European Arachnology 2005 (Deltshev, C. & Stoev, P., eds) Acta zoologica bulgarica, Suppl. No. 1: pp. 191-219.

Spiders of Gaderská and Blatnická Dolina valleys in the southern part of Veľká Fatra Mts., Slovakia (Araneae)

Jaroslav Svatoň1, Peter Gajdoš2

Abstract: During the years 1974-1976 and 1986-1990, a research of epigeic of the Gaderská and Blatnická Dolina valleys was carried out in 71 study sites. Altogether, 3295 spiders belonging to 293 and 28 families were captured in the non-forest, ecotones and forest habitats. Out of the identified species, 32 are listed in the Red List of Spiders of Slovakia and one species ( semiater) has not been previously reported from Slovakia. The occurrence of Meioneta innotabilis is faunistically very important because it was considered an extinct species in Slovakia. Zelotes puritanus has been reported from Slovakia only from this area. The family Linyphiidae was eudominant in species composition (101 species – D=34.5%). Specimens from the families Linyphiidae, Lycosidae and were the most abundant (D=23.4%, 17.4% and 12.3%). Eurytopic psychrophilous and mesophilous spiders represented the highest number of species. Species diversity in the research area is rather high, consisting of approximately 32.5% of the Slovak araneofauna. Four of the study forest habitats (relict origin oak, beech, fir-beech and pine forests) and two non-forest habitats (dry calcareous grassland and subalpine meadows) were evaluated as important habitats for diversity conservation and conservation of threatened spider species (territories with high diversity of spiders, with the highest proportion of endangered and rare species).

Key words: spiders, faunistics, nature conservation, Veľká Fatra National Park, North Slovakia

Introduction The Gaderská and Blatnická Dolina valleys form the southern part of the Veľka Fatra Mts., which are important orographical units in Slovakia. In the zoogeographical classification, the area of the Veľka Fatra Mts. represents the Western Carpathian part of the subprovince of the Carpathian Mountains and the Central European Mountains. There occur of steppe, forest-steppe, cultural steppe, agricultural land, synantropic species, and also species of wetland and marshy biotopes. The submontanous, montanous, boreoalpine, subalpine and alpine species predominate in the area with many rare species, which occur in Slovakia only here. For some species this is the northern border of their distribution in Europe. The Cosmopolitan, Holarctic, Palearctic, Euro-Siberian, Siberian, European, Caucasian, Pontic, Submediterian, Boreo-alpine and Boreal elements of our fauna are manifested here. From the point of view of arachnological research, the territory of the Veľká Fatra Mts., as well as the Gaderská and Blatnická Dolina valleys had received only little attention until 2000. Only a few papers (Svatoň 1981, 1983, Franc 2002) describe the spider fauna of the region. The territory was included in the Natura 2000 network in 2003 and this fact initiated an evaluation of the unpublished araneological material collected from 71 localities in this area in the periods 1974-1976 and 1986-1990. The aims of this study are to present the results of this inventory and to propose a classification of the studied habitats according to their importance from a point of view of the spider fauna conservation.

1 Kernova ulica 8/37, SK-03601 Martin, Slovakia. Е-mail: [email protected] 2 Institute of Landscape Ecology, Nitra Branch, Slovak Academy of Sciences, Akademická 2, SK-94901 Nitra, Slovakia. Е-mail: [email protected]

191 European Arachnology 2005

Material and Methods

Study area The study area comprises the upper part of the Blatnica and Gader Creek catchments (the area of Gaderská and Blatnická Dolina valleys). These valleys lie in the south western part of Veľká Fatra Mts. In accordance with the orographic division of Slovakia (Hromádka 1956) Veľká Fatra Mts. belongs to a zone of the inner Western Carpathians with crystalline core and Mesozoic cover. A predominant part of its geological base consists of limestone and dolomite rocks. This geological substratum allows the occurrence of high number of species which are bound to rocky limestone and dolomitic biotopes with numerous caves, semi-caves, rock hollows and clefts that create suit- able conditions for troglophilous and cavernicolous species. From a biological point of view, the most valuable habitats are inaccessible rock walls, steep slopes and sharp ridges undisturbed by human activities. Natural vertical zoning of the vegetation is here quite often disturbed by follow- ing geomorphological factors: inclination, orientation, relief, wideness and depth of both valleys. This causes species inversion, which means occurrence of cryophilic plant and species at the bottoms of valleys below the zone of beech and oak forest and occurrence of thermophilous species in the top zone of the hills which surround the Gaderská and Blatnická Dolina valleys. Due to different climatic, morphological, geological and vegetation conditions in addition to forestry and farming activities in Veľká Fatra Mts, as well as in the adjacent Turčianska Kot- lina basin, the valleys support a high plant and animal diversity. Altitudinal zoning ranges from 385 m to 1500 m above the sea level, segmentation of terrain, different terrain expositions and different substratum create a vivid mosaic of contrasting sites, biocoenosis with various animal communities bound to these habitat types.

Sampling The spider fauna was studied by different methods such as pitfall trapping, sifting detritus and dead leaves, sweeping herbal layer, beating the trees and shrubs, as well as individual collection under stones, in grass and among leaves. Spiders were fixed in 80% ethanol.

Study sites and habitats Sampling was carried out in 71 study sites of different types of habitats (Appendix 1). The sites were chosen as the most representative habitat types of this region. They represent non-forest habitats (debris and rock formations, caves, dry calcareous meadows, hygrophilous and forest meadows, fen meadows, peat-bogs, subalpine meadows and shrubby habitats) and forest habitats such as alder (Alnetum incanae) forests growing on gravel deposits near creeks, pioneer birch habitat, original azonal oak forests (Quercetum), virgin and secondary beech forests (Fagetum typicum), original fir-beech forests (Abieto-Fagetum), secondary mixed beech-spruce forests (Fageto-Piceetum), original and secondary pine forests (Pineetum), original and secondary spruce forests (Piceetum) and original dwarf pine habitat (Pinion mughi).

Analysis The evaluation of the main studied habitats in the Gaderská and Blatnická Dolina valleys was done on the basis of spider species richness and diversity, mainly by the presence of threatened and rare spider species. For analysis of species data the program CANOCO (TerBraak, Šmilauer 2002) was used. Richness of samples (N1) related to Shannon entropy statistics (H) using the relation 1 H N = e , Shannon’s diversity (H), the maximum achievable value of the diversity (Hmax) (Hmax = log(S), S = number of species) in samples and evenness (J) (J = H/log(S)) were calculated (Table 3).

192 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina

Table 1. List of families: number of captured species and specimens.

Family Species Specimens Number D (%) Number D (%) Atypidae 2 0.68 34 1.03 Pholcidae 1 0.34 1 0.03 Segestriidae 1 0.34 27 0.82 3 1.02 17 0.52 Mimetidae 1 0.34 6 0.18 Theridiidae 19 6.48 53 1.61 Linyphiidae 101 34.47 772 23.43 Tetragnathidae 9 3.07 141 4.28 Araneidae 19 6.48 125 3.79 Lycosidae 30 10.24 572 17.36 Pisauridae 1 0.34 1 0.03 Agelenidae 9 3.07 69 2.09 Cybaeidae 1 0.34 404 12.26 Hahniidae 5 1.71 15 0.46 3 1.02 17 0.52 Amaurobiidae 6 2.05 342 10.38 1 0.34 17 0.52 Anyphaenidae 1 0.34 9 0.27 Liocranidae 3 1.02 61 1.85 Clubionidae 9 3.07 22 0.67 Corinnidae 1 0.34 19 0.58 Gnaphosidae 13 4.44 180 5.46 Zoridae 5 1.71 20 0.61 Heteropodidae 1 0.34 35 1.06 Zodariidae 1 0.34 42 1.27 Philodromidae 8 2.73 52 1.58 Thomisidae 20 6.83 110 3.34 Salticidae 19 6.48 132 4.01 Total 293 100 3295 100

Table 2. Thermo-preference of the spider fauna in the study area: S – number of species, Sp – number of specimens, D – dominance (after Buchar 1983, 1992).

Thermo-preference S D (%) Sp D (%) Thermophilous (T) 47 16.04 419 12.72 Psychrophilous (P) 78 26.62 1405 42.64 Mesophilous (M) 78 26.62 374 11.35 Eurytopic (Non-specific) (N) 87 29.69 1081 32.81 Not determined (?) 3 1.02 16 0.49 Total 293 100 3295 100

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– max 0.54 0.85 0.79 0.84 0.79 0.87 0.75 0.93 0.82 0.62 0.89 0.81 J 3.26 4.25 4.23 3.89 2.08 4.34 2.89 3.56 2.20 3.58 3.69 2.56 max H 1.76 3.62 3.33 3.28 1.63 3.80 2.17 3.31 1.80 2.24 3.28 2.09 (H) 1 5.81 5.13 8.79 6.05 9.37 8.09 37.16 28.06 26.61 44.91 27.25 26.47 N 1 7 5 6 1 8 1 1 0 2 0 19 (1) 3 (1) Σ 1 DD 1 1 2 3 3 5 2 LR 2 8 3 1 1 1 1 – richness of species in samples, (H) – Shannon’s diversity of samples, H samples, of diversity Shannon’s – (H) samples, in species of richness – 1 1 VU 4 1 2 1 EN 4 Categories of threat 1 1 CR 1 1 EX 1 8 9 92 26 70 69 49 77 18 35 36 40 13 S 190 38 57 96 30 51 531 161 307 327 190 256 224 130 Sp 1336 ) ) Pm ) .) ) ) ) AbF ) ) ) ) ) Pic Pic Mix Fag Fag Aln Bet Pin Pin Qu Evaluation of the forest, ecotone and non-forest spider faunas in the study area. Sp – number of specimens, S – number of species, categories of threat: see Evaluation of the forest, ecotone and non-forest spider faunas in the study area. Sp – number of specimens, S – number of species, categories of threat: see Habitat (Or.f.) Origin forests Alder forest ( Oak forest ( Beech forest ( forest ( Fir-beech Spruce forest ( Pine forest ( Dwarf pine forest ( (Se.f.) Secondary forests Birch forest ( Beech forest ( Spruce forest ( Pine forest ( Mixed forest ( Table 3. Table N species, threatened of number total – Σ Slovakia, for species new – (1) 2, Appendix H maximum in samples, J – evenness of samples.

194 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina 1 0.9 0.98 0.92 0.92 0.52 0.97 0.87 0.89 0.83 0.83 J 3.0 3.4 2.48 1.79 4.39 3.26 2.56 4.16 4.51 4.14 3.53 max H 3.0 2.43 1.79 4.03 1.34 2.89 3.61 3.98 3.43 2.84 3.16 (H) 6 1 3.81 11.31 56.07 20.13 18.02 37.15 54.44 30.87 17.03 23.68 N 3 0 0 3 3 2 1 7 3 0 4 22 10 Σ DD 2 1 1 1 LR 9 2 1 1 2 3 2 2 2 2 VU 6 1 1 1 3 1 0 EN 4 3 2 1 1 1 Categories of threat CR 1 1 EX 6 95 12 81 26 13 20 64 91 68 30 34 192 S 6 16 50 88 31 71 273 242 189 338 283 103 1140 Sp S- ) Cv ) ) ) ) Deb Dry Mes Habitat Ecotones O.pine forest S.beech forest S.spruce forest habitats Non-forest Debris and rock-faces ( Caves ( Shrubby habitat (Shr) Dry calcareous grassland ( Subalpine meadows ( alp Forest meadows and mesophilous meadows ( Fen meadows (Fen) Peat bog Table 3. Continued. Table

195 European Arachnology 2005

Many habitats were not evaluated because of the low number of captured specimens and species. For evaluation of the thermo preference of captured species Buchar‘s classification B( uchar 1983, 1992) was used. The nomenclature and systematic order of spiders follow Platnick (2005).

Results

Species composition Altogether 3295 spiders belonging to 293 species were captured in non-forest and forest habitats. The species number in the Gaderská and Blatnická valleys is rather high, representing approxi- mately 32.5 % of the Slovak araneofauna. Of the identified species, 32 species are included in The Red List of Spiders of Slovakia (Gajdoš et al. 1999, Gajdoš, Svatoň 2001) and one species (Centromerus semiater) has not been previously reported from Slovakia. Also the occurrence of the species Meioneta innotabilis was documented, which is important from a faunistic point of view, because this species was considered extinct in Slovakia. Zelotes puritanus has previously been reported from Slovakia only from this area (Gajdoš, Svatoň 1993, Gajdoš et al. 1999, Franc 2002). The composition of the spider fauna in the habitats depends on the habitat types (Appendix 2).

Family composition Twenty eight families were recorded in non-forest and forest ecosystems of the Gaderská and Blatnická Dolina valleys. The families Linyphiidae and Lycosidae were eudominant in the spe- cies composition (101 species – D-34.5%, 30 species – D-10.2%). Specimens from the families Linyphiidae, Lycosidae, Cybaeidae and Amaurobiidae were the most abundant (D - 23.4%, 17.4% 12.3% and 10.4 %) (Table 1).

Thermo-preference The psychrophilous component was represented by 78 species (26.62%) and 1405 specimens (42.64%), while the mesophilous component was represented by 78 species (26.62%). The ther- mophilic fauna was presented in the study area by 47 species (16.04%), occuring mainly in the extremely dry calcareous meadows and azonal oak forests with southern exposition. The largest component of the captured species (87 species) belongs to eurytopic species (29.69%). The re- maining few species can not be categorized (1.02%) (Table 2).

Evaluation of the spider fauna of forest habitats and ecotones In the forest habitats, classified in 7 typological groups Alnetum( incanae, pioneer habitat with Betula pubescens, Quercetum, Fagetum typicum, Abieto-Fagetum, Fageto-Piceetum and Pinion mughi), the species diversity was high (216 species or 73.72% of all captured species, Appendix 2). In these ecosystems we also recorded many species that are significant and important from a faunistic-ecological, as well as from a zoogeographical point of view. The most important captures are: Dasumia carpatica, merens, morulus, Meioneta innotabilis, zimmermanni, and hackmani. In the ecotones the most important spe- cies are: Pardosa sordidata, praeceps, Sitticus zimmermanni, and variegata (Appendix 2). On the basis of the criteria chosen for the habitat evaluation, four of the habitats (relict pine forest (Pinetum dealpinum), relict azonal oak forest (Quercetum), original fir-beech forest, and original beech forest) were considered as important habitats for the spider diversity and also for conservation of threatened species. Of these habitats the relict pine forest has the highest species

196 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina

richness (N1 - 44. 91) and the highest number of threatened species (8 species). Of the ecotone habitats the margin of the secondary spruce forests presents high species richness (N1 - 56.07) with occurrence of 3 threatened species (Table 3).

Evaluation of the spider fauna of non-forest habitat From the non-forest habitats 192 spider species (65.53%) were recorded (Appendix 2). Several spider species bound to the non-forest ecosystems of the Gaderská and Blatnická Dolina valleys, are very important from faunistic-ecological and zoogeographical points of view. Such species are: sarcinatus, Xysticus luctuosus, biscissa, Tapinocyba pallens, etc. Centromerus capucinus, Centromerus serratus, crucifer, improbulus, Peponocranium praeceps and Zelotes puritanus were very rare in the collected material. On the basis of the criteria which have been chosen for the habitat evaluation, two of the studied non- forest habitats (dry calcareous grasslands and sub-alpine meadows) were evaluated as important habitats for conservation of both spider diversity and threatened species. From non-forest habitats the sub-alpine meadows present the highest species richness (N1 - 54.44) and also manifest pres- ence of threatened species (10 species) (Table 3). Discussion The recording of 293 spider species from a restricted area indicates the richeness of the habitats in the studied area. This makes approximately 32.5 % of the Slovak araneofauna, in spite of the fact that still relatively small part of Slovakia has been studied. The compositions of the spider fauna in the studied habitats were quite different, depending mainly on vegetation structure and various environmental factors, as well as on recent utilisation of the landscape. From the investigated non-forest habitats the spider communities of the open dry calcareous grasslands and sub-alpine meadows are especially interesting from nature conservation point of view (areas with high spider species diversity, with the highest proportion of threatened species). The cover of these habitat types is continually decreasing as a result of natural succession, and there is a urgent need for preparation of management plans for their protection.

Acknowledgements: The study was supported by a VEGA (GP 2/5063/25) grant. Received: 25.01.2006 Accepted: 20.05.2006 References

Buchar J. 1983. Die Klassifikation der Spinnenarten Böhmens als ein Hilfsmittel für die Bioindikation der Umwelt. - Fauna Bohemiae septentrionalis, 8: 119-135. Buchar J. 1991. The use of faunistical data for biomonitoring. - Bulletin de la Société neuchâteloise des Sciences Naturelles, 116: 49-57. Franc V. 2002. Contribution to the knowledge on spiders (Araneae) of the Veľká Fatra Mts. - Matthias Belivs University Proceedings, Supplement, 2 (1): 155-163. Gajdoš P., J. Svatoň 1993. The red list of spiders of Slovakia. - Proceedings of the XIV European Colloquium of Arachnology, Catania. Bollettino dell’Accademia Gioenia di Scienze Naturali, 26 (345): 115-133. Gajdoš P., J. Svatoň 2001. Červený (ekosozologický) zoznam pavúkov (Araneae) Slovenska. [Red (Eco- sozological) List of spiders (Araneae) of Slovakia]. - In: Baláž D., K. Marhold, P. Urban (eds.): Červený zoznam rastlín a živočíchov Slovenska. [Red List of plants and animals of Slovakia Nature Conservation]. - Ochrana Prírody, Banská Bystrica, Supplement, 20: 80-86. Gajdoš P., J. Svatoň, K. Sloboda 1999. Katalóg pavúkov Slovenska. [Catalogue of Slovakian spiders]. Ústav krajinnej ekológie SAV, Bratislava, 1, 337 p.; Mapy [Maps], 2, 315 p. Hromádka J. 1956. Orografické třídění Československé republiky. - Sborník československe společnosti zeměpisné, Praha, 61 (3): 161-180, 265-299.

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Platnick N. I. 2005. The World Spider Catalog. Version 6.5. American Museum of Natural History, Wash- ington, http.//research.amnh.org/entomology/spiders/catalog/INTRO1.html. Svatoň J. 1981. Einige neue oder unvollkommen bekannte Spinnenarten aus der Slowakei. - Biológia (Bra- tislava), 36 (2): 167-177. Svatoň J. 1983. Weitere neue oder unvollkommen bekannte Spinnenarten aus der Slowakei. - Biológia (Bratislava), 38 (6): 569-580. Ter Braak C. J. F., P. Šmilauer 2002. CANOCO Reference manual und CanoDraw for Windows User΄s guide: Software for Canonical Community Ordination (version 4.5). Microcomputer Power, Ithaca, NY, USA, 500 p.

Паяци (Araneae) от долините Гадерска и Блатницка в южната част на планината Велка Фатра (Словакия)

Я. Сватон, П. Гайдош

(Резюме) В периода 1974-1976 и 1986-1990 г. е проведено изследване на епигейните паяци в 71 стации в долините Гадерска и Блатницка, намиращи се в южната част на планината Велка Фатра (Словакия). Уловени са 3295 екземпляра от 293 вида и 28 семейства, установени в екотонни или изцяло горски хабитати. Тридесет и два вида присъстват в Червения списък на паяците на Словакия, а един вид – Centromerus semiater – е нов за фауната на Словакия. Видовото разнообразие в долините Гадерска и Блатницка е високо и представлява 32.5% от словашката аранеофауна. Важни фаунистични приноси са намирането на Meioneta innotabilis – вид, смятан за изчезнал, и Zelotes puritanus, който е намиран само в този район на страната. Семейство Linyphiidae е еудоминантно (101 вида – D=34.5%), а най-много екземпляри са уловени от семействата Linyphiidae, Lycosidae и Cybaeidae (съответно D=23.4%, 17.4% и 12.3%). Евритопните, влаголюбивите и мезофилните паяци са представени с най-много видове. Четири от изследваните горски хабитати – реликтни дъбови, букови и борови гори, както и сухите пасища и субалпийските ливади, са важни за опазването на високото видово разнообразие и на големият брой застрашени видове.

198 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina

Appendix 1. List of the studied sites with habitat types.

Locality Habitat Altitude Original forests Blatnická dolina alder forest (Alnetum) 542 Dedošova dolina beech forest (Fagetum typicum) 650 Gaderská dolina alder forest (Alnetum), gravel sediment 512 Horné piesky flowery beech forest Fagetum( dealpinum) 850 Horné piesky relict pine forest (Pinetum dealpinum) 987 Chládkove úplazy relict pine forest (Pinetum dealpinum) 1228 Kamenný úplaz relict pine forest (Pinetum dealpinum) 725 Mohošov flowery beech forest Fagetum( dealpinum) 750 Mohošov grúň flowery beech forest Fagetum( dealpinum) 1136 Nad Ďurďášovou relict pine forest (Pinetum dealpinum) 997 Nad Mažarnou relict oak forest with lime trees 890 Ostrá relict pine forest (Pinetum dealpinum) 1100 Padva virgin beech-fir forest Fageto-Abietum( ) 950 Pod Mažarnou virgin debris beech forest 780 Smrekov origin dwarf pine habitat 1441 Široká beech forest (Fagetum typicum) 950 Veterné beech forest (Fagetum typicum) 900 Zadná hubná beech forest (Fagetum typicum) 1003 Secondary forests Dedošova dolina secondary spruce forest 650 Gaderská dolina secondary spruce forest 574 Ľubená secondary spruce forest 1024 Nad Mažarnou mixed beech and spruce forest 950 Plešovica secondary pine forest 685 Pod Rovnou pioneer birch growth (15-20 year old) after fire 870 Pod vôdkami secondary spruce forest 650 Predná hubná mixed spruce and pine forest 800 Príkra secondary spruce forest 795 Široká secondary spruce forest 902 Škap secondary spruce forest 770 Žihľavná secondary pine forest 752 Ecotone habitats Dedošova dolina margin of secondary spruce forest 567 Dolné piesky margin of beech forest (Fagetum typicum) 594 Gaderská dolina margin of secondary spruce forest 574 Horné piesky margin of relict pine forest (Pinetum dealpinum) 1005 Ostrá margin of relict pine forest (Pinetum dealpinum) 1103 Rovná dolina margin of beech forest (Fagetum typicum) 595

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Appendix 1. Continued.

Locality Habitat Altitude Shrubby habitat Mohošov dry shrubby meadow 1050 Meadow habitats Horné piesky dry calcareous meadow 1003 Horné piesky xerotherm forest meadow 952 Horný Jasienok xerotherm forest meadow 948 Kamenný úplaz dry calcareous meadow 1203 Kozia skala dry calcareous meadow 1162 Kozia skala xerotherm forest meadow under summit 1121 Mohošov grúň dry forest meadow 1136 Mohošovec dry calcareous meadow 1050

Nad Ďurďášovou dry calcareous meadows on the foot of rock formations 700 Pod Rovnou dry shrubby meadow 879 Široká dry calcareous meadows 950 Veľká Skalná dry calcareous meadow 1297 Vrátna dry forest meadow 785 Blatnica: amfiteater forest meadow along creek 500 Blatnická dolina forest meadow along creek 550 Dedošová, under summit complex of forest meadows after fire 670 Dolné piesky forest meadow 575 Kráĺov grúň subalpine meadow 1360 Kráĺova studňa subalpine meadow 1574 Kráľova studňa mountain meadow 1574 Krížna subalpine meadow 1574 Magura mesophilous forest meadow 1142 Mohošov peat bog 730 Ostrá, summit subalpine meadow 1264 Pod Žihlavnou mesophilous meadow 550 Selenec mesophilous and wet forest meadows 675 Tlstá, summit subalpine meadow 1370 Caves, debris and rock formations Gaderská dolina rock formations 574 Konský dol rock formations 620 Mažarná cave 850 Nad vôdkami rock formations 1200 Smrekov debris in subalpine zone 1441 Široká debris 951

200 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina

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201 European Arachnology 2005 3 3 1 6 2 3 3 3 1 2 8 1 1 1 3 7 Σ 1 1 1 S- alp

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Se.

Ecoton Fag 1 Pin

Or. Fag Ecoton 1 Mix 1 2 1 2 2 Pin 1 1 Pic.

Fag Secondary forests 6 Bet

1 Pm

Pin 1 4 2 1 AbF 2 1 1 5 10 Pic Origin forests 6 Qu

3 Fag

2 1 2 Aln. ,

L .

L .

( id e r (O.P.- ( (O.P.- (O.P.-

W (

, 1875) tripartitus , 1978)

on , 1841) t , 1833) i, 1894 , 1873) , 1875) , 1871) a , 1851) , 1851) v all am b ridg e , S , 1884) , 1884) t, 1962 C , 1881) , 1879) FAMILY / Species FAMILY Continued. est ring est ring lackwall ill e r und ev imon imon och och ulczyń sk am b ridg e am b ridg e am b ridg e ock e W S S S B W K C C K C nigrinus ( alticeps ( arcanus Centromerus capucinus Centromerus ( incilium Centromerus pabulator Centromerus (O.P.- sellarius Centromerus ( semiater Centromerus serratus Centromerus ( brevipes ( Ceratinella brevis 1834) Ceratinella major Ceratinella scabrosa brevisetosum M K L P P P P P P T N N N N N M M M M TP

Red list LR (lc) EN new VU Appendix 2.

203 European Arachnology 2005 7 6 6 3 1 2 2 7 1 6 4 3 2 Σ 71 10 10 1 4 1 1 S- 34 alp

Fen Meadows 1 2 1 Mes 1 Dry

Peat bog 3 Cv

Deb Rocky

Shr

Pin 2 1 4 1 2 4 7 Pic

Se.

Ecoton Fag

Pin

Or. Fag Ecoton 7 1 Mix

Pin 1 21 Pic.

Fag Secondary forests

Bet

Pm

8 1 Pin 1 1 1 AbF 6 4 6 1 Pic Origin forests 1 Qu

1 2 1 1 1 Fag

2 1 2 Aln. , , , ,

id e r id e r

L .

(O.P.- ( (L. W W ( (

l e rck C , 1863) ( lackwall B

, 1834) , 1833) , 1841) , 1841) , 1841) , 1841) , 1879) l, 1833) al am b ridg e , 1834) C , 1869) FAMILY / Species FAMILY Continued. id e r lackwall lackwall lackwall lackwall lackwall lackwall und ev och am b ridg e B B B B S W B B K ( cristatus ( Diplocephalus latifrons (O.P.- Diplocephalus picinus ( concolor 1834) bifrons ( socialis ( acuminata ( Entelecara congenera atra 1833 Erigone dentipalpis 1834) globipes Koch, 1872) Evansia merens 1757) paradoxum Gonatium rubellum ( insignis ( C P P P P P T N N N N N N N M M M TP

Red list LR (nt) VU Appendix 2.

204 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina 8 1 1 4 2 2 4 7 1 1 1 3 1 1 Σ 77 30 4 1 1 S- alp 4 Fen Meadows 1 1 44 Mes 1 2 1 11 Dry 2 Peat bog 4 Cv 2 1

Deb Rocky

Shr

Pin 1 7 Pic

Se.

Ecoton Fag

Pin

Or. Fag Ecoton

Mix 2 1 1 Pin 2 Pic.

Fag Secondary forests 2 Bet

1 Pm

Pin 1 1 4 1 12 AbF 4 1 1 Pic Origin forests 1 Qu

1 1 2 1 1 15 Fag

Aln. , ,

L . ,

id e r

W (O.P.- , 1894 (O.P.- id e r L . (C. est ring ( ( W W ( (

imon S

, 1882) , 1898) , 1833) , 1873) , 1863) , 1875) , 1757) , 1865) , 1866) t , 1929) , 1881) , 1836) FAMILY / Species FAMILY Continued. e ng ulczyń sk i hl e r ulczyń sk i lackwall l e rck hor e ll imon och och am b ridg e am b ridg e S M K O B C K K (T C improbulus Improphantes ( Incestophantes crucifer ( dorsalis 1834) Kaestneria torrentum ( leprosus ( ( triangularis ( rufus 1834) mansuetus gallicus Maso sundevalli 1851) morulus Mecynargus Meioneta affinis ( Meioneta equestris Meioneta fuscipalpa Meioneta innotabilis K C P P P T N N N M M M M M M M M M TP

Red list VU VU DD EN EX Appendix 2.

205 European Arachnology 2005 3 1 1 2 1 3 3 1 Σ 22 16 15 10 14 10 29 15 2 6 1 S- 11 11 alp 1 2 Fen Meadows 2 5 8 1 Mes 2 2 Dry 1 1 15 Peat bog 1 Cv 1 2

Deb Rocky 2 1 Shr

Pin 1 2 3 8 1 1 10 Pic

Se. 1 1 Ecoton Fag

Pin 1 Or. Fag Ecoton

1 1 Mix 1 Pin 1 Pic.

Fag Secondary forests 1 Bet

2 2 Pm

3 Pin 2 1 1 2 AbF 1 2 4 Pic Origin forests 5 1 Qu 4 1 1 1 Fag

1 Aln. , ,

,

id e r id e r

L .

W W

( id e r ( (C.

W

, 1872)

( 1842) , , 1842) , 1881) , 1854) , 1841) , 1841) , 1853) , 1830) , 1851) 1976 , , 1875) all am b ridg e t C , 1836) FAMILY / Species FAMILY Continued. est ring alck e na r alck e na r ulczyń sk i lackwall lackwall lackwall lackwall illidg e und ev ick e r och B W S B F K W W B B Meioneta rurestris georgescuae herbigradus Micrargus ( subaequalis Micrargus ( pusilla ( viaria ( marginella 1834) pusillus 1834) mughi ( Mughiphantes pulcher ( emphana ( 1834) Neriene radiata ( Notioscopus sarcinatus (O.P.- obscurus ( agrestis ( M K P P P P P N N N N N M M M M M M TP

Red list LR (nt) Appendix 2.

206 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina 1 1 1 5 4 1 2 2 4 3 2 1 4 8 Σ 14 10

S- 10 alp 1 4 3 1 Fen Meadows 1 Mes 2 1 Dry 3 Peat bog 1 1 Cv

Deb Rocky 2 Shr

Pin 1 4 2 Pic

Se.

Ecoton Fag

Pin

Or. Fag Ecoton

Mix 1 Pin 2 Pic.

Fag Secondary forests

Bet

Pm

Pin 1 1 1 AbF 5 1 2 Pic Origin forests 1 5 Qu 2 1 1

Fag

1 1 Aln.

,

(L.

ill e r

, 1881) M

, 1871) (O.P.- imon S , 1882) ( , 1850) , 1834) , 1841) , 1841) , 1841) , 1834) , 1939 1851) , 1871) , am b ridg e , 1943 , 1884 C FAMILY / Species FAMILY och v il t Continued. est ring ulczyń sk i lackwall lackwall lackwall lackwall lackwall lackwall ill e r ra am b ridg e imon B B K B W B B B C Oedothorax apicatus ( Oedothorax fuscus ( Oedothorax gibbifer ( Oedothorax gibbosus ( Oedothorax retusus ( alutacius Palliduphantes pallidus (O.P.- Peponocranium praeceps radicicola Koch, 1872) phrygianus (C. L. Koch, 1836) pumila ( Poeciloneta variegata ( pygmaeum ( diceros Saloca кulczynskii rayi K S M ? P P P P P P P P P P N N N M M TP

EN Red list LR (lc) LR (lc) LR (nt) Appendix 2.

207 European Arachnology 2005 1 1 2 6 7 8 1 2 3 1 1 3 Σ 17 27 14 67 1 1 1 3 4 1 1 S- alp

Fen Meadows 1 1 1 Mes 1 1 3 Dry 1 1 Peat bog 1 Cv 1

Deb Rocky

Shr

Pin 7 9 Pic

Se. 1 Ecoton Fag

Pin 2 Or. Fag Ecoton 1 Mix 4 Pin 4 2 9 Pic.

Fag Secondary forests 1 2 Bet

Pm

12 Pin 1 1 AbF 1 4 2 2 1 13 Pic Origin forests 2 3 16 Qu

5 1 1 1 3 19 Fag

4 3 Aln.

,

,

t

C . ( id e r

W esse r (O.P.- (O.P.- ( L

, 1878) 1887) , , 1856) , 1853) , 1854) , 1872) , 1872) , 1851) , 1890) , 1956 , 1836) , 1866) am b ridg e , 1834) , 1834) C FAMILY / Species FAMILY t kau och Continued. id e r est ring id e r e ng ulczyń sk i lackwall lackwall lackwall e r illidg e am b ridg e am b ridg e B B M B K W B W W antic Walckenaeria corniger ( affinis Tapinocyba 1907 biscissa Tapinocyba pallens Tapinocyba alacris Tenuiphantes ( cristatus Tenuiphantes ( flavipes Tenuiphantes ( mengei Tenuiphantes ( tenebricola Tenuiphantes ( zimmermanni Tenuiphantes ( parasiticus ( cristatus ( hackmani Trichoncus 1834) atrotibialis Walckenaeria (O.P.- cucullata Walckenaeria C L . K C M P P P P P P P P T N N N N M M M TP

Red list DD DD LR (nt) VU Appendix 2.

208 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina 9 4 2 3 2 1 8 1 2 Σ 54 10 20 26 19 14 2 3 1 1 1 S- alp

9 2 5 1 Fen Meadows

2 1 2 Mes 2 2 Dry

1 2 Peat bog

54 18 Cv 1

Deb Rocky

Shr

Pin 2 6 1 8 3 8 3 1 Pic

Se.

Ecoton Fag

Pin

Or. Fag Ecoton

Mix 1

Pin

2 Pic.

2 Fag Secondary forests

Bet

3 Pm

Pin 2 1 1 1 AbF

1 1 1 11 Pic Origin forests 3 1

Qu

1

Fag

1

Aln. ,

L .

t r e ill

a L

( , 1802) , 1802) , 1870) , 1836 , 1823 , 1830 , 1830 , 1851) , 1758) , 1763) 1869) , 1757) , , 1868) all all all , 1870 FAMILY / Species FAMILY Continued. est ring alck e na r alck e na r e ng e ng lackwall l e rck inna e u s copoli och lackwall und ev und ev und ev M M W B S C L W W S B K furcillata Walckenaeria ( mitrata Walckenaeria ( nudipalpis Walckenaeria ( obtusa Walckenaeria (9) TETRAGNATHIDAE Meta menardi 1804) Metellina mengei ( Metellina merianae ( Metellina segmentata ( Pachygnatha clercki Pachygnatha degeeri Pachygnatha listeri extensa Tetragnatha ( pinicola Tetragnatha ARANEIDAE (19) Aculepeira ceropegia ( alsine ( S S P P P P P P P N N N N M M M M TP

Red list

Appendix 2.

209 European Arachnology 2005 1 9 2 5 4 1 1 2 2 4 1 4 1 3 Σ 11 55 2 2 S- alp 9 2 1 2 Fen Meadows 1 1 20 Mes 5 2 1 1 1 Dry 3 2 2 1 Peat bog

Cv 1 1

Deb Rocky

Shr

Pin 4 1 1 1 5 1 1 1 2 Pic

Se. 1 Ecoton Fag

Pin

Or. Fag Ecoton

Mix

Pin 2 2 1 1 1 Pic.

Fag Secondary forests 1 1 Bet

Pm

Pin 1 1 1 AbF

Pic Origin forests 8 Qu 1 2 2 1 1 Fag

1 Aln. , , , ,

C . , ( , och

l e rck l e rck

C C

ahn

alla s L . K P H ( alck e na r ( ( W , 1802) , 1905) ( , 1851) , 1803) , 1844) , 1757) , 1757) , 1757 , 1757 , 1874) FAMILY / Species FAMILY och Continued. est ring alck e na r ulczyń sk i l e rck l e rck imon chrank l e rck l e rck C S K W S W C Araneus angulatus 1757 Araneus diadematus Araneus marmoreus Araneus quadratus 1757 Araneus sturmi 1831) Araniella alpica 1869) Araniella cucurbitina ( Araniella inconspicua ( Araniella opisthographa ( prominens Cercidia ( Cyclosa conica 1772) Hypsosinga sanguinea L . K Larinioides folium ( Mangora acalypha ( Nuctenea umbratica ( Zilla diodia 1802) C C P P P T T N N N N N N M M M M M TP

Red list

Appendix 2.

210 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina 3 2 5 2 6 6 5 Σ 11 40 20 45 23 32 58 21 82 5 5 2 4 4 S- 18 14 alp 2 1 Fen Meadows 2 2 1 1 17 10 28 12 Mes 7 2 1 28 Dry 5 Peat bog

Cv

Deb Rocky 1 1 1 Shr

Pin 3 2 7 11 10 Pic

Se.

Ecoton Fag

2 1 2 1 Pin 1 Or. Fag Ecoton

Mix 5 2 1 1 1 16 Pin 3 1 1 Pic.

Fag Secondary forests 3 2 4 Bet

Pm

Pin 4 8 3 AbF 6 Pic Origin forests 2 2 5 6 1 1 45 Qu 2 6 1

Fag

6 8 1 Aln. , , , , , , ,

l e rck l e rck

l e rck ves i l e rck ahn a C C C

C

(C. L. P imon C . L H ( ( ( (

( S ( (

( , 1805) , 1802) 1758) , 1817) , , 1757) , 1757) 1757) , 1834) , 1835) FAMILY / Species FAMILY Continued. t r e ill alck e na r alck e na r l e rck l e rck l e rck a inna e u s och och C C C W W L K Zygiella montana (30) LYCOSIDAE Alopecosa accentuata Alopecosa aculeata ( Alopecosa cuneata 1757) Alopecosa inquilina ( Alopecosa pulverulenta ( Alopecosa sulzeri 1873) Alopecosa taeniata Alopecosa trabalis 1757) figurata Arctosa 1876) maculata Arctosa 1822) Aulonia albimana ( amentata Pardosa 1757) lugubris Pardosa ( monticola Pardosa 1757) palustris Pardosa ( K ( L P P P T T T T N N N N N N M M M TP

Red list

Appendix 2.

211 European Arachnology 2005 2 3 1 1 2 1 2 4 7 1 1 Σ 19 31 22 69 49 3 1 1 1 4 7 S- alp

27 Fen Meadows

3 6 Mes 1 1 10 Dry

1 1 2 Peat bog

Cv

2 2

Deb Rocky

2 Shr

Pin 3 1 1 1 2 1 Pic

Se.

1 1 Ecoton Fag

2 Pin

Or. Fag Ecoton

Mix

3 6 11 Pin

Pic.

Fag Secondary forests

1 5 3 2 Bet

Pm

Pin 1 2 1 2 9 AbF 2 3

13 Pic Origin forests

1 2 16 18 Qu

3 16 Fag

2 2 1 1 1 Aln. , , , , , , , , och , ,

e l e rck hor e ll (O.P.- l e rck hor e ll imon L . C L . K D T l e rck T C

S imon (

( ( ( lackwall ( C . L C ( S copoli ( ( B ( S ( ( 1861) , 1895) , , 1875) , 1870) , 1833) FAMILY / Species FAMILY , 1778) Continued. est ring hor e ll och och ee r am b ridg e T W Pisaura mirabilis K prativaga Pardosa pullata Pardosa 1757) riparia Pardosa saltuaria Pardosa 1870) sordidata Pardosa ( Pirata hygrophilus 1872 Pirata knorri 1763) Pirata latitans 1841) Pirata piscatorius 1757) lutetiana Tricca 1876) robusta Trochosa 1876) ruricola Trochosa spinipalpis Trochosa terricola Trochosa 1856 nemoralis Xerolycosa ( PISAURIDAE (1) 1757) K C G P P P P P T T N N N N N M M M M TP

Red list LR (nt)

Appendix 2.

212 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina 1 1 4 1 8 1 1 2 1 Σ 14 13 15 12 10 404 3 1 1 S- 11 alp 1 4 Fen Meadows 2 1 2 1 21 Mes 5 Dry 1 1 Peat bog 1 1 Cv 1 1 4

Deb Rocky 1 Shr

Pin 3 4 2 16 Pic

Se.

Ecoton Fag

Pin

Or. Fag Ecoton

8 Mix 1 Pin 2 1 5 2 1 107 Pic. 3 1 Fag Secondary forests 1 1 3 1 Bet

1 26 Pm

11 Pin 7 1 1 AbF 1 2 1 39 Pic Origin forests 6 1 1 1 1 Qu 1 1 1 6

54 Fag

99 Aln. , , , , och L .

C . L li ve r

C . L

L . O imon ( C . L

S (

(

C . L

lackwall C . L K

B , 1802) ( 1879 , 1757) 1757) , , 1804) , 1837) , 1843 , 1834 , 1872 , 1868 , 1834) FAMILY / Species FAMILY Continued. alck e na r l e rck l e rck anz e r och och och och och e rman och C W C P H K AGELENIDAE (9) Agelena labyrinthica ( Histopona torpida agrestis Tegenaria ( atrica Tegenaria campestris Tegenaria domestica Tegenaria ( ferruginea Tegenaria ( silvestris Tegenaria denticulata Textrix 1789) CYBAEIDAE (1) Cybaeus angustiarum K HAHNIIDAE (5) Cryphoeca carpathica Cryphoeca silvicola Hahnia nava 1841) Hahnia ononidum 1875 Hahnia pusilla 1841 K K K K ? ? P P P P P T T N N M M M M TP

Red list

Appendix 2.

213 European Arachnology 2005 5 2 1 2 9 Σ 10 72 93 53 17 22 29 121 1 9 7 3 1 7 S- alp 1 Fen Meadows 1 2 5 2 Mes 1 1 7 9 7 13 Dry 2 1 4 Peat bog 1 Cv 1 5

Deb Rocky 1 Shr

1 Pin 1 2 3 1 3 1 Pic

Se.

Ecoton Fag

1 Pin 2 2 Or. Fag Ecoton

2 9 1 4 14 Mix 1 4 2 4 Pin 9 2 1 13 15 Pic. 6 1 12 Fag Secondary forests 6 1 8 Bet

1 1 Pm

1 Pin 1 8 2 4 11 AbF 1 6 1 Pic Origin forests 2 2 1 1 6 1 8 14 Qu 3 1 3 6 9

31 16 15 38 Fag

1 4 1 12 Aln.

, , , ,

,

L .

L . (

est ring hor e ll e ng T

W a b riciu s hor e ll

M F T

( , 1830) , 1830 , 1802) 1768) , 1833) , , 1833) , 1868 , 1855) FAMILY / Species FAMILY Continued. alck e na r alck e na r alck e na r ahn ahn t ro e m och och S W H W H W Amaurobius ferox Amaurobius K K DICTYNIDAE (3) Cicurina cicur 1793) Dictyna pusilla 1856 Dictyna uncinata 1856 AMAUROBIIDAE (6) fenestralis Amaurobius ( ( jugorum Amaurobius Callobius claustrarius ( Coelotes atropos ( inermis Eurocoelotes (1) TITANOECIDAE quadriguttata Titanoeca ( ANYPHAENIDAE (1) Anyphaena accentuata ( LIOCRANIDAE (3) cuprea Agroeca 1873 Apostenus fuscus 1851 P P P P P T T T N N N M M TP

Red list

Appendix 2.

214 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina 9 2 2 1 2 3 1 1 1 1 6 Σ 10 19 34 12 1 1 1 4 3 S- alp 1 Fen Meadows 2 Mes 1 1 1 1 2 1 Dry 1 Peat bog

Cv 3 1 1

Deb Rocky 1 3 2 Shr

Pin 1 1 1 Pic

Se.

Ecoton Fag

1 1 Pin

Or. Fag Ecoton

Mix 1 3 7 Pin 1 Pic.

Fag Secondary forests 4 2 1 Bet

2 2 Pm

1 Pin 1 1 1 3 10 AbF 2 Pic Origin forests 1 1 1 3 15 Qu 1 1 2

Fag

Aln. , , och C . L

ahn L . (

H

O.P.- C . L (

O.P.- C . L K

, 1830) , 1802) , 1802) 1882 , 1834) , , 1862 , 1862 , 1851 , 1856) , 1875 , 1866 , 1843 , 1835) FAMILY / Species FAMILY Continued. alck e na r alck e na r alck e na r lackwall est ring hor e ll ulczyń sk i och och och am b ridg e am b ridg e hor e ll W W B W T C K K Liocranum rupicola ( CLUBIONIDAE (9) Clubiona alpicola K Clubiona comta 1839 Clubiona corticalis ( Clubiona diversa Clubiona genevensis Clubiona neglecta Clubiona subsultans Clubiona terrestris Clubiona trivialis CORINNIDAE (1) festivus Phrurolithus GNAPHOSIDAE (13) Drassodes cupreus ( Drassodes lapidosus ( Drassodes pubescens ( Gnaphosa bicolor 1833) K C T W P P T T N N N N N N N N N M M TP

Red list LR (lc) Appendix 2.

215 European Arachnology 2005 4 3 1 1 2 6 3 3 4 2 8 Σ 18 61 31 35 42 1 1 1 2 1 1 1 9 S- 13 alp

1 Fen Meadows 1 3 2

5 1 Mes 1 1 2 3

1 5 2 Dry

2 Peat bog

1 Cv 1

1

Deb Rocky 1 1 3

1 3 Shr

Pin 1 1 3 2 Pic

Se.

Ecoton Fag

1 Pin

Or. Fag Ecoton

Mix 8

1 5 12 Pin

Pic. 2

Fag Secondary forests 1 6

1 2 Bet

1 1

Pm

Pin 1 5 1 3 4 1 5 7

22 AbF

1

Pic Origin forests 4 9 2 1 3 1 1 1 6

14 Qu

1 4 1 2 1 3 Fag

Aln.

, , L . ,

, ( all L . C . ,

( och ( C . L , 1878 ( , 1878 imon und ev hor e ll C . L L . K S

S T imon ( ulczyń sk i ( (

( imon (

S

K S

, 1802) 1922 , 1861) , , 1837) , 1757) , 1866) , 1866) , 1839) , 1833) FAMILY / Species FAMILY och Continued. alck e na r lackwall l e rck och och och och ham be rlin W B C K K K L . K 1870) 1871) 1878) Haplodrassus umbratilis Micaria fulgens ( Phaeocedus braccatus Zelotes clivicola Zelotes erebeus Zelotes latreillei Zelotes petrensis Zelotes puritanus Zelotes subterraneus ZORIDAE (5) Zora distincta 1915 Zora manicata Zora nemoralis ( Zora pardalis spinimana Zora 1833) HETEROPODIDAE (1) virescens Micrommata ( ZODARIIDAE (1) Zodarion germanicum K C P P T T T T T T T T N N N N N N TP

Red list LR (lc) CR VU

Appendix 2.

216 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina 1 1 1 1 3 7 1 8 2 1 Σ 12 26 17 12 17 1 6 1 5 S- alp

1 Fen Meadows 1 4 2 6 Mes 1 1 1 Dry 1 1 1 Peat bog

Cv

Deb Rocky

2 Shr

Pin 1 1 1 6 1 Pic

Se.

Ecoton Fag

Pin

Or. Fag Ecoton

Mix

6 Pin

1 Pic.

Fag Secondary forests 1 8

1 4 Bet

Pm

Pin 1 7 11 15 AbF

Pic Origin forests 1 4 1 Qu

3 1 Fag

Aln. , ,

, , C . imon

(

C . L S imon

anz e r

S

P

l e rck (

C a b riciu s ( F ( , 1802) , 1802) , 1837) , 1775) , 1837) , 1757) , 1757) , 1757) , 1875) , 1835 FAMILY / Species FAMILY och Continued. alck e na r alck e na r alck e na r e ng l e rck l e rck l e rck a b riciu s och C W C C M W F W K ( ( ( 1875 ( ( ( PHILODROMIDAE (8) aureolus Philodromus cespitum Philodromus collinus Philodromus margaritatus Philodromus vagulus Philodromus Thanatus formicinus Thanatus sabulosus oblongus Tibellus THOMISIDAE (20) Coriarachne depressa Diaea dorsata 1777) Misumena vatia 1757) Misumenops tricuspidatus ( Ozyptila atomaria 1801) Ozyptila blackwalli 1875 Ozyptila claveata ( L . K P T T T T N N N M M M M M M TP M Red list EN

Appendix 2.

217 European Arachnology 2005 2 1 3 2 4 1 8 3 1 2 2 1 4 Σ 11 12 27 2 8 2 4 8 1 2 4 S- alp 1 1 Fen Meadows 1 1 1 1 1 1 Mes 1 2 2 Dry

Peat bog

Cv

Deb Rocky 1 Shr

Pin 1 1 2 Pic

Se.

Ecoton Fag 2 Pin

Or. Fag Ecoton

Mix 4 Pin

Pic.

Fag Secondary forests 4 Bet

3 2 Pm

Pin 1 1 1 3 AbF

1 Pic Origin forests 1 1 4 Qu

4 2 1 Fag

Aln.

, , , , , , , och

ahn l e rck C . L

hor e ll C H imon

C . L

chrank

T ( S L . K (

S lackwall ( ( B Thorell, 1872 , 1802) , 1802) ( , 1834) , 1836) , 1882 , 1833) all , 1805 , 1837 FAMILY / Species FAMILY Continued. alck e na r alck e na r lackwall lackwall und ev och ulczyń sk i och B B W S W K 1875) 1846) 1803) 1757) ( 1875 1870 ( 1832) ( ( ( Ozyptila pullata Ozyptila trux Thomisus onustus Xysticus alpicola Xysticus audax Xysticus bifasciatus Xysticus cristatus Xysticus erraticus Xysticus gallicus Xysticus kochi Xysticus luctator Xysticus luctuosus Xysticus robustus (19) SALTICIDAE Ballus chalybeius Dendryphantes rudis Euophrys frontalis K K P P T T T T T N N N N N N N M M TP

Red list VU LR (lc) Appendix 2.

218 J. Svatoň & P. Gajdoš: Spiders of Gaderská and Blatnická Dolina 5 3 1 5 1 4 1 2 1 1 5 2 1 Σ 12 42 14 3295 7 1 1 S- alp 338 4 7 2 Fen 103 Meadows 1 1 1 14 283 Mes 1 2 1 1 189 Dry 2 1 1 3 71 Peat bog

88 Cv 1

50 Deb Rocky

31 Shr 1 Pin 3 2 1 1 Pic 242

Se. 6 Ecoton Fag

16 Pin 8 Or. Fag Ecoton 1 51 Mix 1 2 Pin 130 1 1 226 Pic.

30 Fag Secondary forests 2 4 96 Bet 2 57 Pm

38 Pin 1 9 1 2 256 AbF 5 Pic 190 Origin forests 5 1 Qu 307

2 1 Fag 327

161 Aln. , , , , ,

, ahn H

oda

( C . L l e rck ahn C . L l e rck P l e rck ( C ( ( C H C . L C . L

C . L C ( ( ( ( ( ( ( 1826) , 1802) , i, 1897 , 1877) , 1832) , 1837) , 1837) , 1837) , 1846) , 1837) FAMILY / Species FAMILY Continued. alck e na r alck e na r ahn imon och och och och ulczyń sk och W H W S K 1757) 1757) 1831) ( ( 1761) 1826) ( 1757) ( K K arcuata Evarcha falcata Evarcha Heliophanus aeneus Heliophanus cupreus Heliophanus flavipes Philaeus chrysops Phlegra fasciata Pseudeuophrys erratica Salticus scenicus Salticus zebraneus Sitticus floricola Sitticus rupicola Sitticus saxicola Sitticus zimmermanni Synageles lepidus petrensis Talavera K K K T o t a l : T P P T T T T N N N N M M M M M M TP

Red list EN EN

Appendix 2.

219