of Vietnam). New Caledonia. Syntypes: IOI unregistered. Comments: Amphiprion tunicatus is not a Paramphiprion hainanensis Wang, 1941: 89. (Type replacement name for Amphiprion percula Cuvier locality, Hainan Island, China). Holotype: ? SSCN. 1830 as stated in the ECoF (Fricke et al., 2020). Distribution: Ryukyu Islands to Gulf of Thailand, Also, the ECoF reported “Valid as Amphiprion Northern Australia, New Caledonia and Solomon Islands. tunicatus Cuvier 1830 (Allen, 1991: 237)”, but Allen Comments: Sciaena unimaculata Meuschen, 1781 listed Amphiprion tunicatus as a synonym of A. reported in the Index to the Zoophylacium percula. Moreover the same records misspelled Gronovianum of Gronovius is not available, as it is percula as perculata. Perca tysonis is nomen oblitum published in a rejected work (Opinion 261). Appeared to permit continued use of percula as a nomen as Coracinus seu Sciaena. Placed in synonymy with protectum (Parenti, 2003: 497). Amphiprion polymnus (Linnaeus 1758) by Allen (1991: 237). Amphiprion perideraion Bleeker, 1855 Amphiprion perideraion Bleeker, 1855b: 437. (Type Amphiprion rubrocinctus Richardson, 1842 locality, Obij major Island, Indonesia). Holotype: Amphiprion rubrocinctus Richardson, 1842: 391. RMNH 6445 (smallest of 4 or 5). Bleeker specimens: (Type locality, Broome, Western Australia). Neotype: NMV 45916 (1). WAM P.4474-001. Originally as Amphiprion? Amphiprion rosenbergii Bleeker, 1859: 16. (Type rubrocinctus. Neotype designated by Allen (1972) as locality, Doreh Bay, off Manokwari, West Papua, Richardson's description was based on a drawing and Indonesia, New Guinea, ca. 0°53'25''S 134°05'29''E). the specimen lost. Holotype: Whereabouts unknown. Amphiprion tricolor Günther, 1862: 8. (Type Amphiprion amamiensis Mori, 1966: 469, Fig. 1. locality, Port Essington, Northern Territory; South (Type locality, Japan). Holotype: FRLM 6196. Australia). Syntypes: (9) BMNH 1844.1.13.? (1), Distribution: Micronesia, Southern Japan to New 1847.7.21.56 (1), 1855.9.19.564-567 (7). Caledonia West to Christmas Island East to Amphiprion ruppelii Castelnau, 1873: 91. (Type Tuamotus. locality, Darwin, Northern Territory, Australia). Holotype: MNHN A-0706. Amphiprion polymnus (Linnaeus, 1758) Distribution: Western Australia and Northern Perca polymna Linnaeus, 1758: 291. (Type locality, Territory; previous records from Tonga and Samoa are "Indis"). Holotype: ZMUU Linn. Coll. 198. misidentifications. Anthias bifasciatus Bloch, 1792: 108, Pl. 316 (Fig. 2). (Type locality, Ambon Island, Molucca Islands, Amphiprion sandaracinos Allen, 1972 Indonesia). Lectotype: ZMB 2768. Paralectotypes: ZMB Amphiprion sandaracinos Allen, 1972: 81, Figs. 29– (1, lost). Lectotype selected by Paepke (1999: 119). 30. (Type locality, Pagbilao Island, Luzon Island, Sciaena jordain Walbaum, 1792: 311. (Type locality, Philippines). Holotype: USNM 130. Paratypes: BPBM Ambon Island, Molucca Islands, Indonesia). No 6804 (1), 10617 (3); USNM 160663–64 (2, 2). types known. Distribution: Ryukyu Islands. to New Guinea and Solomon Perca bifasciata Walbaum, 1792: 352. (No locality Islands.; Western Australia and Christmas Island. stated). No types known. Amphiprion sebae Bleeker, 1853 Lutjanus jourdin Lacepède, 1802: 191, 235. (Type Amphiprion sebae Bleeker, 1853e: 478. (Type locality, Ambon Island, Molucca Islands, Indonesia). locality, Jakarta, Java, Indonesia). Syntypes (9, not No types known. found). Bleeker specimens: AMS B.8268 [from Day] Amphiprion laticlavius Cuvier in Cuvier and (1) Malay, NMV 45937 (1). Valenciennes, 1830: 394, Pl. 132 (Fig. 1). (Type Distribution: North Indian Ocean to Java. locality, New Guinea). Syntypes: MNHN 0000-7959 (2), 0000-8107 (2); SMF 1594. Amphiprion thiellei Burgess, 1981 Premnas leucodesmus Cuvier (ex Kuhl and van Amphiprion thiellei Burgess, 1981: 69, Figs. (Type Hasselt) in Cuvier and Valenciennes, 1830: 409. locality, In the vicinity of Cebu, Philippines). Holotype: (Type locality, Java, Indonesia). No types known. USNM 228445. Paratypes: WAM uncat. (1, missing).

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 47 Parenti 48

Distribution: Western Central Pacific: described from Furcaria Poey, 1860: 194. Feminine. Type species two aquarium dealer specimens believed to have Furcaria puncta Poey, 1860. Type by subsequent originated in the vicinity of Cebu, Philippines; habitat designation. Earliest type designation by Jordan and and true distribution unknown. Evermann (1898: 1545). Objectively invalid; preoccupied by Furcaria Lesson 1838 in Aves, Amphiprion tricinctus Schultz and Welander, 1953 replaced by Demoisellea Whitley 1928. Amphiprion tricinctus Schultz and Welander in Ayresia Cooper, 1863: 73. Feminine. Type species Schultz, 1953: 195, Pl. 9 (Fig. H). (Type locality, Ayresia punctipinnis Cooper, 1863. Type by monotypy. Lagoon, Amen Island, Bikini Atoll, Marshall Islands, Onychognathus Troschel, 1866: 231. Masculine. Pacific). Holotype: USNM 152929. Type species Onychognathus cautus Troschel, 1866. Distribution: Marshall Islands. Type by monotypy. Objectively invalid; preoccupied Genus Azurina Jordan and McGregor, 1898 by Onychognathus Hartlaub, 1859 in Aves, replaced Azurina Jordan and McGregor in Jordan and by Agripopa Whitley 1928. Evermann, 1898: 1544. Feminine. Type species Hoplochromis (subgenus of Chromis) Fowler, 1918: Azurina hirundo Jordan and McGregor, 1898. Type 66. Masculine. Type species Heliases caeruleus by original designation. Cuvier, 1830. Type by original designation. Belochromis Fowler, 1944a: 333. Masculine. Type Agripopa Whitley, 1928b: 296. Feminine. Type species Azurina eupalama Heller and Snodgrass, species Onychognathus cautus Troschel, 1866. Type 1903. Type by original designation. by being a replacement name. Replacement for Eastern Pacific; two species. Onychognathus Troschel 1866, preoccupied by Onychognathus Hartlaub, 1859 in Aves. Azurina eupalama Heller and Snodgrass, 1903 Demoisellea Whitley, 1928b: 295. Feminine. Type Azurina eupalama Heller and Snodgrass, 1903: 198, species Furcaria puncta Poey, 1860. Type by being a Pl. 5. (Type locality, Hood Island, Galápagos Islands). replacement name. Replacement for Furcaria Poey, Holotype: SU 6352. Paratypes: SU 6506 (7), 6538 (9), 1860, preoccupied by Furcaria Lesson 1838 in Aves. 2299–2300 (1, 1, both missing). Dorychromis Fowler and Bean, 1928 (subgenus of Distribution: Galapagos Islands. Heliases) Fowler and Bean, 1928: 31, 60. Masculine. Azurina hirundo Jordan and McGregor, 1898 Type species Heliases analis Cuvier 1830. Type by Azurina hirundo Jordan and McGregor in Jordan and original designation. Evermann, 1898: 1544. (Type locality, Guadelupe Island, Lepidochromis (subgenus of Chromis) Fowler and off Baja California. Lectotype: SU 5706. Paralectotypes: Bean, 1928: 31, 58. Masculine. Type species Chromis (2) BMNH 1898.10.29.104 (1). Lectotype established in lepidolepis Bleeker, 1876. Type by original designation. Jordan and McGregor (1899: 282). Pycnochromis Fowler, 1941: 258. Masculine. Type Distribution: Baja California, Guadalupe and species Pycnochromis vanderbilti Fowler, 1941. Revillagigedo Archipelago. Type by original designation. Thrissochromis Fowler, 1941: 262. Masculine. Type Genus Chromis Cuvier, 1814 species Chromis velox Jenkins, 1901. Type by Chromis Cuvier, 1814: 88. Feminine. Type species original designation. Sparus chromis Linnaeus, 1758. Type by original Serrichromis Fowler, 1943: 77. Masculine. Type species designation. Type designated in subtitle (also by Dascyllus pomacentroides Kendall and Goldsborough, absolute tautonymy). Gender ruled feminine by ICZN 1911. Type by original designation. and name placed on Official List (Opinion 1417). Lepicephalochromis Fowler, 1943: 78. Masculine. Chromis Oken (ex Cuvier), 1817: 1182. Feminine. Type species Chromis cupreus Fowler and Bean Type species Sparus chromis Linnaeus, 1758. Type 1923. Type by original designation. by absolute tautonymy. Based on "Les Chromes" of Siphonochromis Fowler, 1946: 143. Masculine. Type Cuvier, 1816: 266. Objectively invalid; preoccupied species Siphonochromis lepidostethicus Fowler 1946. by Chromis Cuvier, 1814 in fishes. Type by original designation. Heliases Cuvier in Cuvier and Valenciennes, 1830: The genus has not been revised; 109 species are 493. Masculine. Type species Heliases insolatus presently listed here. Cuvier, 1830. Type by subsequent designation. Based on "Les Héliases" Cuvier 1829: 160; Latinized first Chromis abrupta Randall, 2001 as above, with six included species. Type designated Chromis sp. 3 Allen, 1991: 88, Fig. (bottom). by Valenciennes in d’Orbigny (1845: 503). (Marquesas Islands). Heliasus Griffith, 1834: 161. (Unjustified emendation). Chromis abrupta Randall, 2001: 93, Figs. 1–2. (Type Heliastes Lowe, 1838: 177. Masculine. Type species locality, Hana Hevane Bay, Tahuata, Marquesas Heliases insolatus Cuvier, 1830. Type by being an Islands). Holotype: BPBM 10388. Paratypes: AMS unjustified emendation. Unjustified emendation of I.39934-001 (1); BPBM 10351 (7), 11709 (2), 12596 Heliases Cuvier, 1815. (4), 38529 (3); CAS 210416 (1); MNHN 2000-6 (2); Heliazes Valenciennes, 1840: Pl. 33 (Fig. 2). (Unjustified NSMT-P57704 (1); USNM 359630 (1). emendation). Distribution: Marquesas Islands.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Chromis abyssicola Allen and Randall, 1985 Holotype: NCIP 6350. Paratypes: NCIP 6351 (1); Chromis abyssicola Allen and Randall, 1985: 242, Fig. USNM 391796 (2); WAM P.33101-005 (5). 1. (Type locality, Norfolk Island Ridge, ca. 520 km Distribution: Eastern Indian Ocean and Western Pacific: Northwest of n. tip of North Island, New Zealand, Indonesia, Southern Korea, and Southern Japan. 32°28'12"S, 167°28'06"E, 115–122 m). Holotype: QM Chromis albomaculata Kamohara, 1960 I.21398. Paratypes: AMS E.2905 (6), I.23685-007 (1), Chromis albomaculatus Kamohara, 1960: 3, Fig. 1. (Type I.24987-001 (2); BMNH 1985.4.30.1 (1); BPBM locality, Kashiwajima, Kochi Pref., Japan). Holotype: 30352 (2); QM I.13415 (6), I.13418 (5); USNM BSKU 7487. Paratypes: BSKU 8514 [not 8541] (1). 270463 (2); WAM P.28421-001 (2). Distribution: Izu Islands to Okinawa. Distribution: Off Southeastern Australia, Kermadec Ridge and Norfolk Ridge. Chromis alleni Randall, Ida and Moyer, 1981 Chromis alleni Randall, Ida and Moyer 1981: (210) Chromis abyssus Pyle, Earle and Greene, 2008 215, Pl. 1 (Fig. A); Fig. 2H. (Type locality, W. side of Chromis abyssus Pyle, Earle and Greene, 2008: 6, Sesoko Island, Okinawa Island, Ryukyu Islands, Japan, Figs. 1a–1c. (Type locality, off Ngemelis Island, 15 m). Holotype: BPBM 19092. Paratypes: BMNH below and slightly North of Blue Hole caverns, 1979.7.17.1 (1); BPBM 19093 (1), 19119 (1), 19142 7°8'16.49"N, 134°13'18.5"E, Palau Islands, depth (1), 22310 (1), 22618 (1); FSKU 760810 (7); NTUM 110 meters). Holotype: BPBM 40861. Paratypes: 4927 (2); TMBS 741230-A (1), 750700 (1), 751001-B BMNH 2007.10.31.1 (1), BPBM 40855 (3), CAS (1); USNM 220557 (1), WAM P.25516-001 (5). 225755 (1), MNHN 2007-1922 (1), USNM 391136 Distribution: Miyake-jima, Bonin Islands of Japan (2), WAM P.32898-001 (1). and South to Taiwan. Distribution: Palau, Indonesia, and Vanuatu. Chromis alpha Randall, 1988 Chromis acares Randall and Swerdloff, 1973 Chromis alpha Randall, 1988a: 74, Fig. 1. (Type Chromis acares Randall and Swerdloff, 1973: 331, locality, outside reef off W. side of Rimatuu Islet, Figs. 2–3. (Type locality, off tanker buoys at harbor Tetiaroa Atoll, Society Islands, 45 m). Holotype: entrance, Rarotonga, Cook Islands, 20 m). Holotype: BPBM 14993. Paratypes: AMS I.16650-001 (1), BPBM 13823. Paratypes: AMS I.16766-001 (1), I.17474-003 (1), I.19476-021 (2), I.22580-012 (1); I.16772-001 (1); BMNH 1972.12.5.4 (1); BPBM ANSP 134269 (1); BPBM 6898 (4), 11544 (2), 14702 8752 (3), 11049 (14), 11612 (2), 11298 (1), 12172 (2), 14999 (1), 16083 (2), 17506 (1), 18388 (1), 18467 (1), 12296 (1), 13686 (1), 13817–18 (10, 3), 13820 (3), 19979 (1), 30938 (14), 30951 (2), 31159 (2); (1), 13970 (1), 14059 (2), 14659 (1); CAS 15912 (1); MNHN 1976-129 (1); USNM 209903 (1), 223922 (4), MNHN 1972-96 (1); USNM 208430 (1). 273546 (1); WAM PP24939 (5), 25283-001 (1), Distribution: Marianas, Marshalls and Hawaiian 27826-025 (3), 28165-007 (1). Islands; New Caledonia, Rotuma, Samoa, Cook Islands, Distribution: Christmas and Cocos-Keeling islands Austral Islands, Line Islands and Pitcairn Islands. and Indonesia East to Caroline and Tuamotu Islands, Chromis agilis Smith, 1960 North to Yaeyama Islands, South to Northwestern Chromis agilis Smith, 1960: 324, Pls. 26 (Fig. A), 32 Australia and New Caledonia. (Fig. J). (Type locality, Astove Island, W. Indian Chromis alta Greenfield and Woods, 1980 Ocean). Holotype: RUSI 292. Paratypes: RUSI Chromis alta Greenfield and Woods, 1980: 630, Figs. 2337–44 (1, 1, 1, 1, 2, 2, 2, 11). 2–3, 6. (Type locality, Bahía Santa Inez, Gulf of Chromis xutha Randall, 1988b: 54, Figs. 5–6. (Type California, Baja California, Mexico, 72–81 m). locality, Reef front, E. side of Villingili Island, North Holotype: SIO 65-305. Paratypes: BMNH 1978.11.7.1- Malé Atoll, Maldives, Indian Ocean, 8–12 m). 2 (2); BPBM 22428 (2); CAS 39276 (1), 39295 (5), Holotype: BPBM 18864. Paratypes: BMNH 39407 (7); FMNH 75777 (1), 89557 (13); SIO 77-193 1988.6.2.3-4 (2); BPBM 16395 (1), 27278 (4), 28441 (1), 78-296 (2); UA 71-79 (3), 71-88 (1), 75-120 (1); (15), 32704–05 (7, 5); CAS 33135 (15); ROM 54940 USNM 219028 (2). (10); RUSI 27721 (2); USNM 296243 (2); WAM Distribution: Western Baja California and Galapagos P.26517-001 (3). Islands. One record from Cocos Island. Distribution: Western Indian Ocean: Maldives, Chagos Archipelago, Seychelles, Madagascar, Chromis amboinensis (Bleeker, 1871) Comoro Islands, and along the East African coast Heliases amboinensis Bleeker, 1871: 111. (Type between Shimoni, Kenya and Bazaruto, locality, Ambon Island, Moluccas Islands, Indonesia). Mozambique. Holotype: RMNH 6505. Glyphidodon bimaculatus MacLeay, 1883: 271. (Type Chromis albicauda Allen and Erdmann, 2009 locality, Southeastern coast of New Guinea). Chromis albicauda Allen and Erdmann, 2009a: 123, Holotype: AMS I.9236. Figs. 1–2, 4–5. (Type locality, Crystal Bay, Chromis fragoris Whitley, 1964: 185, Fig. 13. (Type 8°42.887'S, 115°27.331'E, Nusa Penida [Penida locality, Lihou Atoll, Coral Sea). Holotype: AMS Island], off East coast of Bali, Indonesia [Badung IB.4935. Paratypes: AMS IB.4936–37 (1, 27), Strait, Eastern Indian Ocean], depth 30 meters). IB.4942–43 (1, 1).

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 49 Parenti 50

Distribution: West Pacific from Yaeyama Islands to Chromis atripes Fowler and Bean, 1928 Queensland east to Marshall Islands and Samoa; Chromis atripes Fowler and Bean, 1928: 43, Pl. 2. Christmas Island and Cocos Islands. (Type locality, Sablayan, Mindoro, Philippines). Holotype: USNM 89952. Paratypes: ANSP 85282 Chromis anadema Motomura, Nishiyama and (3), USNM (55 lots, ca. 176 spec.). Chiba, 2017 Distribution: Southern Japan to Australia West to Chromis anadema Motomura, Nishiyama and Chiba, Christmas Island East to Solomon Islands and New 2017: [3], Figs. 2, 3a [printed version 2018: 179] Caledonia. (Type locality, off west coast of Yoron-jima island, Amami Islands, Ryukyu Islands, Japan, 27°03'21"N, Chromis axillaris (Bennett, 1831) 128°24'39"E, depth 10 meters). Holotype: KAUM-I Heliases axillaris Bennett, 1831: 128. (Type locality, 42229. Paratypes: AMS 21646-008; BPBM 6897 (1), Mauritius). No type in BMNH, type probably lost. 11768 (3), 11566 (3), 16628; CAS 55110 (3), 55933 Distribution: Red Sea, Western Indian Ocean: East (2); KAUM–I. 46170 (1); MNHN 2008-0782 (1), Africa to Réunion and Mauritius (western Mascarenes), 2008-0946 (1); OMNH 32490 (1), USNM 390992 and Walters Shoals. Also recorded from New (1), 404653 (1). Caledonia. Distribution: Ryukyu Islands, Mariana Islands, Chromis bami Randall and McCosker, 1992 Marquesas Islands, Society Islands, Gambier Islands Chromis bami Randall and McCosker, 1992: 334, and Pitcairn Islands. Underwater photographs from Figs. 3–4. (Type locality, about 1 km north of Japan, Saipan, Palau, Ryukyu Islands and Tahiti. Duddie's Cocknuts, Bounty Bay, Pitcairn Island, Chromis analis (Cuvier, 1830) 30.5–36.5 m). Holotype: CAS 75992. Paratypes: Heliases analis Cuvier in Cuvier and Valenciennes, AMS I.31223-001 (2); BMNH 1991.5.20.2-3 (2); 1830: 496. (Type locality, Ambon Island, Moluccas BPBM 12782 (2), 13683 (1), 13695 (2), 16524 (5), Islands, Indonesia). Holotype: MNHN 8164. 16687 (9); CAS 63215 (7); NSMT-P 34680 (3); Heliases macrochir Bleeker, 1853g: 346. (Type USNM 316515 (3). locality, Ambon Island, Molucca Islands, Indonesia). Distribution: Tonga, Pitcairn, and the Austral islands. Holotype: ?RMNH (1 of 10). Chromis bermudae Nichols, 1920. Distribution: Southern Japan to Northern Great Chromis bermudae Nichols, 1920: 60. (Type locality, Barrier Reef, Solomon Islands, Fiji, and Loyalty Bermuda). Holotype: AMNH 7317. Paratypes: Islands. AMNH 7318 (3). Chromis athena Allen and Erdmann, 2008 Distribution: Bermuda endemic. Chromis athena Allen and Erdmann, 2008b: 204, Figs. Comments: Previously regarded as a synonym of the 1–2. (Type locality, Keruo Channel, 00°35.291'S, Brazilian C. flavicauda, C. bermudae has been 130°17.542'E, Fam Group, Raja Ampat Islands, Papua resurrected on the basis of differences in color Barat Province, Indonesia, depth 62 meters). Holotype: pattern and a widely disjunct distribution (Smith- NCIP 6337. Paratypes: NCIP 6346 (1); USNM Vaniz and Collette, 2013: 179). 391635 (1); WAM P.33012-001 (2). Chromis bowesi Arango, Pinheiro, Rocha, Greene, Distribution: West Papua (Indonesia) and Papua New Pyle, Copus, Shepherd and Rocha, 2019 Guinea. Chromis bowesi Arango, Pinheiro, Rocha, Greene, Chromis atrilobata Gill, 1862 Pyle, Copus, Shepherd and Rocha, 2019: 9, Figs. 3a– Chromis (Furcaria) atrilobata Gill, 1862: 149. (Type c. (Type locality, Dive and Trek, Batangas, locality, Cape San Lucas, Baja California, Mexico). Philippines, 13°48'3.52"N, 120°54'38.88"E, depth Holotype: USNM 3675. 120 meters). Holotype: PNM 15359. Paratypes: Distribution: Gulf of California to Peru, including BPBM 41350 (1); CAS 242324 (1), 242278 (1); Galapagos Islands. USNM 440406 (1). Distribution: Verde Island Passage, Philippines to Chromis atripectoralis Welander and Schultz, 1951 Iou-jima Island, Southern Japan. Chromis atripectoralis Welander and Schultz, 1951: 107, Fig. 1. (Type locality, Channel reef, Eman Chromis brevirostris Pyle, Earle and Greene, 2008 Island, Bikini Atoll, Central Pacific). Holotype: Chromis brevirostris Pyle, Earle and Greene, 2008: 10, USNM 112397. Paratypes: BMNH 1952.5.8.1-6 [ex Figs. 2a–2c. (Type locality, Alet Islet, 7°21'15.44"N, USNM] (6); FMNH 44703 (orig. 20, now 14), 44704 149°10'47.03"E, Puluwat Atoll, Caroline Islands, (orig. 10, now 9), 44705 (2), 62624 (1); USNM (44 western Pacific, depth 100–103 meters). Holotype: lots, 166). BPBM 40804. Paratypes: BMNH 2007.10.31.2 (1); Distribution: Madagascar, Seychelles, Mascarenes and BPBM 37671 (1), 37713 (1), 39993 (1), 40422 (3), Maldives East to Tuamotu Archipelago and Gambier 40703 (2); CAS 225756 (2); MNHN 2007-1923 (1); islands, North to Ryukyu Islands, South to Western USNM 391137 (4); WAM P.32899-001 (2). Australia, Lord Howe Island, New Caledonia and Distribution: Western Pacific: Indonesia, Marshall Rapa; absent from the Hawaiian Islands. Islands to Fiji, Caroline Islands, and South to Vanuatu.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Chromis cadenati Whitley, 1951 Chromis insulindicus Bleeker, 1876: 390. (Type Chromis lineatus Cadenat, 1950: 669, Fig. (Type locality, Ambon Island, Molucca Islands, Indonesia). locality, Senegal). Syntypes: (19) MNHN 1950-0073 Holotype: RMNH 6514. [ex IFAN] (3). Objectively invalid; nomen Glyphisodon javanicus Van Hasselt in Bleeker, 1877c: praeoccupatum, preoccupied by Chromis lineatus 156. (Type locality, Java, Indonesia). No types known. Fowler and Bean, 1928. Distribution: Sri Lanka to northwestern Australia north Chromis cadenati Whitley, 1951: 66. Nomen novum. to Philippines. Replacement name for Chromis lineatus Cadenat Chromis circumaurea Pyle, Earle and Greene, 1950, preoccupied by Chromis lineatus Fowler and 2008 Bean, 1928. Chromis circumaurea Pyle, Earle and Greene, 2008: Distribution: Senegal to Ghana. 15, Figs. 3a–3c. (Type locality, Yap Island, Caroline Chromis caudalis Randall, 1988 Islands, Western Pacific, 9°26'3.41"N, 138°02'96"E). Chromis caudalis Randall, 1988a: 76, Fig. 2. (Type Holotype: BPBM 40836. Paratypes: BMNH locality, Alite Reef off Malaita, Solomon Islands, 2007.10.31.3 (1), CAS 225757 (1), MNHN 2007-1924 150 ft). Holotype: BPBM 15613. Paratypes: BPBM (1), USNM 391138 (1), WAM P.32900-001 (1). 9376 (2), ?14329 (1), 15637 (1), 19242 (2), 30948 Distribution: Western Pacific: Marshall Islands and (2), 30950 (4), 31299 (2); USNM 223920–21 (2, 4); Mariana Islands [seen from submersibles], Yap. WAM P.26080-003 (1), P.26099-012 (1). Chromis crusma (Valenciennes, 1833) Distribution: Christmas Island to Line Islands North Heliases crusma Valenciennes in Cuvier and to Philippines South to Papua New Guinea. Valenciennes, 1833: 510. (Type locality, Juan Chromis chromis (Linnaeus, 1758) Fernández Islands; Valparaíso, Chile). Syntypes: Sparus chromis Linnaeus, 1758: 280. (Type locality, MNHN 8161 (1) Juan Fernandez, 8162 (2) Valparaiso. Southern Europe). No types known. Chromis cupreus Fowler and Bean, 1923: 22. (Type Chromis castanea Cuvier, 1814: 88. (Type locality, locality, "Singapore, Maui, Fiji, or Samoa," though Coast of Provence, France, northwestern Mediterranean most likely from either of the last two named). Sea). No types known. Holotype: USNM 83108. Chromis mediterranea Cloquet, 1817: 147. Nomen Distribution: Cabo Blanco (Peru) to Talcahuano (Chile). novum. Unneeded replacement name for Sparus Chromis cyanea (Poey, 1860) chromis Linnaeus, 1758. No types known. Furcaria cyanea Poey, 1860: 196, Pls. 13 (Figs. 21, Chromis castaneus Schinz, 1822: 431. Nomen novum. 22), 14 (Fig. 4). (Type locality, Cuba). Holotype: MCZ Unneeded replacement name for and objective 14670. (Type locality, Poey specimens: MCZ 14667– synonym of Sparus chromis Linnaeus, 1758). No 68 (1, 1). types known. Distribution: Bermuda to Caribbean Sea; quoted also Chaetodon niger Nardo, 1824: 234. (Type locality, on Cape Verde Islands in the Eastern Atlantic (Quero Adriatic Sea, Mediterranean Sea). No types known. et al., 1990). Distribution: Mediterranean, Black Sea, Sea of Azov and off coast of Morocco to Lisbon south to Gulf of Chromis dasygenys (Fowler, 1935) Guinea and possibly Angola, including offshore islands. Abudefduf dasygenys Fowler, 1935: 400, Fig. 34. (Type locality, Durban, Natal, South Africa). Chromis chrysura (Bliss, 1883) Holotype: ANSP 63934. Heliastes chrysurus Bliss, 1883: 56. (Type locality, Distribution: Delagoa Bay to Durban (South Africa). Mauritius). Holotype: MCZ 6094. Chromis isharae Schmidt, 1931: 67, Fig. 2. (Type Chromis degruyi Pyle, Earle and Greene, 2008 locality, Amami-Oshima Island, Ryukyu Islands, Chromis degruyi Pyle, Earle and Greene, 2008: 18, Japan). Holotype: ZIN 23289. Figs. 4a–4c. (Type locality, Kayangel Atoll, Siphonochromis lepidostethicus Fowler, 1946: 145, 8°04'16.64"N, 134°40'54.52"E, Palau Island, depth 85 Figs. 13–14. (Type locality, Aguni Shima, Ryukyu meters). Holotype: BPBM 40842. Paratypes: BMNH Islands, Japan). Holotype: ANSP 72010. 2007.10.31.4 (1), CAS 225758 (1), USNM 391139 Lepicephalochromis westalli Whitley, 1964: 180, (1), WAM P.32901-001 (1). Fig. 11. (Type locality, Kenn Reef, Coral Sea). Distribution: Caroline Islands. Holotype: AMS IB.4975. Paratypes: AMS IB.4946– 47 (1, 13), IB.4976 (23). Chromis delta Randall, 1988 Distribution: Three isolated populations: (1) Southern Chromis delta Randall, 1988a: 78, Fig. 3. (Type Japan to Taiwan; (2) Coral Sea to Fiji and Eastern locality, Reef adjacent to wreck of Japanese ship, 7 mi. Australia; (3) Mauritius and Reunion. West of Honiara, Guadalcanal, Solomon Islands, 38– 48 m). Holotype: BPBM 15584. Paratypes: AMS Chromis cinerascens (Cuvier, 1830) I.17506-004 (2), I.19476-020 (1), I.20441-008 (1), Heliases cinerascens Cuvier in Cuvier and Valenciennes, I.22627-005 (2), I.22632-008 (2); BPBM 9436 (11), 1830: 495. (Type locality, Java, Indonesia). Syntypes: 14695 (9), 16188 (4), 18469 (2), 19358 (1), 23435 (1), MNHN 5642 (1), RMNH 947 (2). 26512 (2), 26527 (1), 26678 (3), 30187 (2), 30252 (2),

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 51 Parenti 52

30953 (3) 30974 (2), 31158 (34), 31461 (1); USNM Chromis fieldi Randall and DiBattista, 2013 268413 (1), 223716 (1); WAM P.26099-026 (1). Chromis fieldi Randall and DiBattista, 2013: 5, Figs. Distribution: Christmas Island, Indo-Australian 4–9. (Type locality, West coast off Medine, north of Archipelago to Solomon Islands, Vanuatu and Fiji. Flic en Flac, Mauritius, depth 30 meters). Holotype: BPBM 20241. Paratypes: Many at BPBM 20066 (2), Chromis dimidiata (Klunzinger, 1871) 41005, 16391, 21723 (4), 28436 (11), 27275 (8), Heliastes dimidiatus Klunzinger, 1871: 529. (Type 35567 (3), 37663; CAS 234256 (2); HUJ 20132 (2); locality, Al-Quseir, Egypt, Red Sea). Holotype: ROM 37174 (2); SAIAB 186323 (2); USNM 275619 SMNS 1765. (2), 364137, 405570 (2), 274833 (10). Distribution: Red Sea. Previous records from the Distribution: Indian Ocean: East Africa, South Indian Ocean are based on C. fieldi Randall and Africa, Socotra, Seychelles, Comoros, Madagascar DiBattista, 2013. and Mascarenes East to Western Indonesia. Chromis dispila Griffin, 1923 Chromis flavapicis Randall, 2001 Chromis dispilus Griffin, 1923: 254, Pl. 25. (Type Chromis sp. 1 Allen, 1991: 88, upper fig. (Marquesas locality, West coast of the North Island of New Islands). Zealand). Holotype: AIM MA 275. Paratypes: AIM Chromis flavapicis Randall, 2001: 99, Figs. 4–6. (3). Non-types: (5) mentioned in footnote. (Type locality, Fatu Hiva, Marquesas Islands). Distribution: Kermadec Islands and New Zealand. Holotype: BPBM 11776. Paratypes: AMS I.39935- Chromis durvillei Quéro, Spitz and Vayne, 2010 001 (1); BMNH 11803 (14), 11928 (3), 38508 (1); Chromis durvillei Quéro, Spitz and Vayne, 2010: CAS 210417 (1); MNHN 2000-7 (1); NSMT-P 322, Figs. 1–2. (Type locality, at surface after the 57705 (1); USNM 359632 (1). eruption of Piton de la fournaise). Holotype: MNHN Distribution: Marquesas Islands. 2008-1180. Paratypes: MNH RUN P567 (7); Chromis flavaxilla Randall, 1994 MHNLR 2009. 11.1 (1); SAM 37256 (1). Chromis flavaxilla Randall, 1994: 40, Figs. 1–3. Distribution: Réunion Island, Western Indian Ocean. (Type locality, Um-aabak, Dahklak Archipelago, Chromis earina Pyle, Earle and Greene, 2008 Eritrea, Ethiopia, Red Sea). Holotype: BPBM 36414. Chromis earina Pyle, Earle and Greene, 2008: 21, Paratypes: BMNH 1994.7.22.2 (1); BPBM 21459 Figs.5a–5c. (Type locality, off western coast of Tutuba (1), 27458 (5), 28001 (3), 33289 (1), 35700 (3), Island, 15°32'39.28"N, 167°16'29.82"E, Espiritu Santo, 36415 (19); HUJ 6370 (2), 12849 (3), 13126 (41), Vanuatu). Holotype: MNHN 2007-1921. Paratypes: 13137 (19); USNM 275705 (67), 275710 (6), 275714 BMNH 2007.10.31.5 (1); BPBM 37674 (1), 37714 (4), (1); WAM P.25991-008 (2), P.26461-004 (9). 40720 (2); CAS 225759 (1); MNHN 2007-1926 (1); Distribution: Red Sea, Gulf of Oman, Persian Gulf. USNM 391140 (1); WAM P.32902-001 (1). Chromis flavicauda (Günther, 1880) Distribution: Western Pacific: Palau, to Papua New Heliastes flavicauda Günther, 1880: 7, Pl. 30 (Fig. D). Guinea, Vanuatu, Fiji, to Raja Ampat Islands. (Type locality, Off Pernambuco, Brazil, Challenger Chromis elerae Fowler and Bean, 1928 sta. 122, 30 fms). Holotype: BMNH 1879.5.14.7. Chromis elerae Fowler and Bean, 1928: 52, Pl. 4. Distribution: Brazil. (Type locality, Pasacao, Ragay Gulf, Philippines). Chromis flavipectoralis Randall, 1988 Holotype: USNM 89955. Paratypes: (16) ANSP Chromis flavipectoralis Randall, 1988b: 52, Figs. 3–4. 85281 (1); USNM 98716–20 (1, 2, 1, 1, 1). (Type locality, N. side of Pulau Putri, Seribu Islands, Distribution: Maldives to Fiji North to Taiwan and North of Jakarta, Java Sea, Indonesia, 8 m). Holotype: Guam. BPBM 18569. Paratypes: BMNH 1988.6.2.1-2 (2); Chromis enchrysura Jordan and Gilbert, 1882 BPBM 18554 (2), 30587 (5); CAS 63207 (2); ROM Chromis enchrysurus Jordan and Gilbert, 1882: 286. 54941 (2); RUSI 27720 (1); USNM 270851 (10), (Type locality, Pensacola, Florida, U.S.A). Syntypes: 270853 (4); WAM P.26506-007 (5), P.26514-002 (1). SU 2590 (1), USNM 30871 (now 2). Distribution: Maldives, Similian Islands (Andaman Distribution: the Gulf of Mexico and southeastern Sea) and Java Sea. United States; the West Indies, Bermuda, and Central Chromis fumea (Tanaka, 1917) America. Replaced in northern South America by Pomacentrus fumeus Tanaka, 1917: 9. (Type locality, Chromis vanbebberae. Fish market in Nagasaki, Japan). Holotype: ZUMT Chromis fatuhivae Randall, 2001 7278. Chromis sp. 4 Allen, 1991: 89, upper fig. (Marquesas Chromis caudofasciata Shen and Chen, 1978: 33, Fig. Islands). 12. (Type locality, off Pa-dou-tzu, Northern Taiwan). Chromis fatuhivae Randall, 2001: 97, Fig. 3. (Type Holotype: NTUM 02882. Paratypes: NTUM 02870 locality, Fatu Hiva, Marquesas Islands). Holotype: (1), 02876 (1). BPBM 11766. Paratypes: BPBM 38687 (1), MNHN Chromis n. sp., Hermes, 1986. 2000-5 (1), USNM 359631 (1). Distribution: Southern Japan to Northern New Zealand Distribution: Marquesas Islands. and Sydney; Eastern Malaysia to Western Australia.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Chromis gunting Arango, Pinheiro, Rocha, Chromis iomelas Jordan and Seale, 1906 Greene, Pyle, Copus, Shepherd and Rocha, 2019 Chromis iomelas Jordan and Seale, 1906: 292. (Type Chromis gunting Arango, Pinheiro, Rocha, Greene, locality, Pago Pago, American Samoa). Holotype: Pyle, Copus, Shepherd and Rocha, 2019: 3, Figs. 1a–c. USNM 51743 (1 of 2). Paratypes: SU 8723 (2), (Type locality, Layaglayag point, Batangas, Philippines, USNM 51743 (1 of 2). 13°41'22.74"N, 120°50'12.46"E, depth 100 meters). Distribution: Coral Sea, New Caledonia, Fiji, Samoa, Holotype: PNM 15357. Paratype: CAS 242328. and Society Islands. Distribution: Verde Island Passage, Philippines. Chromis jubauna Moura, 1995 Chromis hangganan Arango, Pinheiro, Rocha, Chromis jubauna Moura, 1995: 91, Figs. 1–2. (Type Greene, Pyle, Copus, Shepherd and Rocha, 2019 locality, Laje de Santos, 24°19'S, 46°11'W, State of Chromis hangganan Arango, Pinheiro, Rocha, Greene, Sao Paulo, Brazil, 18 m). Holotype: MZUSP 44631. Pyle, Copus, Shepherd and Rocha, 2019: 6, Figs. 2a–b. Paratypes: MZUSP 44632–35 (1, 1, 1, 1), 44636–37 (Type locality, Lubang Island, Philippines, (1, 1), 44638–40 (1, 1, 1), 46031–33 (1, 1, 1). 13°45'44.71"N, 120°07'38.31"E, depth 130 meters). Distribution: Southeastern Brazilian coast. Holotype: PNM 15358. Paratype: CAS 243205. Chromis katoi Iwatsubo and Motomura, 2018 Distribution: Lubang Island, Philippines. Chromis katoi Iwatsubo and Motomura, 2018: 29, Chromis hanui Randall and Swerdloff, 1973 Figs. 1–2. (Type locality, Off Eigo, Hachijo Island, Chromis hanui Randall and Swerdloff, 1973: 338, Izu Islands, Japan, 33°08'N, 139°44'E, depth 18 Figs. 6–7. (Type locality, Base of drop-off, Moku meters). Holotype: KAUM-I. 200400. Paratypes: Manu, Oahu I., Hawaiian Islands, U.S.A., 50 m). KAUM-I 200401–200410 (1, 1, 1, 1, 1, 1, 1, 1, 1, 1). Holotype: BPBM 7966. Paratypes: AMS I.16765-001 Distribution: Izu Islands, Japan. (2); BMNH 1972.12.5.1-2 (2); BPBM 4460 (1), 6351 Chromis kennensis Whitley, 1964 (2), 7405 (40), 7931 (4), 9774 (9), 13764 (1); CAS Chromis kennensis Whitley, 1964: 182, Fig. 12. 15913 (2); MNHN 1972-95 (2); RUSI 2406 (3); (Type locality, Kenn Reef, Coral Sea). Holotype: USNM 208431–32 (4, 5). AMS IB.4973. Distribution: Hawaiian Islands. Distribution: Southwest Pacific Ocean: Coral Sea, Chromis howsoni Allen and Erdmann, 2014 Lord Howe Island, Eastern Australia, New Caledonia Chromis howsoni Allen and Erdmann, 2014: 14, and Tonga. Figs. 1–5. (Type locality, Stewart Reefs, 9° 07.741’S, Chromis klunzingeri Whitley, 1929 149°23.039’E, about 7 km Southeast of Tufi, Oro Heliastes hypsilepis Klunzinger, 1880: 398 (nec Province, Papua New Guinea, depth 17–20 meters). Günther) (misidentification). (Type locality, King Holotype: WAM P.33812-001. Paratypes: AMS George’s Sound, Western Australia). I.46380-001 (4); WAM P.33811-001 (6), 33812-002 Chromis klunzingeri Whitley, 1929: 244. (Type (4), 33088-001 (5); USNM 427095 (4). locality, King George Sound, Western Australia). Distribution: Milne Bay and Oro Provinces, Papua Based on Heliastes hypsilepis Klunzinger 1880. New Guinea. Syntypes: SMNS 2556 (2), 2630 (1), 2690 (7). Chromis hypsilepis (Günther, 1867) Distribution: Recherche Archipelago to Abrolhos Heliastes hypsilepis Günther, 1867: 66. (Type Islands (Western Australia). locality, N.S.W., Australia). Syntypes: BMNH Chromis lepidolepis Bleeker, 1876 1866.6.19.10 (1), 1866.6.19.15-16 (2). Chromis lepidolepis Bleeker, 1876: 389. (Type Distribution: Southeastern Australia, Lord Howe and locality, Timor Island, s. Malay Archipelago). Norfolk Islands, Northern New Zealand. Holotype: RMNH 6515. Chromis insolata (Cuvier, 1830) Dascyllus pomacentroides Kendall and Heliases insolatus Cuvier in Cuvier and Valenciennes, Goldsborough, 1911: 298, Pl. 5 (Fig. 1). (Type 1830: 494, Pl. 137. (Type locality, Martinique Island, locality, Taritari, Gilbert Islands, western Pacific on West Indies). Holotype: MNHN 8160. the equator). Holotype: USNM 65812. Distribution: Bermuda, Bahamas and Caribbean Sea; Dascyllus caudofasciatus Montalban, 1928: 26, Pl. 6 also reported from St. Helena Islands (Quero et al., (Fig. 1). (Type locality, Tambagaan, Bungau and 1990). Sibutu Islands, Sulu Archipelago, Philippines). Syntypes: BSMP (13, apparently destroyed). Chromis intercrusma Evermann and Radcliffe, 1917 Lepidochromis brunneus Smith, 1960: 328, Pl. 32 Chromis intercrusma Evermann and Radcliffe, 1917: (Fig. B). (Type locality, Baixo Pinda, Mozambique, 119, Pl. 11 (Fig. 3). (Type locality, Guanape North Western Indian Ocean). Holotype: SAIAB [formerly Island, Peru). Holotype: USNM 77665 [not 77590]. RUSI] 289. Paratypes: USNM 77590 (3). Distribution: Western Indian Ocean to Line Islands Distribution: Cabo Blanco (Peru) to Tarapaca and Australia to Southern Japan. (Chile).

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 53 Parenti 54

Chromis leucura Gilbert, 1905 Chromis meridiana Greenfield and Woods, 1980 Chromis leucurus Gilbert, 1905: 620, Pl. 77 (Fig. 2). Chromis meridiana Greenfield and Woods, 1980: 634, (Type locality, Avau Channel, between Maui Island Fig. 7. (Type locality, Isla San Felix, Chile, and Lanai Island, Hawaiian Islands, U.S.A., 26°17'30"S, 80°5'40"W, 9.1–12.2 m). Holotype: Albatross sta. 3875, 34–65 fms). Holotype: USNM FMNH 75774. Paratypes: BMNH 1978.11.7.7-10 51587. Paratypes: SU 8623 (1), USNM 51660 (1). (4); BPBM 22426 (2); CAS 42069 (2); FMNH 70829 Distribution: Scattered localities in the Indo-West (7); SIO 65-626 (14), 65-629 (32); USNM 219030 Pacific: Hawaiian Islands; Marquesas Islands, (4). Ryukyu Islands, Mauritius, and Réunion. Distribution: San Felix Islands (Eastern Pacific). Chromis limbata (Valenciennes, 1833) Chromis mirationis Tanaka, 1917 Heliases limbatus Valenciennes in Cuvier and Chromis mirationis Tanaka, 1917: 8. (Type locality, Valenciennes, 1833: 511. (Type locality, Madeira). Off Goto Island, Nagasaki Pref., 31°27'–31°33'N, Holotype: MNHN 1377. 125°52'–126°57'E, Japan). Holotype: ZUMT 3267. Heliazes marginatus Valenciennes, 1843: 27. (Type Tea et al. (2019) located the holotype in the locality, Canary Islands). Syntypes: MNHN A-0148 Department of Zoology, University Museum, (2, dry). University of Tokyo, as ZUMT 3627. Distribution: Eastern tropical Atlantic, including Chromis fraenatus Araga and Yoshino in Masuda, Azores and Canary Islands. Araga and Yoshino, 1975: 286, Pl. 96 (Fig. J). (Type locality, Reef off Minabe, Wakayama Prefecture, Chromis limbaughi Greenfield and Woods, 1980 Japan, depth about 30 meters). Holotype: SMBL Chromis limbaughi Greenfield and Woods, 1980: 72067. Paratypes: SMBL 73391 (1). Type materials 627, Fig. 1. (Type locality, Cabo San Lucas, Baja not located (Tea et al., 2019: 251). California, Mexico, 24.4–30.5 m). Holotype: SIO 61- Chromis megalopsis Allen, 1976: 136, Fig. 1. (Type 233. Paratypes: BMNH 1978.11.7.3-6 (4); BPBM locality, About 40 nautical miles West of Bernier 22427 (4); CAS 42068 (4); FMNH 83863 (6), 84730 Island, 24°59'S, 112°27'E, Western Australia, depth (1), 84731-35 (5); LACM 37928-1 (3); SIO 53-178 71 fathoms). Holotype: WAM P.25207-001. (2), 60-147 (1), 61-233 (13), 61-272 (28), 61-277 (31 Paratypes: WAM P.22343-001 (1). or 32), 61-280 (42); USNM 219029 (4). Distribution: Japan to Australia. Distribution: Gulf of California. Chromis monochroma Allen and Randall, 2004 Chromis lineata Fowler and Bean, 1928 Chromis monochroma Allen and Randall, 2004a: 18, Chromis lineatus Fowler and Bean, 1928: 50, Pl. 3. Fig. 1. (Type locality, Near Maumere, Flores, (Type locality, Pangasinan Island, Philippines). Indonesia, about 8°39'S, 122°18'E, depth 53 meters). Holotype: USNM 89953. Paratypes: USNM 93530 (1). Holotype: BPBM 34070. Paratypes: CAS 219161 Distribution: Indo-Australian Archipelago from (1), USNM 376248 (1), WAM P.32461-001 (1). Christmas Island to Great Barrier Reef and Palau. Distribution: Flores, Indonesia. Chromis lubbocki Edwards, 1986 Chromis multilineata (Guichenot, 1853) Chromis lubbocki Edwards, 1986: 184, Figs. 1, 2d. Heliases multilineatus Guichenot, 1853: 76, Pl. 2 (Type locality, Porto da Faja, Brava Island, Cape (Fig. 2). (Type locality, Havana, Cuba). Syntypes: Verde Islands). Holotype: BMNH 1982.4.2.17. MNHN 8222 (2, poor condition). Paratypes: BMNH 1982.4.2.16 (1), 1982.4.2.18–20 Heliases marginata Castelnau, 1855: 9, Pl. 3 (Fig. 1). (1, 1, 1); INIP 5053 (1), 5316 (1), uncat. (1); RMNH (Type locality, Bahia, Brazil). Syntypes: MNHN 29211–12 (1, 1), 29542 (1). 0000-8221 (2). Name spelled marginata in text, list Distribution: Cape Verde Islands. of figures, and index, marginatus in figure caption; Chromis mamatapara Shepherd, Pinheiro, Phelps, Heliases is masculine so marginatus is correct. Easton, Pérez-Matus and Rocha, 2020 Objectively invalid; preoccupied by Heliazes [sic] Chromis mamatapara Shepherd, Pinheiro, Phelps, marginatus Valenciennes 1843; not replaced. Easton, Pérez-Matus and Rocha, 2020: 327, Figs. 1– Furcaria puncta Poey, 1860: 195. (Type locality, 3. (Type locality, Rapa Niu, Easter Island, Chile, Havana, Cuba). Holotype or described syntype: MCZ Hanga Piko, 27°9'12"S, 109°26'52"W, depth 90 14664. Poey specimens: MCZ 14663 (1), 14665 (1), meters). Holotype: CAS 247107. Paratypes: CAS 14666-69 (4). 247108 (1), USNM 447900 (1). Onychognathus cautus Troschel, 1866: 231, Pl. 5 Distribution: Easter Island, Chile. (Figs. 12–14). (Type locality, Cape Verde Islands). Syntypes: (3) whereabouts unknown. Chromis margaritifer Fowler, 1946 Demoisellea caribbaea Fowler, 1944a: 139, Fig. 221. Chromis dimidiatus margaritifer Fowler, 1946: 140, (Type locality, Old Providence Island, Caribbean Figs. 9–10. (Type locality, Aguni Shima, Ryukyu Sea). Holotype: ANSP 70076. Islands, Japan). Holotype: ANSP 72008. Distribution: Caribbean Sea to Brazil and Eastern Distribution: Christmas Island and Western Australia Atlantic: Cape Verde Islands, Ascension and Saint to Central Pacific. Helena islands.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Comments: Onychognathus cautus Troschel, 1866 is Paratypes: BPBM 19703–05 (1, 1, 1); FSKU 751005 a valid species of Chromis according to Lloris and (1), 751013 (1); ZUMT 53958–59 (1, 1). Rucabado (1990: 846). Wirtz et al. (2013: 128) also Distribution: Japan, Taiwan, and China. listed C. cautus as valid based on Lloris and Comments: Synonymies follows Iwatsubo and Rucabado but mention Edwards (1983) who Motomura (2013). Chromis flavomaculata Kamohara, synonymized cautus with multilineata. Demoisellea 1960 is recorded as valid in the ECoF (Fricke et al, caribbaea Fowler, 1944 is a synonym according to 2020), not accepting the conclusions of Iwatsubo and Smith-Vaniz (pers. comm. 1978 in Böhlke, 1984). Motomura. Perca japonica Bloch, 1792 has been declared nomen oblitum by Iwatsubo and Motomura Chromis nigroanalis Randall, 1988 but the authors did not produce evidence as required Chromis nigroanalis Randall, 1988b: 49, Figs. 1–2. by the article 23.9 of the Code. In particular, prevailing (Type locality, Cave, 25 km South of Malindi, usage must be maintained when the following Watamu National Marine Reserve, Kenya, 25 m). conditions are both met (23.9.1.): the senior synonym Holotype: BPBM 27307. Paratypes: BPBM 27308 or homonym has not been used as a valid name after (1), CAS 63206 (1), USNM 270852 (1), WAM 1899 and the junior synonym or homonym has been P.26517-006 (3). used for a particular taxon, as its presumed valid name, Distribution: Off Kenya, Maldives, and Java Sea. in at least 25 works, published by at least 10 authors in Chromis nigrura Smith, 1960 the immediately preceding 50 years and encompassing Chromis nigrurus Smith, 1960: 325, Pl. 29 (Fig. I); a span of not less than 10 years. Discovering that both Fig. 3. (Type locality, Inhaca, Mozambique). the conditions of 23.9.1 are met imply that the Holotype: RUSI 295. Paratypes: RUSI 713–722 (2, following action is taken in accordance with this 2, 1, 1, 1, 2, 2, 3, 4, 6). Article: Chromis notata (Temminck and Schlegel, Distribution: Western Indian Ocean, Sri Lanka, and 1843) is here qualified by the term nomen protectum Maldive Islands East to Indonesia. and the invalid, older names Perca japonica Bloch, 1792, Cichla macrolepidota Bloch and Schneider, Chromis nitida (Whitley, 1928) 1801 and Sparus niphon Lacepède, 1802 are here Tetradrachmum nitidum Whitley, 1928: 219, Pl. 17 qualified as nomina oblita. Therefore, the following (Fig. 3). (Type locality, Hayman Island, Queensland, list of 25 works is produced: Breder and Rosen (1966), Australia). Holotype: AMS IA.1993. Paratypes: Kawamura (1980), Allen (1991), Shao and Chen AMS IA.1994–96 (1, 1, 4). (1992), Kawase et al. (1993), Calumpong et al. (1994), Gliphidodon bicolor MacLeay, 1882: 365. (Type Lee and Kim (1996), Randall et al. (1997), Chen et al. locality, New Guinea). Holotype: MAMU (whereabouts (1997), Ni and Kwok (1999), Laboute and unknown). Grandperrin (2000), Nakabo (2000), Sadovy and Distribution: Central Great Barrier Reef to Sydney Cornish (2000), Shinohara et al. (2000), Shimizu and Lord Howe Islands. (2001), Nakabo (2002), Choi et al. (2003), Motomura Chromis notata (Temminck and Schlegel, 1843) et al. (2010), Kim and Song (2012), Iwatsubo and Perca japonica Bloch, 1792: 88, Pl. 311 (Fig. 1). (Type Motomura (2013), Song et al. (2013), Song et al. locality, Japan). Holotype: ZMB 14630 (dry, lost). (2015), Kim et al. (2016), Zubak et al. (2017), Cichla macrolepidota Bloch and Schneider, 1801: Iwatsubo and Motomura (2018). 339. (Type locality, Japan). Nomen novum. Unneeded Chromis okamurai Yamakawa and Randall, 1989 new name for Perca japonica Bloch, 1792. Chromis okamurai Yamakawa and Randall, 1989: 299, Sparus niphon Lacepède, 1802: 44, 141. Nomen Figs. 1–3. (Type locality, Okinawa Trough, 25°53.4'N, novum. Unneeded replacement name for Perca 123°59.4'E, 135–175 m). Holotype: BSKU 29624. japonica Bloch, 1792. Distribution: Okinawa. Heliases notatus Temminck and Schlegel 1843: 66. (Type locality, Japan). Lectotype: RMNH 895 (larger Chromis onumai Senou and Kudo, 2007 of 2). Paralectotypes: RMNH 895 (smaller of 2), Chromis onumai Senou and Kudo, 2007: 52, Figs. 1– ZMB 2792 (1). Lectotype designated by Boeseman 4. (Type locality, Eluanbi, Pingtung, Taiwan, 21.52'N, (1947: 71). 120.5'E). Holotype: ASIZP 0062621. Paratypes: Chromis villadolidi Jordan and Tanaka, 1927: 387, ASIZP 0066516 (1), 0066517 (1), 0066704 (1); Pl. 34 (Fig. 1). (Type locality, Sea of Japan, between BPBM 40446 (1); NMMB-P 1204 (2), 1205 (1); Tsushima and Fukuoka, Kyushu, Japan). Holotype: NSMT-P 73060 [ex. KPM-NI 8795] (1). FMNH 59185 [ex CM 8328]. Paratypes: (5) CAS-SU Distribution: Southern Taiwan and Izu Islands, Japan. 23681 (1). Chromis opercularis (Günther, 1867) Chromis flavomaculatus Kamohara 1960: 5, Fig. 2. Heliastes opercularis Günther in Playfair and (Type locality, Susaki, Kochi Pref., Japan). Holotype: Günther, 1867: 84, Pl. 11 (Fig. 2). (Type locality, BSKU 8482. Zanzibar). Holotype: BMNH 1865.3.18.34 (right half Chromis miyakeensis Moyer and Ida, 1976: 189. skin). Paratypes: BMNH 1865.3.18.34 (spirits). (Type locality, Igaya Bay, Miyake-jima, Izu Islands, Distribution: Western Indian Ocean, Andaman Sea, Japan, depth 14 meters). Holotype: ZUMT 53957. and Christmas Island.

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Chromis ovalis (Steindachner, 1900) Distribution: Red Sea to Tanzania and Seychelles Heliastes ovalis Steindachner, 1900a: 176. (Type East to Maldives and Chagos Archipelago. locality, Honolulu, Oahu Island, Hawaiian Islands, Chromis planesi Lecchini and Williams, 2004 U.S.A). Holotype: UMB 158 [RK UMB 4626]. Chromis planesi Lecchini and Williams, 2004: 100, Species later described in detail in Steindachner Figs. 3–5. (Type locality, channels off Northeastern (1900b: 502). Preoccupied by Chromis ovalis tip of Rapa Island on outer edge of deep plateau, Steindachner 1866 (=Tilapia sparrmanii Smith, 27°32.974'S, 144°19.123'W, depth 50–54 meters). 1840, Cichlidae), removed in 1990 by Opinion 1622. Holotype: USNM 375192. Paratypes: MNHN 2003- Chromis velox Jenkins, 1901: 393, Fig. 6. (Type 2681 (2), USNM 375193 (3). locality, Honolulu, Oahu Island, Hawaiian Islands). Distribution: Rapa Island, Polynesia. Holotype: USNM 49698. Paratypes: CAS-SU 5995 (1), 23293 (2), 23309 (3). Chromis punctipinnis (Cooper, 1863) Distribution: Hawaiian Islands. Ayresia punctipinnis Cooper, 1863: 73, Fig. 18. (Type locality, San Diego Bay, California, U.S.A). Holotype: Chromis ovatiformis Fowler, 1946 USNM 5227 [ex Calif. State Coll. 596]. Paratypes: Chromis ovatiformis Fowler, 1946: 142, Figs. 11–12. USNM 5226 (1), perhaps others from San Pedro. (Type locality, Aguni Shima, Ryukyu Islands, Distribution: Monterrey Bay to central Baja California. Japan). Holotype: ANSP 72009. Distribution: Izu and Ryukyu Islands, Taiwan, Chromis pura Allen and Randall, 2004 Philippines, Brunei, and Malaysia. Chromis pura Allen and Randall, 2004a: 21, Figs. 2–3. (Type locality, Pura, Alor Islands, Indonesia, 8°17.7'S, Chromis pacifica Allen and Erdmann, 2020 124°22.4'E, depth 18 meters). Holotype: BPBM Chromis pacifica Allen and Erdmann, 2020: 104, 37348. Paratypes: CAS 219160 (1), MNHN 2004-558, Figs. 1–4, 6A. (Type locality, Holmes Reefs, Coral USNM 376247 (1), WAM P.32460-001 (1). Sea, Australia, depth 5–8 meters). Holotype: WAM Distribution: Indonesia. P.34767-002. Paratypes BPBM 6091 (2), 7111, (6), 8002, (4), 8505, (3), 8966, (12), 11292, (3), 17043 Chromis randalli Greenfield and Hensley, 1970 (2), 40060, (4); WAM P.25129-005 (2), P.32094- Chromis randalli Greenfield and Hensley, 1970: 689, 001, (7), P.34734-002 (2). Fig. 1. (Type locality, West coast of Easter Island, Distribution: Western and Central Pacific: Hawaiian between Hanga Piko and Motu Kaokao, se. Pacific, 8 Islands, Johnston Atoll, Line Islands, Pitcairn Island, m). Holotype: USNM 205094. Paratypes: BC65-459 Oeno Atoll, Gambier Islands, Tuamotu Archipelago, (1), BPBM 6691–94 (7, 5, 2, 1), FMNH 70034 (1). Society Islands, Austral Islands, Rapa, Cook Islands, Distribution: Easter Island, Southeastern Pacific. Phoenix Island, Fiji, Tonga, New Caledonia, Loyalty Islands, Chesterfield Islands, Vanuatu, Coral Sea Chromis retrofasciata Weber, 1913 Chromis retrofasciatus Weber, 1913: 359, Pl. 6 (Fig. islands and reefs, Marshall Islands, Chuuk, Pohnpei, 5). (Type locality, Kur Islands, Siboga sta. 250, 27 Yap, Wake Island, Mariana Islands and North to m). Holotype: ZMA 111416. Ogasawara and Bonin Islands (Japan). Distribution: Indo-Australian Archipelago North to Chromis pamae Randall and McCosker, 1992 Ryukyu Islands East to New Caledonia, Fiji, Loyalty Chromis pamae Randall and McCosker, 1992: 331, Islands and Northern Great Barrier Reef. Figs. 1–2. (Type locality, off Christian's Point, Pitcairn Island, 25°04'S, 130°06'W, 7.5 m). Chromis sanctaehelenae Edwards, 1987 Chromis sanctaehelenae Edwards in Edwards and Holotype: BPBM 16905. Paratypes: AMS I.31222- Glass, 1987: 645, Figs. 2–3. (Type locality, Lemon 001 (3); BMNH 1991.5.20.4-6 (3); BPBM 12958 (1), Valley Bay, St. Helena Island, Saint Helena Island, 13501 (4), 13681 (1), 16457 (10), 16922 (5); NSMT- South Atlantic. Depth 5–7 m). Holotype: BMNH P 34681 (3); USNM 316516 (4). Distribution: Pitcairn Island. 1984.7.16.141. Paratypes: BMNH 1867.10.8.58-59 (2), 1868.3.11.13 (1), 1965.12.1.87 (1), 1969.3.10.7- Chromis pelloura Randall and Allen, 1982 8 (2), 1984.7.16.142-144 (3), 1984.7.16.145-148 (4), Chromis pelloura Randall and Allen, 1982: 16, Figs. 1985.9.18.1 (1), 1985.9.18.2 (1); USNM 277976 (4); 1–3. (Type locality, off Coral World, Eilat, Gulf of ZMUC P56.61 (1). Aqaba, Red Sea, 40 m). Holotype: BPBM 21523. Distribution: Saint Helena Islands. Paratypes: BPBM 27855 (2); CAS 50046 (1); MNHN 1977-869 (2), 1982-73 (1); USNM 231351 Chromis scotochiloptera Fowler, 1918 (1); WAM P.25523-004 (2). Chromis scotochilopterus Fowler, 1918: 61, Fig. 24. Distribution: Gulf of Aqaba, Red Sea. (Type locality, Philippines). Holotype: ANSP 47539. Paratypes: ANSP 47540 (1). Chromis pembae Smith, 1960 Distribution: Indonesia, Malaysia and Philippines. Chromis pembae Smith, 1960: 323, Pl. 31 (Fig. E). (Type locality, Pemba Island, Tanzania). Holotype: Chromis scotti Emery, 1968 RUSI 58. Paratypes: (3) RUSI 2336 (?3 of 4). Chromis scotti Emery, 1968: 49, Figs. 1, 3b, 4(c–d). (Type locality, three mi. south of Alligator Light,

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Monroe Co., Florida Keys, Florida, U.S.A., 16 m). Chromis unipa Allen and Erdmann, 2009 Holotype: USNM 201346. Paratypes: ANSP 108411 Chromis unipa Allen and Erdmann, 2009a: 130, Figs. (2); ROM 15042 (1); UF 13968 (1), 205453 [ex 6–7. (Type locality, Tridacna Reef, 2°29.672'S, UMML 5453] (1); USNM 201347 (1). 134°58.990'E, Western side of Cenderawasih Bay, Distribution: Southern Florida and Caribbean Sea. Papua Barat Province, Western Papua, Indonesia, western Pacific, depth 45–60 meters). Holotype: NCIP Chromis struhsakeri Randall and Swerdloff, 1973 6349. Paratypes: WAM P.33043-002 (4). Chromis struhsakeri Randall and Swerdloff, 1973: Distribution: West Papua, Indonesia. 344, Figs. 10–12. (Type locality, off Haleiwa, Oahu Island, Hawaiian Islands, U.S.A., 125–160 m). Chromis vanbebberae McFarland, Baldwin, Holotype: BPBM 10564. Paratypes: BMNH Robertson, Rocha and Tornabene, 2020 1972.12.5.3 (1); BPBM 13819 (1), 13919 (1), 13821 Chromis vanbebberae McFarland, Baldwin, Robertson, (1); CAS 15911 (1); USNM 208433 (2). Rocha and Tornabene, 2020: 120, Figs. 6–9 (Type Distribution: Hawaiian Islands. locality, 117 m, Substation Curaçao Downline, Bapor Kibra, Curaçao), Holotype. USNM 446947. Paratypes: Chromis ternatensis (Bleeker, 1856) USNM 414901 (1), 414902 (1), 413966 (1), 413947 (1), Heliases caeruleus Cuvier in Cuvier and 430030 (1), 406206 (1), 442658 (1); CAS 247234 (1); Valenciennes, 1830: 497. (Type locality, New UW 200069 (1), 200070 (1). Guinea, Ulea, Micronesia). Syntypes: MNHN 5644 Distribution: off Bermuda, the Florida Keys, the (2) New Guinea. Bahamas, scattered sites in the Caribbean Sea and Heliases ternatensis Bleeker 1856a: 377. (Type south to São Paulo, Brazil, including the offshore locality, Ternate, Moluccas, Indonesia). Syntypes: (3) islands of Rocas Atoll, St. Paul Rocks, Trindade, and ?BMNH 1862.2.28.41 (1), ?NMV 46061 (1), RMNH Fernando de Noronha. 6512 (1 or more of 5). Chromis philippinus Fowler, 1918: 63, Fig. 25. (Type Chromis vanderbilti (Fowler, 1941) locality, Philippines). Holotype: ANSP 47541. Pycnochromis vanderbilti Fowler, 1941: 260, Fig. Paratypes: ANSP 47542-47 (6, in jar with holotype). 12. (Type locality, Waianae, Oahu Island, Hawaiian Distribution: Red Sea and western Indian Ocean to Islands, U.S.A). Holotype: ANSP 69749. Micronesia and Melanesia North to Japan. Distribution: Philippines East to Hawaiian Islands Comments: Heliases caeruleus Cuvier has been and Pitcairn Group, North to Southern Japan, South suppressed (Opinion 1563) in order to conserve C. to Western Australia, Queensland (Australia), Lord ternatensis as the valid name for this species Howe Island, New Caledonia, Kermadec Islands and (Randall et al., 1985). New Zealand, Tonga and Rapa. Chromis tingting Tea, Gill and Senou 2019 Chromis verater Jordan and Metz, 1912 Chromis mirationis (nec Tanaka, 1917) Song et al. Chromis verater Jordan and Metz, 1912: 526, Pl. 71 (2014): 2, Figs 1a–c and 2, Table 1. (misidentification). (Fig. 2). (Type locality, Honolulu market, Oahu Chromis tingting Tea, Gill and Senou, 2019: 251, Figs. Island, Hawaiian Islands, U.S.A). Holotype: USNM 2–8. (Type locality, West side of Sagami Bay, East side 73912. Paratypes: SU 22389 (1). of Izu Peninsula, Shizuoka Prefecture, Japan, depth 58 Distribution: Johnston Atoll and Hawaiian Islands. meters). Holotype: KPM-NI 30479. Paratypes: KPM-NI Chromis viridis (Cuvier, 1830) 18916 (1), 23617 (1), 32613 (1). Pomacentrus viridis Cuvier (ex Ehrenberg) in Cuvier Distribution: North-west Pacific: Japan and probably and Valenciennes, 1830: 420. (Type locality, Korea. Massawa, Eritrea, Red Sea; Guam, Mariana Islands, Chromis torquata Allen, 2018 W. Pacific). No types known. Chromis torquata Allen, 2018: 16, Figs. 1–3, 4A, 5A. Heliases frenatus Cuvier in Cuvier and Valenciennes, (Type locality, Reefs off Grand Baie, Mauritius, 1830: 498. (Type locality, Guam). No types known. Western Indian Ocean, approximately 19°59'S, Heliases lepisurus Cuvier in Cuvier and Valenciennes, 57°34'E, depth 30 meters). Holotype: BPBM 16352. 1830: 498. (Type locality, New Guinea). Syntypes: Paratypes: BPBM. MNHN 0000-8254 (4). Spelled lepidurus in early Distribution: Mauritius and Réunion (Western literature. On Official Index (Opinion 1563). Mascarenes). Dascyllus cyanurus Rüppell, 1838: 127, Pl. 31 (Fig. 4). (Type locality, Massawa, Eritrea, Red Sea). Chromis trialpha Allen and Randall, 1980 Lectotype: SMF 1253. Paralectotypes: SMF 4932-33 Chromis trialpha Allen and Randall, 1980: (5) 33, (2). Lectotype established by Dor (1984: 181). Figs. 18–19. (Type locality, Ras Muhammed, Sinai Glyphisodon bandanensis Bleeker, 1851: 248. (Type Peninsula, Red Sea, 12 m). Holotype: BPBM 17890. locality, Banda Neira, Banda Islands, Indonesia). Paratypes: BMNH 1977.7.18.1 (1); BPBM 17912 Holotype: RMNH 27456 (1 of 49). (5), 18257 (3), 19717 (1), 20433 (2); HUJ 8370 (1); Distribution: Red Sea and East Africa, Mozambique USNM 217562 (1); WAM P.25797-001 (2). Channel, Comoros, Madagascar and Mascarenes East Distribution: Red Sea and Socotra. to Wake Atoll, Marquesas Islands and Tuamotu

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Archipelago, North to Southern Japan and Ryukyu CAS 58492 (1); NSMT-P 41766 (1); ROM 50818 (1); Islands, South to Western Australia, Middleton Reef, RUSI 26038 [not 26039, ex BPBM 31255] (1); New Caledonia and Tonga; not recorded from USNM 278732 (1); WAM P25768-002 (6), P25991- Hawaiian Islands. 006 (1), P26460-008 (2), P26461 (1), P28787-001 (1). Comments: Heliases frenatus Cuvier, 1830 is not Distribution: Jana Island (Persian Gulf) and Gulf of reported in the ECoF (Fricke et al., 2020) but it is Oman. listed in synonymy by Allen (1991:241). Chromis xanthura Bleeker, 1854 Chromis weberi Fowler and Bean, 1928 Heliases xanthurus Bleeker, 1854a: 107. (Type Chromis weberi Fowler and Bean, 1928: 41, Pl. 1. locality, Banda Neira, Banda Islands, Indonesia). (Type locality, Jakarta, Java, Indonesia). Holotype: Holotype: RMNH 6510. USNM 72713. Paratypes: (7) USNM 96955 + 98242 Distribution: Réunion (Mascarenes), Christmas and (4 + 3, combined). Cocos-Keeling islands East to Pitcairn Group, North Chromis simulans Smith, 1960: 326, Fig. 4. (Type to Southern Japan, South to Western Australia, locality, Mozambique, Western Indian Ocean). Queensland (Australia), New Caledonia and Tonga, Holotype: SAIAB [formerly RUSI] 60. not in the Hawaiian Islands. Distribution: Red Sea, Persian Gulf and Western Chromis xouthos Allen and Erdmann, 2005 Indian Ocean to Line Islands and Pitcairn Group Chromis xouthos Allen and Erdmann, 2005: 90, Figs. North to Southern Japan, South to Jervis Bay (New 1–2. (Type locality, Patch reef off Teupinpineung South Wales, Australia) and New Caledonia. Point, 5°47.674'N, 95°20.803'E. Palau Weh, Aceh Chromis westaustralis Allen, 1976 Province, Sumatra, Indonesia, depth 25 meters). Chromis westaustralis Allen, 1976: 139, Fig. 2. (Type Holotype: NCIP 6301. Paratypes: NCIP 6302 (1), locality, Long Island, Wallabi Group, Houtman WAM P.32757-001 (2), USNM 383523 (2). Abrolhos, ca. 28°30'S, 113°47'E, Western Australia, Distribution: Indian Ocean: Socotra, Maldives and depth 2–8 m). Holotype: WAM P.25317-008. Sumatra (Indonesia). Paratypes: AMS I.18475-001 (1); BMNH 1975.9.23.9 Chromis yamakawai Iwatsubo and Motomura, 2013 [ex WAM P.22046] (1); BPBM 19058 (2), 19059 (3), Chromis yamakawai Iwatsubo and Motomura, 2013: 19163 (1); USNM 214702 (2); WAM P.2789-001 (1), 209, Figs. 9–11, 12F. (Type locality, Northwest P.24853-001 (1), P.25307-011 (1), P.25308-006 (2), Pacific Ocean, off North West coast of Yoron-jima P.25317-009 (1), P.25376-002 (1). island, Kagoshima, Japan, 27°03'21"N, 128°24'39"E). Distribution: Albany to North West Cape (western Holotype: NSMT-P 111948. Australia). Paratypes: BPBM 6880 (1), 11989 (1), 35316 (1), Chromis woodsi Bruner and Arnam, 1979 19073 (2); BSKU 8513 (1), 65128 (1), 65129 (1), Chromis woodsi Bruner and Arnam, 1979: 50, Figs. 65130 (1), 69938 (1); FAKU 104285 (1), 104420 (1), 1–2. (Type locality, Somalia coast, W. Indian Ocean, 61749 (1), 61753 (1), 121418 (1); FRLM 10757 (1); 11°24'N, 61°35'E, 75–175 m). Holotype: FMNH HMNH-P 3118 (1), 3505 (1), 4286 (1), 7486 (1); 85011. Paratypes: BPBM 21080 (1); FMNH 84998- HUMZ 3455 (1), 3456 (1), 3457 (1), 3461 (1), 39070 85002 (5), 85012 (1), 85013 (now 0), 85014-16 (3), (1), 46754 (1), 46758 (1), 48235 (1), 48567 (1), 48579 85017-18 (0, 0), 85019-21 (3), 85022 (1); ROM (1); KAUM-I 7211 (1), 7212 (1), 29581 (1), 31023 34301 (1); WAM P.26020-001 (1). (1), 31024 (1), 31025 (1) 40100 (1), 40101 (1), 40161 Distribution: Somalia to Mozambique Channel. (1), 40162, 40163 (1), 40170, 40175 (1), 40177, 40178 (1), 40181, 40183 (1), 40186, 40194 (1), 40198, 40213 Chromis xanthochira (Bleeker, 1851) (1), 40214, 40216 (1), 40217, 40129 (1), 40221; KPM- Heliases xanthochirus Bleeker, 1851: 248. (Type NI 4686 (1), 4687 (1) 5065 (1), 5105 (1), 5106 (1), locality, Banda Neira, Banda Islands, Indonesia). 17973 (1), 18542 (1); KSHS 15340 (1), 15341 (1), Syntypes: (5) ?BMNH 1861.2.28.43 (1), ?NMV 16983 (1), 16984 (1), 16985 (1), 16986 (1); NSMT-P 46062 (1). 3246-004 (1), 29624 (1), 30613 (1), 30811 (1), 30812 Chromis reticulatus Fowler and Bean, 1928: 298, Pl. (1), 32798 (1), 90966 (1), 90967 (1), 92979 (1), 32810 5 (Fig. 1). (Type locality, Taritari, Gilbert Islands, (1), 105526 (1),, URM-P 907 (1), 908 (1), 909 (1), 910 Western Pacific on the equator). Holotype: USNM (1), 911 (1), 1182 (1), 4598 (1), 11761 (1), 22905 (1), 65812. 27135 (1), 30301 (1), 31167 (1), 33815 (1), 37845 (1), Distribution: Christmas Island to Solomon Islands 37901 (1), 38096 (1), 39899 (1), 39900 (1); WAM-P and Northern Great Barrier Reef. 25516-016 (1); USNM 314196 (2), 364157 (1). Chromis xanthopterygia Randall and McCarthy, 1988 Distribution: Japan, Taiwan and Philippines. Chromis xanthopterygia Randall and McCarthy, 1988: Genus Cheiloprion Weber, 1913 134, Figs. 1–3. (Type locality, Coral reef, ne. side of Cheiloprion Weber, 1913: 342. Masculine. Type Jana Island, Persian Gulf, Saudi Arabia, 12 m). species Pomacentrus labiatus Day, 1877. Type by Holotype: BPBM 30436. Paratypes: BMNH original designation. 1888.12.29.174-176 (3), 1986.6.24.1 (1); BPBM Indo-Australian Archipelago; monotypic. 21356 (2), 30308 (2), 30407 (1), 30495 (7), 31255 (1);

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Cheiloprion labiatus (Day, 1877) P.27827-026 (1), P.29624-088 (4), P.31144 (3), Pomacentrus labiatus Day 1877: 384, Pl. 81 (Fig. 2). P.32921-001 (1). (Type locality, Andaman and Nicobar Islands, Bay of Distribution: Eastern Indonesia east to New Ireland Bengal). Syntypes and/or Day specimens: AMS I.149 (Papua New Guinea). (1); BMNH 1889.2.1.3935-3938 (4); NMW 22791 Chrysiptera biocellata (Quoy and Gaimard, 1825) (1); RMNH 896 (1); ZSI 1603-04 (2), 1609 (1, Glyphisodon biocellatus Quoy and Gaimard, 1825: figured specimen). 389. (Type locality, Guam, Mariana Islands, w. Distribution: Andaman Sea, Indonesia to Palau, Pacific). Holotype: MNHN 0000-8361. Philippines, and Solomon Islands. Glyphisodon antjerius Cuvier (ex Kuhl and van Genus Chrysiptera Swainson, 1839 Hasselt) in Cuvier and Valenciennes 1830: 481. Chrysiptera (subgenus of Glyphisodon) Swainson, (Type locality, Anjer, Java, Indonesia). Holotype: 1839: 171, 216. Feminine. Type species Glyphisodon ?RMNH 889., 1830; azureus Cuvier, 1830. Type by monotypy. Glyphisodon zonatus Cuvier in Cuvier and Chrysoptera Agassiz, 1846: 84. Unjustified emendation. Valenciennes, 1830: 483. (Type locality, New Paraglyphidodon Bleeker, 1876: 387. Masculine. Guinea; Vanikoro Island, Santa Cruz Islands, Type species Paraglyphidodon oxycephalus Bleeker, Southwestern Pacific). Syntypes: MNHN 0000-8360 1876c. Type by subsequent designation by Jordan (2) New Guinea, 0000-8337 (3) Vanikoro. (1919a: 384). Also appeared in Bleeker (1877d: 40) Glyphisodon punctulatus Cuvier, 1830: 484. (Type and in Bleeker (1877c: 116) with type Glyphidodon locality, Guam, Mariana Islands, Western Pacific). melas Kuhl and van Hasselt by monotypy. Holotype: MNHN 0000-8339. Glyphidodontops Bleeker, 1877c: 128. Masculine. Glyphisodon balinensis Bleeker, 1849: 9. (Type Type species Glyphidodon azureus Cuvier, 1830 (= locality, Boleling, Northern Bali, Indonesia). Genus Glyphidodon cyaneus Quoy and Gaimard, 1825). spelled as Glyphisodon on page 4 and as Type by being a replacement name. Unneeded Glijphisodon on page 9. No types known. replacement for Chrysiptera Swainson, 1839. Glyphisodon rossii Bleeker, 1854b: 48. (Type Iredaleichthys Whitley, 1928: 296. Masculine. Type locality, Nova Selma, Cocos-Keeling Islands, Indian species Glyphisodon azureus Cuvier, 1830. Type by Ocean). Syntypes: (2) RMNH 6498 (2 of 39). being a replacement name. Unneeded replacement Glyphidodon antjerius var. fasciatus Günther, 1862: for Chrysiptera Swainson, 1839. 51. (Type locality, Ambon Island, Molucca Islands, Tropical Indo-West Pacific; 40 species (Allen, 1991; Indonesia; Borneo). Syntypes: (3) BMNH. Allen and Adrim, 1992; Allen and Rajasuriya, 1994; Glyphidodon cingulatus Kner, 1867: 725 [17], Pl. 4 Randall, 1994; Allen, 1999; Allen and Bailey, 2002; (Fig. 2). (Type locality, Samoa Islands). Holotype: Allen et al., 2015; Allen et al., 2017). Three Whereabouts unknown. Original number of holotype: presumably undescribed species have been listed from MGH 3730. (Type locality, Pl. 4 (Fig. 2), labelled Chagos Archipelago (Winterbottom et al., 1989). Glyphidodon albocinctus. Glyphidodon albocinctus Kner, 1867: Pl. 4 (Fig. 2). Chrysiptera albata Allen and Bailey, 2002 (Type locality, probably Fiji Islands). Holotype: Chrysiptera albata Allen and Bailey, 2002: 40, Fig. ?NMW 21614 (1). 1. (Type locality, Nikumaroro (Garden) Id, Phoenix Heliastes cinctus Playfair, 1868: 864. (Type locality, Island). Holotype: WAM P.32109-001. Paratypes: Seychelles). Holotype: BMNH 1867.8.16.75. WAM P.32109-002 (2). Distribution: Western Indian Ocean to Samoa, Distribution: Known only from the type locality. Marshall and Gilbert Islands north to Ryukyu Islands. Chrysiptera annulata (Peters, 1855) Chrysiptera bleekeri (Fowler and Bean, 1928) Pomacentrus annulatus Peters 1855a: 455. (Type Abudefduf bleekeri Fowler and Bean, 1928: 165, Pl. locality, Mozambique, 15°S). Syntypes: ZMB 2762–63 15. (Type locality, Jolo Light, Philippines, Albatross (3, 3). Also described as new in Peters (1855b: 265). sta. 5145). Holotype: not determined. Paratypes: (5) Glyphidodon westermani Bleeker, 1863: 30. (Type ANSP 85278 (1), USNM 93531 (now 3). Additional locality, Java, Indonesia). Syntypes: (2). material: (4). Distribution: Red Sea south to Durban, including Distribution: Timor, Flores, and Philippines. Mauritius, Réunion, Aldabra, and Madagascar. Chrysiptera brownriggii (Bennett, 1828) Chrysiptera arnazae Allen, Erdmann and Barber, Chaetodon brownriggii Bennett, 1828: unnumbered 2010 page [8], Pl. 8. (Type locality, South coast of Sri Chrysiptera arnazae Allen, Erdmann and Barber, Lanka). No types known. 2010: 63, Figs. 1–3. (Type locality, Pulau Lemon, Glyphisodon albofasciatus Hombron and Jacquinot 0°53.395'S, 134°04.782'E, near Manokwari, Papua in Jacquinot and Guichenot 1853: 49, Pl. 5 (Fig. 4). Barat Province, Indonesia, depth 6–10 meters). (Type locality, Poulolant, Borneo). Holotype: Holotype: NCIP 6334. Paratypes: AMS I.44560-001 MNHN 0000-8331. (3), NCIP 6335 (2), USNM 391578 (3), WAM

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Glyphisodon xanthozona Bleeker, 1853d: 283. (Type Island, southern Malay Archipelago). Holotype: locality, Cauer, Sumatra, Indonesia). Syntypes: (3) MNHN 0000-8364. RMNH 6484 (3 of 34). Glyphisodon azureus Cuvier (ex Quoy and Gaimard) Glyphidodon amabilis De Vis, 1884: 452. (Type in Cuvier and Valenciennes, 1830: 479. (Type locality, South Sea Islands). No type known. locality, Timor Island, southern Malay Archipelago; Distribution: Western Indian Ocean to Western Tonga). Syntypes: MNHN 0000-8365 (1) Timor, Indonesia. 0000-8366 (1) Tonga. Chrysiptera gaimardi Swainson, 1839: 216. Based Chrysiptera burtjonesi Allen, Erdmann and on Quoy and Gaimard 1824–25: Pl. 64 (Fig. 4). No Cahyani, 2017 types known. Chrysiptera burtjonesi Allen, Erdmann and Cahyani, Glyphidodon assimilis Günther, 1862: 52. (Type 2017: 13, Figs. 2–4, 5A, 6A. (Type locality, Solomon locality, Borneo; Philippines; Indian Seas). Syntypes: Islands, Florida Group, Nggela Island, Avi Avi, (4 plus literature citation) BMNH 1857.2.17.3-4 (2 09°07.308'S, 160°12.832'E, depth 2–8 meters). juveniles), Indian Seas, BMNH 1858.4.21.8 (1) Holotype: WAM P.34622-001. Paratypes: USNM Borneo, 1976.5.14.3 (1). 432549 (5), WAM P.25136-001 (2), P.34618-001 Abudefduf turchesius Jordan and Seale, 1907: 28, (12), P.34622-002 (4). Fig. 10. (Type locality, Cavite, Luzon Island, Distribution: Solomon Islands: collected at the Philippines). Holotype: CAS-SU 9245. following localities: Isabel, Choiseul, Makira, Malaita Abudefduf sapphirus Jordan and Richardson, 1908: and nearby Alite Reef, Shortland Islands, New 264, Fig. 10. (Type locality, Ticao Island, Georgia Group, Russell Group, and the Florida Group. Philippines). Holotype: CAS-SU 20207. Paratypes: Chrysiptera caeruleolineata (Allen, 1973) CAS-SU 20212 (2), USNM 61682 (3). Abudefduf caeruleolineatus Allen, 1973a: 35, Fig. 2. Glyphisodon hedleyi Whitley, 1927b: 20, Pl. 1 (Fig. (Type locality, Outer reef slope off middle of 5). (Type locality, Dauco Island Reef, Port Moresby, Kranket Island, ca. 1.5 nautical mi. North of Madang, Papua New Guinea). Holotype: AMS IA.1328. New Guinea, 5°12'S, 145°50'52"E, 40 m). Holotype: Paratypes: AMS I.14545 (1), IA.2764 (1). USNM 207941. Paratypes: AMS I.16480-001 (2), Chrysiptera punctatoperculare Fowler, 1946: 156, BMNH 1972.8.14.4-5 (2), BPBM 13227 (2), MNHN Figs. 25–26. (Type locality, Aguni Shima, Ryukyu 1972-87 (2), USNM 207942 (1). Islands, Japan). Holotype: ANSP 72029. Paratypes: Distribution: Rowley Shoals (Western Australia), ANSP 72030-38 (9). Samoa, Fiji to Ryukyu Islands and Guam. Distribution: Western Australia to Great Barrier Reef and Solomon Islands North to Ryukyu and Mariana Chrysiptera caesifrons Allen, Erdmann and Islands. Kurniasih, 2015 Comments: the status of Glyphisodon azureus Chrysiptera sp. Allen and Erdmann 2012: 595 Cuvier, 1830 is not established in the ECoF (Fricke (Halmahera). et al., 2020), however Allen (1991:242) listed this Chrysiptera caesifrons Allen, Erdmann and nominal species in synonymy with Chrysiptera Kurniasih, 2015: 20, Figs. 1–2 and 7a. (Type locality, cyanea (Quoy and Gaimard, 1825). Ayau Kecil Island, Ayau Atoll, Raja Ampat Islands, West Papua Province, Indonesia, 00°21.342'S, Chrysiptera cymatilis Allen, 1999 130°58.648'W, depth 1.5–2 meters). Holotype: MZB Chrysiptera cymatilis Allen, 1999a: 99, Fig. 1. (Type 22260. Paratypes: MZB 22259 (6), 22258 (4), WAM locality, W. side of Goodenough Island, P.33346-001 (6), P.34295-001 (6). D'Entrecasteaux Group, Milne Bay Prov., Papua New Distribution: Halmahera (Eastern Indonesia) east to Guinea, ca. 9°23'S, 150°6.5'E, 9–15 m). Holotype: New Ireland (Papua New Guinea) and Vanuatu, AMS I.16701-004. Paratypes: AMS I.16701-003 South to Queensland (Australia) and New Caledonia. (11), I.16708-002 (4); WAM P.31362-005 (1). Distribution: D’Entrecasteaux Islands, Papua New Chrysiptera chrysocephala Manica, Pilcher and Guinea. Oakley, 2002 Chrysiptera chrysocephala Manica, Pilcher and Chrysiptera ellenae Allen, Erdmann and Cahyani, Oakley, 2002: 311, Figs. 1–2. (Type locality, Pulau 2015 Layang Layang, Spratley Islands, South China Sea). Chrysiptera oxycephala [nec Bleeker] Allen and Holotype: UMZC 2003.1. Paratypes: (2) BMNH Erdmann, 2012: 593. (Raja Ampat Islands, West 2002.3.2.157 (1), UNIMAS uncat. (1). Papua, Indonesia, lower right photograph only). Distribution: South China Sea and Gulf of Tomini. Chrysiptera ellenae Allen, Erdmann and Cahyani, 2015: 60, Figs. 2–4. (Type locality, Wayag Lagoon, Chrysiptera cyanea (Quoy and Gaimard, 1825) Raja Ampat Islands, West Papua Province, Glyphisodon cyaneus Quoy and Gaimard, 1825: 392, Indonesia, 00°00.058'N, 130°01.149'E, depth 4–12 Pl. 64 (Fig. 3). (Type locality, Timor Island, s. Malay meters). Holotype: MZB 22771. Paratypes: MZB Archipelago). Holotype: MNHN 0000-8365. 22772 (16), USNM 432465 (6), WAM P.33947-001 Glyphisodon uniocellatus Quoy and Gaimard, 1825: (2), P.34336-001 (13). 393, Pl. 64 (Fig. 4). (Type locality, Kupang, Timor

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Distribution: Raja Ampat Islands, West Papua, Chrysiptera hemicyanea (Weber, 1913) Indonesia. Abudefduf hemicyaneus Weber, 1913: 351, Pl. 10 (Fig. 11). (Type locality, Tual, Kei Islands, Siboga Chrysiptera flavipinnis (Allen and Robertson, sta. 258). Syntypes: ZMA 109437 (3). 1974) Distribution: Northwest shelf of Australia and Abudefduf flavipinnis Allen and Robertson, 1974: Indonesia East to New Ireland (Papua New Guinea). 157, Pl. 2. (Type locality, Pixie Reef, Great Barrier Reef, off Cairns, Queensland, Australia, 20 m). Chrysiptera kuiteri Allen and Rajasuriya, 1995 Holotype: AMS I.16481-001. Paratypes: AMS Chrysiptera kuiteri Allen and Rajasuriya, 1995: 283, IB.4788 (1), I.15622-008 (1), I.15625-009 (13), Fig. 1. (Type locality, Near shipwreck off Tulamben, I.15635-002 (1), I.15637-011 (1), I.15682-001 (6), NE. coast of Bali, Indonesia, ca. 8°18'S, 115°37'E, 20 I.15684-005 (6), I.16482-001 (3), I.16636-001 (1); m). Holotype: NCIP 6131. Paratypes: NARA BMNH 1972.8.14.6-7 (2); BPBM 13228 (4); MNHN (F).0015 (1); WAM P.30827-001 (1). 1972-88 (2); plus more at AMS. Distribution: Indonesia, Sri Lanka. Distribution: Papua New Guinea to Eastern Australia Chrysiptera leucopoma (Cuvier, 1830) South to Southern New South Wales, Solomon Glyphisodon leucopomus Cuvier in Cuvier and Islands and Western Coral Sea. Valenciennes, 1830: 480. (Type locality, Strong Chrysiptera galba (Allen and Randall, 1974) Island [Oualan Island], Caroline Islands Holotype: Glyphidodontops galbus Allen and Randall, 1974: MNHN (lost). 40, Fig. 2. (Type locality, South side of Ruea Point, Glyphidodon taenioruptus Cartier, 1874: 101. (Type Rapa Island, 14 m). Holotype: BPBM 11244. locality, Bohol, Philippines). Holotype: Whereabouts Paratypes: AMS I.17347-001 (1), I.17348-001 (3); unknown. BMNH 1973.10.17.2-3 (2); BPBM 11150 (1), 11183 Abudefduf miyakoe Okada and Ikeda, 1937: 83, Fig. 2; (10), 11197 (5), 11221 (9), 11232 (3), 11243 (2), Pl. 6 (Fig. 8). (Type locality, Miyako Island, Ryukyu 13685 (1); USNM 211293 (5). Islands, Japan). Holotype: Okada coll. spec. 2256 Distribution: Cook and Austral Islands, Rapa, (lost). Paratypes: Okada coll. spec. 2257 (1, lost). Gambier and Pitcairn Groups. Chrysiptera caudofasciata Okada and Ikeda, 1939: 182, Fig. 15. (Type locality, Sonai, Iriomote Island, Chrysiptera giti Allen and Erdmann, 2008 Taketomi Town, Yaeyama County, Okinawa Chrysiptera giti Allen and Erdmann, 2008a: 173, Prefecture, southern Ryukyu Islands, Japan). Figs. 1–3. (Type locality, Teluk Sebakor, Holotype: YCM-P 30220 [ex Okada coll. spec. 2604]. 3°16'58.36"S, 132°44'17.16"E, Fak Fak Peninsula, Distribution: Christmas Island and Indonesia East to Papua Barat Province, Indonesia, depth 6–10 meters). Wake and Marquesas islands, North to Southern Holotype: NCIP 6332. Paratypes: NCIP 6333 (3), Japan, South to Western Australia, New South Wales USNM 391568 (2), WAM P.32810-002 (3). (Australia), New Caledonia and Austral Islands. Distribution: West Papua (Indonesia) and Sulawesi. Comments: Chrysiptera leucopoma regarded as a Chrysiptera glauca (Cuvier, 1830) synonym of brownriggii by Allen and Bailey (2002: Glyphisodon glaucus Cuvier in Cuvier and 42) is presently validated as Pacific sibling of C. Valenciennes, 1830: 475. (Type locality, Guam, brownriggii (Bennett 1828), which is now restricted Mariana Islands, W. Pacific). Syntypes: MNHN 8371 to the Indian Ocean (Allen et al., 2015). (2). Chrysiptera maurineae Allen, Erdmann and Glyphisodon modestus Schlegel and Müller, 1840: Cahyani, 2015 23, Pl. 6 (Fig. 2). (Type locality, Java Sea; North Chrysiptera maurineae Allen, Erdmann and Cahyani, Java; South Borneo; West Sumatra, Indonesia). 2015: 64, Figs. 5–6. (Type locality, Pulau Nurage, Syntypes: RMNH 909 (1), 910 (3), 911 (2). Cenderawasih Bay, West Papua Province, Indonesia, Glyphisodon phaiosoma Bleeker, 1849: 9. (Type 3°01.807'S, 134°50.058'E, depth 5–12 meters). locality, Boleling, Northern Bali, Indonesia). No Holotype: MZB 22773. Paratypes: MZB 22774 (5); types known. WAM P.33046–009 (5). Glyphidodon pallidus De Vis, 1884: 452. (Type Distribution: West Papua Province, Indonesia. locality, Banks Group, Vanuatu, Southwestern Pacific). Holotype: QM I.1353. Chrysiptera niger (Allen, 1975) Abudefduf caesio Seale, 1906: 53, Fig. 8. (Type Glyphidodontops niger Allen, 1975a: 89, Pl. 3 (Fig. a). locality, Tubuai, Austral Islands, French Polynesia). (Type locality, Near mouth of Tufi Inlet, Cape Nelson, Holotype: BPBM 742. New Guinea, 9°05'S, 149°19'E, 1–2 m). Holotype: Chrysiptera hollisi Fowler, 1946: 155, Figs. 21–22. AMS I.16708-004. Paratypes: AMS I.16708-005 (5). (Type locality, Shima, Ryukyu Islands, Japan). Distribution: Southeastern New Guinea and Holotype: ANSP 72023. Paratypes: ANSP 72024-26 D'Entrecasteaux Islands (3, with holotype), 72027 (1). Chrysiptera notialis (Allen, 1975) Distribution: Western Indian Ocean to Line Islands Glyphidodontops notialis Allen, 1975a: 91, Pl. 3 (Fig. and Pitcairn Group; Australia to Japan. b). (Type locality, Off Phillip Point, Lord Howe

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 61 Parenti 62

Island, 31°32'S, 159°04'E, 15 m). Holotype: AMS Japan). Holotype: ANSP 72017. Paratypes: ANSP I.17402-008. Paratypes: AMS I.17357-016 (1), 72018-19 (2), 72020 (1), 72021-22 (2), 72090 (1). I.17358-012 (6), I.17374-007 (2), I.17377-014 (3), Distribution: Western Australia, Indo-Australian I.17466-001 (1); BMNH 1973.10.17.4-9 (6); BPBM Archipelago to Ryukyu Islands, Great Barrier Reef 15935–36 (18, 16); MNHN 1973-59 (3); USNM and New Caledonia. 211292 (4). Comments: Based on the examination of the holotype Distribution: Lord Howe and Norfolk Islands, New Chrysiptera personata Fowler, 1946 is here regarded as Caledonia. new junior synonym of Chrysiptera rex (Snyder, 1909). Chrysiptera oxycephala (Bleeker, 1876) Chrysiptera rollandi (Whitley, 1961) Paraglyphidodon oxycephalus Bleeker, 1876: 387. Chromis rollandi Whitley, 1961: 60, Fig. 1. (Type (Type locality, Buro Island and Timor Island, locality, Baie des Citrons, Nouméa, New Caledonia). Indonesia). Holotype RMNH 6506. Paratypes: (3). Holotype: AMS IB.4449. Abudefduf azurepunctatus Fowler and Bean, 1928: Distribution: Andaman Sea to Loyalty Islands, New 149. (Type locality, Romblon Harbor, Philippines). Caledonia, and Great Barrier Reef North to Philippines. Holotype: USNM 89963. Paratypes: (19) ANSP Chrysiptera sheila Randall, 1994 85288 (4), USNM 95277-318 (total 83 spec.). Chrysiptera sheila Randall, 1994: 43, Figs. 5–8. (Type Chrysiptera melanomaculata Aoyagi, 1941a: 180, Fig. locality, South coast, Rahah Bay, Oman, 16°57'N, (Type locality, Palau Island). Holotype: YCM-P 30541 54°49'12"E, Arabian Sea, 0–2 m). Holotype: BPBM [ex FAKU 5000]. Paratypes: YCM-P 30542 (1). 36374. Paratypes: BMNH 1994.7.22.3 (1); BPBM Distribution: Indo-Australian Archipelago. 36072 (6), 36096 (8), 36375 (3); HUJ 17700 (1); Chrysiptera papuensis Allen, Erdmann and Cahyani, USNM 329753 (1); WAM P.30793-001 (2). 2015 Distribution: Arabian Sea and Gulf of Oman. Chrysiptera papuensis Allen, Erdmann and Cahyani, Chrysiptera sinclairi Allen, 1987 2015: 72, Figs. 10–11. (Type locality, Waga Waga, Chrysiptera sinclairi Allen 1987a: 107, Fig. 1. (Type Milne Bay Province, Papua New Guinea, locality, Hayne harbor Lagoon, Los Negros Island, 10°24.429'S, 150°24.650'E, depth 25–10 meters). Manus Island, 2°03'S, 147°26'E, 7–9 m). Holotype: Holotype: WAM P.34426-001. Paratypes: USNM WAM P.27827-056. Paratypes: BMNH 1986.5.7.1-4 432446 (4); WAM P.34320-001 (6). (4); BPBM 30926 (4); KFRS unregistered (28); Distribution: Papua New Guinea. MNHN 1986-401 (4); USNM 278055 (3); WAM Chrysiptera parasema (Fowler, 1918) P.27825-042 (5), P.27827-010 (14), P.27828-002 (5). Abudefduf parasema Fowler 1918: 56, Fig. 22. (Type Distribution: Bismarck Archipelago; doubtful records locality, Philippines). Holotype: ANSP 47533. from Northern Solomon Islands and New Ireland as Paratypes: ANSP 47534-37 (4, in jar with holotype). this species is most likely confused with C. cymatilis Distribution: Solomon Islands, Northern New (Allen, 1999). Guinea, Philippines, and Ryukyu Islands. Chrysiptera springeri (Allen and Lubbock, 1976) Chrysiptera pricei Allen and Adrim, 1992 Glyphidodontops springeri Allen and Lubbock, 1976: Chrysiptera pricei Allen and Adrim, 1992: 103, Fig. 347, Fig. 1. (Type locality, off e. shore of Piru Bay, 1. (Type locality, Ampimoi Bay, near Arareni Point, Tandjung Liang, Ceram, Molucca Islands, Indonesia, 8 Yapen Island, Irian Jaya, Indonesia, ca. 1°53'S, m). Holotype: USNM 213577. Paratypes: BMNH 136°32'E, 2–5 m). Holotype: NCIP 4025. Paratypes: 1974.8.22.1-9 (9); BPBM 18618 (3); USNM 209562 NCIP 4026 (42), WAM P30391-001 (5). (45), 209838 (5); WAM P.25127-001 (3). Distribution: West Papua, Indonesia. Distribution: Moluccas, Flores, and Philippines. Chrysiptera rapanui (Greenfield and Hensley, 1970) Chrysiptera starcki (Allen, 1973) Abudefduf rapanui Greenfield and Hensley, 1970: Abudefduf starcki Allen, 1973a: 32, Fig. 1. (Type 693, Fig. 3. (Type locality, off the W. coast of Ahu locality, outer slope of Osprey Reef, Coral Sea, Akapu, Easter Island, SE. Pacific, 21 m). Holotype: 13°58'S, 146°41'E, 37 m). Holotype: AMS I.16477- USNM 204381. Paratypes: BPBM 6685–87 (1, 3, 1), 001. Paratypes: AMS I.16478-001 (1). 6689 (5); CAS 24690 (5); LACM 6560-11 (4); Distribution: New Caledonia, Loyalty Islands, Fiji, USNM 204382 (2). Coral Sea to Japan. Distribution: Easter Island and Kermadec Islands. Chrysiptera talboti (Allen, 1975) Chrysiptera rex (Snyder, 1909) Glyphidodontops talboti Allen, 1975a: 93, Pl. 3 (Fig. c). Abudefduf rex Snyder, 1909: 601. (Type locality, (Type locality, One Tree Island, Capricorn Group, Great Naha, Okinawa Island, Ryukyu Islands, Japan). Barrier Reef, Queensland, Australia, 23°30'S, 152°05'E, Holotype: USNM 62949. Paratypes: SU 21135 (14), 30 m). Holotype: AMS I.15684-004. Paratypes: AMS USNM 74535 (11). I.15486-038 (1), I.15637-009 (1), I.15647-001 (1), Chrysiptera personata Fowler, 1946: 153, Figs. 19– I.16479-001 (1), I.17464-001 (1), I.17472-001 (3) 20. (Type locality, Aguni Shima, Ryukyu Islands, I.17473-001 (1), I.17475-001 (1), I.17479-002 (1);

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

BMNH 1973.10.17.10-12 (3); BPBM 14564 (1); Genus Dascyllus Cuvier, 1829 MNHN 1973-60 (1); USNM 209745 (2), 210026 (2). Dascyllus Cuvier, 1829: 179. Masculine. Type species Distribution: Thailand to New Guinea, Great Barrier Chaetodon aruanus Linnaeus, 1758. Type by Reef to Tonga and Fiji Islands. monotypy. Pirene Gistel, 1848: IX. Feminine. Type species Chrysiptera taupou (Jordan and Seale, 1906) Chaetodon aruanus Linnaeus 1758. Type by being a Abudefduf taupou Jordan and Seale, 1906: 288, Pl. 43 replacement name. Unneeded replacement for (Fig. 3). (Type locality, Apia, Upolu Island, Western Dascyllus Cuvier, not preoccupied by Dascillus Samoa). Holotype: USNM 51741. Paratypes: BPBM Latreille, 1796 in Coleoptera Dascilloidea, Dascillidae. 5327 [ex USNM 51741] (1); SU 8717 (7), 8720 (3). Objectively invalid; nomen praeoccupatum, Chrysiptera elizabethae Fowler, 1955: 377, Fig. 1. preoccupied by Pirene Haliday, 1833 in Hymenoptera (Type locality, off Makaluva Island, near Suva, Fiji Chalcidoidea, Pteromalidae. Islands). Holotype: ANSP 71972. Paratypes: ANSP Tetradrachmum Cantor, 1849: 1223. Neutral. 71973-84 (12 or 11), ? BMNH (1). Chaetodon aruanus Linnaeus, 1758. Type by being a Distribution: Coral Sea, Vanuatu, Loyalty Islands, replacement name. Unneeded replacement for New Caledonia, Fiji, and Samoa. Dascyllus Cuvier 1829, not preoccupied by Dascylus Chrysiptera traceyi (Woods and Schultz, 1960) Agassiz, 1846 in Coleoptera. Pomacentrus traceyi Woods and Schultz in Schultz Pellochromis (subgenus of Pomacentrus) Fowler and et al., 1960: 114, Fig. 93. (Type locality, Lagoon Bean, 1928: 14. Masculine. Type species Pomacentrus coral head near N. end of Rongelap Island, Rongelap trimaculatus Rüppell, 1829. Type by original Atoll, W. Pacific, 18 ft). Holotype: USNM 141310. designation. Paratypes: USNM 141308–09 (1, 10). Semadascyllus Fowler, 1941: 257. Masculine. Type Distribution: Micronesia (Marshall Islands and species Dascyllus albisella Gill, 1862. Type by Caroline Islands) and Philippines. original designation. Tropical Indo-West Pacific; 11 species (Randall and Chrysiptera tricincta (Allen and Randall, 1974) Randall, 2001; Borsa et al., 2014). Glyphidodontops tricinctus Allen and Randall, 1974: 41, Fig. 3. (Type locality, Fagasa Bay, Tutuila, Dascyllus abudafur (Forsskål, 1775) American Samoa, 75 ft). Holotype: BPBM 11336. Chaetodon abudafur Forsskål, 1775: 65, xiii. (Type Paratypes: AMS I.17349-001 (1), I.17350-001 (1); locality, Jeddah, Saudi Arabia, Red Sea). No types USNM 211291 (1). known. Distribution: Southern Japan, Coral Sea, New Distribution: Red Sea, Indian Ocean: East and South Caledonia, Fiji, and Samoa. Africa, Seychelles, Comoros, Madagascar and Mascarenes East to Sunda Islands (Indonesia). Chrysiptera unimaculata (Cuvier, 1830) Comments: Resurrected from synonymy with D. Glyphisodon unimaculatus Cuvier in Cuvier and aruanus by Borsa et al. (2014). Valenciennes, 1830: 478. (Type locality, Timor Island, S. Malay Archipelago). No types known. Dascyllus albisella (Gill, 1862) Glyphidodon dispar Günther, 1862: 53. (Type Dascyllus albisella Gill, 1862: 149. (Type locality, locality, Ambon Island, Molucca Islands, Indonesia). Honolulu, Oahu Island, Hawaiian Islands, U.S.A). Syntypes: BMNH 1858.4.21.413-414 (2), Holotype: USNM 6274. Paratypes: MCZ 35982 [ex 1858.4.21.423 (1). USNM 6274] (1). Glyphidodon hemimelas Kner, 1868a: 30. (Type Dascyllus edmondsoni Pietschmann, 1934: 100. locality, Kandavu Island, Fiji Islands). Holotype: (Type locality, in corals at Kaaawa, Oahu Island, whereabouts unknown. Appeared in more detail in Hawaiian Islands). Holotype: BPBM 5312. Kner (1868b: 351, Pl. 8, fig. 25). Distribution: Hawaiian Islands and Johnston Islands. Glyphisodon filholi Sauvage, 1879: 207. (Type Dascyllus aruanus (Linnaeus, 1758) locality, Fiji Islands). Syntypes: MNHN 0000-9895 Chaetodon aruanus Linnaeus, 1758: 275. (Type locality, (2). "Habitat in Indiis" [Aru Islands]). No types known. Distribution: Red Sea and Western Indian Ocean to Neotype designation by Fricke (1999: 378) is invalid. Fiji and Tonga North to Ryukyu Islands. Chaetodon arcuanus Gmelin, 1789: 1250. Incorrect Chrysiptera uswanasi Allen, Erdmann and subsequent spelling of Chaetodon aruanus Linnaeus, Cahyani, 2018 1758. Chrysiptera uswanasi Allen, Erdmann and Cahyani, Pomacentrus emamo Lesson, 1831: 190. (Type 2018: 77, Figs. 2–4. (Type locality, Sebakor area, locality, Bora Bora Island, Society Islands). Syntypes: Fakfak Peninsula, West Papua Province, Indonesia, Whereabouts unknown. depth 10 meters). Holotype: MZB 24616. Paratypes: Dascyllus blochi Castelnau, 1875: 34. (Type locality, MZB 24617 (5); WAM P.34830-001 (4). Queensland, Australia). No types known. Distribution: Fakfak Peninsula, West Papua Pomacentrus trifasciatus De Vis, 1884: 452. (Type Province, Indonesia. locality, Probably South Seas). No types known.

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 63 Parenti 64

Pomacentrus devisi Jordan and Seale, 1906: 284. Dascyllus melanurus Bleeker, 1854 Nomen novum. Replacement name for Pomacentrus Dascyllus melanurus Bleeker, 1854a: 109. (Type trifasciatus De Vis 1884 regarded as a nomen locality, Banda Neira, Banda Island; Sumbawa praeoccupatum. Island; Priaman, Ulakan, W. Sumatra, Indonesia). Abudefduf caroli Curtiss, 1938: 117. (Type locality, Syntypes: (8) RMNH 6452 (2, very poor condition, Lagoon near Tautira, Tahiti, Society Islands). No in pieces). types known. Pomacentrus onyx De Vis, 1884: 451. (Type locality, Distribution: Red Sea and western Indian Ocean to South Seas). Syntypes: QM I.99 (1), I.10126–31 (1, Tuamotu Archipelago, Marquesas and Line Islands; 1, 1, 1, 1, 1). Sydney to Japan. Distribution: Indo-Australian Archipelago and Western Caroline Islands. Dascyllus auripinnis Randall and Randall, 2001 Dascyllus auripinnis Randall and Randall, 2001: 62, Dascyllus reticulatus (Richardson, 1846) Figs. 3–4. (Type locality, Tabuaera, Fanning Island, Heliases reticulatus Richardson, 1846: 254. (Type Line Islands). Holotype: BPBM 7537. Paratypes: locality, China Seas). Holotype: BMNH. AMS I.39964-001 (1); ASIZT 60474 (1); BMNH Dascyllus xanthosoma Bleeker, 1851: 247. (Type 2000.3.1.1 (1); BPBM 7703 (3), 16260 (2), 19575 locality, Banda Neira, Banda Islands, Indonesia). (1), 19941 (2); CAS 211523 (1); LACM 54140-1 (1); Holotype: RMNH 6455 (1 of 47). MNHN 2000-0515 (1); NSMT-P 59286 (1); ROM Pomacentrus unifasciatus Kner, 1868: 31. (Type 72216 (1); USNM 360267 (1). locality, Kandavu Island, Fiji Islands). Holotype: Distribution: Line Islands, Phoenix Island, and whereabouts unknown. Appeared in more detail in Penrhyn (Tongarevae), northern Cook Islands, where Kner (1868: 348, Pl. 8, fig. 24). Objectively invalid, is sympatric with D. trimaculatus. nomen praeoccupatum, preoccupied by Pomacentrus unifasciatus Steindachner 1867. Dascyllus carneus Fischer, 1885 Distribution: Cocos-Keeling Islands and Indo- Dascyllus carneus Fischer, 1885: 71, Pl. 2 (Fig. 5)]. Australian Archipelago to Southern Japan, Melanesia (Type locality, Mozambique). Lectotype: ZMH H62 to Polynesia, except Hawaiian Islands, Society [ex ZMH 3827]. Paralectotypes: ZMH H184 [ex Islands and Islands eastward of these groups. ZMH 3827] (2). Type catalog and lectotype designation: Ladiges et al. (1958: 164). Dascyllus strasburgi Klausewitz, 1960 Pomacentrus megalops Liénard in Sauvage, 1891: Dascyllus strasburgi Klausewitz, 1960: 45, Fig. 1. 234. (Type locality, Mauritius, Mascarenes, (Type locality, Haka Puuae Bay, Taipi Bay, Nuku southwestern Indian Ocean). Holotype: NHMI (not Hiva, Marquesas Islands, Pacific). Holotype: USNM found). Based on 'Pomacentre gros yeux' of Liénard 190596. Paratypes: SMF 5111–12 (1, 1); Inst. (1839: 34) and Liénard (1840: 297). Oceanogr. uncat. (5). Dascyllus nigripinnis Regan, 1908: 228, Pl. 24 (Fig. Distribution: Marquesas Islands. 5). (Type locality, Maldives, Indian Ocean). Dascyllus trimaculatus (Rüppell, 1829) Holotype: BMNH 1908.3.23.98. Pomacentrus trimaculatus Rüppell, 1829: 39, Pl. 8 Distribution: Southwestern Indian Ocean to (Fig. 3). (Type locality, Massawa, Eritrea, Red Sea). Andaman Sea; Seribu Islands (Java Sea). Lectotype: SMF 1445. Paralectotypes: SMF 2750 (1) Dascyllus flavicaudus Randall and Allen, 1977 and SMF 5113 (1) Massawa, MNHN 8128 (1) Red Dascyllus flavicaudus Randall [H. A.] and Allen, Sea. Lectotype established by Dor (1984: 184). 1977: 375, Fig. 11. (Type locality, Reef of Haurei Pomacentrus nuchalis Anonymous [Bennett], 1830: Bay, Rapa, ca. 27°45'S, 144°30'W, s. Pacific, 90 ft). 688. (Type locality, Sumatra, Indonesia). Possible Holotype: BPBM 13031. Paratypes: AMS I.19109- type: BMNH 1861.6.4.3. Based on Code 001 (2); BPBM 6886 (2), 7192 (4), 13102 (5), recommendation 51D, the anonymous author is listed 16568 (6), 16646 (5), 16926 (1); CAS 38525–33 (2, in square brackets. 1, 2, 3, 1, 1, 23, 1, 5); MNHN 1976-101 (2); USNM Dascyllus unicolor Bennett, 1831: 127. (Type 216381 (3); WAM P.25578-001 (2). locality, Mauritius, Mascarenes, southwestern Indian Distribution: Society Islands, Tuamotu Archipelago, Ocean). Holotype: BMNH 1855.12.26.474. Pitcairn Group and Rapa. Dascyllus niger Bleeker, 1847: 10. (Type locality, Jakarta, Java, Indonesia). Holotype: RMNH 6453 Dascyllus marginatus (Rüppell, 1829) (largest of 38). Pomacentrus marginatus Rüppell, 1829: 38, Pl. 8 Sparus nigricans Gronow in Gray, 1854: 61. (Type (Fig. 2). (Type locality, Massawa, Eritrea, Red Sea). locality, India). Holotype: BMNH 1853.11.12.103 Syntypes and/or Rüppell specimens: ?BMNH [Gronovius collection] (skin). 1845.10.29.94-95 (2); SMF 1498 (1), 2751 (1), Dascyllus axillaris Smith, 1935: 205, Pl. 22 (Fig. D). 27043 (6); MNHN 8130 (2); NMW 70324 (2). (Type locality, Great Fish Point, South Africa, Distribution: Red Sea and Gulf of Oman. western Indian Ocean). Holotype: SAIAB [formerly RUSI] [ex AMG] 284.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Distribution: Red Sea and western Indian Ocean to Dischistodus notophthalmus (Bleeker, 1853) Oceania, except Hawaiian Islands, Marquesas Islands Pomacentrus notophthalmus Bleeker, 1853c: 137. and Line Islands, where is replaced by D. albisella, D. (Type locality, Ternate, Molucca Islands, Indonesia). strasburgi and D. auripinnis, respectively (Randall Holotype: RMNH 6462. Bleeker specimens: BMNH and Randall, 2001); North to Japan South to Sydney. 1862.2.28.29 (1), NMV 46116 (1). Pomacentrus melanotus Bleeker, 1858: 13. (Type Genus Dischistodus Gill, 1863 locality, Manado, Sulawesi, Indonesia). Holotype: Dischistodus Gill, 1863: 214. Masculine. Type RMNH 27458 (114-mm spec. out of 5). species Pomacentrus fasciatus Cuvier, 1830. Type by Pomacentrus suluensis Seale, 1910: 519. (Type original designation. locality, Sitanki Island, Jolo Archipelago, Indo-Australian Archipelago; 7 species (Allen, 1991; Philippines). Holotype: BSMP 4689 (apparently Sin et al., 1995). destroyed). Dischistodus chrysopoecilus (Schlegel and Müller, Distribution: Great Barrier Reef to Southern Japan. 1840) Comments: ECoF (19 Nov. 2020) was wrong in Pomacentrus chrysopoëcilus Schlegel and Müller, declaring that “the name notophthalmus is apparently 1840: 21, Pl. 5 (Fig. 3). (Type locality, Java, not treated as valid since 1899, so the younger name Indonesia). Holotype: MNHN 8249. Possible types: is in use.” Allen (1991: 111) wrote Dischistodus RMNH D1277–79 (1, 1, 1, all dry), 942 (1). notophthalmus is the name previously used for this Pomacentrus notostigmus Richardson, 1844: species. As a matter of fact Yoshino (1982) used the unnumb. Page, Pl. 44 (Figs. 1–2). (Type locality, combination Dischistodus notophthalmus as a valid Questionably Southern Pacific). Holotype: BMNH species in reviewing Dischistodus species from the 1847.5.10.24. Ryukyu Islands, and before him Allen (1975: 131) Pomacentrus fasciatus var. intermedia Weber, 1913: used the name in his comprehensive account of the 340, Pl. 6 (fig. 4). (Type locality, Sanguisiapo, Sulu, damselfishes of the world. Philippines, Siboga station 93). Holotype: ZMA Dischistodus perspicillatus Cuvier, 1830 114611. Paratypes: ZMA 114612 (2). Pomacentrus pavo var. perspicillatus Cuvier in Distribution: Indo-Australian Archipelago. Cuvier and Valenciennes, 1830: 417. (No locality). Dischistodus darwiniensis (Whitley, 1928) Holotype: RMNH. Dascyllus fasciatus MacLeay, 1878: 361, Pl. 10 (Fig. Pomacentrus trimaculatus Cuvier in Cuvier and 2). (Type locality, Darwin, Northern Territory, Valenciennes, 1830: 427. (Type locality, Jakarta, Australia). Holotype: AMS I.16357-001 [ex MAMU Java, Indonesia). Syntypes: MNHN 0000-9795 (1, F825]. Subjectively invalid; secondarily preoccupied dry), 1986-0661 [ex MNHN 0000-9795] (3, dry). in Pomacentrus by Pomacentrus fasciatus Cuvier, Objectively invalid; preoccupied by Pomacentrus 1830; replaced by Pomacentrus darwiniensis trimaculatus Rüppell, 1829, replaced by Whitley, 1928. Pomacentrus dorsomaculatus Kendall and Pomacentrus darwiniensis Whitley 1928: 297. (Type Goldsborough, 1911. locality, Darwin, Northern Terr. Australia). Pomacentrus bifasciatus Bleeker, 1854a: 330. (Type Holotype: AMS I.16357-001 [ex MAMU F825]. locality, Larantuka, Flores Island, Lesser Sunda Replacement name for Dascyllus fasciatus Macleay, Islands, Indonesia). Holotype: RMNH 6457 (possibly 1878, secondarily preoccupied in Pomacentrus by 1 of 3). Bleeker specimens: BMNH 1862.2.28.108 Pomacentrus fasciatus Cuvier, 1830. (1), NMV 45935-36 (2). Distribution: Apparently distributed only in the Pomacentrus frenatus De Vis, 1885: 874. (Type Northern Territory, Australia. locality, Cardwell, Queensland, Australia). Holotype: Comments: Formerly considered a junior synonym of QM I.1366. D. fasciatus Cuvier by Allen (1991), but then it has Pomacentrus dorsomaculatus Kendall and been considered as valid (Allen, 1993; Sin et al., Goldsborough, 1911: 298 [footnote]. Nomen novum. 1995; Larson et al., 2013). In the ECoF (19 Nov. Replacement name for Pomacentrus trimaculatus 2020) Dascyllus fasciatus MacLeay is erroneously Cuvier 1830, preoccupied by Pomacentrus placed in the synonymy of Dischistodus fasciatus trimaculatus Rüppell 1829. (Cuvier 1830): the ECoF (Fricke et al., 2020) cites Chromis humbug Whitley, 1954: 23, Fig. 1. (Type Larson et al. (2013: 171) to support this opinion but locality, Green Island, off Cairns, Queensland, Larson et al. (2013) correctly places D. fasciatus as a Australia). Holotype: AMS IB.3116. synonym of darwiniensis. Distribution: Great Barrier Reef to China and Mariana Islands westward to Andaman Sea. Dischistodus fasciatus (Cuvier, 1830) Pomacentrus fasciatus Cuvier in Cuvier and Dischistodus prosopotaenia Bleeker, 1852 Valenciennes, 1830: 426, Pl. 134. (Type locality, Pomacentrus prosopotaenia Bleeker, 1852a: 67. Java, Indonesia). Syntypes: MNHN 8141 (1), 8503 (Type locality, Singapore). Holotype: RMNH 6466 (1); RMNH D1280 (1), 941 (2). (144 mm TL out of 13 specimens). Bleeker Distribution: Northwestern Australia to Philippines. specimens: NMV 46111 (1).

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 65 Parenti 66

Pomacentrus interorbitalis Günther, 1862: 30. (Type Chromanthias Whitley, 1935: 233. Masculine. Type locality, East Indian Archipelago). Holotype: BMNH species Chromanthias exilis Whitley, 1935. Type by 1862.2.28.39. original designation. Dischistodus cartieri Bleeker (ex Cartier), 1877c: 82. Distribution: Indo-West Pacific; monotypic. (Type locality, Cebu Island, Philippines). No types Lepidozygus tapeinosoma (Bleeker, 1856) known. Pomacentrus tapeinosoma Bleeker, 1856a: 376. Distribution: Nicobar Islands to Great Barrier Reef (Type locality, Ternate, Moluccas Islands, Indonesia). and Solomon Islands North Taiwan, Ryukyu Islands Lectotype: RMNH 28801. Paralectotypes: (25) and Hainan Island. ?BMNH 1862.2.28.38 (1), ?NMV 46481-42 (2), Dischistodus pseudochrysopoecilus (Allen and RMNH 6456 (16). Lectotype designated by Emery Robertson, 1974) (1983: 1330). Pomacentrus pseudochrysopoecilus Allen and Chromanthias exilis Whitley, 1935: 233. (Type Robertson, 1974: 160, Pls. 4, 5A. (Type locality, off locality, Nauru, Micronesia, South Pacific). N. side of Naura Islet, Fergusson Island, Holotype: AMS I.6681. Paratypes: AMS I.21042-001 D'Entrecasteaux Group, 9°36'S, 150°27'E, 2 m). [ex I.6681] (1). Holotype: BPBM 13608. Paratypes: AMS I.15633- Lepidozygus anthioides Smith, 1956: 888, Pl. 20 (fig. 006 (7), AMS I.16479-001 (3), QM I.6009 (1), C). (Type locality, Astove Island, Aldabra, western USNM 207936 (2). Indian Ocean). Holotype: SAIAB [formerly RUSI] Distribution: Philippines, New Guinea, Solomon 283. Paratypes: SAIAB [formerly RUSI] 935–945 (9, Islands., and Great Barrier Reef. 23, 1, 2, 1, 9, 10, 1, 1, 5, 12). Distribution: Western Indian Ocean to Tuamotu Genus Hemiglyphidodon Bleeker, 1877 Archipelago, Marquesas Islands and Line Islands. Hemiglyphidodon (subgenus of Glyphidodon) Bleeker, 1877c: 91, 93. Masculine. Type species Genus Mecaenichthys Whitley, 1929 Glyphisodon plagiometopon Bleeker, 1852. Type by Mecaenichthys Whitley, 1929: 218. Masculine. Type original designation, species Heliastes immaculatus Ogilby, 1885. Type Ctenoglyphidodon (subgenus of Abudefduf) Fowler, by original designation. 1918: 58. Masculine. Abudefduf melanopselion Distribution: Southeastern Australia; monotypic. Fowler, 1918. Type by original designation. Mecaenichthys immaculatus (Ogilby, 1885) Indo-Australian Archipelago; monotypic. Heliastes immaculatus Ogilby, 1885: 446. (Type Hemiglyphidodon plagiometopon (Bleeker, 1852) locality, Shark Reef, Port Jackson, N.S.W. Australia). Glyphisodon plagiometopon Bleeker, 1852a: 67. Holotype: AMS B.7452. Paratypes: (several) ?AMS (Type locality, Singapore). Holotype: ?BMNH B.7450-51 (1, 1 1862.2.28.70. Bleeker specimens: RMNH 6499 (2). Distribution: Coast of New South Wales. Glyphisodon batjanensis Bleeker, 1855a: 373. (Type Genus Microspathodon Günther, 1862 locality, Batjan, Indonesia). Holotype: ? RMNH Microspathodon Günther, 1862: 35, 57. Masculine. whereabouts unknown. Bleeker specimens: ?BMNH Type species Glyphisodon chrysurus Cuvier, 1830. 1862.2.28.32 (1). Type by monotypy. Abudefduf melanopselion Fowler, 1918: 59, Fig. 23. Pomataprion Gill, 1863: 216. Masculine. Type (Type locality, Philippines). Holotype: ANSP 47538. species Hypsypops dorsalis Gill, 1862. Type by Distribution: Thailand and China to Timor Sea subsequent designation. Type designated by Jordan eastward to Solomon Islands and Great Barrier Reef. and Evermann (1898: 1565). Genus Hypsypops Gill, 1861 Azurella Jordan, 1919b: 341. Feminine. Type species Hypsypops Gill, 1861: 165. Masculine. Type species Pomacentrus bairdii Gill, 1862. Type by original Glyphisodon rubicundus Girard, 1854. Type by designation. monotypy. Centrochromis Norman, 1922: 534. Masculine. Type Distribution: Eastern Pacific; one species. species Glyphidodon rudis Poey, 1860. Type by original designation Hypsypops rubicundus (Girard, 1854) Distribution: Atlantic and Eastern Pacific; four Glyphisodon rubicundus Girard, 1854: 148. (Type species. locality, Monterey, California, U.S.A). Syntypes: MCZ 14825 (1), USNM 484 (2). Microspathodon bairdii (Gill, 1862) Distribution: Monterrey Bay to western Baja Pomacentrus bairdii Gill, 1862: 149. (Type locality, California and Guadalupe. Cape San Lucas, Baja California, Mexico). Syntypes: (orig. 2) ?BMNH 1862.5.21.16 [ex USNM] (1), Genus Lepidozygus Günther, 1862 USNM 3656 (orig. 4, now 2). Lepidozygus Günther, 1862: 15. Masculine. Type Distribution: Gulf of California to Ecuador, including species Pomacentrus tapeinosoma Bleeker, 1856. Revillagigedo and Galapagos Islands. Type by monotypy.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Microspathodon chrysurus (Cuvier, 1830) Neoglyphidodon carlsoni (Allen, 1975) Glyphisodon chrysurus Cuvier in Cuvier and Paraglyphidodon carlsoni Allen, 1975b:179, Valenciennes, 1830: 476. (Type locality, St. Thomas unnumb. Fig. (Type locality, Leeward side of Island). Holotype: MNHN A.8125. Yaukuve Levu Island, Great Astrolabe Lagoon, Fiji Pomacentrus denegatus Poey, 1860: 190. (Type Islands, 1–3 m). Holotype: AMS I.17689-001. locality, Cuba). Syntypes: MCZ 14672 (1), 14675 Paratypes: AMS I.17689-002 (4), I.17690-001 (1), (1). I.17691-001 (1). Glyphisodon rudis Poey, 1860: 191. (Type locality, Distribution: Fiji Islands. Cuba). Poey specimens: MCZ 14787 (2), ZMB 5936 Neoglyphidodon crossi Allen, 1991 (1). Neoglyphidodon crossi Allen, 1991: 118, 221, Pomacentrus niveatus Poey, 1876: 144. (Type unnumb. Fig. (Type locality, Ambon Island, Ambon locality, Cuba). Holotype: ?USNM 37455. Bay, Moluccas Islands, Indonesia, 3–4 m). Holotype: Mycrospathodon [sic] fowleri Silvester, 1915: 216. LIPI NCIP 3997. Paratypes: WAM P.25230-007 (4), (Type locality, Coral reef off Guanica harbor, Puerto USNM 288352 (3). Rico, Caribbean Sea). Holotype: PU 3060. Paratypes: Distribution: Sulawesi and Moluccas. PU (5). Distribution: Bermuda, southern Florida and Neoglyphidodon melas (Valenciennes, 1830) Caribbean Sea to Eastern Brazilian coast. Glyphisodon melas Valenciennes (ex Kuhl and van Hasselt) in Cuvier and Valenciennes, 1830: 472. Microspathodon dorsalis (Gill, 1862) (Type locality, Java, Indonesia). Syntypes: RMNH Hypsypops dorsalis Gill, 1862: 147. (Type locality, D1283 (1, stuffed), D1284 (1, stuffed). Cape San Lucas, Baja California, Mexico). Holotype: Glyphisodon ater Valenciennes (ex Ehrenberg) in USNM. Cuvier and Valenciennes, 1830: 473. (Type locality, Pomacentrus quadrigutta Gill, 1862: 149. (Type Massawa, Eritrea, Red Sea). Syntypes: ZMB 2730 (1). locality, Cape San Lucas, Baja California, Mexico). Glyphisodon melanopus Bleeker, 1856b: 82. (Type Syntypes: BMNH 1862.5.21.17 [ex USNM] (1), locality, Ora Malang, southeastern Java, Indonesia). USNM 3657 (2). Syntypes: (2) BMNH 1862.2.28.25 (1). Microspathodon cinereus Gilbert, 1890: 71. (Type Glyphisodon violaceus Brevoort, 1856: 264. (Type locality, Clarion and Socorro Islands). Holotype: locality, Ryukyu Islands, Japan). No types known. Whereabouts unknown. Glyphisodon xanthonotus Bleeker, 1859b: 357. (Type Microspathodon azurissimus Jordan and Starks in locality, Sancapura, Bawean, Indonesia). Holotype: Jordan, 1895: 478, Pl. 44. (Type locality, Venados RMNH 6493. Islands, near Mazatlán, Sinaloa, western Mexico). Abudefduf rhomaleus Snyder, 1911: 534. (Type Syntypes: CAS-SU 2895 (1), 11610 (1, missing), locality, Naha, Okinawa Island, Ryukyu Islands, 11636 (1, missing). Japan). Holotype: USNM 68233. Paratypes: CAS-SU Distribution: Gulf of California to Colombia, 21422 (1). including Galapagos Islands. Distribution: Red Sea and Western Indian Ocean to Microspathodon frontatus Emery, 1970 Indo-Australian Archipelago and Vanuatu North to Microspathodon frontatus Emery, 1970: 294, Figs. 1, Ryukyu Islands. 3. (Type locality, Annobón Island, Isla la Piramide Neoglyphidodon mitratus Allen and Erdmann, 2012 near town of Ambo, Equatorial Guinea, Gulf of Neoglyphidodon mitratus Allen and Erdmann, 2012: Guinea). Holotype: USNM 202500. Paratypes: 1136, Figs. 1–2. (Type locality, Bairoko Inlet, ANSP 111216 (5); FMNH 74356–8 (24, 5, 6); UF 8°11.008'S, 157°16.622'E, New Georgia Island, 223693 [ex UMML 23693] (9), ?223984 [ex UMML Solomon Islands, depth 10 meters). Holotype: WAM 23984] (2); USNM 202501 (3). P.33480-001. Paratypes: WAM P.33480-002 (2). Distribution: Gulf of Guinea to Ghana, including Distribution: Western Pacific: Palau; Papua New Annobón Island and São Tomé and Principe. Guinea; Solomon Islands. Genus Neoglyphidodon Allen, 1991 Neoglyphidodon nigroris (Cuvier, 1830) Neoglyphidodon Allen, 1991: 246. Masculine. Type Glyphisodon nigroris Cuvier in Cuvier and species Glyphisodon melas Valenciennes, 1830. Type Valenciennes, 1830: 485. (No locality). Holotype: by original designation. MNHN 8285. Distribution: Tropical Indo-West Pacific; nine Glyphisodon behni Bleeker, 1847a: 25. (Type species. locality, Jakarta, Java, Indonesia). Syntypes: BMNH Neoglyphidodon bonang (Bleeker, 1852) 1862.2.28.17 (1), RMNH 6494 (1). Also in Bleeker Glyphisodon bonang Bleeker, 1852d: 582. (Type (1847b: 156). locality, Padang, Sumatra, Indonesia). Syntypes: (2) Glyphisodon xanthurus Bleeker, 1853g: 345. (Type ?BMNH 1862.2.28.30 (1), RMNH 6486 (largest of 5). locality, Ambon Island, Molucca Islands, Indonesia). Distribution: Sri Lanka to Indonesia and Solomon Holotype: RMNH 6491. Bleeker specimens: BMNH Islands. 1862.2.28.31 (1), ?NMV 45853 (1).

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 67 Parenti 68

Abudefduf coracinus Seale, 1910: 520, Pl. 13. (Type Neopomacentrus anabatoides (Bleeker, 1847) locality, Sitanki Island, Jolo Archipelago, Glyphisodon anabatoides Bleeker, 1847a: 28. (Type Philippines). Holotype: BSMP 4908 (destroyed). locality, Jakarta, Java, Indonesia). Syntypes: RMNH Abudefduf filifer Weber, 1913: 348, Fig. 74. (Type 6502 (3 of 12). Bleeker specimens: BMNH locality, Feer, Hoch-Kei Island, Indonesia, Siboga 1862.2.28.18 (1), NMV 45906 (1). Appeared first as station 263). Holotype: ZMA 109435. nomen nudum in Bleeker 1845: 523. Described as Chromis bitaeniatus Fowler and Bean, 1928: 56, Pl. new also in Bleeker 1847b: 157. 6. (Type locality, Maricaban Island, Philippines). Distribution: Indo-Australian Archipelago: East Holotype: USNM 89956. Andaman Sea, Malaysia, Singapore, South China Distribution: Andaman Sea to Northern Australia, Sea, Indonesia and Philippines. Solomon Islands, and Vanuatu North to Ryukyu Islands. Neopomacentrus aquadulcis Jenkins and Allen, 2002 Neoglyphidodon oxyodon Bleeker, 1858 Jenkins and Allen, 2002: 379, Fig. 1. (Type locality, Glyphisodon oxyodon Bleeker, 1858: 14. (Type Mouth of freshwater stream at Lawadi Village, Milne locality, Manado, Sulawesi, Indonesia). Holotype: Bay Province, Papua New Guinea, 10°15.81'S, Whereabouts unknown. Bleeker specimens: BMNH 150°43.14'E, depth 1 meter). Holotype: WAM P.31636- 1862.2.28.20 (1). 001. Paratypes: AMS I.37954-001 (30), WAM Distribution: Indonesia, Philippines, and Ashmore P.31636-002 (9). Reef (Timor Sea). Distribution: Papua New Guinea from near East Cape, the easternmost extension of mainland and Cape Roon on Neoglyphidodon polyacanthus (Ogilby, 1889) the Morobe coast; in the lower, tidal reaches of Glyphidodon polyacanthus Ogilby, 1889: 65. (Type freshwater streams in either pure fresh or brackish water. locality, Lord Howe Island Syntypes: AMS I.1903- I.1910 (1 ea.). Neopomacentrus azysron (Bleeker, 1877) Distribution: Capricorn Group (southern Great Pomacentrus (Pomacentrus) azysron Bleeker, 1877c: Barrier Reef), New Caledonia, Lord Howe Islands, 50. (Type locality, Ambon Island, Moluccas Islands, Norfolk Islands. Indonesia). Holotype: Whereabouts unknown. Also appeared in Bleeker (1877b: Pl. 410). Neoglyphidodon thoracotaeniatus (Fowler and Bean, Abudefduf melanocarpus Fowler and Bean, 1928: 182. 1928) (Type locality, Cabugao Bay, Catanduanes Island, Abudefduf thoracotaeniatus Fowler and Bean, 1928: Philippines). Holotype: USNM 95680 [published as 158, Pl. 14. (Type locality, Biri Channel, Philippines). 89965]. Paratypes: USNM 95678–79 (1, 1). Holotype: USNM 89964. Paratypes: (19) ANSP Distribution: Western Indian Ocean to northern 80427 [ex USNM 95683] (5); Philippines Gov't. [ex Australia, Southern New South Wales (Australia) and USNM 95683] (3); USNM 95683 (now 0, 2 New Caledonia, Northward to Philippines, Taiwan, unaccounted for), 95689 (9), ?95696 (1). Melanesia, except Fiji. Distribution: Indo-Australian Archipelago: Indonesia, the Philippines, New Guinea, and the Solomon Neopomacentrus bankieri (Richardson, 1846) Islands. Glyphisodon bankieri Richardson, 1846: 253. (Type locality, Hong Kong, China Seas). Syntypes: BMNH Genus Neopomacentrus Allen, 1975 1848.3.19.113-114 (2). Neopomacentrus Allen, 1975b: 39, 166. Masculine. Distribution: South China Sea, Java Sea and eastern Type species Glyphisodon anabatoides Bleeker, Queensland to southern Papua New Guinea. 1847. Type by original designation. Original type species misidentified; type fixed by Randall et al. Neopomacentrus cyanomos (Bleeker, 1856) (2005: 92). Pomacentrus cyanomos Bleeker 1856b: 89. (Type Tropical Indo-West Pacific on coral reefs with two locality, Jakarta, Java, Indonesia). Holotype: species (aquadulcis and taeniurus) inhabiting Whereabouts unknown. Bleeker specimens: BMNH brackish and freshwaters; 16 species. 1862.2.28.119. Pomacentrus leucosphyrus Fowler, 1904: 533, Pl. 19 Neopomacentrus aktites Allen, Moore and Allen, 2017 (lower left fig.). (Type locality, Padang, Sumatra, Neopomacentrus aktites Allen, Moore and Allen, 2017: Indonesia). Holotype: ANSP 27673. 3, Figs. 1–4. (Type locality, Western Australia, Dampier Pomacentrus prateri Fowler, 1928: 117, Pl. 2. (Type Archipelago, Flying Foam Passage, depth 2–9 meters). locality, Back Bay, Mumbai, India). Holotype: ANSP Holotype: WAM P.34738-009. Paratypes: WAM 53936. P.26668-007, P.30311-018, P.30312-011, P.30391-010, Distribution: Red Sea, East Africa, Persian Gulf, P.31812-015, P.34736-020, P.34737-007, P.34739-004, Socotra and Madagascar to Philippines, New Ireland P.34742-013, P.34743-009, P.34750–009 (6, 3, 8, 6, 6, (Papua New Guinea) and Vanuatu, North to Southern 7, 2, 4, 1, 5, 1). Japan to Northern Australia, New Caledonia and Distribution: Western Australia from Shark Bay Melanesia, except Fiji. Invasive at Campeche Bank, Northward to the Kimberley coast in the far Northern Gulf of Mexico. In April 2016 it was found to occur portion of the state. both on the reefs and the adjacent oil-loading

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831 platform. During the August 2016 expedition this species from the western Indian Ocean represents a species was recorded in small aggregations (up to senior synonym of the Indo-West Pacific species ~150 fish, but usually about several dozen, including taeniurus. many juveniles) at all dive sites on the reef complex. Neopomacentrus fuliginosus (Smith, 1960) It also formed a large, dense aggregation of many Abudefduf fuliginosus Smith, 1960: 339, Fig. 6. tens of thousands of individuals on the oil-loading (Type locality, Mozambique). Holotype: RUSI 61. platform (Robertson et al., 2016). Paratypes: RUSI 792–793 (1, 2). Neopomacentrus filamentosus (MacLeay, 1882) Distribution: Mozambique to Kenya. Gliphidodon filamentosus Macleay, 1882: 365. (Type Neopomacentrus metallicus (Jordan and Seale, 1906) locality, New Guinea). Holotype: MAMU Abudefduf metallicus Jordan and Seale, 1906: 289, (whereabouts unknown). Fig. 51. (Type locality, Apia, Upolu Island, Western Distribution: Andaman Sea and Indo-Australian Samoa). Holotype: USNM 51742. Paratypes: AMS Archipelago, Thailand to New Caledonia. I.7373 (1), BPBM 5328 [ex USNM 51742] (1), SU Neopomacentrus flavicauda Parenti nomen novum 8721 (9), USNM 51742 (5, with holotype). Neopomacentrus xanthurus Allen and Randall, 1980: Distribution: Tonga, Samoa and Fiji Islands. (6) 54, Figs. 36–37. (Type locality, Suakin harbor, Neopomacentrus miryae Dor and Allen, 1977 off marine lab jetty, Sudan, Red Sea, 0.5 m). Neopomacentrus miryae Dor and Allen, 1977: 183, Holotype: BPBM 17908. Paratypes: BMNH Fig. 1. (Type locality, Dahab, Sinai Peninsula, Gulf 1976.9.20.10-13 (4); BPBM 20432 (1), 20473 (1); of Aqaba, Red Sea, 10 m). Holotype: BPBM 20322. HUJ 8371 (1), MNHN 1976-117 (5); TAU P.3780 Paratypes: BMNH 1976.5.4.1-2 (2); BPBM 14327 (7); USNM 216479 (106); WAM P.25618-001 (5), (1), 14666 (1), 18208 (2); HUJ 4825 (3); MNHN P.25790-005 (2), P.25794-003 (8), P.25795-001 (4). 1976-8 (1); SMF 19678 (3); TAU 3225 (11); USNM Distribution: Southern Red Sea to Gulf of Aden. 215463 (2), 216432 (31); WAM P.25523-002 (1). Comments: Allen (1991) placed Dascyllus xanthurus Distribution: Red Sea and Socotra. Bleeker, 1853 in Neopomacentrus, thereby making Neopomacentrus xanthurus Allen and Randall a Neopomacentrus nemurus (Bleeker, 1857) secondary junior homonym that is herein replaced. I Glyphisodon nemurus Bleeker, 1857b: 73. (Type propose Neopomacentrus flavicauda as a new name locality, Kajeli, Buru Island, Moluccas Islands; Ambon for Neopomacentrus xanthurus Allen and Randall, Island, Moluccas Islands, Indonesia). Syntypes: (3) 1980. Etymology: flavicauda refers to a yellow BMNH 1862.2.28.24 (1), RMNH 6503 (2). caudal fin, as does xanthurus. Distribution: Indo-Australian Archipelago. Neopomacentrus fallax (Peters, 1855) Neopomacentrus sindensis (Day, 1873) Glyphidodon fallax Peters, 1855a: 456. (Type locality, Glyphidodon sindensis Day, 1873a: cclxiii. (Type Mozambique). Syntypes: ?BMNH 1861.5.2.67 (1), locality, Karachi, Pakistan). Syntypes and/or Day ZMB 2739 (3). specimens: AMS I.144 (1) Sind; BMNH Pomacentrus taeniurus Bleeker, 1856c: 51. (Type 1889.2.1.3944-3948 (5); MZUF 4000 (1); NMW locality, Ambon Island, Moluccas Islands, Indonesia). 70271 (1); ZSI F1619 (1), 1620-21 (2). Holotype: Whereabouts unknown. Bleeker specimens: Distribution: Persian Gulf and Arabian Sea. BMNH 1862.2.28.110 (1). Glyphidodon amboinensis Bleeker, 1857a: 72. (Type Neopomacentrus sororius Randall and Allen, 2005 Neopomacentrus sororius Randall and Allen, 2005: locality, Ambon Island, Molucca Islands, Indonesia). 77, Figs. 5–7. (Type locality, Lively Rocks, Holotype: Whereabouts unknown. Trincomalee, Sri Lanka, depth 8 meters). Holotype: Glyphidodon cochinensis Day, 1865: 38. (Type BPBM 18080. Paratypes: BPBM 17113 (1), 27715 locality, Cochin, India). Syntypes (3) and/or Day specimens: BMNH 1870.6.14.6 (1), 1889.2.1.4293-95 (1), 28050 (1), 37801 (1); BMNH 2005.5.59 [sic] (1); (3, dry), 1870.6.14.6 (1) Andamans. USNM 301414 (11), 301447 (6), 301443 (31), 363814 (2), 363820 (151), 382249 (1); WAM Pomacentrus rathbuni Jordan and Snyder, 1901: 754, Pl. 34. (Type locality, near Yokohama, doubtless from P.26504-005 (1), P.26507-013 (1), P.26509-003 (2), Misaki or Boshu, Japan). Holotype: USNM 49706. P.26551-002 (1), P.26515-002 (1). Distribution: East Africa, Comoros and Madagascar Paratypes: CAS-SU 6464 (1), 6659 (4). Pomacentrus inhacae Smith, 1955: 894. (Type East to Sumatra (Indonesia). locality, Inhaca Island, Mozambique, 26°S, 33°E, Neopomacentrus violascens (Bleeker, 1848) western Indian Ocean). Holotype: SAIAB [formerly Pristotis violascens Bleeker 1848: 637. (Type RUSI] 285. Paratypes: many. locality, Bima, Sumbawa Island, Lesser Sunda Distribution: Mozambique north to India, Sri Lanka to Islands, Indonesia). Syntype or holotype: RMNH Philippines and Ryukyu Islands south to southeastern 6463 (1 or more of 11). Australia and Solomon Islands. Dascyllus xanthurus Bleeker, 1853b: 117. (Type Comments: Examination of the syntypes ZMB 2739 locality, Ambon Island, Molucca Islands, Indonesia). reveals that fallax, previously regarded as a valid Holotype: RMNH 6463 (1 of 11).

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 69 Parenti 70

Distribution: Indo-Australian Archipelago: East Parma mccullochi Whitley, 1929 Andaman Sea, Indonesia, Philippines, Papua New Parma mccullochi Whitley, 1929: 228. (Type locality, Guinea East to New Caledonia. Rottnest Island, Western Australia). Holotype: AMS I.13144. Genus Nexilosus Heller and Snodgrass, 1903 Distribution: Recherche Archipelago to Abrolhos Nexilosus Heller and Snodgrass, 1903: 204. Islands (Southwestern Australia). Masculine. Type species Nexilosus albemarleus Heller and Snodgrass, 1903. Type by monotypy. Parma microlepis Günther, 1862 Distribution: Eastern Pacific. One species. Parma microlepis Günther, 1862: 57. (Type locality, Port Jackson, N. S. W., Australia). Syntypes: BMNH Nexilosus latifrons (Tschudi, 1846) 1859.6.3.22 (1), 1859.6.3.25-27 (3). Pomacentrus latifrons Tschudi, 1846: 17. (Type Parma squamipinnis Günther, 1862: 58, 505. (Type locality, Huacho, Peru). Holotype: ZMB 2764. locality, Australia). Syntypes: (3) BMNH 1848.10.25.41a Nexilosus albemarleus Heller and Snodgrass, 1903: (1, stuffed), 1855.9.19.288 (1). 204, Pl. 8. (Type locality, Tagus Cove, Isabela Island Pomacentrus semifasciatus Günther, 1881: 226, Pl. [Albemarle Island], Galápagos Islands). Holotype: 125 (Fig. F). (Type locality, Boston Island, Marshall CAS-SU 6359. Paratypes: CAS-SU 6535 (10 young), Islands). Holotype: BMNH 1873.4.3.97. 12290 (1, missing in 1995), 12291-92 (1, 1), 12293 Glyphidodon (Parma) australis Steindachner, 1867: (1, missing in 1995); USNM 61167 (3, not found in 328. (Type locality, Port Jackson, New South Wales, 1980). Australia). Holotype: NMW 70293. Distribution: Peru to Northern Chile and Galapagos Distribution: Byron Bay (New South Wales) to Port Islands. Phillip Bay (Victoria). Genus Parma Günther, 1862 Parma occidentalis Allen and Hoese, 1975 Parma Günther, 1862: 57. Feminine. Type species Parma occidentalis Allen and Hoese, 1975: 283, Parma microlepis Günther, 1862. Type by subsequent Figs. 1E, 2F, 15, 16. (Type locality, Nancy Cove, designation by Jordan (1919: 318). Rottnest Island, Western Australia). Holotype: WAM Actinochromis Bleeker, 1877: 5, 6, 142, 166. P.10883. Paratypes: CSIRO A 1491-99 (9), A.1508- Masculine. Type species Heliastes lividus Klunzinger, 11 (4), C 2684 (1), C 2686 (1), C 2688-91 (4); WAM 1872. Type by original designation. P.10877 (1), P.10882 (1), P.10988 (1), P.10940 (1), Temperate seas of Australia and New Zealand. Ten P.13743 (1), P.24817 (2). species. Distribution: Cape Leeuwin to Coral Bay (Western Parma alboscapularis Allen and Hoese, 1975 Australia). Parma alboscapularis Allen and Hoese, 1975: 279, Parma oligolepis Whitley, 1929 Figs. 1G, 2G, 13, 14. (Type locality, off Erskine Parma oligolepis Whitley, 1929: 230, Pl. 28 (Fig. 1). Valley Stream, Lord Howe Island, 8 m). Holotype: (Type locality, Cape Moreton, Queensland, Australia). AMS I.17361-008. Paratypes: AMS I.17360-023 (1), Holotype: QM I.2536. Paratypes: QM I.2171 (1), I.17393-009 (1), I.17411-010 (1); CMC F755 (1); I.3429 (1). NMNZ 3755 (2), 4097 (1), 5272 (1). Distribution: Cape Tribulation (Queensland) to Sydney Distribution: Lord Howe, Norfolk, and Kermadec (New South Wales). Islands and New Zealand. Parma polylepis Günther, 1862 Parma bicolor Allen and Larson, 1979 Parma polylepis Günther, 1862: 59. (Type locality, Parma bicolor Allen and Larson, 1979: 12, Figs. 1, Norfolk Island). Holotype: BMNH 1855.9.19.272. 3, 4. (Type locality, Mondrain Island, Recherche Distribution: Capricorn Group to Bass Point (New Archipelago, Western Australia, ca. 34°07'S, South Wales), Lord Howe Islands, Norfolk Islands, 122°15'E, 10–20 m). Holotype: WAM P.26045-001. New Caledonia, and Northern New Zealand. Paratypes: AMS I.20701-001 [ex AMS I.20221] (1); WAM P.25468-001 (1), P.25762-005 (1), P.26045- Parma unifasciata (Steindachner, 1867) 002 (1), P.26046-001 [not P.26046-007] (1). Pomacentrus unifasciatus Steindachner, 1867: 326. Distribution: Recherche Archipelago to Rottnest (Type locality, Port Jackson, N.S.W., Australia). Islands (Southwestern Australia). Holotype: NMW 75807. Distribution: Noosa Head (Queensland) to Montague Parma kermadecensis Allen, 1987 Islands (New South Wales). Parma kermadecensis Allen, 1987b: (266) 267, Figs. 1–4. (Type locality, Te Konui Point Boat Coce, Parma victoriae (Günther, 1863) Raoul Island, Kermadec Islands, ca. 29°17'S, Glyphidodon victoriae Günther, 1863: 115 [2]. (Type 177°53'W, 20 m). Holotype: NMNZ P17849. locality, Victoria, South Australia). Holotype: BMNH. Paratypes: NZOI P967 (1), WAM P.28425-002 (2). Heliastes lividus Klunzinger, 1872: 36. (Type locality, Distribution: Kermadec Islands. Port Phillip, Victoria, Australia). Holotype: SMNS 1716.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Parma viola Whitley, 1929: 231, Pl. 28 (fig. 2). (Type Plectroglyphidodon johnstonianus Fowler and Ball, locality, Tasmania, Australia). Holotype: AMS I.6611. 1924 Distribution: Southern Australia, Tasmania to Plectroglyphidodon johnstonianus Fowler and Ball, Dongara, western Australia. 1924: 271. (Type locality, Johnston Island, Pacific). Holotype: BPBM 3418. Paratypes: ANSP 85858 (1). Genus Plectroglyphidodon Fowler and Ball, 1924 Plectroglyphidodon nitidus Smith, 1956: 892, Pl. 21 Negostegastes Whitley, 1929: 225. Masculine. Type (Fig. B). (Type locality, Aldabra Island, Western species Glyphisodon leucozona Bleeker, 1859. Type Indian Ocean). Holotype: SAIAB [formerly RUSI] by original designation. 291. Paratypes: (numerous). Plectroglyphidodon Fowler and Ball, 1924: 271. Distribution: Western Indian Ocean to Pitcairn Group, Masculine. Type species Plectroglyphidodon Line Islands and Hawaiian Islands; Australia to Japan. johnstonianus Fowler and Ball, 1924. Type by original designation. Plectroglyphidodon lacrymatus (Quoy and Gaimard, Oliglyphisodon Fowler, 1941: 266. Masculine. Type 1825) species Glyphisodon imparipennis Vaillant and Glyphisodon lacrymatus Quoy and Gaimard, 1825: Sauvage, 1875. Type by original designation. 388, Pl. 62 (Fig. 7). (Type locality, Guam, Mariana Pterocyclosoma Fowler, 1941: 264. Neutral. Type Islands, W. Pacific). Holotype: MNHN 0000-8336. species Glyphisodon sindonis Jordan and Evermann, Glyphisodon nivosus Hombron and Jacquinot in 1903. Type by original designation. Jacquinot and Guichenot, 1853: 50, Pl. 5 (Fig. 5). Distribution: Tropical Indo-West Pacific. Ten species. (No locality). No types known. Glyphidodon florulentus Günther, 1862: 49. (Type Plectroglyphidodon dickii (Liénard, 1839) locality, Indian Ocean). Holotype: BMNH 1857.2.17.5. Glyphisodon dickii Liénard, 1839: 35. (Type locality, Distribution: Red Sea and Western Indian Ocean to Mauritius). Holotype: NHMI (not found). Marshall and Society Islands; Australia to Ryukyu Glyphidodon unifasciatus Kner and Steindachner, 1867: Islands; Lord Howe Island, New Caledonia and Rapa. 375, Pl. 2 (Fig. 3). (Type locality, Upolu, Samoa). Syntypes: MGH 1631. Plectroglyphidodon leucozonus (Bleeker, 1859) Distribution: Western Indian Ocean to Tuamotu Glyphisodon leucozona Bleeker, 1859c: 338. (Type Archipelago and Line Islands.; Australia North to Japan. locality, Karangbollong, Java, Indonesia). Syntypes: (5) BMNH 1862.2.28.23 and 26 (2, smaller possible type), Plectroglyphidodon flaviventris Allen and Randall, ?NMV 46437 (1), RMNH 6495 (probably 4 of 6). 1974 Glyphidodon cingulum Klunzinger, 1871: 526. (Type Plectroglyphidodon flaviventris Allen and Randall, locality, Al-Qusair, Red Sea Governorate, Egypt, Red 1974: 44, Fig. 4. (Type locality, Outer reef at Sea). Syntypes: SMF 1208 (2, apparently lost), ZIN Rangiroa, 200 m East of Tiputa Pass, Tuamotu 2649 (1), ZMB 8034 (2). Archipelago, 30 ft). Holotype: BPBM 11258. Abudefduf corneyi Jordan and Dickerson, 1908: 613, Paratypes: AMS I.17351-001 (2); BPBM 13997 (1), Fig. 5. (Type locality, Suva, Fiji Islands). Holotype: 11259-61 (2, 2, 1). USNM 61678. Distribution: Tuamotu Archipelago. Abudefduf atrapinna Seale, 1935: 367, Pl. 22 (Fig. 2). Plectroglyphidodon imparipennis (Vaillant and (Type locality, Sikaina Island, Solomon Islands). Sauvage, 1875) Holotype: CAS 5511. Paratypes: CAS 5512-13 (2, in Glyphisodon imparipennis Vaillant and Sauvage, 1875: one jar). 279. (Type locality, Honolulu, Oahu Island, Hawaiian Chrysiptera yamashinai Okada and Ikeda, 1937: 82, Islands, U.S.A). Holotype: MNHN 000-8040. Fig. 1; Pl. 3 (Fig. 2). (Type locality, Miyako Island, Chromis elaphrus Jenkins, 1903: 457, Fig. 23. (Type Ryukyu Islands, Japan). Holotype: Okada coll. spec. locality, Honolulu, Oahu Island, Hawaiian Islands). 2255 (lost). Holotype: USNM 50703. Paratypes: (8). Abudefduf melanozonatus Aoyagi, 1941b: 234, Fig. Abudefduf iwasaki Okada and Ikeda, 1939: 170, Fig. 7. 35. (Type locality, Kowan, Okinawa Island, Ryukyu (Type locality, Ishigaki Island, Yaeyama Islands, Islands, Japan). Syntypes: YCM-P 30469-70 (1, 1). southern Ryukyu Islands, Japan). Holotype: Okada Distribution: Red Sea and western Indian Ocean to coll. spec. 2589 (lost). Marshall and Pitcairn Group; Australia to Japan. Chrysiptera prughi Fowler, 1946: 156, Figs. 23–24. Plectroglyphidodon phoenixensis (Schultz, 1943) (Type locality, Aguni Shima, Ryukyu Islands, Japan). Abudefduf phoenixensis Schultz 1943: 190, Fig. 15. Holotype: ANSP 72028. (Type locality, Reef of Enderbury Island, Phoenix Oliglyphisodon caeruleomaculatus Fowler, 1946: 158, Island). Holotype: USNM 115782. Paratypes: MCZ Figs. 27–28. (Type locality, Aguni Shima, Ryukyu 37268 [ex USNM 15784] (1); USNM 115783 (3), Islands, Japan). Holotype: ANSP 72039. Paratypes: 115784 (orig. 13, now 12), 115785 (orig. 44, now ANSP 72040 (1, missing). 42), 115786 (9). Distribution: Western Indian Ocean to Pitcairn Group, Distribution: Western Indian Ocean to Johnston, Line Islands and Hawaiian Islands; Indo-Australian Marcus and Wake Islands; Marshall Islands, Fiji, Archipelago to Japan.

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 71 Parenti 72

Samoa, Phoenix, Marquesas Islands, Tuamotu Pomacentrus adelus Allen, 1991 Archipelago and Society Islands. Pomacentrus adelus Allen, 1991: 136, 222, Fig. (Type locality, Ashmore Reef, Timor Sea, 0–4 m). Plectroglyphidodon randalli Allen, 1991 Holotype: WAM P.29042-009. Paratypes: BPBM Plectroglyphidodon randalli Allen 1991: 135, 221, 31804 (5); USNM 288343 (24), 288349 (19); WAM unnumb. Fig. (Type locality, Two km north of Flic P.26505-002 (8), P.27658-029 (17), P.28194-016 (1), en Flac, Mauritius, 2–3 m). Holotype: WAM P.29042-032 (15). P.26519-001. Paratypes: BPBM 31811 (3), WAM Distribution: Andaman Sea and Western Australia to P.26519-003 (4). Vanuatu and New Caledonia North to Philippines. Distribution: Réunion and Mauritius. Pomacentrus agassizii Bliss, 1883 Plectroglyphidodon sagmarius Randall and Earle, Pomacenteus [sic] agassizii Bliss 1883: 53. (Type 1999 locality, Mauritius). Syntypes: MCZ 6099 (2), Plectroglyphidodon sagmarius Randall and Earle, USNM 153556 [ex MCZ 6088] (1). 1999: 339, Figs. 5, 6. (Type locality, northeast of Distribution: Madagascar, Réunion, and Mauritius. Matahumu Point, Nuku Hiva, Marquesas Islands). Distribution: Marquesas Islands. Pomacentrus albiaxillaris Allen, Erdmann and Pertiwi, 2017 Plectroglyphidodon sindonis (Jordan and Evermann, Pomacentrus albiaxillaris Allen, Erdmann and 1903) Pertiwi, 2017a: 51, Figs. 2–3, 4B, 5, 6A, 7B. (Type Glyphisodon sindonis Jordan and Evermann, 1903: locality, Republic of Palau, SE of Koror, Uchelbeluu 188. (Type locality, Honolulu, Oahu Island, Hawaiian Reef, 7°16.680'N, 134°33.272'E). Holotype: ROM Islands, U.S.A). Holotype: USNM 50669. Paratypes: 78468. Paratypes: ROM 101282, 78481, 78483, SU 7610 (2), USNM 126524 [ex USBF 1070, USFC 78501, 78520, 78594, 78656, 78667, 78680, 78689, 2727] (1). 78700, (3, 1, 17, 2, 1, 10, 4, 1, 4, 25, 4); USNM Distribution: Hawaiian Islands. 432515 (6); WAM P.34548-001, P.34549-001 (1, 3). Genus Pomacentrus Lacepède, 1802 Distribution: Palau Archipelago. Pomacentrus Lacepède, 1802: 505. Masculine. Pomacentrus albicaudatus Baschieri-Salvadori, 1955 Type species Chaetodon pavo Bloch, 1787. Type by Pomacentrus albicaudatus Baschieri-Salvadori, 1955: subsequent designation by Guichenot, 1839: 503. 64, Pl. 4 (Fig. 4). (Type locality, Red Sea). Syntypes: Parapomacentrus Bleeker, 1877c: 65. Masculine. MCZR (1) Dahlak, MCZR (15) Dissei, MCZR (2) Type species Pomacentrus polynema Bleeker, 1853. Mersa Halaib. Type by subsequent designation by Jordan, 1919: Distribution: Red Sea. 387. Also appeared in Bleeker (1877d: 39) and if this predates Bleeker (1877c: 65) then type species Pomacentrus albimaculus Allen, 1975 is Glyphisodon bankieri Richardson by monotypy. Pomacentrus albimaculus Allen, 1975a: 96, Pl. 4 Pseudopomacentrus (subgenus of Pomacentrus) (Figs. a–b) (Type locality, near main wharf at Bleeker, 1877c: 39. Masculine. Type species Madang, New Guinea, 5°14'S, 145°45'E, 16 m). Pomacentrus littoralis Cuvier, 1830. Type by Holotype: AMS I.16691-006. Paratypes: AMS subsequent designation by Fowler and Bean (1928: I.16691-007 (3). 65). Also appeared in Bleeker (1877d: 39). Distribution: Northern New Guinea, Taiwan. Pellochromis (subgenus of Pomacentrus) Fowler and Pomacentrus alexanderae Evermann and Seale, 1907 Bean, 1928: 14. Masculine. Type species Pomacentrus alexanderae Evermann and Seale, 1907: Pomacentrus trimaculatus Rüppell, 1829. Type by 90, Fig. 17. (Type locality, Bacon, Sorsogon, Luzon original designation. Island, Philippines). Holotype: USNM 55919. Omopomacentrus (subgenus of Pomacentrus) Paratypes: (should be 6) ANSP 33296 (1), CAS 79635 Fowler, 1944a: 363. Masculine. Type species [ex IU 11594] (1), BSMP (1, apparently destroyed), Pomacentrus acapulcoensis Fowler, 1944. Type by MCZ 37285 [ex USNM 126403] (1), SU 20005 (1), original designation. U.S. Bur. Fish. 4540 (1), USNM 55922 (1). Lepidopomacentrus (subgenus of Pomacentrus) Distribution: Malaysia to Moluccas North to Taiwan Allen, 1975b: 39, 43. Masculine. Type species and Ryukyu Islands. Pomacentrus lepidogenys Fowler and Bean, 1928. Type by original designation. Pomacentrus alleni Burgess, 1981 Tropical Indo-West Pacific; the number of species Pomacentrus alleni Burgess 1981: 68, unnumb. Fig. increased dramatically in the 25 years from 53 (Type locality, Ko Ba Ngu Island, Similan Islands, recorded by Nelson (1994) to 77 by Nelson et al. Thailand, Andaman Sea, 8°40'N, 97°38'E). Holotype: (2016) and 82 presently listed here (Allen, 1991; USNM 228446. Paratypes: USNM 228447 (2), Allen, 1992; Allen, 1995; Allen, 1999; Allen, 2002; WAM P.26506-009 (8). Randall, 2002; Allen and Wright, 2003; Allen et al., Distribution: Similan Islands, Andaman Sea, off 2017, Hibib et al., 2020). Northwestern Thailand and Mentawai Islands, Nias Island and the Padang region of West Sumatra.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Pomacentrus amboinensis Bleeker, 1868 Pomacentrus aurifrons Allen, 2004 Pomacentrus amboinensis Bleeker, 1868: 334. (Type Pomacentrus aurifrons Allen, 2004: 3, Figs. 1–2. locality, Ambon Island, Moluccas Islands, Indonesia). (Type locality, Toiyama Island, Amphlett Group, Syntypes: (3) RMNH 6474 (3 of 4). Milne Bay Province, Papua New Guinea, 9°18.909'S, Pomacentrus dimidiatus Bleeker, 1876: 385. (Type 150°51.100'E, depth 12 meters). Holotype: WAM locality, Timor Island, southern Malay Archipelago; P.32261-001. Paratypes: AMS I.42620-001 (2); WAM Ambon Island, Molucca Islands, Indonesia). Syntypes: P.27827-038 (2), P.29742-001, P.32261-002 (5). RMNH 6475 (4). Distribution: Papua New Guinea to Solomon Islands, Distribution: Indo-Australian Archipelago, Melanesia, Vanuatu and New Caledonia. except Fiji, North to Marshall Islands, Taiwan and Pomacentrus auriventris Allen, 1991 Ryukyu Islands. Pomacentrus auriventris Allen, 1991: 140, 224, Fig. Pomacentrus andamanensis Allen, Erdmann and (Type locality, Flying Fish Cove, Christmas Island, Ningsih, 2020 Indian Ocean, ca. 10°29'S, 105°40'E, 5–10 m). Pomacentrus andamanensis Allen, Erdmann and Holotype: WAM P.26126-006. Paratypes: LIPI NCIP Ningsih, 2020: 40, Figs. 1, 2, 5A–C (Type locality, 4015 (1), 4016-17 (2); WAM P.28994-012 (2). Andaman Sea, Thailand, Surin Islands, 8.6667°, Distribution: Indo-Malayan Archipelago, Christmas 97.6333°, 20–35 m). Holotype: WAM P.26506-001. Island, and Caroline Islands. Paratypes: ROM 71364 (1), 71365 (13), 71367 (2), Pomacentrus australis Allen and Robertson, 1974 71368, (11), 71369 (1), 71370 (1), 71371 (3), 71372 (4), Pomacentrus australis Allen and Robertson, 1974: 71596 (3); WAM P. 226504-002 (4), P.26506-017. 158, Pl. 3. (Type locality, Outer reef off e. side of Distribution: Andaman Islands, Myanmar (Mergui One Tree Island Reef, Capricorn Group, Great Archipelago), west coast of Thailand (including Barrier Reef, 23°30'S, 152°05'E, 16 m). Holotype: Similan and Surin Islands), and the northwestern tip of AMS I.16592-001. Paratypes: AMS IA.6758 (1), Sumatra (Pulau Weh). I.15622-007 (1), I.15625-007 (8), I.15637-010 (10), Pomacentrus aquilus Allen and Randall, 1981 I.15679-005 (2), I.16524-002 (1), I.16592-002 (6), Pomacentrus aquilus Allen and Randall, 1980: (7) I.16593-001 (1); BPBM 13607 (3). 68, Figs. 45–47. (Type locality, Marsa el Mukabeila, Distribution: Great Barrier Reef to Sydney. fringing reef, outer reef flat, Gulf of Eilat, Red Sea, Pomacentrus azuremaculatus Allen, 1991 0.5–2 m). Holotype: BPBM 20480. Paratypes: Pomacentrus azuremaculatus Allen, 1991: 141, 225, BMNH 1976.9.20.1-7 (7); BPBM 14668 (3), 19864 Fig. (Type locality, 5 km southeast of Phuket, (5), 20786 (3); HUJ 8373 (7); MNHN 1976-118 (4); Thailand, 7°45'N, 98°22'E, 5–15 m). Holotype: TAU (=SMNHTAU) P.3312 (8); USNM 215157–59 PMBC 4684. Paratypes: BMNH 1987.6.30.7-8 (2), (52, 13, 10), 216476-77 (20, 5 or 4); WAM P.25527- BPBM 31802 (1), USNM 288344 (14), WAM 010 (3); ZMA 114269 (12). P.26506-004 (3). Distribution: Red Sea and Persian Gulf to Kenya and Distribution: Phuket and Similian Islands (Thailand) Madagascar. and Seribu Islands (Java Sea). Pomacentrus arabicus Allen, 1991 Pomacentrus baenschi Allen, 1991 Pomacentrus arabicus Allen, 1991: 139, 223, Pomacentrus baenschi Allen, 1991: 141, 226, Fig. unnumb. Fig. (Type locality, Khawr Kalbuh, Muscat, (Type locality, Kisiti Reef off Shimoni, Kenya, Oman, 23°37'N, 58°35'E, 2–6 m). Holotype: WAM 4°43'S, 39°22'E, 1–3 m). Holotype: WAM P.26515- P.25786-003. Paratypes: BPBM 31801 (4); WAM 008. Paratypes: BPBM 17614 (1); USNM 288346 P.25786-005 (5), P.25989-005 (3). (1), 288350 (5). Distribution: Gulf of Oman. Distribution: Kenya to Mozambique and Comoro Islands. Pomacentrus armillatus Allen, 1993 Pomacentrus bangladeshius Hibib, Islam, Nahar Pomacentrus armillatus Allen, 1993: 21, Figs. 1–2. and Neogi, 2020 (Type locality, Bohaydulong Island, Bodgaya Pomacentrus bangladeshius Hibib, Islam, Nahar and Islands, Sabah, Borneo, East Malaysia, ca. 4°35'N, Neogi, 2020: 415, Figs. 2, 3. (Type locality, 118°45'E, 15–20 m). Holotype: WAM P.30410-004. Bangladesh, Cox’s Bazar, Teknaf, Saint Martin’s Paratypes: WAM P.30409-001 (7). Island, 20°36’47.0” N 92°19’36.0” E). Holotype: Distribution: Borneo; probably occurs also in FBGN-SAU F1710SM-11. Paratypes: FBGN-SAU peninsular Malaysia, Indonesia, and the Philippines. F1801SM-06 (1), F1802SM-14 (1). Pomacentrus atriaxillaris Allen, 2002 Distribution: Known only from Saint Martin’s Island, Pomacentrus atriaxillaris Allen, 2002: 46, Figs. 1–2. Bangladesh. (Type locality, Mitsio Id, Madagascar). Holotype: Pomacentrus bankanensis Bleeker, 1854 WAM P.32015-001. Paratypes: WAM P.32024-001 (2). Pomacentrus bankanensis Bleeker, 1854c: 513. Distribution: Endemic to islands of North-western (Type locality, Bangka, Indonesia). Holotype: Madagascar.

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 73 Parenti 74

RMNH 27436 [ex RMNH 6468] (1 of 7). Based on Pomacentrus cranei Herre, 1935: 409. (Type locality, Pomacentrus taeniops of Bleeker (not of Cuvier). Tenibuli, Ysabel Island, Solomon Islands). Holotype: Pomacentrus dorsalis Gill, 1859: 147. (Type locality, FMNH 17338. Paratypes: FMNH 17339 (1); CAS- Shimoda, Japan). Holotype: USNM 64123. SU 24439 (1), 24442 (2). Pomacentrus delurus Jordan and Seale, 1905: 783, Distribution: Philippines, New Guinea, Solomon Fig. 6. (Type locality, South Negros, Philippines). Islands and Palau. Holotype: USNM 52052 [not 51052]. Paratypes: Pomacentrus caeruleopunctatus Allen, 2002 CAS-SU 9130 (1). Pomacentrus caeruleopunctatus Allen, 2002: 49, Distribution: Eastern Indian Ocean to Melanesia, Figs. 4–6. (Type locality, Mitsio Id, Madagascar). north to southern Japan. Holotype: WAM P.32015-002. Paratypes: WAM Pomacentrus bellipictus Allen, Erdmann and P.32015-003 (3). Hidayat, 2018 Distribution: Islands of North-western Madagascar. Pomacentrus bellipictus Allen, Erdmann and Pomacentrus caeruleus Quoy and Gaimard, 1825 Hidayat, 2018: 4, Figs. 2–3, 4A. (Type locality, Pomacentrus caeruleus Quoy and Gaimard, 1825: Indonesia, West Papua Province, Kokas District, off 397, Pl. 64 (Fig. 2). (Type locality, Mauritius). Arguni Island, Furum Islet, 2.6425°S, 132.5227°E, Syntypes: MNHN 8250 (1), 8504 (3). depth 1.5–2 meters). Holotype: MZB 24596. Pomacentrus caeruleoluteus Liénard in Sauvage, Paratypes: MZB 24597 (5), USNM 443778 (2), 1891: 430. (Type locality, Mauritius, Mascarenes, WAM P.34829-004 (5). southwestern Indian Ocean). No types known. Distribution: Kokas, West Papua (Indonesia). Pomacentrus pulcherrimus Smith, 1960: 344, Pl. 28 Pomacentrus bintanensis Allen, 1999 (Fig. A). (Type locality, Pinda, Mozambique, western Pomacentrus bintanensis Allen, 1999a: 103, Figs. Indian Ocean). Holotype: SAIAB [formerly RUSI] 5B, 6. (Type locality, Off Point Bintan, North coast 286. Paratypes: SAIAB [formerly RUSI]. of Bintan Island, Riau Islands, Indonesia, 1°11.21'N, Distribution: Western Indian Ocean (South Africa, East 104°27.38'E, 6 m). Holotype: NCIP 6153. Paratypes: Africa, Socotra, Seychelles, Comoros, Madagascar and WAM P.30956-001 (3). Mascarenes) to Maldive Islands. Distribution: Bintan Island, Riau Islands, Indonesia. Pomacentrus callainus Randall, 2002 Pomacentrus bipunctatus Allen and Randall, 2004 Pomacentrus callainus Randall, 2002: 169, Figs. 1– Pomacentrus bipunctatus Allen and Randall, 2004b: 2. (Type locality, Hakaumama'o Reef, Tongatapu, 77, Figs. 1–4. (Type locality, Truk Lagoon, 7°15'N, Tonga, depth 3 meters). Holotype: BPBM 37982. 152°02'E, North of Fefan Island, Caroline Islands, Paratypes: AMS I.41138-001 (1); BPBM 11350 (1), Micronesia, depth 5–6 meters). Holotype: BPBM 37970 (1); CAS 214537 (1); MNHN 2002-0092 (1); 39300. Paratypes: BPBM 17967 (2), 28006 (7), NSMT-P 63012 (1); ROM 48677 (18); USNM 29259 (6); CAS 220647 (3); MNHN 2004-2984 (1); 367494 (1); WAM P.31905.001 (1). USNM 380165 (2); WAM P.32354-001 (3). Distribution: Tonga and Fiji Islands. Distribution: Marshall Islands at Enewetak and Pomacentrus cheraphilus Allen, Erdmann and Kwajalein atolls and Truk in the Eastern Caroline Hilomen, 2011 Islands. Pomacentrus cheraphilus Allen, Erdmann and Pomacentrus brachialis Cuvier, 1830 Hilomen, 2011: 36, Figs. 1–3. (Type locality, East Pomacentrus brachialis Cuvier in Cuvier and Brunei Patches, 5°01.215'N, 114°43.892'E, Brunei, Valenciennes, 1830: 420. (Type locality, Java, depth 12–15 meters). Holotype: WAM P.33030-005. Indonesia). Syntypes: MNHN 8240 (1), RMNH 936 (1). Paratypes: UPLB Z-NS 0491-0500 (9); USNM Pomacentrus melanopterus Bleeker, 1852c: 562. (Type 399428 (5); WAM P.33030-001 (4). locality, Ambon Island, Molucca Islands, Indonesia). Distribution: Brunei and Philippines. Holotype: Whereabouts unknown. Bleeker specimens: Pomacentrus chrysurus Cuvier, 1830 BMNH 1862.2.28.109 (1), NMV 45907 (1). Pomacentrus chrysurus Cuvier (ex Broussonet) in Pseudopomacentrus rainfordi Whitley, 1935: 236. Cuvier and Valenciennes, 1830: 423. (Type locality, (Type locality, Hayman Island, Queensland, South Seas). Holotype: MNHN 8249. Australia). Holotype: AMS IA.6389. Paratypes: AMS Pomacentrus hogoleuensis Hombron and Jacquinot IA.385-87 (1, 1, 3), IA.912 (1), IA.1999 (1), IA.2283– in Jacquinot and Guichenot, 1853: 47, Pl. 5 (fig. 3). 86 (1, 1, 1, 1), IA.6390 (1). (Type locality, Hogolen, Caroline Islands). Syntypes: Distribution: Indo-Australian Archipelago to New MNHN 0000-8501 (2). Caledonia and Loyalty Islands. Pomacentrus rhodonotus Bleeker, 1853d: 282. (Type Pomacentrus burroughi Fowler, 1918 locality, Cauer, Sumatra, Indonesia). Holotype: Pomacentrus burroughi Fowler, 1918: 48, Fig. 19. RMNH 6449. (Type locality, Philippines). Holotype: ANSP 47526. Glyphidodon luteocaudatus Saville-Kent, 1893: 368, Paratypes: ANSP 47527 (1). Color Pl. 16 (Fig. 11). (Type locality, Queensland, Australia). No types known.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Pomacentrus flavicauda Whitley, 1928: 297, Fig. 1. Pomacentrus flavoaxillaris Allen, Erdmann and (Type locality, North-west Islet, Capricorn Group, Pertiwi, 2017a: 59, Figs. 4C, 6B, 7C, 9–11. (Type Queensland, Australia). Holotype: AMS IA.2598. locality, Federated States of Micronesia, Pohnpei, Distribution: Maldive Islands and Sri Lanka to Indo- Ant Atoll, pinnacle reef in SE part of lagoon, Australian Archipelago and Micronesia north to 6°46.104'N, 157°59.767'E, depth 8–10 meters). Ryukyu Islands. Holotype: WAM P.33149-001. Paratypes: BPBM 28291, 28309 (1, 1); USNM 223947 (10); WAM Pomacentrus coelestis Jordan and Starks, 1901 P.33149-002, 34504-001, 34505-001 (7, 4, 4). Pomacentrus coelestis Jordan and Starks, 1901: 383, Distribution: Micronesia. Pl. 21. (Type locality, Wakanoura, Kii, Japan). Holotype: SU 6434. Pomacentrus geminospilus Allen 1993 Distribution: Sri Lanka to Tuamotu Archipelago and Pomacentrus geminospilus Allen, 1993: 23, Figs. 3– Line Islands northward to southern Japan. 4. (Type locality, Pass between Bodgaya and Bohaydulong Islands, Sabah, Borneo, Malaysia, ca. Pomacentrus colini Allen, 1991 4°34'N, 118°46'E, 10–12 m). Holotype: WAM Pomacentrus colini Allen, 1991: 146, 227, unnumb. P.30409-006. Paratypes: WAM P.30409-001 (7). Fig. (Type locality, Motupore Island, off Port Distribution: Malaysia and Philippines. Moresby, Papua New Guinea, 13–16 m). Holotype: WAM P.29356-002. Paratypes: BPBM 31805 (2), Pomacentrus grammorhynchus Fowler, 1918 WAM P.29356-001 (4). Pomacentrus grammorhynchus Fowler, 1918: 44, Distribution: Southern Papua New Guinea. Fig. 17. (Type locality, Philippines). Holotype: ANSP 47518. Pomacentrus cuneatus Allen, 1991 Distribution: Taiwan to Indo-Australian Archipelago Pomacentrus cuneatus Allen, 1991: 146, 228, and Great Barrier Reef. unnumb. Fig. (Type locality, Ambon, off Poka village in Ambon Bay, Molucca Islands, Indonesia, 2 Pomacentrus imitator Whitley, 1964 m). Holotype: LIPI NCIP 3998. Paratypes: BPBM Pseudopomacentrus imitator Whitley, 1964: 175, 19467 (4); LIPI NCIP 3999 (3); WAM P.25232-001 Fig. 9. (Type locality, Lihou Atoll, Coral Sea). (5), P.25244-001 (8). Holotype: AMS IB.4945. Paratypes: AMS IB.4908– Distribution: Ambon, Seribu (off Java), Manado, 09 (1, 1), IB.4944 (1). Sulawesi (Indonesia), Singapore, West Malaysia, Distribution: Coral Sea, New Caledonia, Fiji and Rotuma. Gulf of Thailand, and Luzon (Philippines). Pomacentrus indicus Allen, 1991 Pomacentrus emarginatus Cuvier, 1829 Pomacentrus indicus Allen, 1991: 148, 229, Fig. Pomacentrus emarginatus Cuvier, 1829: 179. (No (Type locality, Villingili Island, North Male Atoll, locality). Cuvier and Valenciennes specimens: Maldives, Indian Ocean, 4°10'S, 73°30'E, 2–15 m). MNHN 9656 (1) Waigeo. Available from footnote 5 Holotype: WAM P.26514-004. Paratypes: BPBM as: "P[omacentrus] emarginatus, Seb., III, xxvii, 26, 18923 (7); FMNH 75008 (36, now 16), 75020 (32 or 27, 28." Also described in Cuvier and Valenciennes 33), 97150 (20); USNM 288347 (5); WAM P.29361- (1830: 422) on specimen from Waigeo. 001 (2), P.26514-007 (5). Distribution: Palau and New Guinea. Distribution: Sri Lanka, Chagos Archipelago, Maldive Islands and Seychelles. Pomacentrus fakfakensis Allen and Erdmann, 2009 Pomacentrus fakfakensis Allen and Erdmann, 2009b: Pomacentrus javanicus Allen, 1991 123, Figs. 2–3. (Type locality, Karawatu Island, Pomacentrus javanicus Allen, 1991: 148, 230, Fig. 4°01.803'S. 133°27.711'E, Papua Barat Province, (Type locality, Pulau Putri, Seribu Islands Indonesia, Indonesia, depth 4–5 meters). Holotype: NCIP 6329. 12 m). Holotype: LIPI NCIP 3995. Paratypes: BPBM Paratypes: NCIP 6336 (2); USNM 391618 (3); WAM 18575 (1), WAM P.25248-010 (2). P.32791-005 (2), P.32954-001 (5). Distribution: Seribu Islands, Java Sea. Distribution: West Papua. Pomacentrus komodoensis Allen, 1999 Pomacentrus flavioculus Allen, Erdmann and Pomacentrus komodoensis Allen, 1999a: 101, Figs. Pertiwi, 2017 3–4. (Type locality, Nw. corner of large rocky island Pomacentrus flavioculus Allen, Erdmann and Pertiwi, off S. end of Komodo Island, Indonesia, ca. 8°44'S, 2017b: 24, Figs. 1–2, 4A. (Type locality, Fiji, Lau 119°25'E, 3–4 m). Holotype: NCIP 6152. Paratypes: Archipelago, Vatoa Island, 19°48'S, 178°15'W). WAM P.30956-001 (1). Holotype: WAM P.30151-001. Paratypes: ROM Distribution: Komodo Island, Indonesia. 48799, 48802, 90814, 90820 (12, 37, 10, 16); USNM Pomacentrus lepidogenys Fowler and Bean, 1928 275233, 333945, 337921 (25, 21, 8); WAM P.30147- Pomacentrus lepidogenys Fowler and Bean, 1928: 001 (10). 98, Pl. 8. (Type locality, Ligpo Point, Balayan Bay, Distribution: Fiji and Tonga. Philippines). Holotype: USNM 89959. Pomacentrus flavoaxillaris Allen, Erdmann and Distribution: Andaman Sea to Great Barrier Reef, Pertiwi, 2017 Melanesia and Tonga.

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Pomacentrus leptus Allen and Randall, 1980 m). Holotype: NMMB-P15602. Paratypes: CAS Pomacentrus leptus Allen and Randall, 1980: 74, 235015 (formerly NMMB-P15605) (6); NMMB- Figs. 48–49. (Type locality, Towartit Reef complex, P15591-606, 15403-4, 15607 (14); USNM 410087 19 km south of Port Sudan, Red Sea, 2–3 m). (2), WAM P.33715-001 (2). Not available from 2013 Holotype: BPBM 19698. Paratypes: BMNH electronic version, which appeared without ZooBank 1977.2.1.1-5 (5); BPBM 17893 (2); HUJ 8369 (1); registration number; available from erratum which MNHN 1977-453 (7), 1977-569 (4); SMF 4946 (1), was published on 17 Sept. 2014 (Liu et al., 2014b). 13716 (5); USNM 213672 (25); WAM P.25786-001 Distribution: Marshall Islands but may have a (4), P.26460-010 [not P.25998-010] (1), P.26461-010 distribution from Palau to the Marshall Islands and [not P.25999-010] (2). south to Bali. Distribution: Gulf of Oman to Gulf of Aden and Pomacentrus microspilus Allen and Randall, 2005 Southern Red Sea. Pomacentrus microspilus Allen and Randall, 2005: Pomacentrus limosus Allen, 1992 96, Figs. 1–3. (Type locality, Nananu-I-Ra Bay, Pomacentrus limosus Allen, 1992: 691, Fig. 1. (Type 17°15.475'N, 178°08.68'E, Viti Levu, Fiji, depth 4–8 locality, Churchill Reef, 15°31'S, 123°17'E, Western m). Holotype: WAM P.32755-001. Paratypes: BPBM Australia, 8.0–8.5 m). Holotype: WAM P.30317-001. 39150 (4); CAS 221591 (7), 251592 (6), 221593 (1), Paratypes: WAM P.30317-002 (1). 221594 (1), 221595 (3), 221596 (23); USNM 383521 Distribution: Churchill Reef, off Northwestern (1); WAM P.32755-002 (2). Australia. Distribution: Fiji Islands. Pomacentrus littoralis Cuvier, 1830 Pomacentrus milleri Taylor, 1964 Pomacentrus littoralis Cuvier (ex Kuhl and van Pomacentrus milleri Taylor, 1964: 261, Pl. 60. (Type Hasselt) in Cuvier and Valenciennes, 1830: 425. (Type locality, Coral reef at Yirrkalla, sta. M48-21, near locality, Java, Indonesia). Syntypes: MNHN 0000- Cape Arnhem, Northern Terr., Australia). Holotype: 9445 (1, dry, headless) Moluccas, RMNH 946 (1). AMS IB.5235. Paratypes: AMS IB.7306 (10); Distribution: Singapore, Indonesia, Philippines and USNM 173623–27 (17, 94, 9, 2, 52). Northern Australia. Distribution: Rottnest Island (Western Australia) to Northern Territory. Pomacentrus maafu Allen and Drew, 2012 Pomacentrus maafu Allen and Drew, 2012: 173, Pomacentrus moluccensis Bleeker, 1853 Figs. 1–3. (Type locality, Dravuni Island, Great Pomacentrus moluccensis Bleeker, 1853b: 118. (Type Astrolabe Reef, Viti Levu, Fiji, depth 3–10 meters). locality, Ambon and Ternate, Moluccas, Indonesia). Holotype: ROM 92387. Paratypes: ROM 46160 (28), Syntypes: (3) ?BMNH 1862.2.28.36 (1), ?NMV 48653 (36), 48645 (46); USNM 298829 (52); WAM 45909-10 (2), RMNH 6 (up to 3 of 19). P32333-003 (9). Pomacentrus popei Evermann and Seale, 1907: 90, Distribution: Fiji Islands and Tonga. Fig. 16. (Type locality, Bacon, Sorsogon, Philippines). Holotype: USNM 55903. Pomacentrus magniseptus Allen, Erdmann and Pomacentrus sufflavus Whitley, 1927b: 18, Pl. 1 (Fig. Pertiwi, 2017 3). (Type locality, Michaelmas Cay, Great Barrier Pomacentrus magniseptus Allen, Erdmann and Reef, off Cairns, Queensland, Australia). Holotype: Pertiwi, 2017a: 64, Figs. 6C, 12–13. (Type locality, AMS IA.2777. Paratypes: (11) AMS I.43609-001 [ex Great Barrier Reef, Northern corner of Escape Reef, AMS IA.2777] (1), IA.388–IA.392 (1, 1, 1, 2), IA.925 Queensland, Australia, approximately 15°50'S, (1), IA.2003 (1), IA.2633 (1). 145°50'E, depth 3–6 meters). Holotype: AMS Distribution: Andaman Sea to Melanesia North to I.22574-010. Paratypes: AMS I.16680-007 (1), Ryukyu Islands. I.17061-006 (2), I.20757-045 (3), I.22574-046 (6), I.22616-007 (10); WAM P.28533-010 (1). Pomacentrus nagasakiensis Tanaka, 1917 Distribution: Great Barrier Reef, Queensland, Pomacentrus nagasakiensis Tanaka, 1917: 9. (Type Australia. locality, Fish market in Nagasaki, Japan). Syntypes: ZUMT 6582 (1), 6583 (1, described spec.). Pomacentrus melanochir Bleeker, 1877 Pomacentrus caudovittatus Schmidt, 1931: 64, Pl. 4 (Fig. Pomacentrus melanochir Bleeker, 1876: 384. (Type 2). (Type locality, Kominato, Amami-Oshima Island, locality, Bali, Flores, Timor, and Ambon Island, Ryukyu Islands, Japan). Syntypes: ZIN 23283 (2). Moluccas Islands, Indonesia). Syntypes: RMNH Pomacentrus arenarius Allen, 1987c: 9, Fig. 1. 6473 (11). (Type locality, Escape Reef, Great Barrier Reef, Distribution: Bali, Flores, Timor, Buru, Sulawesi, Queensland, Australia, 15°50'S, 145°50'E, depth 15 and Ambon. meters). Holotype: WAM P.27465-001. Paratypes: Pomacentrus micronesicus Liu, Ho and Dai, 2014 AMS I.26319-001 (1); BPBM 30942 (1); WAM Pomacentrus micronesicus Liu, Ho and Dai, 2014a: P.24940-001 (1), P.25107-008 (2), P.25371-004 (2), 3, 1F, 2, 3. (Type locality, Kwajalen Atoll, Marshall P.27461-004 (1), P.28161-002 (1). Islands, 9°11'02.87"N, 167°25'04.88"E, depth 5–10 Distribution: Izu Islands to Ryukyu Islands.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Pomacentrus nigriradiatus Allen, Erdmann and Pomacentrus hainanensis Wang, 1941 94. (Type Pertiwi, 2017 locality, Hainan Island, China). Holotype: ? SSCN. Pomacentrus nigriradiatus Allen, Erdmann and Distribution: Western Indian Ocean to Tuamotu Pertiwi, 2017a: 68, Figs. 4D, 6D, 14–15. (Type Archipelago, Marshall Islands, and Gilbert Islands; locality, Upolu Island, off Lotofaga, Samoa, northern Australia to Kerama Islands (Southern 14°01.034'S, 171°50.644'W, depth 12–15 m). Japan). Holotype: WAM P.34491-003. Paratypes: AMS Pomacentrus philippinus Evermann and Seale, I.17468-001 (1),; BPBM 8122 (2), 11308 (1), 11423 1907 (1), 16793 (1), 17462 (2); ROM 66095 (4); USNM Pomacentrus philippinus Evermann and Seale, 1907: 324104 (2), 347550 (6), 360955 (4), 362702 (4), 91, Fig. 18. (Type locality, Bacon, Sorsogon, 373336 (7); WAM P.34491-001 (5), P.25129-002 Philippines). Holotype: USNM 55901. Paratypes: SU (2), P.29368-001 (5). 20009 (1). Distribution: New Caledonia to American Samoa. Distribution: Maldive Islands to Melanesia North to Pomacentrus nigromanus Weber, 1913 Ryukyu Islands. Pomacentrus nigromanus Weber, 1913: 338, Fig. 72. Pomacentrus pikei Bliss, 1883 (Type locality, Ambon Island, Moluccas Islands, Pomacenteus [sic] pikei Bliss, 1883: 54 (Type Indonesia, Siboga sta. 231). Holotype: ZMA 112626. locality, Mauritius). Holotype: MCZ 6089. Distribution: Indo-Australian Archipelago. Distribution: Mauritius and Réunion. Pomacentrus nigromarginatus Allen, 1973 Pomacentrus polyspinus Allen, 1991 Pomacentrus nigromarginatus Allen, 1973a: 38, Fig. Pomacentrus polyspinus Allen, 1991: 155, 230, Fig. 3. (Type locality, outer reef slope of N. end of (Type locality, Ko Hi Island off Phuket, Thailand, Kranket Island, ca. 3 nautical mi. North of Madang, 7°45'N, 98°22'E, depth 4–8 m). Holotype: PMBC New Guinea, 5°11'24"S, 145°50'54"E, 35 m). 4683. Paratypes: WAM P.26504-010 (4). Holotype: USNM 207937. Paratypes: AMS I.16655- Distribution: Phuket Island (Thailand). 001 (1), BMNH 1972.8.14.3 (1), BPBM 13226 (2), USNM 207938 (1). Pomacentrus proteus Allen, 1991 Distribution: Eastern Indian Ocean to Coral Sea Pomacentrus proteus Allen, 1991: 156, 231, Fig. North to Ryukyu Islands. (Type locality, Colombo, near Kinross Swimming Club, Sri Lanka, 6°58'N, 79°52'E, depth 3–10 m). Pomacentrus opisthostigma Fowler, 1918 Holotype: WAM P.26509-011. Paratypes: BPBM Pomacentrus opisthostigma Fowler, 1918: 51, Fig. 31803 (3), WAM P.26509-007 (4). 20. (Type locality, Philippines). Holotype: ANSP Distribution: Sri Lanka. 47528. Paratypes: ANSP 47529-30 (2). Distribution: New Guinea and Philippines. Pomacentrus reidi Fowler and Bean, 1928 Pomacentrus reidi Fowler and Bean, 1928: 99, Pl. 9. Pomacentrus pavo (Bloch, 1787) (Type locality, Tataan Island, Tawi Tawi Islands, Chaetodon pavo Bloch, 1787: 60, Pl. 198 (Fig. 1). Philippines). Holotype: USNM 89960. Paratypes: (Type locality, East Indies). Lectotype: ZMB 2752. (48) ANSP 85294 (2), FMNH 62626 (2), USNM Paralectotypes: ZMB (1, lost). On page 44 of Bloch's 98244 (1), 98741–42 (1, 1), 98744–48 (1, 1, 1, 1, 1), Ichthyologie, volume 6. Lectotype selected by 98751–54 (1, 7, 1, 1), 98756 (1), 98758 (5), 98760– Paepke (1999: 120). 71 (1, 1, 1, 1, now 0, 1, 1, 2, 1, 1, 1, 3). Holocentrus diacanthus Lacepède, 1802: 333, 372. Distribution: Sulawesi North to Philippines South to (Type locality, donated by "la république batave"). Northern Great Barrier Reef and Vanuatu. Syntypes: MNHN 0000-8147 (4). Pomacentrus pavoninus Bleeker, 1853f: 85. (Type Pomacentrus rodriguesensis Allen and Wright, locality, Lawajong, Solor Island, Indonesia). 2003 Holotype: Whereabouts unknown. Pomacentrus rodriguesensis Allen and Wright, 2003: Pomacentrus polynema Bleeker, 1853d: 283. (Type 134, Figs. 1–3. (Type locality, Rodrigues Island, locality, Priaman, Sumatra, Indonesia). Holotype: Indian Ocean, 19°39’16’’S, 63°25’28’’E). Holotype: RMNH 6464. SAIAB 66635. Paratypes: SAIAB 66633 (6), 66634 Pomacentrus furcatus Thiollière in Montrouzier, (3), 66636 (1), 66640 (2); WAM 32237-001 (2). 1857: 479. (Type locality, Woodlark Island, South Distribution: Known only from the type locality: Pacific). No types known. Mascarene endemics. Pomacentrus notatus De Vis, 1884: 451. (Type locality, New Britain Island, Bismarck Archipelago). Pomacentrus saksonoi Allen, 1995 No types known. Pomacentrus saksonoi Allen, 1995: 88, Fig. 4. (Type Pomacentrus suvarovensis Stead, 1907: 4, Pl. 1. locality, West side of Pulau Junnkung, Seribu (Type locality, Anchorage Island, Suwarrow Islands, Islands, Indonesia, ca. 5°34'S, 106°31'E, 12–15 m). Cook Islands, southwestern Pacific). Syntypes: AMS Holotype: NCIP 6134. Paratypes: WAM P.30810- IA.2460-64 (5). 002 (1).

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 77 Parenti 78

Distribution: Seribu Islands (Java Sea) and off 2556 (1); ZMB 7891 (1). Lectotype selected by northern Sulawesi. Fricke (1999: 384). Pomacentrus grandidieri Steindachner, 1891a: 174. Pomacentrus similis Allen, 1991 (Type locality, Madagascar). Syntypes: NMW 70639 Pomacentrus similis Allen, 1991: 157, 232, unnumb. (2). Species illustrated and described as new species Fig. (Type locality, Trincomalee, Sri Lanka, ca. in more detail in Steindachner (1891: 372, Pl. 2, fig. 8°30'N, 81°06'E, 3 m). Holotype: WAM P.26511- 3). 001 [not 16511 as published]. Paratypes: PMBC Pomacentrus scintillans Smith, 1955: 895, Pl. 20 5170 (2), WAM P.26504-003 (1). (Fig. A). (Type locality, Zanzibar, Tanzania, western Distribution: Sri Lanka and Andaman Sea. Indian Ocean). Holotype: SAIAB [formerly RUSI] Pomacentrus simsiang Bleeker, 1856 294. Paratypes: (numerous) SAIAB [formerly RUSI]. Pomacentrus simsiang Bleeker, 1856b: 90. (Type Distribution: Red Sea and western Indian Ocean: locality, Jakarta, Java, Indonesia). Holotype: RMNH East Africa to about 21°S. Madagascar, Mauritius, 27437 [ex RMNH 64668] (1 of 5). Comoros, and Seychelles. Pomacentrus tropicus Seale, 1910: 517, Pl. 12 (Fig. Pomacentrus taeniometopon Bleeker, 1852 1). (Type locality, Sitanki Island, Jolo Archipelago, Pomacentrus taeniometopon Bleeker, 1852b: 283. Philippines). Holotype: BSMP 4737 (destroyed). (Type locality, Wahai, N. Ceram, Indonesia). Paratypes: BSMP 4736 (5, destroyed). Holotype: ?RMNH 27434 [ex RMNH 6468] (1 of 7). Pomacentrus hebardi Fowler, 1918: 46, Fig. 18. Distribution: Indo-Malayan Archipelago, Taiwan, (Type locality, Philippines). Holotype: ANSP 47519. and Ryukyu Islands. Paratypes: ANSP 47520-25 (6). Distribution: Indo-Australian Archipelago North to Pomacentrus trichrourus Günther, 1867 Ryukyu Islands. Pomacentrus trichrourus Günther in Playfair and Günther, 1867: 146, Pl. 17 (Fig. 5). (Type locality, Pomacentrus smithi Fowler and Bean, 1928 Zanzibar). Holotype: BMNH 1865.2.27.47. Possible Pomacentrus smithi Fowler and Bean, 1928: 75, Pl. syntypes: BMNH 1865.2.27.47 (1, in alcohol), 7. (Type locality, Biri Channel, Balicuatro Island, 2004.11.5.6 (1, skin). Spelled trichourus throughout Philippines). Holotype: USNM 89958. Paratypes text and trichrourus on plate. Günther, acting as first and/or additional material: (79) ANSP 80431–32 (2, reviser, in Zoological Record for 1866, selected 3); USNM 97968–86 (1, 1, 1, 3, 6, 1, 2, 3, 1, 1, 13, trichrourus. 12, 1, 1, 4, 2, 1, 2, 1). Distribution: Red Sea, Persian Gulf and East coast of Distribution: Indo-Australian Archipelago: Africa (not recorded from islands of the Western Philippines, Papua New Guinea, Solomon Islands, Indian Ocean). and New Hebrides. Pomacentrus trilineatus Cuvier, 1830 Pomacentrus spilotoceps Randall, 2002 Pomacentrus trilineatus Cuvier (ex Ehrenberg) in Pomacentrus spilotoceps Randall, 2002: 174, Figs. Cuvier and Valenciennes, 1830: 428. (Type locality, 4–5. (Type locality, Hakaumama'o Reef, Tongatapu, Red Sea; Moluccas). Syntypes: MNHN 0000-8142 Tonga, depth 3 meters). Holotype: BPBM 37983. (1) Red Sea; Moluccas specimen lost; ?ZMB 2756 Paratypes: MNHN 2002-093 (1), USNM 367495 (1), (1) Ehrenberg specimen. WAM P.31906.001 (1). Pomacentrus biocellatus Rüppell, 1838: 127, Pl. 31 Distribution: Tonga and Fiji Islands. (Fig. 3). (Type locality, Massawa, Eritrea, Red Sea). Pomacentrus stigma Fowler and Bean, 1928 Syntypes: SMF 1596 (4). Pomacentrus stigma Fowler and Bean, 1928: 100, Pl. Distribution: Red Sea to Mozambique and 10. (Type locality, Romblon Harbor, Philippines). Madagascar. Holotype: USNM 89961 (1 of 2). Paratypes (incl. Pomacentrus tripunctatus Cuvier, 1830 additional material): (32) ANSP 80161 (1); FMNH Pomacentrus tripunctatus Cuvier in Cuvier and 62627 (1); USNM 89961 (1 of 2), 97939 (2), 97940– Valenciennes, 1830: 421. (Type locality, Vanikoro 41 (1, 1), 97944–52 (1, 1, 1, 1, 5, 2, 4, 2, 1), 97954– Island, Santa Cruz Islands, sw. Pacific, 11°37'S, 56 (1, 1, 6), 97958–66 (1, 1, 1, 1, 1, 1, 3, 19, 1), 166°58'E). Syntypes: MNHN 0000-8245 (4). 98826 (1). Pomacentrus vanicolensis Cuvier in Cuvier and Pomacentrus tablasensis Montalban, 1928: 72, Pl. 14 Valenciennes, 1830: 421. (Type locality, Vanikoro (Fig. 2). (Type locality, Tablas, Romblon Province, Island, Santa Cruz Islands, southwestern Pacific). Philippines). Holotype: BSMP (destroyed). Syntypes: MNHN 0000-8270 (4). Distribution: Philippines and Taiwan. Pristotis fuscus Bleeker, 1849: 9. (Type locality, Pomacentrus sulfureus Klunzinger, 1871 Boleling, Northern Bali, Indonesia). No types known. Pomacentrus sulfureus Klunzinger, 1871: 521. (Type Pomacentrus katunko Bleeker, 1852b: 169. (Type locality, Al-Quseir [Kosseir], Egypt, Red Sea). locality, Kupang, Timor Island, southern Malay Lectotype: SMNS 1763. Paralectotypes: BMNH Archipelago). Holotype: RMNH 27433 [ex RMNH 1871.7.15.36 (1); SMNS 3526 (4); ZIN 2550 (1), 6468].

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Pomacentrus montrozieri Thiollière in Montrouzier Paratypes: BPBM 18576 (1), LIPI NCIP 4000 (1), 1857: 478. (Type locality, Woodlark Island, South WAM P.25248-002 (2). Pacific). No types known. Distribution: Seribu Islands (Java Sea). Pomacentrus catunco Peters, 1868: 270. Misspelling Pomacentrus yoshii Allen and Randall, 2004 or unjustified emendation. Pomacentrus yoshii Allen and Randall, 2004b: 83, Pomacentrus bilineatus Castelnau, 1873: 89. (Type Figs. 7–8. (Type locality, Outer reef slope off Calalen locality, Darwin, Northern Territory, Australia). Islet, Majuro Atoll, Caroline Islands, depth 10–15 Syntypes: MNHN (2). meters). Holotype: BPBM 39299. Paratypes: BPBM Pomacentrus punctolineatus Cartier, 1874: 98. (Type 9660 (1); CAS 220648 (2); MNHN 2004-2985 (2); locality, Bohol, Philippines). No types known. USNM 380166 (1), 380167 (1); WAM P.32463-001 Pomacentrus obscurus Alleyne and Macleay, 1877: (2). 343, Pl. 15 (Fig. 2). (Type locality, Torres Strait, Distribution: From the Southern Marshall Islands at Queensland, Australia). Syntypes: AMS I.16358-001 Majuro and Mili Atoll. Despite extensive fieldwork it [ex MAMU F826] (4). Nomen praeoccupatum; was not found in the Northern Marshalls at Enewetak objectively invalid preoccupied by Pomacentrus Atoll. obscurus Thiollière in Montrouzier 1857 [=Stegastes punctatus (Quoy and Gaimard, 1825)]. Genus Pomachromis Allen and Randall, 1974 Pomacentrus elongatus Seale, 1910: 518, Pl. 12 (Fig. Pomachromis Allen and Randall, 1974: 45. Masculine. 2). (Type locality, Limbones Cove at entrance to Manila Type species Abudefduf richardsoni Snyder, 1909. Bay, Luzon Island, Philippines). Holotype: BSMP 2214 Type by original designation. (destroyed in World War II). Paratypes: BSMP 2212 (1) Distribution: Tropical Indo-West Pacific. Four species. and 2213 (1) (destroyed in World War II). Pomacentrus macleayi Whitley, 1928: 221. (Type Pomachromis exilis Allen and Emery, 1973 Pomacentrus exilis Allen and Emery, 1973: 565, Fig. locality, Torres Strait, Queensland, Australia). 1. (Type locality, Piis Island, Truk Group, e. Caroline Nomen novum. Replacement name for Pomacentrus Islands, 30 ft). Holotype: USNM 207935. Paratypes: obscurus Alleyne and Macleay 1877, preoccupied by BMNH 1972.8.14.1-2 (2), BPBM 7780 (1), USNM Pomacentrus obscurus Thiollière 1856. 207934 (2). Distribution: Sri Lanka and Andaman Sea to Distribution: Marshall Islands and Truk Islands Solomon Islands and Vanuatu. (Caroline Islands). Pomacentrus vaiuli Jordan and Seale, 1906 Pomachromis fuscidorsalis Allen and Randall, 1974 Pomacentrus vaiuli Jordan and Seale, 1906: 280, Pl. Pomachromis fuscidorsalis Allen and Randall, 1974: 40 (Fig. 2). (Type locality, Samoa). Holotype: 48, Fig. 5. (Type locality, Teavaraa Pass, Papara, Tahiti, USNM 51740 (1 of 5). Paratypes: MCZ 36007 [ex Society Islands, 20 ft). Holotype: BPBM 6883. USNM 51740] (1); SU 8719 (8); USNM 51740 (4 of Paratypes: AMS I.17352-001 (2); BMNH 1973.10.17.1 5), 126355 (4). (1); BPBM 8436 (2), 14247 (1); USNM 211290 (1). Distribution: Western Australia, Moluccas and Distribution: Society Islands, Tuamotu Archipelago, Melanesia to Samoa, Marshall Islands, and Kiribati; and Pitcairn Group. Northward to Southern Japan. Pomachromis guamensis Allen and Larson, 1975 Pomacentrus vatosoa Frable and Tea, 2019 Pomachromis guamensis Allen and Larson, 1975: Pomacentrus vatosoa Frable and Tea, 2019: 324, Figs. (123) 124, Fig. 1. (Type locality, Off Tanguisson 1–4, 5A1–2. (Type locality, Nosy Faho, Madagascar, Point, Guam, Mariana Islands, w. Pacific, 35 ft). 18°17'32.2"S, 49°24'35.8"E, depth 15–20 meters). Holotype: USNM 213576 [ex UGM 4340]. Holotype: SIO 19-2. Paratypes: UADBA 59997 (1), Paratypes: UGM 4457 (1). ZRC 60713 (2). Distribution: Mariana Islands. Distribution: Northeastern Madagascar, western Indian Ocean. Pomachromis richardsoni (Snyder, 1909) Abudefduf richardsoni Snyder, 1909: 600. (Type Pomacentrus wardi Whitley, 1927 locality, Naha, Okinawa Island, Ryukyu Islands, Pomacentrus wardi Whitley, 1927a: 301, unnumb. Japan). Holotype: USNM 62948. Paratypes: SU Fig. (Type locality, Heron Island, Capricorn Group, 21134 (3), USNM 74589 (6). Queensland, Australia). Holotype: AMS IA.2964. Distribution: From scattered localities in the Indo-West Paratypes: AMS IA.2963 (1). Additional material: Pacific, including Mauritius, Ryukyu Islands, Taiwan, (5) IA.6389 (1). Great Barrier Reef, Loyalty Islands, Fiji, and Samoa. Distribution: Great Barrier Reef to Sydney. Genus Premnas Cuvier, 1816 Pomacentrus xanthosternus Allen, 1991 Premnas Cuvier, 1816: 345. Feminine. Type species Pomacentrus xanthosternus Allen, 1991: 163, 233, Chaetodon biaculeatus Bloch, 1790. Type by monotypy. Fig. (Type locality, Pulau Putri, Seribu Islands Java Northeastern Indian Ocean to Indo-Australian Sea, Indonesia, 2–3 m). Holotype: LIPI NCIP 3996. Archipelago; one species.

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 79 Parenti 80

Premnas biaculeatus (Bloch, 1790) Pristotis coeruleopunctatus Rüppell, 1838: 129. Chaetodon biaculeatus Bloch, 1790: 11, Pl. 219 (Fig. (Type locality, Massawa, Eritrea, Red Sea). Alternate 2). (Type locality, East Indies). No types known. name or misprinting of Pristotis cyanostigma. Perca koelreuteri Walbaum, 1792: 343. (No Distribution: Red Sea and Gulf of Aden. locality). No types known. Based on Koelreuter Pristotis obtusirostris (Günther, 1862) (1764: 346), Pl. 8 (fig. 5). Pomacentrus obtusirostris Günther, 1862: 24. (No Perca trifasciata Walbaum, 1792: 354. (No locality). locality). Syntypes: BMNH 1848.3.18.? (1), BMNH No types known. Based on Koelreuter (1764: 346). 1855.9.19.1541 (1). Lutianus trifasciatus Bloch and Schneider, 1801: Pomacentrus jerdoni Day, 1873b: 237. (Type locality, 568. (No locality). No types known. Madras, India). Syntypes: (3) ZSI 1612 (1). Holocentrus sonnerat Lacepède, 1802: 344, 389. Pomacentrus dolii MacLeay, 1881: 65, Pl. 1 (Fig. 1). (Type locality, Mauritius, Mascarenes, southwestern (Type locality, Port Jackson, New South Wales, Indian Ocean). Holotype: MNHN (not found). Australia). Syntypes: AMS I.16359-001 [ex MAMU Scorpaena aculeata Lacepède, 1801: 258, 268. (No F1164] (2). locality). Syntypes: MNHN 0000-9441 (1, dry), A- Daya jerdoni var. fusca McCulloch, 1921: 171, Pl. 9 8120 (2), A-8121 (1, dry). (Fig. 2). (Type locality, Southern Queensland, Premnas trifasciatus Cuvier in Guérin-Méneville, Australia). Holotype: AMS IA.148. Paratypes: (several) 1828: no page number, Livraison 13. (No locality). AMS IA 144 (1), IA.146 (1); QM I.3579 [ex AMS]. No types known. Text applies to Pl. 26 (no. 29) Chromis virescens Ogilby, 1922: 303, Pl. 19 (Fig. 3). published in 1827. (Type locality, Hervey Bay, Southern Queensland, Premnas unicolor Cuvier in Guérin-Méneville, 1828: Australia). Syntypes: QM I.3476–79 (1, 1, 1, 1) no page number, Livraison 13. (No locality). No Pristotis judithae Tyler, 1966: 1, Fig. 1. (Type locality, types known. Text applies to Pl. 26 (no. 19) off Northern tip of Maché Island, Seychelles, depth 60 published in 1827. feet). Holotype: ANSP 103582. Premnas unicolor Cuvier, 1829: 179. (No locality). Distribution: Persian Gulf and Red Sea to Ryukyu Syntypes: MNHN 0000-9441 (1, dry), A-8121 (1, Islands, Taiwan, Indonesia, New Guinea, and Northern dry), A-8120 (2, dry). Australia. Premnas trifasciatus Cuvier in Cuvier and Valenciennes, 1830: 405. (Type locality, Banda and Genus Similiparma Hensley, 1986 Java, Indonesia). No type known. Described as new Similiparma Hensley, 1986: 858. Feminine. Type and independent of Premnas trifasciatus Cuvier, species Glyphidodon (Parma) hermani Steindachner, 1828. 1887. Type by original designation. [Correct spelling Premnas semicinctus Cuvier in Cuvier and for genus of type species is Glyphisodon]. Valenciennes, 1830: 409, Pl. 133 (Fig. 1). (No Distribution: Tropical Eastern Atlantic. Two species. locality). Syntypes: MNHN 0000-9807 (1). Sargus ensifer Gray, 1854: 66. (Type locality, Indian Similiparma hermani (Steindachner, 1887) Ocean). Syntypes: BMNH 1853.11.12.32 [Gronovius Glyphidodon (Parma) hermani Steindachner, 1887a: 230. (Type locality, Coasts of Cape Verde Islands). coll.] (1, skin). Premnas gibbosus Castelnau, 1875: 34. (Type Holotype: NMW 16955. Species later illustrated and locality, Cape York, Queensland, Australia). described in more detail as new species in Steindachner (1887b: 59, Pl. 3) with the same date as Holotype: MNHN (not found). Premnas epigrammata Fowler, 1904: 532, Pl. 19 21 July 1887. (lower right fig.). (Type locality, Padang, Sumatra, Distribution: Cape Verde Archipelago. Indonesia). Holotype: ANSP 27665. Paratypes: Similiparma lurida (Cuvier, 1830) ANSP 27667 (1). Glyphisodon luridus Cuvier in Cuvier and Distribution: India to New Guinea, Solomon Islands, Valenciennes, 1830: 475. (Type locality, Madeira). Vanuatu and northern Queensland. Holotype: MNHN 5286. Glyphidodon lividus Vinciguerra, 1883: 617. Genus Pristotis Rüppell, 1838 Misspelling for Glyphisodon luridus attributed to Pristotis Rüppell, 1838: 128. Feminine. Type species Broussonet according Günther’s Catalog of fishes Pristotis cyanostigma Rüppell, 1838. Type by volume 4, page 56. (see Comments here below). monotypy. Distribution: Madeira, Azores, Canary Islands, Cape Daya Bleeker, 1877c: 71. Feminine. Type species Verde Islands, and Senegal. Pomacentrus jerdoni Day, 1873. Type by monotypy. Comments: Chaetodon luridus Broussonet is a nomen Distribution: Tropical Indo-West Pacific. Two species. nudum. It was reported by Gmelin (1789: 1269) in a Pristotis cyanostigma Rüppell, 1838 footnote without description based on a manuscript of Pristotis cyanostigma Rüppell, 1838: 128, Pl. 31 Broussonet (see also Cuvier, 1830: 475). Gmelin (Fig. 5). (Type locality, Massawa, Eritrea, Red Sea). wrote: “Gens Chaetodonton numerosissima, quorum Lectotype: SMF 1508. Paralectotypes: SMF 4926-31 plurium non satis curate adhuc definitorum, ideo heic (6). Lectotype established by Dor (1984: 187). omissorum, e.g Parati, militaris, […], luridi, […]

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831 definendiorum et delineandorum spem gratissimam Stegastes acapulcoensis (Fowler, 1944) facit Cl. Broiussonet […]. As a matter of fact Pomacentrus acapulcoensis Fowler, 1944a: 363, Chaetodon luridus is mentioned by Broussonet (1782: Figs. 222–223. (Type locality, Acapulco, Mexico). 36) under the description of Chaetodon faber and he Holotype: ANSP 70281. Paratypes: ANSP 70282 (1). wrote: “Hinc differ a Chaet. canescente, alepidoto, Abudefduf nigripinnis Borodin, 1928: 26, Pl. 3 (Fig. cornuto, acuminato, pinnato, argenteo, striato, 2). (Type locality, Galápagos Islands). Syntypes: aruano, citrinello, ariolatore, javo, baltheato, fabro, VMM 401 (2, disintegrated, discarded). leucuro, specioso, aulico, fugitivo, desertore, Distribution: Baja California to Peru. capistrato, nobili, bello, meleagride, lurido, chrysuro, Stegastes adustus (Troschel, 1865) perdica, gigante, plebeio, strigangulo, rotundo, Pomacentrus adustus Troschel in Müller, 1865: 633. saxatili, ciliari, vagabundo, punctato, rostrato, (Type locality, Caribbean Mexico). No types known. arcuato Pinnae Dorsalis aculeis 9.” Chaetodon luridus Pomacentrus dorsopunicans Poey, 1868: 328. (Type is again mentioned under the description of Chaetodon locality, Havana, Cuba). No types known. Original longirostris (Broussonet, 1782: 42) and he wrote: spelling was dorsopunicaus. Spelled dorso-punicans “Hinc differ a Chaet. canescente, alepidoto, cornuto, in errata (page 465) and index (page 473), both acuminato, pinnato, argenteo, striato, aruano, published later in 1868; dorsopunicans used by citrinello, ariolatore, javo, baltheato, fabro, leucuro, subsequent authors. specioso, aulico, fugitivo, desertore, capistrato, nobili, Pomacentrus obscuratus Poey, 1876: 143. (Type bello, meleagride, lurido, chrysuro, perdica, gigante, locality, Havana, Cuba). Syntypes: MCZ 14681 (4). plebeio, strigangulo, rotundo, saxatili, ciliari, Eupomacentrus rubridorsalis Beebe and Hollister, vagabundo, punctato, rostrato, arcuato, Pinnas 1933: 85, Fig. 16. (Type locality, Chatham Bay, Dorfalis aculeis 11. A Chaet. macrolepidoto, argo, Union Island, Grenadines, British West Indies). corusco, incisore, cyprinaceo, lanceolato, Rostro Holotype: USNM 170922 [ex NYZS]. Paratypes: cylindrico.” Most of these nominal species has been USNM 170923 (1). described by Linnaeus (i.e. acuminatus, alepidotus, Distribution: Southern Florida, Bahamas, and arcuatus, argenteus, argus, aruanus, canescens, Caribbean Sea. capistratus, ciliaris, cornutus, lanceolatus, leucurus, Comments: Both Stegastes adustus (Troschel, 1865) macrolepidotus, pinnatus, punctatus, rostratus, and Stegastes dorsopunicans (Poey, 1868) have been rotundus, saxatilis, striatus, vagabundus, whereas the reported as valid in the literature but Troschel's remnant has been reported by Gmelin (1789: 1269), description was earlier and should be treated as valid. i.e. ariolatoris, aulicus, baltheatus, bellus, chrysurus, Stegastes dorsopunicans erroneously dated to Poey citrinellus, coruscus, cyprinaceus, desertoris, 1863 by Allen (1991: 253). fugitivus, gigantis, javus, luridus, meleagris, nobilis, perdica, plebeius, speciosus, stellatus, strigangulus. Stegastes albifasciatus (Schlegel and Müller, 1840) Additional nominal species were mentioned by Pomacentrus albifasciatus Schlegel and Müller, Broussonet under the description of Chaetodon 1840: 21. (Type locality, NE. coast of Sulawesi, triostegus and they are paratus which differs for Indonesia). Syntypes: RMNH 938 (2). Dentibus apice serratis; unicornis, militaris, javus for Pomacentrus leucopleura Bleeker, 1854b: 85. (Type the Pinnae Dorsalis aculeis 9; lineatus, achillis, locality, Ulakan, Sumatra, Indonesia). Syntypes: defensoris, umbratus, nigricans, and stellatus for RMNH 6458 (2 of several). Corpore cingulato. Wheeler (1986) cited Gmelin as Pomacentrus eclipticus Jordan and Seale, 1906: 282, the first author reporting all of Broussonet’s Fig. 50. (Type locality, Apia, Upolu Island, Samoa). manuscript names, but we can see that these names Holotype: USNM 51739 [not 51735]. Paratypes: MCZ were first reported by Broussonet. They are nomina 36012 [ex USNM 51739] (1); CAS-SU 8718 (30), 8722 nuda, as no distinguishing features have been (14); USNM 51739 (8), 126247 [ex USBF 1259] (4). included. Distribution: Western Indian Ocean to Tuamotu Archipelago and Line Islands and Australia to Genus Stegastes Jenyns, 1840 Ryukyu Islands. Stegastes Jenyns, 1840: 62. Masculine. Type species Stegastes imbricatus Jenyns, 1840. Type by Stegastes altus (Okada and Ikeda, 1937) monotypy. Pomacentrus altus Okada and Ikeda, 1937: 87, Fig. Brachypomacentrus (subgenus of Eupomacentrus) 3; Pl. 4 (Fig. 1). (Type locality, Kowan, Okinawa Bleeker, 1877c: 73. Masculine. Type species Island, Ryukyu Islands, Japan). Holotype: YCM-P Pomacentrus albifasciatus Schlegel and Müller, 1840. 30064 [ex Okada coll. spec. 2295]. An unneeded Type by monotypy. lectotype has been designated by Hayashi (1995: 25). Eupomacentrus Bleeker, 1877c: 73. Masculine. Type Distribution: Southern Japan and Ryukyu Islands. species Pomacentrus lividus Forster in Bloch and Schneider, 1801. Type by monotypy. Stegastes apicalis (De Vis, 1885) Pomacentrus apicalis De Vis, 1885: 874. (Type Distribution: Tropical Indo-Pacific and Atlantic locality, Great Barrier Reef, Australia). Holotype: Oceans; 39 species (Allen, 1991; de Souza et al., QM I.691. 2017).

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Distribution: Cape York (Queensland) to Sydney. Stegastes fasciolatus (Ogilby, 1889) Pomacentrus niomatus De Vis, 1884: 451. (Type Stegastes arcifrons (Heller and Snodgrass, 1903) locality, probably South Sea Islands). Holotype: QM Pomacentrus arcifrons Heller and Snodgrass, 1903: I.111. 202, Pl. 7. (Type locality, Barrington Island, Pomacentrus fasciolatus Ogilby, 1889: 64. (Type Galápagos Island). Holotype: SU 6356. Paratypes: locality, Lord Howe Island). Syntypes: AMS I.1897- BMNH 1901.6.28.212-221 [ex SU] (17); MCZ 1902 (7). Original with types as I.1897-I.1902. 29456 (2); SU 6419 (13), 6536 (now 3), 51109 (2), Pomacentrus inornatus Jordan and Seale, 1906: 281. 51120 (1); ?USNM 50047–48 (4, 20), 153624 [ex Unjustified emendation of Pomacentrus niomatus De MCZ 29456] (1). Vis, 1884. Distribution: Galápagos Islands, Coco Islands, and Pomacentrus jenkinsi Jordan and Evermann, 1903: Malpelo Islands. 189. (Type locality, Hawaiian Islands). Holotype: Stegastes aureus (Fowler, 1927) USNM 50671. Paratypes: (10) ANSP 24228 (1); Pomacentrus aureus Fowler, 1927: 22, Fig. 3. (Type MCZ 28908 (1); ?CAS-SU 7942 (2), ?7479-80 (4); locality, Howland Island, central Pacific). Holotype: USNM 126552 (8). BPBM 3425. Paratypes: ANSP 109209 (1). Eupomacentrus paschalis Whitley, 1929: 225. (Type Distribution: New Caledonia, Phoenix, Samoa, locality, Easter Island). Holotype: Whereabouts Tuamotu Archipelago, and Marquesas Islands. unknown. Pomacentrus vanderbilti Fowler, 1941: 263, Fig. 14. Stegastes baldwini Allen and Woods, 1980 (Type locality, Waianae, Oahu Island, Hawaiian Stegastes baldwini Allen and Woods, 1980: 183, Fig. Islands). Holotype: ANSP 69752. Paratypes: ANSP 7. (Type locality, Clipperton Island, ca. 10°18'N, 69753-56 (4). 109°13'W, ca. 0–1.5 km north of wreck on east side Pomacentrus atrilabiatus Fowler, 1946: 146, Figs. of island, e. Pacific). Holotype: USNM 114944. 15–16. (Type locality, Aguni Shima, Ryukyu Islands, Paratypes: ANSP 75182 (181), BMNH 1957.1.21.1- Japan). Holotype: ANSP 72011 (with paratype). 10 (10), CAS 20715 (137), FMNH 61882 (54), SU Paratypes: ANSP 72012 (1). 49169 [ex W56-241] (93), USNM 114945 (38 or 37), Pseudopomacentrus navalis Whitley, 1964: 174, Fig. WAM P.25547-001 [ex ANSP 75182] (10). 8. (Type locality, Kenn Reef, Coral Sea [Queensland, Distribution: Clipperton Islands (Eastern Pacific). Australia, Southwestern Pacific]). Holotype: AMS Stegastes beebei (Nichols, 1924) IB.4970. Paratypes: AMS IB.4967–69 (1, 1, 1), Eupomacentrus beebei Nichols, 1924: 63. (Type locality, IB.4971 (1). Eden, Indefatigable [Santa Cruz] Island, Galápagos Pomacentrus craticulus Smith, 1965: 31, Pl. 7 (Fig. Islands). Holotype: AMNH 8270 [ex NYZS]. B). (Type locality, Durban, KwaZulu-Natal, Distribution: Galápagos Islands. Regarded as a Southwestern Indian Ocean, depth 2–3 fathoms). subspecies of S. leucorus by Allen (1991). Holotype: SAIAB [formerly RUSI] 293. Distribution: Three distinct populations: (1) Stegastes diencaeus (Jordan and Rutter, 1897) Hawaiian Islands; (2) West Pacific and Eastern Eupomacentrus diencaeus Jordan and Rutter, 1897: Indian Ocean; (3) Western Indian Ocean and Cocos- 116. (Type locality, Kingston, Jamaica. Lectotype: Keeling Islands. SU 4969. Paralectotypes: SU 14901 [ex SU 4969] Comments: Pomacentrus niomatus De Vis, 1884 was (1). Lectotype designated by Greenfield and Woods emended without reason to P. inornatus by Jordan 1974: 10. and Seale (1906) and as P. inornatus De Vis, 1884 Eupomacentrus mellis Emery and Burgess, 1974: was listed in synonymy with fasciolatus (Allen, 880, Figs. 1, 2A, 4A. (Type locality, 0.2 kilometers 1991). The original combination Pomacentrus southwest of Alligator Reef Light, Florida Keys, niomatus De Vis, 1884 is a valid name and threatens U.S.A., depth 7 meters). Holotype: USNM 202179. Pomacentrus fasciolatus Ogilby, 1889. The former is Paratypes: UF 205436 [ex UMML 5436] (1), 205607 a senior synonym and it has not been used as a valid [ex UMML 5607] (1), 207013 [ex UMML 7013] (1), name since 1899, while the latter has been used as a 219715 [ex UMML 19715] (1), 219960 [ex UMML valid name in over 25 publications by more than 10 19960] (2); USNM 202180 (2). authors in the last 50 years. Therefore, under Article Distribution: Southern Florida, Bahamas, and 23.9 of the Code, Pomacentrus fasciolatus Ogilby, Caribbean Sea. 1889 must be regarded as a nomen protectum and Stegastes emeryi (Allen and Randall, 1974) Pomacentrus niomatus De Vis, 1884 must be Eupomacentrus emeryi Allen and Randall, 1974: regarded as nomen oblitum. To give evidence that the 37, Fig. 1. (Type locality, off Christian's Point, conditions of the Article 23.9.1.2. are met the Pitcairn Island, 40–60 ft). Holotype: BPBM 14245. following list of publications is produced: Breder and Paratypes: AMS I.17346-001 (2); BPBM 13540 (5), Rosen (1966), Hobson (1974), Allen (1975), Brock 14246 (3). (1977), Chang et al. (1978), Chang et al. (1983), Distribution: Pitcairn Group, Oeno and Ducie Atolls, Allen and Emery (1985), Francis et al. (1987), Allen and Tuamotu Archipelago. (1991), Francis (1993), Kulbicki et al. (1994), Irving

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831 et al. (1995), Donaldson (1996), Winterbottom and Öhman (2003), Mundy (2005), Motomura et al. Anderson (1997), Fricke (1999), Nakabo (2000), (2010), Eddy (2011), Allen and Erdmann (2012), Cole (2001), Allen and Adrim (2003), Garpe and Fricke et al. (2014), Frédéric and Parmentier (2016). Stegastes flavilatus (Gill, 1862) Pomacentrus leucorus Gilbert, 1892: 554. (Type Pomacentrus flavilatus Gill, 1862: 148. (Type locality, Socorro Island, Revilla Gigedo Islands, off w. locality, Cape San Lucas, Baja California, Mexico). Mexico). Syntypes: (numerous) BMNH 1901.6.28.202- Holotype: not identified. Possible unique holotype in: 211 (13); SU 329 (3); USNM 48248–49 (1, 2), 124989 USNM 3671 (3), MCZ 36023 (1). [ex USBF 447] (2). Pomacentrus gilli Gilbert and Starks, 1904: 141, Pl. Pomacentrus elaimelas Fowler, 1944a: 381, Figs. 225– 22 (Fig. 44). (Type locality, Panama). Holotype: 227. (Type locality, Mazatlán, Sinaloa, western CAS-SU 6803. Paratypes: (12) BMNH 1903.5.15.85 Mexico). Holotype: ANSP 70261. (1), MCZ 29453 (1), CAS-SU 6906 (5), USNM Distribution: Revillagigedo Archipelago and Guadalupe 50388 (1), ZMB 15993 [ex SU] (1). Island. Distribution: Western Baja California and lower Gulf Stegastes leucostictus (Müller and Troschel, 1848) of California to Ecuador. Pomacentrus leucostictus Müller and Troschel in Stegastes fuscus (Cuvier, 1830) Schomburgk, 1848: 674. (Type locality, Barbados, Pomacentrus fuscus Cuvier in Cuvier and West Indies). Syntypes: ZMB 2760–61 (1, 1), 5730 Valenciennes, 1830: 432. (Type locality, Brazil). (2). Appeared first as above; reprinted in Ann. Mag. Holotype: MNHN 0000-8281. Nat. Hist. 1848: 18. Stegastes trindadensis Gasparini, Moura and Sazima, Pomacentrus atrocyaneus Poey, 1860: 190. (Type 1999: 5, Figs. 2–3. (Type locality, Enseada dos locality, Havana, Cuba). No types known. Portugueses, Ilha da Trinidade, 20°30' S, 29°19' W, Pomacentrus caudalis Poey, 1868: 328. (Type off State of Espírito Santo, Brazil). Holotype: locality, Havana, Cuba). Syntypes: MCZ 14682 (7). MZUSP 51243. Paratypes: MZUSP 51245–46 (1, 1), Pomacentrus analis Poey, 1868: 327. (Type locality, 51429–30 (10, 1), 51432 (4); UFPB 3547 (1); ZUEC Havana, Cuba). Syntypes: MCZ 14676 (3), 14678–79 2688–89 (1, 1), 3031 (1), 3155–56 (1, 1). (3, 1). Distribution: Continental Brazil and Trindade Island Eupomacentrus nepenthe Nichols, 1921: 1, Fig. 1. of Southern Brazil. (Type locality, Berry Islands, Bahamas, in rock pools). Holotype: AMNH 7768. Stegastes gascoynei (Whitley, 1964) Distribution: Southern Florida, Bahamas, and Pseudopomacentrus gascoynei Whitley, 1964: 173, Caribbean Sea. Fig. 7. (Type locality, Kenn Reef, Coral Sea). Holotype: AMS IB.4987. Paratypes: AMS IB.4856 Stegastes limbatus (Cuvier, 1830) (1), IB.4950 (1), IB.4984–86 (1, 1, 1), IB.4988–89 Glyphisodon limbatus Cuvier in Cuvier and (1, 1). Valenciennes, 1830: 477. (Type locality, Réunion Distribution: Capricorn Group to Northern Tasman Island). Holotype: MNHN 8487. Sea; Lord Howe Islands, Norfolk Islands to Northern Pomacentrus pristiger Valenciennes in Cuvier and New Zealand and New Caledonia. Valenciennes, 1833: 506. (Type locality, Mauritius). Holotype: MNHN 0000-8242 (poor condition). Stegastes imbricatus Jenyns, 1840 Pomacentrus madagascariensis Sauvage, 1882: 174. Stegastes imbricatus Jenyns, 1840: 63, 165, Pl. 9 (Type locality, Madagascar). Holotype: MNHN A-1996. (Fig. 2). (Type locality, Quail Island, Porto Praya, Pomacentrus (Brachypomacentrus) ater Bleeker, (ex Cape Verde Islands Holotype: BMNH 1917.7.14.45. Liénard), 1879: 5, Pl. 2. (Type locality, Mauritius). ? Pomacentrus hamyi Rochebrune, 1880: 165. (Type Lectotype: ZMH 17035. locality, Casamance River, Senegal). Holotype: lost. Pomacentrus ater Liénard in Sauvage, 1891: 428. Distribution: Canary Islands, Capo Verde Islands and (Type locality, Mauritius, Mascarenes, southwestern Senegal to Angola. Indian Ocean). No types known. Objectively invalid; Stegastes insularis Allen and Emery, 1985 nomen praeoccupatum, preoccupied by Pomacentrus Stegastes insularis Allen and Emery, 1985: 19, Pl. 2 ater Bleeker, 1879. (Figs. F, G). (Type locality, off Ethel Beach, Distribution: Madagascar, Réunion, and Mauritius. Christmas Island, e. Indian Ocean, 3–6 m). Holotype: Stegastes lividus (Bloch and Schneider, 1801) WAM P.26083-044. Paratypes: AMS I.22798-001 Chaetodon lividus Forster in Bloch and Schneider, (1); BMNH 1982.3.12.6 (1); BPBM 7114–16 (10, 7, 1801: 235. (Type locality, Pacific). No types known. 9); MNHN 1982-125 (1); ROM 39311 (5); SMF Stegastes robertsoni Randall, 2001: 103, Figs. 7–9. 17586 (1); USNM 231752 (1); WAM P.10868 (1), (Type locality, Taiohae Bay, west side at head of P.10873 (1), P.26083-002 (6), P.26088-005 (4), bay, Nuku Hiva, Marquesas Islands, depth 5 meters). P.26093-006 (1), P.26110-010 (4), P.26120-006 (1). Holotype: BPBM 38679. Paratypes: AMS I.39936 Distribution: Christmas Island and Marcus Islands (1); BPBM 10413 (6), 12565 (11), 38685 (3); CAS (Western Pacific). 210418 (1); MNHN 2000-8 (1); NSMT-P 57706 (1); Stegastes leucorus (Gilbert, 1892) USNM 89765 (17), 359633 (1).

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Distribution: Marquesas Islands with other records as Stegastes otophorus (Poey, 1860) misidentification of Stegastes punctatus. Pomacentrus otophorus Poey, 1860: 188. (Type locality, Cuba). Described syntype: USNM 0 (1). Stegastes lubbocki Allen and Smith, 1992 Distribution: Jamaica, Atlantic coast of Panama, and Stegastes lubbocki Allen and Smith, 1992: 114, Fig. Cuba. 1. (Type locality, Pratt Point, Ascension Island, 7°57'S, 14°22'W). Holotype: WAM P.30283-001. Stegastes partitus (Poey, 1868) Paratypes: BMNH 1992.4.6.1-8 (8); USNM 320806 Pomacentrus partitus Poey, 1868: 327 (Type (4); WAM P30279-001 (7), P30280-001 (1), P30281- locality, Havana, Cuba). Holotype: MCZ 14680. 001 (1), P30282-001 (2), P30284-001 (16). Pomacentrus freemani Beebe and Tee-Van, 1928: Distribution: Ascension Island. 196, unnumb. Fig. (Type locality, Sand Cay, Port-au- Prince Bay, Haiti, West Indies). Holotype: USNM Stegastes luteobrunneus (Smith, 1960) 170898 [ex NYZS 7269]. Paratypes: USNM 170897 Pomacentrus luteobrunneus Smith, 1960: 343, Pls. [ex NYZS 7347] (6), 170898 [ex NYZS 7269] (1). 26 (Fig. F), 27 (Fig. F). (Type locality, Aldabra Distribution: Southern Florida, Bahamas, and Island, Seychelles, Western Indian Ocean). Holotype: Caribbean Sea. SAIAB [formerly RUSI] 65. Paratypes: SAIAB [formerly RUSI] 18553 (1). Stegastes pelicieri Allen and Emery, 1985 Distribution: Red Sea, Western Indian Ocean: East Stegastes pelicieri Allen and Emery 1985: 27, Pl. 3 Africa to Madagascar. (Figs. G, H). (Type locality, Off Flic en Flac village, Mauritius, 6 m). Holotype: WAM P.26522-002. Stegastes marginatus (Jenkins, 1901) Paratypes: BMNH 1982.3.12.7 (1); BPBM 27983 (2); Eupomacentrus marginatus Jenkins, 1901: 391, Fig. MNHN 1982-126 (1); RUSI 1345 (2); USNM 231753 5. (Type locality, Honolulu, Oahu Island, Hawaiian (1); WAM P.26520-002 (3), P.26522-001 (2). Islands). Holotype: USNM 49700. Paratypes: CAS- Distribution: Mauritius and possibly Réunion. SU 2470 (3), 6094 (1), 7480 (1), 7942 (2), 23331 (now 19); ?FMNH 4257-59 (3). Stegastes pictus (Castelnau, 1855) Distribution: Hawaiian Islands. Pomacentrus pictus Castelnau, 1855: 9, Pl. 2 (Fig. 1). (Type locality, Bahia, Brazil). Holotype: MNHN Stegastes nigricans (Lacepède, 1802) 0000-8280. Holocentrus nigricans Lacepède (ex Commerson), Distribution: Brazil. 1802: 332, 367. (No locality, probably Mauritius). No types known. Based on Commerson manuscript. Stegastes planifrons (Cuvier, 1830) Pomacentrus scolopseus Quoy and Gaimard, 1825: Pomacentrus planifrons Cuvier in Cuvier and 398. (Type locality, Mauritius). Holotype: not found. Valenciennes, 1830: 431. (Type locality, Martinique Pomacentrus taeniops Cuvier in Cuvier and Island, West Indies). Holotype: MNHN 547. Valenciennes, 1830: 423. Partim. (Type locality, Distribution: Southern Florida, Bahamas, and Mauritius; Bora Bora; Guam, Mariana Islands, Caribbean Sea. Western Pacific). Syntypes: MNHN 0000-1378 (1), Stegastes punctatus (Quoy and Gaimard, 1825) 0000-8144 (1), 0000-8244 (1), 0000-8246 (1), 0000- Pomacentrus punctatus Quoy and Gaimard, 1825: 8247 (1). 395, Pl. 64 (Fig. 1). (Type locality, Mauritius). Pomacentrus scolopsis Günther, 1862: 28. Holotype: MNHN 0000-8505. Unjustified emendation of Pomacentrus scolopseus Pomacentrus prosopotaenioides Bleeker, 1852b: Quoy and Gaimard, 1825. 286. (Type locality, Ambon Island, Molucca Islands, Pomacentrus subniger De Vis, 1885: 873. (Type Indonesia). Syntypes: RMNH 6470 (2 of 22). locality, Cardwell, Queensland, Australia). Holotype: Pomacentrus cyanospilos Bleeker, 1853a: 709. (Type QM I.80. locality, Wahai, Ceram, Indonesia). Syntypes: (9) Abudefduf teemai Aoyagi, 1941b: 230, Fig. 33. (Type RMNH 6470 (some of 22). locality, Miyako Island, Ryukyu Islands, Japan). Pomacentrus obscurus Thiollière in Montrouzier Holotype: not at YCM, apparently lost. 1857: 478. (Type locality, Woodlark Island, South Distribution: Red Sea and Western Indian Ocean to Pacific). No types known. Oceania eastward to Line Islands., Tuamotu Pomacentrus vitianus Sauvage, 1879: 206. (Type Archipelago, and Marquesas Islands excluding locality, Fiji Islands). Syntypes: MNHN 0000-9897 (4). Hawaiian Islands; also Australia to Ryukyu Islands. Distribution: Red Sea and Western Indian Ocean to Stegastes obreptus (Whitley, 1948) Oceania Eastward to Tuamotus Islands, excluding Pomacentrus obreptus Whitley, 1948: 275. (Type Hawaiian Islands; also Australia to Ryukyu Islands. locality, Monte Bello Islands, Western Australia). Stegastes rectifraenum (Gill, 1862) Holotype: AMS IB.1571. Paratypes: AMS IB.1572 Pomacentrus rectifraenum Gill, 1862: 148. (Type (1). locality, Cape San Lucas, Baja California, Mexico). Distribution: India, Sri Lanka to Western Australia Syntypes: (6) BMNH 1866.5.21.15 [ex USNM] (1), North to Ryukyu Islands. USNM 3670 (4).

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Pomacentrus analigutta Gill in Günther, 1862: 27. Distribution: Gulf of Mexico, Bahamas, Caribbean (Type locality, Cape San Lucas, Baja California). Sea to Southern Brazil. Syntypes: MCZ 35984 [ex USNM 3674] (1), USNM Comments: Further studies may reveal that the 3674 (orig. 20, now 13). Caribbean form is a cryptic species that has been Distribution: Gulf of California. confused with S. variabilis. Stegastes redemptus (Heller and Snodgrass, 1903) Stegastes xanthurus (Poey, 1860) Pomacentrus redemptus Heller and Snodgrass, 1903: Pomacentrus xanthurus Poey, 1860: 190. (Type 200, Pl. 6. (Type locality, Clarion Island, Revilla locality, Havana, Cuba). Syntypes and/or Poey Gigedo [Revillagigedo] Islands, off W. Mexico). specimens: MCZ 151392 [ex MCZ 14677a] (3). Holotype: SU 6358. Paratypes: SU (6, missing). Distribution: Florida Keys and the Gulf of Mexico to Distribution: Revillagigedo Archipelago and Cabo Northern South America. San Lucas (Baja California). Genus Teixeirichthys Smith, 1953 Stegastes rocasensis (Emery, 1972) Teixeirichthys Smith, 1953: 13. Masculine. Type Eupomacentrus rocasensis Emery, 1972: 330, Figs. species Teixeirichthys mossambicus Smith, 1953. 1–3. (Type locality, Lagoon off Atol das Rocas, n. Type by original designation. Brazil, 3°56'S, 33°49'W, 0.3 m). Holotype: USNM Wangia Fowler, 1954: 54. Feminine. Type species 202177. Paratypes: ANSP 109422 (4); FMNH 74077 Pomacentrus formosanus Fowler and Bean, 1922. (12); UF 19286 [ex TABL 100816] (orig. 7, now 4), Type by original designation. Not preoccupied by 223693 [ex UMML 23693] (26); USNM 202178 (4), Wanga Chen (1943) in Mollusca, Gastropoda, 208561 [ex TABL 100816] (3). Prosobranchia, Cerithioidea. Replaced by Wangietta Stegastes sanctipauli Lubbock and Edwards, 1981: Whitley (1965). 144, Fig. 3. (Type locality, North side of Belmonte Wangietta Whitley, 1965: 26. Feminine. Type Islet, Saint Paul's Rocks, 0°56' N, 29°22' W, Atlantic, species Pomacentrus formosanus Fowler and Bean, 5–10 m). Holotype: MZUSP 14604. Paratypes: 1922. Type by being a replacement name. Unneeded BMNH 1980.1.22.3-4 (2), 1980.1.22.5-6 (1, 1); replacement name for Wangia Fowler, 1954, thought MHNG 2014.25-30 (6); MZUSP 14605-09 (5); SMF to be preoccupied by Wanga Chen, 1943 in 15679 (6). Mollusca. Distribution: Saint Paul's Rocks (mid Atlantic) and Tropical Indo-West Pacific. One species. One Atol das Rocas (250 Km off Northern Brazil). undescribed species has been reported from Norfolk Comments: Recent studies support the conclusion Island by Francis (1993). that S. sanctipauli is a junior synonym of S. Comments: Fowler, probably unaware of rocasensis, however they do not exclude that the two Teixeirichthys Smith, erected the genus Wangia to nominal species may represent species in statu accommodate the distinctive species Pomacentrus nascendi (de Souza et al., 2017). formosanus. Whitley (1965) erroneously reported Wangia Fowler, 1954 to be preoccupied, but the Stegastes sanctaehelenae (Sauvage, 1879) original name is actually Wanga Chen, 1943 (see Pomacentrus sanctaehelenae Sauvage 1879: 207. References). Whitley’s error has been reproduced in (Type locality, Lemon Valley Bay, Saint Helena the ECoF by Fricke et al. (2020). Island, South Atlantic, 5–7 m. [original locality was St. Helena]). Neotype: MNHN 1986-450 designated Teixeirichthys jordani (Rutter, 1897) by Edwards and Glass 1987: 653. Pomacentrus jordani Rutter, 1897: 77. (Type Distribution: Saint Helena Islands. locality, Swatow, Shantou, coast of se. China). Holotype: SU 1760. Stegastes uenfi Novelli, Nunan and Lima, 2000 Pomacentrus polylepis Regan, 1908: 229, Pl. 24 (Fig. Stegastes uenfi Novelli, Nunan and Lima, 2000: 2, 3). (Type locality, Cargados Carajos, Indian Ocean, Fig. 1. (Type locality, Arraial do Cabo, Prainha depth 20–30 fathoms). Syntypes: BMNH beach, 22°57'S, 42°10'W, Rio de Janeiro, Brazil, 1908.3.23.99-100 (2). depth 3 meters). Holotype: MNRJ 17239. Paratypes: Pomacentrus formosanus Fowler and Bean, 1922: MNRJ 17235–37 (1, 1, 2), 17241 (3). 46, Fig. 4. (Type locality, Kaohsiung [Takao], Distribution: Brazil. Taiwan). Holotype: USNM 76644. Stegastes variabilis (Castelnau, 1855) Teixeirichthys mossambicus Smith, 1953: 14, Fig. 1. Pomacentrus variabilis Castelnau 1855: 9, Pl. 3 (Fig. (Type locality, Mozambique Channel, Western Indian 3). (Type locality, Bahia, Brazil). Syntypes: MNHN Ocean). Holotype: SAIAB [formerly RUSI] 66. 0000-8135 (2). Distribution: Red Sea and coasts of East Africa, Pomacentrus flaviventris Troschel in Müller 1865: India, Sri Lanka to Ryukyu Islands and Southern 633. (Type locality, Atlantic). No types known. Japan. Eupomacentrus chrysus Bean, 1908: 32. (Type locality, White's Flat Channel, Bermuda). Holotype: FMNH 5025.

Journal of Animal Diversity (2021), 3 (1): 37–109 | www.jad.lu.ac.ir 85 Parenti 86

Nominal species incertae sedis South Pacific. Placed in synonymy with A. vaigiensis. Abudefduf chengi Lin, 1934: 683, Fig. 2. (Type locality, Fuchow, Fukien, China). Holotype: FESC Glyphisodon geant Liénard, 1839: 35. Vernacular (now at PRFRI) 16. Species inquirendae in name. Chrysiptera. Glyphisodon gigas Allen (ex Liénard), 1991: 234. Name published in synonymy in Allen and without Abudefduf trilineatus Wang, 1941: 100. (Type diagnosis. locality, Hainan Island, China). Holotype: SSCN. Typesetting error as trileniatus on page 100, but Pomacentrus kumkum Montrouzier, 1857: 478. species is correctly spelled as trilineatus in abstract Manuscript name mentioned in passing in synonymy and in summary on page 119. Species inquirendae under two species; Woodlark Island, South Pacific. (Allen, 1991: 254). Nannapogon polynesiae Fowler, 1938: 143, Pl. 9 Acknowledgements (Fig. 21). (Type locality, Papeete, Tahiti, Society I thank Kyle R. Luckenbill from the Academy of Islands). Holotype: ANSP 68399. Paratypes: ANSP Natural Sciences of Drexel University for providing 68400 (1), 68401-14 (14), 68415 (1, missing). the photos of the holotype of Chrysiptera personata Species based on a young specimen (Randall and Edda Aßel, Collection Manager, Ichthyology, annotation). Museum für Naturkunde, Berlin for providing the Pomacentrus albolineatus Montalban, 1928: 70, Pl. photographs of the Syntypes: ZMB 2738 Glyphisodon 14 (Fig. 1). (Type locality, Bungau Island, Sulu sculptus and ZMB 2739 Glyphisodon fallax. I thank Archipelago, Philippines). Lectotype: CAS-SU Ronald Fricke for help in retrieving some original 25571. Paralectotypes: BSMP (1, apparently papers and the Biodiversity Heritage Library for destroyed). Lectotype selected by Böhlke 1953: 83. allowing a huge amount of old books and journals to Species inquirenda. (Allen, 1991: 254). be freely available through the internet. I also would like to thank two anonymous reviewers for their Pomacentrus modestus Castelnau, 1875: 35. (Type valuable comments on this work. locality, Gulf of Carpentaria, Queensland, Australia). Holotype: MNHN A-0702. Species inquirenda. Conflict of interest (Allen, 1991: 254). Possibly a Neopomacentrus. The author declares that there are no conflicting Pomacentrus pingi Wang, 1941: 93. (Type locality, issues related to this review article. Hainan Island, China). Holotype: ? SSCN. Species inquirenda. (Allen, 1991: 254). References Pomacentrus profundus De Vis, 1885: 873. (Type locality, Great Barrier Reef, Australia). Holotype: Akimichi, T. and Sauchomal, S. (1982). Satawalese ?QM I.4518 (apparently not the type). Species fish names. Micronesica, 18(2): 1–34. inquirenda. Status uncertain (Allen, 1991: 254). Alcala, A. C., Ingles, J. A. and Bucol, A. A. (2008). Review of the biodiversity of southern Philippines Unavailable names seas. The Philippine Scientist, 45: 1–61. Abudefduf multifasciatus Seale, 1906: 53. No https://doi.org/10.3860/psci.v45i0.991 distinguishing features. Allen G. R. (1975a). Four new damselfishes Glyphisodon angulatus Bleeker, 1845: 523. No (Pomacentridae) from the southwest Pacific. distinguishing features; from Batavia, Java. Placed in Proceedings of the Linnean Society of New South synonymy with Chromis cinerascens (Cuvier, 1830) Wales, 99 (pt 2): 87–99. (Allen 1991: 239). Allen, G. R. (1972). The Anemonefishes. Their Chromis fourmanoiri Rivaton et al., 1990. Nomen classification and biology. T. F. H. Publications, nudum. Reported without authorship on page 63 of Neptune City, NJ. 1–288. Allen and Randall, 1974 and on page 86 in the Allen, G. R. (1973a). Three new species of deep- checklist of fishes from New Caledonia prepared by dwelling damselfishes (Pomacentridae) from the Rivaton et al. (1989); not reported by Allen (1991). south-west Pacific Ocean. Australian Zoologist, Not in the ECoF. 18 (pt 1): 31–42. Chromis profunda Allen and Randall, 1974 nomen Allen, G. R. (1973b). Amphiprion leucokranos, a nudum. From a checklist of fishes from New new species of pomacentrid fish, with notes on Caledonia prepared by Rivaton et al., 1990; other anemonefishes of New Guinea. Pacific apparently a species to be described. Science, 27 (4): 319–326. Glyphisodon biniar Montrouzier, 1857: 477. Name Allen, G. R. (1975b). Damselfishes of the South mentioned in passing in synonymy under Seas. T.F.H. Publ., Inc., Neptune City, New Glyphisodon waigiensis? from Woodlark Island, Jersey. 1–240, pls.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

Allen, G. R. (1976). Two new species of Allen, G. R. (2018). Chromis torquata, a new species damselfishes (Pomacentridae) from Western of damselfish (Pomacentridae) from Mauritius Australia. Records of the Western Australian and Réunion. aqua, International Journal of Museum, 4 (pt 2): 133–144. Ichthyology, 24 (1): 15–22. Allen, G. R. (1987a). Chrysiptera sinclairi, a new Allen, G. R. and Adrim, M. (1992). A new species of species of damselfish from the tropical western damselfish (Chrysiptera: Pomacentridae) from Pacific Ocean. Revue française d'Aquariologie Irian Java, Indonesia. Records of the Western Herpétologie, 13 (4): 107–110. Australian Museum, 16: 103–108. Allen, G. R. (1987b). A new species of pomacentrid Allen, G. R. and Adrim, M. (2000). Amblypomacentrus fish with notes on other damselfishes of the clarus, new species of damselfish (Pomacentridae) Kermadec Islands. Records of the Western from the Banggai Islands, Indonesia. Records of the Australian Museum, 13 (pt 2): 263–273. Western Australian Museum, 20: 51–55. Allen, G. R. (1987c). New Australian fishes. Part 3. Allen, G. R. and Adrim, M. (2003). Coral reef fishes A new species of Pomacentrus (Pomacentridae). of Indonesia. Zoological Studies, 42 (1): 1–72. Memoirs of the Museum of Victoria, 48 (1): 9–11. Allen, G. R. and Bailey, S. (2002). Chrysiptera Allen, G. R. (1989). Freshwater Fishes of Australia. albata, a new species of damselfish from Phoenix T.F.H. Publications, Neptune City, New Jersey, Island. aqua, Journal of Ichthyology and Aquatic 240 pp. Biology, 6 (1): 39–43. Allen, G. R. (1991). Damselfishes of the world. Allen, G. R. and Drew, J. A. (2012). A new species Mergus Publisher, Melle, Germany. 271 pp. of damselfish (Pomacentrus; Pomacentridae) Allen, G. R. (1992). A new species of damselfish from Fiji and Tonga. aqua, International Journal (genus Pomacentrus) from north-western of Ichthyology, 18 (3): 171–180. Australia. Records of the Western Australian Allen, G. R. and Emery, A. R. (1973). Pomacentrus Museum, 15 (4): 691–695. exilis, a new species of damselfish from the Allen, G. R. (1993). Two new species of central-west Pacific. Copeia, 1973 (3): 565–568. damselfishes (Pomacentrus), with comments on https://doi.org/10.2307/1443121 the validity of two additional pomacentrid fishes. Allen, G. R. and Emery, A. R. (1985). A review of Revue française d'Aquariologie Herpétologie, 20 the pomacentrid fishes of the genus Stegastes (no. 1): 21–26. from the Indo-Pacific, with descriptions of two Allen, G. R. (1993). Two new species of damselfishes new species. Indo-Pacific Fishes, 3: 1–31. (Pomacentrus) with comments on the validity of Allen, G. R. and Erdmann, M. V. (2005). Chromis two additional pomacentrid fishes. Revue française xouthos, a new species of damselfish (Pomacentridae) d'Aquariologie Herpétologie, 20: 21–26. from the East Andaman Sea and central Indian Allen, G. R. (1995). Two new species of damselfish Ocean. aqua, Journal of Ichthyology and Aquatic (Pomacentridae) from Indonesian Seas. Revue Biology, 10 (3): 89–94. française d'Aquariologie Herpétologie, 21 (3–4): Allen, G. R. and Erdmann, M. V. (2008a). A new 86–90. species of damselfish (Pomacentridae: Chrysiptera) Allen, G. R. (1997). Marine fishes of tropical from western New Guinea and the Togean Islands, Australia and south-east Asia. Western Australian Indonesia. aqua, International Journal of Museum, Perth. 292 pp. Ichthyology, 13 (3–4): 171–178. Allen, G. R. (1999a). Three new species of Allen, G. R. and Erdmann, M. V. (2008b). A new damselfishes (Pomacentridae) from Indonesia and species of damselfish (Chromis: Pomacentridae) eastern Papua New Guinea. Revue française from the Raja Ampat Islands, Papua Barat d'Aquariologie Herpétologie, 25 (3/4): 99–105. Province, Indonesia. aqua, International Journal of Ichthyology, 14 (4): 203–208. Allen, G. R. (1999b). Altrichthys, a new genus of damselfishes (Pomacentridae) from Philippine Allen, G. R. and Erdmann, M. V. (2009a). Two new Seas with description of a new species. Revue species of damselfishes (Pomacentridae: française d'Aquariologie Herpétologie, 26 (1–2): Chromis) from Indonesia. aqua, International 23–28. Journal of Ichthyology, 15 (3): 121–134. Allen, G. R. (2002). Descriptions of two new Allen, G. R. and Erdmann, M. V. (2009b). A new species species of damselfishes from Madagascar. of damselfish (Pomacentrus: Pomacentridae) from aqua, International Journal of Ichthyology, 6 western New Guinea. Records of the Western (2): 45–52. Australian Museum, 25 (pt 2): 121–126. Allen, G. R. (2004). Pomacentrus aurifrons, a new https://doi.org/10.18195/issn.0312-3162.25(2).2009.121-126 species of damselfish (Pomacentridae) from Allen, G. R. and Erdmann, M. V. (2012). Reef fishes Melanesia. Zootaxa, 399 (1): 1–7. of the East Indies. Volumes I–III. Tropical Reef https://doi.org/10.11646/zootaxa.399.1.1 Research, Perth Australia. [v. I: x + 1–424 + end

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note; v. II: 425–855; v. III: preface, map, contents Pacific pomacentrid genus Amblyglyphidodon, and 857–1260; including Appendix 1 (new species with descriptions of two new species. aqua, descriptions) and Appendix II (addendum)]. Journal of Ichthyology and Aquatic Biology, 5 Allen, G. R. and Erdmann, M.V. (2014). A new (4): 139–152. species of damselfish (Chromis: Pomacentridae) Allen, G. R. and Randall, J. E. (2004a). Two new from Papua New Guinea. Journal of the Ocean species of damselfishes (Pomacentridae: Chromis) Science Foundation, 11: 13–22. from Indonesian seas. aqua, Journal of Ichthyology https://doi.org/10.5281/zenodo.1044523. and Aquatic Biology, 9 (1): 17–24. Allen, G. R. and Erdmann, M.V. (2020). Chromis Allen, G. R. and Randall, J. E. (2004b). Two new pacifica, a new Pacific Ocean damselfish distinct species of damselfishes (Pomacentridae) from from Indian Ocean Chromis agilis (Teleostei: Micronesia. aqua, Journal of Ichthyology and Pomacentridae). Journal of the Ocean Science Aquatic Biology, 9 (2): 75–87. Foundation, 35: 102–117. Allen, G. R. and Randall, J. E. (2005). A new species https://doi.org/10.5281/zenodo.3988552 of damselfish (Pomacentrus: Pomacentridae) Allen, G. R., Erdmann, M.V. and (2020). from Fiji. aqua, Journal of Ichthyology and Pomacentrus andamanensis, a new species of Aquatic Biology, 10 (3): 95–101. damselfish (Teleostei: Pomacentridae) from the Allen, G. R. and Robertson, D. R. (1974). Description Andaman Sea, Indian Ocean. Journal of the of four new damselfishes (Pomacentridae) from Ocean Science Foundation, 36: 38–48. Papua New Guinea and eastern Australia. Records https://doi.org/10.5281/zenodo.4340748 of the Australian Museum, 29 (4): 153–167. Allen, G. R. and Hoese, D. F. (1975). A review of the Allen, G. R. and Smith, K. N. (1992). A new species pomacentrid fish genus Parma, with description of damselfish (Pomacentridae: Stegastes) from of two new species. Records of the Western Ascension Island, Atlantic Ocean. Records of the Australian Museum, 3 (4): 261–293. Western Australian Museum, 16: 113–117. Allen, G. R. and Larson, H. K. (1975). Pomachromis Allen, G. R. and Smith-Vaniz, W. F. (1994). Fishes guamensis, a new species of damselfish of the Cocos (Keeling) Islands. Atoll Research (Pomacentridae) from the Mariana Islands. Bulletin, 412: 1–21. Micronesica, 11 (1): 123–126. Allen, G. R. and Steene, R. C. (1988). Fishes of Allen, G. R. and Larson, H. K. (1979). Parma bicolor, Christmas Island Indian Ocean. Christmas Island a new species of damselfish from southwestern Natural History Association, Christmas Island, Australia. Revue française d'Aquariologie Indian Ocean, 6798, Australia. 197 pp. Herpétologie, 6 (1): 11–14. Allen, G. R. and R. Swainston (1988). The marine Allen, G. R. and Lubbock, R. (1976). Glyphidodontops fishes of north-western Australia. A field guide springeri, a new species of damselfish for anglers and divers. Western Australian (Pomacentridae) from the Philippine and Molucca Museum, Perth. vi + 201, 70 Pls. Islands. Proceedings of the Biological Society of Allen, G. R. and Woods, L. P. (1980). A review of Washington, 88 (31): 345–350. the damselfish genus Stegastes from the eastern Allen, G. R. and Rajasuriya, A. (1994). Chrysiptera Pacific with description of a new species. kuiteri, a new species of damselfish Records of the Western Australian Museum, 8 (Pomacentridae) from Indonesia and Sri Lanka. (2): 171–198. Records of the Western Australian Museum, 17 Allen, G. R. and Wright, J. E. (2003). Description of (pt 3): 283–286. a new species of damselfish (Pomacentridae: Allen, G. R. and Randall, J. E. (1974). Five new Pomacentrus) from Rodrigues Island, Indian species and a new genus of damselfishes (family Ocean. aqua, Journal of Ichthyology and Aquatic Pomacentridae) from the South Pacific Ocean. Biology, 7 (4): 133–138. Tropical Fish Hobbyist, 22 (9): 36–46, 48–49. Allen, G. R., Drew, J. A. and Fenner, D. (2010). Allen, G. R. and Randall, J. E. (1980). A review of Amphiprion pacificus, a new species of the damselfishes (Teleostei: Pomacentridae) of anemonefish (Pomacentridae) from Fiji, Tonga, the Red Sea. Journal of Zoology, 29 (1/3): 1–98. Samoa, and Wallis Island. aqua, International https://doi.org/10.1080/00212210.1980.10688486 Journal of Ichthyology, 16 (3): 129–138. Allen, G. R. and Randall, J. E. (1985). A new species Allen, G. R., Drew, J. A. and Kaufman, L. (2008). of damselfish (Pomacentridae) from eastern Amphiprion barberi, a new species of Australia and the Norfolk Island Ridge. Records anemonefish (Pomacentridae) from Fiji, Tonga, of the Western Australian Museum, 12 (pt 2): and Samoa. aqua, International Journal of 241–245. Ichthyology, 14 (3): 105–114. Allen, G. R. and Randall, J. E. (2002). A review of Allen, G. R., Erdmann, M. V. and Barber, P. H. the leucogaster species complex of the Indo- (2010). A new species of damselfish

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

(Chrysiptera: Pomacentridae) from Papua New Australia. Journal of the Ocean Science Guinea and eastern Indonesia. aqua, International Foundation, 29: 1–10. Journal of Ichthyology, 16 (2): 61–70. http://dx.doi.org/10.5281/zenodo.1066344 Allen, G. R., Erdmann, M. V. and Cahyani. N. K. D. Alleyne, R. H and Macleay, W. (1877). The (2015). Review of the Chrysiptera oxycephala ichthyology of the Chevert expedition. complex of damselfishes (Pomacentridae) with Proceedings of the Linnean Society of New South descriptions of three new species from the East Wales, 1 (pts 3–4): 261–281, 321–359. Indian Archipelago. Journal of the Ocean Science Anonymous [Bennett, E. T.] (1830). Class Pisces. Pp. Foundation, 17: 56–84. 686–694. (In: Memoir of the life and public http://dx.doi.org/10.5281/zenodo.891435 services of Sir Thomas Stamford Raffles.. By his Allen, G. R., Erdmann, M. V. and Cahyani. N. K. D. Widow [Lady Stamford Raffles]. John Murray, (2017). A new species of damselfish London. 701 pp. (Chrysiptera: Pomacentridae) from coral reefs of Aoyagi, H. (1941). One new species of the the Solomon Islands. Journal of the Ocean Pomacentridae, Pisces, from the Palau Islands. Science Foundation, 28: 10–21. Dobutsugaku Zasshi, Zoological Magazine http://dx.doi.org/10.5281/zenodo.891041 Tokyo, 53 (3): 180–181. Allen, G. R., Erdmann, M. V. and Cahyani. N. K. D. Aoyagi, H. (1941b). The damsel fishes found in the (2018). Chrysiptera uswanasi, a new waters of Japan. Biogeographica, Tokyo, 4 (1): microendemic species of damselfish (Teleostei: 157–279. Pomacentridae) from West Papua Province, Arango, B. G., Pinheiro, H. T., Rocha, C., Greene, B. Indonesia. Journal of the Ocean Science D., Pyle, R. L., Corpus, J. M., Shepherd, B. and Foundation, 31: 74–86. Rocha, L. A. (2019). Three new species of http://dx.doi.org/10.5281/zenodo.1482015 Chromis (Teleostei, Pomacentridae) from Allen, G. R., Erdmann, M. V. and Hidayat, N. I. mesophotic coral ecosystems of the Philippines. (2018). Pomacentrus bellipictus, a new ZooKeys, 835: 1–15. microendemic species of damselfish (Pisces: https://doi.org/10.3897/zookeys.835.27528 Pomacentridae) from the Fakfak Peninsula, West Baschieri–Salvadori, F. (1955). Spedizione subacquea Papua, Indonesia. Journal of the Ocean Science italiana nel Mar Rosso. Ricerche zoologiche. IX. Foundation, 30: 1–10. Pomacentridi. Rivista di Biologia Coloniale, 15: http://dx.doi.org/10.5281/zenodo.1246885 57–68. Allen, G. R., Erdmann, M. V. and Hilomen, V. V. Bauchot, M. L., Desoutter, M. and Allen, G. R. (2011). A new species of damselfish (Pomacentrus: (1978). Catalogue critique des types des poissons Pomacentridae) from Brunei and the Philippines. du Muséum national d'Histoire Naturelle. Bulletin Aqua, International Journal of Ichthyology, 17 (1): du Muséum National d'Histoire Naturelle (Série 35–42. 3: Zoologie), 1: 1–56. Allen, G. R., Erdmann, M. V. and Kurniasih, E. M. Bean, T. H. (1908). Descriptions of new Bermudan (2015). Chrysiptera caesifrons, a new species of fishes. Proceedings of the Biological Society of damselfish (Pomacentridae) from the south– Washington, 19: 29–33. western Pacific Ocean. Journal of the Ocean Science Foundation, 15: 16–32. Beebe, W. and Hollister, G. (1933). New species of http://dx.doi.org/10.5281/zenodo.891445 fish from the West Indies. Zoologica, Scientific Contributions of the New York Zoological Allen, G. R., Erdmann, M. V. and Pertiwi, P. D. Society, 12 (9): 83–88. (2017a). Descriptions of four new species of damselfishes (Pomacentridae) in the Pomacentrus Beebe, W. and Tee-Van, J. (1928). The fishes of philippinus complex from the tropical western Port.au-Prince Bay, Haiti, with a summary of the Pacific Ocean. Journal of the Ocean Science known species of marine fishes of the island of Foundation, 25: 47–76. Haiti and Santo Domingo. Zoologica, Scientific http://dx.doi.org/10.5281/zenodo.317395 Contributions of the New York Zoological Society, 10 (1): 1–279. Allen, G. R., Erdmann, M. V. and Pertiwi, P. D. (2017b). Pomacentrus flavioculus, a new species Bennett E. T. (1828–30). A selection from the most of damselfish from Fiji and Tonga (Teleostei: remarkable and interesting fishes found on the Pomacentridae). Journal of the Ocean Science coast of Ceylon. First Edition. Eduard Bulletin. Foundation, 28: 22–33. London, viii + 30 unnumbered pp., Pls. 1–30. http://dx.doi.org/10.5281/zenodo.896910 Bennett E. T. (1831). The exhibition of the collection of fishes formed at the Mauritius by Mr. Telfair, Allen, G. R., Moore, G. I. and Allen, M. G. (2017). portions of which had been brought before the Neopomacentrus aktites, a new species of Committee at the meeting in April, was resumed. damselfish (Pisces: Pomacentridae) from Western From among them Mr. Bennett pointed out more

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particularly the following species which he Bleeker, P. (1852b). Bijdrage tot de kennis der believed to have been previously undescribed. ichthijologische fauna van de Moluksche Proceedings of the Committee of Science and Eilanden. Visschen van Amboina en Ceram. Correspondence of the Zoological Society of Natuurkundig Tijdschrift voor Nederlandsch London, 1830–31 (1): 126–128. Indië, 3 (2): 229–309. Bennett, E. T. (1832). Observations on a collection of Bleeker, P. (1852c). Nieuwe bijdrage tot de kennis der fishes from the Mauritius presented by Mr. ichthyologische fauna van Amboina. Natuurkundig Telfair with character of new genera and species. Tijdschrift voor Nederlandsch Indië, 3 (4): 545–568. Proceedings of the Zoological Society of London, Bleeker, P. (1852d). Diagnostische beschrijvingen 1831: 165–169. van nieuwe of weinig bekende vischsoorten van Bennett, J. W. (1830). A Selection from the Most Sumatra. Tiental I – IV. Natuurkundig Tijdschrift Remarkable and Interesting Fishes Found on the voor Nederlandsch Indië, 3 (4): 569–608. Coast of Ceylon. London. First edition. Part 3: Bleeker, P. (1853a). Nieuwe bijdrage tot de kennis der 16–30 (unnumbered pages: 1–10 publ. in 1828, ichthyologische fauna van Ceram. Natuurkundig 11–15 in 1829, 16–30 in 1830). Tijdschrift voor Nederlandsch Indië, 3 (5): 689–714. Bernardi, G., Longo, G. C. and Quiros, T. E. A. L. Bleeker, P. (1853b). Derde bijdrage tot de kennis der (2017). Altrichthys alelia, a new brooding ichthyologische fauna van Amboina. Natuurkundig damselfish (Teleostei, Perciformes, Pomacentridae) Tijdschrift voor Nederlandsch Indië, 4 (1): 91–130. from Busuanga Island, Philippines. ZooKeys, 675: Bleeker, P. (1853c). Bijdrage tot de kennis der 45–55. ichthyologische fauna van Ternate. Natuurkundig https://doi.org/10.3897/zookeys.675.12061 Tijdschrift voor Nederlandsch Indië, 4 (1): 131– Betancur–R, R., Wiley, E.O., Arratia, G., Acero, A., 140. Bailly, N., Miya, M., Lecointre, G. and Orti, G. Bleeker, P. (1853d). Diagnostische beschrijvingen (2017). Phylogenetic classification of bony fishes. van nieuwe of weinig bekende vischsoorten van BMC Evolutionary Biology, 17 (162): 1–40. Sumatra. Tiental V–X. Natuurkundig Tijdschrift https://doi.org/10.1186/s12862–017–0958–3 voor Nederlandsch Indië, 4 (2): 243–302. Bleeker, P. (1845). Bijdragen tot de geneeskundige Bleeker, P. (1853e). Diagnostische beschrijvingen Topographie van Batavia. Generisch overzicht van nieuwe of weinig bekende vischsoorten van der Fauna. Natuur- en Geneeskundig Archief voor Batavia. Tiental I–IV. Natuurkundig Tijdschrift Neȇrlandsch Indië, 2 (3): 505–528. voor Nederlandsch Indië, 4: 451–516. Bleeker, P. (1847a). Labroideorum ctenoideorum Bleeker, P. (1853f). Bijdrage tot de kennis der bataviensium diagnoses et adumbrationes. ichthyologische fauna van Solor. Natuurkundig [Overzigt der te Batavia voorkomende soorten Tijdschrift voor Nederlandsch Indië, 5 (1): 67–96. van Kamschubbige Labroieden, met beschrijving van 5 nieuwe soorten.]. Verhandelingen van het Bleeker, P. (1853g). Vierde bijdrage tot de kennis der Bataviaasch Genootschap van Kunsten en ichthyologische fauna van Amboina. Natuurkundig Wetenschappen, 21 (art. 3): 1–33. Tijdschrift voor Nederlandsch Indië, 5 (3): 317–352. Bleeker, P. (1847b). Pharyngognathorum Siluroideorumque Bleeker, P. (1854a). Derde bijdrage tot de kennis der species novae Javanenses. Natuur- en Geneeskundig ichthyologische fauna van de Banda-eilanden. Archief voor Neȇrlandsch Indië, 4 (2): 155–169. Natuurkundig Tijdschrift voor Nederlandsch Indië, 6 (1): 89–114. Bleeker, P. (1848). A contribution to the knowledge of the ichthyological fauna of Sumbawa. Journal Bleeker, P. (1854b). Bijdrage tot de kennis der of the Indian Archipelago and Eastern Asia ichthyologische fauna van de Kokos-eilanden. (Singapore), 2 (9): 632–639. Natuurkundig Tijdschrift voor Nederlandsch Indië, 7 (1): 37–48. Bleeker, P. (1849). Bijdrage tot de kennis der ichthyologische fauna van het eiland Bali, met Bleeker, P. (1854c). Nieuwe tientallen diagnostische beschrijving van eenige nieuwe species. beschrijvingen van nieuwe of weinig bekende Verhandelingen van het Bataviaasch Genootschap vischsoorten van Sumatra. Natuurkundig Tijdschrift van Kunsten en Wetenschappen, 22: 1–11. voor Nederlandsch Indië, 5 (3): 495–534. Bleeker, P. (1851). Bijdrage tot de kennis der Bleeker, P. (1855a). Bijdrage tot de kennis der ichthyologische fauna van de Banda–eilanden. ichthyologische fauna van Batjan. Natuurkundig Natuurkundig Tijdschrift voor Nederlandsch Tijdschrift voor Nederlandsch Indië, 7 (3): 359–378. Indië, 2 (2): 225–261. Bleeker, P. (1855b). Bijdrage tot de kennis der Bleeker, P. (1852a). Bijdrage tot de kennis der ichthyologische fauna van het eiland Groot-Obij. ichthyologische fauna van Singapore. Natuurkundig Natuurkundig Tijdschrift voor Nederlandsch Tijdschrift voor Nederlandsch Indië, 3 (1): 51–86. Indië, 9 (3): 431–438.

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831

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Whitley, G. P. (1954). New locality records for some Ocean. Ichthyological Bulletin J.L.B. Smith Australian fishes. Proceedings of the Royal Institute, (66): 1–28. Zoological Society of New South Wales, 1952–53: Winterbottom R., Emery A.R., Holm E. (1989). An 23–30. annotated checklist of the fishes of the Chagos Whitley, G. P. (1961). Fishes from New Caledonia. Archipelago, central Indian Ocean. Royal Ontario Proceedings of the Royal Zoological Society of Museum Life Science Contributions, 145: 1–226, New South Wales, 79th Annual Report for the Pls. 1–8. years 1958–59: 60–65. Wirtz, P., A. Brito, J. M. Falcón, R. Freitas, R. Whitley, G. P. (1964). Fishes from the Coral Sea and Fricke, V. Monteiro, F. Reiner and O. Tariche the Swain Reefs. Records of the Australian (2013). The coastal fishes of the Cape Verde Museum, 26 (5): 145–195. Islands – new records and an annotated check– Whitley, G. P. (1965). Some fish genera scrutinized. list. Spixiana (München), 36 (1): 113–142. Proceedings of the Royal Zoological Society of Yamakawa, T. and Randall, J. E. (1989). Chromis New South Wales, 1964–65: 25–26. okamurai, a new damselfish from the Okinawa Wibowo, K., Koeda, K., Muto, N. and Motomura, Trough, Japan. Japanese Journal of Ichthyology, H. (2018). Abudefduf nigrimargo, a new species 36 (3): 299–302. of damselfish (Perciformes: Pomacentridae) Yoshino, T. (1982). Damselfishes of the genus from Taiwan. Ichthyological Research, 65 (4): Dischistodus found in the Ryukyu Islands. 471–481. Bulletin of the College of Science / University of https://doi.org/10.1007/s10228–018–0634–7 the Ryukyus, 3333: 69–74. Wibowo, K., Toda, M. and Motomura, H. (2017). Zajonz, U., Lavergne, E., Bogorodsky, S. V., Saeed, Validity of Abudefduf caudobimaculatus Okada F. N., Aideed, M. S. and Krupp, F. (2019). and Ikeda 1939 and synonymies of Abudefduf Coastal fish diversity of the Socotra Archipelago, vaigiensis (Quoy and Gaimard 1825) (Perciformes: Yemen. Zootaxa, 4636 (1): 1–108. Pomacentridae). Ichthyological Research, 65 (1): https://doi.org/10.11646/zootaxa.4636.1.1 78–91. Zubak, I., Kruschel, C. and Schultz, S. T. (2017). https://doi.org/10.1007/s10228–017–0594–3 Predators structure fish communities in Posidonia Winterbottom, R. and Anderson, R.C. (1997). A oceanica meadows: meta–analysis of available revised checklist of the epipelagic and shore data across the Mediterranean basin. Marine fishes of the Chagos Archipelago, Central Indian Ecology Progress Series, 566: 145–157.

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Appendix Systématique et Évolution, Laboratoire d'Ichthyologie Générale et Appliquée, Paris, France; MNRJ, Museu The list of institutions where the type material is Nacional, Departamento de Vertebrados, Setor de located. Ictiología, Universidade Federal do Rio de Janeiro, AFAQ, Queensland Amateur Fishing Clubs Rio de Janeiro, Brazil; MZB, Bogor Zoological Association Inc., Australia; AIM, Auckland Museum Museum / Museum Zoologicum Bogoriense, Institute, Auckland, New Zealand; AMNH, American Cibinong, West Java, Indonesia; MZUSP, Museum of Natural History, New York, U.S.A; AMS, Universidade de São Paulo, Museu de Zoologia, São Australian Museum, Sydney, New South Wales, Paulo, Brazil; NCIP, Pusat Penelitian dan Australia; ANSP, Academy of Natural Sciences of Pengembangan Oseanologi, Indonesian Institute of Drexel University, Philadelphia, Pennsylvania, U.S.A; Sciences (LIPI), Jakarta, Java, Indonesia; NHRM, ASIZP (or ASIZT), Academia Sinica, Biodiversity Naturhistoriska Riksmuseet, Department of Vertebrate Research Center, Taipei, Taiwan; BMNH, Natural Zoology, Ichthyology Section, Stockholm, Sweden; History Museum, London, U.K; BPBM, Bishop NMMB, Pisces Collection, National Museum of Museum, Honolulu, Hawai'i, U.S.A; BSKU, Kochi Marine Biology and Aquarium, Pingtung, Taiwan; University, Department of Natural Science, Faculty of NMNZ, Museum of New Zealand Te Papa Science, Kochi, Japan; BSMP, Department of Tongarewa, Wellington; NMV, Museum Victoria, Agriculture, Bureau of Science, Manila, Philippines; Ichthyology, Melbourne, Victoria, Australia; NMW, CAS, California Academy of Sciences, San Francisco, Naturhistorisches Museum, Zoologische Abteilung, California, U.S.A; CMC, Canterbury Museum, Fische, Wien, Austria; NSMT, National Museum of Christchurch, New Zealand; CSIRO, Commonwealth Nature and Science, Zoology Department, Division of Scientific and Industrial Research Organisation, Fishes, Tsukuba, Japan; NTUM, National Taiwan National Collections and Marine Infrastructure, University, Institute of Zoology, Taipei, Taiwan; Hobart, Tasmania, Australia; FBGN-SAU, Aquatic NYZS, New York Zoological Society, New York, Bioresource Research Lab., Department of Fisheries U.S.A; NZOI, National Centre for Coasts and Oceans, Biology and Genetics, Sher-e-Bangla Agricultural National Institute of Water and Atmospheric Research University, Dhaka, Bangladesh; FMNH, Division of (NIWA) Taihoro Nukurangi, Wellington, New Fishes, Department of Zoology, Field Museum of Zealand; OMNH, Osaka Museum of Natural History, Natural History, Chicago, Illinois, U.S.A; FRLM, Laboratory of Zoology, Osaka, Japan; PMBC, Phuket Fisheries Research Laboratory, Mie University, Mie- Marine Biological Centre, Phuket, Thailand; PNM, ken, Japan; FSKU, Kitasato University, School of National Museum of the Philippines, Manila, Luzon, Fisheries Sciences, Iwate Pref., Japan; HMNH, Hiwa Philippines; PRFRI, Pearl River Fishery Research Museum for Natural History, Shobara, Japan; HUJ, Institute, Chinese Academy of Fishery Sciences, The Hebrew University, Faculty of Science, Guangzhou, Guangdong, China; QM, Queensland Department of Zoology, Zoological Museum; HUMZ, Museum, Centre for Biodiversity, Fishes, Brisbane, Hokkaido University, Laboratory of Marine Zoology, Queensland, Australia; RMNH, Rijksmuseum van Faculty of Fisheries, The Hokkaido University Natuurlijke Histoire, Leiden, Netherlands; ROM, Museum, Hakodate, Hokkaido, Japan; IFAN, Institut Royal Ontario Museum, Department of Natural Fondamental d'Afrique Noire, Dakar, Senegal; IOI, History, Toronto, Ontario, Canada; RUSI, Rhodes Institut Océanographique de l'Indo-Chine de Cauda, University, J.L.B. Smith Institute of Ichthyology; Nha Trang, Vietnam; KAUM, Kagoshima University SAIAB, South African Institute for Aquatic Museum, Ichthyology, Kagoshima, Japan; KFRS, Biodiversity, National Research Foundation, Kanudi Fisheries Research Station, National Fisheries Grahamstown, South Africa; SAM, South African Authority, Konedobu, Papua New Guinea; KPM, Museum, Cape Town, South Africa; SIO, Scripps Kanagawa Prefectural Museum of Natural History, Institution of Oceanography, Marine Vertebrate Odawara, Kanagawa, Japan; LACM, Natural History Collection, La Jolla, California, U.S.A; SMBL, Kyoto Museum of Los Angeles County, Los Angeles, University, Seto Marine Biology Laboratory, California, U.S.A; LIPI, Lembaga Biologi Nasional, Wakayama Prefecture, Japan; SMF, Senckenberg Bogar, Indonesia; MAMU, University of Sydney, Forschungsinstitut und Naturmuseum, Abteilung Macleay Museum, Sydney, New South Wales, Marine Zoologie, Sektion Ichthyologie, Frankfurt am Australia; MCZ (or MCZH), Museum of Main, Hessen, Germany; SMNS, Staatliches Museum Comparative Zoology, Harvard University, für Naturkunde in Stuttgart, Stuttgart, Baden- Ichthyology Department, Cambridge, Massachusetts, Württemberg, Germany; SSCN, Museum of the U.S.A; MGH, Museum Godeffroy, Hamburg, Biological Laboratory, Chinese Academy of Sciences, Germany; MHNG, Museum d'Histoire Naturelle, Nanjing, China; SU, Stanford University. The Département d'Herpétologie et Ichthyologie, Ville de collection now resides at CAS; TAU, The Steinhardt Genève, Genève, Switzerland; MHNLR, Museum Museum of Natural History, George S. Wise Faculty d'Histoire Naturelle, La Rochelle, France; MNH, of Life Sciences; TMBS, Tatsuo Tanaka Memorial Hungarian National Museum, Budapest, Hungary; Biological Laboratory, Ako, Miyake-jima, Izu Islands, MNHN, Muséum National d'Histoire naturelle, Japan; UGM, University of Guam, College of Natural

Journal of Animal Diversity (2021) | © Lorestan University Press An annotated checklist of damselfishes, Family Pomacentridae Bonaparte, 1831 and Applied Sciences, Western Pacific Tropical Washington, Seattle; WAM, Western Australian Research Center, Mangilao, Guam; UMB, Museum, Department of Aquatic Zoology, Überseemuseum, Bremen, Germany; UMML, Ichthyology, Perth, Western Australia, Australia; University of Miami, Dorothy H. and Lewis YCM, Yokosuka City Museum, Yokosuka City, Rosenstiel School of Marine and Atmospheric Kanagawa, Japan; ZIN, Laboratory of Ichthyology, Science, Division of Marine Biology and Fisheries, Zoological Institute, Russian Academy of Sciences, St. Miami, Florida, U.S.A; UMZC, University Museum Petersburg, Russia; ZMA, Universiteit van of Zoology, Cambridge, U.K; UNIMAS, University of Amsterdam, Faculty of Science, Zoölogisch Museum, Malaysia Sarawak, Kota Samarahan, Sarawak, Amsterdam, The Netherlands; ZMB, Museum für Malaysia; UPLB, University of the Philippines Los Naturkunde, Leibniz-Institut für Evolutions- und Baños, Animal Biology Division, Institute of Biodiversitätsforschung, Berlin, Germany; ZMH, Biological Sciences, Laguna, Philippines; UPMSI, Zoologisches Institut und Zoologisches Museum der Marine Science Institute, University of the Philippines, Universität Hamburg; ZMUC, Københavns Quezon City; URM, University of the Ryukyus, Universitet, Zoologisk Museum, Vertebrater, Faculty of Science, Department of Chemistry, Biology Fiskesamlingen, Copenhagen, Denmark; ZMUU, and Marine Science, Nishihara, Okinawa, Japan; Uppsala Universitet, Zoologiska Museet, Uppsala, USNM, Smithsonian Institution National Museum of Sweden; ZSI, Zoological Survey of India, Kolkata, Natural History, Department of Vertebrate Zoology, India; ZUMT, Department of Zoology, University Division of Fishes, Washington D.C., U.S.A; UW, Museum, University of Tokyo, Tokyo, Japan. School of Aquatic and Fishery Sciences, University of

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