British & Irish Botany 2(3): 169-189, 2020

List of vascular plants endemic to Britain, Ireland and the Channel Islands 2020

Timothy C.G. Rich Cardiff, U.K.

Corresponding author: Tim Rich: [email protected]

This pdf constitutes the Version of Record published on 31st August 2020

Abstract A list of 804 plants endemic to Britain, Ireland and the Channel Islands is broken down by country. There are 659 taxa endemic to Britain, 20 to Ireland and three to the Channel Islands. There are 25 endemic sexual species and 26 sexual subspecies, the remainder are mostly critical apomictic taxa. Fifteen endemics (2%) are certainly or probably extinct in the wild.

Keywords: England; Northern Ireland; Republic of Ireland; Scotland; Wales.

Introduction This note provides a list of vascular plants endemic to Britain, Ireland and the Channel Islands, updating the lists in Rich et al. (1999), Dines (2008), Stroh et al. (2014) and Wyse Jackson et al. (2016). The list includes endemics of subspecific rank or above, but excludes infraspecific taxa of lower rank and hybrids (for the latter, see Stace et al., 2015). There are, of course, different taxonomic views on some of the taxa included. Nomenclature, taxonomic rank and endemic status follows Stace (2019), except for Hieracium (Sell & Murrell, 2006; McCosh & Rich, 2018), Ranunculus auricomus group (A. C. Leslie in Sell & Murrell, 2018), Rubus (Edees & Newton, 1988; Newton & Randall, 2004; Kurtto & Weber, 2009; Kurtto et al. 2010, and recent papers), Taraxacum (Dudman & Richards, 1997; Kirschner & Štepànek, 1998 and recent papers) and Ulmus (Sell & Murrell, 2018). Ulmus is included with some reservations, as many taxa are largely vegetative clones which may occasionally reproduce sexually and hence may not merit species status (cf. Stace, 2019). There are two recent endemics described from Scotland, Alchemilla sciura and the diploid Dryopteris affinis subsp. cluthensis (Lynes, 2019; Church et al., 2019). Two recently- described Scottish Alchemilla endemics (Hogarth, 2020) require valid publication before being accepted. I differ from Stace (2019) in recognising Cochlearia atlantica but not accepting C. micacea or C. officinalis subsp. scotica, or Anthyllis vulneraria subsp. corbierei which is simple genetic variation (Rich, 2006). Dryopteris kerryensis is here accepted at species rank and Sorbus avonensis in now also treated as a species as the population forms a distinct clone in the Avon Gorge of about 42 individuals (L. Houston, unpublished data, pers. comm. 2020).

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The political regions to which species are endemic are given to indicate geographic responsibilities and priorities; these are Channel Islands, England, Isle of Man, Scotland or Wales or for any combination of these as Britain, and Northern Ireland or the Republic of Ireland or where both as Ireland, and where in both Britain and Ireland as Britain & Ireland. Fifteen endemics are certainly or probably extinct in the wild but there is little recent data for many other taxa which could also be extinct. The list is given in Appendix 1 and the number of taxa endemic to each region is summarised in Tab. 1. The endemics are mostly critical apomictic taxa; Alchemilla (2), Dryopteris (1), Hieracium (332), Limonium (12), Ranunculus auricomus group (58), Rubus (194), Sorbus (38) and Taraxacum (53). There are 25 sexual species and 26 sexual subspecies.

Table 1. Number of endemics in each region

Geographic Region No. endemics Total Channel Islands 3 England 287 Isle of Man 0 Scotland 189 Wales 59 Britain 121 Total for Britain 659 Northern Ireland 0 Republic of Ireland 14 Ireland 6 Total for Ireland 20 Britain & Ireland 125 Overall Total 804

Acknowledgements I would like to thank Ian Denholm, Hendrik Devriese, Dave Earl, Libby Houston, Alan Leslie, David McCosh, Rob Randall, John Richards, Fred Rumsey, Clive Stace, Abraham van der Beek and Mike Wyse Jackson.

References Church, A.R., Evans, A.J., Golding, R., Rumsey, F.J. & Viane, R.L.L. 2019. Dryopteris affinis subsp. cluthensis: A new taxon in the Dryopteris affinis complex (Dryopteridaceae). Fern Gazette, 21: 85-95. Dines, T. 2008. A vascular plant Red Data List for Wales. Salisbury: Plantlife International. Dudman, A. & Richards, A.J. 1997. Dandelions of Great Britain and Ireland. London: Botanical Society of the British Isles. Edees, E.S. & Newton, A. 1988. Brambles of the British Isles. London: Ray Society. Hogarth, B.G. 2020. Alchemilla A field guide to the lady’s-mantles of the British Isles. Scotland: Amenta Publishing. Kirschner, J. & Štepànek, J. 1998. A monograph of Taraxacum sect. Palustria. Pruhonice: Institute of Botany, Academy of Sciences of Czech Republic.

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Kurtto, A. & Weber, H.E. 2009. Rubus. In: Kurtto, A., ed.: Rosaceae. Euro+Med Plantbase - the information resource for Euro-Mediterranean plant diversity. [accessed April 2020]. Available at http://ww2.bgbm.org/EuroPlusMed/. Kurtto, A. & Weber, H.E., Lampinen, R. & Sennikov, A.N., eds. 2010. Atlas Flora Europaeae. Distribution of Vascular plants in Europe. 15. Rosaceae (Rubus). Helsinki: The Committee for Mapping the Flora of Europe & Societas Biologica Fennica Vanamo. Lynes, M. 2019. Alchemilla sciura (Rosaceae), a new species of Lady’s-mantle. British & Irish Botany 1: 335-341. McCosh, D.J. & Rich, T.C.G. 2018. Atlas of British and Irish Hawkweeds (Pilosella Hill and Hieracium L.). 2nd ed. Harpenden: Botanical Society of Britain and Ireland. Newton, A. & Randall, R.D. 2004. Atlas of British and Irish Brambles. London: Botanical Society of the British Isles. Rich, T.C.G. 2006. A simple genetic base for the spreading and appressed hair types in Anthyllis vulneraria L. (Fabaceae). Watsonia, 26: 69. Rich, T.C.G., Hutchinson, G., Randall, R.D. & Ellis, R.G. 1999. List of plants endemic to the British Isles. BSBI News, 80: 23-27. Sell, P.D. & Murrell, G. 2006. Flora of Great Britain and Ireland. Volume 4. Cambridge: Cambridge University Press. Sell, P.D. & Murrell, G. 2018. Flora of Great Britain and Ireland. Volume 1. Cambridge: Cambridge University Press. Stace, C.A., Preston, C.D. & Pearman, D.A. 2015. Hybrid flora of the British Isles. Bristol: Botanical Society of Britain and Ireland. Stace, C.A. 2019. New flora of the British Isles. 4th ed. Middlewood Green, Suffolk: C & M Floristics. Stroh, P.A., Leach, S. J., August, T.A., Walker, K.J., Pearman, D.A., Rumsey, F.J., Harrower, C.A., Fay, M.F., Martin, J.P., Pankhurst, T., Preston, C.D. & Taylor, I. 2014. A Vascular Plant Red List for England. Bristol: Botanical Society of Britain and Ireland. Wyse Jackson, M., FitzPatrick, Ú., Cole, E., Jebb, M., McFerran, D., Sheehy Skeffington, M. & Wright, M. 2016. Ireland Red List No. 10: Vascular Plants. Dublin: National Parks and Wildlife Service, Department of Arts, Heritage, Regional, Rural and Gaeltacht Affairs.

Copyright retained by author(s). Published by BSBI under the terms of the Creative Commons Attribution 4.0 International Public License. ISSN: 2632-4970 https://doi.org/10.33928/bib.2020.02.169

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Appendix 1. List of vascular plants endemic to Britain, Ireland and the Channel Islands broken down by country.

Species Distribution summary Alchemilla minima England Alchemilla sciura Scotland Arenaria ciliata subsp. hibernica Ireland Arenaria norvegica subsp. anglica England Bromus interruptus England (Extinct) Calamagrostis scotica Scotland Centaurium intermedium England Centaurium tenuiflorum subsp. England but further studies in NW anglicum Europe required Cerastium fontanum subsp. scoticum Scotland Cochlearia atlantica Britain & Ireland Coincya monensis subsp. monensis Britain Coincya wrightii England Cotoneaster cambricus Wales Dactylorhiza incarnata subsp. coccinea Britain & Ireland but further studies in NW Europe required Dactylorhiza incarnata subsp. gemmana Britain & Ireland but further studies in NW Europe required Dactylorhiza incarnata subsp. pulchella Britain & Ireland but further studies in NW Europe required Dactylorhiza kerryensis Republic of Ireland Dactylorhiza traunsteinerioides Britain & Ireland Dryopteris affinis subsp. cluthensis Scotland Dryopteris kerryensis Britain & Ireland Epipactis dunensis Britain & Ireland Epipactis sancta England Erythranthe peregrina Scotland Euphrasia cambrica Wales Euphrasia campbelliae Scotland Euphrasia heslop-harrisonii Scotland Euphrasia marshallii Scotland Euphrasia officinalis subsp. anglica Britain & Ireland Euphrasia pseudokerneri Britain & Ireland Euphrasia rivularis Britain Euphrasia rotundifolia Scotland Euphrasia vigursii England Fumaria capreolata subsp. babingtonii Britain & Ireland Fumaria occidentalis England Fumaria purpurea Britain & Ireland Gentianella amarella subsp. anglica Britain Gentianella amarella subsp. occidentalis Wales Helianthemum oelandicum subsp. England

172 levigatum Geranium purpureum subsp. forsteri England and Channel Islands Gymnadenia borealis Britain & Ireland but further studies in NW Europe required Herniaria ciliolata subsp. ciliolata England and Channel Islands Herniaria ciliolata subsp. subciliata Channel Islands Hieracium acamptum England (Extinct) Hieracium aequiserratum England Hieracium aggregatifolium Scotland Hieracium aggregatum Scotland Hieracium amaurostictum Scotland Hieracium amnicola England Hieracium ampliatiforme England Hieracium ampliatum Britain & Ireland Hieracium amydrostictum Scotland Hieracium anfractiforme Scotland Hieracium anglicum Britain & Ireland Hieracium anglorum Britain Hieracium anguinum Britain Hieracium angustatiforme Wales Hieracium angustisquamum Britain & Ireland Hieracium apheles Wales Hieracium aphyllopodioides Scotland Hieracium apiculatidens England Hieracium argentatum Republic of Ireland Hieracium argentiforme Scotland Hieracium argutifolium Britain & Ireland Hieracium aristidens Scotland Hieracium armadalense Scotland Hieracium arnsidense England Hieracium arranense Scotland Hieracium arvonense Wales Hieracium ascendentidens Scotland Hieracium asteridiophyllum Wales Hieracium atriglandulosum Scotland Hieracium attenboroughianum Wales Hieracium attenuatifolium Scotland Hieracium auratiflorum Britain Hieracium australius Scotland Hieracium backhousei Scotland Hieracium bakerianum England Hieracium basalticola Ireland Hieracium basicrinum Scotland Hieracium beebyanum Britain & Ireland Hieracium benhopense Scotland Hieracium bettyhillense Scotland

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Hieracium boreoanglicum Britain Hieracium boswellii Britain Hieracium breacense Scotland (Extinct) Hieracium breadalbanense Scotland Hieracium breconense Wales Hieracium breconicola Wales Hieracium breve Scotland Hieracium brigantum England Hieracium britanniciforme Britain Hieracium britannicoides Wales Hieracium britannicum England Hieracium cacuminum Wales Hieracium caesiopilosum Britain Hieracium caesitium England Hieracium caledonicum Scotland Hieracium calenduliflorum Scotland Hieracium callistophyllum Scotland Hieracium calvum Scotland Hieracium cambrense Britain Hieracium cambricogothicum Wales (Extinct) Hieracium cambricum Wales Hieracium camptopetalum Scotland Hieracium candelabrae England Hieracium cantianum England Hieracium carenorum Scotland Hieracium carneddorum Wales Hieracium centripetale Scotland Hieracium cerinthiforme Britain & Ireland Hieracium charitodon Wales Hieracium chloranthum Scotland Hieracium chrysolorum Britain Hieracium cillense Wales Hieracium cinderella Britain & Ireland Hieracium clivicola Wales (+England?) Hieracium clovense Scotland Hieracium completum Scotland Hieracium cravoniense Britain Hieracium crebridens England Hieracium crebridentiforme England Hieracium cremnanthes Scotland Hieracium cumbriense England Hieracium cuneifrons Britain Hieracium cuspidens Scotland Hieracium cyathis Britain Hieracium cyclicum Britain & Ireland Hieracium cymbifolium Britain

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Hieracium daedalolepioides Britain Hieracium dalense England Hieracium dasythrix Scotland Hieracium deargicola Scotland Hieracium decolor England Hieracium deganwyense Britain & Ireland Hieracium dentulum Britain Hieracium dewari Scotland Hieracium dicella Britain Hieracium difficile Scotland Hieracium dilectum Scotland Hieracium dipteroides Scotland Hieracium discophyllum Wales Hieracium diversidens Scotland Hieracium dolorosum Scotland Hieracium dowardense Britain Hieracium drummondii Scotland Hieracium duriceps Britain & Ireland Hieracium eboracense Britain Hieracium ebudicum Scotland Hieracium einichense Scotland Hieracium elevatum Wales Hieracium elongatifolium Scotland Hieracium eminentiforme Britain Hieracium erubescens Britain & Ireland Hieracium eucallum Scotland Hieracium eustales Scotland Hieracium eustomon Britain Hieracium eximium Scotland Hieracium farrense Scotland Hieracium filisquamum England Hieracium fissuricola England Hieracium flocculipubens Britain & Ireland Hieracium flocculosiforme Britain & Ireland Hieracium flocculosum Britain & Ireland Hieracium fratrum Scotland Hieracium fucatifolium Scotland Hieracium fulvocaesium Scotland Hieracium glanduliceps England Hieracium glandulidens Scotland Hieracium glevense Britain Hieracium globosiflorum Scotland Hieracium gothicoides Scotland Hieracium gracilifolium Scotland Hieracium grampianum Scotland Hieracium graniticola Scotland

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Hieracium gratum Scotland Hieracium griffithii Wales (Extinct) Hieracium grovesii Scotland Hieracium hanburyi Scotland Hieracium hartii Republic of Ireland Hieracium hastiforme Scotland Hieracium hebridense Britain & Ireland Hieracium hethlandiae Scotland (Extinct) Hieracium hibernicum Ireland Hieracium holophyllum Britain Hieracium holosericeum Britain Hieracium hyparcticoides Scotland Hieracium hypochaeroides Britain & Ireland Hieracium hypophalacrum Scotland Hieracium inaequilaterum England Hieracium insigne Scotland Hieracium inspissatum Wales Hieracium insulare Scotland Hieracium iricum Britain & Ireland Hieracium irregularidens Scotland Hieracium isabellae Scotland Hieracium itunense England Hieracium jaculifolium England Hieracium jonesianum England Hieracium kennethii Scotland Hieracium kentii England Hieracium kingshousense Scotland Hieracium kinkellense Scotland Hieracium kinrossense Scotland Hieracium kintrawense Scotland Hieracium klingrahoolense Scotland Hieracium lacinifolium England Hieracium laetificum Scotland Hieracium lagganense Scotland Hieracium lakelandicum England Hieracium langwellense Britain & Ireland Hieracium larigense Scotland Hieracium lepiduloides Britain & Ireland Hieracium leptodon Scotland Hieracium leyanum Britain Hieracium leyi Britain Hieracium linguans Wales Hieracium lingulatum Scotland Hieracium lintonense Britain Hieracium lintonii Britain Hieracium maccoshianum Scotland

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Hieracium macrocarpum Scotland Hieracium maculoides England Hieracium magniceps Scotland (Extinct?) Hieracium mammidens England Hieracium mariae Scotland Hieracium maritimum Britain & Ireland Hieracium marshallii Scotland Hieracium megapodium England Hieracium melanochloricephalum Scotland Hieracium melanoglochin Scotland Hieracium memorabile Scotland Hieracium micracladium Scotland Hieracium milesii Scotland Hieracium mirandum England Hieracium mucronellum Scotland Hieracium mundum Scotland Hieracium naviense England Hieracium neocoracinum Wales Hieracium neomarginatum Scotland Hieracium nidense Wales Hieracium nigrifactum Scotland Hieracium nitidum Scotland Hieracium northroense Scotland Hieracium notabile Scotland Hieracium obovatifolium England Hieracium ochthophilum Scotland Hieracium oenophyllum Scotland Hieracium optimum Scotland Hieracium orcadense Britain & Ireland Hieracium orimeles Britain Hieracium orithales Scotland Hieracium ornatilorum England Hieracium ovaliforme Scotland Hieracium oxybeles Britain & Ireland Hieracium oxyodus Scotland (Extinct?) Hieracium pachyphylloides Britain Hieracium pauculidens Scotland Hieracium peccense England Hieracium pensum Scotland Hieracium pentaploideum Scotland Hieracium peroblongum England Hieracium perscitum Scotland Hieracium perthense Scotland Hieracium petrocharis Scotland Hieracium pictorum Scotland Hieracium piligerum Britain

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Hieracium placerophylloides Britain Hieracium pollinarioides Scotland Hieracium pollinarium Scotland Hieracium portlandicum England Hieracium praesigne Britain Hieracium praethulense Scotland Hieracium probum Scotland Hieracium prominentidens England Hieracium promontoriale Britain Hieracium protentum Scotland Hieracium proximum Britain & Ireland Hieracium pruinale Scotland Hieracium pseudacroleucum Britain Hieracium pseudanglicoides Scotland Hieracium pseudanglicum Scotland Hieracium pseudintegratum England Hieracium pseudocurvatum Scotland Hieracium pseudoleyi Wales Hieracium pseudopetiolatum Scotland Hieracium pseudosarcophyllum Scotland Hieracium pseudosubcyaneum England Hieracium pseudozetlandicum Scotland Hieracium pugsleyi Scotland Hieracium pulchrius Britain Hieracium pusillifolium Wales (Extinct) Hieracium radyrense Wales Hieracium raveniorum Scotland Hieracium reayense Britain Hieracium rectulum Wales Hieracium repandulare Wales Hieracium reticulatiforme Britain Hieracium rhombicum Britain Hieracium riddelsdellii Wales Hieracium rivale Scotland Hieracium robertsii Wales Hieracium ronasii Scotland Hieracium ronayense Scotland Hieracium rubicundiforme Britain & Ireland Hieracium rubiginosum Britain & Ireland Hieracium sanguineum Britain & Ireland Hieracium sannoxense Scotland Hieracium saxorum Britain & Ireland Hieracium scabrisetum Britain Hieracium scarpicum Scotland Hieracium scoticum Scotland & Ireland Hieracium scottii Scotland

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Hieracium scullyi Republic of Ireland Hieracium senescens Scotland & Ireland Hieracium shoolbredii Scotland Hieracium siluriense Wales Hieracium silvaticoides Britain & Ireland Hieracium sinuans Scotland Hieracium sinuolatum England Hieracium snowdoniense Wales Hieracium solum Scotland Hieracium sommerfeltii Britain & Ireland Hieracium sowadeense Scotland Hieracium sparsifrons Republic of Ireland Hieracium spenceanum Scotland Hieracium stenolepiforme England Hieracium stenopholidium Britain Hieracium stenophyes Scotland Hieracium stewartii Britain & Ireland Hieracium stictum England Hieracium strictiforme Britain Hieracium subamplifolium Britain & Ireland Hieracium subbritannicum Britain Hieracium subcrinellum Britain Hieracium subcrocatum Britain & Ireland Hieracium subcyaneum England Hieracium subduriceps Scotland Hieracium subglobosum Scotland Hieracium subgracilentipes England Hieracium subhirtum Scotland Hieracium subintegrifolium England (Extinct) Hieracium subminutidens Wales Hieracium submutabile Britain Hieracium subplanifolium Britain Hieracium subprasinifolium England Hieracium subrubicundum Scotland & Ireland Hieracium subscoticum Scotland Hieracium substrigosum Britain Hieracium subtenue Scotland Hieracium subtenuifrons Scotland Hieracium subtruncatum Scotland Hieracium subumbellatiforme Scotland Hieracium surrejanum England Hieracium tavense Wales Hieracium tenuifrons Scotland Hieracium thalassinum Scotland Hieracium tottoense Scotland Hieracium triangularifolium Scotland

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Hieracium tricolorans England Hieracium uiginskyense Britain & Ireland Hieracium uistense Scotland Hieracium uisticola Scotland Hieracium umbellatum subsp. Britain & Ireland bichlorophyllum Hieracium umbellatum subsp. ogwenii Wales Hieracium vagense Britain Hieracium vagicola England Hieracium varifolium Scotland Hieracium vennicontium Britain Hieracium veterascens Scotland Hieracium vinicaule Scotland Hieracium vinifolium Scotland Hieracium vorlichense Scotland Hieracium westii Scotland Hieracium zetlandicum Scotland Limonium binervosum subsp. anglicum England Limonium binervosum subsp. cantianum England Limonium binervosum subsp. mutatum England Limonium binervosum subsp. saxonicum England Limonium britannicum subsp. England britannicum Limonium britannicum subsp. celticum Britain Limonium britannicum subsp. England coombense Limonium britannicum subsp. Britain transcanalis Limonium dodartiforme England Limonium loganicum England Limonium paradoxum Wales Limonium parvum Wales Limonium procerum subsp. cambrense Wales Limonium procerum subsp. devoniense England Limonium procerum subsp. procerum England Limonium recurvum subsp. humile Britain Limonium recurvum subsp. Britain & Ireland portlandicum Limonium recurvum subsp. Republic of Ireland pseudotranswallianum Limonium recurvum subsp. recurvum England Limonium transwallianum Wales Linum perenne subsp. anglicum England Luzula multiflora subsp. hibernica Ireland Milium vernale subsp. sarniense Channel Islands Pilosella flagellaris subsp. bicapitata Scotland

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Primula scotica Scotland Ranunculus acutissimus England Ranunculus aequilaterus England Ranunculus angularis England Ranunculus angustiscutatus England Ranunculus argillicola England Ranunculus armingfordensis England Ranunculus attenuilobus England Ranunculus brewisiae England Ranunculus calvitorus England Ranunculus cantabrigiensis England Ranunculus combertonensis England Ranunculus completus England Ranunculus conspicuidens England Ranunculus contiguilobus England Ranunculus convexilobus England Ranunculus crassilobus England Ranunculus curvilobus England Ranunculus dalensis England Ranunculus densipilus England Ranunculus dolichodontus England Ranunculus flammula subsp. minimus Britain & Ireland Ranunculus flammula subsp. scoticus Britain & Ireland Ranunculus gongylodes England Ranunculus gracilidens England Ranunculus gracililobus England Ranunculus gracilistylus England Ranunculus henslowii England Ranunculus hertfordensis England Ranunculus horwoodii England Ranunculus imbricatus England Ranunculus mammidens England Ranunculus medioviridis England Ranunculus multidens A.C. Leslie, nom. England illeg. Ranunculus multilobus England Ranunculus notauricomus England Ranunculus nubiliviridis England Ranunculus obesicaulis England Ranunculus obovatilobatus England Ranunculus parvidens England Ranunculus paucipetalus England Ranunculus petaloideus England Ranunculus prominentilobus England Ranunculus pubitorus England Ranunculus rectilobus England

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Ranunculus rotundilobus England Ranunculus sepulcralis England Ranunculus spathulilobus England Ranunculus stipitatilobus England Ranunculus stipitatus England Ranunculus surrejanus England Ranunculus symmetricidens England Ranunculus toftensis England Ranunculus totipetalus England Ranunculus unipetalus England Ranunculus vadincisus England Ranunculus validicaulis England Ranunculus varipetalus England Ranunculus viridissimus England Ranunculus waltersii England Ranunculus wrayensis England Rubus acclivitatum Britain & Ireland Rubus accrescens Britain Rubus acutifrons Britain Rubus adamsii England Rubus aghadergensis Britain & Ireland Rubus ahenifolius England Rubus albionis Britain & Ireland Rubus anglofuscus Britain Rubus altiarcuatus Britain, Ireland & Channel Isles Rubus amplificatus Britain & Ireland Rubus angloserpens England Rubus angusticuspis Britain Rubus aquarum Wales Rubus ariconiensis Britain & Ireland Rubus aristisepalus Britain Rubus armipotens Britain Rubus avaloniensis England Rubus averyanus England Rubus babingtonianus England Rubus babingtonii Britain Rubus bagnallianus England Rubus bakerianus England Rubus bartonii Britain & Ireland Rubus biloensis Britain Rubus bloxamianus England Rubus boudiccae Britain & Ireland Rubus breconensis Wales Rubus brevistaminosus England Rubus briggsianus Britain, Ireland, Isle of Man, Channel Isles

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Rubus briggsii England Rubus britannicus England Rubus bucknallii England Rubus calvatus Britain & Ireland Rubus cambrensis Britain Rubus canterburiensis England Rubus carnkiefensis England Rubus castrensis England Rubus celticus Wales (Ireland?) Rubus cerdicii England Rubus chrysoxylon Britain Rubus cinerosiformis Britain & Ireland Rubus cinerosus England & Ireland Rubus clausentinus England Rubus confertiflorus England Rubus conspersus England Rubus cordatifolius Channel Islands Rubus cornubiensis England Rubus cotteswoldensis England Rubus crespignyanus England Rubus criniger England & Isle of Man Rubus crudelis England Rubus cumbrensis Britain Rubus curvidens Britain Rubus curvispinosus England Rubus daltrii England Rubus dasycoccus Britain Rubus daveyi England and Channel Islands Rubus decussatus England Rubus devoniensis England Rubus distractiformis Britain & Ireland Rubus diversiarmatus England Rubus dobuniensis England (Extinct?) Rubus dunensis Britain & Ireland Rubus durescens England Rubus durotrigum England Rubus eboracensis Britain Rubus ebudensis Scotland Rubus echinatoides Britain & Ireland Rubus effrenatus Britain (Ireland?) Rubus errabundus Britain & Ireland Rubus erythrops England Rubus euanthinus Britain Rubus furnarius England Rubus furvicolor Scotland Rubus fuscicaulis Britain & Ireland

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Rubus fuscicortex Britain Rubus gallofuscus Britain Rubus gariannensis England Rubus glanduliger England Rubus griffithianus Britain & Ireland Rubus halsteadensis Britain Rubus hantonensis England Rubus hebridensis Britain & Ireland Rubus herefordensis England Rubus hesperius Ireland & Isle of Man Rubus heterobelus England Rubus hibernicus Britain & Ireland Rubus hindii Britain & Ireland Rubus hirsutissimus Britain Rubus hylocharis Britain & Ireland Rubus hyposericeus England Rubus iceniensis England Rubus incurvatiformis Britain Rubus infestior England Rubus infestisepalus England Rubus informifolius England Rubus intensior Britain Rubus iodnephes England Rubus iricus Britain & Ireland, Channel Isles Rubus iscanus Britain Rubus lacustris England Rubus lamburnensis Britain & Ireland, Channel Isles Rubus largificus Britain & Ireland Rubus lasiodermis England Rubus latifolius Britain & Ireland Rubus laxatifrons England Rubus lentiginosus Britain & Ireland Rubus lettii Britain & Ireland Rubus leucandriformis England Rubus lintonii England Rubus londinensis England Rubus longifrons England (Extinct?) Rubus longus Britain & Ireland Rubus ludensis Britain Rubus malvernicus Britain & Ireland Rubus marshallii England Rubus melanocladus Britain Rubus mercicus England Rubus merlinii Wales Rubus metallorum England Rubus milesianus England (if native; introduced in

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Ireland) Rubus mollissimus England Rubus monensis Wales Rubus morganwgensis Britain Rubus moylei Britain & Ireland Rubus mucronatiformis England (Ireland?) Rubus mucronatoides Britain Rubus newbouldianus England Rubus newbouldii Britain Rubus newbridgensis England Rubus newtonii Britain Rubus norvicensis Britain & Ireland, Channel Isles Rubus obesifolius England Rubus obscuriflorus England Rubus orbus Britain Rubus ordovicum Britain & Ireland Rubus painteri England & Ireland Rubus pallidisetus Britain & Ireland Rubus pampinosus Britain Rubus pascuorum Britain & Ireland Rubus percrispus Britain & Ireland, Channel Isles (Extinct?) Rubus perdigitatus Britain Rubus permundus England Rubus pervalidus England Rubus pictorum Britain Rubus pliocenicus England Rubus poliodes England Rubus polyoplus Britain & Ireland Rubus porphyrocaulis England Rubus powellii England Rubus pseudoplinthostylus England Rubus pullifolius England Rubus purchasianus Britain Rubus putneiensis England Rubus pydarensis England and Channel Islands Rubus regillus England Rubus riddelsdellii Britain & Ireland Rubus robiae Britain & Ireland Rubus rossensis Britain & Ireland Rubus rotundifolius England (Extinct?) Rubus rubriflorus England Rubus rubristylus Britain & Ireland, Isle of Man Rubus sagittarius England Rubus salteri England Rubus scoticus Britain & Ireland

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Rubus sectiramus England Rubus segontii Britain & Ireland Rubus semiglaber Wales Rubus silurum Britain (Ireland?) Rubus sneydii England Rubus spadix England Rubus stanneus England Rubus subintegribasis England Rubus surrejanus Britain (introduced to Ireland?) Rubus tardus England Rubus tavensis Wales Rubus tenuiarmatus Britain Rubus trelleckensis Wales Rubus tresidderi England Rubus triangularis England Rubus trinovantium England Rubus troiensis Britain Rubus turritus England Rubus vagensis Britain Rubus varvicensis England Rubus villicauliformis Britain, Ireland and Channel Islands Rubus villosior England Rubus vindomensis England Rubus waddellii Ireland & Isle of Man Rubus warrenii Britain & Ireland Rubus watsonii England Rubus wedgwoodiae England Rubus wirralensis Britain, Ireland & Isle of Man Rubus wolley-dodii England Rumex acetosa subsp. hibernicus Britain & Ireland Saxifraga rosacea subsp. hartii Republic of Ireland Scleranthus perennis subsp. prostratus England Senecio cambrensis Britain Senecio eboracensis England (Extinct) Sisyrinchium bermudiana Ireland (if different to North American taxon; naturalised in Britain) Sorbus admonitor England Sorbus anglica (including S. waltersii) Britain & Ireland Sorbus arranensis Scotland Sorbus arvonicola Wales Sorbus avonensis England Sorbus bristoliensis England Sorbus cambrensis Wales Sorbus cheddarensis England Sorbus cuneifolia Britain Sorbus devoniensis Britain & Ireland

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Sorbus eminens Britain Sorbus eminentiformis Britain Sorbus eminentoides England Sorbus evansii England Sorbus greenii England Sorbus herefordensis England Sorbus hibernica Ireland Sorbus lancastriensis England Sorbus leighensis England Sorbus leptophylla Wales Sorbus leyana Wales Sorbus margaretae England Sorbus minima Wales Sorbus parviloba England Sorbus porrigentiformis (including S. Britain humphreyana) Sorbus pseudofennica Scotland Sorbus pseudomeinichii Scotland Sorbus richii England Sorbus rupicoloides England Sorbus saxicola England Sorbus scannelliana Republic of Ireland Sorbus spectans England Sorbus stenophylla Wales Sorbus stirtoniana Wales Sorbus subcuneata England Sorbus vexans England Sorbus whiteana England Sorbus wilmottiana England Spartina anglica Britain but widely planted elsewhere Taraxacum acutum England Taraxacum aesculosum Britain & Ireland Taraxacum amarellum Republic of Ireland Taraxacum amicorum England Taraxacum atrocollinum Britain & Ireland Taraxacum berthae Britain Taraxacum breconense Wales Taraxacum caledonicum Scotland Taraxacum calophyllum Scotland Taraxacum cambricum Britain & Ireland Taraxacum celticum Britain & Ireland Taraxacum cherwellense England Taraxacum chrysoglossum Scotland Taraxacum claudiae Scotland Taraxacum clovense Scotland Taraxacum cophocentrum Britain & Ireland

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Taraxacum cornubiense England Taraxacum degelii Britain & Ireland Taraxacum elegantifrons Republic of Ireland Taraxacum fulvicarpum Britain & Ireland Taraxacum gaelorum Republic of Ireland Taraxacum geirhildae Scotland Taraxacum haworthianum Britain & Ireland Taraxacum hesperium Britain & Ireland Taraxacum hexhamense Britain & Ireland Taraxacum hibernicola Republic of Ireland Taraxacum hirsutissimum Scotland Taraxacum inane Britain & Ireland Taraxacum inclinorum Republic of Ireland Taraxacum inopinatum Britain & Ireland Taraxacum lancastriense Britain Taraxacum luteum Britain & Ireland Taraxacum margettsii England Taraxacum nigridentatum Britain Taraxacum oellgaardii Britain Taraxacum olgae Britain Taraxacum orcadense Scotland Taraxacum palmeri Scotland Taraxacum palustrisquameum Britain & Ireland Taraxacum pankhurstianum Scotland Taraxacum porteri Britain Taraxacum pseudonordstedtii England Taraxacum richardsianum Britain Taraxacum ronae Britain & Ireland Taraxacum rufofructum Scotland Taraxacum sahlinianum Britain & Ireland Taraxacum scoticum Britain Taraxacum serpenticola Scotland Taraxacum stenacrum Britain & Ireland Taraxacum subbracteatum Britain & Ireland Taraxacum subnaevosum Britain Taraxacum tanylepis Scotland Taraxacum webbii Republic of Ireland Tephroseris integrifolia subsp. maritima Wales Ulmus acuminatissima England Ulmus acutissima England Ulmus alta England Ulmus anglosaxonica England Ulmus angustiformis Britain Ulmus asymmetrica England Ulmus atrovirens England Ulmus cantabrigiensis England

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Ulmus chaterorum Wales Ulmus coriaceifolia England Ulmus coritana England Ulmus cornubiensis England Ulmus crassa England Ulmus crenata England Ulmus cuneiformis Britain Ulmus curvifolia England Ulmus daveyi Britain Ulmus diversifolia England Ulmus elegantissima Britain Ulmus exoniensis England Ulmus gyrophylla Britain Ulmus hollandica Britain Ulmus incisa England Ulmus longicaudata Britain Ulmus longidens Britain Ulmus longidentata England Ulmus madingleyensis England Ulmus microdon England Ulmus minor England Ulmus mossii Britain Ulmus multidentata England Ulmus obesidens England Ulmus oblanceolata England Ulmus occidentalis Britain Ulmus peninsularis England Ulmus platyphylla England Ulmus plotii Britain Ulmus prionophylla England Ulmus prominentidens England Ulmus pseudelegantissima England Ulmus pseudobovata England Ulmus pseudocoritana England Ulmus rasilis England Ulmus rhombifolia England Ulmus sativa England Ulmus serrata England Ulmus serratifrons England Ulmus serratula England Ulmus sowerbyi England Ulmus sylvatica England Ulmus vegeta Britain Ulmus wheatleyi Britain

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