(Orchidaceae): Coelogyne Lindl. Comprises Approximately (1907
BLUMEA 45 (2000) 275-318 Revision of Coelogyne section Verrucosae (Orchidaceae): a new sectional delimitation based on morphological and molecular evidence S.E.C. Sierra B. Gravendeel & E.F. de Vogel Nationaal Herbarium Nederland, Universiteit Leiden branch, P.O. Box 9514, 2300 RA Leiden, The Netherlands Summary Section Verrucosae Pfitzer & Kraenzl. ofthe Lindl. is revised genus Coelogyne using morphological and molecular data. Eight species are recognized, including two new ones ( C. marthae and C. ver- One is reduced Four included several authors in rucosa). name to synonymy. species formerly by sect. Verrucosae (C. brachyptera, C. papillosa, C. parishii and C. virescens) are excluded. A total data the evidence analysis ofmorphologicalcharacters and ITS and matK sequence supports mono- phyly of the section as here recognized. Coelogyne virescens (sect. Brachypterae) is identified as nearest neighbour to the species of sect. Verrucosae. The number of sterile bracts on the rhachis and the of the ornamentation the be informative shape on epichile appear to phylogenetically characters, in contrast with the inflorescence type, ovary indumentum and number ofkeels on the hypochile. Key words . Coelogyne sect. Verrucosae, matK, orchids, phylogeny, ribosomal ITS, systematics. Introduction The orchid genus Coelogyne Lindl. comprises approximately 200 species, distributed from Southeast Asia to the south-western Pacific Islands. Pfitzer & Kraenzlin (1907), in their revision of subtribe Coelogyninae, subdivided the genus into 14 different maintainedthis section. sections, among which is sect. Verrucosae. All later authors According to Pfitzer & Kraenzlin sect. Verrucosae consists of plants with large pseudobulbs and leaves, with very large flowers having an unusual combination of with black and with keels and of In colours (green markings), a lip a mass papillae.
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